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diff --git a/.gitattributes b/.gitattributes new file mode 100644 index 0000000..6833f05 --- /dev/null +++ b/.gitattributes @@ -0,0 +1,3 @@ +* text=auto +*.txt text +*.md text diff --git a/30297-0.txt b/30297-0.txt new file mode 100644 index 0000000..2d23fec --- /dev/null +++ b/30297-0.txt @@ -0,0 +1,398 @@ +*** START OF THE PROJECT GUTENBERG EBOOK 30297 *** + +UNIVERSITY OF KANSAS PUBLICATIONS + +MUSEUM OF NATURAL HISTORY + +Volume 14, No. 17, pp. 483-491, 2 figs. +March 2, 1964 + + +Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, +From the Chadronian and Orellan + + +BY + +WILLIAM A. CLEMENS, JR. + + +UNIVERSITY OF KANSAS +LAWRENCE +1964 + + +UNIVERSITY OF KANSAS PUBLICATIONS, MUSEUM OF NATURAL HISTORY + +Editors: E. Raymond Hall, Chairman, Henry S. Fitch, +Theodore H. Eaton, Jr. + +Volume 14, No. 17, pp. 483-491, 2 figs. +Published March 2, 1964 + +UNIVERSITY OF KANSAS +Lawrence, Kansas + +PRINTED BY +HARRY (BUD) TIMBERLAKE, STATE PRINTER +TOPEKA, KANSAS + +1964 + + +29-8587 + + + + +Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, +From the Chadronian and Orellan + +BY + +WILLIAM A. CLEMENS, JR. + + +Introduction + + +The family Apatemyidae has a long geochronological range in North +America, beginning in the Torrejonian land-mammal age, but is +represented by a relatively small number of fossils found at a few +localities. Two fossils of Orellan age, found in northeastern Colorado +and described here, demonstrate that the geochronological range of the +Apatemyidae extends into the Middle Oligocene. Isolated teeth of +_Sinclairella dakotensis_ Jepsen, part of a sample of a Chadronian +local fauna collected by field parties from the Webb School of +California, are also described. + + I thank Mr. Raymond M. Alf, Webb School of California, + Claremont, California, and Dr. Peter Robinson, University of + Colorado Museum, Boulder, Colorado, for permitting me to + describe the fossils they discovered. Also Dr. Robinson made + available the draft of a short paper he had prepared on the + tooth found in Weld County, Colorado; his work was + facilitated by a grant from the University of Colorado + Council on Research and Creative Work. I also gratefully + acknowledge receipt of critical data and valuable comments + from Drs. Edwin C. Galbreath, Glenn L. Jepsen, and Malcolm + C. McKenna who is currently revising the Paleocene + apatemyids and studying the phylogenetic relationships of + the family. The prefixes of catalogue numbers used in the + text identify fossils in the collections of the following + institutions: KU, Museum of Natural History, The University + of Kansas, Lawrence; Princeton, Princeton Museum, Princeton, + New Jersey; RAM-UCR, Raymond Alf Museum, Webb School of + California, Claremont, California (the permanent repository + for these specimens will be the University of California, + Riverside); and UCM, University of Colorado Museum, Boulder, + Colorado. The system of notations for teeth prescribed for + use here is as follows: teeth in the upper half of the + dentition are designated by a capital letter and a number; + thus M2 is the notation for the upper second molar; teeth in + the lower half of the dentition are designated by a + lower-case letter and a number; thus p2 is the notation for + the lower second premolar. + + + + +Family APATEMYIDAE Matthew, 1909 + +Genus =Sinclairella= Jepsen, 1934 + +=Sinclairella dakotensis= Jepsen, 1934 + + +The type of the species, Princeton no. 13585, was discovered in +Chadronian strata of the upper part of the Chadron Formation cropping +out in Big Corral Draw, approximately 13 miles south-southwest of +Scenic, in southwestern South Dakota (Jepsen, 1934, p. 291). Detailed +descriptions of the type specimen are given in papers by Jepsen (1934) +and Scott and Jepsen (1936). Isolated teeth of Chadronian age referable +to _Sinclairella dakotensis_ have been discovered subsequently at a +locality in Nebraska and fossils of Orellan age, also referable to _S. +dakotensis_, have been collected at two localities in Colorado. The +sample from each locality is described separately. + + +Sioux County, northwestern Nebraska + + _Material._--RAM-UCR nos. 381, left M1; 598, left m2; 1000, + right m1; 1001, right m2; 1079, right m2; 1674, right M2; + and 3013, left m2. + + _Locality and stratigraphy._--These Chadronian fossils were + discovered by Raymond Alf and members of his field parties + in several harvester ant mounds built in exposures of the + Chadron Formation in Sec. 26, T 33 N, R 53 W, Sioux County, + Nebraska (Alf, 1962, and Hough and Alf, 1958). This is UCR + locality V5403. The collectors carefully considered the + possibility that some of the fossils found in the ant mounds + were collected from younger strata by the harvester ants and + concluded this was unlikely (Alf, personal communication). + + _Description and comments._--The cusps of RAM-UCR no. 381, a + left M1, are sharp and the wear-facets resulting from + occlusion with the lower dentition are small. The paraconule + is a low, ill-defined cusp on the anterior margin of the + crown; a metaconule is not present. A smooth stylar shelf is + present labial to the metacone. The crown was supported by + three roots. There are no interradicular crests. + + The crown of RAM-UCR no. 1674, a right M2, is heavily + abraded and many morphological details of the cusps have + been destroyed. Low interradicular crests linked the three + roots of the tooth with a low, central prominence. As was + the case with RAM-UCR no. 381, no significant differences + could be found in comparisons with illustrations of the + teeth preserved in Princeton no. 13585. + + RAM-UCR nos. 598, 1001, 1079, and 3013 all appear to be + m2's. The talonids of these teeth are not elongated, their + trigonids have quadrilateral outlines, and the paraconids + are small but prominent, bladelike cusps. The trigonid of + RAM-UCR 1000 is elongated and the paraconid is a minute + cusp; the tooth closely resembles the m1 of the type of + _Sinclairella dakotensis_. + + +Logan County, northeastern Colorado + + _Material._--KU no. 11210 (fig. 1), a fragment of a left + maxillary containing P4 and M1-2. + + _Locality and stratigraphy._--The fossil was found in the + center of the W-1/2, Sec. 21, T 11 N, R 53 W, Logan County, + Colorado, "... in the bed below _Agnotocastor_ bed, Cedar + Creek Member...." (Ronald H. Pine, 1958, field notes on file + at the University of Kansas). The bed so defined is part of + unit 3 in the lower division of the Cedar Creek Member, as + subdivided by Galbreath (1953:25) in stratigraphic section + XII. The fauna obtained from unit 3 is of Orellan age. + +[Illustration: FIG. 1. _Sinclairella dakotensis_ Jepsen, KU no. 11210, +fragment of left maxillary with P4 and M1-2; Orellan, Logan County, +Colorado; drawings by Mrs. Judith Hood: a, labial view; b, occlusal +view; both approximately × 9.] + + _Description and comments._--P4 of KU no. 11210 has a large + posterolingual cusp separated from the main cusp by a + distinct groove, which deepens posteriorly. The + posterolingual cusp is supported by the broad posterior + root. P4 of the type specimen of _Sinclairella dakotensis_ + is described (Jepsen, 1934, p. 392) as having an oval + outline at the base of the crown, and a small, + posterolingual cusp. A chip of enamel is missing from the + posterior slope of the main cusp of the P4 of KU no. 11210. + The anterior slope of the main cusp is flattened, possibly + the result of wear, and there is no evidence of a groove + like that present on the P4 of the type specimen. + + Only a few differences were found between the molars + preserved in KU no. 11210 and their counterparts in the type + specimen. A stylar shelf is present labial to the metacone + of M1 of KU no. 11210, but, unlike the type, its surface is + smooth and there is no evidence of cusps. Of the three small + stylar cusps on the stylar shelf of M2 the smallest is in + the position of a mesostyle. The M2 lacks a chip of enamel + from the lingual surface of the hypocone. Unlike the M2 of + Princeton no. 13585, in occlusal view the posterior margin + of the M2 of KU no. 11210 is convex posterior to the + metacone. The anterior edge of the base of the zygomatic + arch of KU no. 11210 was dorsal to M2. The shallow oval + depression in the maxillary dorsal to M1 might be the result + of post-mortem distortion. + + The molars preserved in KU no. 11210 and their counterparts + in the type specimen do not appear to be significantly + different in size (table 1) or morphology of the cusps. The + only difference between the two specimens that might be of + classificatory significance is the difference in size of the + posterolingual cusp of P4. At present the range of + intraspecific variation in the morphology of P4 has not been + documented for any species of apatemyid. The evolutionary + trend or trends of the apatemyids (McKenna, 1960, p. 48) for + progressive reduction of function of p4 probably were + paralleled by similar trends in the evolution of the P4. If + so, the intraspecific variation in the morphology of P4 + could be expected to be somewhat greater than that of the + upper molars, for example. The morphological difference + between the P4's of the type of _Sinclairella dakotensis_ + and KU no. 11210 is not extreme and does not exceed the + range of intraspecific variation that could be expected for + this element of the dentition. The close resemblances in + size and morphology between the M1-2 of Princeton no. 13585 + and KU no. 11210 also favor identification of the latter as + part of a member of an Orellan population of _Sinclairella + dakotensis_. + + +Weld County, northeastern Colorado + +[Illustration: FIG. 2. _Sinclairella dakotensis_ Jepsen, UCM no. 21073, +right M2; Orellan, Weld County, Colorado; drawing by Mrs. Judith Hood: +occlusal view, approximately × 9.] + + _Material._--UCM no. 20173 (fig. 2), is a right M2. + + _Locality and stratigraphy._--The tooth was discovered at + the Mellinger locality, Sec. 17, T 11 N, R 65 W, Weld + County, Colorado. The Mellinger locality is in the Cedar + Creek Member, White River Formation, and its fauna is + considered to be of Orellan age (Patterson and McGrew, 1937, + and Galbreath, 1953). + + _Description and comments._--UCM no. 21073, which is more + heavily abraded than KU no. 11210, shows no evidence of a + stylar cusp either anterolabial to the metacone or in the + position of a mesostyle. A small stylar cusp is present + anterolabial to the paracone. A notch that appears to have + been cut through the enamel of the posterolabial corner of + the crown could have received the parastylar apex of M3. A + similar notch is not present on the M2 of KU no. 11210 nor + indicated in the illustrations of the M2 of Princeton no. + 13585. The coronal dimensions of UCM no. 21073 (table 1) do + not appear to differ significantly from those of the M2's of + KU no. 11210 and the type specimen of _Sinclairella + dakotensis_. + + +Comments + +With the discovery of Orellan apatemyids the geochronological range of +the family in North America is shown to extend from the Torrejonian +through the Orellan land-mammal ages. The discoveries reported here +enlarge the Oligocene record of apatemyids to include not only the type +specimen of _Sinclairella dakotensis_, a skull and associated mandible +from South Dakota, but also seven isolated teeth, representing at least +two individuals, from a Chadronian fossil locality in Nebraska and one +specimen from each of two Orellan fossil localities in northeastern +Colorado. Simpson (1944:73, and 1953:127) presented tabulations of the +published records of American apatemyids and suggested the data +indicated the populations of these mammals were of small size +throughout the history of the family. The few pre-Oligocene occurrences +of apatemyids described subsequently (note McKenna, 1960, figs. 3-10, +and p. 48) and occurrences described here tend to reinforce Simpson's +interpretation. This interpretation may have to be modified to some +degree, however, when current studies of collections of pre-Oligocene +apatemyids are completed (McKenna, personal communication). + +Although information concerning the evolutionary trends of American +apatemyids has been published, no data on the morphological variation +in a population are available in the literature. An adequate basis for +evaluating the significance of the morphological differences between +the P4's of Princeton no. 13585 and KU no. 12110 coupled with the +similarities of their M1-2's is lacking. In the evolution of American +apatemyids the P4 underwent reduction in size and, apparently, +curtailment of function. This history suggests the range of +morphological variation of P4 in populations of _Sinclairella +dakotensis_ could be expected to be greater than that of the molars and +encompass the morphological differences between the P4's of Princeton +no. 13585 and KU no. 12110. The difference in age of the Chadronian and +Orellan fossils does not constitute proof that they pertain to +different species. Although the identification is admittedly +provisional until more fossils including other parts of the skeleton +are discovered, the Orellan fossils described here are referred to +_Sinclairella dakotensis_. + + +TABLE 1.--MEASUREMENTS (IN MILLIMETERS) OF TEETH OF SINCLAIRELLA +DAKOTENSIS JEPSEN. + +========================================================================== + | P4 | M1 | M2 +-----------------------+------------+------------------+------------------ + |length|width|length[1]|width[1]|length[1]|width[1] +-----------------------+------+-----+---------+--------+---------+-------- +Princeton no. 13585[2] | 2.1 | 1.1 | 4.0 | 3.7 | 3.4 | 4.7 +RAM no. 381 | | | 4.1 | 3.5 | | +RAM no. 1674 | | | | | 3.4 | 4.2 +KU no. 11210 | 2.4 | 1.6 | 3.9 | 3.5 | 3.8 | 4.1+ +UCM no. 21073 | | | | | 3.6 | 4.1 +-----------------------+------+-----+---------+--------+---------+-------- + | m1 | m2 + +---------+--------+---------+-------- + | length | width | length | width + +---------+--------+---------+-------- +Princeton no. 13585[3] | 3.5 | 2.4 | 3.7 | 2.8 +RAM no. 1000 | 3.5 | 2.2 | | +RAM no. 598 | | | 3.8 | 2.6 +RAM no. 1001 | | | 3.6+ | 2.6 +RAM no. 1079 | | | 4.0 | 2.8 +RAM no. 3013 | | | 3.6 | 2.8 +------------------------------------+---------+--------+---------+-------- + +[Footnote 1: Length defined as maximum dimension of the labial half of +the crown measured parallel to a line drawn through the apices of +paracone and metacone. Width defined as maximum coronal dimension +measured along line perpendicular to line defined by apices of paracone +and metacone.] + +[Footnote 2: Dimensions provided by Dr. Glenn L. Jepsen.] + +[Footnote 3: Dimensions taken from Jepsen (1934:300).] + + + + +Literature Cited + +ALF, R. + 1962. A new species of the rodent _Pipestoneomys_ from the + Oligocene of Nebraska. Breviora, Mus. Comp. Zool., no. 172, + pp. 1-7, 3 figs. + +GALBREATH, E. C. + 1953. A contribution to the Tertiary geology and paleontology + of northeastern Colorado. Univ. Kansas Paleont. Cont., + Vertebrata, art. 4, pp. 1-120, 2 pls., 26 figs. + +HOUGH, J., and ALF, R. + 1958. A Chadron mammalian fauna from Nebraska. Journ. Paleon. + 30:132-140, 4 figs. + +JEPSEN, G. L. + 1934. A revision of the American Apatemyidae and the description + of a new genus, _Sinclairella_, from the White River + Oligocene of South Dakota. Proc. Amer. Philos. Soc., + 74:287-305, 3 pls., 4 figs. + +MCKENNA, M. C. + 1960. Fossil Mammalia from the early Wasatchian Four Mile fauna, + Eocene of northwest Colorado. Univ. California Publ. in + Geol. Sci., 37:1-130, 64 figs. + +MATTHEW, W. D. + 1909. The Carnivora and Insectivora of the Bridger Basin, Middle + Eocene. Mem. Amer. Mus. Nat. Hist., 9:289-567, pls. 42-52, + 118 figs. + +PATTERSON, B. and MCGREW, P. O. + 1937. A soricid and two erinaceids from the White River Oligocene. + Geol. Ser., Field Mus. Nat. Hist., 6:245-272, figs. 60-74. + +SCOTT, W. B. and JEPSEN, G. L. + 1936. The mammalian fauna of the White River Oligocene--Part I. + Insectivora and Carnivora. Trans. Amer. Philos. Soc., n. s., + 28:1-153, 22 pls., 7 figs. + +SIMPSON, G. G. + 1944. Tempo and mode in evolution. New York: Columbia Univ. Press, + xviii + 237 pp., 36 figs. + + 1953. The major features of evolution. New York: Columbia Univ. + Press, xx + 434 pp., 52 figs. + +_Transmitted June 24, 1963._ + + + + + +End of the Project Gutenberg EBook of Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, From the Chadronian and Orellan, by William A. Clemens + +*** END OF THE PROJECT GUTENBERG EBOOK 30297 *** diff --git a/30297-8.txt b/30297-8.txt new file mode 100644 index 0000000..5b06997 --- /dev/null +++ b/30297-8.txt @@ -0,0 +1,795 @@ +The Project Gutenberg EBook of Records of the Fossil Mammal Sinclairella, +Family Apatemyidae, From the Chadronian and Orellan, by William A. Clemens + +This eBook is for the use of anyone anywhere at no cost and with +almost no restrictions whatsoever. You may copy it, give it away or +re-use it under the terms of the Project Gutenberg License included +with