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Seventh and Revised Edition. +Crown 8vo, cloth, 5s.</p> +</blockquote> + +<blockquote> +<p class="advert">“We have no space to draw particular attention to the +multitude of good things to be found in this cheerful and +unaffected little book, from every page of which there breathes +a sincerity which is necessary to the enjoyment of such stories +as these.”—<i>Saturday Review.</i></p> + +<p class="advert">“The book is one which may be warmly recommended for +the simplicity with which it is written and the power of +observation which it displays.”—<i>Athenæum.</i></p> +</blockquote> + +<hr> +<blockquote> +<p class="adverttitle"><b>MORE ABOUT WILD NATURE.</b> With +Illustrations by the Author. Third Edition. +Crown 8vo, cloth, 5s.</p> +</blockquote> + +<blockquote> +<p class="advert">“No better book can be given to juvenile naturalists.”—<i>Graphic.</i></p> + +<p class="advert">“When this delightful book, full of observation, of tenderness, +and of humour, has been read, it becomes a friend, to be +visited often and cherished much.”—<i>World.</i></p> +</blockquote> +<hr> +<blockquote> +<p class="adverttitle"><b>INMATES OF MY HOUSE AND GARDEN.</b> +Illustrated by <span class="smcap">Theo. Carreras</span>. Crown 8vo, +cloth, 3s. 6d.</p> +</blockquote> + +<blockquote> +<p class="advert">“This is a charming book, and prettily illustrated.”—<i>Spectator.</i></p> + +<p class="advert">“A charming series of sketches.”—<i>Times.</i></p> +</blockquote> +<hr> +<p class="center">LONDON: T. FISHER UNWIN.</p> + + +<div class="chapter"> +<figure class="figcenter illowp49" id="i_004" style="max-width: 62.5em;"> + <img class="w100" src="images/i_004.jpg" alt="Plant with upright stem and +long elongated leaves growing in a hanging basket. From the ends of the +leaves hang down beaker-like outgrowths"> + <figcaption> + <p>NEPENTHES RAFFLESIANA.</p> + </figcaption> +</figure> +</div> + + +<div class="chapter"> +<p class="center" style="font-size: 150%;">GLIMPSES<br> +INTO PLANT-LIFE</p> +</div> +<p class="center" style="font-size: 80%;"><i>AN EASY GUIDE TO THE STUDY<br> +OF BOTANY</i></p> + +<p class="center" style="font-size: 60%; margin-top: 5.0em">BY</p> +<p class="center">MRS. BRIGHTWEN, F.E.S.</p> +<p class="center" style="font-size: 80%;"> +<i>Author of “Wild Nature Won by Kindness,” &c.</i></p> + +<p class="center" style="font-size: 90%; margin-top: 5.0em"> +WITH ILLUSTRATIONS BY THE AUTHOR AND<br> +THEO. CARRERAS</p> + +<p class="center blackletter" style="margin-top: 5.0em"> +London</p> +<p class="center">T. FISHER UNWIN</p> +<p class="center" style="font-size: 60%;">PATERNOSTER SQUARE</p> +<p class="center" style="font-size: 60%;">1897</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + + + + + <p class="center" style="margin-top: 10em; margin-bottom: 10em;"> + <span style="font-size:110%; font-weight: bold;"> + [<i>All rights reserved.</i>] + </span> + </p> +</div> + +<hr class="chap x-ebookmaker-drop"> + +<div class="chapter"> +<p class="center blackletter"> +To</p> +<p class="center gesperrt" style="font-size: 120%">SIR JOSEPH DALTON HOOKER,</p> +<p class="center" style="font-size: 90%">K.C.S.I., P.P.R.S., D.C.L., LL.D., ETC.,</p> +<p class="center" style="font-size: 90%"><i>Formerly Director of the Royal Gardens, Kew</i>. +</p> +</div> + + +<p style="margin-left: 2.0em"> + <span class="smcap">Dear Sir Joseph Hooker</span>,— +</p> + +<p style="text-indent: 5.0em">In conversation with you I have often been impressed +by your conviction of the importance of inducing young +people to observe the elementary facts of botany, and I have heard +you express your admiration of the efforts made in this direction, in +an earlier generation, by that distinguished botanist, Prof. Henslow. +You have assured me of your sense of the value of independent +observations made by young students, for themselves, in the simplest +and clearest language.</p> + +<p>By your own example the studies of beginners have often been +led in this direction, but in this little book, which I have ventured +to produce, and of which you have kindly accepted the dedication, +I have not attempted to compete with even your least ambitious +flights.</p> + +<p>All I have endeavoured to do is to prepare the minds of young +people for the study of botany by explaining in the simplest +language some of the elementary phenomena of plant-life. It +is an humble experiment, but made, as I believe, on lines which +are novel so far as they go, and essentially practical.</p> + +<p>If, as is too likely to be the case, I have fallen into any technical +errors, your good nature must not be held responsible for my fault.</p> + +<p class="right"> + <span style="margin-right: 7.5em;">Believe me to be,</span><br> + <span style="margin-right: 2.5em;">Yours very sincerely,</span><br> + ELIZA BRIGHTWEN. +</p> + +<p><i>April, 1897.</i></p> + + + +<hr class="chap x-ebookmaker-drop"> + +<div class="chapter"> +<p class="center"> +<i>⁂ The greater portion of this work has</i><br> +<i>appeared in serial form in “The</i><br> +<i>Girl’s Own Paper.”</i> +</p> + +<p><span class="pagenum" id="Page_9">[Pg 9]</span></p> +</div> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + <h2 class="nobreak" id="PREFACE"> + PREFACE + </h2> +</div> +<hr class="r5"> + +<div> +<img class="drop-cap" + src="images/i_009.jpg" + style="width: 12%; height: auto;" + alt="W"> +</div> +<p class="drop-cap"><span class="upper-case">When</span> I was a +young girl I can well +remember how much I longed for +some simple book that would help +me to learn, not merely the name of +a plant and what class and order it belonged +to, but something about its life-history.</p> + +<p>It seemed very wonderful that, if I put seeds +into the ground, dry and dead as they looked, +I might feel sure young plants would presently +come up; that, if I planted an acorn, a young +oak-tree would in due time be seen; but how all +this came to pass I could not discover. I had +access to an excellent library, but although I +searched there hour after hour, and found many +a learned book about plants, they might as well +have been written in Sanskrit for all I could +<span class="pagenum" id="Page_10">[Pg 10]</span>understand of their scientific pages. There seemed +nothing suited to the mind of a thoughtful child, +and although, since that long-ago time, endless +books have been written for young readers and +thinkers, delightful books, too, which meet the +needs of those who desire information and instruction, +I have not hitherto met with one that +makes the careful study of plant-life really interesting +and practicable for those young people +who may not have a teacher to help them in +their study.</p> + +<p>It has been my aim in the little volume I +now venture to send forth to my young friends, +known and unknown, to supply this deficiency. +I want to enable them to share the joy of spending +hours in a garden learning to understand the +structure of plants. I want to make them able, +when they see a bud, or a root, or a twig, to +know what the history of that object is, how it +comes to have the shape it takes, how it developed +into its present condition, and what its +next form will be.</p> + +<p>The possession of the simple facts which I have +tried to make plain to every intelligence in the +<span class="pagenum" id="Page_11">[Pg 11]</span>following pages will turn a country walk from a +useless lounge into a lively object-lesson, delightful, +from beginning to end, alike to teacher and +taught. Nor will I apologise for the simple +language which I have used, for my design has +been, while taking advantage of all the latest discoveries +of science, to use no terms and introduce +no ideas which cannot be made intelligible +to a thoughtful child. In the hope that even so +humble an effort as this may not be without a +use in enlarging and quickening a sense of that +infinite harmony which runs through every part +of the Creator’s marvellous plan of nature, I put +forth, not without a full sense of its inadequacy, +this little volume. It has, at any rate, given me +an excuse for endless hours of pleasure within +the precincts of my own woods and garden.</p> + +<p>I have to acknowledge great indebtedness to +Mr. J. W. Odell, F.R.H.S., whose wide knowledge +of botanical science has been of essential service +in ensuring, as far as possible, the accuracy of +my statements.</p> + +<p class="right"> + <span style="margin-right: 1.5em;">ELIZA BRIGHTWEN.</span> +</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum"><a id="Page_12"></a><a id="Page_13"></a>[Pg 13]</span></p> + + + <h2 class="nobreak" id="CONTENTS"> + CONTENTS + </h2> +</div> +<hr class="r5"> + +<table> +<tr> +<td class="tdc" colspan="3">CHAPTER I.</td> +</tr> +<tr> +<td></td> +<td></td> +<td class="tdr" style="font-size: 70%;">PAGE</td> +</tr> +<tr> +<td class="tdl" colspan="2"><span class="smcap">Adaptation</span></td> +<td class="tdr"><a href="#CHAPTER_I">25</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Plant collections—General view of vegetable growth—Lowest + forms of plant life—Water buttercup—Vallisneria—Water-lily—Mare’s-tail—Sea-weeds—Cacti—Orchids—Tillandsia—Seed + dispersion. +</td> +<td></td> +</tr> +<tr> + + +<td class="tdc extrahigh" colspan="3">CHAPTER II.</td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Roots</span></td> +<td class="tdr"><a href="#CHAPTER_II">47</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Root fibres—Hygroscopic water—Root hairs—The evil + of stagnant water—Binding roots—Carex arenaria—Psamma + arenaria—Cheddar cliffs—Correlation of roots—Land, + water, and air roots—Laurel root in well—Horizontal + stems—Poa bulbosa—Earth-nut—Potato—Onion—Air + roots—Hoya—Aëroids—Parasitic roots—Yellow + rattle—Clover dodder—Flax dodder—Mistletoe—Root-cap—Office + of the root—Growing mustard-seed—Strength + of roots. +</td> +<td></td> +</tr> + +<tr> + +<td class="tdc extrahigh" colspan="3">CHAPTER III. +<span class="pagenum" id="Page_14">[Pg 14]</span></td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Tree Stems</span></td> +<td class="tdr"><a href="#CHAPTER_III">73</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Expansion of bark—Shedding bark—Repairing injury—Epidermis—Fibro-vascular + bundles. +</td> +<td></td> +</tr> +<tr> + + +<td class="tdc extrahigh" colspan="3">CHAPTER IV.</td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Leaves</span></td> +<td class="tdr"><a href="#CHAPTER_IV">97</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + A leaf a digestive organ—Petiole and blade venation—Monocotyledons—Dicotyledons—Radical + leaves—Phyllotaxis—Epidermis—Stomata—Influence + of foliage upon climate—Eucalyptus—Mesophyll—Chlorophyllon—Pallisade + tissue—Protoplasm—Absorption of carbon-dioxide—Jewel-weed—Alchemilla—Growing + wheat—Vertical + leaves of eucalyptus cork layer—Fall of the leaf—Useful + products of trees. +</td> +<td></td> +</tr> +<tr> + + +<td class="tdc extrahigh" colspan="3">CHAPTER V.</td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Buds</span></td> +<td class="tdr"><a href="#CHAPTER_V">123</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Buds formed in summer—Hollow stalks of plane-tree—Terminal + and axillary buds—Spiral arrangement—Pollard + willow—Suckers—Dormant buds—Bulbils on lily stem—Protection + from cold—Leaf and flower buds—Folding + of embryo leaves and flowers. +</td> +<td></td> +</tr> +<tr> + + +<td class="tdc extrahigh" colspan="3">CHAPTER VI.</td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Flowers</span></td> +<td class="tdr"><a href="#CHAPTER_VI">149</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Floral envelopes—Buttercup—-Gamopetalous—Polypetalous—Epipetalous—Apocarpous—Woodsorrel—Syncarpous + pistil—Geranium-coloured bracts—Composite + flowers—Fading flowers—Flower expansion—Flower + dissection—Cruciferous plants—Papilionaceous flowers—Honey + glands—Entomophilous flowers—Catkins—Monœcious + and diœcious flowers—Arum—Protection + from insects and moisture. +</td> +<td></td> +</tr> +<tr> + +<td class="tdc extrahigh" colspan="3">CHAPTER VII. +<span class="pagenum" id="Page_15">[Pg 15]</span></td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Pollination</span></td> +<td class="tdr"><a href="#CHAPTER_VII">175</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Cross pollination in dog’s mercury and hazel—Pin-eyed + primrose—Maiden pink—Wind pollination—Araucaria—Pollination + by moths—Nottingham catchfly—Attraction + of scent—Bartsia—St. John’s wort—Japanese toad-lily—Pollination + by humming-birds. +</td> +<td></td> +</tr> +<tr> + + +<td class="tdc extrahigh" colspan="3">CHAPTER VIII.</td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Fertilisation</span></td> +<td class="tdr"><a href="#CHAPTER_VIII">193</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Pollen grains—Micropyle—Nucellus—Embryo-sac—Experiment + with lily pollen—Changes after fertilisation—Growth + of embryo—Albumen—Arillus in spindle-tree—Yew + and nutmeg—Persistent style of clematis—Ovules—Changes + in ovary of oak and datura. +</td> +<td></td> +</tr> +<tr> + + +<td class="tdc extrahigh" colspan="3">CHAPTER IX.</td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Fruit</span></td> +<td class="tdr"><a href="#CHAPTER_IX">209</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Wide meaning of the word fruit—Diversity in appearance + and character of various fruits—Dehiscent and indehiscent + pericarp—Epicarp—Chestnut involucre—Mesocarp and + endocarp—Pericarp—Achene—Strawberry an apocarpous + fruit—Pineapple—Irritating hairs—Resinous protection of + cones—Coiled stem of cyclamen—Seed-pod of ivy-leaved + toad-flax hidden in wall crevices. +</td> +<td></td> +</tr> +<tr> + + +<td class="tdc extrahigh" colspan="3">CHAPTER X.</td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Dispersion of Fruits and Seeds</span></td> +<td class="tdr"><a href="#CHAPTER_X">227</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Dispersion by hooks and spines—Martynia—Testa—Di + Quaglia—Burdock—Bedstraw—Dispersion by winged + seeds or achenes—Sycamore, pinus tribe, birch—Dispersion + by silky down or hairs—Dandelion, goat’s-beard, +<span class="pagenum" id="Page_16">[Pg 16]</span> groundsel—Willow-herb—Bird agency—Seas and rivers—Dispersion + by elastic force—Pansy—Balsam—Furze—Broom—Squirting + cucumber—Dispersion by hygrometric + sensitiveness—Barley—Feather-grass—Mexican insect—Dispersion + by sticky glands—Linnæa Borealis—Salvia + glutinosa—Plumbago—Ground-nut self-buried. +</td> +<td></td> +</tr> +<tr> + + +<td class="tdc extrahigh" colspan="3">CHAPTER XI.</td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Germination</span></td> +<td class="tdr"><a href="#CHAPTER_XI">249</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Testa and its various forms—Collomia—Axis, tigellum, + hypocotyle—Centre of growth—Double embryo—Three + conditions required for germination—Varying time in + different plants—Cedar of Lebanon—Railway-bank flora—Broad + beans—Cress seed—Tillandsia—Collecting seedling + trees—Fern and moss spores—Prothallium—Archegonium—Antheridium + germ cells—Capsular fruit of + mosses—Protonema. +</td> +<td></td> +</tr> +<tr> + + +<td class="tdc extrahigh" colspan="3">CHAPTER XII.</td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Physiology of Plants</span></td> +<td class="tdr"><a href="#CHAPTER_XII">273</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Processes of plant growth—Nutrition—Water and gas as + plant food—Water culture—Osmosis—Experiment + showing absorption—Nitrates—Insectivorous plants able + to absorb nitrogen to some extent—Second experiment + on absorption—Preparation of plant food—Water bouquet—Transpiration—Use + of Stomates—Respiration—Seeds + give off carbon-dioxide—Effect of light, darkness, and + heat—Reproduction—Protococcus—-Strawberry runner—Asexual + and sexual reproduction—Effect of cold upon + seeds. +</td> +<td></td> +</tr> +<tr> + + +<td class="tdc extrahigh" colspan="3">CHAPTER XIII.</td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Insectivorous Plants</span></td> +<td class="tdr"><a href="#CHAPTER_XIII">295</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Special purpose of each plant organ—Various modes by + which plants entrap insects—Sundew—Venus fly-trap—Trapping +<span class="pagenum" id="Page_17">[Pg 17]</span> insects by sticky hairs—By viscid glands—By + pitchers containing fluid—Sarracenias—Roridula—Utricularia—Nepenthes—Pinguicula. +</td> +<td></td> +</tr> +<tr> + + +<td class="tdc extrahigh" colspan="3">CHAPTER XIV.</td> +</tr> +<tr> + +<td class="tdl" colspan="2"><span class="smcap">Habit of Growth in Plants</span></td> +<td class="tdr"><a href="#CHAPTER_XIV">313</a></td> +</tr> +<tr> +<td></td> +<td class="tdj"> + Tropical forest—Perching orchids not parasites—Mistletoe—Yellow + rattle—Saprophytes—Murderer fig-tree—Mutualism—Corn + blue-bottle—Clover and bacteria—Symbiosis + in white poplar and fungus—Sea anemone and + algæ—Bryony tendril—Mycetozoa. +</td> +<td></td> +</tr> +<tr> + +<td class="tdl extrahigh" colspan="2"><span class="smcap">Glossary</span></td> +<td class="tdr extrahigh"><a href="#GLOSSARY_OF_SCIENTIFIC_WORDS">335</a></td> +</tr> +</table> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum"><a id="Page_18"></a><a id="Page_19"></a>[Pg 19]</span></p> + + + <h2 class="nobreak" id="LIST_OF_ILLUSTRATIONS"> + LIST OF ILLUSTRATIONS + </h2> +</div> +<hr class="r5"> + +<table> +<tr> +<td class="tdl"><span class="allsmcap">NEPENTHES RAFFLESIANA</span></td> +<td class="tdr"><a href="#i_004"><i>Frontispiece</i></a></td> +</tr><tr> +<td></td> +<td class="tdr" style="font-size: 70%;">PAGE</td> +</tr><tr> +<td class="tdl"><span class="allsmcap">WATER BUTTERCUP</span></td> +<td class="tdr"><a href="#i_036">36</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">VALLISNERIA</span></td> +<td class="tdr"><a href="#i_037">37</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">CATTLEYA WALKERIANA</span> (<i>A Brazilian Orchid</i>)</td> +<td class="tdr"><a href="#i_041">41</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">YOUNG SCOTCH FIR GROWING IN HOUSE-LEEK</span></td> +<td class="tdr"><a href="#i_044">44</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">CREEPING GRASS</span></td> +<td class="tdr"><a href="#i_056">56</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">POA BULBOSA</span></td> +<td class="tdr"><a href="#i_057">57</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">LILY BULBILS</span></td> +<td class="tdr"><a href="#i_060">60</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">CLOVER DODDER</span></td> +<td class="tdr"><a href="#i_064">64</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">SECTION OF ROOT CAP</span></td> +<td class="tdr"><a href="#i_068">68</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">TURKEY OAK BARK</span></td> +<td class="tdr"><a href="#i_076">76</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">SCOTCH FIR BARK</span></td> +<td class="tdr"><a href="#i_077">77</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">SNAKE-BARK MAPLE</span></td> +<td class="tdr"><a href="#i_078">78</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">TURKEY OAK STEM</span> (<i>struck by lightning</i>)</td> +<td class="tdr"><a href="#i_079">79</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">PLANE TREE BARK</span></td> +<td class="tdr"><a href="#i_080">80</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">HORSE-CHESTNUT BARK</span></td> +<td class="tdr"><a href="#i_081">81</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">WHITE POPLAR BARK</span></td> +<td class="tdr"><a href="#i_082">82</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">SYCAMORE BARK</span> +<span class="pagenum" id="Page_20">[Pg 20]</span></td> +<td class="tdr"><a href="#i_083">83</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">TREE-FERN BARK</span></td> +<td class="tdr"><a href="#i_084">84</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">STEM OF YELLOW WATER-LILY</span></td> +<td class="tdr"><a href="#i_087a">87</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">STEM OF WHITE WATER-LILY</span></td> +<td class="tdr"><a href="#i_087b">87</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">GROUP OF BEECH-TREES WITH INTERLACING STEMS +AND ROOTS</span></td> +<td class="tdr"><a href="#i_093">93</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">CONVOLVULUS</span></td> +<td class="tdr"><a href="#i_094">94</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">NETTED VEINS</span></td> +<td class="tdr"><a href="#i_100">100</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">MONOCOTYLEDON</span><a id="tn-001"></a></td> +<td class="tdr"><a href="#i_101">101</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">DICOTYLEDON</span></td> +<td class="tdr"><a href="#i_102">102</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">TAMARIND SEEDLING</span></td> +<td class="tdr"><a href="#i_103">103</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">STOMATA</span></td> +<td class="tdr"><a href="#i_107">107</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">LEAF SECTION</span></td> +<td class="tdr"><a href="#i_109">109</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">YOUNG SHOOT OF EUCALYPTUS</span></td> +<td class="tdr"><a href="#i_116">116</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">MATURE FORM OF EUCALYPTUS LEAVES</span></td> +<td class="tdr"><a href="#i_117">117</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">OAK IN WINTER</span></td> +<td class="tdr"><a href="#i_127">127</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">OAK IN SUMMER</span></td> +<td class="tdr"><a href="#i_129">129</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">HORSE-CHESTNUT</span></td> +<td class="tdr"><a href="#i_135">135</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">YOUNG BEECH</span></td> +<td class="tdr"><a href="#i_138">138</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">UNFOLDING ARUM LEAF</span></td> +<td class="tdr"><a href="#i_140a">140</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">UNFOLDING LEAVES OF HART’S-TONGUE FERN</span></td> +<td class="tdr"><a href="#i_140c">140</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">PEAR LEAF</span></td> +<td class="tdr"><a href="#i_140b">140</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">BUDS OF WAYFARING TREE</span></td> +<td class="tdr"><a href="#i_145">145</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">PRIMROSE</span></td> +<td class="tdr"><a href="#i_153">153</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">POINSETTIA</span></td> +<td class="tdr"><a href="#i_157">157</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">WINTER CHERRY</span></td> +<td class="tdr"><a href="#i_160">160</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">WALLFLOWER</span></td> +<td class="tdr"><a href="#i_162">162</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">SWEET-PEA</span></td> +<td class="tdr"><a href="#i_165">165</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">BIRCH FRUIT</span></td> +<td class="tdr"><a href="#i_168">168</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">WILD ARUM</span> +<span class="pagenum" id="Page_21">[Pg 21]</span></td> +<td class="tdr"><a href="#i_170">170</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">PRIMROSE</span></td> +<td class="tdr"><a href="#i_179">179</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">MAIDEN PINK</span></td> +<td class="tdr"><a href="#i_181">181</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">ARISTOLOCHIA</span></td> +<td class="tdr"><a href="#i_186">186</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">STAPELIA</span></td> +<td class="tdr"><a href="#i_188">188</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">HYPERICUM</span></td> +<td class="tdr"><a href="#i_189">189</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">TOAD-LILY</span></td> +<td class="tdr"><a href="#i_191">191</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">POLLEN-GRAINS</span></td> +<td class="tdr"><a href="#i_196">196</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">WHITE-LILY<a id="tn-025"></a> PISTIL. SECTION OF PISTIL</span></td> +<td class="tdr"><a href="#i_197">197</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">POLLEN TUBE</span></td> +<td class="tdr"><a href="#i_198">198</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">SECTION OF COCOA-NUT</span></td> +<td class="tdr"><a href="#i_203">203</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">SPINDLE-TREE</span></td> +<td class="tdr"><a href="#i_204">204</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">NUTMEG AND MACE</span></td> +<td class="tdr"><a href="#i_205">205</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">CLEMATIS OR TRAVELLERS’ JOY</span></td> +<td class="tdr"><a href="#i_206">206</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">SECTION OF PEACH</span></td> +<td class="tdr"><a href="#i_216">216</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">POPPY CAPSULE</span></td> +<td class="tdr"><a href="#i_217">217</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">WOODY PEAR</span></td> +<td class="tdr"><a href="#i_218">218</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">PINE-CONES</span></td> +<td class="tdr"><a href="#i_223">223</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">SEED-POD OF MARTYNIA</span></td> +<td class="tdr"><a href="#i_230">230</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">BIRCH SEED</span></td> +<td class="tdr"><a href="#i_232">232</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">PARACHUTE</span></td> +<td class="tdr"><a href="#i_234">234</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">DANDELION SEED</span></td> +<td class="tdr"><a href="#i_235">235</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">GOAT’S-BEARD</span></td> +<td class="tdr"><a href="#i_236">236</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">COCOS-DE-MER</span></td> +<td class="tdr"><a href="#i_238">238</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">BROOM AND SWEET-PEA PODS</span></td> +<td class="tdr"><a href="#i_242">242</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">STIPA PINNATA (FEATHER GRASS)</span></td> +<td class="tdr"><a href="#i_244">244</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">BIGNONIA SEED</span></td> +<td class="tdr"><a href="#i_252">252</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">DOUBLE EMBRYO OF ORANGE</span></td> +<td class="tdr"><a href="#i_255">255</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">BROAD BEANS</span></td> +<td class="tdr"><a href="#i_258">258</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">GROWING MUSTARD SEEDS</span> +<span class="pagenum" id="Page_22">[Pg 22]</span></td> +<td class="tdr"><a href="#i_261">261</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">BEECH COTYLEDONS</span></td> +<td class="tdr"><a href="#i_264">264</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">ACORN</span></td> +<td class="tdr"><a href="#i_266">266</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">HORSE-CHESTNUT</span></td> +<td class="tdr"><a href="#i_267">267</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">YOUNG DATE-PALM</span></td> +<td class="tdr"><a href="#i_269">269</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">TRANSFUSION DIAGRAM</span><a id="tn-002"></a></td> +<td class="tdr"><a href="#i_278">278</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">SKELETON LEAF</span></td> +<td class="tdr"><a href="#i_284">284</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">SUNDEW</span></td> +<td class="tdr"><a href="#i_300">300</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">VENUS FLY-TRAP</span></td> +<td class="tdr"><a href="#i_304">304</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">SARRACENIA FLAVA</span><a id="tn-003"></a></td> +<td class="tdr"><a href="#i_306">306</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">BLADDERWORT</span></td> +<td class="tdr"><a href="#i_308">308</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">PITCHER OF NEPENTHES RAFFLESIANA</span></td> +<td class="tdr"><a href="#i_309">309</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">BUTTERWORT</span></td> +<td class="tdr"><a href="#i_311">311</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">PERCHING ORCHID</span></td> +<td class="tdr"><a href="#i_317">317</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">RAFFLESIA ARNOLDII</span></td> +<td class="tdr"><a href="#i_320">320</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">GIANT COW-PARSNIP</span> (<i>Heracleum Giganteum</i>)</td> +<td class="tdr"><a href="#i_325">325</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">CORN BLUE-BOTTLE</span></td> +<td class="tdr"><a href="#i_326">326</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">BRYONY TENDRIL</span></td> +<td class="tdr"><a href="#i_331">331</a></td> +</tr><tr> +<td class="tdl"><span class="allsmcap">TRICHIA THROWING OUT SPORES</span></td> +<td class="tdr"><a href="#i_332">332</a></td> +</tr> +</table> +<div class="chapter"> +<p><span class="pagenum" id="Page_23">[Pg 23]</span></p> + +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“To me be Nature’s volume broad display’d,</div> + <div class="verse indent1">And to peruse its all-instructing page;</div> + <div class="verse indent1">Or, haply catching inspiration thence</div> + <div class="verse indent1">Some easy passage raptur’d to translate,</div> + <div class="verse indent1">My sole delight.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">Thomson.</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum"><a id="Page_24"></a><a id="Page_25"></a>[Pg 25]</span></p> + + + <h2 class="nobreak" id="CHAPTER_I"> + CHAPTER I + <br> + <i>ADAPTATION</i></h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_26">[Pg 26]</span></p> + + +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“My heart is awed within me, when I think</div> + <div class="verse indent1">Of the great miracle that still goes on</div> + <div class="verse indent1">In silence round me—the perpetual work</div> + <div class="verse indent1">Of Thy creation, finished, yet renewed</div> + <div class="verse indent1">For ever. Written on Thy works, I read</div> + <div class="verse indent1">The lesson of Thine own eternity.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">Bryant.</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_27">[Pg 27]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_I_1"> + CHAPTER I + <br> + <br> + <span style="font-size: 70%;">ADAPTATION</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_027.jpg" + style="width: 12%; height: auto;" + alt="T"> +</div> +<p class="drop-cap"><span class="upper-case">The</span> +study of plants appears to me +to be one of the most delightful and +instructive that can be taken up by +young people. It has this advantage over +many other pursuits that it can be carried +on almost everywhere, for, even if the student’s +lot is to live in a town, there are generally botanic +gardens within reach, and visits paid in the +country are made the more enjoyable when some +special study can be carried on in the daily +walks.</p> + +<p>Then collections of dried leaves and flowers +can be formed during the summer, and the arrangement +and classification of these will provide +pleasant winter occupation.</p> + +<p><span class="pagenum" id="Page_28">[Pg 28]</span></p> + +<p>I fear that many young people are apt to +consider botany a very dry study. They are +naturally repelled by the long words and many +technical terms used in describing plants.</p> + +<p>It has long been my belief that the study of +botany should be approached through the garden +rather than the schoolroom, beginning with a +country ramble which should be an object-lesson +opening out endless paths for future study.</p> + +<p>Our Heavenly Father has given us a beautiful +world to live in, and, when our eyes have once +been opened to observe what lies around us, +nature becomes like an exquisite book of pictures, +always revealing to us something new and wonderful +as we turn over each fresh page.</p> + +<p>It is suited to all ages; the baby child begins +by gathering daisies and buttercups, while older +children make wild-flower collections and perhaps +work in their own little gardens watching the +growth of seeds and slips.</p> + +<p>The beauty of ferns and mosses is sure to +lead to some painstaking study of those fascinating +growths.</p> + +<p>Later on the fact that all trees have flowers +<span class="pagenum" id="Page_29">[Pg 29]</span>comes as a surprise to the unobservant, and thus, +when rightly guided, young people can hardly +fail to love a pursuit that promises such endless +sources of interest.</p> + +<p>In the chapters that will follow on the subject +of plant life, I do not purpose to write for quite +young children, as my hope is that older readers +will explain what is written, and make it interesting +to the little ones as they walk in gardens +and fields, giving as it were object-lessons on +buds, leaves, and flowers, and training young +minds to search for themselves into the wonders +that lie around them.</p> + +<p>How much there is to learn about, even in +the simplest things, some of the succeeding +chapters will endeavour to show, for example:</p> + +<p>How young plants grow out of seeds;</p> + +<p>How those seeds are dispersed;</p> + +<p>How much is folded up in a bud;</p> + +<p>How flowers are formed;</p> + +<p>How the bark splits off different trees.</p> + +<p>Any one of these subjects would need very +careful, patient observation truly to understand +it.</p> + +<p><span class="pagenum" id="Page_30">[Pg 30]</span></p> + +<p>I stand as it were only on the threshold of +scientific research, and look with wonder at the +work of such a student as Darwin, who gave +twenty long years to observation of the common +earth-worm before he wrote his deeply interesting +book upon it. Again, we see Sir John Lubbock +giving years of his life to the growing of seeds +and their seed leaves, in order to learn exactly +how plants begin their life, and two very thick +volumes are required to contain the vast amount +of information he has thus obtained.</p> + +<p>These two examples will suffice to show that +the minutest objects in nature are worthy of +reverent attention, and if these chapters tend to +awaken young people to a perception of this +fact and act as a humble guide to new lines +of thought, I shall feel that they have not been +written in vain.</p> + +<p>I fear it is impossible to explain the processes +nature is carrying on in the plant-world without +occasionally using scientific words, but, when I +am obliged to do so I shall try to explain their +meaning,⁠<a id="FNanchor_1_1" href="#Footnote_1_1" class="fnanchor">[1]</a>, and when once we rightly understand +<span class="pagenum" id="Page_31">[Pg 31]</span>an exact expression we soon begin to use it, +because it is more convenient and often saves +repeating a long sentence.</p> + + +<p>I would ask my readers to try and obtain +from their gardens and fields the various objects +mentioned at the close of each chapter, and +compare them with the plates, learning all about +them as they read the letterpress.</p> + +<p>This will, I feel sure, add much interest to the +study, for having something to collect and examine +tends to lighten mental work and enables +us better to understand descriptive writing.</p> + +<p>In this introductory chapter I will simply take +a general view of vegetable growth and its +adaptation to the situation in which it is found.</p> + +<p>In many respects plants require the same +conditions as animals, birds, and insects; they +must have air, food, moisture and light in order +to attain healthy growth, and although they differ +from animals in being usually stationary, their +life is carried on in a very similar way. Let us +take a forest tree as a type.</p> + +<p>It is anchored in the soil by its roots which +are its feeding organs; through them it draws +<span class="pagenum" id="Page_32">[Pg 32]</span>up various kinds of nourishment from the earth +in which it stands.</p> + +<p>The roots by several chemical processes render +the elements they have taken up from the soil +fit for the nourishment of the tree; they send +it up through the stem and branches into the +leaves, and these being the breathing organs +have essential work to do in receiving from the +air, and giving out again, certain gases which +contribute largely to maintain the life and vigour +of the tree. Thus it grows year by year, producing +annually its flowers and seed, which is +the end and aim of all plant life.</p> + +<p>We can trace another analogy with animal life, +in the necessity for pure sweet air, plants +growing in a vitiated or smoke-laden atmosphere +soon showing unmistakable signs of weakness. +The stunted hedges and trees on the fringe of +London always remind me of the poor, ill-grown +children of the slums.</p> + +<p>Besides the plant life which we see around us in +the shape of trees, shrubs, and flowers, there are +lower and perhaps still more wonderful forms of +vegetable life affording endless fields of study.</p> + +<p><span class="pagenum" id="Page_33">[Pg 33]</span></p> + +<p>Mosses, lichens, and fungi we are familiar +with everywhere in the country, but below these +again are such growths as the green stain⁠<a id="FNanchor_2_2" href="#Footnote_2_2" class="fnanchor">[2]</a> which +makes the tree trunks in moist places as brilliant +in colour as the leaves themselves. Looked at +through a lens we see the colour arises from a +growing plant of extremely simple form, little +more in fact than a succession of cells, each living +and increasing “after its kind.”</p> + +<p>Again, if we consider the process of fermentation, +we find that when it is set up in a cask of +wine its action is due to the growth of a minute +vegetable that feeds upon the alcohol and sugar, +and by robbing the wine of those two elements +turns it into vinegar or acetic acid.</p> + +<p>A somewhat similar growth causes the thick +jelly-like substance we sometimes find in our inkglass +when it has been allowed to remain too long +without renewal; the minute germs floating in the +air have found the ink suitable to them, and thus +their mycelium⁠<a id="FNanchor_3_3" href="#Footnote_3_3" class="fnanchor">[3]</a> begins to form at the bottom of +the glass, to the great discomfort of the writer.</p> + +<p>The yeast with which our bread is fermented is +<span class="pagenum" id="Page_34">[Pg 34]</span>another of these minute plants, and consists of +oval cells which multiply with great rapidity when +placed in a pan of flour, and kept in a warm +atmosphere.</p> + +<p>By the careful study of these lower forms of +vegetable life, Pasteur, Koch, Frankland, and +others have discovered and classified the germs +or microbes,⁠<a id="FNanchor_4_4" href="#Footnote_4_4" class="fnanchor">[4]</a> as they are called, which give rise to +various diseases. In books upon the subject, their +different shapes are figured as they appear when +immensely magnified, so that we can see that which +will give rise to consumption, erysipelas, or cholera, +and one reads with deep wonderment of all that +science has ascertained of late years as to the +presence in the air of these seeds of disease +which are ever floating more or less around us. +But for the restraining hand of God, it appears +as if universal sickness and death would be our +fate.</p> + +<p>Leaving these lower forms of growth, we may +consider the three divisions into which plants are +naturally classed as to their duration of life.</p> + +<p>Annuals are those which grow and flower, and +<span class="pagenum" id="Page_35">[Pg 35]</span>form their seeds in one year, within which their +life-history is closed.</p> + +<p>Biennials produce leaves only in the first year; +by their aid they lay up stores of nutriment in the +form of tuberous roots, on this food they can exist +through the winter, produce flowers the following +summer, perfect their seeds, and then die.</p> + +<p>To this class we owe such useful plants as the +carrot, parsnip, beetroot, and many others which +afford us such nourishing vegetable diet.</p> + +<p>Perennial plants live on for an indefinite number +of years, flowering annually, in some cases dying +down to the root in autumn, and producing fresh +foliage the following year.</p> + +<p>Water plants seldom have a fixed root, but remain +floating, borne up and kept in position by the +water, their roots being the means by which, in +conjunction with the leaves, they derive nourishment +from air and water. It is well worth while +to observe the two forms of leaves in the water +buttercup. Those on the surface are three-lobed, +flat, and round, they absorb from the air such gases +as the plant requires; while the leaves beneath +the surface are divided into threads so as to offer +<span class="pagenum" id="Page_36">[Pg 36]</span>no obstruction to the flow of water and enable the +plant to collect needful food from the water. It +can vary the form of its leaves according to its +requirements, since in running streams it may +often be found with the hair-like leaves only.</p> + +<figure class="figcenter illowp72" id="i_036" style="max-width: 62.5em;"> + <img class="w100" src="images/i_036.jpg" alt="Plant growing under water, with +a part floating. The leaves under water have a very fine, strongly branched shape, +the floating ones are more rounded and lobed. Above the water are some flowers, +each having a crown of 5 leaflets."> + <figcaption> + <p>WATER BUTTERCUP.</p> + </figcaption> +</figure> + +<p>On the other hand, if its seeds are sown in moist +earth, the seedlings will grow and develop those +flat leaves only which are characteristic of land +plants. This water buttercup, therefore, gives us a +<span class="pagenum" id="Page_37">[Pg 37]</span>wonderful example of adaptation to surrounding +influences.</p> + +<figure class="figcenter illowp38" id="i_037" style="max-width: 45.25em;"> + <img class="w100" src="images/i_037.jpg" alt="Plant with very elongated +leaves. A few thin stems grow winding to the surface."> + <figcaption> + <p>VALLISNERIA.</p> + </figcaption> +</figure> + +<p>Adaptation is remarkably shown in the Vallisneria, +a grass-like water-plant, found in Southern +Europe;⁠<a id="FNanchor_5_5" href="#Footnote_5_5" class="fnanchor">[5]</a> it grows in freshwater lakes, rooted in the +<span class="pagenum" id="Page_38">[Pg 38]</span>mud, and yet its flowers need to be fertilised in the +air. In order to effect this, the small male flowers +detach themselves from their stems, and, rising +through the water, float about upon its surface. +The female flowers are borne on a stalk, spirally +twisted, so that it can uncoil and allow the flower +to reach the top of the water whether it be deep +or shallow. There the two kinds of flowers meet, +the seeds are formed and the stem coils up again +and brings the capsule below the surface, where +it gradually matures.</p> + +<p>The water-lily can grow a long or short stem +as the depth of the water may require to enable +its leaves to lie flat upon the surface. I have +gathered lily flowers in my lake with stems from +four to five feet long, where the plant happened +to be growing in deep water.</p> + +<p>In such plants as the mare’s-tail (<i>Hippuris +vulgaris</i>), we find the stem specially adapted to +a submerged life. Growing out of mud at the +bottom of a stream the plant upholds its slender +stalks by two different methods. Inside the epidermis +(or outer skin) a strand of rather tough +tissue running through the centre gives flexible +<span class="pagenum" id="Page_39">[Pg 39]</span>support, whilst the rest of the space is filled up +with very large air cells, which give such buoyancy +to the stems that even if they are three feet in +length they are kept upright in the water, rising +ten or twelve inches above the surface. It is a +valuable as well as a curious plant, as it has the +property of absorbing the gases emitted by stagnant +water, and tends thus to purify the air.</p> + +<p>The same power of adaptation is to be found +in sea-weeds. Those growing on rocky shores +having short fronds covered with fructification, +while out at sea, ribbons of oar-weed may be found +many yards in length, formed, like the gulf-weed, +of tough texture to bear the friction of waves and +storms.</p> + +<p>If we were travelling in a Mexican desert, we +should find those remarkable plants which can be +so well studied in the cactus-house at Kew Gardens. +Bearing in mind that for many months the plant +must do without a drop of rain, or in fact without +moisture of any kind, it has been necessary that +the leaf-surface should be reduced to prevent loss +of moisture by evaporation, and so spines take +the place of leaves, and the stems are encased +<span class="pagenum" id="Page_40">[Pg 40]</span>in a thick leathery skin, which protects the plant +from the burning heat of the sun. Very little +moisture escapes through this thick green epidermis; +therefore when rain falls the plants receive +and store up their liquid food, and live sparingly +upon it during the long periods of drought, which +last for three-quarters of the year. Some of these +cacti, as we see them at Kew, are tall, straight-stemmed +plants, others low-growing rounded +masses, little spiny cushions, almost like vegetable +hedgehogs.</p> + +<p>In the arid prairies of Texas, advantage is taken +of the watery stores of the cactus, for when other +supplies fail, its fleshy stems are cut open, and +horses and cows greedily devour the succulent +food, which answers the purpose of drink, as well +as affording nutritious fodder.</p> + +<p>Our British spurge-plants have green leaves, +a thin epidermis, and all the ordinary characters +of the plants of a temperate region, but by comparing +them with the spurges found in Madeira, +we see how climate causes adaptation to differing +conditions. One of these spurges growing in my +greenhouse has a tall column-like stem, no leaves, +<span class="pagenum" id="Page_41">[Pg 41]</span>and a thick leathery skin, which would enable it +to bear a hot, dry climate. It thus mimics the +giant cacti of Mexico.</p> + +<figure class="figcenter illowp60" id="i_041" style="max-width: 61.125em;"> + <img class="w100" src="images/i_041.jpg" alt="Plant with tuber formation +directly above the roots. An oval-shaped leaf with smooth edges grows +from a few tubers. The plant has 1 large flower with 6 petals"> + <figcaption> + <p><span class="allsmcap">CATTLEYA WALKERIANA</span> (<i>a Brazilian Orchid</i>).</p> + </figcaption> +</figure> + +<p>We may trace another contrast in our common +groundsel and the large succulent groundsels of the +Cape and the Canary Isles, with their thick fleshy +leaves, the difference in form and texture being +simply an expression of the wonderful modification +due to climate.</p> + +<p><span class="pagenum" id="Page_42">[Pg 42]</span></p> + +<p>The lovely tribe of orchids make the same +provision for long periods of drought. Many of +the species live in countries where the rainy season +lasts about six months, and is succeeded by as +many months of dryness and heat.</p> + +<p>The air-plants we obtain from these countries +have large pseudo-bulbs, that is, the stems are +enlarged so as to be storehouses of nutriment upon +which the plant exists, and by means of which +it brings out the gorgeous flowers which make +Brazilian forests such fairylands of beauty; every +tree-branch being laden with parasitic orchids, +their lovely blossoms lasting month after month +without the aid of rain or dew, because Nature +has provided each plant with its special store of +food, and has thus adapted it to the position it is +created to adorn.</p> + +<p>Another of these perching-plants is <i>Tillandsia +Usneoides<a id="tn-017"></a></i>, known in Florida as Spanish moss, and +often called “old man’s beard.” It hangs from +the tree-branches in tufts, like grey hair, and grows +in such profusion that it is collected and used for +stuffing cushions. This curious plant has no roots, +but simply hangs from the branches, and lives like +<span class="pagenum" id="Page_43">[Pg 43]</span>the orchids by absorbing water from the moist air +in the humid forests where it is found.</p> + +<p>The absorption by the long, hanging, grey roots +of the orchids in one case, and by the finely-divided +leaves and stems in the other, are both instances +of the wonderful way in which Nature “adapts” +the parts of a plant to its requirements.</p> + +<p>It often happens that seeds, blown hither and +thither by the wind, chance to fall upon places +which are quite unsuitable to their mode of growth; +then we have an opportunity of seeing how their +power of adaptation enables them to triumph over +almost insuperable difficulties.</p> + +<p>I have observed a tiny plant of groundsel +growing out of a chink in a wall where there +was scarcely any soil from which it could derive +nourishment, contriving to live on, however, and +make the best of its hard lot. Its stem, which +should have been a foot high, could only attain +about two inches, and instead of dozens of leaves +it had but four, and yet it survived and even +produced two small flowers, thus touchingly displaying +its power of adaptation.</p> + +<figure class="figcenter illowp60" id="i_044" style="max-width: 62.5em;"> + <img class="w100" src="images/i_044.jpg" alt="The top of a wall on which +a group of rosette-shaped plants grows. Between those plants grows a +small fir tree"> + <figcaption> + <p>YOUNG SCOTCH FIR GROWING IN HOUSE-LEEK.