1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
297
298
299
300
301
302
303
304
305
306
307
308
309
310
311
312
313
314
315
316
317
318
319
320
321
322
323
324
325
326
327
328
329
330
331
332
333
334
335
336
337
338
339
340
341
342
343
344
345
346
347
348
349
350
351
352
353
354
355
356
357
358
359
360
361
362
363
364
365
366
367
368
369
370
371
372
373
374
375
376
377
378
379
380
381
382
383
384
385
386
387
388
389
390
391
392
393
394
395
396
397
398
399
400
401
402
403
404
405
406
407
408
409
410
411
412
413
414
415
416
417
418
419
420
421
422
423
424
425
426
427
428
429
430
431
432
433
434
435
436
437
438
439
440
441
442
443
444
445
446
447
448
449
450
451
452
453
454
455
456
457
458
459
460
461
462
463
464
465
466
467
468
469
470
471
472
473
474
475
476
477
478
479
480
481
482
483
484
485
486
487
488
489
490
491
492
493
494
495
496
497
498
499
500
501
502
503
504
505
506
507
508
509
510
511
512
513
514
515
516
517
518
519
520
521
522
523
524
525
526
527
528
529
530
531
532
533
534
535
536
537
538
539
540
541
542
543
544
545
546
547
548
549
550
551
552
553
554
555
556
557
558
559
560
561
562
563
564
565
566
567
568
569
570
571
572
573
574
575
576
577
578
579
580
581
582
583
584
585
586
587
588
589
590
591
592
593
594
595
596
597
598
599
600
601
602
603
604
605
606
607
608
609
610
611
612
613
614
615
616
617
618
619
620
621
622
623
624
625
626
627
628
629
630
631
632
633
634
635
636
637
638
639
640
641
642
643
644
645
646
647
648
649
650
651
652
653
654
655
656
657
658
659
660
661
662
663
664
665
666
667
668
669
670
671
672
673
674
675
676
677
678
679
680
681
682
683
684
685
686
687
688
689
690
691
692
693
694
695
696
697
698
699
700
701
702
703
704
705
706
707
708
709
710
711
712
713
714
715
716
717
718
719
720
721
722
723
724
725
726
727
728
729
730
731
732
733
734
735
736
737
738
739
740
741
742
743
744
745
746
747
748
749
750
751
752
753
754
755
756
757
758
759
760
761
762
763
764
765
766
767
768
769
770
771
772
773
774
775
776
777
778
779
780
781
782
783
784
785
786
787
788
789
790
791
792
793
794
795
796
797
798
799
800
801
802
803
804
805
806
807
808
809
810
811
812
813
814
815
816
817
818
819
820
821
822
823
824
825
826
827
828
829
830
831
832
833
834
835
836
837
838
839
840
841
842
843
844
845
846
847
848
849
850
851
852
853
854
855
856
857
858
859
860
861
862
863
864
865
866
867
868
869
870
871
872
873
874
875
876
877
878
879
880
881
882
883
884
885
886
887
888
889
890
891
892
893
894
895
896
897
898
899
900
901
902
903
904
905
906
907
908
909
910
911
912
913
914
915
916
917
918
919
920
921
922
923
924
925
926
927
928
929
930
931
932
933
934
935
936
937
938
939
940
941
942
943
944
945
946
947
948
949
950
951
952
953
954
955
956
957
958
959
960
961
962
963
964
965
966
967
968
969
970
971
972
973
974
975
976
977
978
979
980
981
982
983
984
985
986
987
988
989
990
991
992
993
994
995
996
997
998
999
1000
1001
1002
1003
1004
1005
1006
1007
1008
1009
1010
1011
1012
1013
1014
1015
1016
1017
1018
1019
1020
1021
1022
1023
1024
1025
1026
1027
1028
1029
1030
1031
1032
1033
1034
1035
1036
1037
1038
1039
1040
1041
1042
1043
1044
1045
1046
1047
1048
1049
1050
1051
1052
1053
1054
1055
1056
1057
1058
1059
1060
1061
1062
1063
1064
1065
1066
1067
1068
1069
1070
1071
1072
1073
1074
1075
1076
1077
1078
1079
1080
1081
1082
1083
1084
1085
1086
1087
1088
1089
1090
1091
1092
1093
1094
1095
1096
1097
1098
1099
1100
1101
1102
1103
1104
1105
1106
1107
1108
1109
1110
1111
1112
1113
1114
1115
1116
1117
1118
1119
1120
1121
1122
1123
1124
1125
1126
1127
1128
1129
1130
1131
1132
1133
1134
1135
1136
1137
1138
1139
1140
1141
1142
1143
1144
1145
1146
1147
1148
1149
1150
1151
1152
1153
1154
1155
1156
1157
1158
1159
1160
1161
1162
1163
1164
1165
1166
1167
1168
1169
1170
1171
1172
1173
1174
1175
1176
1177
1178
1179
1180
1181
1182
1183
1184
1185
1186
1187
1188
1189
1190
1191
1192
1193
1194
1195
1196
1197
1198
1199
1200
1201
1202
1203
1204
1205
1206
1207
1208
1209
1210
1211
1212
1213
1214
1215
1216
1217
1218
1219
1220
1221
1222
1223
1224
1225
1226
1227
1228
1229
1230
1231
1232
1233
1234
1235
1236
1237
1238
1239
1240
1241
1242
1243
1244
1245
1246
1247
1248
1249
1250
1251
1252
1253
1254
1255
1256
1257
1258
1259
1260
1261
1262
1263
1264
1265
1266
1267
1268
1269
1270
1271
1272
1273
1274
1275
1276
1277
1278
1279
1280
1281
1282
1283
1284
1285
1286
1287
1288
1289
1290
1291
1292
1293
1294
1295
1296
1297
1298
1299
1300
1301
1302
1303
1304
1305
1306
1307
1308
1309
1310
1311
1312
1313
1314
1315
1316
1317
1318
1319
1320
1321
1322
1323
1324
1325
1326
1327
1328
1329
1330
1331
1332
1333
1334
1335
1336
1337
1338
1339
1340
1341
1342
1343
1344
1345
1346
1347
1348
1349
1350
1351
1352
1353
1354
1355
1356
1357
1358
1359
1360
1361
1362
1363
1364
1365
1366
1367
1368
1369
1370
1371
1372
1373
1374
1375
1376
1377
1378
1379
1380
1381
1382
1383
1384
1385
1386
1387
1388
1389
1390
1391
1392
1393
1394
1395
1396
1397
1398
1399
1400
1401
1402
1403
1404
1405
1406
1407
1408
1409
1410
1411
1412
1413
1414
1415
1416
1417
1418
1419
1420
1421
1422
1423
1424
1425
1426
1427
1428
1429
1430
1431
1432
1433
1434
1435
1436
1437
1438
1439
1440
1441
1442
1443
1444
1445
1446
1447
1448
1449
1450
1451
1452
1453
1454
1455
1456
1457
1458
1459
1460
1461
1462
1463
1464
1465
1466
1467
1468
1469
1470
1471
1472
1473
1474
1475
1476
1477
1478
1479
1480
1481
1482
1483
1484
1485
1486
1487
1488
1489
1490
1491
1492
1493
1494
1495
1496
1497
1498
1499
1500
1501
1502
1503
1504
1505
1506
1507
1508
1509
1510
1511
1512
1513
1514
1515
1516
1517
1518
1519
1520
1521
1522
1523
1524
1525
1526
1527
1528
1529
1530
1531
1532
1533
1534
1535
1536
1537
1538
1539
1540
1541
1542
1543
1544
1545
1546
1547
1548
1549
1550
1551
1552
1553
1554
1555
1556
1557
1558
1559
1560
1561
1562
1563
1564
1565
1566
1567
1568
1569
1570
1571
1572
1573
1574
1575
1576
1577
1578
1579
1580
1581
1582
1583
1584
1585
1586
1587
1588
1589
1590
1591
1592
1593
1594
1595
1596
1597
1598
1599
1600
1601
1602
1603
1604
1605
1606
1607
1608
1609
1610
1611
1612
1613
1614
1615
1616
1617
1618
1619
1620
1621
1622
1623
1624
1625
1626
1627
1628
1629
1630
1631
1632
1633
1634
1635
1636
1637
1638
1639
1640
1641
1642
1643
1644
1645
1646
1647
1648
1649
1650
1651
1652
1653
1654
1655
1656
1657
1658
1659
1660
1661
1662
1663
1664
1665
1666
1667
1668
1669
1670
1671
1672
1673
1674
1675
1676
1677
1678
1679
1680
1681
1682
1683
1684
1685
1686
1687
1688
1689
1690
1691
1692
1693
1694
1695
1696
1697
1698
1699
1700
1701
1702
1703
1704
1705
1706
1707
1708
1709
1710
1711
1712
1713
1714
1715
1716
1717
1718
1719
1720
1721
1722
1723
1724
1725
1726
1727
1728
1729
1730
1731
1732
1733
1734
1735
1736
1737
1738
1739
1740
1741
1742
1743
1744
1745
1746
1747
1748
1749
1750
1751
1752
1753
1754
1755
1756
1757
1758
1759
1760
1761
1762
1763
1764
1765
1766
1767
1768
1769
1770
1771
1772
1773
1774
1775
1776
1777
1778
1779
1780
1781
1782
1783
1784
1785
1786
1787
1788
1789
1790
1791
1792
1793
1794
1795
1796
1797
1798
1799
1800
1801
1802
1803
1804
1805
1806
1807
1808
1809
1810
1811
1812
1813
1814
1815
1816
1817
1818
1819
1820
1821
1822
1823
1824
1825
1826
1827
1828
1829
1830
1831
1832
1833
1834
1835
1836
1837
1838
1839
1840
1841
1842
1843
1844
1845
1846
1847
1848
1849
1850
1851
1852
1853
1854
1855
1856
1857
1858
1859
1860
1861
1862
1863
1864
1865
1866
1867
1868
1869
1870
1871
1872
1873
1874
1875
1876
1877
1878
1879
1880
1881
1882
1883
1884
1885
1886
1887
1888
1889
1890
1891
1892
1893
1894
1895
1896
1897
1898
1899
1900
1901
1902
1903
1904
1905
1906
1907
1908
1909
1910
1911
1912
1913
1914
1915
1916
1917
1918
1919
1920
1921
1922
1923
1924
1925
1926
1927
1928
1929
1930
1931
1932
1933
1934
1935
1936
1937
1938
1939
1940
1941
1942
1943
1944
1945
1946
1947
1948
1949
1950
1951
1952
1953
1954
1955
1956
1957
1958
1959
1960
1961
1962
1963
1964
1965
1966
1967
1968
1969
1970
1971
1972
1973
1974
1975
1976
1977
1978
1979
1980
1981
1982
1983
1984
1985
1986
1987
1988
1989
1990
1991
1992
1993
1994
1995
1996
1997
1998
1999
2000
2001
2002
2003
2004
2005
2006
2007
2008
2009
2010
2011
2012
2013
2014
2015
2016
2017
2018
2019
2020
2021
2022
2023
2024
2025
2026
2027
2028
2029
2030
2031
2032
2033
2034
2035
2036
2037
2038
2039
2040
2041
2042
2043
2044
2045
2046
2047
2048
2049
2050
2051
2052
2053
2054
2055
2056
2057
2058
2059
2060
2061
2062
2063
2064
2065
2066
2067
2068
2069
2070
2071
2072
2073
2074
2075
2076
2077
2078
2079
2080
2081
2082
2083
2084
2085
2086
2087
2088
2089
2090
2091
2092
2093
2094
2095
2096
2097
2098
2099
2100
2101
2102
2103
2104
2105
2106
2107
2108
2109
2110
2111
2112
2113
2114
2115
2116
2117
2118
2119
2120
2121
2122
2123
2124
2125
2126
2127
2128
2129
2130
2131
2132
2133
2134
2135
2136
2137
2138
2139
2140
2141
2142
2143
2144
2145
2146
2147
2148
2149
2150
2151
2152
2153
2154
2155
2156
2157
2158
2159
2160
2161
2162
2163
2164
2165
2166
2167
2168
2169
2170
2171
2172
2173
2174
2175
2176
2177
2178
2179
2180
2181
2182
2183
2184
2185
2186
2187
2188
2189
2190
2191
2192
2193
2194
2195
2196
2197
2198
2199
2200
2201
2202
2203
2204
2205
2206
2207
2208
2209
2210
2211
2212
2213
2214
2215
2216
2217
2218
2219
2220
2221
2222
2223
2224
2225
2226
2227
2228
2229
2230
2231
2232
2233
2234
2235
2236
2237
2238
2239
2240
2241
2242
2243
2244
2245
2246
2247
2248
2249
2250
2251
2252
2253
2254
2255
2256
2257
2258
2259
2260
2261
2262
2263
2264
2265
2266
2267
2268
2269
2270
2271
2272
2273
2274
2275
2276
2277
2278
2279
2280
2281
2282
2283
2284
2285
2286
2287
2288
2289
2290
2291
2292
2293
2294
2295
2296
2297
2298
2299
2300
2301
2302
2303
2304
2305
2306
2307
2308
2309
2310
2311
2312
2313
2314
2315
2316
2317
2318
2319
2320
2321
2322
2323
2324
2325
2326
2327
2328
2329
2330
2331
2332
2333
2334
2335
2336
2337
2338
2339
2340
2341
2342
2343
2344
2345
2346
2347
2348
2349
2350
2351
2352
2353
2354
2355
2356
2357
2358
2359
2360
2361
2362
2363
2364
2365
2366
2367
2368
2369
2370
2371
2372
2373
2374
2375
2376
2377
2378
2379
2380
2381
2382
2383
2384
2385
2386
2387
2388
2389
2390
2391
2392
2393
2394
2395
2396
2397
2398
2399
2400
2401
2402
2403
2404
2405
2406
2407
2408
2409
2410
2411
2412
2413
2414
2415
2416
2417
2418
2419
2420
2421
2422
2423
2424
2425
2426
2427
2428
2429
2430
2431
2432
2433
2434
2435
2436
2437
2438
2439
2440
2441
2442
2443
2444
2445
2446
2447
2448
2449
2450
2451
2452
2453
2454
2455
2456
2457
2458
2459
2460
2461
2462
2463
2464
2465
2466
2467
2468
2469
2470
2471
2472
2473
2474
2475
2476
2477
2478
2479
2480
2481
2482
2483
2484
2485
2486
2487
2488
2489
2490
2491
2492
2493
2494
2495
2496
2497
2498
2499
2500
2501
2502
2503
2504
2505
2506
2507
2508
2509
2510
2511
2512
2513
2514
2515
2516
2517
2518
2519
2520
2521
2522
2523
2524
2525
2526
2527
2528
2529
2530
2531
2532
2533
2534
2535
2536
2537
2538
2539
2540
2541
2542
2543
2544
2545
2546
2547
2548
2549
2550
2551
2552
2553
2554
2555
2556
2557
2558
2559
2560
2561
2562
2563
2564
2565
2566
2567
2568
2569
2570
2571
2572
2573
2574
2575
2576
2577
2578
2579
2580
2581
2582
2583
2584
2585
2586
2587
2588
2589
2590
2591
2592
2593
2594
2595
2596
2597
2598
2599
2600
2601
2602
2603
2604
2605
2606
2607
2608
2609
2610
2611
2612
2613
2614
2615
2616
2617
2618
2619
2620
2621
2622
2623
2624
2625
2626
2627
2628
2629
2630
2631
2632
2633
2634
2635
2636
2637
2638
2639
2640
2641
2642
2643
2644
2645
2646
2647
2648
2649
2650
2651
2652
2653
2654
2655
2656
2657
2658
2659
2660
2661
2662
2663
2664
2665
2666
2667
2668
2669
2670
2671
2672
2673
2674
2675
2676
2677
2678
2679
2680
2681
2682
2683
2684
2685
2686
2687
2688
2689
2690
2691
2692
2693
2694
2695
2696
2697
2698
2699
2700
2701
2702
2703
2704
2705
2706
2707
2708
2709
2710
2711
2712
2713
2714
2715
2716
2717
2718
2719
2720
2721
2722
2723
2724
2725
2726
2727
2728
2729
2730
2731
2732
2733
2734
2735
2736
2737
2738
2739
2740
2741
2742
2743
2744
2745
2746
2747
2748
2749
2750
2751
2752
2753
2754
2755
2756
2757
2758
2759
2760
2761
2762
2763
2764
2765
2766
2767
2768
2769
2770
2771
2772
2773
2774
2775
2776
2777
2778
2779
2780
2781
2782
2783
2784
2785
2786
2787
2788
2789
2790
2791
2792
2793
2794
2795
2796
2797
2798
2799
2800
2801
2802
2803
2804
2805
2806
2807
2808
2809
2810
2811
2812
2813
2814
2815
2816
2817
2818
2819
2820
2821
2822
2823
2824
2825
2826
2827
2828
2829
2830
2831
2832
2833
2834
2835
2836
2837
2838
2839
2840
2841
2842
2843
2844
2845
2846
2847
2848
2849
2850
2851
2852
2853
2854
2855
2856
2857
2858
2859
2860
2861
2862
2863
2864
2865
2866
2867
2868
2869
2870
2871
2872
2873
2874
2875
2876
2877
2878
2879
2880
2881
2882
2883
2884
2885
2886
2887
2888
2889
2890
2891
2892
2893
2894
2895
2896
2897
2898
2899
2900
2901
2902
2903
2904
2905
2906
2907
2908
2909
2910
2911
2912
2913
2914
2915
2916
2917
2918
2919
2920
2921
2922
2923
2924
2925
2926
2927
2928
2929
2930
2931
2932
2933
2934
2935
2936
2937
2938
2939
2940
2941
2942
2943
2944
2945
2946
2947
2948
2949
2950
2951
2952
2953
2954
2955
2956
2957
2958
2959
2960
2961
2962
2963
2964
2965
2966
2967
2968
2969
2970
2971
2972
2973
2974
2975
2976
2977
2978
2979
2980
2981
2982
2983
2984
2985
2986
2987
2988
2989
2990
2991
2992
2993
2994
2995
2996
2997
2998
2999
3000
3001
3002
3003
3004
3005
3006
3007
3008
3009
3010
3011
3012
3013
3014
3015
3016
3017
3018
3019
3020
3021
3022
3023
3024
3025
3026
3027
3028
3029
3030
3031
3032
3033
3034
3035
3036
3037
3038
3039
3040
3041
3042
3043
3044
3045
3046
3047
3048
3049
3050
3051
3052
3053
3054
3055
3056
3057
3058
3059
3060
3061
3062
3063
3064
3065
3066
3067
3068
3069
3070
3071
3072
3073
3074
3075
3076
3077
3078
3079
3080
3081
3082
3083
3084
3085
3086
3087
3088
3089
3090
3091
3092
3093
3094
3095
3096
3097
3098
3099
3100
3101
3102
3103
3104
3105
3106
3107
3108
3109
3110
3111
3112
3113
3114
3115
3116
3117
3118
3119
3120
3121
3122
3123
3124
3125
3126
3127
3128
3129
3130
3131
3132
3133
3134
3135
3136
3137
3138
3139
3140
3141
3142
3143
3144
3145
3146
3147
3148
3149
3150
3151
3152
3153
3154
3155
3156
3157
3158
3159
3160
3161
3162
3163
3164
3165
3166
3167
3168
3169
3170
3171
3172
3173
3174
3175
3176
3177
3178
3179
3180
3181
3182
3183
3184
3185
3186
3187
3188
3189
3190
3191
3192
3193
3194
3195
3196
3197
3198
3199
3200
3201
3202
3203
3204
3205
3206
3207
3208
3209
3210
3211
3212
3213
3214
|
*** START OF THE PROJECT GUTENBERG EBOOK 60949 ***
Transcriber’s notes:
The text of this e-book has been preserved as in the original,
including inconsistent capitalisation and hyphenation. Archaic and
inconsistent spellings have also been preserved except where obviously
misspelled in the original. A list of spelling corrections and
inconsistencies is appended at the end. Archaic phraseology has been
retained. Incorrect paragraph numbering on p. 17 has been corrected,
and several missing full stops have been inserted.
