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diff --git a/43928-0.txt b/43928-0.txt new file mode 100644 index 0000000..e184956 --- /dev/null +++ b/43928-0.txt @@ -0,0 +1,3707 @@ +*** START OF THE PROJECT GUTENBERG EBOOK 43928 *** + +Transcriber's Note: Italic text is denoted by _underscores_ and bold +text by =equal signs=. + + + + +THE BRITISH WOODLICE. + + + + +This monograph first appeared in the "Essex Naturalist" (Volume XIV., +1905-6) and has been republished by special arrangement with the +Council of the Essex Field Club. + + + + +THE BRITISH WOODLICE. + +BEING + +A MONOGRAPH OF THE TERRESTRIAL ISOPOD CRUSTACEA + +OCCURRING IN THE BRITISH ISLANDS. + +BY + +WILFRED MARK WEBB, F.L.S., + +_Lecturer on Biology and Nature Study to the Surrey County Council, +Honorary Secretary of the Selborne Society, Sometime, Senior Assistant +Lecturer on Biology to the Essex County Council, and Editor of the +Journal of Malacology, Joint Author of_ "Eton Nature Study and +Observational Lessons." + +AND + +CHARLES SILLEM. + +WITH TWENTY-FIVE PLATES AND FIFTY-NINE FIGURES IN THE TEXT. + + LONDON: + DUCKWORTH & CO., + 3, HENRIETTA STREET, COVENT GARDEN. + 1906. + + + + +PREFACE. + + +In Professor Sars' "Crustacea of Norway," quite a number of the British +species of woodlice are figured in detail and described in English, but +few copies of this fine work are to be met with in our country. The +Rev. Canon Norman has from time to time published notes on the British +species in "The Annals and Magazine of Natural History;" these are, +however, scattered, and contain but few figures, while other literature +that exists is out of date. Under these circumstances, we have thought +that the following account and figures of all the British species would +be useful to those anxious to work at the woodlice, and might also +encourage others to pay attention to the distribution and habits of the +interesting tribe to which they belong. + +The writers would welcome any corrections or additions in view of a +second edition. + + W.M.W. + C.S. + +ODSTOCK, HANWELL, _December, 1905_. + + + + +CONTENTS. + + + PAGE + + Introduction 1 + Geological history 1 + External structure and appendages 2 + Alimentary canal 6 + Circulatory system 7 + Excretory system 7 + Nervous system 8 + Reproductive organs 8 + Development 9 + Habits and Economic considerations 12 + Local names 15 + Methods of collections and preservation 16 + Classification 17 + Scheme of classification and synopsis of generic characters 18 + British Species 19 + Section--Ligiæ 19 + Family--Ligiidæ 19 + Genus--Ligia Fabricius 19 + _Ligia oceanica_ Linzé 19 + Genus--Ligidium Brandt 21 + _Ligidium hypnorum_ Cuvier 21 + Family--Trichoniscidæ 22 + Genus--Trichoniscus Brandt 22 + _Trichoniscus pusillus_ Brandt 22 + _Trichoniscus vividus_ Koch 23 + _Trichoniscus roseus_ Koch 24 + Genus--Trichoniscoides, Sars 25 + _Trichoniscoides albidus_ Budde-Lund 25 + Genus--Haplophthalmus Schobl 26 + _Haplophthalmus mengii_ Zaddach 26 + _Haplophthalmus danicus_ Budde-Lund 27 + Family--Oniscidæ 27 + Genus--Oniscus Linné 27 + _Oniscus asellus_ Linné 27 + Genus--Philoscia Latreille 29 + _Philoscia muscorum_ Scopoli 29 + _Philoscia couchii_ Kinahan 30 + Genus--Platyarthrus Brandt 30 + _Platyarthrus hoffmannseggii_ Brandt 30 + Genus--Porcellio Latreille 32 + _Porcellio scaber_ Latreille 32 + _Porcellio pictus_ Brandt and Ratzeburg 33 + _Porcellio dilatatus_ Brandt 33 + _Porcellio rathkei_ Brandt 34 + _Porcellio laevis_ Latreille 35 + _Porcellio ratzeburgii_ Brandt 36 + Genus--Metoponorthus Budde-Lund 37 + _Metoponorthus pruinosus_ Brandt 37 + _Metoponorthus cingendus_ Kinahan 38 + Genus--Cylisticus Schnitzler 38 + _Cylisticus convexus_ De Geer 39 + Family--Armadillidiidæ 40 + Genus--Armadillidium Brandt 40 + _Armadillidium nasatum_ Budde-Lund 40 + _Armadillidium vulgare_ Latreille 41 + _Armadillidium pulchellum_ Zencker 42 + _Armadillidium depressum_ Brandt 43 + Distribution of species 43 + Conclusion 43 + Bibliography 49 + + + + +LIST OF FIGURES. + + +PLATES I.-XXV. with a List, will be found at the end of the Book. + + +FIGURE. PAGE. + + 1. Parts of the body (_Oniscus asellus_) 2 + 2. The first antenna (_Oniscus asellus_) 2 + 3. The second antenna (_Oniscus asellus_) 3 + 4. The underside of the head (_Oniscus asellus_) 3 + 5. The mandibles (_Oniscus asellus_) 4 + 6. The first maxillae (_Oniscus asellus_) 4 + 7. The second maxillae (_Oniscus asellus_) 4 + 8. The fused maxillipeds (_Oniscus asellus_) 4 + 9. The "upper lip" (_Oniscus asellus_) 5 + 10. The "lower lip" (_Oniscus asellus_) 5 + 11. A typical thoracic segment (_Oniscus asellus_) 5 + 12. The fifth thoracic segment of a female + (_Oniscus asellus_) 5 + 13. The underside of the abdomen of a female + (_Oniscus asellus_) 6 + 14. A typical abdominal appendage + (_Oniscus asellus_) 6 + 15. The first abdominal appendage of the male + (_Oniscus asellus_) 6 + 16. The second abdominal appendage of the male + (_Oniscus asellus_) 6 + 17. The alimentary canal (_Oniscus asellus_) 7 + 18. The circulatory system (_Oniscus asellus_) 7 + 19. The nervous system (_Oniscus asellus_) 8 + 20. Female reproductory organs (_Oniscus asellus_) 8 + 21. The male reproductory organs (_Oniscus asellus_) 9 + 22. The fertilized egg (_Porcellio scaber_) after Roule 10 + 23. The fertilized egg seen in section + (_Porcellio scaber_) after Roule 10 + 24 to 31. The development of a woodlouse + (_Porcellio scaber_) after Roule 10 + 32. Embryo of the woodlouse showing the three divisions of the + intestine separately developed (_Porcellio scaber_) + after Roule 11 + 33. Embryo of the woodlouse showing traces of the segments + (_Porcellio scaber_) after Roule 11 + 34. An embryo woodlouse ready to be hatched + (_Porcellio scaber_) after Roule 11 + 35. Flagellum and last peduncular joint of the antenna of + _Ligia oceanica_ 12 + 36. Flagellum and last peduncular joint of the antenna of + _Ligidium hypnorum_ 13 + 37. Flagellum and last peduncular joint of the antenna of + _Trichoniscus pusillus_ 23 + 38. Flagellum and last peduncular joint of the antenna of + _Trichoniscus vividus_ 24 + 39. Flagellum and last peduncular joint of the antenna of + _Trichoniscus roseus_ 24 + 40. Flagellum and last peduncular joint of the antenna of + _Trischoniscoides albidus_ 25 + 41. Flagellum and last peduncular joint of the antenna of + _Haplophthalmus mengii_ 26 + 42. Flagellum and last peduncular joint of the antenna of + _Haplophthalmus danicus_ 27 + 43. Flagellum and last peduncular joint of the antenna of + _Oniscus asellus_ 28 + 44. Flagellum and last peduncular joint of the antenna of + _Philoscia muscorum_ 29 + 45. Flagellum and last peduncular joint of the antenna of + _Philoscia couchii_ 30 + 46. Flagellum and last peduncular joint of the antenna of + _Platyarthrus hoffmannseggii_ 31 + 47. Flagellum and last peduncular joint of the antenna of + _Porcellio scaber_ 32 + 48. Flagellum and last peduncular joint of the antenna of + _Porcellio pictus_ 33 + 49. Flagellum and last peduncular joint of the antenna of + _Porcellio dilatatus_ 34 + 50. Flagellum and last peduncular joint of the antenna of + _Porcellio rathkei_ 35 + 51. Flagellum and last peduncular joint of the antenna of + _Porcellio laevis_ 35 + 52. Flagellum and last peduncular joint of the antenna of + _Porcellio ratzeburgii_ 36 + 53. Flagellum and last peduncular joint of the antenna of + _Metoponorthus pruinosus_ 37 + 54. Flagellum and last peduncular joint of the antenna of + _Metoponorthus cingendus_ 38 + 55. Flagellum and last peduncular joint of the antenna of + _Cylisticus convexus_ 39 + 56. Flagellum and last peduncular joint of the antenna of + _Armadillidium nasatum_ 40 + 57. Flagellum and last peduncular joint of the antenna of + _Armadillidium vulgare_ 41 + 58. Flagellum and last peduncular joint of the antenna of + _Armadillidium pulchellum_ 42 + 59. Flagellum and last peduncular joint of the antenna of + _Armadillidium depressum_ 43 + + + + +THE BRITISH WOODLICE. + + +=Introduction.=--Having finished a somewhat exhaustive list of the land +and fresh-water molluscs of Essex,[1] one of the present writers felt +that if he were to make any further contributions of importance to a +knowledge of the fauna of that interesting county, he must turn his +attention to some other group of animals. It seemed most fitting that +some creatures should be chosen which are commonly met with during the +search for molluscs. Centipedes, millepedes, and woodlice fulfilled +these conditions, and all were collected, but as only seventeen +species of woodlice had at the time been found in England, it was +deemed advisable to study these in detail to begin with. The present +contribution is the result of the undertaking, and we have thought that +a general consideration of the British Woodlice, with careful drawings +from nature of all the species now known from this country, ought to +lead to a more general study of these interesting creatures and their +habits. + +=Position in the scheme of classification.=--The Woodlice belong to +an immense group of invertebrate animals known as the Arthropoda, the +bodies of which are segmented and provided with jointed appendages for +purposes of walking, swimming, and feeding. Of this group, two large +divisions are recognized. The first contains the forms which breathe +by means of air-tubes, such as the Insects; and the second has been +constituted for Crustacea, which breathe by means of gills. The latter +are, of course, adapted more especially for a life in water, but here +and there we come across examples so modified that they can exist in +air. The land-crabs are a case in point, and so are the Woodlice. These +belong to an order which contains many fresh-water and marine species, +known as the Isopoda. + +=Geological history.=--The known history of the order is a long one, +for remains occur in the Old Red Sandstone (Devonian) of Herefordshire, +and in the Coal Measures. (79)[2]. A form which has been named +_Archæoniscus brodiei_, and is said to be referable to the recent +family Aegidae which is found in some numbers in the Purbeck Beds +(Upper Jurassic), of this country (47). Fossil Isopods have also been +recorded from the Oolite and from the Oligocene (Isle of Wight). + +Turning to the Woodlice proper, we find that they first make their +appearance in the Miocene (of Oenigen and Baden), and occur also +in amber (79); while examples of genera, such as _Oniscus_ and +_Porcellio_, have been discovered in late Tertiary deposits (47). + +=External structure and appendages.=--Woodlice agree in being of a +somewhat oval form, and their bodies are arched, the curve varying in +different genera and species. A _head_ is to be distinguished; behind +this comes the _thorax_ of seven segments which are often considerably +broader than the six succeeding ones which form the _abdomen_ (see fig. +1.) + +[Illustration: FIG. 1.--PARTS OF THE BODY. (_Oniscus asellus._)] + +The head carries two _large antennae_ (fig. 3) which are very evident, +and a careful search with a lens will reveal a second and minute pair +(the _smaller antennae_) situated between the base of the others, and +really anterior to them. (figs. 2 and 4.) + +[Illustration: FIG. 2.--THE FIRST ANTENNA. (_Oniscus asellus._)] + +The larger antennae are customarily bent at certain points, and we can +distinguish a terminal part, or _flagellum_, and a basal part, the +_peduncle_ (fig. 3). The number of joints in these structures, which +varies in different genera and species, forms a useful classificatory +character, and the relative length of the component parts is of +considerable value in distinguishing species. + +[Illustration: FIG. 3.--THE SECOND ANTENNA. (_Oniscus asellus._)] + +There are four pairs of mouth appendages--namely the jaws or +_mandibles_ (fig. 5), the _first maxillae_ (fig. 6), the _second +maxilla_ (fig. 7), and the _maxillipeds_ (fig. 8). When the head is +examined from the underside the last of these organs will be seen +first, covering in the others. + +[Illustration: FIG. 4.--THE UNDERSIDE OF THE HEAD. (_Oniscus +asellus._)] + +A small median plate attached to the front of the head has been called +"_the upper lip_" (fig. 9), while inside the mouth appendages is a +little bilobed structure "_the lower lip_" (fig. 10). + +[Illustration: FIG. 5.--THE MANDIBLES. (_Oniscus asellus._)] + +Before leaving the external features of the head, we must allude to +the pair of _eyes_ which are usually present, though never raised on +stalks. In the Common Woodlouse (_Oniscus asellus_, from which all +our figures to illustrate structure have been made), as in many other +species, the eyes are compound (fig. 4), but in some forms these are +simple. + +[Illustration: FIG. 6.--THE FIRST MAXILLAE. (_Oniscus asellus._)] + +[Illustration: FIG. 7.--THE SECOND MAXILLAE. (_Oniscus asellus._)] + +[Illustration: FIG. 8.--THE FUSED MAXILLIPEDS. (_Oniscus asellus._)] + +Each of the seven joints of the thorax bears a pair of _walking legs_ +(fig. 11), and in the female at the time when the eggs are laid, a pair +of plates (fig. 12) arises on segments II. to V. These plates together +form a brood pouch, in which the eggs are carried (fig. 12) until +they are hatched, and in which the young ones remain for some time +afterwards. + +When we examine the abdomen, we find that the appendages are +plate-like, with the exception of the last pair (fig. 13), and they all +agree in having two divisions, an arrangement which would prove awkward +in limbs used for walking or feeling. + +[Illustration: FIG. 9.--THE "UPPER LIP." (_Oniscus asellus._)] + +[Illustration: FIG. 10.--THE "LOWER LIP." (_Oniscus asellus._)] + +[Illustration: FIG. 11.--A TYPICAL THORACIC SEGMENT. (_Oniscus +asellus._)] + +[Illustration: FIG. 12.--THE FIFTH THORACIC SEGMENT OF A FEMALE. +(_Oniscus asellus._)] + +The inner plate (or endopodite) is in structure a _gill_, but the blood +that passes through it, is enabled to take up oxygen from moist air, +while the outer division (or exopodite) acts as a protecting cover +(fig. 14). In _Porcellio_, air-tubes (_tracheae_) may be present (see +below). + +[Illustration: FIG. 13.--THE UNDERSIDE OF THE ABDOMEN OF A FEMALE. +(_Oniscus asellus._)] + +In the male, the first two pairs of abdominal appendages are specially +modified, the inner divisions (endopodites) being long and pointed +(figs. 15 and 16). The last pair, or tail appendages, in the male are +often considerably larger than in the female, and the form of these +structures is sometimes of value in classification. + +[Illustration: FIG. 14.--A TYPICAL ABDOMINAL APPENDAGE. (_Oniscus +asellus._)] + +[Illustration: FIG. 15.--THE FIRST ABDOMINAL APPENDAGE OF THE MALE. +(_Oniscus asellus._)] + +[Illustration: FIG. 16.--THE SECOND ABDOMINAL APPENDAGE OF THE MALE. +(_Oniscus asellus._)] + +=Alimentary canal.=--The main portion of the alimentary system is, +practically speaking, a straight tube (fig. 17). Its first part (not +shown in the figure) is a narrow gullet, which after passing through +the nerve collar dilates to form a sort of stomach. Into this the +secretion of four digestive glands is poured by two ducts. These glands +have a somewhat striking appearance, being yellow tubes spirally +coiled, and they end blindly. From the stomach the intestine runs to +the hinder end of the body and passes under the heart. + +=Circulatory system.=--The blood being aërated in the abdominal +appendages, we find that the heart is situated towards the hinder end +of the body (fig. 18). Three main arteries supply the thorax and head, +while the blood is brought from the gills to the heart. + +=Excretory system.