</p> + </figcaption> +</figure> + +<p>Another more remarkable instance which occurs +<span class="pagenum" id="Page_44">[Pg 44]</span>to me was that of a seedling Scotch fir, which had +rooted itself in a lump of house-leek on the top +of a garden wall. For eight years the young tree +managed to live and grow, until it became a symmetrical +<span class="pagenum" id="Page_45">[Pg 45]</span>well-branched fir-tree, almost twelve inches +high. By a supreme effort it produced a crop of +miniature cones, and soon after it died from +drought and starvation, the wonder being that it +could have lived so long upon the modicum of +food the barren wall supplied, besides having to +endure at times periods of scorching heat as well +as drought. The chief interest in this example is +centred in the fact that as soon as fruit-bearing +has been attained, then, and not till then, the little +tree died, showing how persistently under all +hindrances and difficulties a plant will endeavour +to carry out the purpose of its creation.</p> + +<p>We have seen in these instances some striking +examples of the way in which plant-life is adapted +to its surroundings. Our examples have been +such as are easy of attainment, and such as we +can verify with our own eyes; but even more +wonderful are the adaptations hidden away in +the recesses of the plant, and as we progress in +our study these arrangements of cells and tissues +will be revealed to us. In order however to see +them, and to understand their true significance, +we must proceed step by step to study the parts +<span class="pagenum" id="Page_46">[Pg 46]</span>of an ordinary plant; because it is only by first +mastering all we can of one part of a plant, and +then comparing that part with other plants, that +we can hope to gain real knowledge. Accordingly +in our next chapter we shall take the root as our +starting-point, and ascertain its functions and uses, +and the part it has to play in the economy of the +plant.</p> + +<p>Specimens to be obtained:—Green stain on treebark +(<i>Protococcus</i>); yeast; annual, biennial, and +perennial plants; water buttercup leaves; vallisneria; +water-lily stems; mare’s-tail plant; cacti; +spurge; orchids; tillandsia; plants growing in +wall crevices.</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_47">[Pg 47]</span></p> + + + <h2 class="nobreak" id="CHAPTER_II"> + CHAPTER II + <br> + <i>ROOTS</i> + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_48">[Pg 48]</span></p> + +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“While thus through all the stages thou hast push’d</div> + <div class="verse indent1">Of treeship—first a seedling, hid in grass;</div> + <div class="verse indent1">Then twig; then sapling; and, as century roll’d</div> + <div class="verse indent1">Slow after century, a giant bulk</div> + <div class="verse indent1">Of girth enormous, with moss-cushion’d root</div> + <div class="verse indent1">Upheaved above the soil.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">Cowper.</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_49">[Pg 49]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_II_1"> + CHAPTER II + <br> + <br> + <span style="font-size: 70%;">ROOTS</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_049.jpg" + style="width: 12%; height: auto;" + alt="L"> +</div> +<p class="drop-cap"><span class="upper-case">Let</span> +us begin our study of roots by +considering the way in which plants +obtain their nourishment from the +earth, and are kept in an upright position +by means of their root-fibres. These +being out of sight, we may easily not be familiar +with this part of the economy of plant life, but we +shall soon see what important duties the roots have +to fulfil, and how much they vary in character and +appearance according to the soil, the climate, and +the work they are required to do. The greater +number of annual plants (those which live only +one year) have fibrous roots, and of these we can +find examples almost everywhere. A piece of +groundsel or tuft of grass will answer our purpose. +<span class="pagenum" id="Page_50">[Pg 50]</span>On pulling it out of the ground we see a bunch of +whitish threads or fibres springing from the crown +of the plant (which is the junction between the +stem and the root), and on these slender fibres are +hairs which are really the active part of the root, +for it is only through these hairs that the rootlets +are able to absorb the liquid from the soil, the +fibres simply acting as channels to convey the +watery nourishment to the stem and leaves.</p> + +<p>Common earth consists of small particles of +mineral substances such as flint, chalk, or iron, and +also of such vegetable matter as decayed leaves +and rotten wood.</p> + +<p>The spaces between the particles are more or +less filled with air, each mineral particle being +enveloped with a film of water. However dry the +soil may appear, this will always be found to be +the case. It may be tested by weighing in an +agate balance some dry soil on a summer’s day. +There is a very delicate instrument called a +hygroscope, which can tell us when there is the +slightest amount of moisture in the air, and a +clever German writer, Von Sachs,⁠<a id="FNanchor_6_6" href="#Footnote_6_6" class="fnanchor">[6]</a> has termed this +<span class="pagenum" id="Page_51">[Pg 51]</span>film of water, which gathers round earth-particles, +hygroscopic water. It has been ascertained by +careful experiment that it is only on this delicate +watery film that the root-hairs of plants are able to +feed. As these hairs drain away the hygroscopic +film it is always being renewed by the free water +which comes from rain and dew. The free water +of the soil is constantly passing from the surface to +the subsoil, and by this action plant-food, in the +form of soluble earth salts, is presented to the +roots. The passage of the water is of the highest +service to the roots, since the warm air follows the +water through the soil, and helps to oxidise the +mineral particles; these are thus rendered soluble, +and are taken up by the fine films of water, and +so indirectly the roots are fed. If, however, there +is no outlet for the water and the soil becomes +water-logged this beneficial action is retarded, and +to land-roots the water is hurtful.</p> + +<p>We can now understand why stagnant water +in the ground is so injurious to plant-life, as it +prevents the needful air from coming into contact +with the roots, and this is the reason why farmers +are careful to remove the surplus water from their +<span class="pagenum" id="Page_52">[Pg 52]</span>fields by thorough drainage and ploughing. Roots +adapt themselves very wonderfully to their situation.</p> + +<p>This piece of grass, which we are examining, +if it grew in sandy soil, would have its root-fibres +covered with a downy growth to enable them the +more readily to absorb every particle of moisture +in the sand. Dr. Bonar speaks of the date-palm +as having this same characteristic. “These palm +roots are peculiarly fitted to obtain every drop of +water that the sand contains; they consist of long +fleshy strings or ropes, shooting straight down +into the sand, in numbers quite beyond our +reckoning, and extending over a large circle.”</p> + +<p>The tendency of fibrous roots to bind sand +together is taken advantage of on many of our +sea-coasts, where the sand blows inland and +renders acres of ground sterile and useless. +There, if the <i>Carex arenaria</i> (a kind of sedge) is +planted, its roots will spread far and wide, interlacing +and creeping through the sandy soil, until +in time the latter becomes solid and no longer +drifts inland.</p> + +<p>An allied species of grass, <i>Psamma arenaria</i> (or +<span class="pagenum" id="Page_53">[Pg 53]</span>marrem grass) grows abundantly at Bournemouth, +and wishing to ascertain how far one of its +underground stems extended, with some amount of +patience I disinterred about six or seven feet of it +in a bank on the sea-shore where it was accessible. +As it seemed to have no end, I could not ascertain +its entire length.</p> + +<p>Another instance of root growth adapting itself +to situations occurs to me. In visiting the +Cheddar Cliffs in Somersetshire I was struck by +the beauty of a plant which grew here and there +out of the crevices of the rocks. Its tufts of vivid +green leaves looked so healthy and vigorous I +could not help wondering how it could obtain +moisture enough to produce such foliage, placed +as it was high up on the dry face of a rock.</p> + +<p>Failing to reach its roots in any other way, I +climbed up to a spot where I could remove some +of the horizontal layers of stone. At last I lifted a +flat piece of rock just above one of those plants, +and there I saw at a glance the secret of its +vigorous growth:</p> + +<p>The roots had spread out over the surface of the +stone for a distance of eight or nine inches in a +<span class="pagenum" id="Page_54">[Pg 54]</span>perfectly flat layer of fine fibrous rootlets no +thicker than a sheet of paper; these would doubtless +suck up abundant moisture whenever the rain +beat upon the rocks, and there, pressed closely +between the two layers of stone the plant has its +water-supply stored up, and is enabled to look +fresh and green when other vegetation is suffering +from drought.</p> + +<p>In plant-life there is a marvellous variety in +root-structure. Roots differ much, not only in +form, but in texture and duration of life, so that to +gain a true knowledge of them we must carefully +examine those of herbs, shrubs, and trees, and +observation will soon teach us the fact that there +exists a close correlation between the form and +texture of the root and the size and character of +the plant. The external shape will depend principally +upon whether a tap-root is developed or no. +Such, for instance, as the carrot and the dock are +those of the true tap-root character. Of branching +roots we may find endless modifications amongst +ordinary field or garden flowers from the fibrous +roots of the little groundsel to the large fleshy +tubers of the dahlia. Between these two types +<span class="pagenum" id="Page_55">[Pg 55]</span>there are others of an intermediate kind, but it is +possible to recognise amongst common plants the +roots belonging to one or other of the types I +have described. For the purposes of study we +may broadly group roots into classes according to +their method of collecting and absorbing food. +Thus we find one group growing in soil and +feeding upon the soluble earth salts and moisture +of the soil. Another group will be found growing +in water, like the water-lily and pond weeds. A +third group simply hangs down in space from +some perching plant like the tropical orchid, whilst +a fourth and very small group consists of parasitic +roots, of which a very common example is the +mistletoe. We will now study each of these +groups separately.</p> + +<p>I have already spoken of some kinds of fibrous +roots, and may add that if the root of a land plant +is immersed in water, it will after a time develop a +different kind of fibre, capable of receiving nourishment +from water instead of earth. I remember +seeing an instance of this in the case of a laurel +bush which grew near a well in our garden. We +had occasion to examine the water, and found that +<span class="pagenum" id="Page_56">[Pg 56]</span>the laurel had thrown down its roots below the +surface, where they grew luxuriantly, finely subdivided, +of a delicate ivory white, owing to the +absence of light, and more than a yard in length. +They had adapted themselves to the duty of +absorbing water only, but had we replanted them +in earth they would have withered, from their +unfitness to take up the hygroscopic water of +which I have already spoken. On the other hand, +if the seeds of a plant formed to live in the water, +such, for instance, as the water-lily, are sown in +ordinary soil, they adapt themselves to the new +conditions, and are able to live on the hygroscopic +water they find around the particles of earth.</p> + +<figure class="figcenter illowp93" id="i_056" style="max-width: 62.5em;"> + <img class="w100" src="images/i_056.jpg" alt="Three tufts of grass, sprouting +from a stem that grows horizontally"> + <figcaption> + <p>CREEPING GRASS.</p> + </figcaption> +</figure> + +<p>Some plants send out a horizontal stem (culm) +along the ground, with a bud and some roots +<span class="pagenum" id="Page_57">[Pg 57]</span>growing out of it at regular intervals. Each of +these joints (or nodes) takes root and forms a +separate plant. What are called strawberry +runners are stems of this kind, and so are the +creeping stems of <i>Potentilla reptans</i>.</p> + +<figure class="figcenter illowp45" id="i_057" style="max-width: 42.875em;"> + <img class="w100" src="images/i_057.jpg" alt="A plant, consisting of bulbs +from which roots grow downward and stems and long elongated leaves grow upward."> + <figcaption> + <p>POA BULBOSA.</p> + </figcaption> +</figure> + +<p>I once found a plant of the latter growing on +a low wall, and, as I imagine, because it desired +to reach the ground and root itself there, it had +thrown down a stem a yard and a half long with +eight young plants growing upon it at intervals +<span class="pagenum" id="Page_58">[Pg 58]</span>ready to form so many colonies when they should +reach the ground.</p> + +<p>One may frequently find stems of various +grasses running along the ground, and taking +root at each joint. I have one such spray in my +herbarium, with twelve young plants upon it at +regular intervals.</p> + +<p>Some plants store up nourishment in their roots, +as may be seen in one of our common seaside +grasses (<i>Poa bulbosa</i>); this soon withers after +flowering, and becoming uprooted, its bulbs, which +are like small round cheeses strung together, may +be seen blowing about in the wind.</p> + +<p>With such a provision as this, the parent plant +is able to bear extremes of cold and drought.</p> + +<p>It is well for us that plants have this power of +storing up their food underground, for to it we +owe such useful tubers as the potato and Jerusalem +artichoke.</p> + +<p>One of our native plants, the earth-nut (<i>Bunium +flexuosum</i>), has a single round tuber which is eatable +when roasted, and is often dug up by children. +Long ago, when England was liable to famines, +even this small tuber was valued as a means of +<span class="pagenum" id="Page_59">[Pg 59]</span>eking out the labourer’s daily meal. It is worth +while to examine the curious divided tubers of +some of our common orchises, such as the spotted +orchis (<i>O. maculata</i>), or the meadow orchis (<i>O. +morio</i>). The tuber which produces the leaves and +flowers withers away at the end of the summer, +but it leaves behind it a second tuber in which is +stored up the nourishment required to enable it to +bring forth leaves and flowers in the following +spring.</p> + +<p>Tubers are in reality underground stems which +have thickened into rounded balls to contain plant +food.</p> + +<figure class="figcenter illowp100" id="i_060" style="max-width: 62.5em;"> + <img class="w100" src="images/i_060.jpg" alt="Part of the stem of a plant +with oval-shaped leaves with articulate veins. In the axils of the leaves grow +small rounded bulbs. A detail of one of the bulbs shows it has rootlets and +some compacted leaves"> + <figcaption> + <p>LILY BULBILS.</p> + </figcaption> +</figure> + +<p>If we examine a potato we shall see that it contains +true buds in the little hollows on its surface; +these are called “eyes,” and each of them if sown +in the ground will produce a new potato plant. If +a potato is left in a damp cellar, each of these eyes +will send out a stem, thus proving that the “eye” +has the nature of a bud. If we cut the potato in +half we shall see it is of an even substance mainly +composed of starch, but if we halve an onion it will +be found to consist of rings or layers of a thick +fleshy nature, which proves it to be a bulb and not +<span class="pagenum" id="Page_60">[Pg 60]</span>a tuber. The onion is like a large bud growing +underground, instead of on a tree branch. We can +prove how similar the onion and the bud are, by +searching on a lily stem for buds or bulbils, which +are often produced in the axils of the leaves; if we +plant such a bud it will throw out fibres and become +a bulbous-rooted plant. Some of our native grasses +seem to have a singular power of adapting themselves +<span class="pagenum" id="Page_61">[Pg 61]</span>to their position. For instance, the common +Timothy grass (<i>Phleum pratense</i>), which usually +lives by means of a fibrous root, can, if needful, +produce a bulb which enables it to keep living in +a very dry place, but if removed to a wet soil it +returns to a fibrous root. Other grasses have been +observed to alter their root-growth in the same way, +adapting themselves to their surroundings.</p> + + +<h3>Air Roots.</h3> + +<p>These absorb the watery vapour of the air; they +cannot adapt themselves to live in earth, but under +certain conditions they can put forth other kinds of +roots that are partially adapted for growing in soil.</p> + +<p>I may here give some personal observations +about a certain <i>Hoya</i> plant that came into my +possession so long ago as 1855. This muchenduring +plant lived in a hanging basket for many +years, in the dry air of a sitting-room. Its leaves +were sometimes shrivelled from lack of water, and +it never had vigour enough to produce flowers. At +last, after enduring this life for twenty years, it was +placed in a stove-house where the moist heat suited +its requirements. Then it flowered charmingly, and +<span class="pagenum" id="Page_62">[Pg 62]</span>even now is showing a further degree of enterprise +by growing a bunch of fibrous roots at the end of +one of its stems. I imagine it intends to plant +itself into another pot standing near. I am watching +it with much curiosity, because if it does this, +the old plant will prove that it has a high degree +of intelligence, and that although it remained +quiescent for so many years, it was only from lack +of opportunity to do more than quietly endure its +privations.</p> + +<p>In tropical countries, some plants and trees such +as <i>Monstera</i> and <i>Philodendron</i> send down slender +aerial<a id="tn-016"></a> roots called lianes, many hundred feet in +length.</p> + +<p>In the Aëroid House at Kew, I remember seeing +these lianes coming down from the roof of the +house in search of water and earthy nourishment. +It seemed like actual intelligence that directed +these roots to a tank of water twenty-five feet +distant from their starting-point above. Whilst we +are considering this subject, I may mention the +curious root action of a kind of fig-tree growing in +the tropics which is sometimes known by the name +of the “Murderer.” Its seed often falls, or is +<span class="pagenum" id="Page_63">[Pg 63]</span>dropped by birds, amongst the leaves in the head +of a palm-tree, there it begins to grow and forms +root after root, gradually descending the stem of +the tree and clasping it so tightly that at last the +palm is strangled and falls to the ground carrying +its destroyer with it, where it roots and grows into +a tree.</p> + + +<h3>Parasitic Roots.</h3> + +<p>As in human society there are thievish characters +who live by preying upon their neighbours, so in +vegetable society we find quite a number of different +plants growing at the expense of others, inserting +their roots into the stems and roots of trees instead +of drawing their nourishment from the ground. +Careful distinction must be drawn between such +plants as ivy, virginian creeper, clematis, lichens, &c., +which simply grow and climb on the bark of trees, +and the true parasites which are nourished by the +juices of the trees and plants into which their routs +penetrate.</p> + +<figure class="figcenter illowp100" id="i_064" style="max-width: 62.5em;"> + <img class="w100" src="images/i_064.jpg" alt="Three separate pictures. To the +left there are 3 germinating seeds, out of which grow thin threads. In the middle +there is a stem of a clover plant with several trefoil leaves and a composite +flower. The stem is entwined by a thread-like structure, bearing several groups +of very small flowers. To the right is a clover seedling with 2 trefoil leaves, +next to which a seed germinates from which a thread grows that has started to +reach for the clover."> + <figcaption> + <p>CLOVER DODDER.</p> + </figcaption> +</figure> + +<p>Some plants are only partially parasitic, such as +the cow-wheat (<i>Melampyrum</i>) and the yellow rattle +(<i>Rhinanthus</i>). These represent a very deceitful +<span class="pagenum" id="Page_64">[Pg 64]</span>kind of growth. To all appearance the plants are +getting an honest living, the leaves are perfectly +green and capable of performing all the duties of +<span class="pagenum" id="Page_65">[Pg 65]</span>leaves, and yet, if we remove a little of the soil the +plant will be found to be attached to, and growing +from the roots of some strong kind of grass, and is +deriving its nourishment from the food collected +by those grass roots.</p> + +<p>Yellow rattle grows abundantly in undrained +marshy fields, where it is easy to obtain the plant, +so as to examine its mode of growth. We may then +go on to a clover-field and seek for that true parasite +and most troublesome enemy to the farmer, +the clover-dodder (<i>Cuscuta trifolii</i>). Its seeds are +frequently mixed with the clover, and when +sown they germinate on the surface, but the +little thread-like stem, instead of entering the +ground, feels about in the air until it reaches a +young clover-plant. It soon clasps its victim +with its fast-growing stem; as the clover grows +the dodder coils around it and is carried away +from the ground.</p> + +<p>As the wiry stem gains strength, it developes a +series of suckers that eat into the clover stem and +rob it of the food it has collected; it lives, flowers, +and grows at the other’s expense. The rate of +growth of the dodder exceeds that of the clover, so +<span class="pagenum" id="Page_66">[Pg 66]</span>that the latter is both exhausted and choked by its +snake-like enemy.</p> + +<p>I once sowed a patch of flax in a garden, and not +knowing that it too had a parasitic enemy, I was +greatly puzzled to find quantities of pinkish threads +growing out of the flax stems. These threads bore +round bunches of tiny flowers. All this was very +pretty and interesting, but it resulted in my patch +of flax becoming a mass of interlacing threads and +dying a miserable death, fairly strangled by the +flax dodder. Another species of <i>Cuscuta epilinum</i>, +grows on furze and also on heather, it having the +twine-like stems by which dodder may readily be +known.</p> + +<p>We are all familiar with the mistletoe, its leathery +leaves and its white berries.</p> + +<p>This plant grows out of the branches of poplar, +hawthorn, and apple, and very occasionally upon +the oak.</p> + +<p>In France and Belgium the custom of bordering +the fields with single rows of Lombardy poplars +seems to favour the growth of mistletoe, for its large +green bunches form quite a feature in the landscape, +and cannot fail to be observed by the traveller as he +<span class="pagenum" id="Page_67">[Pg 67]</span>journeys in the railway train. I have been told that +mistletoe is sufficiently abundant to be used in +Normandy as cattle-food.</p> + +<p>If a mistletoe-berry is gently pressed upon a +young branch of an apple-tree, its own viscid juice +will cause it to adhere, and before long it germinates +and sends its roots into the tissues of the tree; +as it grows, it fuses with them, and derives all its +root nourishment from the substances in the branch. +Of course the tree is weakened by this parasite, the +sucking roots of which disturb the flow of the sap; +woody knots are apt to form, and not unfrequently +the branch is killed by the intruder which has +fastened upon it.</p> + +<p>Having touched upon the four principal kinds of +roots, we will now take a single root-fibre and +examine it more closely. It seems scarcely possible +that such a brittle, feeble thread should be able to +penetrate into the ground and make its way +amongst stones and sharp-edged fragments of earth +without being bruised or torn. The chief friction is +borne by the growing-point, and this always has for +its protection a root-cap; the section of the growing-point +of root-fibre given in the plate shows the outer +<span class="pagenum" id="Page_68">[Pg 68]</span>skin, called the epidermis, and over that is the root-cap +shaped like a thimble formed of small cells. +As they are worn away outside and become dead +tissue, owing to friction with the soil, the cells are +constantly being renewed from within. The root +is thus enabled to grow and perform its part in +maintaining the life of the plant. The presence of +this root-cap and the absence of +leaves are the marks by which a true +root is known and distinguished from +an underground stem. With a small +lens one can see this extinguisher-like +cap protecting the extreme point of +the root, and it is well to examine a +variety of specimens, and see how they +differ slightly in size and shape.</p> + +<figure class="figleft w20" id="i_068"> + <img class="w100" src="images/i_068.jpg" alt="A microscopic picture of plant +tissue. At the top are rows of cuboid cells. These rows narrow down to the +bottom. That part is covered by a cap of more oval cells."> + <figcaption> + <p>SECTION OF + ROOT CAP.</p> + </figcaption> +</figure> + +<p>The one especial office of the root is to absorb +liquid nourishment from the soil for the benefit of +the plant, and, as I have already explained, this is +done mainly through the hairs which grow upon +the fibres of the roots. For instance, there is no +absorption in old tree-roots, such as we sometimes +see above the ground, nor in carrots, turnips, and +parsnips, but thrown out from such bulbous plants +<span class="pagenum" id="Page_69">[Pg 69]</span>are the fibres and their hairs which enable them to +grow to maturity. We may naturally inquire how +the solid materials in the soil, which are needful to +the growth of the stem and leaves, can possibly be +taken up by these extremely minute hairs.</p> + +<p>We may look upon the earth as being a sort of +storehouse<a id="tn-019"></a> of indigestible, unprepared plant-food +which must be altered in its character before it will +be fit for absorption by the roots. Some substances, +such as sugar, will readily dissolve in water; others, +such as starch and sand, are insoluble, but the effect +of rain-water and atmospheric air passing through +the soil, converts this insoluble dormant food into +soluble active food.</p> + +<p>The root-hairs convey this food to the small +fibres, and through them as channels it passes on +to larger ones, until it reaches the stem and goes to +feed the growing leaves and flowers.</p> + +<p>In order to remain in a healthy state, roots must +absorb oxygen gas, and for this reason gardeners, +when they find the soil growing caked and hard on +the surface, dig and rake the flower-borders in order +that air may freely permeate the soil and find access +to the roots of the plants.</p> + +<p><span class="pagenum" id="Page_70">[Pg 70]</span></p> + +<p>Roots appear to be endowed with certain remarkable +attributes, about which learned books have +been written of late, giving the result of patient +investigation as to their power of movement, the +way in which they are affected by gravitation, the +influence of light, and other forces.</p> + +<p>The experiments of Darwin and other scientists +have revealed very singular facts about the movements +of plants. The term used to describe their +motion is one we must learn, as it frequently appears +in botanical works. Circumnutation we may translate +as wavering around, and it well describes the +curious way in which rootlets, for instance, are +always moving slowly from one side to the other, +describing a kind of oval zig-zag track through the +earth. The fibres appear to have a discriminating +power, enabling them to choose convenient crevices +through which to penetrate hard soil, to avoid stone, +and to seek out any attractive food which lies in +their way.</p> + +<p>As soon as roots emerge from the seed they at +once turn from the light and seek to bury themselves +in the earth; the plumule from which the +leaves will spring has exactly the reverse tendency, +<span class="pagenum" id="Page_71">[Pg 71]</span>and invariably seeks the light and grows upwards. +This can be proved by growing some mustard seeds +on a piece of flannel about the size of a shilling, +floating it on water in a saucer exposed to light +from a single window; as soon as the leaves appear, +they will lean towards the light, whilst the roots +will point towards the dark part of the room. +If a germinating seed is even placed with the +root uppermost, and the plumule pointing downwards, +it will very speedily right itself, the stem +will turn and grow up, and the root will seek +the ground.</p> + +<p>The amazing strength of growing tree roots can +be imagined when we watch a tree in full leaf +during a high wind. As the terrific force of the +gale sways the trunk backwards and forwards the +roots are subjected to an enormous strain. Like +great india-rubber cables they give and retract, and +when the wind subsides we find the trunk as rigid +as ever.</p> + +<p>If my readers will seek for the specimens +enumerated below, and compare them with the +remarks made in this chapter, they will have such +a general idea of the functions of roots as will, I +<span class="pagenum" id="Page_72">[Pg 72]</span>trust, enable them to enjoy the study of more +advanced works upon the subject.</p> + +<p>Specimens to be obtained and compared with +the descriptions in this chapter:—Sedge or marrem +grass growing on a sandy sea coast; plants growing +between layers of stone; tree roots at the edge of +a pond; strawberry runners; a plant of <i>Potentilla +reptans</i>; creeping grasses; <i>Poa bulbosa</i> roots from +the seaside; potato. Earth nuts; lily bulbils; +Timothy grass; cow-wheat (<i>Melampyrum</i>); yellow +rattle (<i>Rhinanthus</i>); clover dodder (<i>Cuscuta trifolii</i>); +flax dodder (<i>Cuscuta epilinum</i>); mistletoe. Root +fibres of various plants. Mustard seed sown on +flannel.</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_73">[Pg 73]</span></p> + + + <h2 class="nobreak" id="CHAPTER_III"> + CHAPTER III + <br> + <i>TREE STEMS</i> + + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_74">[Pg 74]</span></p> +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“If thou art worn and hard beset</div> + <div class="verse indent1">With sorrows, that thou wouldst forget,</div> + <div class="verse indent1">If thou wouldst read a lesson, that will keep</div> + <div class="verse indent1">Thy heart from fainting and thy soul from sleep,</div> + <div class="verse indent1">Go to the woods and hills! No tears</div> + <div class="verse indent1">Dim the sweet look that Nature wears.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">Longfellow.</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_75">[Pg 75]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_III_1"> + CHAPTER III + <br> + <br> + <span style="font-size: 70%;">TREE STEMS</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_075.jpg" + style="width: 12%; height: auto;" + alt="A"> +</div> +<p class="drop-cap"><span class="upper-case">A walk</span> +through a wood on a bright +day in February will afford us many +interesting intuitions about the growth +of trees.</p> + +<p>We are apt to think of winter as a +dead season, and long for summer days once +more, that we may pursue our botanical studies; +but as soon as February begins there is already a +secret work going on within the tree-stems, the sap +is rising from the roots, and this ascent is easily to +be traced if we look carefully at the trunks of those +trees, such as the oak, elm, and others, which have +rugged bark. The wood within is swelling; fresh +layers of material will, a little later on, be added to +the inner side of the bark as a result of this ascent +of the sap.</p> + +<p><span class="pagenum" id="Page_76">[Pg 76]</span></p> + +<p>As the bark is hard and inelastic, it cannot +expand in proportion, and therefore has to crack +and split in yielding to the internal pressure. If +we look for these fresh cracks, we shall see the +clean new bark within, which, before long, will +harden and become of the same shade of grey as +the rest of the stem.</p> + +<figure class="figcenter illowp56" id="i_076" style="max-width: 48.9375em;"> + <img class="w100" src="images/i_076.jpg" alt="Photograph of part of a tree trunk, +covered by rough bark."> + <figcaption> + <p>TURKEY OAK BARK.</p> + </figcaption> +</figure> + +<p>It is at this season, too, that the plane-tree sheds +off its fragments of bark in greatest quantity, as +<span class="pagenum" id="Page_77">[Pg 77]</span>one may plainly see in the London squares, where +this tree grows so remarkably well. Its stem is +always peeling more or less throughout the year, +and possibly that fact may be one of the reasons +of its flourishing so well in the midst of smoke +and fog.</p> + +<figure class="figcenter illowp54" id="i_077" style="max-width: 48.0625em;"> + <img class="w100" src="images/i_077.jpg" alt="Photograph of part of a tree trunk, +covered bu very rough bark, having deep grooves and some pellets that come loose."> + <figcaption> + <p>SCOTCH FIR BARK.</p> + </figcaption> +</figure> + +<p>Trees shed their bark in many different ways.</p> + +<p>A reference to the illustrations will show the +concentric rings of the horse-chestnut, the square +<span class="pagenum" id="Page_78">[Pg 78]</span>pieces of the sycamore, which are due to the cleavage +being both vertical and horizontal, the hexagonal +shape of the divisions of the Scotch fir, the +rugged bark of the Turkey oak, the sycamore and +other species.</p> + +<figure class="figcenter illowp56" id="i_078" style="max-width: 47.125em;"> + <img class="w100" src="images/i_078.jpg" alt="Photograph of part of two tree +trunks, having a striped bark."> + <figcaption> + <p>SNAKE-BARK MAPLE.</p> + </figcaption> +</figure> + +<p>Where a woodpecker or a nuthatch has bored +a hole into the living wood of a tree-stem, it is +interesting to watch how the injury is repaired. +New bark begins to form at the edges of the +wound, and to this a layer is added each year, until +<span class="pagenum" id="Page_79">[Pg 79]</span>at last the hole is filled up, and only a scar is left +to show where it once existed.</p> + +<figure class="figcenter illowp58" id="i_079" style="max-width: 48.0625em;"> + <img class="w100" src="images/i_079.jpg" alt="Photograph of part of a tree trunk +where the bark is damaged, showing the wood underneath."> + <figcaption> + <p><span class="allsmcap">TURKEY OAK STEM</span> (<i>struck by lightning</i>).</p> + </figcaption> +</figure> + +<p>I have been able to watch this repairing process +going on for twelve years in the case of a Turkey +oak, which was injured by lightning. I was watching +the progress of the storm from one of our +upper windows, and happened to be looking at +this particular tree in the park, when out of a lurid +cloud above it, a streak of forked lightning +<span class="pagenum" id="Page_80">[Pg 80]</span>descended upon the tree, and rent off the bark of +one side from the top to the bottom, carrying away +portions of it to a distance of fifty feet or more, +leaving a white gash which looked pitiful enough +for many months. Year by year a wave of new +bark rolls on, covering the bare place by slow +degrees, but it is never destined to be quite healed +in this case, for the inner wood was killed to some +extent by the lightning, so it has become a home +<span class="pagenum" id="Page_81">[Pg 81]</span>for the boring beetles, who are riddling it with +holes wherein to lay their eggs.</p> + +<figure class="figcenter illowp54" id="i_080" style="max-width: 47.875em;"> + <img class="w100" src="images/i_080.jpg" alt="Photograph of part of a tree trunk +with a smooth bark, having dark and light patches. From the dark patches peaces of +bark are coming loose."> + <figcaption> + <p>PLANE-TREE BARK.</p> + </figcaption> +</figure> + +<figure class="figcenter illowp56" id="i_081" style="max-width: 47.875em;"> + <img class="w100" src="images/i_081.jpg" alt="Photograph of part of a tree trunk +with many fine grooves, creating a pattern of pellets."> + <figcaption> + <p>HORSE-CHESTNUT BARK.</p> + </figcaption> +</figure> + +<p>Such a tree becomes a happy hunting ground +for the woodpecker, who is attracted by the insect +diet he finds there. The large holes he makes in +getting at his prey will let in the rain, so that after +a time the moist rotten wood forms a suitable +place for various fungoid growths, and all these +agencies work together for the destruction of the +<span class="pagenum" id="Page_82">[Pg 82]</span>wood until the tree becomes a hollow stem, and the +leafage above is solely produced by the sap carried +upward by the bark.</p> + +<figure class="figcenter illowp55" id="i_082" style="max-width: 48.25em;"> + <img class="w100" src="images/i_082.jpg" alt="Photograph of part of a tree trunk +with a smooth bark, containing many diamond-shaped darker patches."> + <figcaption> + <p>WHITE POPLAR BARK.</p> + </figcaption> +</figure> + +<figure class="figcenter illowp55" id="i_083" style="max-width: 48.0625em;"> + <img class="w100" src="images/i_083.jpg" alt="Photograph of part of a tree trunk +with a bark having many horizontal grooves and a few vertical ones, creating +the appearance of many rectangular scales."> + <figcaption> + <p>SYCAMORE BARK.</p> + </figcaption> +</figure> + +<p>Let us inquire a little more carefully into the +formation of a tree-stem, and the different parts of +which it consists. Some rather hard names are +given to the four principal parts of a tree trunk +but, by reference to the plate, and by knowing the +meaning of the names, I hope they will soon be +<span class="pagenum" id="Page_83">[Pg 83]</span>mastered, and then our future walks in the woods +will be fuller of interest than ever, when once we +understand something about the hidden work that +is being carried on in those grand old trunks +around us. A tree may be compared to a large +manufactory. As we stand outside the building +we see the brick walls and the roof, and smoke is +coming out of the chimneys. We know that a +great deal of work is being done inside, and carts +<span class="pagenum" id="Page_84">[Pg 84]</span>are leaving its doors laden with the products of the +machinery within, but how the work is done we +cannot tell from the outside. We perhaps desire +to obtain this knowledge, and under the guidance +of the manager, we are taken from room to room +and see the marvellous processes by which raw +material is converted into exquisite fabrics, or it +may be clay is turned into priceless china or +porcelain. We leave the building full of wonder +at the things we have seen, and those particular +<span class="pagenum" id="Page_85">[Pg 85]</span>manufactures will ever afterwards be invested with +a special interest for us, because we have seen +with our own eyes how they are produced.</p> + +<figure class="figcenter illowp56" id="i_084" style="max-width: 48.5em;"> + <img class="w100" src="images/i_084.jpg" alt="Photograph of part of a tree trunk +covered with many oval an heart-shaped dark patches, close to one another."> + <figcaption> + <p>TREE-FERN BARK.</p> + </figcaption> +</figure> + +<p>Just in the same way we shall look upon trees +in a new light, if we are able in some measure to +follow the processes nature is carrying on in them +year by year so as to ensure the foliage, flower, and +fruit, which minister so much to our pleasure and +profit.</p> + +<p>The four names we must learn about in order to +understand the formation of wood are these. First +the outer bark, called epidermis, from two Greek +words <i>epi</i> upon, and <i>derma</i> the skin. <i>Cortex</i>, a +Latin word meaning bark. Fibro-vascular bundles; +this long phrase refers to certain threads or fibres +which exist in stems and give them toughness and +elasticity. From such fibres in the flax plant we +obtain linen, and from the hemp fibres ropes are +made. <i>Fibro</i> comes from the Latin <i>fibra</i>, a thread +or fibre; and <i>vasculum</i> is Latin for a little vessel; +we know the word better, perhaps, in another +sense as <i>vasculum</i>, the tin box in which botanists +place their plant collections.</p> + +<p>These thread-like vessels are well called bundles, +<span class="pagenum" id="Page_86">[Pg 86]</span>because they exist in little masses in the substance +of the stem.</p> + +<p>Most young people know what is called King +Charles’s Oak in the stem of the brake fern, +so plainly seen when it is cut across with a +penknife. The dark markings are the ends of the +fibro-vascular bundles which happen to resemble +an oak tree in form, though some think them more +like an eagle with outstretched wings, so the fern +is named <i>Pteris aquilina</i>, from <i>aquila</i>, an eagle.</p> + +<p>The fourth word is pith, the white substance in +the centre of the stem, which can readily be seen +by dividing a piece of elder branch, when the +middle will be found full of white pith.</p> + +<p>When we have these four parts of the stem +clearly in our minds it will be possible to go on +with our study and learn about the spaces between, +which are filled with different kinds of cells.</p> + +<p>The honey-comb formed by bees consists of +small cells, little hollow spaces in which they store +the honey or bee-food. Woody structure consists +largely of cells of various shapes to contain sap +and other substances. A beautiful specimen of +cell net-work may be obtained by placing a thin +<span class="pagenum" id="Page_87">[Pg 87]</span>slice of either white or yellow water-lily stem on a +piece of glass and, holding it up to the light, a fine +sort of lace-work will be seen. These are the cells +which convey air and water through the stem up +to the leaves and flowers. Or if we examine a +flower petal with a magnifying glass we shall find +it to be entirely composed of minute cells.</p> + +<div class="col50"> +<figure class="figcenter illowp100" id="i_087a" style="max-width: 37.875em;"> + <img class="w100" src="images/i_087a.jpg" alt="Cross section of a stem showing +many white circles, that grow smaller to the periphery. Dispersed are some darker spots."> + <figcaption> + <p>STEM OF YELLOW + WATER-LILY.</p> + </figcaption> +</figure> +</div> + <div class="col50"> +<figure class="figcenter illowp85" id="i_087b" style="max-width: 31.1875em;"> + <img class="w100" src="images/i_087b.jpg" alt="Cross section of a stem showing some +large and several smaller white circles, in a compact dark matrix."> + <figcaption> + <p>STEM OF WHITE + WATER-LILY.</p> + </figcaption> +</figure> +</div> +<p>In these little spaces are stored very many +and very different materials, all necessary to the +growth of a tree; we shall try and learn about +them by degrees; at present we must endeavour +to obtain a clear idea of their structure.</p> + +<p>A tree-stem increases in size yearly by the +growth of fresh cells within the outer bark, and +this active increase of tissue is due mainly to what +is called the cambium layer, which is developed +only in the spring and summer and does not exist +<span class="pagenum" id="Page_88">[Pg 88]</span>in winter; it forms bast, or phloëm, on the outer +side next the bark, and on the inner side next the +pith it creates woody tissue.</p> + +<p>Our English lime tree has a layer of fibre beneath +the bark which is worth examination; it +is the same in character, but not so wide or strong, +as the bast which we import from Russia in mats +to protect vegetation from frosts. Squirrels are +very fond of this soft material; they strip it +cleverly off the branches of our lime trees to form +a warm lining for their nests.</p> + +<p>It is easily found by cutting the outer bark off +any small branch of lime within reach, when we +can peel off the inner layer of bast, or phloëm, as +botanists call it.</p> + +<p>The phloëm from the lace-bark tree of the West +Indies is like the finest possible net-work, and is +used for many ornamental purposes. <i>Liber</i> (Latin +for the inner rind of a tree) is another term applied +to this cell formation.</p> + +<p>The study of different forms of woody fibre will +be found most interesting.</p> + +<p>I obtained one of my best specimens of it by +placing a very old Swedish turnip in water for +<span class="pagenum" id="Page_89">[Pg 89]</span>some months until the soft parts had melted away +and only the round ball of fibre remained. If any +one wishes to follow my example I would suggest +placing the turnip and its pan of water in some +outhouse where its perfume will not incommode +any one. A maid came to me one day with a sad +account of a fearful smell which had been noticed +for some time in a lumber-room at the top of the +house, and very naturally she thought that the +plumber should be called in to remedy the evil. I +had almost forgotten my interesting skeleton, but +in due time I traced the odour to its right cause, +and the turnip was banished to a distant spot, +where many washings and some soaking in +chloride of lime changed it into a really beautiful +specimen of woody fibre. I possess now only a +quarter of it, for botanists have so earnestly +begged for pieces of it that I have been persuaded +to share it with them.</p> + +<p>I have sometimes picked up on the seashore old +cabbage-stems bleached to a delicate ivory white, +forming really beautiful instances of woody fibre. +These we can prepare for ourselves, if desired, by +soaking the stems in water until they can be +<span class="pagenum" id="Page_90">[Pg 90]</span>brushed perfectly clean, and then bleached by +mixing a little chloride of lime in water and +letting them soak in it till they are white and +free from odour.</p> + +<p>In a manufactory there must of necessity be a +series of windows on the different floors, not only +to let in light but for purposes of ventilation. +Now, the processes of tree-growth are carried on +without light in the stem, but air is necessary, and +it is supplied by means of small apertures called +lenticels. These are not open holes, but are more +like gratings which admit a small amount of air +through loosely-packed cells.</p> + +<p>These lenticels are the small brown specks +which may be traced in great numbers on the +young branches of almost any tree. They remain +open through the spring and summer, admitting +the needful air to the interior of the bark, but +when the tree-growth is over for the season, and +air is no longer needed, a layer of cork forms +within the lenticel which entirely shuts it up and +keeps out the wintry cold. Thus it remains sealed +up till, by the growth of the cambium layer in the +following spring, the corky barrier is split open +and air is again admitted.</p> + +<p><span class="pagenum" id="Page_91">[Pg 91]</span></p> + +<p>These lenticels are nature’s ventilators, opening +and shutting in this curious way in order that the +manufacture which is going on beneath the bark +may receive from the outer air the various gases +essential to the work which is being carried on +within.</p> + +<p>I have said that many and various things are +stored in the stem-cells of trees. It would occupy +too much space to attempt to make anything like +a complete list of the liquids and solids which are +obtained from trees, but I will enumerate a few of +those with which we are familiar from their usefulness +in every-day life.</p> + +<p>Turpentine is obtained from various kinds of +firs—the Scotch fir, larch, and others. Burgundy +pitch from the spruce fir. A kind of tar is also +prepared from Scotch fir and larch. From various +kinds of cinchona we obtain quinine, so valuable +as a remedy for fever. Camphor is a product of a +Chinese tree. Tannin, by which skins are converted +into leather, is obtained from the bark of +the oak-tree. A kind of sugar is made from the +sap of the maple, which is largely used in America. +Gum arabic and a great number of gums used in +<span class="pagenum" id="Page_92">[Pg 92]</span>medicine are produced by foreign trees of various +kinds. The interior pith of a West Indian palm +tree produces the sago of commerce.</p> + +<p>Stems, like every other part of a plant, are to be +seen in endless variety when we come to examine +them for ourselves. In common garden plants +such as the calceolaria and petunia, the consistence +is soft, and such stems are known as herbaceous; +these generally die down in autumn. Roses and +rhododendrons have stems of a harder and more +rigid character, and seem to be intermediate between +the soft herbaceous stems and tall tree trunks.</p> + +<p>If in some country ramble we resolve to make +the trunks and bark of trees our study, we shall +find much that is interesting and well worthy of +observation.⁠<a id="FNanchor_7_7" href="#Footnote_7_7" class="fnanchor">[7]</a></p> + +<figure class="figcenter illowp100" id="i_093" style="max-width: 62.5em;"> + <img class="w100" src="images/i_093.jpg" alt="Photograph of a group of tree trunks +growing from a web of entangled roots."> + <figcaption> + <p>GROUP OF BEECH-TREES WITH INTERLACING STEMS AND ROOTS.