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+<!DOCTYPE html PUBLIC "-//W3C//DTD XHTML 1.0 Strict//EN"
+ "http://www.w3.org/TR/xhtml1/DTD/xhtml1-strict.dtd">
+
+<html xmlns="http://www.w3.org/1999/xhtml">
+ <head>
+ <meta http-equiv="Content-Type" content="text/html;charset=UTF-8" />
+ <title>
+ The Project Gutenberg eBook of A New Doglike Carnivore, Genus Cynarctus,
+From the Clarendonian, Pliocene, of Texas, by E. Raymond Hall And Walter W. Dalquest.
+ </title>
+ <style type="text/css">
+
+
+ p { margin-top: .75em;
+ text-align: justify;
+ margin-bottom: .75em;
+ }
+ h1 { text-align: center; line-height: 1.5; clear: both; }
+
+ h2,h3 { text-align: center; clear: both; }
+
+ p.title { text-align: center; text-indent: 0;
+ font-weight: bold;
+ line-height: 1.4; margin-bottom: 3em; }
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+ margin-right: auto;
+ clear: both;
+ }
+
+ body{margin-left: 10%;
+ margin-right: 10%;
+ }
+
+ .pagenum { /* uncomment the next line for invisible page numbers */
+ /* visibility: hidden; */
+ position: absolute;
+ left: 92%;
+ font-size: smaller;
+ text-align: right;
+ } /* page numbers */
+
+ .blockquot{margin-left: 5%; margin-right: 10%;}
+
+ .center {text-align: center;}
+ .smcap {font-variant: small-caps;}
+
+ .caption {font-weight: bold;}
+
+ .figcenter {margin: auto; text-align: center;}
+
+ </style>
+ </head>
+<body>
+<div>*** START OF THE PROJECT GUTENBERG EBOOK 30260 ***</div>
+
+<hr style="width: 65%;" />
+<p class="title"><span class="smcap">University of Kansas Publications</span><br />
+
+<span class="smcap">Museum of Natural History</span><br />
+<br />
+Volume 14, No. 10, pp. 135-138, 2 figs.<br />
+
+April 30, 1962</p>
+<hr style="width: 65%;" />
+
+<h1>A New Doglike Carnivore, Genus Cynarctus,<br />
+From the Clarendonian, Pliocene, of Texas<br /></h1>
+
+<p class="title"><small>BY</small><br /><br />
+
+<big>E. RAYMOND HALL and WALTER W. DALQUEST</big><br /><br /></p>
+
+
+<p class="title"><span class="smcap">University of Kansas</span><br />
+
+<span class="smcap">Lawrence</span><br />
+
+1962</p>
+<hr style="width: 65%;" />
+
+
+<p class="center"><span class="smcap">University of Kansas Publications, Museum of Natural History</span><br />
+<br />
+Editors: E. Raymond Hall, Chairman, Henry S. Fitch,<br />
+Theodore H. Eaton, Jr.<br />
+<br />
+<br />
+Volume 14, No. 10, pp. 135-138, 2 figs.<br />
+Published April 30, 1962<br />
+<br />
+<span class="smcap">University of Kansas</span><br />
+Lawrence, Kansas<br />
+<br />
+<small>PRINTED BY</small><br />
+
+<small>JEAN M. NEIBARGER, STATE PRINTER</small><br />
+
+<small>TOPEKA, KANSAS</small><br />
+
+<small>1962</small><br />
+
+<small>29-2890</small><br />
+</p>
+<hr style="width: 65%;" /><p><span class="pagenum"><a name="Page_137" id="Page_137">[Pg 137]</a></span></p>
+
+
+
+
+<h2>
+A New Doglike Carnivore, Genus Cynarctus,<br />
+From the Clarendonian, Pliocene, of Texas<br />
+</h2>
+
+<p class="center"><small>BY</small><br />
+
+<small>E. RAYMOND HALL and WALTER W. DALQUEST</small></p>
+
+
+<p>A study of a right maxilla bearing P3-M1 and part of a right
+mandibular ramus bearing m2 (see figures) reveals the existence of
+an unnamed species of cynarctine carnivore. It may be known as:</p>
+
+<h3><b>Cynarctus fortidens</b> new species</h3>
+
+<div class="blockquot"><p><i>Holotype.</i>&mdash;Right maxilla bearing P3, P4, and M1, No. 11353 KU; bluff
+on west side of Turkey Creek, approximately 75 feet above stream, Raymond
+Farr Ranch, Center NE, NE, S. 48 Blk. C-3, E. L. and R. R. Ry. Co.,
+Donley County, Texas [approximately 6.5 miles north and 1 mile east of
+Clarendon], Clarendon fauna, Early Pliocene age. Obtained by W. W. Dalquest,
+on June 25, 1960.</p>
+
+<p><i>Referred material.</i>&mdash;Fragment of right lower mandible bearing m2, No.
+11354 KU (see fig. 2), found about two feet horizontally distant from the
+holotype in the same stratum as the holotype and on the same date by the
+same collector (a staff member of the Department of Biology of Midwestern
+University, Wichita Falls, Texas).</p></div>
+
+<p class="figcenter" style="width: 600px;">
+<img src="images/image001.png" width="600" height="387" alt="Fig. 1. Cynarctus fortidens, No. 11353 KU (Midwestern Univ. No. 2044).
+Lateral view of holotype × 1, and occlusal view of check-teeth × 2.
