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| author | nfenwick <nfenwick@pglaf.org> | 2025-02-07 23:06:08 -0800 |
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| committer | nfenwick <nfenwick@pglaf.org> | 2025-02-07 23:06:08 -0800 |
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diff --git a/56507-0.txt b/56507-0.txt new file mode 100644 index 0000000..1c86e00 --- /dev/null +++ b/56507-0.txt @@ -0,0 +1,35629 @@ +*** START OF THE PROJECT GUTENBERG EBOOK 56507 *** + + + + + + + + +Transcriber's note: The Errata (after the List of Plates) have been worked +into the main text. All other apparent mistakes have been retained as +printed. Text enclosed by underscores is in italics (_italics_); page +numbers enclosed by curly braces (example: {25}) have been incorporated to +facilitate the use of the Index.. + + * * * * * + + + + +THE GEOGRAPHICAL + +DISTRIBUTION OF ANIMALS + +_WITH A STUDY OF THE RELATIONS OF LIVING AND EXTINCT FAUNAS AS ELUCIDATING +THE PAST CHANGES OF THE EARTH'S SURFACE._ + +BY + +ALFRED RUSSEL WALLACE, + +AUTHOR OF "THE MALAY ARCHIPELAGO," ETC. + +WITH MAPS AND ILLUSTRATIONS. + +_IN TWO VOLUMES.--VOLUME II._ + +London: MACMILLAN AND CO. 1876. + +[_The Right of Translation and Reproduction is Reserved._] + + + + + LONDON: + R. CLAY, SONS, AND TAYLOR, PRINTERS, + BREAD STREET HILL. + + + + + CONTENTS OF THE SECOND VOLUME. + + PART III. (_continued_). + + ZOOLOGICAL GEOGRAPHY: A REVIEW OF THE CHIEF FORMS OF + ANIMAL LIFE IN THE SEVERAL REGIONS AND SUB-REGIONS, + WITH THE INDICATIONS THEY AFFORD OF GEOGRAPHICAL MUTATIONS. + + CHAPTER XIV. + + THE NEOTROPICAL REGION. + + General Zoological Features of the Neotropical Region (p. 5)--Distinctive + Characters of Neotropical Mammalia (p. 6)--Of Neotropical Birds (p. 7)-- + Neotropical Reptiles (p. 9)--Fresh-water Fishes (p. 12)--Insects (p. 13) + --Coleoptera (p. 15)--Land Shells (p. 19)--Marine Shells (p. 20)-- + Brazilian Sub-region (p. 21)--Its Mammalia (p. 23)--Its Birds (p. 24)-- + Islands of Tropical South America, Galapagos (p. 29)--Chilian Sub-region + (p. 36)--Birds (p. 38)--Reptiles and Amphibia (p. 40)--Fresh-water + Fishes (p. 42)--Lepidoptera (p. 42)--Coleoptera (p. 44)--Islands of South + Temperate America (p. 49)--Mexican Sub-region (p. 51)--Mammalia and + Birds (p. 52)--Reptiles and Fishes (p. 54)--Insects (p. 55)--Relations of + the Mexican Sub-region to the North and South American Continents (p. 57) + --Islands of the Mexican Sub-region (p. 59)--The Antillean Sub-region + (p. 60)--Its Mammalia (p. 62)--Its Birds (p. 64)--Table of the Resident + Land Birds of the Antilles (p. 68)--Reptiles (p. 72)--Insects (p. 73)-- + Land Shells (p. 75)--Past History of the Antilles (p. 78)--Summary of the + Past History of the Neotropical Region (p. 80)--Table I. Families of + Animals inhabiting the Neotropical Region (p. 85)--Table II. Genera of + Terrestrial Mammalia and Birds of the Neotropical Region (p. 91) + 1-113 + + CHAPTER XV. + + THE NEARCTIC REGION. + Zoological Characteristics of the Nearctic Region (p. 115)--List of + Typical Nearctic Genera of Land Birds (p. 118)--Summary of Nearctic + Vertebrata (p. 120)--Insects (p. 122)--Terrestrial and Fluviatile + Mollusca (p. 124)--The Californian Sub-region (p. 127)--The Rocky + Mountain Sub-region (p. 129)--The Alleghany Sub-region (p. 131)--The + Bermudas (p. 134)--The Canadian Sub-region (p. 135)--Greenland + (p. 138)--Table I. Families of Animals inhabiting the Nearctic Region + (p. 140)--Table II. Genera of Terrestrial Mammalia and Birds of the + Nearctic Region (p. 145) 114-153 + + CHAPTER XVI. + + SUMMARY OF THE PAST CHANGES AND GENERAL RELATIONS OF THE SEVERAL REGIONS + 154-164 + + PART IV. + + GEOGRAPHICAL ZOOLOGY: A SYSTEMATIC SKETCH OF THE CHIEF FAMILIES OF LAND + ANIMALS IN THEIR GEOGRAPHICAL RELATIONS. + + INTRODUCTION 167-169 + + CHAPTER XVII. + + THE DISTRIBUTION OF THE FAMILIES AND GENERA OF MAMMALIA. + + Primates (p. 170)--General Remarks on the Distribution of Primates + (p. 179)--Chiroptera (p. 181)--Remarks on the Distribution of Chiroptera + (p. 185)--Insectivora (p. 186)--General Remarks on the Distribution of + Insectivora (p. 191)--Carnivora (p. 192)--General Remarks on the + Distribution of the Carnivora (p. 204)--Cetacea (p. 207)--Sirenia + (p. 210)--Ungulata (p. 211)--General Remarks on the Distribution of the + Ungulata (p. 226)--Proboscidea (p. 227)--Hyracoidea (p. 228)--Rodentia + (p. 229)--General Remarks on the Distribution of the Rodentia (p. 243)-- + Edentata (p. 244)--General Remarks on the Distribution of the Edentata + (p. 247)--Marsupialia (p. 248)--General Remarks on the Distribution of + Marsupialia (p. 253)--Monotremata (p. 253) 170-254 + + CHAPTER XVIII. + + THE DISTRIBUTION OF THE FAMILIES AND GENERA OF BIRDS. + + Passeres (p. 255)--General Remarks on the Distribution of the Passeres + (p. 299)--Picariæ (p. 302)--General Remarks on the Distribution of the + Picariæ (p. 322)--Psittaci (p. 324)--General Remarks on the Distribution + of the Psittaci (p. 329)--Columbæ (p. 331)--General Remarks on the + Distribution of the Columbæ (p. 335)--Gallinæ (p. 337)--General Remarks + on the Distribution of Gallinæ (p. 344)--Opisthocomi (p. 345)--Accipitres + (p. 345)--General Remarks on the Distribution of the Accipitres (p. 351) + --Grallæ (p. 351)--General Remarks on the Distribution of the Grallæ + (p. 362)--Anseres (p. 363)--General Remarks on the Distribution of the + Anseres (p. 367)--Struthiones (p. 368)--Struthious Birds recently Extinct + (p. 369)--General Remarks on the Distribution of the Struthiones (p. 370) + 255-371 + + CHAPTER XIX. + + THE DISTRIBUTION OF THE FAMILIES AND GENERA OF REPTILES AND AMPHIBIA. + + Ophidia (p. 372)--General Remarks on the Distribution of Ophidia (p. 386) + --Lacertilia (p. 388)--General Remarks on the Distribution of Lacertilia + (p. 403)--Rhyncocephalina (p. 405)--Crocodilia (p. 405)--General Remarks + on the Distribution of Crocodilia (p. 406)--Chelonia (p. 407)--Remarks on + the Distribution of Chelonia (p. 410)--Amphibia, Pseudophidia (p. 411)-- + Urodela (p. 411)--Anura (p. 414)--General Remarks on the Distribution of + Amphibia (p. 422) 372-423 + + CHAPTER XX. + + THE DISTRIBUTION OF THE FAMILIES OF FISHES, WITH THE RANGE OF SUCH GENERA + AS INHABIT FRESH WATER. + + Acanthopterygii (p. 424)---Acanthopterygii Pharyngognathi (p. 437)-- + Anacanthini (p. 439)--Physostomi (p. 441)--Lophobranchii (p. 456)-- + Plectognathi (p. 457)--Sirenoidei (p. 458)--Ganoidei (p. 458)-- + Chondropterygii (p. 460)--Cyclostomata (p. 463)--Leptocardii (p. 464)-- + Remarks on the Distribution of Fishes (p. 464) 424-467 + + CHAPTER XXI. + + THE DISTRIBUTION OF SOME OF THE MORE IMPORTANT FAMILIES AND GENERA OF + INSECTS. + + Lepidoptera (p. 470)--General Remarks on the Distribution of the Diurnal + Lepidoptera and Sphingidea (p. 483)--Coleoptera (p. 486)--Cicindelidæ + (p. 486)--Carabidæ (p. 488)--Lucanidæ (p. 492)--Cetoniidæ (p. 494)-- + Buprestidæ (p. 495)--Longicornia (p. 498)--General Observations on the + Distribution of Coleoptera (p. 502) 468-503 + + CHAPTER XXII. + + AN OUTLINE OF THE GEOGRAPHICAL DISTRIBUTION OF MOLLUSCA. + + Cephalopoda (p. 505)--Gasteropoda (p. 507)--Pulmonifera (p. 512)--General + Observations on the Distribution of Land Mollusca (p. 522)--Pteropoda + (p. 531)--Brachiopoda (p. 532)--Conchifera (p. 533)--General Remarks on + the Distribution of Marine Mollusca (p. 537) 504-539 + + CHAPTER XXIII. + + SUMMARY OF THE DISTRIBUTION AND LINES OF MIGRATION OF THE SEVERAL + CLASSES OF ANIMALS. + + Mammalia (p. 540)--Lines of Migration of the Mammalia (p. 544)--Birds + (p. 545)--Reptiles (p. 547)--Amphibia (p. 548)--Fresh-water Fishes + (p. 549)--Insects (p. 550)--Terrestrial Mollusca (p. 551)--Conclusion + (p. 552) 540-553 + + GENERAL INDEX 557 + + + + +MAPS AND ILLUSTRATIONS IN VOL. II. + + + + _To face page_ + + 1. Map of the Neotropical Region 3 + + 2. Plate XIV. A Brazilian Forest with Characteristic Mammalia 24 + + 3. Plate XV. A Scene on the Upper Amazon, with some Characteristic + Birds 28 + + 4. Plate XVI. The Chilian Andes, with Characteristic Animals 40 + + 5. Plate XVII. A Scene in Cuba, with Characteristic Animals 67 + + 6. Map of the Nearctic Region 115 + + 7. Plate XVIII. Scene in California with some Characteristic Birds 128 + + 8. Plate XIX. The North American Prairies with Characteristic + Mammalia 130 + + 9. Plate XX. A Canadian Forest with Characteristic Mammalia 136 + + + + + + +ERRATA IN VOL. II. + +As in Vol. I. mis-spellings are not given here, being mostly corrected in +the Index. + + + Page 111, No. 642, _for_ 1 _read_ 2. + " 111, No. 643, _for_ 15 _read_ 9. + " 267, line 7, _add_ Borneo. + " 276, line 10, _for_ 16 Genera _read_ 11 Genera. + " " 8 lines from foot, for _Drepanornis_ read _Neodrepanis_. + " 291, 5 lines from foot, for _Sayornis_ read _Empidias_. + + + + +THE + +GEOGRAPHICAL DISTRIBUTION +OF ANIMALS. + +PART III. (_continued._) + +_ZOOLOGICAL GEOGRAPHY:_ + +_A REVIEW OF THE CHIEF FORMS OF ANIMAL LIFE IN THE SEVERAL REGIONS AND +SUB-REGIONS, WITH THE INDICATIONS THEY AFFORD OF GEOGRAPHICAL MUTATIONS._ + +[Illustration: NEOTROPICAL REGION] + + + + +{3}CHAPTER XIV. + +THE NEOTROPICAL REGION. + + +This region, comprehending not only South America but Tropical North +America and the Antilles, may be compared as to extent with the Ethiopian +region; but it is distinguished from all the other great zoological +divisions of the globe, by the small proportion of its surface occupied by +deserts, by the large proportion of its lowlands, and by the altogether +unequalled extent and luxuriance of its tropical forests. It further +possesses a grand mountain range, rivalling the Himalayas in altitude and +far surpassing them in extent, and which, being wholly situated within the +region and running through eighty degrees of latitude, offers a variety of +conditions and an extent of mountain slopes, of lofty plateaus and of deep +valleys, which no other tropical region can approach. It has a further +advantage in a southward prolongation far into the temperate zone, +equivalent to a still greater extension of its lofty plateaus; and this +has, no doubt, aided the development of the peculiar alpine forms of life +which abound in the southern Andes. The climate of this region is +exceptionally favourable. Owing to the lofty mountain range situated along +its western margin, the moisture-laden trade winds from the Atlantic have +free access to the interior. A sufficient proportion of this moisture +reaches the higher slopes of the Andes, where its condensation gives rise +to innumerable streams, which cut deep ravines and carry down such an +amount of sediment, that they have formed the vast plains of the Amazon, of +{4}Paraguay, and of the Orinooko out of what were once, no doubt, arms of +the sea, separating the large islands of Guiana, Brazil, and the Andes. +From these concurrent favourable conditions, there has resulted that +inexhaustible variety of generic and specific forms with a somewhat limited +range of family and ordinal types, which characterise neotropical zoology +to a degree nowhere else to be met with. + +Together with this variety and richness, there is a remarkable uniformity +of animal life over all the tropical continental portions of the region, so +that its division into sub-regions is a matter of some difficulty. There +is, however, no doubt about separating the West Indian islands as forming a +well-marked subdivision; characterised, not only by that poverty of forms +which is a general feature of ancient insular groups, but also by a number +of peculiar generic types, some of which are quite foreign to the remainder +of the region. We must exclude, however, the islands of Trinidad, Tobago, +and a few other small islands near the coast, which zoologically form a +part of the main land. Again, the South Temperate portion of the continent, +together with the high plateaus of the Andes to near the equator, form a +well-marked subdivision, characterised by a peculiar fauna, very distinct +both positively and negatively from that of the tropical lowland districts. +The rest of Tropical South America is so homogeneous in its forms of life +that it cannot be conveniently subdivided for the purposes of a work like +the present. There are, no doubt, considerable differences in various parts +of its vast area, due partly to its having been once separated into three +or more islands, in part to existing diversities of physical conditions; +and more exact knowledge may enable us to form several provinces or perhaps +additional sub-regions. A large proportion of the genera, however, when +sufficiently numerous in species, range over almost the whole extent of +this sub-region wherever the conditions are favourable. Even the Andes do +not seem to form such a barrier as has been supposed. North of the equator, +where its western slopes are moist and forest-clad, most of the genera are +found on both sides. To the south of this line its western valleys are arid +and its lower plains almost deserts; and thus the absence of a {5}number of +groups to which verdant forests are essential, can be traced to the +unsuitable conditions rather than to the existence of the mountain barrier. +All Tropical South America, therefore, is here considered to form but one +sub-region. + +The portion of North America that lies within the tropics, closely +resembles the last sub-region in general zoological features. It possesses +hardly any positive distinctions; but there are several of a negative +character, many important groups being wholly confined to South America. On +the other hand many genera range into Mexico and Guatemala from the north, +which never reach South America; so that it is convenient to separate this +district as a sub-region, which forms, to some extent, a transition to the +Nearctic region. + +_General Zoological Features of the Neotropical Region._--Richness combined +with isolation is the predominant feature of Neotropical zoology, and no +other region can approach it in the number of its peculiar family and +generic types. It has eight families of Mammalia absolutely confined to it, +besides several others which are rare elsewhere. These consist of two +families of monkeys, Cebidæ and Hapalidæ, both abounding in genera and +species; the Phyllostomidæ, or blood-sucking bats; Chinchillidæ and Caviidæ +among rodents; besides the greater part of the Octodontidæ, Echimyidæ and +Cercolabidæ. Among edentata, it has Bradypodidæ, or sloths, Dasypodidæ, or +armadillos, and Myrmecophagidæ, or anteaters, constituting nearly the +entire order; while Procyonidæ, belonging to the carnivora, and +Didelphyidæ, a family of marsupials, only extend into the Nearctic region. +It has also many peculiar groups of carnivora and of Muridæ, making a total +of full a hundred genera confined to the region. Hardly less remarkable is +the absence of many wide-spread groups. With the exception of one genus in +the West Indian islands and a _Sorex_ which reaches Guatemala and Costa +Rica, the Insectivora are wholly wanting; as is also the extensive and +wide-spread family of the Viverridæ. It has no oxen or sheep, and indeed no +form of ruminant except deer and llamas; neither do its vast forests and +grassy plains support a single form of non-ruminant ungulate, except the +tapir and the peccary. + +{6}_Birds._--In birds, the Neotropical region is even richer and more +isolated. It possesses no less than 23 families wholly confined within its +limits, with 7 others which only extend into the Nearctic region. The names +of the peculiar families are: Cærebidæ, or sugar-birds; Phytotomidæ, or +plant-cutters; Pipridæ, or manakins; Cotingidæ, or chatterers; +Formicariidæ, or ant-thrushes; Dendrocolaptidæ, or tree-creepers; +Pteroptochidæ; Rhamphastidæ, or toucans; Bucconidæ, or puff-birds; +Galbulidæ, or jacamas; Todidæ, or todies; Momotidæ, or motmots; +Steatornithidæ, the guacharo, or oil-bird; Cracidæ, or curassows; Tinamidæ, +or tinamous; Opisthocomidæ, the hoazin; Thinocoridæ; Cariamidæ; Aramidæ; +Psophiidæ, or trumpeters; Eurypygidæ, or sun-bitterns; and Palamedeidæ, or +horned-screamers. The seven which it possesses in common with North America +are: Vireonidæ, or greenlets; Mniotiltidæ, or wood-warblers; Tanagridæ, or +tanagers; Icteridæ, or hang-nests; Tyrannidæ, or tyrant-shrikes; +Trochilidæ, or