</p> + </figcaption> +</figure> + +<p>The Lombardy poplar, with its tall bending stem, +the graceful willow and the silver birch, contrast +strongly with the thick and sturdy trunks of the +elm and oak. Even these two differ, the wood of +<span class="pagenum" id="Page_93">[Pg 93]</span>the elm being short and brittle, whilst that of the +oak is hard and flexible. Again, we may note the +slender drawn-up stems of trees growing thickly +<span class="pagenum" id="Page_94">[Pg 94]</span>together in a wood, where light and air are in a +measure shut out, and compare them with other +specimens standing in a park in free air and light. +There we see trees, growing as nature intended, with +grand sturdy trunks and welldeveloped +branches spreading out +on all sides. Lastly, in this chapter, +we may note the climbing +stems; these are especially numerous +and diversified in their +manner of growth. Almost every +part is modified and adapted to +assist the stem to climb. The +common ivy develops upon the +surface of its stem numerous +rootlets, and by their clasping +nature the ivy is enabled to +ascend the smoothest tree-trunk. +The hop and the convolvulus +climb by means of their habit of twining around +some rigid stem or twig. Then the peas and +vetches send out little clasping tendrils in the +place of leaflets, whilst that lovely ornament of +the hedges—traveller’s joy—climbs by occasionally +<span class="pagenum" id="Page_95">[Pg 95]</span>using the leaf stalk for a clasping holdfast. Not +less interesting are the plants that climb by means +of their hooks; the common bramble is of this +kind; it scrambles over the hedge in a very enterprising +and aggressive manner, while its spines +and hooks effectually prevent it from slipping back.</p> + +<figure class="figleft w20" id="i_094" style="max-width: 27.875em;"> + <img class="w100" src="images/i_094.jpg" alt="Stem or stick entwined +by a thin stem from which heart-shaped leaved grow and some flowers. The leaves +show their veins. The flowers are funnel-shaped and have some 5 or 6 petals."> + <figcaption> + <p>CONVOLVULUS.</p> + </figcaption> +</figure> + +<p>A very highly developed organ of climbing +is that to be found upon the stems of the +small Virginian creeper (<i>Ampelopsis Veitchii</i>). +On the points of its small tendrils we shall discover +little globular, crimson-coloured pads, which, +when pressed against a tree or wall, secrete +a kind of vegetable glue. This fixes the +tendril and enables the weak slender stem to +climb upwards. In these instances, as well as +others mentioned earlier in the chapter, we +have again evidences of how wonderfully plants +are adapted to their wants and environment.</p> + +<p>Things to observe and collect:—The various +ways in which trees shed their bark; how trees +repair holes in the stem; fibres in flax stem +and in hemp; specimens easily obtained by +sowing linseed and hemp-seed; section of brake-fern +stem; section of elder stem; thin section of +<span class="pagenum" id="Page_96">[Pg 96]</span>white or yellow water-lily stem; flower petal; +piece of bast matting; West Indian lace-bark; +turnip and cabbage stalk prepared as specimens of +woody fibre; lenticels on various trees; suitable +leaves for skeletonising—holly, magnolia, tulip-tree, +pear, poplar, aspen, mahonia, plum and +maple; suitable capsules—poppy, stramonium, +henbane, winter-cherry, campanula, and the calyces +of the yellow-rattle.</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_97">[Pg 97]</span></p> + + + <h2 class="nobreak" id="CHAPTER_IV"> + CHAPTER IV + <br> + <i>LEAVES</i> + + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_98">[Pg 98]</span></p> +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent20">“These naked shoots</div> + <div class="verse indent0">Barren as lances, among which the wind</div> + <div class="verse indent0">Makes wintry music, sighing as it goes,</div> + <div class="verse indent0">Shall put their graceful foliage on again,</div> + <div class="verse indent0">And more aspiring, and with ampler spread,</div> + <div class="verse indent0">Shall boast new charms, and more than they have lost.</div> + <div class="verse indent0">Then each, in its peculiar honours clad,</div> + <div class="verse indent0">Shall publish even to the distant eye,</div> + <div class="verse indent0">Its family and tribe.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">Cowper.</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_99">[Pg 99]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_IV_1"> + CHAPTER IV + <br> + <br> + <span style="font-size: 70%;">LEAVES</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_099.jpg" + style="width: 12%; height: auto;" + alt="W"> +</div> +<p class="drop-cap"><span class="upper-case">We</span> +have learned in the previous chapters +that the roots are the means by +which a plant gathers out of the earth +the various constituents which are needful +to maintain its life.</p> + +<p>The leaves have also to do their part in collecting +from the air such gases as are required to +effect the processes carried on within the substance +of the leaf.</p> + +<p>The leaf is really the digestive organ of the +plant; it feeds, breathes, and gives off in the form +of vapour any excess of water not required for its +work. For these purposes sunlight and air are +necessary.</p> + +<p>A leaf consists of a stalk, called a petiole, and +the flat green part, which we may call the blade.</p> + +<p><span class="pagenum" id="Page_100">[Pg 100]</span></p> + +<figure class="figleft w30" id="i_100" style="max-width: 33.4375em;"> + <img class="w100" src="images/i_100.jpg" alt="An oval shaped leaf +with a pointed tip showing its veins. Those are arranged in a feather-like +branching structure, sprouting from a central vein that is placed straight from +the base to the tip. Near the edge the veins connect to each other."> + <figcaption> + <p>NETTED VEINS.</p> + </figcaption> +</figure> + +<p>If we hold a leaf up to the light we see a +network of veins, and it is by their help the leaf +becomes a broad expansion of tissue, so exposed +that it gets the fullest possible benefit from the +sunlight and air. This fibrous +network gives strength to the +leaf, and answers to the bones +in animal structure.</p> + +<p>The fibro-vascular bundles, +which we see in the stem, go +up through the petiole, and +branch out in a beautiful and +regular manner. We may observe +this arrangement very +clearly in a skeleton leaf, the +midrib<a id="tn-020"></a> forming a backbone to +the whole structure, while the +smaller veins tend off to the +edge of the leaf, and then +overlap so as to form a +system of girders supporting the edge, and preventing +the wind from tearing the delicate tissues +into shreds.</p> + +<p>The arrangement of leaf network is called +<span class="pagenum" id="Page_101">[Pg 101]</span>venation, and by a glance at it we can at once +see to which of the great divisions in botany a +plant belongs. If the fibres are straight and run +parallel to each other without being netted, then +we know the leaf is that of a plant which begins +its life with only one seed-leaf; such +are all the species of corn and grass, +bulbs, palm-trees, bananas, and others.</p> + +<figure class="figright w20" id="i_101" style="max-width: 20.1875em;"> + <img class="w100" src="images/i_101.jpg" alt="A seed from which a young plant +emerges. At the bottom there are some roots. The plant has one leaf, that forms +a sheath around the small, growing stem."> + <figcaption> + <p>MONOCOTYLEDON.</p> + </figcaption> +</figure> + +<p>The long name applied to this +division of plants must be explained, +as it is a term we cannot +do without, and I must own it looks +formidable until we understand its +meaning.</p> + +<p>The first leaf that comes out of a +seed is called a cotyledon, from +<i>kotúle</i>, a cavity, or cup. The Greek +for one is <i>mónos</i>, so plants with one +seed-leaf are called monocotyledons.</p> + +<p>If we sow a date-stone or a few seeds of Indian +corn in moist soil they will grow readily, and +afford us nice little specimens of a one-seed leaf-plant.</p> + +<p>If we see that a leaf has netted veins, then we +<span class="pagenum" id="Page_102">[Pg 102]</span>know its seed produced two leaves at first,⁠<a id="FNanchor_8_8" href="#Footnote_8_8" class="fnanchor">[8]</a> so +plants belonging to this great division are called<a id="tn-018"></a> +dicotyledons.</p> + +<p>In order to watch the growth of two-leaved +seedlings, we may select a broad bean, or some +<span class="pagenum" id="Page_103">[Pg 103]</span>of the seeds out of tamarind jam; either will +grow readily in a pot of earth, if it is placed in +a sunny window, or near a stove, and kept moist.</p> + +<figure class="figcenter illowp50" id="i_102" style="max-width: 60.0625em;"> + <img class="w100" src="images/i_102.jpg" alt="A seed, from which one long root +grows downward. Equally long is the stem that grows upward. The stem has four +leaves, grouped in pairs. The bottom pair have elongated shapes with smooth edges and +veins that are arranged longitudinally. The upper pair have more of a heart shape +with serrated edges. Their veins are arranged in a feather-like structure."> + <figcaption> + <p>DICOTYLEDON.</p> + </figcaption> +</figure> + +<p>Orange and lemon pips may sometimes be +found sprouting within the fruit, and either of +these seeds will germinate, and form charming +little evergreen plants to brighten a town window-ledge.</p> + +<figure class="figright w30" id="i_103" style="max-width: 34.6875em;"> + <img class="w100" src="images/i_103.jpg" alt="A germinating seed, from +which a root grows into the soil. The seed is lifted above the ground. It +consists of 2 lobes and in between are 2 small, feather-like leaves."> + <figcaption> + <p>TAMARIND SEEDLING.</p> + </figcaption> +</figure> + +<p>Now we need not be afraid of those two long +words which are used to describe +one-leaved and two-leaved +seedlings, since we +know their meaning, and it +will be interesting when we +come across some new plant +to see to which division it +belongs, because knowing that will mean knowing +a great deal besides.</p> + +<p>All our English trees (with the exception of the +firs, which have many seed-leaves) are dicotyledons; +they increase their stems from the outside, +and are therefore called exogens, and most of our +plants belong to this division.</p> + +<p>The monocotyledons increase from the centre, +<span class="pagenum" id="Page_104">[Pg 104]</span>that is to say, the second leaf grows out of the +first, and the third leaf and its stem grow out of +the sheath of the second leaf, and so on; and this +is the law of their growth, whether they be corn +plants or palm-trees. These sheathing leaves and +the straight veins will always enable us to recognise +a one-seed leaf-plant at sight.</p> + +<p>The development of the stem has a marked +influence upon the arrangement of the leaves; +these, in such plants as the cyclamen, sundew, or +primrose, are said to be radical; that is, growing +from the root. Close observation will reveal the +cause to be the non-development of the internodes, +the leaves being crowded upon a very short, suppressed +stem, and thus we get the beautiful little +rosettes we find in the daisy and plantain. When +the stem is of greater length the leaves are ranged +at definite intervals, the spaces between the leaves +(the internodes) varying in length in proportion to +the size of the leaf. Small leaves are thus much +thicker upon the tree than larger ones. This will +readily be seen if we compare a branch of sycamore +with one of elm, the former having its large +leaves much further apart than the latter.</p> + +<p><span class="pagenum" id="Page_105">[Pg 105]</span></p> + +<p>Then, also, the arrangement of leaves upon the +stem (<i>phyllotaxis</i>) varies much. If we take a spray +of beech we shall find that its buds are placed +alternately on either side of the stem, so that the +third bud is exactly below the first, and the second +bud is in a line with the fourth, and so on. This +is also the plan of the elm, hazel, lime, hornbeam, +and many other trees. In the alder and whitebeam +the buds occur in three rows, and in some of +the willows in series of eight.</p> + +<p>The leaves of the horse-chestnut are borne in +pairs on alternate sides of the stem, and this plan +is common to a number of plants, especially those +of the type of the dead nettle and speedwell.</p> + +<p>Quite a distinct arrangement is that to be found +in the woodruff and bedstraws, where the leaves +are placed in a ring (a whorl) at regular intervals on +the stem.</p> + +<p>The botanical student should carefully observe +the differing methods of leaf arrangement, since, +as branches are developed from buds, the varying +order in their position must naturally modify the +general aspect of a tree, and has also much +physiological importance. We shall find that +<span class="pagenum" id="Page_106">[Pg 106]</span>buds are so placed that each leaf shall receive +its full share of sunlight and air, for it needs this +position in order to enable it to carry out the +wonderful work of assimilation which it has to +perform.</p> + +<p>The upper surface of a leaf is covered by a thin +layer of cells, known as the epidermis (or skin); +this does not prevent the light from falling +through, and its outer surface is protected by a +thickening, known as the cuticle. This is of great +use in controlling the escape of moisture, otherwise +the leaf would soon shrivel up in a hot sun. +In a young seedling leaf the cuticle is not developed, +and it can therefore breathe out moisture +very rapidly; later on, when the cuticle is formed, +it controls the escape of moisture, which can then +only exude through the under surface of the leaf.</p> + +<p>We can easily peel off a portion of the skin +from the under surface of the leaf, and if we place +it in a little water between two pieces of glass +and look at it in a microscope we shall see that +it consists of an extremely thin layer of cells, +with numbers of little openings called stomata +(from the Greek <i>stoma</i>, a mouth), answering +<span class="pagenum" id="Page_107">[Pg 107]</span>somewhat to the lenticels to be found in young +tree-stems, only those are solely for the admission +of air, while these little mouths are to let in and +out not only air, but water, vapour, and oxygen.</p> + +<p>These stomata look like little crescent-shaped +slits with a curved cell on either side, and as +they curve more or less, the mouths are opened +or shut as the plant may require. These little +mouths play a very important +part in the economy of the leaf, +and they exist in immense quantities +on its under surface.</p> + +<figure class="figright w30" id="i_107" style="max-width: 27.625em;"> + <img class="w100" src="images/i_107.jpg" alt="A microscopic image showing +vertical lines, that picture cellular borders.There are 3 openings between +the cells that have an elliptical shape and each are bordered by 2 banana-shaped +structures that join at the tips and thus enclose the opening."> + <figcaption> + <p>STOMATA.</p> + </figcaption> +</figure> + +<p>It has been calculated that a +million stomata exist on a single +leaf of the lime tree. When the +root has taken up more moisture than is required, +then it is the office of these pores, or stomata, in +the leaf to give out this extra water in the form +of vapour, and we can thus see how the action of +leaves must influence climate. If forests are recklessly +cut down, the bare country, with no foliage +to throw moisture into the air, may become an +almost barren desert, and again in marshy places, +where the air is too damp, a wise reduction in the +<span class="pagenum" id="Page_108">[Pg 108]</span>number of trees may alter the climate to a healthy +condition.</p> + +<p>Remarkable results have been obtained by planting +the Australian gum-tree, <i>Eucalyptus globulus</i>; +it thrives well in malarious places, and at once +produces a marked hygienic change in the air. A +Monsieur Gimbert relates that “A farm some +twenty miles from Algiers was noted for its pestilential +air, and in the spring of 1867, 13,000 +eucalyptus trees were planted there, since which +time not a single case of fever has occurred.</p> + +<p>“The gum-tree grows rapidly and absorbs as +much as ten times its weight of water from the +soil, and emits camphoraceous antiseptic vapour +from its leaves. It is therefore often called the +fever-destroying tree.”</p> + +<p>Experiments have been made to try and find +out how much moisture is really given out by +leaves. It was found that a sunflower three and a +half feet high, with a leaf expanse of over five +thousand inches, exhaled one pint of liquid in the +course of the day.</p> + +<p>No wonder, therefore, that trees tend to make +the air damp.</p> + +<p><span class="pagenum" id="Page_109">[Pg 109]</span></p> + +<p>Each stomate leads into air spaces between the +cells, and is thus connected with the interior of the +leaf.</p> + +<figure class="figcenter illowp78" id="i_109" style="max-width: 53.875em;"> + <img class="w100" src="images/i_109.jpg" alt="Microscopic image showing several +layers of tissue. The top layer consists of square cells that are tightly +connected. The second layer consist of long, elongated rectangular cells, grouped +with their long sides next to each other and their short base to the top layer. +These cells contain dark particles. The third layer consists of shorter +rectangular cells, also with their longer sides connected to each other and with +their shorter base to the layer above. These cells too contain particles, that +are somewhat larger that in the layer above and have a lighter colour. +The following layer consists of irregularly shaped cells that from a spongy +tissue with large gaps between the cells. These cells contain 2 types of particles: +one type similar to the ones in the layer above, and a second type that is very +dark. Finally the bottom layer consists, like the top layer, of square, +interconnected cells, a bit smaller though. At one point an opening in this layer +can be seen. This opening is bordered by 2 very small cells."> + <figcaption> + <p>LEAF SECTION.</p> + </figcaption> +</figure> + +<p>The tissue and cells of a leaf (bifacial)⁠<a id="FNanchor_9_9" href="#Footnote_9_9" class="fnanchor">[9]</a> can be +understood by reference to the accompanying diagram. +Between the upper and under surfaces of +a leaf there is a layer, more or less thick, of soft +green tissue known as <i>mesophyll</i>, and if we hold a +leaf to the sunlight we shall see the veins traversing +this tissue.</p> + +<p>The upper part of the mesophyll consists of +elongated cells arranged at right angles to the +<span class="pagenum" id="Page_110">[Pg 110]</span>surface, and placed so evenly parallel to each other +that they have been compared to the pales of a +fence, and are called palisade tissue. These cells +contain a quantity of the green substance called +<i>chlorophyllon</i> (from <i>chloros</i>, green, and <i>phyllon</i>, a +leaf), so named because to this bright green substance +we owe all the lovely verdure of our woods +and gardens.</p> + +<p>Below this palisade tissue is another of quite a +different form, consisting of large spongy cells, and +therefore known as spongy tissue.</p> + +<p>In its intercellular spaces are stored those +secretions which make certain herbs, such as +thyme, marjoram, and others so fragrant when +bruised.</p> + +<p>The chemical changes which are ever going on +in these various layers, require a constant supply +of the outer air, and this is secured by the little +openings, called stomata, on the under surface of +the leaf, which have been already described; these +constitute the breathing apparatus of the leaf, for +they open and shut, and regulate the supply of +air into little air chambers, from which it passes +into the structure of the plant.</p> + +<p><span class="pagenum" id="Page_111">[Pg 111]</span></p> + +<p>Before going any further I must try and explain +a little about the wonderful substance called protoplasm.⁠<a id="FNanchor_10_10" href="#Footnote_10_10" class="fnanchor">[10]</a></p> + +<p>If we have ever watched a potter at work, we +know he takes a lump of clay and moulds it +according to his purpose, into a rough pot, or a +lovely vase; now protoplasm seems to be just +such a foundation material from which the Divine +Creator causes animal and vegetable forms to proceed. +<i>First material</i> seems to me to be a term +that actually expresses the meaning of the word +protoplasm.</p> + +<p>It lines the cell walls of leaves, it is capable of +forming fresh cells, it can absorb moisture and +other matters, it contracts and expands, it has +power of movement, as one may readily see when +a portion of a leaf is placed in a microscope, so as +to show the grains of bright green chlorophyll +circulating in the lining of each little cell.</p> + +<p>Learned volumes would be needed to explain +the nature of protoplasm, so I must be content +with these simple facts about its nature, and proceed +to the chemical action going on in leaves.</p> + +<p><span class="pagenum" id="Page_112">[Pg 112]</span></p> + +<p>In ordinary atmosphere there is a very small +quantity of a gas called carbon-dioxide.⁠<a id="FNanchor_11_11" href="#Footnote_11_11" class="fnanchor">[11]</a> The +leaves absorb this gas from the air, and because +there is so little of it, each tree needs to spread out +an immense amount of foliage, that it may drink +in, by its means, all the carbon-dioxide that can +possibly be obtained.</p> + +<p>When this gas comes in contact with the +chlorophyll in a leaf, one part of the oxygen is set +free, and returns to the air in a pure condition, +thus making it more healthy for us to breathe; +then the carbon and the remaining oxygen combine +with water in the leaf cells, and form starch, +the leaves retain the carbon, to build up their own +structure; it enters indeed so largely into the +composition of vegetable substance, that in some +cases if we could burn one hundred parts of it, fifty +parts of the ashes would prove to be carbon or +charcoal.</p> + +<p>In a rough sort of way we may see for ourselves +how much carbon there is in woody fibre, by lighting +an ordinary wooden match and letting it burn +<span class="pagenum" id="Page_113">[Pg 113]</span>itself out; the black portion that remains will be a +piece of charcoal not very much smaller than the +original match.</p> + +<p>Of course, in the process of burning, the match +has lost the resin, and other organic substances +which were stored up in the cells of the wood; all +these have passed into the air, and only the carbon +remains. If, however, instead of this slow manner +of combustion, we had set light to a whole box +of matches so that it burnt fiercely, the flame +would have been strong enough to consume the +charcoal, and nothing would then be left but +mineral ashes.</p> + +<p>When charcoal burners are at work in a forest, +we may see them making a stack of wood, which +they cover with a thick layer of clay so that the +wood may burn away very slowly; in this case +the charcoal will be left in the same way as when +we burnt the single match.</p> + +<p>As long as the upper side of leaves are soaking +in the sunlight, starch is being formed, as I have +described; but during the night the starch thus +formed is dissolved, and passing through the leaf +fibres finds its way into every part of the plant, +<span class="pagenum" id="Page_114">[Pg 114]</span>either to be used in forming new tissue, or else +to be stored up for future use.</p> + +<p>The net-work of veins act as a service of tiny +pipes, to convey the liquids up and down the +petiole (leaf-stalk).</p> + +<p>Generally the water given off from the stomata +is in the form of vapour; but in some plants +drops of water exude from the apex or point of the +leaf through the water pores. In <i>Saxifraga crustata</i>, +there are pores round the edges of the +leaves, through which water, highly charged with +lime and other salts, passes out, and as it evaporates +a white deposit of lime remains which is quite +visible in the form of a frosted edging to the +leaves.</p> + +<p>There is an American plant called the jewel-weed, +which shows to perfection this power of distilling +drops of water. I will quote a short description +of its appearance at night-fall.</p> + +<p>“Upon the approach of twilight, each leaf droops +as if wilted, and from the notches along the edge, +the crystal beads begin to grow until its border is +hung full with its gems. It is Aladdin’s lantern +that you see among a bed of these succulent pale +<span class="pagenum" id="Page_115">[Pg 115]</span>green plants, for the spectacle is like dreamland.”⁠<a id="FNanchor_12_12" href="#Footnote_12_12" class="fnanchor">[12]</a></p> + +<p>A very similar effect may be observed if we +visit a plant of Lady’s-mantle (<i>Alchemilla vulgaris</i>) +at early morning after a warm dewless night; each +leaf will be found beautifully decked with dewdrops +at equal distances round the edge of the +leaves where the pores have exuded the moisture +with which they are charged.</p> + +<p>Nasturtium and fuchsia may also be examined +for this purpose and will show exudation from +their leaf pores.</p> + +<p>If a small quantity of wheat is grown in some +cocoa fibre, it will illustrate this power of giving +off water, for when the little blades are a few +inches high, they will be found each morning +tipped with a large dewdrop, the result of exudation +during the night.</p> + +<p>In countries where the sun is intensely hot, if +the leaves of trees were to be exposed to its full +power, they would probably wither, and vegetation +would perish.</p> + +<figure class="figcenter illowp69" id="i_116" style="max-width: 62.5em;"> + <img class="w100" src="images/i_116.jpg" alt="A stem with 8 leaves, grouped +in pairs. The leaves grow smaller toward the tip. They are of a rounded shape +with smooth edges. Veins are clearly visible. At the tip is a bud. The leaves +directly underneath seam to grow small stalks with buds from there axils."> + <figcaption> + <p>YOUNG SHOOT OF EUCALYPTUS.</p> + </figcaption> +</figure> + +<p>Against this danger some trees are enabled to +<span class="pagenum" id="Page_116">[Pg 116]</span>make special provision, by changing the form of +their leaves, and their mode of hanging on the +branch. In Australia, for instance, where the sun +is almost vertical, the acacias and eucalyptus trees, +instead of holding their leaves flat or horizontally +as trees do in England, so that they may catch +every ray of sunlight, avoid the heat as much as +possible, by holding them edgeways to the light.</p> + +<p><span class="pagenum" id="Page_117">[Pg 117]</span></p> + +<figure class="figcenter illowp100" id="i_117" style="max-width: 62.5em;"> + <img class="w100" src="images/i_117.jpg" alt="Photograph of a branch with +leaves and flowers. The leaves are oval, elongated, with sharp tips. The flowers, +some still in buds, some flowering, some finished already, have a round shape with +a multitude of hairlike parts."> + <figcaption> + <p>MATURE FORM OF EUCALYPTUS LEAVES.</p> + </figcaption> +</figure> + +<p><span class="pagenum" id="Page_118">[Pg 118]</span></p> + +<p>While eucalyptus trees are young, and partially +shaded by surrounding vegetation, their leaves are +flat and oval, and English seedlings of this tree +usually retain such leaves from five to ten years, +our climate not being hot enough to require the +mature form of leaf which hangs vertically, and is +of an entirely different form.</p> + +<p>Reference to the plates will show a young shoot +of <i>Eucalyptus globulus</i> and a branch of the older +leaves, with their edges only exposed to sunlight.</p> + +<p>The curved sickle-shaped leaves of the eucalyptus +afford very little shade to the traveller in +Australia for this reason, that only fine intercrossing +lines of shadow are seen on the ground. +To make this clear, let my readers take a sheet +of notepaper out of doors on a sunny day and +hold it perfectly flat, so as to expose it to all the +sunlight it can receive upon its surface, as if it +were a growing beech-leaf, and it will throw a +large shadow on the ground. Then hold it edgeways +to the sun, and it will form the kind of thin +line of shadow that would be cast by a mature +eucalyptus leaf.</p> + +<p>Preparation for the fall of the leaf begins in +<span class="pagenum" id="Page_119">[Pg 119]</span>spring, when a fine line or ridge may be traced +just below the junction of the leaf with the stem. +This dark line is in reality a thin layer of cork, +which, during the summer months, continues to +grow inwards to form in due time a covering +for the bare place on the stem that will be left +when the leaf falls off; this is called the leaf-scar.</p> + +<p>It is interesting to watch this line, growing more +and more visible as the year goes on.</p> + +<p>Another curious fact is, that some of the starch +which the leaves have been making during the +summer, becomes stored up in autumn at the base +of the leaf-stalk, so as to afford nourishment to the +bud which will arise out of the axil of the leaf. +When a weak solution of iodine is applied to it, +this starch turns blue, and in this way its presence +can be ascertained.</p> + +<p>The fall of the leaf appears to take place mainly +because the starch has the effect of softening the +cells of the leaf-stalk; as it dries up it loses its +hold of the twig, and either the wind or a slight +frost will suffice to bring the leaves down to the +ground in showers.</p> + +<p><span class="pagenum" id="Page_120">[Pg 120]</span></p> + +<p>Another reason for their fall is, that their year’s +work is done. Like good servants, they have been +hard at work all through the summer and autumn +months, taking in stores of nourishment for the +benefit of the tree, and giving out volumes of +oxygen, so helpful for the maintenance of human +life. They have secured and laid up sufficient +nutriment for the development of the next year’s +buds, and having done this, their special office +being at an end, they fall beneath the tree to +become leaf-mould, which, in its turn when fully +decayed, will yield nourishing elements to be +carried by the winter and spring rains to the tree +roots.</p> + +<p>I might add many more useful objects which +we owe to trees, and I commend it to my young +readers as an instructive study to try and make +out a complete list of the useful products of our +English trees. I imagine we do not yet know +all that might be obtained from them, new discoveries +continue to reveal their value in the way +of medicines; for instance, the crystals of the +willow (called salicine) are now frequently prescribed +as a remedy for rheumatism. Euonomine +<span class="pagenum" id="Page_121">[Pg 121]</span>and many others might be included amongst the +valuable gifts which nature has stored in the cells +of tree-stems.</p> + +<p>Specimens to be obtained:—Leaves with straight +veins, such as grass or corn, hyacinth, narcissus, or +any bulbous plants; leaves with netted veins, such +as oak, ivy, vine, &c.; monocotyledon seedlings; +dicotyledon seedlings. Leaf-skin to be examined +through a microscope, in order to see stomata, +chlorophyll, network, and cells. Examine waterpores +in leaves when exuding moisture. Observe +shadows thrown by leaves held flat and edgeways +to the sun. Compare young and old eucalyptus +leaves. Observe line of cork below leaf-scars.</p> + + +<p class="subtext">Leaves can be made into beautiful skeletons by soaking a good +many together in a pan of soft water until the upper and under +surfaces of the leaves are sufficiently decayed to be easily removed +by a soft brush; the fibre which is left can then be bleached with +chloride of lime. When mounted with fine wire these skeleton +leaves form an interesting group to place under a glass shade.</p> + + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum"><a id="Page_122"></a><a id="Page_123"></a>[Pg 123]</span></p> + + + <h2 class="nobreak" id="CHAPTER_V"> + CHAPTER V + <br> + <i>BUDS</i> + + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_124">[Pg 124]</span></p> +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“Oh! who can speak the joys of spring’s young morn,</div> + <div class="verse indent2">When wood and pasture open on his view,</div> + <div class="verse indent1">When tender green buds blush upon the thorn</div> + <div class="verse indent2">And the first primrose dips its leaves in dew.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">Clare.</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_125">[Pg 125]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_V_1"> + CHAPTER V + <br> + <br> + <span style="font-size: 70%;">BUDS</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_125.jpg" + style="width: 12%; height: auto;" + alt="H"> +</div> +<p class="drop-cap"><span class="upper-case">How</span> +we watch for the buds as tokens +of the coming spring! We delight +to see them daily growing larger, and +opening out their leafy treasures to the +sun.</p> + +<p>The re-clothing of the trees has always an +element of wonder about it; the change is truly +a resurrection; a few days of warm sunshine +and gentle rain, and then the dry, dead-looking +branches begin to bud and blossom as if by a +miracle.</p> + +<p>We may, however, trace the secret of this +sudden change, if we look back to the processes +Nature was carrying on during the previous +summer, and we may learn from her many a +useful lesson of foresight and preparation.</p> + +<p><span class="pagenum" id="Page_126">[Pg 126]</span></p> + +<p>If during the summer we lift up the branch +of any deciduous tree and search amongst the +leaves, we shall find that the buds for the following +year are already there, waiting to be developed +in due time. When the leaves turn dry and sere, +they fall off and leave the buds to be hardened +and matured by the rain, snow, and frost of +winter.</p> + +<p>Certain species of Turkey oak, young beeches, +hornbeams in hedges, and other trees, appear to +retain their leaves, to some extent, as if to protect +the buds until the rising sap in spring loosens +their hold upon the branches, and makes them +fall off.</p> + +<p>The plane-tree appears to be an exception to +most trees in the curious protection it affords its +young buds. If we search for these in the summer +or autumn, they are not to be found, for the leaf-stalk +is so swollen and hollow at the base as to +enclose the bud within it; even when the leaf falls +off, the bud is covered by a tough outer case +coated with resin, and within are many fur-lined +scales. When these are removed we see the tiny +leaves wrapped in silky coverings, and when the +<span class="pagenum" id="Page_127">[Pg 127]</span>warmth of spring enables them to expand, their +under surfaces have such a thick coat of down +that the plane is known in some countries as the +cotton-tree. From its fruit being in the form of +hanging circular balls, its name in America is the +button-wood. The need of this special protection +against cold is shown by the fact that if severe +frost returns after the leaves have expanded, they +<span class="pagenum" id="Page_128">[Pg 128]</span>frequently shrivel and perish. Some Japanese +maples have the same arrangement of hollow leaf-stalks +to contain the buds.</p> + +<figure class="figcenter illowp100" id="i_127" style="max-width: 62.5em;"> + <img class="w100" src="images/i_127.jpg" alt="A large, leafless tree +on the top of a hill."> + <figcaption> + <p>OAK IN WINTER.</p> + </figcaption> +</figure> + +<p>When buds are situated at the end of a branch +they are called terminal, and their office is to +increase the length of the branch.</p> + +<p>When they grow in the axil of a leaf (that is, +where the leaf-stalk joins the stem) then they are +called axillary, and as they grow out and form +fresh stems and leaves, the branch broadens on +either side.</p> + +<figure class="figcenter illowp72" id="i_129" style="max-width: 59.0em;"> + <img class="w100" src="images/i_129.jpg" alt="A large tree bearing leaves +together with other ones in a fenced-off meadow."> + <figcaption> + <p>OAK IN SUMMER.</p> + </figcaption> +</figure> + +<p>Seeing that the branches of a tree thus spring +from the buds, it follows that the position and +development of the buds upon the stem, as we +tried to show in the last chapter, have much to do +with the ultimate shape of the tree. The development +of the axil buds, as well as of the terminal +bud, gives rise to a branched tree like the oak; +these buds, however, are often erratic, and in some +trees the terminal bud of the shoot is often suppressed +and the axil buds grow with extra vigour, +whilst in other instances the terminal bud grows +strongly and the axil buds either grow feebly or +are altogether suppressed. In the bamboos, palms, +<span class="pagenum" id="Page_129">[Pg 129]</span>and sugar-cane we get good examples of this +terminal bud-growth, the axillary buds being +suppressed; the suckers that grow from the axils +of the lower leaves of the palm are often evidence +of the presence of axillary buds, although they +are, as a rule, dormant. We are all more or less +familiar with the character of ordinary forest trees, +the rounded outline of the oak, the slender sprays +of the birch, the spreading branches of the beech, +<span class="pagenum" id="Page_130">[Pg 130]</span>but perhaps we may not have remarked how much +these variations of form are due to the position of +the buds upon the branches. We will suppose +that on a winter’s day we are looking at the +tracery of some elm-branches against the sky; +the form of each branch shows that the terminal +bud in this tree usually ceases to grow, and allows +the lateral shoots to increase in length, and take +its place; this produces short, twiggy branches, +and a stem which makes a tall tree rather than +a wide-spreading one. The horse-chestnut, again, +produces its flowers in the terminal buds; this +arrests their growth, and side shoots have to grow +on instead, thus usually giving height rather than +breadth to the tree. We may note the differing +outline of the willow, birch, and many others +where the terminal buds do not cease to grow, +but each year continue to add to the breadth as +well as the height of the tree.</p> + +<p>In pine-trees the buds are produced at the ends +of the branches, and several shoots proceed from +one bud.</p> + +<p>The spiral arrangement of leaves is well seen +in a young coniferous shoot, also in the flower-bud, +<span class="pagenum" id="Page_131">[Pg 131]</span>and especially in the fir-cone itself, in which +an ever-varying double spiral can be traced.</p> + +<p>Loudon remarks, “The perfection of a fir consists +in height rather than in lateral expansion; +buds are produced very sparingly and nearly +always at the extremities of the shoots. Provision +is thus made for the upward growth of +the tree more than for side expansion.”</p> + +<p>When we speak of a coniferous shrub having +lost its leader, we mean that the terminal bud on +the topmost shoot having been broken off, one +or more of the lower branches must rise up and +take its place, and thus lateral buds in time +become terminal and grow upright instead of +sideways.</p> + +<p>A silver fir, that I have been observing +for years past, bears such a crop of heavy +cones on its slender upper branches that the +leader is almost invariably broken off by the +weight, and the lateral shoots have to take its +place, to the great detriment of the central stem, +which is twisted and curved out of shape by the +efforts the tree makes to repair its terminal +shoot.</p> + +<p><span class="pagenum" id="Page_132">[Pg 132]</span></p> + +<p>In other trees, again, the unfolding of all the +buds is nearly simultaneous, but in the fir tribe +the bud which terminates the summit of the tree +and is destined to form its leading shoot and +increase its height is developed last; this delay +seems a provision of nature for the safety of the +most important shoot which the tree can produce, +ensuring its height rather than its breadth, and +the production of timber by the preservation of +its permanent trunk rather than by its temporary +branches.</p> + +<p>If a willow is deprived of the upper part of +its stem and so made a pollard tree, it develops +a bushy head of small stems which spring from +the other buds thrown out to repair the loss of +the central stem. This pollarding is often resorted +to in order to obtain wood of the right +kind for basket-making, and young ash trees are +thus treated, so that slender rods suitable for +hop-poles and tool-handles may spring from the +lopped stem.</p> + +<p>When buds are found growing on any other +part of a plant except those just mentioned, +they are called adventitious buds. These may +<span class="pagenum" id="Page_133">[Pg 133]</span>be found growing on the edges of the leaves +of the marsh tway-blade; they also spring out of +the flat surface of the fronds of the viviparous +fern.</p> + +<p>Under favourable conditions every part of a +plant will produce buds, and, taking advantage +of this fact, florists increase their stock of succulent +plants by putting the leaves on a wet surface, +which induces them quickly to send out buds and +roots. Such plants as begonias, gloxinias, hoyas, +and sedums are readily increased by this mode +of propagation. Underground stems will often +send out buds, and they produce the underwood +from the stumps of fallen trees.</p> + +<p>We are all familiar with the suckers of trees +which spring up in our lawns and gravel paths +often many yards away from the parent tree; +these all arise from active buds on underground +stems. Gardeners are always careful to prune +away such growths at the base of their wall-fruit-trees, +since they rob their valuable peaches, +nectarines, and apricots of strength and nourishment. +These well-named “suckers” spring from +the common stock upon which the choice fruit-trees +<span class="pagenum" id="Page_134">[Pg 134]</span>were grafted, as one may see by gathering +a leaf from a sucker and comparing it with a +peach or nectarine leaf.</p> + +<p>On the oak, chestnut, lime, beech, and other +trees there are sometimes to be found dormant +buds in the form of rounded knobs covered +with bark and increasing in size with the growth +of the tree; these, in the event of other buds +perishing, will start into active growth and do +their part in preserving the life of the tree.</p> + +<p>Such woody balls when found on the oak are +worth examination, as when divested of their +bark they show exquisite structure of woody fibre.</p> + +<p>The small bulbils we find in the axils of lily +stems, on the cuckoo-flower, on <i>Dentaria bulbifera</i>, +and on some species of Allium, are all adventitious +buds, which drop off in due time and become +young plants.</p> + +<p>They are in many respects similar to bulbs, +and if we cut one in half and compare it with +a divided hyacinth we shall see that they both +consist of over-lapping scales. In the onion +these scales are fleshy and succulent, but in +most tree buds they are dry, hard membranes.</p> + +<p><span class="pagenum" id="Page_135">[Pg 135]</span></p> + +<p>The pear and magnolia buds are secured +against wintry cold by woolly linings to the +scales, and in the horse-chestnut they are covered +with a kind of resin which renders them impervious +to moisture.</p> + +<figure class="figcenter illowp78" id="i_135" style="max-width: 62.5em;"> + <img class="w100" src="images/i_135.jpg" alt="A large bare tree in a meadow +with other bare trees in the background. In the foreground there are some +lighter patches at the soil."> + <figcaption> + <p>HORSE-CHESTNUT.</p> + </figcaption> +</figure> + +<p><span class="pagenum" id="Page_136">[Pg 136]</span></p> + +<p>It requires a careful use of the microscope to +trace all that a bud contains; I will therefore +quote the words of a German naturalist +who dissected a horse-chestnut bud gathered in +winter, and found that it contained sixty flowers. +It would be interesting to select a terminal +flower-bud of this tree; by taking it carefully to +pieces one might, with patience and using a +powerful lens (or a microscope if one is available), +see for ourselves a good deal of what the +writer describes:—</p> + +<p>“Having removed the outer scales, seventeen +in number, cemented together by a gummy substance +to render the bud waterproof<a id="tn-004"></a>, I discovered +four leaves surrounding a spike of flowers, so +clearly visible when magnified that I not only +counted the number of flowers, but could discern +the pollen on the stamens.”</p> + +<p>The winter covering of a bud, both the inner +and outer scales, are only a temporary protection +in order to keep out moisture and keep in warmth, +so that as the sun begins to gain power, the +gummy covering of the bud melts and yields +to the expanding pressure from within, when one +<span class="pagenum" id="Page_137">[Pg 137]</span>after another the sticky scales fall off, showing the +young leaves with their soft woolly surfaces; +these leaves rapidly unfold and hang droopingly +until the midribs gain strength enough to hold +them upright.</p> + +<p>Evelyn remarked that, “As soon as the leading +shoot of the horse-chestnut has come out of the +bud, it continues to grow so fast as to be able to +form its whole summer’s shoot, sometimes eighteen +inches long, in about three weeks. After this it +grows but little more in length, only thickens, +becomes strong and woody, and forms the buds +for next year’s shoot.”</p> + +<p>Buds have always been to me a most interesting +subject of study; there is much variety of +character in them, and to those who observe them +closely they reveal in the autumn and winter +what the tree is purposing to do in the following +season.</p> + +<p>A beech-tree on my lawn bears its nuts only +every second or third summer, and in the previous +autumn I can always tell whether the squirrels are +likely to be well off for food in the coming year, +by observing the size and shape of the buds. +<span class="pagenum" id="Page_138">[Pg 138]</span>Those which contain the flowers are round and +bulky, whilst the leaf-buds are long and slender.</p> + +<figure class="figcenter illowp56" id="i_138" style="max-width: 46.3125em;"> + <img class="w100" src="images/i_138.jpg" alt="A bare tree, standing amidst +some shrubs. In the background some fir trees."> + <figcaption> + <p>YOUNG BEECH.</p> + </figcaption> +</figure> + +<p>Embryo flowers are disposed in the buds in +different ways. The woodsorrel<a id="tn-021"></a> is rolled into a +spiral, rose-petals are placed one within the other, +the pink is folded in five divisions, and others are +pleated and fluted into the smallest possible space. +Perhaps of all others the bud of the great Oriental +poppy is the best example of exquisite packing. +Early on a summer’s morning you may see its +<span class="pagenum" id="Page_139">[Pg 139]</span>huge green hairy bud at the end of a stem several +feet in length, and whilst you are looking the +sepals or calyx leaves suddenly divide and fall +off, the mass of vivid scarlet petals crumpled into +a thousand folds begins to open out, and before +long the glorious flower, which is often as much as +seven inches across, holds itself erect in majestic +beauty.</p> + +<p>Those who possess a tulip-tree will find its +opening buds reward examination. The leaves +are folded in half and bent double, a pair of +leaf-scales enclosing each of the true leaves. One +may unpack the entire bud until we come to +leaves almost too minute to be discerned.</p> + +<p>Young sycamore-trees often have buds of large +size and brilliant crimson colour; the foldings of +their leaves are very intricate, and form an interesting +contrast to those of the tulip-tree. Hart’s-tongue +fern, arum, and pear leaves afford three very remarkable +modes of folding in the bud.</p> +<div class="col50"> +<figure class="figcenter illowp45" id="i_140a" style="max-width: 28.75em;"> + <img class="w100" src="images/i_140a.jpg" alt="A rolled-up leaf, with a +pointed tip, yielding a cigar-like shape."> + <figcaption> + <p>UNFOLDING ARUM + LEAF.</p> + </figcaption> +</figure> + </div> +<div class="col50"> +<figure> + <figure class="figcenter illowp57" id="i_140b" style="max-width: 29.625em;"> + <img class="w100" src="images/i_140b.jpg" alt="A leaf that is rolled-up from +both sides, yielding two small cylinders alongside the stalk."> + <figcaption> + <p>PEAR LEAF.</p> + </figcaption> + </figure> + <figure class="figcenter illowp46" id="i_140c" style="max-width: 32.6875em;"> + <img class="w100" src="images/i_140c.jpg" alt="Two leaves rolled-up lengthwise, +each yielding a short cylinder, perpendicular to the stalk."> + <figcaption> + <p>UNFOLDING LEAVES + OF HART’S-TONGUE + FERN.</p> + </figcaption> + </figure> +</figure> +</div> + +<p>Another point of character in buds is of considerable +importance to the horticulturist, namely, the +fact that in some cases the value of the flowers +produced varies with the position of the buds. For +<span class="pagenum" id="Page_140">[Pg 140]</span>instance, the blossoms produced from the crownbud⁠<a id="FNanchor_13_13" href="#Footnote_13_13" class="fnanchor">[13]</a> +of certain chrysanthemums are poor and pale +in colour compared with those grown on the side +<span class="pagenum" id="Page_141">[Pg 141]</span>shoots; the latter are therefore retained and +fostered, so that from them flowers of the finest +description may be obtained.</p> + +<p>In cultivating fruit-trees it is found needful not +only to prune away redundant branches which +bear leaves only, but also where strong woody +roots are promoting the growth of leaf-buds, they +also have to be pruned, so that the check thus +given to the growth of the tree may result in the +formation of fibrous roots, which will tend to the +production of flower-buds and a resulting crop of +fruit.</p> + +<p>I have often observed that the transplantation +of trees leads to their throwing out flowers in the +succeeding year. This was notably the case with +an avenue of deodars which had overgrown my +carriage drive; they received a considerable check +in being transplanted, but in the following year +their branches were covered with male catkins and +some few cones succeeded.</p> + +<p>For this reason the removal of fruit-trees is not +unfrequently resorted to, as a means of inducing +fruit-bearing.