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-The Project Gutenberg EBook of Many-Storied Mountains, by Greg Beaumont
-
-This eBook is for the use of anyone anywhere in the United States and most
-other parts of the world at no cost and with almost no restrictions
-whatsoever. You may copy it, give it away or re-use it under the terms of
-the Project Gutenberg License included with this eBook or online at
-www.gutenberg.org. If you are not located in the United States, you'll have
-to check the laws of the country where you are located before using this ebook.
-
-Title: Many-Storied Mountains
- The Life of Glacier National Park
-
-Author: Greg Beaumont
-
-Release Date: July 19, 2017 [EBook #55150]
-
-Language: English
-
-Character set encoding: UTF-8
-
-*** START OF THIS PROJECT GUTENBERG EBOOK MANY-STORIED MOUNTAINS ***
-
-
-
-
-Produced by Stephen Hutcheson, MFR and the Online
-Distributed Proofreading Team at http://www.pgdp.net
-
-
-
-
-
-
- ★GPO:1978-261-215/3
- For sale by the
- Superintendent of Documents,
- U.S. Government Printing Office,
- Washington, DC 20402.
- Stock Number 024-005-00709-1.
-
- Library of Congress Cataloging In Publication Data
-
- Beaumont, Greg, 1943-
- Many-storied mountains.
-
- (Natural history series)
- 1. Natural history—Montana—Glacier National Park.
- 2. Glacier National Park. I. Title. II. Series: Natural history series
- (Washington, D.C.)
- OH105 M9B43 500.9′786′52 78-606071
-
-
-
-
- Many-storied Mountains
- The Life of Glacier National Park
-
-
- Written and
- photographed by
- Greg Beaumont
-
- 1978
- Natural History Series
- Division of Publications
- National Park Service
- U.S. Department of the Interior
-
- [Illustration: The landform of Glacier National Park is a monument
- to the power of moving ice. This view from Stoney Indian Pass is
- startlingly different from the scene of a million years ago, when
- the glaciers of the Pleistocene were sculpturing the land. Only the
- higher peaks would then have been visible above the blanket of ice
- that flowed like a slow-moving river down into the Mokowanis Valley
- and out into the Great Plains beyond.]
-
-
-About This Book
-
-This natural history of the mountain wilderness called Glacier National
-Park is not a guidebook, but provides an overview of the ecology of the
-region. At the same time, it is a personal statement, revealing one
-individual’s response to this rugged, delicate land.
-
-For their consistent cooperation and helpfulness, I wish especially to
-thank Chief Naturalist Ed Rothfuss and his capable staff. Technical and
-field assistance came from many; for special thanks, I would like to
-single out Art Sedlack, Francis Singer, Bert Gildart, Walt Martin, Craig
-Kuchel, and Danny On. The manuscript profited greatly from the criticism
-of Douglas Chadwick, to whom I am deeply grateful.
-
- —G.B.
-
-
-The National Park Service Division of Publications gratefully
-acknowledges the financial support given this book project by the
-Glacier Natural History Association, Inc., West Glacier, Mont.
-
-
-
-
- Contents
-
-
- Song of the High Peaks 1
- Cycles and Seasons 5
- Bedrock: The First Story 6
- The Rising of the Sun and the Running of the Deer: A Glacier
- Year 39
- Plant-and-Animal Communities 43
- Over Going-to-the-Sun Road 44
- Groves and Grasslands: The Prairie Sea 46
- The Forest 70
- Scrub Forest 105
- Tundra 109
- Water Communities 114
- Shooting Stars 121
- Appendix 125
- Pictorial Features
- The Mountains of Glacier 10
- The Forests of Glacier 48
- The Vital Predator 78
- Protective Coloration 84
- _Ursus arctos horribilus:_ The Vulnerable King 88
- Bald Eagles and Kokanee Salmon: A Recent Gathering 92
- A Triumph of Many Colors 96
- Fire Succession: Key to Continuity 100
-
-
-Illustrations by Celia Strain/Morgan-Burchette Assoc.
-
- [Illustration: Soaring eagle]
-
-
-
-
- Song of the High Peaks
-
-
-April again and the wind turns on the Great Plains. Wedges of geese,
-high and determined, began this storm of spring, their voices sharp as
-the morning frost. Sicklebills cry to claim the land, sandhill cranes
-wheel and talk overhead, and everywhere the killdeer shout.
-Pasqueflowers push the bleak soil aside, beginning the westward rush
-that I must join, seeking again the sight of mountains.
-
-In Glacier National Park the land is folded up. On the east, Chief
-Mountain, Curly Bear, and Rising Wolf break the prairie’s hold. When the
-early French fur trappers saw these peaks glistening in the distance
-with summer-long snows and perpetual ice, they named this region “the
-land of shining mountains.” But for all the ice and snow that reflect
-the summer sun, the park’s present glaciers are but snowflakes compared
-to the mighty rivers of ice that carved this land. Glaciation, the
-magnificent sculptor, left its bold signature everywhere, and this park
-honors with its name the force that shaped it.
-
-But the essential excitement of this land is more than cliff face,
-spire, and sudden storm. It comes to you when you realize that here is
-an aggregation of dramatically differing life zones, where a day’s walk
-can easily take you from prairie and forest to treelimit and tundra;
-where a dense forest of redcedar and hemlock, similar to the rain
-forests of the Pacific coast, exists a score of kilometers from the
-great prairie sea.
-
-Or it comes when you discover that these mountains—young and sharp with
-shadows, snow-jeweled and newly gowned with forests—are chiseled from
-the oldest unaltered sedimentary rocks on earth.
-
-I come from the prairie and love its broad strokes; I’ve learned to hear
-the singing in the grass and to see those long, slow seasons soar the
-level horizons like gliding hawks. But here I learned to match my days
-against a wild earth, and in me grew the mysterious need to know a
-mountain from its every side. Mountains that wear the dawn like yellow
-hats, repeated in the named and nameless lakes. Mountains that stretch
-the storms between them and balance rainbows ridge to ridge.
-
-I must see again the secret forest places, where the paleflowered
-wood-nymphs hover like a breath, and know once more the endless meadows
-painted camas blue.
-
-I need the perfect freedom of this land, to be able to say, _today I
-will climb Siyeh_: to stand, for a time, on the rugged shoulders of this
-upright earth.
-
- [Illustration: The sharp spire of little Matterhorn and the broad
- face of Mt. Edwards loom above Going-to-the-Sun Road in the upper
- McDonald Valley. During warm days in spring the valleys of the park
- resound with the thunder of avalanches.]
-
- [Illustration: Twilight view.]
-
-
-
-
- Cycles and Seasons
-
-
-Bedrock: The First Story
-
-On the trail that connects the Logan Pass visitor center to Hidden Lake
-overlook there is a shallow pond. Near Hidden Pass, it collects its
-meltwater from the Continental Divide and sends it down the shallow
-gorge that drains the Hanging Gardens; as a waterfall it plunges into
-the upper St. Mary Valley where it becomes Reynolds Creek; joined by
-other tributaries, it continues its long journey to Hudson Bay.
-
-The surface of this pond is seldom still, for the wind treats it like a
-sea. Because the water is shallow, the wave action wrinkles the bottom
-mud into ripple patterns, mimicking the churning waves.
-
-I like to come here early in the morning. Sometimes, arriving before the
-wind awakes, I catch reflections of the surrounding mountains. Beyond
-the low bench of Logan Pass the Garden Wall begins, running northward
-with the Divide. In the eastern valley the pitched peak of
-Going-to-the-Sun hunkers in the morning light like a tensed warrior. To
-the south, the incisor Bearhat, beautiful cloud cutter of Hidden Lake
-Valley, juts above the nearby saddle of the pass. But over this place,
-standing as fresh monuments to an age of ice, tower the cliffs of
-Clements and the pyramid Reynolds.
-
-I am sitting on a wedge of red rock. Its surface exhibits a wrinkled
-pattern identical to the ripples in the soft mud of the shallow pond.
-The distance is not great; with a stick I could reach out and touch the
-mud. Yet this represents a gulf no bird can fly, for between the ripples
-of this rock and the ripples of this mud lie billions of vanished
-mornings, a constellation of years.
-
-These red, green, tan, white, black and purple bands of rock that layer
-Glacier’s mountains comprise the oldest unaltered sedimentary rocks on
-Earth. They were laid down in Precambrian time, more than a billion
-years ago, when life was just beginning, as the deposits of an inland
-sea.
-
-For millions of years, sand, mud and carbonates washed into the ancient
-sea, compressing the lower layers into mudstones and limestones,
-building up a sediment thickness that may have been as much as 10,000
-meters (_see_ metric conversion table on page 136).
-
-When we look at the sharp contours of Glacier’s mountains, we see the
-evidence of uplift, overthrust and glaciation. But on the geologic clock
-these are recent events, a mere eyeblink of time ago. For the vast
-majority of years, the rocks lay undisturbed and level beneath the sea
-and land.
-
-To understand better the tremendous time scale these rocks represent, we
-need a way to visualize the vast collection of years. If we were to make
-a movie of these geologic events, we would first need to determine how
-many years each minute should represent. Since the Pleistocene lasted
-about 3,000,000 years (its four ice ages sculpting the present muscle of
-this land), let us make each minute portray a million years. To
-chronicle these rocks we will then need a film 60 hours long!
-
-Not until the fifty-seventh hour of our film will the Mesozoic lowlands
-begin to bulge with the coming Rocky Mountain chain. During the long
-preceding hours we would have seen little else but sea—withdrawing,
-advancing, deep and shallow; yellow, green, and brown with great
-colonies of algae. Unseen below the water, lava has spilled out
-occasionally on the sea bottom; once, it intruded between the rock
-layers below, forming the conspicuous, 60-meter-thick band of black
-diorite that we see today on many mountain faces in Glacier.
-
-During this time of initial uplift an amazing process is going on deep
-underground. A major fault has developed, fracturing the buckled layers
-of rock. A vast mountain plate begins to slide eastward, over-riding and
-submerging the rock layers to the east and opening the wide trench that
-is today the North Fork Valley. Known as the Lewis Overthrust, this
-gigantic earth-force has created an unusual situation: ancient rock
-strata lying atop recent rock strata.
-
-Now less than 3 minutes of film remain. The arrival of the ice is
-imminent. We look at the landscape of featureless mountains and wonder
-at the dramatic difference that this last 3 million years will make. We
-do not see the familiar forests and lakes, the savage peaks, and the
-broad, deep valleys of this present land. These mountains are gentle,
-arid, and shallow-valleyed. The vague outlines are there; we recognize
-the general alignments of the drainage systems, the bloated domes from
-which sharp peaks will be cut. The mountains are connected to one
-another by blunt ridges and smooth saddles, and the shadows they cast
-are dull, dunelike.
-
-Suddenly the ice is there, filling the landscape, with only the
-mountaintops protruding. Four times in these last 3 minutes of film the
-ice sheets advance and retreat, each time leaving an altered landscape.
-Strange lakes and forests fill the gaps between the glacial invasions.
-Then we see the mountains we now know come into being rapidly, as if the
-land were being hacked into shape by giant cleavers.
-
-After this flicker of Pleistocene time, the film ends, the forests
-return, and familiar lakes shine beneath the sun again—these lakes and
-forests we had thought to be timeless.
-
-
-■
-Up springs the morning wind from Hidden Valley, making the nearby alpine
-fir branches whiz with its passing and shattering the perfect reflection
-of Bearhat Peak on the pond. From where I sit, it is a short distance to
-Hidden Pass; so I leave the pond and walk to the overlook to see again
-the fine basin quarried by an ancient glacier.
-
-Hidden Lake, deep, far below, so blue, fits into its cliffed, crooked
-valley like a polished boomerang. Closely ringed by ridge and
-peak—distant Sperry Glacier and pointed Gunsight peering up from the
-southern jumble, and broad Bearhat impossibly close—this lovely lake is
-almost lost amid such sharp proclamations of rock. Its outlet gorge
-gives a narrow view across the angled, hidden valleys of Avalanche and
-McDonald, past the pyramid of Stanton, to the low, faraway undulations
-of the Whitefish Range.
-
-Glaciation is a cruel master of mountains, biting deeply into their bulk
-and leaving sheer, spectacular contours when the glaciers disappear. The
-landforms here attest to their power, everywhere exhibiting the effects
-of glaciation.
-
-In eating back the mountain headwall, alpine glaciers formed rounded
-depressions, called cirques. Unlike the narrow clefts left by running
-water, these broad, deep basins look as though they were made by
-ice-cream scoops gouging into the rock. Hidden, Ptarmigan, Iceberg, and
-Avalanche Lakes sit in well-developed cirque basins, and many mountains
-are dimpled by the beginnings of other cirques—the conspicuous
-amphitheater on the south shoulder of Heaven’s Peak, for example.
-
-Occupying all major drainage systems, glaciers modified the contour of
-the valleys, changing them from their narrow, stream-cut V-shapes into
-broad U-shapes. Into these wide main valleys, waterfalls plunge from
-higher, smaller valleys. Like rivers, flowing glaciers have tributaries.
-Lacking the ice mass and cutting power of the main glaciers, these
-tributary ice fingers could not bite as deeply into the bedrock. When
-the ice melted, hanging valleys were left stranded high above the main
-valley floor. Hidden Lake sits in one of these hanging valleys, and from
-it Hidden Creek plunges 750 meters into Avalanche Basin toward McDonald
-Creek.
-
-On my many previous visits to this pass I have been too busy enjoying
-the wildflowers, the weather, or the scenery to realize what an open
-textbook of glaciation is everywhere displayed.
-
-I stand here on a small saddle of a pass. Wherever glaciers met, passes,
-or cols, were created. A high, notched pass like this one (or
-Swiftcurrent or Gunsight) reveals recent connections. Broad, lower
-passes, such as Logan, resulted where the ice early overran the mountain
-ridge and had a chance to work longer.
-
-Where two glaciers worked on opposing sides of a ridge and failed to
-meet, they formed an arête—a thin, steep-walled remnant resembling a saw
-blade. Another ice age would probably consume the park’s many thin
-arêtes, such as the Garden Wall and Ptarmigan Wall; but it would also
-create new ones from existing ridges.
-
-Further testimony to the sculpting power of ice is presented by Mt.
-Reynolds, looming to the east. The most dramatic feature of a glaciated
-landscape is the pyramid-shaped mountain called a horn—and Reynolds is a
-perfect example. Horns were formed when three or more glaciers cloaked
-the mountain, excavating its sides toward its core and gradually
-transforming its original domed shape into a sheer-sided peak. Glacier
-has many remarkable horns, from the sleek spire of St. Nicholas in the
-south to exquisite Kinnerly in the northern Kintla valley.
-
-Sperry Glacier stares back at me from the flank of Gunsight. Glaciers
-found in the park today are not remnants of the last ice phase, which
-ended here about 8,000 years ago, but are newly formed, having come into
-existence some 4,000 years ago. They reflect a cooling trend in the
-present climate.
-
-Shrinking steadily from their period of greatest extent in the middle of
-the last century, these modern glaciers finally stabilized in the late
-1940s and since then have shown only a slight increase in area.
-
-Movement distinguishes glaciers from icefields, and the movement of ice
-is a force on as well as a feature of a landscape. A glacier excavates
-by abrading and plucking at the rock. Alternately melting and freezing,
-ice at the headwalls plucks out blocks of rock. Ultimately the rocks are
-deposited along the sides or at the feet of the glacier as moraine
-debris. But as they move in the grip of the ice, they constantly abrade
-the rock surfaces they encounter. Polished rock beds of past glaciers
-show striations—grooves gouged by rock fragments imbedded in the moving
-ice.
-
-Flow rate of a glacier depends upon the thickness of the ice and the
-degree of slope. Under tremendous pressure, ice becomes plastic, like
-thick taffy. Unlike kilometer-thick continental glaciers, which may move
-a hundred meters a day, small alpine glaciers seldom progress more than
-two or three centimeters per day.
-
-Although a glacier moves, it gets nowhere if in a state of
-equilibrium—when annual melting equals annual accumulation. Snow mass
-gained at the sun-shielded headwall is usually lost as melt at the
-exposed snout. Glaciers such as Sexton or Weasel Collar, whose snouts
-perch on cliff edges, also lose mass by calving. Thunder you hear on a
-late-summer day near such a glacier may actually be the sound of ice
-pushed off from the lip of a cliff.
-
-Walking back to the visitor center, I suddenly stop where the trail
-skirts the steep moraine of Mt. Clements. From the opposite side of the
-moraine five mountain goats have appeared. Spotting me on the trail
-below, they also halt. But before I can get to my camera they are off in
-a stiff-legged gallop, running in single file along the crest of the
-moraine to the distant safety of the mountain face.
-
-Moraines are ridges of rock debris piled up along the edges and
-terminuses of glaciers. Like a bracelet lying against the wall of this
-mountain, the circle of steeply piled debris marks the extent of a
-small, recently vanished glacier. Ghost of the power that once resided
-here, a stagnant icefield lies beneath the confining walls of the
-moraine. The recent accumulation of these rock fragments is a mighty
-accomplishment, attesting to the force of moving ice.
-
- _continued on p. 38_
-
- [Illustration: The Mountains of Glacier
-
- Lying astride the Continental Divide in the Northern Rockies,
- Glacier is above all else a mountain park. The special beauty of its
- lakes, streams, and forests derives from the microclimates and
- varied topography and soil produced by mountain-building and
- mountain-eroding forces.]
-
- [Illustration: Overthrust Mountains
-
- 1 A hypothetical block of the Earth’s crust in the region of Glacier
- National Park as it existed more than 60 million years ago. The two
- layers shown actually represent many strata of sedimentary rocks.
-
- 2 Lateral pressure begins to force the rock layers to buckle.
-
- 3 A large fold has been created, forcing the rock strata to double
- over and overturning some layers. A break, or _fault_, is forming at
- the plane of greatest stress.
-
- 4 The break has been completed and the strata west of the fault have
- slid eastward, up and over the rocks east of the fault.
-
- 5 The Glacier landscape today. Throughout the millions of years
- during which the folding, faulting, and overthrusting have been
- taking place, the process of erosion has continued; a thousand
- meters of stratified rocks have been worn away, so that only a
- remnant of the overthrust layers can be seen today. Because
- Glacier’s eastern slope represents the eroded face of the overthrust
- block, the mountain range rises precipitously from the prairie, with
- no foothills breaking the abrupt transition from open prairie to
- mountain valley.]
-
- [Illustration: The peaks in this photograph (a view to the northwest
- from Marias Pass) are remnants of the overthrust block, which moved
- eastward. The dividing line between the light-colored rocks and the
- gray talus slopes beneath them is the Lewis Overthrust Fault.]
-
- [Illustration: Glaciation
-
- 1 This is how the landscape in this region might have appeared
- before the onset of the Pleistocene, millions of years ago. Note the
- stream-eroded, V-shaped valleys. The climate at that time was dry.
-
- 2 Glaciers began to form high on the peaks, crept downward, and
- joined to form larger glaciers.
-
- 3 After many centuries of glaciation, tributary glaciers have cut
- back into the peaks, forming basins called _cirques_. Thick
- glaciers, moving rapidly and carrying rock fragments, have abraded
- the main valleys’ floors and sides, widening and deepening the
- valleys into characteristic U-shapes.]
-
- V-shaped Valley
- Tributary Glacier
- Unglaciated Peak
- Headwall
- Meltwater Stream
- Nose of Glacier
- Crevasse
-
- [Illustration: 4 In the present landscape, free of all but remnant
- glaciers, small lakes called _tarns_ occupy many of the cirque
- basins; and waterfalls plunge into the main valleys from higher,
- shallower, tributary valleys, called _hanging valleys_. _Alluvial
- cones_—recent accumulations of rock debris—have begun to build along
- the valley walls. In the main valley, a _moraine_ (a deposit of rock
- materials left by the retreating glacier) has formed a dam that
- holds back a large lake.
-
- During all this time, all parts of the terrain not buried under ice
- and snow have been weathered and eroded by nonglacial forces. Thus
- the contours of the jagged peaks and sheer cliffs have been
- softened.]
-
- Unglaciated V-shaped Valley
- U-shaped Valley
- Hanging Valley
- Cirque
- Tarn
- Alluvial Cone
- Moraine
- Morainal Lake
-
- [Illustration: Glacial landforms can be identified in this view of
- the Mokowanis Valley.]
-
- [Illustration: A Divided Climate
-
- Because of an eastward flow of cool, moist Pacific air masses, the
- climate of northwestern Montana, including the western portion of
- Glacier National Park, differs from that of other portions of
- Montana. As a result of increased precipitation, Glacier’s western
- valleys support a rich flora, more typical of the Pacific Northwest.
-
- West
-
- Moist Pacific air
-
- As the moisture-laden Pacific winds are pushed up the windward
- slopes of Glacier’s mountains, the air cools and water vapor
- condenses, forming fog or clouds. Rain or snow begins to fall as the
- air continues to rise and cool. By the time the air mass reaches the
- crest and flows down the leeward slopes, most of the moisture has
- been lost.
-
- Western slopes average about 70 cm. of precipitation at elevations
- between 900 and 1,100 m. Upper elevations average 200 to 250 cm.,
- mostly in the form of snow; and 300 to 500 cm. is common.
-
- East
-
- Dry chinook winds
-
- Eastern slopes, under the influence of Continental air masses,
- receive less annual precipitation. West Glacier’s annual average is
- 66.5 cm. Babb, a small town east of the park, averages 49.3 cm.