humming-birds; and Conuridæ, or macaws. Most of these +families abound in genera and species, and many are of immense extent; such +as Trochilidæ, with 115 genera, and nearly 400 species; Tyrannidæ, with +more than 60 genera and nearly 300 species; Tanagridæ, with 43 genera and +300 species; Dendrocolaptidæ with 43 genera and more than 200 species; and +many other very large groups. There are nearly 600 genera peculiar to the +Neotropical region; but in using this number as a basis of comparison with +other regions we must remember, that owing to several ornithologists having +made the birds of South America a special study, they have perhaps been +more minutely subdivided than in the case of other entire tropical regions. + +_Distinctive Characters of Neotropical Mammalia._--It is important also to +consider the kind and amount of difference between the various animal forms +of this region and of the Old World. To begin with the Quadrumana, all the +larger American monkeys (Cebidæ) differ from every Old World group in the +possession of an additional molar tooth in each jaw; and it is in this +group alone that the tail is developed into a prehensile organ of wonderful +power, adapting the animals to a purely arboreal life. Four of the genera, +comprising more than half the {7}species, have the prehensile tail, the +remainder having this organ either short, or lax as in the Old World +monkeys. Other differences from Old World apes, are the possession of a +broad nasal septum, and a less opposable thumb; and the absence of +cheek-pouches, ischial callosities, and a bony ear-tube. The Hapalidæ, or +marmozets, agree with the Cebidæ in all these characters, but have others +in addition which still more widely separate them from the Simiidæ; such as +an additional premolar tooth, acute claws, and thumb not at all opposable; +so that the whole group of American monkeys are radically different from +the remainder of the order. + +The Procyonidæ are a distinct family of Carnivora, which make up for the +scarcity of Mustelidæ in South America. The Suidæ are represented by the +very distinct genus _Dicotyles_ (Peccary) forming a separate sub-family, +and differing from all other genera in their dentition, the absence of tail +and of one of the toes of the hind feet, the possession of a dorsal gland, +and only two mammæ. The rodents are represented by the Chinchillidæ and +Caviidæ, the latter comprising the largest animals in the order. The +Edentata are almost wholly confined to this region; and the three families +of the sloths (Bradypodidæ), armadillos (Dasypodidæ), and ant-eaters +(Myrmecophagidæ), are widely separated in structure from any Old World +animals. Lastly, we have the opossums (Didelphyidæ), a family of +marsupials, but having no close affinity to any of the numerous Australian +forms of that order. We have already arrived at the conclusion that the +presence of marsupials in South America is not due to any direct +transference from Australia, but that their introduction is comparatively +recent, and that they came from the Old World by way of North America (vol. +i., p. 155). But the numerous and deep-seated peculiarities of many other +of its mammalia, would indicate a very remote origin; and a long-continued +isolation of South America from the rest of the world is required, in order +to account for the preservation and development of so many distinct groups +of comparatively low-type quadrupeds. + +_Distinctive Characters of Neotropical Birds._--The birds which are +especially characteristic of this region, present similar distinctive +features. In the enormous group of Passerine {8}birds which, though +comprising nearly three-fourths of the entire class, yet presents hardly +any well-marked differences of structure by which it can be subdivided--the +families confined to America are, for the most part, more closely related +to each other than to the Old World groups. The ten families forming the +group of "Formicaroid Passeres," in our arrangement (vol. i., p. 94), are +characterised by the absence of singing muscles in the larynx, and also by +an unusual development of the first primary quill; and seven of this series +of families (which are considered to be less perfectly developed than the +great mass of Old World passeres) are exclusively American, the three +belonging to the Eastern hemisphere being of small extent. Another group of +ten families--our "Tanagroid Passeres," are characterised by the abortion +or very rudimentary condition of the first quill; and of these, five are +exclusively American, and have numerous genera and species, while only two +are non-American, and these are of small extent. On the other hand the +"Turdoid Passeres," consisting of 23 families and comprising all the true +"singing-birds," is poorly represented in America; no family being +exclusively Neotropical, and only three being at all fully represented in +South America, though they comprise the great mass of the Old World +passeres. These peculiarities, which group together whole series of +families of American birds, point to early separation and long isolation, +no less surely than the more remarkable structural divergences presented by +the Neotropical mammalia. + +In the Picariæ, we have first, the toucans (Rhamphastidæ); an extraordinary +and beautiful family, whose enormous gaily-coloured bills and long +feathered tongues, separate them widely from all other birds. The Galbulidæ +or jacamars, the motmots (Momotidæ), and the curious little todies (Todidæ) +of the Antilles, are also isolated groups. But most remarkable of all is +the wonderful family of the humming-birds, which ranges over all America +from Tierra del Fuego to Sitka, and from the level plains of the Amazon to +above the snow-line on the Andes; which abounds both in genera, species, +and individuals, and is yet strictly confined to this continent alone! How +vast must have been the time required to develop those beautiful and +{9}highly specialized forms out of some ancestral swift-like type; how +complete and long continued the isolation of their birthplace to have +allowed of their modification and adaptation to such divergent climates and +conditions, yet never to have permitted them to establish themselves in the +other continents. No naturalist can study in detail this single family of +birds, without being profoundly impressed with the vast antiquity of the +South American continent, its long isolation from the rest of the land +surface of the globe, and the persistence through countless ages of all the +conditions requisite for the development and increase of varied forms of +animal life. + +Passing on to the parrot tribe, we find the peculiar family of the +Conuridæ, of which the macaws are the highest development, very largely +represented. It is in the gallinaceous birds however that we again meet +with wholly isolated groups. The Cracidæ, including the curassows and +guans, have no immediate relations with any of the Old World families. +Professor Huxley considers them to approach nearest to (though still very +remote from) the Australian megapodes; and here, as in the case of the +marsupials, we probably have divergent modifications of an ancient type +once widely distributed, not a direct communication between the southern +continents. The Tinamidæ or tinamous, point to a still more remote +antiquity, since their nearest allies are believed to be the Struthiones or +ostrich tribe, of which a few representatives are scattered widely over the +globe. The hoazin of Guiana (Opisthocomus) is another isolated form, not +only the type of a family, but perhaps of an extinct order of birds. +Passing on to the waders, we have a number of peculiar family types, all +indicative of antiquity and isolation. The _Cariama_ of the plains of +Brazil, a bird somewhat intermediate between a bustard and a hawk, is one +of these; the elegant _Psophia_ or trumpeter of the Amazonian forests; the +beautiful little sun-bittern of the river banks (_Eurypyga_); and the +horned screamers (_Palamedea_), all form distinct and isolated families of +birds, to which the Old World offers nothing directly comparable. + +_Reptiles._--The Neotropical region is very rich in varied forms of reptile +life, and the species are very abundant. It has six {10}altogether peculiar +families, and several others which only range into the Nearctic region, as +well as a very large number of peculiar or characteristic genera. As the +orders of reptiles differ considerably in their distributional features, +they must be considered separately. + +The snakes (Ophidia) differ from all other reptiles, and from most other +orders of vertebrates, in the wide average distribution of the families; so +that such an isolated region as the Neotropical possesses no peculiar +family, nor even one confined to the American continent. The families of +most restricted range are--the Scytalidæ, only found elsewhere in the +Philippine islands; the Amblycephalidæ, common to the Oriental and +Neotropical regions; and the Tortricidæ, most abundant in the Oriental +region, but found also in the Austro-Malay islands and Tropical South +America. Sixteen of the families of snakes occur in the region, the +Colubridæ, Amblycephalidæ, and Pythonidæ, being those which are best +represented by peculiar forms. There are 25 peculiar or characteristic +genera, the most important being _Dromicus_ (Colubridæ); _Boa_, +_Epicrates_, and _Ungalia_ (Pythonidæ); _Elaps_ (Elapidæ); and +_Craspedocephalus_ (Crotalidæ). + +The lizards (Lacertilia) are generally more restricted in their range; +hence we find that out of 15 families which inhabit the region, 5 are +altogether peculiar, and 4 more extend only to N. America. The peculiar +families are Helodermidæ, Anadiadæ, Chirocolidæ, Iphisiadæ, and +Cercosauridæ; but it must be noted that these all possess but a single +genus each, and only two of them (Chirocolidæ and Cercosauridæ) have more +than a single species. The families which range over both South and North +America are Chirotidæ, Chalcidæ, Teidæ, and Iguanidæ; the first and second +are of small extent, but the other two are very large groups, the Teidæ +possessing 12 genera and near 80 species; the Iguanidæ 40 genera and near +150 species; the greater part of which are Neotropical. There are more than +50 peculiar or highly characteristic genera of lizards, about 40 of which +belong to the Teidæ and Iguanidæ, which thus especially characterize the +region. The most important and characteristic genera are the following; +_Ameiva_ (Teidæ); _Gymnopthalmus_ (Gymnopthalmidæ); {11}_Celestus_ and +_Diploglossus_ (Scincidæ); _Sphærodactylus_ (Geckotidæ); _Liocephalus_, +_Liolæmus_, _Proctotretus_, and many smaller genera (Iguanidæ). The three +extensive Old World families Varanidæ, Lacertidæ, and Agamidæ, are absent +from the entire American continent. + +In the order Crocodilia, America has the peculiar family of the alligators +(Alligatoridæ), as well as several species of true crocodiles +(Crocodilidæ). The Chelonia (tortoises) are represented by the families +Testudinidæ and Chelydidæ, both of wide range; but there are six peculiar +genera,--_Dermatemys_ and _Staurotypus_ belonging to the former +family,--_Peltocephalus_, _Podocnemis_, _Hydromedusa_, and _Chelys_, to the +latter. Some of the Amazon river-turtles of the genus _Podocnemys_ rival in +size the largest species of true marine turtles (Cheloniidæ), and are +equally good for food. + +_Amphibia._--The Neotropical region possesses representatives of sixteen +families of Amphibia of which four are peculiar; all belonging to Anoura or +tail-less Batrachians. The Cæciliadæ or snake-like amphibia, are +represented by two peculiar genera, _Siphonopsis_ and _Rhinatrema_. Tailed +Batrachians are almost unknown, only a few species of _Spelerpes_ +(Salamandridæ) entering Central America, and one extending as far south as +the Andes of Bogota in South America. Tail-less Batrachians on the other +hand, are abundant; there being 14 families represented, of which +4,--Rhinophryndæ, Hylaplesidæ, Plectromantidæ, and Pipidæ are peculiar. +None of these families contain more than a single genus, and only the +second more than a single species; so that it is not these which give a +character to the South American Amphibia-fauna. The most important and best +represented families are, Ranidæ (true frogs), with eleven genera and more +than 50 species; Polypedatidæ (tree-frogs) with seven genera and about 40 +species; Hylidæ (tree-frogs) with eight genera and nearly 30 species; +Engystomidæ (toads) (5 genera), Bombinatoridæ (frogs), (4 genera), +Phryniscidæ and Bufonidæ (toads), (each with 2 genera), are also fairly +represented. All these families are widely distributed, but the Neotropical +genera are, in almost every case, peculiar. + +{12}_Fresh-water fishes._--The great rivers of Tropical America abound in +fish of many strange forms and peculiar types. Three families, and three +sub-family groups are peculiar, while the number of peculiar genera is +about 120. The peculiar families are Polycentridæ, with two genera; +Gymnotidæ, a family which includes the electric eels, (5 genera); and +Trygonidæ, the rays, which are everywhere marine except in the great rivers +of South America, where many species are found, belonging to two genera. Of +the extensive family Siluridæ, three sub-families Siluridæ anomalopteræ, S. +olisthopteræ, and S. branchiolæ, are confined to this region. The larger +and more important of the peculiar genera are the following: _Percilia_, +inhabiting Chilian and _Percichthys_ South Temperate rivers, belong to the +Perch family (Percidæ); _Acharnes_, found only in Guiana, belongs to the +Nandidæ, a family of wide range in the tropics; the Chromidæ, a family of +exclusively fresh-water fishes found in the tropics of the Ethiopian, +Oriental and Neotropical regions, are here represented by 15 genera, the +more important being _Acara_ (17 sp.), _Heros_ (26 sp.), _Crenicichla_ (9 +sp.), _Satanoperca_ (7 sp.). Many of these fishes are beautifully marked +and coloured. The Siluridæ proteropteræ are represented by 14 genera, of +which _Pimelodus_ (42 sp.), and _Platystoma_ (11 sp.), are the most +important; the Siluridæ stenobranchiæ by 11 genera, the chief being _Doras_ +(13 sp.), _Auchenipterus_ (9 sp.), and _Oxydoras_ (7 sp.). The Siluridæ +proteropodes are represented by 16 genera, many of them being among the +most singular of fresh-water fishes, clothed in coats of mail, and armed +with hooks and serrated spines. The following are the most +important,--_Chætostomus_ (25 sp.), _Loricaria_ (17 sp.), _Plecostonus_ (15 +sp.) and _Callichthys_ (11 sp.). The Characinidæ are divided between +Tropical America and Tropical Africa, the former possessing about 40 genera +and 200 species. The Haplochitonidæ are confined to South America and +Australia; the American genus being _Haplochiton_. The Cyprinodontidæ are +represented by 18 genera, the most important being, _Pæcilia_ (16 sp.), +_Girardinus_ (10 sp.), and _Gambusia_ (8 sp.) The Osteoglossidæ, found in +Australian and African rivers, are represented in South America by the +peculiar _Arapaima_, the "pirarucu" of the {13}Amazon. The ancient +Sirenoidei, also found in Australia and Africa, have the _Lepidosiren_ as +their American representative. Lastly, _Ellipisurus_ is a genus of rays +peculiar to the fresh waters of South America. We may expect these numbers +to be largely increased and many new genera to be added, when the extensive +collections made by Agassiz in Brazil are described. + +_Summary of Neotropical Vertebrates._--Summarizing the preceding facts, we +find that the Neotropical region possesses no less than 45 families and +more than 900 genera of Vertebrata which are altogether peculiar to it; +while it has representatives of 168 families out of a total of 330, showing +that 162 families are altogether absent. It has also representatives of 131 +genera of Mammalia of which 103 are peculiar to it, a proportion of 4/5; +while of 683 genera of land-birds no less than 576 are peculiar, being +almost exactly 5/6 of the whole. These numbers and proportions are far +higher than in the case of any other region. + + +_Insects._ + +The Neotropical region is so excessively rich in insect life, it so abounds +in peculiar