=--The excretory organs consist of a (_a_) pair of +so-called "shell glands," which are considered to be the equivalents +of the excretory tubes or nephridia of annelid worms. In the woodlouse +these excretory organs open on the second pair of maxillae. They are +composed of a tube (_sacculus_) closed at one end and more or less bent +upon itself (5, p. 261) which communicates with a _labyrinth_ that +is provided with an excretory orifice. Matters are eliminated by the +_epithelial cells_ [the histology has been described and figured in +_Ligidium hypnorum_ (66)], which are very large in _Ligia oceanica_. + +[Illustration: Fig. 17.--THE ALIMENTARY CANAL. (_Oniscus asellus._)] + +[Illustration: Fig. 18.--THE CIRCULATORY SYSTEM. (_Oniscus asellus._)] + +(_b_) Masses of cellules in the head, very greatly developed in _Ligia +oceanica_ (but numbering scarcely more than ten in _Oniscus asellus_), +which have no external opening. They also function as excretory organs +(5, p. 263), and have been called "cephalic nephrocytes." + +[Illustration: FIG. 19.--THE NERVOUS SYSTEM. (_Oniscus asellus._)] + +(_c_) Other "branchial nephrocytes" are situated on the dorsal surface +between the last thoracic and the first abdominal segments, as well as +between those that follow, with the exception of the last two; they are +in distinct patches, one on each of the middle line in _Ligia_, but +more or less continuous in _Oniscus_ (5, p. 265). + +(_d_) The digestive glands have also been shown to be excretory (5, p. +270). + +=Nervous system.=--The nervous system consists of _paired ganglia_ +in the head, above the alimentary canal which send off nerves +(_commissures_) that meet below, to form a _double nerve cord_ with +ganglia at intervals (see fig. 19). + +[Illustration: FIG. 20.--FEMALE REPRODUCTORY ORGANS. (_Oniscus +asellus._)] + +=Reproductive organs.=--In the female there are a pair of _ovaries_ in +the positions shewn in fig. 20; and _ducts_ run to the underside of the +fifth thoracic segment. + +The openings are very difficult to identify, and Lereboullet (39, p. +113) was unable to find them. It is obvious that the openings must be +underneath the plates that form the egg pouch, and as a change of skin +is required to set these free, it would appear that at ordinary seasons +the ducts from the ovaries are closed. The writers have been able to +determine from external examination of specimens which had moulted and +were about to lay eggs, that the oviducts at such time open to the +inside of the base of each walking leg on the fifth segment. In similar +specimens the oviducts were also followed to the opening from within. +The _brood pouch_ has already been described. + +The male organs consist of six _testes_ arranged in two pairs, each +of which is provided with a _reservoir_ (see fig. 21). The efferent +ducts from the two reservoirs unite at the base of the thorax to form a +common duct (or "penis"). + +[Illustration: FIG. 21.--THE MALE REPRODUCTORY ORGANS. (_Oniscus +asellus._)] + +=Development.=--The eggs, in the common species of woodlice, at +least, are laid at the beginning of summer, and are retained in the +brood pouch, where they undergo their development. The process has +been recently traced with great care by Professor Louis Roule (58) in +_Porcellio scaber_ and the description which follows is based upon his +researches. + +As, practically speaking, the larval stages are passed within the egg, +and there is no free embryo differing in form from the parent, it is +necessary for the young creatures to be well supplied with nutritive +material. In fact, the bulk of the large egg is made up of _food-yolk_, +on the outside of which the _formative protoplasm_ is disposed in +irregular patches. In the fertilized ovum, one of the latter, which +lies in a particular position at the end, is found to be larger than +the others (see fig. 22). It contains the nucleus of the egg-cell (see +fig. 23) and is called the _cicatricula_. This is the only portion of +the egg which divides and produces _nucleated cells_. It is these which +gradually spread all over the surface of the food-yolk, forming a layer +known as the _blastoderm_, which is at first but one cell thick (see +figs. 24, 26, and 28). + +Before, however, the food-yolk is quite closed in, a differentiation +into two layers--the _pro-ectoderm_ and _pro-endoderm_--takes place +(see fig. 25) and rudiments of the first two pairs of _appendages_ +appear (see fig. 26). Moreover, the cells of the ectoderm change their +shape and begin to multiply at two points to form the beginnings of the +cerebral ganglia and the nerve cord respectively. + +[Illustration: FIG. 22.--THE FERTILIZED EGG (_Porcellio scaber_), AFTER +ROULE.] + +[Illustration: FIG. 23.--THE FERTILIZED EGG SEEN IN SECTION (_Porcellio +scaber_), AFTER ROULE.] + +As the blastoderm closes over the food-yolk, two more appendages arise +and these are soon followed by others (see fig. 28). A depression +appears at the point where the blastoderm closed and internally the +pro-endoderm or inner layer is differentiated into two--the _endoderm +proper_ and the _mesoderm_ (see fig. 29). The former begins to grow +so that its edges unite to form the middle part of the intestine (see +fig. 29) seen from the outside in fig. 30. The depression already +mentioned grows deeper, forming a tube which is the hind portion of the +_intestine_, while at the anterior end of the embryo the front part of +the intestine is similarly formed (see fig. 30). By this time also all +the nineteen appendages have made their appearance and the mesoderm, +(which has grown considerably, to form the beginnings of the muscles) +has sent prolongations into each of them. About this time, spaces (see +fig. 31) are formed in the muscular mesoderm which are all that remain +of the _true body cavity_ characteristic of animals above the level of +the jelly fish, and in these spaces the blood ultimately circulates. + +_SURFACE VIEWS._ + +[Illustration: FIG. 24.] + +[Illustration: FIG. 26.] + +[Illustration: FIG. 28.] + +[Illustration: FIG. 30.] + +_OPTICAL SECTIONS._ + +[Illustration: FIG. 25.] + +[Illustration: FIG. 27.] + +[Illustration: FIG. 29.] + +[Illustration: FIG. 31.] + +THE DEVELOPMENT OF A WOODLOUSE (_Porcellio scaber_), AFTER ROULE. + +Figs. 24, 26, 28, 30, are Surface Views, and figs. 25, 27, 29, 31, +which indicate slightly later stages respectively than the others, are +of egg seen in Optical Section. + +The body next alters somewhat in shape and the three divisions of +the intestine approach one another (see fig. 32) previous to their +junction. As may be imagined during these processes the food-yolk has +gradually been used up and the space which it occupied taken by the +internal organs, which we have mentioned. + +[Illustration: FIG. 32.--EMBRYO OF THE WOODLOUSE SHOWING THE THREE +DIVISIONS OF THE INTESTINE SEPARATELY DEVELOPED (_Porcellio scaber_), +AFTER ROULE.] + +[Illustration: FIG. 33.--EMBRYO OF THE WOODLOUSE SHOWING TRACES OF THE +SEGMENTS (_Porcellio scaber_), AFTER ROULE.] + +[Illustration: FIG. 34.--AN EMBRYO WOODLOUSE READY TO BE HATCHED +(_Porcellio scaber_), AFTER ROULE.] + +In the last stages of the development the appendages become larger +still, the _heart_ makes its appearance, segmentation of the body is +completed, and except that the seventh pair of walking legs are as yet +rudimentary the woodlouse is completed. It is only after hatching that +the pair of legs mentioned, attain to their normal length. + +The process of segmentation of the egg and the formation of its layers +lasts about a fortnight, while the completion of the development +proceeds much more rapidly, for another three weeks bring it to an end. + +After the first moult or change of skin the last pair of walking +legs makes its appearance, and Mr. James B. Casserley [whose work +one of us (75) has described elsewhere] found when keeping a number +of the common pill-woodlouse (_Armadillidium vulgare_) in captivity +that his specimens did not subsequently change their skins more than +once in the six months during which he had them under observation. He +also noted that the crustaceans go on growing after they are sexually +mature. As his specimens grew older, Mr. Casserley noticed that their +colour became darker, and a curious point recorded by him is that two +examples of the same age may change their skins at the same time, +and while one may have afterwards nearly twice as many markings, on +the other very few at all will be seen. The time required for the +growth of a woodlouse from the size of a pin's head to that of an +adult example--say three-quarters-of-an-inch long--must be fairly +considerable, taking into account the fact that any appreciable +increase in size can only occur at a moult and Mr. Casserley's +observations as to the infrequency of the process in _Armadillidium +vulgare_. (See p. 14.) + +=Habits and Economic Considerations.=--The construction of the +breathing organs of woodlice, and the necessity which exists for these +to be kept moist, restricts the habitats of the animals considerably. +Woodlice are found under stones and logs, beneath the bark of dead and +rotten trees, among decaying vegetable matter as well as living grass +and moss in damp or wet situations. When looking for some of the common +species under the bark of fallen trees it is surprising to notice that +the crustaceans may be entirely absent from many trunks, while when +another is examined which seems to differ very slightly, if at all, in +condition or situation, they are found in swarms. There is no doubt +but that the habits of woodlice would well repay the attention of +naturalists, who are now recognizing that besides anatomy as such, and +the classification which a knowledge of structure permits, there is the +equally important consideration of the creatures as they live their own +life and affect that of others. It is not our object to give a detailed +account of the ecology of British woodlice, but rather to provide a +basis from which it may be approached. Nevertheless a few general +remarks may not come amiss. Many points in the life-history of woodlice +may no doubt be learned by keeping them in captivity and there is just +sufficient difficulty in doing this successfully to give an interest to +the matter. + +Apart from a supply of proper food, we take it that the chief object to +be attained is the provision of the amount of moisture required by the +particular species under examination, together with a sufficient supply +of air. + +A great many interesting observations can be thus carried out, such +as those of Mr. Casserley, to which allusion has already been made. +The process of moulting for instance is well worth watching, and +although specimens with half their coat changed may be found in remote +corners, yet the whole course of the moult can be seen much better +in the case of captive woodlice. The following account is taken from +Mr. Casserley's description (75) of what happens in the case of +_Armadillidium vulgare_:--The approach of the moult is indicated by +the appearance of a white border on each segment of the body, which +becomes gradually more marked, while at the same time the animal is +seen to be less active and often makes a small burrow in which to +hide. Sometimes a sheltered corner against a stone is looked upon as +affording sufficient protection, but in either case each woodlouse +keeps to the place originally chosen. About ten days after the white +lines have become visible the animal appears to be divided into two. +Its skin is becoming loose and little movement can take place at the +joints of its body with the exception of that between the fourth and +fifth thoracic segments where the skin will ultimately break. The +woodlouse spends a day or two in this condition and then, by suddenly +walking forward, frees itself from the covering of the hinder portion +of its body. The three last pairs of walking legs are carefully pulled +out from the old skin, which now appears perfectly white, and at the +same time the lining of the hind portion of the alimentary canal (hind +gut) is also shed. After putting the tender half of his body well into +his corner or burrow the woodlouse proceeds to eat the part of his skin +that he has cast. The creature has now a very odd appearance. His front +half with the exception of the white edges is as it was before, the +rest of him instead of a light slaty blue, and is very soft as well as +proportionately a little larger. + +In three days or so the tail end becomes hard and attains the normal +colour. Then the old skin from the front half is pushed off and the +creature becomes practically defenceless, so much so in fact, that any +of his species that happen to find him will attack him and eat all his +front half, rejecting, however, his now hardened tail-end. + +Provided that the moulting woodlouse has survived (and in captivity, to +ensure this, he must be isolated), after three days his jaws will be +sufficiently hardened to allow of his eating, and usually he first of +all devours the second half of his cast skin. The operation of moulting +does not occupy quite so long a time in the case of young examples. +Specimens half-an-inch long do not moult more than once in six months +and show but little increase in size after the process. + +Woodlice do not appear to live on either animal or vegetable food +alone, but adopt a mixed diet. It is, however, owing to their attacks +upon cultivated plants that the creatures are looked upon as pests by +the horticulturalist. The animals feed either in the night or in the +very early morning, on seedlings, orchid tubers, mushrooms, or anything +that comes to hand. Few of the accounts, however, of their ravages, +mention that the crustaceans have been caught absolutely in the act +of doing the damage ascribed to them. Some careful inquiries have +nevertheless enabled us to discover several observers who have watched +woodlice feeding. Mr. F. V. Theobald, of Wye College, and one of the +students at Swanley Horticultural College are among the number. The +former has also given us an account of the methods, out of many tried, +which he has found most successful for getting rid of the crustaceans. +Out of doors trapping with moss, sacking or horse-dung is best. In +glass houses, fumigation with hydro-cyanic acid gas has cleared them +out, and poison baits, especially potatoes cut and soaked in white +arsenic, have done some good. Stable manure is especially favourable to +these creatures, particularly when it is used "long": in this condition +it should therefore be avoided. + +It is interesting to note how the woodlice in winter simply remain +where they happen to be so long as there is sufficient moisture, though +they are ready to run about as rapidly, for a time, as in summer, +should they happen to be disturbed. + +No doubt many points of inter-relation between woodlice and other +animals remain to be discovered. Mr. John W. Odell tells us that on +Exmoor, in the open, he found no _Armadillidia_, though other forms +occurred under nine out of every ten stones that he turned over, and +here the smaller species of ants also abounded. Close to stone walls +_Armadillidia_ were to be seen to the exclusion of all other genera, +and this state of affairs was ascribed by Mr. Odell to the presence of +swarms of the large wood-ants which he considers would make short work +of any woodlice that could not protect themselves by rolling up. + +We ought not to conclude this account without mentioning the fact that +woodlice once played an important part in medicine. + +Doctor Fernie (28) gives some interesting extracts with regard to the +hog-louse and the woodlouse. The latter he seems to have identified +quite correctly as _Oniscus asellus_. He calls the former, however, +indiscriminately, "the common armadillo" (which is the old name for +the pill-woodlice now known as _Armadillidium_), "the pill millipede" +and "_Glomeris marginata_." The last two names are those of another +creature, not a crustacean, which when it is rolled up can be very +easily mistaken for an _Armadillidium_, though, when it uncurls, it +will be seen to have many more than seven pairs of legs. The local +appellations applied to the hog-louse by Doctor Fernie, and his remarks +with regard to its commonness, tend to show that it is _Armadillidium +vulgare_, to which he really refers, and the use of which in medicine +was commonly general. + +Hog-lice were prescribed for scrofulous diseases and obstructions of +the liver and digestive organs, among other things, and the London +College of Physicians directed that the creatures should be prepared +by suspending them in a thin canvas bag placed within a covered vessel +over the steam of hot spirit or wine, so that being killed by the +spirit they might become friable. Hog-lice and Woodlice were also +administered alive, while the former were also put down the throats of +cows "to promote the restoration" of their cud, hence their name of +"cud-worm." There seems to be considerable evidence that even in modern +times Woodlice have had considerable remedial effect which depends upon +"an alkalescent fluid" contained in them. + +=Local Names.