- Frequent high winds east of the Divide further reduce moisture
- through evaporation.
-
- Exposed to Arctic air masses flowing down from Canada, locations
- east of the Divide also suffer more severe winter conditions than do
- protected western valleys. Average January temperature is -5°C at
- West Glacier, -8° at Babb.
-
- Moreover, 80 percent of the winter days in the western portion of
- the park are overcast, a condition almost identical to that of
- Seattle, Wash. This serves to moderate winter temperatures and to
- minimize evaporation.]
-
- [Illustration: Moss campion and mountain forget-me-not colonize a
- fellfield. Fellfields are rocky alpine sites that are slightly less
- than 50% bare rock, interspersed with such plant pioneers as cushion
- plants, mosses, and lichens.]
-
- [Illustration: High lakes generally occupy cirque basins. These
- depressions in the valley bedrock, quarried by glaciers, are deepest
- near the headwall where ice thickness was greatest. Cold and deep,
- and ice-free only weeks each year, tarns cannot support vascular
- plants or vertebrates. Iceberg Lake, shut off from the sun most of
- the year by the encompassing 1,000-meter walls of Mt. Wilbur and the
- Ptarmigan Wall, is never completely free of ice.]
-
- [Illustration: Lake McDonald, 16 kilometers long, 2 kilometers wide,
- and 134 meters deep, is the largest lake in the park. Its basin is
- the classic U-shaped glacial valley. Forested lateral moraines on
- either shore gently rise 600 meters above lake level.
- Going-to-the-Sun Road snakes along the eastern shore, and Logan Pass
- lies near the center of the photograph, behind the peaks of the
- Lewis Range.]
-
- [Illustration: Subjected to the drying and shaping effects of wind
- both winter and summer, this Douglas-fir, growing in the prairie
- community near St. Mary, will attain neither the symmetrical shape
- nor the great size of the Douglas-firs growing in moister, more
- sheltered sites on the western slopes of the Continental Divide.]
-
- [Illustration: Freeze-and-thaw cycles continually fracture and
- loosen rocks along joints, making them subject to removal by the
- actions of water, gravity, and avalanche. The resulting fans of rock
- debris (talus cones) indicate the extent of erosion since the
- withdrawal of the Pleistocene glaciers.]
-
- [Illustration: Although moving water is an agent of erosion—the
- primary destructive force of mountain masses—it also permits life.
- Even small watercourses, such as this freshet, abound with plant and
- invertebrate life.]
-
- [Illustration: Going-to-the-Sun Mountain, towering above the St.
- Mary Valley, from the trail to Siyeh Pass. The coniferous forest at
- its base and the alpine tundra plants at its summit are closely
- juxtaposed in space; but if these two communities grew at the same
- elevation they would be separated by thousands of kilometers. Hiking
- from St. Mary Lake up to Siyeh Pass is going, in effect, from
- Montana to the Arctic Circle; but here the life zones are compressed
- and sharply divided rather than extended and overlapping.]
-
- [Illustration: Setting moon and snow shelf near the summit of
- Heavens Peak. Note stratification of Precambrian sediments.]
-
- [Illustration: Western redcedars line the shores of Lake McDonald.
- Because of prevailing air currents from the Pacific coast, winters
- in the protected western valleys are moist and comparatively mild,
- and this deep body of water freezes over an average of only one
- winter in four.]
-
- [Illustration: Moose often follow the spring snowmelt upwards to the
- headwaters of drainages. This bull will remain at Thunderbird Pond,
- at the base of Brown Pass, until autumn, when it will return to its
- Waterton Valley wintering ground.]
-
- [Illustration: Because of the high reproductive capacity of insects
- and small mammals, if all their offspring survived the earth’s plant
- life would be consumed within one year. This is prevented by natural
- controls such as predation and parasitism. The American kestrel
- (“sparrow hawk”) feeds primarily on large insects and on small
- rodents such as the meadow vole here.]
-
- [Illustration: Gray jays are found in the deep coniferous forests of
- the park. In some parks gray jays, or “camp robbers,” loiter about
- campgrounds and picnic areas begging or stealing food. In Glacier,
- however, they are seldom noticed as they search out seeds, berries,
- and insects.]
-
- [Illustration: A generalized predator, the coyote will eat almost
- anything, from berries to carrion. When man eliminated most of the
- coyote’s enemies and competitors, including the wolf, grizzly, and
- cougar, it enlarged its range to fill the void. Intelligent and
- social, the coyote thrives despite man’s persecution. Although most
- numerous in the prairie community, it ranges up to timberline.]
-
- [Illustration: The spruce grouse is a year-round resident of the
- spruce/fir and lodgepole communities. It forages on the ground for
- seeds and insects, in winter turning to needles. Several other
- species of grouse occupy different habitats in Glacier.]
-
- [Illustration: Chipmunks are found in every community, from prairie
- to tundra, in Glacier. Each of the park’s three very similar species
- has its preferred habitat. The diurnal counterpart to nocturnally
- active mice, which have the same diet of seeds, berries and
- occasional insects, chipmunks adapt easily to the presence of people
- and become nuisances if encouraged by handouts. Feeding rodents is
- dangerous and is harmful to them. By altering their diets and
- blunting their cautious instincts, daily exposure to “free lunches”
- makes the animals less fit to face the harsh realities of their
- natural environment.]
-
- [Illustration: Unlike whitetail deer, which remain in lowland areas
- all year, mule deer range upward into high meadows during the
- summer. The bucks, especially, are wanderers and travel together.
- Velvet antlers, worn during the time of summer sociability, presage
- the autumn contests to come.]
-
- [Illustration: The checkerspot butterfly belongs to the most diverse
- group of animals on the planet—the insects, whose importance can
- hardly be overestimated. They not only help recycle nutrients in the
- living community and provide an abundant food base for other
- lifeforms, but are instrumental in pollinating most of the earth’s
- terrestrial plants.]
-
- [Illustration: Alpine vegetation must be able to survive freezing
- temperature during the growing season, since winter conditions are
- possible even in summer. Early bloomers, such as the glacier lily,
- endure repeated snowfalls during the unstable weather conditions of
- June.]
-
- [Illustration: Unlike mountain goats, these bighorn rams will desert
- the alpine zone at the approach of winter; they will join other
- bighorns congregating in the lower valleys.]
-
- [Illustration: In November bighorn sheep rams end their summer-long
- isolation from the ewes, move down from the higher slopes, and begin
- a bloodless but taxing ritual of strength and endurance to determine
- the harem master. The sharp reports of clashing horns may carry for
- kilometers, and the contests continue for weeks until the dominant
- ram emerges. (Note the Many Glacier hotel complex in the valley
- below.)]
-
- [Illustration: Hummingbirds, like shrews and other small-bodied,
- warm-blooded animals, exist at the theoretical thresh-hold of life.
- Because of their small size, body volume is not large enough in
- relation to surface area to prevent a rapid loss of body heat. To
- compensate for this, metabolic rates must be high; food is rapidly
- processed and used up. Thus, since fat reserves are not practical on
- such small animals, they must eat at frequent intervals.
-
- Two species of hummingbirds—the rufous and the calliope—are found in
- Glacier. Pictured is a female rufous (which weighs about the same as
- a dime) landing on its lichen decorated nest to feed its two young
- on a protein-rich mixture of nectar and small insects.]
-
- [Illustration: The insect-eating yellowthroat prefers moist
- habitats. Unlike many of its treetop-dwelling relatives, this tiny
- (10-11 cm.) warbler is usually seen near or on the ground.]
-
- [Illustration: Bands of bighorn ewes and lambs do not summer as high
- as the rams and are often encountered in the scrub-forest zone. Note
- the gnarled limber pine in the foreground of this photograph taken
- on the south face of Altyn Peak.]
-
-Reaching the mountain wall, the goats scramble upward to a ledge,
-sending scree streams pouring from several clefts. Encountering a
-narrow, steep snowbank, they do not hesitate but continue across the
-slope. Above the rock fingers of this peak the gathering clouds grow
-black. A sudden crack of thunder hurries me down the trail.
-
-Although geologically young, the Rocky Mountains in Glacier are composed
-of soft sedimentary rocks that are easily assailed by the many agents of
-weathering and erosion. If not rejuvenated by continual uplift, these
-magnificent peaks will glimmer but briefly in the long memory of the
-planet.
-
-Already the sharp countenance of this land is being softened by the
-ongoing forces of erosion. Chief among these is water, which attacks the
-mountains everywhere. In addition, frost action continually exploits
-rock fractures, breaking down blocks of rock into talus and scree.
-Avalanche and rockfall sweep down the slopes. Layers of softer rock
-erode quickly, undercutting more resistant rock and creating overhangs
-which gravity, in time, will collapse.
-
-The lashing rain catches me on this sun-and-storm-contested pass. Ice,
-gravity, wind, and especially water—all attack a land that dares the
-clouds.
-
-
-The Rising of the Sun and the Running of the Deer: A Glacier Year
-
-As if to make up for the days-long darkness of this last blizzard, the
-peaks today wear snow plumes—long, graceful trails of white, curving up
-into an ice-blue sky. Yesterday the snow-mad wind raced through the
-forest. Today the motionless trees are cloaked in heavy, glistening
-robes, the leafless aspen and young larch bent down.
-
-Moderate snowfall helps many plants and animals survive the winter. For
-ground dwellers it provides insulation from the wildly fluctuating
-winter temperatures encountered east of the Divide, protecting the
-hibernators and providing cover for the many small mammals that remain
-active during the winter. Wind-swept ground freezes deep; but under a
-mantle of snow life-sustaining heat is trapped, permitting many animals
-to survive and allowing the work of decomposers to continue.
-
-But this has been a winter of too much snow and too many temperature
-extremes. The heavy snowpack has forced the sharp-hoofed deer to yard up
-in great numbers; unable to range freely in deep snow, they are forced
-into smaller and smaller confines where their numbers allow them to
-break and maintain trails. But in time they exhaust the food supply.
-Younger deer, unable to reach the increasingly higher browse line,
-starve first. Then the does, heavy with unborn fawns, grow weak and fall
-to predators. So the imprisoned herds dwindle quickly this year,
-sometimes less than a kilometer from plentiful browse.
-
-Deep snow is also death for many seed-eating birds. As they are unable
-to scratch for food, their body furnaces quickly fail, and during a
-night of cold wind their fluffed corpses drop into the snow.
-
-Exposed to the noon sun, the snow surface thaws; when refrozen, it is
-restructured to crystalline ice. If snow repeatedly thaws and freezes,
-an ice barrier is formed, shutting off vital air exchange. Plants are
-then subject to rot, and micro-animal life is smothered. Travel beneath
-the snow is made more difficult for mice and shrews and they are
-deprived of food and cover. Under such conditions their numbers rapidly
-decline.
-
-But while many starve in a winter of deep snow, others benefit. The
-exposed traffic of small mammals is to the owl’s advantage. Foxes and
-coyotes more easily run down rabbits and hares on crusted snow. Deer
-and, to a lesser extent, wapiti and moose—their hoofs punching through
-the snowpack—swiftly tire in deep snow and become helpless before cougar
-or wolf, whose lighter weight is supported by the crust.
-
-Grim as this winter’s toll becomes, enough will survive to begin the
-process of renewal in spring. Last winter, a season of light snow, was a
-time of hardship for predators. The deer remained strong, the wapiti
-remote on high, windswept ridges, and the small mammals hidden.
-
-Only the water ouzel, winter after winter, seems not to notice the
-hardships of the season. Lord of his small world of open water, he sings
-in February, wading and swimming his diminished stream to find a
-never-failing supply of water insects and small fish. It is a voice of
-spring—glad, wild, continual as the moving water—an incongruous song in
-this winter-shrouded land.
-
-But with the growing stature of the sun, the grip of winter softens. The
-firs and spruce send their loads of snow sliding to the ground. Streams
-begin to sing again and soon the lakes increase, the booming of
-splitting ice breaking the silence of the valleys. Avalanches thunder
-down the steeper slopes, carrying trees to the swollen streams. Rivers
-hiss and rage, speeding the debris along. A spring that comes too
-suddenly will bring flood to lower elevations.
-
-Snow geese thread through the valleys, and ground squirrels tunnel up
-through snow to find invasions of birds returning from the south. Soon
-the three-petaled wakerobins appear, chasing the snowline up the ridges.
-Glacier lilies and Calypso orchids are next, and with the shooting stars
-spring arrives.
-
-The melting snow releases a new group of animals to populate the
-winter-thinned land. Up come chipmunks. Bears reappear. Young red
-squirrels, helpless and blind, squirm in their nest holes. Hidden dens
-rustle with pups and kits. Soon warm days will bring them out and the
-business of learning to cope with their world will begin.
-
-All life responds irresistibly to the growing strength of the Sun.
-Cottonwood, willow, and maple come into flower and unfold new leaves;
-green needle clusters spot the limbs of larches that in winter had
-seemed lifeless snags among the other conifers. Beneath the soil of
-prairie, meadow, and forest, in the mud of lakes and ponds, other life
-stirs; armies of insects, spiders, crustaceans, amphibians, and fish
-will strive to complete their life cycles against the formidable odds of
-a predatory world.
-
-Spring reaches higher up the mountains, the lowlands passing into
-summer. Wapiti and mountain sheep follow the rising tide of succulent
-browse up to the high meadows. In forest, grove, and meadow and along
-the stream new fledglings appear—thrush, vireo, hummingbird, waxwing,
-harlequin duck, bluebird, osprey, and flicker—as holes, nests, and
-cavities brim with begging mouths.
-
-In the alpine meadows, where snow overlaps the spring and winter follows
-hard behind the summer, the growing season is short and the climate
-unstable. Sensing the stronger light, flowers push up impatiently
-through the snow and hasten into bloom. Pikas and marmots scurry and
-sunbathe among the rocks of scree slopes.
-
-Summer matures in ripening huckleberries, and the bears that grazed the
-spring grasses now gorge themselves fat. Dry days of August bring
-probing lightning, threatening the forests with fire.
-
-Sweeps of beargrass reach their climax now in the highest meadows. In
-dizzy succession wildflowers set seed. Fat and sluggish, marmots and
-ground squirrels disappear beneath the rocks. The golden eagle must
-search longer each day to find prey within its vast domain.
-
-Autumn lingers in the valleys and on the flanks of low ridges. The
-morning sun glints on hoarfrost, firing the yellow leaves of larch,
-aspen, birch, maple, and cottonwood, and shines on the blood-red berries
-of mountain-ash. Soon a night of killing frost will bring down the
-corpses of insects and spiders by the millions. The reptiles and
-amphibians, being cold-blooded animals, seem out of place in this
-long-wintered land. Unable to maintain body temperatures appreciably
-above their surroundings, they are the first to seek the protection of
-hibernation, collecting in dens or burying themselves beneath the ooze
-of pond bottoms.
-
-Songbirds gather and leave the valleys. The harsh cries of jays sound
-ominous now in the forest. Only the chickadees seem to ignore the long
-tree shadows; their ceaseless conversations carry through the leafless
-underbrush as they busily search for seed.
-
-Velvet has gone to bone, and in these final noon-warm days the rut runs
-through the land. It begins in the valleys in September with the
-joustings of deer and moose and the buglings of bull wapiti puncturing
-the forest silence. By November the higher meadows ring with the
-collisions of bighorn rams who compete for ewes by smashing together
-their massive, curled horns. On high slopes mountain goat billies
-posture and swagger; head to tail, they circle, threatening each other
-with dagger-like horns.
-
-From Flathead Lake, 100 stream kilometers to the south, kokanee salmon
-return to spawn in the clear, cold shallows of McDonald Creek. Gathering
-bald eagles surround the stream, again and again lifting vulnerable fish
-from pool and riffle. Perched by the hundreds along the stream course,
-their white heads and tails glistening against the dark trees, they
-stand out like lanterns strung for a banquet.
-
-Now the stinging wind comes down from the peaks and shuts the lakes.
-Life slows or sleeps. Ptarmigan, snowshoe hare, and longtail weasel, all
-wearing winter white, seek shelter and food in a silent land where
-spring and yellow lilies seem forever lost.
-
-
-■
-All life faces one ultimate challenge: to survive or not, to reproduce
-or fail, to bring one’s kind to tomorrow’s sun or vanish forever. This
-land is harsh. To survive in nature demands skill in the individual,
-excellence in the species, and a chance from the environment.
-
- [Illustration: The mink, a solitary predator associated with
- low-elevation watercourses, preys on anything it can catch and
- subdue.]
-
-
-
-
- Plant-and-Animal Communities
-
-
-Over Going-to-the-Sun Road
-
-I like to begin with St. Mary, a lake the whitecaps love to run. From
-the far passes the several winds gather and collect, arranging long
-lines of white waves for the race downlake. Past the purple scree of
-Mahtotopa and Little Chief they go, white as the headdress of
-Going-to-the-Sun Mountain, colliding, collapsing along the promontory
-snares about the Narrows. Onward they press, spreading out and setting
-sail for the straight rush to the final shore where a line of
-cottonwoods sings with a sound like applause.
-
-Across the lake the timbered ridge starkly contrasts the finger of
-prairie that claims the north shore. This is a flower-glad place, a
-meeting-ground for mountain and prairie plants. Along the road the
-grassland holds the conifers back, allowing only scattered clumps of
-aspens.
-
-Finally, at Rising Sun, beneath the shadow of Goat Mountain, the prairie
-ends and wind-seasoned Douglas-firs announce the coming forest.
-
-There’s excitement now, with the prairie heat gone, the wind scent raw
-with fir and high meadows, honed by waterfall and tall, dank rock. Our
-mountain thirst is never extinguished, and a road that tightens down to
-cliff face and sudden turn brings back to our blood the ancient need to
-go to the highest place.
-
-There is sword-edged Citadel, and the snow-flanked spike of Fusillade
-holding court like a queen in this valley of peaks; then the dome of
-Jackson and the Gunsight notch. Our eyes are kept high, transfixed at
-last by looming Heavy Runner and the distant promise of Reynolds.
-
-Looking for mountain goats, we scan the walls around the sweep of Siyeh
-Bend, catching a glimpse of the trail that crosses the scree to hidden
-Piegan Pass.
-
-Beargrass heads lean out above the road like old men conferring on the
-view. The purple trumpets of penstemon crowd the rocks, and spots of
-Indian paintbrush lead like a blood-trail to the higher slopes.
-
-Intoxicated now, feeling the fresh full force of the wind from Logan
-Pass, we race on. We hardly notice the struggle of the forest in
-Reynolds Creek far below, how it thins and loses strength in its own
-hard climb. We sweep past it on the broad magnificence of this pass.
-
-Level but a moment, the road dips to a shelf on the headwall above Logan
-Creek and swings over the great sculptured cliff of the Garden Wall. For
-several kilometers this masterpiece of a road glides down a constant
-grade, squeezed between rock face and space, twisting into tight
-drainages—a road for storm lovers, wet with spray and snow-seep, its
-quick turns concealing sudden winds.
-
-Mighty, snow-robed Heaven’s Peak appears, taking our attention from the
-Pass-group mountains and the hanging valley that spills Birdwoman Falls.
-Northward is the great array of peaks encircling distant Flattop,
-jumbles of mountains and glaciers. How are we to notice the forest far
-below?
-
-Not until we have passed the Loop and are moving past the blackened
-snags of a recent burn do we realize the stature of this forest. The
-long road down will take us into a valley much deeper than any on the
-eastern side. Near Avalanche Creek are trees we have seen nowhere else
-in the park—giant western redcedars, western hemlocks with their nodding
-tops, monstrous black cottonwoods with bark so deeply furrowed that it
-looks hewn by hatchet.
-
-We take a long ride down the valley, past the low pyramid of Mt.
-Stanton, final peak in the Livingston Range. Near the outlet end of Lake
-McDonald, birch and aspen again appear in numbers, and the road enters a
-crowded stand of lodgepole pine.
-
-Our memories cluttered with mountains, waterfalls, and snowfields, we do
-not quite realize the significance of this 80-kilometer journey. We have
-crossed the boundaries of several different plant-and-animal
-communities, spanning a range of climate that would be encountered on a
-5,000-kilometer north-south journey at sea level.
-
-At first glance the various trees, wildflowers, and animals seem
-randomly distributed, scattered about like the distant mountains. But
-mountainous terrain represents an organized high-rise approach to life.
-From the lowest, most protected valley to the highest wind-and-ice-cut
-summit, the life-forms align themselves, each according to its own
-climatic tolerance.
-
-Here too can be seen the great cycles of nature: fire and regrowth, the
-building of soil and its erosion, the incessant duel of the eaters and
-the eaten.
-
-In the following sections we will spend some time in these various
-communities, from prairie to tundra.
-
-
-Groves and Grasslands: The Prairie Sea
-
-There is something about spring on the prairie that gets me up before
-dawn. I like to watch the seasons change their guard over the landscape,
-from the wintry cold of pre-dawn dark to the spring-scented morning air
-to the hot summer-foretaste of the noon May sun.
-
-Hoarfrost surrounds these patches of pasqueflowers, blue goblets on
-downy stems. On this windless night, frost has formed everywhere,
-reclaiming for a time its vast winter range, sparkling over the green
-handiworks of spring.
-
-But the god of the growing grasslands is the sun, and it now proclaims
-itself, stretching out to make the mountains shine. With its assault the
-frost collapses, becoming bright beads on grass tip and leaf joint by
-which a beetle might refresh itself.