this eBook or online at www.gutenberg.org + + +Title: Records of the Fossil Mammal Sinclairella, Family Apatemyidae, From the Chadronian and Orellan + +Author: William A. Clemens + +Release Date: October 20, 2009 [EBook #30297] + +Language: English + +Character set encoding: ISO-8859-1 + +*** START OF THIS PROJECT GUTENBERG EBOOK FOSSIL MAMMAL SINCLAIRELLA *** + + + + +Produced by Chris Curnow, Joseph Cooper, Diane Monico, and +the Online Distributed Proofreading Team at +https://www.pgdp.net + + + + + + + + + + + +UNIVERSITY OF KANSAS PUBLICATIONS + +MUSEUM OF NATURAL HISTORY + +Volume 14, No. 17, pp. 483-491, 2 figs. +March 2, 1964 + + +Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, +From the Chadronian and Orellan + + +BY + +WILLIAM A. CLEMENS, JR. + + +UNIVERSITY OF KANSAS +LAWRENCE +1964 + + +UNIVERSITY OF KANSAS PUBLICATIONS, MUSEUM OF NATURAL HISTORY + +Editors: E. Raymond Hall, Chairman, Henry S. Fitch, +Theodore H. Eaton, Jr. + +Volume 14, No. 17, pp. 483-491, 2 figs. +Published March 2, 1964 + +UNIVERSITY OF KANSAS +Lawrence, Kansas + +PRINTED BY +HARRY (BUD) TIMBERLAKE, STATE PRINTER +TOPEKA, KANSAS + +1964 + + +29-8587 + + + + +Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, +From the Chadronian and Orellan + +BY + +WILLIAM A. CLEMENS, JR. + + +Introduction + + +The family Apatemyidae has a long geochronological range in North +America, beginning in the Torrejonian land-mammal age, but is +represented by a relatively small number of fossils found at a few +localities. Two fossils of Orellan age, found in northeastern Colorado +and described here, demonstrate that the geochronological range of the +Apatemyidae extends into the Middle Oligocene. Isolated teeth of +_Sinclairella dakotensis_ Jepsen, part of a sample of a Chadronian +local fauna collected by field parties from the Webb School of +California, are also described. + + I thank Mr. Raymond M. Alf, Webb School of California, + Claremont, California, and Dr. Peter Robinson, University of + Colorado Museum, Boulder, Colorado, for permitting me to + describe the fossils they discovered. Also Dr. Robinson made + available the draft of a short paper he had prepared on the + tooth found in Weld County, Colorado; his work was + facilitated by a grant from the University of Colorado + Council on Research and Creative Work. I also gratefully + acknowledge receipt of critical data and valuable comments + from Drs. Edwin C. Galbreath, Glenn L. Jepsen, and Malcolm + C. McKenna who is currently revising the Paleocene + apatemyids and studying the phylogenetic relationships of + the family. The prefixes of catalogue numbers used in the + text identify fossils in the collections of the following + institutions: KU, Museum of Natural History, The University + of Kansas, Lawrence; Princeton, Princeton Museum, Princeton, + New Jersey; RAM-UCR, Raymond Alf Museum, Webb School of + California, Claremont, California (the permanent repository + for these specimens will be the University of California, + Riverside); and UCM, University of Colorado Museum, Boulder, + Colorado. The system of notations for teeth prescribed for + use here is as follows: teeth in the upper half of the + dentition are designated by a capital letter and a number; + thus M2 is the notation for the upper second molar; teeth in + the lower half of the dentition are designated by a + lower-case letter and a number; thus p2 is the notation for + the lower second premolar. + + + + +Family APATEMYIDAE Matthew, 1909 + +Genus =Sinclairella= Jepsen, 1934 + +=Sinclairella dakotensis= Jepsen, 1934 + + +The type of the species, Princeton no. 13585, was discovered in +Chadronian strata of the upper part of the Chadron Formation cropping +out in Big Corral Draw, approximately 13 miles south-southwest of +Scenic, in southwestern South Dakota (Jepsen, 1934, p. 291). Detailed +descriptions of the type specimen are given in papers by Jepsen (1934) +and Scott and Jepsen (1936). Isolated teeth of Chadronian age referable +to _Sinclairella dakotensis_ have been discovered subsequently at a +locality in Nebraska and fossils of Orellan age, also referable to _S. +dakotensis_, have been collected at two localities in Colorado. The +sample from each locality is described separately. + + +Sioux County, northwestern Nebraska + + _Material._--RAM-UCR nos. 381, left M1; 598, left m2; 1000, + right m1; 1001, right m2; 1079, right m2; 1674, right M2; + and 3013, left m2. + + _Locality and stratigraphy._--These Chadronian fossils were + discovered by Raymond Alf and members of his field parties + in several harvester ant mounds built in exposures of the + Chadron Formation in Sec. 26, T 33 N, R 53 W, Sioux County, + Nebraska (Alf, 1962, and Hough and Alf, 1958). This is UCR + locality V5403. The collectors carefully considered the + possibility that some of the fossils found in the ant mounds + were collected from younger strata by the harvester ants and + concluded this was unlikely (Alf, personal communication). + + _Description and comments._--The cusps of RAM-UCR no. 381, a + left M1, are sharp and the wear-facets resulting from + occlusion with the lower dentition are small. The paraconule + is a low, ill-defined cusp on the anterior margin of the + crown; a metaconule is not present. A smooth stylar shelf is + present labial to the metacone. The crown was supported by + three roots. There are no interradicular crests. + + The crown of RAM-UCR no. 1674, a right M2, is heavily + abraded and many morphological details of the cusps have + been destroyed. Low interradicular crests linked the three + roots of the tooth with a low, central prominence. As was + the case with RAM-UCR no. 381, no significant differences + could be found in comparisons with illustrations of the + teeth preserved in Princeton no. 13585. + + RAM-UCR nos. 598, 1001, 1079, and 3013 all appear to be + m2's. The talonids of these teeth are not elongated, their + trigonids have quadrilateral outlines, and the paraconids + are small but prominent, bladelike cusps. The trigonid of + RAM-UCR 1000 is elongated and the paraconid is a minute + cusp; the tooth closely resembles the m1 of the type of + _Sinclairella dakotensis_. + + +Logan County, northeastern Colorado + + _Material._--KU no. 11210 (fig. 1), a fragment of a left + maxillary containing P4 and M1-2. + + _Locality and stratigraphy._--The fossil was found in the + center of the W-1/2, Sec. 21, T 11 N, R 53 W, Logan County, + Colorado, "... in the bed below _Agnotocastor_ bed, Cedar + Creek Member...." (Ronald H. Pine, 1958, field notes on file + at the University of Kansas). The bed so defined is part of + unit 3 in the lower division of the Cedar Creek Member, as + subdivided by Galbreath (1953:25) in stratigraphic section + XII. The fauna obtained from unit 3 is of Orellan age. + +[Illustration: FIG. 1. _Sinclairella dakotensis_ Jepsen, KU no. 11210, +fragment of left maxillary with P4 and M1-2; Orellan, Logan County, +Colorado; drawings by Mrs. Judith Hood: a, labial view; b, occlusal +view; both approximately × 9.] + + _Description and comments._--P4 of KU no. 11210 has a large + posterolingual cusp separated from the main cusp by a + distinct groove, which deepens posteriorly. The + posterolingual cusp is supported by the broad posterior + root. P4 of the type specimen of _Sinclairella dakotensis_ + is described (Jepsen, 1934, p. 392) as having an oval + outline at the base of the crown, and a small, + posterolingual cusp. A chip of enamel is missing from the + posterior slope of the main cusp of the P4 of KU no. 11210. + The anterior slope of the main cusp is flattened, possibly + the result of wear, and there is no evidence of a groove + like that present on the P4 of the type specimen. + + Only a few differences were found between the molars + preserved in KU no. 11210 and their counterparts in the type + specimen. A stylar shelf is present labial to the metacone + of M1 of KU no. 11210, but, unlike the type, its surface is + smooth and there is no evidence of cusps. Of the three small + stylar cusps on the stylar shelf of M2 the smallest is in + the position of a mesostyle. The M2 lacks a chip of enamel + from the lingual surface of the hypocone. Unlike the M2 of + Princeton no. 13585, in occlusal view the posterior margin + of the M2 of KU no. 11210 is convex posterior to the + metacone. The anterior edge of the base of the zygomatic + arch of KU no. 11210 was dorsal to M2. The shallow oval + depression in the maxillary dorsal to M1 might be the result + of post-mortem distortion. + + The molars preserved in KU no. 11210 and their counterparts + in the type specimen do not appear to be significantly + different in size (table 1) or morphology of the cusps. The + only difference between the two specimens that might be of + classificatory significance is the difference in size of the + posterolingual cusp of P4. At present the range of + intraspecific variation in the morphology of P4 has not been + documented for any species of apatemyid. The evolutionary + trend or trends of the apatemyids (McKenna, 1960, p. 48) for + progressive reduction of function of p4 probably were + paralleled by similar trends in the evolution of the P4. If + so, the intraspecific variation in the morphology of P4 + could be expected to be somewhat greater than that of the + upper molars, for example. The morphological difference + between the P4's of the type of _Sinclairella dakotensis_ + and KU no. 11210 is not extreme and does not exceed the + range of intraspecific variation that could be expected for + this element of the dentition. The close resemblances in + size and morphology between the M1-2 of Princeton no. 13585 + and KU no. 11210 also favor identification of the latter as + part of a member of an Orellan population of _Sinclairella + dakotensis_. + + +Weld County, northeastern Colorado + +[Illustration: FIG. 2. _Sinclairella dakotensis_ Jepsen, UCM no. 21073, +right M2; Orellan, Weld County, Colorado; drawing by Mrs. Judith Hood: +occlusal view, approximately × 9.] + + _Material._--UCM no. 20173 (fig. 2), is a right M2. + + _Locality and stratigraphy._--The tooth was discovered at + the Mellinger locality, Sec. 17, T 11 N, R 65 W, Weld + County, Colorado. The Mellinger locality is in the Cedar + Creek Member, White River Formation, and its fauna is + considered to be of Orellan age (Patterson and McGrew, 1937, + and Galbreath, 1953). + + _Description and comments._--UCM no. 21073, which is more + heavily abraded than KU no. 11210, shows no evidence of a + stylar cusp either anterolabial to the metacone or in the + position of a mesostyle. A small stylar cusp is present + anterolabial to the paracone. A notch that appears to have + been cut through the enamel of the posterolabial corner of + the crown could have received the parastylar apex of M3. A + similar notch is not present on the M2 of KU no. 11210 nor + indicated in the illustrations of the M2 of Princeton no. + 13585. The coronal dimensions of UCM no. 21073 (table 1) do + not appear to differ significantly from those of the M2's of + KU no. 11210 and the type specimen of _Sinclairella + dakotensis_. + + +Comments + +With the discovery of Orellan apatemyids the geochronological range of +the family in North America is shown to extend from the Torrejonian +through the Orellan land-mammal ages. The discoveries reported here +enlarge the Oligocene record of apatemyids to include not only the type +specimen of _Sinclairella dakotensis_, a skull and associated mandible +from South Dakota, but also seven isolated teeth, representing at least +two individuals, from a Chadronian fossil locality in Nebraska and one +specimen from each of two Orellan fossil localities in northeastern +Colorado. Simpson (1944:73, and 1953:127) presented tabulations of the +published records of American apatemyids and suggested the data +indicated the populations of these mammals were of small size +throughout the history of the family. The few pre-Oligocene occurrences +of apatemyids described subsequently (note McKenna, 1960, figs. 3-10, +and p. 48) and occurrences described here tend to reinforce Simpson's +interpretation. This interpretation may have to be modified to some +degree, however, when current studies of collections of pre-Oligocene +apatemyids are completed (McKenna, personal communication). + +Although information concerning the evolutionary trends of American +apatemyids has been published, no data on the morphological variation +in a population are available in the literature. An adequate basis for +evaluating the significance of the morphological differences between +the P4's of Princeton no. 13585 and KU no. 12110 coupled with the +similarities of their M1-2's is lacking. In the evolution of American +apatemyids the P4 underwent reduction in size and, apparently, +curtailment of function. This history suggests the range of +morphological variation of P4 in populations of _Sinclairella +dakotensis_ could be expected to be greater than that of the molars and +encompass the morphological differences between the P4's of Princeton +no. 13585 and KU no. 12110. The difference in age of the Chadronian and +Orellan fossils does not constitute proof that they pertain to +different species. Although the identification is admittedly +provisional until more fossils including other parts of the skeleton +are discovered, the Orellan fossils described here are referred to +_Sinclairella dakotensis_. + + +TABLE 1.--MEASUREMENTS (IN MILLIMETERS) OF TEETH OF SINCLAIRELLA +DAKOTENSIS JEPSEN. + +========================================================================== + | P4 | M1 | M2 +-----------------------+------------+------------------+------------------ + |length|width|length[1]|width[1]|length[1]|width[1] +-----------------------+------+-----+---------+--------+---------+-------- +Princeton no. 13585[2] | 2.1 | 1.1 | 4.0 | 3.7 | 3.4 | 4.7 +RAM no. 381 | | | 4.1 | 3.5 | | +RAM no. 1674 | | | | | 3.4 | 4.2 +KU no. 11210 | 2.4 | 1.6 | 3.9 | 3.5 | 3.8 | 4.1+ +UCM no. 21073 | | | | | 3.6 | 4.1 +-----------------------+------+-----+---------+--------+---------+-------- + | m1 | m2 + +---------+--------+---------+-------- + | length | width | length | width + +---------+--------+---------+-------- +Princeton no. 13585[3] | 3.5 | 2.4 | 3.7 | 2.8 +RAM no. 1000 | 3.5 | 2.2 | | +RAM no. 598 | | | 3.8 | 2.6 +RAM no. 1001 | | | 3.6+ | 2.6 +RAM no. 1079 | | | 4.0 | 2.8 +RAM no. 3013 | | | 3.6 | 2.8 +------------------------------------+---------+--------+---------+-------- + +[Footnote 1: Length defined as maximum dimension of the labial half of +the crown measured parallel to a line drawn through the apices of +paracone and metacone. Width defined as maximum coronal dimension +measured along line perpendicular to line defined by apices of paracone +and metacone.] + +[Footnote 2: Dimensions provided by Dr. Glenn L. Jepsen.] + +[Footnote 3: Dimensions taken from Jepsen (1934:300).] + + + + +Literature Cited + +ALF, R. + 1962. A new species of the rodent _Pipestoneomys_ from the + Oligocene of Nebraska. Breviora, Mus. Comp. Zool., no. 172, + pp. 1-7, 3 figs. + +GALBREATH, E. C. + 1953. A contribution to the Tertiary geology and paleontology + of northeastern Colorado. Univ. Kansas Paleont. Cont., + Vertebrata, art. 4, pp. 1-120, 2 pls., 26 figs. + +HOUGH, J., and ALF, R. + 1958. A Chadron mammalian fauna from Nebraska. Journ. Paleon. + 30:132-140, 4 figs. + +JEPSEN, G. L. + 1934. A revision of the American Apatemyidae and the description + of a new genus, _Sinclairella_, from the White River + Oligocene of South Dakota. Proc. Amer. Philos. Soc., + 74:287-305, 3 pls., 4 figs. + +MCKENNA, M. C. + 1960. Fossil Mammalia from the early Wasatchian Four Mile fauna, + Eocene of northwest Colorado. Univ. California Publ. in + Geol. Sci., 37:1-130, 64 figs. + +MATTHEW, W. D. + 1909. The Carnivora and Insectivora of the Bridger Basin, Middle + Eocene. Mem. Amer. Mus. Nat. Hist., 9:289-567, pls. 42-52, + 118 figs. + +PATTERSON, B. and MCGREW, P. O. + 1937. A soricid and two erinaceids from the White River Oligocene. + Geol. Ser., Field Mus. Nat. Hist., 6:245-272, figs. 60-74. + +SCOTT, W. B. and JEPSEN, G. L. + 1936. The mammalian fauna of the White River Oligocene--Part I. + Insectivora and Carnivora. Trans. Amer. Philos. Soc., n. s., + 28:1-153, 22 pls., 7 figs. + +SIMPSON, G. G. + 1944. Tempo and mode in evolution. New York: Columbia Univ. Press, + xviii + 237 pp., 36 figs. + + 1953. The major features of evolution. New York: Columbia Univ. + Press, xx + 434 pp., 52 figs. + +_Transmitted June 24, 1963._ + + + + + +End of the Project Gutenberg EBook of Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, From the Chadronian and Orellan, by William A. Clemens + +*** END OF THIS PROJECT GUTENBERG EBOOK FOSSIL MAMMAL SINCLAIRELLA *** + +***** This file should be named 30297-8.txt or 30297-8.zip ***** +This and all associated files of various formats will be found in: + https://www.gutenberg.org/3/0/2/9/30297/ + +Produced by Chris Curnow, Joseph Cooper, Diane Monico, and +the Online Distributed Proofreading Team at +https://www.pgdp.net + + +Updated editions will replace the previous one--the old editions +will be renamed. + +Creating the works from public domain print editions means that no +one owns a United States copyright in these works, so the Foundation +(and you!) can copy and distribute it in the United States without +permission and without paying copyright royalties. 