</p> + +<p>So much vigour is stored up in the bud, that it +<span class="pagenum" id="Page_142">[Pg 142]</span>will bear being removed from one tree and inserted +in the stem of another, within which it will grow +and become a part of the living tree. This is one +of the means by which we have obtained such an +infinite variety of roses; the buds from choice +species being readily made to grow upon strong +briar stocks, and thus one may also see roses of +several different colours blossoming on the same +stem. Choice varieties of fruit-trees are cultivated +in the same way by means of buds inserted in the +bark.</p> + +<p>Having observed how flowers are arranged in +the bud, we may go on to dissect incipient leaves +and learn how they are placed.⁠<a id="FNanchor_14_14" href="#Footnote_14_14" class="fnanchor">[14]</a> We shall find +that the frond of the hart’s-tongue fern is rolled up +from the tip, the arum gracefully curved lengthwise. +Pear leaves are rolled from side to side +towards the middle, and so is the primrose, but the +reverse way. Beautiful examples of curled leaves +may also be seen in the water-lily and banana.</p> + +<p>In grasses the first leaves are equivalent to budscales, +and protect those which continue to grow +from the centre, each one sheathing out of the +<span class="pagenum" id="Page_143">[Pg 143]</span>previous leaf after the manner of monocotyledons.⁠<a id="FNanchor_15_15" href="#Footnote_15_15" class="fnanchor">[15]</a></p> + +<p>The colouring of buds is one of the lovely +features of spring. Seen against the blue of the +sky, the coral red of the lime, sycamore, and +Japanese maple buds, cannot be passed by without +notice. The whitebeam has a beauty of its own +for its buds are large and white with downy coverings, +giving promise of the future leaves which are +so light-coloured underneath, that the effect when +they are blown aside by the wind is curious and +beautiful. The Germans call it <i>mehl-baum</i> or meal-tree, +from its whitish downy leaves.</p> + +<p>The variegated vine, sometimes seen in greenhouses, +has exquisite buds of pinkish crimson, with +bright yellow stipules. By way of contrast, I once +placed some sprays of it in a glass with twigs of +purple hazel which are of a deep claret brown; +they were not only opposite in colour, but curiously +different in habit, the vine holding its bud erect, +and the hazel as persistently drooping. These +variations lead me again to remark that, to a close +observer, buds will be found to differ much in +<span class="pagenum" id="Page_144">[Pg 144]</span>character and to be well worthy of close attention.</p> + +<p>I will mention some trees whose buds are specially +remarkable for beauty of form whilst unfolding. +The mountain ash has very graceful leaves when +just emerging from the bud; they show on their +upper and under surfaces two distinct shades of +green.</p> + +<p>The unfolding weigelia buds are extremely +pretty in shape, the leaves being pointed and +delicately curled.</p> + +<p>I need hardly mention the beech; nothing can +be more exquisite than a spray of its opening +buds with their silky fringed young leaves and +crimson leaf-scales. I look forward every spring +to the joy of watching the unfolding of these +caskets.</p> + +<p>A warm shower or two and some sunny days +cause them to expand with a rapidity which seems +magical, and one almost regrets to find the beauty +of the buds in their early stages so quickly passing +away. The ash attracts notice by its jet-black +buds, and the wayfaring tree by the delicate venation +of its young leaves.</p> + +<p><span class="pagenum" id="Page_145">[Pg 145]</span></p> + +<p>I cannot refrain from mentioning another beautiful +effect arising from young buds in the case of a +<i>Picea nobilis glauca</i>, which long name simply means +a sea-green silver-fir, standing on our lawn. In +the summer its terminal buds are a very pale sea-green, +and as they grow and +are seen against the dark green +of the rest of the foliage the +effect is very curious, as though +each branch had become tipped +with frosted silver.</p> + +<figure class="figright w30" id="i_145" style="max-width: 31.4375em;"> + <img class="w100" src="images/i_145.jpg" alt="Three stems each with several +small leaves. The leaves are more or less folds and have very pronounced +veins."> + <figcaption> + <p>BUDS OF WAYFARING + TREE.</p> + </figcaption> +</figure> + +<p>The soft silky buds of the +willow, and especially those of +the low growing sallow which +are gathered as “palm” for +church decoration, are amongst +the welcome signs of early +spring. The sallow has its +male blossoms on one tree, +but not far away we shall +find the female tree bearing +the flowers which will eventually produce the seeds. +We may therefore seek for three kinds of buds, +those which produce the flowers on each tree, and +<span class="pagenum" id="Page_146">[Pg 146]</span>the others which will clothe the tree with leaves +when the blossoms are over.</p> + +<p>This chapter shows us how much there is to +instruct the student of nature during the winter as +well as the summer months.</p> + +<p>I have but indicated a very few out of the many +lines of study which may be taken up; one could +write essay after essay upon the growth of a single +hedgerow, but all I can hope to do in simple +chapters of this kind is to throw out hints and +indications, and trust that my young readers may +find their interest sufficiently excited by what they +have read, to lead them on to fuller, deeper study of +each point touched upon.</p> + +<p>Nature is an inexhaustible storehouse of +wonders, and the further we explore, the more +our eyes are opened to see the vistas that lie +before us, branching out in various directions.</p> + +<p>This special path of botanical study is one that, +more or less, can be pursued at intervals, as opportunity +may offer through life, and as it adds much +pleasure to leisure hours, I specially commend it +to my young readers.</p> + +<p>Specimens to be obtained and compared with +<span class="pagenum" id="Page_147">[Pg 147]</span>the descriptions in this chapter:—Search for buds +in summer; plane-tree buds; Japanese maples; +terminal and axillary buds; observe shape and +outline of trees; buds of coniferous trees; fir cone; +fir-tree that has lost its leading shoot; pollard +willow, and other trees, buds on marsh tway-blade, +and viviparous fern; buds on underground stems; +suckers from wall-fruit trees; dormant buds or +knobs on tree-stems; bulbils on lily, dentaria +and allium; horse-chestnut terminal bud; observe +leading shoot of horse-chestnut in early summer; +flower and leaf buds on beech; various flower-buds; +Oriental poppy; tulip-tree buds; various +leaf-buds unfolding; colouring of Japanese maple, +lime, and sycamore buds; whitebeam; variegated +vine; purple hazel; mountain ash; spray of beech +buds; ash buds; <i>Picea nobilis glauca</i>; willow and +sallow, male and female flower buds and leaf +buds; bamboo.</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum"><a id="Page_148"></a><a id="Page_149"></a>[Pg 149]</span></p> + + + <h2 class="nobreak" id="CHAPTER_VI"> + CHAPTER VI + <br> + <i>FLOWERS</i> + + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_150">[Pg 150]</span></p> +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“Your voiceless lips, O flowers, are living preachers;</div> + <div class="verse indent1">Each cup a pulpit, every leaf a book</div> + <div class="verse indent1">Supplying to the fancy numerous teachers</div> + <div class="verse indent14">From loneliest nook.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">Horace Smith.</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_151">[Pg 151]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_VI_1"> + CHAPTER VI + <br> + <br> + <span style="font-size: 70%;">FLOWERS</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_151.jpg" + style="width: 12%; height: auto;" + alt="I"> +</div> +<p class="drop-cap"><span class="upper-case">Instead</span> +of looking at flowers as bright +and beautiful objects made to be a +source of continual delight in our +daily lives, though such they truly are, +we will rather now, for purposes of study, +consider them as the means by which the plant +carries out the purpose of its creation, namely, to +perfect its seed and thus perpetuate its species.</p> + +<p>In the life-history of shrubs, trees, and plants we +find this is their one aim, and that everything else +is subservient to it.</p> + +<p>The stamens and pistil being of essential importance +in forming the seed, we find them placed for +safety in the centre of the flower; folded round +them are the petals or coloured parts of the flower, +<span class="pagenum" id="Page_152">[Pg 152]</span>and outside these again are the green sepals, or +leaves of the calyx.</p> + +<p>These two sets of enfolding leaves are called +“floral envelopes,” because they fold over and +protect the central organs, the stamens and the +pistil.</p> + +<p>We will select a buttercup as a type, and taking +it to pieces we will try to learn the names and +uses of its various parts.</p> + +<p>The outside is a greenish-yellow cup which is +called the calyx.</p> + +<p>The divisions of this little green cup are called +sepals, and their office is to protect the five bright +yellow leaves within, which are called petals when +we speak of them singly, but, taken all together, +form the corolla.</p> + +<p>In the buttercups the petals are all separate, but +if we look at a primrose we shall see that the +corolla is in one piece, united in a tube; so also is +the calyx.</p> + +<p>The botanical term for a corolla thus formed is +<i>gamopetalous</i>, a long word but easily understood +when we know that <i>gamos</i> means united; a flower +with petals in one instead of many divisions is +<span class="pagenum" id="Page_153">[Pg 153]</span>more easily referred to by this word than if we +had each time to express it by a sentence.</p> + +<figure class="figcenter illowp50" id="i_153" style="max-width: 40.875em;"> + <img class="w100" src="images/i_153.jpg" alt="A flower, viewed from the top. +It has 5 petals. In the centre is a pentagon formed by small stripes. In the +centre of the pentagon is a circular shape. The flower as a whole has a +circular shape."> + <figcaption> + <p>PRIMROSE.</p> + </figcaption> +</figure> + +<p>Gathering a newly-opened flower, we can see at +a glance that the sepals are placed quite below the +central green organs of the flower, and that they +are in no way influenced by the petals; we also see +that the petals are entirely separate from the other +parts of the flower, and we learn, as the result of +our examination, that the parts of the buttercup +are <i>free</i>. To express this botanically we prefix +the word “poly” to the words sepals and petals, +and so we get <i>polysepalous</i>, meaning that the sepals +are quite free and distinct, and <i>polypetalous</i> referring +to the same condition of the petals.</p> + +<p>Now, having removed the petals and sepals, we +<span class="pagenum" id="Page_154">[Pg 154]</span>can proceed to study the other parts of the +flower.</p> + +<p>First we find a great number of little yellowish +stalks tipped with tiny pouches; these are the +stamens, and in the little pouches (anthers) the +yellow powder termed pollen is developed. We +will carefully take away these stamens, and note in +so doing that they are all distinct and all sprung +below the green central part. Like the sepals and +petals, we find the stamens are free and uninfluenced +by the other parts. If we again compare +this with a primrose-flower we shall find a +difference; the stamens of the primrose spring +from the petals and are therefore called <i>epipetalous</i> +(<i>epi</i> upon, a petal). Again in the sweet-pea or +scarlet runner we find the stalks of the stamens +are all joined together. We now have left upon +the flower-stalk the little central green parts +previously mentioned; there are quite a number +of them; each one is distinct from its neighbour +and is free. These bodies are known as carpels, +they are large at one end and taper to a curved +point at the other, the broad end being attached to +the stem. Collectively these carpels constitute the +<span class="pagenum" id="Page_155">[Pg 155]</span><i>pistil</i>, and because the carpels are apart and free it +is said to be <i>apocarpous</i>.</p> + +<p>The flower of the little woodsorrel (<i>Oxalis +acetosella</i>) will help us to understand better the +arrangement of the carpels. If we take away the +sepals, petals and stamens, we shall have only the +carpels left, and these are five in number. They +are in the same position as those of the buttercup, +but they are not separate, they are joined by their +inner surfaces. We can plainly see that this is the +case, since each carpel is distinctly outlined and +there are five little tapering ends (stigmas). The +pistil in this case is said to be <i>syncarpous</i>.</p> + +<p>Names are given to express some quality, and +they often draw our attention to interesting facts +about the plant’s mode of growth or the place +where it is found; for instance, the pretty blue +nemophila is so called from <i>nemos</i>, a grove, and +<i>philo</i>, I love, because it delights in shady places.</p> + +<p>Geranium is derived from <i>geranos</i>, a crane, +because the fruit of some of the species resemble +the beak of that bird.</p> + +<p>Some plants are named after famous botanists, +as <i>Linnæa</i> after Linnæus.</p> + +<p><span class="pagenum" id="Page_156">[Pg 156]</span></p> + +<p>Others derive their names from their mode of +growth, as stone-crop, which is called sedum, from +<i>sedo</i>, I sit, the plant having scarcely any stalk, and +sitting, as it were, on walls and rocks.</p> + +<p>These instances will show that it is well worth +while to study names and learn their meanings, as +they often throw so much light upon the history of +a plant.</p> + +<p>In the flowers of bulbous plants we find that the +calyx and petal leaves are frequently alike in +colour and texture; in that case the three sepals +and three petals, of which they usually consist, are +spoken of as a perianth.</p> + +<p>In looking at the brilliant colouring of a flower +we should hardly imagine that the petals have to +some extent the nature of leaves, and under certain +conditions may be changed to the green colour +and form of ordinary leaves.</p> + +<p>In very wet seasons we may sometimes find +rose-buds with the sepals of the calyx developed +into perfect green leaves. The floral envelopes +therefore possess the nature of true leaves.</p> + +<figure class="figcenter illowp100" id="i_157" style="max-width: 62.5em;"> + <img class="w100" src="images/i_157.jpg" alt="A tree branch with some +20 leaves, all growing from the end. The leaves are elongated, oval, with sharp +tips. From the end of the branch some flowers grow. In detail a flower consists +of a bulbous form with a lip-like structure. At the top of the bulb grows a bud +from which curly threads appear."> + <figcaption> + <p>POINSETTIA.</p> + </figcaption> +</figure> + +<p>The brilliant scarlet so-called flowers of the +poinsettia are really coloured bracts, the true +<span class="pagenum"><a id="Page_157"></a><a id="Page_158"></a>[Pg 158]</span>flower being the small inconspicuous blossom in +the centre.</p> + +<p>In the chapter on leaves we saw that bracts are +those small imperfectly-shaped leaves in the axils +of which flowers are placed. They are usually +green, but may be also brilliantly tinted as in the +mauve-coloured Bougainvillia, the bright violet +spikes of the <i>Salvia Hormineum</i>, and also pure +white as in the spathe of the arum.</p> + +<p>By special cultivation flowers can be made +double, for excess of nourishment will cause the +plant to multiply its petals. Instead of the five +pink petals of the wild rose we find one of our +garden roses bearing as many as eighty or a +hundred petals.</p> + +<p>Double flowers but rarely produce seeds, because +the stamens and pistil have been turned into petals, +and as there is no need to attract insects for +fertilising purposes, there is no secretion of honey, +and therefore we scarcely ever see honey-bees in +double flowers; they are wise enough to know +that their visits to them would be in vain.</p> + +<p>In composite flowers such as asters and sunflowers +the change, when they are double, occurs +in several ways.</p> + +<p><span class="pagenum" id="Page_159">[Pg 159]</span></p> + +<p>The centre may become filled with florets similar +to those in the outside ring, or the florets in the +middle may become larger or of a different colour.</p> + +<p>These various changes may be readily observed +in the cultivated chrysanthemums, in which every +form and variety of flowering can be traced.</p> + +<p>When the pollen has reached the pistil the +flower begins to fade, because its end has been +attained; nature, however, has such variety in +even the smallest of her operations that the +passing away of a flower is accomplished in +different ways. In the primrose the corolla withers +and drops to the ground. The flower of the +spiderwort, one of our common garden plants, +becomes pulpy as it fades, in this way resembling +the pineapple<a id="tn-022"></a> plant, the flower of which eventually +becomes the luscious succulent fruit.</p> + +<p>The poppy is proverbial for its fleeting petals, +which scarcely last more than a few hours, a +passing wind soon scattering them far and wide.</p> + +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“Pleasures are like poppies spread,</div> + <div class="verse indent1">You seize the flower, its bloom is shed!</div> + <div class="verse indent1">Or like the snow-fall in the river,</div> + <div class="verse indent1">A moment white—then melts for ever.” (<span class="smcap">Burns.</span>)</div> + </div> + </div> +</div> + +<p>Some flowers, as the hydrangea, have persistent +<span class="pagenum" id="Page_160">[Pg 160]</span>petals, which simply lose their brilliant tints and +become tough and brown.</p> + +<figure class="figcenter illowp100" id="i_160" style="max-width: 62.5em;"> + <img class="w100" src="images/i_160.jpg" alt="A branch bearing four fruits, +that are heart shaped and show heavy veins. They are hanging from bent stalks. +In detail a cross section is shown. There is the stalk, ending in a circular shape. +Dotted lines indicate the heart-shape of the outer structure."> + <figcaption> + <p>WINTER CHERRY.</p> + </figcaption> +</figure> + +<p>The calyx of the physalis or winter cherry +<span class="pagenum" id="Page_161">[Pg 161]</span>continues to grow after the flowers are fertilised +until the round balloon-like bag is formed in +which the fruit is enclosed.</p> + +<p>We will now examine the parts of a flower +separately, beginning with the calyx.</p> + +<p>In the buttercup the calyx consists of one whorl +or ring of five sepals.</p> + +<p>In the strawberry there are two whorls of sepals, +and in the cotton plant there are three whorls +forming its green calyx.</p> + +<p>There are also variations in the mode of flower +expansion.</p> + +<p>As a poppy-bud opens it detaches its calyx +from the stem, and the sepals fall off (the calyx is +therefore called caducous, a term which means +ready to drop off).</p> + +<p>Many flowers retain the calyx until the petals +wither and it falls off with them. It is then called +a deciduous calyx.</p> + +<p>Others again have a permanent calyx, so that +when, as in the primrose, the corolla withers and +drops off, the sepals close over the seed-vessel and +protect it until the seeds are matured; this would +be called botanically a persistent calyx.</p> + +<p><span class="pagenum" id="Page_162">[Pg 162]</span></p> + +<p>The best way to learn the names of the different +parts of a flower is to pull it carefully to pieces +and arrange the separate organs on a thin card. +They can be tacked on to the card with a stitch +or two of fine thread, and +when the lesson is over, if the +card is placed between sheets +of blotting-paper under a +weight, the flower dissections +will dry and be useful for +reference later on.</p> + +<p>Each separate part of the +flower should have its name +neatly written beneath it, so +that when a good many different +flowers have been thus +dissected they may be compared +and the variations in +form and position duly noted.</p> + +<p>A wallflower will be a +good subject for our dissection.</p> + +<p>At the back of the petals we first take off the +calyx, which consists of four divisions called +sepals. We then pull off the four yellow petals, +<span class="pagenum" id="Page_163">[Pg 163]</span>and as they are placed in the form of a cross it +shows that this plant is a crucifer, or cross-bearer, +one of a very large natural order, <i>Cruciferæ</i>,⁠<a id="FNanchor_16_16" href="#Footnote_16_16" class="fnanchor">[16]</a> none +of which are poisonous and very many are useful +food-plants, such as cabbage, turnip, watercress, +and cauliflower. Now there remain six stamens—four +long and two shorter ones; these last rise +outside of and alternate with two nectaries or +honey-glands.</p> + +<figure class="figleft w30" id="i_162" style="max-width: 31.625em;"> + <img class="w100" src="images/i_162.jpg" alt="A stem bearing three small leaves +at the base. Upward there are about 10 flowers, in different stages of ripeness. +At the top there are still buds, in the middle they are in full bloom and further +down blooming is over. Each flower has 4 equally sized petals. They emerge from +an elongated calyx, that sits on a short stalk."> + <figcaption> + <p>WALLFLOWER.</p> + </figcaption> +</figure> + +<p>The stems of the stamens are called filaments, +from <i>filum</i>, a thread; and the upper part, containing +yellow powder, is called the anther, the proper +name for the powder itself being pollen.</p> + +<p>In the centre of the flower is the pistil, the lower +part of which is the ovary, the part of a flower +which contains the ovules, and is so named from +<i>ovum</i>, an egg.</p> + +<p>The stem part of the pistil is called the style +and the top of it is the stigma.</p> + +<p>Such simple words as I have given must be +learned, else we cannot understand botanical +descriptions, and if this page is studied whilst we +have the flowers in our hands it will not be difficult +<span class="pagenum" id="Page_164">[Pg 164]</span>to identify each separate organ; when these are +once arranged on a card with the name of each +part written beneath it, we shall have attained +some very useful information ready for future +study.</p> + +<p>In the buttercup flower all the five petals are +the same size and shape; therefore, like hundreds +of other evenly-formed blossoms, it would be +described as “regular”; but if we take a sweet-pea, +balsam, or monkshood-flower and examine its +separate petals, we shall find they vary very much +in form, and they are known as “irregular” +flowers.</p> + +<p>The sweet-pea is a type of a large order of +plants producing what are called butterfly-shaped +flowers, and <i>papilio</i> being Latin for a butterfly, +they are therefore called papilionaceous flowers. +If we learn clearly about the various parts of such +a flower we shall henceforth be able to recognise it +at a glance.</p> + +<p>In the sweet-pea we find a broad petal at the +back of the flower which is called the standard, +beneath it are the two side petals called wings, and +within them is the keel, so named because it is +<span class="pagenum" id="Page_165">[Pg 165]</span>shaped like the bottom of a ship. Within the keel +lie the stamens and pistil—the most important +parts of the flower, and to protect them from injury +the standard is so formed as to catch the wind like +a sail and turn the blossom round so that this +broad petal shelters the keel from rain.</p> + +<figure class="figcenter illowp63" id="i_165" style="max-width: 51.8125em;"> + <img class="w100" src="images/i_165.jpg" alt="A bunch of 4 flowers, one +of which still in bud, the others fully flowering. Each flower has a complex form +because the petals are of different shapes and sizes. On top is one large petal, +lower down are 2 smaller petals, arranged alongside each other. At the bottom is a +smaller structure formed out of 1 or 2 petals. Each flower has a small, flat +calyx and sits on a thin stalk."> + <figcaption> + <p>SWEET-PEA.</p> + </figcaption> +</figure> + +<p>In our next ramble out of doors it will be well +to try and gather all the specimens we can find of +this order of plants. If it be in summer or autumn +we shall soon collect a handful of these butterfly-shaped +flowers.</p> + +<p>On a common we shall find broom, furze, restharrow, +<span class="pagenum" id="Page_166">[Pg 166]</span>vetches, tares, trefoil, clover, saintfoin, and +other plants. In the garden and greenhouse we +shall see many more species belonging to this +class.</p> + +<p>Having shown the difference between a regular +and irregular flower, we will now proceed to notice +how irregularity is caused.</p> + +<p>If we pull off one of the buttercup petals and +look at the base of it, we shall see a small pouch +which contains honey; it is called a nectary or +honey gland, and the position of this gland has +much to do with the shape of the flower.</p> + +<p>As each petal of the buttercup has a nectary at +its base it follows that, all the petals being the same +size and shape, the flower is perfectly regular—like +a small golden cup. Now in other flowers we +shall find the nectary very large and confined to +one petal or sepal only, and this results in the +flower having an irregular shape. Gather a violet, +examine and compare the petals; four of them +will be found to be nearly alike, but the lower petal +is much larger because it has grown into a tube +(called a spur) to secrete honey, and I need hardly +say that the honey is intended to attract the bees +<span class="pagenum" id="Page_167">[Pg 167]</span>so that the flower may be enabled to produce +fertile seed. The enlargement of the lower petal +gives the flower an irregular shape, and the same +thing happens in the monkshood and many other +flowers, where both the petals and sepals are +thrown out of shape to form nectaries. In the +orchid family this influence may be traced to a +wonderful degree. The contrivances for insuring +the fertilisation of their flowers are so many and +various that books of the greatest interest have +been written on that subject alone.</p> + +<p>In the flowers we have hitherto noticed, both +stamens and pistils are found, the petals are +coloured, honey-glands exist, and some specimens +also possess a powerful scent.</p> + +<p>Such flowers are obviously very attractive to +insects, and on that account they are called by +modern botanists, entomophilous, which long word +means that they are beloved by insects.</p> + +<p>In sharp contrast to these gay and conspicuous +flowers we may observe the very simple catkins of +the birch, <i>Betula alba</i>. If we examine a twig of +this tree in spring, we shall find two very distinct +kinds of flowers (or catkins, as tree-blossoms ought +<span class="pagenum" id="Page_168">[Pg 168]</span>properly to be called), one a stiff green spike +standing upright, and the other longer and of +yellowish colour, always to be found hanging +down.</p> + +<p>The former consists of a number of scales +arranged on a central stem, and in the axil of each +scale is the little pistil, with its pointed and divided +stigmas. This catkin, later on, becomes the fruit +of the tree, and sheds out with every passing breeze +its little winged fruits, which are carried far and +wide and often sow themselves in rocky crevices, +and appear able to grow and flourish with only a +modicum of soil.</p> + +<figure class="figcenter illowp100" id="i_168" style="max-width: 53.8125em;"> + <img class="w100" src="images/i_168.jpg" alt="Two pictures of a seed, one +natural size, one magnified. The magnification shows a grain in the centre +with two wings at the sides. The grain is elliptic, with pointed ends. The +wings are rounded."> + <figcaption> + <p> + BIRCH FRUIT.</p> + </figcaption> +</figure> + +<p>The pendulous catkin is very soft and loose, and +on the inner surface of its scales we find the +stamens, which in due time will shed from their +anthers the fertilising pollen. Here then we see +flowers which are not so attractive to insects, +<span class="pagenum" id="Page_169">[Pg 169]</span>flowers in which the stamens and pistils are +separated and developed in different catkins, and +such flowers are termed monœcious, from <i>monos</i>, +single, and <i>oikos</i>, a house.</p> + +<p>The most interesting feature of these tree-blossoms +is their fertilisation by the wind; the +slightest puff of air liberates little clouds of pollen +from the loose swinging anthers; these pollen +grains become entangled in the upright catkins +bearing the pistils, and the future seed thus +becomes fruitful. There are many trees and plants +which are thus fertilised by the agency of the wind, +and they are termed by botanists <i>anemophilous</i>, +from the Greek words <i>anemos</i>, wind, and <i>philos</i>, +beloved by.</p> + +<p>In the common bryony of the hedges, we get +another example of a green inconspicuous flower. +Gather a few sprays of this in early summer, +taking care to keep the specimens of each plant +separate. Take up one specimen and you will find +each flower has a small green calyx, a minute +corolla, and five little stamens; not one pistil can +we find on the spray.</p> + +<p>The flowers on the next spray look very +<span class="pagenum" id="Page_170">[Pg 170]</span>similar, but in them there are no stamens, the +centre of each flower being occupied by a small +pistil, and thus we learn that there are two distinct +sexes in the bryony plant, the one bearing only +staminate flowers, and the other +producing those bearing only +pistils. Such plants are termed +diœcious, from <i>di</i>, two, and <i>oikos</i>, +a house.</p> + +<figure class="figleft w30" id="i_170" style="max-width: 28.25em;"> + <img class="w100" src="images/i_170.jpg" alt="A cross section of a flower. +Calyx and petals are shown schematically. They build a flask-like shape. From the +bottom grows an intricate structure. At the base is a bulbous shape built from +smaller forms, giving it the appearance of a blackberry. On top of that is a +ball, with a pattern of small circles. On top of that is a club-like shape, about +2 times the size of the bulbs beneath it combined. At the base of the club grow +hairs."> + <figcaption> + <p>WILD ARUM.</p> + </figcaption> +</figure> + +<p>One of the earliest spring +flowers is the arum of the hedges, +known to village children as +“lords and ladies.” Accustomed +as we are to bright-hued flowers +if in our gardens and fields, it is +somewhat difficult to recognise +that the pale-green sheath of +the arum is a flower at all. It +consists of a beautifully-folded +spathe or bract, curving over at +the top, and if we remove that we find a central +stalk bearing a number of little naked flowers, +arranged in the order shown in the plate.</p> + +<p>First, below the club-like apex, a few hairs tending +<span class="pagenum" id="Page_171">[Pg 171]</span>downwards, then the anthers containing pollen, +and below these the pistils with protruding stigmas. +The whole stalk is termed a spadix.</p> + +<p>The outer green spathe forms a kind of prison, +into which flies are enticed by the somewhat fetid +odour which is exhaled by the flower. The flies +easily creep in past the circle of hairs, which, as +they point downwards, do not prevent their +entrance, but, once in, these hairs are like a +<i>chevaux-de-frise</i>, and hinder the escape of the +insects. The flies in all probability carry upon +their wings pollen from some other arum flower, +and in their efforts to escape they brush off this +pollen upon the stigmas, which thus become +fertilised. When this has taken place the stigmas +throw out a sweet juice upon which the insects +feed; the anthers now shed out their pollen, with +which the flies become covered; the hairs meanwhile +have withered, and thus the flies, having done +their appointed work in fertilising the flower, are +free to crawl out and perform the same office for +some neighbouring plant.</p> + +<p>We have not space to do more than allude to +certain plants, whose flowers never open and are +<span class="pagenum" id="Page_172">[Pg 172]</span>self-fertilised. The common violet, for instance +produces, in addition to its well-known fragrant +flowers, certain inconspicuous blossoms, hidden +under the leaves and close to the root, very seldom +noticed by any but botanists, and known to them +as cleistogamic flowers; these are fertile, and +always produce seed. Other such plants are the +woodsorrel and sundews.</p> + +<p>It is interesting to observe the various ways in +which flowers are protected from browsing animals, +snails, and caterpillars by thorns, spines, prickles, +and spiny bracts. The teasel secretes water in the +bracts around its stem, which prevents ants from +ascending to the flowers, and in many plants we +may see quantities of small insects caught by a +sticky gum exuded from the leaves and twigs.</p> + +<p>Many delicate plants entirely alter the position +of their flowers in order to protect them from rain. +On a sunny day the wood-anemone holds its little +snowy cup so as to receive the full sunlight, but on +a damp day every blossom is closed and held +downwards. We may observe this in the poppy, +the blue-anemone, and nearly all composite flowers.</p> + +<p>These are merely hints scattered over a wide +<span class="pagenum" id="Page_173">[Pg 173]</span>field of study, which some readers may like to +follow out.</p> + +<p>Objects to collect and examine:—Buttercup +flowers, seed-vessel of wild-geranium, stonecrop +growing on walls, flowers of bulbous plants, flowers +of poinsettia, bougainvillia, salvia hormineum, +arum. Examine various chrysanthemum flowers, +sunflowers, asters and woodsorrel. Difference +between hydrangea and poppy flowers, winter +cherry (physalis); prepare flower dissections. +Examine various cruciferous flowers and pea-shaped +flowers, regular and irregular flowers, birch +catkins, wild arum flowers, cleistogamous flowers, +protection of flowers, position of flowers.</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum"><a id="Page_174"></a><a id="Page_175"></a>[Pg 175]</span></p> + + + <h2 class="nobreak" id="CHAPTER_VII"> + CHAPTER VII + <br> + <i>POLLINATION</i> + + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_176">[Pg 176]</span></p> +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent6">“When summer shines,</div> + <div class="verse indent0">The bee transports the fertilising meal</div> + <div class="verse indent0">From flower to flower, and even the breathing air</div> + <div class="verse indent0">Wafts the rich prize to its appointed use.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">Cowper.</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_177">[Pg 177]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_VII_1"> + CHAPTER VII + <br> + <br> + <span style="font-size: 70%;">POLLINATION</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_177.jpg" + style="width: 12%; height: auto;" + alt="W"> +</div> +<p class="drop-cap"><span class="upper-case">We</span> +now come to the consideration of the +real function of the flower of a plant. +In whatever form it is developed, +whether as a gay and fragrant blossom, +in a dull foul-smelling structure like the +arum, or as a green inconspicuous little floret like +the grass, its main office is to reproduce itself by +the formation of seed. We will first glance at +some of the wonderful agencies that actively help +in this work.</p> + +<p>There are at least three distinct processes +necessary for the complete formation of a perfect +seed, and we must, I fear, persuade ourselves to +learn some of the long words by which botanists +speak of these processes. They are known as +<span class="pagenum" id="Page_178">[Pg 178]</span>pollination, fertilisation, and the growth of the +ovule. There is so much to be said about the +first subject, that I must leave the two latter for +a succeeding chapter.</p> + +<p>Before seed can be formed it is necessary that +the powder contained in the anthers, which is +called pollen, should be transferred from those +anthers to the stigma or upper part of the pistil, +and this transference is called pollination. If we +examine a tulip or, better still, a buttercup, we +find the anthers and stigmas so near together +that the transfer of the dust-like pollen to the +sticky-looking stigmas can easily take place. +This would be called an instance of self-pollination, +but although cases of this kind do +occur in nature, they are not at all common. +As a rule, in order to ensure what is called +cross-pollination, the transfer of the pollen of one +flower to the stigma of another, many wonderful +and interesting arrangements exist even in some +of our commonest flowers.</p> + +<p>Cross-pollination must be the case in such plants +as dog’s mercury, because we find in a colony of +these plants—so frequently seen by the roadside—that +<span class="pagenum" id="Page_179">[Pg 179]</span>some plants have flowers with stamens only, +and others containing only pistils. Again, in the +hazel we may see how impossible it is for self-pollination +to take place, as, if we examine the +pistils, we find that they consist of scales bearing +stamens and pollen only, whilst somewhere close +by, on the same stem, hangs the pretty little red +flower which possesses the pistil and forked stigma. +If seed is to be formed in either of these flowers +and in many others similarly arranged, then the +pollen of one flower must be transferred to the +stigma of the other.</p> + +<figure class="figcenter illowp72" id="i_179" style="max-width: 56.625em;"> + <img class="w100" src="images/i_179.jpg" alt="Two flowers is cross section. +Both have a funnel-like shape. In both sections the anthers en pistils are shown. +In the left one the pistil is short, so it only reaches halfway the funnel. +The anthers are in the opening. In the right on the pistil is long and reaches +the opening. The anthers are halfway the funnel."> + <figcaption> + <p>PRIMROSE.</p> + </figcaption> +</figure> + +<p>There are interesting facts to be learned about +the common primrose. When we examine a little +bunch of these flowers we find quite half of them +are what children call pin-eyed, meaning that the +<span class="pagenum" id="Page_180">[Pg 180]</span>stigma, which is at the end of a long pistil, is like +the head of a pin in the throat of the primrose.</p> + +<p>Looking at the sketch, we see at once that self-pollination +is hindered by the fact that the anthers +in this flower being at the bottom of the tube, the +pollen they contain must be transferred by some +direct agency before it can come in contact with +any stigma. Now let us examine the other flowers +in our primrose nosegay; we find the stamens in +these are placed in the mouth of the tube, and the +pistil is quite short and low down in position. At +first sight it appears as if the pollen would fall +directly upon the pistil, since the stamens are +above that organ, but this is not exactly what +happens; the pollen of this particular form of +flower is shed before the stigma is mature, so +that when it has reached maturity the pollen +is all gone.</p> + +<p>The arrangement of nature is as follows. An +insect attracted by the sweet-smelling bank of +primroses will visit the flowers, thrusting its +proboscis down a pin-eyed flower until in so +doing its head has been dusted with the pollen +of the stamens; then withdrawing from that +<span class="pagenum" id="Page_181">[Pg 181]</span>flower the insect visits another near by, possibly +one with a short pistil; the pollen on its head +is now rubbed off and falls upon the stigma below +and pollinates it, for that is the term used when +this act takes place.</p> + +<figure class="figcenter illowp93" id="i_181" style="max-width: 62.5em;"> + <img class="w100" src="images/i_181.jpg" alt="Two flowers, both funnel-shaped. +Each has 5 petals of equal size, arranged around the opening. In the left one +the stigma is shown that has 2 parts, each a thread-like structure that grow +out of the opening, to opposite sides of the flower. In the right one the +anthers are shown: a group a small bud-like shapes standing on thin stalks, right +in the opening of the flower."> + <figcaption> + <p>MAIDEN PINK.</p> + </figcaption> +</figure> + +<p>The pretty maiden-pink will help us still more +clearly to understand how cross-pollination is promoted +in flowers containing both stamens and +pistils. Select a flower that has just opened, the +petals of which are spreading and fringed, whilst +from the centre of the flower a cluster of stamens +projects with the pollen mature and easily shaken +<span class="pagenum" id="Page_182">[Pg 182]</span>out of the anther lobes; the pistil is concealed in +the long tube, and in this stage there is no sign +of stigma. In a short time, however, if we examine +the flower again, we shall find the stamens have +shrivelled up, and in their place a forked stigma +appears, as shown in the sketch. Here again it is +obvious that the fact of the stamens ripening first +and expending their energy before the pistil is ripe +must mean, that in order to secure seed the pollen +from some younger flower must be transferred, +probably also by insect agency. It will give fresh +interest to our garden rambles if we remember that +the bees and flies we see hovering over the flowers +are not only collecting honey or feasting upon it, +but are also performing a very important office for +the benefit of the plants they are visiting.</p> + +<p>We may now proceed to notice the various +agencies for the conveyance of pollen between +flowers.</p> + +<p>These agencies are water, wind, insects and birds.</p> + +<p>In an earlier chapter I gave an account of the +<i>Vallisneria spiralis</i>, which will serve as a type of +a water-pollinated flower.</p> + +<p>Those pollinated by wind are, as I have said in +<span class="pagenum" id="Page_183">[Pg 183]</span>a previous chapter, called anemophilous (<i>anemos</i>, +wind, and <i>philos</i>, loving). They are usually of +small size and inconspicuous character, with very +little or no scent, and devoid of colour; these are +characteristics that are not always associated in the +same species; thus in the hazel, which is a wind-pollinated +flower, we find a bright yellow catkin (so +well known to children as lambs’ tails) and a small +but bright red pistil.</p> + +<p>Let us notice, however, how wonderfully these +plants are adapted for this method of pollination; +the stamens are usually hanging, and the pollen, +produced in great quantities, is easily set free by +the slightest breath of wind. The stigma of the +hazel, of different grasses and of sedges are both +forked and plumed, so that pollen grains floating in +the air are readily intercepted.</p> + +<p>The firs and pines are excellent examples of +wind-pollinated trees. I remember once possessing +a ripe male cone of the <i>Araucaria imbricata</i>, and +ascertaining that it contained as much as a wine-glassful +of pollen. Speaking about this fact to the +gardener at the Pinetum at Dropmore, I was +shown how this fertilising dust from the great +<span class="pagenum" id="Page_184">[Pg 184]</span>Araucaria (which was planted there in 1830) was +carried by the wind for an amazing distance to a +female tree on the other side of the garden, pollinating +its cones so that they produced fertile seeds. +In some of the Canadian pine forests, the trees +shed forth such quantities of pollen in the flowering +season that the ground becomes perfectly yellow. +The early settlers, being unable to account +for the strange phenomenon in any other way, +ascribed it to showers of sulphur descending from +the clouds. Even in our own country the foliage +and undergrowth in the neighbourhood of fir +woods is often thickly coated with the yellow dust +falling from the male catkins of the trees; the +structure of the pollen grains is such that they float +very buoyantly, each grain being provided with two +air bladders. I may mention in passing that this +apparently wasted pollen affords a rich feast to +endless species of bees and flies, and is in many +cases stored up by them as food for their young +grubs. The various adaptations for wind pollination +will perhaps be better understood if we glance +at the attractions which flowers offer to birds and +insects.</p> + +<p><span class="pagenum" id="Page_185">[Pg 185]</span></p> + +<p>Colour serves to render flowers attractive to +insects, and to make them conspicuous; the bracts, +petals, and sepals of flowers are usually of some +light or dark colour quite distinct from the green +tone of the foliage.</p> + +<p>It has been ascertained also that plants which +are pollinated by night-flying moths generally have +white or light-yellow flowers so as to be easily seen +in twilight.</p> + +<p>One of the most interesting of these night-pollinated +flowers is <i>Silene nutans</i>, the Nottingham +catchfly. In the daytime the five narrow petals +are curled up and look dead and withered, but as +night comes on they change their position, and the +flower has the expanded shape of an alpine pink. +In this open condition it is visited by the moths +which, flying from one flower to another, transfer +the pollen, and thus accomplish at night what more +frequently occurs in the sunlight; at daybreak the +petals roll up once more, and one would again suppose +the flower to be dead; but no, it will continue +to open at nightfall until some moth finally succeeds +in pollinating its blossom. A small species +of moth⁠<a id="FNanchor_17_17" href="#Footnote_17_17" class="fnanchor">[17]</a> visits this catchfly in order to deposit its +<span class="pagenum" id="Page_186">[Pg 186]</span>eggs; these, by means of a very long ovipositor, it +places in the ovary, and in that somewhat inflated +cavity they produce microscopic caterpillars which +find shelter and nutriment in the strange nest. +<span class="pagenum" id="Page_187">[Pg 187]</span>When the caterpillars arrive at maturity they +escape by biting a hole in the wall of the capsule, +and creeping out, they seek for a suitable place in +which to turn to chrysalides.</p> + +<figure class="figcenter illowp80" id="i_186" style="max-width: 62.5em;"> + <img class="w100" src="images/i_186.jpg" alt="A stem with 4 leaves and a flower. +The leaves are grouped in pairs. In each par one leaf is small and sits directly +on the stem. The other is large, heart-shaped and sits on a stalk. The flower +sits on a stalk that emerges from the axil of the upper pair of leaves. It has the +form of an antique clay pipe, or a saxophone. The base is small and directed +downward, but then it widens and bends upward. The petals have a pattern of small +irregular spots, like a leopard skin."> + <figcaption> + <p>ARISTOLOCHIA.</p> + </figcaption> +</figure> + +<p>Scentless flowers usually have some equivalent +form of attraction, such as honey, brilliant colour +pollen in abundance, or the grouping of a number +of small florets, in order to secure a conspicuous +effect as in the ox-eye daisy, or hedge parsley.</p> + +<p>Strong and varied odours are great helps to +ensure pollination by insects. The bee-tribe and +moths and butterflies are specially attracted by the +sweet scents of roses, violets, carnations, and sweet-peas, +and the powerful odour emitted by such +flowers as the evening primrose, tobacco, and night-flowering +rocket as evening comes on tends to +guide the nocturnal moths to these and similar +flowers. An odour may, of course, be pleasant to +an insect which to us would be simply intolerable. +The arum of the hedges, and those curious plants, +the aristolochias and stapelias, all emit scents of +the most fœtid description, as we think, but flies, on +the contrary, are attracted by thousands, and +hold apparently joyous revels in the blossoms +<span class="pagenum" id="Page_188">[Pg 188]</span>which they are pollinating by their frequent +visits.</p> + +<figure class="figcenter illowp80" id="i_188" style="max-width: 62.5em;"> + <img class="w100" src="images/i_188.jpg" alt="A stem covered with +scale-like leaves from which a bud and a flower grow, both on stalks. +The bud is round, a little flattened and with a point where the petals meet +in the centre. The flower is shown from above. It has 6 equally formed petals. +Each is rounded with a sharp tip. The petals have a pattern of dark spots and lines +perpendicular to their axis. The petals are arranged around a centre that is about +half the diameter of the flower as a whole. The centre is dotted as well and in the +midst the is a star shape that contains pistil and anthers."> + <figcaption> + <p>STAPELIA.</p> + </figcaption> +</figure> + +<p>A little care and patience in watching the visits +of insects to different flowers will soon be rewarded +<span class="pagenum" id="Page_189">[Pg 189]</span>by a perception of the tastes and likings of insect +life, and we shall gradually learn to expect to see +certain insects on the flowers they specially frequent.</p> + +<figure class="figright w30" id="i_189" style="max-width: 34.75em;"> + <img class="w100" src="images/i_189.jpg" alt="A stalk with at the end +2 elongated leaves, a pistil that sits on a stalk of its own and a group of 8 +anthers. Each anther has a 2-lobed shape and sits on a thin stalk."> + <figcaption> + <p>HYPERICUM.</p> + </figcaption> +</figure> + +<p>I would call attention to the interesting fact that +if one agency fails to effect pollination, another is +adopted in order to attain the desired end. Thus, +when the flowers of the +common bartsia first open, +they are visited by insects; +but, in the later stages of +flowering, the pollen is blown +out by the wind, and the +neighbouring stigmas thus +become pollinated. We see +in the arrangement of the +flower of the St. John’s wort +(<i>Hypericum</i>) a perfect type +of this provision against any +possible failure of pollination. The stigma is surrounded +by groups of stamens of unequal length; +those in the centre nearest to the stigma are as long +as the style itself, whilst those on the outside are +short, and these shed their pollen first, whilst those +in close contact with the stigma shed their contents +<span class="pagenum" id="Page_190">[Pg 190]</span>last. Thus we find that if insects fail to effect +cross-pollination by means of the short and early +opened stamens, it is secured by means of the +longer stamens whose anthers are in close contact +with the stigma. Again, when we stand under a +sycamore tree, we may see that the green tassel-like +flowers are having their pollen dispersed both by +wind and bees.