=--Among the local names by which these creatures are +known are those of "sow bug," "lucre pig" (Berkshire), "carpenter" +and "chiselhog" (Berkshire). Doctor Fernie (28) gives a number of +others:--"thrush-louse," "tiggyhog," "cheslip," "kitchenball," +"chiselbob," "lugdor," "palmer," and "cudworm." In the eastern +counties the same writer notes that they are known as "old-sows" or +"St. Anthony's hogs" while the Welsh call them "little grey-hogs," "the +little old women of the wood" or "grammar-sows," grammar signifying +a shrivelled up old dame. _Oniscus asellus_ was sometimes called +"socchetre," "church louse," and "chinch." + +=Methods of Collection and Preservation.=--Woodlice should be collected +straightway into tubes or bottles half filled with 30 per cent. +methylated spirit.[3] Woodlice dropped into this weak spirit become +gradually narcotised and die, and they remain limp enough for purposes +of examination or to allow, of their legs and antennæ being set out +during the process of mounting. Specimens to be kept permanently should +be placed in 70 per cent. alcohol. For storage purposes the specimens +of each species from a given locality should be put together into a +small flat bottomed tube such as is used for pillules by apothecaries +or specially made for natural history purposes. A paper label on +which the name, locality, date of capture and any other necessary +particulars have been written with dark lead pencil, is not affected by +the spirit. The tubes may be corked, though if not frequently examined +all the spirit may evaporate, and cause the specimens to be spoilt. A +safer method is to plug the tubes with cotton wool and keep all those +containing a given species or specimens from a particular locality +beneath the surface of spirit in a large wide-mouthed bottle, into +which first of all some cotton wool has been put to prevent the tubes +from coming into sudden contact with the glass at the bottom. For show +purposes in museums, specimens taken direct from 30 per cent. spirit +should be mounted on slips of opal glass by means of gum-tragacanth +which has been powdered and shaken up in spirit before having water +added to it. The slips can be exhibited in glass tubes, six inches high +by one across, or in narrow stoppered museum jars. A variation of the +method is to mount the animals on clear glass and to place behind them +another strip of any colour that may be preferred. + +=Classification.=--The various genera of woodlice are connected +together so closely, by intermediate forms, that their division into +families is, to a very great extent, arbitrary. Bate and Westwood +described but a single family Oniscidæ (I), though they distinguished +two sub-families:--Ligiinæ, which included the forms with many joints +to the flagellum of the antenna, and Oniscinæ, which contained the rest. + +Since then the pill-woodlice have been thought by some to be +sufficiently different from the other genera to warrant their +separation, and three families namely, Ligiidæ, Oniscidæ, and +Armadillidæ have been recognized, as for instance by Dr. Scharff (63). + +A fourth family--Trichoniscidæ--has been added by Professor G. O. Sars, +who in his _Crustacea of Norway_ (59) alludes to the division of the +tribe into the sections Ligiæ and Onisci and has adopted the following +classification:-- + +_Order_--=ISOPODA.= + +_Tribe_--ONISCOIDA. + + _Family I._--LIGIIDAE. + _Ligia._ + _Ligidium._ + + _Family II._--TRICHONISCIDÆ. + _Trichoniscus._ + _Trichoniscoides._ + _Haplophthalmus._ + + _Family III._--ONISCIDÆ. + _Oniscus._ + _Philoscia._ + _Platyarthrus._ + _Porcellio._ + _Metoponorthus._ + _Cylisticus._ + + _Family IV._--ARMADILLIDIIÆ. + _Armadillidium._ + +All the genera described by Professor Sars are represented in the +British Islands. + + * * * * * + +Below is a scheme of classification and synopsis of the characters of +British genera of woodlice which we have compiled in order to render +easy the determination of the genus to which any particular specimen +may belong. + +_SCHEME OF CLASSIFICATION AND SYNOPSIS OF GENERIC CHARACTERS._ + +_Order_--=ISOPODA.= + +_Tribe_--ONISCOIDA. + + +_Section I._--=LIGIÆ.= + + _The Two Divisions of the Tail Appendages alike in Shape._ + + (A.)--Flagellum with 10 or more joints; tail + appendages wholly visible; head without + lateral lobes LIGIIDAE. + (1.)--Abdomen broad; body large; + habitat, the sea-shore _Ligia._ + (2.)--Abdomen narrow; habitat, wet + moss _Ligidium._ + + (B.)--Flagellum with less than 10 joints; head + with small lateral lobes, tail appendages + partly covered TRICHONISCIDÆ. + (3.)--Abdomen narrow; eyes compound; + flagellum usually with more + than 3 joints _Trichoniscus._ + (4.)--Abdomen narrow; eyes simple or + wanting; flagellum with 4 joints _Trichoniscoides._ + (5.)--Abdomen broad (comparatively); + eyes simple; back with + longitudinal ridges; flagellum + with 3 joints _Haplophthalmus._ + + +_Section II._--=ONISCI.= + + _The Outer Divisions of the Tail Appendages Broader than the Inner._ + + (A.)--Tail appendages projecting when the animal + is walking ONISCIDÆ. + (a.)--Unable to roll up into a complete ball. + (6.)--Flagellum with 3 joints; abdomen + broad; head, with lateral lobes _Oniscus._ + (7.)--Flagellum with 3 joints; abdomen + narrow; head without lateral + lobes _Philoscia._ + (8.)--Flagellum with 1 joint; eyes + wanting; abdomen broad; habitat, + ant's nests _Platyarthrus._ + (9.)--Flagellum with 2 joints; abdomen + broad; frontal lobe projecting _Porcellio_. + (10.)--Flagellum with 2 joints; abdomen + narrow _Metoponorthus._ + (b.)--Able to roll up into a complete ball. + (11.)--Flagellum with 2 joints; antennae + folded together over the thorax + when the animal is rolled up + into a ball _Cylisticus._ + + (B.)--Tail appendages not projecting when the + animal walking ARMADILLIDIIDÆ. + (12.)--Flagellum with 2 joints; antennae + hidden or carried at the sides + of the head when the animal is + rolled up into a ball _Armadillidium._ + +=British Species.=--Naturalists in this country paid little attention +to the recognition or description of Woodlice, until the latter half of +the nineteenth century. + +In 1857 Kinahan read a paper before the British Association (32) in +which he described fourteen species of woodlice from the British +Islands, and eleven years later when Bate and Westwood published +their book (1), the number had risen to seventeen. One of the species +(_Oniscus fossor_), however, was doubtful, and although Dr. Scharff +in 1894 (63) rejected it, his list contained also seventeen species, +for in the meantime the Rev. T. R. R. Stebbing had found _Ligidium +hypnorum_ in Surrey (70). + +Since then the Rev. Canon Norman, Dr. Scharff, the Rev. T. R. R. +Stebbing, and one of the present writers, have added other species, as +will be seen from the following pages, in which all those found, up to +the present time in the British Islands are described and figured. + +We shall now consider in detail the British genera and species of +woodlice and give their synonymy and distribution. + +_Order_--=ISOPODA.= + +_Tribe--ONISCOIDA._ + + +_Section_--=LIGIÆ.= + +THE TWO DIVISIONS OF THE TAIL APPENDAGES ALIKE IN SHAPE. + +_Family_--LIGIIDÆ. + +=Flagellum with ten or more joints; tail appendages wholly visible; +head without lateral lobes.= + +_Genus_--=LIGIA= Fabricius, 1798 (27), p. 301. + +_Abdomen broad; body large; habitat, the sea-shore._ + +The genus _Ligia_ agrees with _Ligidium_ alone, in that the flagellum +of the larger antennæ has more than ten joints. In both genera, there +are no lateral lobes to the head, and the tail appendages are wholly +visible from the upper surface of the body. The latter in _Ligia_ is, +however, very many times bigger than in _Ligidium_ and shows no abrupt +decrease in the width of its segments when the abdomen is reached. + +=Ligia oceanica= Linné (The Quay-louse). PLATE I. + + 1767 _Oniscus oceanicus_ Linné (43), p. 1061. + 1793 _Cymothoa oceanica_ Fabricius (26), p. 509. + 1815 _Ligia scopulorum_ Leach (38), p. 374. + 1868 _Ligia oceanica_ Bate and Westwood (1), p. 444. + 1898 _Ligia oceanica_ Sars (59), 156, pl. LXX. + +There is but one British species of _Ligia_, and this, the largest +member of the whole tribe to be met with in these Islands, usually +attains a length of two centimetres, while adult males may be nearly +half as long again. It is the _Oniscus oceanicus_ of Linnæus and lives +on the sea shore, where it may be found at low tide beneath stones and +rubbish in the crevices of timber. _Ligia_ forms a connecting link +between the woodlice proper and the many Isopods which actually live in +the sea. + +The colour of the animals is a greenish grey, and the compound eyes +are almost black, so that they are very conspicuous; there are from +eleven to fourteen joints to the flagellum of the outer antennae and +this feature, taken in conjunction with the large size and habitat, is +sufficient to identify the species in question. + +On the coast of Essex the name "quay-lowders" is given to these +crustaceans, "lowder" being apparently an old plural of louse. + +[Illustration: FIG. 35.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Ligia oceanica_.] + +It is worthy of mention that Mr. Webb, when in charge of the Marine +Biological Station at Brightlingsea, examined a very large male +specimen of _Ligia oceanica_, in which the maxillæ were duplicated and +consisted of four pairs instead of two. + +_BRITISH LOCALITIES:--_ + + _England_: Brightlingsea; (W.M.W.): Maldon; (W.M.W. from R.M.): + Southend; (J.A.M.): Whitstable; (W.M.W.): Herne Bay; Margate; + Dover; Folkestone; (J.A.M.) + + _Scotland_: Shetland to Cornwall; (Norman, 49). + + _Ireland_: East Coast; West Glengariff; Castletown; Berehaven; + Bundoran; (Scharff, 63). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Spain; (12): Denmark; Prussia; Norway; + Faroe Islands; Belgium; (59). + + _Africa_: Morocco; (16). + +_Genus_--=LIGIDIUM= Brandt, 1833 (3), p. 173. Zia, Koch (34). + +_Abdomen narrow; habitat, wet moss._ + +In _Ligidium_ there are numerous joints to the flagellum, lateral +lobes are absent from the head, and the tail appendages are completely +to be seen. All the segments of the abdomen are distinctly narrower +than those of the thorax and in this it agrees with _Trichoniscus_, +_Trichoniscoides_, _Philoscia_, and _Metoponorthus_. In these, however, +the flagellum has never more than seven joints, the tail appendages (as +in all genera but _Ligia_ and _Ligidium_) are partially hidden by the +last segment, and in all the four but _Philoscia_ there are lobes to +the head. + +=Ligidium hypnorum=, Cuvier. PLATE II. + + 1792 _Oniscus hypnorum_ Cuvier (9), pl. XXVI., figs. 3-5. + 1793 _Oniscus agilis_ Persoon, quoted by Koch in Panzer (51), + part 5, pl. XXIV. + 1830 _Ligia hypnorum_ Bosc (2), p. 179. + 1833 _Ligidium persoonii_ J. F. Brandt (3), p. 174, pl. IV., + figs. 6-7. + 1840 _Zia agilis_ Koch (34), part 34, pls. XXII. and XXIII. + 1844 _Ligidium personii_ Zaddach (77), p. 17. + 1853 _Ligidium personii_ Lereboullet (39), p. 14, pl. I., fig. 1, + pl. II., figs. 20-31. + 1857 _Ligidium personii_ Kinahan (32), p. 275, pl. XXI., fig. 14, + pl. XXII., fig. 9. + 1873 _Zia saundersii_ Stebbing (70), p. 286. + 1873 _Ligidium agile_ Norman (48), p. 419. + 1885 _Ligidium hypnorum_ Budde-Lund (8), p. 254. + 1898 _Ligidium hypnorum_ G. O. Sars (59), p. 158, pl. LXXI. + +This species, which like the last, is the only British representative +of its genus, was added to our fauna in 1873 by the Rev. Thomas R. R. +Stebbing (70) who found specimens in the neighbourhood of Copthorne +Common, Surrey. Up to the present time, when we are pleased to announce +that we discovered it in the spring of 1902 at Warley in Essex, +_Ligidium hypnorum_ has not been recorded from any other place in the +British Islands. + +[Illustration: FIG. 36.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Ligidium hypnorum_.] + +As the name of the species implies, it lives in wet situations and in +its turn connects _Ligia_ with the forms which inhabit drier places. +_Ligidium hypnorum_ might be mistaken for _Philoscia muscorum_, but as +already pointed out in the generic description, the latter has but a +few (three) joints to the flagellum, instead of from ten to thirteen. +From _Ligia_, the species under consideration is distinguished by its +small size, narrow abdomen, and habitat. + +_BRITISH LOCALITIES:--_ + + _England_: Warley, Essex; (W.M.W.): Copthorne Common, Surrey; + (Stebbing, 70). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Sweden; Denmark; Germany; (59): Turkey; (8). + +_Family_--TRICHONISCIDÆ. + +=Flagellum with less than ten joints; head with lateral lobes; tail +appendages partly hidden.