+
+Fig. 2. Cynarctus fortidens, No. 11354 KU (Midwestern Univ. No. 2045).
+Lateral view of right lower mandible and m2 × 1 and oblique occlusal view
+of m2 × 2." title="Fig. 1. Cynarctus fortidens, No. 11353 KU (Midwestern Univ. No. 2044).
+Lateral view of holotype × 1, and occlusal view of check-teeth × 2.
+
+Fig. 2. Cynarctus fortidens, No. 11354 KU (Midwestern Univ. No. 2045).
+Lateral view of right lower mandible and m2 × 1 and oblique occlusal view of m2 × 2." />
+<span class="caption">Fig. 1. Cynarctus fortidens, No. 11353 KU (Midwestern Univ. No. 2044).<br />
+Lateral view of holotype × 1, and occlusal view of check-teeth × 2.<br />
+<br />
+Fig. 2. Cynarctus fortidens, No. 11354 KU (Midwestern Univ. No. 2045).<br />
+Lateral view of right lower mandible and m2 × 1 and oblique occlusal view
+of m2 × 2.</span>
+</p>
+<p><span class="pagenum"><a name="Page_138" id="Page_138">[Pg 138]</a></span></p>
+
+<div class="blockquot"><p><i>Diagnosis.</i>&mdash;Size large (see measurements); no accessory cusp between
+protocone and paracone of fourth upper premolar; first upper molar longer than
+broad and lacking cingulum on part of tooth lingual to protocone.</p>
+
+<p><i>Comparisons.</i>&mdash;From <i>Cynarctus crucidens</i> Barbour and Cook (see page 225
+of Two New Fossil Dogs of the Genus Cynarctus from Nebraska. Nebraska
+Geol. Surv., 4(pt. 15):223-227, 1914; also pages 330 and 338 of Dental
+Morphologie of the Procyonidae with a Description of Cynarctoides, Gen.
+Nov. Geol. Ser. Field Mus. Nat. Hist., 6:323-339, 10 figs., October 31, 1938)
+<i>C. fortidens</i> differs in lacking, instead of having, an accessory cusp between
+the protocone and paracone of the fourth upper premolar and in lacking, instead
+of having, a cingulum on the part of P4 that is internal (lingual) to the
+protocone.</p></div>
+
+<div class="blockquot"><p><i>Remarks.</i>&mdash;The lower jaw and its second molar seem to be from an individual
+significantly larger than the holotype. Possibly the lower jaw and
+upper jaw are from two species but the lower jaw probably is from a male
+and the upper jaw from a female of the same species.</p>
+
+<p>Reasons for regarding <i>Cynarctus</i> as belonging to the family Canidae instead
+of to the family Procyonidae have been stated recently in detail by E. C.
+Galbreath (Remarks on <i>Cynarctoides acridens</i> from the Miocene of Colorado.
+Trans. Kansas Acad. Sci., 59(3):373-378, 1 fig., October 31, 1956) and need
+not be repeated here. Although some uncertainty remains as to the familial
+position of <i>Cynarctus</i>, we favor Galbreath's view that the genus belongs in
+the family Canidae.</p>
+
+<p>The holotype of <i>Cynarctus crucidens</i> is from Williams Canyon, Brown
+County, Nebraska. According to C. B. Schultz (<i>in litt.</i>, December 6, 1961),
+Williams Canyon is a tributary of Plumb Creek; the upper part of the Valentine
+formation and the younger lower part of the Ash Hollow formation are
+exposed in Williams Canyon; which one of these formations yielded the holotype
+of <i>C. crucidens</i> is unknown.</p>
+
+<p>On the basis of the correlation chart (Pl. 1 in Nomenclature and Correlation
+of the North American Continental Tertiary. Bull. Geol. Soc. Amer.,
+52(pt. 1):1-48, 1941) by H. E. Wood 2nd <i>et al.</i>, <i>C. fortidens</i> and <i>C. crucidens</i>
+are equivalent in age or <i>C. fortidens</i> is the younger.</p>
+
+<p>The rounded summits of the principal cusps of the teeth of <i>C. fortidens</i>
+suggests that it was mainly frugivorous instead of carnivorous&mdash;more frugivorous
+by far than the living gray fox, <i>Urocyon cinereoargenteus</i>, that is
+known to eat substantial amounts of fruits and berries. Indeed, no other
+canid that we know of has teeth so much adapted to a frugivorous diet as
+are those of <i>C. fortidens</i>. Its degree of adaptation to a frugivorous diet is
+more than in the procyonid genus <i>Nasua</i> but less than in the procyonid genus
+<i>Bassaricyon</i>.</p></div>
+
+<div class="blockquot"><p><i>Measurements</i> (of crowns) of <i>C. fortidens</i>.&mdash;P3-M1, length, 25.8 (millimeters);
+P4-M1, 18.9; P3, length, 6.2; P3, breadth, 2.8; P4, length of outer
+border, 9.3; P4, breadth, 7.05; M1, length, 9.7; M1, breadth, 9.3; m2, length,
+10.3; m2, breadth, 6.6; depth of mandible at posterior end of m2, 17; thickness
+of mandible, 7.1.</p></div>
+
+
+<p><i>Transmitted February 21, 1962.</i></p>
+
+
+<p class="center"><small>29-2890</small></p>
+
+<div>*** END OF THE PROJECT GUTENBERG EBOOK 30260 ***</div>
+</body>
+</html>
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