-
-Spring is best perceived ant-level, at its ground beginnings, where the
-bright yellow-green tips of new grass shoots reclaim the winter-blighted
-land. I look closely at a drag line of spider silk; a necklace of
-dewdrops slides down, collects to a moment’s greatness, in which I
-briefly see a curved horizon, the morning sunburst, and myself, before
-it falls away.
-
-Getting up from my prone position, my belly damp from the prairie earth,
-I startle a whitetail jackrabbit; bounding high, it zigzags off. The
-commotion disturbs a distant badger, which faces about from its diggings
-to confront danger in whatever form it might take. It swings its snout
-to scent the air. Somewhat uncertainly, it returns to the business of
-hunting, then hesitates, swings about once more and waits, myopic,
-patient.
-
-Satisfied at last, the spurt of the now distant rabbit lost in its
-brain, the creature snorts a defiance at the mystery and resumes its
-morning gopher hunt.
-
-Overhead a marsh hawk skims past, its flight erratic as a butterfly’s.
-Far away a magpie rattles at the passing hawk and takes flight, briefly
-flashing black and white.
-
-
-■
-It is easy to see only pieces in the natural puzzle—a badger throwing
-dirt, horned larks dipping into wind, black ants dragging the rosette of
-a dead spider—and be satisfied with the scattered scenes. But at last,
-to make it meaningful, we must complete the picture. There is that
-special joy in discovering larger schemes: green plants utilizing
-sunlight; a rabbit building its days at the plants’ expense; the falcon
-tearing the rabbit meat for its young; magpies picking at the fallen
-falcon; and then, in the end, all returning to the earth.
-
-Here on the prairie, as in every plant-and-animal association, the
-ancient drama repeats itself over and over; the distant tundra is a
-drastically different stage with different actors, but the cycle is the
-same. Life depends upon the interaction of all its many forms. Unseen
-bacteria are as necessary to the land as green grass; the meadow vole
-and the coyote are as much a part of the prairie as the grasses.
-
-The secret of life rests in the wonder of photosynthesis. Only green
-plants can manufacture food from the earth’s raw minerals. This is the
-vital first step upon which the great pyramid of animal and plant life
-is built. Using energy from the sun, green plants combine water and
-carbon dioxide to synthesize sugar, and give off oxygen as a by-product.
-The caterpillar takes its energy from the plant tissue, converting to
-protein the sugar and minerals in its body. The caterpillar is then food
-for a spider or other predator. A yellow warbler may take the spider and
-in turn be ambushed by the prairie falcon. Thus the energy produced by
-the plant travels through the food chain. When the prairie falcon dies,
-scavengers—including insects and other invertebrates, birds, and
-mammals—redistribute its wealth among themselves; the rest is decomposed
-by bacteria. Thus, eventually, the nutrients on which the plants depend
-return to the soil.
-
-When we look at any living organism, whether it is plant, herbivore,
-carnivore, parasite, scavenger, or decomposer, we are soon made aware of
-its associations with other living things, each puzzle piece leading us
-to another and another. We begin to see a picture whole—the fox, meadow
-mouse, grasshopper, bunchgrass, and sparrow hawk—all interlocked.
-
-Geologically speaking, grasslands are a recent development. As the Rocky
-Mountains were being uplifted, the prevailing warm, moist climate began
-to change. The rising mountain mass intercepted moisture-laden winds
-that blew in from the Pacific, creating a rain shadow that lengthened
-eastward as the mountains rose higher. A continental climate,
-characterized by severe winters and dry, wildfire summers gradually took
-shape, extinguishing the great forests that had grown across the
-continent’s interior. Herbaceous plants, which had been evolving amid
-the forests, inherited the land.
-
-Unlike trees, grasses die back to the ground each winter, hoarding their
-life-germ beneath the protecting soil. Growing not from the tip but from
-the joints, grasses regenerate quickly after fire or grazing. Suspension
-of the normal metabolic processes enable the grasses to go dormant and
-thus survive periods of severe heat and drought.
-
-Although the great prairie sea washes up against Glacier’s eastern
-boundary, with estuaries probing into the mountain valleys on the drier,
-south-facing slopes, the grassland community comprises less than 5
-percent of the land area of Glacier National Park. This includes the
-puddles of prairie west of the Divide that interrupt the dense
-coniferous forests along the North Fork of the Flathead River.
-
-From the pasqueflowers that bloom in early May to the asters and
-goldenrod of September, these summer-long gardens of grasses and flowers
-lean with the wind. Here are timothy, oatgrass and the
-bunchgrasses—rough fescue, bluebunch fescue, and bluebunch wheatgrass.
-Among the grasses bloom bitterroot, blue camas, lupine, gaillardia,
-balsamroot, cinquefoil, sticky geranium, and wild rose.
-
- _continued on p. 68_
-
- [Illustration: The Forests of Glacier
-
- From the lush redcedar-hemlock forest in the McDonald Valley to the
- subalpine fir, whitebark pine, and Engelmann spruce struggling for
- existence near treeline, the forests of Glacier reflect the
- conditions of temperature, exposure, soil, and drainage prevailing;
- and each forest has its characteristic association of understory
- trees and shrubs, herbaceous ground cover, and vertebrate and
- invertebrate animal life.]
-
- [Illustration: Life Zones
-
- Many physical and climatic factors determine the range of Glacier’s
- plant-and-animal communities. Boundaries between communities are
- seldom sharply defined, but rather merge together in broad zones of
- transition.
-
- With elevation gain, average daily temperature drops at the rate of
- 5° per 900 meters. Precipitation, wind velocity, and evaporation
- loss increase. Soil thins. These factors, along with others such as
- fire frequency, north or south exposure, and availability of
- moisture, combine to determine the range of each community.
-
- In the forest community below 1,800 meters, Douglas-fir, lodgepole
- pine, and western larch predominate. In the valleys, Engelmann
- spruce and subalpine fir are found. The somewhat lower and much
- better watered western valleys of the park support western redcedar
- and western hemlock.
-
- Treeline is the upper limit to which the tolerances of trees to
- environmental conditions permit them to grow. Because there are so
- many controlling factors (wind, temperature, exposure to sunlight,
- snow cover, etc.) treeline in the diagram is only approximate. In
- Glacier it averages 2,000 meters. Avalanche chutes or sheer cliff
- walls may suppress it to below 1,500 meters; on protected slopes it
- may be as high as 2,150 meters.
-
- At the eastern edge of the park below 1,200 meters, the forest gives
- way to the prairie community, composed mostly of soft-stemmed plants
- adapted to the conditions of low precipitation that prevail here in
- the rainshadow of the mountain range. Clumps of aspen, found in the
- prairie in sheltered spots, occur here in the transition zone
- between prairie and forest.]
-
- [Illustration: A Mountain Profile
-
- This diagram represents the eastward-facing slope of a hypothetical
- mountain near the eastern boundary of Glacier National Park. Its
- life communities are somewhat different from those of mountain
- slopes at the western edge, chiefly because of the differential in
- annual precipitation.
-
- Illustration: Here, above approximately 2,750 meters, in a realm of
- ice, snow, and barren rock, there is little life.
-
- Alpine tundra
-
- Below 2,750 meters and above 2,000 meters, depending on other
- factors such as exposure to sun and wind and steepness of terrain,
- exists the alpine tundra community, with vegetation similar to that
- of the vast, essentially level, treeless zones of the Arctic.
-
- Scrub-forest
-
- Roughly between 1,800 and 2,000 meters, the dominant vegetation is
- scrub-forest. Trees here are stunted; except in sheltered spots they
- are more or less prone rather than upright. Net growth is slow, not
- only because of the short growing season but also because of the
- pruning effect of icy mountain winds. Very few tree species can
- survive in this harsh habitat.
-
- Coniferous forest
-
- In the forest community below 1,800 meters, Douglas fir, lodgepole
- pine, and western larch predominate. In the valleys, Engelmann
- spruce and subalpine fir are found. The somewhat lower and much
- better watered western valleys of the park support western redcedar
- and western hemlock. See page 54
-
- Prairie
-
- At the eastern edge of the park below 1,200 meters, the forest gives
- way to the prairie community, composed mostly of soft-stemmed plants
- adapted to the conditions of low precipitation that prevail here in
- the rainshadow of the mountain range. Clumps of aspen, found in the
- prairie in sheltered spots, occur here in the transition zone
- between prairie and forest.]
-
- [Illustration: The Forest Community
-
- A forest is organized vertically like an apartment house or office
- building, with layers corresponding to stories. The _canopy_ is the
- branches and foliage of tall trees that form a roof over the
- community. Below the canopy are the _understory_ trees: young
- individuals of the canopy species; and small, shade-tolerant trees
- that will never become part of the canopy. Beneath the understory
- branches is the _shrub layer_, occupied by knee-high-to-man-high
- woody plants; beneath that is the _herb layer_, where most of the
- ferns, wildflowers, grasses, and smaller woody plants grow. The
- _forest floor_ is the zone of mosses, mushrooms, creeping plants,
- and forest litter (leaves, twigs, needles, feathers, bits of bark,
- animal droppings, etc.). The forest has a “basement,” too,
- interlaced by plant roots, mycelia of fungi, and tunnels of myriad
- animals.
-
- Each layer of the forest has its characteristic animal species, but
- most forage over more than one level. Some nest in one story and
- feed in another. The red squirrel races back and forth from the
- forest floor to the highest branches.
-
- The forest community also has a socio-economic organization. Every
- animal (and plant) takes up space and consumes a portion of the
- available nutrients. Each has a place in the community food
- chain—as, for example, _herbivore_, _carnivore_, or _scavenger_.
- Each directly or indirectly affects all the other organisms.
-
- The Forest Community
-
- The role of a species in the community, like the job and social
- function of a person, is its _niche_. Similar species of animals
- have different niches, thus lessening competition for food and
- living space. Thrushes hunt close to the ground; vireos and kinglets
- hunt among the branches; flycatchers snap up airborne insects. The
- flicker feeds upon insects, excavates nesting holes that are later
- occupied by other species such as squirrels and owls, and is preyed
- upon by the great horned owl; its niche is _insect exterminator /
- food for carnivores / homebuilder_. The great horned owl, hunting
- mammals, birds, and reptiles by night, preys on species different
- from those hunted by the goshawk, and thus occupies a parallel
- niche. When it dies, its remains, like those of other animals, are
- decomposed and return to the soil.]
-
- Canopy
- Great Horned Owl
- Yellow-bellied Sapsucker
- Understory
- Flying Squirrel
- Shrub Layer
- Ruffed Grouse
- Herb Layer
- Red Squirrel
- Western Toad
- Forest Floor
- Shorttail Weasel
- Scavenging Insects
- Deer Mouse
- Garter Snake
- Soil Layer
- Ground Squirrel
- Earthworm
- Masked Shrew
-
- [Illustration: Sun, Green Plants, and Animals
-
- The sun is the source of energy for any plant-and-animal community.
- Green plants draw nitrogen and minerals from the soil, and in a
- process called photosynthesis use sunlight to convert raw materials
- (carbon dioxide and water) into carbohydrates (sugar, starch,
- cellulose), giving off oxygen as a by-product. Besides burning
- oxygen, animals depend on plants for food.
-
- Green Plants, trees and shrubs, grasses and sedges, wildflowers,
- ferns, mosses, algae and lichens—are fed upon by animals, which are
- unable to manufacture their own food.
-
- The Redback Vole, like other rodents, pikas and hares, seed-eating
- birds, grazing and browsing hoofed animals, and herbivorous insects,
- derives its energy from the seeds and other parts of green plants
- that it eats.
-
- The Garter Snake, feeding upon the vole, is dependent upon plants
- even though it does not eat them.
-
- The Great Horned Owl, preying upon the garter snake, is one more
- step removed from the green plants—but still dependent on them.
-
- Scavengers such as carrion beetles feed upon the carcass of the owl;
- the remains are then attacked by Decomposers, primarily bacteria,
- that break down the animal tissues into basic organic compounds.
-
- The Soil, enriched by the minerals and carbon and nitrogen compounds
- added to it by the decomposers (and by other processes such as fire)
- supports new green plant growth.
-
- Thus energy derived from the sun flows through the ecosystem in a
- food chain. A plant-and-animal community is a complex, interlocking
- web of such food chains.]
-
- Sun
- Green Plants
- Redback Vole
- Garter Snake
- Great Horned Owl
- Scavengers, Decomposers
- Soil
-
- [Illustration: A Pyramid of Numbers
-
- Necessarily, the number of plants in an ecosystem far exceeds the
- number of plant eaters, and the number of prey species must exceed
- the number of predators. During its lifetime, a golden eagle will
- consume a vast number of lesser animals. The combined mass of prey
- animals necessary to sustain an eagle greatly outweighs the eagle
- itself. Ecologists refer to this proportional relationship of mass
- between each link in the food chain as the _pyramid of numbers_.
-
- The diagram represents a numbers pyramid for the alpine zone.
- Because of its limiting environment, the alpine zone supports a
- lesser plant mass than the forest zone. As a result, the carrying
- capacity of the alpine is less than that of the forest.
-
- 1 Kilo
-
- _Tertiary_ (third-order) _consumers_ are the predators (Golden
- Eagle, Swainson’s Hawk, etc.) that feed upon other predators.
- Because of the 90% loss of energy at each level of the food chain,
- there will be very few hawks and eagles in comparison to the numbers
- of marmots.
-
- 10 Kilos
-
- _Secondary consumers_ are the predators (weasels, shrews,
- carnivorous insects and birds, etc.) that eat herbivores. The
- animals at this level of the pyramid are often—though not
- always—larger than the animals they feed upon. But they are much
- less numerous, because it takes many prey animals to sustain one
- predator.
-
- 100 Kilos
-
- _Primary consumers_ (plant eaters, or herbivores) convert plant
- tissue into animal flesh. In the process about 90% of the energy
- stored as plant food is lost, mostly as heat energy. In the alpine
- community the herbivores include pikas, marmots, ground squirrels,
- and ptarmigan, as well as herbivorous insects.
-
- 1,000 Kilos
-
- _Producers_ are the green plants at the base of the food pyramid,
- manufacturing food for the animals of the alpine community. The
- _biomass_ (total weight) of each level of the food chain is ten
- times (more or less) the weight of the stage above it: 1,000 kilos
- of green plants will produce only 100 kilos of primary consumers.]
-
- [Illustration: Great horned owls are the nocturnal equivalent of
- Cooper’s hawks and goshawks in the low-elevation forests of the
- park. Large and powerful, they are capable of taking prey as big as
- skunks. This young bird, disturbed on its day roost, clacked its
- bill and fluffed its feathers in a menacing manner.]
-
- [Illustration: The only sizable mature stand of ponderosa pine found
- within the park is along the North Fork truck trail. A scattering of
- old ponderosas growing at the lower end of Lake McDonald suggests
- that at one time ponderosa forests were more extensive in this
- region than at present.]
-
- [Illustration: A black bear near treelimit. Bears will eat almost
- anything, from ants to carrion, grass to garbage. Color phases
- include brown and blonde bears. Unlike the larger, more aggressive
- grizzly, which ranges out onto the plains, black bears are strictly
- forest creatures.]
-
- [Illustration: The water ouzel, or dipper, a creature of fast
- mountain water, is admirably outfitted to cope with its demanding
- environment. Stubby wings, chunky body, short tail, and oily plumage
- allow it to walk under water, where it scavenges for aquatic insect
- larvae and small fish. In flying up- and down-stream, ouzels never
- shortcut but follow the winding streamcourse.
-
- As long as there is open water, the dipper suffers no hardship from
- the mountain winter. Then, when the land is shut down and lakes are
- frozen over, this little bird carries on in its mountain-stream
- habitat, plunging into the cold water to find food, and pausing
- occasionally to sing.]
-
- [Illustration: Ouzels construct their nests of living moss on cliff
- faces or ledges where constant spray keeps the moss moist. At
- fledging, the four young of this nest in Avalanche Gorge tumbled one
- by one into the torrent below, to be collected by the adults in
- quieter water downstream. Within a day they appeared to have
- mastered the underwater gymnastics and were feeding on their own.]
-
- [Illustration: From their lowland wintering grounds, wapiti move up
- to higher elevations in spring. Summer range in the park is
- abundant, but winter range is limited; as a result, wapiti have a
- tendency to increase their populations beyond the carrying capacity
- of available winter range. In a severe winter many starve. But in a
- balanced ecosystem such loss is not waste, for the carrion helps
- sustain scavengers; it is an important initial food source for bears
- emerging from hibernation.]
-
- [Illustration: Cedar waxwings nest in moist areas of low valleys
- where fruits and berries are abundant. Although they also subsist on
- insects (which they can capture on the wing), their weakness for
- fruit is so pronounced that the birds will sometimes gorge
- themselves until rendered incapable of flight.]
-
- [Illustration: The Columbian ground squirrel is found at all park
- elevations, from prairie to alpine meadow. Hibernation occupies
- almost three-quarters of its five-year lifespan. Unlike other park
- ground squirrels, it lives in colonies. Although not as tightly
- structured as a prairie dog town, the association is beneficial to
- all members in that danger is readily detected.]
-
- [Illustration: The tundra community is encountered above Preston
- Park on the Siyeh Pass trail. Mt. Reynolds, a classic example of a
- horn, dominates the distant Logan Pass area.]
-
- [Illustration: Camas blooms in the prairie community along the Red
- Eagle road. An important staple, camas bulbs were gathered as food
- by Indians.]
-
-Conspicuous also are many insects—including grasshoppers; flies; ants,
-wasps and bees; butterflies and moths; bugs; and beetles—which fulfill
-important roles as herbivores, carnivores, and scavengers while also
-acting as pollinators for flowering plants and providing an abundant
-food source for other animals.
-
-Below the ground are the tunnels. Burrowing is an important means of
-survival on the open prairie, and life underground is extensive. Some of
-the animals are rarely seen—the northern pocket gopher, for example,
-with a diet of underground insects, grubs, worms, and roots, spends most
-of its life tunneling just below the surface. Others, like the badger,
-leave their burrows during the day to dig for rodents. Most conspicuous
-of the burrowing animals in the park’s grasslands is the Columbian
-ground squirrel. Its alert upright stance has earned it the nickname
-“picket pin.” When danger approaches from the air or on land, its shrill
-alarm whistle passes the warning to others of its kind.
-
-Where prairie and forest meet, a never-ending struggle for dominion is
-waged. The isolated patches of prairie that dot the North Fork Valley
-near Polebridge hold the great forest of the park’s northwest region at
-bay.
-
-This broad valley, floored with coarse glacial outwash and terraced
-downward to the deep channel of the North Fork River, presents a graphic
-battleground between grass and tree. Lining the upper terraces, from
-which they glower down on the dry, well drained grass flats like a line
-of warriors, are the Douglas-fir, western larch, and ponderosa pine.
-Seedling trees continually invade the prairie. But most perish early,
-their shallow roots no match for the extensive root systems of the
-fast-growing, moisture-greedy grasses. If encouraged by a series of wet
-summers, however, the young lodgepoles quickly gain stature. They had
-made significant inroads at Big Prairie when the disastrously dry summer
-of 1967 killed most of these 15-year-old pioneer trees.
-
-These North Fork grasslands and the immediately surrounding lodgepole
-pine forests are an important spring range. Deer, wapiti, and
-grizzly—and, in the wetter areas, moose—graze or browse here. And here,
-low on the western slopes of the Livingston Range, are the park’s only
-stands of ponderosa pine, a tree that prefers warm, dry habitats. As a
-result, at low elevations it often merges with the prairie community.
-
-Groves of aspen colonize the eastern prairies in areas where there is
-sufficient water and protection from wind. These aspen parklands are
-important havens for animals. Wherever two differing communities
-interact, a phenomenon known as “edge effect” occurs. Here wildlife
-exists in abundance; the animals that favor forest cover mingle freely
-with those that prefer open areas. Aspen groves—supporting grasses,
-herbs, and shrubs beneath their thin canopies—are favored haunts for
-grouse, varying hare, deer, and wapiti, all of which find among the
-trees abundant food, shelter and concealment. Populations of insects,
-small mammals, and birds, which are high for the same reasons, attract a
-wide range of predators.
-
-Isolated aspen groves are characteristically dome-shaped. Because aspens
-are capable of reproducing themselves vegetatively, the grove slowly
-expands outward from the parent tree. As a result, most of these groves
-are either exclusively male or exclusively female.
-
-Since quick-growing aspens provide a bountiful food source for beaver,
-streams near these trees are often dammed by the rodents flooding
-lowlands and creating additional habitat in the form of willow flats.
-Another “edge effect” is established, attracting animals found near
-water. Waterfowl, marsh birds, moose, mink, muskrat, skunks, amphibians,
-and many others find such areas to their liking.
-
-
-■
-Before the appearance of the white man, these eastern prairies were a
-paradise for animals. Once, on the summit of Rising Wolf, light-headed
-from the climb and the view of endless prairie, I fancied that I saw
-that vast, undisturbed animal panorama spread before me.
-
-Principally there were the bison, darkening the uneven land. Pronghorn
-bands flashed white on ridgetops, and moose moved through the long
-fingers of willow that extended eastward with the rivers. Caribou and
-wolves inhabited the shadows. Among vast cities of prairie dogs, swift
-fox and grizzly roamed. There were the clamorings of sandhill crane, and
-white clouds of trumpeter swans.
-
-This land, endowed with a wealth of wild grass, wore its wilderness
-well.