= + +_Genus_--=TRICHONISCUS= Brandt, 1833 (3), p. 174. + +_Abdomen narrow; eyes compound; flagellum, usually with more than three +joints._ + +In _Trichoniscus_ the flagellum may have from seven to four (rarely +three) joints. As in _Trichoniscoides_ and _Haplophthalmus_ there are +lateral lobes to the head, though these are not very pronounced; the +body is also of small size, the abdomen narrow with both divisions of +the tail appendages equally so, and almost of the same length though +slightly covered by the last segment. The compound eyes distinguish +_Trichoniscus_ from the two genera named, and from _Platyarthrus_, +while its small size and the character of its tail-parts mark it out +from all others. + +=Trichoniscus pusillus= Brandt. Plate III. + + 1833 _Trichoniscus pusillus_ Brandt (3), p. 174, pl. IV., fig. 9. + 1838 _Itea riparia_ Koch (34), part 22, pl. XVII. + 1844 _Itea lævis_ Zaddach (77), p. 16. + 1857 _Philougria celer_ Kinahan (32), p. 281, pl. XXII., figs. 1-4. + 1858 _Philougria riparia_ Kinahan (33), pp. 191 and 198, pl. XXIII., + fig. 1. + 1868 _Philougria riparia_ Bate and Westwood (1), p. 456. + 1898 _Trichoniscus pusillus_ Sars (59), p. 161, pl. LXXII., fig. 1. + +This tiny species is found commonly amongst the roots of the herbage in +very moist places. It presents a horny translucent appearance and is +of a reddish brown colour. It runs with considerable speed, and when +it is moving, the white irregular lines with which it is beset are +not evident. _Trichoniscus pusillus_ is very much like _Trichoniscus +vividus_ in colour but the latter species is nearly twice as big and +has from five to seven joints to the flagellum, while the former has +never more than four. _Trichoniscus roseus_ is also much larger and +its bright red colour (which it loses, however, when preserved in +alcohol) is another means of distinguishing it from the species under +consideration. + +Professor Sars in his _Crustacea of Norway_ (p. 162) describes from +Christiania, under the name of _Trichoniscus pygmæus_, a still smaller +species. As this may possibly be discovered in this country a brief +comparison between it and _Trichoniscus pusillus_ may be of value. +The former reaches a length of but two millimetres; it is "whitish, +semi-pellucid with a few light brown pigmentary ramifications across +the segments and a double row of irregular opaque patches along the +middle of its back" (p. 163). Its body is covered with minute tubercles +and there are only three joints to the flagellum; its movements are by +no means rapid. + +[Illustration: FIG. 37.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Trichoniscus pusillus_.] + +The body of _Trichoniscus pusillus_ is smooth and polished. It has four +joints to the flagellum--Dr. Scharff (63) says three or four--and it +moves quickly. + +_BRITISH LOCALITIES:--_ + + _England_: Brightlingsea; Warley; (W.M.W.): Epping Forest; (Bate + and Westwood, 1): Hanwell; Southall; Kew Gardens; Langley; Burnham + Beeches; Dropmore; Skirmett; Bluebell Hill, Maidstone; (W.M.W.): + Chislehurst; Plymouth; Polperro; Looe; (Bate and Westwood, 1): + Hertfordshire; Northumberland; Durham; (Norman, 49): Exeter; + (Parfitt, 53). + + _Scotland_: Edinburgh; (Scott, 68): Cumbrae; (Robertson, 57). + + _Ireland_: Connemara; (Norman, 49): Dublin; Wexford; Cork and + Kerry; (Percival Wright _teste_ Bate and Westwood, 1): Tyrone; + Waterford; Portlaw; Kilkenny; Wicklow; (Kinahan, 33). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Spain; (15): Italy; (19): Norway; Sweden; + Denmark; Germany; (59). + + _Africa_: Algeria; Tunis; Azores; (24). + + _America_: Niagara; North America; (59). + +=Trichoniscus vividus=, Koch. PLATE IV. (from a spirit specimen). + + 1840 _Itea vivida_ Koch (34), part 34, pl. IV. + 1858 _Philougria vivida_ Kinahan (33), pp. 197 and 198, pl. XXIII., + fig. 2. + 1868 _Philougria vivida_ Bate and Westwood (1), Vol. II., pp. 458 + and 459, figs. + +This species is claret-brown in colour and under a lens it is seen +to be marbled with white, indeed in appearance it is much like +_Trichoniscus pusillus_ though twice the size. There are important +differences between the two species as regards the number of joints to +the flagellum. These vary from five to seven in _Trichoniscus vividus_ +while in the other, as already pointed out, there are not more than +four. The body is practically speaking smooth for it bears only very +small tubercles, widely separated. In _Trichoniscus vividus_ the +antennæ lack the bristles which characterise those of the other species +in the genus. The species under consideration was discovered by Dr. +Kinahan in March, 1858, at Portlaw, Co. Waterford and is active even +amongst the snow. + +[Illustration: FIG. 38.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Trichoniscus vividus_.] + +_BRITISH LOCALITIES:--_ + + _Ireland_: Portlaw, Co. Waterford; (Kinahan, 33): Cappagh, Co. + Waterford; (Scharff, Irish Nat., Vol. IX., p. 158): Borris, Co. + Carlow; (Scharff, 64.) + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: Spain; (12). + +=Trichoniscus roseus= Koch. PLATE V. + + 1838 _Itea rosea_ Koch (34), part 122, pl. XVI. + 1858 _Philougria rosea_ Kinahan (33), pp. 197 and 199, pl. XXIII., + fig. 3. + 1858 _Philougria rosea_ Bate and Westwood (1), p. 460. + 1898 _Trichoniscus roseus_ Sars (59), p. 163, pl. LXXIII., fig. 1. + +The third British species of _Trichoniscus_ is of a deep pink colour +and has a light yellow stripe down the back (in some habitats the +animals are said to be quite white). Arranged in transverse rows upon +the body are large tubercles, each of which under strong magnification +will be found to end in a tiny hair. It is distinguished from +_Trichoniscus pusillus_ by the larger size of its body, which is +also comparatively broader, and from _Trichoniscus vividus_ by the +four joints of the flagellum of its antennæ which latter have strong +bristles upon them. In the former species there are five or more joints +to the flagellum and the antennæ, though hairy, lack the bristles. +_Trichoniscus roseus_ is to be looked for in old gardens. + +[Illustration: FIG. 39.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Trichoniscus roseus_.] + +_BRITISH LOCALITIES:--_ + + _England_: Warley; (W.M.W.): Maldon; (W.M.W. from R.M.): Stanmore; + Hanwell; Ealing; Wimbledon; (W.M.W.): Berkhamsted; Torquay; + (Norman, 49): Plymouth; (Bate and Westwood, 1 and B.M.,): + Grassendale, near Liverpool; (R.W.): Newtownards; (R.W., Irish + Nat, 1904, p. 260.) + + _Scotland_: Tarbert; (Scot, 68). + + _Ireland_: Dublin; Ballyfinder, Co. Down; (Scharff, 63): + Templeogue; Dundrum; Blackrock; Rathgar, Co. Dublin; Bray, Co. + Wicklow; (R.F.S.): Oakleigh; Kerry; (R.W.): Belfast; (Welch, Irish + Nat., 1896, p. 213.): At the grave of Josiah Welch (grandson of + John Knox), Castle Upton; Richhill, Co. Armagh; Castleconnell + Ferry; (R.W.): Glenade House, Co. Antrim; (R.W. from R. Ll. + Praeger). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Spain; (12): Italy; (59): Denmark; + Germany; Holland; (39): Dalmatia; (18). + + _Africa_: Algeria; Tunis; (24). + +_Genus_--=TRICHONISCOIDES=, Sars, 1898 (59), p. 164. + +_Abdomen narrow; eyes simple; (or wanting); flagellum, with four +joints._ + +The members of this genus are very much like those of _Trichoniscus_. +In the latter, however, the hinder legs are longer in proportion and +the eyes are compound. + +=Trichoniscoides albidus= Budde-Lund. PLATE VI. + + 1879 _Trichoniscus albidus_ Budde-Lund (7) p. 9. + 1898 _Trichoniscoides albidus_ Sars (59), p. 165, pl. LXXIII., + fig. 2. + +[Illustration: FIG. 40.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Trichoniscoides albidus_.] + +We are able to include this species, as a specimen was found by Mr. +Webb at Eton Wick in the summer of 1899. It is one of a number of +species which the Rev. Canon Norman (49, p. 18) suggested as likely to +be British. It is the only representative of its genus, which does not +differ in any very important characters from the others in the family. +The narrow elongated body will serve to separate it from _Trichoniscus +vividus_ and _Trichoniscus roseus_, but on account of its size, which +is much the same as that of _Trichoniscus pusillus_ and the two British +species of _Haplophthalmus_, it will be advisable to give some further +points of distinction. From the first its white colour will serve +to differentiate it; the other two lack the narrow abdomen seen in +_Trichoniscoides albidus_. Moreover, not one of the three shows the +serrations on the side plates which characterise the species under +consideration. _Platyarthrus hoffmannseggii_ is small and white and +the edges of its side plates are toothed, but it is oval in shape, +possesses no eyes, and its stout antennæ have but a single joint to the +flagellum instead of four. On the Continent this species has been found +in rich soil. + +_BRITISH LOCALITIES:--_ + + _England_: Eton; (Stebbing, 71a): Sunderland; (Brady, 50a). + + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; Wimereux and Lyons, Forêt (25): Norway; + Denmark; (59). + +_Genus_--=HAPLOPHTHALMUS= Schöbl, 1850 (66), p. 449. + +_Abdomen broad (comparatively); eyes simple; flagellum with three +joints; back with longitudinal ridges._ + +The body of _Haplophthalmus_ is long in proportion to its width, but +there is no abrupt decrease in the breadth of the abdomen as seen in +_Trichoniscus_ and _Trichoniscoides_. The eyes are simple as in the +latter genus and the lateral lobes of the head are rather large, while +the side plates of the body are well separated. + +=Haplophthalmus mengii= Zaddach. PLATE VII. + + 1844 _Itea mengii_ Zaddach (77), p. 16. + 1860 _Haplophthalmus elegans_ Schöbl (66), p. 449. + 1885 _Haplophthalmus mengii_ Budde-Lund (8), p. 250. + 1898 _Haplophthalmus mengii_ Sars (59), p. 167, pl. LXXIV., + fig. 1. + +The Rev. Canon Norman discovered two specimens of this species in +Ireland in June, 1900 (50); in the previous year one of us (Mr. Webb) +found a single example at Eton Wick. + +The main differences between the members of this genus and their allies +are set forth in the generic description and incidentally elsewhere, so +we shall content ourselves with giving the distinctive points of the +two British species. _Haplophthalmus mengii_ has a number of raised +longitudinal ribs on each segment of the thorax, the outer ridges being +somewhat broken. There are also two prominent ribs upon the third +segment of the abdomen. + +[Illustration: FIG. 41.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Haplophthalmus mengii_.] + +_BRITISH LOCALITIES:--_ + + _England_: Eton; (Stebbing, 71a): Sunderland; (Brady, 50a). + + _Ireland_: Corcumroe Abbey; Co. Clare (Norman, 50). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Norway; Prussia; Germany; Bohemia; (59). + +=Haplophthalmus danicus= Budde-Lund. PLATE VIII. + + 1870 _Haplophthalmus elegans_ Budde-Lund (6), p. 228 + (not _Haplophthalmus elegans_ Schöbl). + 1879 _Haplophthalmus danicus_ Budde-Lund (7), p. 9. + 1881 _Haplophthalmus mengii_ Weber (76), p. 192, pl. V., figs. 7-9 + (not _Itea mengii_ Zaddach). + 1885 _Haplophthalmus danicus_ Budde-Lund (8), p. 250. + 1898 _Haplophthalmus danicus_ Sars (59), p. 168, pl. LXXIV., fig. 2. + +This species was added to the British list by the Rev. Canon Norman +(49), who found a colony in his garden at Berkhamsted. It has rows of +tubercles on its thorax instead of ridges, and there are no ribs at all +upon the abdomen. The front of the head projects further comparatively +and forms a more acute point than in _Haplophthalmus mengii_ and it is +not so purely white in colour as the latter species. + +[Illustration: FIG. 42.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Haplophthalmus danicus_.] + +_BRITISH LOCALITIES:--_ + + _England_: Warley Place; (W.M.W. from Miss Willmott): Queen's + Cottage, Kew Gardens; Stanmore; Hanwell, garden at Odstock, + Bennett's Nurseries; (W.M.W.): Berkhamsted; (Norman, 49): + Sunderland; (Brady, 50a). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Denmark; Holland; Germany; (Dollfus, Feu + de Jeun, Nat., April, 1896): Norway; (Sars, 59). + + +_Section_--=ONISCI.= + +THE OUTER DIVISIONS OF THE TAIL APPENDAGES BROADER THAN THE INNER ONES. + +_Family_-ONISCIDÆ. + +=Tail appendages projecting when the animal is walking.= + + (1.) Unable to roll up into a complete ball. + +_Genus_-=ONISCUS= Linné 1746 (41), p. 360. + +_Flagellum, with three joints; abdomen broad; head with lateral lobes._ + +The characters given above taken in conjunction with the size of the +animals will serve to distinguish the members of this genus. + +=Oniscus asellus= Linné (The "Common Slater.") PLATE IX. + + 1761 _Oniscus asellus_ Linné (41), p. 500, No. 2058. + 1792 _Oniscus murarius_ Cuvier (9), p. 22, pl. XXVI. + 1838 _Oniscus fossor_ Koch (34), part 22, pl. XXII. + 1868 _Oniscus asellus_ Bate and Westwood (1), p. 468. + 1868 _Oniscus fossor_ Bate and Westwood (1), pp. 471-2. + 1898 _Oniscus asellus_ Sars (59), p. 171, pl. LXXV. + +_Oniscus asellus_ is one of the largest of our woodlice and it is also +probably the commonest, though _Porcellio scaber_ is in many places +quite as abundant. The body of _Oniscus_ is broad and expanded and the +colour is usually a slate grey with yellowish markings more or less +regularly arranged. + +From the genus _Porcellio_ the species with which we are concerned +is at once distinguished by the three jointed flagellum. _Porcellio_ +has but two joints and has, besides, a prominent lobe projecting from +the middle of the head, which is not seen in _Oniscus_. _Philoscia_, +although it has three joints to the flagellum, has a narrow abdomen +and lacks entirely the lateral lobes which are a feature of the other +genera of Oniscidæ. + +_Oniscus fossor_ of Koch (34) was recognized by Kinahan and by Bate and +Westwood as a species. Dr. Scharff submitted specimens to Professor +Budde-Lund who found no differences between them and _Oniscus asellus_. +The former (63) mentions, however, that the characteristics of the +supposed species are those of young examples of _Oniscus asellus_, and +Professor Sars (59, p. 173) seems to be of the same opinion. Many young +examples of _Oniscus asellus_ that we have examined have a curious +whitish transverse band owing to the light colour of the dorsal plates +of the first abdominal segments. The flagellum also does not seem to +shew in young animals a distinct division into three joints. + +[Illustration: FIG. 43.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Oniscus asellus_.] + +_BRITISH LOCALITIES:--_ + + _England_: High Beach, Epping, including an albino; Maldon; + Brightlingsea; Iver; Hanwell; Eton; Kew; Pamber Forest; + Kingston-on-Soar; Bluebell Hill, Maidstone; (W.M.W.): + Lynmouth; (W.M.W. from J.T.C.). + + _Scotland_: (Scharff, 63). Dinnet, Aberdeenshire; (W.M.W. from + Madame Christen). + + _Ireland_: (Scharff, 63). Yellow form with black spots, Donegal (R.W.) + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: Almost throughout; (12): France; (25): Spain; (12): + Sweden Norway; Denmark; Germany; Holland; Italy; Iceland; (59): + Faroe Islands; Thorsharn; (R.F.S.) + + _Africa_: Azores; (24). + + _America_: Greenland; (59): North America; (Budde-Lund). + +_Genus_--=PHILOSCIA= Latreille, 1804 (37), p. 43. + +_Flagellum with three joints; abdomen narrow; head without lateral +lobes._ + +If any further differences of an obvious kind be required to +distinguish _Philoscia_ from _Oniscus_, one at least will be found in +the much greater development of the hinder legs in the former genus. + +=Philoscia muscorum= Scopoli. PLATE X. + +[Not of Lereboullet, which is an _Oniscus_, see Bate and Westwood (1).] + + 1763 _Oniscus muscorum_ Scopoli (67), p. 415. + 1793 _Oniscus sylvestris_ Fabricius (26), p. 397. + 1793 _Oniscus agilis_ Koch in Panzer (51), part 9, pl. XXIV. + 1833 _Philoscia marmorata_ Brandt (3), p. 183. + 1838 _Ligia melanocephala_ Koch (34), part 22, pl. XVIII. + 1847 _Zia melanocephala_ Koch (38), part 40, pl. I. p. 212. + 1868 _Philoscia muscorum_ Bate and Westwood (1), p. 450. + 1898 _Philoscia muscorum_ Sars (59), p. 174, pl. LXXVI., fig. 1. + +This species lives chiefly at the roots of grass and under the stones +or sticks that lie among it. _Philoscia muscorum_ has a very smooth and +shining body, and its long legs enable it to move very rapidly. The +ground colour of its dorsal surface varies from light yellow to deep +brown. There are characteristic dark markings down the middle of the +thorax and on the sides, between which are lighter patches. In dark +coloured specimens the markings are by no means so evident. + +[Illustration: FIG. 44.