groups, in forms of exquisite beauty, and in an endless +profusion of species, that no adequate idea of this branch of its fauna can +be conveyed by the mere enumeration of peculiar and characteristic groups, +to which we are here compelled to limit ourselves. Our facts and figures +will, however, furnish data for comparison; and will thus enable those who +have some knowledge of the entomology of any other country, to form a +better notion of the vast wealth of insect life in this region, than a more +general and picturesque description could afford them. + +_Lepidoptera._--The Butterflies of South America surpass those of all other +regions in numbers, variety and beauty; and we find here, not only more +peculiar genera and families than elsewhere, but, what is very remarkable, +a fuller representation of the whole series of families. Out of the 16 +families of butterflies in all parts of the world, 13 are found here, and 3 +of these are wholly peculiar--Brassolidæ, Heliconidæ, and Eurygonidæ, with +a fourth, Erycinidæ, which only extends into the Nearctic {14}region; so +that there are 4 families peculiar to America. These four families comprise +68 genera and more than 800 species; alone constituting a very important +feature in the entomology of the region. But in almost all the other +families there are numbers of peculiar genera, amounting in all to about +200, or not far short of half the total number of genera in the +world--(431). We must briefly notice some of the peculiarities of the +several families, as represented in this region. The Danaidæ consist of 15 +genera, all peculiar, and differing widely from the generally sombre-tinted +forms of the rest of the world. The delicate transparent-winged Ithomias of +which 160 species are described, are the most remarkable. _Melinæa_, +_Napeogenes_, _Ceratina_ and _Dircenna_ are more gaily coloured, and are +among the chief ornaments of the forests. The Satyridæ are represented by +25 peculiar genera, many of great beauty; the most remarkable and elegant +being the genus _Hætera_ and its allies, whose transparent wings are +delicately marked with patches of orange, pink, or violet. The genus +_Morpho_ is perhaps the grandest development of the butterfly type, being +of immense size and adorned with the most brilliant azure tints, which in +some species attain a splendour of metallic lustre unsurpassed in nature. +The Brassolidæ are even larger, but are crepuscular insects, with rich +though sober colouring. The true Heliconii are magnificent insects, most +elegantly marked with brilliant and strongly contrasted tints. The +Nymphalidæ are represented by such a variety of gorgeous insects that it is +difficult to select examples. Prominent are the genera _Catagramma_ and +_Callithea_, whose exquisite colours and symmetrical markings are unique +and indescribable; and these are in some cases rivalled by _Agrias_ and +_Prepona_, which reproduce their style of coloration although not closely +allied to them. The Erycinidæ, consisting of 59 genera and 560 species, +comprise the most varied and beautiful of small butterflies; and it would +be useless to attempt to indicate the unimaginable combinations of form and +colour they present. It must be sufficient to say that nothing elsewhere on +the globe at all resembles them. In Lycænidæ the world-wide genus _Thecla_ +is wonderfully developed, and the South {15}American species not only +surpass all others in size and beauty, but some of them are so gorgeous on +the under surface of their wings, as to exceed almost all the combinations +of metallic tints we meet with in nature. The last family, Hesperidæ, is +also wonderfully developed here, the species being excessively numerous, +while some of them redeem the character of this generally sober family, by +their rich and elegant coloration. + +In the only other group of Lepidoptera we can here notice, the Sphingina, +the Neotropical region possesses some peculiar forms. The magnificent +diurnal butterfly-like moths, _Urania_, are the most remarkable; and they +are rendered more interesting by the occurrence of a species closely +resembling them in Madagascar. Another family of day-flying moths, the +Castniidæ, is almost equally divided between the Neotropical and Australian +regions, although the genera are more numerous in the latter. The American +Castnias are large, thick-bodied insects, with a coarse scaly surface and +rich dull colours; differing widely from the glossy and gaily coloured +Agaristas, which are typical of the family in the East. + +_Coleoptera._--This is so vast a subject that, as in the case of the +regions already treated, we must confine our attention to a few of the more +important and best known families as representatives of the entire order. + +Cicindelidæ.--We find here examples of 15 out of the 35 genera of these +insects; and 10 of these genera are peculiar. The most important are +_Oxychila_ (11 sp.), _Hiresia_ (14 sp.), and _Ctenostoma_ (26 sp.). +_Odontochila_ (57 sp.) is the most abundant and characteristic of all, but +is not wholly peculiar, there being a species in the Malay archipelago. +_Tetracha_, another large genus, has species in Australia and a few in +North America and Europe. The small genus _Peridexia_ is divided between +Brazil and Madagascar,--a somewhat similar distribution to that of _Urania_ +noticed above. One genus, _Agrius_, is confined to the southern extremity +of the continent. + +Carabidæ.--Besides a considerable number of cosmopolitan or wide-spread +genera, this family is represented by more than 100 genera which are +peculiar to the Neotropical region. The {16}most important of these are +_Agra_ (150 sp.), _Ardistonus_ (44 sp.), _Schizogenius_ (25 sp.), +_Pelecium_ (24 sp.), _Calophena_ (22 sp.), _Aspidoglossa_ (21 sp.), and +_Lia_, _Camptodonotus_, _Stenocrepis_, and _Lachnophorus_, with each more +than 12 species. These are all tropical; but there are also a number of +genera (26) peculiar to Chili and South Temperate America. The most +important of these are _Antarctia_ (29 sp,), all except two or three +confined to South Temperate America; _Scelodontis_ (10 sp.), mostly +Chilian; _Feronomorpha_ (6 sp.) all Chilian; and _Tropidopterus_ (4 sp.), +all Chilian. _Helluomorpha_ (18 sp.), is confined to North and South +America; _Galerita_, _Callida_, and _Tetragonoderus_, are large genera +which are chiefly South American but with a few species scattered over the +other tropical regions, _Casnonia_ and _Lebia_ are cosmopolite, but most +abundant in South America. _Pachyteles_ is mostly South American but with a +few species in West Africa; while _Lobodonotus_ has one species in South +America and two in Africa. + +Lucanidæ.--The Neotropical species of this family almost all belong to +peculiar genera. Those common to other regions are _Syndesus_, confined to +Tropical South America and Australia, and _Platycerus_ which is Palæarctic +and Nearctic, with one species in Brazil. The most remarkable genus is +undoubtedly _Chiasognathus_, confined to Chili. These are large insects of +metallic green colours, and armed with enormous serrated mandibles. The +allied genera, _Pholidotus_ and _Sphenognathus_, inhabit Tropical South +America. _Streptocerus_ confined to Chili, is interesting, as being allied +to the Australian _Lamprima_. The other genera present no remarkable +features; but _Sclerognathus_ and _Leptinoptera_ are the most extensive. + +Cetoniidæ.--These magnificent insects are but poorly represented in +America; the species being mostly of sombre colours. There are 14 genera, +12 of which are peculiar. The most extensive genus is _Gymnetis_, which, +with its allies _Cotinis_ and _Allorhina_, form a group which comprehends +two-thirds of the Neotropical species of the family. The only other genera +of importance are, _Inca_ (7 sp.), remarkable for their large size, and +being the only American group in which horns are developed on the head; +{17}and _Trigonopeltastes_ (6 sp.), allied to the European _Trichius_. The +non-peculiar genera are, _Stethodesma_, of which half the species are +African and half tropical American; and _Euphoria_, confined to America +both North and South. + +Buprestidæ.--In this fine group the Neotropical region is tolerably rich, +having examples of 39 genera, 18 of which are peculiar to it. Of these, the +most extensive are _Conognatha_ and _Halecia_, which have a wide range over +most parts of the region; and _Dactylozodes_, confined to the south +temperate zone. Of important genera which range beyond the region, +_Dicerca_ is mainly Nearctic and Palæarctic; _Cinyra_ has a species in +North America and one in Australia; _Curis_ is divided between Chili and +Australia; the Australian genus _Stigmodera_ has a species in Chili; +_Polycesta_ has a species in Madagascar, two in the Mediterranean region, +and a few in North America; _Acherusia_ is divided between Australia and +Brazil; _Ptosima_ has one species in south temperate America, the rest +widely scattered from North America to the Philippines; _Actenodes_ has a +single species in North America and another in West Africa; _Colobogaster_ +has two in West Africa, one in Java and one in the Moluccas. The relations +of South America and Australia as indicated by these insects has already +been sufficiently noticed under the latter region. + +Longicornia.--The Neotropical Longicorn Coleoptera are overwhelming in +their numbers and variety, their singularity and their beauty. In the +recent Catalogue of Gemminger and Harold, it is credited with 516 genera, +489 of which are peculiar to it; while it has only 5 genera in common +(exclusively) with the Nearctic, and 4 (in the same way) with the +Australian region. Only the more important genera can be here referred to, +under the three great families into which these insects are divided. + +The Prionidæ are excessively numerous, being grouped in 64 genera, more +than double the number possessed by any other region; and 61 of these are +peculiar. The three, common to other regions, are, _Parandra_ and +_Mallodon_, which are widely distributed; and _Ergates_, found also in +California and Europe. The most remarkable genera are, the +magnificently-coloured _Psalidognathus_ and _Pyrodes_; the large and +strangely marked {18}_Macrodontia_; and _Titanus_, the largest insect of +the entire family. + +Of the Cerambycidæ there are 233 genera, exceeding by one-half, the number +in any other region; and 225 of these are peculiar. Only 2 are common to +the Neotropical and Nearctic regions exclusively, and 3 to the Neotropical +and Australian. The most extensive genera are the elegant _Ibidion_ (80 +sp.); the richly-coloured _Chrysoprasis_ (47 sp.); the prettily-marked +_Trachyderes_ (53 sp.); with _Odontocera_ (25 sp.); _Criodon_ (22 sp.); and +a host of others of less extent, but often of surpassing interest and +beauty. The noteworthy genera of wide range are, _Oeme_ and _Cyrtomerus_, +which have each a species in West Africa, and _Hammatocerus_, which has one +in Australia. + +The Lamiidæ have 219 genera, and this is the only tropical region in which +they do not exceed the Cerambycidæ. This number is almost exactly the same +as that of the Oriental genera, but here there are more peculiar groups, +203 against 160 in the other region. The most extensive genera are +_Hemilophus_ (80 sp.), _Colobothea_ (70 sp.), _Acanthoderes_ (56 sp.), +_Oncoderes_ (48 sp.), _Lepturgus_ (40 sp.), _Hypsioma_ (32 sp.), and +_Tæniotes_ (20 sp.). _Macropus longimanus_, commonly called the harlequin +beetle, is one of the largest and most singularly-marked insects in the +whole family. _Leptostylus_ has a single species in New Zealand; +_Acanthoderes_ has one species in Europe, W. Africa, and Australia, +respectively; _Spalacopsis_ has a species in W. Africa; _Pachypeza_ is +common to S. America and the Philippines; _Mesosa_ is Oriental and +Palæarctic, but has one species on the Amazon; _Apomecyna_ ranges through +the tropics of the Eastern Hemisphere, but has two species in S. America; +_Acanthocinus_ has one species in Tasmania, and the rest in South America, +North America, and Europe; _Phæa_ is wholly Neotropical, except two species +in the Philippine Islands. + +_General Conclusions as to the Neotropical Insect-fauna._--Looking at the +insects of the Neotropical region as a whole, we are struck with the vast +amount of specialty they present; and, considering how many causes there +are which must lead to the dispersal of insects, the number of its groups +which are scattered {19}over the globe is not nearly so great as we might +expect. This points to a long period of isolation, during which the various +forms of life have acted and reacted on each other, leading to such a +complex yet harmoniously-balanced result as to defy the competition of the +chance immigrants that from time to time must have arrived. This is quite +in accordance with the very high antiquity we have shown most insect-forms +to possess; and it is no doubt owing to this antiquity, that such a +complete diversity of _generic_ forms has been here brought about, without +any important deviation from the great _family_ types which prevail over +the rest of the globe. + + +_Land Shells._--The Neotropical region is probably the richest on the globe +in Terrestrial Mollusca, but this is owing, not to any extreme +productiveness of the equatorial parts of the continent, where almost all +other forms of life are so largely developed, but to the altogether +exceptional riches of the West India Islands. The most recent estimates +show that the Antilles contain more species of land shells than all the +rest of the region, and almost exactly as many as all continental America, +north and south. + +Mr. Thomas Bland, who has long studied American land shells, points out a +remarkable difference in the distribution of the Operculated and +Inoperculated groups, the former being predominant on the islands, the +latter on the continent. The Antilles possess over 600 species of +Operculata, to about 150 on the whole American continent, the genera being +as 22 to 14. Of Inoperculata the Antilles have 740, the Continent 1,250, +the genera being 18 and 22. The proportions of the two groups in each +country are, therefore: + + West India Islands. American Continent. + + Operculata Gen. 22 Sp. 608 14 151 + Inoperculata " 18 " 737 22 1251 + +The extensive family of the Helicidæ is represented by 22 genera, of which +6 are peculiar. _Spiraxis_ is confined to Central America and the Antilles; +_Stenopus_ and _Sagda_ are Antillean only; _Orthalicus_, _Macroceramus_, +and _Bulimulus_ have a wider range, the last two extending into the +southern United {20}States. Important and characteristic genera are, +_Glandina_, in all the tropical parts of the region; _Cylindrella_, in +Central America and the Antilles; _Bulimus_, containing many large and +handsome species in South America; _Stenogyra_, widely spread in the +tropics; and _Streptaxis_, in Tropical South America. + +Among the Operculata, the Aciculidæ are mostly Antillean, two genera being +peculiar there, and one, _Truncatella_, of wide distribution, but most +abundant in the West Indian Islands. The Cyclostomidæ are represented by 15 +genera, 9 being peculiar to the region, and 5 of these (belonging to the +sub-family Licinidæ) to the Antilles only. Of these peculiar genera +_Cistula_ and _Chondropoma_ are the most important, ranging over all the +tropical parts of the region. Other important genera are _Cyclotus_ and +_Megalomastoma_; while _Cyclophorus_ also occurs all over the region. The +Helicinidæ are mostly Neotropical, six out of the seven genera being found +here, and four are peculiar. _Stoastoma_, is one of the largest genera; +and, with _Trochatella_ and _Alcadia_, is confined to the Antilles, while +the wide-spread _Helicina_ is most abundant there. + +The Limacidæ, or Old World slugs, are absent from the region, their place +being taken by the allied family, Oncidiadæ. + +_Marine Shells._--We go out of our usual course to say a few words about +the marine shells of this region, because their distribution on the two +sides of the continent is important, as an indication of the former +separation of North and South America, and the connection of the Atlantic +and Pacific Oceans. It was once thought that no