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Clemens, Jr. + </title> + <style type="text/css"> + + + p { margin-top: .75em; + text-align: justify; + margin-bottom: .75em; + } + h1 { text-align: center; line-height: 1.5; clear: both; } + + h2,h3 { text-align: center; clear: both; } + + p.title { text-align: center; text-indent: 0; + font-weight: bold; + line-height: 1.4; margin-bottom: 3em; } + + hr { width: 33%; + margin-top: 2em; + margin-bottom: 2em; + margin-left: auto; + margin-right: auto; + clear: both; + } + + table {margin-left: auto; margin-right: auto;} + + table, td, th {border: solid thin;} + thead, tfoot, th { font-variant: small-caps; + text-align: center; + font-weight: normal;} + + body{margin-left: 10%; + margin-right: 10%; + } + + .pagenum { /* uncomment the next line for invisible page numbers */ + /* visibility: hidden; */ + position: absolute; + left: 92%; + font-size: smaller; + text-align: right; + } /* page numbers */ + + .blockquot{margin-left: 5%; margin-right: 10%;} + .center {text-align: center;} + .smcap {font-variant: small-caps;} + + .caption {font-weight: bold;} + + .figcenter {margin: auto; text-align: center;} + + .figleft {float: left; clear: left; margin-left: 0; margin-bottom: 1em; margin-top: + 1em; margin-right: 1em; padding: 0; text-align: center;} + + .footnotes {border: dashed 1px;} + .footnote {margin-left: 10%; margin-right: 10%; font-size: 0.9em;} + .footnote .label {position: absolute; right: 84%; text-align: right;} + .fnanchor {vertical-align: super; font-size: .8em; text-decoration: none;} + + .i4 {display: block; margin-left: 2.5em; + padding-left: 2.5em; text-indent: -2.5em;} + + + </style> + </head> +<body> +<div>*** START OF THE PROJECT GUTENBERG EBOOK 30297 ***</div> + +<hr style="width: 65%;" /> +<p class="title"><span class="smcap">University of Kansas Publications</span><br /> + +<span class="smcap">Museum of Natural History</span><br /> +<br /> +Volume 14, No. 17, pp. 483-491, 2 figs.<br /> +March 2, 1964</p> + +<hr style="width: 65%;" /> + +<h1>Records of the Fossil Mammal<br /> +Sinclairella, Family Apatemyidae,<br /> +From the Chadronian and Orellan<br /></h1> + +<p class="title"><small>BY</small><br /><br /> + +<big>WILLIAM A. CLEMENS, JR.</big><br /><br /></p> + + +<p class="title"><span class="smcap">University of Kansas</span><br /> + +<span class="smcap">Lawrence</span><br /> + +1964</p> +<hr style="width: 65%;" /> + + +<p class="center"><span class="smcap">University of Kansas Publications, Museum of Natural History</span><br /> +<br /> +Editors: E. Raymond Hall, Chairman, Henry S. Fitch,<br /> +Theodore H. Eaton, Jr.<br /> +<br /> +<br /> +Volume 14, No. 17, pp. 483-491, 2 figs.<br /> +Published March 2, 1964<br /> +<br /> + +<span class="smcap">University of Kansas</span><br /> +Lawrence, Kansas<br /> +<br /> +<small>PRINTED BY</small><br /> + +<small>HARRY (BUD) TIMBERLAKE, STATE PRINTER</small><br /> + +<small>TOPEKA, KANSAS</small><br /> + +<small>1964</small><br /> +<br /> +<small>29-8587</small><br /> +</p> +<hr style="width: 65%;" /> + + + +<p><span class="pagenum"><a name="Page_485" id="Page_485">[Pg 485]</a></span></p> + +<h2>Records of the Fossil Mammal<br /> +Sinclairella, Family Apatemyidae,<br /> +From the Chadronian and Orellan<br /> +</h2> + +<p class="title"><small>BY</small><br /> + +WILLIAM A. CLEMENS, JR.<br /> +</p> + + +<h3>Introduction</h3> + + +<p>The family Apatemyidae has a long geochronological range in +North America, beginning in the Torrejonian land-mammal age, but +is represented by a relatively small number of fossils found at a few +localities. Two fossils of Orellan age, found in northeastern Colorado +and described here, demonstrate that the geochronological +range of the Apatemyidae extends into the Middle Oligocene. Isolated +teeth of <i>Sinclairella dakotensis</i> Jepsen, part of a sample of a +Chadronian local fauna collected by field parties from the Webb +School of California, are also described.</p> + +<div class="blockquot"><p>I thank Mr. Raymond M. Alf, Webb School of California, Claremont, +California, and Dr. Peter Robinson, University of Colorado Museum, Boulder, +Colorado, for permitting me to describe the fossils they discovered. Also Dr. +Robinson made available the draft of a short paper he had prepared on the +tooth found in Weld County, Colorado; his work was facilitated by a grant +from the University of Colorado Council on Research and Creative Work. I +also gratefully acknowledge receipt of critical data and valuable comments +from Drs. Edwin C. Galbreath, Glenn L. Jepsen, and Malcolm C. McKenna +who is currently revising the Paleocene apatemyids and studying the phylogenetic +relationships of the family. The prefixes of catalogue numbers used +in the text identify fossils in the collections of the following institutions: KU, +Museum of Natural History, The University of Kansas, Lawrence; Princeton, +Princeton Museum, Princeton, New Jersey; RAM-UCR, Raymond Alf Museum, +Webb School of California, Claremont, California (the permanent repository +for these specimens will be the University of California, Riverside); and UCM, +University of Colorado Museum, Boulder, Colorado. The system of notations +for teeth prescribed for use here is as follows: teeth in the upper half of the +dentition are designated by a capital letter and a number; thus M2 is the notation +for the upper second molar; teeth in the lower half of the dentition are +designated by a lower-case letter and a number; thus p2 is the notation for the +lower second premolar.</p></div> + + + +<hr style="width: 65%;" /><p><span class="pagenum"><a name="Page_486" id="Page_486">[Pg 486]</a></span></p> +<h3>Family <span class="smcap">Apatemyidae</span> Matthew, 1909<br /> + +Genus <b>Sinclairella</b> Jepsen, 1934<br /> + +<b>Sinclairella dakotensis</b> Jepsen, 1934<br /></h3> + + +<p>The type of the species, Princeton no. 13585, was discovered in +Chadronian strata of the upper part of the Chadron Formation +cropping out in Big Corral Draw, approximately 13 miles south-southwest +of Scenic, in southwestern South Dakota (Jepsen, 1934, +p. 291). Detailed descriptions of the type specimen are given in +papers by Jepsen (1934) and Scott and Jepsen (1936). Isolated +teeth of Chadronian age referable to <i>Sinclairella dakotensis</i> have +been discovered subsequently at a locality in Nebraska and fossils +of Orellan age, also referable to <i>S. dakotensis</i>, have been collected +at two localities in Colorado. The sample from each locality is +described separately.</p> + + +<h3>Sioux County, northwestern Nebraska</h3> + +<div class="blockquot"><p><i>Material.</i>—RAM-UCR nos. 381, left M1; 598, left m2; 1000, right m1; 1001, +right m2; 1079, right m2; 1674, right M2; and 3013, left m2.</p> + +<p><i>Locality and stratigraphy.</i>—These Chadronian fossils were discovered by +Raymond Alf and members of his field parties in several harvester ant mounds +built in exposures of the Chadron Formation in Sec. 26, T 33 N, R 53 W, Sioux +County, Nebraska (Alf, 1962, and Hough and Alf, 1958). This is UCR +locality V5403. The collectors carefully considered the possibility that some +of the fossils found in the ant mounds were collected from younger strata by +the harvester ants and concluded this was unlikely (Alf, personal communication).</p> + +<p><i>Description and comments.</i>—The cusps of RAM-UCR no. 381, a left M1, +are sharp and the wear-facets resulting from occlusion with the lower dentition +are small. The paraconule is a low, ill-defined cusp on the anterior margin +of the crown; a metaconule is not present. A smooth stylar shelf is present +labial to the metacone. The crown was supported by three roots. There are +no interradicular crests.</p> + +<p>The crown of RAM-UCR no. 1674, a right M2, is heavily abraded and +many morphological details of the cusps have been destroyed. Low interradicular +crests linked the three roots of the tooth with a low, central prominence. +As was the case with RAM-UCR no. 381, no significant differences +could be found in comparisons with illustrations of the teeth preserved in +Princeton no. 13585.</p> + +<p>RAM-UCR nos. 598, 1001, 1079, and 3013 all appear to be m2's. The +talonids of these teeth are not elongated, their trigonids have quadrilateral outlines, +and the paraconids are small but prominent, bladelike cusps. The trigonid +of RAM-UCR 1000 is elongated and the paraconid is a minute cusp; the +tooth closely resembles the m1 of the type of <i>Sinclairella dakotensis</i>.</p></div> + +<p><span class="pagenum"><a name="Page_487" id="Page_487">[Pg 487]</a></span></p> +<h3>Logan County, northeastern Colorado</h3> + +<div class="blockquot"><p><i>Material.</i>—KU no. 11210 (fig. 1), a fragment of a left maxillary containing +P4 and M1-2.</p> + +<p><i>Locality and stratigraphy.</i>—The fossil was found in the center of the W-1/2, +Sec. 21, T 11 N, R 53 W, Logan County, Colorado, " ... in the bed below +<i>Agnotocastor</i> bed, Cedar Creek Member...." (Ronald H. Pine, 1958, +field notes on file at the University of Kansas). The bed so defined is part of +unit 3 in the lower division of the Cedar Creek Member, as subdivided by +Galbreath (1953:25) in stratigraphic section XII. The fauna obtained from +unit 3 is of Orellan age.</p></div> + +<p class="figcenter" style="width: 545px;"> +<img src="images/image005.png" width="545" height="600" alt="Fig. 1. Sinclairella dakotensis Jepsen, KU no. 11210, fragment of left maxillary +with P4 and M1-2; Orellan, Logan County, Colorado; drawings by Mrs. Judith +Hood: a, labial view; b, occlusal view; both approximately × 9." title="Fig. 1. Sinclairella dakotensis Jepsen, KU no. 11210, fragment of left maxillary" /> +<span class="caption">Fig. 1. Sinclairella dakotensis Jepsen, KU no. 11210, fragment of left maxillary +with P4 and M1-2; Orellan, Logan County, Colorado; drawings by Mrs. Judith +Hood: a, labial view; b, occlusal view; both approximately × 9.</span> +</p> + +<div class="blockquot"><p><i>Description and comments.</i>—P4 of KU no. 11210 has a large posterolingual +cusp separated from the main cusp by a distinct groove, which deepens posteriorly.<span class="pagenum"><a name="Page_488" id="Page_488">[Pg 488]</a></span> +The posterolingual cusp is supported by the broad posterior root. P4 +of the type specimen of <i>Sinclairella dakotensis</i> is described (Jepsen, 1934, p. +392) as having an oval outline at the base of the crown, and a small, posterolingual +cusp. A chip of enamel is missing from the posterior slope of the main +cusp of the P4 of KU no. 11210. The anterior slope of the main cusp is +flattened, possibly the result of wear, and there is no evidence of a groove like +that present on the P4 of the type specimen.</p> + +<p>Only a few differences were found between the molars preserved in KU no. +11210 and their counterparts in the type specimen. A stylar shelf is present +labial to the metacone of M1 of KU no. 11210, but, unlike the type, its surface +is smooth and there is no evidence of cusps. Of the three small stylar +cusps on the stylar shelf of M2 the smallest is in the position of a mesostyle. +The M2 lacks a chip of enamel from the lingual surface of the hypocone. Unlike +the M2 of Princeton no. 13585, in occlusal view the posterior margin of the +M2 of KU no. 11210 is convex posterior to the metacone. The anterior edge +of the base of the zygomatic arch of KU no. 11210 was dorsal to M2. The +shallow oval depression in the maxillary dorsal to M1 might be the result of +post-mortem distortion.</p> + +<p>The molars preserved in KU no. 11210 and their counterparts in the type +specimen do not appear to be significantly different in size (table 1) or morphology +of the cusps. The only difference between the two specimens that +might be of classificatory significance is the difference in size of the posterolingual +cusp of P4. At present the range of intraspecific variation in the morphology +of P4 has not been documented for any species of apatemyid. The +evolutionary trend or trends of the apatemyids (McKenna, 1960, p. 48) for +progressive reduction of function of p4 probably were paralleled by similar +trends in the evolution of the P4. If so, the intraspecific variation in the morphology +of P4 could be expected to be somewhat greater than that of the upper +molars, for example. The morphological difference between the P4's of the +type of <i>Sinclairella dakotensis</i> and KU no. 11210 is not extreme and does not +exceed the range of intraspecific variation that could be expected for this element +of the dentition. The close resemblances in +size and morphology between the M1-2 of Princeton +no. 13585 and KU no. 11210 also favor identification +of the latter as part of a member of an +Orellan population of <i>Sinclairella dakotensis</i>.</p></div> + + +<h3>Weld County, northeastern Colorado</h3> + +<p class="figleft" style="width: 342px;"> +<img src="images/image006.png" width="342" height="375" alt="Fig. 2. Sinclairella dakotensis +Jepsen, UCM no. 21073, right M2; Orellan, Weld County, Colorado; +drawing by Mrs. Judith Hood: occlusal view, approximately × 9." title="Fig. 2. Sinclairella dakotensis +Jepsen, UCM no. 21073, right M2" /> +<span class="caption">Fig. 2. Sinclairella dakotensis +Jepsen, UCM no. 21073, right M2; Orellan, Weld County, Colorado; +drawing by Mrs. Judith Hood: occlusal view, approximately × 9.</span> +</p> + +<div class="blockquot"><p><i>Material.</i>—UCM no. 20173 (fig. 2), is a right M2.</p> + +<p><i>Locality and stratigraphy.</i>—The tooth was discovered +at the Mellinger locality, Sec. 17, T 11 +N, R 65 W, Weld County, Colorado. The Mellinger +locality is in the Cedar Creek Member, +White River Formation, and its fauna is considered +to be of Orellan age (Patterson and McGrew, +1937, and Galbreath, 1953).</p> + +<p><i>Description and comments.</i>—UCM no. 21073, +which is more heavily abraded than KU no. 11210, +shows no evidence of a stylar cusp either anterolabial<span class="pagenum"><a name="Page_489" id="Page_489">[Pg 489]</a></span> +to the metacone or in the position of a mesostyle. A small stylar cusp +is present anterolabial to the paracone. A notch that appears to have been cut +through the enamel of the posterolabial corner of the crown could have received +the parastylar apex of M3. A similar notch is not present on the M2 of KU no. +11210 nor indicated in the illustrations of the M2 of Princeton no. 13585. The +coronal dimensions of UCM no. 21073 (table 1) do not appear to differ significantly +from those of the M2's of KU no. 11210 and the type specimen of +<i>Sinclairella dakotensis</i>.</p></div> + + +<h3>Comments</h3> + +<p>With the discovery of Orellan apatemyids the geochronological +range of the family in North America is shown to extend from the +Torrejonian through the Orellan land-mammal ages. The discoveries +reported here enlarge the Oligocene record of apatemyids +to include not only the type specimen of <i>Sinclairella dakotensis</i>, a +skull and associated mandible from South Dakota, but also seven +isolated teeth, representing at least two individuals, from a Chadronian +fossil locality in Nebraska and one specimen from each of +two Orellan fossil localities in northeastern Colorado. Simpson +(1944:73, and 1953:127) presented tabulations of the published records +of American apatemyids and suggested the data indicated the +populations of these mammals were of small size throughout the +history of the family. The few pre-Oligocene occurrences of apatemyids +described subsequently (note McKenna, 1960, figs. 3-10, +and p. 48) and occurrences described here tend to reinforce Simpson's +interpretation. This interpretation may have to be modified +to some degree, however, when current studies of collections of pre-Oligocene +apatemyids are completed (McKenna, personal communication).</p> + +<p>Although information concerning the evolutionary trends of American +apatemyids has been published, no data on the morphological +variation in a population are available in the literature. An adequate +basis for evaluating the significance of the morphological differences +between the P4's of Princeton no. 13585 and KU no. 12110 coupled +with the similarities of their M1-2's is lacking. In the evolution of +American apatemyids the P4 underwent reduction in size and, apparently, +curtailment of function. This history suggests the range +of morphological variation of P4 in populations of <i>Sinclairella dakotensis</i> +could be expected to be greater than that of the molars +and encompass the morphological differences between the P4's of +Princeton no. 13585 and KU no. 12110. The difference in age of +the Chadronian and Orellan fossils does not constitute proof that +they pertain to different species. Although the identification is admittedly<span class="pagenum"><a name="Page_490" id="Page_490">[Pg 490]</a></span> +provisional until more fossils including other parts of the +skeleton are discovered, the Orellan fossils described here are referred +to <i>Sinclairella dakotensis</i>.