</p> + +<p>We cannot draw hard-and-fast lines in nature, +for although a special end may be kept in view, the +various means and adaptations by which it is +attained are a continual source of admiration and +wonder to the reverent student of nature.</p> + +<p>We have already seen that there are all kinds of +devices by which the pollen of one flower may be +made sure to reach the stigma of another; but, if +by any means this crossing fails, if the weather is +such that insects are scarce, or other conditions +cause failure, then, in the case of many flowers, +most curious contrivances are provided to secure +seed by self-pollination. Truly this is one of the +most beautiful of God’s wonders in floral construction. +One of the gems of my own flower garden +is a lovely little Japanese toad-lily (<i>Tricyrtis hirta</i>). +<span class="pagenum" id="Page_191">[Pg 191]</span>In this flower there are three styles which stand +well above the stamens; the points of the styles +are bent over as in the plate, and the stigmatic surface +grows mature before the anthers shed their +pollen; if, however, no insect visits the flowers, pollination +is effected in the following way. The +styles bend down and place their forked points in +direct contact with the open anther-lobes (as shown +in drawing), the style assuming +almost the form of a semicircle. +This is done very deliberately, +for it is often fully +a week before the act is complete.</p> + +<figure class="figright w30" id="i_191" style="max-width: 31.0625em;"> + <img class="w100" src="images/i_191.jpg" alt="2 drawings of a pistil and anther. +In both the pistil is on top. It is a large form, bent downward at the end. The +anther is lower and consists of a 2-lobed structure at the end of a straight stalk, +almost perpendicular to the base of the pistil. In the left drawing the pistil +is bent to the extreme, thus making contact with the anther. In the right drawing +the pistil is bent less, and there is no contact."> + <figcaption> + <p>TOAD-LILY.</p> + <p><i>Stigma and Stamen.</i></p> + </figcaption> +</figure> + +<p>Pollination is effected in tropical countries not +only by insects of many kinds, but by the lovely +tribes of humming-birds which abound in those +regions. Their slender, curved beaks are specially +adapted to penetrate the honey-laden flowers with +long-tubed blossoms, which could only be pollinated +by some such agency.</p> + +<p>Those who are within reach of the Natural History +Museum at South Kensington may there see a +gallery filled with exquisite specimens of humming-birds, +<span class="pagenum" id="Page_192">[Pg 192]</span>arranged in cases, and some of the birds are +shown as they appear in life, hovering over tropical +flowers, drawing honey from their hanging blossoms, +and performing the useful office of transferring the +pollen from one flower to another, thus ensuring +the fertilisation of the seed.</p> + +<p>I might go on multiplying examples of the +various methods by which seed is rendered fertile, +but perhaps enough has been said to show what +hidden force exists in flowers to enable them to +attain the end for which they mainly exist, namely, +the perpetuation of their species by means of seed.</p> + +<p>Specimens to be obtained and compared with +the descriptions in this chapter:—Buttercup flower, +dog’s mercury, hazel catkins, primrose flowers, +male blossoms of pine trees in June, Nottingham +catchfly, ox-eye daisy, bartsia, St. John’s wort +flowers, and Japanese toad-lily.</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_193">[Pg 193]</span></p> + + + <h2 class="nobreak" id="CHAPTER_VIII"> + CHAPTER VIII + <br> + <i>FERTILISATION</i> + + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_194">[Pg 194]</span></p> +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent34">“The men</div> + <div class="verse indent0">Whom Nature’s works can charm, with God Himself</div> + <div class="verse indent0">Hold converse.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">Akenside.</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_195">[Pg 195]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_VIII_1"> + CHAPTER VIII + <br> + <br> + <span style="font-size: 70%;">FERTILISATION</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_195.jpg" + style="width: 12%; height: auto;" + alt="H"> +</div> +<p class="drop-cap"><span class="upper-case">Having</span> +now considered some of the +many wonderful arrangements by +which the pollen of plants is dispersed, +we will endeavour by tracing the course +of the pollen-grains after they reach the +stigma, to learn what is meant by the term “fertilisation +of the ovules.” These are the minute +specks contained in the ovary which are to become +seeds, and by means of which the plant will +eventually reproduce itself.</p> + +<p>To the naked eye the yellow pollen we see on +the anthers of flowers appears as small grains; +but, when magnified, these grains are seen to be +singularly beautiful, each little sphere having on +<span class="pagenum" id="Page_196">[Pg 196]</span>its surface a chequered network and delicately +sculptured patterns.</p> + +<p>The forms, too, are as varied as the ornamentation.</p> + +<p>Some plants have triangular grains, some oval-shaped +and others many-sided.</p> + +<figure class="figcenter illowp100" id="i_196" style="max-width: 62.5em;"> + <img class="w100" src="images/i_196.jpg" alt="Pollen grains, numbered 1–7. +Nr. 1 has the shape of a rounded cylinder, with small buds at the circumference in +the middle. +Nr. 2 is rounded with a skin consisting of hexagons. +Nr. 3 is perfectly round, with small circular dots on the outside. +Nr. 4 is perfectly round, with a pattern of pentagonal ribs on the surface. +Nr. 5 consists of three semi-round shapes, stuck to each other in a line. So +one is in the middle of the other two. The one in the middle is deformed into an +trapezoid shape. It contains a small dot on the surface. The whole arrangement is +not completely linear, but bent somewhat. +Nr. 6 is elongated, with a stripe lengthwise. +Nr. 7 is triangular, with little spikes on the surface."> + <figcaption> + <p>POLLEN-GRAINS.</p> + <p> + 1 <i>Morina.</i> + 2 <i>Cobea.</i> + 3 <i>Convolvulus.</i> + 4 <i>Dianthus.</i> + 5 <i>Pinus.</i><br> + 6 <i>Albucca.</i> + 7 <i>Buphthalmum.</i></p> + </figcaption> +</figure> + +<p>I have given a few examples, and would +specially call attention to the pollen-grains of +the Pinus tribe (fir-trees), to which I alluded in +the last chapter. These are remarkably buoyant, +owing to the two little bladders with which they +are furnished.</p> + +<figure class="figcenter illowp100" id="i_197" style="max-width: 62.5em;"> + <img class="w100" src="images/i_197.jpg" alt="Two drawings of a pistil, one +from the outside, one in cross section. The outside shows the pistil growing on +a stalk. At the base is a bulbous shape, somewhat elongated, followed by a +long shaft, ending in a shape consisting of 3 smaller bulbs, slightly wider +than the shaft. In cross section the shaft and the bulb at the base are hollow. +The bulb contains a small stalk with 12 circular forms along the length of it. +At the top of the shaft is a small black circular form, with a thin thread into +the hollow of the shaft that ends in one of the circular shapes at the base."> + <figcaption> + <p style="display: flex; justify-content: center;"> + <span style="text-align: left;">WHITE-LILY PISTIL.</span> + <span style="text-align: right;">SECTION OF PISTIL.</span></p> + </figcaption> +</figure> + + +<p>Now we are going to watch this yellow dust +performing its appointed office in the central +organ of a flower. In order to do so we will take +<span class="pagenum"><a id="Page_197"></a><a id="Page_198"></a>[Pg 198]</span>a white garden lily, and remove the petals, sepals, +and stamens, leaving only the pistil, which, as +shown in the drawing, consists of three parts, the +club-like stigma, a very long style, and its base the +ovary, which contains three cavities. In these +last we see a number of small, colourless spore-like +bodies termed ovules (from <i>ovum</i>, an egg), each +consisting of an outer coat, and a mass of cells in +the centre called the <i>nucellus</i>.</p> + +<figure class="figleft w20" id="i_198" style="max-width: 24.0625em;"> + <img class="w100" src="images/i_198.jpg" alt="A microscopic image of a cross- +section. It shows a round, hollow shape with thick walls and an opening at the top. +In the centre is a peanut-shaped object, with a black spot at the side directed +to the opening. A tube enters through the opening and contacts the peanut-shape +at the black spot."> + <figcaption> + <p>POLLEN TUBE.</p> + </figcaption> +</figure> + +<p>An opening exists at one end +of each ovule called the micropyle +(meaning a little gate or entrance), +and this opening leads down into +the middle of the nucellus, where +lies what we may call the life-principle, +but what is known in botany +as the embryo-sac.</p> + +<p>We need the aid of a microscope to enable us +to see how the pollen exerts its influence upon the +ovules.</p> + +<p>If we place a drop of very weak sugar and +water upon a slip of grass, and sprinkle over it +some pollen grains of the common white lily, +then allowing the slide to remain for a few +<span class="pagenum" id="Page_199">[Pg 199]</span>hours in a dark place, it will be fit for our +purpose.</p> + +<p>When placed in the microscope we shall observe +that many of the grains will have thrown out long +thread-like tubes, and this is just what happens +when pollen falls upon the viscid stigma of the +lily. Referring to the section of a lily pistil we +see that a pollen grain has rested on the stigma, +and, excited into growth by the sweetish fluid +which holds it there, it sends down a slender tube +through the centre of the pistil, which is lined with +a very delicate loose tissue of cells filled with +starch, oils and food-materials. The pollen-tube +is stimulated and fed by this nourishment stored +up in the conducting tissue, and on it goes until, +passing through the micropyle, it enters the +embryo-sac of one of the ovules, adheres to it, and +renders it fertile.</p> + +<p>Only one grain is shown in the drawing for the +sake of clearness, but of course each ovule is +sought out and fertilised by a pollen-tube. With +infinite variation this process takes place in every +flower, so that even the commonest weed affords +evidence of the marvellous provisions made by +<span class="pagenum" id="Page_200">[Pg 200]</span>an All-Wise Creator for the preservation of +species.</p> + +<p>The time occupied by the passage of the pollen-tube +varies considerably. In the fir tribe it takes +nearly twelve months, in the hazel-nut and orchis +it requires several weeks, whilst in many other +plants the whole process is completed in a few +hours.</p> + +<p>One of the first results of fertilisation is a rapid +withering of the style and flower; the great end +of the flowering period has been attained, and so +without further expense of energy the bright petals +die away.</p> + +<p>At the same time other external changes take +place, which are obvious to every observer of +nature. The lower end of the pistil, known as +the ovary, begins its second growth, and in a short +time swells into a large structure, the shape of +which varies much in different species of plants. +Finally, the ovary changes colour and develops +other characteristics quite different from its +former conditions. These characters have reference +to the distribution of its seeds, and in our +chapter on fruits we shall learn something about +<span class="pagenum" id="Page_201">[Pg 201]</span>the interesting botanical significance of the various +hard and soft fruits, and see how they all arise +from fertilisation.</p> + +<p>Take, for example, the flower of an apple +immediately after fertilisation is effected. The +petals fall off, the styles shrivel up and the ovary +rapidly enlarges; the tube of the calyx becomes +fleshy, and finally the well-formed apple is +produced. The change, however, does not end +here; in this stage of development the little apple +is bitter and is charged with a vegetable acid. As +the fruit grows on, however, this acid changes into +sweet juice varying in flavour according to the +species of apple.</p> + +<p>Now let us examine the interior of the ovary +and see what changes have arisen as a consequence +of fertilisation.</p> + +<p>The egg cell which has received the pollen +grain becomes filled with an embryo, whilst the +thin delicate coat of the ovule develops into strong +seed-coats.</p> + +<p>The embryo is the first germ of the young plant +that is to be. It is a tiny speck indeed in its +beginning, but deeply interesting to us when we +<span class="pagenum" id="Page_202">[Pg 202]</span>realise that, because it possesses life, it will grow +on and on, and result, according to its species, +either in a plant but a few inches in height, or +in a grand forest-tree which may give shelter +to man and animals for hundreds of years.</p> + +<p>The naked eye can scarcely trace any indications +of form in the embryo, but when dissected +and examined with a lens it is seen to consist of a +tiny plant, root, stem and leaves (cotyledons).</p> + +<p>The size of the embryo in comparison with the +other part of the seed is a point which should be +observed.</p> + +<p>As the embryo develops it absorbs the special +nutrient or reserve tissue that exists in all ovules; +a bean embryo, for example, rapidly absorbs all +the nucellus of the ovule, so that at length the +seed-coat<a id="tn-005"></a> contains nothing but the embryo, the +two cotyledons of which are thick and filled with +stores of food for the first growth of the seed.</p> + +<p>I would advise students to plant a few broad +beans in a little damp cocoa fibre, and carefully +watch their growth. It is advisable to dissect +these beans successively at different stages, so as +to watch the development of the radicle (root) and +<span class="pagenum" id="Page_203">[Pg 203]</span>plumule (young leaf-bud). Place the seed in what +position we may, the radicle will always find its +way down into the earth, while the plumule obeys +its vegetable instinct, and rises into the air. The +embryo of the castor-oil bean and that of the +cocoa-nut do not, however, use up all the nutritive +matter in the ovule as the broad bean does, so +that when the seed is ripe we find inside it, not +only the embryo, but also a quantity of cheesy +matter known as <i>albumen</i>, and seeds of this kind +are hence called <i>albuminous</i>, whilst peas, beans +and hazel-nuts are classed as <i>ex-albuminous</i> (without +albumen).</p> + +<figure class="figcenter illowp72" id="i_203" style="max-width: 51.9375em;"> + <img class="w100" src="images/i_203.jpg" alt="A section showing an oval form +with several layers. The outside layer is about one quarter of the total volume. +It's called 'Pericarp'. The second layer is much thinner, and very dense. +No name is attached to it. The third layer is of a lighter shade and also +about one quarter. It's called 'nucellus'. The central circle is empty. +In the third layer a small elongated shape is drawn, oriented radially, +and called 'embryo'."> + <figcaption> + <p>SECTION OF COCOA-NUT.</p> + </figcaption> +</figure> + +<p>An interesting development consequent upon +<span class="pagenum" id="Page_204">[Pg 204]</span>fertilisation is a growth which occurs in some +plants from the base of the ovule. The pretty +red coverings of the seeds of the spindle-tree, and +the bright berry-like structure on the seeds of +the yew-tree are examples of this growth, which +is known botanically as an aril (from <i>arillus</i>, a +wrapper). In the willows this aril is a very +lovely covering of silky hairs, these serve to float +the seeds on the atmosphere at every puff of +wind.</p> + +<figure class="figcenter illowp80" id="i_204" style="max-width: 62.1875em;"> + <img class="w100" src="images/i_204.jpg" alt="A small branch with 7 flowers, +some open, some still closed. Each flower sits on a stalk and is built from 4 +lobes, that open to form the calyx. Within the calyx are small petals."> + <figcaption> + <p>SPINDLE-TREE.</p> + </figcaption> +</figure> + +<p>The pretty lace-like covering on the nutmeg is +another example of an aril, better known to us in +the form of the fragrant spice called mace.</p> + +<p>The style, which in most plants dies as soon as +the ovules are fertilised, is in other cases persistent, +<span class="pagenum" id="Page_205">[Pg 205]</span>as in the hedge-climber called travellers’ joy. +The white, feathery-looking seeds owe their +special character to the persisting styles, which, +after fertilisation, grow into the bunches of fluffy +seeds, which hang in profusion on hedges in the +country.</p> + +<figure class="figcenter illowp100" id="i_205" style="max-width: 62.5em;"> + <img class="w100" src="images/i_205.jpg" alt="A short branch with 3 leaves and +a nut. The leaves are elongated with pointed ends, and sit on short stalks. The nut +also sits on a stalk and has an outer shell, which is partly opened and shows its +contents: a shape with ribs and grooves. This contents is shown separately."> + <figcaption> + <p>NUTMEG AND MACE.</p> + </figcaption> +</figure> + +<p><span class="pagenum" id="Page_206">[Pg 206]</span></p> + +<figure class="figcenter illowp85" id="i_206" style="max-width: 62.5em;"> + <img class="w100" src="images/i_206.jpg" alt="The drawing is in two parts. +The top part shows a bunch of threads at the top of a stalk. The threads +pointing in every direction. The bottom part shows a single thread, that appears +to be covered by short hairs to give it a feathery appearance. At the base of the +thread is a small globule, less than one tenth of the length of the hair."> + <figcaption> + <p>CLEMATIS OR TRAVELLERS’ JOY.</p> + </figcaption> +</figure> + +<p>I will conclude this chapter with a reference to +a change of quite a different character. Not unfrequently, +<span class="pagenum" id="Page_207">[Pg 207]</span>fertilisation results in the suppression +of certain chambers in the ovary, and in the +consequent failure of the development of the +ovules.</p> + +<p>A cross-section of a young oak ovary shows a +three-chambered structure, each cavity containing +two ovules, so that the ovary in this stage contains +six ovules in three chambers. Soon after the +act of fertilisation, one of the fertilised ovules takes +the lead in growth, starves the other five ovules, +and, as it grows, pushes the partitions of the other +chambers aside, and gradually fills up the whole +interior, converting it into a one-celled structure. +This change happens also in the birch; its two-chambered +ovary becomes one; and in the lime, +though at first it has a many-chambered ovary, +yet in the ripened fruit there is rarely more than +one to be found.</p> + +<p>In a few plants, changes of quite an opposite +character take place. In the ovary of the datura,⁠<a id="FNanchor_18_18" href="#Footnote_18_18" class="fnanchor">[18]</a> +for instance, we find two cells; after fertilisation, +two false or spurious partitions are developed, +dividing the original two-celled structure into four +<span class="pagenum" id="Page_208">[Pg 208]</span>parts, and as a consequence we get a four-chambered +fruit. The same change takes place +in some of the pea family.</p> + +<p>Specimens to be observed:—Examine pollen +grains, with lens or microscope, dissect white lily, +flower-pollen on glass slide. Observe changes +in growing apple, plant broad beans, castor-oil +seeds, and maize; examine spindle-tree berries +(<i>euonymus</i>), yew-tree berries, willow seeds, nutmeg, +and mace; travellers’ joy (clematis), section +of oak ovary in the pistillate flower. Examine +birch catkins and lime-tree flowers. Datura seed-vessel.</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_209">[Pg 209]</span></p> + + + <h2 class="nobreak" id="CHAPTER_IX"> + CHAPTER IX + <br> + <i>FRUIT</i> + + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_210">[Pg 210]</span></p> +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“Here, as I steal along the sunny wall,</div> + <div class="verse indent1">Where Autumn basks, with fruit empurpled deep,</div> + <div class="verse indent1">My pleasing theme continual prompts my thought;</div> + <div class="verse indent1">Presents the downy peach; the shining plum;</div> + <div class="verse indent1">The ruddy, fragrant nectarine; and dark,</div> + <div class="verse indent1">Beneath his ample leaf, the luscious fig.</div> + <div class="verse indent1">The vine, too, here her curling tendrils shoots,</div> + <div class="verse indent1">Hangs out her clusters, glowing to the south,</div> + <div class="verse indent1">And scarcely wishes for a warmer sky.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">James Thomson.</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_211">[Pg 211]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_IX_1"> + CHAPTER IX + <br> + <br> + <span style="font-size: 70%;">FRUIT</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_211.jpg" + style="width: 12%; height: auto;" + alt="I"> +</div> +<p class="drop-cap"><span class="upper-case">If</span> +we are shown a collection of delicious +apples, pears, grapes, peaches and +cherries, we form a very appreciative +opinion of the use and function of fruit, +but that opinion is somewhat modified +when we are shown a basket of poppy-heads, +acorns, the light downy seeds of the thistle, the +small dry carpels of the buttercup or the winged +fruits of the maple. We usually connect the term +fruit with some luscious product of the vinery or +kitchen-garden, and we may include as such the +brightly-coloured berries of the hawthorn and +wild rose, which are so conspicuous on trees and +hedges in autumn; but if we examine the subject +<span class="pagenum" id="Page_212">[Pg 212]</span>botanically we shall have to widen our ordinary +conception of the term.</p> + +<p>There is probably no part of a plant so difficult +to understand as its fruit, and this difficulty is due +to those many changes which I described in my +last chapter. A very general definition of fruit is +that it consists of the ripened ovary, and this will +be found to be correct in a great number of cases, +but this term is not exactly wide enough to +express the general formation of all fruit. In +some cases it is composed of the ripened ovary with +the parts of the stalk or the original flower, enlarged +or incorporated in the structure of the fruit, +but in other specimens we find the ovary, although +present, very little enlarged, and playing but a +minor part in the ultimate character of the mature +fruit.</p> + +<p>No fact seems so emphatic to the observant +botanist as that which upsets his artificial rules +and classifications of plants and the parts of plants. +We say, for instance, that fruit is the ripened ovary, +and yet directly we leave our books and go out to +study botany in the fields and woods, we find a +large group of fruits perfectly innocent of any such +<span class="pagenum" id="Page_213">[Pg 213]</span>structure. The firs and pines have no organ of +this kind, and yet their fruits are most important +and extremely interesting. Scarcely any part of a +plant varies so much in different species as the +fruit does. Although leaves may be found of +every size and shape, they still have some general +similarity of form, but we hasten to observe what +an immense contrast there is between the huge +<i>Musa</i> fruit (banana) and that of the oak (acorn), +although the former is, compared to the latter, +but a poor weakly plant.</p> + +<p>Again, let us note the difference between the +cocoa-nut palm fruit, a nut, which with its outer +husk is almost as large as a peck measure, and +that of the St. John’s wort or any other of our +native wild flowers.</p> + +<p>These differences in size have their counterparts +in other directions. We generally think of fruits +as being soft, luscious, and pleasant to the taste. +Many fruits of delightful colour and texture are, +however, bitter as gall, and possess highly noxious +qualities. I well remember gathering a plateful of +rich purple berries from a plant I discovered in +one of my childish rambles and carrying them +<span class="pagenum" id="Page_214">[Pg 214]</span>home as a great prize; I was not a little disappointed +when I learned that they were the +poisonous fruits of the deadly nightshade; their +deceitful resemblance to plums, as well as the +berries of the woody nightshade to red currants, +make these two of our most dangerous native +plants.</p> + +<p>As offering very distinct contrasts to the above, +we may note the dry membranous fruits of many +of our forest-trees, the hard nuts of the hazel and +walnut and the leathery husk of the chestnut. +Again, the shape of fruits is wonderfully diversified. +We have round and oval apples, plums, and +gooseberries; the linear seed-pods of the cabbage, +cauliflower, wallflower, peas and beans, and other +plants in endless varieties of forms.</p> + +<p>There are contrasts again in the smooth surface +of some fruits and the hairy coats of others where +the roughness is due to hooks, prickles and other +contrivances. How different, too, is the airy +pappus of the dandelion to those heavy fruits +which drop like stones and are to be found lying +exactly beneath the branches where they have +ripened.</p> + +<p><span class="pagenum" id="Page_215">[Pg 215]</span></p> + +<p>These differences in external form are multiplied +when we examine fruit more minutely. We shall +then find a useful dividing line in the manner in +which fruits allow their seeds to escape. In one +large division the fruit when perfectly ripe splits +open and allows the seed to fall out; such fruits +are termed dehiscent (from <i>dehisco</i>, I gape). In +the other division the fruit remains closed, and the +substance of it must decay before the seeds can +escape and grow; these are classed as indehiscent +(I gape not). Before referring to a few examples +of each division we will endeavour to distinguish +clearly the various parts of a fruit and learn their +proper botanical names.</p> + +<p>We must be careful not to confound the seed +and the coats of the ovary; the latter is termed the +pericarp (<i>peri</i>, around, <i>karpos</i>, a fruit). In some +fruits this pericarp is developed into distinct coats, +or layers. In a peach, for instance, the outer coat +is rough and hairy, this is called the epicarp (1) (<i>epi</i>, +upon, <i>karpos</i>, a fruit); the middle coat is the +succulent delicious fruit, and is known as the mesocarp +(2) (<i>mesos</i>, middle, <i>karpos</i>, fruit), whilst the inner +coat is the hard stone, or endocarp (3) (<i>endon</i>, within, +<span class="pagenum" id="Page_216">[Pg 216]</span><i>karpos</i>, fruit), and inside it lies the kernel, or true +seed. As a type of a dehiscent fruit we may +select a pea-pod; here we get no division of the +coats into distinct parts, the pericarp is dry and +tough, and when perfectly ripe it bursts open, +and allows the seeds to escape.</p> + +<figure class="figcenter illowp80" id="i_216" style="max-width: 59.5em;"> + <img class="w100" src="images/i_216.jpg" alt="A cross section of a fruit, +containing a stone. The stone has deep grooves. Parts of the fruit are +numbered. The outer surface has number 1, the part below has number 2 and +the stone number 3."> + <figcaption> + <p>SECTION OF PEACH.</p> + </figcaption> +</figure> + +<p>It would be very interesting to make a collection +of various seed-vessels, and note the immense +variety of ways in which the seeds find their way +out of the dry capsules. A poppy-head, campanula +and antirrhinum sprays, henbane, columbine, +stramonium, and many other plants afford +good examples.</p> + +<p><span class="pagenum" id="Page_217">[Pg 217]</span></p> + +<figure class="figright w30" id="i_217" style="max-width: 33.9375em;"> + <img class="w100" src="images/i_217.jpg" alt="A complex structure at the +end of a stalk. It is a bulbous shape connected to the stalk by a thin neck, +somewhat resembling an upside-down bottle. The top of it has an opening, +made by a ring of small lips around the circumference. The opening is +covered by a lid, that has a pattern of ribs radiating from its centre."> + <figcaption> + <p>POPPY CAPSULE.</p> + </figcaption> +</figure> + +<p>The woody pear is the hard fruit of a New +Holland plant which splits open to release the +seeds. The horse-chestnut is a conspicuous instance +of a dehiscing fruit, the rough prickly part is the +pericarp, and when the fruit is mature this splits +open and allows the two large chestnuts (seeds) +to escape. In the sweet-chestnut +we get an altogether +different structure. +If we pick up one of its spiny +burrs, we hold in our hand +what is called in botany an +involucre (from <i>involucrum</i>, +a cover), that is, a number +of bracts which have grown +together and formed an +outer case to the fruit. The +acorn-cup is an involucre, and +we may find other good examples in composite +flowers and those of the umbelliferæ. The small +green whorl in which a daisy-flower is set is, therefore, +not a calyx, but an involucre consisting of +minute bracts grown together. The true fruit of +the sweet-chestnut is enclosed in a mass of spiny +<span class="pagenum" id="Page_218">[Pg 218]</span>bracts, and thus differs entirely from the pericarp +of the horse-chestnut; if we wish to speak of it +correctly we must call it either a cupule or involucre. +We will now select a few examples of +fruits that are indehiscent.</p> + +<p>On the outside of an +orange we find the yellow +coat of the pericarp, +next to it is the white +mesocarp, and inside is +the juicy endocarp,⁠<a id="FNanchor_19_19" href="#Footnote_19_19" class="fnanchor">[19]</a> in +which the seeds are embedded. +When an orange +falls to the ground these +different coats simply +decay, and the seeds are +aided in their efforts to +grow by the succulent +flesh of the fruit, which affords them moisture +and nutriment. The hazel-nut is a fruit of +another texture altogether. The hard shell is the +pericarp, and the one or two seeds within it must +<span class="pagenum" id="Page_219">[Pg 219]</span>remain enclosed there until the shell decays and the +kernels can germinate and become new plants.</p> + +<figure class="figleft w30" id="i_218" style="max-width: 40.3125em;"> + <img class="w100" src="images/i_218.jpg" alt="A fruit, partly burst open, that +consists of 4 equal parts, joined at the centre. Every part has a broad base and +narrows down at the top. In the middle are some seeds."> + <figcaption> + <p>WOODY PEAR.</p> + </figcaption> +</figure> + +<p>In the currant, gooseberry, and date we find +examples of indehiscent fruits with a sweet fleshy +pericarp. In the date there is only one seed in +each fruit, and a curious thin endocarp can be +observed enveloping the solitary seed. Many +allied species, as well as the date, possess this +sweet pericarp, which must decay in order to +liberate the seeds, and in the case of succulent +fruits the process is frequently assisted by the +fruit-eating birds.</p> + +<p>It may be well to draw attention to the very +simple kind of fruit possessed by the buttercup +and other similar plants. It is a dry membranous +pericarp, and inside one seed exists free from the +pericarp; this remains closed, like other forms of +the indehiscent types, and technically this fruit is +known as an achene (from <i>achanes</i>, not gaping), +and it is well named, as it remains closed until +decay enables the growing radicle to break through +the pericarp and enter the ground. The curious +after-development of the strawberry fruit is worth +a little careful study.</p> + +<p><span class="pagenum" id="Page_220">[Pg 220]</span></p> + +<p>This flower is known as apocarpous (<i>apo</i>, apart, +<i>karpos</i>, fruit), consisting of a number of distinct +ovaries each with one ovule; these ovaries when +ripe are exactly like the achenes of the buttercup, +but they are developed upon a receptacle which, +when fertilisation has taken place, begins to dilate +and swell, with the result that the little achenes +are gradually scattered over the surface of a large +fleshy receptacle which, as it nears its time of perfection, +becomes of a most tempting crimson +colour. The little seed-like dots we notice on the +strawberry are distinct and perfect fruits embedded +in a sweet succulent floral receptacle. Thus we +find that the strawberry, speaking botanically, is +not a berry, but a collection of achenes, the term +“berry” being usually restricted to such fruits as +the currant and gooseberry. For this reason the +strawberry and the common fig are sometimes +termed spurious fruits, for in these the soft pulpy +flesh is really the receptacle and the little round +so-called seeds are the true fruit.</p> + +<p>There is a very different formation in the pineapple, +since this fruit is the development of an +entire spike of flowers; these in their early stage +<span class="pagenum" id="Page_221">[Pg 221]</span>are crowded together on the flower stalk, but as +time goes on they coalesce and fuse, with their +ovaries, bracts, and receptacles, into a succulent +mass, the various parts of which can be well defined +if we cut a section through a pineapple before it +is quite ripe.</p> + +<p>This chapter may fittingly conclude with a brief +reference to the ultimate purpose of these varied +forms and textures of fruit, for that they each have +their special work, and that there is a meaning for +every form, is a truism we may accept without +doubt. The fruit is in reality the storehouse for +the seeds, the latter being the vital part of the +plant. If we review the life-history of a plant, +first its producing flowers, then the special and +intricate processes of pollination and fertilisation, +and subsequently the growth of that wonderful +little part, the ovule, into a seed, and further if we +reflect that the whole strength of the plant has +been concentrated on producing that seed, we +shall then comprehend the true significance of +fruit.</p> + +<p>The seed is first stored up in the recesses of the +ovary; clearly then the ovary, which subsequently +<span class="pagenum" id="Page_222">[Pg 222]</span>becomes the fruit (pericarp), is intended to protect +the seeds, and it is interesting to note some of the +various ways in which this protection is afforded. +Take first the soft and sweet fruits so plentiful in +the autumn; this edible sweet flesh is not developed +until the seeds are quite ripe. All through +the period of growth and ripening the pericarp +is hard or stringy or it may be also sour or acid. +This is especially true of hedgerow fruit, such as +crab-apples, sloes, and wild pears, texture and juice +alike affording complete protection.</p> + +<p>Again, such fruit as the walnut and chestnut +are protected by their rough covering and hard +shells, and many others have their outer coats +covered with prickles and spines for the same +reason. The most extreme case is perhaps that +of <i>Mucuna pruriens</i>, a leguminous climber found +in the tropics; this has downy pods not unlike +those of a sweet-pea, and these pods are covered +with brownish hairs which, if incautiously touched, +enter the pores of the skin and cause a most +intolerable irritation; a truly formidable protection +this to the seed.</p> + +<p>Let me now point out how the seed is protected +<span class="pagenum" id="Page_223">[Pg 223]</span>in some of the pine family (firs), where there is +no pericarp. During the growth and development +of the pine seeds, the woody cone is rich in resin, +and should an enterprising nuthatch try to peck +out the seeds, he finds his beak covered with the +resin and his effort baffled.</p> + +<figure class="figcenter illowp80" id="i_223" style="max-width: 62.5em;"> + <img class="w100" src="images/i_223.jpg" alt="Two pine cones, one closed and the +other open. Between them, an enlarged seed. The closed one is somewhat oval, with +a pointed end. It is covered in diamond-shaped scales. The open one shows that these +scales are the ends of little sticks that are attached to a common centre. +By opening, these sticks are bent apart. The seed has a small, dark kernel with +a very light wing attached. The wing has one straight edge, while the other is bent, +almost part of a circle. The seed sits at the end of the wing."> + <figcaption> + <p>PINE-CONES<a id="tn-006"></a>.</p> + </figcaption> +</figure> + +<p>Protection is also afforded to the seed by the +movements of fruit after fertilisation, and of this +the cyclamen flower affords a good illustration. +As soon as fertilisation has taken place the flower +stalk coils up like a watch-spring, and the seed-pod +is thus placed safely beneath the leaves to +ripen.</p> + +<p>In crevices of old walls we may often find that +<span class="pagenum" id="Page_224">[Pg 224]</span>charming little wilding, the ivy-leaved toad-flax; +it has a highly intelligent method of protecting +its seeds. When the flower is fertilised its stalk +bends its point round to the wall, and places the +tiny ovary in a cranny of the brickwork to +mature and ripen its seeds. These are but two +instances, out of hundreds, of plants whose fruits +are protected by what we call instinctive movements.</p> + +<p>It is of essential importance to young seedlings +that they should have sufficient soil, light, and air, +to ensure their healthy growth. To begin life +directly under the leaves of the parent plant is +to court failure and starvation, and so we find +in the fruit that wonderful provisions are made +to ensure the dispersion of the seed when it leaves +the parent plant, and so endless are the contrivances +for the dispersion of fruits and seeds, +that it will be needful to devote the next chapter +entirely to that subject.</p> + +<p>Objects to collect and examine:—Compare +various fruits, fir-cone, banana, acorn, seeds, and +berries, &c. Examine a peach and pea-pod. +Collect seed-vessels, horse-chestnut, sweet-chestnut, +<span class="pagenum" id="Page_225">[Pg 225]</span>daisy-flower, orange, hazel-nut, date-fruit, strawberry, +pineapple.</p> + +<p>Observe seed coverings, pine-cones, cyclamen +stems after flowering, seed capsules of ivy-leaved +toad-flax in wall crevices.</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum"><a id="Page_226"></a><a id="Page_227"></a>[Pg 227]</span></p> + + + <h2 class="nobreak" id="CHAPTER_X"> + CHAPTER X + <br> + <i>DISPERSION OF FRUITS AND SEEDS</i> + + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_228">[Pg 228]</span></p> +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“Who gave the thistle’s feather’d seed its plumes,</div> + <div class="verse indent1">That wing-like waft it on each gentle breeze</div> + <div class="verse indent1">To sterile yet to it congenial soils,</div> + <div class="verse indent1">Investing them with purple beauty, rife</div> + <div class="verse indent1">With fragrant treasures for the wild bees’ store?”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">T. L. Meritt.</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_229">[Pg 229]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_X_1"> + CHAPTER X + <br> + <br> + <span style="font-size: 70%;">DISPERSION OF FRUITS AND SEEDS</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_229.jpg" + style="width: 12%; height: auto;" + alt="I"> +</div> +<p class="drop-cap"><span class="upper-case">I purpose</span> +in this chapter to explain +some of the many remarkable ways in +which plants are enabled to scatter +their fruits and seeds. The chief agencies +which assist in carrying out this purpose +are wind, animals, birds, running water, and +moisture in the atmosphere. We shall find that +many seeds are furnished with certain outgrowths +and peculiarities which are specially adapted to +the action of these agencies, with the result that +such seeds are distributed far and wide. We will +first examine some of those fruits which are scattered +by animals; this end is generally attained +by means of hooks and curved spines on the outside +of the fruit.</p> + +<p><span class="pagenum" id="Page_230">[Pg 230]</span></p> + +<p>Perhaps one of the most remarkable instances of +this class is the seed-pod (or capsule) of the Martynias. +During the visit of the Prince of Wales to +India, a panther killed in one of the shooting +excursions was found to have quantities of long-hooked +seeds attached to his skin: these must have +been brushed from a plant of <i>Martynia proboscidea</i>, +which has sharp curved horns three or four inches +long.</p> + +<figure class="figcenter illowp64" id="i_230" style="max-width: 52.3125em;"> + <img class="w100" src="images/i_230.jpg" alt="The seed is of an oval shape, +with 2 pronounced ribs running in the length and is is covered by numerous +little thorns. At the front end, the seed has a hook. At the rear end, 2 large +outgrows appear, about twice the size of the seed. These point upwards and at the +end are bent forward, to give them the appearance of antlers."> + <figcaption> + <p>SEED-POD OF MARTYNIA.</p> + </figcaption> +</figure> + +<p>Another species called by the Italians <i>Testa di +<span class="pagenum" id="Page_231">[Pg 231]</span>Quaglia</i>, or quail’s head, sows itself in a similar +manner by clinging to moving objects.</p> + +<p>Many common hedgerow plants have their fruits +armed with quite formidable hooks, so that creeping +or flying creatures may be made unwittingly the +means of distributing the fruits. The burdock +is a most persistent plant in this respect, each +of its numerous fruits being covered with long +hooks which successfully retain their hold of our +clothing if we happen to brush past the plant when +covered with its troublesome burrs. Other examples +are the rough seeds of the forget-me-not, +agrimony, enchanter’s nightshade—a great pest in +gardens—and all the bedstraw tribe.</p> + +<p>These plants, we may observe, are low-growing +and herbaceous, quite distinct in the matter of +position from the tall trees and shrubs which depend +upon the wind to scatter their seeds.</p> + +<p>We are all familiar with the winged fruits of the +sycamore; they are to be seen in early autumn. +The clusters are first of a pale green, and then the +seeds⁠<a id="FNanchor_20_20" href="#Footnote_20_20" class="fnanchor">[20]</a> often attain a flush of pale crimson which +<span class="pagenum" id="Page_232">[Pg 232]</span>adds much to the picturesque beauty of the tree. +The equinoctial gales separate the seeds from +their stalks, and away they go far and wide, borne +up by the delicate membrane attached to the seed +which catches the wind, and is carried by it to a +great distance from the parent tree. In the same +way the winged keys of the ash, being very light, +are borne by the autumn gales to strange habitats, +so that the tree may often be found growing on +church towers, in ruins, and on crags inaccessible +to man.</p> + +<figure class="figcenter illowp80" id="i_232" style="max-width: 55.5625em;"> + <img class="w100" src="images/i_232.jpg" alt="Two pictures of a seed, one +natural size, one magnified. The magnification shows a grain in the centre +with two wings at the sides. The grain is elliptic, with pointed ends. The +wings are rounded."> + <figcaption> + <p> + BIRCH SEED.</p> + </figcaption> +</figure> + +<p>The pinus tribe of trees have seeds with wings +slightly<a id="tn-008"></a> twisted so that, if we hold up a dry fir-cone, +the seeds descend from it with a whirling +motion like small shuttlecocks.</p> + +<p>The winds which blow strongly in mountainous +places carry these seeds before them, and are thus +ever renewing the pine-forests by sowing the products +<span class="pagenum" id="Page_233">[Pg 233]</span>of their cones on bare tracts of land. The +lightest of all tree seeds is that of the birch; it is +gifted with two wings or membranes, so that it +floats in the air before the lightest breeze, and this +may account for the wide distribution of the tree +which has been found growing from Mount Etna +to Iceland and Greenland. I may give an instance +of a common which, twenty years ago, was covered +only by furze, broom, and brake-fern; about fourteen +years since, a shower of birch seed must have +been strewn over the ground, and now it has become +a wood, shutting out the distant views and +quite altering the character of the landscape.</p> + +<p>The wind again is the agency for the dispersion +of the seeds of such plants as the common groundsel; +here it may not be uninteresting to note the +beautiful provision made in regard to the buoyancy +of the seeds. These winged structures which the +wind so lightly blows into the air must attain a +certain altitude from which they can be successfully +launched, and therefore we find that a large +class of low-growing plants have their seeds furnished +with accessories in the form of light silky +down or hairs.</p> + +<p><span class="pagenum" id="Page_234">[Pg 234]</span></p> + +<figure class="figcenter illowp59" id="i_234" style="max-width: 48.5625em;"> + <img class="w100" src="images/i_234.jpg" alt="A sail, in the shape of the +upper part of a globe. From the edge, many lines, at equal distances, hang down, +and are connected to a much smaller ring. From this ring hangs a basket, again +connected by lines and in the basket a person is sketchily drawn."> + <figcaption> + <p>PARACHUTE.</p> + </figcaption> +</figure> + +<figure class="figcenter illowp56" id="i_235" style="max-width: 47.375em;"> + <img class="w100" src="images/i_235.jpg" alt="A seed in the shape of an ellipse, +elongated and somewhat bent, with several fine ribs running in the length. It +stands with its longs axis vertically. At the top end it has some small spikes. +At the top is is connected to a thin stalk, about twice the length of the seed. +This stalk has at its top an array of hairs, almost as long as the stalk, growing +more or less horizontally, in all directions around a circle."> + <figcaption> + <p>DANDELION SEED.</p> + </figcaption> +</figure> + +<p>Most of the plants known as <i>compositæ</i> have +their seeds thus feathered, and amongst them are +those plagues of the farmer, the thistle, dandelion, +goat’s-beard and others. The dandelion may serve +as our example, and I would advise my readers to +watch carefully the variations of position in the +flowering stems. Whilst the flower is still fully +expanded the stalk remains in an upright position +so that it is conspicuous and likely to attract +the notice of insect visitors. After the florets +are fertilised it gradually lowers itself until it +<span class="pagenum" id="Page_235">[Pg 235]</span>lies on the ground under the leaves for a period +of ten or twelve days. During this time the +seed-vessel matures and ripens, then the stalk +rises to the erect position once more, and the +beautiful downy globe expands into a soft fluffy +ball of seeds hanging so loosely that the first +breeze carries them away, and their descent into +the ground is curiously provided for. Persons have +sometimes alighted on the earth from a balloon by +means of a parachute, a machine which closely +resembles an open umbrella with a car at the lower +<span class="pagenum" id="Page_236">[Pg 236]</span>end. Now the dandelion seed descends in a similar +manner, touching the ground first with its lower +end, the weight of the seed enabling it to drop into +some hole in the soil, and the spiny projections at +the upper end preventing the feathery part of the +seed from dragging it out again. The common +goat’s-beard is perhaps the most beautiful English +example of fruit with a downy pappus. A single +flower will produce a sphere as large as a cricketball, +and each seed is furnished with a starlike +<span class="pagenum" id="Page_237">[Pg 237]</span>crown of branched feathers which the wind can +bear away to a considerable distance.</p> + +<figure class="figcenter illowp63" id="i_236" style="max-width: 53.6875em;"> + <img class="w100" src="images/i_236.jpg" alt="Part of a stem, that end in a +ball. The ball is formed of some 20 little, opened up umbrellas, that through +their shafts are connected to the end of the stem. At the connection point, +each of the shafts has a thickening."> + <figcaption> + <p>GOAT’S BEARD.</p> + </figcaption> +</figure> + +<p>The handsome willow-herb, which adds so much +colour and beauty to our river banks, bears its seed +in long, narrow pods, and these, when ripe, split +up into five segments which, curling back as they +open, leave the downy seeds free to be carried off +by the passing breeze.</p> + +<p>Bird agency in seed dispersion is a most interesting +subject, and one can but admire the wonderful +way in which the services of winged creatures +are made available.</p> + +<p>Succulent berries and sticky fruits are highly +attractive to many kinds of birds, and whilst they +revel upon the sweet, soft flesh of the berry, the +seeds which they swallow with it are enabled to +resist the action of digestion by a hard covering +which protects the kernel until the shell shall +decay and allow the seed to germinate. In this +way I find my garden in early spring quite thickly +strewn with the seeds of the Irish ivy, always a +favourite food of the common wood-pigeon which +is so frequently to be heard cooing in my woods.