VETERINARY SCIENCE ASSOCIATION
OF AMERICA
NOTES ON
VETERINARY ANATOMY
BY
CHARLES J. KORINEK, V. S.
_Graduate of the Ontario Veterinary College, in affiliation with
the University of Toronto, Canada. Hon. Member of the Ontario
Veterinary Medical Society. Ex. State Veterinarian for
Oregon. Ex. President of the Oregon State Veterinary Medical
Board of Examiners. Author of “The Veterinarian”. Principal
of the Veterinary Science Association of America. Sixteen
years of Practical Experience as a Veterinary Surgeon._
[Illustration]
PUBLISHED BY THE
VETERINARY SCIENCE ASSOCIATION
OF AMERICA
COPYRIGHT 1917
by the
VETERINARY SCIENCE ASSOCIATION
OF AMERICA
TABLE OF ILLUSTRATIONS.
Page
Plate I. Skeleton of the Horse 7
Plate II. Muscles of the Horse, superficial layer 29
Plate III. Muscles of the Horse, deep layer 31
Plate IV. Digestive and Urinary Organs of the Horse 37
Plate V. Heart and Chief Blood Vessels of the Horse 69
Plate VI. Nervous System of the Horse 73
Plate VII. Structure of the Horse’s Foot 78
TABLE OF CONTENTS.
Page
Chapter I. Osteology, or Structure of the Bones 8
Chapter II. Arthrology, or Study of the Joints 21
Chapter III. Myology, or Study of the Muscular System 28
Chapter IV. Splanchnology, or Study of the Digestive,
Respiratory, Urinary and Generative Systems 38
Chapter V. Angiology, or Study of the Blood-Vascular
System, as the Heart, Arteries, Veins,
Lymphatic Vessels and Glands 67
Chapter VI. Neurology, or Study of the Nervous Tissue,
as the Brain, Spinal Cord and Their Nerves 73
Chapter VII. Aesthesiology, or the Study of the Organs of
Special Sense, as the Ear, Eye and Organs of
Sensation, as Skin, Hair, Foot, etc. 75
Chapter VIII. Embryology or Development of the Foetus 81
PREFACE
There are a number of excellent works on Veterinary Anatomy, and many
of them will amply repay the student for the time taken to master
them, but for quick reference none seem to contain the wants of the
veterinary practitioner and student for which this humble little work
is primarily intended.
It has been my endeavor to briefly describe each organ as found in the
healthy animal; its functions, etc., in a condensed yet complete form.
I am positive that the student or veterinary practitioner will find its
pages highly instructive as well as profitable and interesting.
In compiling this work a few authorities have been consulted and
quoted, while it has not been practical to give individual credit for
the use of ideas and language, a general acknowledgement is here made:
Veterinary Science, Hodgins and Haskett.
Veterinary Anatomy, Strangeways.
CHARLES J. KORINEK, V. S.
DESCRIPTIVE ANATOMY
Osteology is a term applied to that section of descriptive anatomy
which treats of the bones. _Arthrology_, to the consideration of the
joints or modes of union between the bones, while by _Myology_ is
meant the doctrine of the muscular system. _Splanchnology_ treats of
the viscera, _Angiology_ of the circulatory and absorbent systems.
_Neurology_ deals with the nervous system. _Aesthesiology_ with
the organs of sense; while _Embryology_, as before stated is the
consideration of the animal frame at periods preceding its birth.
In this work the various departments are discussed in the order
here given. The structures which are the subject of the first three
divisions are sometimes classed together as the _Organs of Locomotion_;
for bones form the frame work of the body and often act as levers; the
joints connect the bones, permitting more or less motion between them;
while the muscles move the bones, and so produce motion of a part of
the body--or it may be locomotion, or change of situation in the entire
frame.
In the study of comparative Anatomy the terms _analogy_ and _homology_
are frequently met with. Although these words are unfrequently used
indiscriminately, the following differences should be noted. Organs
are said to be analogous when, though differing in structure, they
perform the same function; but when their functions are different,
which, in the broad sense, they correspond in structure or form, they
are said to be homologous. Thus the middle finger of the human hand is
the homologue of the anterior (front) digit of a horse, because they
have the same general structure, and relation to the rest of the limb;
but as the functions they perform are quite dissimilar, they cannot be
termed analogous. Again, the lungs of a mammal are analogous to the
gills of a fish, for, though they differ widely in structure, position
and form, and are therefore not homologous, their ultimate use is
the same--each of them being an apparatus in which is carried on the
process of purifying the blood.
DISSECTION.
Students must dissect as many animals as possible, so as to familiarize
themselves with the _frame work or structure_, and the location of the
_digestive, nervous and blood systems_, as it will aid materially in
the art or process of determining the nature of various diseases.
During cool weather an animal for dissecting purposes can be kept for a
considerable length of time without preservatives
It is well to have a copy of Anatomy at hand when dissecting for
it will show the location of the various organs and explain their
functions.
[Illustration: PLATE I.
SKELETON OF HORSE--AFTER MEGNIN.]
EXPLANATION OF PLATE I
SKELETON OF THE HORSE
1. Skull, or skeleton of the head.
2. Cervical vertebræ or neck bones.
3. Dorsal vertebræ or back bones.
4. Lumbar vertebræ or loin bones.
5. Sacral vertebræ or rump bones.
6. Coccygeal vertebræ or tail bones.
7. Pelvic or hip bones.
8. Sternum or breast bone.
9. Ribs.
10. Scapula or shoulder blade.
11. Humerus or shoulder bone.
12. Radius or bone of the fore-arm.
13. Ulna or bone of the fore-arm.
14. Carpus or bones of the knee.
15. Os Melacarpi Magnus, metacarpal, or cannon bone.
16. Ossa Melacarpi Parva, or splint bones.
17. Proximal Phalanx, os suffraginis, or large pastern bone.
18. Great Sesamoid Bones.
19. Medium Phalanx, os coronæ, or small pastern bone.
20. Distal Phalanx, os pedis, or coffin bone.
21. Os Naviculare, small sesamoid, or shuttle bone.
(This bone can be plainly seen Plate VII).
22. Femur, or thigh bone.
23. Patella, or stifle bone.
24. Tibia, or leg bone.
25. Fibula. (This bone is little developed in the horse.)
26. Tarsus or hock bones.
27. Metatarsus, or os metatarsi magnus.
28. Ossa Metatarsi Parva, or splint bones of the hind leg.
Names of joints placed according to numbers.
I. Shoulder Joint.
II. Elbow Joint.
III. Carpus or knee joint.
IV. Fetlock Joint.
V. Pastern Joint.
VI. Coffin Joint.
VII. Hip Joint.
VIII. Stifle Joint.
IX. Tarsus or hock joint.
CHAPTER I.
OSTEOLOGY.
STRUCTURE OF THE BONES--Bones are hard, yellow-white, insensitive
objects, which form the skeleton and give attachment to soft structures
(muscles, tendons and ligaments); they are of various sizes, forms and
densities. In the limbs the bones are ordinarily more or less long,
circular bodies, with expanded ends, effectually supporting the body,
supplying leverage and attachment for soft structures, and forming the
basis of all joints. Where cavities, such as the cranium, chest, and
pelvic, enclosing the organs requiring protection and support, the
bones tend to assume a flat, expanded form.
Living bone is bluish pink, insensitive, and elastic; on exposure to
air it becomes diseased and blackened, very sensitive and painful; (the
teeth excepted) which are harder and of a higher specific gravity than
any other bone formation.
Bones are composed of two kinds of substance--animal, which makes the
bone tough and flexible; earthy, which makes it hard and fragile.
In young animals the animal matter forms about one-half of the bone
substance; in the adult, it diminishes to about a third, while in old
animals it is still less; hence the bones of very old animals are
brittle, more liable to fracture and harder to mend.
Bones in a six-year-old horse contained, Phosphate of Lime, 54.37 per
cent; Carbonate of Lime, 12.00 per cent; Phosphate of Magnesia, 1.83
per cent; Soluble Salts, 0.70 per cent, or mineral matter, 68.90 per
cent. While they contain Cartilage, 27.99 per cent; Fat, etc., 3.11 per
cent, or animal matter 31.10 per cent.
In bone tissue there are two modifications of texture, the _compact_
and the _cancellated_. The former--hard, dense, and ivory-like, is
always situated externally; the latter porous and spongy lies within.
Although the compact tissue appears uniformly dense, and destitute of
porosity, yet, if we transversely sectate the shaft of a long bone, and
examine it under the microscope, by transmitted light, it is found to
contain numerous round openings. These are called _Haversian canals_.
They transmit bloodvessels, and run in a longitudinal or slightly
oblique direction, opening on either the outer or inner surface of the
bone.
The external (outer) surface of every bone is covered by a tough,
fibrous, inelastic membrane called periosteum, which can be seen by
examining the bone of an animal which has recently died. The only
exception to this is at the joints where one bone articulates with
another, and where a tendon or muscle plays over a bone; here we find
its place taken by articular cartilage. By its strength it sometimes
retains bones in contact after an oblique fracture; in the young, it
is thicker and more vascular than in the adult. Blood-vessels which
penetrate the periosteum pass directly to the bone; the outer surface
of the bone is always studded with numerous _foramina_ through which
these enter.
The periosteum owing to its inelasticity, is, when inflamed, the seat
of intense pain; and should any part of it be stripped off, there is
every probability of the denuded bone dying and separating.
CONTENTS OF BONE.--Red marrow is found in the extremities or near the
ends of bones, white marrow is found in the shaft.
CLASSES OF BONES.--Bones are classed as long, flat, and irregular.
Long or cylindrical bones are found in the limbs or extremities, and
serve as levers and pillars for traveling and to support the body.
Descriptively, a long bone is divisible into a center or _shaft_ and
two ends or extremities. Flat bones are found where visceral organs
need protection. As the shoulder or scapula and ribs, to protect the
heart and lungs; pelvic or hip bones, to protect the rectum and
urinary and genital organs; also the cranial bones, to protect the
vital organ called the brain. Irregular bones are found in the spinal
column and in the joints, such as the knee or carpus, hock or tarsus,
where great strength is required. They usually possess many angles and
indentations, with surfaces for articulation and tendonous attachment,
and consist of a thin, dense, external (outer) case of compact bone
enclosing cancellated tissue. In proportion to their size they present
a much larger extent of articular surface and greater mechanical
strength than any other class.
1. SKULL, or skeleton of the head, the most anterior (forward) part
of the horse’s skeleton, articulates with the first cervical vertebra
(or atlas), from which it is suspended by its posterior (or back part)
extremity, its anterior (forward) extremity being free. Its position
varies with the attitude of the animal; but in our descriptions we
shall always suppose it to be placed in a horizontal position.
In the young animal the skull is composed of a number of bones, all of
which, with the exception of the lower jaw, the teeth, the bones of the
tongue, and ossicles of the ear, become united by ossification (growing
together) in the adult.
In speaking of the different points of importance in connection with
the head bones of the horse, and other animals, suppose a cross or
longitudinal section of the head is made. It will be noticed that it is
full of cavities or sinuses. The uses of these are to lighten the head
and also to warm the air as it passes into the nostrils, on its way
down to the lungs. It will also be found full of foramen or small holes
through which the nerves from the brain and various blood vessels pass
to the organs situated in the head, such as the tongue, lips and the
various glands in and around the head.
Then there are the cavities in which the eyes are situated, one on each
side of the head, called the orbital fossa. In examining this fossa you
will find a small opening or foramen, through which the optic nerve
passes in coming from the brain to the eye. This is the nerve of sight.
Then the most important part of all to consider is the cranial cavity
in which that very important organ is situated called the brain, which
controls all the various functions and movements of the body.
Another important point is the situation of the ear drum. It is
situated in the hardest bone found in the whole skeleton, called the
petrosal. The nerve that gives the function called hearing comes from
the brain down to the petrosal bone and enters by a small foramen or
hole into the drum of the ear to give hearing. This nerve is called the
auditory nerve.
2. CERVICAL VERTEBRAE OR NECK BONES.--These bones are seven in number.
The first and second bones proceeding from the head receive special
names. The first one is called the atlas, from which the head is
suspended and to which it is attached; it somewhat resembles the body
of a bird with wings extended. The second bone receives the name
of dentata. This is the bone which allows the head to turn in any
direction, hence it is sometimes called the axis or pivot of the neck.
Between these two bones, on the upper surface, is the only place where
the spinal cord is not covered with bone, a spot about three-fourths of
an inch in diameter. The next four bones receive no name, and are about
the same in size and length. The last, or the seventh bone, is only
about one half the length of the preceding ones and receives no special
name.
3. DORSAL VERTEBRAE OR BACK-BONE.--Dorsal bones are eighteen in number.
The chief point of interest about them are the height of the spines on
the upper surface of the bones. These large spines form the withers of
the horse, as will be noticed in the skeleton. On either side of these
bones the ribs are attached, 18 pairs corresponding with the number of
bones in this region.
4. LUMBAR VERTEBRAE (or the bones which form the skeleton of
the loins). These bones are six in number, and they are situated
immediately above the kidneys.
5. SACRAL VERTEBRAE OR RUMP BONES.--There are five of these bones in
the young horse, but in the adult they unite as a single bone, somewhat
triangular-shape. These are situated between the upper hip bones and
help to form the rump. Beneath these bones the bladder is situated.
6. COCCYGEAL VERTEBRAE OR TAIL BONES.--These are 18 to 20 in number.
There is no complete canal, like in the previous vertebraes, for
enclosure of the spinal cord.
7. PELVIC OR HIP BONES.--The pelvic bones are flat, but somewhat
irregularly-shaped, and they form the sides, floor, and part of the
roof of the pelvic cavity. Above they are connected with the sacrum,
and below united to each other, in the adult, by ossification (union
of bone). In the young animal, as above stated, and especially in the
foetus, each side consists of three parts, which retain their names of
ilium, ischium, and pubis, even after union by ossification. They all
three meet in the acetabulum, or articular cavity for the femur or hip
bone.
8. STERNUM OR BREAST BONE.--This bone is small and short in the horse
and is situated on the lower surface of the chest cavity. The principle
points to be noted in this bone are that of its softness, and that the
first eight pairs of ribs are attached to it on either side. This bone,
in the horse resembles the keel and cut-water of a boat.
9. RIBS.--In the horse the ribs usually number eighteen on each side.
They extend in a series of arches of varying curvature from the dorsal
vertebrae above, towards the sternum and sides of the abdomen below.
Their shape, in a great measure, determines the conformation or
shape of the thorax or chest cavity; they protect its contents, and
materially aid in its contraction and expansion. They are continued
downward and forward by a small piece of cartilage or gristle, and are
just slightly attached to the breast bone or sternum; these are called
the false ribs. Note that, starting with the first rib, they get longer
until the ninth rib is reached; they then get shorter, the last rib
being only a few inches long.
10. SCAPULA OR SHOULDER BLADE.--The scapula is a flat bone situated on
the antero-lateral (front side) surface of the thorax or chest cavity,
it is triangular in shape, the base being turned upwards. The inferior
or lower extremity articulates with the humerus or shoulder bone.
11. HUMERUS OR SHOULDER BONE.--The humerus is a long bone extending
from the scapula to the radius and ulna in an oblique direction
downwards and backwards. Like all long bones it possesses a shaft and
two extremities and two articular surfaces.
12. RADIUS, OR BONE OF THE FORE-ARM.--The radius is a long bone, and
occupies a vertical position between the humerus and the carpus.
13. ULNA, OR BONE OF THE FORE-ARM.--The ulna is an irregular bone,
in form triangular, with the base uppermost, and is placed on the
supero-posterior or (upper and back) part of the radius.
14. CARPUS, OR KNEE.--The carpus, knee or wrist, as it is incorrectly
termed in quadrupeds, is composed of seven and often of eight small,
irregular bones arranged in two rows of three each, one above the
other, the seventh being at the back of the three in the upper row,
and the eight, when present, in a similar position with respect to the
lower row. Their names are as follows: scaphoid, lunar bone, cuneiform
bone, trapezium, trapezoid, os magnus, unciform, pisiform, which is not
always present.
15. OS METACARPI MAGNUS, METACARPAL OR CANNON BONE.--The large
metacarpal, or cannon bone, is a long, straight bone, placed in a
vertical direction. Its superior or upper extremity articulates with
the carpus and its inferior or lower extremity articulates with the os
suffraginis and the two sesamoids.