-
-
-The Forest
-
-On Gunsight Pass, the rain lancing down, I found a sharpedged rock that
-split the continent in two. On both sides the rain rivulets ran down, a
-fraction of an inch determining the stream’s destination: Pacific or
-Atlantic.
-
-The Continental Divide is a mighty barrier, a line of consequence that
-does more than determine watersheds. Its effect in Glacier is dramatic,
-as a look at the forests will reveal.
-
-Obstructing the eastward flow of the moisture-laden Pacific winds, the
-Divide extracts a heavy annual tribute of precipitation from the air
-mass, forcing it to rise up the mountain chain, where it cools and
-condenses. Chief benefactors are the low western valleys, which respond
-with a lush growth of Pacific coastal-type forests.
-
-The eastern valleys, however, deprived of abundant annual moisture and
-exposed to the wind and temperature ravages of the prairie’s continental
-climate, support a dramatically different kind of forest. Here Engelmann
-spruce and subalpine fir are the climax trees, contrasted with such
-trees as the western redcedar and western hemlock of the mild and moist
-McDonald valley.
-
-Elevation exerts an additional restriction on the distribution of tree
-species. Since climatic conditions vary with change in elevation—lower
-temperatures resulting in shorter growing seasons, and increased wind
-exposure resulting in greater loss of moisture through evaporation—we
-would expect to find the forest composition change as we ascend a
-mountain slope. In Glacier, eastern valleys average 240 meters higher
-than western, and thus even if they had more moisture they would not
-sustain the redcedars and hemlocks. All plants have range limits, some
-narrow, some broad; and they excel where their particular set of
-preferences as to moisture, soil, sunlight, and wind exposure are best
-met. On sites that do not meet their optimum requirements, they face
-being crowded out by species better adapted to the prevailing
-conditions.
-
-Physical features of the land determine vegetation also. Certain trees
-prefer the moist areas along a streambed—the great black cottonwoods,
-for example. And on steep hillsides, avalanches prevent the growth of
-climax trees, permitting instead only shrubby, pliant
-growth—mountain-ash, mountain maple, alder, menziesia.
-
-Forest communities are named for their dominant tree species. Thus, an
-area in which Douglas-fir dominates is called a “Douglas-fir forest.”
-Glacier does have forests in which Douglas-fir is the climax species;
-these are chiefly dry areas, below 1,800 meters, with south and west
-exposures. But we usually associate the park with its Engelmann
-spruce-subalpine fir forests, found extensively between 1,200 and 2,100
-meters, and with the western redcedar-western hemlock forests in the
-McDonald valley.
-
-Because forests mature slowly and change is usually imperceptible, we
-are tempted to think of them as static and eternal. But since a forest
-is a community of living things, it responds to changes in the
-environment. Subtle physical or climatic changes, such as a rising or
-falling water table or a slight increase or decrease in annual
-precipitation, will favor some species of trees and hinder others,
-eventually altering the composition of the forest.
-
-Other changes are more dramatic. Most notable of these is fire.
-
-
- From Fire to Forest
-
-Heat lightning, glimmering soundless behind the western peaks. Then the
-first low rumble. At first the flashing had been from cloud to cloud,
-but now, as the storm nears, the first ground-spears appear, lighting up
-the night. Here is a big storm, many-celled, engulfing more and more
-territory beneath its angry bulk. Lightning dances into the dry August
-forest. In their towers the lookouts stay awake.
-
-Close strike and a flare-up! The ridge snag burns like a Roman candle,
-sending bright embers down. Valley, ridge, and peak blink on and off
-with blue light as the storm roars like night-firing artillery.
-
-Passing overhead, the low cloud belly brings a sudden lash of rain. But
-it is not enough: tomorrow will mean long hours of fire watch.
-
-The next day dawns clear, a morning of heavy dew. The ridge strikes did
-not ignite the forest. Inspecting the storm path, aircraft and lookouts
-find no evidence of fire.
-
-But two days later, in a morning of high wind, thin smoke plumes rise
-upward. Smoldering in the thick duff of the forest floor, a lingering
-hot spot explodes with the fanning wind. It quickly spreads from a
-hectare to ten while the quadrants are called in and the hot-shot crews
-dispatched; then to a hundred, bringing in the smoke jumpers and
-mobilizing the vast fire-control network. A thousand hectares, perhaps
-ten thousand might burn this week of big fires.
-
-In the resulting skeleton forest, the scene of devastation is almost
-overpowering: life seems forevermore excluded from this blackened ruin.
-But fire is nothing new to forest communities. We may think fire demonic
-because it takes from our life span this block of mature forest, a sight
-we will never again see in this place. But nature does not operate in
-terms of human time scales. This forest is simply pushed back closer to
-its starting point, to begin again its long progression toward a climax
-vegetation cover.
-
-
- Forest Succession
-
-Through a series of complex vegetation stages, each characterized by
-different herbs, trees, and shrubs, the forest slowly returns to the
-type of vegetation best suited to the physical and climatic conditions
-of the site; this is called a climax community. The fact that most of
-Glacier’s forests are in some stage of recovery from fire accounts in
-part for the mosaic of forest cover found here.
-
-The forest of Huckleberry Mountain on the Camas Creek road was consumed
-in the 1967 fire. By 1969, among the charred, lifeless trunks of the
-former forest, lush grass and sunloving fireweed, thistle, and
-paintbrush were growing. And by 1974 lodgepole pine seedlings along the
-road were a meter or two high. Lodgepole is a fast-growing tree that
-requires full sun to germinate. Forest fire is necessary for the
-regeneration of these trees: the intense heat causes the tightly closed
-cones to open, releasing the seeds that will establish the forest. So
-young pines developed among fireweed, spiraea, willow, and mountain
-maple shrubs.
-
-The lodgepole forest near the western entrance to the park has been
-developing since 1929, when fire destroyed the redcedar-hemlock forest
-in the area between Apgar and West Glacier. Beneath the scattered spires
-of old larch that survived the burn, the lodgepoles have now grown up,
-forming a canopy that shades the forest floor. Because lodgepole live
-only about 80 years and will not germinate in shade, this forest will
-not exist long. Shade-tolerant Douglas-fir, white pine, Engelmann spruce
-and western redcedar seedlings are now taking hold. But the physical
-characteristics of this area—the climate, terrain, and soil—are
-ultimately most favorable for western redcedar and hemlock; and unless
-other disruptions intervene, this area will eventually again become a
-dense redcedar-hemlock forest.
-
-But this will not happen quickly. The soil after hundreds of years of
-collecting debris will again become rich and moist. Young hemlocks will
-germinate on and near decaying logs. When old larches, firs, and pines
-fall, the slow-growing redcedars and hemlocks will take their places in
-the canopy.
-
-Forest succession is a more complicated story than this; it is a
-fascinating study involving herbs, shrubs, small and large trees, and
-animal populations. From location to location it will vary; only in its
-broad outlines is it predictable. It is based on the observation that,
-given time, a forest—or any other plant community—will progress until it
-reaches climax—that is, the stage that will perpetuate itself.
-
-
-■
-How then are we to think about fire? Increasingly, experts are concerned
-not so much with fire suppression as with fire management. For
-suppression has at least three disadvantages: it allows the accumulation
-of unburned fuels that can result in “fire storms” when they are finally
-ignited; an undiversified climax forest is more vulnerable to disease
-than is a mixed forest; and a dense forest canopy discourages shrub
-growth, an important food source for deer, wapiti, moose, and smaller
-animals.
-
-As the well-being of the deer herd depends on the predators that thin
-its numbers, so the long-term well-being of the forest depends on fire
-to rejuvenate it periodically. We must realize that wilderness is
-identified with fire, landslide, avalanche, windfall, and flood. Nature
-not only has learned to cope with these agents of change—she depends
-upon them for maintaining the delicate balances between landscape and
-life. There is in the business of nature, after all, more than the
-pleasing of man’s eye.
-
-
- Spruce Morning
-
-Of all times to get a rock in my boot! I had just started out, the
-morning was still cool in this eastern valley, and the heavy pack was
-not yet biting into my shoulders. Sitting down beside the trail, I
-leaned the pack against the base of an old spruce and began unlacing.
-
-I could hear the scratching of the red squirrel descending to
-investigate, but I didn’t look up until it let go with long indignant
-chatter at finding its territory invaded. I plunked out the pebble and
-began relacing my boot. Cautiously the squirrel came down, pausing
-frequently to scold, its lower jaw quivering with rage and exposing
-yellow rodent teeth. Neighboring squirrels joined in and soon the trees
-danced with flicking tails.
-
-Down the squirrel came, almost to the ground, then raced back up the
-tree, stopping at each lateral branch to deliver a vocal broadside.
-Finding no danger to themselves, the other squirrels soon quit the
-uproar and went about their morning business. I was beginning to suspect
-that I was committing some graver offense than the mere exercise of
-squatters’ rights—perhaps I threatened its cache of fir cones. Then into
-the corner of my vision shot another form, streaking soundless as a
-shadow; the squirrel also saw it—but too late. With a thin terrified
-squeak, the rodent started to go higher; but the pine marten was above
-it. The squirrel quickly reversed itself, sending bits of bark showering
-down.
-
-As the squirrel leaped from the tree in desperation, the marten overtook
-it in mid-air; they came down together. Clamping the limp creature
-firmly in its jaws, the marten strode up the incline of a fallen spruce.
-Before it hopped off onto a shelf of higher ground to disappear, it
-looked briefly back at me. I fancied I could read, fixed in its eyes, a
-certain recognition of my having distracted its prey.
-
-A breeze made me shiver, snapping me back from that swift vision of
-luxuriant fur, that blinding grace which flashed its orange throat-patch
-through the trees, and I realized I was sweating. For a moment I had
-been that squirrel, eyes wide with terror, seeing fate bear down, and
-powerless before the natural order of things.
-
-The incident got the other squirrels singing again; but the confidence
-was gone, and soon it was quiet. What dreams do squirrels dream, I
-wondered, looking around. I saw that place more clearly then, having
-been caught between a marten and its prey. I saw each spruce: its age,
-its condition, the onslaughts it had borne; the beargrass coming up in
-an opening; and down the trail a meadow that was yellow, white, and red
-with sulphur plant, mariposa, and Indian paintbrush. Bees, flies,
-spiders, and butterflies worked that little garden tucked among the
-crowding trees. Countless forms of life beneath the soil and bark, in
-tunnel, crevice, hole, and pocket, working unseen to sustain their
-lives, and somehow, when all were added up, maintaining the forest as
-well.
-
-A flicker called, its loud _Klee-yer_ breaking the forest hush. Birds,
-mammals, plants, insects—all hide together here, their lives so
-skillfully embroidered that no loose thread exists that my mind might
-grasp to unravel and understand the work.
-
-The forest had once been a place that obstructed my view, a great blank
-to stride through, a few hours of necessary blur before the high lake or
-pass was reached. Now I was quite content to remain awhile beneath these
-great-boled trees.
-
-
-■
-A forest, like the mountains themselves, supports various levels of
-life. The floor and substratum are a great processing plant where
-bacteria, fungi, and insects work, decomposing the plant and animal
-litter, recycling dead and discarded tissue back to simpler organic
-compounds, gases, and minerals, thereby providing sustenance for growing
-plants. As spiders, shrews, wrens, and thrushes seem to know, there is
-good hunting on the forest floor.
-
-Just above the forest floor is the herb layer, a seasonal layer of
-growth including flowers, mushrooms, grasses, and other small plants.
-
-Above that grows the shrub layer, then the understory of young trees
-awaiting their chance to take a place in the forest’s canopy high above.
-From the swaying canopy, exposed to the full force of sun and wind, to
-the dim, moist floor, the forest provides a wide range of habitat.
-
-Relatively few animals live in the treetops. The almost incessant motion
-makes nesting too hazardous for birds. Red squirrels venture up to cut
-cones in the canopy, but store their booty and make their nests farther
-down.
-
-In the mid-range between canopy and understory, goshawks and Cooper’s
-hawks nest. Woodpeckers, nuthatches, and sapsuckers forage on the tree
-trunks and nest in cavities they excavate or appropriate. Red squirrels
-and the nocturnal flying squirrels create a major traffic here, along
-with the martens and owls that hunt them.
-
-The understory and shrub layers house the greatest numbers of nesting
-birds. Here the effects of storm and rain are minimized and protective
-cover is greatest. Vireos, thrushes, warblers, hummingbirds, bluebirds,
-flycatchers, and others can be found among the tangle of this sometimes
-impenetrable layer.
-
-The most populated area, the forest floor, supports an astonishing
-abundance of organisms. Below the busy traffic of mice, shrews, and
-larger animals is a bewildering array of insects and other
-invertebrates. The attrition rate in the litter of the forest floor—a
-continual battleground difficult to comprehend—is enormous. The smaller
-the organism, the greater its numbers are likely to be. This humus-rich,
-moist soil teems with bacteria, and a handful will contain surprising
-numbers of small spiders, pseudo-scorpions and almost microscopic mites.
-
-Each year some two to three thousand kilograms, dry weight, of falling
-material litter an average hectare of forest. All this plant and animal
-waste—twigs, leaves, limbs, fallen trees, feathers, hair, feces, and
-carcasses—is processed by the armies of decomposers that thrive on the
-forest floor. With the aid of larger creatures that break up the plant
-and animal tissue, most microscopic bacteria are able to decompose from
-a hundred to a thousand times their own weight every day.
-
-Few trees die of old age in the forest. The seedling mortality rate is
-necessarily high, since far greater numbers of seeds germinate each year
-than can reach maturity. Of those that do, many fall victim to the
-ever-present dangers of disease, insect infestation, windfall, stream
-erosion, and fire. Insects alone present a formidable threat to trees,
-for they have evolved every means of attack—chewing and mining leaves,
-boring into twigs, eating cambium and heartwood, sucking sap, triggering
-galls. If the insect world did not police itself, aided by spiders,
-insectivorous birds and other animals, forests and other plantlife would
-quickly fade before the chewing, boring, sucking horde.
-
-
-■
-Through the trees the light on Citadel shows the morning slipping by. As
-I start to get up I see a garter snake sliding out into the dusty trail,
-seeking the sun-warmed earth. Moving slowly, alert for danger, it probes
-the air frequently with its sensitive tongue. But against the
-lightcolored duff its dark shape offers a fine target, begging attack. A
-chipmunk, watching from a nearby lookout stump, twitches its tail
-nervously over its back, curious—perhaps suspicious—at the sight of a
-snake. Very slightly the snake’s head goes up, its tongue flickering.
-For a few seconds reptile and rodent regard each other. Then the
-chipmunk drops back soundlessly into its hollow stump, and the snake
-lowers its head onto warm ground.
-
-Some day soon, a sparrowhawk or weasel will interrupt the snake’s
-morning sun-bath. The snake will fuel bird or mammal for a time, as
-mice, fledgling birds, and insects now sustain the snake. The chipmunk
-too, rummaging nearby, lives in shadows of talon and tooth.
-
-Until that time of sharp encounter, each has its own niche, a way of
-life, a shaft of sun, and food enough.
-
-
- A Walk in the Redcedar Forest
-
-Climax! The word takes on a true significance here, among these
-broad-based trees. When you enter this forest the road noise does not
-follow far—as, when you walk into a cave and turn a corner, sound and
-light are left behind. There is a surprising spaciousness, a feeling of
-openness in a mature western redcedar forest. With scant understory and
-the canopy so far above and everywhere complete, it seems like some
-vast, high-ceilinged catacomb, pillared by the huge, shaggy-barked
-cedars and the deeply scored trunks of the black cottonwoods. The floor
-is strewn with fallen giants in magnificent disarray, uplifted roots
-still grasping fractured rock.
-
-A rainy day is a good time to walk a cedar trail, when the dull light
-seems to shine from the wet moss, making the underleaves of devil’s-club
-and Rocky Mountain maple glow. Wind and rain, like light, penetrate with
-difficulty the latticework of this canopy; thin lines of fog develop
-over the bogs. The air is fresh with growing plants, snow-cold still
-when the first spring flowers appear.
-
-Fiddleheads of unfolding lady ferns line the trail in May, pushing up
-from the hub of last year’s leveled, lifeless fronds. Beds of trillium
-shine their white, three-pointed flowers like flashlights in the dark
-recesses. Unlike the small, hidden calypso orchid, which bears its
-purple spikes and yellow throat low above the moss, the trilliums make
-no secret of spring growth. They are bold, handsome plants, broad-leaved
-and tall, with waxy white petals that tinge to purple in their
-month-long bloom.
-
-Moss covers everything. Boulders are green and weightless-looking,
-resilient and topped with miniature forests of cedar seedlings. Ancient
-fallen trees are disguised with blankets of moss, sprouting hemlock here
-and there. The rich greens that characterize Glacier’s summers seem to
-begin here amid the moisture-glossed leaves of twinflower, bunchberry
-and bead-lily.
-
-Later, the spiders will spin thousands of kilometers of gossamer
-filament among the trees. The orb-weavers will hang their webs high and
-low, suspended in every opening. Walking through the forest then, you
-will see shafts of sunlight whirling in the higher webs until they seem
-like tops set spinning among the treetrunks.
-
-Indianpipes, the “ghost flowers” that need no light to grow, will break
-through the forest soil. Like mushrooms, with fruiting bodies that are
-nourished by underground mycelia, these saprophytes absorb their
-nutrients from a fungus that covers their roots.
-
-Receiving an average of about 18 centimeters more annual precipitation
-than forests east of the Divide, Glacier’s redcedar-hemlock community
-hoards its moisture. Its dense growth and the surrounding mountain walls
-inhibit the circulation of drying winds. Mosses and ferns transpire
-their moisture, which you can feel; place your hand close, and you will
-sense a coolness like the air exuding from an ice cave. Draped from the
-tree limbs are long filaments of squawhair and goatsbeard, black and
-grey lichen strands that flourish in the damp air.
-
-Except for the black bear, few large animals inhabit the deep forest.
-Grizzlies find better forage in meadows or along the forest edge. Since
-shade discourages shrubby undergrowth, deer and wapiti will search
-elsewhere for browse. In summer, wapiti, grizzlies, and mule deer bucks
-tend to wander up into high meadows.
-
-Contrasted to the noisy, conspicuous birds of the prairie—meadowlarks
-and bobolinks—birds of the forest seem elusive and secretive. Although
-numerous, the varied thrushes, Townsend’s solitaires, and Swainson’s
-thrushes are seldom seen; but when approached, they fly silently off and
-are swallowed by the forest shadow.
-
-There seems to be serenity in a mature forest, as though the struggle
-for life is somehow suspended, the needs of the animals here less
-urgent, muffled. The towering redcedar forest seems to be no battlefield
-at all, but rather a monument to what Earth can do.
-
-
- The Perpendicular Night
-
-Behind Avalanche campground a trail leads back toward Lake McDonald
-Lodge. I decided to follow it one June evening, to experience the
-sensation of the deep forest changing into night. With the nearby
-mountain wall intercepting the sun, dusk comes early to this valley. On
-the prairie, night passes across the landscape in an even line,
-forthright as a waxing tide; you can almost feel the globe in its
-turning from the sun. There is reassurance in the night’s coming, its
-steady purple doming over the sky.
-
-But here darkness seems to sprout from the earth. It collects beneath
-the hemlock clumps, bridges the creekbottoms. It seems to flit from
-place to place. You look about, uneasy, trying to catch it here or
-there, but always miss its infiltrations. It captures the narrow
-clearings when you look away; pockets of tree-darkness join together,
-forcing the light upward until the tree-tops seem impossibly bright and
-distant.
-
-Through the trees I could see a dozen fires dance in the growing shadow,
-wood-smoke and camp sounds filling the air. Turning uptrail, I felt a
-reluctance to leave the presence of those fires—a senseless feeling, but
-strong. A growing forest-dread impelled me almost physically backward to
-those circles of firelight. I felt the need to be near a fire, to be
-reassured by heat and light. Fire was our greatest friend, our greatest
-weapon. With it we beat the long ages of ice and held the forest gloom
-away. There was no harm here, only silence; yet the longer I walked,
-with beard-moss hanging down like daggers all around, the more I craved
-the comradeship of fire.
-
- _Continued on p. 104_
-
- The Vital Predator
-
- The merciless law of predation might at first thought seem cruel;
- but the predator plays a vital part in maintaining the balance of
- the biotic community. Without the controlling factor of predation,
- prey species quickly enlarge their populations. If plant eaters are
- not checked, the resulting excess population exceeds the carrying
- capacity of the range. Food supply rapidly diminishes. In a damaged
- range, competition and stress result, usually culminating in a
- massive die-off through starvation and disease.
-
- Ironically, predators thus provide a service to their prey. First to
- fall to the predator are the old, the diseased, the unwary, and the
- young. By removing many young and old deer from a typical herd,
- cougars lessen competition among the deer for choice range, thus
- tending to keep herbivore numbers at parity with the land’s carrying
- capacity. Only the strongest and wariest deer survive, ensuring that
- the fittest will continue the species. When man upsets this delicate
- balance—destroying predators in the hope of increasing numbers of
- game animals—the result is ecological disaster. In the 1930s, in a
- misguided attempt to “preserve” the whitetail deer herds of the
- park’s North Fork area, many coyotes and cougars were exterminated.
- In 1935 alone, 50 cougars were killed. Relieved of the pressure of
- predation, the deer flourished. In a few years, however, the
- normally adequate range was severely overbrowsed. Suffering also
- from this imbalance were wapiti (“elk”) and moose, ungulates that
- share the winter range with deer.