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Philoscia muscorum_.] + +_BRITISH LOCALITIES:--_ + + _England_: High Beach, Epping; Warley; (W.M.W.): Maldon; (W.M.W. + from R.M.): Kew; Langley; Hanwell, yellow variation; Bluebell + Hill, Maidstone; (W.M.W.): Liphook; (C.S.): Pamber Forest; + Kingston-on-Soar; (W.M.W.) + + _Scotland_: (Scott, 68). + + _Ireland_: Almost throughout; (Scharff, 63). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Spain; (12): Sicily; (19): Hertsogovinia; + (22): Sweden; (21): Norway; Denmark; Prussia; Germany; Holland; + Poland; Austria; Italy; (59): Sardinia; (21). + + _Africa_: Algeria; Tunis; (24). + +=Philoscia couchii= Kinahan. PLATE XI. + + 1858 _Philoscia couchii_ Kinahan (33), p. 195, pl. XXIII., fig. 4. + 1868 _Philoscia couchii_ Bate and Westwood (1), p. + 1885 _Ligidium couchii_ Budde-Lund (8), p. 257. + 1885 _Philoscia longicornis_ Budde-Lund (8), p. 221. + 1897 _Philoscia couchii_ Dollfus (21), p. 72, pl. I., fig. 1. + +_Philoscia couchii_ is an inhabitant of the sea-side; it is smaller +than the last species, its colour to the naked eye is a uniform +lead-grey, and its antennæ are very large (compared with its size) and +hairy. + +This species was discovered by Professor Kinahan when in the company of +Messrs. Bate and Westwood near Polperro in Cornwall in the year 1858, +and dried specimens presented by him are in the British Museum (Natural +History). + +_BRITISH LOCALITIES:--_ + + _England_: Talland Cove; Polperro; (Bate and Westwood, 1): Salcombe, + Devon; (Norman, 49): Meadefoot, Torquay; (Stebbing in 49). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Spain; (12): Sicily; (19): Sebastopol; + (Norman, 49). + + _Africa_: Azores; Canaries; Morocco; Algiers; Tunis: Egypt + Senegal; (24). + + _Atlantic Isles_: Canaries; Azores; (21). + + _Asia_: Syracuse; Bazone (18). + +[Illustration: FIG. 45.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Philoscia couchii_.] + +_Genus_--=PLATYARTHRUS= Brandt, 1833 (3), p. 174. + +[_Typhloniscus_ Schöbl (66), p. 279.] + +_Flagellum with one joint; eyes wanting; abdomen broad; habitat, ants' +nests._ + +The broad body, which is much flattened, and the very thick +antennæ distinguish _Platyarthrus_ from the other small woodlice +(Trichoniscidæ). + +=Platyarthrus hoffmannseggii= Brandt. PLATE XII. + + 1833 _Platyarthrus hoffmannseggii_ Brandt (3), p. 174, pl. IV., + fig. 10. + 1844 _Itea crassicornis_ Koch (34), part 36, pl. V. + 1860 _Typhloniscus steinii_ Schöbl (66), p. 282. + 1868 _Platyarthrus hoffmannseggii_ Bate and Westwood (1), p. 464. + 1898 _Platyarthrus hoffmannseggii_ Sars (59), p. 175, pl. LXXVI., + fig. 2. + +Up to the present this is the only woodlouse which has been found in +the nests of British ants. It is small and oval, its colour is white, +and its body is covered with tubercles. The edges of its side plates +are toothed, its flagellum has but a single joint and it has no eyes. + +Miss Kate Hall tells us that, if very hungry, ants in captivity will +kill and eat _Platyarthrus_. With regard to its own food, Lord Avebury +has favoured us with the opinion that it lives on the spores of the +lower plants, such as would be found in the ants' nest. + +[Illustration: FIG. 46.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Platyarthrus hoffmannseggii_.] + +_BRITISH LOCALITIES:--_ + + _England_: Warley; Hanwell; West Drayton; Langley; + Kingston-on-Soar; Bluebell Hill, Maidstone; (W.M.W.): Berkhamsted; + Salcombe; Devon; Cheddar Cliffs, Somerset; (Norman, 49): Ide, near + Exeter; (Parfitt, 53): Torquay; (Stebbing in 49); Lulworth Cove; + (Rev. A. R. Hogan _teste_ Bate and Westwood, 1): Hammersmith; + Oxford; Berry Head, Torquay; Plymouth; (Bate and Westwood, 1): In + the nest of _Myrmica rubra_, Newton Ferrers (E. E. Lowe). + + _Scotland_: Banff; (Thomas Edward in 49). + + _Ireland_: Leixlip, Co. Dublin; Lismore, Co. Waterford; Glengariff, + Co. Cork; (Scharff, 63): Bagenalstown, Co. Carlow; (64). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (28): Spain; (12): Denmark; Germany; Holland; + Bohemia; Austria; Tyrol; Helvetia; (59). + + * * * * * + + NOTE.--In the genera which follow, air-tubes or air-cavities + (tracheæ) are present in the outer plates of the abdominal + appendages, 1 and 2, or 1 to 5. The appendages in question have + in consequence a milk-white appearance in the living animal + owing to the fact that the enclosed air reflects white light. + Considerable interest attaches to the study of these tracheæ, + which have the same function as those of insects, but which have + been independently developed. To emphasise the latter fact the + structures are often termed "pseudotracheæ." + +_Genus_--=PORCELLIO= Latreille, 1804 (37), p. 45. + +_Flagellum, with two joints; abdomen, broad; frontal lobe projecting._ + +_Porcellio_ is easily separated from the previous genera--_Oniscus_, +_Philoscia_, and _Platyarthrus_--by its two-jointed flagellum. The +fact that the abdomen is not abruptly narrowed separates it from +_Metoponorthus_, which also lacks the prominent frontal lobe so +characteristic of _Porcellio_. The species of this genera might be +confused with _Cyclisticus_ which has two joints to the flagellum and +a broad abdomen, but the latter genus has the power of rolling itself +into a ball, while its frontal lobe is very small, and the first +segment of its thorax is comparatively larger than in any species of +_Porcellio_. + +=Porcellio scaber= Latreille. PLATE XIII. + + 1804 _Porcellio scaber_ Latreille (37), p. 45. + 1818 _Oniscus granulatus_ Lamark (36), p. 261. + 1818 _Porcellio nigra_ Say (62), p. 432. + 1840 _Porcellio brandtii_ Milne-Edwards (46), p. 168. + 1840 _Porcellio dubius_ Koch (34), part 34, pl. VIII. + 1847 _Porcellio asper_ Koch (35), p. 207, pl. VIII., fig. 98. + 1857 _Porcellio montezumæ_ Saussure (60), p. 207. + 1865 _Porcellio paulensis_ Heller (31), p. 136, p. XII., fig. 5. + 1868 _Porcellio scaber_ Bate and Westwood (1), p. 475. + 1876 _Porcellio graniger_ Miers (44), p. 223. + 1885 _Porcellio graniger_ Budde-Lund (8), p. 149. + 1898 _Porcellio scaber_ Sars (59), p. 176, pl. LXXVII. + +The body of _Porcellio scaber_ is densely covered with tubercles. Its +colour is usually of a very dark grey, but at times it is quite red or +variegated with yellow. Albino specimens have been recorded. The two +joints of the flagellum are of the same length and together equal that +of the last joint of the peduncle. Air-tubes are present in the outer +plates of the first two abdominal appendages. + +[Illustration: FIG. 47.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Porcellio scaber_.] + +_BRITISH LOCALITIES:--_ + + _England_: High Beach, Epping; Warley; Brightlingsea; (W.M.W.): + Maldon; (W.M.W. from R.M.): Langley; Kew; Skirmett; Pamber Forest; + (W.M.W.): Liphook; (C.S.): Stoke-on-Trent; Kingston-on-Soar; + (W.M.W.) + + _Scotland_: Dinnet (W.M.W. from Madame Christen). + + _Ireland_: Common everywhere; (Scharff, 63.) + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: Throughout; (59): France; (28): Spain; (15): Iceland; + (59): Faroe Isles--Thorsharn and Naalsoe--(R.F.S. and B.M., N. + Annadale). + + _America_: Greenland; North America; Sandwich Isles; (B.M.); + Mexico; (59): St. Paul; St. Croix; (59); Ascension; Tristan + d'Acunha; (23) + + _Asia_: Ceylon; Kamtschatka: (23). + + _Australia_: Melbourne; Sydney; Tasmania; New Zealand; (B.M., + Chilton). + + _Africa_: Azores; Canaries; Cape of Good Hope; (24). + +=Porcellio pictus= Brandt and Ratzeburg. PLATE XIV. + + 1833 _Porcellio pictus_ Brandt and Razteburg (4), p. 78, pl. 12, + fig. 5. + 1839 _Porcellio melanocephalus_ Koch (34), part 28, pl. XVIII. + 1853 _Porcellio melanocephalus_ Schnitzler (65), p. 24. + 1856 _Porcellio mixtus_ Fitch (29), p. 120. + 1868 _Porcellio pictus_ Bate and Westwood (1), p. + 1898 _Porcellio pictus_ Sars (59), p. 177. pl. LXXVII., fig. 1. + +There are tubercles on the body of _Porcellio pictus_, which is a +striking looking animal. Its head is black with the lateral lobes +curved outwards; there is a dark band down the middle of the back and +commonly two others on each side, with more or less conspicuous yellow +markings between. + +The distal (terminal) joint of the flagellum is but half the length of +the proximal one and the last peduncular joint is longer than the two +combined. + +The abdominal appendages--1 and 2--are provided with air-tubes. + +[Illustration: FIG. 48.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Porcellio pictus_.] + +_BRITISH LOCALITIES:--_ + + _England_: Maldon; (W.M.W. from R.M.): Chislehurst; (Bate and + Westwood, 1): Cooper's Hill, near Cheltenham; (Norman, 49): + Exeter; (Parfitt, 53): Kent; (Bate and Westwood, 1.) + + _Scotland_: Between Leith and Portobello; (Scott, 68): Cumbrae + (Scott, 68a): Ayrshire; (Boyd in Norman, 49): Banff; (T. Edwards + in Norman, 49). + + _Ireland_: Dublin; Belfast; (Bate and Westwood, 1): Galway; + Maryborough; Queen's Co., Castel; and Caher Co. Tipperary; (R.F.S.) + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): North, West-Central, and East Europe; (8): + Sweden; Norway; Denmark; Germany; Hungary; Russia; (59). + + _North America_: (8). + +=Porcellio dilatatus= Brandt. PLATE XV. + + 1833 _Porcellio dilatatus_ Brandt and Ratzeburg (4), p. 78, pl. 12., + fig. 6. + 1840 _Porcellio scaber_ Milne-Edwards (not Latreille) (46), p. 167. + 1868 _Porcellio dilatatus_ Bate and Westwood (1), p. + 1898 _Porcellio dilatatus_ Sars (59), p. 179, pl. LXXVII., fig. 2. + +The fact that _Porcellio dilatatus_ is more than half as broad as it is +long, at once distinguishes it from the other species of _Porcellio_. +It is tuberculated and of somewhat a lighter grey than _Porcellio +scaber_ usually is. The two species agree in having the two joints of +the flagellum equal, but the last peduncular joint, as in _Porcellio +pictus_, is longer than the flagellum. + +As in the two preceding species, air-tubes are found in the outer +plates of the appendages on the first two abdominal segments. +_Porcellio dilatatus_ is to be looked for near houses. + +[Illustration: FIG. 49.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Porcellio dilatatus_.] + +_BRITISH LOCALITIES:--_ + + _England_: Maldon; (W.M.W. from R.M.): Eton; (Stebbing from + W.M.W., 71a): Berkhamsted; (Norman, 50): Headley, Surrey; Ventnor; + (Stebbing in Norman, 49). + + _Ireland_: Dublin; (Scharff, 63): Dundrum; (Scharff in Norman, + 50): Galway; Roundstone; (R.F.S.): Belfast; (C. W. Buckle, Irish + Nat., Vol. XI. (1902), p. 43). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Spain; (12 ): Denmark; Norway; Germany; + Poland; Holland; (59). + + _Africa_: Madeira; Azores; (24). + + _Australia_: New Guinea; (59). + +=Porcellio rathkei= Brandt. PLATE XVI. + + 1833 _Porcellio rathkei_ Brandt (3), p. 177, fig. 10. + 1833 _Porcellio ferrugineus_ Brandt (3), p. 178. + 1840 _Porcellio trilineatus_ Koch (34), part 34, pl. IX. + 1853 _Porcellio trivittatus_ Lereboullet (39), p. 54, pl. I., + figs. 13 and 14. + 1853 _Porcellio tetramoerus_ Schnitzler (65), p. 24. + 1853 _Porcellio striatus_ Schnitzler (65), p. 24. + +There is often a light band down the back and one on either side of it +near the margin in _Porcellio rathkei_ (especially in the males), with +other more irregularly arranged light patches between. Unlike the three +species previously considered, the present one has a smooth body. The +distal joint of the flagellum is the longer, and the flagellum itself +is equal in length to the last joint of the peduncle. + +Some specimens found by Mr. Webb in 1899 at Eton were submitted to Mr. +Stebbing, and since then the former has found _Porcellio rathkei_ to +be pretty generally distributed in West Middlesex, where the species +appears to frequent the open fields. + +Air-tubes occur in abdominal appendages 1 to 5 and the white appearance +of all of these at once serves to distinguish the living animal from +_Porcellio scaber_ in which the first two pairs of abdominal appendages +alone are white. + +_BRITISH LOCALITIES:--_ + + _England_: Eton; (Stebbing, 71a): Lane End; (Stebbing, from the + Misses Johnston, 71a): Acton; Ealing; Hanwell; Southall; Northolt; + Greenford; West Drayton; Mortlake; (W.M.W.); Sunderland; (Brady, + 50a). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Bosnia; Servia; (22): Hertzogovania + (B.M.); Norway; Northern, Western, and Middle Europe, everywhere; + (59): Corfu (B.M.) + + _Asia_: Transcaucasia; (59). + + _North America_: (59). + +[Illustration: FIG. 50.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Porcellio rathkei_.] + +=Porcellio laevis= Latreille. PLATE XVII. + + 1804 _Porcellio laevis_ Latreille (37), p. 46. + 1827 _Porcellio degeerii_ Savigny and Audouin (61), p. 289. + 1833 _Porcellio cucercus_ Brandt (3), p. 177. + 1833 _Porcellio syriacus_ Brandt (3), p. 178. + 1833 _Porcellio musculus_ Brandt (3), p. 180. + 1833 _Porcellio cinerascens_ Brandt (3), p. 178. + 1833 _Porcellio dubius_ Brandt (3), p. 178. + 1837 _Porcellio poeyi_ Guérin (30), p. 6. + 1844 _Porcellio urbicus_ Koch (34), part 36, pl. IV. + 1847 _Porcellio flavipes_ Koch (35), p. 206, pl. VIII., fig. 97. + 1853 _Cylisticus laevis_ Schnitzler (65), p. 25. + 1857 _Porcellio cubensis_ Saussure (60), p. 307. + 1857 _Porcellio sumichtasli_ Saussure (60), p. 307. + 1857 _Porcellio cotillæ_ Saussure (60), p. 307. + 1857 _Porcellio aztecus_ Saussure (60), p. 307. + 1857 _Porcellio mexicanus_ Saussure (60), p. 307. + +Another smooth species is _Porcellio laevis_. The colour of its body +is light grey with irregular white markings. The large size of this +species and the very long tail-appendages of the males are features +which will help to identify it. The distal joint of the flagellum is +slightly the longer and as in the last species (_P. rathkei_) the +flagellum is equal in length to the last joint of the peduncle. The +chief habitats for this species are among vegetable rubbish near human +dwellings. + +Only the first two abdominal appendages contain air-tubes. + +[Illustration: FIG. 51.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Porcellio laevis_.] + +_BRITISH LOCALITIES:--_ + + _England_: Maldon; (W.M.W. from R.M.): Ipswich (1892); Hanwell; + Wimbledon; (W.M.W.): Kent; (Kinahan, 32). + + _Ireland_: Dublin; (Bate and Westwood, 1): Blackrock Dundrum; Co. + Dublin; Galway; (R.F.S.) + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Spain; (15): Sicily; (19): Hertzogovania; + (22): Sweden; Denmark; Germany; Belgium; Austria; Italy; Dalmatia; + Greece; Turkey; (59); Corfu; (B.M.): Inca, Majorca (23); + (B.M.--Pocock and Thomas.) + + _Asia_: Syria; Turkestan; (21). + + _Africa_: Morocco; Algeria; Tunis; Tripoli; Senegal; Egypt; (23). + + _Atlantic Isles_: Bermudas; Azores; Canaries; Cape Vera; Madeira; + (24). + + _America_: North America; Mexico; Peru; Brazil; Chili; West + Indies; Pacific Islands; (59); Sandwich Isles; (B.M.) + +=Porcellio ratzeburgii= Brandt. PLATE XVIII. + + 1833 _Porcellio ratzeburgii_ Brandt, (3), p. 178. + 1839 _Porcellio nemorensis_ Koch (34), part 28, pl. XIX. + 1839 _Porcellio lugubris_ Koch (34), part 28, pl. XX. + 1853 _Porcellio quercum_ Schnitzler (65), p. 24. + 1898 _Porcellio ratzeburgii_ Sars (59), p. 182, pl. LXXX, fig. 1. + +There are granulations on the middle of the segments in _Porcellio +ratzeburgii_ and the sides of its body are more nearly parallel than +in the other species of _Porcellio_; the frontal lobe is, practically +speaking, semicircular and the lateral plates of the thorax flank the +head to a considerable extent. As in _Porcellio pictus_, the dark band +is in the middle of the back. The distal joint of the flagellum is +nearly twice as long as the proximal, and the flagellum is shorter than +the last joint of the peduncle. This species was added to the British +list by Mr. Webb (74) in 1898. + +_Porcellio ratzeburgii_ agrees with _Porcellio rathkei_ in having +air-tubes in the first five abdominal appendages. + +[Illustration: FIG. 52.