16. OSSA METACARPI PARVA, OR SPLINT BONES.--The two small metacarpal,
or splint bones, tuberous in form (marked or covered with projections)
at the carpus and tapering distally (lower portion), are attached to
the large bone, one on each side of its posterior (back) surface, by
ligaments in the young animal, and by ossification (union of bone) in
most grown up, and in all old animals.
17. PROXIMAL PHALANX, OS SUFFRAGINIS, OR LARGE PASTERN BONE.--The os
suffraginis, or large pastern bone, passes obliquely downwards and
forwards, and articulates with the cannon bone above, and the median
phalanx or os coronae below. It belongs to the class of long bones.
18. GREAT SESAMOID BONES.--These bones, two in number, are placed side
by side at the postero-inferior (back and lower) part of the metacarpus
and postero-superior (back and upper) part of the os suffraginis; they
are irregular in shape, their back parts are covered with cartilage,
for the passage of the flexor tendons of the digit or the last four
bones of the limb. Their superior or upper surface is roughened, and
their sides, which are grooved for the reception of the suspensory
ligament. (This is called the fetlock joint.)
19. MEDIAN PHALANX, OS CORONAE, OR SMALL PASTERN BONE.--The second
phalanx, os coronae, or small pastern bone, has no marrow canal, and
belongs to the class of irregular bones. It is inclined, like the os
suffraginis, obliquely downwards and forwards, and is partly covered
by the hoof. The upper portion of this bone articulates with the os
suffraginis and the two sesamoids, the lower part articulates with the
os pedis.
20. DISTAL PHALANX, OS PEDIS, OR COFFIN BONE.--The third, or ungual
phalanx, os pedis, or coffin bone, is an irregular bone situated within
the hoof, and, when in a healthy state, corresponding somewhat to it in
shape, being semilunar in form, with the convexity to the front. This
bone is very hard and porous, having many openings for the transmission
of arteries and veins. In this bone we notice the wall, the sole, the
tendonous surface, the articular surface, and the alae or wings. This
bone articulates with os coronae superiorly (upper) and posteriorly it
articulates with the os navicular bone.
21. OS NAVICULARE.--The naviculare, third or small sesamoid, or shuttle
bone, is an irregular bone, situated with its long axis transversely,
behind and below the os coronae, and behind the os pedis, with both
of which it articulates, the articulation of the three forming the
so-called _coffin_ joint. The lower surface of this bone is important
as it is covered with cartilage, and together form a kind of a pully
over which plays the great flexor perforans muscle. The remaining
portion of the anterior limb or front leg, will be considered under
anatomy of the foot. The navicular bone can be plainly seen on Plate
VII, anatomy of the foot.
22. FEMUR OR THIGH BONE.--The os femoris, femur or thigh bone, the
largest, thickest, and strongest bone in the body, belongs to the class
of long bones, and is placed in a direction obliquely downwards and
forwards, articulating with the cup-shaped cavity in the pelvic or hip
bones superiorly (or upper), and with the tibia and patella inferiorly
(or below). This bone is roughened for the attachment of the powerful
muscles of the hip.
23. PATELLA, OR STIFLE BONE.--This, the kneepan or stifle bone, is
placed in front of the pully-shaped groove of the femur. It is very
compact, its front surface being irregular, round and very much
roughened, for ligamentous attachment, and its posterior (or back)
surface very smooth to articulate with the groove in the femur,
presenting two depressions divided by a ridge, the inner being the
larger, and in the fresh state enlarged still more by projecting
cartilaginous lip, or elevation. The patella increases the power of the
hind leg, and it is this bone that causes stifle joint lameness when it
slips out of the groove in which it glides normally.
24. TIBIA, OR LEGBONE.--The tibia or leg bone, is a long bone, larger
at its upper than its lower end, situated between the femur and the
astragalus, slanting downwards and backwards. This bone is three-sided,
possessing outer, inner and back surfaces, all of which are wider above
than below. This bone gives attachment to the flexor muscles of the hip.
25. FIBULA.--This is a long slender bone, little developed in the
horse, and is an appendage to the tibia, being attached to the outer
side of that bone, and extending from its head to its lower third, to
which it is affixed by a ligament; the space between the two bones is
called the tibial arch. The fibula gives attachment to the peroneus
muscle, the muscle that is supposed to be severely contracted when an
animal is affected with string-halt.
26. TARSUS, OR HOCK BONES.--The tarsus, or hock, corresponding to the
ankle-joint of a man, is composed of six irregular compact bones,
situated between the lower end of the tibia and the superior or upper
extremity of the metatarsus; they are arranged in two series; one
consisting of the cuboid and three cuneiform bones, the magnum, medium,
and parvum, corresponds to the lower row or carpal bones; the other
upper series consists of the astragalus and calcaneum; the first,
forming with the bone above the mobile portion of the joint, may be
said to correspond to the upper row of carpal bones, while the latter,
being the lever bone, corresponds to the trapezium. These bones, like
those of the carpus, are thickly covered with cartilage on their
articular surfaces, which acts as a protection against concussion.
It is these bones that become diseased and united when an animal is
affected with bone spavin. The calcaneum bone which forms the prominent
part, termed the point of the hock, and corresponds to the heel-bone
of man. This bone gives attachment to the calcaneo-cuboid ligament, and
it is this ligament that is sprained or ruptured in curb of the hock.
28. METATARSUS, OR OS METATARSI MAGNUM.--This bone presents the same
general appearance as the large metacarpal or cannon bone, from which
it differs principally in being about one-sixth longer and flattened
from side to side. It is rounded and more prominent in front. This
bone articulates above with the tarsus bones, and its lower portion
articulates with the os suffraginis.
28. OSSA METATARSI PARVA, OR SPLINT BONES OF THE HIND LEG.--These also
present the same general form as the small metacarpal bones of the
front leg; the outer is the longest and largest, and has the largest
head, with two surfaces which articulates with the cuboid bone; the
inner one has also three articular surfaces, two for small, and one for
the middle cuniform bones. The two surfaces on each head articulate
with corresponding ones on the large metatarsal bone. The remaining
bones of the hind extremity, viz.: the three phalanges, with their
three accessory bones, so closely resemble the corresponding bones
of the fore extremity, that it seems at first sight difficult to
distinguish one from the other; the chief differences being, that the
first phalanx of the hind extremity is longer; its upper end larger,
and its lower end smaller, than in the fore extremity.
COMPARATIVE OSTEOLOGY.
In this section we shall endeavor to point out where the skeleton of
the domesticated animals, other than the horse, differ from the typical
skeleton of the latter in any important particular. The following
descriptions are therefore in all cases comparisons, where comparison
is possible, between the typical skeleton and the skeleton in question.
RUMINANTIA.
(Cud Chewing Animal.)
In this class we take the ox as the animal which represents the best
for our purpose, the family of ruminating or cud-chewing animals, as
cattle, sheep and goats.
THE SKULL OR CRANIUM.--In the skull of the ox an important feature is
the development of the frontal or forehead bone, which extends from
below the eyes to the back of the skull, forming the entire forehead
and crest or top, in the middle of which is the forehead tuberosity or
knob-like elevation, which is very large in hornless animals. Springing
from the sides of the top are two processes, varying in size and shape,
but corresponding to the shape of the horns, which they support.
VERTEBRAE.--The true vertebral column is made up of 26 bones divided
into seven cervical or neck-bones, 13 dorsal or back-bones, and six
lumbar or loin bones.
CERVICAL OR NECK-BONES.--The bodies of these bones are shorter than
those of the horse, but same in number.
DORSAL OR BACK-BONES.--These bones are longer than in the horse, but
have the same general form. It gives attachment to 13 ribs on each
side in the same manner as those of the horse.
LUMBAR OR LOIN BONES.--The lumbar vertebrae are longer and thicker than
in the horse, their bodies being more round on the sides and lower
surfaces.
SACRUM, OR RUMP BONES.--The sacrum is larger and more arched, and the
upper surface more round than in the horse.
COCCYGEAL OR TAIL BONES.--The tail bones are from 15 to 20 in number,
are strong and rougher than those of the horse.
PELVIS OR HIP BONES.--The pelvis is larger, but presents the same
general appearance as in the horse.
In studying the bones of the limbs in ruminants, the only point of
difference is in bones below the knee. The large metacarpal bone
presents a vertical groove down its front, which marks the original
division of the bone into two bones. The lower extremity is divided by
a deep groove into two articulations, each resembling the single one
of the horse. The outer one being always the smaller, a rudimentary
metacarpus is placed on the back and outer surface. The pasterns and
sesamoids in either limb are double, one set forming each digit; they
are small and narrow, the coffin bone resembles half of that of the
horse, equally divided.
In the aged ruminant, two bones are commonly found in the heart,
and may be termed the cardiac bones. They are found related with
auriculo-ventricular rings. In shape they present three angles, three
borders, and two surfaces. The left bone is somewhat smaller than the
right.
OMNIVORA.
Animals eating both vegetable and animal food.
The omnivora are represented in veterinary anatomy by the hog.
CARNIVORA.
An order, suborder, or family of mammals, especially wild animals,
including the dog and cat, etc. Animals that eat flesh.
In this order our description alludes mostly to the dog.
AVES.
The scope of the present work will allow only very brief treatment of
this part of the subject.
Of or pertaining to Birds, Poultry, Etc.
The bone tissue or substance of birds are exceedingly compact and hard,
white in color; and some of the bones are pneumatic, or contain air
instead of marrow, notably these are bones of the skull, the sternum
or breast bone, and the upper bones of the limbs. There are a great
variety in the amount of pneumaticity or air possessed by the skeleton
of different species, but it is not necessarily determined by the
flying power of the animal.
CHAPTER II.
ARTHROLOGY.
What is known regarding the articulations of bones of joints. The
several bones which form the skeleton are united by means of certain
soft structures, forming a number of articulations or joints, the study
of which is termed arthrology.
Before considering the different forms of joints, it will be advisable
to describe briefly the various tissue, other than bone, which enter
into and contributes towards their formation. These are chiefly
cartilage, connective and elastic substance or tissues, and fat.
In health, one bone never comes directly in contact with another,
cartilage or fibrous tissue being always interposed; an exception to
this exists in the adult skull, most of the bones of which become
firmly united by ossification of the interposed soft material.
CARTILAGE.--Cartilage, known also by the familiar name of gristle, is
a firm, bluish-white elastic animal substance, somewhat transparent,
resilient, and flexible, possessing great cohesive power. That which
forms the original basis of the bony framework is termed temporary,
and that which persists in the adult, permanent cartilage; the former
disappears as it is replaced by bone, but the latter, of which alone we
have to treat here, never under normal circumstances become ossified.
Cartilage consists of corpuscles or cells, usually embedded in an
intercellular substance.
The articular cartilage is important as it encrusts the articular
surfaces of bones, helping to form joints by supplying smooth, elastic
cushions, which diminishes both concussion and friction.
CONNECTIVE TISSUE (White fibrous tissue.)--In one form or another this
tissue is found in all parts of the body. The chief varieties are the
areolar and the fibrous; the former serving as a connecting medium, and
support to the various organs, and to the structures of which they are
formed. It appears as a loose, transparent mesh, its interwoven bundles
forming spaces termed the areolar or cells.
Connective tissue contains nerves and blood-vessels, for the supply of
neighboring structures as well as for its own nourishment. When healthy
it is little sensitive to pain.
YELLOW ELASTIC TISSUE.--This differs from the white or connective
tissue in being yellow, elastic, and not so tough or strong. Its fibers
are usually large but when mixed with the white tissue in tendons the
size diminishes.
Yellow elastic tissue is found nearly pure in the ligamentum nuchae,
and tunica abdominis, the coats of the largest arteries and elsewhere.
The lungs contain a large quantity of this tissue.
When white fibrous tissue is boiled gelatine is obtained, which is not
the result of boiling the yellow elastic tissue.
ADIPOSE TISSUE.--Fat or adipose tissue consists of cells containing an
oily material, and arranged in isolated groups, or slightly separated
by meshes of areolar tissue. It is found in many parts of the body, and
varies greatly in quantity; in joints it occurs between the ligaments,
and serves the purpose of a packing material, while in the form of
medulla or marrow it occupies the cavities of bones. In fat cell a
nucleus is very rarely visible.
LIGAMENTS.--Ligaments are dense, fibrous, connecting structures. They
exist in most articulations, and are made up principally of white
fibrous tissue. There are two kinds--capsular or bursal, and funicular
or binding ligaments.
Capsular ligaments are membranous structures enclosing true joints.
They consist of a dense interlacement of fibers attached to bones,
round the edges of the articular cartilages; some regard them as a
continuation of the periosteum or the covering of bone. They are never
closely applied, their use being to form cavities round the joints,
enclosing and protecting the synovial or lubricating apparatus inside.
Funicular or binding ligaments consist of rounded or flattened cords,
or bands of fibrous tissue, passing from one bone to another, firmly
attached to roughened portions of their surfaces. They hold the bones
in their places, at the same time allowing the requisite amount of
motion in the joints. Ligaments which are situated between bones are
often termed interosseous. Annular ligaments are those which bind
down and protect the tendons of muscles in certain joints, converting
grooves in the bones into channels or tubes which are lined with
synovial membrane, and through which the tendons play.
Some ligaments are composed almost entirely of yellow elastic tissue,
such as the ligamentum nuchae (the elastic ligament of the neck) and
the ligaments connecting the vertebral arches (arches between the bones
of the spinal column).
SYNOVIAL MEMBRANES.--These are thin membranes lining the capsular
ligaments of joints, or they are interposed elsewhere between
structures which move one upon another, and which would otherwise be
injured by the friction. Near the borders of articular cartilages
the membrane is generally found as a projecting fold, the projection
being due to a small pad of fat, interposed between the membrane and
the capsular ligament. These projections were once erroneously termed
synovial glands, but their use is probably to assist in forcing the
synovia between the opposing surfaces of cartilage.
SYNOVIA.--Synovia or joint oil, is a viscid, transparent fluid,
colorless, or pale yellow, physically resembling oil, but it contains
very little fatty material, consisting chiefly of albumen, salts
and water; it is secreted by the cells of the inner surface of the
synovial membrane. When an animal is in active exertion, there is a
greater demand for joint oil than when at rest, consequently there is
an increased secretion of it.
CLASSES OF JOINTS.--Joints may be divided into three
classes--Immovable, movable and mixed.
MOTION IN JOINTS.--The following terms express the various movements
allowed by joints--_Extension_ tends to bring two bones as nearly into
a straight line as the structure of the joint will permit; _flexion_ is
the reverse of this, and diminishes the angle that extension increases;
_abduction_ expresses the outward movement of a limb or bone from the
central line of the body; _adduction_ is the reverse action; rotation
signifies the partial _revolution_ of a bone or number of bones, as it
were, on their own axis; _circumduction_ implies the movement of the
lower end of a bone or limb, where it describes a curve, as the bow
of a circle, _ellipse_, etc. The term gliding explains itself, and is
peculiar to movable and other joints having no frictional surfaces.
Important points of joints to be considered will be described on Plate
I by Roman figures to correspond with those given below.
JOINTS OF THE FRONT LEG.
I. SHOULDER JOINT.--This joint is formed by the lower end of the
scapula, or shoulder blade, and the upper end of humerus or shoulder
bone. This is a ball-and-socket joint, and is held in its place by
ligaments and muscles. This belongs to the class of rotation joints,
its action is outward, inward, backward and forward.
II. ELBOW JOINT.--This joint is formed by the lower part of the humerus
and the upper portion of the radius and ulna. It is covered with
ligaments and muscles similar to the above mentioned joint. This is
a hinge joint the action of which is only forward and backward, or
flexion and extension. It has no lateral movement.
III. CARPUS OR KNEE JOINT.--This joint has been explained very
thoroughly in the previous chapter. There are three separate articular
surfaces in this joint. This joint has a large capsular ligament,
and has binding ligaments one above the other, the seventh being at
the back of the three in the upper row, and the eight, when present,
in a similar position with respect to the lower row. Thus there are
three separate articular surfaces in this joint. The upper surface of
the carpus articulates with the radius; this forms the hinge joint
of the knee and is where flexion and extension takes place. Another
articulation between the upper and lower carpus bones give slight
motion to the knee, but not so much as the upper articular surface.
Between the lower surface of the lower row of carpus bones and the
upper extremity of the metacarpal, or cannon bones, is another
articular surface which gives motion to the knee. This joint has a
large capsular ligament, and has binding ligaments inside and outside.
In addition to the flexion and extension movement the knee joint
possesses, it also serves as a cushion and relieves concussion of the
fore limb.
IV. FETLOCK JOINT.--This joint is formed above by the lower portion
of the metacarpal, or cannon bone, and below by the part of the os
suffraginis or the large pastern bone. At the back of this joint
there are two small bones called the sesamoid bones. This joint has a
capsular and two lateral binding ligaments. The motion of this joint is
a backward and forward or flexion and extension, same as that of the
knee.
V. PASTERN JOINT.--This joint is situated just above the hoof, and is
formed by the lower end of the os suffraginis or large pastern bone
and the upper end of the os coronae or small pastern bone. It has a
capsular and lateral binding ligaments, same as the fetlock joint. It
also has the same motion as the fetlock joint, and is often the seat of
what is termed a high ringbone.
VI. COFFIN JOINT.--This joint is situated within the hoof. It is formed
above by the lower end of the os coronae or small pastern bone, and
below by the upper surface of the os pedis or coffin bone. Immediately
behind this joint, and articulating with the two mentioned bones, is
the navicular, or shuttle bone--it gets its name from its likeness to
the shuttle of a sewing machine. This bone when diseased is the seat of
navicular disease, or coffin-joint lameness. The action of this joint
is very slight forward and backward.
JOINTS OF THE HIND LEG.
VII. HIP JOINT. The hip joint is formed by the pelvis and femur bone.
This is a true ball-and-socket joint similar to the shoulder joint. It
has a capsular, and is called the round ligament, in the joint, holding
the head or ball on the femur in the socket of the pelvis. This can be
seen plainly by examining the joint. This round ligament is important,
as it often becomes strained, being the seat of hip joint lameness. It
is also held together by the strong muscles of the hip. Its motion is
rotation similar to that of the shoulder joint.