-
- Some predators are more specialized than others. The Canada lynx,
- for example, has oversize feet, an adaptation that helps it move
- across deep snow without breaking the surface. As a result, it is an
- efficient predator of the snowshoe hare, another large-footed
- animal. Relying on this adaptation, the lynx feeds almost
- exclusively on snowshoe hares. Consequently, its numbers inevitably
- fluctuate with the 10-year “boom and bust” cycle of the snowshoe.
-
- The coyote, on the other hand, is a generalized predator, exploiting
- whatever prey is currently abundant. Should mice or ground squirrels
- be in short supply, it will subsist on anything from grasshoppers to
- berries until favored prey again becomes available. (Animals that
- normally eat both plant and animal food are referred to as
- omnivores.) Generalized predators are thus better equipped to
- survive temporary ecological imbalances, maintaining their numbers
- at relatively consistent levels from year to year.
-
- Carnivores all, the animals on these pages illustrate various
- adaptations for capturing prey.
-
- [Illustration: The population of the Canada lynx, which is widely
- distributed in Glacier’s coniferous forests, fluctuates in cycles.
- The lynx is abundant or scarce depending on the population condition
- of its chief prey, the equally cyclic snowshoe hare.]
-
- [Illustration: The cougar, which feeds primarily on deer, requires a
- large territory. Because of its strength, stealth, and speed,
- American folklore has given this wary cat a false reputation as a
- man-stalker.]
-
- [Illustration: The red fox depends largely on a well-developed sense
- of smell to locate its prey; it also relies on its keen eyesight,
- speed, and agility to capture mice, hares, birds, and whatever else
- it can run down or surprise.]
-
- [Illustration: To feed its demanding young, the Swainson’s thrush
- hunts for insects along the forest floor and in the dense
- underbrush. This thrush relies on its secretive behavior to protect
- its nest near the ground from detection by other predators.]
-
- [Illustration: Armed with enlarged forelegs, the crab spider waits
- on or near flowers to ambush visiting bees, flies, or other insects.
- Its venom produces a quick kill, allowing it to attack insects many
- times its own size.]
-
- [Illustration: The spotted frog is a large-mouthed predator that not
- only eats water striders and other insects but also gulps down
- smaller frogs and small fish.]
-
- Protective Coloration
-
- To escape extermination, each species must in some manner foil its
- enemies. Protective coloration is one of the more common adaptations
- helping to do this. Most animals resemble their environment to some
- extent. The conspicuous markings of some, like the bitter-tasting
- monarch butterfly or the striped skunk, seem to function as a
- warning to prospective predators that it is in their best interest
- to look elsewhere for a meal.
-
- Some animals, such as the white-tailed ptarmigan and the snowshoe
- hare, have seasonal changes in plumage or pelage, wearing white in
- winter and brown in summer. Even predators, such as longtail and
- shorttail weasels, benefit from seasonal camouflage. Protective
- coloration makes them less noticeable to prey species and to larger
- predators.
-
- Many insects, too, change coloration with the season. Bright green
- grasshoppers of early summer become more brown with each molt,
- matching the changes in the surrounding vegetation.
-
- _Obliterative shading_ is especially important to animals that
- frequent more than one habitat. Seen from above, turtles match their
- dark background; from below, because of their lighter underbody
- shading they blend into the bright skylight.
-
- _Disruptive coloration_ aids in breaking up an animal’s outline.
- Butterflies and moths commonly have disruptive wing markings. The
- distinctive shapes of eyes can be concealed. Eye coloration may
- mimic body color—as in the green katydid—or the eye may continue
- disruptive body markings.
-
- Ground-nesting birds are especially vulnerable to attack. Their eggs
- tend to be heavily blotched with earthy colors, making them less
- conspicuous. Chicks also carry these disruptive colorations on natal
- down.
-
- Most mammals, with coats of brown or gray, are inconspicuous when
- motionless. Deer fawns are endowed with speckled coats, mimicking
- the sun-flecked forest floor; this disruptive coloration, coupled
- with absence of scent and their instinctive “freezing” behavior,
- makes it difficult for predators to detect them.
-
- The whitetail deer not only uses its white “flag” to warn others in
- the herd of danger; it also allows a pursuing predator to use it as
- a target. When the tail is suddenly dropped—abruptly obliterating
- the bright white patch—the deer seems to disappear into its dim
- surroundings.
-
- Since overly conspicuous animals are prone to predation, natural
- selection favors development of appropriate camouflage.
-
- [Illustration: For such ground-dwelling birds as the white-tailed
- ptarmigan, camouflage is an important survival adaptation. The
- ptarmigan changes its plumage to match its surroundings: it is white
- in winter, speckled in summer. Moving slowly and refraining from
- flight, it is less likely than more-active birds to be detected by
- sharp-eyed, motion-conscious predators.]
-
- [Illustration: Birds that when hatched are covered with down and are
- able to move about freely are called _precocial_. They are less
- dependent upon their parents than are _altricial_ young, which are
- naked and helpless when they hatch; but they must rely heavily on a
- resemblance to their surroundings for survival during their first
- flightless weeks. This spruce grouse chick, which blends into its
- sunflecked forest-floor habitat, is an example of a precocial bird.]
-
- [Illustration: The bold disruptive pattern of the killdeer chick’s
- plumage helps this precocial bird avoid detection in its
- open-prairie environment. This adaptation, coupled with the chick’s
- instinct to freeze at the approach of danger, ensures that enough
- young will survive to perpetuate the species.]
-
- _Ursus arctos horribilus_: The Vulnerable King
-
- At the apex of the food pyramid, this great beast is unquestionably
- the king of Glacier’s biotic community. Yet the long-range future of
- the grizzly bear is uncertain. With the grizzly exterminated from
- most of its former range—which once extended into the midcontinent
- and south into Mexico—its numbers have dwindled in proportion to its
- diminished range. Present concentrations in the contiguous United
- States remain in and around Yellowstone and Glacier National Parks.
- Probably fewer than 200 of these magnificent creatures live in
- Glacier National Park.
-
- Grizzlies are easily distinguished from the more common black bear.
- In addition to larger size and heavier build, grizzlies have a
- characteristic shoulder hump; long, conspicuous claws; and a broad,
- concave face that gives them a “dished-in” appearance. Fur is
- usually brown; like the fur of the black bear, however, color may
- range from black to yellowish. Light tipped hairs make the fur
- appear frosted, giving rise to the nickname, “silvertip.”
-
- Grizzlies, popularly considered arch predators, are more accurately
- described as omnivores. Carrion, grasses, cow parsnip, and several
- species of berries, bulbs, and tubers make up a grizzly’s diet,
- along with insects, small mammals, and an occasional ungulate that
- it can catch. As a result, grizzlies play several roles in the
- biotic community, functioning as herbivore, scavenger, and predator.
-
- Ranging widely in all life zones, grizzlies follow the spring
- snowmelt up to the alpine meadows, returning to lower elevations to
- hibernate from November until April. One to three cubs are born in
- midwinter during hibernation. Since the maternal bond lasts two
- years, a sow will accept a mate only every other year. Mortality of
- subadults is high, resulting principally from competition among the
- bears themselves. As with most animals, range—habitat—appears to be
- the limiting factor of grizzly populations.
-
- The grizzly is normally shy and fearful of man—but highly
- unpredictable. Wounded or sick bears, sows defending cubs, young
- adults, and bears that have become conditioned to human scent are
- the most dangerous. As humans continue to encroach on grizzly
- territory, odds of confrontation also increase. Recent fatalities
- and personal injuries inflicted by grizzlies pose a vexing problem
- to the National Park Service, which is charged with visitor safety
- on the one hand and protection of the park’s remaining grizzly
- population on the other. Continuing study of grizzly ecology and
- increasingly enlightened bear management programs will, it is hoped,
- allow man and bear to co-exist in a wilderness both require.
-
- [Illustration: Grizzlies are fond of succulent spring grasses.]
-
- [Illustration: Traversing all life zones in the park, the grizzly is
- a true opportunist, eating anything from ants and berries to
- wapiti.]
-
- [Illustration: Seldom will a grizzly exceed 225 kilograms in
- Glacier. This is a young adult.]
-
- Bald Eagles and Kokanee Salmon: A Recent Gathering
-
- In 1916 the kokanee salmon, a small, land-locked form of the Pacific
- coast species, was planted in the Flathead drainage. With the first
- planting augmented by additional stockings, the fish thrived in
- cold, deep Flathead Lake, and, to a lesser extent, in Lake McDonald.
- The salmon fed almost exclusively on zooplankton.
-
- By the mid-1930s, salmon runs were becoming established. The outlet
- of Lake McDonald provides an ideal spawning site for the salmon. The
- fast-flowing water is clear, cold, and shallow, and the creek bed is
- gravelly.
-
- Averaging 0.3 meters in length and weighing less than a half-kilo,
- the 4-year-old adult salmon cease feeding and begin to migrate. Many
- thousands swim the 100 kilometers from Flathead Lake to McDonald
- Creek. Males appear in the creek first, arriving in late September,
- and are soon followed by the females.
-
- Using her tail to dig a redd (a shallow nest depression), the female
- deposits about 650 eggs. After fertilization by the male, the eggs
- are covered over. The adults die within three weeks after spawning,
- their bodies exhausted from the rigorous migration journey and the
- weeks-long lack of sustenance.
-
- Egg fatalities are high, due to stream erosion and disturbance by
- other spawning salmon. Hatching in late March, the fry work their
- way out of the gravel and migrate downstream.
-
- Attracted to the 75,000-150,000 salmon concentrated in a 3-kilometer
- stretch of shallow water, bald eagles begin gathering at McDonald
- Creek in October. It is not known where the eagles come from or
- where they go after the spawning run. Glacier has fewer than 20
- summer-resident bald eagles, and these are distributed among the
- remote lakes of the North Fork area.
-
- In 1939, 37 bald eagles were counted along the creek. By 1969, 373
- were reported, representing approximately 10 percent of that year’s
- estimated winter population for the contiguous United States. Since
- 1960, the count has averaged 240 birds. (In 1977 there were 444.)
-
- Eagles feed by swooping down to pluck salmon from the water or by
- wading out to grab a fish stranded on a shallow riffle. An eagle may
- consume as many as six fish a day. Immature birds are not as adept
- at catching fish and may harry adults or other immatures into
- releasing their catch.
-
- [Illustration: From its vantage point, this mature bald eagle
- examines the waters of McDonald Creek. Average weight is 5.7
- kilograms; average wingspan is 2.2 meters. Females are slightly
- larger than males.]
-
- [Illustration: This immature bald eagle lacks the familiar white
- head and tail of the adult birds. It will not acquire those markings
- until it is several years old.]
-
- [Illustration: Breeding male and female kokanee salmon are easily
- distinguishable; as spawning time approaches, they change
- appearance. The dark gray backs turn red; heads become green, and
- the males develop humped backs and hooked jaws.]
-
- [Illustration: Swooping upward with a fish, a mature eagle heads for
- a convenient perch to consume its catch. A strategically located
- tree may contain 30 birds.]
-
- A Triumph of Many Colors
-
- Grassland, meadow, tundra, or any other area in Glacier suitable for
- plant growth and supplied with abundant sunlight produces an
- extravagance of wildflowers. This display of various shapes and
- colors is neither an accident nor a mere decoration of nature. Nor
- would Earth’s recent explosion of mammal and bird species have been
- possible without the evolution of flowering plants.
-
- Two hundred million years ago, early in the Age of Reptiles,
- angiosperms (flowering plants) had not yet evolved. Plant
- reproduction still relied on spores and cones. Then, during the
- Cretaceous Period, the last sediments were being laid down in the
- inland sea that covered most of Montana. (It was these sediments
- that the ancient Precambrian rocks of Glacier’s mountains later
- overrode, forming the Lewis Overthrust.) During this period the
- evolutionary miracle occurred: flowering plants—grasses, vines,
- shrubs, broadleaf trees, wildflowers—inherited the Earth.
-
- The timing was important. As Earth’s tropical climate gradually
- changed to temperate extremes during this period, the domination of
- cold-blooded dinosaurs ended and the moisture-demanding coniferous
- forests that had covered the earth in green monotony began to
- shrink. Angiosperms provided a solution to the ecological void:
- grasses and forbs grew where trees no longer could. Most important,
- relationships evolved between this new class of plants and the
- relatively few species of insects then existing.
-
- Insects began to use the pollen of flowering plants; the
- angiosperms, in turn, evolved bright petals and nectar that
- exploited visiting insects for the plants’ own reproductive
- purposes. This partnership allowed insects to diversify rapidly,
- evolving new, specialized forms such as bees, moths, and
- butterflies. As a result, predatory forms of insects and arachnids
- also rapidly diversified.
-
- The most dramatic change, however, involved warm-blooded birds and
- mammals, whose high rates of metabolism required high-energy fuels.
- Unlike gymnosperm seeds, which contain no protective covering,
- angiosperm seeds are surrounded by a fruit. The development of these
- highly nutritious seeds, and the attendant explosion of insect
- species, ensured survival of the newly evolved birds.
-
- As birds diversified into seed-eaters, insectivores, and carnivores,
- mammals, then uncertain little ratlike creatures darting among the
- feet of dinosaurs, began a rapid rise to dominance; grasslands
- promoted an explosion of herbivorous and carnivorous species.
-
- The evolution of angiosperms, and the animal revolution it made
- possible, came with amazing speed. Most significant, it was a vital
- first step upon which the meteoric rise of man depended.
-
- [Illustration: Indian paintbrush is common at all elevations below
- tundra. It may be white, yellow, orange, pink or red. The actual
- flowers, inconspicuous and green, are surrounded by brilliantly
- colored bracts. Semi-parasitic on other plants, paintbrush is
- normally found growing in conjunction with other wildflowers; its
- roots steal sustenance from neighboring plants.]
-
- [Illustration: Yellow stonecrop, widely distributed in forest and
- scrub-forest zones, is one of the park’s few plants having succulent
- leaves, an adaptation that helps it survive in such situations as
- dry, rocky outcrops.]
-
- [Illustration: The Calypso orchid grows in the cool, shadowed forest
- where light is dim. It lives in partnership with certain fungi that
- exist about the orchid’s roots and seem to help nourish it.]
-
- [Illustration: Silky lupine, a legume, has nitrogen-fixing nodules
- on its roots, thus allowing it to grow in nitrogen-poor soil. It is
- widely distributed in grassland and forest communities.]
-
- Fire Succession: Key to Continuity
-
- Most of Glacier’s fires are lightning-caused. Strikes may flare up
- immediately; or fires may smolder in the forest duff for days until
- fanned into flame by wind. _Ground fires_ may race through the
- forest understory, causing minor damage; or they may bridge the
- understory and reach the canopy, thus becoming rapidly spreading
- _crown fires_. Under certain conditions, uncontrollable infernos may
- develop, generating terrific winds and heat. These rare
- conflagrations are called _fire storms_.
-
- Every type of forest habitat has _climax vegetation_—trees and
- shrubs that are best suited to the site and thus maintain themselves
- indefinitely if not disrupted.
-
- After a major fire, habitat conditions are usually so altered that
- the site must pass through several _seral stages_ before conditions
- are such that climax vegetation can return. A _sere_ is a series of
- plant communities that follow one another in orderly fashion until
- climax conditions are again reached.
-
- [Illustration: Lightning fires occur most often during the hot, dry
- weeks of late summer.]
-
- [Illustration: When the forest is dry, lightning often causes quick
- flare-ups.]
-
- [Illustration: The forest may continue to burn for days after the
- main conflagration has passed.]
-
- [Illustration: After a major fire, sun-loving grasses, shrubs, and
- wildflowers quickly invade the former forest. Deer and wapiti
- benefit from these new food sources.]
-
- [Illustration: Lodgepole pine, a pioneer species quick to take over
- burned areas at lower elevations, grows rapidly. These trees are
- five years old.]
-
- [Illustration: This is a Glacier National Park forest 80 years after
- a major fire.]
-
-Sudden hammering made me jump. Above the forest darkness, a pileated
-woodpecker leaned out from a high larch snag, braced against the trunk
-by its specialized, stiff tail feathers. This was the first time I had
-seen this big white-and-black bird, the “cock-of-the-woods.” There was
-ample evidence of his work: the deep, oblong excavations in the trunk
-and the pile of large wood chips at its base, both characteristic of
-this species. Again he hammered, and I could see the chips falling.
-After a little edge-work around the hole, he extracted a grub and flew
-off, yammering against the advancing dark.
-
-Near a stream I stopped to sit down, to listen to the water and maybe
-catch sight of some small animal. Across the narrow defile, from a slope
-dense with young hemlock, came the buzzing note of a varied thrush.
-Several notes followed, all on a different pitch, all drawn out, level
-and clear; the quality was pure but songless, disjointed, deliberate,
-like someone testing the reed of a strange woodwind. There seemed no
-gladness in the heart of this thrush. The song was dark, haunting,
-lonely.
-
-On the trail ahead I could make out a bird hopping rapidly along. After
-passing the spot I could hear its song. There couldn’t be a hundred
-meters between us, yet it seemed to be coming from a great distance. I
-listened for as long as it would sing. I tried to hear it for what it
-was, a male Swainson’s thrush proclaiming its territory. But the
-ethereal, flute-like phrases seemed an evensong made not for man’s ears
-but only for the forest itself.
-
-I hurried on after the bird had ceased. It was getting dark beneath the
-trees, but I was beginning to be aware of creatures underfoot, the mad
-dartings of shrew and vole, more imagined than seen. When a deer mouse
-jumped away I got out my flashlight. Soon the beam caught a woodrat
-sitting atop a fallen log. The light didn’t bother him in the least; as
-I approached, he picked up his bushy tail in his forepaws. Whiskers
-twitching, he looked more caricature than real. Then he bounded off the
-log with graceful, arching hops, and disappeared into the night.
-
-Against a patch of sky that appeared in a clearing, I could make out
-bats, circling and dipping like swallows. Locating a hovering moth, I
-kept the light beam on it until it vanished into a furry streak of
-silence. It was time to head back.
-
-By now it had become utterly dark within the trees, a moonless,
-sightless, alien world, given over to the marble-black eyes of the small
-night mammals and the creatures that hunt them. I thought of the
-strange, unseen societies of the flying squirrels, the nocturnal
-counterparts of red squirrels; of the great-horned owls, inspecting the
-same ground the goshawks scanned during the day. Perhaps a foraging red
-fox moved through the darkness nearby, or a coyote on night patrol.
-
-The flashlight beam probed ahead along the trail. The exposed roots were
-given unnatural shading and they seemed to thicken and squirm as I
-approached. On either side the tree trunks appeared to step backward
-from the dim glow of the light. I felt lost in this night, thinking of
-the great darkness in all the timbered ridges that ran westward from the
-Divide. In this vast cathedral of crowded tree and peak, night was stood
-on end, the stars shrunken to a circle overhead, as if seen from the
-bottom of a well. Mouselike, shivering, insignificant in this
-wilderness, I scurried back to find a fire and fill my empty senses with
-its heat and snap and light, holding off the fright of night and
-thinking of tomorrow’s sun.
-
-
-Scrub-Forest
-
-The crowning beauty of Glacier—the high, cirqueheld meadows that scent
-the wind with wildflower and waterfall—belongs to the zone of
-scrub-forest.
-
-At Logan Pass you are introduced to the highlands. Here an exquisite
-upland basin holds the Hanging Gardens, a wildflower-clothed gradient
-laced with stair-step bogs and lines of wind-bent subalpine fir. In the
-dawn sun, before the first engine noise, it shines unbroken, dewbright
-and sagging like a spider web secured to the circle of surrounding
-peaks.
-
-This is the region the hiker remembers best. The tall mountains wear
-this zone close to the cliffs, and the trails encounter it near the
-passes or follow it for long, level stretches, as along the Garden Wall.
-I remember Preston Park and Fifty Mountain, the fire-touched bench of
-Granite Park and the first sight of Sperry chalet, built on a brow of
-rock at the upper reach of trees. But most of all I remember the
-terrible waterfall that becomes Bowman Creek, the plunge of nearly a
-kilometer that drains the magnificent upland bench called
-Hole-in-the-Wall.
-
-
- Hole-in-the-Wall
-
-September. The season is growing late, the meadow-rue dying and the
-leaves of the wild strawberry failing at last. Everywhere the red
-contagion of autumn surrounds the vital green. The lower valleys have
-lost the whistle of ground squirrels. They sun themselves no longer
-these late, mild days. Ripe, sluggish, and hawk-vulnerable, they sensed
-the need of hibernation.
-
-It has been eight years since I last visited Hole-in-the-Wall, but I
-retain its dimensions and hear its dozen waterfalls at will. Once you
-have seen this basin you have a measure by which to judge the high
-country and a thirst for the meadows at tree-line.
-
-In Glacier, treelimit ranges between 1,850 and 2,300 meters, depending
-on local conditions. The upper limit of tree growth—rarely an even,
-horizontal line—is generally an indistinct band running erratically
-across a mountain’s face: a tension zone reflecting variations in wind
-and sun exposure, degree of slope, snowpack accumulations, and the
-presence of adequate soil and water.