</p> + + + + + +<div class="center"><span class="smcap">Table 1.—Measurements (in millimeters) of Teeth of Sinclairella<br /> +dakotensis Jepsen.</span><br /><br /> + +<table border="1" cellpadding="4" cellspacing="0" summary="teeth"> +<tr><td align="left"> </td><th colspan="2">P4</th><th colspan="2">M1</th><th colspan="2">M2</th></tr> +<tr><td align="left"> </td><td align="center">length</td><td align="center">width</td><td align="center">length<a name="FNanchor_1_2" id="FNanchor_1_2"></a><a href="#Footnote_1_1" class="fnanchor">[1]</a></td><td align="center">width<a name="FNanchor_1_3" id="FNanchor_1_3"></a><a href="#Footnote_1_1" class="fnanchor">[1]</a></td><td align="center">length<a name="FNanchor_1_4" id="FNanchor_1_4"></a><a href="#Footnote_1_1" class="fnanchor">[1]</a></td><td align="center">width<a name="FNanchor_1_1" id="FNanchor_1_1"></a><a href="#Footnote_1_1" class="fnanchor">[1]</a></td></tr> +<tr><td align="left">Princeton no. 13585<a name="FNanchor_2_2" id="FNanchor_2_2"></a><a href="#Footnote_2_2" class="fnanchor">[2]</a></td><td align="center"> 2.1</td><td align="center"> 1.1</td><td align="center"> 4.0</td><td align="center"> 3.7</td><td align="center"> 3.4</td><td align="center"> 4.7</td></tr> +<tr><td align="left">RAM no. 381</td><td align="center"></td><td align="center"></td><td align="center"> 4.1</td><td align="center"> 3.5</td><td align="center"> </td><td align="center"> </td></tr> +<tr><td align="left">RAM no. 1674</td><td align="center"></td><td align="center"></td><td align="center"></td><td align="center"></td><td align="center"> 3.4</td><td align="center"> 4.2</td></tr> +<tr><td align="left">KU no. 11210</td><td align="center"> 2.4</td><td align="center"> 1.6</td><td align="center"> 3.9</td><td align="center"> 3.5</td><td align="center"> 3.8</td><td align="center"> 4.1+</td></tr> +<tr><td align="left">UCM no. 21073</td><td align="center"></td><td align="center"></td><td align="center"></td><td align="center"></td><td align="center"> 3.6</td><td align="center"> 4.1</td></tr> +<tr><td colspan="3" rowspan="2"> </td><td colspan="2">m1</td><td colspan="2">m2</td></tr> +<tr><td>length</td><td>width</td><td>length</td><td>width</td></tr> +<tr><td colspan="3" align="left">Princeton no. 13585<a name="FNanchor_3_3" id="FNanchor_3_3"></a><a href="#Footnote_3_3" class="fnanchor">[3]</a></td><td align="center"> 3.5</td><td align="center"> 2.4</td><td align="center"> 3.7</td><td align="center"> 2.8</td></tr> +<tr><td colspan="3" align="left">RAM no. 1000</td><td align="center"> 3.5</td><td align="center"> 2.2</td><td align="center"> </td><td align="center"> </td></tr> +<tr><td colspan="3" align="left">RAM no. 598</td><td align="center"></td><td align="center"></td><td align="center"> 3.8</td><td align="center"> 2.6</td></tr> +<tr><td colspan="3" align="left">RAM no. 1001</td><td align="center"></td><td align="center"></td><td align="center"> 3.6+</td><td align="center"> 2.6</td></tr> +<tr><td colspan="3" align="left">RAM no. 1079</td><td align="center"></td><td align="center"></td><td align="center"> 4.0</td><td align="center"> 2.8</td></tr> +<tr><td colspan="3" align="left">RAM no. 3013</td><td align="center"></td><td align="center"></td><td align="center"> 3.6</td><td align="center"> 2.8</td></tr> +</table></div> + + +<div class="footnote"><p><a name="Footnote_1_1" id="Footnote_1_1"></a><a href="#FNanchor_1_1"><span class="label">[1]</span></a> Length defined as maximum dimension of the labial half of the crown measured +parallel to a line drawn through the apices of paracone and metacone. Width defined as +maximum coronal dimension measured along line perpendicular to line defined by apices of +paracone and metacone.</p></div> + +<div class="footnote"><p><a name="Footnote_2_2" id="Footnote_2_2"></a><a href="#FNanchor_2_2"><span class="label">[2]</span></a> Dimensions provided by Dr. Glenn L. Jepsen.</p></div> + +<div class="footnote"><p><a name="Footnote_3_3" id="Footnote_3_3"></a><a href="#FNanchor_3_3"><span class="label">[3]</span></a> Dimensions taken from Jepsen (1934:300).</p></div> + + + +<p><span class="pagenum"><a name="Page_491" id="Page_491">[Pg 491]</a></span></p><hr style="width: 65%;" /> +<h2>Literature Cited</h2> + +<div class="blockquot"><p><span class="smcap">Alf, R.</span></p> + +<p><span class="i4">1962. A new species of the rodent <i>Pipestoneomys</i> from the Oligocene of +Nebraska. Breviora, Mus. Comp. Zool., no. 172, pp. 1-7, 3 figs.</span></p> + +<p><span class="smcap">Galbreath, E. C.</span></p> + +<p><span class="i4">1953. A contribution to the Tertiary geology and paleontology of northeastern +Colorado. Univ. Kansas Paleont. Cont., Vertebrata, art. 4, +pp. 1-120, 2 pls., 26 figs.</span></p> + +<p><span class="smcap">Hough, J.</span>, and <span class="smcap">Alf, R.</span></p> + +<p><span class="i4">1958. A Chadron mammalian fauna from Nebraska. Journ. Paleon. 30: +132-140, 4 figs.</span></p> + +<p><span class="smcap">Jepsen, G. L.</span></p> + +<p><span class="i4">1934. A revision of the American Apatemyidae and the description of a +new genus, <i>Sinclairella</i>, from the White River Oligocene of South +Dakota. Proc. Amer. Philos. Soc., 74:287-305, 3 pls., 4 figs.</span></p> + +<p><span class="smcap">McKenna, M. C.</span></p> + +<p><span class="i4">1960. Fossil Mammalia from the early Wasatchian Four Mile fauna, +Eocene of northwest Colorado. Univ. California Publ. in Geol. +Sci., 37:1-130, 64 figs.</span></p> + +<p><span class="smcap">Matthew, W. D.</span></p> + +<p><span class="i4">1909. The Carnivora and Insectivora of the Bridger Basin, Middle +Eocene. Mem. Amer. Mus. Nat. Hist., 9:289-567, pls. 42-52, 118 +figs.</span></p> + +<p><span class="smcap">Patterson, B.</span> and <span class="smcap">McGrew, P. O.</span></p> + +<p><span class="i4">1937. A soricid and two erinaceids from the White River Oligocene. Geol. +Ser., Field Mus. Nat. Hist., 6:245-272, figs. 60-74.</span></p> + +<p><span class="smcap">Scott, W. B.</span> and <span class="smcap">Jepsen, G. L.</span></p> + +<p><span class="i4">1936. The mammalian fauna of the White River Oligocene—Part I. Insectivora +and Carnivora. Trans. Amer. Philos. Soc., n. s., 28:1-153, +22 pls., 7 figs.</span></p> + +<p><span class="smcap">Simpson, G. G.</span></p> + +<p><span class="i4">1944. Tempo and mode in evolution. New York: Columbia Univ. Press, +xviii + 237 pp., 36 figs.</span></p> + +<p><span class="i4">1953. The major features of evolution. New York: Columbia Univ. +Press, xx + 434 pp., 52 figs.</span></p> + +<p><i>Transmitted June 24, 1963.</i></p></div> + +<div>*** END OF THE PROJECT GUTENBERG EBOOK 30297 ***</div> +</body> +</html> diff --git a/30297-h/images/image005.png b/30297-h/images/image005.png Binary files differnew file mode 100644 index 0000000..a1e4620 --- /dev/null +++ b/30297-h/images/image005.png diff --git a/30297-h/images/image006.png b/30297-h/images/image006.png Binary files differnew file mode 100644 index 0000000..f691f3f --- /dev/null +++ b/30297-h/images/image006.png diff --git a/30297.txt b/30297.txt new file mode 100644 index 0000000..71b1815 --- /dev/null +++ b/30297.txt @@ -0,0 +1,795 @@ +The Project Gutenberg EBook of Records of the Fossil Mammal Sinclairella, +Family Apatemyidae, From the Chadronian and Orellan, by William A. Clemens + +This eBook is for the use of anyone anywhere at no cost and with +almost no restrictions whatsoever. You may copy it, give it away or +re-use it under the terms of the Project Gutenberg License included +with this eBook or online at www.gutenberg.org + + +Title: Records of the Fossil Mammal Sinclairella, Family Apatemyidae, From the Chadronian and Orellan + +Author: William A. Clemens + +Release Date: October 20, 2009 [EBook #30297] + +Language: English + +Character set encoding: ASCII + +*** START OF THIS PROJECT GUTENBERG EBOOK FOSSIL MAMMAL SINCLAIRELLA *** + + + + +Produced by Chris Curnow, Joseph Cooper, Diane Monico, and +the Online Distributed Proofreading Team at +https://www.pgdp.net + + + + + + + + + + + +UNIVERSITY OF KANSAS PUBLICATIONS + +MUSEUM OF NATURAL HISTORY + +Volume 14, No. 17, pp. 483-491, 2 figs. +March 2, 1964 + + +Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, +From the Chadronian and Orellan + + +BY + +WILLIAM A. CLEMENS, JR. + + +UNIVERSITY OF KANSAS +LAWRENCE +1964 + + +UNIVERSITY OF KANSAS PUBLICATIONS, MUSEUM OF NATURAL HISTORY + +Editors: E. Raymond Hall, Chairman, Henry S. Fitch, +Theodore H. Eaton, Jr. + +Volume 14, No. 17, pp. 483-491, 2 figs. +Published March 2, 1964 + +UNIVERSITY OF KANSAS +Lawrence, Kansas + +PRINTED BY +HARRY (BUD) TIMBERLAKE, STATE PRINTER +TOPEKA, KANSAS + +1964 + + +29-8587 + + + + +Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, +From the Chadronian and Orellan + +BY + +WILLIAM A. CLEMENS, JR. + + +Introduction + + +The family Apatemyidae has a long geochronological range in North +America, beginning in the Torrejonian land-mammal age, but is +represented by a relatively small number of fossils found at a few +localities. Two fossils of Orellan age, found in northeastern Colorado +and described here, demonstrate that the geochronological range of the +Apatemyidae extends into the Middle Oligocene. Isolated teeth of +_Sinclairella dakotensis_ Jepsen, part of a sample of a Chadronian +local fauna collected by field parties from the Webb School of +California, are also described. + + I thank Mr. Raymond M. Alf, Webb School of California, + Claremont, California, and Dr. Peter Robinson, University of + Colorado Museum, Boulder, Colorado, for permitting me to + describe the fossils they discovered. Also Dr. Robinson made + available the draft of a short paper he had prepared on the + tooth found in Weld County, Colorado; his work was + facilitated by a grant from the University of Colorado + Council on Research and Creative Work. I also gratefully + acknowledge receipt of critical data and valuable comments + from Drs. Edwin C. Galbreath, Glenn L. Jepsen, and Malcolm + C. McKenna who is currently revising the Paleocene + apatemyids and studying the phylogenetic relationships of + the family. The prefixes of catalogue numbers used in the + text identify fossils in the collections of the following + institutions: KU, Museum of Natural History, The University + of Kansas, Lawrence; Princeton, Princeton Museum, Princeton, + New Jersey; RAM-UCR, Raymond Alf Museum, Webb School of + California, Claremont, California (the permanent repository + for these specimens will be the University of California, + Riverside); and UCM, University of Colorado Museum, Boulder, + Colorado. The system of notations for teeth prescribed for + use here is as follows: teeth in the upper half of the + dentition are designated by a capital letter and a number; + thus M2 is the notation for the upper second molar; teeth in + the lower half of the dentition are designated by a + lower-case letter and a number; thus p2 is the notation for + the lower second premolar. + + + + +Family APATEMYIDAE Matthew, 1909 + +Genus =Sinclairella= Jepsen, 1934 + +=Sinclairella dakotensis= Jepsen, 1934 + + +The type of the species, Princeton no. 13585, was discovered in +Chadronian strata of the upper part of the Chadron Formation cropping +out in Big Corral Draw, approximately 13 miles south-southwest of +Scenic, in southwestern South Dakota (Jepsen, 1934, p. 291). Detailed +descriptions of the type specimen are given in papers by Jepsen (1934) +and Scott and Jepsen (1936). Isolated teeth of Chadronian age referable +to _Sinclairella dakotensis_ have been discovered subsequently at a +locality in Nebraska and fossils of Orellan age, also referable to _S. +dakotensis_, have been collected at two localities in Colorado. The +sample from each locality is described separately. + + +Sioux County, northwestern Nebraska + + _Material._--RAM-UCR nos. 381, left M1; 598, left m2; 1000, + right m1; 1001, right m2; 1079, right m2; 1674, right M2; + and 3013, left m2. + + _Locality and stratigraphy._--These Chadronian fossils were + discovered by Raymond Alf and members of his field parties + in several harvester ant mounds built in exposures of the + Chadron Formation in Sec. 26, T 33 N, R 53 W, Sioux County, + Nebraska (Alf, 1962, and Hough and Alf, 1958). This is UCR + locality V5403. The collectors carefully considered the + possibility that some of the fossils found in the ant mounds + were collected from younger strata by the harvester ants and + concluded this was unlikely (Alf, personal communication). + + _Description and comments._--The cusps of RAM-UCR no. 381, a + left M1, are sharp and the wear-facets resulting from + occlusion with the lower dentition are small. The paraconule + is a low, ill-defined cusp on the anterior margin of the + crown; a metaconule is not present. A smooth stylar shelf is + present labial to the metacone. The crown was supported by + three roots. There are no interradicular crests. + + The crown of RAM-UCR no. 1674, a right M2, is heavily + abraded and many morphological details of the cusps have + been destroyed. Low interradicular crests linked the three + roots of the tooth with a low, central prominence. As was + the case with RAM-UCR no. 381, no significant differences + could be found in comparisons with illustrations of the + teeth preserved in Princeton no. 13585. + + RAM-UCR nos. 598, 1001, 1079, and 3013 all appear to be + m2's. The talonids of these teeth are not elongated, their + trigonids have quadrilateral outlines, and the paraconids + are small but prominent, bladelike cusps. The trigonid of + RAM-UCR 1000 is elongated and the paraconid is a minute + cusp; the tooth closely resembles the m1 of the type of + _Sinclairella dakotensis_. + + +Logan County, northeastern Colorado + + _Material._--KU no. 11210 (fig. 1), a fragment of a left + maxillary containing P4 and M1-2. + + _Locality and stratigraphy._--The fossil was found in the + center of the W-1/2, Sec. 21, T 11 N, R 53 W, Logan County, + Colorado, "... in the bed below _Agnotocastor_ bed, Cedar + Creek Member...." (Ronald H. Pine, 1958, field notes on file + at the University of Kansas). The bed so defined is part of + unit 3 in the lower division of the Cedar Creek Member, as + subdivided by Galbreath (1953:25) in stratigraphic section + XII. The fauna obtained from unit 3 is of Orellan age. + +[Illustration: FIG. 1. _Sinclairella dakotensis_ Jepsen, KU no. 11210, +fragment of left maxillary with P4 and M1-2; Orellan, Logan County, +Colorado; drawings by Mrs. Judith Hood: a, labial view; b, occlusal +view; both approximately x 9.] + + _Description and comments._--P4 of KU no. 11210 has a large + posterolingual cusp separated from the main cusp by a + distinct groove, which deepens posteriorly. The + posterolingual cusp is supported by the broad posterior + root. P4 of the type specimen of _Sinclairella dakotensis_ + is described (Jepsen, 1934, p. 392) as having an oval + outline at the base of the crown, and a small, + posterolingual cusp. A chip of enamel is missing from the + posterior slope of the main cusp of the P4 of KU no. 11210. + The anterior slope of the main cusp is flattened, possibly + the result of wear, and there is no evidence of a groove + like that present on the P4 of the type specimen. + + Only a few differences were found between the molars + preserved in KU no. 11210 and their counterparts in the type + specimen. A stylar shelf is present labial to the metacone + of M1 of KU no. 11210, but, unlike the type, its surface is + smooth and there is no evidence of cusps. Of the three small + stylar cusps on the stylar shelf of M2 the smallest is in + the position of a mesostyle. The M2 lacks a chip of enamel + from the lingual surface of the hypocone. Unlike the M2 of + Princeton no. 13585, in occlusal view the posterior margin + of the M2 of KU no. 11210 is convex posterior to the + metacone. The anterior edge of the base of the zygomatic + arch of KU no. 11210 was dorsal to M2. The shallow oval + depression in the maxillary dorsal to M1 might be the result + of post-mortem distortion. + + The molars preserved in KU no. 11210 and their counterparts + in the type specimen do not appear to be significantly + different in size (table 1) or morphology of the cusps. The + only difference between the two specimens that might be of + classificatory significance is the difference in size of the + posterolingual cusp of P4. At present the range of + intraspecific variation in the morphology of P4 has not been + documented for any species of apatemyid. The evolutionary + trend or trends of the apatemyids (McKenna, 1960, p. 48) for + progressive reduction of function of p4 probably were + paralleled by similar trends in the evolution of the P4. If + so, the intraspecific variation in the morphology of P4 + could be expected to be somewhat greater than that of the + upper molars, for example. The morphological difference + between the P4's of the type of _Sinclairella dakotensis_ + and KU no. 11210 is not extreme and does not exceed the + range of intraspecific variation that could be expected