VIII. STIFLE JOINT.--This joint is formed above by the lower end of the
femur, or hip bone, and the upper end of the tibia or thigh bone. These
two bones in front form a pulley-like surface on which the patella,
or stifle bone, is situated. This bone, when the joint is in motion,
glides up and down over the pulley-like surface. It sometimes becomes
displaced, and this is termed dislocation of the patella or stifle
bone. This is an important point to notice about this joint. It has a
capsular and lateral, or binding ligament, which hold the stifle bone
to its place as it plays upon the pulley-like process of this joint.
The action of this joint is only forward and backward or flexion and
extension. It has no lateral or side motion.
IX. TARSUS OR HOCK JOINT.--This joint contains six bones. The two upper
bones, one of which is a pulley-like bone placed in front, and the
other placed behind, forms that portion of the hock which is called
the point of the hock to which the muscles of the gambe are attached.
It can be easily seen or felt. The upper surface of these two bones
articulate with the lower portion of the tibia or thigh bone, and forms
a true articulation of the hock joint. This part is what gives most of
the motion to the joint. Below these are three other small, irregular
bones, placed one upon the other, having an articular surface between
them. Immediately behind these three small bones is what is called the
cuboid bone. This bone also articulates with the three small irregular
bones, helping to form the articular surface of the hock. The lower
articular surface helps to give a small amount of motion to the joint.
This joint is the seat of bone or bog spavin due to severe sprains, or
poorly conformed joints.
Fetlock, pastern and coffin joints of the hind leg are so closely
allied to those of the fore leg that it is not worth while
discriminating between them.
CHAPTER III.
MYOLOGY.
The branch of anatomy which treats of the muscular system is called
Myology.
Muscles are the active organs of motion, or locomotion, each being
separated from the other by a thin delicate membrane made up of
connective tissue, which forms a sheath for the muscle. A muscle is
divided into two parts, viz.: muscular and tendinous. The muscular
part is the larger of the two. It is sometimes called the belly of
the muscle or flesh. This part is known as muscular tissue, and has a
reddish, meaty color. At both ends of the muscle there is a tendinous
part, or the hard, white portion of the muscle which becomes attached
to the bone. All muscles are attached to two or more places of
different bones, and when contracted, the joints of the body are moved.
They are well supplied with nerves, which give strength and feeling,
and also well supplied with small blood vessels, from which the muscle
is fed. Muscles are found in separate groups, all of which have
different actions to perform. There are two kinds of muscles--voluntary
and involuntary. The voluntary muscles are under the control of the
will of the animal; example--the muscles of the head, neck, back, hip
and legs. The involuntary muscles are beyond control of the animal,
and will act even though the animal were asleep--such as those of the
heart, the large muscular curtain which separates the chest cavity from
the abdominal cavity, which is one of the great muscles of breathing;
also the muscles around the chest which assist in breathing.
[Illustration: PLATE II.
MUSCLES OF THE HORSE--SUPERFICIAL LAYER--AFTER MEGNIN.]
EXPLANATION OF PLATE II.
MUSCLES OF THE HORSE.
This illustration shows the superficial muscles of the body after the
skin and panniculus carnosis muscle has been carefully removed. This
muscle is spread over the greater part of the body, which is related
externally with the skin; internally with the superficial layer of
muscles. Its action corrugates the skin, and thus enables the animal
to expel or shake off insects and irritating bodies, its use being
thus protective to some extent; it also supports and binds down the
superficial muscles.
SUPERFICIAL LAYER.
The panniculus and tunica abdominalis are removed.
1. Abducens.
2. Retrahentes muscles.
2´. Attollens maximus.
2´´. Attollens anticus.
3. Temporalis.
4. Nasalis longus.
5. Orbicularis palpebrarum.
6. Levator labii superioris alæque nasi.
7. Dilatator naris lateralis.
8. Orbicularis oris.
9. Zygomaticus.
9´. Buccinator.
10. Depressor labii inferioris.
11. Masseter.
12. Levator humeri.
13. Trapezius cervicalis.
14. Trapezius dorsalis.
15. Latissimus dorsi.
18. Pectoralis parvus.
19. Pectoralis magnus.
20. Sterno-maxillaris.
23. Subscapulo-hyoideus.
24. Antea-spinatus.
25. Teres externus.
26. Postea-spinatus.
30. Caput medium of the triceps.
31. Caput magnum extensor brachii.
33. Extensor metacarpi magnus.
34. Humeralis obliquus.
35. Extensor pedis.
36. Flexor metacarpi externus and medius.
37. Flexor metacarpi internus.
46. Cervical
47. Dorsal serratus magnus.
48. Rhomboideus longus.
49. Superficialis costarum.
50. Splenius.
52. Intercostales.
54. Tensor fasciæ latæ.
55. Triceps abductor femoris.
56. Gluteus externus.
61. Biceps rotator tibialis.
62. Rectus femoris.
63. Vastus externus.
65. Gastrocnemius externus.
68. Flexor pedis perforans.
69. Peroneus.
70. Extensor pedis.
71. Flexor metatarsi.
72. Flexor pedis accessorius.
74. Obliquus abdominis externus.
VOLUNTARY MUSCLES.--The voluntary muscles are in groups. The first
muscle we will call the student’s attention to, after removing the
skin, is the panniculus carnosus (not shown in Plate I), which is a
thin muscle, and almost entirely covering the body, which is sometimes
accidentally removed by a careless person in skinning the animal. The
action of this muscle is to shake the skin when flies or other objects
bother the horse. This muscle is not shown in Plate I it having been
removed in order to show the more important ones.
HEAD MUSCLES.--At the head there is a group of muscles which assist in
chewing, or masticating, the food.
GULLET OR PHARYNX MUSCLES.--Around the throat is another set of
muscles, sometimes called the muscles of the gullet, or pharynx, which
assist in swallowing.
NECK MUSCLES.--The neck muscles are divided into two groups, one on
each side. The action of these is to raise and lower the head, also to
turn the neck and head from side to side.
MUSCLES OF THE BACK.--The muscles of the back are generally divided
into two groups, one above the spinal column and the other below. The
muscles above the spine assist the animal in running, jumping and
rearing. The muscles below the spine are sometimes called the psoae, or
lumbar, muscles, situated below the lumbar bones, or the bones of the
small of the back. The action of these muscles is to assist the animal
in getting up. These muscles are important, for when paralyzed the
horse cannot use his hindquarters. Below these muscles are the kidneys.
HIP OR GLUTEAL MUSCLES.--The muscles of the hip are very large, filling
in around the hip bones. The action of these is much the same as those
of the back, as they assist in jumping, running, rearing and in flexing
and extending the hind leg.
[Illustration: PLATE III.
MUSCLES OF THE HORSE--DEEP LAYER--AFTER MEGNIN.]
EXPLANATION OF PLATE III.
MUSCLES OF THE HORSE.
_Deep Layer._
1. Temporalis.
1. Stylo-maxillaris.
2. Rectus capitis anticus major.
3. Sterno-thyro-hyoideus.
4. Sterno-maxillaris.
5. The Trachea.
6. Scalenus.
7. Splenius.
8. Funicular part of ligamentum nuchæ.
9. Rhomboideus longus.
10. Cervical.
15. Costal serratus magnus.
11. Cartilage of prolongation.
12. Rhomboideus brevis.
13. Transversalis costarum.
14. Longissimus dorsi.
15. Serratus Magnus.
16. External intercostals.
17. Internal intercostals.
18. Rectus abdominis.
19 19. Pectoralis magnus.
20. Postea-spinatus minor.
21. Flexor brachii.
22. Humeralis obliquus.
22´. Caput parvum (of triceps extensor brachii).
23. Extensor suffraginis.
24. Extensor metacarpi magnus divided.
25. Extensor metacarpi obliquus.
25´. Its tendon.
26, 28. Flexor pedis perforans and perforatus.
29. Obliquus abdominis internus.
30. Gluteus maximus.
31. Erector coccygis.
32. Curvator coccygis.
33. Depressor coccygis.
34. Rectus femoris.
35. Vastus externus.
36. Part covered by triceps abductor.
37. Biceps rotator tibialis.
38. Gastrocnemius externus.
39. Plantaris.
40. Flexor pedis perforans.
41. Peroneus.
42. Flexor metatarsi.
43. Extensor pedis (cut across).
TAIL OR COCCYGEAL MUSCLES.--Here there are four that are important,
one situated on the upper side of the tail when straight out, the
action of which is to raise the tail; two, one on each side of the
tail, have the power of drawing the tail to either side; the fourth is
situated under the tail and is the smallest one of the four. Its action
is to draw the tail down.
ABDOMINAL MUSCLES.--The abdominal or belly muscles, are four large,
flat muscles on each side of the abdomen. The outer edge of these
muscles is attached to the outer ends of the false ribs, also to the
processes of the lumbar bones and the outer angles of the pelvic or
hip bones. They unite below to what is called the linea alba, a hard,
white fibrous cord. They pass back in the center of the belly and are
attached to the front of the pelvic bones, called the lower bones of
the pelvic cavity. About ten inches from where it is attached here,
passing forward, is a small slit or hole, which is called the navel,
or umbilical opening. Here the navel vessels pass in and out during
the foetus life, or before the colt is foaled. This is a point of
importance to note, for sometimes at the time of birth this opening
does not close and allows the bowels to come down and form what is
known as umbilical or navel rupture.
Before finishing the description of this group of muscles a very large,
important ligament should be noted, which is found spread all over the
abdomen of the horse. It is of a yellowish color and about one-eighth
of an inch thick, attached in front to the back of the breast bone and
to the pelvic bones behind. This is the first structure seen after
removing the skin from the abdomen. This ligament gives great support
to the organs contained in the abdominal cavity. The action, or uses,
of the abdominal muscles are to support the organs contained in the
abdominal cavity, to flex the back-bone and assist in passing of the
feces. In the mare these muscles assist in parturition, or foaling.
SHOULDER MUSCLES.--The shoulder muscles are very large and powerful.
There are only three of great importance. Two situated on the outside
of the scapula or shoulder blade are important, as they are muscles
affected in the disease called shoulder sweeny. The other important one
is that which passes down over the shoulder joint through the groove
or pulley-like surface on the humerus, or shoulder bone. This is a
long, powerful muscle, attached above to the lower end of the scapula,
or shoulder blade, passing down through the groove mentioned, and is
attached to the upper and front part of the radius or fore arm bone.
Its chief point of importance rests in its action in raising the front
leg, where it passes over the pulley-like surface mentioned, when it
becomes injured or diseased; it is the seat of shoulder joint lameness.
MUSCLES OF THE FRONT LEG.--Muscles of the front leg, from the shoulder
down, are divided into two separate kinds, the extensor and flexor
muscles. The extensor muscles are those which bring the leg forward.
These muscles above are attached to the bones around the elbow joint,
passing down in front of the arm bones. About three inches above the
knee they become changed into the tendinous part of the muscles, or
what is called the cords of the leg. Some of them are attached to the
bones about the knee joint, while others pass over the front of the
joint and are held down to their place by a band or ligament, forming
a loop, as it were, for the tendinous portion of the muscle to glide
into when the leg is in action. Each one of these loops through which
the muscles pass are supplied with a synovial membrane to secrete the
synovia, or oil, which lubricate it during action, the same as in the
joint. This is a point of importance, as sometimes, on account of
injury or strain of this part of the joint through which the muscles
play there may be found a small, puffy enlargement containing oil
secreted by the synovial membrane. This disease is called bursal
enlargement.
The flexor muscles are situated at the back part of the leg, attached
above to the back part of the elbow joint, passing downwards at the
back part of the leg. About two or three inches above the back part
of the knee joint they become tendinous, and from there down to the
back part of the coffin bone, where two of the principle muscles are
attached; these form what is known as the back tendons, or cords, of
the leg. Some of them become attached to the back part of the knee,
same as the muscles on the front part of the leg, while the other two
principal tendons pass through a loop formed by ligaments, the same as
those mentioned in the front part of the knee. In tracing these tendons
down from the knee to the fetlock, notice that they pass through
another larger loop or sheath formed at the back of the fetlock, where
some of the fibers are attached, while others continue down at the back
part of the pastern bones, and are attached to the os pedis or coffin
bone. These tendons are important, because when they are strained the
fact is spoken of as the strain of the back tendons. The action of
these muscles is to flex the leg, bend the knee, pastern joints and
fetlock.
MUSCLES OF THE HIND LEG.--These are also divided into two groups,
extensor and flexor. The extensor muscles are situated in front of
the hind leg. They are attached above, around the stifle joint, and
pass downward in front of the tibia, or thigh bone, one being attached
to the front part of the hock. The other passes through sheaths, or
loops, supplied by a synovial membrane, formed by ligaments, to hold
the muscles firm in front while the leg is in action. In tracing them
down, in front of the shin bone to the fetlock, note that they pass
through loops, or sheaths, and continue down in front of the pastern
bones to where they are attached. The action of these is to bring the
leg forward.
The flexor muscles of the hind legs are attached, above, around the
back part of the stifle joint. In tracing them down it will be found
that they become tendonous. Two of the principal ones pass down to
that part of the hock joint, which forms the point known as the cap.
These form what is called the gambe of the leg, and are partly attached
at the point of the hock, and other part passing down to the fetlock
joint through a loop, or sheath, along the back part of the pastern
bones, and are attached to the os pedis or coffin bone of the foot.
This muscle, from the hock down, forms one of the back tendons of the
hind leg. Another important muscle is found passing down underneath
those already mentioned, through a loop, or sheath, at the back part of
the hock, where it is supplied with a synovia sac. This is a point of
importance, because when it becomes sprained it is the seat of what is
called thoroughpin. It then passes down the back part of the shin bone
beneath the other tendons already mentioned, through the loop at the
fetlock to the back part of the os pedis or coffin bone, where it is
attached. The action of these muscles are to flex or bend the fetlock
and raise the hock joint in traveling.
INVOLUNTARY MUSCLES.--Involuntary muscles, or muscles not under the
control of the will. The first to notice are those of breathing or
respiration. They form a group situated about the chest in such a way
as to be the means of increasing or decreasing the size of the chest
cavity. When these muscles expand the chest cavity is enlarged, causing
the air to rush into the lungs, known an inspiration. On the other
hand, when these muscles contract the air is expelled from the lungs,
known as expiration.
The diaphragm is a muscular curtain which separates the chest from the
abdominal cavity. It also assists greatly in drawing the air in, when
it contracts. This muscle also assists in passing faeces, and in the
mare foaling. It separates the heart and lungs from the bowels, liver
and stomach. This muscle should be carefully examined by the students.
It can be seen by opening any dead animal.
There is one muscle which is both voluntary and involuntary. It is
situated in the penis, surrounding the urethra, or the tube, which
carries the urine from the bladder to the penis in the male animal.
Its action is voluntary while the animal is passing urine. It is
involuntary during sexual intercourse, forcing the semen down through
the penis.
[Illustration: PLATE IV.
DIGESTIVE AND MALE URINARY APPARATUS OF A HORSE--AFTER MEGNIN.]
EXPLANATION OF PLATE IV
DIGESTIVE APPARATUS OF THE HORSE
1. Mouth.
2. Pharynx.
3. Œsophagus.
4. Diaphragm.
5. Spleen.
6. Stomach (left sac).
7. Duodenum.
8. Liver (upper extremity).
9. Great colon.
10. Cæcum.
11. Small intestine.
12. Floating colon.
13. Rectum.
14. Anus.
15. Left kidney and ureter.
16. Bladder.
17. Urethra.
_a._ Hard palate.
_b._ Tongue.
_c._ Soft palate.
_d._ Trachea.
_e._ Pulmonary artery (divided).
_f._ Heart.
_g._ Posterior aorta.
CHAPTER IV.
SPLANCHNOLOGY.
Study of large interior organs in any of the four great bodily cavities
especially those in the abdomen.
DIGESTIVE ORGANS OF THE HORSE.--The digestive organs comprise the
alimentary canal and the accessories by which the alimentary matter is
received and subjected to specific actions, which adapt it for purposes
of nutrition. Digestion, therefore, embraces the collective operations
and changes which the food undergoes in the alimentary canal.
The whole digestive track from the mouth to the anus which is situated
just below the tail, is sometimes called the alimentary canal.
The mouth is an oval cavity at the commencement of the alimentary or
digestive canal. In front of the mouth are the lips, one above and
one below; at the sides are the cheeks. The mouth is lined with what
is known as the mucous membrane, in which are several small openings,
from the glands, which are situated about the mouth. Through these the
saliva is poured. On the upper part of the mouth the mucous membrane
is thrown into ridges, or folds, from 18 to 20 in number. This is a
point of importance in connection with bleeding a horse with lampas. It
is never safe to bleed back of the third bar because there is a large
artery which runs down through the roof of the mouth and enters the
hole in the bone just before it reaches this bar. The tongue, which has
the chief nerves of the sense of taste, is situated in the mouth; this
organ also has a very important part to perform in masticating the food
and mixing it with saliva. The teeth, which also take a very active
part in the masticating of food, are dealt with under the heading of
“Teeth.”
SALIVARY GLANDS.--These glands secrete the saliva that is poured into
the mouth while the animal is eating. There are only three pairs of
much importance. One large pair, one on each side of the throat below
the ears, known as the parotid glands, fill up the space between the
jaw bone and the neck. This pair has tubes passing around and under the
lower jaw and up into the cheek muscles entering the mouth opposite
the fourth molar tooth. These tubes, known as steno’s ducts are about
an eighth of an inch in diameter and convey the saliva from the
glands into the mouth. The next pair are situated under the pair just
mentioned. Their tubes enter into the bottom part of the mouth. The
third pair are situated under the tongue, one on each side. They pour
their secretion into the mouth by several small openings near the front
under the tongue. This can be seen by examining the under surface of
the tongue closely. This is very important fluid in connection with the
digesting of the food.
The gullet is a cavity situated just back of the mouth. It is chiefly
made up of muscles which perform the act of swallowing the food. It is
lined with the continuation of the mucous membrane of the mouth.
The oesophagus, or the continuation of the gullet, is a tube extending
from the gullet to the stomach, and is used to convey the food to
that organ. It is made up of two coats, the muscular and the mucous.