-
-Subalpine fir, whitebark pine, and Engelmann spruce do not relinquish
-easily their upward climb; where conditions become severe, their growth
-is retarded and their stature dwarfed. Deformed and pruned by wind,
-their leaders winter-killed when they outreach the protection of the
-winter snowpack, trees become shrubs, forced to hug the ground. Size
-belies age in these elfin forests, or krummholz, where the growing
-season is painfully brief and progress is always uncertain. A twisted,
-gnarled little bush, more snag than live branch, bearing a single cone
-or two, may be senior by a century to the giants of its race in the
-valley below, which yearly shower the ground with an abundant crop of
-cones.
-
-This time I will come from Goathaunt, passing Lakes Janet and Francis,
-reaching Brown Pass from the east, and camp in the spectacular garden
-between Brown and Boulder Passes.
-
-Meadows and rock slides break the forest as the trail gains elevation
-and distance through the valley. The spruce and fir thin out rapidly at
-the valley head, the trail climbing the grassy slope to low, broad Brown
-Pass. Below the pass is Thunderbird Pond, which receives the meltwater
-from a glacier high on a shelf of Thunderbird Mountain and is bordered
-by a low jungle of willow. In the water stands a bull moose, its heavy,
-fully formed antlers ready for the season’s impending business.
-
-I was hoping again to see Cassin’s finches and Audubon’s warblers on the
-pass; but the fir grove is quiet. Sitting down to rest and listen, I
-become aware of a strange silence. No birds sing or flit among the
-trees, no alarms pass back and forth among alert ground squirrels. There
-is no wind—an odd condition for the Continental Divide. This place seems
-to be holding its breath. High overhead, a veil of cirrus cloud arranges
-long spears across the sky.
-
-Moving off the pass, along the dome of Mt. Chapman, I experience anew
-the old excitement of this high country. Abruptly the gorge of Bowman
-valley opens up, revealing the twisting blue snake of Bowman Lake far
-down the narrow, cliff-imprisoned valley. Here again are the northern
-titans—Numa, Peabody, Boulder, Thunderbird, and Rainbow; and Carter,
-with its high glacier baring blue ice teeth to the sun.
-
-It is not the climb that makes your heart pound now; the trail is
-suddenly narrow and cliff-defiant, cut by the plunging waters of
-snowbanks far above. These are splendid peaks, unmatched in a land of
-muscled, brutal earth. Even the air seems to retain the scent of glacier
-work.
-
-At last the view of Hole-in-the-Wall, a staircase cirque excavated
-between the gigantic spread ribs of Mt. Custer. The slopes of beargrass
-are seed-spotted and gaunt now, the white fullness gone. Western
-pasqueflowers have accomplished their magic transformation; known in
-this season as old man’s beard, they nod their tufts of grizzled
-seedhead silk in the wind. Red and yellow monkeyflowers bloom yet,
-crowding along the many stream courses, and waterloving sedges and
-mosses surround pools of collected water on the broad horseshoe tiers.
-
-A spur trail drops down into the campground on the last ledge. Through a
-cleft in its lip plummets the gathered water of the basin. From the
-valley below, the waterfall appears to be springing from a hole in the
-headwall, giving this basin its name. Down, down, down, roars the water
-where once a mighty glacier ground its teeth.
-
-I leave until later the making of camp; by now the sharp shadows of
-Boulder Peak stab the valley forest and are beginning the upward assault
-of Thunderbird.
-
-Around the basin headwalls, last winter’s snowbanks remain formidable.
-Snow caves send out meltwater torrents. Glacier lilies and patches of
-spring beauty line their fringes. Pasqueflowers bloom in pockets. Here,
-among the asters of August, bloom also the first flowers of spring,
-shooting up as the snowbanks shrink, making these spots of snow-free
-ground a patchwork of May and July, August and June. The shrubs that
-line the furious water are willows, still bud-swollen this tenth day of
-September. The coming days will bring a sharp surprise.
-
-Winter will soon stop the melting of this snow. Could it be that I am
-seeing the first year of a reawakening ice age? If so, each year the
-snowfields would grow thicker and broader, connecting the shelves into
-one ice mass again, lilies and willows entombed, the summer heat failing
-to rescue them, until the ice at last began to slide, stripping the soil
-and once more plucking at living rock.
-
-Then these dwarfed fir, which cling precariously to the cliffs and hide
-behind the backs of boulders, would be in more danger than they were
-from their recent antagonists. Engulfed by ice, they would know the
-shearing wind no more. Their skeletons would rain down into the valley
-below, signalling another long forest retreat. But they have waited out
-the mountain ice before and would send their seeds again to this valley,
-changed however it might be, as they have always done.
-
-Evening brings out two sleek mule deer does. As they graze, their large
-ears stand erect, sorting out the lesser sounds from the ceaseless roar
-of water. Both raise their heads and point their ears, statue straight,
-at the scuttle of a porcupine. A noise among the rocks draws a backward
-glance and focus of those ears. I would like the sensitivity of such
-fine equipment, to hear what deer have always heard.
-
-Setting about the business of camp, I wonder about those animals that
-watched me for a while, then moved off, having seen a tent go up before.
-With the appearance of the moon the wind increases and they test the air
-more often now. Do they have visions of cougar or grizzly with every
-snap the wind delivers?
-
-In summer these high meadows see a surprising variety of animal life.
-Briefly out of hibernation are marmots and the handsome golden-mantled
-ground squirrels. Mice, voles, shrews, and woodrats run among the
-shadows, feeding on the season’s feast of seeds and insects. A nightmare
-for these are the fierce little weasels that haunt the rocks.
-
-Tracks of cougar and wolverine are sometimes seen, often teasingly
-fresh; to glimpse either of these elusive predators is to taste the
-finest wine of wilderness.
-
-Before the berry season, grizzlies grub the meadows for the tasty bulbs
-of glacier lilies and the tubers of spring beauty; often distracted by
-the scent of a ground squirrel in its burrow, they sometimes make a huge
-excavation for a small reward.
-
-White-crowned sparrows sing in July from the low tops of the battered
-trees, though their nests are on the ground below. Grey-crowned rosy
-finches patrol the drier ground for seeds while water pipits hunt
-insects in the wet areas. High above, a golden eagle scans the basin
-again, circling slowly before following a ridge south to sight another
-likely slope in its 10,000-hectare territory.
-
-The moon shines through the tent top. The wind, blowing more violently
-now, shivers the nylon and interrupts the voice of the waterfall. I have
-followed the pasqueflower run from the April prairies here to its
-highest bloom near treeline. I think about the triangular seed pods of
-the glacier lilies, colonies of steep-throated blue gentians, and the
-season’s last glory of goldenrod. Indian paintbrush, from white to fire
-red, blazes the slopes that light the fringes of sleep.
-
-I awake to a determined rain, the moon gone and the tent shuddering with
-wind-blast. I try not to think of the steel-cold air, and slip into a
-fitful sleep that seems an endless treadmill of rocky trail.
-
-Stiff and unrefreshed, I look out into the dawnless morning. The tip of
-Thunderbird is detached from its base by grey clouds swirling at its
-throat. A wave of sleet slants down, dancing on the rocks, chanting
-triumph over the buried, bent, and broken flowers of yesterday.
-
-So I must make my escape, short of Boulder Pass. Unattainable now,
-invisible above the cirque, that high pass grows in my memory. This
-testament to what a glacier can do, to the struggle of trees and the
-life-pioneers that invade such harsh places, is at my feet but shrouded
-with snow. My hands grow stiff and numb in the blunt work of packing up.
-
-I had wished to see Kinnerly Peak again, rising from the western Kintla
-valley, and walk along black ledges of the lava that floors the pass.
-Beyond it grows a grove of subalpine larch, stately, seldom encountered,
-the least common tree species in Glacier. Confined to this narrow zone
-between forest and alpine, it reaches up tall and proud, impervious to
-the gruelling climate that makes cowering shrubs of other trees.
-
-But all must wait another year, for this season comes down hard. And the
-will of winter is to erase whatever summer had devised.
-
-
-Tundra
-
-Porcelain-cold, the November sun dawns in the southeast sky. The ledges,
-ice-encrusted, layered with sleet from a recent squall, whistle the cold
-morning wind aside. Rattling down, a slide of rock plunges off the final
-ledge, seconds passing before the hollow sounds of impact clatter back.
-Like an apparition of winter itself, white beard bent sideways by the
-wind, a mountain goat steps to the precipice edge. Looking out across
-the vast white void, its long belly hair and pantaloons streaming with
-the ceaseless wind, this strange animal, product of some unfathomable
-ingenuity, hesitates but a moment; dropping down from step to invisible
-step along the sheer rock face, fracturing the ice glaze as it goes, it
-turns a wall and disappears. A nimble, eight-months-old kid follows.
-
-Blinking and twisting in the dull light, the shower of shattered ice
-clinks softly downward against rock, fading away like the short summers
-of this place.
-
-But while the wind chants winter, life has made a passage here, and also
-waits, hidden in seed and root and den.
-
-
-■
-The nanny and her kid have bedded down now, looking across the deep,
-snowy basin below. Their ledge shines with the first spear of sunlight.
-
-Far below the pass that connects Mount Siyeh to the snow-giants Matahpi
-and Going-to-the-sun, three male white-tailed ptarmigan emerge from
-their night’s huddle within a snowbank and step out to peck at an
-exposed mat of willow. Ptarmigan, the only birds on the winter tundra,
-wear white plumage in this season, helping to camouflage them in the
-snow, just as their mottled brown summer plumage makes them difficult to
-detect among bare rocks. There are few predators here to hunt them now,
-but they move with habitual slowness; quick movement can be fatal when
-summer brings numerous eyes to scan the slopes. With legs and feet
-heavily feathered and sharp claws to scratch for food beneath the snow,
-the ptarmigan live at truce with winter. When blizzards rage between the
-peaks, they nestle together in snow dens, beyond the reach of the winds.
-Ptarmigan hens winter lower in taller willow thickets, but the males
-prefer to take their winter as high as possible.
-
-Now they crouch behind the wind-deflecting rocks, dozing in the meager
-warmth of the morning sun.
-
-Near the snowless summit crags, a flash of brown fur zigzags among the
-rocks. That would be a pika. Only for a moment does it show itself, so
-quickly does it move.
-
-Also called the rock rabbit, the diminutive pika belongs to the order of
-hares and rabbits. Resembling a small guinea pig, this sturdy creature
-spurns hibernation as a way to beat the challenge of winter. Instead, it
-spends the summer laying in a store of hay for the lean season,
-spreading cut grass to cure upon the rocks and tending its “haystacks,”
-on which its survival hangs.
-
-Winter is a great peril to small mammals. Their small bodies, because of
-a large surface area in relation to volume, retain heat poorly, and
-their high metabolic fires consume calories quickly. Great amounts of
-energy are required to sustain an active animal in rough terrain,
-placing further demands on the animal’s capacity to survive the cold.
-The pika may need to stack as much as 25 kilos of hay; to keep its
-furnace burning during winter it will have to fuel its stomach almost
-hourly.
-
-Small animals of cold climates often show distinctive body adaptations.
-On the pika the small, rounded ears lie flat along the head, the tail is
-inconspicuous, the legs are short; heat loss from exposed surfaces is
-thus reduced. Fur insulates the soles of the pika’s feet while at the
-same time providing good traction on steep rock faces.
-
-Hidden below these rocks are the hibernating marmots and the sleeping
-ground squirrels. Beneath the snow the mice, shrews, and pocket gophers
-struggle on with their lives. But above ground, directly confronting
-this arctic climate, are the pika, the ptarmigan, and the mountain goat.
-
-A triumph of adaptation, the mountain goat faces the winter day without
-benefit of either the pika’s den or the ptarmigan’s snow roost.
-
-The nanny and kid descend from their ledge to search out browse at
-treeline with other members of a loose band—yearlings, young males,
-other nannies with kids. At the fringes of the band a solitary adult
-billy only grudgingly associates with other members of his kind; for
-this is the season of rut.
-
-Not really goats at all, these relatives of the European mountaineering
-chamois are insulated from the wind by coats of long, hollow-haired fur
-overlying woolly underfur. They are stocky, stiff-legged, and
-deliberate, able to negotiate the walls and pinnacles with their
-superbly adapted hoofs. The unique design of these hoofs gives the
-animal great traction and stability on precarious crags. Opening towards
-the front, the cleft between the two hoofs spreads each outward as the
-animal descends a slope, helping to grip the rocky surface. In addition,
-the large, rough, and pliant sole of each foot conforms to the bare
-rock, increasing traction.
-
-There is little need for the goat to leave its steep sanctuaries; it can
-subsist on lichens and mosses if browse is not available. It depends on
-the inaccessibility of the cliffs for its security. Accidents,
-avalanches, and rockfall are greater enemies than predators. Golden
-eagles sometimes attempt to knock newborn kids from ledges and a young
-goat quickly retreats under its nanny when an eagle soars by. With the
-protection of sharp spike horns and a terrifying terrain, adult goats
-seldom fall victim to cougar or grizzly.
-
-It will be a long time before the snow releases this land and wapiti,
-bighorn, grizzly, and cougar wander back into these high basins. In this
-winter minimum of life, the spring songs of rosy finches, water pipits
-and white-crowned sparrows seem an impossible extravagance.
-
-
-■
-I am drawn to the spring tundra—to the vigor and tenacity of its sparse
-life—where survival itself seems ceremony enough. But it is a strange
-world, where a man is out of perspective. Here the plant cover is
-carpet-high, and distance, for the lack of trees, tricks the eye. Here
-the wind, snow, and sun quickly burn skin, and the intense light,
-reflected from snowbanks, stabs at the eyes. Almost instantly, a
-sandwich is sucked dry of its moisture. The desiccating wind probes the
-ears until it seems at last to pierce your brain. Except for fearful
-mountain walls the only shadow is your own. Animals seem somehow remote
-and unknowable, as if seen through glass. A day on the tundra and you
-feel the want of a company of trees.
-
-Yet once exposed, you acquire a craving for the look of tundra. Nowhere
-else is there such an impatience for spring—the flowers rush into bloom;
-the male water pipit soars, its skylark song crystal sharp in the thin
-air. The nesting birds are restless, for sun-days and warm days are few,
-precious, and quickly spent. Insects and spiders abound—flying about the
-peaks or crawling among the rocks.
-
-Summer brings bands of bighorn rams up from the valley to explore the
-highest meadows. Though not so sure-footed as the goats, they too have
-hoofs adapted to climbing steep faces, and they walk the slopes not far
-below the goats.
-
-Marmots, which whistle sharply when threatened, spend their days
-sunbathing and grazing; they must fill out their now loose-hanging fur
-coats with life-sustaining fat for the coming winter.
-
-Alpine animals are blessed with mobility and can choose their weathers,
-retreating to burrow, den, or rock-harbor to escape the worst fury of
-storms. But what about the plants, rooted forever in one spot, assaulted
-by an untempered sun and a drying wind, and facing the almost daily
-threat of freeze and storm?
-
-Alpine plants, through their design and growing habits, have adapted
-themselves to the rigorous demands of this climate in many ways. Most
-plants are perennial: there just aren’t enough days or nutrients
-available for the growing of entire plants each year from seed. And they
-have the ability to grow and carry on photosynthesis at temperatures
-just above freezing, thus extending their season. In this zone,
-temperatures are rarely above 15° C; the mean summer temperature is
-about 10° C. But a flower such as the alpine buttercup, which is found
-at treeline or above, can grow through several centimeters of snow; heat
-given off during the plant’s respiration will create an opening through
-which it can emerge.
-
-Plants have various adaptations to meet the demands of the alpine
-environment. Yellow stonecrop, not restricted to this zone, is
-nevertheless able to survive here because of its fleshy succulence and a
-waxy covering that prevents water loss. On some plants, protective hairs
-covering leaves and stems help retard the burning effects of wind and
-sun. Often this pubescent foliage looks more grey than green, for the
-soft hairs mute the color.
-
-Cushion growth is another alpine adaptation. The moss campion cushion,
-covered with delicate pink flowers, grows to about one-third of a meter
-across and only 3 to 5 centimeters high. Spreading out close to the
-ground, the plant avoids the major violence of the wind and hoards
-moisture like a sponge.
-
-The dryad, growing abundantly on the windy sweep of Siyeh Pass, shows
-alpine adaptations in several ways. The energy of the mature plant is
-channeled primarily into reproduction: its large flower, supported by a
-short stem, matures quickly; and it produces many seeds, ensuring
-germination of a few. An evergreen, it begins to synthesize water and
-carbon dioxide into food as soon as the snow is gone; and its rolled
-leaves prevent rapid evaporation. It grows as a low and woody mat that
-year by year extends itself through the production of new shoots that
-carpet the rock. Mat growth has the advantage of retaining dead plant
-material and capturing wind-blown grains of soil, allowing the plant
-slowly to enlarge its soil base.
-
-Compared to the forest, the heartbeat of the tundra is painfully slow.
-Here a plant may grow for a quarter of a century before it has acquired
-the reserves necessary for flowering. Contrasted with the progress on
-the tundra, forest succession races by with dizzying speed. Yet
-imperceptible as the change may be the alpine plant community also
-passes from pioneer to climax.
-
-Beyond the limit of other plants, lichens thrive, encrusting rocks with
-their rainbow colors. A lichen is actually a primitive and highly
-successful association between a fungus and an alga, working together
-for mutual benefit. The fungus protects the delicate alga, trapping and
-holding moisture; the green alga, in turn, produces enough food to
-sustain the needs of the fungus.
-
-Generating rock-disintegrating acids that help secure this partnership
-to the rock, lichens, along with physical weathering, help break down
-the rocks into soil particles. Collected in pockets by run-off or wind,
-rudimentary soil is slowly invaded by cushion plants. After centuries of
-colonization by these, while the meager soil is deepened and enriched
-and moisture retention is increased, other plants move in, climaxing at
-last in hardy grasses and sedges. As in the forest, pioneer species
-change the environment to their detriment, creating a habitat better
-suited to other species.
-
-Although it will progress with geologic slowness, the rocky ground of
-Siyeh Pass—its plant cover presently scant and wind-rowed by frost-heave
-and relentless wind—will in time develop grasses and sedges, the climax
-vegetation of the alpine meadows.
-
-
-■
-Simplicity rules the alpine zone. Here life is reduced to bare
-essentials. Chief controlling force is climate; but the plants and
-animals that live here are well adapted. Compared to the lower realms,
-where both competition and predation are fierce, life here looks secure.
-
-There is a penalty to simplicity. In the lowland, the long food chains
-and diversity of species, the long growing season, and the abundant food
-supply give the forest an adjustment mechanism and healing power not
-found on the critically balanced tundra. The greater the variety in a
-plant and animal community, the greater the stability. So in the alpine
-world there exists a paradox: the most durable life forms constitute the
-most fragile community.
-
-
-The Water Communities
-
-Snowfields begin again their summer-long melt. The alpine stream, vocal
-again, collects its water from a thousand places. Miniature gorges drain
-the meadow, gurgling with the sparkle and rush of meltwater in the
-lengthening spring days.
-
-Gathering volume, the stream seems to hurry faster; at the first rock
-staircase, it begins to sing. I follow the gully downward, drawn like
-the water. There is excitement in the growing dash and roar, a wind-gust
-sweeping spray into the air. A rainbow appears, holding steady to the
-swirling cloud of spray, then doubles and abruptly disappears.
-
-At the first great plunge the water lunges outward over the lip. Like
-glass at shattering, long shards lance out. But the wind feathers the
-sharp edges as they fall.
-
-The close thunder of a waterfall beats at your head, and your mind must
-shout to think. Here is water, a most amazing and most important
-substance. Perhaps some of this same water was once part of the ancient
-sea in which was laid down the mudstone of this ledge; was once drunk by
-dinosaurs; has coursed the globe countless times; and has flowed in this
-very stream before. In solid, liquid, or gaseous form, it goes through
-its own cycle. Together with sunlight, water makes possible and
-maintains all life on Earth.
-
-
- Ouzel Music
-
-A glacier might cling to a winter snow a hundred years and turn it to
-ice, a blue tool to rasp and pluck at rocks, before letting it go.
-Lingering summer snowfields might delay its passage for a time. But the
-water always wins at last, becoming, in one decisive instant, liquid
-again, and beginning its long journey to the sea. Plants and dry air
-will intercept some of its molecules, sending them back into the
-atmosphere to bloom as fog and cloud; but as rain, snow, or dew, these
-are soon commissioned to the land again.
-
-Water is so familiar to us that we seldom think about it. We know that
-fish swim in the lower lakes, and we are vaguely aware of the
-bewildering assortment of life-forms abounding in a pond. But life
-begins in the streams.
-
-Even cups of cold meltwaters, scooped out of a rivulet only a few meters
-away from its snowbank source, contain some life. Snow algae, which grow
-on the snowbank surface, often sufficiently dense to give the snow a
-distinctive red complexion, are released into the meltwater. In summer,
-small invertebrate life can be discovered in the standing pools of even
-the highest cirque.
-
-But conditions are not good for the development of complete aquatic food
-chains in the streams and lakes of higher elevations. Alpine lakes, or
-tarns, support little visible life. Often flanked by high ridges and
-peaks, many tarns receive scant direct sunlight during the day. Since
-these lakes occupy basins that capture tremendous amounts of snowfall,
-the snowbanks persist in the mountain shadows, and summer makes little
-progress in warming the water. Iceberg Lake, for example, is seldom free
-of floating ice, and its temperature never rises above 4° C in summer,
-even at the surface.
-
-Moving out of the cirque lakes, water is soon churning again, dashing
-downward many hundreds of meters to the valleys below, in rapids,
-cascades, and breathless waterfalls. Not surprisingly, few plants and
-animals are adapted to life in fast-moving water.