for + this element of the dentition. The close resemblances in + size and morphology between the M1-2 of Princeton no. 13585 + and KU no. 11210 also favor identification of the latter as + part of a member of an Orellan population of _Sinclairella + dakotensis_. + + +Weld County, northeastern Colorado + +[Illustration: FIG. 2. _Sinclairella dakotensis_ Jepsen, UCM no. 21073, +right M2; Orellan, Weld County, Colorado; drawing by Mrs. Judith Hood: +occlusal view, approximately x 9.] + + _Material._--UCM no. 20173 (fig. 2), is a right M2. + + _Locality and stratigraphy._--The tooth was discovered at + the Mellinger locality, Sec. 17, T 11 N, R 65 W, Weld + County, Colorado. The Mellinger locality is in the Cedar + Creek Member, White River Formation, and its fauna is + considered to be of Orellan age (Patterson and McGrew, 1937, + and Galbreath, 1953). + + _Description and comments._--UCM no. 21073, which is more + heavily abraded than KU no. 11210, shows no evidence of a + stylar cusp either anterolabial to the metacone or in the + position of a mesostyle. A small stylar cusp is present + anterolabial to the paracone. A notch that appears to have + been cut through the enamel of the posterolabial corner of + the crown could have received the parastylar apex of M3. A + similar notch is not present on the M2 of KU no. 11210 nor + indicated in the illustrations of the M2 of Princeton no. + 13585. The coronal dimensions of UCM no. 21073 (table 1) do + not appear to differ significantly from those of the M2's of + KU no. 11210 and the type specimen of _Sinclairella + dakotensis_. + + +Comments + +With the discovery of Orellan apatemyids the geochronological range of +the family in North America is shown to extend from the Torrejonian +through the Orellan land-mammal ages. The discoveries reported here +enlarge the Oligocene record of apatemyids to include not only the type +specimen of _Sinclairella dakotensis_, a skull and associated mandible +from South Dakota, but also seven isolated teeth, representing at least +two individuals, from a Chadronian fossil locality in Nebraska and one +specimen from each of two Orellan fossil localities in northeastern +Colorado. Simpson (1944:73, and 1953:127) presented tabulations of the +published records of American apatemyids and suggested the data +indicated the populations of these mammals were of small size +throughout the history of the family. The few pre-Oligocene occurrences +of apatemyids described subsequently (note McKenna, 1960, figs. 3-10, +and p. 48) and occurrences described here tend to reinforce Simpson's +interpretation. This interpretation may have to be modified to some +degree, however, when current studies of collections of pre-Oligocene +apatemyids are completed (McKenna, personal communication). + +Although information concerning the evolutionary trends of American +apatemyids has been published, no data on the morphological variation +in a population are available in the literature. An adequate basis for +evaluating the significance of the morphological differences between +the P4's of Princeton no. 13585 and KU no. 12110 coupled with the +similarities of their M1-2's is lacking. In the evolution of American +apatemyids the P4 underwent reduction in size and, apparently, +curtailment of function. This history suggests the range of +morphological variation of P4 in populations of _Sinclairella +dakotensis_ could be expected to be greater than that of the molars and +encompass the morphological differences between the P4's of Princeton +no. 13585 and KU no. 12110. The difference in age of the Chadronian and +Orellan fossils does not constitute proof that they pertain to +different species. Although the identification is admittedly +provisional until more fossils including other parts of the skeleton +are discovered, the Orellan fossils described here are referred to +_Sinclairella dakotensis_. + + +TABLE 1.--MEASUREMENTS (IN MILLIMETERS) OF TEETH OF SINCLAIRELLA +DAKOTENSIS JEPSEN. + +========================================================================== + | P4 | M1 | M2 +-----------------------+------------+------------------+------------------ + |length|width|length[1]|width[1]|length[1]|width[1] +-----------------------+------+-----+---------+--------+---------+-------- +Princeton no. 13585[2] | 2.1 | 1.1 | 4.0 | 3.7 | 3.4 | 4.7 +RAM no. 381 | | | 4.1 | 3.5 | | +RAM no. 1674 | | | | | 3.4 | 4.2 +KU no. 11210 | 2.4 | 1.6 | 3.9 | 3.5 | 3.8 | 4.1+ +UCM no. 21073 | | | | | 3.6 | 4.1 +-----------------------+------+-----+---------+--------+---------+-------- + | m1 | m2 + +---------+--------+---------+-------- + | length | width | length | width + +---------+--------+---------+-------- +Princeton no. 13585[3] | 3.5 | 2.4 | 3.7 | 2.8 +RAM no. 1000 | 3.5 | 2.2 | | +RAM no. 598 | | | 3.8 | 2.6 +RAM no. 1001 | | | 3.6+ | 2.6 +RAM no. 1079 | | | 4.0 | 2.8 +RAM no. 3013 | | | 3.6 | 2.8 +------------------------------------+---------+--------+---------+-------- + +[Footnote 1: Length defined as maximum dimension of the labial half of +the crown measured parallel to a line drawn through the apices of +paracone and metacone. Width defined as maximum coronal dimension +measured along line perpendicular to line defined by apices of paracone +and metacone.] + +[Footnote 2: Dimensions provided by Dr. Glenn L. Jepsen.] + +[Footnote 3: Dimensions taken from Jepsen (1934:300).] + + + + +Literature Cited + +ALF, R. + 1962. A new species of the rodent _Pipestoneomys_ from the + Oligocene of Nebraska. Breviora, Mus. Comp. Zool., no. 172, + pp. 1-7, 3 figs. + +GALBREATH, E. C. + 1953. A contribution to the Tertiary geology and paleontology + of northeastern Colorado. Univ. Kansas Paleont. Cont., + Vertebrata, art. 4, pp. 1-120, 2 pls., 26 figs. + +HOUGH, J., and ALF, R. + 1958. A Chadron mammalian fauna from Nebraska. Journ. Paleon. + 30:132-140, 4 figs. + +JEPSEN, G. L. + 1934. A revision of the American Apatemyidae and the description + of a new genus, _Sinclairella_, from the White River + Oligocene of South Dakota. Proc. Amer. Philos. Soc., + 74:287-305, 3 pls., 4 figs. + +MCKENNA, M. C. + 1960. Fossil Mammalia from the early Wasatchian Four Mile fauna, + Eocene of northwest Colorado. Univ. California Publ. in + Geol. Sci., 37:1-130, 64 figs. + +MATTHEW, W. D. + 1909. The Carnivora and Insectivora of the Bridger Basin, Middle + Eocene. Mem. Amer. Mus. Nat. Hist., 9:289-567, pls. 42-52, + 118 figs. + +PATTERSON, B. and MCGREW, P. O. + 1937. A soricid and two erinaceids from the White River Oligocene. + Geol. Ser., Field Mus. Nat. Hist., 6:245-272, figs. 60-74. + +SCOTT, W. B. and JEPSEN, G. L. + 1936. The mammalian fauna of the White River Oligocene--Part I. + Insectivora and Carnivora. Trans. Amer. Philos. Soc., n. s., + 28:1-153, 22 pls., 7 figs. + +SIMPSON, G. G. + 1944. Tempo and mode in evolution. New York: Columbia Univ. Press, + xviii + 237 pp., 36 figs. + + 1953. The major features of evolution. New York: Columbia Univ. + Press, xx + 434 pp., 52 figs. + +_Transmitted June 24, 1963._ + + + + + +End of the Project Gutenberg EBook of Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, From the Chadronian and Orellan, by William A. Clemens + +*** END OF THIS PROJECT GUTENBERG EBOOK FOSSIL MAMMAL SINCLAIRELLA *** + +***** This file should be named 30297.txt or 30297.zip ***** +This and all associated files of various formats will be found in: + https://www.gutenberg.org/3/0/2/9/30297/ + +Produced by Chris Curnow, Joseph Cooper, Diane Monico, and +the Online Distributed Proofreading Team at +https://www.pgdp.net + + +Updated editions will replace the previous one--the old editions +will be renamed. + +Creating the works from public domain print editions means that no +one owns a United States copyright in these works, so the Foundation +(and you!) can copy and distribute it in the United States without +permission and without paying copyright royalties. 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Anyone seeking to utilize +this eBook outside of the United States should confirm copyright +status under the laws that apply to them. diff --git a/README.md b/README.md new file mode 100644 index 0000000..9e44300 --- /dev/null +++ b/README.md @@ -0,0 +1,2 @@ +Project Gutenberg (https://www.gutenberg.org) public repository for +eBook #30297 (https://www.gutenberg.org/ebooks/30297) diff --git a/old/30297-8.txt b/old/30297-8.txt new file mode 100644 index 0000000..5b06997 --- /dev/null +++ b/old/30297-8.txt @@ -0,0 +1,795 @@ +The Project Gutenberg EBook of Records of the Fossil Mammal Sinclairella, +Family Apatemyidae, From the Chadronian and Orellan, by William A. Clemens + +This eBook is for the use of anyone anywhere at no cost and with +almost no restrictions whatsoever. You may copy it, give it away or +re-use it under the terms of the Project Gutenberg License included +with this eBook or online at www.gutenberg.org + + +Title: Records of the Fossil Mammal Sinclairella, Family Apatemyidae, From the Chadronian and Orellan + +Author: William A. Clemens + +Release Date: October 20, 2009 [EBook #30297] + +Language: English + +Character set encoding: ISO-8859-1 + +*** START OF THIS PROJECT GUTENBERG EBOOK FOSSIL MAMMAL SINCLAIRELLA *** + + + + +Produced by Chris Curnow, Joseph Cooper, Diane Monico, and +the Online Distributed Proofreading Team at +https://www.pgdp.net + + + + + + + + + + + +UNIVERSITY OF KANSAS PUBLICATIONS + +MUSEUM OF NATURAL HISTORY + +Volume 14, No. 17, pp. 483-491, 2 figs. +March 2, 1964 + + +Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, +From the Chadronian and Orellan + + +BY + +WILLIAM A. CLEMENS, JR. + + +UNIVERSITY OF KANSAS +LAWRENCE +1964 + + +UNIVERSITY OF KANSAS PUBLICATIONS, MUSEUM OF NATURAL HISTORY + +Editors: E. Raymond Hall, Chairman, Henry S. Fitch, +Theodore H. Eaton, Jr. + +Volume 14, No. 17, pp. 483-491, 2 figs. +Published March 2, 1964 + +UNIVERSITY OF KANSAS +Lawrence, Kansas + +PRINTED BY +HARRY (BUD) TIMBERLAKE, STATE PRINTER +TOPEKA, KANSAS + +1964 + + +29-8587 + + + + +Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, +From the Chadronian and Orellan + +BY + +WILLIAM A. CLEMENS, JR. + + +Introduction + + +The family Apatemyidae has a long geochronological range in North +America, beginning in the Torrejonian land-mammal age, but is +represented by a relatively small number of fossils found at a few +localities. Two fossils of Orellan age, found in northeastern Colorado +and described here, demonstrate that the geochronological range of the +Apatemyidae extends into the Middle Oligocene. Isolated teeth of +_Sinclairella dakotensis_ Jepsen, part of a sample of a Chadronian +local fauna collected by field parties from the Webb School of +California, are also described. + + I thank Mr. Raymond M. Alf, Webb School of California, + Claremont, California, and Dr. Peter Robinson, University of + Colorado Museum, Boulder, Colorado, for permitting me to + describe the fossils they discovered. Also Dr. Robinson made + available the draft of a short paper he had prepared on the + tooth found in Weld County, Colorado; his work was + facilitated by a grant from the University of Colorado + Council on Research and Creative Work. I also gratefully + acknowledge receipt of critical data and valuable comments + from Drs. Edwin C. Galbreath, Glenn L. Jepsen, and Malcolm + C. McKenna who is currently revising the Paleocene + apatemyids and studying the phylogenetic relationships of + the family. The prefixes of catalogue numbers used in the + text identify fossils in the collections of the following + institutions: KU, Museum of Natural History, The University + of Kansas, Lawrence; Princeton, Princeton Museum, Princeton, + New Jersey; RAM-UCR, Raymond Alf Museum, Webb School of + California, Claremont, California (the permanent repository + for these specimens will be the University of California, + Riverside); and UCM, University of Colorado Museum, Boulder, + Colorado. The system of notations for teeth prescribed for + use here is as follows: teeth in the upper half of the + dentition are designated by a capital letter and a number; + thus M2 is the notation for the upper second molar; teeth in + the lower half of the dentition are designated by a + lower-case letter and a number; thus p2 is the notation for + the lower second premolar. + + + + +Family APATEMYIDAE Matthew, 1909 + +Genus =Sinclairella= Jepsen, 1934 + +=Sinclairella dakotensis= Jepsen, 1934 + + +The type of the species, Princeton no. 13585, was discovered in +Chadronian strata of the upper part of the Chadron Formation cropping +out in Big Corral Draw, approximately 13 miles south-southwest of +Scenic, in southwestern South Dakota (Jepsen, 1934, p. 291). Detailed +descriptions of the type specimen are given in papers by Jepsen (1934) +and Scott and Jepsen (1936). Isolated teeth of Chadronian age referable +to _Sinclairella dakotensis_ have been discovered subsequently at a +locality in Nebraska and fossils of Orellan age, also referable to _S. +dakotensis_, have been collected at two localities in Colorado. The +sample from each locality is described separately. + + +Sioux County, northwestern Nebraska + + _Material._--RAM-UCR nos. 381, left M1; 598, left m2; 1000, + right m1; 1001, right m2; 1079, right m2; 1674, right M2; + and 3013, left m2. + + _Locality and stratigraphy._--These Chadronian fossils were + discovered by Raymond Alf and members of his field parties + in several harvester ant mounds built in exposures of the + Chadron Formation in Sec. 26, T 33 N, R 53 W, Sioux County, + Nebraska (Alf, 1962, and Hough and Alf, 1958). This is UCR + locality V5403. The collectors carefully considered the + possibility that some of the fossils found in the ant mounds + were collected from younger strata by the harvester ants and + concluded this was unlikely (Alf, personal communication). + + _Description and comments._--The cusps of RAM-UCR no. 381, a + left M1, are sharp and the wear-facets resulting from + occlusion with the lower dentition are small. The paraconule + is a low, ill-defined cusp on the anterior margin of the + crown; a metaconule is not present. A smooth stylar shelf is + present labial to the metacone. The crown was supported by + three roots. There are no interradicular crests. + + The crown of RAM-UCR no. 1674, a right M2, is heavily + abraded and many morphological details of the cusps have + been destroyed. Low interradicular crests linked the three + roots of the tooth with a low, central prominence. As was + the case with RAM-UCR no. 381, no significant differences + could be found in comparisons with illustrations of the + teeth preserved in Princeton no. 13585. + + RAM-UCR nos. 598, 1001, 1079, and 3013 all appear to be + m2's. The talonids of these teeth are not elongated, their + trigonids have quadrilateral outlines, and the paraconids + are small but prominent, bladelike cusps. The trigonid of + RAM-UCR 1000 is elongated and the paraconid is a minute + cusp; the tooth closely resembles the m1 of the type of + _Sinclairella dakotensis_. + + +Logan County, northeastern Colorado + + _Material._--KU no. 11210 (fig. 1), a fragment of a left + maxillary containing P4 and M1-2. + + _Locality and stratigraphy._--The fossil was found in the + center of the W-1/2, Sec. 21, T 11 N, R 53 W, Logan County, + Colorado, "... in the bed below _Agnotocastor_ bed, Cedar + Creek Member...." (Ronald H. Pine, 1958, field notes on file + at the University of Kansas). The bed so defined is part of + unit 3 in the lower division of the Cedar Creek Member, as + subdivided by Galbreath (1953:25) in stratigraphic section + XII. The fauna obtained from unit 3 is of Orellan age. + +[Illustration: FIG. 1. _Sinclairella dakotensis_ Jepsen, KU no. 11210, +fragment of left maxillary with P4 and M1-2; Orellan, Logan County, +Colorado; drawings by Mrs. Judith Hood: a, labial view; b, occlusal +view; both approximately × 9.] + + _Description and comments._--P4 of KU no. 11210 has a large + posterolingual cusp separated from the main cusp by a + distinct groove, which deepens posteriorly. The + posterolingual cusp is supported by the broad posterior + root. P4 of the type specimen of _Sinclairella dakotensis_ + is described (Jepsen, 1934, p. 392) as having an oval + outline at the base of the crown, and a small, + posterolingual cusp. A chip of enamel is missing from the + posterior slope of the main cusp of the P4 of KU no. 11210. + The anterior slope of the main cusp is flattened, possibly + the result of wear, and there is no evidence of a groove + like that present on the P4 of the type specimen. + + Only a few differences were found between the molars + preserved in KU no. 11210 and their counterparts in the type + specimen. A stylar shelf is present labial to the metacone + of M1 of KU no. 11210, but, unlike the type, its surface is + smooth and there is no evidence of cusps. Of the three small + stylar cusps on the stylar shelf of M2 the smallest is in + the position of a mesostyle. The M2 lacks a chip of enamel + from the lingual surface of the hypocone. Unlike the M2 of + Princeton no. 13585, in occlusal view the posterior margin + of the M2 of KU no. 11210 is convex posterior