The former contains fibers which, when once the food enters the tube,
contract behind it, forcing it down to the stomach. Its lining is a
continuation of the mucous membrane of the mouth and the gullet. In
tracing the oesophagus or tube down the neck from the gullet, note
that it passes down the left side of the neck, entering the thoracic,
or chest cavity, between the lungs over the heart through the large
muscular curtain known as the diaphragm, then enters the stomach an
inch or two after passing the diaphragm.
THE STOMACH.--This organ is very small in the horse in comparison with
that of the ox. It holds only about four gallons, and is situated
just back of the curtain which separates it from the lungs. It lies
mostly to the left side. The walls of the stomach being composed of
three coats. That on the outside is called the serous membrane, a name
applied to membranes which line closed cavities, such as the abdominal
cavity. The inside lining is a continuation of the mucous membrane
lining the organs before mentioned. The lining in the left part of
the stomach or the part where the food is prepared for digestion is
the same color as that of the mouth. The lining of the right part
of the stomach, which is the true digestive part, is of a deep red
color resembling velvet, and when placed under a microscope has the
appearance of a honey-comb. When the stomach is empty this membrane is
thrown into loose folds. Several small openings may be noticed through
which the gastric juice and pepsin from the glands, situated in the
walls of the stomach, enters. These are very important fluids as they
assist greatly in digesting the food. The third coat is known as the
muscular coat already mentioned. Its action is to give the stomach a
churning motion, rolling the food around and mixing it with the juices.
The opening to the stomach is guarded by a valve which prevents the
food from passing back through the gullet. There is also a valve at
the opening of the bowels, preventing any coarse, undigested food
from entering them. The stomach is held in its place by five large
ligaments, and is well supplied by blood-vessels and nerves. Digestion
of the food takes place very quickly in the horse in comparison to
other animals. Frequently a change of food or working too soon after
eating will interfere with the digestion, thus setting up what is known
as indigestion. This is a very painful disease in the horse. After the
food is acted upon by the juices in the stomach it changes into what
is known as chyme, which passes into the bowels.
THE BOWELS.--They are divided into two parts--the large and small.
The small bowels are 72 feet in length, about one inch in diameter,
and are made up of three coats, same as the stomach. The serous coat
on the outside contains small glands which secrete an oily material
to lubricate the outside of the bowels, which comes in contact with
the inner wall of the abdominal cavity. The muscular coat, made up of
muscular fibers, is situated between the other two coats, the same
as in the stomach; its action is to contract the bowels, giving them
motion to convey the food along through them. The mucous coat is a
continuation of the mucous coat of the stomach. Along this coat are
found small glands known as villi lacteal; these absorb the nourishment
from the food as it passes along through the bowels and pours it into
the blood. The small intestines or bowels are attached on the upper
side to what is known as the mesentery, which is attached above to the
roof of the abdominal cavity. It can be seen in any of the smaller
animals upon examination. About six inches from the stomach, in the
bowels, are found two openings. One of these receive the hepatic duct,
a tube for the purpose of carrying the bile from the liver to be poured
in on the food as it passes through the bowels. The other opening is
for the duct of the gland known as the pancreas. It secretes a clear
fluid known as the pancreatic juice. These juices act on the food
in the first part of the small intestines, changing it into chyle.
After this, the action of the rest of the intestines is to absorb the
nourishment out of the food as it is passing back. The small intestines
and stomach, when in a healthy condition, should be found empty one
hour after food has been eaten. The small bowels or intestines are
situated mostly on the left side just behind the stomach.
The large bowels have three coats, the same as the small ones. The
first part of the large bowels is known as the blind bowel or caecum,
and is about three feet in length; this is generally the first thing to
protrude when opening a horse’s abdominal cavity. Its use is to act as
a reservoir to hold the water and fluid of the food; from this organ
the water and fluid parts are mostly taken up into the system. The next
part of the large bowels is known as the large colon; it lays along
the floor of the abdominal cavity, is about nine feet in length and
is doubled on itself three times. In this bowel the solid part of the
food is found. Here digestion is brought about by the contraction and
expansion of the muscles of the bowel and the nourishment taken from
it, after which it is worked back out of this bowel and enters what is
known as the floating colon. This is about ten feet in length and about
two inches in diameter, or double the size of the small bowel. It is
thrown into folds or pleats, and as that portion of the food containing
no nourishment passes through it is worked into balls which pass back
and are emptied into the rectum or back bowel. This is situated at the
back part of the abdominal and pelvic cavity back of the small bowels
or intestines and like them, is suspended by a fold paritoneum.
The rectum or back bowel is sometimes known as the straight bowel.
It is about 18 inches long and forms the last part of the bowels or
intestines. Its coats are a continuation of those of the large bowel,
but each is thicker and heavier. Above this bowel are the bones of the
sacrum, below it, in the horse the bladder and other small glands.
Below the rectum of the mare are situated the womb and the vagina, the
latter being the passage into the womb from the outside. The bones
which help to form the pelvic cavity are situated at the sides, and
at the back immediately under the tail is what is known as the annus.
The use of the rectum is to hold the balls as they pass back from the
floating colon. When the rectum becomes so full that there is pressure
on the sides of the wall thus stimulating the nerves the muscular coat
contracts and forces the contents back towards the annus. At the same
time the muscles of the annus dilate, causing the faeces to pass out.
ACCESSORY ORGANS OF DIGESTION.--In the abdominal region these organs
are the Liver, the Pancreas, and the Spleen.
The Liver is the largest gland in the body and is situated between the
stomach and the diaphragm. The liver of the horse weighs from ten to
twelve pounds. It is of a dark brown color, well supplied with blood
and nerves, and is held in place by several strong ligaments. A bitter,
greenish colored fluid called the bile is secreted from the liver and
emptied into the digestive system, where it plays an important part
in its action on the food. There is no gall bladder in the horse, but
simply a tube passing from the liver to the small bowel into which it
empties the bile about six inches back of the stomach. It is important
to note that it is in this tube that gall stones sometimes collect.
The Pancreas is another very important gland. It is of a grayish, fatty
color, and may be found near the roof of the abdominal cavity in front
of the kidneys. The Pancreas secretes a clear, colorless fluid called
the pancreatic juice. This fluid, like the bile from the liver, also
plays an important part in the digestion of food. It is carried down
from the Pancreas by a duct or tube emptying into the small bowel just
back of that of the liver.
The Spleen is a long, flat gland about fifteen inches in length,
situated along the left side of the stomach and to which it is closely
attached. It is of a grayish red color and feels quite soft. It is
ductless, there being no secretion passing from it. The function of
this gland is not clearly understood, but by many it is supposed to
regulate the temperature of the stomach during the process of digestion
and to act as a reservoir for the blood. Some speak of it as the
burying ground of the red corpuscles. It is well supplied with blood
vessels and nerves and weighs about two pounds in an average sized
horse.
RUMINANTIA.
DIGESTIVE SYSTEM.--The lips of an ox are thick and hard. The upper has
no hair on it and varies in color with the color of the animal. When
cattle are in good health this space is always moist.
The cheeks on the inside are covered by many small rough processes,
which give them a very rough appearance.
The Tongue of the ox is stronger than that of the horse and is more
movable. It is very thick and heavy at the back, pointed at the front
end and the upper part of it is very rough. It is by means of the
tongue that the ox takes most of the food into the mouth.
The Salivary glands are similar to those of the horse.
The Teeth differ very much from those of the horse.
The ox has no front teeth in the upper part of his mouth, their place
being taken by a pad of cartilage or gristle. This pad takes the place
of the upper row of front teeth. The lower row of teeth press against
it when the animal is cropping grass. This accounts for the fact that
cattle do not do as well on short grass as horses.
The front teeth in the lower jaw also differ from those of the horse.
They are eight in number, chisel-shaped, and are loosely set in the gum.
The molars, or back teeth, are similar to those of the horse, only
they are smaller and not so smooth on their upper surface. The ox has
twenty-four molars or back teeth, and eight incisors or front teeth,
making thirty-two in all.
The Gullet of the ox is much larger than that of the horse.
The Oesophagus or tube, which carries the food down from the mouth to
the stomach is well developed, the fibers in it being very strong and
possessing a double action. When the animal is eating they carry the
food from the mouth to the stomach, and when chewing the cud they act
the very opposite, carrying the food from the stomach back into the
mouth.
THE STOMACH.--The student will do well to give some time to the study
of this important organ of the ox, as it is very frequently the seat of
disease.
The stomach has a capacity of fifty-two gallons and is divided into
four separate and distinct compartments: the Rumen or Paunch, the
Reticulum or Honey Comb, the Omasum or Many-plies, and the Abomasum or
the true Stomach. In the first three of these the food undergoes a sort
of preparatory process, while in the fourth the process of digestion is
complete.
The Rumen or paunch is very large, and in an aged animal fills
three-quarters of the abdominal cavity. It lies up against the left
side of the wall of the abdomen, where it is attached and held to its
place by the ligaments. Its situation being an important matter, as
many diseases of the rumen, or paunch, are first noticed on the left
side. Tapping for bloating is always done on the left side because
of this fact. The walls of the paunch of an ox resemble those of the
stomach of the horse, but are not so sensitive, and stand a great deal
of abuse before inflammation sets in. The paunch has two openings, both
of which are at the front; through one the food enters, while through
the other it passes out into the next division.
The Reticulum, or honey comb, is the smallest division and resembles a
honey comb in appearance. This part has little to do with preparing the
food. It is provided with two openings, one in front, where the food
enters, the other at the back, where it passes through into the third
division. In the reticulum, or honey comb, the food is softened further
by the water that the animal drinks which passes directly into the
second division. The food is here pressed into balls and prepared to be
forced back into the mouth to be further masticated.
The Omasum, or many-plies, is the second largest division of the
stomach. When full it is ovoid in shape. It is placed just behind the
second division and at the right side of the paunch. The inside is full
of folds, or layers of membrane, into which all the coarse parts of the
food pass and roll about until it is fine and well prepared to pass
into the last division. When this part of the stomach becomes deranged
and the food becomes dry and hard between the folds, the disease called
impaction of the many-plies, or dry murrain, is the result.
The Abomasum, or fourth stomach, is the true digestive part. In it the
food is completely digested. The walls are redder in color than those
of the three first divisions and contain the glands which secrete the
acids and gastric juices. This stomach has two openings, one through
which the food enters and the other through which it passes into the
small bowels.
The Bowels, or intestines, of the ox are divided into large and small
bowels. This, together with their structure and action resembles that
of the horse. The small bowels are only half the size of the horse,
being about one-half inch in diameter, and about one hundred and fifty
feet in length. The large bowels are not nearly so long as those of the
horse and are thirty-five feet in length.
The Liver of the ox resembles that of the horse, except that it is
provided with a gall bladder which resembles a pear in shape. This acts
as a sac in which to store the gall during the time it is not required
in digestion. When digestion is going on the wall of the gall sac
contracts and forces the gall down to the food. The other glands, the
pancreas and spleen, resemble those of the horse. The juices from these
glands have the same action in cattle as they have in the horse.
RUMINATION, OR CHEWING THE CUD.--Food when first taken into the mouth
of a ruminant is but lightly masticated and mixed with the saliva from
the salivary glands, after which it is swallowed, passing through the
oesophagus into the rumen or paunch (first stomach). This division
acts as a reservoir or storehouse for food thus eaten quickly. When
the animal has time, so to speak, he lies down or stands quietly and
completes the process of mastication of his food by chewing the cud.
This peculiar act is performed as follows: After being softened and
moistened by warmth, the food passes from the rumen or paunch into
the second division--the reticulum, honey comb or second stomach. In
this small globe-like compartment the food is moistened and compressed
into pellets--the cud. By a peculiar reverse action of the oesophagus
or gullet these pellets are taken back into the mouth for further
mastication or chewing. When re-mastication is completed it is again
swallowed, but this time it passes directly into the omasum, many-plies
or third stomach, and thence to the fourth or true stomach.
PROCESS OF DIGESTION AFTER RUMINATION.--Following rumination which,
strictly speaking, is the first step in the digesting process, the food
passes into the third stomach. The fine parts pass right along to the
fourth stomach while the coarser parts are drawn between the folds of
the membrane in this division and worked about until it is fine and
ready to pass into the fourth stomach, where it becomes fully digested
by the action of acids and gastric juices which are secreted in this
part. It then passes into the small bowels, and is acted upon by the
bile from the liver and the pancreatic juice from the pancreas. These
juices are emptied into the first part of the small bowels through
little ducts or tubes, which lead from the glands down to the bowels,
just on the same principle as that of the horse. After this, throughout
the rest of the bowels, the nourishment of the food is taken into the
system by means of little glands which are situated in the coating of
the bowels. The nourishment when once in the blood goes to supply the
different parts of the body, while the part containing no nourishment
or undigested passes off through the back bowels in the form of feces.
TEETH.
Teeth are objects situated in the upper and lower jaws. They are made
of the same tissues as bone but contain 10-1/2 per cent. more of earthy
salts. This fact accounts for their extreme hardness. Unlike bone they
can stand exposure to air and friction without becoming diseased. Teeth
are used to masticate or chew the food, and because of the constant
change in their formation and appearance they serve as a guide in
telling the age of the horse.
There are three hard structures that enter into the formation of the
teeth--Dentine or Ivory, Enamel, and Cementum, or Crusta Petrosa.
The Dentine, or ivory, is situated in the upper part around the pulp or
nerve cavity. It is of yellowish color and largely supplied with nerves
which pass through it from the pulp cavity.
The Enamel is the hardest substance of the tooth, and covers the
outside of all the exposed part. This substance is characterized by its
whiteness and, unlike the dentine, contains no blood vessels or nerves.
If part of the enamel is broken off it is never replaced, and the tooth
below the broken part generally becomes decayed.
The Cementum, or crusta petrosa, is found in the fang or root and the
parts situated below the gum. It is the softest part of the tooth.
Teeth may be simple or compound. Simple as in the dog, where the entire
exposed surface is covered by a solid cap of enamel, which alone is in
wear; compound or complex, as in the horse, where various tissues are
in wear. A tooth consists of the following anatomical parts: The body,
or crown, that part above the gum; the table, the part that comes into
wear on the top; the neck, the part to which the gums are attached; and
the fangs or roots, the parts situated down in the bone.
There are three kinds of teeth found in the horse, the incisors, the
canine and the molars.
The Incisors, or front teeth, situated in the front part of the mouth
just inside the lips, are twelve in number, six above and six below.
The Canine, or bridle teeth, are often absent in the mare. They are
four in number, two in the upper and two in the lower jaw, one on
each side about two inches back from the incisor teeth. They are from
a quarter to three-quarters of an inch above the gum, are round and
pointed and of no particular use. They resemble the eye teeth of other
animals.
The Molars, or back teeth, are twenty-four in number, six on each side
in the upper and six on each side in the lower jaw. With these the food
is ground and masticated.
Wolf Teeth are two small, round, pointed temporary teeth which vary in
size in different animals, situated one on each side in front of the
molars or back teeth in the upper jaw.
TEMPORARY AND PERMANENT TEETH.--The horse has two sets of teeth. The
milk are temporary and are those that the colt sheds; while those that
come in and remain without being shed are called the permanent teeth.
The cutting of the teeth of the foal varies, but at or within nine
days after birth he has four front teeth, two in the center above and
two below, and in the back part of the mouth twelve molars. At from
seven to nine weeks four more incisors or front teeth appear, one at
each side of the two center teeth in each jaw. At nine months he gets
the last of his milk or temporary teeth, these being the corner teeth,
two in the upper side and two in the lower side of the jaw. At this
time he has his full set of milk temporary teeth, consisting of twelve
molars or grinders and twelve incisors or front teeth, six above and
six below, making twenty-four in all. As the colt advances in age he
sheds all these teeth. He then commences to get permanent teeth. When
the age of one year is reached, four permanent molars appear, two in
each jaw, one on each side, behind the three temporary teeth. At two
years of age he gets four more back molars, one on each side of each
jaw. When the age of two years and nine months is reached the two
middle teeth of the temporary incisors, or front teeth of each jaw
fall out, and are replaced by two permanent incisors in each jaw; thus
at the age of three years these four permanent incisors are up and in
wear. At this age, the first eight molars, two on each side of each
jaw, are shed and replaced by eight permanent molars. At four years of
age he sheds four more front or incisor teeth next to those shed at
three years, and these are replaced by four more permanent incisors or
front teeth. At this age, too, he sheds the four remaining temporary
molars, or grinders, which are replaced by four more permanent molars.
He also gets four more permanent molars at the back of the mouth. Thus
at the age of four years the colt has a full set of permanent molars,
consisting of six on each side of each jaw, making twenty-four in all.
At five years of age he sheds the four remaining temporary incisors or
front teeth, which are replaced by four permanent incisors, known as
the corner teeth. It is important to become familiar with the time at
which the colt sheds his different teeth, for sometimes the caps or
shells of the teeth do not fall off when they should. These should be
watched, for they greatly interfere with feeding and should be removed
with forceps. At five years of age the canine or bridle teeth make
their appearance; thus at the age of five years the colt has all his
teeth or what is known as a full mouth.
The following table shows the various changes taking place in the
mouth of the horse from the time of birth up to the age of five years:
Hence the horse has--
--Incisors-- Canine --Molars--
Age Temporary Permanent Temporary Permanent
At or soon after birth 4 0 0 12 0
9 weeks 8 0 0 12 0
1 year 12 0 0 12 4
2 years 12 0 0 12 8
3 years 8 4 0 4 16
4 years 4 8 0 0 24
5 years 0 12 4 0 24=40
The table given below indicates the various changes which occur in
the mouths of ruminants, and more particularly in the mouth of the ox:
RUMINANTS
--Incisors-- --Molars--
Age Temporary Permanent Age Temporary Permanent
At or soon after
birth 4 0 12 0
2 weeks 6 0 1 year 12 4
3 weeks 8 0 2 years 8 12
2 years 6 2 3 years 4 16
3 years 4 4
4 years 2 6 4 and 5 years 0 24=32
5 years 0 8
A table giving the number and variety of teeth as they occur in
the domestic animals and in man may be serviceable as one of handy
reference, and is herewith appended:
Incisors Molars Canine Bicuspid Total
4 6 2 4
Man -- -- -- -- = 32
4 6 2 4
6 12 2 0
Horse -- -- -- -- = 40
6 12 2 0
0 12 0 0
Ox -- -- -- -- = 32
8 12 0 0
6 12 2 0
Dog -- -- -- -- = 42
6 14 2 0
6 14 2 0
Pig -- -- -- -- = 44
6 14 2 0
6 8 2 0
Cat -- -- -- -- = 30
6 6 2 0
The dental formula of the ox is the same as that of all ruminants.