species of shells were +common to the two sides of the Central American Isthmus, and Dr. Mörch +still holds that opinion; but Dr. Philip Carpenter, who has paid special +attention to the subject, considers that there are at least 35 species +absolutely identical, while as many others are so close that they may be +only varieties. Nearly 70 others are distinct but representative species. +The genera of marine mollusca are very largely common to the east and west +coasts, more than 40 being so named in the lists published by Mr. Woodward. +The West Indian Islands being a rich shell district, produce a number of +peculiar forms, and the west coast of {21}South America is, to some extent, +peopled by Oriental and Pacific genera of shells. On the west coast there +is hardly any coral, while on the east it is abundant, showing a difference +of physical conditions that must have greatly influenced the development of +mollusca. When these various counteracting influences are taken into +consideration, the identity or close affinity of about 140 species and 40 +genera on the two sides of the Isthmus of Panama becomes very important; +and, combined with the fact of 48 species of fish (or 30 per cent. of those +known) being identical on the adjacent coasts of the two oceans (as +determined by Dr. Günther), render it probable that Central America has +been partially submerged up to comparatively recent geological times. Yet +another proof of this former union of two oceans is to be found in the +fossil corals of the Antilles of the Miocene age, which Dr. Duncan finds to +be more allied to existing Pacific forms, than to those of the Atlantic or +even of the Caribbean Sea. + + +NEOTROPICAL SUB-REGIONS. + +In the concluding part of this work devoted to geographical zoology, the +sub-regions are arranged in the order best adapted to exhibit them in a +tabular form, and to show the affinities of the several regions; but for +our present purpose it will be best to take first in order that which is +the most important and most extensive, and which exhibits all the peculiar +characteristics of the region in their fullest development. We begin +therefore with our second division. + + +_II. Tropical South-America, or the Brazilian Sub-region._ + +This extensive district may be defined as consisting of all the tropical +forest-region of South America, including all the open plains and pasture +lands, surrounded by, or intimately associated with, the forests. Its +central mass consists of the great forest-plain of the Amazons, extending +from Paranaiba on the north coast of Brazil (long. 42° W.) to Zamora, in +the province of Loja (lat. 4° S., long. 79° W.), high up in the Andes, on +the west;--a distance in a straight line of more than 2,500 English miles, +{22}along the whole of which there is (almost certainly) one continuous +virgin forest. Its greatest extent from north to south, is from the mouths +of the Orinooko to the eastern slopes of the Andes near La Paz in Bolivia +and a little north of Sta. Cruz de la Sierra (lat. 18° S.), a distance of +about 1,900 miles. Within this area of continuous forests, are included +some open "campos," or patches of pasture lands, the most important +being,--the Campos of the Upper Rio Branco on the northern boundary of +Brazil; a tract in the interior of British Guiana; and another on the +northern bank of the Amazon near its mouth, and extending some little +distance on its south bank at Santarem. On the northern bank of the +Orinooko are the Llanos, or flat open plains, partly flooded in the rainy +season; but much of the interior of Venezuela appears to be forest country. +The forest again prevails from Panama to Maracaybo, and southwards in the +Magdalena valley; and on all the western side of the Andes to about 100 +miles south of Guayaquil. On the N.E. coast of Brazil is a tract of open +country, in some parts of which (as near Ceara) rain does not fall for +years together; but south of Cape St. Roque the coast-forests of Brazil +commence, extending to lat. 30° S., clothing all the valleys and hill sides +as far inland as the higher mountain ranges, and even penetrating up the +great valleys far into the interior. To the south-west the forest country +reappears in Paraguay, and extends in patches and partially wooded country, +till it almost reaches the southern extension of the Amazonian forests. The +interior of Brazil is thus in the position of a great island-plateau, +rising out of, and surrounded by, a lowland region of ever-verdant forest. +The Brazilian sub-region comprises all this forest-country and its included +open tracts, and so far beyond it as there exists sufficient woody +vegetation to support its peculiar forms of life. It thus extends +considerably beyond the tropic in Paraguay and south Brazil; while the +great desert of Chaco, extending from 25° to 30° S., lat. between the +Parana and the Andes, as well as the high plateaus of the Andean range, +with the strip of sandy desert on the Pacific coast as far as to about 5° +of south latitude, belong to south temperate America, or the sub-region of +the Andes. + +{23}Having already given a sketch, of the zoological features of the +Neotropical region as a whole, the greater part of which will apply to this +sub-region, we must here confine ourselves to an indication of the more +important groups which, on the one hand, are confined to it, and on the +other are absent; together with a notice of its special relations to other +regions. + +_Mammalia._--Many of the most remarkable of the American monkeys are +limited to this sub-region; as _Lagothrix_, _Pithecia_, and _Brachyurus_, +limited to the great Amazonian forests; _Eriodes_ to south-east Brazil; and +_Callithrix_ to tropical South America. All the marmosets (Hapalidæ) are +also confined to this sub-region, one only being found at Panama, and +perhaps extending a little beyond it. Among other peculiar forms, are 8 +genera of bats; 3 peculiar forms of wild dog; _Pteronura_, a genus of +otters; _Inia_, a peculiar form of dolphin inhabiting the upper waters of +the Amazon; tapirs of the genus _Tapirus_ (a distinct genus being found +north of Panama); 4 genera of Muridæ; _Ctenomys_, a genus of Octodontidæ; +the whole family of Echimyidæ, or spiny rats, (as far as the American +continent is concerned) consisting of 8 genera and 28 species; _Chætomys_, +a genus of Cercolabidæ; the capybara (_Hydrochoerus_) the largest known +rodent, belonging to the Caviidæ; the larger ant-eaters (_Myrmecophaga_); +sloths of the genus _Bradypus_; 2 genera of armadillos (Dasypodidæ); and +two peculiar forms of the opossum family (Didelphyidæ). No group that is +typically Neotropical is absent from this sub-region, except such as are +peculiar to other single sub-regions and which will be noticed accordingly. +The occurrence of a solitary species of hare (_Lepus braziliensis_) in +central Brazil and the Andes, is remarkable, as it is cut off from all its +allies, the genus not being known to occur elsewhere on the continent +further south than Costa Rica. The only important external relation +indicated by the Mammalia of this sub-region is towards the Ethiopian +region, 2 genera of Echimyidæ, _Aulacodes_ and _Petromys_, occurring in +South and South-east Africa. + +_Plate IV. Characteristic Neotropical Mammalia._--Our illustration +represents a mountainous forest in Brazil, the part of South America where +the Neotropical Mammalia are perhaps best {24}developed. The central and +most conspicuous figure is the collared ant-eater, (_Tamandua +tetradactyla_), one of the handsomest of the family, in its conspicuous +livery of black and white. To the left are a pair of sloths (_Arctopithecus +flaccidus_) showing the curious black spot on the back with which many of +the species are marked, and which looks like a hole in the trunk of a tree; +but this mark seems to be only found on the male animal. The fur of many of +the sloths has a greenish tinge, and Dr. Seemann remarked its resemblance +to the _Tillandsia usneoides_, or "vegetable horsehair," which clothes many +of the trees in Central America; and this probably conceals them from their +enemies, the harpy-eagles. On the right are a pair of opossums (_Didelphys +azaræ_), one of them swinging by its prehensile tail. Overhead in the +foreground are a group of howling monkeys (_Mycetes ursinus_) the largest +of the American Quadrumana, and the noisiest of monkeys. The large hollow +vessel into which the hyoid bone is transformed, and which assists in +producing their tremendous howling, is altogether unique in the animal +kingdom. Below them, in the distance, are a group of Sapajou monkeys +(_Cebus_ sp.); while gaudy screaming macaws complete the picture of +Brazilian forest life. + +_Birds._--A very large number of genera of birds, and some entire families, +are confined to this sub-region, as will be seen by looking over the list +of genera at the end of this chapter. We can here only notice the more +important, and summarize the results. More than 120 genera of Passeres are +thus limited, belonging to the following 12 families: Sylviidæ (1), +Troglodytidæ (2), Coerebidæ (4), Tanagridæ (26), Fringillidæ (8), Icteridæ +(5), Pteroptochidæ (3), Dendrocolaptidæ (12), Formicariidæ (16), Tyrannidæ +(22), Cotingidæ (16), Pipridæ (10). Of the Picariæ there are 76 peculiar +genera belonging to 9 families, viz., Picidæ (2), Rhamphastidæ (1), +Cuculidæ (1), Bucconidæ (2), Galbulidæ (5), Momotidæ (2), Podargidæ (1), +Caprimalgidæ (4), Trochilidæ (58). There are 3 peculiar genera of Psittaci, +8 of Gallinæ, the only genus of Opisthocomidæ, 3 of Accipitres, 1 of +Rallidæ, _Psophia_ and _Eurypyga_ types of distinct families, and 1 genus +of Ardeidæ, Palamedeidæ, and Anatidæ respectively. The preceding +enumeration shows how very rich this sub-region is in peculiar types of all +the most characteristic American families, such as the Tanagridæ, +Tyrannidæ, Cotingidæ, Formicariidæ, Trochilidæ, and Galbulidæ. A +considerable proportion of the genera of the Chilian and Mexican +sub-regions also occur here, so that out of about 680 genera of Neotropical +land-birds more than 500 are represented in this sub-region. + + + +Plate XIV. + +[Illustration] + +A BRAZILIAN FOREST, WITH CHARACTERISTIC MAMMALIA. + +{25}Without entering minutely into the distribution of species it is +difficult to sub-divide this extensive territory with any satisfactory +result.[1] The upland tract between the Amazon and Orinooko, which may be +termed Guiana, was evidently once an island, yet it possesses few marked +distinctive features. Brazil, which must have formed another great island, +has more speciality, but the intermediate Amazonian forests form a perfect +transition between them. The northern portion of the continent west of the +Orinooko has more character; and there are indications that this has +received many forms from Central and North America, and thus blended two +faunas once more distinct than they are now. The family of wood-warblers +(Mniotiltidæ) seems to have belonged to this more northern fauna; for out +of 18 genera only 5 extend south of the equator, while 6 range from Mexico +or the Antilles into Columbia, some of these being only winter immigrants +and no genus being exclusively South American. The eastern slopes of the +Andes constitute, however, the richest and best marked province of this +sub-region. At least 12 genera of tanagers (Tanagridæ) are found here only, +with an immense number of Fringillidæ,--the former confined to the forests; +the latter ranging to the upland plains. The ant-thrushes (Formicariidæ) on +the other hand seem more abundant in the lowlands, many genera being +peculiar to the Amazonian forests. The superb chatterers (Cotingidæ) also +seem to have their head-quarters in the forests of Brazil and Guiana, and +to have thence spread {26}into the Amazonian valley. Guiana still boasts +such remarkable forms as the cardinal chatterer (_Phoenicocercus_), the +military chatterer (_Hæmatoderus_), as well as _Querula_, _Gymnoderus_, and +_Gymnocephalus_; but the first three pass to the south side of the Lower +Amazon. Here also belong the cock of the rock (_Rupicola_), which ranges +from Guiana to the Andes, and the marvellous umbrella-birds of the Rio +Nigro and Upper Amazon (_Cephalopterus_), which extends across the +Ecuadorean Andes and into Costa Rica. Brazil has _Ptilochloris_, +_Casiornis_, _Tijuca_, _Phibalura_, and _Calyptura_; while not a single +genus of this family, except perhaps _Heliochæra_, is confined to the +extensive range of the Andes. Almost the same phenomena are presented by +the allied Pipridæ or manakins, the greater part of the genera and species +occurring in Eastern South America, that is in Brazil, Guiana, and the +surrounding lowlands rather than in the Andean valleys. The same may be +said of the jacamars (Galbulidæ) and puff-birds (Bucconidæ); but the +humming-birds (Trochilidæ) have their greatest development in the Andean +district. Brazil and Guiana have each a peculiar genus of parrots; Guiana +has three peculiar genera of Cracidæ, while the Andes north of the equator +have two. The Tinamidæ on the other hand have their metropolis in Brazil, +which has two or three peculiar genera, while two others seem confined to +the Andes south of the equator. The elegant trumpeters (Psophiidæ) are +almost restricted to the Amazonian valley. + +Somewhat similar facts occur among the Mammalia. At least 3 genera of +monkeys are confined to the great lowland equatorial forests and 1 to +Brazil; _Icticyon_ (Canidæ) and _Pteronura_ (Mustelidæ) belong to Guiana +and Brazil; and most of the Echimyidæ are found in the same districts. The +sloths, ant-eaters, and armadillos all seem more characteristic of the +eastern districts than of the Andean; while the opossums are perhaps +equally plentiful in the Andes. + +The preceding facts of distribution lead us to conclude that the highlands +of Brazil and of Guiana represent very ancient lands, dating back to a +period long anterior to the elevation of the Andean range (which is by no +means of great geological {27}antiquity) and perhaps even to the elevation +of the continuous land which forms the base of the mountains. It was, no +doubt, during their slow elevation and the consequent loosening of the +surface, that the vast masses of debris were carried down which filled up +the sea separating the Andean chain from the great islands of Brazil and +Guiana, and formed that enormous extent of fertile lowland forest, which +has created a great continent; given space for the free interaction of the +distinct faunas which here met together, and thus greatly assisted in the +marvellous development of animal and vegetable life, which no other +continent can match. But this development, and the fusion of the various +faunas into one homogeneous assemblage must have been a work of time; and +it is probable that most of the existing continent was dry land before the +Andes had acquired their present altitude. The blending of the originally +distinct sub-faunas has been no doubt assisted by elevations and +depressions of the land or of the ocean, which have alternately diminished +and increased the land-area. This would lead to a crowding together at one +time, and a dispersion at others, which would evidently afford opportunity +for many previously restricted forms to enter fresh areas and become +adapted to new modes of life. + +From the preceding sketch it will appear, that the great sub-region of +Tropical South America as here defined, is really formed of three +originally distinct lands, fused together by the vast lowland Amazonian +forests. In the class of birds sufficient materials exist for separating +these districts; and that of the Andes contains a larger series of peculiar +genera than either of the other sub-regions here adopted. But there are +many objections to making such a sub-division here. It is absolutely +impossible to define even approximate limits to these divisions--to say for +example where the "Andes" ends and where "Brazil" or "Amazonia" or "Guiana" +begins; and the unknown border lands separating these are so vast, that +many groups, now apparently limited in their distribution, may prove to +have a very much wider range. In mammalia, reptiles, and insects, it is +even more difficult