</p> + +<p>The seeds of aquatic plants often cling to the +<span class="pagenum" id="Page_238">[Pg 238]</span>feathers of birds that visit pieces of inland water, +and are widely distributed by them in their flight +from one lake to another.</p> + +<figure class="figcenter illowp100" id="i_238" style="max-width: 62.5em;"> + <img class="w100" src="images/i_238.jpg" alt="2 ellipses of equal size, a bit +flattened lengthwise at one side and connected at this flattened side. Their +surface is striped along the length axis."> + <figcaption> + <p>COCOS-DE-MER<a id="tn-009"></a>.</p> + </figcaption> +</figure> + +<p>Darwin has shown by careful experiment that +the mud clinging to the feet of various birds almost +always contains seeds. A wounded partridge had +a ball of earth weighing six and a half ounces +adhering to its legs. From this earth Darwin +reared no less than eighty-two separate plants of +five distinct species. Seas and rivers also do their +part in dispersing seeds. The huge nuts of the +<i>Cocos-de-mer</i> palm, which grows only upon the +<span class="pagenum" id="Page_239">[Pg 239]</span>Seychelles Islands, are often thrown upon very +distant shores. This nut is said to take ten +years to come to perfection; it is exceedingly +hard, and sometimes weighs as much as forty +pounds. The common cocoa-nut is often found +growing on the shores of coral and other islands +in the Pacific Ocean, and owes its position there to +the buoyant nature of the nut, which floats uninjured +in the sea until it finds a resting-place and +a home on some atoll or island. In this way the +cocoa-palm has spread to such an extent that it is +now perhaps the only palm common to the western +and eastern hemispheres. West Indian seeds +and fruits have even been thrown upon the Norwegian +coasts, and, but for the unsuitability of the +climate, there is little doubt that tropical trees and +plants might sometimes be found growing even so +far north. It is obvious that the seeds of all +vegetation on the banks of rivers, small running +streams and lakes, must be liable to very wide +distribution. Darwin made many interesting experiments +as to the length of time seeds could +retain their vitality when floating in fresh or salt +water. Ripe hazel nuts germinated after being +<span class="pagenum" id="Page_240">[Pg 240]</span>ninety days in water. An asparagus plant with +mature berries, when dried, floated for eighty-five +days, and the seeds afterwards grew vigorously. +Out of ninety-four plants experimented upon, +eighteen floated for more than a month and some +for three months, their germinating power not +being destroyed. In quite a large number of +species the plants themselves possess the means +necessary to distribute the seed. It is true the +distance traversed by each seed may not be great, +but it is sufficient to give the seed a new field of +growth. This power varies in different species. It +is perhaps best defined as elastic force, and in the +majority of cases the seed is actually thrown away +from the parent plant by the expenditure of this +force. The seed-pod is generally in a state of +tension, due to the gradual drying up of the +tissues. Then a puff of wind, a slight blow, or +even a change in the atmospheric condition of the +air, gives the final impetus, causing the pod to +burst with such force that seeds are thrown out in +all directions. The fibro-vascular cords are often +found crossing the pod in an oblique direction, or +even in a spiral manner, so that finally, as they +<span class="pagenum" id="Page_241">[Pg 241]</span>shorten through dryness, they act upon the walls +of the legume and we see the result in such dried +pods as those of the sweet pea, broom, and +laburnum.</p> + +<figure class="figcenter illowp80" id="i_242" style="max-width: 62.5em;"> + <img class="w100" src="images/i_242.jpg" alt="A drawing of two pods. At +the bottom there is one that consists of two halves that are very elongated and +bent somewhat. One halve bends to the left, the other to the right. The one +bending to the left shows the outside of the pod, that is rather smooth and +covered with file lines. The other hale shows the pod's inside, where the seeds +are lying, 8 of them, evenly distributed along the axis of the pod. One of the +seed is beside the pod. The seeds are round, with circular lines within. The upper +drawing shows 2 pods, connected to a stem. Both pods are open showing their +two halves separated. Each of the halves is coiled up, almost like a +corkscrew. In one of the pods some seeds are shown, again rounded, with circular +lines within."> + <figcaption> + <p>BROOM AND SWEET-PEA PODS.</p> + </figcaption> +</figure> + +<p>The pansy has a three-valved seed-pod, and as +it dries the edges of the valves press upon the +polished, hard-shelled seeds and they are squirted +out with a jerk to a distance of several feet. I +was once greatly puzzled by a strange, crackling +sound in my room, and after a few minutes’ search +I discovered it was caused by a fusillade of pansy +seeds striking against the sides of a small box in +which I had placed the capsules to ripen. It is +worthy of notice that the capsule hangs down to +protect the seed-valves from rain; but when the +seeds are matured the capsule rises to an upright +position so that they may be projected far and +wide. A conspicuous example of the elastic force +of which I have spoken is seen in the British +balsam, <i>Impatiens Noli-me-tangere</i> (touch-me-not). +When its seeds are mature, the valves of the +capsule curl up in a spiral form with such force as +to project both themselves and the seeds through +the air many feet from the plant dropping the +<span class="pagenum" id="Page_242">[Pg 242]</span>seeds by the way. On a hot summer’s day one +may hear the dispersion of seeds! The furze and +broom pods, the sweet peas, and especially fir tree +cones, make quite a loud report as they split and +scatter their contents. The tension causing these +explosions is in some cases brought about by the +fluids inside the fruit. This is the case with the +squirting cucumber, which, when fully ripe, is so +<span class="pagenum" id="Page_243">[Pg 243]</span>distended with fluids that the slightest touch or +movement is sufficient to cause it to break away +from its stalk, and then the whole contents are +ejected with great force, so that the seed is thrown +some distance. The extent of dispersion is very +limited in those plants that are dependent upon +the varying moisture of the air. Such plants are +usually furnished with special awn-like⁠<a id="FNanchor_21_21" href="#Footnote_21_21" class="fnanchor">[21]</a> appendages; +these are hygroscopic⁠<a id="FNanchor_22_22" href="#Footnote_22_22" class="fnanchor">[22]</a> in their nature, and +the difference in the amount of moisture in the air +lengthens and contracts these apparently moving +organs. When the seeds fall from an ear of barley +they lie thickly strewn around the bottom of the +stem, and, were they to take root there, they must +inevitably choke each other; but each awn is +thickly set with bristles, and as the morning sun +shortens and the evening dew lengthens the hair-like +awn, the prickles only allow the awn to move +in one direction, and the seed which is attached to +it is slowly but surely drawn many inches away. +What is popularly called the dancing oat is another +curious example of this hygrometric property. If +a dry seed (or oat) is placed for a moment in +<span class="pagenum" id="Page_244">[Pg 244]</span>water, and then laid on a smooth table, it will be +seen to wave its long horns as if they were the +<span class="pagenum" id="Page_245">[Pg 245]</span>antennæ of an insect, and to turn over and over +until it has progressed some inches from the point +where it was first placed. In <i>Avena elator</i> (the tall +oat grass) and <i>Stipa pinnata</i> the awns are bent +sharply just as they emerge from the flowers, the +part below the bend being like a corkscrew and +highly sensitive to moisture, relaxing and contracting +according to the amount of moisture in +the air, with the result that the seed travels along +the ground. By the help of the long awn it can +pass over small obstacles, such as stones or clods, +the movement resembling that of a lever.</p> + +<figure class="figcenter illowp100" id="i_244" style="max-width: 62.5em;"> + <img class="w100" src="images/i_244.jpg" alt="A seed, in the shape of an +elongated ellipse, covered with hairs that all point to one end. At that end +a very long thread, about 20 times the length of the seed, is attached. The +first quarter of the thread, connected to the seed, is slightly coiled, giving +it the appearance of a corkscrew. This part is bare, while the rest of the +thread is covered in hairs, that grow perpendicular to the thread itself."> + <figcaption> + <p>STIPA PINNATA (FEATHER GRASS).</p> + </figcaption> +</figure> + +<p>I must here guard my readers against those +movements that are caused by some insect larva. +The so-called jumping bean imported from Mexico +is now so well known that it may be taken as a +type of these curious movements due, not to the +seed itself, but to the efforts of an imprisoned +insect, the grub of a small moth which passes its +larval stage inside the hard-shelled seed of a kind +of euphorbia.</p> + +<p>In conclusion, we may glance at a small group +of plants that develop sticky glands for the purposes +of dispersion.</p> + +<p><span class="pagenum" id="Page_246">[Pg 246]</span></p> + +<p>That charming Alpine plant <i>Linnæa borealis</i> +has a pair of bracts closely adherent to the fruit +and these bracts are covered with stalked glands +of a sticky nature, so that when an animal, bird, +or even a passing moth brushes against the little +fruits they stick to the intruder and are thus borne +away. Now it may perhaps occur to the thoughtful +reader that the Linnæa seed-vessel, being part +of a growing plant, would not readily break off +with a slight touch, but it is another instance of +that consummate skill and arrangement that is so +apparent to the close observer. In the stalk of the +little fruit there is a special separating layer⁠<a id="FNanchor_23_23" href="#Footnote_23_23" class="fnanchor">[23]</a> +(analogous to that of the falling leaf which we +noted in a previous chapter), and at this point the +fruit readily separates if the slightest pressure is +brought to bear upon it. This example is typical +of what takes place in such plants as <i>Salvia +glutinosa</i> and <i>Plumbago capensis</i> and <i>Rosea</i>. As a +contrast to these various modes of dispersion I +may mention those seed-vessels which are actually +buried by the plants themselves, such as the +ground-nut, ivy-leaved toad-flax<a id="tn-023"></a>, and others. We +<span class="pagenum" id="Page_247">[Pg 247]</span>must bear in mind that these plants usually have +aerial flowers in addition to those matured underground, +and that these aerial flowers produce fruits +which are subject to dispersion. We may therefore +conclude that the underground seeds are to ensure +the continuance of the plant when the ordinary +methods have perhaps partially failed. My +readers may each autumn find an endless source +of wonder and interest in the thousands of differing +fruits and seed-vessels which may be obtained +in any hedgerow and field; and by careful observation +they may yet learn many new facts and be +ever adding to their store of knowledge by gathering +and comparing the fruits and their dispersion, +as shown in the types sketched in this chapter.</p> + +<p>Objects to collect and examine:—Fruits and +seed-vessels of martynia, burdock, forget-me-not, +agrimony, enchanter’s nightshade, bedstraw, samara +of sycamore, ash-keys, pinus seeds, birch seeds, +dandelion and goat’s-beard seeds, pods of sweet-pea, +broom, laburnum, and pansy. Seed-vessels of +balsam, wild oat, feather-grass, <i>Linnæa borealis</i>, +salvia, plumbago, ground-nut, and ivy-leaved +toad-flax.</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum"><a id="Page_248"></a><a id="Page_249"></a>[Pg 249]</span></p> + + + <h2 class="nobreak" id="CHAPTER_XI"> + CHAPTER XI + <br> + <i>GERMINATION</i> + + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_250">[Pg 250]</span></p> +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“O Source unseen of life and light,</div> + <div class="verse indent1">Thy secrecy of silent might</div> + <div class="verse indent2">If we in bondage know,</div> + <div class="verse indent1">Our hearts, like seeds beneath the ground,</div> + <div class="verse indent1">By silent force of life unbound,</div> + <div class="verse indent2">Move upward from below.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">T. T. Lynch</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_251">[Pg 251]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_XI_1"> + CHAPTER XI + <br> + <br> + <span style="font-size: 70%;">GERMINATION</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_251.jpg" + style="width: 12%; height: auto;" + alt="H"> +</div> +<p class="drop-cap"><span class="upper-case">Having</span> +considered the processes which +lead up to the formation of seed, we +may now investigate the life-history of +a seed and its various forms.</p> + +<p>Like fruits, seeds differ much in their +outward shape. In size alone we find a great +contrast between the dust-like seeds of the orchids +and the huge seeds of the cocoa-nut-palm, while +between those two extremes we may note every +gradation of size. In other respects, also, the seed +offers no less variety of form and covering than +the fruit, such variations having relation to the +particular mode of dispersion and germination. +The outer skin or coat of a seed, called the <i>testa</i>, +offers a very interesting field of study, and such +seeds as the poppy and <i>silene</i> with beautiful network, +<span class="pagenum" id="Page_252">[Pg 252]</span>the <i>bignonia</i> and <i>pinus</i> with membraneous +wings, the cotton-plant seed with long hairs, and +the <i>collomia</i> with hairs that are resolved into +mucilage when wetted, are all worth special study. +When a small portion of <i>collomia</i> seed is moistened +and placed in a microscope one may see the rapid +change being effected; that which had been a hard +dry atom suddenly throws out coils of gum, like +watch springs, and a novice is led to ask, “Is the +thing alive?” so full of motion does the object +appear.</p> + +<figure class="figcenter illowp72" id="i_252" style="max-width: 51.5625em;"> + <img class="w100" src="images/i_252.jpg" alt="A shape, an almost circular +ellipse with edges that aren't completely smooth. This ellipse is lying with +it's long axis horizontally. In the middle of this ellipse, below the centre, is +a shape that consists of two ellipses joined at their long side.That shape is +surrounded by a roughly circular shape. And that shape again is surrounded +by a shape that has many rays emanating from it, thus resembling a star."> + <figcaption> + <p>BIGNONIA SEED.</p> + </figcaption> +</figure> + +<p>We may regard a seed under various aspects. +As a special means of continuing the life of a +plant, one of its modes of reproduction, as a special +means of tiding a plant over a season that would +<span class="pagenum" id="Page_253">[Pg 253]</span>be fatal to its life in its ordinary condition of +leafage, in the seed we have the germ of the future +plant, a reproduction of its parent. This germ or +embryo is lethargic or hibernating like many +animals which exist throughout the winter in a +dormant condition, yet still continue to be living +vital bodies waiting for some special influence to +come into play, and ready to resume all the activity +of a growing organism. The construction of a +seed is simple; inside the coat or <i>testa</i> we find +the embryo with or without a special supply of +albumen; if the seed is ex-albuminous, then we +may expect to meet with thick, fleshy seed-leaves +especially stored with this substance. The embryo +contains all the essential parts of the plant, the +root, stem, and leaves; the root in the seed state +is called the radicle, and is that part of the embryo +which usually points towards the micropyle; this +radicle forms one end of the first shoot which +comes out of a seed, the other end terminating in +the stem or plumule. This first shoot is known by +three names—axis, <i>tigellum</i>, or hypocotyle. The +<i>tigellum</i> in many plants gives rise to a special +structure; thus in the cyclamen it forms the tuber, +<span class="pagenum" id="Page_254">[Pg 254]</span>and the greater part of the “roots” of radishes +and turnips is due to it. In other instances it is +a mere collar forming a slightly thickened surface +between the base of the cotyledon and the radicle. +The <i>tigellum</i> is in reality a centre of growth, as +may easily be shown by cutting off an inch of the +upper part of a well-grown carrot and placing the +slice in a saucer of water; before long a crown of +young leaves will spring up and will continue to +grow and flourish as long as the plant food +contained in the slice is sufficient to maintain the +leafage. In botanical language we have thus been +growing carrot leaves from this <i>tigellum</i>.</p> + +<p>The embryo varies very much in the relative +position of its parts. Thus the embryo of the +reed-mace is straight in the <i>tigellum</i> of the embedding +albumen. In contrast to this is the curved +embryo of the deadly nightshade and the spiral +embryo of the hop.</p> + +<figure class="figright w40" id="i_255" style="max-width: 39.6875em;"> + <img class="w100" src="images/i_255.jpg" alt="A germinating seed from which +two roots grow downward and two stems upward. The stems each have two or three +oval-shaped leaves near the top."> + <figcaption> + <p>DOUBLE EMBRYO OF ORANGE.</p> + </figcaption> +</figure> + +<p>The seeds of the orange often contain two embryos, +which is rather a rare occurrence in the vegetable +world. Before we can trace the future of these +parts we must attain a clear idea of the change +the seed undergoes when it germinates. In the +<span class="pagenum" id="Page_255">[Pg 255]</span>whole of our studies our attention has been drawn +to no process so deeply interesting and yet so +mysterious as that of the breaking into life of the +seed. There are three conditions that promote the +process of germination: warmth, moisture, and +air. When these three +conditions are present and +the seed is healthy, growth +begins, and its first stage +is the absorption by the +seed of moisture; this, +combined with warmth +and the oxygen of the air, +sets up a change in the +contents of the seed. We +have already seen that +seeds are of a dry and +starchy nature, and in this +condition they are insoluble +and unfit to be active plant food. The +change that ensues results in this starchy matter +being converted into sugar which is soluble; then +the parts of the embryo begin to unfold, first the +radicle and finally the plumule are developed. +<span class="pagenum" id="Page_256">[Pg 256]</span>In this early stage these parts live entirely upon +the contents of the seed, just as a young chick is +developed and nourished upon the albumen of the +egg.</p> + +<p>The temperature requisite for germination varies +according to the species; those of us who possess +gardens know to our cost at what a low temperature +such plants as chickweed, bittercress, groundsel, +and some of the speedwells grow; as long as +the thermometer is above freezing-point these +troublesome weeds will make their appearance in +our flower borders. Sach’s experiments on germination +tend to show that wheat and barley begin +to grow below five degrees centigrade, whilst +French beans and maize germinate at nine degrees +centigrade.</p> + +<p>Some plants start into growth very quickly. +Garden cress, vegetable marrows, and some grasses +appear above ground a few days after they are +sown, whilst other seeds, enclosed in a hard, woody +seed-case, will require twelve months to germinate. +This was the case with a seed taken out of a cedar +cone brought from Mount Lebanon; I vainly +watched for the young plant, and when a year had +<span class="pagenum" id="Page_257">[Pg 257]</span>passed by the pot was thrown aside on a rubbish +heap. Shortly after I was passing by and observed +a fir-cotyledon growing on the heap, and this +proved to be the long-desired young cedar-plant.</p> + +<p>Seeds have the power to retain their vitality for +years, especially those of the <i>Leguminosæ</i>, but I +believe the stories of Egyptian mummy wheat +germinating are scarcely to be believed. A good +object-lesson upon this subject is furnished by a +newly-made railway cutting; here we may always +find growing upon the freshly-turned soil quite a +crop of plants which have sprung from seeds that +in the course of years have become embedded in +the earth, it may be at so great a depth as to +preclude the admission of air or prevent one of +the necessary conditions of germination. When, +however, the underlayer of soil is brought to the +surface and exposed to light, air, and moisture, the +seeds are able to grow.</p> + +<figure class="figcenter illowp100" id="i_258" style="max-width: 62.5em;"> + <img class="w100" src="images/i_258.jpg" alt="Two drawings of seedlings. +In both drawings a dotted line indicates the soil level. Both seedlings +have a root and a small stem with at their left and right sides a round +lobe. In the left drawing, the stem reaches above the soil level and has +some tiny leaves. In the right drawing the stem is bent at the soil level +and no leaves are visible."> + <figcaption> + <p>BROAD BEANS.</p> + </figcaption> +</figure> + +<p>To this we owe the richness of our railway-bank +flora, and many a rare plant may be discovered +there which cannot be found elsewhere in the +neighbourhood. We will now in imagination +conduct a few simple experiments that we may +<span class="pagenum" id="Page_258">[Pg 258]</span>learn something of the behaviour of seeds during +their early stages of growth. Each seed that we +thus study may be regarded by us as a type of +many others. First, then, we will sow, in a few +pots, about a dozen broad beans; before doing so +we may notice on the seed the black stripe or +ridge known as the <i>hilum</i>; this is the scar showing +where the seed was attached to the pod, and at +one end of it is the micropyle (<i>small gate</i>). If we +remove the skin of the seed we shall observe the +two fleshy cotyledons or seed leaves, a tiny point +which is the rudimentary root, and, lying close to +the inner face of the cotyledon, the slightly curved +<span class="pagenum" id="Page_259">[Pg 259]</span>plumule. After the beans had been sown a few +days and carefully watered, we may take up two +or three for examination. At first we may only +see the radicle just emerging from the little hole +at the end of the <i>hilum</i>, but if we wait, say, eight +or nine days, we shall get a further development.</p> + +<p>Before digging up our seed we will see if any +others are peeping through the soil. Yes, here is +one, just an arched kind of shoot, no leaves, only +the bow of the arch pushing up the particles of +the soil, so that the point of the shoot is clearly +still below the ground. Now, taking up a seed we +notice that the radicle has penetrated some way +down into the soil, and with a pocket lens we are +able to see a little higher than the tip of the root +quite a crop of delicate little root-hairs. The +cotyledons are still enclosed in the tough skin, but +the upward growth of the <i>tigellum</i> is acting on +them like a lever, and we can now plainly see that +it is this <i>tigellum</i> that, by its upward growth, is +penetrating the soil, and in so doing is drawing +the cotyledons from the seed coat. All this time +the delicate plumule is kept out of danger by the +arched shape of the <i>tigellum</i> and the folding of the +<span class="pagenum" id="Page_260">[Pg 260]</span>cotyledons. Leaving our seeds for a day or two +longer we find a further change. The plumule +has been carried up beyond the soil-level and has +begun to expand into leafage. It is interesting to +note how the curved <i>tigellum</i>, pushing through the +soil first, effectually guards the plumule from +injury arising from contact with rough particles +of earth; the cotyledons remain just below the +soil-level and we see that the <i>tigellum</i> is thickening +and forming a distinct connecting branch +between the new shoots and the fleshy seed leaves; +these latter are full of plant food, and the plumule +is supplied from this storehouse of nutriment until +the first leaves are formed and are able to decompose +carbon-dioxide for the nourishment of +the plantlet. The seed-leaves in this case do not +perform this function, but act simply as storehouses.</p> + +<p>Our next seed example will be the familiar +mustard plant. These we may sow in two lots, +the first we only need to sprinkle upon some fine +soil and the second may be sown in a shallow drill +and covered with fine earth.</p> + +<figure class="figcenter illowp72" id="i_261" style="max-width: 55.625em;"> + <img class="w100" src="images/i_261.jpg" alt="Three drawings of germinating +seeds, arranged from left to right. Each seed is depicted as a circle lying on +the soil. From the left one a thin root extends to the right, which bends downward +and reaches the soil. The middle one also has a root extending into the soil +and is lifted upward a bit. The right one is resting on the soil again and has a +root that extends deeper into the soil and has developed secondary roots."> + <figcaption> + <p>GROWING MUSTARD SEEDS.</p> + </figcaption> +</figure> + +<p>The first sowing will quickly germinate, and the +<span class="pagenum" id="Page_261">[Pg 261]</span>movement of the radicle which pushes out of the +micropyle may be understood by reference to the +appended diagram. In it we see the white thread-like +radicle emerging from the seed coat; it turns +very quickly towards the ground and pushes +directly into the soil. Here I must direct my +readers’ attention to one of those minute arrangements +which, though apparently insignificant +enough if we fail to study the context, is really +an evidence of the infinite perfection, care, and +wisdom of the Creator in even such a tiny detail +as the springing up of a mustard seed. As the +seed lies upon the ground, the lengthening radicle, +while it penetrates the ground, has a tendency to +force the seed into the air (as shown in the illustration), +and were it allowed to do so the seedling +would soon shrivel up and die. This catastrophe +is, however, averted by the development upon the +radicle of quite a crop of fine white root-hairs; +these adhere closely to the minute particles of the +<span class="pagenum" id="Page_262">[Pg 262]</span>soil, and are thus enabled to counteract the force +exerted by the tip of the radicle; the latter pushes +through the ground without uplifting the seed. +This action can be watched and the growth of the +root-hairs observed by means of a pocket lens and +by the exercise of that virtue, most necessary for +all young naturalists—patience.</p> + +<p>Returning to the seeds that were sown under +the soil, we find they have germinated; the radicle +is pushing downwards, and just above the soil-level +we may see the short curved <i>tigellum</i>. This +very quickly straightens itself, and then we observe +that the cotyledons have been drawn out +of the seed-coats and are displayed as two green +leaves, which in a few days will be an inch or two +above the ground, owing to the growth of the +<i>tigellum</i>. Here we get quite a departure from the +bean seed, whose cotyledons were <i>hypogean</i> (under +the earth), those of the mustard being <i>epigean</i> +(upon the earth). There is also another point of +difference; the mustard cotyledons are green, they +contain chlorophyll corpuscles, have stomates, and +so can perform all the functions of the normal +green leaf; thus they help at once to feed the +<span class="pagenum" id="Page_263">[Pg 263]</span>young plantlet by decomposing the carbon-dioxide<a id="tn-012"></a> +of the air and forming starch, whilst in contrast to +this we learnt that the seed-leaves of the bean +were storehouses only. We are now sufficiently +acquainted with the functions of the seed to be +able to appreciate the variations of the <i>testa</i>, or +seed-coat. In numerous instances the spines, +prickles, hairs, and other growths on the surface +have, in addition to their use in dispersing the +seed, an essential purpose in holding the seed in +its rightful position. We will take cress as our +next example, since it may be regarded as a type +of all smooth seeds. Cress seed remains intact +until water comes in contact with it; then it +becomes slimy by the liberation of a mucilaginous +cement from the outer coat layer; this is, of +course, highly adhesive, and thus the seeds are +fixed firmly into the soil.</p> + +<p>Another example is that of the little epiphyte +(mentioned in our first chapter), <i>Tillandsia usneoides</i><a id="tn-010"></a>, +or old man’s beard. When the seeds leave the +capsule they are furnished with silky hairs, which +enable the tiny little structures to float through the +air; they soon come in contact with the bark of +<span class="pagenum" id="Page_264">[Pg 264]</span>trees, and then the little hairs cling to the rough +surface. In this position the seeds germinate, and +are held firmly in their place by the tightly-clasping +silken strands.</p> + +<figure class="figcenter illowp72" id="i_264" style="max-width: 62.5em;"> + <img class="w100" src="images/i_264.jpg" alt="Three drawings of seedlings, +two at the top and one below. The upper left one shows a short stem growing +from the soil with two leaves of a roughly rounded, triangular shape. The leaves +are squeezed together and covered by a small cap. The upper right one shows +the same situation, now without the cap and the leaves are more expanded, +however still squeezed together. The one below shows a stem with three +pairs of leaves. The lowest pair is spread out horizontally and has a rounded +shape. The middle pair is spread out horizontally too, but their shape is more +oval with a sharp tip. They have pronounced veins in a feather-like structure. +The top pair are much smaller and are of the same shape as the middle one, +but they are growing more upward. The stem ends in a bud."> + <figcaption> + <p>BEECH COTYLEDONS.</p> + </figcaption> +</figure> + +<p>Hardly any pursuit is more delightful than the +collecting and drying of seedling trees; a ramble +<span class="pagenum" id="Page_265">[Pg 265]</span>through the woods in early summer will reveal +many specimens under or near the outskirts of the +foliage. Under the beeches we shall soon light +upon the nuts of last year coming up through the +moist, rotting soil, in the form of two broad, green +seed-leaves. As they often retain the dry, three-cornered +seed-husk upon them, we can easily see +that they are young beeches; otherwise, the +cotyledon leaves being so unlike the perfect form, +it might be rather difficult to distinguish the +species. These seedlings have germinated somewhat +like the bean seed, the radicle has grown +downward, and the curved <i>tigellum</i>, pushing upwards, +has drawn the cotyledons out of the seed-coat. +We may notice with surprise through how +small an aperture the cotyledons have been +pushed, and still they are uninjured, a fact that is +due to their being folded up like a fan in the seed-husk. +As soon as the <i>tigellum</i> reaches light and +air it straightens out, and the flat seed leaves, which +are at first of the palest green, soon deepen in +colour, and are working away preparing food for +the growth of the young plumule which springs up +from between the cotyledons, crowned with two +<span class="pagenum" id="Page_266">[Pg 266]</span>perfect young beech-leaves. This is all the baby-tree +can do the first year. We can distinguish the +second-year seedlings by their woody stem, brown +leaf-scales, and silken-fringed young beech-leaves.</p> + +<figure class="figleft w30" id="i_266" style="max-width: 32.375em;"> + <img class="w100" src="images/i_266.jpg" alt="A seed in the shape of an ellipse. +From the left-pointing end grows a root that has developed several secondary +roots. The connection between root and seed is twofold. From between these two +connections a small stem grows upward, bearing very small leaf-like structures +near its tip."> + <figcaption> + <p>ACORN.</p> + </figcaption> +</figure> + +<p>We shall not find cotyledons on the young oak, +horse-chestnut, or sweet-chestnut +seedlings, because +these remain normally below +the ground (hypogean), +forming a storehouse of +nutriment for the young +tree. It is interesting to +watch the growth of an +acorn when placed in damp +moss in a saucer. After a +few weeks the acorn will +have absorbed water, and +the leathery seed-coat will +burst at the pointed end; +through this rent the radicle will protrude, fibres +will be found growing upon the root, the +<i>tigellum</i> is thick, and just where the stalks of +the cotyledons are joined to it the plumule +emerges as from a sheath. The plumule is in +<span class="pagenum" id="Page_267">[Pg 267]</span>no hurry to develop leaves; its first growth is +provided for by the rich supply of food within the +acorn. If, however, we look carefully at its little +stem, we shall observe upon its surface a few +scattered scales, each with a rudimentary bud in +its axil. When the shoot has attained a height of +three or four inches it develops its first green leaf, +and by the end of its first summer about six will +have been formed. A collection of these seedling +trees, carefully dried⁠<a id="FNanchor_24_24" href="#Footnote_24_24" class="fnanchor">[24]</a> and neatly arranged in a +blank book, with the English and Latin names +to each, a note of the age of the seedling, the +<span class="pagenum" id="Page_268">[Pg 268]</span>spot where it was obtained, and the date, will in +time form a pleasant memento of forest rambles, +and, probably, may lead to further studies of a +similar kind.</p> + +<figure class="figcenter illowp72" id="i_267" style="max-width: 59.9375em;"> + <img class="w100" src="images/i_267.jpg" alt="A large, almost round seed. +From the bottom grows a root, undulating horizontally. It's a but longer +than the size of the seed and has developed many secondary roots, growing +in all directions. A bit below the point where the root starts from the seed, +a stems originates. This stem is about half the length of the root, grows +horizontally in the other direction and is bent upward toward its tip. At the tip +it bears a bud."> + <figcaption> + <p>HORSE-CHESTNUT.</p> + </figcaption> +</figure> + +<p>To make the collection complete there should +be some seedlings of the other great division of +plants, namely, the plants with one seed-leaf +(monocotyledons). A few date-stones will supply +these specimens; they should be sown in moist +earth and placed either in a greenhouse or on a +sunny window-ledge, where their growth can be +watched.</p> + +<p>Their germination is quite different from that of +the other seeds we have described, and if a number +of seeds are sown the different stages can be seen +as in the accompanying figure.</p> + +<p>One long cotyledon is pushed out from the seed, +the free end is like a sheath. The part nearest the +seed forms a structure resembling a rolled-up +stalk; from the former roots are developed, whilst +from the rolled-up stalk or sheath grows the next +formed leaf, and each successive leaf is sheathed +like its predecessor. This arrangement can be +well seen in young growing grasses which can be +<span class="pagenum" id="Page_269">[Pg 269]</span>taken to pieces and examined. I shall conclude +this chapter with a brief reference +to the spores or so-called seeds of +ferns and mosses.</p> + +<figure class="figright w20" id="i_269" style="max-width: 22.6875em;"> + <img class="w100" src="images/i_269.jpg" alt="A young leaf growing +from a sheath. The leaf is very elongated and has longitudinal veins. +The sheath comes out of a small rosette that is connected via a thin +stalk to a seed. The seed is ellipse shaped with pointed ends and has a +longitudinal groove. From the rosette extends a thin root downward."> + <figcaption> + <p>YOUNG DATE-PALM.</p> + </figcaption> +</figure> + +<p>These are essentially different +from the seeds that have formed +our study in the earlier part of +this chapter, they do not contain +an embryo. Let us first notice +fern-spores, which we shall find in +abundance at the back of maiden-hair +and other fern fronds; they +are contained in little brown +patches known as spore cases +(<i>sporangium</i>, from <i>spora</i>, a spore, +and <i>aggeion</i>, a vessel). If we collect +some of these and sow them on +some very fine damp earth, keeping +it at the same time shaded +and warm, the spores will soon +germinate. We shall not find a +radicle this time as the result of +growth, but in its stead a flat +expansion of green tissue (prothallium, +<span class="pagenum" id="Page_270">[Pg 270]</span>Gr. <i>protos</i>, first, <i>thallos</i>, a branch) +growing upon the earth like an exceedingly +delicate leaf. From the underside of this green +film a few very fine root-like hairs (rhizoids, Gr. +<i>rhiza</i>, a root) are developed; very soon with a +microscope we shall be able to discern upon the +surface of this structure a few little projections. +In one of these is developed a flask-shaped mass +of cells (archegonium, Gr. <i>archegonos</i>, first of a +race), in the other (antheridium, diminutive of Gr. +<i>anthera</i>, an anther) some minute bodies (antherozoides, +Gr. <i>anthera</i> and <i>zooid</i>, a minute life) with +tails; these escape from the covering and wriggle +about very much like tiny animalcules until finally +they come into contact with the flask-shaped opening +before mentioned. These tailed structures are +something like pollen grains in their function, only +they differ from pollen grains, which are passive, +by being endowed with the power of motion; the +result of their fusion with the flask-like body is to +fertilise the germ cell (oospore, Gr. <i>oon</i>, an egg) in +that structure, and from the germ cell so fertilised +is developed an embryo from which at once springs +the young fern plant. The first leaf grows from +<span class="pagenum" id="Page_271">[Pg 271]</span>the upper part of the embryo and from the lower +part is developed the “foot,” a little connecting-link +between the green prothallus and the baby +fern which serves to nurse the little plant until two +or more leaves have been produced; the roots also +grow from the same part of the embryo. I +imagine that fern spores could be grown and +watched through all their various stages even by +those of my readers who dwell in towns, as a bell +glass would maintain the requisite dampness and +shelter the young ferns from smoky air.</p> + +<p>Lastly I will describe an even simpler form of +spore development. At any season of the year we +may find the capsule fruit of mosses (Calyptra, Gr. +<i>Kaluptra</i>, a veil), a very common one being the +hair moss (Polytrichum, Gr. <i>Polutrichos</i>, having +much hair), borne upon long wiry stalks. Inside +the capsule we shall find a large quantity of small +greenish bodies; these are the spores, which of +course fall out when the spore-case is blown by the +wind, and being light are easily carried away and +at length find a resting-place in some damp nook +or shady bank. In such a place they find the +conditions necessary for their germination, which +<span class="pagenum" id="Page_272">[Pg 272]</span>is not unlike the same process in other seeds and +spores we have studied. The result is very simple. +A fine, silky, thread-like body (protonema, Gr. +<i>protos</i>, first, and <i>nema</i>, a thread) is developed; +when this has attained a fair size, a little moss +plant begins to grow upon its surface exactly as +we see a bud grow upon a tree-branch, and it is +upon this moss plant that the organs of reproduction +are produced. We have now come to the end +of our study of seeds.</p> + +<p>An endless source of interest to the student of +nature is opened up to view by carefully observing +the beginning of all vegetable life, and the seed or +spore of the commonest weed or fern will teach us +lessons that should ever make us mindful of the +wonderful mystery of life and its genesis.</p> + +<p>Objects to collect and examine:—Poppy, silene, +and collomia seeds. Examine <i>tigellum</i> of cyclamen, +radish, and carrot. Sow broad beans, mustard, and +cress seed. Collect seedling trees. Sow date-stones. +Examine fern and moss spores.</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_273">[Pg 273]</span></p> + + + <h2 class="nobreak" id="CHAPTER_XII"> + CHAPTER XII + <br> + <i>THE PHYSIOLOGY OF PLANTS</i> + + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_274">[Pg 274]</span></p> +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“Lo! on each seed, within its tender rind,</div> + <div class="verse indent1">Life’s golden threads in endless circles wind;</div> + <div class="verse indent1">Maze within maze the lucid webs are roll’d,</div> + <div class="verse indent1">And, as they burst, the living flames unfold.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="author"> + <span class="smcap">Erasmus Darwin</span>, <i>The Botanic Garden</i>. + </span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_275">[Pg 275]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_XII_1"> + CHAPTER XII + <br> + <br> + <span style="font-size: 70%;">THE PHYSIOLOGY OF PLANTS</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_275.jpg" + style="width: 12%; height: auto;" + alt="I"> +</div> +<p class="drop-cap"><span class="upper-case">In</span> +this chapter I will endeavour to +present to my readers a concise view +of the nature and method of the various +processes that go on continually in the +growing plant.</p> + +<p>These processes were incidentally referred to in +our examination of the character of the various +organs of the plant. Thus, in dealing with the +root, we spoke of its physiology so far as +concerned the absorption of water by its root-hairs. +In the leaf, we touched upon the correlation +between the shape and arrangement of the +leaf tissues and the part the leaf plays in the +economy of the plant. The physiology of the +<span class="pagenum" id="Page_276">[Pg 276]</span>reproductive organs, again, we briefly explained +in connection with their natural history.</p> + +<p>In order to arrange our studies systematically, +we may divide the physiology or function of plants +into groups, and, taking each group separately +study their effect on the plant.</p> + +<p>We may then divide the functions of plants into</p> + +<p style="margin-left: 10.0em; text-indent: 0.0em;"> + Nutrition,<br> + Assimilation, and<br> + Reproduction. +</p> + +<p>The first teaches us how a plant feeds and what +it feeds upon; the second, how the food is +prepared by the plant so as to enable it to use +this food for growth and to store some of it away +for future use. The third group deals with the +various means adopted by plants for multiplying +and increasing the species.</p> + +<p>Plants, like animals, must <i>feed</i> and <i>breathe</i> in +order to live; the food of plants, however, differs +from that of animals in being more simple and +elementary.</p> + +<p>Plant food is of two kinds, water and gas. +Water is an actual necessity to the plant, both as +a direct food and as a medium to convey inorganic +<span class="pagenum" id="Page_277">[Pg 277]</span>food. If we burn some wood to a white ash and +then analyse it, six inorganic elements will always +be found—potassium, magnesium, calcium, iron, +phosphorus, and sulphur. These substances have +been proved by experimental water-culture⁠<a id="FNanchor_25_25" href="#Footnote_25_25" class="fnanchor">[25]</a> to +be indispensable to plant-life; others are found in +larger or smaller quantities, but they are not, +judging by experimental tests, essential to plant +life. These inorganic elements do not enter the +plant as such, but in the form of salts dissolved in +water; the phosphorus and sulphur as phosphates +and sulphates. Exactly how these salts and other +elements are absorbed will be best learnt from a +simple experiment.</p> + +<p><span class="pagenum" id="Page_278">[Pg 278]</span></p> + +<p>We must first provide a large glass jar three parts +full of clear water. Then a lamp chimney, to the +bottom of which a piece of membrane (which any +butcher will supply) has been affixed, should be +partly filled with water +coloured by sulphate of copper, +and then suspended in +the glass jar. Through a cork +fitted to the top of the lamp +chimney a long tube should +be inserted. The fluid in the +lamp-glass will be seen to rise +in the tube shortly after the +experiment is made, and the +clean water in the large jar +will become slightly coloured.</p> + +<figure class="figleft w30" id="i_278" style="max-width: 30.75em;"> + <img class="w100" src="images/i_278.jpg" alt="A cylinder standing +vertically, covered with a lid. Through the lid a cork protrudes. +Through the cork extends a thin tube. At the bottom side this tube ends +in a small flask that is hanging in the cylinder. The bottom of the flask +is made of foil, fastened by a thin ring. The cylinder is filled with a +liquid up to a level reaching the neck of the flask. The flask contains +a darker liquid the extends upward through the tube, up to a level above +the lid. The upper border of that liquid is shown."> + <figcaption> + <p>TRANSFUSION DIAGRAM.</p> + </figcaption> +</figure> + +<p>This experiment teaches us +that liquids have the power of +passing through a membrane; +this power is known as diffusion, or <i>osmosis</i>. Further, +we notice that the clear fluid passes into the +coloured water more rapidly than the heavy +coloured water passes out.</p> + +<p>Now the fine <i>root hairs</i> of a growing plant are +<span class="pagenum" id="Page_279">[Pg 279]</span><i>membranes</i>, having the same property as the +membrane we placed on the lamp shade; inside +the root hairs there exists heavy dense cell sap, +outside are the films of hygroscopic water containing +(dissolved) inorganic salts, and this water +passes in through the membrane of the root, whilst +a very little of the cell sap passes out into the soil, +the quantity passing in being greatly in excess of +that which escapes.</p> + +<p>When once the crude water of the soil is inside, +it is soon passed along to the stem and leaves by +the pressure of more water coming in, and by what +is called <i>capillary power</i>, this power we may easily +see if we dip a fine tube into water, when at once +the water will rise up some distance into the tube. +I have pointed out that plant food is gaseous as +well as aqueous.</p> + +<p>Oxygen is absorbed by the root very freely from +the soil, and, therefore, farmers and gardeners +frequently plough and stir the soil of fields and +gardens so that the roots may obtain a supply of +this needful gas.</p> + +<p>Let us now endeavour to see how the gaseous +food is taken into the plant. In order to do so +<span class="pagenum" id="Page_280">[Pg 280]</span>we must remember that the gases necessary for +plant food form part of the air we breathe; this +air is made up of two-thirds nitrogen, one-third +oxygen, with a small and varying, but always +present, quantity of carbon-dioxide, and of these +the latter is the most essential to the life of +plants.</p> + +<p>We have learnt in our study of the leaf how it, +by the aid of the green chlorophyll granules, and +under the influence of sunlight, absorbs this carbon-dioxide<a id="tn-011"></a> +and effects certain changes in it. One of +the most essential elements in the growth of plants +is <i>nitrogen</i>; this we have just seen constitutes two-thirds +of the air we breathe, but the plant is unable +to make use of it in this free form; that is to say, +although the leaf can freely absorb carbon-dioxide +it cannot absorb nitrogen; it has to be taken in +by the roots of ordinary plants in the form of +nitrates, that is, in conjunction with some other +element. There is, however, an important exception +to this rule; for what are called the +insectivorous plants have the power to absorb +nitrogen under certain conditions. These will be +explained in the succeeding chapter. We can +<span class="pagenum" id="Page_281">[Pg 281]</span>now summarise the processes of nutrition. The +roots absorb water containing earthy salts as well +as oxygen gas. The leaves absorb gaseous food +in the form of carbon-dioxide, and I may add +sometimes water vapour. There are two simple +experiments that my readers can make which +will prove these statements, and will give them +a greater interest in the somewhat dry details of +vegetable physiology. Our first experiment to +show the absorptive power of roots is taken from +Sir Joseph Hooker’s Primer on Botany.</p> + +<p>“Take up three plants of the buttercup carefully +by the roots; leave one (No. 1) on the table; place +another (No. 2) with its roots in water; hang the +third (No. 3) upside down over a tumbler of water +with a few of the leaves in the water, but the root +exposed. In due time No. 1 will have faded; +No. 2 will be quite fresh; No. 3 will have the +parts not in the water faded. No. 1 shows that +water contained in the plant has evaporated from +its surface; No. 2 that the water has been absorbed +by the root and conveyed to the leaves; No. 3 +that the immersed leaves have not supplied the +other portions of the plant with water.”