RESPIRATORY SYSTEM OF THE HORSE.
ORGANS OF RESPIRATION.--By the action of these organs certain chemical
and physical changes take place in the blood, the chief of these
consisting of absorption of oxygen from, and giving off carbonic
acid to the atmospheric air, the former change being necessary for
the elaboration of the fluid, the latter for the elimination of a
substance which, if retained, would prove injurious. The organs of
respiration are invariably adapted to the wants of the animal and the
medium in which it lives. Thus insects breathe by air-tubes, opening
on the surface of the body; in the oyster breathing is performed by
fringes; in fishes by gills; in the mammalia by means of elastic
air-receptacles, called lungs, which are enclosed in special cavities,
and communicate with the atmosphere by means of an air-tube.
In the horse, who breathes only through his nose, the organs of
respiration are the nostrils, nasal-chambers, larynx, trachea, and in
the thoracic cavity, the bronchi, bronchial tubes, and the lungs.
NOSTRILS.--The nostrils are two openings, one on each side of the nose.
They are held open by the aid of cartilage and muscles. About one and
one-half inches up the nostril on the under side is a small opening
about the size of a grain of shot. Through this opening the duct or
tube which carries the tears down from the eyes empties into the nose.
The nostril is lined with a thin, delicate skin which changes into
mucous membrane as it passes up into the chambers of the head.
NASAL CHAMBERS.--These give passage to the air from the nostril into
the larynx. There are two of these chambers, divided in the center by a
thin partition or cartilage called the septum nasi. These communicate
with the sinuses of the head. The horse cannot breathe through the
mouth on account of the formation of the throat, and this compels him
to always breathe through the nostrils. This is a point which should be
remembered.
LARYNX.--This is a cavity made of cartilage. It gives passage to the
air and also the organ of voice. It is situated in the floor of the
gullet. This cavity has an opening on its upper side, guarded by a
valve, which is always opened except when the animal is swallowing
food or water. When the food is being swallowed it passes over the
valve which closes the opening while the food passes over it. This is
important, for if the valve does not close properly, thus allowing
either food or water to drop into the windpipe, the animal will have a
fit of coughing. This is sometimes referred to as “the food going down
the wrong way.” On the outside of this cavity of cartilage are found
several small muscles which help to hold it in its place. It is lined
inside by a continuation of the same membrane as that of the chambers
of the head. The vocal cords which come into play when the animal is
whinnying are found along the inside of this cavity. These cords are
not nearly so well marked as in the human being, and if they or the
cartilage of the larynx become affected it generally gives rise to the
disease called wind-broken or roaring.
TRACHEA (Windpipe).--This is a tube which conveys the air down from
the larynx to the bronchial tubes in the lungs. It is made up of forty
or fifty rings of cartilage which are united to each other by strong
elastic ligaments. They give to the windpipe its flexibility, that is,
the power to bend in any direction almost like a piece of elastic. From
the larynx the windpipe enters the chest where it terminates into two
small tubes, one going to the right lung and one to the left. These are
called the bronchial tubes.
BRONCHIAL TUBES AND AIR CELLS.--These are made up of the same material
as that of the windpipe, but are only about half the size. After
passing into the substance of the lungs they break up into small tubes
which pass all through the lungs and terminates into what is known as
the air-cells. These small tubes and air cells are lined inside by a
very thin mucous membrane, a continuation of the membrane lining the
other organs already mentioned. Just inside this thin mucous membrane
is found the capillary network of the lungs, and while the blood is
slowly passing through this network of vessels it gives off to the air
in the air cells carbonic acid gas and takes in the oxygen from the
pure air while it is in the lungs.
LUNGS.--Lungs are the most important organs of respiration. They are
spongy, yellowish organs, two in number, one situated on the right
side, and the other on the left. The right lung is the largest because
of the left one having a hollow in its side for the heart. The lungs
are separated by a partition known as the mediastinum, by the heart
which is in the folds of this partition, and also the large blood
vessels and oesophagus. They are made up of light elastic tissue and
are full of air cells and tubes. While the animal is alive they are
very large and fill up nearly the whole chest cavity, but after death
they collapse and are not nearly so large. Between the lungs and
the ribs is found a serous membrane called the pleura or the lining
membrane of the chest. It is made up of two folds, one being attached
around the outer surface of the lungs, while the other is attached to
the ends of the ribs at the side and at the back to the large curtain
which separates the lungs from the bowels. The little glands situated
in this membrane secrete an oily fluid which serves to lubricate these
parts while the lungs are working in the chest so as not to cause
friction. When this membrane becomes inflamed from a chill or injury it
sets up the disease called pleurisy.
RESPIRATION.--The number of respirations per minute varies with the
different classes of animals; as a rule, the larger the animal the
slower the respiration.
The horse 8 to 10
Cattle 12 to 15
Sheep and goats 12 to 20
The dog 15 to 20
Swine 10 to 15
The rate of breathing is increased from the process of digestion
immediately after eating, or may increase from exercise.
RESPIRATORY SYSTEM OF THE OX.
The nostrils are narrow and capable of little dilation compared to
those of the horse. The nasal chambers differ chiefly in there being
an additional turbinated bone. The nasal chambers communicate with the
mouth, therefore cattle can breathe through the mouth to a certain
extent. The larynx is simpler in construction, the true vocal cords
being only slightly developed. The trachea or windpipe presents no
important variation. We may note the presence of a third bronchus,
which passes to the right lung to supply a lobe which is wanting in the
horse.
The Thoracic Cavity is relatively smaller in the ruminants, and the
pleurae present a very important deviation from the arrangement found
in the solipede--viz., the back or posterior mediastinum is imperforate
and strong, completely separating one pleural sac from the other. This
arrangement exists in all the domesticated mammals but the solipede.
The left lung is divided into two lobes, the right into four, the front
one recurving over and almost covering the front of the heart. The
interlobular or cellular tissue is exceedingly thick, the separation
between the lobules being distinctly visible. This arrangement explains
perfectly the special nature of pneumonic lesions in the large
ruminants.
URINARY SYSTEM OF THE HORSE.
The organs of this system secrete the urine from the blood, and excrete
or expel it from the body. These organs are chiefly the kidneys,
ureters, bladder and urethra. The urine, which is a watery fluid, is
secreted by the kidneys, and carried off by their ducts, the ureters,
to a special reservoir, the bladder, where it accumulates and from
which it is finally expelled at intervals through the urethra.
The kidneys are two compound tubular glands, one on the right side and
one on the left side, and are situated just below the small of the back
(sublumbar region), the right one being the fartherest ahead. In shape
they are long and narrow and resemble the liver in color. In cutting
one of the kidneys open, it is found to be full of glands and tubes,
which secrete the urine from the blood while it is passing through the
kidneys. These tubes pass to the center of the kidneys, where they
empty the urine into what is called the pelvis. The glands are largely
supplied with blood vessels and nerves. The use of the kidneys are to
secrete the urine from the blood, which contains a large amount of what
is known as ureaic acid, and if not taken out of the blood by these
glands, acts as a poison to the system.
The Ureters are tubes which carry the urine down from the pelvis of
the kidney to the bladder. They are two in number, one situated on the
right side of the pelvic cavity and the other on the left side, close
to the walls--they enter on each side at the upper surface of the
bladder. They are only about one-sixteenth of an inch in diameter.
The Bladder is situated in the pelvic cavity. When it is full it
sometimes stretches out into the abdominal cavity. It consists of a
body and neck. The body is the large part, and is placed in front;
the neck being at the back part of the bladder. This is where the
urine passes out of the bladder. The bladder is made up of three
coats, somewhat similar to that of the bowels. The serous coat is a
continuation of the serous coat found in the abdominal cavity lining
the bowels. The inside is lined with mucous membrane which is thrown
into folds when the bladder is empty. Another coat is found between
the two membranes above mentioned, called the muscular coat, the
action of which is to contract the bladder when the animal wants to
urinate. The bladder is held in by ligaments. The rectum lies above the
bladder, which in the horse rests on the floor of the pelvic cavity.
Its position in the mare differs from that of the horse. Instead of
the rectum or back bowel being immediately above it, as in the horse,
the womb is just above the bladder or between it and the rectum. The
bladder acts as a reservoir in which to store the urine until it is
full; it then presses on the walls and nerves, giving a peculiar
sensation to these parts, causing the walls to contract, forcing the
urine into a tube which carries it from the body. This is called the
urethra. The neck of the bladder is simply an opening at the back part,
and is guarded by a valve which prevents the urine from dripping out
except when the animal is passing its urine or water.
The Urethra is the tube which carries the urine from the bladder out
of the body. It is situated much differently in the mare than in the
horse. In the mare it is very short, passing from the neck of the
bladder along below the womb and vagina, which is the passage from the
outside into the neck of the womb. It opens up into the underside of
this passage about four inches in from the outside. This opening is
guarded by a small, thin valve, and can be felt by passing the finger
along the under side of the passage which leads into the womb. In the
horse this tube is a great deal longer than in the mare. It commences
at the bladder, passes along below the rectum or back bowel to just
below the anus. Here this tube bends downward and forward and passes
into the penis, continuing down to the end where it terminates. Its
purpose is to carry the urine from the bladder out of the body and
to perform certain actions in connection with the genital organs. Its
lining is a continuation of the membrane of the bladder.
URINARY ORGANS OF THE OX.
The chief difference in these organs occurs in the kidneys, which in
the ox are larger, and in place of being smooth, like those of the
horse, are rough, resembling a bunch of grapes. The bladder and the
urinary organs resemble those of the horse.
GENERATIVE SYSTEM OF THE HORSE.
Animals possess the faculty of reproducing or propagating their
species, and this function may be non-sexual or sexual, the former
being confined to certain lowly-organized classes of animals.
In all the higher animals the generation of a new being is dependent
upon two individuals, a male and a female, the female furnishing a
germ, or ovum, the male a fecundating fluid, or sperm, which animates
the germ and renders it fit for development.
Both the ovum of the female and the sperm of the male are secretions
of glands, which are termed the genital glands, male and female; and
in either sex the generative system may be said to consist of these
glands, with certain accessory organs. The act of coition brings the
two secretions into contact.
We have two systems of genital organs to consider--the male and the
female.
MALE.--The genital organs of the horse are as follows: The scrotum or
bag, the testicles, the spermatic cord, the vesiculae seminales or
pouches which hold the semen or sperm, the urethra, the penis and the
sheath.
The Scrotum is a sac or bag which contains the testicles. It is
situated between the hind legs, and is covered on the outside by a
very fine, soft skin. Passing up in the center under the sheath the
scrotum is a well marked line in the skin called median raphe. This
can be plainly seen when the horse is on his back. It continues up,
gradually getting fainter until it reaches the anus. Under the skin
are layers of white fascia or tissue which can be seen by cutting
through the scrotum. There is a partition in the scrotum separating the
two testicles. The size of the scrotum is affected very much by the
weather. In cold weather its fibres contract, causing it to get very
much smaller, while in warm weather the fibres relax, causing it to
become very much larger. The scrotum contains, supports and protects
the testicles.
The Testicles are the glands which secrete the semen or sperm. They
are two in number, one situated on the left side and the other on the
right. They are oval in shape, and are attached above to the spermatic
cord. Before the animal is born the testicles are situated in the
abdominal cavity and attached to the serous membrane which has already
been spoken of in connection with that cavity. At or about the time of
birth, there takes place what is known as the descent of the testicles
into the scrotum. In their downward course they pass through a slit or
small opening at the back part of the muscles of the abdomen, where
they are attached to the under part of the hip bone. These slits or
openings are known as the inguinal rings. They can be felt in the horse
by pressing the fingers well up into the groins. The descent of the
testicles is an important point to remember. If the testicle does not
descend into the scrotum the horse is known as a ridgling. In this case
the testicle is not found in the scrotum. At the front part of the
testicle there is a small ridge called the globus major and at the back
of it is another smaller ridge called the globus minor. Passing between
these two ridges is another well marked ridge called the epididymis.
These can be easily seen by examining the testicle after the animal is
altered or castrated. The substance of the testicle is made up of small
glands and fine tubes.
These tubes, as they pass towards the back of the testicle, form into
larger tubes and finally unite to form one called the vas deferens.
The Spermatic Cords, or the cords of the testicles, are attached
above to the inguinal rings or openings mentioned before. They are
about five or six inches long and have the testicles attached to them
below. In each cord is found a small muscle which goes by the name
of the spermatic muscle, the rest of the cord being made up of the
spermatic artery, veins and nerves. Running up at the back of these
cords is found a tube called the vas deferens. Around the spermatic
cords and testicles is a serous membrane, one layer being attached to
the testicle and cord, while the other is closely attached around the
inside of the scrotum. In this membrane are small glands which secrete
an oily fluid to lubricate the parts, preventing friction when they
are jolted about in the scrotum. This fluid flies out as soon as the
scrotum is cut. This is important, because sometimes from a slight
injury the glands will secrete a large amount of this fluid, thus
causing the scrotum to look large and swollen. This disease is known as
hydrocele or water in the scrotum.
VAS DEFERENS.--These tubes are two in number and are situated just
behind the spermatic cords. They are about an eighth of an inch in
diameter and quite hard. They carry the semen up the back part of the
spermatic cord through the inguinal rings, before mentioned. They
pass backward and upwards, one on each side, to the upper part of the
bladder, where they empty into two small pouches or sacs, called the
vesiculae seminales. These store up the semen as it is secreted by the
testicles, and when full present the appearance of a pear.
VESICULAE SEMINALES.--These sacs or pouches are situated at the upper
side, over the neck of the bladder, one on each side. They have the
vas deferens emptying into them at the front end, while at the back
end of each is a small opening that leads out into another small tube
which passes backward and empties into the urethra, mentioned before as
carrying the urine out from the bladder. These sacs or pouches store
up the semen or sperm of the horse. During sexual intercourse, these
pouches contract and force the semen through the little tubes mentioned
out into the urethra, leading down through the penis.
The Penis is the main organ of sexual intercourse. Its substance
is formed of what is known as erectile tissue, which under certain
circumstances becomes enormously distended with blood. Passing up the
under side there is, what has already been mentioned, the urethra, or
the tube, which carries the urine out of the body, and also in the act
of intercourse carries the semen. This is used for two purposes, as we
have already mentioned.
The Sheath is a loose process of skin which passes downward from the
scrotum, generally from about five to six inches, according to the
size of the animal. It is attached to each side, leaving a hole or
opening in the center through which the penis passes. The outside of
the sheath is covered by a thin, delicate skin similar to that of the
scrotum. It is lined inside by a membrane containing many small glands,
which secrete a thick, dark fluid to lubricate this passage. Sometimes
this fluid collects in here and has the appearance of tar. This is
important, for when it collects to a large extent the sheath should be
washed.
The Semen or sperm of the horse is a light fluid, which, when
examined under a microscope is found to contain small objects called
spermatozoa. These move about, and when in the womb meet the ovum of
the female, which is secreted by a gland called the ovary. When these
two small objects unite, they form the foetus, or what may be called
the animal in its first stage of development.
FEMALE GENITAL ORGANS.
The female genital organs, or the organs of the mare, are very much
different from those of the horse. They are known as follows: The
ovaries, the fallopian tubes, or tubes which carry the ovum from the
ovaries to the uterus or womb, the uterus or womb, the vagina, and the
vulva.
The Ovaries in the mare correspond to the testicles in the horse. Each
is about the size of a pigeon’s egg, and resembles it much in shape.
They are held in place by ligaments, and at the back part are provided
with tubes leading from them called the fallopian tubes. The ovaries
secrete the ovum or germ. This is a very minute body, which, when
examined under the microscope, is found to be only 1-150 of an inch in
diameter.
The Fallopian Tubes are two canals, one on each side. They pass
backward and upward, and enter the front part of the uterus or womb.
These small tubes are simply used to carry the germ or ovum up from the
ovaries and empty into the uterus or womb.
The Uterus or Womb is a muscular sac situated in the pelvic cavity,
bounded above by the rectum, below by the bladder, and on either side
by the walls of the pelvic cavity. It is divided into what is known
as a body and a neck. The body of the womb is very small, being only
about four to six inches in length and a couple of inches in diameter
when the animal is not pregnant. Near the front end, at the upper side
there are openings by which the ovum enters. When the animal becomes
pregnant, the body of the womb becomes enlarged and passes forward
and to the left side of the abdominal cavity. It continues to enlarge
as the time of pregnancy passes on, until the foetus, or young, has
attained its full size. After the mare has had her young, the womb
begins to get smaller until it attains its natural size again. The
womb is very largely supplied with blood vessels and nerves. This is
especially so when the animal is pregnant, as it takes a large amount
of blood to nourish the foetus, or the young animal, before birth. It
is made up of three coats. The inner is called mucous membrane, and
in the mare, while pregnant, is covered with numerous processes about
the size of peas to which the placenta or after-birth of the foal is
attached. The muscular coat is next to that of the mucous coat, and
lies between the outer and inner coats of the womb. It is made up of
muscular fibres, and is strong and thick in the womb, much thicker
than it is in the bowels or other organs, already mentioned. This coat
supports and protects the foetus, or young, while being carried in
the womb, and at the time of parturition, or what is commonly known
as foaling, this coat also comes in use. It contracts the womb very
forcibly on the foal, while the neck of the womb lies open, thus
helping to force the foal out of the womb. This is important as the
contraction of this coat produces what is known as labor pains. Lying
outside, and covering around the womb, is a serous coat, a continuation
of the serous coat of the bowels. The womb is held in place by strong
ligaments attached to the sides, and from there to the hip bones. These
are called broad ligaments. At the back part of the womb is the neck.
It consists of an opening, formed by a projection about the size of an
egg. This has a hard, gritty feeling when the animal is not in season.