to maintain such divisions, so that on the whole it +seems better to treat the entire area as one sub-region, {28}although +recognizing the fact of its zoological and geographical diversity, as well +as its vast superiority over every other sub-region in the number and +variety of its animal forms. + +The reptiles, fishes, mollusca, and insects of this sub-region have been +sufficiently discussed in treating of the entire region, as by far the +larger proportion of them, except in the case of land-shells, are found +here. + +_Plate XV. Characteristic Neotropical Birds._--To illustrate the +ornithology of South America we place our scene on one of the tributaries +of the Upper Amazon, a district where this class of animals is the most +prominent zoological feature, and where a number of the most remarkable and +interesting birds are to be found. On the left we have the umbrella-bird +(_Cephalopterus ornatus_), so called from its wonderful crest, which, when +expanded, completely overshadows its head like an umbrella. It is also +adorned with a long tassel of plumes hanging from its breast, which is +formed by a slender fleshy tube clothed with broad feathers. The bird is as +large as a crow, of a glossy blue-black colour, and belongs to the same +family as the exquisitely tinted blue-and-purple chatterers. Flying towards +us are a pair of curl-crested toucans (_Pteroglossus beauharnaisii_), +distinguished among all other toucans by a crest composed of small black +and shining barbless plumes, resembling curled whalebone. The general +plumage is green above, yellow and red beneath, like many of its allies. To +the right are two of the exquisite little whiskered hummers, or +"frill-necked coquettes," as they are called by Mr. Gould, (_Lophornis +gouldi_). These diminutive birds are adorned with green-tipped plumes +springing from each side of the throat, as well as with beautiful crests, +and are among the most elegant of the great American family of +humming-birds, now numbering about 400 known species. Overhead are perched +a pair of curassows (_Crax globulosa_), which represent in America the +pheasants of the Old World. There are about a dozen species of these fine +birds, most of which are adorned with handsome curled crests. That figured, +is distinguished by the yellow caruncular swellings at the base of the +bill. The tall crane-like bird near the water is one of the trumpeters, +(_Psophia leucoptera_), elegant birds with silky plumage peculiar to the +Amazon valley. They are often kept in houses, where they get very tame and +affectionate; and they are useful in catching flies and other house +insects, which they do with great perseverance and dexterity. + + + +Plate XV. + +[Illustration] + +A FOREST SCENE ON THE UPPER AMAZON, WITH SOME CHARACTERISTIC BIRDS. + + +{29}_Islands of Tropical South America._ + +These are few in number, and, with one exception, not of much interest. +Such islands as Trinidad and Sta. Catherina form parts of South America, +and have no peculiar groups of animals. The small islands of Fernando +Noronha, Trinidad, and Martin Vaz, off the coast of Brazil, are the only +Atlantic islands somewhat remote from land; while the Galapagos Archipelago +in the Pacific is the only group whose productions have been carefully +examined, or which present features of special interest. + +_Galapagos Islands._--These are situated on the equator, about 500 miles +from the coast of Ecuador. They consist of the large Albemarle island, 70 +miles long; four much smaller (18 to 25 miles long), named Narborough, +James, Indefatigable, and Chatham Islands; four smaller still (9 to 12 +miles long), named Abingdon, Bindloes, Hood's, and Charles Islands. All are +volcanic, and consist of fields of black basaltic lava, with great numbers +of extinct craters, a few which are still active. The islands vary in +height from 1,700 to 5,000 feet, and they all rise sufficiently high to +enter the region of moist currents of air, so that while the lower parts +are parched and excessively sterile, above 800 or 1,000 feet there is a +belt of comparatively green and fertile country. + +These islands are known to support 58 species of Vertebrates,--1 quadruped, +52 birds and 5 reptiles, the greater part of which are found nowhere else, +while a considerable number belong to peculiar and very remarkable genera. +We must therefore notice them in some detail. + +_Mammalia._--This class is represented by a mouse belonging to the American +genus _Hesperomys_, but slightly different from any found on the continent. +A true rat (_Mus_), slightly differing from any European species, also +occurs; and as there can be little doubt that this is an escape from a +ship, somewhat {30}changed under its new conditions of life (the genus +_Mus_ not being indigenous to the American continent), it is not +improbable, as Mr. Darwin remarks, that the American mouse may also have +been imported by man, and have become similarly changed. + +_Birds._[2]--Recent researches in the islands have increased the number of +land-birds to thirty-two, and of wading and aquatic birds to twenty-three. +All the land birds but two or three are peculiar to the islands, and +eighteen, or considerably more than half, belong to peculiar genera. Of the +waders 4 are peculiar, and of the swimmers 2. These are a rail (_Porzana +spilonota_); two herons (_Butorides plumbea_ and _Nycticorax pauper_); a +flamingo (_Phoenicopterus glyphorhynchus_); while the new aquatics are a +gull (_Larus fuliginosus)_, and a penguin (_Spheniscus mendiculus_). + +The land-birds are much more interesting. All except the birds of prey +belong to American genera which abound on the opposite coast or on that of +Chili a little further south, or to peculiar genera allied to South +American forms. The only _species_ not peculiar are, _Dolichonyx +oryzivorus_, a bird of very wide range in America and of migratory habits, +which often visits the Bermudas 600 miles from North America,--and _Asio +accipitrinus_, an owl which is found almost all over the world. The only +genera not exclusively American are _Buteo_ and _Strix_, of each of which a +peculiar species occurs in the Galapagos, although very closely allied to +South American species. There remain 10 genera, all either American or +peculiar to the Galapagos; and on these we will remark in systematic order. + +1. _Mimus_, the group of American mocking-thrushes, is represented by three +distinct and well-marked species. 2. _Dendroeca_, an extensive and +wide-spread genus of the wood-warblers (Mniotiltidæ), is represented by one +species, which ranges over the greater part of the archipelago. The genus +is especially abundant in Mexico, the Antilles, and the northern parts of +{31}tropical America, only one species extending south as far as Chili. 3. +_Certhidea_, a peculiar genus originally classed among the finches, but +which Mr. Sclater, who has made South American birds his special study, +considers to belong to the _Coerebidæ_, or sugar-birds, a family which is +wholly tropical. Two species of this genus inhabit separate islands. 4. +_Progne_, the American martins (Hirundinidæ), is represented by a peculiar +species. 5. _Geospiza_, a peculiar genus of finches, of which no less than +eight species occur in the archipelago, but not more than four in any one +island. 6. _Camarhynchus_ (6 sp.) and 7. _Cactornis_ (4 sp.) are two other +peculiar genera of finches; some of the species of which are confined to +single islands, while others inhabit several. 8. _Pyrocephalus_, a genus of +the American family of tyrant-flycatchers (Tyrannidæ), has one peculiar +species closely allied to _T. rubineus_, which has a wide range in South +America. 9. _Myiarchus_, another genus of the same family which does not +range further south than western Ecuador, has also a representative species +found in several of the islands. 10. _Zenaida_, an American genus of +pigeons, has a species in James Island and probably in some of the others, +closely allied to a species from the west coast of America. + +It has been already stated that some of the islands possess peculiar +species of birds distinct from the allied forms in other islands, but +unfortunately our knowledge of the different islands is so unequal and of +some so imperfect, that we can form no useful generalizations as to the +distribution of birds among the islands themselves. The largest island is +the least known; only one bird being recorded from it, one of the +mocking-thrushes found nowhere else. Combining the observations of Mr. +Darwin with those of Dr. Habel and Prof. Sundevall, we have species +recorded as occurring in seven of the islands. Albemarle island has but one +definitely known species; Chatham and Bindloe islands have 11 each; +Abingdon and Charles islands 12 each; Indefatigable island and James island +have each 18 species. This shows that birds are very fairly distributed +over all the islands, one of the smallest and most remote (Abingdon) +furnishing as many as the much larger Chatham Island, which is also the +nearest {32}to the mainland. Taking the six islands which seem tolerably +explored, we find that two of the species (_Dendroeca aureola_ and +_Geospiza fortis_) occur in all of them; two others (_Geospiza strenua_ and +_Myiarchus magnirostris_) in five; four (_Mimus melanotis_, _Geospiza +fuliginosa_, _G. parvula_, and _Camarhynchus prosthemelas_) in four +islands; five (_Certhidea olivacea_, _Cactornis scandens_, _Pyrocephalus +nanus_), and two of the birds of prey, in three islands; nine (_Certhidea +fusca_, _Progne concolor_, _Geospiza nebulosa_, _G. magnirostris_, +_Camarhynchus psittaculus_, _C. variegatus_, _C. habeli_ and _Asio +accipitrinus_) in two islands; while the remaining ten species are confined +to one island each. These peculiar species are distributed among the +islands as follows. James, Charles and Abingdon islands, have 2 each; +Bindloes, Chatham, and Indefatigable, 1 each. The amount of speciality of +James Island is perhaps only apparent, owing to our ignorance of the fauna +of the adjacent large Albemarle island; the most remote islands north and +south, Abingdon and Charles, have no doubt in reality most peculiar +species, as they appear to have. The scarcity of peculiar species in +Chatham Island is remarkable, it being large, very isolated, and the +nearest to the mainland. There is still room for exploration in these +islands, especially in Albemarle, Narborough, and Hood's islands of which +we know nothing. + +_Reptiles._--The few reptiles found in these islands are very interesting. +There are two snakes, a species of the American genus _Herpetodryas_, and +another which was at first thought to be a Chilian species (_Psammophis +Temminckii_), but which is now considered to be distinct. Of lizards there +are four at least, belonging to as many genera. One is a species of +_Phyllodactylus_, a wide-spread genus of Geckotidæ; the rest belong to the +American family of the Iguanas, one being a species of the Neotropical +genus _Leiocephalus_, the other two very remarkable forms, _Trachycephalus_ +and _Oreocephalus_ (formerly united in the genus _Amblyrhynchus_). The +first is a land, the second a marine, lizard; both are of large size and +very abundant on all the islands; and they are quite distinct from any of +the very numerous genera of Iguanidæ, spread all over the American +continent. The last {33}reptile is a land tortoise (_Testudo nigra_) of +immense size, and also abundant in all the islands. Its nearest ally is the +equally large species of the Mascarene Islands; an unusual development due, +in both cases, to the absence of enemies permitting these slow but +continually growing animals to attain an immense age. It is believed that +each island has a distinct variety or species of tortoise. + +_Insects_.--Almost the only insects known from these islands are some +Coleoptera, chiefly collected by Mr. Darwin. They consist of a few peculiar +species of American or wide-ranging genera, the most important being, a +_Calosoma_, _Poecilus_, _Solenophorus_, and _Notaphus_, among the Carabidæ; +an _Oryctes_ among the Lamellicornes; two new genera of obscure Heteromera; +two Curculionidæ of wide-spread genera; a Longicorn of the South American +genus _Eburia_; and two small Phytophaga,--a set of species highly +suggestive of accidental immigrations at rare and distant intervals. + +_Land-Shells._--These consist of small and obscure species, forming two +peculiar sub-genera of _Bulimulus_, a genus greatly developed on the whole +West coast of America; and a single species of _Buliminus_, a genus which +ranges over all the world except America. As in the case of the birds, most +of the islands have two or three peculiar species. + +_General Conclusions._--These islands are wholly volcanic and surrounded by +very deep sea; and Mr. Darwin is of opinion, not only that the islands have +never been more nearly connected with the mainland than at present, but +that they have never been connected among themselves. They are situated on +the Equator, in a sea where gales and storms are almost unknown. The main +currents are from the south-west, an extension of the Peruvian drift along +the west coast of South America. From their great extent, and their +volcanoes being now almost extinct, we may assume that they are of +considerable antiquity. These facts exactly harmonize with the theory, that +they have been peopled by rare accidental immigrations at very remote +intervals. The only peculiar _genera_ consist of birds and lizards, which +must therefore have been the earliest {34}immigrants. We know that small +Passerine birds annually reach the Bermudas from America, and the Azores +from Europe, the former travelling over 600, the latter over 1000 miles of +ocean. These groups of islands are both situated in stormy seas, and the +immigrants are so numerous that hardly any specific change in the resident +birds has taken place. The Galapagos receive no such annual visitants; +hence, when by some rare accident a few individuals of a species did +arrive, they remained isolated, probably for thousands of generations, and +became gradually modified through natural selection under completely new +conditions of existence. Less rare and violent storms would suffice to +carry some of these to other islands, and thus the archipelago would in +time become stocked. It would appear probable, that those which have +undergone most change were the earliest to arrive; so that we might look +upon the three peculiar genera of finches, and _Certhidea_, the peculiar +form of Coerebidæ, as among the most ancient inhabitants of the islands, +since they have become so modified as to have apparently no near allies on +the mainland. But other birds may have arrived nearly at the same time, and +yet not have been much changed. A species of very wide range, already +adapted to live under very varied conditions and to compete with varied +forms of life, might not need to become modified so much as a bird of more +restricted range, and more specialized constitution. And if, before any +considerable change had been effected, a second immigration of the same +species occurred, crossing the breed would tend to bring back the original +type of form. While, therefore, we may be sure that birds like the finches, +which are profoundly modified and adapted to the special conditions of the +climate and vegetation, are among the most ancient of the colonists; we +cannot be sure that the less modified form of tyrant-flycatcher or +mocking-thrush, or even the unchanged but cosmopolitan owl, were not of +coeval date; since even if the parent form on the continent has been +changed, successive immigrations may have communicated the same change to +the colonists. + +The reptiles are somewhat more difficult to account for. We know, however, +that lizards have some means of dispersal over {35}the sea, because we find +existing species with an enormous range. The ancestors of the +_Amblyrhynchi_ must have come as early, probably, as the earliest birds; +and the same powers of dispersal have spread them over every island. The +two American genera of lizards, and the tortoises, are perhaps later +immigrants. Latest of all were the snakes, which hardly differ from +continental forms; but it is not at all