</p> + +<p><span class="pagenum" id="Page_282">[Pg 282]</span></p> + +<p>The second function, assimilation, depends upon +several processes that together go to make up the +work of digestion and preparing plant food. These +processes are transpiration, respiration, and evolution +of oxygen; the latter process is associated +with the feeding of the leaf—that is, the absorption +of carbon-dioxide. This compound gas is under +the influence of sunlight, and by the agency of +the green colouring granules, decomposed into +carbon-monoxide and oxygen; the latter is +eliminated, whilst the carbon and a part of the +oxygen is retained, and with the absorbed water +is converted into material that the plant can use +for the purpose of increasing its structure.</p> + +<p>By a very simple experiment we can prove the +escape of oxygen from the foliage of plants. A few +sprays of such leaves as laurustinus, bay, arbor +vitæ, and maiden-hair fern should be tied firmly +to a piece of stone. We should have ready a +soup-plate, a glass shade, and a tub full of fresh +spring water (one large enough to allow the shade +to be held upright under the water). When all is +ready, place the bunch of leaves and stone in the +glass shade held horizontally, and gradually sink +<span class="pagenum" id="Page_283">[Pg 283]</span>it under the water till the shade is quite full; +place the soup-plate at the open end where the +shade is, and slowly raise the glass until it is +upright, and then it can be lifted out and placed +on a table in a window where the sun or bright +light can reach it. The bubbles of oxygen will +soon begin to form along all the edges of the +leaves and the jewelled effect of the bouquet will +be very curious and beautiful. It is hardly needful +to say the stone is simply required to keep the +group in an upright position. By the following +day there will be a large bubble of oxygen collected +in the upper part of the shade, eliminated +from the leaves by the aid of chlorophyll and +sunlight.</p> + +<p>These changes resulting in assimilation are +always in correlation with the process known as +<i>transpiration</i>. The root is continually taking in +fluids charged with inorganic salts; these are +by the water conveyed to the leaves by means +of the network of veins, which we know by the +term fibro-vascular bundles. These, as we may +see in skeleton leaves, traverse the entire substance +of the leaves where the salts are used up in the +<span class="pagenum"><a id="Page_284"></a><a id="Page_285"></a>[Pg 285]</span>constructive work of the plant. The water is not +all wanted; part of it passes off in the form of +vapour. Transpiration, then, is the passing off +of this water.</p> + +<figure class="figcenter illowp100" id="i_284" style="max-width: 54.0625em;"> + <img class="w100" src="images/i_284.jpg" alt="A picture showing in great +detail the netted structure of the veins of a leaf. The leaf's shape is an +elongated ellipse with pointed ends. Its edges are a bit serrated. The main +vein grows from the bottom end of the leaf to the top. Secondary veins grow +at regular distances outward to the edges. This is filled up with a mesh +of smaller veins."> + <figcaption> + <p>SKELETON LEAF.</p> + </figcaption> +</figure> + +<p>We can easily see this process going on if we +place a few tropæolum leaves in a cool tumbler, +and then expose the tumbler to sunlight. In a +short time the sides of the glass will show a film +of moisture due to the transpiration of the leaves. +This process takes place more freely in a warm +temperature than in cool conditions; consequently, +in hot weather there is rapid transpiration, and as +the water is parted with more cell sap passes into +the leaves and stems, and so the plant is kept cool. +We can now see the great use of the little pores +known as <i>Stomates</i>; these are found mainly on +the under surface, and it is principally through +these pores that the leaf transpires.</p> + +<p>We must now carefully note the fact that all +growing parts of the plant take up oxygen and +give off carbon-dioxide. This power which is +common to all life is known as <i>respiration</i>. It +is a process that cannot be observed in daylight +in green plants because this respiration is feeble, +<span class="pagenum" id="Page_286">[Pg 286]</span>and also because the opposite power of assimilation +is so strong that the action of breathing is +obscured. In the absence of sunlight, however, +it can be observed, as also it may be traced in +connection with parts of the plant other than the +green leaves. Seeds, for example, during their +earlier growth (germination) give off carbon-dioxide +freely by respiration. This we can prove +for ourselves by taking a large glass jar holding +about two or three quarts; fill this about half full +of beans that have been well soaked in water so as +to swell them and induce them to commence +germination. Close the jar with a tight fitting +cork; after six or seven hours the presence of +carbon-dioxide may be easily seen. Have ready +a small phial of clear lime water, and with a piece +of twine let this down into the jar without spilling +its contents; allow it to remain there some +minutes, keeping at the same time the top closed +with a handkerchief. We shall see that the +clear lime water will after a short time become +cloudy or milky; this is due to the carbon-dioxide, +liberated by the seeds, forming chalk +with the calcium of the lime water, the chalk being +<span class="pagenum" id="Page_287">[Pg 287]</span>insoluble and easily seen. Now take out the +phial and let it stand, well covered, when the +chalk in the form of a fine precipitate will be +seen at the bottom of the phial. If desired, a +second experiment can be made with the same +jar by lowering into it a lighted taper; we shall +find it will go out owing to the presence of the +carbon-dioxide; as this gas does not support +combustion our lighted taper is quickly extinguished.</p> + +<p>We can see from these experiments that respiration +goes on in the growing plant and that this +process is independent of chlorophyll. It is an +essential part of the life of all plants, and my +readers who may perhaps wonder why it is +that two such opposite processes as I have +described are both carried on in the plant must +remember that in the main the feeding process +which depends on sunlight and the presence +of chlorophyll is carried on in the <i>daytime</i>, +whilst respiration is practically counteracted +in the daytime by the vigorous intake of carbon-dioxide. +At night when the rays of light cease +and no longer enable the plant to feed, the respiration +<span class="pagenum" id="Page_288">[Pg 288]</span>is evident. Briefly, we learn that <i>in light</i> the +plant gains in weight, whilst <i>in darkness</i> (by respiration) +it loses. The green plant can only construct +growing material out of simple substances in light, +having no power to do so in the dark.</p> + +<p>Heat is just as needful to plant-life; it must +be above freezing point, and a somewhat high +temperature is necessary to set in motion all +those chemical processes that I have briefly +described.</p> + +<p>At a low temperature the work of assimilation +and other processes are arrested; on the other +hand, a rise in temperature increases the activity +of these processes.</p> + +<p>We now come to the third function called +reproduction. We have seen in connection with +the food of plants how they convert inorganic +material into organic. This one fact is significant +of the great office of plant-life in nature; animal-life +could not exist without its help. Plant-life +may be said to prepare the food of animal-life, and +retain that balance of gases in the atmosphere +necessary to healthy respiration. How important +then it is that all kinds of herbs, trees and plants +<span class="pagenum" id="Page_289">[Pg 289]</span>should multiply and be fruitful, life of any sort is +of limited duration, and subject to all the vicissitudes +of accident, constitution, and climate, and +so we find that plants have been endowed with +wonderful powers of reproduction in order that the +earth may be constantly clothed with vegetation, +necessary for the life of man and all animal +nature.</p> + +<p>By reproduction I want my readers to clearly +understand the power possessed by the individual +plant to multiply its kind or species; and this +power is carried into effect in a variety of ways +in different species. These various methods of +reproduction then will occupy the concluding +pages of this chapter. The protoplasm (or life +principle) of any individual plant is endowed +with the power of giving rise to an entirely new +individual. This is accomplished in one of two +ways. In the first by cells forming a part of the +plant, but yet not specially modified for the purpose +of reproduction. This mode of increase is +known as vegetative reproduction. We will illustrate +it by two examples widely apart. Many +lowly plants like protococcus (the bright green +<span class="pagenum" id="Page_290">[Pg 290]</span>substance which so beautifully colours tree trunks +in moist situations) and yeast, are formed of one +cell only, and when such cells attain their full size +they simply divide into two or more cells which +grow, and finally attain maturity when the process +is repeated.</p> + +<p>The other example is that known as the strawberry +“runner,” this, as we know, is only an +elongated stem bearing at the end a bunch of +leaves, and from the base of the leaves a few roots, +the whole being a new plant which may be +removed from the parent and grown in some other +place.</p> + +<p>These, then, are examples of vegetative reproduction, +and my readers can discover for themselves +many other instances in the garden.</p> + +<p>The plan of propagation by “cuttings” is simply +the gardener’s practical application of vegetative +reproduction.</p> + +<p>The second mode of increase is by special +reproductive cells, which are set free by the parent +plants and become new individuals. The second +mode is common to all plant-life, and in it two +distinct processes can be observed. We often see +<span class="pagenum" id="Page_291">[Pg 291]</span>on a decayed pear or apple a patch of brown +mould (mucor). If we examine it with a lens we +see a little forest of tiny erect stalks, and upon the +apex of each is a round ball containing reproductive +cells, each of these, which are called spores (the +spore-case being called the sporangium), contains +protoplasm, which is endowed with the power of +giving rise to a new individual mould.</p> + +<p>This process is typical of what is common to +ferns, and many other cryptogamic plants, and is +called <i>asexual reproduction</i>.</p> + +<p>The second form is that in which two such +spore-like organs as we have noticed in the mould, +fuse together and form a spore capable of giving +rise to a new plant.</p> + +<p>This is known as <i>sexual reproduction</i>, and is +dependent upon the fact that the protoplasm of +either of the two organs is incapable of +giving rise to a new individual plant, and that +they must come in contact and fuse organically +before a new plant can be formed. This process +of fusion I have in an earlier chapter described as +fertilisation. The pollen grain, the fertilising +agent, is one of the reproductive cells, and the +<span class="pagenum" id="Page_292">[Pg 292]</span>other, the ovule, is the cell that has to be fertilised. +After this there is the subsequent development of +the ovule into the seed, and in this seed we may +recognise a plant in embryo endowed with powers +not possessed by its parent, that enables it to +resist extremes of heat and cold which would +result in many cases in death to the parent +plant. By way of experiment some seeds have +been subjected to 40 degrees of cold, and yet +have not lost their germinating power, whilst, on +the other hand, it is known that seeds of some +plants growing in sandy deserts lie baking in the +sun for many months in a temperature of over 70 +degrees, and yet begin to grow as soon as moisture +reaches them.⁠<a id="FNanchor_26_26" href="#Footnote_26_26" class="fnanchor">[26]</a></p> + +<p>Things to be observed or collected:—Experiments +to be made in order to show diffusion, +<span class="pagenum" id="Page_293">[Pg 293]</span>transpiration, and respiration, collection of +oxygen from water bouquet. Carbon-dioxide +from germinating beans. Observe—</p> + +<p style="margin-left:3.0em; text-indent: 0.0em;"> + Blue mould on fruit.<br> + Strawberry runner.<br> + Rooted cuttings.<br> + Stamens and pistil of any flowering plant. +</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum"><a id="Page_294"></a><a id="Page_295"></a>[Pg 295]</span></p> + + + <h2 class="nobreak" id="CHAPTER_XIII"> + CHAPTER XIII + <br> + <i>INSECTIVOROUS PLANTS</i> + + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_296">[Pg 296]</span></p> +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“Beyond, the moorland has its wealth</div> + <div class="verse indent2">Of pink and purple, blue and gold;</div> + <div class="verse indent1">Heather and gorse, whose breath gives health,</div> + <div class="verse indent2">And ling, a hive of bees that hold:—</div> + <div class="verse indent1">And when there’s moisture in the brake,</div> + <div class="verse indent2">The clammy sundew’s glistening glands</div> + <div class="verse indent1">’Mid carmine foliage boldly make</div> + <div class="verse indent2">Slaves of invading insect bands.”</div> + </div> + </div> +</div> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_297">[Pg 297]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_XIII_1"> + CHAPTER XIII + <br> + <br> + <span style="font-size: 70%;">INSECTIVOROUS PLANTS</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_297.jpg" + style="width: 12%; height: auto;" + alt="T"> +</div> +<p class="drop-cap"><span class="upper-case">The</span> +statement in our previous chapter +that the leaf has no power to absorb +nitrogen, has to be received with +a certain exceptions. These exceptions +are discovered in a large group of plants, +having little or no botanical relationship, +and widely separated as regards their geographical +distribution and habit of growth. The term +insectivorous (insect-eating) has been applied to +these by eminent botanists who have studied their +habits and mode of growth. We may, as a +preliminary to our study, summarise the main +features of these interesting plants, because I wish +my readers to see in them an extension and +elaboration of the various processes we have tried +<span class="pagenum" id="Page_298">[Pg 298]</span>to investigate in plant-life, and not a mere description +of a few vegetable wonders. Rather would I +point out that in studying these deviations from +the ordinary type, as elsewhere, the young botanist +should try to arrive at some explanation of these +peculiarities, bearing always in mind that every +part of the plant is created for some special purpose. +This train of thought, if brought to bear +upon our botanical study will prevent our regarding +the contrivances of these insectivorous plants +as mere freaks of nature, which appears to me +to be a low and unworthy view to take of such +delicate and wonderful structures.</p> + +<p>Occasionally, it is true, we meet with monstrosities, +in the formation of which we fail to see any +hidden purpose; but even here by careful observation +we shall probably be able to perceive that it +is the result of some injury or the accompaniment +of disease from which plant-life is no more free +than animal-life is.</p> + +<p>Let us now trace the features that are common +to the plants which form the subject of this +chapter.</p> + +<p>Perhaps their most interesting function is that +<span class="pagenum" id="Page_299">[Pg 299]</span>of catching and retaining insects. This is accomplished +in various ways, by viscid fluids which +imprison small flies, as in the leaves of the sundew +and other plants; by movements in the leaves, +as in the Venus fly-trap; by a combination of +both, as in the butterwort; or by special pitfalls +and traps, as in the pitcher plants, sarracenias, +bladderwort, and cephalotus. Having caught their +prey, these plants dissolve it by means of an acid +secretion; the dissolved animal-life is then +absorbed and appropriated for the purposes of +vegetable growth. Not all these processes are +carried on by insect-eating plants. In some, for +example, the secretion of dissolving acid is not +very apparent, in others the absorbing glands are +not fully developed; but, briefly, the above +features are those possessed by this singular class +of plants, and there is every reason to believe that +powers of this kind are more widely spread than +is usually supposed.</p> + +<p>We will now notice a few types in detail.</p> + +<p>The sundew (<i>Drosera rotundifolia</i>) is the pretty +and poetic name of a plant which may often be +found on boggy moors. It is barely an inch in +<span class="pagenum" id="Page_300">[Pg 300]</span>height, a mere rosette of leaves shaped like a +battledore, radiating from a very short root stock, +and bearing, in early summer, a central flower-stalk +from four to six inches high, furnished with +a few tiny white flowers. The whole plant lies +close to the ground, and is +often embedded in bogmoss, +and, were it not for +the bright colour of the +leaves⁠<a id="FNanchor_27_27" href="#Footnote_27_27" class="fnanchor">[27]</a> and their sparkling +dewy effect, it would be +a difficult plant to find. +With the naked eye we +can see that the leaves +are covered with hairs, and +a lens will show still more +plainly that these hairs +have each a club-like +end bearing a gummy +fluid, in appearance not unlike glycerine. These +globules of fluid sparkle in the sun; hence the +name of sundew and the botanical name of +<i>drosera</i>, from the Greek “<i>aroseros</i>,” or dewy.</p> + +<figure class="figleft w40" id="i_300" style="max-width: 37.9375em;"> + <img class="w100" src="images/i_300.jpg" alt="A plant that has a rosette +of round leaves. Each leaf is covered with long hairs. From the rosette +a short, strongly branched root grows downward en two shoots grow upward. +The shoots bear at the end a row of buds, the lowest ones have opened into +flowers on short stalks. A leaf is shown in detail. It is round with smooth edges +and has a very short stalk. The hairs are long, and have little dots at their +tips."> + <figcaption> + <p>SUNDEW.</p> + </figcaption> +</figure> + +<p><span class="pagenum" id="Page_301">[Pg 301]</span></p> + +<p>Leaves with glandular hairs are not rare +amongst our wild plants, and if this was the +only character that the sundew possessed it would +not be specially noticeable. It is, however, the +unusual structure and behaviour of these hairs +that claims our notice. The term tentacle is a +not inappropriate one to apply to these “hairs.” +A leaf of sundew, with all its tentacles standing +out at different angles from the surface of the leaf, +and each point armed with a drop of viscid fluid, +is an effective arrangement for catching insects. +The bright glistening drops are a fatal attraction +to flies, gnats, and other small insects. When they +alight upon the points of the tentacles they soon +find that they are held prisoners. In their efforts +to get free they entangle themselves more and +more on the slimy points of the treacherous hairs. +If we watch the tentacles after a fly has been +caught, it will soon be seen that the hairs are +bending over and closely pressing down the +wretched captive. This folding over occupies +four or five hours from the time the capture +is made. The glands also begin to give out an +increased amount of gummy secretion, and this +<span class="pagenum" id="Page_302">[Pg 302]</span>flow kills the insect by stopping up its breathing +pores, so that literally it dies of suffocation. The +fluid not only increases in quantity, but becomes +acid, and its effect is to dissolve the insect and +render it soluble; the dissolved parts are then +absorbed by the glands and digested. This +interesting process can be watched quite easily +by carefully taking up a few plants of sundew +with some of the bog-soil and moss in which they +were growing and placing them in a glass dish, +where they will continue for months in perfect +health if kept very wet and covered with a bell +glass.</p> + +<p>I once lighted on some magnificent sundew +growing on boggy land near Woolmer Forest. +Whilst taking up some roots of it I was persistently +attacked by a stinging fly, and, my hands +being occupied, I could not well defend myself. +Happily the sundew acted a friendly part! I was +carrying a tuft of it in my hand when, looking +down, I saw my tormenting fly was securely caught +upon its leaves. Somehow one always feels compassion +for the unfortunate, and I confess I tried +to rescue the captive, but the creature’s wings and +<span class="pagenum" id="Page_303">[Pg 303]</span>legs were already so glued together by the viscid +dew that it was impossible to release it, and I +realised more than ever how effective the sundew +is as a fly-trap.</p> + +<p>In transplanting specimens of drosera great care +should be taken that the leaves are untouched, +else, being sticky, they will cling together and lose +their delicate beauty. Every few days the plants +may be fed, and happily they are quite willing to +accept very minute pieces of raw beef, so that flies +need not be sacrificed in the cause of science. +The little “beafeater” must not be fed a second +time until the hairs have uncurled and the leaf has +fully expanded, showing that the last meal has +been digested. I have kept a large pan of sundew +in great beauty for about four months in summer, +and when the glass was taken off and bright sunshine +lit up the jewelled leaves the effect was +lovely, and a magnifying glass showed the structure +of the leaves and the prismatic colouring of +the dew-tipped hairs.</p> + +<figure class="figcenter illowp49" id="i_304" style="max-width: 48.0em;"> + <img class="w100" src="images/i_304.jpg" alt="A plant that has a rosette of +leaves. The leaves are elongated. At their end they have a special structure +built of consisting of two rounded shapes on the main vein of the leaf. These +shapes have many long spikes along the edges and e few in the central part. In +some of the leaves these shapes are folded together along the vein, whereby +the spikes along the edges intertwine. From between these folds, insects +can be seen to protrude. From the rosette grows one stalk, carrying several +flowers. Each flower has 5 separate petals and a multitude of anthers."> + <figcaption> + <p>VENUS FLY-TRAP.</p> + </figcaption> +</figure> + +<p>The Venus fly-trap is an exotic member of the +insectivorous family. Its leaves are remarkably +like an ordinary spring rat-trap. A glance at the +<span class="pagenum" id="Page_304">[Pg 304]</span>drawing will show its formation. On the two lobes +of the leaf are a row of stiff bristles occupying the +precise position of the teeth of a rat-trap. The +inner surface of the leaves is of a reddish colour, +due to its being thickly covered with minute red +glands; on each lobe there are three stiff hairs. If +a fly alighted on the leaf and walked across its +surface, it would touch one of these hairs, and no +matter how light the touch might be, the hairs are +<span class="pagenum" id="Page_305">[Pg 305]</span>so sensitive they would convey the signal to the +hinge of the lobes, and they would instantly rise +up and clasp the fly, eventually crushing it to +death. Then would follow, as in the case of the +sundew, the emission of acrid secretion and the +absorption and digestion of the insect.</p> + +<p>Insect-destroying plants are numerous in the +vegetable world. They may be roughly divided +into three groups, although there is no strict line +of demarcation between them. First, those like +the red lychnis and others, which, by means of +sticky hairs, catch and kill small insects, an +operation that, so far as we know, results in no +special good to the plant. Then there are those, +like the sundew, which catch, kill, and digest the +insect for food; whilst the third group consists of +plants which catch and kill insects, but have no +digestive process. Decomposition of the captured +insects takes place, but the absorption which goes +on is simply that of the liquid products of +decomposition, the latter process resulting from +the insects being immersed in fluid. To this +latter group belong the pitcher plants (<i>Nepenthes</i>) +and sarracenias. These last are North American +<span class="pagenum" id="Page_306">[Pg 306]</span>plants of peculiar structure and appearance. The +leaf is folded and modified into a tunnel-shaped +tube differing in form in the various species. In +all there is a kind of cap or lid to the tube, so +that rain is kept out. In one or two species the +lid is so arranged that the mouth is exposed. In +the bottom of these tubes there is usually a +quantity of somewhat slimy fluid. The inner +face of the lid and surface just inside the rim of +the tube is smooth, usually of a bright shining +<span class="pagenum" id="Page_307">[Pg 307]</span>colour and covered with minute honey-secreting +glands, a most attractive lure for insects. Below +this honeyed surface the character of the sides of +the tube changes completely; for, down to the +fluid, it is covered with stiff hairs all pointing +downwards. Now we see how the trap is set. +The honey just inside the tube is attractive, and +the insect feeding finds it very easy to descend the +tube; the smooth surface offers no foothold, and +the downward pointed hairs prevent it from returning, +until at last the insect becomes engulfed in the +pool of water at the bottom of the tube. In this +fluid insects generally accumulate, decompose, and +become liquid manure.</p> + +<figure class="figcenter illowp48" id="i_306" style="max-width: 44.375em;"> + <img class="w100" src="images/i_306.jpg" alt="From the soil emerge six +elongated beaker shaped structures with a large lip at the end. From the +midst of these beakers grows one flower on a stem of about the same height. +The stem is bent at the end, so the flower hangs downward. It has 5 hanging +petals that emerge from a calyx."> + <figcaption> + <p>SARRACENIA FLAVA.</p> + </figcaption> +</figure> + +<p>In Georgia and North Florida these sarracenias +are found in the swamps in large quantities attaining +one to two feet in height, their great tubes half-filled +with insects showing their value in tending +to reduce the swarms of flies which abound in +such localities. We can see from these characteristics +of the sarracenia a link between the +insect-eating plants which have a true digestive +process and ordinary plants that obtain their +food in part direct from the soil. The sarracenia +<span class="pagenum" id="Page_308">[Pg 308]</span>is simply making an attempt to collect nitrogenous +food by the aid of its form and sweet +secretions; thus it lures on flies and other insects +to their doom, which to the plant means an +increased supply of liquid manure for its nourishment.</p> + +<figure class="figcenter illowp42" id="i_308" style="max-width: 43.375em;"> + <img class="w100" src="images/i_308.jpg" alt="The drawing shows two parts +of a plant, not connected. In the background is an upright stem carrying 9 +buds and flowers; to the top there are buds, more downward are the op en flowers. +The flowers grow on small stalks, that emerge from a tiny leaflet. Flowers are +asymmetrical: from sideways the petals at the bottom are largest and covered by +the petal on top. The second part of the plant, in the foreground, is a stem that +starts vertically but then bends horizontal. From this point on it has many +side-branches. Every side-branch is branched as well and from these hang small +bulbs."> + <figcaption> + <p>BLADDERWORT.</p> + </figcaption> +</figure> + +<p>Between the two types of insectivorous plants +and ordinary plants there are endless varieties. +The largest known species of “fly-catcher” is the +<span class="pagenum" id="Page_309">[Pg 309]</span><i>Roridula dentata</i> of South Africa, which attains +a height of six feet, with leaves similar to the +sundew in character. So efficient are these leaves +in catching flies that the Boers hang up branches +in their rooms as fly-traps.</p> + +<p>The smallest insect-eating plant is probably +the bladderwort (<i>Utricularia vulgaris</i>), +a rootless water plant with +minute bladders on small thread-like +leaves. The bladders only +open inwards, so that when an +insect pushes against the opening +or valve it easily enters, and cannot +get out again. The bladder +contains water, but the insect +quickly consumes the oxygen in +it, and consequently dies, and +when decayed its substance is +absorbed by glands on the inner +surface of the bladder.</p> + +<figure class="figright w30" id="i_309" style="max-width: 25.6875em;"> + <img class="w100" src="images/i_309.jpg" alt="A vertical stalk from which, +at the bottom end, grows a beaker-like structure. This beaker ends in a large +lip that at some distance hangs over the opening of the beaker. The surface +of the beaker and the lip is covered in dark, irregular spots. There are two +wing-like structures on the beaker, starting at the end point of the stalk +and stretching up to the opening. These wings have many hairs along their edges."> + <figcaption> + <blockquote> + PITCHER OF NEPENTHES + RAFFLESIANA. + </blockquote> + </figcaption> +</figure> + +<p>Perhaps the most attractive of the group of +plants we are considering is the pitcher plant +or Nepenthes. It grows commonly in Borneo +and Ceylon. The pitcher is a direct development +<span class="pagenum" id="Page_310">[Pg 310]</span>of the midrib of the leaf. It varies in +size from the little thimble-like pitcher of +<i>Nepenthes gracilis</i> to the large jug-like receptacles +of <i>Nepenthes Rafflesiana</i>⁠<a id="FNanchor_28_28" href="#Footnote_28_28" class="fnanchor">[28]</a> and others, each capable +of holding nearly a pint of fluid. The pitchers +are furnished with a lid overhanging the mouth +of the receptacle, this is kept open by a thick +rim. This rim and the under-surface of the lid +both secrete a sweet fluid which is attractive to +insects, and from the rim and opening of the +mouth a smooth surface directs the ill-fated flies +to the sweet sticky fluid always found at the bottom +of the pitcher, out of which they rarely come alive.</p> + +<p>Another of our native plants exhibiting these +insectivorous habits is the butterwort (<i>Pinguicula</i>). +Like the sundew it is a mere rosette of radical +leaves, having upturned margins and a very +succulent pellucid appearance. These leaves are +covered with glands which exude a viscid kind +of fluid like that on the tentacles of the sundew. +This natural birdlime catches and holds small flies, +midges, and other tiny flying creatures, as well as +crawling insects. The presence of these insects +<span class="pagenum" id="Page_311">[Pg 311]</span>on the leaf appears to stimulate it to further +secretion which must, of course, lessen the chances +of the insect’s escape, and as a further barrier to +prevent its creeping away, the edges of the leaf +begin slowly to curve inwards, +so that the caught +insect is imprisoned in the +folds of the leaf. The acid +secretion which now exudes +from the glands soon dissolves +all the nitrogenous +and soft parts of the insect, +which are taken up by the +absorptive glands of the +leaf. There are many +other plants, of which I +have not space to make +mention, although they are +full of interest, as owing to +their curious structure, it is probable that insectivorous +habits might also be ascribed to them. +The field of study is a wide one, and throws +much light upon the physiology of plants as +well as the relationship between the plant and +<span class="pagenum" id="Page_312">[Pg 312]</span>animal world. I would suggest to my young +readers, as a practical means of knowing more +of this subject, to try and grow for themselves +the sundew, pinguicula, and sarracenia.</p> + +<figure class="figright w30" id="i_311" style="max-width: 38.0em;"> + <img class="w100" src="images/i_311.jpg" alt="A rosette of leaves, elongated +with curly edges. From the bottom of the rosette grows a short, branched root. +From the top grow three stalks, each bearing one flower or bud. With open +flowers, the tips of the stalks are bent, so the flowers hang down. The three +petals at the bottom are larger than the two on top."> + <figcaption> + <p>BUTTERWORT.</p> + </figcaption> +</figure> + +<p>The two first can be found, as I have already +said, on boggy moors in England, and the latter +plant can be obtained from any florist. All can +be successfully grown in a greenhouse or garden +frame, and studying their growth and habits in +this way will teach the young botanist far more +agreeably than learning only from books.</p> + +<p>At Kew there is always a fine collection of these +insectivorous plants to be seen in vigorous growth, +whilst at the South Kensington Natural History +Museum (Botanical Department) there are some +highly interesting cases illustrating the life history +of these remarkable plants.</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_313">[Pg 313]</span></p> + + + <h2 class="nobreak" id="CHAPTER_XIV"> + CHAPTER XIV + <br> + <i>HABIT OF GROWTH IN PLANTS</i> + + + </h2> +</div> +<div class="chapter"> +<p><span class="pagenum" id="Page_314">[Pg 314]</span></p> +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“Some clothe the soil that feeds them, far diffused</div> + <div class="verse indent1">And lowly creeping, modest and yet fair,</div> + <div class="verse indent1">Like virtue, thriving most where little seen;</div> + <div class="verse indent1">Some, more aspiring, catch the neighbour shrub</div> + <div class="verse indent1">With clasping tendrils, and invest his branch,</div> + <div class="verse indent1">Else unadorn’d, with many a gay festoon</div> + <div class="verse indent1">And fragrant chaplet, recompensing well</div> + <div class="verse indent1">The strength they borrow with the grace they lend.”</div> + </div> + </div> +</div> + +<p class="right"> + <span class="smcap author">Cowper.</span> +</p> +</div> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_315">[Pg 315]</span></p> + + + <p class="nobreak ph2" id="CHAPTER_XIV_1"> + CHAPTER XIV + <br> + <br> + <span style="font-size: 70%;">HABIT OF GROWTH IN PLANTS</span> + </p> +</div> + +<div> +<img class="drop-cap" + src="images/i_315.jpg" + style="width: 12%; height: auto;" + alt="M"> +</div> +<p class="drop-cap"><span class="upper-case">My</span> +readers have possibly noticed that in +the previous chapters my aim has been +to describe the various organs of a +plant, and that I have tried to show +not merely the botanical meaning of +the many differences in the organs of allied +species, but to point out also how these structures +are adapted to help the plant to multiply itself. +The object of this final chapter is to take a +more general view of plant-life, and to give some +idea of the different habits of plants; how in their +struggle to grow and reproduce themselves they +form such habits as tend to assist them in this +effort, and also how entirely, in some cases, they +differ from our ordinary conception of plant-growth.</p> + +<p><span class="pagenum" id="Page_316">[Pg 316]</span></p> + +<p>We have already seen how beautifully plants are +adapted to the life they have to lead, how they are +specially fitted to grow in some particular place +and climate, and now I will ask my readers to +study with me certain of the varying habits of +plant-life. A typical plant of an ordinary kind +grows, of course, in the earth, produces root, +stem, and leaves, and finally flowers, which are +the origin of fruits and seed; by the latter the +plant is again produced, and by this circular +action the continuity of that particular plant is +maintained.</p> + +<p>Let us now, in imagination, peep into a tropical +forest. On its outskirts we shall see the prototypes +of our typical plant; but inside there are also others +of quite a different aspect, and the first to attract +our attention would probably be the curious orchids +perched upon the tree-branches. Their mode of +growth differs greatly from that of a normal plant, +for they are merely attached to the branches by +means of clasping rootlets, which do not in any +way extract sap from the tree to which they are +clinging.</p> + +<p>The moisture they need is collected by the leaves +<span class="pagenum" id="Page_317">[Pg 317]</span>and hanging rootlets from the humid atmosphere +of the forest. These plants that have acquired a +perching habit sometimes grow to an immense size, +and where they do so vegetable <i>débris</i> accumulates +about their lower leaves and roots to such an +extent that it serves to supply them with needful +food.</p> + +<figure class="figcenter illowp100" id="i_317" style="max-width: 62.5em;"> + <img class="w100" src="images/i_317.jpg" alt="A rosette of leaves, irregularly +shaped, somewhat oval, that clings to a stick by a bunch of roots. From the +rosette grow 2 stalks that carry a multitude of flowers and buds. The flowers +have an intricate shape: there are five petals arranged starlike. In the centre +of that star there are two crescents in between which are pistils and anthers."> + <figcaption> + <p>PERCHING ORCHID.</p> + </figcaption> +</figure> + +<p>This habit of growth is not confined to the lovely +<span class="pagenum" id="Page_318">[Pg 318]</span>orchids; mosses, lichens, ferns, and many other +plants have acquired a similar mode of growth, +and the various ways by which they attach themselves +to the bearer plants would form an interesting +subject of investigation. It is a not uncommon +error to regard these perching plants as parasites, +but this term is properly used for plants which +actually feed upon the branches of the trees +where they grow, and of course seriously injure +the trees by so doing. The orchids, on the other +hand, do not in any way injure the branch upon +which they rest. Robert Louis Stevenson in one +of his later poems has, with a poet’s license, which +in this case is contrary to fact, described the perching +orchid thus—</p> + +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“For in the groins of branches, lo!</div> + <div class="verse indent1">The cancers of the orchid grow.”</div> + </div> + </div> +</div> + +<p>This inaccurate observation is, however, more +than atoned for by the wonderful impression +Stevenson has given us of the character of woodland +strife, the ceaseless struggle for light and air +which goes on in tropical forests.</p> + +<p>In studying the parasites as a group of plants +<span class="pagenum" id="Page_319">[Pg 319]</span>associated by the same habit of growth, we are led +to the conclusion that there is some difference after +all in the morality of plants! Here, for example, +we are confronted with a group of plants that differ +entirely from those we have hitherto examined. +The mistletoe, which is the commonest type, is +certainly lower in the social plant-scale than the +perching orchid, the latter with its leaves and +rootlets being enabled to earn its own living, +while the mistletoe sends its roots down into the +soft sap of the branch upon which it is growing +and—there is no other name for it—steals its +means of living and growing from the substance +of the poor tree upon which it preys. It is true +it does, in a half-hearted kind of way, assimilate a +little gaseous food for itself, but the sickly metallic +hue of its leaves is evidence that even in this respect +it is shirking its proper duties of nutrition.</p> + +<figure class="figcenter illowp100" id="i_320" style="max-width: 62.5em;"> + <img class="w100" src="images/i_320.jpg" alt="The base of a tree, around +which some climbing plants grow. Parts of these plants are on the soil as well. +From these parts buds and flowers grow. One flower is open. It's size is +about the same as the circumference of the tree trunk. The 5 petals are round +and of equal size. They lie flat on the soil. They are attached to a ring +in the centre. Within the ring are a multitude of small buds. The petals +are covered by small cross-like lightly coloured spots."> + <figcaption> + <p>RAFFLESIA ARNOLDII.</p> + </figcaption> +</figure> + +<p>If we desire to study the curious habits of +parasitic plants, the two examples referred to in +a previous chapter, the clover-dodder and the +yellow rattle, will afford good examples, the +latter plant being easily obtainable in fields +where the pasture is poor and scanty. Very +<span class="pagenum"><a id="Page_320"></a><a id="Page_321"></a>[Pg 321]</span>curious are the modifications and contrivances +developed by plants which have acquired this +habit of parasitism, especially amongst such weird +tropical species as <i>Rafflesia</i>, a huge parasite growing +on the <i>Cissus</i> in Sumatra. When the leaves and +flowers of the cissus have withered, then here and +there a huge knob protrudes from the stem or root, +and this grows in time to an immense stemless +flower, measuring more than three feet across, its +cup frequently containing as much as twelve pints +of liquid, and the weight of the whole flower being +said to be about fifteen pounds.</p> + +<p>Differing a little in habit from the parasites are +the <i>saprophyte</i> plants, which live on decaying vegetation. +The little brown leafless orchid called the +bird’s-nest orchis is of this character, as well as the +equally curious coral-root orchis. These plants, +as well as many other parasites, are destitute of +chlorophyll, and are therefore dependent on organic +material for food; this they obtain either as we +have seen from living plants or from decaying +organic matter. In their efforts to obtain a +needful supply of light and air, some plants +assume climbing habits, using as supports other +<span class="pagenum" id="Page_322">[Pg 322]</span>trees and plants, to the very obvious disadvantage +of the latter. We can well understand how, in a +tropical forest, the weak-climbing plants strive to +pass out of the shaded recesses and force their way +to the tops of the slower growing trees, in order to +obtain the share of light, moisture, and air which +are essential to their existence. Very vividly has +the late Mr. Louis Stevenson described such a scene +in a tropical forest—</p> + +<div class="poetry-container"> + <div class="poetry"> + <div class="stanza"> + <div class="verse indent0">“The hooked liana in his gin</div> + <div class="verse indent1">Noosed his reluctant neighbours in;</div> + <div class="verse indent1">There the green murderer throve and spread,</div> + <div class="verse indent1">Upon his smothering victims fed,</div> + <div class="verse indent1">And wantoned on his climbing coil.</div> + <div class="verse indent1">Contending roots fought for the soil</div> + <div class="verse indent1">Like frighted demons; with despair</div> + <div class="verse indent1">Competing branches pushed for air.”</div> + </div> +<hr class="poem"> + + <div class="stanza"> + <div class="verse indent0">“So hushed the woodland warfare goes</div> + <div class="verse indent1">Unceasing; and the silent foes</div> + <div class="verse indent1">Grapple and smother, strain and clasp</div> + <div class="verse indent1">Without a cry, without a gasp.”</div> + </div> + </div> +</div> + +<p>I may explain that the “murderer” alluded to +is a species of fig-tree which, in its early youth +climbs up the trunks of other trees, and by means +of its clasping roots so constricts their stems that +they ultimately perish.</p> + +<p><span class="pagenum" id="Page_323">[Pg 323]</span></p> + +<p>In pleasing contrast to this phase of vegetable +growth is the habit which indicates to us something +of mutual help and co-operation. In the <i>Compositæ</i> +we find many instances of a habit of growth that +bears distinctly upon this “help-one-another” mode +of life. A common daisy will serve as a type-flower +of this kind. The little head is a colony of +flowers, but so close is the association of its individual +florets that it is usual to regard it as one +flower rather than a distinct inflorescence composed +of numerous separate and distinct flowers.</p> + +<p>In order to understand the mutualism displayed +by this little flower, we must remember that it is +an insect-fertilised blossom, and, therefore, insects +must be attracted to it. If we carefully dissect a +flower-head we shall find first a ring of strap-shaped +flowers on the outside, constituting the ray florets—these +are imperfect;⁠<a id="FNanchor_29_29" href="#Footnote_29_29" class="fnanchor">[29]</a> but placed side by side on +the outer edge they become conspicuous; then we +find in the centre of the flower-head a number of +tiny yellow flowers, each one containing stamens +and pistils. What wee things they are, and if they +were developed singly how inconspicuous they +<span class="pagenum" id="Page_324">[Pg 324]</span>would be! When, however, they are grouped side +by side in the centre, and further, when the outer +florets are of a different colour and shape, what a +beautiful and symmetrical whole they make! Truly +this is another rendering of the maxim, Union is +strength. From a different point of view the +arrangement is equally interesting. The white and +pink tipped florets of the ray are not capable of +bearing seed, and yet we see how they help those +florets that are perfect by their attractive appearance; +then at night or on a cold rainy day these +same ray florets bend over and completely cover up +the florets in the centre which are busy producing +seed. My readers will find a rich field of investigation +open before them in studying the flowers of the +daisy family, and finding out for themselves how +the florets are grouped together, and to what extent +this principle of co-operation can be traced.⁠<a id="FNanchor_30_30" href="#Footnote_30_30" class="fnanchor">[30]</a></p> + +<figure class="figcenter illowp100" id="i_325" style="max-width: 62.5em;"> + <img class="w100" src="images/i_325.jpg" alt="Part of a garden in +which many flowers of different kinds, before a wall of shrubs and trees, grow. +In the centre is a group of high plants with large flowering umbrellas +and deeply cut leaves."> + <figcaption> + <p><span class="allsmcap">GIANT COW-PARSNIP</span> (<i>Heracleum Giganteum</i>).</p> + </figcaption> +</figure> + +<p>Students will find the corn blue-bottle especially +<span class="pagenum"><a id="Page_325"></a><a id="Page_326"></a>[Pg 326]</span>interesting; the large outer florets contain no +organs of reproduction, but still they are brightly +coloured and highly attractive to bees; the inner +florets with their protruding stigmas and anthers, +are much smaller; they are the seed-bearers, and +cannot fail to receive pollination when the bee +alights on the flower-head, allured by the showy +outer florets, which apparently exist solely that +they may draw insects to visit the unattractive +flowers of the disc.</p> + +<figure class="figcenter illowp48" id="i_326" style="max-width: 41.75em;"> + <img class="w100" src="images/i_326.jpg" alt="A bud and a flower. The +bud is oval and covered in scales. From the top a group of small petals +emerges. The flower shows this group of petals further developed. They +form a bunch of little flowers of their own. Each of these has six petals +that are arranged in a star shape. These flowers are arranged in a circle. +In the centre of this circle is a group of small pistils and anthers."> + <figcaption> + <p>CORN BLUE-BOTTLE.</p> + </figcaption> +</figure> + +<p>The direct influence of the separate parts of a +<span class="pagenum" id="Page_327">[Pg 327]</span>plant upon one another, and the very distinct habit +of associating together that they may attain some +end such as the visits of insects, leads us to consider +two other aspects of plant-life, both of which +are so full of interest that no botanical work can +now be considered complete without some reference +to the matter. If we carefully dig up a clover +plant or a broad bean and examine the little rootlets +we shall observe some small knobs or swellings +upon them. These swellings are only found here +and there on some of the roots, so that their +presence is not a normal condition. Placing one of +these knobs under a powerful microscope, we shall +find it to be not ordinary root tissue but a substance +teeming with countless numbers of rod-like or +rounded atoms which botanists who have investigated +the subject tell us are bacteria, <i>i.e.</i>, inconceivably +small one-celled plants which are often +the cause of terrible diseases. But some of these +mysterious organisms, on the other hand, are +capable of beneficial results. It has of late been +clearly proved that leguminous plants having these +colonies of bacteria on their roots possess the +power of assimilating the free nitrogen that forms +<span class="pagenum" id="Page_328">[Pg 328]</span>such a large proportion of atmospheric air. When +therefore a farmer sows his wheat in a field previously +occupied by clover he finds the clover roots +left in the soil contribute the best possible supply +of nitrogen to the wheat crop. This seems a +remarkable fact, since vegetable physiologists have +hitherto insisted upon the fact that plant-life is +unable to make use of the free nitrogen of the air. +The other instance of strange habit is that of a +symbiosis,⁠<a id="FNanchor_31_31" href="#Footnote_31_31" class="fnanchor">[31]</a> which exists between certain trees on +the one hand and the threads of spawn of some +fungi on the other. If the roots of the white +poplar are examined minutely, quite a mantle of +whitish threads will be found covering the growing +point. It is said by that eminent botanist, Professor +Kerner, and by others that, as the roots are +developed from the young seedling-tree, they are +enclosed in the meshes of the fungus, and that this +particular fungus is always a close associate of the +roots as they grow in all directions. This fact we +can see when we dig up the roots, but the most +striking part of the story is this, that between this +<span class="pagenum" id="Page_329">[Pg 329]</span>fungus root and the roots of the tree there is an +organic connection, a division of labour which +results in the tree receiving from the thread-like +filaments of the fungus (<i>hyphæ</i>) both moisture and +certain food stuffs from the ground, whilst the +fungus gets in return such organic food as the tree +has produced by means of its green leaves. Such +cases as these present to us a manner of growth +that is akin to social habit, and, strange as the +union may appear, the circumstance is by no means +uncommon in the vegetable kingdom. Stranger +still perhaps is the union that is sometimes to be +found between plants and some member of the +animal world, of which union I shall give an +example. On one of the larger species of sea-anemones +(<i>Anthea cereus</i>) are small yellowish spots, +which at one time were supposed to form part of +the animal itself. But now the spots turn out to +be vegetable cells, which can be isolated and +induced to continue growing after the death of +the anemone. The yellow spots are small algæ, +and are furnished with chlorophyll. We must not +regard the algæ as parasites on the sea-anemone, +because they split up the carbon-dioxide under the +<span class="pagenum" id="Page_330">[Pg 330]</span>influence of sunlight, and by so doing supply the +anemone with oxygen for respiration, whilst the +starch formed in the protoplasm of the algæ passes +by diffusion into the anatomy of the animal. The +transaction does not end here; the algæ in all +probability receives nitrogenous substances in +return, so that there is a mutual interchange.