The neck at this time is closed. The neck of the womb is under control
of the muscle around it, and this muscle is under control of the nerves
of the womb. When a mare comes in season this muscle is relaxed to a
certain extent, thus allowing the neck to open wide enough for the
passage of a couple of fingers. By working around it with the fingers
at this period it can be forced wide enough to admit a man’s hand. If
the mare is put to the horse at this time and becomes pregnant or with
foal, the muscles in the neck of the womb contract, firmly closing
it. It remains closed until the time of foaling. When, at the time
of foaling, the labor pains come on, the muscle in the neck dilates,
allowing the neck of the womb to open large enough for the foal to
pass out. The neck of the womb can be felt easily by oiling the hand
and passing it into the passage of the womb. It will be noticed, too,
that the neck spoken of projects into the passage.
THE VAGINA AND THE VULVA.--These two organs together make up the
passage which leads into the womb from the outside. In the young mare
they are separated by a thin curtain, or partition, made up of mucous
membrane. This curtain is found about four inches from the outside,
and is known as the hymen. It is destroyed, or should be, when the
mare is first put to the horse, although it is broken down other ways,
and in some cases it will disappear of its own accord. The part of
the passage in front of the hymen is called the vagina. This passage,
in structures, resembles the womb, but is not so strong. There are
numerous glands situated along the inner lining which secretes a fluid
to lubricate it. The principle use of this organ is to guide the penis
during sexual intercourse, and at the time of foaling serves as a
passage for the foal. That part of the passage behind the hymen is
known as the vulva. It is about four inches long and about two or three
inches high, varying according to the size of the mare. In front, it
is separated from the vagina by the hymen membrane. It resembles the
vagina in structure and is also provided with little glands in its
inner membrane to secrete fluid to lubricate the passage. At the back
part of the vulva or around the outside is what is known as the lips
of the vulva, one on each side of the opening. The outside of the lips
is covered by a very fine skin. Just below the skin, they consist of
erectile tissue, which is the same kind of tissue as that of the penis
of the horse. This tissue is found more abundantly in the lips of the
vulva of the young mare than in those of an old mare. The opening
between these lips is situated just below the anus, or the opening
where the back bowel ends. At the back part of the vulva, on the under
side, is an opening, or hole, about large enough for the passage of
a man’s finger. Through this hole the tube leading from the bladder
enters into the passage and allows the urine to pass into the vulva,
through which it runs out of the body. The clitoris is situated on the
upper side of this passage, just inside the lips. It can be seen in
the mare when she works the vulva after passing urine. Just below the
clitoris are found two or three small glands which secrete the fluid
that passes away when the mare is horsing.
Mammary Glands, or what is known as the mare’s udder or bag, are two
in number, situated between the thighs. In the young mare they are
very small, but after the mare is pregnant a few months these glands
enlarge, until at foaling time they attain their largest size. They are
covered outside by a thin, smooth skin. The substance of them consists
of small glands and tubes retain or hold milk until it is drawn away
from the bag either by milking or by the young animals sucking. During
the time of suckling the young, the glands are largely supplied with
blood, from which the milk is secreted. On the under side of each
gland is found the teat, or that part taken hold of by the young when
sucking. The end of the teat is pierced by several small holes, through
which the milk passes.
GENITAL ORGANS OF THE BULL.
The testicles are ovoid in shape and well developed, its long axis
being nearly vertical; the membrane which separate the two testicles is
very strong.
The spermatic cord and artery are small compared with those of the
horse.
The penis is long and pointed, and has an S-shaped curve in it just
below the pelvic bones; this curve can be felt by feeling just behind
the bag.
The sheath is long and runs further forward on the belly. It has a tuft
of hair on the point of it. During the time the bull is serving the
S-shaped part of the penis is straightened out by the action of the
protracter muscles, and drawn back into the S-shaped curve by retractor
muscles.
The urethra is completely enveloped by the fibrous sheath.
GENITAL ORGANS OF THE COW.
The ovaries of the cow are comparatively smaller than those of the
mare, but resemble them in structure.
The uterus or womb of the cow somewhat resembles that of the mare, but
the inner membrane is different, being covered with sixty or eighty
mushroom-like bodies about the size of a pigeon’s egg, more flattened
out. These bodies receive the name of cotyledons; to these the placenta
or afterbirth is attached--a very important point with which every
person interested should be familiar. These may be felt by examining a
cow soon after calving. The passage from the womb of the cow is shorter
than that of the mare, but is formed on the same principles.
The mammary glands constitute an organ termed the _udder_, which is
composed of two symmetrical halves, placed one against the other. Each
half is again divided into two distinct glands, each with its own
teat, so that the udder consists of four mammae and four teats; behind
this there may be two small rudimentary teats. In the center of each
quarter, just at the base of the teat, is a large cavity, the general
receptacle of all the milk ducts. From this cavity, which is sometimes
large enough to contain a quart, proceeds down the center of the teat
one defined canal from which the milk is drawn.
In the small ruminants as the sheep and goat there are two mammae and
two teats, constructed like those of the cow.
CHAPTER V.
ANGIOLOGY.
Blood-Vascular System of the Horse.
Under this heading we describe the organs of circulation, by the action
of which certain fluids are propelled through the body. It is customary
to divide this branch of the subject into two sections, considering
respectively the blood-vascular and lymphatic systems.
BLOOD-VASCULAR SYSTEM.--This involves the consideration of the
blood, a fluid which supplies nutriment to the tissues and receives
effete material from them; the heart, a muscular organ which, by its
contraction, initiates the motion of the blood; the arteries, a series
of tubes which convey the blood from the heart to all parts of the
body; the veins, tubes which return that fluid to the heart; and the
capillaries, minute tubes joining the small arteries and veins.
BLOOD.--Blood is a fluid tissue, which nourishes all living structures,
being the medium by which nutritive material is conveyed to, and effete
or waste material conveyed away from the solid tissues. It is an
opaque, thickish, clammy liquid, with a peculiar odor, sickly saline
taste, and alkaline reaction. Its color varies in different parts of
the same animal, that in the arteries being bright red or scarlet,
while the blood in the veins is of a dark purplish hue.
When examined microscopically, the blood is found to consist of
minute corpuscles, and a clear, transparent, yellow fluid, the liquor
sanguinis, or plasma, in which the corpuscles float. The corpuscles are
of two kinds, the red and the white or colorless; the former, by far
more numerous, vary in proportion. Red corpuscles vary in shape, but in
all mammals (animals that suckle their young) are more or less flat,
the families excepted where they are oval, as in birds, reptiles, and
fish, which are also nucleated. Their average diameter in the horse,
ox or sheep is about 1/4000th part of an inch, their average thickness
being about one-fourth of this. Each surface is depressed towards its
center, hence the corpuscle is appropriately described as a bi-concaved
disc.
The white corpuscles are larger than the red, round in shape, and
nucleated.
The liquor sanguinis is pale and clear, and consists of water, fibrin,
albumen, fatty compounds, extracts, odoriferous and saline matters.
The serum is a thin, transparent liquid, of a pale-straw or yellow
color, consisting of the liquor sanguinis deprived of fibrin. It
contains nearly 90 per cent of water, is always slightly alkaline,
and coagulates when heated, owing to the large quantity of albumen it
contains. Fibrin is a white, stringy elastic substance, which, when the
blood is in circulation, is in solution, and cannot be distinguished
from the other constituents of the plasma.
HEART.--The heart is the principal organ of circulation; it weighs
about six and one-half pounds in the average horse and acts as a force
pump to force the blood through the arteries. It is composed of strong
muscular tissue, which acts involuntarily, and is situated between the
lungs, which are divided by what is known as the mediastinum. This is
a division between the lungs made up of two folds, the heart being
between them. The bottom or apex, of the heart is downward and rests
just above the breast-bone; the upper part, or base is directed upward
and to the left side, the left lung having a depression on its inner
surface for the heart to work in. There is a covering or sack around
the heart which helps to protect and support it in its place. It is
attached above to the back-bone, and below to the bones of the breast.
This sack is made up of fibrous tissue and is of a whitish appearance;
inner surface is smooth, and supplied with numerous small glands which
secrete an oily substance called serous fluid. This lubricates the
outer surface of the heart and the inner surface of the sack so that
in action it does not irritate the walls or surfaces. The cavity of
the heart is divided into two parts, the right and left sides; each of
these parts is again subdivided. The upper cavity is called auricle and
the lower cavity ventricle; thus there are the right and left ventricle
and right and left auricle. The right auricle communicates with the
right ventricle by an opening in the septum or partition on the right
side of the heart. This opening is guarded by a valve to keep the blood
from flowing back into the auricle. The left auricle communicates with
the left ventricle, same as on the right side. The right side of the
heart is sometimes called the venous side and contains only venous or
impure blood. The left side is sometimes called the arterial side. It
contains pure blood only. This side of the heart is very much stronger
and thicker than the right side.
[Illustration: PLATE V.
THE HEART AND THE CHIEF BLOOD VESSELS OF THE HORSE--AFTER MEGNIN.]
EXPLANATION OF PLATE V
BLOOD-VASCULAR SYSTEM OF THE HORSE
1. Heart, right ventricle.
2. Heart, left ventricle.
3. Heart, left auricle.
4. Pulmonary artery.
5. Pulmonary veins.
6. Anterior aorta.
7. Carotid artery.
8. Glosso-facial artery.
9. Left brachial artery.
10. Dorsal artery.
11. Superior cervical artery.
12. Vertebral artery.
12´. Internal thoracic artery.
13. Humeral artery.
14. Radial artery.
14´. Cubital artery.
15. Great metacarpal artery.
16. Ungual branches.
17. Posterior aorta.
18. Coeliac axis.
19. Mesenteric arteries.
20. Renal artery (left).
21. Small testicular artery.
22. Posterior vena cava.
23. Portal vein.
23´. Hepatic circulation.
24. External iliac artery.
25. Internal iliac artery.
26. Lateral sacral artery.
27. Femoral artery.
28. Posterior tibial artery.
28´. Anterior tibial artery.
28´´. Femoro-popliteal artery.
29. Metatarsal vessels.
30. Venous plexus of the foot.
31. Internal saphenic vein.
32. Cephalic vein.
33. Jugular vein.
ARTERIES.--Arteries are tubes the purpose of which is to convey the
blood from the heart. For this reason it is apparent that all arteries
carry pure arterial blood with but one exception. The pulmonary
artery carries the blood from the right ventricle to the lungs, and
consequently carries impure or venous blood. Each time the left
ventricle contracts it causes a wave, as it were, to pass all through
the arteries. This contraction takes place when in a healthy condition
about 36 to 42 times every minute and gives rise to what is known as
the pulse. This wave, or beating, may be detected at any point where
the artery is situated so closely to the surface as to affect the
outside of the body sufficient to be felt by placing the finger on the
point; consequently the pulse may be counted at any of these points.
Place your forefinger on the lower edge of your own lower jaw directly
under the corner of your mouth. At this point an artery passes out over
the jaw bone and therefore runs very close to the surface, making it
quite possible to feel the wave caused by the contraction of your own
heart, quite distinctly. Near this point on the jaw of the horse the
pulse is most conveniently felt and counted.
The walls of the arteries are composed of elastic tissue and after
death are always lying open. Blood is never found in them after death
because they continue to contract sufficiently long enough to force all
the blood through them.
VEINS.--Veins are tubes in construction not so strong as the
arteries--the purpose of which is to convey the blood from all parts
of the body to the heart. The heart wave does not affect the veins,
and consequently the pulse cannot be detected by placing the finger
on an exposed portion of one of them. It is also apparent that all
veins carry impure or venous blood with but one exception, viz., the
pulmonary vein, the purpose of which is to conduct the purified blood
from the lungs to the heart.
CAPILLARIES.--The small arteries terminate in a system of minute
vessels--the capillaries--which are interposed between the termination
of the arteries and the commencement of the veins, forming plexuses
(network) which vary much in arrangement. Their average diameter is
about 2/1000ths of an inch, varying in different construction of the
organs, smallest in the brain and mucous membrane of the intestines,
larger in the skin, in glands, and the interior of bones. All arteries
do not terminate in capillaries, an exception being in erectile
tissue of the penis, where arteries end in cells or cavities placed
at the origin of the veins. As the blood passes slowly through these
capillaries, the nourishment is absorbed from it through their very
thin walls to supply the tissues of the body. When the blood passes
through this capillary network it again enters into large vessels
called the veins, which carry it on its way back to the heart.
COURSE OF THE BLOOD.--We have seen that the heart is divided into a
right or venous, and a left or arterial portion. The blood is pumped
by the heart to all parts of the body, through the arteries, passing
through the capillary system, where it parts with its nourishment, is
collected and returned to the heart by the veins, is again pumped by
the heart to the lungs, where it is purified and returned to the heart
to again commence the circuit as before.
LYMPHATIC SYSTEM.--The lymphatic or absorbent system is closely
connected with the blood-vascular system, and is made up of very fine
minute tubes and glands. These convey from the tissues of the body a
clear fluid known as lymph, and pours it into the blood of the veins as
it is on its way back to the heart. These glands are found all through
the body; for instance, there is a large group inside the thigh or
stifle joint of the horse, and another large group inside the shoulder.
It is important to note these, as they sometimes become inflamed and
the leg is swollen. They are then the seat of the disease called weed
in the leg, or lymphangitis.
[Illustration: PLATE VI.
_1. Brain._
_2. Spinal cord._
_3. Brachial plexus._
_4. Sacrolumbar plexus._
_5. Pneumogastric._
_6. Sciatic._
_7. Sympathetic System._
_8. Solar plexus._
NERVOUS SYSTEM OF A HORSE--AFTER MEGNIN.]
CHAPTER VI.
NEUROLOGY.
This illustration shows where the brain, spinal canal and the principal
nerves of the horse are located.
STUDY OF THE NERVES.--The nervous system is a very important set of
organs controlling the motion of the various members of the body and
supplying the different senses of feeling, seeing, hearing, smelling
and tasting. The two principal organs of the nervous system are the
brain and spinal cord.
The brain is the center of the whole nervous system, and is situated in
the cranial cavity, surrounded by three delicate membranes, the outer
one being attached to the inner wall of the bones forming the brain
cavity. The brain contains several important nerves called cranial
nerves, which are given off from the brain and pass down through the
various foramen or openings in the head to supply the different organs
situated there. The optic nerve passes down to the eye, giving the
sense of sight. The auditory nerve passes down to the drum of the ear
to give the sense of hearing. The olfactory nerves, which give the
sense of smell, are situated in the mucous membrane lining the nose.
The nerves passing down to the tongue give the sense of taste. Other
nerves pass down to the lips, teeth, mouth and face, giving motion and
feeling to the parts mentioned. Others pass down to the pharynx or
gullet, giving it the power of swallowing.
The spinal cord passes from the brain through the openings in the
bones of the back, which gives off numerous small nerves that supply
the muscles of the back with motion and feeling. Nearly opposite the
shoulder blade the spinal cord gives off a large trunk of nerves,
portion of which supplies the heart and lungs with nervous power.
This is a point of importance, for if the spinal cord becomes injured
in front of these nerves immediate death is the result. The other
portions of this trunk of nerves supply the shoulder, chest and muscles
of the front legs. Passing backward along the spinal cord is found the
sympathetic system of nerves, which go to supply the bowels, stomach,
liver, kidneys, and other organs situated in the abdominal cavity.
Coming backwards along the spinal cord to about opposite the hip bones,
is another set of nerves, one of which goes to supply the rectum, or
back bowels. Others go to the generative and urinary organs where they
assist in performing their functions. Other nerves pass to the small
organs situated in the pelvic cavity; some of these nerves pass down to
the hind legs, supplying them with nervous power. The remainder of the
nerves go to supply the tail.
The difference between the nervous system of a horse and other animals
is not worth mentioning.
CHAPTER VII.
AESTHESIOLOGY.
The study of organs of special sense, the ear, eye and organs of
special sensation, skin, hair, foot, etc.
THE EAR.
The apparatus of hearing is composed of three parts the outer, middle,
and inner ear; the two first being accessory for the collection and
transmission of sounds, and the latter the essential organ which
receives the impressions thus conveyed.
The inner part, or drum, of the ear, is situated in the hardest bone
of the body, called the petrosal. The nerve which passes into the drum
of the ear and gives the sense of hearing, is called the auditory
nerve. From the drum a small opening passes out into the outer part of
the ear; this is the portion which is seen on top of the head. It is
made up of a membrane known as the cartilage, which gives the ear its
stiffness. This cartilage is covered by a fine, delicate skin, covered
on the outside by fine, short hair. Situated on the inner side of the
outer ear are numerous long hairs projecting outward, the use of which
is to keep foreign bodies from dropping into the ear. The ear is moved
backward and forward by small muscles which are attached around it.
THE EYE.
The apparatus of vision comprises the essential organ, the globe of
the eye or eyeball, and its accessory parts or appendages. The eyeball
is situated in the orbital fossa, mentioned in chapter on the bones of
the head. It is chiefly made up of several coats around the outside,
and in the center by the humours of the eye. On the inner side of these
coats is a thin membrane called the retina, which contains the branches
of the optic nerve. This receives the reflections of objects as they
pass through the humours of the eye and from which the sensation
passes along the optic nerve to the brain. The oblong opening seen
in the middle of the eye is known as the pupil. If a horse be led
from a dark stall into the light and the pupils of the eyes watched
closely, it will be noticed that they get smaller, but on returning
it to the stall the pupils will be noticed to dilate or get larger;
thus it is seen that the pupils do not always remain the same size.
The chief use of the pupil is to gauge the sight. At the back part of
the eye are several muscles attached from around the eye to the bones
in the fossa. These muscles move the eye and assist in holding it to
its place. Around the front part are two movable curtains, one above
and the other below, called eyelids, the use of which is to open and
close the eye, and also to protect it from injuries. Around the free
border of the eyelids are what is known as the eyelashes, which keep
foreign substances from falling into the eye. Situated in the inner
angle is what is known as the haw of the eye; this membrane also helps
to protect it. In the corner of this angle is a small duct or opening,
through which a fluid called the tears passes down into the nasal
tubes, from whence it is carried down through the bones of the head and
emptied into the under part of the nostril or nose. A small gland is
situated on the upper part of the eye. This gland secretes the tears
which lubricates the eyes. The color of the eye is generally brown, but
in some cases it is white. It is then called a moon eye.
THE SKIN.