-
-Algae can be found covering streambed rocks and stranded, water-polished
-tree trunks. Securely attached by holdfasts, these small plant forms
-survive the rigorous stream flow that would destroy the larger vascular
-plants. Several species exist, from microscopic forms to branched
-filamentous algae whose long hairlike strands wave in the current.
-
-A surprising number of insects live on the stream bottom, finding a
-measure of protection from the current in the jumble of rocks.
-Underwater beetles live under the gravel or among the debris at the
-stream-edge, or cling to stones and sticks. Scurrying and creeping among
-the rock-crannies are the larvae of stoneflies, mayflies, and
-caddisflies. These and the small fish that venture up from lower lakes
-are the food of the water ouzel, a creature that loves the places where
-the waters thunder.
-
-
-■
-The noise of the water is overpowering. A slip into this boiling rage
-would mean quick death. Looking 10 meters across the dim, mist-slippery,
-water-scoured canyon, I see a young water ouzel peering out of its
-unique nest, on the lookout for its parents. Clouds of spray keep the
-nest of living moss continually wet; but this bird is waterproofed with
-an oily plumage and keeps its vigil at the nest opening. Peering into
-the torrent below, then upstream and downstream, it awaits patiently the
-delivery of the next meal.
-
-With the approach of one of the adults, three other heads crowd the
-opening, begging yellow mouths agape. Flying low, the ouzel parent zeros
-through the heavy spray, alighting on a slippery boulder below the nest
-ledge. Preparing to fly up to the nest with its load of insect larvae,
-the ouzel spots me across the water. At its sharp _jigic, jigic_ alarm,
-the bills of the young snap instantly shut. Nervously the bird regards
-my close presence, dipping its entire body rapidly up and down, as if
-keeping time with the surging torrent.
-
-Discovering no danger, the dusky blue-grey bird bobs more slowly. The
-other adult, returning from an upstream forage, alights on the same
-rock, occasioning a new outcry from the fledglings. Each in turn, the
-parent birds fly up to feed their young, beating their wings to maintain
-their position at the perchless nest. Not pausing to regard me further,
-they split the stream between them again, one flying upstream and one
-down, to continue the hunt. Blinking and shaking the collected mist from
-its bill, the single young sentry renews its watch.
-
-
- In Shallow Waters
-
-Life abounds in the shallow lakes and ponds. Calm, protected John’s Lake
-offers a fine example of how a complex aquatic plant-and-animal
-community can exist in balance in a confined space. The water teems with
-the microscopic algae, protozoans, and rotifers that sustain the barely
-visible zooplankton. Dancing, flitting, hopping, and swaying through the
-water, these zooplankton in turn support the larger plankton-eating
-animals.
-
-Dragonflies and damselflies shoot past, crackling their wings, and perch
-in the bog grass. Looking into the shallow water, you will see a wealth
-of small animal life. A spotted frog swims into view, floating to the
-surface beside a lily pad so that its eyes protrude above the water.
-
-The ribbonlike form of a leech swims across the bottom toward deeper
-water. Looking closer, you see that the water swarms with bizarre
-shapes—water boatmen propelling themselves with oarlike appendages, a
-gliding mayfly nymph, then a predacious diving beetle surfacing,
-grasping a bubble of air beneath its shiny brown wing plates and
-disappearing downward again—the bubble’s edge shining silver—into the
-brown bottom debris. Suddenly a whirligig beetle sets the surface to
-spinning, wrinkling the view below.
-
-Everywhere in the water there is animal life, forms that are attached,
-free-swimming, crawling on the bottom, and clinging to or swimming on
-the surface film. The gray, slimy encrustation on a sunken log looks
-like a covering of lichen but is really a freshwater sponge, a colonial
-animal that feeds by filtering minute plankton from the water. Another
-attached creature is the barely visible hydra; this twig-shaped
-predator, related to marine jellyfish, captures water fleas and other
-small animals in its several poisonous tentacles.
-
-Water beetles, backswimmers, water boatmen, and many other creatures
-move about more or less freely in the water, propelling themselves along
-with jerky movements. Suspended between surface and bottom are the
-zooplankton, the tiny water fleas, cyclops, daphnia, and others, which
-feed by filtering minute algae. On the bottom and below live scavenging
-worms. Water striders skate on the surface film.
-
-Along the shore, frogs, salamanders, garter snakes, and water shrews are
-hunting. Dabbling and diving ducks patrol about, tipping or submerging
-for the bottom plants. Moose tracks circle the muddy shore. Because it
-produces vegetation abundantly, John’s Lake sustains a great diversity
-of animal life.
-
-
- Beaver Ponds
-
-Fully 10 percent of all the present meadow area in the Rocky Mountains
-is estimated to have been created by beaver, the only animal besides man
-that engineers extensive changes in the environment to suit its own
-needs.
-
-When beavers dam a stream, they set in motion another form of
-succession. If the resulting backwater floods a forest area, the trees
-are soon killed, creating a broad opening in the forest canopy.
-Water-associated plants and shrubs quickly invade the pond and
-shoreline, creating favorable habitat for waterfowl, moose, blackbirds,
-amphibians, wading birds, warblers, marsh hawks, and a score of other
-animals.
-
-After many years the water becomes shallow, filling in with silt and
-plant debris. When the beavers abandon the site, the dam may rupture for
-lack of maintenance and the pond will rapidly drain. Or it may continue
-to hold, delaying for several more years its slow conversion to meadow.
-Stimulated by the nutrient-rich mud, the water grasses, sedges, and
-shrubs finally choke the water with their accumulating debris,
-transforming the area into a bog.
-
-Gradually the ground firms as more humus is created and more silt is
-trapped. The area becomes meadow, supporting grasses, sedges, and other
-flowering plants. Trees begin to reinvade the drier ground, and
-eventually the meadow reverts to forest. Centuries may be required to
-see this cycle through, from forest to pond, to bog, to meadow, to
-forest again. At each stage many of the animal inhabitants change: the
-song of the western robin and the chatter of a red squirrel in the
-original, pre-beaver forest give way to the croak of a heron; the heron
-is replaced by the insect-and-berry-eating cedar waxwing; the waxwing is
-followed by the tree-dwelling western robin and red squirrel.
-
-
- Lakes Cold and Deep
-
-Seeming to skate on its own reflection, a spotted sandpiper comes in low
-over the quiet water, wingtips almost touching the surface of the lake.
-It alights at the shore and folds its wings. Amid the rounded rocks,
-this plain but elegant little shorebird is all but swallowed up.
-Teetering constantly on long legs, it sets off along the water’s edge,
-pecking here and there, coming closer and closer, never forgetting to
-stop and curtsy, as if acknowledging, while hurrying offstage, the
-applause of an audience.
-
-As it draws near, several water striders skate away from the shore. A
-stonefly, scuttling between two rocks, is deftly speared. So large a
-morsel makes the bird pause and rough its feathers, then scamper into
-the water to take a drink. Teetering again, it passes in front of me and
-continues down the shore, where I soon lose sight of it rounding a rocky
-point.
-
-I am sitting at the foot of Lake McDonald, watching the darkness gather
-over the valley, seeing the last light slide upward to the tips of the
-distant mountains. As daylight dissolves, this long fleet of familiar
-peaks seems almost to glide toward darkness, slow and silent as sailing
-ships.
-
-The sheet of motionless water stretches many kilometers away between
-tree-covered moraines. The water is deep and cold. No emergent plants
-line the barren shore. It would seem that no life, except for the single
-gull that rests on the water far away, exists in this nearly
-thousand-meter-high lake.
-
-
-■
-Considering the great volume of Glacier’s large, deep lakes, the life
-they support is indeed meager. A large part of the reason lies with the
-nature of their shores, where almost no plants grow. A combination of
-factors prevents the development of a lush shoreline growth.
-
-Contoured like bathtubs, these steep-sided lakes exhibit narrow or
-non-existent shoreline shallows, which are vital for the production of
-rooted plants. Strong wave action and extensive seasonal fluctuations in
-the level of these natural reservoirs prevent the development of
-emergent water plants in locations where they might otherwise be
-expected.
-
-Since sunlight cannot penetrate to the bottom of these deep lakes, they
-are deprived of bottom-anchored plants in midlake as well. As a result,
-herbivorous animal life must depend almost wholly on algal growth. Wave
-action inhibits the spread of free-floating algae by washing much of it
-onto the shore. Deep lakes are also low in available oxygen, preventing
-the development of bottom decomposers, which would rapidly release
-nutrients as they break down the accumulating debris washed into the
-lake. Without a steady supply of nutrients, plant growth is retarded.
-
-Since the food chain depends upon green plants, the ability of a lake to
-support higher animals such as fish depends upon its ability first to
-produce adequate plant growth. The production of one kilo of trout
-requires that a lake produce about 1,000 kilos of plants to support 100
-kilos of herbivorous invertebrates, which are eaten by 10 kilos of
-carnivorous insects, on which the trout feed.
-
-Compared to smaller shallow lakes, which teem with visible life, cold,
-deep, nutrient-poor lakes such as McDonald appear to be watery deserts.
-Yet because of their great volume—Lake McDonald contains 5 or 6 cubic
-kilometers of water—these large lakes do sustain significant numbers of
-fish. Of the 22 kinds of fishes found within the park, most are
-coldwater species. Trout, whitefish, grayling, suckers, minnows, and
-carp fill the roles of herbivore, carnivore, and scavenger. Agile,
-highly mobile, and acutely sensitive, fish represent the most successful
-total adaptation to the aquatic environment.
-
-Through the stocking of nonnative species, including plantings in
-formerly fish-free lakes, the natural aquatic communities of many of
-Glacier’s lakes and streams have been permanently modified.
-
-Aquatic food chains are not confined to the water. Ospreys, ducks,
-mergansers, otter, mink, and many other semi-aquatic or terrestrial
-birds and mammals utilize the plants and animals of the water. In fall,
-a remarkable spectacle occurs along the outlet of Lake McDonald.
-Attracted to the kokanee salmon concentrations, which run from Flathead
-Lake to spawn and die in these clear, shallow waters, bald eagles
-collect to exploit the vulnerable fish. In 1977, 444 eagles were counted
-in one census. This food resource is also exploited by grizzlies,
-coyotes, skunks, gulls, loons, and other animals. On occasion, even
-white-tail deer have been observed swallowing salmon!
-
- [Illustration: Sunset]
-
-
-
-
- Shooting Stars
-
-
-This park is very special. The people who know it well feel proprietary
-toward its mountains, scattered lakes, and glaciers. Perhaps it is the
-arrangement of the land, an unsurpassed concentration of American
-wilderness. Time and again I have thought, as I regarded some aspect of
-this country, _yes, this is exactly right_—almost, it would seem, as if
-some magic existed that could translate thought and emotion into rock
-and bark.
-
-Glacier remains largely unexploited, bearing still the aspect of the
-Earth the Indians knew for 500 generations—a land where it is yet
-possible to feel a sense of discovery, sense that a single man matters.
-On too many mountains, man has tarnished whatever he has touched; but
-here the land has shed, as a fir sloughs snow, a long succession of
-traders, trappers, explorers, hunters, surveyors, prospectors, loggers,
-settlers, and tourists.
-
-You may walk the same trail a dozen times and not tire of the view. I
-have given up wondering why. I know only that these are mountains a man
-might grow old with, and that mountain-fever never diminishes but only
-changes its look, as a forest does over many years.
-
-Repeatedly I have noticed that this park creates an instant bond between
-strangers. A certain pause intrudes at the first mention of Glacier
-National Park, and a look of distance comes, as Red Eagle becomes real
-again, or the wind at Firebrand is remembered, or the flowers of
-Fifty-Mountain converge once more upon the senses.
-
-Never are we quenched. If a goshawk rushes past, straining upward with
-its squirming load of ground squirrel, forever afterward our blood
-demands more. The sight of a wolverine running is not enough. Nor the
-magnificent assemblage of bald eagles feasting on November salmon. More
-days of this: mountain goats leaping impossible ledges, wave tracks from
-a beaver reaching out on dawn water. There are messages here, loud as
-kingfishers. The land has languages, stories to tell.
-
-But in wilderness there is no moral, save that it must continue. For all
-our probings and plottings we discover no adequate interpretation of the
-forces we find swirling about us. A larch you must touch to know; your
-neck must feel the ache of too much looking up. Watch its treepoint
-pirouette. Then, looking back at the world level, you will find that you
-have lost all answers. We have learned the art of building bridges,
-cataloging plants, predicting what a shrew might do. Of the essential
-mystery, we know nothing.
-
-For nature assigns no “roles” to its creatures; there is no “reason” for
-a forest fire, which burns mightily but with no intent. Life’s only
-“purpose” is the feeding of life, and the beauty we see therein is but
-its lack of guarantee: for the chipmunk and the weasel, and the man who
-measures his life to theirs, no assurance of long days and tempered
-seasons, abundant seeds, ample meat. In wilderness there is mystery yet,
-unsimplified, not reduced, resplendent and immense.
-
-Whatever the conclusion of this planet, however many the acts to follow
-in this consuming drama—mountains coming up, mountains going down,
-forests, lakes, and seas skimming past like wind-driven scud clouds
-before a storm—at least in the scant shadow of this present age there is
-an achievement of sorts. For now, with this creature man, such things as
-mountains can be loved. And men have memories to fill.
-
-Tomorrow I will look for shooting stars—purple spring flowers that point
-their fire down, always down toward the center of the Earth, as if to
-give in their brief term beneath the sun a tribute to this most
-excellent mystery.
-
-Today I can say nothing more, neck-sore now from looking at larchtops
-swaying with the wind of this splendid morning.
-
- [Illustration: Shooting star.]
-
- [Illustration: Mountain goats.]
-
-
-
-
- Appendix
-
-
-Mammals of Glacier National Park
-
-Distribution information was obtained from _Meet the Mammals of
-Waterton-Glacier International Peace Park_, by Robert C. Gildart (see
-Reading List). Nomenclature follows, for the most part, _a Field Guide
-to Mammals_, by William H. Burt and Richard P. Grossenheider.
-
- Key to symbols:
- E—occurs east of Continental Divide (spruce-fir forest; aspen;
- bunchgrass meadows)
- W—occurs west of Continental Divide
- (redcedar-hemlock-lodgepole-fir-larch forest; some meadows)
- A—occurs in alpine areas (above upper edge of continuous forest)
- R—rare in Glacier National Park
-
- Shrews
- Masked shrew, _Sorex cinereus_
- E, W, coniferous forests, meadows, pond and stream edges
- Vagrant shrew, _Sorex vagrans_
- E, W, A, moist forests and grasslands, marsh and stream edges
- Northern water shrew, _Sorex palustris_
- E, W, stream edges
-
- Bats
- Little brown myotis, _Myotis lucifugus_
- E, W, coniferous forests, often around buildings, caves; nocturnal
- Long-eared myotis, _Myotis evotis_
- E, W, A, R, coniferous forests, meadows; nocturnal
- Long-legged myotis, _Myotis volans_
- E, W, A, coniferous forests, meadows; nocturnal
- Big brown bat, _Eptesicus fuscus_
- E, W, coniferous forests; often around buildings, caves; nocturnal
- Silver-haired bat, _Lasionycteris noctivagans_
- E, W, coniferous forests; meadows; nocturnal
- Hoary bat, _Lasiurus cinereus_
- E, W, coniferous forests; mostly nocturnal
-
- [Illustration: Cougar]
-
- Cats
- Bobcat, _Lynx rufus_
- E, open forests, brushy areas
- Lynx, _Lynx canadensis_
- E, W, coniferous forests
- Cougar, _Felis concolor_
- E, W, coniferous forests
-
- Raccoon, bears
- Raccoon, _Procyon lotor_
- E, W, R, open forests, stream bottoms
- Black bear, _Ursus americanus_
- E, W, A, forests, slide areas, alpine meadows
- Grizzly, _Ursus arctos_
- E, W, A, forests, slide areas, alpine meadows
-
- [Illustration: Coyote]
-
- Canines
- Red Fox, _Vulpes vulpes_
- E, grasslands, open forest
- Coyote, _Canis latrans_
- E, W, A, forests, grasslands
- Gray wolf, _Canis lupus_
- E, W, R, coniferous forests
-
- [Illustration: Wolverine]
-
- [Illustration: Longtail weasel]
-
- Mustelids
- Striped skunk, _Mephitis mephitis_
- E, W, open forests, grasslands
- Badger, _Taxidea taxus_
- E, W, grasslands
- River otter, _Lutra canadensis_
- E, W, R, rivers, lakes
- Wolverine, _Gulo gulo_
- E, W, A, coniferous forests, alpine meadows
- Least weasel, _Mustela rixosa_
- E, R, open forests, grasslands
- Shorttail weasel, _Mustela erminea_
- E, W, A, coniferous forests, meadows
- Longtail weasel, _Mustela frenata_
- E, W, A, open forests, meadows
- Mink, _Mustela vison_
- E, W, creek and lake edges
- Marten, _Martes americana_
- E, W, A, coniferous forests
- Fisher, _Martes pennanti_
- E, W, R, coniferous forests
-
- Lagomorphs
- Pika, _Ochotona princeps_
- E, W, A, rockslides
- Snowshoe hare, _Lepus americanus_
- E, W, coniferous forests
- Whitetail jackrabbit, _Lepus townsendii_
- E, W, R, grasslands
-
- Squirrels
- Hoary marmot, _Marmota caligata_
- E, W, A, rocky areas, alpine meadows
- Richardson ground squirrel, _Spermophilus richardsonii_
- E, R, grasslands
- Columbian ground squirrel, _Citellus columbianus_
- E, W, A, open woodlands, grasslands, alpine meadows
- Thirteen-lined ground squirrel, _Spermophilus tridecemlineatus_
- E, R, grasslands
- Golden-mantled squirrel, _Spermophilus lateralis_
- E, W, A, high, open forests; rocky areas
- Least chipmunk, _Eutamias minimus_
- E, W, A, high, open forests; brushy, rocky areas; alpine meadows
- Yellow pine chipmunk, _Eutamias amoenus_
- E, W, open forests; brushy, rocky areas
- Redtail chipmunk, _Eutamias ruficaudus_
- E, W, open forests; brushy, rocky areas
- Red squirrel, _Tamiasciurus hudsonicus_
- E, W, coniferous forests
- Northern flying squirrel, _Glaucomys sabrinus_
- E, W, coniferous forests; nocturnal
-
- Pocket gophers
- Northern pocket gopher, _Thomomys talpoides_
- E, W, A, meadows
-
- [Illustration: Beaver]
-
- Beaver
- Beaver, _Castor canadensis_
- E, W, streams, lakes
-
- Voles and kin
- Deer mouse, _Peromyscus maniculatus_
- E, W, A, forests, grasslands, alpine meadows
- Bushytail woodrat, _Neotoma cinerea_
- E, W, A, rocky areas, old buildings
- Northern bog lemming, _Synaptomys borealis_
- W, R, coniferous forests
- Mountain phenacomys, _Phenacomys intermedius_
- E, W, A, coniferous forests, alpine meadows
- Boreal redback vole, _Clethrionomys gapperi_
- E, W, coniferous forests
- Meadow vole, _Microtus pennsylvanicus_
- E, W, open forests, meadows; along streams; marshy areas
- Longtail vole, _Microtus longicaudus_
- E, W, coniferous forests, grasslands
- Water vole, _Arvicola richardsoni_
- E, W, A, high-elevation stream and lake edges
- Muskrat, _Ondatra zibethica_
- W, streams, lakes, marshy areas
- Western jumping mouse, _Zapus princeps_
- E, W, A, grasslands, alpine meadows
-
- Deer
- Wapiti (American elk), _Cervus canadensis_
- E, W, A, open forests, meadows
- Mule deer, _Odocoileus hemionus_
- E, W, A, open forests, meadows, often at high elevations
- Whitetail deer, _Odocoileus virginianus_
- E, W, coniferous forests, meadows, creek and river bottoms
- Moose, _Alces alces_
- E, W, coniferous forests, lakes, slow streams, marshy areas
-
- [Illustration: Mountain goat]
-
- Bovids
- Mountain goat, _Oreamnos americanus_
- E, W, A, high peaks and meadows
- Bighorn, _Ovis canadensis_
- E, A, open mountainous areas
-
-
-Reptiles and Amphibians of Glacier National Park
-
-Note: This check list is based upon actual specimens in the Park and
-other collections, according to Dr. Royal Brunson, Montana State
-University.
-
- Reptiles
- Great Basin Garter Snake, _Thamnophis elegans vagrans_
- A large garter snake of mountainous areas, usually with large spots.
-
- Great Plains Red-sided Garter Snake, _Thamnophis ordinoides
- parietalis_
- Dorsal stripes varying from yellow to blue or black. Usually found
- near water.
-
- Hypothetical List:
- Rubber Boa, _Charina bottae utahensis_
- May occur in rock slides or, possibly, in forested areas, on either
- side of the Divide.
-
- Gopher Snake, _Pituophis catenifer sayi_
- May occur along eastern boundary (Great Plains).
-
- Yellow-bellied Blue Racer, _Coluber constrictor mormon_
- May occur on eastern boundary of Park along border of Great Plains.
-
- Painted Turtle, _Chrysemys picta_
- May occur in ponds and sluggish waters from Upper Sonoran Zone to
- Canadian Zone.