to the + metacone. The anterior edge of the base of the zygomatic + arch of KU no. 11210 was dorsal to M2. The shallow oval + depression in the maxillary dorsal to M1 might be the result + of post-mortem distortion. + + The molars preserved in KU no. 11210 and their counterparts + in the type specimen do not appear to be significantly + different in size (table 1) or morphology of the cusps. The + only difference between the two specimens that might be of + classificatory significance is the difference in size of the + posterolingual cusp of P4. At present the range of + intraspecific variation in the morphology of P4 has not been + documented for any species of apatemyid. The evolutionary + trend or trends of the apatemyids (McKenna, 1960, p. 48) for + progressive reduction of function of p4 probably were + paralleled by similar trends in the evolution of the P4. If + so, the intraspecific variation in the morphology of P4 + could be expected to be somewhat greater than that of the + upper molars, for example. The morphological difference + between the P4's of the type of _Sinclairella dakotensis_ + and KU no. 11210 is not extreme and does not exceed the + range of intraspecific variation that could be expected for + this element of the dentition. The close resemblances in + size and morphology between the M1-2 of Princeton no. 13585 + and KU no. 11210 also favor identification of the latter as + part of a member of an Orellan population of _Sinclairella + dakotensis_. + + +Weld County, northeastern Colorado + +[Illustration: FIG. 2. _Sinclairella dakotensis_ Jepsen, UCM no. 21073, +right M2; Orellan, Weld County, Colorado; drawing by Mrs. Judith Hood: +occlusal view, approximately × 9.] + + _Material._--UCM no. 20173 (fig. 2), is a right M2. + + _Locality and stratigraphy._--The tooth was discovered at + the Mellinger locality, Sec. 17, T 11 N, R 65 W, Weld + County, Colorado. The Mellinger locality is in the Cedar + Creek Member, White River Formation, and its fauna is + considered to be of Orellan age (Patterson and McGrew, 1937, + and Galbreath, 1953). + + _Description and comments._--UCM no. 21073, which is more + heavily abraded than KU no. 11210, shows no evidence of a + stylar cusp either anterolabial to the metacone or in the + position of a mesostyle. A small stylar cusp is present + anterolabial to the paracone. A notch that appears to have + been cut through the enamel of the posterolabial corner of + the crown could have received the parastylar apex of M3. A + similar notch is not present on the M2 of KU no. 11210 nor + indicated in the illustrations of the M2 of Princeton no. + 13585. The coronal dimensions of UCM no. 21073 (table 1) do + not appear to differ significantly from those of the M2's of + KU no. 11210 and the type specimen of _Sinclairella + dakotensis_. + + +Comments + +With the discovery of Orellan apatemyids the geochronological range of +the family in North America is shown to extend from the Torrejonian +through the Orellan land-mammal ages. The discoveries reported here +enlarge the Oligocene record of apatemyids to include not only the type +specimen of _Sinclairella dakotensis_, a skull and associated mandible +from South Dakota, but also seven isolated teeth, representing at least +two individuals, from a Chadronian fossil locality in Nebraska and one +specimen from each of two Orellan fossil localities in northeastern +Colorado. Simpson (1944:73, and 1953:127) presented tabulations of the +published records of American apatemyids and suggested the data +indicated the populations of these mammals were of small size +throughout the history of the family. The few pre-Oligocene occurrences +of apatemyids described subsequently (note McKenna, 1960, figs. 3-10, +and p. 48) and occurrences described here tend to reinforce Simpson's +interpretation. This interpretation may have to be modified to some +degree, however, when current studies of collections of pre-Oligocene +apatemyids are completed (McKenna, personal communication). + +Although information concerning the evolutionary trends of American +apatemyids has been published, no data on the morphological variation +in a population are available in the literature. An adequate basis for +evaluating the significance of the morphological differences between +the P4's of Princeton no. 13585 and KU no. 12110 coupled with the +similarities of their M1-2's is lacking. In the evolution of American +apatemyids the P4 underwent reduction in size and, apparently, +curtailment of function. This history suggests the range of +morphological variation of P4 in populations of _Sinclairella +dakotensis_ could be expected to be greater than that of the molars and +encompass the morphological differences between the P4's of Princeton +no. 13585 and KU no. 12110. The difference in age of the Chadronian and +Orellan fossils does not constitute proof that they pertain to +different species. Although the identification is admittedly +provisional until more fossils including other parts of the skeleton +are discovered, the Orellan fossils described here are referred to +_Sinclairella dakotensis_. + + +TABLE 1.--MEASUREMENTS (IN MILLIMETERS) OF TEETH OF SINCLAIRELLA +DAKOTENSIS JEPSEN. + +========================================================================== + | P4 | M1 | M2 +-----------------------+------------+------------------+------------------ + |length|width|length[1]|width[1]|length[1]|width[1] +-----------------------+------+-----+---------+--------+---------+-------- +Princeton no. 13585[2] | 2.1 | 1.1 | 4.0 | 3.7 | 3.4 | 4.7 +RAM no. 381 | | | 4.1 | 3.5 | | +RAM no. 1674 | | | | | 3.4 | 4.2 +KU no. 11210 | 2.4 | 1.6 | 3.9 | 3.5 | 3.8 | 4.1+ +UCM no. 21073 | | | | | 3.6 | 4.1 +-----------------------+------+-----+---------+--------+---------+-------- + | m1 | m2 + +---------+--------+---------+-------- + | length | width | length | width + +---------+--------+---------+-------- +Princeton no. 13585[3] | 3.5 | 2.4 | 3.7 | 2.8 +RAM no. 1000 | 3.5 | 2.2 | | +RAM no. 598 | | | 3.8 | 2.6 +RAM no. 1001 | | | 3.6+ | 2.6 +RAM no. 1079 | | | 4.0 | 2.8 +RAM no. 3013 | | | 3.6 | 2.8 +------------------------------------+---------+--------+---------+-------- + +[Footnote 1: Length defined as maximum dimension of the labial half of +the crown measured parallel to a line drawn through the apices of +paracone and metacone. Width defined as maximum coronal dimension +measured along line perpendicular to line defined by apices of paracone +and metacone.] + +[Footnote 2: Dimensions provided by Dr. Glenn L. Jepsen.] + +[Footnote 3: Dimensions taken from Jepsen (1934:300).] + + + + +Literature Cited + +ALF, R. + 1962. A new species of the rodent _Pipestoneomys_ from the + Oligocene of Nebraska. Breviora, Mus. Comp. Zool., no. 172, + pp. 1-7, 3 figs. + +GALBREATH, E. C. + 1953. A contribution to the Tertiary geology and paleontology + of northeastern Colorado. Univ. Kansas Paleont. Cont., + Vertebrata, art. 4, pp. 1-120, 2 pls., 26 figs. + +HOUGH, J., and ALF, R. + 1958. A Chadron mammalian fauna from Nebraska. Journ. Paleon. + 30:132-140, 4 figs. + +JEPSEN, G. L. + 1934. A revision of the American Apatemyidae and the description + of a new genus, _Sinclairella_, from the White River + Oligocene of South Dakota. Proc. Amer. Philos. Soc., + 74:287-305, 3 pls., 4 figs. + +MCKENNA, M. C. + 1960. Fossil Mammalia from the early Wasatchian Four Mile fauna, + Eocene of northwest Colorado. Univ. California Publ. in + Geol. Sci., 37:1-130, 64 figs. + +MATTHEW, W. D. + 1909. The Carnivora and Insectivora of the Bridger Basin, Middle + Eocene. Mem. Amer. Mus. Nat. Hist., 9:289-567, pls. 42-52, + 118 figs. + +PATTERSON, B. and MCGREW, P. O. + 1937. A soricid and two erinaceids from the White River Oligocene. + Geol. Ser., Field Mus. Nat. Hist., 6:245-272, figs. 60-74. + +SCOTT, W. B. and JEPSEN, G. L. + 1936. The mammalian fauna of the White River Oligocene--Part I. + Insectivora and Carnivora. Trans. Amer. Philos. Soc., n. s., + 28:1-153, 22 pls., 7 figs. + +SIMPSON, G. G. + 1944. Tempo and mode in evolution. New York: Columbia Univ. Press, + xviii + 237 pp., 36 figs. + + 1953. The major features of evolution. New York: Columbia Univ. + Press, xx + 434 pp., 52 figs. + +_Transmitted June 24, 1963._ + + + + + +End of the Project Gutenberg EBook of Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, From the Chadronian and Orellan, by William A. Clemens + +*** END OF THIS PROJECT GUTENBERG EBOOK FOSSIL MAMMAL SINCLAIRELLA *** + +***** This file should be named 30297-8.txt or 30297-8.zip ***** +This and all associated files of various formats will be found in: + https://www.gutenberg.org/3/0/2/9/30297/ + +Produced by Chris Curnow, Joseph Cooper, Diane Monico, and +the Online Distributed Proofreading Team at +https://www.pgdp.net + + +Updated editions will replace the previous one--the old editions +will be renamed. + +Creating the works from public domain print editions means that no +one owns a United States copyright in these works, so the Foundation +(and you!) can copy and distribute it in the United States without +permission and without paying copyright royalties. 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Clemens, Jr. + </title> + <style type="text/css"> + + + p { margin-top: .75em; + text-align: justify; + margin-bottom: .75em; + } + h1 { text-align: center; line-height: 1.5; clear: both; } + + h2,h3 { text-align: center; clear: both; } + + p.title { text-align: center; text-indent: 0; + font-weight: bold; + line-height: 1.4; margin-bottom: 3em; } + + hr { width: 33%; + margin-top: 2em; + margin-bottom: 2em; + margin-left: auto; + margin-right: auto; + clear: both; + } + + table {margin-left: auto; margin-right: auto;} + + table, td, th {border: solid thin;} + thead, tfoot, th { font-variant: small-caps; + text-align: center; + font-weight: normal;} + + body{margin-left: 10%; + margin-right: 10%; + } + + .pagenum { /* uncomment the next line for invisible page numbers */ + /* visibility: hidden; */ + position: absolute; + left: 92%; + font-size: smaller; + text-align: right; + } /* page numbers */ + + .blockquot{margin-left: 5%; margin-right: 10%;} + .center {text-align: center;} + .smcap {font-variant: small-caps;} + + .caption {font-weight: bold;} + + .figcenter {margin: auto; text-align: center;} + + .figleft {float: left; clear: left; margin-left: 0; margin-bottom: 1em; margin-top: + 1em; margin-right: 1em; padding: 0; text-align: center;} + + .footnotes {border: dashed 1px;} + .footnote {margin-left: 10%; margin-right: 10%; font-size: 0.9em;} + .footnote .label {position: absolute; right: 84%; text-align: right;} + .fnanchor {vertical-align: super; font-size: .8em; text-decoration: none;} + + .i4 {display: block; margin-left: 2.5em; + padding-left: 2.5em; text-indent: -2.5em;} + + + </style> + </head> +<body> + + +<pre> + +The Project Gutenberg EBook of Records of the Fossil Mammal Sinclairella, +Family Apatemyidae, From the Chadronian and Orellan, by William A. Clemens + +This eBook is for the use of anyone anywhere at no cost and with +almost no restrictions whatsoever. You may copy it, give it away or +re-use it under the terms of the Project Gutenberg License included +with this eBook or online at www.gutenberg.org + + +Title: Records of the Fossil Mammal Sinclairella, Family Apatemyidae, From the Chadronian and Orellan + +Author: William A. Clemens + +Release Date: October 20, 2009 [EBook #30297] + +Language: English + +Character set encoding: ISO-8859-1 + +*** START OF THIS PROJECT GUTENBERG EBOOK FOSSIL MAMMAL SINCLAIRELLA *** + + + + +Produced by Chris Curnow, Joseph Cooper, Diane Monico, and +the Online Distributed Proofreading Team at +https://www.pgdp.net + + + + + + +</pre> + + + + + + + + + + +<hr style="width: 65%;" /> +<p class="title"><span class="smcap">University of Kansas Publications</span><br /> + +<span class="smcap">Museum of Natural History</span><br /> +<br /> +Volume 14, No. 17, pp. 483-491, 2 figs.<br /> +March 2, 1964</p> + +<hr style="width: 65%;" /> + +<h1>Records of the Fossil Mammal<br /> +Sinclairella, Family Apatemyidae,<br /> +From the Chadronian and Orellan<br /></h1> + +<p class="title"><small>BY</small><br /><br /> + +<big>WILLIAM A. CLEMENS, JR.</big><br /><br /></p> + + +<p class="title"><span class="smcap">University of Kansas</span><br /> + +<span class="smcap">Lawrence</span><br /> + +1964</p> +<hr style="width: 65%;" /> + + +<p class="center"><span class="smcap">University of Kansas Publications, Museum of Natural History</span><br /> +<br /> +Editors: E. Raymond Hall, Chairman, Henry S. Fitch,<br /> +Theodore H. Eaton, Jr.<br /> +<br /> +<br /> +Volume 14, No. 17, pp. 483-491, 2 figs.<br /> +Published March 2, 1964<br /> +<br /> + +<span class="smcap">University of Kansas</span><br /> +Lawrence, Kansas<br /> +<br /> +<small>PRINTED BY</small><br /> + +<small>HARRY (BUD) TIMBERLAKE, STATE PRINTER</small><br /> + +<small>TOPEKA, KANSAS</small><br /> + +<small>1964</small><br /> +<br /> +<small>29-8587</small><br /> +</p> +<hr style="width: 65%;" /> + + + +<p><span class="pagenum"><a name="Page_485" id="Page_485">[Pg 485]</a></span></p> + +<h2>Records of the Fossil Mammal<br /> +Sinclairella, Family Apatemyidae,<br /> +From the Chadronian and Orellan<br /> +</h2> + +<p class="title"><small>BY</small><br /> + +WILLIAM A. CLEMENS, JR.<br /> +</p> + + +<h3>Introduction</h3> + + +<p>The family Apatemyidae has a long geochronological range in +North America, beginning in the Torrejonian land-mammal age, but +is represented by a relatively small number of fossils found at a few +localities. Two fossils of Orellan age, found in northeastern Colorado +and described here, demonstrate that the geochronological +range of the Apatemyidae extends into the Middle Oligocene. Isolated +teeth of <i>Sinclairella dakotensis</i> Jepsen, part of a sample of a +Chadronian local fauna collected by field parties from the Webb +School of California, are also described.</p> + +<div class="blockquot"><p>I thank Mr. Raymond M. Alf, Webb School of California, Claremont, +California, and Dr. Peter Robinson, University of Colorado Museum, Boulder, +Colorado, for permitting me to describe the fossils they discovered. Also Dr. +Robinson made available the draft of a short paper he had prepared on the +tooth found in Weld County, Colorado; his work was facilitated by a grant +from the University of Colorado Council on Research and Creative Work. I +also gratefully acknowledge receipt of critical data and valuable comments +from Drs. Edwin C. Galbreath, Glenn L. Jepsen, and Malcolm C. McKenna +who is currently revising the Paleocene apatemyids and studying the phylogenetic +relationships of the family. The prefixes of catalogue numbers used +in the text identify fossils in the collections of the following institutions: KU, +Museum of Natural History, The University of Kansas, Lawrence; Princeton, +Princeton Museum, Princeton, New Jersey; RAM-UCR, Raymond Alf Museum, +Webb School of California, Claremont, California (the permanent repository +for these specimens will be the University of California, Riverside); and UCM, +University of Colorado Museum, Boulder, Colorado. The system of notations +for teeth prescribed for use here is as follows: teeth in the upper half of the +dentition are designated by a capital letter and a number; thus M2 is the notation +for the upper second molar; teeth in the lower half of the dentition are +designated by a lower-case letter and a number; thus p2 is the notation for the +lower second premolar.</p></div> + + + +<hr style="width: 65%;" /><p><span class="pagenum"><a name="Page_486" id="Page_486">[Pg 486]</a></span></p> +<h3>Family <span class="smcap">Apatemyidae</span> Matthew, 1909<br /> + +Genus <b>Sinclairella</b> Jepsen, 1934<br /> + +<b>Sinclairella dakotensis</b> Jepsen, 1934<br /></h3> + + +<p>The type of the species, Princeton no. 13585, was discovered in +Chadronian strata of the upper part of the Chadron Formation +cropping out in Big Corral Draw, approximately 13 miles south-southwest +of Scenic, in southwestern South Dakota (Jepsen, 1934, +p. 291). Detailed descriptions of the type specimen are given in +papers by Jepsen (1934) and Scott and Jepsen (1936). Isolated +teeth of Chadronian age referable to <i>Sinclairella dakotensis</i> have +been discovered subsequently at a locality in Nebraska and fossils +of Orellan age, also referable to <i>S. dakotensis</i>, have been collected +at two localities in Colorado. The sample from each locality is +described separately.</p> + + +<h3>Sioux County, northwestern Nebraska</h3> + +<div class="blockquot"><p><i>Material.</i>—RAM-UCR nos. 381, left M1; 598, left m2; 1000, right m1; 1001, +right m2; 1079, right m2; 1674, right M2; and 3013, left m2.</p> + +<p><i>Locality and stratigraphy.</i>—These Chadronian fossils were discovered by +Raymond Alf and members of his field parties in several harvester ant mounds +built in exposures of the Chadron Formation in Sec. 26, T 33 N, R 53 W, Sioux +County, Nebraska (Alf, 1962, and Hough and Alf, 1958). This is UCR +locality V5403. The collectors carefully considered the possibility that some +of the fossils found in the ant mounds were collected from younger strata by +the harvester ants and concluded this was unlikely (Alf, personal communication).