</p> + +<p>These are but one or two of the many wonderful +phases of vegetable life, and I hope by thus briefly +sketching a few of them my readers will be +stimulated into a greater desire to explore God’s +marvellous works in nature. There is an endless +succession of such wonders to be investigated, but +in order to find them we need a careful spirit of +observation, passing nothing by without trying to +learn something of its life history. Every hedgerow +is full of delightful problems which will reward the +interested student. A single field has been found +to contain as many as fifty different species of +plants, and every month of the year will present +a new aspect of life. In the early spring we have +the germinating seed and the tiny growing moss. +A little later the opening buds with their wealth of +interesting points to study, then the unfolding of +<span class="pagenum" id="Page_331">[Pg 331]</span>the leaves and the gradual development of the +flower. Here and there a climbing plant will +engage our attention, its mode of climbing, its +modification of part or parts to enable it successfully +to overcome difficulties, its acceptance of help +by the way—as in the case of a bryony tendril I +once came across which cleverly attached itself to +a minute hole in a laurel leaf—these and many +<span class="pagenum" id="Page_332">[Pg 332]</span>other items will interest us in our walks if we keep +our eyes open.</p> + +<figure class="figcenter illowp45" id="i_331" style="max-width: 45.4375em;"> + <img class="w100" src="images/i_331.jpg" alt="A small piece of a branch +from which grows a stalk sideways ending in the remains of a flower. In the +axil of this stalk sits a small bud. From the same point on the branch emerges +a thin thread, heavily coiled. This thread has grown through a hole in a leaf, +also shown in the drawing. The leaf is elongated, somewhat oval, with lightly +serrated edges."> + <figcaption> + <p>BRYONY TENDRIL.</p> + </figcaption> +</figure> + +<p>Then, as summer slowly passes away and autumn +approaches, the fruits will engage our attention; +their forms and shapes and modes of dispersion +will afford ample subjects for study.</p> + +<figure class="figcenter illowp64" id="i_332" style="max-width: 49.0625em;"> + <img class="w100" src="images/i_332.jpg" alt="On a stick sits a group of +about 10 shapes in the form of calyxes. From the central shape emerges +a bundle of bent hairs and several small dots."> + <figcaption> + <p>TRICHIA THROWING OUT SPORES.</p> + </figcaption> +</figure> + +<p>Winter, too, still brings its store of pleasure for +the young botanist. Nature is not dead—she only +sleeps. Nay, unless there is hard frost and deep +snow the field for observation is just as wide and +the harvest as plentiful as at any other season. +Look on the old apple-trees and see what a host +of tiny plantlets there is there to glean. Here are +pale-green bearded moss and lichens, there a branch, +perhaps, lies on the ground dead and decaying, +under whose mouldering bark, if we have keen +<span class="pagenum" id="Page_333">[Pg 333]</span>eyes, we may discover tiny tufts of the <i>Mycetozoa</i>, +whose capsules, under the microscope (and in some +cases even with the naked eye) are seen to give off +clouds of spores, actually thrown out by the active +movements of fine waving threads, a sight never +to be forgotten when it has been watched under +favourable circumstances. Winter is also rich in +its harvest of mushroom-like fungi; these will well +repay a little study. We shall be led to note their +form, colour, mode, and habit of growth, how they +affect certain trees and soils, and the important +difference of some kinds being eatable and others +virulently poisonous; the mere book student can +know very little of the keen pleasure enjoyed by +those who thus think about what they see, and are +ever adding to their stock of knowledge by personal +observation. I may close with some true +and beautiful thoughts by one⁠<a id="FNanchor_32_32" href="#Footnote_32_32" class="fnanchor">[32]</a> who is herself a +reverent student of the book of nature.</p> + +<p>“No pleasure is more sure and none less costly +than that of watching day by day the signs of the +coming spring; than the delight of seeing unexpectedly +<span class="pagenum" id="Page_334">[Pg 334]</span>the first primrose, and of finding that the +anemones and hyacinths are pushing their way to +the sunshine. Year by year the miracle of springtime, +when the green leaves are shaken forth from +the hard bud is more miraculous. Summer after +summer the lilies are fairer, the wild roses more +exquisite, and on through the seasons the varying +pleasures succeed one another. These things never +pall; and if the time should come when we can no +longer go out to the hills and woods to welcome +the spring and revel in the bounty of summer we +know that the past is not lost. The fair remembrance +of the flowers of the field is safe in our +hearts, and will ‘flash upon that inward eye which +is the bliss of solitude.’”</p> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + +<p><span class="pagenum" id="Page_335">[Pg 335]</span></p> + + + <h2 class="nobreak" id="GLOSSARY_OF_SCIENTIFIC_WORDS"> + <i>GLOSSARY OF SCIENTIFIC WORDS + USED IN THIS VOLUME</i> + + </h2> +</div> + + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> +<p><span class="pagenum"><a id="Page_336"></a><a id="Page_337"></a>[Pg 337]</span></p> + + <p class="nobreak ph2" id="GLOSSARY"> + GLOSSARY. + + </p> +</div> + + +<p><span class="smcap">A clear</span> definition of scientific terms involves an +exact knowledge of several languages, and when +translated into technical phraseology these definitions<a id="tn-013"></a> +often appear to me to be as difficult to a +simple comprehension as the original words they +purport to explain.</p> + +<p>I have endeavoured therefore, in this glossary, +to put scientific terms into plain words as clearly +as was consistent with the facts, and not by any +means to attempt a really exhaustive scientific +definition.</p> + + +<h3>A</h3> + +<p class="glossary"><i>Absciss</i>—A term applied to a layer of separating cells.</p> + +<p class="glossary"><i>Absorption</i>—Taking in food by diffusion.</p> + +<p class="glossary"><i>Accessory</i>—Anything additional.</p> + +<p class="glossary"><i>Acetic</i>—Applied to an acid, sour.</p> + +<p class="glossary"><i>Achene</i>—A small dry indehiscent fruit with a leathery coat.</p> + +<p class="glossary"><i>Adaptation</i>—As applied to plant-life meaning the structure +of the plant becoming most fitted to its environment.</p> + +<p class="glossary"><i>Adventitious</i>—Not developed in regular order.</p> + +<p class="glossary"><span class="pagenum" id="Page_338">[Pg 338]</span></p> + +<p class="glossary"><i>Aerial</i>—Inhabiting or existing in the air.</p> + +<p class="glossary"><i>Æstivation</i>—The arrangement of the parts of the flower in +the bud.</p> + +<p class="glossary"><i>Albumen</i>—Reserve material contained in the seed, analogous +to the white of an egg.</p> + +<p class="glossary"><i>Alchemilla</i>—A genus of rosaceous plants with small green +flowers.</p> + +<p class="glossary"><i>Allium</i>—The onion genus.</p> + +<p class="glossary"><i>Altitude</i>—Height.</p> + +<p class="glossary"><i>Ampelopsis</i>—A genus of climbing plants allied to the vine +whose leaves are brilliantly coloured in autumn.</p> + +<p class="glossary"><i>Anemophilous</i>—Pollinated by the wind.</p> + +<p class="glossary"><i>Animalcule</i>—Microscopic insect life.</p> + +<p class="glossary"><i>Annual</i>—A plant whose duration of life is one season: +<i>Ex.</i> mignonette.</p> + +<p class="glossary"><i>Anthea</i>—A genus of sea-anemones.</p> + +<p class="glossary"><i>Anther</i>—The dilated end of the stamen in which the pollen +grains are developed.</p> + +<p class="glossary"><i>Antheridium</i>—The case containing the antherozoids in +cryptogamic plants.</p> + +<p class="glossary"><i>Antherozoides</i>—The male cell, or active member in fertilisation +of cryptogams.</p> + +<p class="glossary"><i>Antirrhinum</i>—The snap-dragon genus.</p> + +<p class="glossary"><i>Antiseptic</i>—Counteracting decay or putrefaction.</p> + +<p class="glossary"><i>Apocarpous</i>—Applied to the pistil when the carpels are +distinct or when the pistil consists of one carpel.</p> + +<p class="glossary"><i>Appendages</i>—Something hanging or appended, extra.</p> + +<p class="glossary"><i>Aquatic</i>—Relating to water.</p> + +<p class="glossary"><i>Araucaria</i>—The generic name of the monkey-puzzle tree.</p> + +<p class="glossary"><i>Archegonium</i>—The flask-shaped organ containing the female +cell in the cryptogams.</p> + +<p class="glossary"><i>Arid</i>—Dry and waterless.</p> + +<p class="glossary"><i>Arillus</i>—An out-growth from the funicle (or seed-coat).</p> + +<p class="glossary"><i>Aristolochia</i>—A genus of climbing plants with curious +“prison” flowers which attract and retain insects.</p> + +<p class="glossary"><span class="pagenum" id="Page_339">[Pg 339]</span></p> + +<p class="glossary"><i>Arum</i>—A genus of poisonous plants with an inflorescence +consisting of spadix and spathe.</p> + +<p class="glossary"><i>Asexual</i>—Not sexual.</p> + +<p class="glossary"><i>Asparagus</i>—A genus of edible vegetables and climbing +plants.</p> + +<p class="glossary"><i>Assimilation</i>—The conversion of crude food into protoplasm.</p> + +<p class="glossary"><i>Aster</i>—The generic name of the Michaelmas daisies.</p> + +<p class="glossary"><i>Avena</i>—The generic name of the oat.</p> + +<p class="glossary"><i>Awn</i>—The beard of barley and other corn.</p> + +<p class="glossary"><i>Axillary</i>—Growing in the axil of the leaf.</p> + + +<h3>B</h3> + +<p class="glossary"><i>Bacteria</i>—Minute one-celled living atoms, the cause of most +contagious diseases.</p> + +<p class="glossary"><i>Bamboo</i>—A giant grass.</p> + +<p class="glossary"><i>Banana</i>—The fruit of the genus Musa.</p> + +<p class="glossary"><i>Bark</i>—The rough external part of a stem.</p> + +<p class="glossary"><i>Barm</i>—Same as yeast.</p> + +<p class="glossary"><i>Bast</i>—The fibrous tissue between the bark and the wood of +a dicotyledonous stem.</p> + +<p class="glossary"><i>Begonia</i>—A genus of plants with bright flowers and oblique +or one-sided leaves.</p> + +<p class="glossary"><i>Betula</i>—Generic name of the birch-tree.</p> + +<p class="glossary"><i>Biennial</i>—A plant whose duration of life is two seasons: +<i>Ex.</i> Beetroot.</p> + +<p class="glossary"><i>Bifacial</i>—With upper and lower sides structurally different: +<i>Ex.</i> laurel leaf.</p> + +<p class="glossary"><i>Bignonia</i>—A genus of flowering climbing plants.</p> + +<p class="glossary"><i>Blade</i>—The broad part of the leaf.</p> + +<p class="glossary"><i>Bougainvillia</i>—A genus of climbing tropical plants with +bright pink bracts and small yellowish flowers.</p> + +<p class="glossary"><i>Bulb</i>—A dormant bud surrounded with fleshy scales.</p> + +<p class="glossary"><i>Bulbils</i>—Small bulbs.</p> + +<p class="glossary"><span class="pagenum" id="Page_340">[Pg 340]</span></p> + +<p class="glossary"><i>Bunium</i>—A genus of tuberous Umbelliferæ, earthnut.<a id="tn-014"></a></p> + +<p class="glossary"><i>Buoyant</i>—Light, able to float in air or water.</p> + +<p class="glossary"><i>Button-wood</i>—A term applied in America to the plane +tree.</p> + + +<h3>C</h3> + +<p class="glossary"><i>Cacti</i>—A family of succulent plants usually devoid of leaves.</p> + +<p class="glossary"><i>Caducous</i>—Quickly dropping off.</p> + +<p class="glossary"><i>Calceolaria</i>—A genus of herbaceous garden plants with +pouched flowers.</p> + +<p class="glossary"><i>Calcium</i>—An element present in all calcareous rocks.</p> + +<p class="glossary"><i>Calyptra</i>—The hood of a moss-fruit.</p> + +<p class="glossary"><i>Calyx</i>—The outer whorl of the flower or floral envelope, +cup-shaped.</p> + +<p class="glossary"><i>Cambium-layer</i>—A layer of active growing tissue.</p> + +<p class="glossary"><i>Campanula</i>—A genus of Alpine and herbaceous plants with +bell-shaped flowers.</p> + +<p class="glossary"><i>Camphor</i>—A drug obtained by dry distillation of the leaves +and stems of Camphora officinarum.</p> + +<p class="glossary"><i>Capillary</i>—Fine and minute, hair-like.</p> + +<p class="glossary"><i>Carbon-dioxide</i>—Symbol CO<sub>2</sub>. A gas existing in small quantities +in the air, otherwise called carbonic-acid gas.</p> + +<p class="glossary"><i>Carbon-monoxide</i>—A poisonous gas whose molecule is composed +of one atom of carbon and one atom of oxygen.</p> + +<p class="glossary"><i>Carex</i>—A genus of sedge-like plants.</p> + +<p class="glossary"><i>Carpel</i>—A pistillate leaf, one of the component parts of the +pistil.</p> + +<p class="glossary"><i>Caterpillar</i>—The form of an insect after it is hatched, first +stage.</p> + +<p class="glossary"><i>Catkin</i>—A spike of staminate or pistillate flowers usually +pendulous.</p> + +<p class="glossary"><i>Checkered</i>—Outlined into a square-like pattern.</p> + +<p class="glossary"><i>Chevaux de frise</i>—An obstacle consisting of iron spikes set +in a framework of iron.</p> + +<p class="glossary"><i>Chlorophyll</i>—The green colouring matter of leaves and stems.</p> + +<p class="glossary"><i>Cholera</i>—A contagious disease.</p> + +<p class="glossary"><span class="pagenum" id="Page_341">[Pg 341]</span></p> + +<p class="glossary"><i>Chrysalis</i>—<i>pl.</i> Chrysalides. The form assumed by some +insects before they reach the winged state.</p> + +<p class="glossary"><i>Chrysanthemum</i>—A genus of showy flowering plants belonging +to the Compositæ.</p> + +<p class="glossary"><i>Cinchona</i>—A genus of trees yielding quinine.</p> + +<p class="glossary"><i>Circumnutation</i>—The rotating motion made by the growing +point of the stem and leaf.</p> + +<p class="glossary"><i>Cissus</i>—A genus of vine-like plants often with brilliant +coloured leaves.</p> + +<p class="glossary"><i>Climatic</i>—Influenced by a climate.</p> + +<p class="glossary"><i>Coalesce</i>—To fuse, cohering of parts not usually joined.</p> + +<p class="glossary"><i>Cocos-de-mer</i>—The large double cocoa-nut tree of the +Seychelles Isles.</p> + +<p class="glossary"><i>Collomia</i>—A genus of plants whose seeds are remarkable +for the spiral fibres which expand elastically when +wetted.</p> + +<p class="glossary"><i>Compositæ</i>—A group of plants having an inflorescence +of florets arranged upon a common receptacle or +head.</p> + +<p class="glossary"><i>Concentric</i>—A number of rings having a common centre.</p> + +<p class="glossary"><i>Cone</i>—The hard woody fruits of the fir-tree.</p> + +<p class="glossary"><i>Coniferous</i>—Fir-like, or cone-like; belonging to the cone-bearing +family.</p> + +<p class="glossary"><i>Continuity</i>—Unbroken succession.</p> + +<p class="glossary"><i>Corolla</i>—The second whorl of the floral envelope usually +brightly coloured.</p> + +<p class="glossary"><i>Corpuscles</i>—Grains or granular.</p> + +<p class="glossary"><i>Correlation</i>—<i>i.e.</i>, connection, interdependence.</p> + +<p class="glossary"><i>Cortex</i>—The bark or outer covering of stems.</p> + +<p class="glossary"><i>Cotyledon</i>—A seed leaf.</p> + +<p class="glossary"><i>Cruciferæ</i>—A group of plants having their petals arranged +crosswise, with six stamens two of which are longer +than the others.</p> + +<p class="glossary"><i>Cryptogamic</i>—Relating to flowerless plants.</p> + +<p class="glossary"><i>Culm</i>—The straw-like stems of the grasses.</p> + +<p class="glossary"><span class="pagenum" id="Page_342">[Pg 342]</span></p> + +<p class="glossary"><i>Cuscuta</i>—The dodder genus, parasitic upon flax and +clovers, &c.</p> + +<p class="glossary"><i>Cuticle</i>—The exterior and thickened part of the epidermis.</p> + +<p class="glossary"><i>Cyclamen</i>—Dwarf primulaceous plants with shortened stems +(corms).</p> + + +<h3>D</h3> + +<p class="glossary"><i>Dahlia</i>—A genus of tuberous-rooted plants.</p> + +<p class="glossary"><i>Darlingtonia</i>—A genus of Californian plants related to the +side-saddle plants.</p> + +<p class="glossary"><i>Datura</i>—The generic name of the thorn-apple.</p> + +<p class="glossary"><i>Débris</i>—Remains, rubbish.</p> + +<p class="glossary"><i>Deciduous</i>—Applied to plants, the leaves of which fall off in +autumn.</p> + +<p class="glossary"><i>Dehiscent</i>—Splitting open when ripe.</p> + +<p class="glossary"><i>Deodar</i>—A tree allied to the cedar of Lebanon.</p> + +<p class="glossary"><i>Dentaria</i>—A cruciferous plant bearing bulbils in the axils +of the leaves.</p> + +<p class="glossary"><i>Diagrammatic</i>—Drawn to illustrate a statement.</p> + +<p class="glossary"><i>Dicotyledon</i>—A plant whose embryo has two primary seed-leaves.</p> + +<p class="glossary"><i>Diffusion</i>—The intermingling of fluids (gases or liquids).</p> + +<p class="glossary"><i>Diœcious</i>—When the pistillate flowers and staminate flowers +are borne upon separate plants of the same species.</p> + +<p class="glossary"><i>Dispersion</i>—Scattering.</p> + +<p class="glossary"><i>Drosera</i>—The generic name of the sundews.</p> + + +<h3>E</h3> + +<p class="glossary"><i>Elastic</i>—Springy.</p> + +<p class="glossary"><i>Embryo</i>—The future plant contained in the substance of +the seed.</p> + +<p class="glossary"><i>Embryo-sac</i>—The cavity in the substance of the nucellus, +containing the egg-cell, which after fertilisation +becomes the embryo.</p> + +<p class="glossary"><i>Endocarp</i>—The inside layer of the pericarp.</p> + +<p class="glossary"><i>Entomophilous</i>—Pollinated by insects.</p> + +<p class="glossary"><i>Epicarp</i>—The outside layer of the pericarp.</p> + +<p class="glossary"><span class="pagenum" id="Page_343">[Pg 343]</span></p> + +<p class="glossary"><i>Epidermis</i>—A layer of generally flattened cells forming the +skin of the plant.</p> + +<p class="glossary"><i>Epigean</i>—Developed like the cotyledons of mustard, above +ground.</p> + +<p class="glossary"><i>Epipetalous</i>—Growing upon the petals.</p> + +<p class="glossary"><i>Erysipelas</i>—A disease of the blood causing a red eruption.</p> + +<p class="glossary"><i>Eucalyptus</i>—The generic name of the Australian blue gum +tree.</p> + +<p class="glossary"><i>Euonomin</i>—A dry extract made from the root-bark of +Euonymus altro-purpureus, a North American shrub.</p> + +<p class="glossary"><i>Euonymus</i>—A genus of shrubs and hedgerow trees.</p> + +<p class="glossary"><i>Euphorbia</i>—The spurge genus.</p> + +<p class="glossary"><i>Exogen</i>—Growing by addition to outside of wood and inside +of bark, synonymous with dicotyledon.</p> + + +<h3>F</h3> + +<p class="glossary"><i>Fermentation</i>—Changes that take place in wort when barm +or yeast is added, or when fluids are exposed to the +air. <i>See</i> Yeast.</p> + +<p class="glossary"><i>Fertilised</i>—Completion of the act of fertilisation, <i>i.e.</i>, fusion +of the male element contained in the pollen tube with +the egg cell of the ovule.</p> + +<p class="glossary"><i>Fibrous</i>—Meaning a structure of fine loose filaments or +hairs, <i>i.e.</i>, young rootlets.</p> + +<p class="glossary"><i>Fibro-vascular</i>—A compound tissue of fibres and vessels.</p> + +<p class="glossary"><i>Filament</i>—A thread-like fibre.</p> + +<p class="glossary"><i>Flaccid</i>—Want of firmness, soft and lax.</p> + +<p class="glossary"><i>Fructification</i>—The fruit system of a plant.</p> + +<p class="glossary"><i>Fuchsia</i>—A genus of exotic flowering plants having a +petaloid calyx.</p> + +<p class="glossary"><i>Function</i>—As applied to plant-life, meaning the use and lifework +of the members of a plant.</p> + +<p class="glossary"><i>Fungoid</i>—Growth like a fungus.</p> + + +<h3>G</h3> + +<p class="glossary"><i>Gamopetalous</i>—Petals united.</p> + +<p class="glossary"><span class="pagenum" id="Page_344">[Pg 344]</span></p> + +<p class="glossary"><i>Gamosepalous</i>—Sepals united.</p> + +<p class="glossary"><i>Genesis</i>—Creation, production.</p> + +<p class="glossary"><i>Germinate</i>—The change of the seed from the dormant state +to the active growing stage.</p> + +<p class="glossary"><i>Gloxinia</i>—A genus of popular hothouse plants with large +handsome flowers.</p> + + +<h3>H</h3> + +<p class="glossary"><i>Habitat</i>—The natural abode of a plant.</p> + +<p class="glossary"><i>Herbaceous</i>—Applied to plants which do not form a hard +woody stem.</p> + +<p class="glossary"><i>Herbarium</i>—A collection of dried plants.</p> + +<p class="glossary"><i>Hexagonal</i>—A six-sided and angled figure.</p> + +<p class="glossary"><i>Hibernating</i>—Sleeping, a dormant condition.</p> + +<p class="glossary"><i>Hilum</i>—The black scar on a bean seed.</p> + +<p class="glossary"><i>Hippuris</i>—A genus of aquatic flowering plants.</p> + +<p class="glossary"><i>Horizontal</i>—Parallel to the horizon level.</p> + +<p class="glossary"><i>Hoya</i>—A genus of tropical climbing plants.</p> + +<p class="glossary"><i>Hyacinths</i>—Bulbous plants.</p> + +<p class="glossary"><i>Hydrangea</i>—A genus of flowering shrubs.</p> + +<p class="glossary"><i>Hygienic</i>—Relating to the preservation of health.</p> + +<p class="glossary"><i>Hygrometric</i>—Moisture and its influence.</p> + +<p class="glossary"><i>Hygroscopic</i>—Applied to the film of water surrounding the +particles of the soil.</p> + +<p class="glossary"><i>Hypericum</i>—The generic name of the St. John’s wort.</p> + +<p class="glossary"><i>Hyphæ</i>—Filaments or threads of the fungus spawn.</p> + +<p class="glossary"><i>Hypogean</i>—Development of the cotyledons under ground.</p> + + +<h3>I</h3> + +<p class="glossary"><i>Impatiens</i>—The generic name of the balsam.</p> + +<p class="glossary"><i>Impervious</i>—Not to be penetrated by water.</p> + +<p class="glossary"><i>Insectivorous</i>—Catching and killing insects, plants that have +this power and can absorb the decomposed insects.</p> + +<p class="glossary"><i>Insoluble</i>—Substances that do not dissolve in water.</p> + +<p class="glossary"><span class="pagenum" id="Page_345">[Pg 345]</span></p> + +<p class="glossary"><i>Intercellular</i>—Spaces between the cells.</p> + +<p class="glossary"><i>Internode</i>—The space between two nodes.</p> + +<p class="glossary"><i>Involucre</i>—A whorl of bracts.</p> + +<p class="glossary"><i>Iodine</i>—A soluble substance extracted from kelp and used +as a test for starch.</p> + + +<h3>L</h3> + +<p class="glossary"><i>Laburnum</i>—Yellow-flowered trees allied to the Pea +family (Leguminosæ).</p> + +<p class="glossary"><i>Legume</i>—The dehiscent fruit of the pea family, a pod.</p> + +<p class="glossary"><i>Leguminosæ</i>—A family of plants having for their fruit a +legume or pod, <i>i.e.</i>, Pea, Laburnum.</p> + +<p class="glossary"><i>Lenticels</i>—Minute pores in the bark.</p> + +<p class="glossary"><i>Liane</i>—A hanging root or stem.</p> + +<p class="glossary"><i>Liber</i>—The inner bark, same as phloëm.</p> + +<p class="glossary"><i>Linnæa</i>—A genus of dwarf trailing plants.</p> + +<p class="glossary"><i>Luscious</i>—Sweet and succulent.</p> + + +<h3>M</h3> + +<p class="glossary"><i>Magnesium</i>—The metallic base of magnesia.</p> + +<p class="glossary"><i>Magnolia</i>—A genus of flowering shrubs and trees.</p> + +<p class="glossary"><i>Mahonia</i>—A genus of evergreen shrubs belonging to the +barberry family.</p> + +<p class="glossary"><i>Martynia</i>—A genus of plants having capsules with long +curved hooks.</p> + +<p class="glossary"><i>Melampyrum</i>—A genus of dwarf flowering plants partly +parasitic.</p> + +<p class="glossary"><i>Membranous</i>—Thin and destitute of green colour usually +applied to bracts.</p> + +<p class="glossary"><i>Mesocarp</i>—The central layer of the pericarp.</p> + +<p class="glossary"><i>Mesophyll</i>—The ground tissue of the leaf.</p> + +<p class="glossary"><i>Metabolism</i>—Changes which take place in protoplasm and +which it causes in other substances.</p> + +<p class="glossary"><i>Microbes</i>—A term applied to one-celled plant atoms, like +bacteria.</p> + +<p class="glossary"><span class="pagenum" id="Page_346">[Pg 346]</span></p> + +<p class="glossary"><i>Micropyle</i>—A small pore in the coats of the ovule through +which the pollen tube passes.</p> + +<p class="glossary"><i>Modicum</i>—Moderate sized; a small quantity.</p> + +<p class="glossary"><i>Monocotyledon</i>—A plant whose seed is furnished with one +seed leaf.</p> + +<p class="glossary"><i>Monœcious</i>—Applied to a plant when the stamens and pistil +are in distinct flowers.</p> + +<p class="glossary"><i>Monstera</i>—A genus of climbing aroids with edible fruit.</p> + +<p class="glossary"><i>Mucilaginous</i>—Sticky, gumlike, secreting mucilage.</p> + +<p class="glossary"><i>Mucuna</i>—A genus of Brazilian Leguminosæ, yielding the +cowage (consisting of intensely irritating hairs), of the +Materia Medica.</p> + +<p class="glossary"><i>Mutualism</i>—Interchange of some advantage, botanically +applied to the union of two dissimilar plants which +live in <i>close</i> contact with each other to their mutual +benefit.</p> + +<p class="glossary"><i>Mycelium</i>—The root-like colourless filaments of fungi.</p> + +<p class="glossary"><i>Mycetozoa</i>—A term applied to the slime-fungi.</p> + + +<h3>N</h3> + +<p class="glossary"><i>Nectary</i>—A honey secreting gland or spur.</p> + +<p class="glossary"><i>Nemophila</i>—A genus of dwarf annual flowering plants.</p> + +<p class="glossary"><i>Nepenthes</i>—A genus of plants having as a prolongation of +the midrib of the leaves, ascidia or pitchers.</p> + +<p class="glossary"><i>Nocturnal</i>—Happening by night.</p> + +<p class="glossary"><i>Node</i>—The exact point on the stem from which the leaf is +developed.</p> + +<p class="glossary"><i>Normal</i>—Regular, unaffected by any modification.</p> + +<p class="glossary"><i>Noxious</i>—Hurtful or poisonous.</p> + +<p class="glossary"><i>Nucellus</i>—The internal tissue of the ovule within which the +embryo-sac is embedded.</p> + +<p class="glossary"><i>Nutrition</i>—The process and function of taking in food for +the purpose of growth and to replace waste.</p> + +<p class="glossary"><span class="pagenum" id="Page_347">[Pg 347]</span></p> + + +<h3>O</h3> + +<p class="glossary"><i>Orchis</i>—A genus of the orchid family growing in the soil.</p> + +<p class="glossary"><i>Osmosis</i>—The passage of fluids through a membrane.</p> + +<p class="glossary"><i>Ovary</i>—The ovule case, that part of the carpel that bears +ovules.</p> + +<p class="glossary"><i>Ovule</i>—The structure which after fertilisation forms the +seed.</p> + +<p class="glossary"><i>Ovum</i>—The egg cell of the ovule.</p> + +<p class="glossary"><i>Oxalis</i>—The generic name of the wood sorrel.</p> + +<p class="glossary"><i>Oxygen</i>—A gas, one of the constituents of the atmosphere.</p> + + +<h3>P</h3> + +<p class="glossary"><i>Palisade-tissue</i>—A tissue of oblong cells placed side by side +at right angles to the flat surface of the leaf.</p> + +<p class="glossary"><i>Papilionaceous</i>—Butterfly shaped.</p> + +<p class="glossary"><i>Pappus</i>—A light hairy development from the calyx of some +plants.</p> + +<p class="glossary"><i>Parasitic</i>—The habit of growing upon and deriving nourishment +from another plant.</p> + +<p class="glossary"><i>Pellucid</i>—Shining and transparent.</p> + +<p class="glossary"><i>Perennial</i>—Plants that live for an indefinite period.</p> + +<p class="glossary"><i>Perianth</i>—A term used when there is no distinction between +calyx and corolla.</p> + +<p class="glossary"><i>Pericarp</i>—The ripened walls of the ovary constituting the +structure of the fruit.</p> + +<p class="glossary"><i>Persistent</i>—Applied to the parts of the flower that remain +on for some time.</p> + +<p class="glossary"><i>Petunia</i>—A genus of Brazilian Solanaceæ.</p> + +<p class="glossary"><i>Philodendron</i>—A genus of aroids usually climbers.</p> + +<p class="glossary"><i>Phleum</i>—A grass.</p> + +<p class="glossary"><i>Phloëm</i>—The inner bark, containing sieve-tubes.</p> + +<p class="glossary"><i>Phosphate</i>—A salt formed by the union of phosphoric acid +with some base.</p> + +<p class="glossary"><i>Phyllotaxis</i>—The law of leaf arrangement.</p> + +<p class="glossary"><i>Physalis</i>—The generic name of the winter cherry.</p> + +<p class="glossary"><span class="pagenum" id="Page_348">[Pg 348]</span></p> + +<p class="glossary"><i>Physiological</i>—Having reference to the function or life work +of the plant.</p> + +<p class="glossary"><i>Picea</i>—A genus of the Conifer family.</p> + +<p class="glossary"><i>Pinetum</i>—A garden devoted to the culture of pine-trees.</p> + +<p class="glossary"><i>Pinguicula</i>—The generic name of the butterworts.</p> + +<p class="glossary"><i>Pinus</i>—A genus of the Conifer family.</p> + +<p class="glossary"><i>Pistil</i>—The female part of the flower consisting of ovary, +style, and stigma.</p> + +<p class="glossary"><i>Pistillate</i>—Applied to flowers having the pistil only.</p> + +<p class="glossary"><i>Pith</i>—The soft tissue in the centre of the stem.</p> + +<p class="glossary"><i>Plumbago</i>—The generic name of the leadworts, small +flowering plants and shrubs.<a id="tn-015"></a></p> + +<p class="glossary"><i>Plumule</i>—The first stem shoot of the germinating seed.</p> + +<p class="glossary"><i>Poa</i>—A grass.</p> + +<p class="glossary"><i>Poinsettia</i>—A genus of Mexican plants having bright scarlet +bracts and small flowers.</p> + +<p class="glossary"><i>Pollard</i>—A tree trunk with its branches cut short.</p> + +<p class="glossary"><i>Pollen</i>—The fertilising or male part of the flower.</p> + +<p class="glossary"><i>Pollination</i>—The act of conveying the pollen from the +stamen to the stigma.</p> + +<p class="glossary"><i>Polypetalous</i>—Separate or many petals.</p> + +<p class="glossary"><i>Polysepalous</i>—Separate or many sepals.</p> + +<p class="glossary"><i>Polytrichum</i>—The generic name of the hair moss.</p> + +<p class="glossary"><i>Potassium</i>—The metallic base of potash.</p> + +<p class="glossary"><i>Proboscis</i>—The feeling and feeding organ of an insect.</p> + +<p class="glossary"><i>Prothallus</i>—The first growth when the spore of a fern +germinates.</p> + +<p class="glossary"><i>Protococcus</i>—A genus of unicellular plants forming a green +stain upon trees, &c.</p> + +<p class="glossary"><i>Protonema</i>—The first growth of the moss-spore.</p> + +<p class="glossary"><i>Protoplasm</i>—A highly complex substance forming the essential +part of all living cells, and to which all life growth +is due.</p> + +<p class="glossary"><i>Prototypes</i>—First forms of plant-life.</p> + +<p class="glossary"><i>Psamma</i>—A genus of the grass family.</p> + +<p class="glossary"><span class="pagenum" id="Page_349">[Pg 349]</span></p> + +<p class="glossary"><i>Pseudo-bulb</i>—A swollen stem common in the epiphytic +orchids.</p> + +<p class="glossary"><i>Pteris</i>—The generic name of the bracken fern.</p> + + +<h3>Q</h3> + +<p class="glossary"><i>Quiescent</i>—Inactive, dormant.</p> + +<p class="glossary"><i>Quinine</i>—An alkaloid extracted from the cinchona trees.</p> + + +<h3>R</h3> + +<p class="glossary"><i>Radicle</i>—The first formed root when a seed germinates.</p> + +<p class="glossary"><i>Rafflesia</i>—A genus of brown leafless parasites.</p> + +<p class="glossary"><i>Receptacle</i>—That part of the stalk on which the flower is +developed.</p> + +<p class="glossary"><i>Resin</i>—A secretion from certain trees which hardens on +exposure.</p> + +<p class="glossary"><i>Respiration</i>—The process of breathing.</p> + +<p class="glossary"><i>Rhinanthus</i>—The generic name of the yellow-rattle (a root +parasite).</p> + +<p class="glossary"><i>Rhododendron</i>—A genus of popular flowering shrubs and +dwarf trees.</p> + +<p class="glossary"><i>Root-cap</i>—A loose covering of tissue that protects the extreme +point of the growing root.</p> + +<p class="glossary"><i>Root-hairs</i>—The delicate unicellular hairs found on the +young root.</p> + + +<h3>S</h3> + +<p class="glossary"><i>Salicine</i>—A substance obtained from the bark of willows, +soluble in water and alcohol, and crystallising in bright +white needles.</p> + +<p class="glossary"><i>Salvia</i>—A genus of labiate plants.</p> + +<p class="glossary"><i>Samara</i>—Winged fruit.</p> + +<p class="glossary"><i>Saprophyte</i>—Plants that live upon decaying organic matter.</p> + +<p class="glossary"><i>Sarracenia</i>—The generic name of the North American +side-saddle plants.</p> + +<p class="glossary"><i>Saxifraga</i>—A genus of dwarf Alpine plants.</p> + +<p class="glossary"><span class="pagenum" id="Page_350">[Pg 350]</span></p> + +<p class="glossary"><i>Scales</i>—Rudimentary leaves.</p> + +<p class="glossary"><i>Secretion</i>—Applied to substances like resin and honey, the +production of assimilation and metabolism.</p> + +<p class="glossary"><i>Sedum</i>—A genus of succulent Alpine plants.</p> + +<p class="glossary"><i>Soluble</i>—Any substance that dissolves in water.</p> + +<p class="glossary"><i>Spadix</i>—The inflorescence of the Aroideæ.</p> + +<p class="glossary"><i>Spathe</i>—The bract of the Aroideæ.</p> + +<p class="glossary"><i>Sporangium</i>—The spore-case of some of the cryptogamia.</p> + +<p class="glossary"><i>Spurious</i>—False.</p> + +<p class="glossary"><i>Stapelia</i>—A genus of succulent plants, very poisonous and +fœtid.</p> + +<p class="glossary"><i>Starch</i>—Colourless grains, a product of assimilation in the +leaf.</p> + +<p class="glossary"><i>Stigma</i>—The receptive part of the pistil.</p> + +<p class="glossary"><i>Stipa</i>—A genus of the grass family.</p> + +<p class="glossary"><i>Stipules</i>—Small outgrowths at the base of the petiole.</p> + +<p class="glossary"><i>Stomata</i>—Minute pores in the epidermis of the leaf or green +stem.</p> + +<p class="glossary"><i>Sulphate</i>—A salt formed by the combination of sulphuric +acid with some base.</p> + +<p class="glossary"><i>Sycamore</i>—The plane tree of Scotland, Acer pseudoplatanus.</p> + +<p class="glossary"><i>Symbiosis</i>—Mutualism, a living for one another, interchange +of benefits by united growth.</p> + +<p class="glossary"><i>Syncarpous</i>—United carpels.</p> + + +<h3>T</h3> + +<p class="glossary"><i>Tannin</i>—A substance widely diffused through the leaves +and stems of plants, of an astringent character.</p> + +<p class="glossary"><i>Tap-root</i>—A root that forms an unbranched tapering axis: +<i>Ex.</i>, carrot.</p> + +<p class="glossary"><i>Tendril</i>—A coiled or hooked filament modified to assist +plants to climb.</p> + +<p class="glossary"><i>Tentacles</i>—The glandular and feeler-like hairs of the +sundew.</p> + +<p class="glossary"><i>Terminal</i>—At the apex or end.</p> + +<p class="glossary"><span class="pagenum" id="Page_351">[Pg 351]</span></p> + +<p class="glossary"><i>Testa</i>—The skin of seed.</p> + +<p class="glossary"><i>Tigellum</i>—The first stalk of the seed bearing the cotyledons.</p> + +<p class="glossary"><i>Tillandsia</i>—A New World genus of perching or epiphytic +plants.</p> + +<p class="glossary"><i>Tissue</i>—A group of cells; having a common origin.</p> + +<p class="glossary"><i>Tormentilla</i>—A genus of small creeping rosaceous plants.</p> + +<p class="glossary"><i>Transpiration</i>—The giving off of water vapour from the +surface of leaves and stems.</p> + +<p class="glossary"><i>Tricyrtis</i>—The generic name of the toad-lily.</p> + +<p class="glossary"><i>Tuber</i>—A fleshy root or succulent underground stem.</p> + + +<h3>U</h3> + +<p class="glossary"><i>Umbelliferæ</i>—A group of plants having an umbellate arrangement +of the inflorescence or flower-head.</p> + + +<h3>V</h3> + +<p class="glossary"><i>Vallisneria</i>—A genus of aquatic flowering plants.</p> + +<p class="glossary"><i>Valved</i>—Having valves, <i>e.g.</i>, anther of the barberry.</p> + +<p class="glossary"><i>Vapour</i>—Gas into which most liquids and solids are converted +by heat.</p> + +<p class="glossary"><i>Vasculum</i>—A little vessel or box for collecting botanical +and other specimens.</p> + +<p class="glossary"><i>Venation</i>—The arrangement of veins in a leaf.</p> + + +<h3>W</h3> + +<p class="glossary"><i>Weigelia</i>—A genus of flowering shrubs allied to Honeysuckle.</p> + +<p class="glossary"><i>Whorl</i>—An arrangement of leaves or parts of the flower +in rings.</p> + +<p class="glossary"><i>Wort</i>—Sweet unfermented new beer.</p> + + +<h3>Y</h3> + +<p class="glossary"><i>Yeast</i>—A unicellular plant that sets up fermentation under +certain conditions.</p> + +<hr class="chap x-ebookmaker-drop"> +<div class="chapter"> + + + <h2 class="nobreak" id="FOOTNOTES"> + FOOTNOTES. + + </h2> +</div> + +<div class="footnote"><p><a id="Footnote_1_1" href="#FNanchor_1_1" class="label">[1]</a> See glossary at the end of the book.</p></div> + +<div class="footnote"><p><a id="Footnote_2_2" href="#FNanchor_2_2" class="label">[2]</a> <i>Protococcus.</i></p></div> + +<div class="footnote"><p><a id="Footnote_3_3" href="#FNanchor_3_3" class="label">[3]</a> First form of fungoid growth.</p></div> + +<div class="footnote"><p><a id="Footnote_4_4" href="#FNanchor_4_4" class="label">[4]</a> Small living atoms.</p></div> + +<div class="footnote"><p><a id="Footnote_5_5" href="#FNanchor_5_5" class="label">[5]</a> It can generally be met with at naturalists’ shops where aquaria +are sold.</p></div> + +<div class="footnote"><p><a id="Footnote_6_6" href="#FNanchor_6_6" class="label">[6]</a> Author of “Vegetable Physiology.”</p></div> + +<div class="footnote"><p><a id="Footnote_7_7" href="#FNanchor_7_7" class="label">[7]</a> For instance, I have noticed some curious examples of trees +growing together. A Turkey oak and Silver fir in my own grounds +are closely united at the base. The fir-seed and the acorn must +have germinated in such close proximity that the stems have almost +grown into each other. The group of beeches shown in the plate +gives another example of interlacing stems and roots.</p></div> + +<div class="footnote"><p><a id="Footnote_8_8" href="#FNanchor_8_8" class="label">[8]</a> The Maranta and a few other plants are exceptions to this rule.</p></div> + +<div class="footnote"><p><a id="Footnote_9_9" href="#FNanchor_9_9" class="label">[9]</a> That is, a leaf like the beech or sycamore, having an upper and +under surface; vertical leaves, like the iris, have palisade tissue on +both sides.</p></div> + +<div class="footnote"><p><a id="Footnote_10_10" href="#FNanchor_10_10" class="label">[10]</a> Greek: <i>proto</i>, “first”; <i>plasma</i>, “anything moulded.”</p></div> + +<div class="footnote"><p><a id="Footnote_11_11" href="#FNanchor_11_11" class="label">[11]</a> Carbon meaning charcoal, and dioxide meaning two parts +oxygen. Sometimes called carbonic-acid gas.</p></div> + +<div class="footnote"><p><a id="Footnote_12_12" href="#FNanchor_12_12" class="label">[12]</a> “Sharp Eyes,” by Wm. Hamilton Gibson.</p></div> + +<div class="footnote"><p><a id="Footnote_13_13" href="#FNanchor_13_13" class="label">[13]</a> The uppermost bud of the central shoot.</p></div> + +<div class="footnote"><p><a id="Footnote_14_14" href="#FNanchor_14_14" class="label">[14]</a> Venation.</p></div> + +<div class="footnote"><p><a id="Footnote_15_15" href="#FNanchor_15_15" class="label">[15]</a> One seed leaf plants.</p></div> + +<div class="footnote"><p><a id="Footnote_16_16" href="#FNanchor_16_16" class="label">[16]</a> All cross-shaped flowers do not, however, belong to this order.</p></div> + +<div class="footnote"><p><a id="Footnote_17_17" href="#FNanchor_17_17" class="label">[17]</a> <i>Dianthræcia albimacula.</i></p></div> + +<div class="footnote"><p><a id="Footnote_18_18" href="#FNanchor_18_18" class="label">[18]</a> Thorn-apple.</p></div> + +<div class="footnote"><p><a id="Footnote_19_19" href="#FNanchor_19_19" class="label">[19]</a> Strictly speaking, the endocarp of the orange is a thin membrane, +and the pulp grows from it and fills up the ovary cavities.</p></div> + +<div class="footnote"><p><a id="Footnote_20_20" href="#FNanchor_20_20" class="label">[20]</a> In botany the fruit of the sycamore, maple, ash, &c., is called a +<i>samara</i>, and is properly speaking a winged <i>achene</i>.</p></div> + +<div class="footnote"><p><a id="Footnote_21_21" href="#FNanchor_21_21" class="label">[21]</a> The beard of corn.</p></div> + +<div class="footnote"><p><a id="Footnote_22_22" href="#FNanchor_22_22" class="label">[22]</a> Sensitive to moisture.</p></div> + +<div class="footnote"><p><a id="Footnote_23_23" href="#FNanchor_23_23" class="label">[23]</a> Called botanically an “absciss layer.”</p></div> + +<div class="footnote"><p><a id="Footnote_24_24" href="#FNanchor_24_24" class="label">[24]</a> They merely need to be placed between sheets of blotting +paper, which should be dried daily and kept in a press +or under a weight for a few days until the specimens are fit +to be placed in a book.</p></div> + +<div class="footnote"><p><a id="Footnote_25_25" href="#FNanchor_25_25" class="label">[25]</a> Testing the effect of plant food by water-culture is carried out in +the following manner. Six large jars are filled with distilled water. +In No. 1 all the six elements above mentioned are placed in small +quantities, so as to form a weak solution. In No. 2 only five of +them are added to the water, and in each succeeding jar one +element is left out. A seedling plant which has been germinated +on damp sand is suspended in each jar in such a manner that the +leaves are in the air and the roots in the water without the seed +touching the liquid. The growth of the young plants is carefully +observed, and the result is found to be that No. 1 will grow and +flourish, finding all its needful food in the water, whilst the rest of +the seedlings will show plainly by their feeble and starved condition +that, the food elements being absent, they cannot build up their +stems and leaves, and must eventually perish.</p></div> + +<div class="footnote"><p><a id="Footnote_26_26" href="#FNanchor_26_26" class="label">[26]</a> From “Nat. Hist. of Plants,” p. 554: “It has been proved +experimentally that seeds which have been deprived by calcium +chloride of as much water as possible are not killed even at the +boiling point of water.” Careful experiment has shown that there +are three stages of activity in the life and work of a plant—(1) A +<i>minimum or zero</i>, at which the processes are just possible; (2) a +<i>medium stage</i> or <i>optimum point</i> where the activity is the greatest; +and (3) a <i>maximum stage of heat</i> where <i>growth is arrested</i>. So +that we learn that plant-life can suffer from too high a temperature +as well as that which is too low.</p></div> + +<div class="footnote"><p><a id="Footnote_27_27" href="#FNanchor_27_27" class="label">[27]</a> On sunny heaths they are often of a rich crimson tint.</p></div> + +<div class="footnote"><p><a id="Footnote_28_28" href="#FNanchor_28_28" class="label">[28]</a> See Frontispiece.</p></div> + +<div class="footnote"><p><a id="Footnote_29_29" href="#FNanchor_29_29" class="label">[29]</a> Barren.</p></div> + +<div class="footnote"><p><a id="Footnote_30_30" href="#FNanchor_30_30" class="label">[30]</a> A single flower of the Heracleum giganteum would not be +specially noticeable, but when hundreds of them are grouped together +in a huge umbelliferous head they form a most striking object, +as may be seen in the plate. I have often watched the swarms of +flies, beetles, and bees visiting these attractive blossoms on sunny +days, and the great umbels of seed in autumn showed how effectually +the insects had carried out their work of fertilisation.</p></div> + +<div class="footnote"><p><a id="Footnote_31_31" href="#FNanchor_31_31" class="label">[31]</a> A word meaning two plants living together and deriving mutual +benefit.</p></div> + +<div class="footnote"><p><a id="Footnote_32_32" href="#FNanchor_32_32" class="label">[32]</a> Miss Blanche Atkinson, member of the Barmouth Branch of +the Selborne Society.</p></div> + +<div class="chapter"> +<p><span class="pagenum" id="Page_352">[Pg 352]</span></p> + +<p class="center blackletter"> +The Gresham Press,</p> +<p class="center">UNWIN BROTHERS,</p> +<p class="center"><span style="font-size: 90%;">WOKING AND LONDON.</span></p> +</div> +<div class="transnote chapter"> +<h2 class="nobreak">Transcriber’s notes</h2> +<p> +The text contains some inconsistencies in the usage of hyphens. Only +those have been corrected where the majority was spelled otherwise. All +corrections made are listed below. +</p> +<p> +The asterism (⁂) on page 8 should be inverted, but this was not +possible to achieve since no such symbol exists. +</p> +<h3>Corrections</h3> + +<ul> +<li>Page <a href="#tn-001">20</a> In the list of illustrations hyphens removed from “MONO-COTYLEDON” and “DI-COTYLEDON”, to match the captions of the illustrations;</li> +<li>Page <a href="#tn-025">21</a> In the list of illustrations “WHITE LILY PISTIL” changed to “WHITE-LILY PISTIL” and “POLLEN-TUBE” changed to “POLLEN TUBE”, to match the captions of the illustrations;</li><li>Page <a href="#tn-002">22</a> In the list of illustrations “DIAGRAM OF TRANSFUSION” changed to “TRANSFUSION DIAGRAM”, to match the caption of the illustration;</li> +<li>Page <a href="#tn-003">22</a> In the list of illustrations “FLAVA” added behind “SARRACENIA”, to match the caption of the illustration;</li> +<li>Page <a href="#tn-017">42</a> “Another of these perching-plants is <i>Tillandsia Usnoides</i>” changed to “Another of these perching-plants is <i>Tillandsia Usneoides</i>”;</li> +<li>Page <a href="#tn-016">62</a> “aërial” in “send down slender aërial roots” changed to “aerial”, to match the spelling in the rest of the text;</li> +<li>Page <a href="#tn-019">69</a> In “We may look upon the earth as being a sort of store-house”, “store-house” changed to “storehouse” to match the spelling in the +rest of the text;</li> +<li>Page <a href="#tn-020">100</a> In “in a skeleton leaf, the mid-rib”, “mid-rib” changed to “midrib”, to match the spelling in the rest of the text;</li> +<li>Page <a href="#tn-018">102</a> one occurrence of “called” removed from “belonging to this + great division are called called dicotyledons”;</li> +<li>Page <a href="#tn-004">136</a> “to render the bud waterpoof” changed to “to render the bud waterproof”;</li> +<li>Page <a href="#tn-021">138</a> In “wood-sorrel is rolled into a spiral”, “wood-sorrel” is changed to “woodsorrel”, to match the spelling in the rest of the text;</li> +<li>Page <a href="#tn-022">159</a> In “resembling the pine-apple plant”, the hyphen is removed from “pine-apple”, to match the spelling in the rest of the text;</li> +<li>Page <a href="#tn-005">202</a> “so that at length the seed-coats” changed to “so that at length the seed-coat”;</li> +<li>Page <a href="#tn-006">223</a> “PINE CONES” in the caption of the illustration, change to “PINE-CONES” (in the list of illustrations too) to match the spelling in the rest of the text and “fir-cones”;</li> +<li>Page <a href="#tn-008">232</a> “have seeds with wings lightly twisted” changed to “have seeds with wings slightly twisted”;</li> +<li>Page <a href="#tn-009">238</a> “COCOS DE MER” in the caption of the illustration is hyphenated (in the list of illustrations too) to match the spelling in the rest of the text;</li> +<li>Page <a href="#tn-023">246</a> “ivy-leafed toadflax” changed to “ivy-leafed toad-flax” to match the spelling in the rest of the text;</li> +<li>Page <a href="#tn-010">263</a> “the little epiphyte (mentioned in our first chapter), <i>Tillandsia usnoides</i>” changed to “the little epiphyte (mentioned in our first chapter), <i>Tillandsia usneoides</i>”;</li> +<li>Page <a href="#tn-012">263</a> “by decomposing the carbon dioxide” changed to “by decomposing the carbon-dioxide” to match the spelling in the rest of the text;</li> +<li>Page <a href="#tn-011">280</a> “absorbs this carbon dioxide and effects” changed to “absorbs this carbon-dioxide and effects” to match the spelling in the rest of the text;</li> +<li>Page <a href="#tn-013">337</a> “into technical phraseology these defininition” changed to “into technical phraseology these definitions”;</li> +<li>Page <a href="#tn-014">340</a> comma after “tuberous Umbelliferæ, earthnut” changed to period;</li> +<li>Page <a href="#tn-015">348</a> period added after “flowering plants and shrubs”.</li> +</ul> +</div> +<div style='text-align:center'>*** END OF THE PROJECT GUTENBERG EBOOK 79061 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