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Porcellio ratzeburgii_.] + +_BRITISH LOCALITIES:--_ + + _England_: Warley; Brightlingsea; young examples (W.M.W.): Maldon; + young examples (W.M.W. from R.M.) + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: Trafoi St. Martini, and Capitello, in the Tyrol; + (Norman, 50); East Alps, very common; Val-de-Joux; Massif de la + Chartreuse Vaulnaveys (25): Bosnia; (22): Norway; Central Europe; + Upper Pfaltz; Bohemia; Saxony; Rhaetia; (59). + +_Genus_--=METOPONORTHUS= Budde-Lund, 1879 (7), p. 4. _Porcellionides_ +Miers, 1876 (44), p. 98. + +_Flagellum, with two joints; abdomen, narrow; frontal lobe not +developed._ + +The hinder legs of _Metoponorthus_ are proportionately longer than in +any other Oniscidæ save _Philoscia_. Both genera have a narrow abdomen, +but _Philoscia_ has an extra joint to the flagellum, and shows no sign +of lateral lobes to the head. + +=Metoponorthus pruinosus= Brandt. PLATE XIX. + + 1833 _Porcellio pruinosus_ Brandt (3), p. 181. + 1840 _Porcellio truncatus_ Milne-Edwards (46), p. 173. + 1840 _Porcellio maculicornis_ Koch (34), part 34, pl. XVI. + 1853 _Porcellio frontalis_ Lereboullet (39), p. 63, pl. I, fig. 17. + 1868 _Porcellio pruinosus_ Bate and Westwood (1), p. 487. + 1877 _Porcellio (Porcellioides) flavo-vittatus_ Miers (45), p. 669, + pl., LXVIII., fig. 4. + 1898 _Metoponorthus pruinosus_ Sars (59), p. 184, pl. LXXX., fig. 2. + +Undamaged specimens of _Metoponorthus pruinosus_ are of a beautiful +bluish-grey colour, owing to a "bloom" which is easily brushed off, +revealing a dark reddish-brown tint beneath it. The antennæ are long +and have white markings upon them. + +Air-tubes occur in the first two abdominal appendages. + +[Illustration: FIG. 53.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Metoponorthus pruinosus_.] + +_BRITISH LOCALITIES:--_ + + _England_: Maldon; (W.M.W. from R.M.): Hanwell; Eton Wick; Kew; + Ipswich; Stoke-on-Trent; (W.M.W.): Chiselhurst; Oxford; (Bate and + Westwood): Berkhamsted; Burnmoor; Durham; (Norman, 49): Exeter; + (Parfitt, 53); Torquay; (B.M.--T.R.R.S.) + + _Scotland_: Banff; (Thomas Edwards in Norman, 49) + + _Ireland_: Dublin; (Kinahan, 32): Foyle District; Donegal; Galway; + Clonbrock, Co. Galway; Mornington, Co. Meath; Santry; Gleeson + Park; Dundrum, Co. Dublin; Bray; (R.F.S.) + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: Practically all the Countries of Europe are given in + Dollfus' list; (23). + + _Asia_: Japan; China; Syria; Ceylon; Sumatra; Celebes; + Phillipines; Caucasus; Himalayas; (23): Christmas Island; (B.M.) + + _Africa_: Generally distributed; Madagascar; Seychelles; (23). + + _Atlantic Isles_; (23). + + _America_: North and South, almost everywhere, to judge from M. + Dollfus' list; (23). + + _Australia_: New Caledonia; (23). + +=Metoponorthus cingendus= Kinahan. PLATE XX. + + 1857 _Porcellio cingendus_ Kinahan (32), p. 279, pl. XIX., + figs. 1468-9. + 1868 _Porcellio cingendus_ Bate and Westwood (1), p. 489. + 1885 _Metoponorthus simplex_ Budde-Lund (8), p. 188. + +The colour of _Metoponorthus cingendus_ is steel blue with red or +yellowish spots. It has a raised line across each thoracic segment and +its abdomen is narrower than in _Metoponorthus pruinosus_. + +[Illustration: FIG. 54.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Metoponorthus cingendus_.] + +_BRITISH LOCALITIES:--_ + + _England_: Salcombe, Devon; (Norman, 49): South Devon; (Stebbing + in 49). + + _Ireland_: Dublin; (B.M. from Kinahan); Mountain Districts of + Dublin, Wicklow, and Cork; Coast of Kerry; Arran Islands; Achill, + Co. Mayo; Roundstone, Co. Galway; Mallow, Caef Island; Glandore; + Brock Haven, Co. Cork; Killoughrim Forest, Co. Wexford; Kenmare, + Co. Kerry; (R.F.S.). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Spain; (12). + + (2.) Able to roll up into a ball. + +_Genus_--=CYLISTICUS= Schnitzler, 1853 (65), p. 24. + +_Flagellum, with two joints; abdomen broad; frontal lobe, very small._ + +The characters given immediately above are almost those of _Porcellio_ +with which _Cylisticus_ might, perhaps, be confounded. The latter +has the power, however, of rolling itself into a ball, and the first +segment of the thorax is comparatively larger than in any species of +_Porcellio_, indeed the side plates of the segment in question entirely +flank the head. These features, as well as the straight sides of the +body and the arched back, connect _Cylisticus_ with _Armadillidium_, +from which the former is, however, at once separated by its long +pointed tail appendages. + +=Cylisticus convexus= De Geer. PLATE XXI. + + 1778 _Oniscus convexus_ De Geer (10), p. 553, pl. XXXV., fig. 11. + 1833 _Porcellio spinifrons_ Brandt (3), p. 177. + 1836 _Porcellio laevis_ Koch (34), part 6, pl. I. + 1853 _Porcellio armadilloides_ Lereboullet (39), p. 65. pl. I., + fig. 18. + 1853 _Cylisticus laevis_ Schnitzler (65), p. 25. + 1868 _Porcellio armadilloides_ Bate and Westwood (1), p. 485. + 1898 _Cylisticus convexus_ Sars (59), p. 186, pl. LXXXI. + +There is but a single species of _Cylisticus_ found in this country, +so that it is not necessary for us to go into much further detail with +regard to it. _Cylisticus convexus_ has the two joints of the flagellum +about equal, and they together in turn closely approximate in length to +the last joint of the peduncle. Mr. Stebbing says, in a letter, that +British examples do not appear to have the "white tail-piece" seen in +Continental ones. It is not noticeable in the preserved specimens which +we have seen from Berkhamsted and Leixlip, but it is very evident in +the living ones found at Hanwell and Maidstone. + +The abdominal appendages 1 to 5 are provided with air-tubes. + +[Illustration: FIG. 55.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Cylisticus convexus_.] + +_BRITISH LOCALITIES:--_ + + _England_: Maldon; (W.M.W. from R.M.): Hanwell; Bluebell Hill, + Maidstone; Eton; (W.M.W.): Berkhamsted; Portland; (Norman, 49). + + _Scotland_: Salisbury Crags; Edinburgh; Lanarkshire; Rothesay; + (Scott, 68): Killwinning; (John Smith _fide_ Robertson, 57): + Highgate; (Bate and Westwood, 1). + + _Ireland_: Leixlip, Co. Dublin; Tempo, Co. Fermanagh; Goresbridge, + Co. Kilkenny; (R.F.S.) + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (25): Sweden; Norway; Denmark; Germany; Bohemia; + Holland; Belgium; Turkey; Caucasus; (59). + + _North America_; (59). + +_Family_--ARMADILLIDIIDÆ. + +=Tail appendages not projecting when the animal is walking.= + +_Genus_--=ARMADILLIDIUM= Brandt, 1833 (3), p. 184. + +_Flagellum, with two joints; outer division of the tail appendages +expanded and broader at the hinder end._ + +The members of the genus _Armadillidium_ are more likely to be +confounded, by the uninitiated, with the "Pill-millipedes" than with +other Woodlice. Excepting _Cylisticus_ (which has long pointed tail +appendages) no other British forms have the power of rolling themselves +up into a complete ball. The very arched body is characteristic of +_Armadillidium_, and so is the groove into which the basal joints of +the antennæ fit when the creatures curl up. + +The first two abdominal appendages only are provided with air-tubes. + +=Armadillidium nasatum= Budde-Lund. PLATE XXII. + + 1885 _Armadillidium nasatum_ Budde-Lund (8), p. 51. + 1892 _Armadillidium nasatum_ Dollfus (14), p. 10, fig. 12. + 1899 _Armadillidium nasatum_ Norman (misprinted _Porcellidium_) (49), + p. 57. pl. VI., figs. 5-8. + +_Armadillidium nasatum_ has a narrow but very prominent frontal lobe, +which is almost square and curves somewhat upwards and backwards. The +joints of the flagellum are approximately equal, and are together of +the same length as the last peduncular joint. + +The telson is as long as it is broad at the base, and tapers to a +roundish point, while its sides are slightly incurved. + +[Illustration: FIG. 56.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Armadillidium nasatum_.] + +The outer divisions of the tail appendages are considerably longer than +broad, and are more or less paddle-shaped. + +It will be noticed that the slope from thorax to telson is more gentle +than in the common species, _Armadillidium vulgare_, and the first +thoracic segment is not so greatly developed. Consequently the species +which we are considering does not produce a perfect sphere, and the +antennæ are not hidden when it rolls up. It is interesting to compare +this species with _Cylisticus convexus_. The surface of the body is +smooth, and its colour is a delicate brownish grey with more or less +distinct rows of darker markings. + +_BRITISH LOCALITIES:--_ + + _England_: Maldon; (W.M.W. from R.M.): Bluebell Hill, Maidstone + (W.M.W.); Clifton, banks of the Avon; (W.M.W. from J.T.C. 1900): + Leigh Woods, Clifton; Tunbridge Wells; South Devon; (Stebbing in + 49); Cheddar Cliffs, Somerset; (Norman, 49). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; (28): Spain; (12): Italy; (23). + +=Armadillidium vulgare= Latreille. PLATE XXIII. + + 1804 _Armadillo vulgaris_ Latreille (37), p. 48. + 1804 _Armadillo variegatus_ Latreille (37). + 1853 _Armadillo ater_ Schnitzler (65), p. 48. + 1816 _Armadillo maculatus_ Risso (56), p. 158. + 1818 _Armadillo pillularis_ Say (62), p. 432. + 1825 _Armadillo pustulosus_ Dermarest (11), p. 323, pl. XLIX. + 1830-4 _Armadillidium commutatum_ Brandt and Ratzeburg (4), p. 81, + pl. XIII., fig. 123. + 1833 _Armadillidium zenckeri_ Brandt (3), p. 185. + 1839 _Armadillo trivialis_ Koch (34), part 28, pl. XIV. + 1898 _Armadillidium vulgare_ Sars (59), p. 189, pl. LXXXII. + +The common pill woodlouse is _Armadillidium vulgare_. Its frontal +lobe is not large, though it is broad, while its margin where it +joins the head is rounded and slightly recurved. The proximal joint +of the flagellum is somewhat the shorter and the two together, as in +_Armadillidium nasatum_, are of about the same length as the last joint +of the peduncle. + +The telson has the form of a triangle with the angles truncated and is +about as long as it is broad at the base. The outer divisions of the +tail appendages are considerably broader than they are long. + +The species can roll itself up into a very perfect sphere, and when +it assumes this form its antennæ are hidden beneath the much expanded +lateral plates of the first thoracic segment. + +[Illustration: FIG. 57.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Armadillidium vulgare_.] + +The body is smooth, shiny, and strongly arched. Its colour varies very +considerably, generally it is of a slaty-grey, but yellow markings are +often present to a greater or less extent. In a specimen before us +(from Bluebell Hill, Maidstone) the head is of a uniform dark grey, +the sides of the thorax are yellow, while the back is mottled with the +same colour. The abdomen, including the telson, is also yellow with the +exception only of the tail appendages, which are dark grey. + +_BRITISH LOCALITIES:--_ + + _England_: Warley; (W.M.W): Maldon; (W.M.W. from R.M.): + Brightlingsea; Hanwell; Mortlake; Bluebell Hill, Maidstone; + Langley; Skirmett; Pamber Forest; Kingston-on-Soar; Ipswich; + (W.M.W.); Lynmouth; (W.M.W. from J.T.C.) + + _Scotland_: (Scott, 68.) + + _Ireland_: Ardrahan; (Norman, 50): Borris, Co. Carlow; Glandare, + Terneay, Co. Cork; Courtstown, Co. Wexford; Cappagh, Co. + Waterford; Castel, Co. Tipperary; (R.F.S.) + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: Throughout; (23). + + _Asia_: Damascus; (23). + + _Africa_: Algeria; (23). + + _Atlantic Isles_: (23). + + _America_: North and South; (23). + + _Australia_: Melbourne (64 quoting Budde-Lund); New Zealand; (23). + +=Armadillidium pulchellum= Zencker. PLATE XXIV. + + 1799 _Oniscus pulchellus_ Zencker (78) (quoted by Koch in Panzer), + part 62, pl. XXI. + 1833 _Armadillidium pulchellum_ Brandt (3), p. 188. + 1861 _Armadillo maculatus_ Sill (69), p. 5. + 1870 _Armadillidium pictum_ Plateau (not Brandt) (55), p. 116. + 1898 _Armadillidium pulchellum_ Sars (59), p. 191, pl. LXXXIII., + fig. 4. + +The smallest British species is _Armadillidium pulchellum_. The frontal +lobe projects so as to make the head somewhat triangular. The antennæ +are very short and the distal joint of the flagellum is three times the +length of the other, while the two together are not as long as the last +peduncular joint. + +[Illustration: FIG. 58.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Armadillidium pulchellum_.] + +The telson is truncated at the end so that it is by no means as long as +it is broad at the base, and the outer divisions of the tail appendages +are in similar proportion. + +The colour of the body (which is smooth) is dark brown with four +important series of light patches running down the back and less marked +variegations between them. + +_BRITISH LOCALITIES:--_ + + _England_: Matlock; (T.R.R.S.) Arnside; Westmorland (Brady, 50a). + + _Ireland_: Ballymote, (Irish Nat., May, 1901), Sligo; (Scharff). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: Vosges; Switzerland; Pyrenees; (23): Forest de Soignes; + Belgium; (14). + +=Armadillidium depressum= Brandt. PLATE XXV. + + 1833 _Armadillidium depressum_ Brandt (3), p. 82. pl. XII., + figs, 4, 5, 6, C, D. + 1892 _Armadillidium depressum_ Dollfus (14), p. 17-18. + +The frontal lobe in _Armadillidium depressum_ is very prominent and +much recurved. The antennæ are fairly long and while the two joints +of the flagellum are nearly equal they are together not so long as +the last peduncular joint. + +[Illustration: FIG. 59.--FLAGELLUM AND LAST PEDUNCULAR JOINT OF THE +ANTENNA OF _Armadillidium depressum_.] + +The telson is slightly longer than it is broad at the base, and its +sides are incurved. As in _Armadillidium vulgare_ and _Armadillidium +pulchellum_ the outer divisions of the tail appendages are broader +than they are long. + +The body is flatter than in the other species and bears tubercles; its +colour is a slate-grey with yellowish markings. The first thoracic +segment is well developed and the head appears as if almost completely +imbedded in it. + +_BRITISH LOCALITIES:--_ + + _England_: Clifton, banks of the Avon; (W.M.W. from J.T.C., 1900): + Shirehampton, near Bristol; (Stebbing in 49): Clifton; (Dollfus + from Miers 14). + +_FOREIGN DISTRIBUTION:--_ + + _Europe_: France; Italy; (25): Asia Minor; (14, quoting Brandt). + +=Distribution of Species.= There are not sufficient records at present +to enable us to draw any conclusions as to the general distribution of +Woodlice in the British Isles, but it is hoped that more attention will +be given to these creatures, and that before long there may be other +material available. + +=Conclusion.