</p> + +<p><i>Description and comments.</i>—The cusps of RAM-UCR no. 381, a left M1, +are sharp and the wear-facets resulting from occlusion with the lower dentition +are small. The paraconule is a low, ill-defined cusp on the anterior margin +of the crown; a metaconule is not present. A smooth stylar shelf is present +labial to the metacone. The crown was supported by three roots. There are +no interradicular crests.</p> + +<p>The crown of RAM-UCR no. 1674, a right M2, is heavily abraded and +many morphological details of the cusps have been destroyed. Low interradicular +crests linked the three roots of the tooth with a low, central prominence. +As was the case with RAM-UCR no. 381, no significant differences +could be found in comparisons with illustrations of the teeth preserved in +Princeton no. 13585.</p> + +<p>RAM-UCR nos. 598, 1001, 1079, and 3013 all appear to be m2's. The +talonids of these teeth are not elongated, their trigonids have quadrilateral outlines, +and the paraconids are small but prominent, bladelike cusps. The trigonid +of RAM-UCR 1000 is elongated and the paraconid is a minute cusp; the +tooth closely resembles the m1 of the type of <i>Sinclairella dakotensis</i>.</p></div> + +<p><span class="pagenum"><a name="Page_487" id="Page_487">[Pg 487]</a></span></p> +<h3>Logan County, northeastern Colorado</h3> + +<div class="blockquot"><p><i>Material.</i>—KU no. 11210 (fig. 1), a fragment of a left maxillary containing +P4 and M1-2.</p> + +<p><i>Locality and stratigraphy.</i>—The fossil was found in the center of the W-1/2, +Sec. 21, T 11 N, R 53 W, Logan County, Colorado, " ... in the bed below +<i>Agnotocastor</i> bed, Cedar Creek Member...." (Ronald H. Pine, 1958, +field notes on file at the University of Kansas). The bed so defined is part of +unit 3 in the lower division of the Cedar Creek Member, as subdivided by +Galbreath (1953:25) in stratigraphic section XII. The fauna obtained from +unit 3 is of Orellan age.</p></div> + +<p class="figcenter" style="width: 545px;"> +<img src="images/image005.png" width="545" height="600" alt="Fig. 1. Sinclairella dakotensis Jepsen, KU no. 11210, fragment of left maxillary +with P4 and M1-2; Orellan, Logan County, Colorado; drawings by Mrs. Judith +Hood: a, labial view; b, occlusal view; both approximately × 9." title="Fig. 1. Sinclairella dakotensis Jepsen, KU no. 11210, fragment of left maxillary" /> +<span class="caption">Fig. 1. Sinclairella dakotensis Jepsen, KU no. 11210, fragment of left maxillary +with P4 and M1-2; Orellan, Logan County, Colorado; drawings by Mrs. Judith +Hood: a, labial view; b, occlusal view; both approximately × 9.</span> +</p> + +<div class="blockquot"><p><i>Description and comments.</i>—P4 of KU no. 11210 has a large posterolingual +cusp separated from the main cusp by a distinct groove, which deepens posteriorly.<span class="pagenum"><a name="Page_488" id="Page_488">[Pg 488]</a></span> +The posterolingual cusp is supported by the broad posterior root. P4 +of the type specimen of <i>Sinclairella dakotensis</i> is described (Jepsen, 1934, p. +392) as having an oval outline at the base of the crown, and a small, posterolingual +cusp. A chip of enamel is missing from the posterior slope of the main +cusp of the P4 of KU no. 11210. The anterior slope of the main cusp is +flattened, possibly the result of wear, and there is no evidence of a groove like +that present on the P4 of the type specimen.</p> + +<p>Only a few differences were found between the molars preserved in KU no. +11210 and their counterparts in the type specimen. A stylar shelf is present +labial to the metacone of M1 of KU no. 11210, but, unlike the type, its surface +is smooth and there is no evidence of cusps. Of the three small stylar +cusps on the stylar shelf of M2 the smallest is in the position of a mesostyle. +The M2 lacks a chip of enamel from the lingual surface of the hypocone. Unlike +the M2 of Princeton no. 13585, in occlusal view the posterior margin of the +M2 of KU no. 11210 is convex posterior to the metacone. The anterior edge +of the base of the zygomatic arch of KU no. 11210 was dorsal to M2. The +shallow oval depression in the maxillary dorsal to M1 might be the result of +post-mortem distortion.</p> + +<p>The molars preserved in KU no. 11210 and their counterparts in the type +specimen do not appear to be significantly different in size (table 1) or morphology +of the cusps. The only difference between the two specimens that +might be of classificatory significance is the difference in size of the posterolingual +cusp of P4. At present the range of intraspecific variation in the morphology +of P4 has not been documented for any species of apatemyid. The +evolutionary trend or trends of the apatemyids (McKenna, 1960, p. 48) for +progressive reduction of function of p4 probably were paralleled by similar +trends in the evolution of the P4. If so, the intraspecific variation in the morphology +of P4 could be expected to be somewhat greater than that of the upper +molars, for example. The morphological difference between the P4's of the +type of <i>Sinclairella dakotensis</i> and KU no. 11210 is not extreme and does not +exceed the range of intraspecific variation that could be expected for this element +of the dentition. The close resemblances in +size and morphology between the M1-2 of Princeton +no. 13585 and KU no. 11210 also favor identification +of the latter as part of a member of an +Orellan population of <i>Sinclairella dakotensis</i>.</p></div> + + +<h3>Weld County, northeastern Colorado</h3> + +<p class="figleft" style="width: 342px;"> +<img src="images/image006.png" width="342" height="375" alt="Fig. 2. Sinclairella dakotensis +Jepsen, UCM no. 21073, right M2; Orellan, Weld County, Colorado; +drawing by Mrs. Judith Hood: occlusal view, approximately × 9." title="Fig. 2. Sinclairella dakotensis +Jepsen, UCM no. 21073, right M2" /> +<span class="caption">Fig. 2. Sinclairella dakotensis +Jepsen, UCM no. 21073, right M2; Orellan, Weld County, Colorado; +drawing by Mrs. Judith Hood: occlusal view, approximately × 9.</span> +</p> + +<div class="blockquot"><p><i>Material.</i>—UCM no. 20173 (fig. 2), is a right M2.</p> + +<p><i>Locality and stratigraphy.</i>—The tooth was discovered +at the Mellinger locality, Sec. 17, T 11 +N, R 65 W, Weld County, Colorado. The Mellinger +locality is in the Cedar Creek Member, +White River Formation, and its fauna is considered +to be of Orellan age (Patterson and McGrew, +1937, and Galbreath, 1953).</p> + +<p><i>Description and comments.</i>—UCM no. 21073, +which is more heavily abraded than KU no. 11210, +shows no evidence of a stylar cusp either anterolabial<span class="pagenum"><a name="Page_489" id="Page_489">[Pg 489]</a></span> +to the metacone or in the position of a mesostyle. A small stylar cusp +is present anterolabial to the paracone. A notch that appears to have been cut +through the enamel of the posterolabial corner of the crown could have received +the parastylar apex of M3. A similar notch is not present on the M2 of KU no. +11210 nor indicated in the illustrations of the M2 of Princeton no. 13585. The +coronal dimensions of UCM no. 21073 (table 1) do not appear to differ significantly +from those of the M2's of KU no. 11210 and the type specimen of +<i>Sinclairella dakotensis</i>.</p></div> + + +<h3>Comments</h3> + +<p>With the discovery of Orellan apatemyids the geochronological +range of the family in North America is shown to extend from the +Torrejonian through the Orellan land-mammal ages. The discoveries +reported here enlarge the Oligocene record of apatemyids +to include not only the type specimen of <i>Sinclairella dakotensis</i>, a +skull and associated mandible from South Dakota, but also seven +isolated teeth, representing at least two individuals, from a Chadronian +fossil locality in Nebraska and one specimen from each of +two Orellan fossil localities in northeastern Colorado. Simpson +(1944:73, and 1953:127) presented tabulations of the published records +of American apatemyids and suggested the data indicated the +populations of these mammals were of small size throughout the +history of the family. The few pre-Oligocene occurrences of apatemyids +described subsequently (note McKenna, 1960, figs. 3-10, +and p. 48) and occurrences described here tend to reinforce Simpson's +interpretation. This interpretation may have to be modified +to some degree, however, when current studies of collections of pre-Oligocene +apatemyids are completed (McKenna, personal communication).</p> + +<p>Although information concerning the evolutionary trends of American +apatemyids has been published, no data on the morphological +variation in a population are available in the literature. An adequate +basis for evaluating the significance of the morphological differences +between the P4's of Princeton no. 13585 and KU no. 12110 coupled +with the similarities of their M1-2's is lacking. In the evolution of +American apatemyids the P4 underwent reduction in size and, apparently, +curtailment of function. This history suggests the range +of morphological variation of P4 in populations of <i>Sinclairella dakotensis</i> +could be expected to be greater than that of the molars +and encompass the morphological differences between the P4's of +Princeton no. 13585 and KU no. 12110. The difference in age of +the Chadronian and Orellan fossils does not constitute proof that +they pertain to different species. Although the identification is admittedly<span class="pagenum"><a name="Page_490" id="Page_490">[Pg 490]</a></span> +provisional until more fossils including other parts of the +skeleton are discovered, the Orellan fossils described here are referred +to <i>Sinclairella dakotensis</i>.</p> + + + + + +<div class="center"><span class="smcap">Table 1.—Measurements (in millimeters) of Teeth of Sinclairella<br /> +dakotensis Jepsen.</span><br /><br /> + +<table border="1" cellpadding="4" cellspacing="0" summary="teeth"> +<tr><td align="left"> </td><th colspan="2">P4</th><th colspan="2">M1</th><th colspan="2">M2</th></tr> +<tr><td align="left"> </td><td align="center">length</td><td align="center">width</td><td align="center">length<a name="FNanchor_1_2" id="FNanchor_1_2"></a><a href="#Footnote_1_1" class="fnanchor">[1]</a></td><td align="center">width<a name="FNanchor_1_3" id="FNanchor_1_3"></a><a href="#Footnote_1_1" class="fnanchor">[1]</a></td><td align="center">length<a name="FNanchor_1_4" id="FNanchor_1_4"></a><a href="#Footnote_1_1" class="fnanchor">[1]</a></td><td align="center">width<a name="FNanchor_1_1" id="FNanchor_1_1"></a><a href="#Footnote_1_1" class="fnanchor">[1]</a></td></tr> +<tr><td align="left">Princeton no. 13585<a name="FNanchor_2_2" id="FNanchor_2_2"></a><a href="#Footnote_2_2" class="fnanchor">[2]</a></td><td align="center"> 2.1</td><td align="center"> 1.1</td><td align="center"> 4.0</td><td align="center"> 3.7</td><td align="center"> 3.4</td><td align="center"> 4.7</td></tr> +<tr><td align="left">RAM no. 381</td><td align="center"></td><td align="center"></td><td align="center"> 4.1</td><td align="center"> 3.5</td><td align="center"> </td><td align="center"> </td></tr> +<tr><td align="left">RAM no. 1674</td><td align="center"></td><td align="center"></td><td align="center"></td><td align="center"></td><td align="center"> 3.4</td><td align="center"> 4.2</td></tr> +<tr><td align="left">KU no. 11210</td><td align="center"> 2.4</td><td align="center"> 1.6</td><td align="center"> 3.9</td><td align="center"> 3.5</td><td align="center"> 3.8</td><td align="center"> 4.1+</td></tr> +<tr><td align="left">UCM no. 21073</td><td align="center"></td><td align="center"></td><td align="center"></td><td align="center"></td><td align="center"> 3.6</td><td align="center"> 4.1</td></tr> +<tr><td colspan="3" rowspan="2"> </td><td colspan="2">m1</td><td colspan="2">m2</td></tr> +<tr><td>length</td><td>width</td><td>length</td><td>width</td></tr> +<tr><td colspan="3" align="left">Princeton no. 13585<a name="FNanchor_3_3" id="FNanchor_3_3"></a><a href="#Footnote_3_3" class="fnanchor">[3]</a></td><td align="center"> 3.5</td><td align="center"> 2.4</td><td align="center"> 3.7</td><td align="center"> 2.8</td></tr> +<tr><td colspan="3" align="left">RAM no. 1000</td><td align="center"> 3.5</td><td align="center"> 2.2</td><td align="center"> </td><td align="center"> </td></tr> +<tr><td colspan="3" align="left">RAM no. 598</td><td align="center"></td><td align="center"></td><td align="center"> 3.8</td><td align="center"> 2.6</td></tr> +<tr><td colspan="3" align="left">RAM no. 1001</td><td align="center"></td><td align="center"></td><td align="center"> 3.6+</td><td align="center"> 2.6</td></tr> +<tr><td colspan="3" align="left">RAM no. 1079</td><td align="center"></td><td align="center"></td><td align="center"> 4.0</td><td align="center"> 2.8</td></tr> +<tr><td colspan="3" align="left">RAM no. 3013</td><td align="center"></td><td align="center"></td><td align="center"> 3.6</td><td align="center"> 2.8</td></tr> +</table></div> + + +<div class="footnote"><p><a name="Footnote_1_1" id="Footnote_1_1"></a><a href="#FNanchor_1_1"><span class="label">[1]</span></a> Length defined as maximum dimension of the labial half of the crown measured +parallel to a line drawn through the apices of paracone and metacone. Width defined as +maximum coronal dimension measured along line perpendicular to line defined by apices of +paracone and metacone.</p></div> + +<div class="footnote"><p><a name="Footnote_2_2" id="Footnote_2_2"></a><a href="#FNanchor_2_2"><span class="label">[2]</span></a> Dimensions provided by Dr. Glenn L. Jepsen.</p></div> + +<div class="footnote"><p><a name="Footnote_3_3" id="Footnote_3_3"></a><a href="#FNanchor_3_3"><span class="label">[3]</span></a> Dimensions taken from Jepsen (1934:300).</p></div> + + + +<p><span class="pagenum"><a name="Page_491" id="Page_491">[Pg 491]</a></span></p><hr style="width: 65%;" /> +<h2>Literature Cited</h2> + +<div class="blockquot"><p><span class="smcap">Alf, R.</span></p> + +<p><span class="i4">1962. A new species of the rodent <i>Pipestoneomys</i> from the Oligocene of +Nebraska. Breviora, Mus. Comp. Zool., no. 172, pp. 1-7, 3 figs.</span></p> + +<p><span class="smcap">Galbreath, E. C.</span></p> + +<p><span class="i4">1953. A contribution to the Tertiary geology and paleontology of northeastern +Colorado. Univ. Kansas Paleont. Cont., Vertebrata, art. 4, +pp. 1-120, 2 pls., 26 figs.</span></p> + +<p><span class="smcap">Hough, J.</span>, and <span class="smcap">Alf, R.</span></p> + +<p><span class="i4">1958. A Chadron mammalian fauna from Nebraska. Journ. Paleon. 30: +132-140, 4 figs.</span></p> + +<p><span class="smcap">Jepsen, G. L.</span></p> + +<p><span class="i4">1934. A revision of the American Apatemyidae and the description of a +new genus, <i>Sinclairella</i>, from the White River Oligocene of South +Dakota. Proc. Amer. Philos. Soc., 74:287-305, 3 pls., 4 figs.</span></p> + +<p><span class="smcap">McKenna, M. C.</span></p> + +<p><span class="i4">1960. Fossil Mammalia from the early Wasatchian Four Mile fauna, +Eocene of northwest Colorado. Univ. California Publ. in Geol. +Sci., 37:1-130, 64 figs.</span></p> + +<p><span class="smcap">Matthew, W. D.</span></p> + +<p><span class="i4">1909. The Carnivora and Insectivora of the Bridger Basin, Middle +Eocene. Mem. Amer. Mus. Nat. Hist., 9:289-567, pls. 42-52, 118 +figs.</span></p> + +<p><span class="smcap">Patterson, B.</span> and <span class="smcap">McGrew, P. O.</span></p> + +<p><span class="i4">1937. A soricid and two erinaceids from the White River Oligocene. Geol. +Ser., Field Mus. Nat. Hist., 6:245-272, figs. 60-74.</span></p> + +<p><span class="smcap">Scott, W. B.</span> and <span class="smcap">Jepsen, G. L.</span></p> + +<p><span class="i4">1936. The mammalian fauna of the White River Oligocene—Part I. Insectivora +and Carnivora. Trans. Amer. Philos. Soc., n. s., 28:1-153, +22 pls., 7 figs.</span></p> + +<p><span class="smcap">Simpson, G. G.</span></p> + +<p><span class="i4">1944. Tempo and mode in evolution. New York: Columbia Univ. Press, +xviii + 237 pp., 36 figs.</span></p> + +<p><span class="i4">1953. The major features of evolution. New York: Columbia Univ. +Press, xx + 434 pp., 52 figs.</span></p> + +<p><i>Transmitted June 24, 1963.</i></p></div> + + + + + + + + +<pre> + + + + + +End of the Project Gutenberg EBook of Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, From the Chadronian and Orellan, by William A. 