The skin is a membrane or external casing of the body. The skin itself
consists of two layers covered with hair, fine or coarse, long or
short, according to its position or purpose which nature intended it to
serve. The outer layer is called the epidermis, the inner the dermis.
THE EPIDERMIS.--The epidermis is the outer layer. It is not supplied
with nerves and blood vessels, its purpose being to protect the inner
layer. This layer undergoes a continual process of being made up and
passing away in dandruff.
THE DERMIS.--The dermis or true skin lies under the epidermis. It is
well supplied with nerves and blood vessels, part of the nerves being
the nerves of touch. This fact accounts for its becoming so very
sensitive and painful when through injury of any kind the outer layer
is scraped off. It is attached to the body by a layer of white tissue
known as the areolor tissue, this being that which is cut through
when the animal is being skinned. The thickness of the skin varies in
different parts of the body, being thinnest in the under parts. The
sweat glands are situated in the dermis.
THE HAIR.
There are three kinds of hair on the horse--the common, the finest of
the three, covers most of the body; that of the mane and tail, coarse
and long; and that growing on the muzzle or nose and lips, long and
usually black, known as tactile or cat hairs.
On the inside of the front legs, just above the knee, and on the inside
of the hind legs, above the hock, are rough, horny spots. These are
called chestnuts.
EXPLANATION OF PLATE VII
ANATOMY OF THE FOOT.--This illustration represents the foot of a horse
sawed from above the fetlock down through the center of the foot. It
shows the structure of the foot, the name of each part being given
according to number.
1. Lower end of large metacarpal, or cannon bone.
2. Bursa, which secretes the joint oil that lubricates the place where
the tendon, or cord, on the front of the leg passes down over the front
of the fetlock joint. This is important as it sometimes gets injured
and becomes enlarged. It is then called a bursal enlargement, and is of
the same nature as a wind gall.
3. Fetlock joint.
4. Os suffraginis, or large pastern bone.
5. Pastern joint. This joint is important; when diseased it is the seat
of a high ringbone.
6. Os coronae or small pastern bone.
7. Coffin joint. This joint is important, for when it is diseased it is
known as a low ringbone.
8. Wall of the hoof.
9. Os pedis, or coffin bone.
10. Sensitive wall, or quick of the foot.
11. Sensitive sole, or quick of the foot.
12. Frog of the foot, or horney frog.
13. Plantar cushion, or fatty frog.
14. Navicular bone. This is also important, for when diseased it is the
seat of navicular, or coffin joint lameness.
15. Back tendons below the fetlock.
16. Sesamoid, or fetlock bones.
17. Skin.
18. Back tendons above the fetlock.
[Illustration: PLATE VII.
CROSS SECTION OF THE FOOT OF THE HORSE.]
FOOT.--In equine anatomy the word “foot” implies the hoof, together
with the bones and soft structures contained therein. Many of these
objects have already been described, so that our description here will
be confined almost to the hoof and the structures with which it comes
into immediate contact internally.
WALL OF THE FOOT.--The wall is that part of the hoof seen when the
foot is resting flat on the ground. It is divided into the toe, the
quarters, the heels and the bars. The toe forms the front, and is the
thickest and strongest part of the wall. The quarters are situated at
the side. The walls are not nearly so thick here as at the toe, but are
almost straight up and down. The heels are situated at the back part
of the foot. From the heel is a process of hoof, which looks like a
bar, passing forward between the frog and the sole of the foot; this
can be seen plainly by raising up the foot. There is one of these at
each side of the frog. They act as braces to the heel and the quarters
of the wall; these are called the bars. Covering the outside of the
wall is a fine membrane called the periople, which gives the hoof its
polished appearance. This can be seen best when the hoof is well washed
off, as it is after traveling through wet grass. This membrane keeps
the moisture in the hoof and protects it from water. This is a point
of importance in shoeing horses, as it is very injurious to file the
wall too much. Around the top part of the wall, where it unites with
the skin, is a groove which contains a white band, called the coronary
substance, or band. This nourishes the wall of the hoof, or, in other
words, it is from this that the wall of the hoof grows. The under part
of the wall, or that which rests on the ground in the unshod animal,
is called the spread of the foot. On the inside of the wall, attaching
it to the bone of the foot called the os pedis, is the part called the
quick, or sensitive laminae. It is important to note this when driving
nails in shoeing. The nail should not be driven into this membrane, nor
should it be even pressed upon, for it is very sensitive. When a nail
has been driven so as to injure the membrane it is a common expression
to say, “You have pricked that horse’s foot.”
SOLE OF THE FOOT.--The sole is a thick plate of horn which helps to
form the under part of the hoof. It is situated between the inner
border of the under part of the wall already mentioned and the front of
the frog. The under part of the sole is concave, or hollowed out. The
upper part of the sole is attached to the under surface of the os pedis
bone, or bone of the foot, by a membrane called the quick, or sensitive
sole--this membrane is merely a continuation of the sensitive laminae.
The outer part of the sole is attached to the inner part of the wall.
When pared down a white ring is seen where the sole and the wall unite.
At the back part of the sole there is a notch the shape of the letter
V; in this notch the frog is situated. It is important to remember when
shoeing never to let the shoe rest on any part of the sole; neither
is it well to pare off too much of the barky-looking substance of the
sole, as this helps to keep the moisture in the foot. When this is
taken off it allows the moisture to escape and the hoof becomes dry and
contracted.
FROG OF THE FOOT.--The frog is the important spongy horn found in
the V-shaped notch in the back of the sole. It is wide at the back
and helps to form the heel of the foot; the pointed part in front is
called the apex of the frog. The under part of the frog is triangular
in shape and has a hollow in it called the cleft of the frog. There is
a hollow at each side of the frog, between it and the bars, called the
commissures of the frog. On the upper part is a membrane, known as the
sensitive frog, which attaches it to the under part of the os pedis, or
foot bone. This membrane is simply a continuation of the sensitive sole
spoken of in connection with the sole. The back part of the frog is the
widest part and spreads out to form the heel.
The study of the foot of the horse is of the greatest practical
importance, owing to the many diseases and injuries to which it is
liable. It resolves itself here into the consideration of the hoof or
horny case, and the parts contained within it.
CHAPTER VIII.
EMBRYOLOGY.
THE DEVELOPMENT OF THE YOUNG.--We must here first speak of the ovum or
germ, which is secreted by the ovary of the mare. Every time she comes
in season (which occurs every three weeks during the hot weather) this
ovum passes down the tubes before the womb as before mentioned, where
it remains a few days and then dies if she is not put to the horse; but
if, during the time this ovum is in the womb she is put to the horse
and one of the spermatozoa from the semen of the horse comes in contact
with it (the ovum) and a union of these takes place, then the rest of
the semen dies and passes away, and the neck of the womb contracts
gradually until it is perfectly tight. These two little bodies begin
to grow when united and form the foetus, or foal. The foetus may be
for convenience divided into three parts, viz.: the foetus proper, the
navel string, and the placenta. The placenta is the part which is found
covering the foal and is attached to the little pea-like elevations
on the inside of the womb. This covering is found to be full of small
blood vessels which finally unite to form two larger vessels, known as
the navel veins. These carry the blood up through the navel opening of
the foal and then to its heart. By the action of the heart it is forced
all through the body of the foal and returned again to the heart. It is
then forced down another artery to the navel opening, along the navel
cord, into the placenta again, where it is distributed through the
small blood vessels. When the blood comes down this cord from the foal
it is in its impure state, and while it is passing through these small
vessels in the placenta it comes very close to the small blood vessels
in the womb. The blood is cleansed and nourished from the blood of its
mother by a process similar to that which was spoken of when describing
the lungs. The foetus, or foal, does not grow so fast the first month
as it does later on. At the age of seventeen weeks the first hair
appears on the lips and the tip of the tail. Between the thirty-fifth
and the fortieth week the foal begins to show signs of life, and is
completely covered with hair. After this time it grows very rapidly
and may be seen moving around by watching the flank closely. The mare
carries her foal eleven months, but in some cases an aged mare has been
known to carry her foal over twelve months. In rare cases young mares
may lack a few days of eleven months.
The Average Periods of Gestation of Domestic Animals.
Mare 11 months
Ass 12 months
Cow 9 months
Sheep 5 months
Goat 5 months
Sow 3-1/2 months
Bitch 9 weeks
Cat 8 weeks
Note--A mare having been served by a stallion may occasionally in the
course of four or five weeks, manifest a desire for a second visit from
the male; is again served and conceives both times. This is known as
superfoetation. Such a case has been recorded by a veterinarian; the
animal in question, a mare, giving birth to a horse colt and a mule
colt, both dead. The mare had been covered by a jack and subsequently
by a horse.
SIGNS OF PREGNANCY.--The veterinarian is occasionally called upon to
give an opinion as to the pregnancy or non-pregnancy, of an animal and
consequently should familiarize himself with the various indications
which tend to prove the absence, or presence, of this condition. As a
rule, when the mare conceives, heat, or the desire for the male, is
no longer observable, and, on being led to the horse, she not only
refuses to receive his caresses, but assumes the offensive, viciously
striking and biting at him until led away. Soon the hair becomes more
glossy, and the mare becomes quieter in disposition. This change is
usually well marked in mares that are of a vicious disposition. The
abdomen gradually enlarges as pregnancy advances, the right side being
a little larger than the left. This enlargement is especially well
marked in the cow. In some cases the beating of the foetal heart may
be heard with the assistance of the stethoscope. Such an examination
is, however, very likely to give rise to mistakes. After the eighth
month well-marked symptoms of pregnancy are manifested, the belly at
this time being considerably distended, the back sinking, etc. Before
this time it is, however, impossible to make a positive statement as to
the condition of the animal except by making a very close and thorough
examination per rectum. The rectum should be cleared out by means of an
enema (injection) of tepid water; the hand and arm should be well oiled
and passed into the rectum. The region of the uterus being reached, an
examination may be made of its condition. As the time for parturition
approaches, the ligaments relax to a greater or less degree, and
a well-marked depression or sinking in the lumbosacral region may
be observed, the udder or bag enlarges, and milk is secreted.
The secretion of milk sometimes appears long before the time of
parturition, and has frequently been noticed to take place in animals
that have never been bred. As a rule the animal shows slight uneasiness
for a day or two before parturition, slight abdominal pain, etc., being
manifested. About this time the vulva becomes larger, and presents more
or less tumefaction. There may also be observed a flow of mucous taking
place from the vulva for a day or two before parturition.
INDEX
HORSE
Descriptive Anatomy, 5
Dissection, 6
BONES
Osteology
Back Bones, 11
Breast Bone, 12
Cannon Bone, 13
Carpus, 13
Cervical Vertebrae, 11
Classes of Bones, 9
Coccygeal Vertebrae, 12
Coffin Bone, 14
Contents of Bones, 9
Distal Phalanx, 14
Dorsal Vertebrae, 11
Femur, 15
Fibula, 16
Fore-arm Bone (large), 13
Fore-arm Bone (small), 13
Hip Bones, 12
Hock Bones, 16
Humerus, 12
Knee, 13
Loin Bones, 11
Lumbar Vertebrae, 11
Median Phalanx, 14
Metacarpal, 13
Metatarsus, 17
Neck Bones, 11
Os Coronae, 14
Os Metacarpi Magnus, 13
Os Metatarsi Magnum, 17
Os Naviculare, 15
Os Pedis, 14
Os Suffraginis, 14
Ossa Metatarsi Parva, 17
Ossa Metacarpi Parva, 14
Pastern Bone (large), 14
Pastern Bone (small), 14
Patella, 15
Pelvic Bones, 12
Proximal Phalanx, 14
Radius, 13
Ribs, 12
Rump Bones, 12
Sacral Vertebrae, 12
Scapula, 13
Sesamoid Bones, 14
Shoulder Blade, 13
Shoulder Bone, 12
Shuttle Bone, 15
Skeleton, 7
Skull, 10
Splint Bones (fore limb), 14
Splint Bones (hind limb), 17
Sternum, 12
Stifle Bone, 15
Structure of Bones, 8
Tail Bones 12
Tarsus, 16
Thigh Bone, 15
Tibia, 16
Ulna, 13
JOINTS
ARTHROLOGY
Adipose Tissue, 22
Cartilage, 21
Connective Tissue, 21
Elastic Tissue (Yellow), 22
Fat, 22
Ligaments, 22
Synovial Membranes, 23
Synovia, 23
Joints
Carpus, 25
Classes of Joints, 24
Coffin Joint, 26
Elbow Joint, 24
Fetlock Joint, 25
Hip Joint, 26
Hock Joint, 26
Joints of the Front Leg, 24
Joints of the Hind Leg, 26
Knee Joint, 25
Motion in Joints, 24
Pastern Joint, 25
Shoulder Joint, 24
Stifle Joint, 26
Tarsus, 26
MYOLOGY
Abdominal Muscles, 32
Back Muscles, 30
Coccygeal Muscles, 30
Deep Layer of Muscles, 31
Gluteal Muscles, 30
Gullet Muscles, 30
Head Muscles, 30
Hip Muscles, 30
Involuntary Muscles, 35
Leg Muscles (front), 33
Leg Muscles (hind), 34
Muscles, 28
Neck Muscles, 30
Pharynx Muscles, 30
Shoulder Muscles, 32
Superficial Layer of Muscles, 29
Tail Muscles, 30
Voluntary Muscles, 30
SPLANCHNOLOGY
Air Cells, 53
Bladder, 56
Bowels, 41
Breathing, Normal, 55
Bronchial Tubes, 53
Canine Teeth, 49
Cementum (or crusta petrosa), 48
Dentine (or ivory), 48
Digestive Apparatus, 37
Enamel, 48
Fallopian Tubes, 62
Female Genital Organs, 62
Generative, 58
Incisor Teeth, 49
Kidneys, 56
Larynx, 53
Liver, 43
Lungs, 54
Male, 58
Mammary Glands, 65
Molar Teeth, 49
Nasal Chambers, 52
Nostrils, 52
Organs of Respiration, 52
Ovaries, 62
Pancreas, 43
Penis, 61
Permanent Teeth, 49
Rectum, 42
Respiration, 54–55
Respiratory System of the Horse, 52
Salivary Glands, 39
Scrotum, 58
Semen, 61
Sheath, 61
Spleen, 43
Sperm, 61
Spermatic Cords, 60
Stomach, 40
Teeth, 48
Teeth Tables (comparative), 51
Temporary Teeth, 49
Testicles, 59
Ureters, 56
Urethra, 57
Urinary System, 56
Uterus, 62
Vagina, 64
Vas Deferens, 60
Vesiculae Seminales, 60
Vulva, 64
Windpipe or Trachea, 53
Wolf Teeth, 49
Womb, 62
ANGIOLOGY
Arteries, 70
Blood, 67
Blood-vascular System, 67
Capillaries, 71
Circulation, 69
Corpuscles, 67
Course of Blood, 71
Heart, 68
Liquor Sanguinis, 68
Lymphatic System, 72
Veins, 71
NEUROLOGY
Brain, 73
Spinal Cord, 73
AESTHESIOLOGY
Anatomy of the Foot, 78
Dermis, 77
Ear, 75
Epidermis, 77
Eye, 75
Foot Sole, 80
Foot Frog, 80
Foot Wall, 79
Frog of Foot, 80
Hair, 77
Skin, 76
Sole of Foot, 80
Wall of Foot, 79
EMBRYOLOGY
Development of Young, 81
Gestation of Domestic Animals, 82
Pregnancy, 82
COMPARATIVE ANATOMY
Abomasum, 46
Air Cells, 53
Aves, 20
Back Bones, 18
Bowels, 46
Bronchial Tubes, 53
Carnivora, 19
Cervical, 18
Chewing Cud, 46
Coccygeal, 18
Comparative Osteology, 17
Cud Chewing, 46
Digestive System, 44
Dorsal, 18
Fourth Stomach, 46
Genital Organs of the Bull, 65
Genital Organs of the Cow, 66
Gullet, 44
Hip Bones, 19
Honey Comb, 45
Larynx, 53
Liver, 46
Loin Bones, 18
Lumbar, 18
Lungs, 54
Mammary Glands, 66
Many-plies, 45
Neck Bones, 18
Normal Breathing, 55
Nostrils, 55
Oesophagus, 44
Omasum, 45
Omnivora, 19
Paunch, 45
Pelvis Bones, 19
Penis, 65
Process of Digestion, 47
Respirations per Minute of Different Animals, 54
Respiratory System of the Ox, 55
Reticulum, 45
Rumen, 45
Ruminantia, 44
Ruminantia (cud chewing animals), 18
Rumination, 47
Rump Bones, 18
Sacrum, 18
Salivary, 44
Sheath, 65
Skull, 18
Spermatic Cord, 65
Stomach, 45
Tail Bones, 18
Teeth, 44
Teeth, Front, 44
Teeth, Molars, 44
Testicles, 65
Thoracic Cavity (lung cavity), 55
Tongue, 44
Trachea, 53
Urethra, 66
Uterus, 66
Urinary System of the Ox, 58
Vertebrae, 18
Windpipe, 53
Womb, 66
PRESS OF
SWEENEY, VARNEY & STRAUB
PORTLAND, OREGON
Spelling_inconsistencies:
after-birth/afterbirth
anus/annus
areolor/areolar
blood vessels/blood-vessels/bloodvessels
Œsophagus/oesophagus
pully/pulley
sweeney/sweeny
tendinous/tendonous
vertebræ/vertebrae/vertebraes
principle/principal not used strictly in accordance with present day usage
Spelling corrections:
angilogy → angiology
Corpulscles → Corpuscles
cusion → cushion
Dessection → Dissection
discribing → describing
feltock → fetlock
femer → femur
Fermoro-popliteal → Femoro-popliteal
form → from
Forth → Fourth
heapitic → hepatic
lumlosacral → lumbosacral
mammelia → mammalia
Many Plies → Many-plies
mostened → moistened
muscles plays → muscles play
oposite → opposite
pannisulus → panniculus
planter → plantar
prefrom → perform
protracters → protracter
skining → skinning
thench → thence
through → though
vasiculae → vesiculae
Voscular → Vascular
End of Project Gutenberg's Notes on Veterinary Anatomy, by Charles J. Korinek
*** END OF THE PROJECT GUTENBERG EBOOK 60949 ***
|