= At the beginning it was mentioned that the present work +grew out of an investigation into the fauna of Essex, and in order +to show what may be expected when places are explored in which no +collecting has been done, we may briefly indicate the results which we +obtained in the county in question. + +It was not long before a species new to Britain--to wit, _Porcellio +ratzeburgii_--was found (74). This discovery was mentioned by Mr. +Stebbing in the _Victoria County History of Essex_ (p. 71), and he +prophesied that most of the British species then would be met with in +the county. We may safely claim to have shown that his prediction was +true, for we have been able to record in the preceding pages no less +than sixteen other species, as will be seen from the following lists:-- + +_WOODLICE RECORDED FROM ESSEX._ + + 1. _Ligia oceanica_ + 2. _Ligidium hypnorum_ + 3. _Trichoniscus pusillus_ + 4. _Trichoniscus roseus_ + 5. _Haplophthalmus danicus_ + 6. _Oniscus asellus_ + 7. _Philoscia muscorum_ + 8. _Platyarthrus hoffmannseggii_ + 9. _Porcellio scaber_ + 10. _Porcellio pictus_ + 11. _Porcellio dilatatus_ + 12. _Porcellio laevis_ + 13. _Porcellio ratzeburgii_ + 14. _Metoponorthus pruinosus_ + 15. _Cylisticus convexus_ + 16. _Armadillidium nasatum_ + 17. _Armadillidium vulgare_ + +Of these _Ligidium hypnorum_ calls for special mention, as it had not +been found in this country since Mr. Stebbing discovered it in Surrey +in 1873. Several of the Porcellios and _Cylisticus convexus_ have been +met with in but few places, and the same may be said of _Armadillidium +nasatum_. + +Of the British species not as yet found in Essex _Trichoniscus vividus_ +has at present only been recorded from Ireland; _Philoscia couchii_ and +_Armadillidium depressum_ have not been collected except in the extreme +south west of England, while _Metoponorthus cingendus_ has hitherto +only been noticed in Devonshire and Ireland. The other four species, +with the exception of _Porcellio rathkei_, which is well distributed +in west Middlesex (and might have been expected to occur in Essex), +are still rare. In fact, for _Trichoniscoides albidus_ but two British +localities are known; for _Haplophthalmus mengii_ three (two in England +and one in Ireland); while _Armadillidium pulchellum_ has only been +recorded from two or three places. + +In other counties quite as satisfactory results were obtained as in +Essex--a systematic search in Buckinghamshire brought to light at Eton +three species which at the time had not been recorded from the British +Isles, while in Middlesex, no less than a dozen species were found at +Hanwell. + +Since part of this contribution was printed our attention has been +drawn to some notes by the Rev. Canon Norman and Professor G. S. Brady +(50a). These bear out the remarks which have already been made, for +among the species found by Professor Brady in the north of England +were _Trichoniscoides albidus_, _Haplophthalmus mengii_, _Porcellio +rathkei_, and _Armadillidium pulchellum_. In one of Canon Norman's +previous papers (50--1903) he claims to have added the second species +to the British list, and in the notes in question a similar claim is +made with regard to the first and third. It should, however, be pointed +out that all three of them were found in Buckinghamshire in 1899 by Mr. +Webb, and that they were exhibited at the Nature Study Exhibition held +in London in August, 1902. A specimen of _Armadillidium pulchellum_ +from Matlock was sent to us by the Rev. T. R. R. Stebbing in January, +1904, and was found, we understand, some considerable time previously. + +It only remains for us to express our hearty thanks to the numerous +friends and correspondents who have given us their ready help. The +names of these have been printed in the text, but we would like to +mention more particularly Dr. Calman, of the British Museum (Natural +History), Monsieur Adrian Dollfus, Mr. Roland Matthams, the Rev. Canon +Norman, Dr. Scharff, the Rev. T. R. R. Stebbing, and Miss Willmott. + + +FOOTNOTES: + +[1] "The Non-Marine Molluscs of Essex," by Wilfred Mark Webb; ESSEX +NATURALIST, Vol. x. (1897), pp. 27-48 and 65-81. + +[2] The numbers in brackets refer to papers mentioned in the +Bibliography at the end. + +[3] It should be pointed out that the methylated spirit now sold in the +shops contains mineral naphtha and goes milky on the addition of water. +Permission can be obtained from Somerset House to buy what is still +called "ordinary methylated spirit," but at present five gallons has to +be purchased at one time. + + + + +BIBLIOGRAPHY. + + +Explanation of the initials given in the text:-- + + B.M.--Specimens in the British Museum (Natural History). + J.T.C.--John Thomas Carrington. + J.A.M.--James A. Murie. + R.M.--Roland Matthams. + R.F.S.--R. F. Scharff. + C.S.--Charles Sillem. + T.R.R.S.--The Rev. Thomas R. R. Stebbing. + W.M.W.--Wilfred Mark Webb. + R.W.--R. Welch. + + (1) BATE, C. SPENCE, and WESTWOOD, J.C.: _A history of the British + Sessile-eyed Crustacea_, London, 1868. + + (2) BOSC, L. A. G.: _Manuel de l'histoire Naturelle des + Crustacés_, Vol. II. Paris, 1830. + + (3) BRANDT, J. F.: "Conspectus Monographiae Crustaceorum + Oniscodorum Latreillei," _Bull. Soc. Nat., Moscow._ Vol. VI. + (1833), pp. 171-193. + + (4) BRANDT, J. F., and RATZEBURG, J. T. C.: _Medizinische + Zoologie._ Vol. II., Berlin, 1830-1834. + + (5) BRUNTZ, L.: "Contribution à l'Etude de l'Excretion chez les + Arthropodes." _Archives de Biologie._ Vol. XXII. (Nov., 1903), + pp. 215-422, pls. I.-VIII. + + (6) BUDDE-LUND, G.: "Danmarks Isopode Landkrebsdyr," + _Naturhistorisk Tidsskrift._ Part 3, vol. VII. (1870, + pp. 217-245). + + (7) BUDDE-LUND, G.: _Prospectus generum specierumque Crustaceorum + Isopodum terrestrium._ Copenhagen, 1879. + + (8) BUDDE-LUND, G.: _Crustacea Isopoda Terrestria._ Hauniae, 1885. + + (9) CUVIER, G.: "Memoires sur les Cloportes terrestres." _Journ. + d'hist. nat._ Vol. II. 1792. + + (10) DE GEER, C.: _Memoires pour servir à l'histoire des + insectes._ Vol. VII., Stockholm, 1778. + + (11) DESMAREST, A.G.: _Considérations générales sur la classe des + Crustacés._ Paris, 1825. + + (12) DOLLFUS, A.: "Catalogue raisonné des Isopodes terrestres de + l'Espagne." _Ann. Soc. Espan. Hist. Nat._ Vol. XXI. (1892), + pp. 161-190. + + (13) DOLLFUS, A.: "Sur la Distribution Geographique des Isopodes + terrestres dans la region des Basses-Pyrenees." _Assoc. + Francaise pour l'avancement des Sciences. Rep. Congress de + Pau._, 1892. + + (14) DOLLFUS, A.: "Le Genre Armadillidium." _Feu. des Jeunes + Naturalistes_, Ser. 3. May 1892. + + (15) DOLLFUS, A.: "Catalogue raisonné des Isopodes terrestres de + l'Espagne." Premier supplement, _Ann. Soc. Espan. Hist. Nat._ + Vol. XXII. (1893), pp. 47-51. + + (16) DOLLFUS, A.: "Isopodes Terrestres in 'Voyage de M. Ch. + Allmand aux Iles Canaries.'" _Mem. Soc. Zool. de France._ + Vol. VI. (1893), p. 46. + + (17) DOLLFUS, A.: "Sur la Distribution Geographique des + Armadilliens en Europe." _Compte-Rendu troisieme Congres. + Internal. de Zool._ Leyden, Sept. 1895 (1896), pp. 356-358. + + (18) DOLLFUS, A.: "Les Isopodes terrestres du Nord de l'Afrique du + Cap. Blanc a Tripoli." _Mem. Soc. Zool. Trans._ Vol. IX. + (1896), pp. 523-553. + + (19) DOLLFUS, A.: _Crustacés Isopodes de la Sicile._ 1896. + + (20) DOLLFUS, A.: "On West Indian Isopod Crustaceans." _Proc. + Zool. Soc. Lond._ 1896, pp. 388-400. + + (21) DOLLFUS, A.: "Tableau Iconographique des _Philoscia_ + d'Europe." _Feu. des Jeun. Natural._ Ser. 3, 1897, pp. 70-73 + and 91-95. + + (22) DOLLFUS, A.: "Land Isopods der Balkan region." _Wissenschaft + Mittheilung aus Bosnien und der Hercegovina._ Vol. IV. (1896). + + (23) DOLLFUS, A.: "Les Crustacés Isopodes Terrestres à grande + dispersion," _Feu des Jeun. Natural._ Ser. 3 (Oct. 1897). + + (24) DOLLFUS, A.: "Sur la Distribution Geographique des Isopodes + Terrestres dans l'Afrique Septentrional du Senegal à Obock." + _Proc. Inter. Congres. Zool._ Cambridge, 1898, pp. 249-259. + + (25) DOLLFUS, A.: "Catalogue des Crustacés Isopodes Terrestres de + France." _Feu. des Jeun. Natural._ Ser. 3 (October 1899). + + (26) FABRICIUS, J.C.: _Entomologia systematica._ Vol. II., + Hauniae, 1793. + + (27) FABRICIUS, J. C.: Supplement to the last, 1798. + + (28) FERNIE, Dr.: _Animal Simples_, 1899. + + (29) FITCH, Asa.: _Noxious Insects of New York._ Albany, 1856. + + (30) GUERIN, F.E.: "Sur _Porcellio Poeyi_." _Ann. Sci. Soc. Entom. + de France._ Vol. VI. (Paris, 1837, in Bull. ent., page 6). + + (31) HELLER, C.: _Reise der Novara; Crustacea._ 1865. + + (32) KINAHAN, J.R.: "Analysis of certain genera of terrestrial + Isopoda." _Nat. Hist. Rev._ Vol. IV. (1857). + + (33) KINAHAN, J.R.: "On the genera _Philoscia_, _Itea_ and + _Philougria_." _Nat. Hist. Rev._ Vol. V. 1858. + + (34) KOCH, C.L.: _Deutschlands Crustaceen, Myriapoden and + Arachniden._ Regensburg, 1835-1844. + + Koch's descriptions and figures were published in _Deutschlands + Crustaceen_ and appeared also in Panzer's _Faunæ Insectorum + Germanicæ_ continued by Herrich-Schäffer (which see). The + number of the parts of the latter which correspond to those + of the former are placed in square brackets after them in the + list here given. Part 6 (1836) [139]; Part 22 (1838) [162]; + Part 28 (1839) [178]; Part 34 (1840) [180]; Part 36 (1844) + [186]. Some of the dates quoted will be found to differ from + those usually given (in Budde-Lund (8, p. 7) for instance), + and printed on the parts of the copy in the Zoological + Society's Library. Our authority for this change is Mr. C. D. + Sherborne, who bases his dates upon reviews which he has + discovered. + + (35) KOCH, C. L.: _System der Myriapoden mit den Verzeichnissen + und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und + Arachniden._ Regensburg, 1847. + + (36) LAMARCK, J. B.: _Histoire naturelle des animaux sans + vertébres._ Vol. V., 1818. + + (37) LATREILLE, P. A.: _Histoire Naturelle des Crustacés et des + Insectes._ Vol. VII., Paris, 1804. + + (38) LEACH, W. E.: "Tabular view of the external characters of + four classes of animals, which Linné arranged under Insectes." + _Trans. Linn. Soc. Lond._ Vol. XI., 1815. + + (39) LEREBOULLET, A.: "Memoire sur les Crustacés de la famille des + Cloportides qui habitent les environs de Strasbourg." _Mem. Soc. + Mus. Hist. Nat. Strasbourg._ Vol. IV., 1853, pp. 130, pl. X. + + (40) LEUCKART, R.: _Verzeichnisz der zur Fauna Helgolands + gehörenden wirbellosen Seethiere._ Braunsweig, 1847. + + (41) LINNÉ, C.: _Fauna Suecica._ First edition, Holmiæ, 1746. + + (42) LINNÉ, C.: _Fauna Suecica._ Second edition, Holmiæ, 1761. + + (43) LINNÉ, C.: _Systema Naturae_, ed. 12, 1767. + + (44) MIERS, E. J.: "Catalogue of New Zealand Crustacea." _Ann. + Mag. Nat. Hist., Ser. 4._, Vol. XVII., 1876. + + (45) MIERS, E. J.: "On a collection of Crustacea (Decapoda and + Isopoda), chiefly from South America." _Proc. Zool. Soc._, + 1877, p. 653-678. + + (46) MILNE-EDWARDS, H.: _Histoire naturelle des Crustacés._ Vol. + III. Paris, 1840. + + (47) NICHOLSON and LYDDEKER: _Manual of Palæontology_, 1889, Vol. + II., p. 559. + + (48) NORMAN, A. M.: "Note on the discovery of _Ligidium agile_, + Persoon (_Zia saundersii_ Stebbing)." _Ann. Mag. Nat. Hist. + Ser. 4_, Vol. XI., p. 419. + + (49) NORMAN, A. M.: "British Land Isopoda." _Ann. Mag. Nat. Hist. + (Ser. 7)_, Vol. III. (1899), pp. 71-78, pl. VI. + + (50) NORMAN, A. M.: Continuation of the last. Vol. XI., 1903, pp. + 309-372. + + (50a) NORMAN, A.M.: and BRADY C.S.: "British Land Isopoda." Second + Supplement _Ann. Mag. Nat. Hist. (Ser. 7)_, Vol. XIV. (1904), + pp. 449-450. + + (51) PANZER, G. W. F.: _Faunæ Insectorum Germanicæ initia, oder + Deutschlands Insecten._ Parts 1-110, Nurnberg, 1793-1813. + (See Koch.) + + (52) PANZER, G. W. F.: Continuation by Herrich-Schäffer, Parts + 111-190, 1829-1844. (See Koch.) + + (53) PARFITT, E.: "The Fauna of Devon." Sessile-Eyed Crustacea. + _Trans. Devon Assoc. Sci._, Sept. 1873. + + (54) PERSOON: No paper upon Woodlice by this author is known, but + he is quoted in Panzer's _Deutschlands Insecten_ (51), which + see. + + (55) PLATEAU, F.: "Crustacés Isopodes Terrestres." _Bull. Acad. + Roy. de Belgique, Ser. 2._, Vol. XXIX., No. 2 (1870), p. 112. + + (56) RISSO, A.: _Histoire Naturelle des Crustacés des environs de + Nice._ Paris, 1816. + + (57) ROBERTSON, D.: "Catalogue of the Amphipoda and Isopoda of + the Firth of Clyde." _Trans. Nat. Hist. Soc. Glasgow_, Vol. II. + (1888), pp. 9-99. + + (58) ROULE, LOUIS: "Etudes sur le Development de Crustacés." _Ann. + Sci. Nat._, Vol. XVIII. (1895), pp. 1-156, pls. I.-X. + + (59) SARS, G. O.: _An account of the Crustacea of Norway._ Vol. + II., 1896-1899. + + (60) SAUSSURE, H. de: "Diagnoses de quelques crustacés nouveaux + des Antilles et du Mexique." _Rev. et Mag. de Zool. (Ser. 2)_, + Vol. IX. (1857), pp. 304-308. + + (61) SAVIGNY, J. C., and AUDOUIN, V.: _Description de l'Egypte._ + Vol. XXII., 1827. + + (62) SAY, T.: "An account of the Crustacea of the United States." + _J. Acad. Nat. Sci. Philadelphia_, Vol. I., pt. II., 1818. + + (63) SCHARFF, R. F.: "The Irish Woodlice." _Irish Naturalist_, + Vol. III., 1894, pp. 4-7 and 25-29, pl. II. + + (64) SCHARFF, R. F.: "The Woodlice of Co. Carlow." _Irish + Naturalist_, 1895, p. 319. + + (65) SCHNITZLER, H. J.: _De Oniscineis agri Bonnensis._ (Thesis.), + Cologne, 1853. + + (66) SCHOBL, J.: "_Typhloniscus_, eine neue blinde Gattung der + Crustacea Isopoda." _Sitzungsberichte der math. naturw. Acad. + Wiss. Wien._ Vol. XL. (1860), pp. 279-330. + + (67) SCOPOLI, J. A.: _Entomologia Carniolica._ Vindibonæ, 1763. + + (68) SCOTT, T.: "The Land and Freshwater Crustacea of the District + around Edinburgh." _Proc. R. Phys. S. Edin._, Vol. XI. + (1890-91), p. 75. + + (68a) SCOTT, T.: _British Association Hand Book on the Natural + History of Glasgow._ 1901; _Isopoda_, pp. 335 and 336. + + (69) SILL, VICTOR: "Beitrag zur Kentniss der Crustaceen, + Arachniden and Myriapoden Siebenburgens." _Verhandl. u. + Mittheil. des Siebenburghischen Ver. für Naturwiss. zu + Hermannstadt_, Vol. XII. (1861), p. 1-11. + + (70) STEBBING, T. R. R.: "On a Crustacean of the Genus Zia." _Ann. + Mag. Nat. Hist. Ser. 4._, Vol. XI. (1873). + + (71) STEBBING, T. R. R.: _The Victoria County History of Essex_ + (1903); Crustacea, pp. 27-28. + + (71a) STEBBING, T. R. R.: _The Victoria County History of + Buckinghamshire_ (1905); Crustacea. + + (72) VEJDOVSKYF: "Zur Morphologie der Antennen und Schalendruse + der Crustaceen." _Zeit. Wiss. Zool._, Vol. LXIX., p. 378. + + (73) VERHOEF, K. W.: "Ueber Palæarktische Isopoden." _Zool. Anz._, + Vol. XXIV. + + (74) WEBB, WILFRED MARK: "The occurrence in Essex of a species + of Woodlouse (Isopoda) new to Britain (_Porcellio ratzeburgii_, + Brandt)." _Essex Naturalist_, Vol. XI. (1899), p. 127. + + (75) WEBB, WILFRED MARK: "Notes on Woodlice" (Including extracts + from a paper read before the North London Natural History + Society by James B. Casserley). _Science Gossip_, Vol. VI., + New Series (1900), pp. 295-296. + + (76) WEBER, MAX.: "Uber einige neue Isopoden der Niederlandischen + Fauna." _Tijdschr der Niederland Dierk veren_, Vol. V. (1881), + pp. 167-196, pl. V. + + (77) ZADDACH, E. G.: _Synopses crustaceorum Prussicorum prodromus + Regiomonti_, 1844. + + (78) ZENKER, C. D.: No paper upon woodlice by this author is + known, but he is quoted in Panzer's _Deutschlands Insecten_ + (51), which see. + + (79) ZITTEL, K. A. von: _Textbook of Palæontology_. English + Translation 1900, p. 668. + + + + +INDEX. + + + Abdomen, 2, =2= + + Aegidae, 2 + + Air cavities, in abdominal appendages, 31 + + Air tubes, 1, 6, 31, 33, 34, 35, 36, 37, 39 + + _albidus, Trichoniscoides_, description of, 25 + + Alcohol, 16 + + Antennae, large, 2, =3= + + " smaller, 2, =3= + + Alimentary canal, 6, =7= + + Ants, 31 + + " wood, 14 + + Anus, 31, inset facing =10= + + Appendages, 2 + + " abdominal, 5, =6= + + _Archaeoniscus brodiei_, 2 + + Armadillidæ, 17 + + " characters of, 18, 40 + + _Armadillidium_, 15, 38 + + " _commutatum_, 41 + + " _depressum_, 44 + + " " description of, 43 + + " " flagellum of, =43= + + _Armadillidium nasatum_, 41, 44, Plate xxii. + + " " description of, 40 + + " " flagellum of, =40= + + _Armadillidium