-
- Western Skink, _Eumeces skiltonianus_
- May occur in Transition Zone along western border of Park.
-
- [Illustration: Northern Alligator Lizard]
-
- Northern Alligator Lizard, _Gerrhonotus coeruleus principis_
- May occur in Transition Zone along western border of Park.
-
- Amphibians
- Tiger Salamander, _Ambystoma tigrinum melanostrictum_
- Ground color either black or bluish-black, with large spots or
- blotches of yellow.
-
- Long-toed Salamander, _Ambystoma macrodactylum_
- Ground color black or dark brown; wide band of yellow extends from
- back of head to tip of tail.
-
- Northwestern Toad, _Bufo boreas boreas_
- Widely distributed over entire Park. (Also known as Columbian,
- Northern, or Western Toad.)
-
- [Illustration: Western Spotted Frog]
-
- Western Spotted Frog, _Rana pretiosa pretiosa_
- Widely distributed over entire Park. (Also known as Western or Pacific
- Frog.)
-
- Green Frog, _Rana clamitans_
- One specimen, from Bowman Lake. (Chicago Natural History Museum)
-
- Tailed Frog, _Ascaphus truei_
- Should be fairly common, although it is not often taken.
-
- Pacific Tree-toad, _Hyla regilla_
- Small size and disks on fingers and toes identify this species. Common
- throughout Park.
-
-
-Fishes of Glacier National Park
-
-Classification and common scientific names are from: “A List of Common
-and Scientific Names of Fishes from the United States and Canada,”
-American Fisheries Society Publication No. 2, 1960.
-
- Key to symbols:
- N Species native to at least one major drainage of the Park.
- I Non-native species, having been introduced into Park waters by man.
- S A species of sporting qualities and valued for recreational angling.
- 1 Waterton Drainage
- 2 Belly River Drainage
- 3 Swiftcurrent Drainage
- 4 St. Mary Drainage
- 5 Two Medicine Drainage
- 6 Middle Fork Flathead River Drainage (exclusive of McDonald Valley)
- 7 McDonald Valley Drainage
- 8 North Fork Flathead River Drainage
-
- [Illustration: Lake Trout]
-
- Family _Salmonidae_ (trouts, whitefishes, and grayling)
- Lake Whitefish, _Coregonus clupeaformis_ (I) (1, 2, 3, 4, 7)
- Pygmy Whitefish, _Prosopium coulteri_ (N) (7)
- Mountain Whitefish, _Prosopium williamsoni_ (N) (S) (1, 2, 3, 4, 5, 6,
- 7, 8)
- Kokanee (Sockeye) Salmon, _Oncorhyncus nerka_ (I) (S) (3, 7, 8)
- Cutthroat Trout, _Salmo clarki_ (N) (S) (1, 2, 3, 4, 5, 6, 7, 8)
- Rainbow Trout, _Salmo gairdneri_ (I) (S) (1, 2, 3, 4, 5, 7)
- Brook Trout, _Salvelinus fontinalis_ (I) (S) (1, 2, 3, 4, 5, 6, 7)
- Dolly Varden, _Salvelinus malma_ (N) (S) (1, 2, 3, 4, 6, 7, 8)
- Lake Trout, _Salvelinus namaycush_ (N) (S) (1, 2, 4, 5, 7, 8)
- Arctic Grayling, _Thymallus arcticus_ (I) (S) (2, 8)
-
- Family _Esocidae_ (pikes)
- Northern pike, _Esox lucius_ (N) (S) (1, 2, 3)
-
- [Illustration: Redside Shiner]
-
- Family _Cyprinidae_ (minnows and carps)
- Longnose Dace, _Rhinichthys cataractae_ (N) (2, 3, 4, 5, 6, 7, 8)
- Northern Pearl Dace, _Margariscus margarita_ (N) (3, 5)
- Redside Shiner, _Richardsonius balteatus_ (N) (7, 8)
- Streamline Chub, _Hybopsis dissimilis_ (N) (1, 3)
- Northern Squawfish, _Ptychocheilus oregonensis_ (N) (7, 8)
-
- [Illustration: White Sucker]
-
- Family _Catostomidae_ (suckers)
- White Sucker, _Catostomus commersoni_ (N) (1, 2, 3, 4, 5)
- Largescale Sucker, _Catostomus macrocheilus_ (N) (6, 7, 8)
- Longnose Sucker, _Catostomus catostomus_ (N) (1, 2, 3, 4, 5, 6, 7, 8)
-
- Family _Gadidaie_ (codfishes and hakes)
- Burbot, _Lota lota_ (N) (S) (1, 4)
-
- Family _Cottidae_ (sculpins)
- Mottled sculpin, _Cottus bairdi_ (N) (5, 6, 7, 8)
- Spoonhead sculpin, _Cottus ricei_ (N) (1, 2, 3, 4)
-
-
-Birds of Glacier National Park
-
- Key to symbols:
- E—occurs on east side of the park (east of the Divide)
- W—occurs on west side of the park (west of the Divide)
- A—occurs in alpine areas
- ab—abundant
- c—common
- u—uncommon
- r—rare
- i—introduced
- a—accidental
-
- [Illustration: Common Loon]
-
- Loons
- Common Loon E, W, ab
- Arctic Loon?
- Red-throated Loon?
-
- [Illustration: Western Grebe]
-
- Grebes
- Red-necked Grebe E, W, c
- Horned Grebe E, W, ab
- Eared Grebe E, W, c
- Western Grebe E, W, u
- Pied-billed Grebe E, W, r
-
- Pelicans, Cormorants
- White Pelican E, W, u
- Double-crested Cormorant E, r
-
- [Illustration: Great Blue Heron]
-
- [Illustration: American Bittern]
-
- Herons, Bitterns
- Great Blue Heron E, W, c
- Black-crowned Night Heron a
- American Bittern, W, r
-
- [Illustration: Mallard]
-
- [Illustration: Wood Duck]
-
- [Illustration: Ruddy Duck]
-
- Swan, Geese, Ducks
- Whistling Swan E, W, ab
- Trumpeter Swan E, W, r
- Canada Goose E, W, c
- Snow Goose E, W, c
- Ross’ Goose E, W, r
- Mallard E, W, ab
- Gadwall E, W, r
- Pintail E, W, c
- Green-winged Teal E, W, c
- Blue-winged Teal E, W, u
- Cinnamon Teal E, W, u
- European Widgeon E, W, c
- American Widgeon E, W, ab
- Northern Shoveler E, W, c
- Wood Duck E, W, r
- Redhead E, W, c
- Ring-necked Duck E, W, u
- Canvasback E, W, u
- Lesser Scaup E, W, c
- Greater Scaup?
- Common Goldeneye E, W, c
- Barrow’s Goldeneye E, W, ab
- Bufflehead E, W, u
- Harlequin Duck E, W, c
- White-winged Scoter E, W, r
- Ruddy Duck E, W, c
- Hooded Merganser E, W, u
- Common Merganser E, W, ab
- Red-breasted Merganser, E, W, u
-
- [Illustration: Cooper’s Hawk]
-
- [Illustration: Marsh Hawk]
-
- Vultures, Hawks, Eagles
- Turkey Vulture E, W, r
- Goshawk E, W, c
- Sharp-shinned Hawk E, W, u
- Cooper’s Hawk E, W, u
- Red-tailed Hawk E, W, c
- Red-shouldered Hawk a
- Swainson’s Hawk E, W, c
- Rough-legged Hawk E, W, r
- Ferruginous Hawk E, W, u
- Golden Eagle, E, W, A, c
- Bald Eagle, E, W, ab
- Marsh Hawk E, W, ab
- Osprey E, W, ab
- Prairie Falcon E, W, A, r
- Peregrine Falcon E, W, r
- American Kestrel E, W, c
-
- [Illustration: Sharp-tailed Grouse]
-
- Grouse, Ptarmigans
- Blue Grouse, E, W, ab
- Spruce Grouse E, W, ab
- Ruffed Grouse E, W, ab
- Sharp-tailed Grouse E, r
- White-tailed Ptarmigan A, c
- Willow Ptarmigan ?
- Ring-necked Pheasant E, W, r, i
- Gray Partridge E, W, r, i
-
- Cranes
- Sandhill Crane E, r
-
- [Illustration: American Coot]
-
- Rails, Coots
- Sora E, W, r
- American Coot E, W, ab
-
- [Illustration: Greater Yellowlegs]
-
- Shorebirds
- Killdeer E, W, c
- Black-bellied Plover E, r
- Common Snipe E, W, c
- Long-billed Curlew E, r
- Upland Sandpiper E, r
- Spotted Sandpiper E, W, A, ab
- Solitary Sandpiper E, r
- Willet, E, r
- Pectoral Sandpiper E, r
- Baird’s Sandpiper E, W, r
- Lesser Yellowlegs, E, W r
- Greater Yellowlegs E, W, r
- American Avocet E, W, u
- Northern Phalarope E, W, r
- Wilson’s Phalarope E, W, u
- Black Turnstone ?
- Long-billed Dowitcher E, W, r
-
- [Illustration: Herring Gull]
-
- Gulls, Terns
- Herring Gull E, W, r
- California Gull E, W, ab
- Ring-billed Gull E, W, c
- Franklin’s Gull E, W, c
- Bonaparte’s Gull E, u
- Forster’s Tern E, W, u
- Common Tern E, r
- Caspian Tern a
- Black Tern E, W, u
-
- [Illustration: Mourning Dove]
-
- Doves, Pigeons
- Band-tailed Pigeon E, W, r
- Mourning Dove E, W, c
- Rock Dove E, W, r, i
-
- [Illustration: Great Horned Owl]
-
- Owls
- Screech Owl E, W, r
- Great Horned Owl E, W, ab
- Snowy Owl E, W, u
- Hawk Owl E, W, u
- Pygmy Owl E, W, ab
- Barred Owl E, W, c
- Great Gray Owl E, W, u
- Long-eared Owl E, W, r
- Short-eared Owl, E, W, c
- Boreal Owl E, W, r
- Saw-whet Owl E, W, u
-
- [Illustration: Common Nighthawk]
-
- Nighthawks, Swifts
- Common Nighthawk E, W, ab
- Black Swift E, W, u
- Vaux’s Swift E, W, ab
- White-throated Swift W, A, r
-
- Hummingbirds
- Broad-tailed Hummingbird E, W, r
- Rufous Hummingbird E, W, A, ab
- Calliope Hummingbird E, W, A, ab
- Black-chinned Hummingbird E, W, r
-
- [Illustration: Belted Kingfisher]
-
- Kingfishers
- Belted Kingfisher E, W, ab
-
- Woodpeckers
- Common Flicker E, W, ab
- Pileated Woodpecker E, W, ab
- Red-headed Woodpecker E, W, r
- Lewis’ Woodpecker E, W, c
- Yellow-bellied Sapsucker E, W, ab
- Williamson’s Sapsucker E, W, u
- Hairy Woodpecker E, W, ab
- Downy Woodpecker E, W, ab
- Black-backed Three-toed Woodpecker E, W, ab
- Northern Three-toed Woodpecker E, W, ab
-
- [Illustration: Ash-throated Flycatcher]
-
- Flycatchers
- Eastern Kingbird E, W, ab
- Western Kingbird E, W, u
- Ash-throated Flycatcher a
- Say’s Phoebe E, W, r
- Willow Flycatcher E, W, c
- Hammond’s Flycatcher E, W, ab
- Olive-sided Flycatcher E, W, ab
- Western Flycatcher E, r
- Western Wood Peewee E, W, c
-
- Larks
- Horned Lark E, W, A, ab
-
- [Illustration: Barn Swallow]
-
- Swallows
- Violet-green Swallow E, W, A, ab
- Tree Swallow E, W, ab
- Bank Swallow E, W, ab
- Rough-winged Swallow E, W, u
- Barn Swallow E, W, u
- Cliff Swallow E, W, A, ab
-
- [Illustration: Common Crow]
-
- Jays, Magpies, Crows
- Gray Jay E, W, ab
- Blue Jay E, W, r
- Steller’s Jay E, W, ab
- Black-billed Magpie E, W, ab
- Common Raven E, W, A, ab
- Common Crow E, W, ab
- Clark’s Nutcracker E, W, A, ab
-
- Chickadees
- Black-capped Chickadee E, W, ab
- Mountain Chickadee E, W, ab
- Boreal Chickadee E, W, r
- Chestnut-backed Chickadee E, W, u
-
- Nuthatches, Creepers
- White-breasted Nuthatch E, W, u
- Red-breasted Nuthatch E, W, ab
- Brown Creeper E, W, ab
-
- [Illustration: Winter Wren]
-
- Dippers, Wrens
- Dipper E, W, A, ab
- House Wren E, W, u
- Winter Wren E, W, ab
- Long-billed Marsh Wren a
- Rock Wren E, W, u
-
- Catbirds, Thrashers
- Gray Catbird E, W, u
-
- [Illustration: Mountain Bluebird]
-
- Thrushes, Bluebirds, Solitaires
- American Robin E, W, A, ab
- Varied Thrush E, W, ab
- Hermit Thrush E, W, ab
- Swainson’s Thrush E, W, ab
- Veery E, W, c
- Western Bluebird E, W, r
- Mountain Bluebird E, W, A, ab
- Townsend’s Solitaire E, W, A, ab
-
- Kinglets
- Golden-crowned Kinglet E, W, ab
- Ruby-crowned Kinglet E, W, ab
-
- Pipits
- Water Pipit E, W, A, ab
-
- [Illustration: Cedar Waxwing]
-
- Waxwings
- Bohemian Waxwing E, W, ab
- Cedar Waxwing E, W, ab
-
- Shrikes
- Loggerhead Shrike E, W, r
- Northern Shrike E, W, r
-
- [Illustration: Starling]
-
- Starlings
- Starling E, W, c, i
-
- [Illustration: Red-eyed Vireo]
-
- Vireos
- Solitary Vireo E, W, ab
- Red-eyed Vireo E, W, ab
- Warbling Vireo E, W, ab
-
- Warblers
- Black and White Warbler W, r
- Tennessee Warbler E, W, r
- Orange-crowned Warbler E, W, r
- Nashville Warbler E, W, r
- Yellow Warbler E, W, ab
- Yellow-rumped Warbler E, W, ab
- Townsend’s Warbler E, W, ab
- Northern Waterthrush E, W, ab
- MacGillivray’s Warbler E, W, ab
- Common Yellowthroat E, W, ab
- Wilson’s Warbler E, W, ab
- American Redstart E, W, ab
- Yellow-breasted Chat ?
-
- [Illustration: House Sparrow]
-
- Weaver Finches
- House Sparrow E, W, r, i
-
- Blackbirds, Orioles
- Bobolink E, r
- Western Meadowlark E, W, u
- Red-winged Blackbird E, W, ab
- Northern Oriole E, W, r
- Brewer’s Blackbird E, W, u
- Rusty Blackbird E, W, r
- Yellow-headed Blackbird E, r
- Common Grackle E, r
- Brown-headed Cowbird E, W, c
-
- [Illustration: Evening Grosbeak]
-
- Tanagers, Grosbeaks
- Western Tanager E, W, ab
- Evening Grosbeak E, W, ab
- Pine Grosbeak E, W, ab
- Black-headed Grosbeak E, W, r
-
- [Illustration: American Goldfinch]
-
- Finches, Sparrows, Buntings
- Lazuli Bunting E, W, c
- Lark Bunting E, W, r
- Snow Bunting E, W, c
- Cassin’s Finch E, W, A, ab
- Gray-crowned Rosy Finch E, W, A, ab
- American Goldfinch E, W, u
- Common Redpoll E, W, c
- Pine Siskin E, W, A, ab
- Red Crossbill E, W, ab
- White-winged Crossbill E, W, u
- Rufous-sided Towhee E, W, u
- Green-tailed Towhee E, W, r
- Savannah Sparrow E, W, c
- LeConte’s Sparrow E, W, u
- Vesper Sparrow E, W, ab
- Tree Sparrow E, W, r
- Chipping Sparrow E, W, A, ab
- Brewer’s Sparrow E, W, r
- Harris’ Sparrow E, W, r
- White-crowned Sparrow E, W, A, ab
- Fox Sparrow E, W, A, ab
- Lincoln’s Sparrow E, W, A, c
- Song Sparrow E, W, ab
- Dark-eyed Junco E, W, c
- McCown’s Longspur E, c
- Lapland Longspur E, W, c
- Chestnut-collared Longspur E, c
-
-
-
-
- Suggested Reading
-
-
-Alexander, Taylor R. and George S. Fichter, _Ecology_ (a Golden guide).
- Western Publishing Co., Inc., Racine, Wis. 1973.
-
-Alt, David D. and Donald W. Hyndman, _Rocks, Ice and Water, the Geology
- of Waterton-Glacier Park_. Mountain Press Publishing Co.,
- Missoula, Mont. 1973.
-
-Baker, William, et. al., _Wildlife of the Northern Rocky Mountains_.
- Naturegraph Co., Healdsburg, Calif. 1961.
-
-Borland, Hal, _The History of Wildlife in America_. National Wildlife
- Federation, Washington, D.C. 1975.
-
-Brooks, Maurice, _The Life of The Mountains_. McGraw-Hill, New York.
- 1967.
-
-Costello, David F., _The Mountain World_. Thomas Y. Crowell Co., New
- York. 1975.
-
-Craighead, John J., et. al., _A Field Guide to Rocky Mountain
- Wildflowers_. Houghton Mifflin Co., Boston. 1963.
-
-Dobie, J. Frank, _The Voice of the Coyote_. Little, Brown and Co.,
- Boston. 1950.
-
-Farb, Peter, _Face of North America_. Harper and Row, New York. 1963.
-
-Gildart, Robert C., _Meet the Mammals of Waterton-Glacier_. Glacier
- Natural History Association, Inc. Thomas Printing, Inc.,
- Kalispell, Mont. 1975.
-
-McCormick, Jack, _The Life of the Forest_. McGraw-Hill, New York. 1966.
-
-Milne, Lorus and Margery Milne, _The Balance of Nature_. Alfred A.
- Knopf, Inc., New York. 1960.
-
-Nelson, Alan G., _Wildflowers of Glacier National Park_. Nelson, Great
- Falls, Mont. 1970.
-
-Peattie, Donald Culross, _A Natural History of Western Trees_. Bonanza
- Books, New York. 1953.
-
-Ruhle, George C., _Roads and Trails of Waterton-Glacier Parks_. John W.
- Forney, Minneapolis, Minn. 1972.
-
-Shea, David S., _Animal Tracks of Glacier National Park_. Special
- Bulletin No. 11, Glacier Natural History Association, Inc., West
- Glacier, Mont., 1969.
-
-Storer, John H., _The Web of Life_. Devin-Adair Co., Old Greenwich,
- Conn. 1953.
-
-Zwinger, Ann H. and Beatrice E. Willard, _Land Above the Trees_. Harper
- and Row, New York. 1972.
-
- [Illustration: WATERTON LAKES NATIONAL PARK—GLACIER NATIONAL PARK]
-
- Using Metrics
-
- As we go to press with this book, the United States is in the early
- stages of conversion to the metric system of measurement, and though
- we urge you to think metric—for most of the world does—we provide
- this table to help you understand the measurements given in the
- book.
-
- To convert from to multiply by
-
- Millimeters Sixteenth-inches 0.6301
- Centimeters Inches 0.3937
- Meters Feet 3.2808
- Kilometers Miles 0.6214
- Hectares Acres 2.4711
- Hectares Square miles 0.00386
- Grams Troy Ounces 0.0322
- Kilograms Pounds 2.2046
- Degrees—Celsius Degrees—Fahrenheit 1.8, and add 32
-
- [Illustration: Temperature Conversion Chart]
-
- [Illustration: Length Conversion Chart]
-
- Drawings from David S. Shea, _Animal Tracks of Glacier National
- Park_
-
- [Illustration: red fox,
- hind foot, in mud
- 53 mm.]
-
- [Illustration: mule deer,
- adult buck, in snow
- 72 mm.]
-
- [Illustration: badger,
- left front foot, in mud
- 43 mm.]
-
- [Illustration: coyote,
- hind foot, in snow
- 63 mm.]
-
- About the Author
-
- Greg Beaumont’s interest in Glacier National Park dates from 1963,
- when he was a summer employee at Lake McDonald Lodge. In 1966 he and
- his wife were fire-control lookouts on Numa Ridge in the Bowman
- Valley. Now a free-lance writer-photographer, he lives with his
- family in Lincoln, Nebraska.
-
- National Park Service
- U.S. Department of the Interior
-
- As the Nation’s principal conservation agency, the Department of the
- Interior has responsibility for most of our nationally owned public
- lands and natural resources. This includes fostering the wisest use
- of our land and water resources, protecting our fish and wildlife,
- preserving the environmental and cultural values of our national
- parks and historical places, and providing for the enjoyment of life
- through outdoor recreation. The Department assesses our energy and
- mineral resources and works to assure that their development is in
- the best interests of all our people. The Department also has a
- major responsibility for American Indian reservation communities and
- for people who live in Island Territories under U.S. administration.
-
-
-
-
- Transcriber’s Notes
-
-
-—Retained publication information from the printed edition: this eBook
- is public-domain in the country of publication.
-
-—Corrected a few palpable typos.
-
-—Included a transcription of the text within some images.
-
-—In the text versions only, text in italics is delimited by
- _underscores_.
-
-
-
-
-
-
-
-End of the Project Gutenberg EBook of Many-Storied Mountains, by Greg Beaumont
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