improbable that these latter, as +well as the peculiar American mouse, have been early human importations. +Snakes are continually found on board native canoes whose cabins are +thatched with palm leaves; and a few centuries would probably suffice to +produce some modification of a species completely isolated, under +conditions widely different from those of its native country. Land-shells, +being so few and small, and almost all modifications of one type, are a +clear indication of how rare are the conditions which lead to their +dispersal over a wide extent of ocean; since two or three individuals, +arriving on two or three occasions only during the whole period of the +existence of the islands, would suffice to account for the present fauna. +Insects have arrived much more frequently; and this is in accordance with +their habits, their lower specific gravity, their power of flight, and +their capacity for resisting for some time the effects of salt water. + +We learn, then, from the fauna of these islands, some very important facts. +We are taught that tropical land-birds, unless blown out of their usual +course by storms, rarely or never venture out to sea, or if they do so, can +seldom pass safely over a distance of 500 miles. The immigrants to the +Galapagos can hardly have averaged a bird in a thousand years. We learn, +that of all reptiles lizards alone have some tolerably effective mode of +transmission across the sea; and this is probably by means of currents, and +in connection with floating vegetation. Yet their transmission is a far +rarer event than that of land-birds; for, whereas three female immigrants +will account for the lizard population, at least eight or ten ancestors are +required for the birds. Land serpents can pass over still more rarely, as +two such transmissions would have sufficed to stock the islands with their +snakes; and it is not certain that either of these occurred without the aid +of man. {36}It is doubtful whether mammals or batrachians have any means of +passing, independently of man's assistance; the former having but one +doubtfully indigenous representative, the latter none at all. The +remarkable absence of all gay or conspicuous flowers in these tropical +islands, though possessing a zone of fairly luxuriant shrubby vegetation, +and the dependence of this phenomenon on the extreme scarcity of insects, +has been already noticed at Vol. I. p. 461, when treating of a somewhat +similar peculiarity of the New Zealand fauna and flora. + + +_I. South Temperate America, or the Chilian Sub-region._ + +This sub-region may be generally defined as the temperate portion of South +America. On the south, it commences with the cold damp forests of Tierra +del Fuego, and their continuation up the west coast to Chiloe and northward +to near Santiago. To the east we have the barren plains of Patagonia, +gradually changing towards the north into the more fertile, but still +treeless, pampas of La Plata. Whether this sub-region should be continued +across the Rio de la Plata into Uruguay and Entre-rios, is somewhat +doubtful. To the west of the Parana it extends northward over the Chaco +desert, till we approach the border of the great forests near St. Cruz de +la Sierra. On the plateau of the Andes, however, it must be continued still +further north, along the "paramos" or alpine pastures, till we reach 5° of +South latitude. Beyond this the Andes are very narrow, having no double +range with an intervening plateau; and although some of the peculiar forms +of the temperate zone pass on to the equator or even beyond it, these are +not sufficiently numerous to warrant our extending the sub-region to +include them. Along with the high Andes it seems necessary to include the +western strip of arid country, which is mostly peopled by forms derived +from Chili and the south temperate regions. + +_Mammalia._--This sub-region is well characterised by the possession of an +entire family of mammalia having Neotropical affinities--the Chinchillidæ. +It consists of 3 genera--_Chinchilla_ (2 sp.), inhabiting the Andes of +Chili and Peru as far as 9° south latitude, and at from 8,000 to 12,000 +feet altitude; _Lagidium_ (3 sp.), ranging over the Andes of Chili, Peru, +and South Ecuador, {37}from 11,000 to 16,000 feet altitude; and +_Lagostomus_ (1 sp.), the "viscacha," confined to the pampas between the +Uruguay and Rio Negro. Many important genera are also confined to this +sub-region. _Auchenia_ (4 sp.), including the domesticated llamas and +alpacas, the vicugna which inhabits the Andes of Peru and Chili, and the +guanaco which ranges over the plains of Patagonia and Tierra del Fuego. +Although this genus is allied to the Old World camels, it is a very +distinct form, and its introduction from North America, where the family +appear to have originated, may date back to a remote epoch. _Ursus +ornatus_, the "spectacled bear" of the Chilian Andes, is a remarkable form, +supposed to be most allied to the Malay bear, and probably forming a +distinct genus, which has been named _Tremarctos_. Four genera of +Octodontidæ are also peculiar to this sub-region, or almost so; +_Habrocomus_ (1 sp.) is Chilian; _Spalacopus_ (2 sp.) is found in Chili and +on the east side of the southern Andes; _Octodon_ (3 sp.) ranges from Chili +into Peru and Bolivia; _Ctenomys_ (6 sp.) from the Straits of Magellan to +Bolivia, with one species in South Brazil. _Dolichotis_, one of the Cavies, +ranges from Patagonia to Mendoza, and on the east coast to 37½° S. +latitude. _Myopotamus_ (1 sp.), the coypu (Echimyidæ), ranges from 33° to +48° S. latitude on the west side of the Andes, and from the frontiers of +Peru to 42° S. on the east side. _Reithrodon_ and _Acodon_, genera of +Muridæ, are also confined to Temperate South America; _Tolypeutes_ and +_Chlamydophorus_, two genera of armadillos, the latter very peculiar in its +organization and sometimes placed in a distinct family, are found only in +La Plata and the highlands of Bolivia, and so belong to this sub-region. +_Otaria_, one of the "eared seals" (Otariidæ), is confined to the coasts of +this sub-region and the antarctic islands. Deer of American groups extend +as far as Chiloe on the west, and the Straits of Magellan on the east +coast. Mice of the South American genera _Hesperomys_ and _Reithrodon_, are +abundant down to the Straits of Magellan and into Tierra del Fuego, Mr. +Darwin having collected more than 20 distinct species. The following are +the genera of Mammalia which have been observed on the shores of the +Straits of Magellan, those marked * extending into Tierra del Fuego: +{38}*_Pseudalopex_ (two wolf-like foxes), _Felis_ (the puma), _Mephitis_ +(skunks), _Cervus_ (deer), *_Auchenia_ (guanaco), *_Ctenomys_ (tucu-tucu), +*_Reithrodon_ and *_Hesperomys_ (American mice). + +_Birds._--Three families of Birds are confined to this +sub-region,--Phytotomidæ (1 genus, 3 sp.), inhabiting Chili, La Plata, and +Bolivia; Chionididæ (1 genus, 2 sp.) the "sheath-bills," found only at the +southern extremity of the continent and in Kerguelen's Island, which with +the other antarctic lands perhaps comes best here; Thinocoridæ (2 genera, 6 +species) an isolated family of waders, ranging over the whole sub-region +and extending northward to the equatorial Andes. Many genera are also +peculiar: 3 of Fringillidæ, and 1 of Icteridæ; 9 of Dendrocolaptidæ, 6 of +Tyrannidæ, 3 of Trochilidæ, and 4 of Pteroptochidæ,--the last four South +American families. There is also a peculiar genus of parrots +(_Henicognathus_) in Chili; two of pigeons (_Metriopelia_ and _Gymnopelia_) +confined to the Andes and west coast from Peru to Chili; two of Tinamous, +_Tinamotes_ in the Andes, and _Calodromus_ in La Plata; three of +Charadriidæ, _Phægornis_, _Pluvianellus_, and _Oreophilus_; and _Rhea_, the +American ostriches, inhabiting all Patagonia and the pampas. Perhaps the +Cariamidæ have almost as much right here as in the last sub-region, +inhabiting as they do, the "pampas" of La Plata and the upland "campos" of +Brazil; and even among the wide-ranging aquatic birds, we have a peculiar +genus, _Merganetta_, one of the duck family, which is confined to the +temperate plateau of the Andes. + +Against this extensive series of characteristic groups, all either of +American type or very distinct forms of Old World families, and therefore +implying great antiquity, we find, in mammalia and birds, very scanty +evidence of that direct affinity with the north temperate zone, on which +some naturalists lay so much stress. We cannot point to a single +terrestrial genus, which is characteristic of the north and reappears in +this south temperate region without also occurring over much of the +intervening land. _Mustela_ seems only to have reached Peru; _Lepus_ is +isolated in Brazil; true _Ursus_ does not pass south of Mexico. In birds, +the northern groups rarely go further south than Mexico or the Columbian +Andes; and the only case of discontinuous {39}distribution we can find +recorded is that of the genus of ducks, _Camptolæmus_, which has a species +on the east side of North America and another in Chili and the Falkland +Islands, but these, Professor Newton assures me, do not properly belong to +the same genus. Out of 30 genera of land-birds collected on the Rio Negro +in Patagonia, by Mr. Hudson, only four extend beyond the American +continent, and the same exclusively American character applies equally to +its southern extremity. No list appears to have been yet published of the +land-birds of the Straits of Magellan and Tierra del Fuego. The following +is compiled from the observations of Mr. Darwin, the recent voyage of +Professor Cunningham, and other sources; and will be useful for comparison. + + TURDIDÆ. + 1. Turdus falklandicus. + + TROGLODYTIDÆ. + 2. Troglodytes magellanicus. + + FRINGILLIDÆ. + 3. Chrysomitris barbata. + *4. Phrygilus gayi. + *5. " aldunatii. + 6. " fruticeti. + *7. " xanthogrammus. + 8. Zonotrichia pileata. + + ICTERIDÆ. + 9. Sturnella militaris. + 10. Curæus aterrimus. + + HIRUNDINIDÆ. + 11. Hirundo meyeni. + + TYRANNIDÆ. + 12. Tænioptera pyrope. + 13. Myiotheretes rufiventris. + 14. Muscisaxicola mentalis. + 15. Centrites niger. + 16. Anæretes parulus. + 17. Elainea griseogularis. + + DENDROCOLAPTIDÆ. + 18. Upucerthia dumetoria. + *19. Cinclodes patagonicus. + *20. " fuscus. + *21. Oxyurus spinicauda. + + PTEROPTOCHIDÆ. + *22. Scytalopus magellanicus. + + PICIDÆ. + *23. Campephilus magellanicus. + 24. Picus lignarius. + + ALCEDINIDÆ. + 25. Ceryle stellata. + + TROCHILIDÆ. + 26. Eustephanus galeritus. + + CONURIDÆ. + 27. Conurus patagonus. + + VULTURIDÆ. + 28. Cathartes aura. + 29. Sarcorhamphus gryphus. + + FALCONIDÆ. + 30. Circus macropterus. + 31. Buteo erythronotus. + 32. Geranoaëtus melanolencus. + 33. Accipiter chilensis. + 34. Cerchneis sparverius. + 35. Milvago albogularis. + 36. Polyborus tharus. + + STRIGIDÆ. + 37. Asio accipitrinus. + 38. Bubo magellanicus. + 39. Pholeoptynx cunicularia. + 40. Glaucidium nana. + 41. Syrnium rufipes. + + STRUTHIONIDÆ. + 42. Rhea darwinii. + +{40}In the above list the species marked * extend to Tierra del Fuego. It +is a remarkable fact that so many of the species belong to genera which are +wholly Neotropical, and that the specially South American families of +Icteridæ, Tyrannidæ, Dendrocolaptidæ, Pteroptochidæ, Trochilidæ, and +Conuridæ, should supply more than one-third of the species; while the +purely South American genus _Phrygilus_, should be represented by four +species, three of which abound in Tierra del Fuego. + +_Plate XVI. A Scene in the Andes of Chili, with characteristic +Animals._--The fauna of South Temperate America being most fully developed +in Chili, we place the scene of our illustration in that country. In the +foreground we have a pair of the beautiful little chinchillas (_Chinchilla +lanigera_), belonging to a family of animals peculiar to the sub-region. +There are only two species of this group, both confined to the higher +Andes, at about 8000 feet elevation. Coming round a projecting ridge of the +mountain, are a herd of vicunas (_Auchenia vicugna_), one of that peculiar +form of the camel tribe found in South America and confined to its +temperate and alpine regions. The upper bird is a plant-cutter (_Phytotoma +rara_), of sober plumage but allied to the beautiful chatterers, though +forming a separate family. Below, standing on a rock, is a plover-like +bird, the _Thinocorus orbignianus_, which is considered to belong to a +separate family, though allied to the plovers and sheath-bills. Its habits +are, however, more those of the quails or partridges, living inland in dry +and desert places, and feeding on plants, roots, and insects. Above is a +condor, the most characteristic bird of the high Andes. + +_Reptiles and Amphibia._--These groups show, for the most part, similar +modifications of American and Neotropical forms, as those we have seen to +prevail among the birds. Snakes do not seem to go very far south, but +several South American genera of Colubridæ and Dendrophidæ occur in Chili; +while _Enophrys_ is peculiar to La Plata, and _Callorhinus_ to Patagonia, +both belonging to the Colubridæ. The Elapidæ do not extend into the +temperate zone; but _Craspedocephalus_, one of the Crotalidæ, occurs at +Bahia Blanca in Patagonia (Lat. 40° S.) + + + +Plate XVI. + +[Illustration] + +THE CHILIAN ANDES, WITH CHARACTERISTIC ANIMALS. + +{41}Lizards are much more numerous, and there are several peculiar and +interesting forms. Three families are represented; Teidæ by two +genera--_Callopistes_ peculiar to Chili, and _Ameiva_ which ranges over +almost the whole American continent and is found in Patagonia; _Geckotidæ_ +by four genera, two of which,--_Caudiverbera_ and _Homonota_--are peculiar +to Chili, while _Sphærodactylus_ and _Cubina_ are Neotropical, the former +ranging to Patagonia, the latter to Chili; and lastly the American family +Iguanidæ represented by eight genera, no less than six being peculiar, (or +almost so,) to the South temperate region. These are _Leiodera_, +_Diplolæmus_ and _Proctrotretus_, ranging from Chili to Patagonia; +_Leiolæmus_, from Peru to Patagonia; _Phrymaturus_, confined to Chili, and +_Ptygoderus_ peculiar to Patagonia and Tierra del Fuego. The other two +genera, _Oplurus_ and _Leiosaurus_, are common to Chili and tropical South +America. + +Tortoises appear to be scarce, a species of _Hydromedusa_ only being +recorded. Of the Amphibia, batrachia (frogs and toads) alone are +represented, and appear to be tolerably abundant, seventeen species having +been collected by Mr. Darwin in this sub-region. Species of the South +American genera _Phryniscus_, _Hylaplesia_, _Telmatobius_, _Cacotus_, +_Hylodes_, _Cyclorhamphus_, _Pleurodema_, _Cystignathus_, and _Leiuperus_, +are found in various localities, some extending even to the Straits of +Magellan,---the extreme southern limit of both Reptilia and Amphibia, +except one lizard (_Ptygoderus_) found by Professor Cunningham in Tierra +del Fuego. There are also four peculiar genera, _Rhinoderma_ belonging to +the Engystomidæ; _Alsodes_ and _Nannophryne_ to the Bombinatoridæ; +_Opisthodelphys_ to the Hylidæ; and _Calyptocephalus_ to the Discoglossidæ. + +It thus appears, that in the Reptiles all the groups are typically +American, and that most of the peculiar genera belong to families which are +exclusively American. The Amphibia, on the other hand, present some +interesting external relations, but these are as much with Australia as +with the North temperate regions. The Bombinatoridæ are indeed Palæarctic, +but a larger proportion are Neotropical, and one genus inhabits New +Zealand. The Chilian genus _Calyptocephalus_ is allied to Australian +tropical genera. {42}The Neotropical genera of Ranidæ, five of which extend +to Chili and Patagonia, belong to a division which is Australian and +Neotropical, and which has species in the Oriental and Ethiopian regions. + +_Fresh-water Fishes._--These present some peculiar forms, and some very +interesting phenomena of distribution. The genus _Percilia_ has been found +only in the Rio de Maypu in Chili; and _Percichthys_, also belonging to the +perch family, has five species confined to the fresh waters of South +Temperate America, and one far away in Java. _Nematogenys_ (1 sp.) is +peculiar