pictum_, 42 + + " _pulchellum_, 43, 44, 45, Plate xxxiv. + + " " description of, 42 + + " " flagellum of, =42= + + _Armadillidium vulgare_, 15, 40, 43, 44, Plate xxiii. + + " " description of, 41 + + " " flagellum of, =41= + + " " moulting of, 12, 13 + + " _zenckeri_, 41 + + _Armadillo ater_, 41 + + " _maculatus_, 41, 42 + + " _pillularis_, 41 + + " _pustulosus_, 41 + + " _trivialis_, 41 + + " _variegatus_, 41 + + " _vulgaris_, 41 + + Arthropoda, 1 + + Arteries, 7, =7= + + _asellus, Oniscus_, description of, 27 + + Avebury, Lord, 31 + + + Baden, Miocene of, 2 + + Bate and Westwood, 17, 19, 30 + + Blastoderm, 10 + + Body cavity, 11, inset facing =10= + + Brady, Professor C. S., 45 + + Branchial nephrocytes, 8 + + British Association, 19 + + Brood pouch, 4, =5=, 9 + + Budde-Lund, 28 + + + Calman, Dr., 45 + + Carpenter, 15 + + Cells, nucleated, 10, inset facing =10= + + Cephalic nephrocytes, 8 + + Cheslip, 15 + + Chiselbob, 15 + + Church louse, 16 + + Cicatricula, 10, inset facing =10= + + _cingendus, Metoponorthus_, description of, 38 + + Circulatory system, 6, 7 + + Classification, 17 + + Coal measures, 1 + + Collection of Woodlice, methods of, 16 + + Commissures, 8, =8= + + Conclusion, 43 + + _convexus, Cylisticus_, description of,39 + + _couchii, Philoscia_, description of, 30 + + Crabs, land, 1 + + Crustacea, 1 + + Cud worm, 15, 16 + + _Cylisticus_, 40 + + " characters of, 17, 18, 38 + + _Cylisticus convexus_, 44, Plate xxi. + + " " description of, 39 + + " " flagellum of, =39= + + " _laevis_, 35, 39 + + _Cymothoa oceanica_, 20 + + + _danicus, Haplophthalmus_, description of, 27 + + _depressum, Armadillidium_, description of, 43 + + Development, 9 + + Digestive glands, 6, =7= + + _dilatatus, Porcellio_, description of, 33 + + Distribution of species, 43 + + Dollfus, Adrian, 45 + + + Economic considerations, 12 + + Egg, 9, 10, =10=, =11=, inset facing =10= + + " segmentation of, 11, inset facing =10= + + Embryo, 10, =11= + + Endoderm, 10, inset facing =10= + + Essex, Woodlice recorded from, 44 + + Excretory organs, 7 + + Exhibition of species new to Britain, 45 + + Exmoor, woodlice and ants on, 14 + + External structure, 2 + + + Flagellum, 3, =3= + + Food, 13, 14 + + + Ganglia, cerebral 8 + + Geological history, 1 + + Genera, 17 + + Gill, 5, =5= + + Gills, 7, =6= + + _Glomeris marginata_, 15 + + Grammar sows, 16 + + Growth, time required for, 12 + + Gullet, 6 + + Gut, fore, inset facing =10= + + " mid, inset facing =10= + + + Habits, 12 + + Hall, Miss Kate M., 31 + + _Haplophthalmus_, 22 + + " characters of, 17, 18, 26 + + " _danicus_, 44, Plate viii. + + " " description of, 27 + + " " flagellum of, =27= + + " _elegans_, 26 + + " _mengii_, 44, 45, Plate vii. + + " " description of, 26 + + " " flagellum of, =26= + + Head, 2, =2=, =3= + + Heart, 6, 7, =7=, 11 + + Herefordshire, Old Red Sandstone, 1 + + History, geological, 1 + + _hoffmannseggii, Platyarthrus_, description of, 30 + + Hog-louse, 15 + + Hogs, little grey, 16 + + Hydro-cyanic acid gas for destroying woodlice, 14 + + _hypnorum, Ligidium_, description of, 21 + + + Insects, 1 + + Intestine, 6, =7= + + " formation of, in embryo, 10, 11 + + Isle of Wight, Oligocene of, 2, Oolite of, 2 + + Isopoda, 1, 19 + + _Itea crassicornis_, 30 + + " _laevis_, 22 + + " _mengii_, 26 + + " _rosea_, 24 + + " _riparia_, 22 + + " _vivida_, 23 + + + Jurassic, Upper, 2 + + + Kinahan, Professor, 19, 28, 30 + + Kitchenball, 15 + + + Labels, 16 + + _laevis, Porcellio_, description of, 35 + + _Ligia_, 21 + + " branchial nephrocytes, 8 + + " characters of, 17, 18, 19, 20 + + " _hypnorum_, 21 + + " _melanocephala_, 29 + + " _oceanica_, 44, Plate i. + + " " description of, 20 + + " " excretory organs of, 7, 8 + + " " flagellum of, 20, =20= + + " _scopulorum_, 20 + + Ligiae, 17 + + " characters of, 19 + + Ligiidae, characters of, 17, 18, 19 + + _Ligidium_, 19 + + " characters of, 17, 18, 21 + + " _hypnorum_, 44, Plate ii. + + " " description of, 21 + + " " excretory organs of, 7 + + " " in Surrey, 19 + + " _persoonii_, 21 + + Lip, the lower, 4, =5= + + " the upper, 4, =5= + + Lucre pig, 15 + + Lugdor, 15 + + + Mandibles, 3, =4= + + Manure, from stables, favourable to woodlice, 14 + + Maxillae, first, 3, =4= + + " second, 3, =4= + + " openings of excretory organs on second, 7 + + Maxillipeds, 3, =4= + + _mengii, Haplophthalmus_, description of, 26 + + Mesoderm, 10, inset facing =10= + + Methylated spirit, 16 + + _Metoponorthus_, 21 + + " characters of, 18, 37 + + " _cingendus_, description of, 38, Plate xx. + + " " flagellum of, =38= + + " _pruinosus_, 44, Plate xix. + + " " description of, =37= + + " " flagellum of, 37 + + " _simplex_, 38 + + Miocene, 2 + + Middlesex, West, 35 + + Millipede, pill, 15, 40 + + Moult, 11 + + Moulting, process of, 12, 13, 14 + + Mouth appendages, 3, =4= + + _muscorum, Philoscia_, description of, 29 + + + Names, local, 15 + + _nasatum, Armadillidium_, description of, 40 + + Nephrocytes, branchial, 8 + + " cephalic, 8 + + Nerve cord, 8, =8= + + Nervous system, 8 + + Norman, Rev. Canon A. M., 25, 27, 45 + + + _oceanica, Ligia_, description of, 20 + + Oenigen, Miocene of, 2 + + Old-sows, 16 + + Old women of the wood, little, 16 + + Oligocene, 2 + + Onisci, 17 + + " characters of, 27 + + Oniscidae, 19 + + _Oniscus_, 28 + + " branchial nephrocytes, 8 + + " characters of, 17, 18, 27 + + " _agilis_, 21 + + " _asellus_, 4, 15, 28, 44, Plate ix. + + _Oniscus asellus_, Albino in Epping Forest, 28 + + " " description of, 27 + + " " excretory organs of, 8 + + " " flagellum of, =28= + + " " young examples of, 28 + + " _convexus_, 39 + + " _fossor_, 19, 28 + + " _granulatus_, 32 + + " _murarius_, 27 + + " _muscorum_, 29 + + " _oceanicus_, 20 + + " _pulchellum_, 42 + + " _sylvestris_, 29 + + Oolite, 2 + + Ovaries, 8, =8= + + Oviducts, 8, =8= + + + Palmer, 16 + + Penis, 9, =9= + + Peduncle, 3, =3= + + _Philoscia_, 21, 28 + + " characters of, 17, 18, 29 + + " _couchii_, 44, Plate xi. + + " " description of, 30 + + " " flagellum of, =30= + + " _marmorata_, 29 + + " _muscorum_, 21, 44, Plate x. + + " " description of, 20 + + " " flagellum of, =29= + + _Philougria celer_, 29 + + " _riparia_, 22 + + " _rosea_, 24 + + " _vivida_, 23 + + Physicians, college of, rules for preparing woodlice for + medicinal use, 18 + + _pictus, Porcellio_, description of, 33 + + Pill millipede, 15, 40 + + _Platyarthrus_, 22 + + " characters of, 17, 18, 39 + + " _hoffmannseggii_, 44, Plate xii. + + " " description of, 30 + + " " flagellum of, =31= + + _Porcellio_, 6, 28, 38 + + " characters of, 7, 18, 32 + + " _armadilloides_, 39 + + " _asper_, 32 + + " _aztecus_, 35 + + " _brandtii_, 32 + + " _cinerascens_, 35 + + " _cingendus_, 38 + + " _cotillae_, 35 + + " _cubensis_, 35 + + " _degeerii_, 35 + + " _dilatatus_, 34, 44, Plate xv. + + " " description of, 33 + + " " flagellum of, =34= + + " _dubius_, 32, 33 + + " _ferrugineus_, 34 + + " _flavipes_, 35 + + " _flavo-vittatus_, 37 + + " _frontalis_, 37 + + " _graniger_, 32 + + " _laevis_, 39, 44, Plate xvii. + + " " description of, 35 + + " " flagellum of, =35= + + " _lugubris_, 36 + + " _maculicornis_, 37 + + " _melanocephalus_, 33 + + " _mexicanus_, 35 + + " _mixtus_, 33 + + " _montezumae_, 32 + + " _musculus_, 35 + + " _nemorensis_, 36 + + " _nigra_, 32 + + " _paulensis_, 32 + + " _pictus_, 34, 44, Plate xiv. + + " description of, 33 + + " flagellum of, =33= + + " _poeyi_, 35 + + " _pruinosus_, 37 + + " _quercum_, 36 + + " _rathkei_, 35, 36, 44, 45 + + " " description of, 34, Plate xvi. + + " " flagellum of, =35= + + " _ratzeburgii_, 44, Plate xviii. + + " " description of, 36 + + " " flagellum of, =36= + + " _scaber_, 28, 44, Plate xiii. + + " " description of, 32 + + " _scaber_, development of, 9 + + " " flagellum of, 32 + + " " Milne Edwards, 33 + + " _spinifrons_, 39 + + " _striatus_, 34 + + " _sumichtasti_, 35 + + " _syriacus_, 35 + + " _tetramoerus_, 34 + + " _trilineatus_, 34 + + " _trivittatus_, 34 + + " _truncatus_, 37 + + " _urbicus_, 35 + + _Porcellionides_, Miers, 37 + + Preservation of Woodlice, methods of, 16 + + Pro-ectoderm, 10, inset facing =10= + + Pro-endoderm, 10, inset facing =10= + + Protoplasm, formative, 10 + + _pruinosus, Metoponorthus_, description of, 37 + + pseudotracheae, 31 + + _pulchellum, Armadillidium_, description of, 42 + + _pusillus, Trichoniscus_, description of, 22 + + Purbeck Beds, 2 + + + Quay-louse, 20 + + Quay lowders, 20 + + + _rathkei, Porcellio_, description of, 34 + + _ratzeburgii, Porcellio_, description of, 36 + + Reproductive organs, 8, =8=, =9= + + _roseus, Trichoniscus_, description of, 24 + + Roule, Professor Louis, 9 + + + Sandstone, old red, 1 + + Sars, Professor G. O., 17, 23 + + _scaber, Porcellio_, description of, 32 + + Scharff, Dr. R. F., 17, 19, 23, 45 + + Seminal reservoir, 9, =9= + + Socchetre, 16 + + Sow bug, 15 + + Species, distribution of, 43 + + St. Anthony's hogs, 16 + + Stebbing, the Rev. T. R. R., 19, 21, 35, 44, 45 + + Swanley Horticultural College, 14 + + + Tertiary deposits, 2 + + Testes, 9, =9= + + Thorax, 2, 3 + + Thrush louse, 15 + + Tiggyhog, 15 + + Tracheae, 6, 31 + + Trichoniscidæ, characters of, 18, 22 + + _Trichoniscoides_, 21, 22, 26 + + " characters of, 17, 18, 25 + + " _albidus_, 44, 45 + + " " description of, 25 + + " " flagellum of, =25= + + _Trichoniscus_, 21, 22, 26 + + " characters of, 17, 18, 22 + + " _pusillus_, 22, 23, 24, 25, 44, Plate iii. + + " " description of, 22 + + " " flagellum of, =23= + + " _pygmaeus_, description of, 23 + + " _roseus_, 23, 25, 44, Plate v. + + " " description of, 24 + + " " flagellum of, =24= + + " _vividus_, 22, 25, 44, Plate iv. + + " " description of, 23 + + " " flagellum of, =24= + + Tubes for specimens, 16 + + _Typhloniscus steinii_, 30 + + + _vividus, Trichoniscus_, description of, 23 + + _vulgare, Armadillidium_, description of, 41 + + + Walking legs, 4, 5 + + Webb, Mr. Wilfred Mark, 20, 25, 26, 35, 36, 45 + + Welsh names for Woodlice, 16 + + Westwood, Bate and, 17, 19, 30 + + White arsenic for destroying Woodlice, 14 + + Willmott, Miss, 45 + + Wood-ants, 15 + + Woodlice, in captivity, 12 + + " methods for getting rid of, 24 + + " position of, 1 + + " storage of specimens, 16 + + " Welsh names for, 16 + + Woodlouse, common, 4 + + + Yolk, food, 10, 11, inset facing =10= + + + _Zia agilis_, 21 + + " _melanocephala_, 29 +$ + + + + +LIST OF PLATES. + + + PLATE + + _Ligia oceanica_ Linné I. + + _Ligidium hypnorum_ Cuvier II. + + _Trichoniscus pusillus_ Brandt III. + + _Trichoniscus vividus_ Koch IV. + + _Trichoniscus roseus_ Koch V. + + _Trichoniscoides albidus_ Budde-Lund VI. + + _Haplophthalmus mengii_ Zaddach VII. + + _Haplophthalmus danicus_ Budde-Lund VIII. + + _Oniscus asellus_ Linné IX. + + _Philoscia muscorum_ Scopoli X. + + _Philoscia couchii_ Kinahan XI. + + _Platyarthrus hoffmannseggii_ Brandt XII. + + _Porcellio scabier_ Latreille XIII. + + _Porcellio pictus_ Brandt and Ratzeburg XIV. + + _Porcellio dilatatus_ Brandt XV. + + _Porcellio rathkei_ Brandt XVI. + + _Porcellio laevis_ Latreille XVII. + + _Porcellio ratzeburgii_ Brandt XVIII. + + _Metoponorthus pruinosus_ Brandt XIX. + + _Metoponorthus cingendus_ Kinahan XX. + + _Cylisticus convexus_ De Geer XXI. + + _Armadillidium nasatum_ Budde-Lund XXII. + + _Armadillidium vulgare_ Latreille XXIII. + + _Armadillidium pulchellum_ Zencker XXIV. + + _Armadillidium depressum_ Brandt XXV. + +[Illustration: _PLATE I._ + +LIGIA OCEANICA Linné. THE QUAY-LOUSE. + +_Length, two to three centimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE II._ + +LIGIDIUM HYPNORUM Cuvier. + +_Length, nine millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE III._ + +TRICHONISCUS PUSILLUS Brandt. + +_Length, four millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE IV._ + +TRICHONISCUS VIVIDUS Koch. + +_Length, eight millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE V._ + +TRICHONISCUS ROSEUS Koch. + +_Length, five millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE VI._ + +TRICHONISCOIDES ALBIDUS Budde-Lund. + +_Length, four millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE VII._ + +HAPLOPHTHALMUS MENGII Zaddach. + +_Length, three to four millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE VIII._ + +HAPLOPHTHALMUS DANICUS Budde-Lund. + +_Length, three to four millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE IX._ + +ONISCUS ASELLUS Linné (The common slater). + +_Length, sixteen millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE X._ + +PHILOSCIA MUSCORUM Scopoli. + +_Length, nine millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XI._ + +PHILOSCIA COUCHII Kinahan. + +_Length, nine millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XII._ + +PLATYARTHRUS HOFFMANNSEGGII Brandt. + +_Length, three millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XIII._ + +PORCELLIO SCABER Latreille. + +_Length, fourteen millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XIV._ + +PORCELLIO PICTUS Brandt and Ratzeburg. + +_Length, thirteen millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XV._ + +PORCELLIO DILATATUS Brandt. + +_Length, fifteen millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XVI._ + +PORCELLIO RATHKEI Brandt. + +_Length, twelve millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XVII._ + +PORCELLIO LAEVIS Latreille. + +_Length, sixteen millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XVIII._ + +PORCELLIO RATZEBURGII Brandt. + +_Length, eleven millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XIX._ + +METOPONORTHUS PRUINOSUS Brandt. + +_Length, nine millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XX._ + +METOPONORTHUS CINGENDUS Kinahan. + +_Length, six millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XXI._ + +CYLISTICUS CONVEXUS De Geer. + +_Length, twelve millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XXII._ + +ARMADILLIDIUM NASATUM Budde-Lund. + +_Length, fifteen millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XXIII._ + +ARMADILLIDIUM VULGARE Latreille. + +_Length, fifteen millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XXIV._ + +ARMADILLIDIUM PULCHELLUM Zencker. + +_Length, five millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + +[Illustration: _PLATE XXV._ + +ARMADILLIDIUM DEPRESSUM Brandt. + +_Length, fifteen millimetres._ + +Charles Sillem, del. ad nat. F. W. Reader, sculpt.] + + + * * * * * + + +Transcriber's Notes + +Obvious typographical errors have been corrected. Inconsistent +accents, punctuation, and hyphenation are as in the original text. + +The "æ" ligature is used interchangeably with "ae" throughout. + +Some taxonomic names may have changed since 1906. + +The following misprints and misspellings have been noted or corrected: + +CONTENTS:- +Page vii: "Family--Ligidæ " changed to "Family--Ligiidæ". +Page vii: "Haplopthalmus mengii" changed to "Haplophthalmus mengii". +Page viii: "Armydillidium vulgare" changed to "Armadillidium vulgare". + +MAIN PART OF BOOK:- +Page 2: "Aegidae is found" changed to "Aegidae which is found". +Page 16: "naptha" changed to "naphtha" in footnote. +Page 28: "Thornsharn" changed to "Thorsharn". +Page 29: "Philoscia marmorala" changed to "Philoscia marmorata". +At the end of the following lines, there is a missing page reference: + Page 30: "1868 _Philoscia couchii_ Bate and Westwood (1), p." + Page 33: "1868 _Porcellio pictus_ Bate and Westwood (1), p." + Page 33: "1868 _Porcellio dilatatus_ Bate and Westwood (1), p." + +INDEX:- +Page 51: "Ligidae, characters of" changed to "Ligiidae, characters of". + + + + + +End of the Project Gutenberg EBook of The British Woodlice, by +Wilfred Mark Webb and Charles Sillem + +*** END OF THE PROJECT GUTENBERG EBOOK 43928 *** |