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Clemens + +This eBook is for the use of anyone anywhere at no cost and with +almost no restrictions whatsoever. You may copy it, give it away or +re-use it under the terms of the Project Gutenberg License included +with this eBook or online at www.gutenberg.org + + +Title: Records of the Fossil Mammal Sinclairella, Family Apatemyidae, From the Chadronian and Orellan + +Author: William A. Clemens + +Release Date: October 20, 2009 [EBook #30297] + +Language: English + +Character set encoding: ASCII + +*** START OF THIS PROJECT GUTENBERG EBOOK FOSSIL MAMMAL SINCLAIRELLA *** + + + + +Produced by Chris Curnow, Joseph Cooper, Diane Monico, and +the Online Distributed Proofreading Team at +https://www.pgdp.net + + + + + + + + + + + +UNIVERSITY OF KANSAS PUBLICATIONS + +MUSEUM OF NATURAL HISTORY + +Volume 14, No. 17, pp. 483-491, 2 figs. +March 2, 1964 + + +Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, +From the Chadronian and Orellan + + +BY + +WILLIAM A. CLEMENS, JR. + + +UNIVERSITY OF KANSAS +LAWRENCE +1964 + + +UNIVERSITY OF KANSAS PUBLICATIONS, MUSEUM OF NATURAL HISTORY + +Editors: E. Raymond Hall, Chairman, Henry S. Fitch, +Theodore H. Eaton, Jr. + +Volume 14, No. 17, pp. 483-491, 2 figs. +Published March 2, 1964 + +UNIVERSITY OF KANSAS +Lawrence, Kansas + +PRINTED BY +HARRY (BUD) TIMBERLAKE, STATE PRINTER +TOPEKA, KANSAS + +1964 + + +29-8587 + + + + +Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, +From the Chadronian and Orellan + +BY + +WILLIAM A. CLEMENS, JR. + + +Introduction + + +The family Apatemyidae has a long geochronological range in North +America, beginning in the Torrejonian land-mammal age, but is +represented by a relatively small number of fossils found at a few +localities. Two fossils of Orellan age, found in northeastern Colorado +and described here, demonstrate that the geochronological range of the +Apatemyidae extends into the Middle Oligocene. Isolated teeth of +_Sinclairella dakotensis_ Jepsen, part of a sample of a Chadronian +local fauna collected by field parties from the Webb School of +California, are also described. + + I thank Mr. Raymond M. Alf, Webb School of California, + Claremont, California, and Dr. Peter Robinson, University of + Colorado Museum, Boulder, Colorado, for permitting me to + describe the fossils they discovered. Also Dr. Robinson made + available the draft of a short paper he had prepared on the + tooth found in Weld County, Colorado; his work was + facilitated by a grant from the University of Colorado + Council on Research and Creative Work. I also gratefully + acknowledge receipt of critical data and valuable comments + from Drs. Edwin C. Galbreath, Glenn L. Jepsen, and Malcolm + C. McKenna who is currently revising the Paleocene + apatemyids and studying the phylogenetic relationships of + the family. The prefixes of catalogue numbers used in the + text identify fossils in the collections of the following + institutions: KU, Museum of Natural History, The University + of Kansas, Lawrence; Princeton, Princeton Museum, Princeton, + New Jersey; RAM-UCR, Raymond Alf Museum, Webb School of + California, Claremont, California (the permanent repository + for these specimens will be the University of California, + Riverside); and UCM, University of Colorado Museum, Boulder, + Colorado. The system of notations for teeth prescribed for + use here is as follows: teeth in the upper half of the + dentition are designated by a capital letter and a number; + thus M2 is the notation for the upper second molar; teeth in + the lower half of the dentition are designated by a + lower-case letter and a number; thus p2 is the notation for + the lower second premolar. + + + + +Family APATEMYIDAE Matthew, 1909 + +Genus =Sinclairella= Jepsen, 1934 + +=Sinclairella dakotensis= Jepsen, 1934 + + +The type of the species, Princeton no. 13585, was discovered in +Chadronian strata of the upper part of the Chadron Formation cropping +out in Big Corral Draw, approximately 13 miles south-southwest of +Scenic, in southwestern South Dakota (Jepsen, 1934, p. 291). Detailed +descriptions of the type specimen are given in papers by Jepsen (1934) +and Scott and Jepsen (1936). Isolated teeth of Chadronian age referable +to _Sinclairella dakotensis_ have been discovered subsequently at a +locality in Nebraska and fossils of Orellan age, also referable to _S. +dakotensis_, have been collected at two localities in Colorado. The +sample from each locality is described separately. + + +Sioux County, northwestern Nebraska + + _Material._--RAM-UCR nos. 381, left M1; 598, left m2; 1000, + right m1; 1001, right m2; 1079, right m2; 1674, right M2; + and 3013, left m2. + + _Locality and stratigraphy._--These Chadronian fossils were + discovered by Raymond Alf and members of his field parties + in several harvester ant mounds built in exposures of the + Chadron Formation in Sec. 26, T 33 N, R 53 W, Sioux County, + Nebraska (Alf, 1962, and Hough and Alf, 1958). This is UCR + locality V5403. The collectors carefully considered the + possibility that some of the fossils found in the ant mounds + were collected from younger strata by the harvester ants and + concluded this was unlikely (Alf, personal communication). + + _Description and comments._--The cusps of RAM-UCR no. 381, a + left M1, are sharp and the wear-facets resulting from + occlusion with the lower dentition are small. The paraconule + is a low, ill-defined cusp on the anterior margin of the + crown; a metaconule is not present. A smooth stylar shelf is + present labial to the metacone. The crown was supported by + three roots. There are no interradicular crests. + + The crown of RAM-UCR no. 1674, a right M2, is heavily + abraded and many morphological details of the cusps have + been destroyed. Low interradicular crests linked the three + roots of the tooth with a low, central prominence. As was + the case with RAM-UCR no. 381, no significant differences + could be found in comparisons with illustrations of the + teeth preserved in Princeton no. 13585. + + RAM-UCR nos. 598, 1001, 1079, and 3013 all appear to be + m2's. The talonids of these teeth are not elongated, their + trigonids have quadrilateral outlines, and the paraconids + are small but prominent, bladelike cusps. The trigonid of + RAM-UCR 1000 is elongated and the paraconid is a minute + cusp; the tooth closely resembles the m1 of the type of + _Sinclairella dakotensis_. + + +Logan County, northeastern Colorado + + _Material._--KU no. 11210 (fig. 1), a fragment of a left + maxillary containing P4 and M1-2. + + _Locality and stratigraphy._--The fossil was found in the + center of the W-1/2, Sec. 21, T 11 N, R 53 W, Logan County, + Colorado, "... in the bed below _Agnotocastor_ bed, Cedar + Creek Member...." (Ronald H. Pine, 1958, field notes on file + at the University of Kansas). The bed so defined is part of + unit 3 in the lower division of the Cedar Creek Member, as + subdivided by Galbreath (1953:25) in stratigraphic section + XII. The fauna obtained from unit 3 is of Orellan age. + +[Illustration: FIG. 1. _Sinclairella dakotensis_ Jepsen, KU no. 11210, +fragment of left maxillary with P4 and M1-2; Orellan, Logan County, +Colorado; drawings by Mrs. Judith Hood: a, labial view; b, occlusal +view; both approximately x 9.] + + _Description and comments._--P4 of KU no. 11210 has a large + posterolingual cusp separated from the main cusp by a + distinct groove, which deepens posteriorly. The + posterolingual cusp is supported by the broad posterior + root. P4 of the type specimen of _Sinclairella dakotensis_ + is described (Jepsen, 1934, p. 392) as having an oval + outline at the base of the crown, and a small, + posterolingual cusp. A chip of enamel is missing from the + posterior slope of the main cusp of the P4 of KU no. 11210. + The anterior slope of the main cusp is flattened, possibly + the result of wear, and there is no evidence of a groove + like that present on the P4 of the type specimen. + + Only a few differences were found between the molars + preserved in KU no. 11210 and their counterparts in the type + specimen. A stylar shelf is present labial to the metacone + of M1 of KU no. 11210, but, unlike the type, its surface is + smooth and there is no evidence of cusps. Of the three small + stylar cusps on the stylar shelf of M2 the smallest is in + the position of a mesostyle. The M2 lacks a chip of enamel + from the lingual surface of the hypocone. Unlike the M2 of + Princeton no. 13585, in occlusal view the posterior margin + of the M2 of KU no. 11210 is convex posterior to the + metacone. The anterior edge of the base of the zygomatic + arch of KU no. 11210 was dorsal to M2. The shallow oval + depression in the maxillary dorsal to M1 might be the result + of post-mortem distortion. + + The molars preserved in KU no. 11210 and their counterparts + in the type specimen do not appear to be significantly + different in size (table 1) or morphology of the cusps. The + only difference between the two specimens that might be of + classificatory significance is the difference in size of the + posterolingual cusp of P4. At present the range of + intraspecific variation in the morphology of P4 has not been + documented for any species of apatemyid. The evolutionary + trend or trends of the apatemyids (McKenna, 1960, p. 48) for + progressive reduction of function of p4 probably were + paralleled by similar trends in the evolution of the P4. If + so, the intraspecific variation in the morphology of P4 + could be expected to be somewhat greater than that of the + upper molars, for example. The morphological difference + between the P4's of the type of _Sinclairella dakotensis_ + and KU no. 11210 is not extreme and does not exceed the + range of intraspecific variation that could be expected for + this element of the dentition. The close resemblances in + size and morphology between the M1-2 of Princeton no. 13585 + and KU no. 11210 also favor identification of the latter as + part of a member of an Orellan population of _Sinclairella + dakotensis_. + + +Weld County, northeastern Colorado + +[Illustration: FIG. 2. _Sinclairella dakotensis_ Jepsen, UCM no. 21073, +right M2; Orellan, Weld County, Colorado; drawing by Mrs. Judith Hood: +occlusal view, approximately x 9.] + + _Material._--UCM no. 20173 (fig. 2), is a right M2. + + _Locality and stratigraphy._--The tooth was discovered at + the Mellinger locality, Sec. 17, T 11 N, R 65 W, Weld + County, Colorado. The Mellinger locality is in the Cedar + Creek Member, White River Formation, and its fauna is + considered to be of Orellan age (Patterson and McGrew, 1937, + and Galbreath, 1953). + + _Description and comments._--UCM no. 21073, which is more + heavily abraded than KU no. 11210, shows no evidence of a + stylar cusp either anterolabial to the metacone or in the + position of a mesostyle. A small stylar cusp is present + anterolabial to the paracone. A notch that appears to have + been cut through the enamel of the posterolabial corner of + the crown could have received the parastylar apex of M3. A + similar notch is not present on the M2 of KU no. 11210 nor + indicated in the illustrations of the M2 of Princeton no. + 13585. The coronal dimensions of UCM no. 21073 (table 1) do + not appear to differ significantly from those of the M2's of + KU no. 11210 and the type specimen of _Sinclairella + dakotensis_. + + +Comments + +With the discovery of Orellan apatemyids the geochronological range of +the family in North America is shown to extend from the Torrejonian +through the Orellan land-mammal ages. The discoveries reported here +enlarge the Oligocene record of apatemyids to include not only the type +specimen of _Sinclairella dakotensis_, a skull and associated mandible +from South Dakota, but also seven isolated teeth, representing at least +two individuals, from a Chadronian fossil locality in Nebraska and one +specimen from each of two Orellan fossil localities in northeastern +Colorado. Simpson (1944:73, and 1953:127) presented tabulations of the +published records of American apatemyids and suggested the data +indicated the populations of these mammals were of small size +throughout the history of the family. The few pre-Oligocene occurrences +of apatemyids described subsequently (note McKenna, 1960, figs. 3-10, +and p. 48) and occurrences described here tend to reinforce Simpson's +interpretation. This interpretation may have to be modified to some +degree, however, when current studies of collections of pre-Oligocene +apatemyids are completed (McKenna, personal communication). + +Although information concerning the evolutionary trends of American +apatemyids has been published, no data on the morphological variation +in a population are available in the literature. An adequate basis for +evaluating the significance of the morphological differences between +the P4's of Princeton no. 13585 and KU no. 12110 coupled with the +similarities of their M1-2's is lacking. In the evolution of American +apatemyids the P4 underwent reduction in size and, apparently, +curtailment of function. This history suggests the range of +morphological variation of P4 in populations of _Sinclairella +dakotensis_ could be expected to be greater than that of the molars and +encompass the morphological differences between the P4's of Princeton +no. 13585 and KU no. 12110. The difference in age of the Chadronian and +Orellan fossils does not constitute proof that they pertain to +different species. Although the identification is admittedly +provisional until more fossils including other parts of the skeleton +are discovered, the Orellan fossils described here are referred to +_Sinclairella dakotensis_. + + +TABLE 1.--MEASUREMENTS (IN MILLIMETERS) OF TEETH OF SINCLAIRELLA +DAKOTENSIS JEPSEN. + +========================================================================== + | P4 | M1 | M2 +-----------------------+------------+------------------+------------------ + |length|width|length[1]|width[1]|length[1]|width[1] +-----------------------+------+-----+---------+--------+---------+-------- +Princeton no. 13585[2] | 2.1 | 1.1 | 4.0 | 3.7 | 3.4 | 4.7 +RAM no. 381 | | | 4.1 | 3.5 | | +RAM no. 1674 | | | | | 3.4 | 4.2 +KU no. 11210 | 2.4 | 1.6 | 3.9 | 3.5 | 3.8 | 4.1+ +UCM no. 21073 | | | | | 3.6 | 4.1 +-----------------------+------+-----+---------+--------+---------+-------- + | m1 | m2 + +---------+--------+---------+-------- + | length | width | length | width + +---------+--------+---------+-------- +Princeton no. 13585[3] | 3.5 | 2.4 | 3.7 | 2.8 +RAM no. 1000 | 3.5 | 2.2 | | +RAM no. 598 | | | 3.8 | 2.6 +RAM no. 1001 | | | 3.6+ | 2.6 +RAM no. 1079 | | | 4.0 | 2.8 +RAM no. 3013 | | | 3.6 | 2.8 +------------------------------------+---------+--------+---------+-------- + +[Footnote 1: Length defined as maximum dimension of the labial half of +the crown measured parallel to a line drawn through the apices of +paracone and metacone. Width defined as maximum coronal dimension +measured along line perpendicular to line defined by apices of paracone +and metacone.] + +[Footnote 2: Dimensions provided by Dr. Glenn L. Jepsen.] + +[Footnote 3: Dimensions taken from Jepsen (1934:300).] + + + + +Literature Cited + +ALF, R. + 1962. A new species of the rodent _Pipestoneomys_ from the + Oligocene of Nebraska. Breviora, Mus. Comp. Zool., no. 172, + pp. 1-7, 3 figs. + +GALBREATH, E. C. + 1953. A contribution to the Tertiary geology and paleontology + of northeastern Colorado. Univ. Kansas Paleont. Cont., + Vertebrata, art. 4, pp. 1-120, 2 pls., 26 figs. + +HOUGH, J., and ALF, R. + 1958. A Chadron mammalian fauna from Nebraska. Journ. Paleon. + 30:132-140, 4 figs. + +JEPSEN, G. L. + 1934. A revision of the American Apatemyidae and the description + of a new genus, _Sinclairella_, from the White River + Oligocene of South Dakota. Proc. Amer. Philos. Soc., + 74:287-305, 3 pls., 4 figs. + +MCKENNA, M. C. + 1960. Fossil Mammalia from the early Wasatchian Four Mile fauna, + Eocene of northwest Colorado. Univ. California Publ. in + Geol. Sci., 37:1-130, 64 figs. + +MATTHEW, W. D. + 1909. The Carnivora and Insectivora of the Bridger Basin, Middle + Eocene. Mem. Amer. Mus. Nat. Hist., 9:289-567, pls. 42-52, + 118 figs. + +PATTERSON, B. and MCGREW, P. O. + 1937. A soricid and two erinaceids from the White River Oligocene. + Geol. Ser., Field Mus. Nat. Hist., 6:245-272, figs. 60-74. + +SCOTT, W. B. and JEPSEN, G. L. + 1936. The mammalian fauna of the White River Oligocene--Part I. + Insectivora and Carnivora. Trans. Amer. Philos. Soc., n. s., + 28:1-153, 22 pls., 7 figs. + +SIMPSON, G. G. + 1944. Tempo and mode in evolution. New York: Columbia Univ. Press, + xviii + 237 pp., 36 figs. + + 1953. The major features of evolution. New York: Columbia Univ. + Press, xx + 434 pp., 52 figs. + +_Transmitted June 24, 1963._ + + + + + +End of the Project Gutenberg EBook of Records of the Fossil Mammal +Sinclairella, Family Apatemyidae, From the Chadronian and Orellan, by William A. Clemens + +*** END OF THIS PROJECT GUTENBERG EBOOK FOSSIL MAMMAL SINCLAIRELLA *** + +***** This file should be named 30297.txt or 30297.zip ***** +This and all associated files of various formats will be found in: + https://www.gutenberg.org/3/0/2/9/30297/ + +Produced by Chris Curnow, Joseph Cooper, Diane Monico, and +the Online Distributed Proofreading Team at +https://www.pgdp.net + + +Updated editions will replace the previous one--the old editions +will be renamed. + +Creating the works from public domain print editions means that no +one owns a United States copyright in these works, so the Foundation +(and you!) can copy and distribute it in the United States without +permission and without paying copyright royalties. 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