to Chili; _Trichomycterus_ reaches 15,000 feet elevation in the +Andes,--both belonging to the Siluridæ; _Chirodon_ (2 sp.), belonging to +the Characinidæ, is peculiar to Chili; and several other genera of the same +family extend into this sub-region from Brazil. The family _Haplochitonidæ_ +has a remarkable distribution; one of its genera, Haplochiton (2 sp.), +inhabiting Tierra del Fuego and the Falkland Islands, while the other, +_Prototroctes_, is found only in South Australia and New Zealand. Still +more remarkable is _Galaxias_ (forming the family Galaxidæ), the species of +which are divided between Temperate South America, and Australia, Tasmania, +and New Zealand; and there is even one species (_Galaxias attenuatus_) +which is found in the Chatham Islands, New Zealand, and Tasmania, as well +as in the Falkland Islands and Patagonia. _Fitzroya_ (1 sp.) is found only +at Montevideo; _Orestias_ (6 sp.) is peculiar to Lake Titicaca in the high +Andes of Bolivia; _Jenynsia_ (1 sp.) in the Rio de la Plata--all belonging +to the characteristic South American family of the Cyprinodontidæ. + +_Insects._--It is in insects more than in any other class of animals, that +we find clear indications of a not very remote migration of northern forms, +along the great mountain range to South Temperate America, where they have +established themselves as a prominent feature in the entomology of the +country. The several orders and families, however, differ greatly in this +respect; and there are some groups which are only represented by +modifications of tropical forms, as we have seen to be almost entirely the +case in birds and reptiles. + +{43}_Lepidoptera._--The butterflies of the South Temperate Sub-region are +not numerous, only about 29 genera and 80 species being recorded. Most of +these are from Chili, which is sufficiently accounted for by the general +absence of wood on the east side of the Andes from Buenos Ayres to South +Patagonia. The families represented are as follows: Satyridæ, with 11 +genera and 27 species, are the most abundant; Nymphalidæ, 2 genera and 8 +species; Lemoniidæ, 1 genus, 1 species; Lycænidæ, 3 genera, 8 species; +Pieridæ, 6 genera, 14 species; Papilionidæ, 2 genera, 8 species; Hesperidæ, +4 genera, 13 species. One genus of Satyridæ (_Elina_) and 2 of Pieridæ +(_Eroessa_ and _Phulia_) are peculiar to Chili. The following are the +genera whose derivation must be traced to the north temperate +zone:--_Tetraphlybia_, _Neosatyrus_, and 3 allied genera of 1 species each, +were formerly included under _Erebia_, a northern and arctic form, yet +having a few species in South Africa; _Argyrophorus_, allied to _Æneis_, a +northern genus; _Hipparchia_, a northern genus yet having a species in +Brazil;--all Satyridæ. The Nymphalidæ are represented by the typical north +temperate genus _Argynnis_, with 7 species in Chili; _Colias_, among the +Pieridæ, is usually considered to be a northern genus, but it possesses +representatives in South Africa, the Sandwich Islands, Malabar, New +Grenada, and Peru, as well as Chili, and must rather be classed as +cosmopolitan. These form a sufficiently remarkable group of northern forms, +but they are accompanied by others of a wholly Neotropical origin. Such are +_Stibomorpha_ with 6 species, ranging through South America to Guatemala, +and _Eteona_, common to Chili and Brazil (Satyridæ); _Apodemia_ (Lemoniidæ) +confined to Tropical America and Chili. _Hesperocharis_ and _Callidryas_ +(Pieridæ), both tropical; and _Thracides_ (Hesperidæ) confined to Tropical +America and Chili. Other genera are widely scattered; as, _Epinephile_ +found also in Mexico and Australia; _Cupido_, widely spread in the tropics; +_Euryades_, found only in La Plata and Paraguay, allied to South American +forms of _Papilio_, to the Australian _Eurycus_, and the northern +_Parnassius_; and _Heteropterus_, scattered in Chili, North America, and +Tropical Africa. We find then, among butterflies, a large north-temperate +element, {44}intermingled in nearly equal proportions with forms derived +from Tropical America; and the varying degrees of resemblances of the +Chilian to the northern species, seems to indicate successive immigrations +at remote intervals. + +_Coleoptera._--It is among the beetles of South Temperate America that we +find some of the most curious examples of remote affinities, and traces of +ancient migrations. The Carabidæ are very well represented, and having been +more extensively collected than most other families, offer us perhaps the +most complete materials. Including the Cicindelidæ, about 50 genera are +known from the South Temperate Sub-region, the greater part from Chili, but +a good number also from Patagonia and the Straits of Magellan. Of these +more than 30 are peculiar, and most of them are so isolated that it is +impossible to determine with precision their nearest allies. + +The only remarkable form of Cicindelidæ is _Agrius_, a genus allied to the +_Amblycheila_ and _Omus_ of N.W. America. Two genera of Carabidæ, +_Cascellius_ and _Baripus_, are closely allied to _Promecoderus_, an +Australian genus; and another, _Lecanomerus_, has one species in Chili and +the other in Australia. Five or six of the peculiar genera are undoubtedly +allied to characteristic Palæarctic forms; and such northern genera as +_Carabus_, _Pristonychus_, _Anchomenus_, _Pterostichus_, _Percus_, +_Bradycellus_, _Trechus_, and _Bembidium_, all absent from Tropical +America, give great support to the view that there is a close relation +between the insects of the northern regions and South Temperate America. A +decided tropical element is, however, present. _Tropopterus_ is near +_Colpodes_, a Tropical and South American genus; _Mimodromius_ and +_Plagiotelium_ are near _Calleida_, a South American genus; while +_Pachyteles_, _Pericompsus_, _Variopalpus_, and _Calleida_ are widely +spread American groups. The preponderance of northern forms seems, however, +to be undoubted. + +Six Carabidæ are known from Juan Fernandez, 3 being identical with Chilian +species and 3 peculiar. As the island is 350 miles from the mainland, we +have here a proof of how readily insects may be transported great +distances. + +{45}The Palæarctic affinity of the South Temperate Carabidæ may be readily +understood, if we bear in mind the great antiquity of the group, and the +known long persistence of generic and specific forms of Coleoptera; the +facility with which they may be transported to great distances by gales and +hurricanes, either on land or over the sea; and, therefore, the probability +that suitable stations would be rapidly occupied by species already adapted +to them, to the exclusion of those of the adjacent tracts which had been +specialised under different conditions. If, for example, we carry ourselves +back to the time when the Andes had only risen to half their present +altitude, and Patagonia had not emerged from the ocean (an epoch not very +remote geologically), we should find nearly all the Carabidæ of South +America, adapted to a warm, and probably forest-covered country. If, then, +a further considerable elevation of the land took place, a large temperate +and cold area would be formed, without any suitable insect inhabitants. +During the necessarily slow process of elevation, many of the tropical +Carabidæ would spread upwards, and some would become adapted to the new +conditions; while the majority would probably only maintain themselves by +continued fresh immigrations. But, as the mountains rose, another set of +organisms would make their way along the highest ridges. The abundance and +variety of the North Temperate Carabidæ, and their complete adaptation to a +life on barren plains and rock-strewn mountains, would enable them rapidly +to extend into any newly-raised land suitable to them; and thus the whole +range of the Rocky Mountains and Andes would obtain a population of +northern forms, which would overflow into Patagonia, and there, finding no +competitors, would develope into a variety of modified groups. This +migration was no doubt effected mainly, during successive glacial epochs, +when the mountain-range of the Isthmus of Panama, if moderately increased +in height, might become adapted for the passage of northern forms, while +storms would often carry insects from peak to peak over intervening forest +lowlands or narrow straits of sea. If this is the true explanation, we +ought to find no such preponderant northern element in groups which {46}are +proportionally less developed in cold and temperate climates. Our further +examination will show how far this is the case. + +Lucanidæ.--Only four genera are known in the sub-region. Two are peculiar, +_Chiasognathus_ and _Streptocerus_, the former allied to Tropical American, +the latter to Australian genera; the other two genera are exclusively South +American. + +Cetoniidæ.--These seem very scarce, only a few species of the Neotropical +genus _Gymnetis_ reaching Patagonia. + +Buprestidæ.--These are rather numerous, many very beautiful species being +found in Chili. Nineteen genera are represented in South Temperate America, +and 5 of these are peculiar to it; 3 others are South American genera; 2 +are Australian, and the remainder are wide-spread, but all are found also +in Tropical America. The only north-temperate genus is _Dicerca_, and even +this occurs also in the Antilles, Brazil, and Peru. Of the peculiar genera, +the largest, _Dactylozodes_ (26 sp.), has one species in South Brazil, and +is closely allied to _Hyperantha_, a genus of Tropical America; +_Epistomentis_ is allied to _Nascis_, an Australian genus; _Tyndaris_ is +close to _Acmoeodera_, a genus of wide range and preferring desert or dry +countries. The other two are single species of cosmopolitan affinities. On +the whole, therefore, the Buprestidæ are unmistakeably Neotropical in +character. + +Longicorns.--Almost the whole of the South Temperate Longicorns inhabit +Chili, which is very rich in this beautiful tribe. About 75 genera and 160 +species are known, and nearly half of the genera are peculiar. Many of the +species are large and handsome, rivalling in beauty those of the most +favoured tropical lands. Of the 8 genera of Prionidæ 6 are peculiar, but +all are allied to Tropical American forms except _Microplophorus_, which +belongs to a group of genera spread over Australia, Europe, and Mexico. The +Cerambycidæ are much more abundant, and their affinities more interesting. +Two (_Syllitus_ and _Pseudocephalus_) are common to Australia and Chili. +Twenty-three are Neotropical; and among these _Ibidion_, _Compsocerus_, +_Callideriphus_, _Trachyderes_, and _Xylocharis_, are best represented. +Twenty are {47}altogether peculiar, but most of them are more or less +closely allied to genera inhabiting Tropical America. Some, as the handsome +_Cheloderus_ and _Oxypeltus_, have no close allies in any part of the +world. _Holopterus_, though very peculiar, shows most resemblance to a New +Zealand insect. _Sibylla_, _Adalbus_, and _Phantagoderus_, have Australian +affinities; while _Calydon_ alone shows an affinity for north-temperate +forms. One species of the northern genus, _Leptura_, is said to have been +found at Buenos Ayres. + +The Lamiidæ are less abundant. Nine of the genera are Neotropical. Two +(_Apomecyna_ and _Exocentrus_) are spread over all tropical regions. Ten +genera are peculiar; and most of these are related to Neotropical groups or +are of doubtful affinities. Only one, _Aconopterus_, is decidedly allied to +a northern genus, _Pogonochærus_. It thus appears, that none of the Lamiidæ +exhibit Australian affinities, although these are a prominent feature in +the relations of the Cerambycidæ. + + + +It is evident, from the foregoing outline, that the insects of South +Temperate America, more than any other class of animals, exhibit a +connection with the north temperate regions, yet this connection is only +seen in certain groups. In Diurnal Lepidoptera and in Carabidæ, the +northern element is fully equal to the tropical, or even preponderates over +it. We have already suggested an explanation of this fact in the case of +the Carabidæ, and with the butterflies it is not more difficult. The great +mass of Neotropical butterflies are forest species, and have been developed +for countless ages in a forest-clad tropical country. The north temperate +butterflies, on the other hand, are very largely open-country species, +frequenting pastures, mountains, and open plains, and often wandering over +an extensive area. These would find, on the higher slopes of mountains, a +vegetation and conditions suited to them, and would occupy such stations in +less time than would be required to adapt and modify the forest-haunting +groups of the American lowlands. In those groups of insects, however, in +which the conditions of life are nearly the same as regards both temperate +and tropical species, the superior {48}number and variety of the tropical +forms has given them the advantage. Thus we find that among the Lucanidæ, +Buprestidæ, and Longicorns, the northern element is hardly perceptible. +Most of these are either purely Neotropical, or allied to Neotropical +genera, with the admixture, however, of a decided Australian element. As in +the case of the Amphibia and fresh-water fishes, the Australian affinity, +as shown by insects, is of two kinds, near and remote. We have a few genera +common to the two countries; but more commonly the genera are very +distinct, and the affinity is shown by the genera of both countries +belonging to a group peculiar to them, but which may be of very great age. +In the former case, we must impute some of the resemblance of the two +faunas to an actual interchange of forms within the epoch of existing +genera--a period of vast and unknown duration in the class of insects; +while in the latter case, and perhaps also in many of the former, it seems +more in accordance with the whole of the phenomena, to look upon most of +the instances as survivals, in the two southern temperate areas, of the +relics of groups which had once a much wider distribution. That this is the +true explanation, is suggested by the numerous cases of discontinuous and +scattered distribution we have had to notice, in which every part of the +globe, without exception, is implicated; and there is a reason why these +survivals should be rather more frequent in Australia and temperate South +America, inasmuch as these two areas agree in the absence of a considerable +number of otherwise cosmopolitan vertebrate types, and are also in many +respects very similar in climatic and other physical conditions. The +preponderating influence of the organic over the physical environment, as +taught by Mr. Darwin, leads us to give most weight to the first of the +above-mentioned causes; to which we may also impute such undoubted cases of +survival of ancient types as the Centetidæ of the Antilles and +Madagascar--both areas strikingly deficient in the higher vertebrate forms. +The probable mode and time of the cross migration between Australia and +South America, has been sufficiently discussed in our chapter on the +Australian region, when treating of the origin and affinities of the New +Zealand fauna. + + +{49}_Islands of the South Temperate Sub-region._ + +These are few, and of not much zoological interest. Tierra del Fuego, +although really an island, is divided from the mainland by so narrow a +channel that it may be considered as forming part of the continent. The +guanaco (_Auchenia huanaco_) ranges over it, and even to small islands +further south. + +_The Falkland Islands._--These are more important, being situated about 350 +miles to the east of Southern Patagonia; but the intervening sea is +shallow, the 100 fathom line of soundings passing outside the islands. We +have therefore reason to believe that they have been connected with South +America at a not distant epoch; and in agreement with this view we find +most of their productions identical, while the few that are peculiar are +closely allied to the forms of the mainland. + +The only indigenous Mammals are a wolf-like fox (_Pseudalopex antarcticus_) +said to be found nowhere else, but allied to two other species inhabiting +Southern Patagonia; and a species of mouse, probably one of the American +genera _Hesperomys_ or _Reithrodon_. + +Sixty-seven species of Birds have been obtained in these islands, but only +18 are land-birds; and even of these 7 are birds of prey, leaving only 11 +Passeres. The former are all common South American forms, but one species, +_Milvago australis_, seems peculiar. The 11 Passeres belong to 9 genera, +all found on the adjacent mainland. Three, or perhaps four, of the species +are however peculiar. These are _Phrygilus melanoderus_, _P. +xanthogrammus_, _Cinclodes antarcticus_, and _Muscisaxicola macloviana_. +The wading and swimming birds are of little interest, except the penguins, +which are greatly developed; no less than eight species being found, five +as residents and three as accidental visitors. + +No reptiles are known to inhabit these islands. + +_Juan Fernandez._--This island is situated in the Pacific Ocean, about 400 +miles west of Valparaiso in Chili. It is only a few miles in extent, yet it +possesses four land-birds, excluding the powerful Accipitres. These are +_Turdus falklandicus_; _Anæretes {50}fernandensis_, one of the Tyrannidæ; +and two humming-birds, _Eustephanus fernandensis_ and _E. galeritus_. The +first is a widespread South Temperate species, the two next are peculiar to +the island, while the last is a Chilian species which ranges south to +Tierra del Fuego. But ninety miles beyond this island lies another, called +"Mas-a-fuero," very much smaller; yet this, too, contains four species of +similar birds; one, _Oxyurus mas-a-fueræ_, allied to the wide-spread South +Temperate _O. spinicauda_, and _Cinclodes fusus_, a South Temperate +species--both Dendrocolaptidæ; with a humming-bird, _Eustephanus leyboldi_, +allied to the species in the larger island. The preceding facts are taken +from papers by Mr. Sclater in the _Ibis_ for 1871, and a later one in the +same journal by Mr. Salvin (1875). The former author has some interesting +remarks on the three species of humming-birds of the genus _Eustephanus_, +above referred to. The Chilian species, _E. galeritus_, is green in both +sexes. _E. fernandensis_ has the male of a fine red colour and the female +green, though differently marked from the female of _E. galeritus_. _E. +leyboldi_ (of Mas-a-fuera) has the male also red and the female green, but +the female is more like that of _E. galeritus_, than it is like the female +of its nearer ally in Juan Fernandez. Mr. Sclater supposes, that the +ancient parent form of these three birds had the sexes alike, as in the +present Chilian bird; that a pair (or a female having fertilised ova) +reached Juan Fernandez and colonised it. Under the action of sexual +selection (unchecked by some conditions which had impaired its efficacy on +the continent) the male gradually assumed a brilliant plumage, and the +female also slightly changed its markings. Before this change was completed +the bird had established an isolated colony on Mas-a-fuera; and here the +process of change was continued in the male, but from some unknown cause +checked in the female, which thus remains nearer the parent form. Lastly +the slightly modified Chilian bird again reached Juan Fernandez and exists +there side by side with its strangely altered cousin. + +All the phenomena can thus be accounted for by known laws, on the theory of +very rare accidental immigrations from the {51}mainland. The species are +here so very few, that the greatest advocate for continental extensions +would hardly call such vast causes into action, to account for the presence +of these three birds on so small and so remote an island, especially as the +union must have continued down to the time of existing species. But if +accidental immigration has sufficed here, it will also assuredly have +sufficed where the islands are larger, and the chances of reaching them +proportionately greater; and it is because an important principle is here +illustrated on so small a scale, and in so simple a manner as to be almost +undeniable, that we have devoted a paragraph to its elucidation. + +A few Coleoptera from Juan Fernandez present analogous phenomena. All +belong to Chilian genera, while a portion of them constitute peculiar +species. + +Land-shells are rather plentiful, there being about twenty species +belonging to seven genera, all found in the adjacent parts of South +America; but all the species are peculiar, as well as four others found on +the island of Mas-a-fuera. + + +_III. Tropical North America, or the Mexican Sub-region._ + +This sub-region is of comparatively small extent, consisting of the +irregular neck of land, about 1,800 miles long, which connects the North +and South American continents. Almost the whole of its area is mountainous, +being in fact a continuation of the great range of the Rocky Mountains. In +Mexico it forms an extensive table-land, from 6,000 to 9,000 feet above the +sea, with numerous volcanic peaks from 12,000 to 18,000 feet high; but in +Yucatan and Honduras, the country is less elevated, though still +mountainous. On the shores of the Caribbean Sea and Gulf of Mexico, there +is a margin of low land from 50 to 100 miles wide, beyond which the +mountains rise abruptly; but on the Pacific side this is almost entirely +wanting, the mountains rising almost immediately from the sea shore. With +the exception of the elevated plateaus of Mexico and Guatemala, and the +extremity of the peninsula of Yucatan, the whole of Central America is +clothed with forests; and as its surface is much broken up into hill and +valley, and the volcanic {52}soil of a large portion of it is very fertile, +it is altogether well adapted to support a varied fauna, as it does a most +luxuriant vegetation. Although many peculiar Neotropical types are absent, +it yet possesses an ample supply of generic and specific forms; and, as far +as concerns birds and insects, is not perhaps inferior to the richest +portions of South America in the number of species to be found in equal +areas. + +Owing to the fact that the former Republic of Mexico comprised much +territory that belongs to the Nearctic region, and that many Nearctic +groups extend along the high-lands to the capital city of Mexico itself, +and even considerably further south, there is much difficulty in +determining what animals really belong to this sub-region. On the +low-lands, tropical forms predominate as far as 28° N. latitude; while on +the cordilleras, temperate forms prevail down to 20°, and are found even +much farther within the tropics. + +_Mammalia._--Very few peculiar forms of Mammalia are restricted to tropical +North America; which is not to be wondered at when we consider the small +extent of the country, and the facility of communication with adjacent +sub-regions. A peculiar form of tapir (_Elasmognathus bairdi_) inhabits +Central America, from Panama to Guatemala, and, with _Myxomys_, a genus of +Muridæ, are all at present discovered. _Bassaris_, a remarkable form of +Procyonidæ, has been included in the Nearctic region, but it extends to the +high-lands of Guatemala. _Heteromys_, a peculiar genus of Saccomyidæ or +pouched rats, inhabits Mexico, Honduras, Costa Rica, and Trinidad. Five +genera of monkeys extend here,--_Ateles_, _Mycetes_, _Cebus_, +_Nyctipithecus_, and _Saimiris_; the two former alone reaching Mexico, the +last only going as far as Costa Rica. Other typical Neotropical forms are +_Galera_, the tayra, belonging to the weasel family; _Nasua_, the +coatimundi; _Dicotyles_, the peccary; _Cercolabes_, the tree porcupine; +_Dasyprocta_, the agouti; _Cælogenys_, the paca; _Choloepus_, and +_Arctopithecus_, sloths; _Cyclothurus_, an ant-eater; _Tatusia_, an +armadillo; and _Didelphys_, oppossum. Of Northern forms, _Sorex_, _Vulpes_, +_Lepus_, and _Pteromys_ reach Guatemala. + +_Birds._--The productiveness of this district in bird life, may {53}be +estimated from the fact, that Messrs. Salvin and Sclater have catalogued +more than 600 species from the comparatively small territory of Guatemala, +or the portion of Central America between Mexico and Honduras. The great +mass of the birds of this sub-region are of Neotropical families and +genera, but these are intermingled with a number of migrants from temperate +North America, which pass the winter here; with some northern forms on the +high-lands; and with a considerable number of peculiar genera, mostly of +Neotropical affinities. + +The genera of birds peculiar to this sub-region belong to the following +families:--Turdidæ (2 genera); Troglodytidæ (1 gen.); Vireonidæ (1 gen.); +Corvidæ (2 gen.); Ampelidæ (1 gen.); Tanagridæ (1 gen.); Fringillidæ (2 +gen.); Icteridæ (1 gen.); Formicariidæ (2 gen.); Tyrannidæ (2 gen.); +Cotingidæ (1 gen.); Momotidæ (1 gen.); Trogonidæ (1 gen.); Trochilidæ (14 +gen.); Conuridæ (1 gen.); Cracidæ (2 gen.); Strigidæ (1 gen.); in all 37 +genera of land-birds. The Neotropical families that do not extend into this +sub-region are, Pteroptochidæ; the sub-family _Furnariinæ_ of the +Dendrocolaptidæ; the sub-family _Conophaginæ_ of the Tyrannidæ; the +sub-family _Rupicolinæ_ of the Cotingidæ; Phytotomidæ; Todidæ; +Opisthocomidæ; Chionididæ; Thinocoridæ; Cariamidæ; Psophiidæ; Eurypygidæ; +Palamedeidæ; and Struthionidæ. On the other hand Paridæ, Certhiidæ, +Ampelidæ, and Phasianidæ, are northern families represented here, but which +do not reach South America; and there are also several northern genera and +species, of Turdidæ, Troglodytidæ, Mniotiltidæ, Vireonidæ, Fringillidæ, +Corvidæ, Tetraonidæ, and Strigidæ, which are similarly restricted. Some of +the most remarkable of the Neotropical genera only extend as far as Costa +Rica and Veragua,--countries which possess a rich and remarkable fauna. +Here only are found an umbrella bird, (_Cephalopterus glabricollis_); a +bell bird (_Chasmorhynchus tricarunculatus_); and species of _Dacnis_ +(Ceroebidæ), _Buthraupis_, _Eucometis_, _Tachyphonus_ (Tanagridæ), +_Xiphorhynchus_ (Dendrocolaptidæ); _Hypocnemis_ (Formicariidæ); +_Euscarthmus_ (Tyrannidæ); _Attila_ (Cotingidæ); _Piprites_ (Pipridæ); +_Capito_, _Tetragonops_ (Megalæmidæ); _Selenidera_ (Rhamphastidæ); +_Neomorphus_ {54}(Cuculidæ); _Monasa_ (Bucconidæ); many genera of +Trochilidæ; and _Nothocercus_ (Tinamidæ); none of which extend further +north. A considerable number of the peculiar genera noted above, are also +found in this restricted area, which is probably one of the richest +ornithological districts on the globe. + +_Reptiles._--These are much less known than the preceding classes, but they +afford several peculiar and interesting forms. Snakes are perhaps the least +remarkable; yet there are recorded 4 peculiar genera of Calamariidæ, 1 of +Colubridæ, 1 of Homalopsidæ, 3 of Dipsadidæ; while _Boa_ and _Elaps_ are in +common with South America. Lizards are much more specially developed. +_Chirotes_, one of the Amphisbænians, is confined to Mexico and the +southern part of the Nearctic region; _Heloderma_ forming a peculiar +family, Helodermidæ, is Mexican only; _Abronia_ and _Barissia_ (Zonuridæ) +are also Mexican, as is _Siderolampus_ belonging to the Scincidæ, while +_Blepharactitis_ (same family) inhabits Nicaragua; _Brachydactylus_, one of +the geckoes, is from Costa Rica; while _Phymatolepis_, _Lamanctus_, +_Corytheolus_, _Cachrix_, _Corythophanes_ and _Chamæleopsis_, all belonging +to the Iguanidæ, are confined to various parts of the sub-region. In the +same family we have also the Antillean, _Cyclura_, and the Nearctic +_Phrynosoma_ and _Tropidolepis_, as well as the wide-spread American genus +_Anolius_. + +Among the tortoises, _Staurotypus_, allied to _Chelydra_, is found in +Mexico and Guatemala; and another genus, _Claudius_, has been lately +described from Mexico. + +_Amphibia._--These are chiefly Batrachians; _Rhinophryna_ (forming a +peculiar family) being confined to Mexico; _Triprion_, a genus of Hylidæ, +inhabiting Yucatan, with _Leyla_ and _Strabomantis_ (Polypedatidæ) found +only in Costa Rica and Veragua, are peculiar genera. The Salamandridæ, so +abundant in the Nearctic region, are represented by a few species of +_Amblystoma_ and _Spelerpes_. + +_Fresh-water fish._--Since the British Museum catalogue was published, a +valuable paper by Dr. Günther, in the Transactions of the Zoological +Society for 1868, furnishes much additional information on the fishes of +Central America. In that part of the region south of Mexico, 106 species of +fresh-water fishes are {55}enumerated; and 17 of these are found in streams +flowing into both the Atlantic and Pacific Oceans. On the whole, 11 +families are represented among the fresh-water fish, and about 38 genera. +Of these, 14 are specially Nearctic,--_Amiurus_ (Siluridæ); _Fundulus_ +(Cyprinodontidæ); _Sclerognathus_ (Cyprinidæ); and _Lepidosteus_ +(Ganoidei). A much larger number are Neotropical; and several Neotropical +genera, as _Heros_ and _Poecilia_, are more largely developed here than in +any other part of the region. There are also a considerable number of +peculiar genera;--_Petenia_, _Theraps_, and _Neotrophus_ (Chromides); +_Ælurichthys_ (Siluridæ); _Chalcinopsis_ (Characniidæ); _Characodon_, +_Belonesox_, _Pseudoxiphophorus_, _Platypoecilus_, _Mollienesia_, and +_Xiphophorus_ (Cyprinodontidæ). A few peculiar Antillean forms are also +present; as _Agonostoma_ (Mugilidæ); _Gambusia_ and _Girardinuus_ +(Cyprinodontidæ). The other families represented are Percidæ (1 genus); +Pristopomatidæ (2 gen.); Gobiidæ (1 gen.); Clupeidæ (2 gen.); and Gymnotidæ +(1 genus). + +On the whole the fish-fauna is typically Neotropical, but with a small +infusion of Nearctic forms. There are a considerable proportion of peculiar +genera, and almost all the species are distinct from those of other +countries. The predominant family is that of the Cyprinodontidæ, +represented by 12 genera; and the genus _Heros_ (Chromidæ) has here its +maximum development, containing between thirty and forty species. Dr. +Günther considers that a number of sub-faunas can be distinguished, +corresponding to some extent, with the islands into which the country would +be divided by a subsidence of about 2,000 feet. The most important of these +divisions is that separating Honduras from Costa Rica, and as it also +divides a very marked ornithological fauna we have every reason to believe +that such a division must have existed during the latter portion of the +tertiary epoch. We shall find some farther evidence of this division in the +next class. + +_Insects._--The butterflies of various parts of Central America and Mexico, +having been largely collected, offer us some valuable evidence as to the +relations of this sub-region. Their general character is wholly +Neotropical, about one half of the {56}South American genera being found +here. There are also a few peculiar genera, as, _Drucina_ (Satyridæ); +_Microtia_ (Nymphalidæ); _Eumæus_ (Lycænidæ); and _Eucheira_ (Pieridæ). +_Clothilda_ (Nymphalidæ) is confined to this sub-region and the Antilles. +The majority of the genera range over the whole sub-region from Panama to +Mexico, but there are a considerable number, comprising many of the most +characteristic South American forms, which do not pass north of Costa Rica +or Nicaragua. Such are _Lycorea_, _Ituna_, _Thyridia_, _Callithomia_, +_Oleria_ and _Ceratina_,--all characteristic South American groups of +Danaidæ; _Pronophila_ and _Dynastor_ (Satyridæ); _Protogonius_, _Pycina_, +_Prepona_, _Nica_, _Ectima_ and _Colænis_ (Nymphalidæ); _Eurybia_ and +_Methonella_ (Nemeobiidæ); _Hades_, and _Panthemos_ (Erycinidæ). + +_Coleoptera._--These present some interesting features, but owing to their +vast number only a few of the more important families can be noticed. + +Cicindelidæ.--The only specially Neotropical genera recorded as occurring +in this sub-region, are _Ctenostoma_ and _Hiresia_, both reaching Mexico. + +Carabidæ.--Several genera are peculiar. _Molobrus_ is found in all parts of +the sub-region, while _Onychopterygia_, _Phymatocephalus_, and +_Anisotarsus_ are Mexican only. There are about 20 South American genera, +most of which extend to Mexico, and include such characteristic Neotropical +forms as _Agra_, _Callida_, _Coptodera_, _Pachyteles_, _Ardistomus_, +_Aspidoglossa_, _Stenocrepis_, and _Pelecium_. + +Lucanidæ.--Of this important family there is, strange to say, not a single +species recorded in Gemminger and Harold's catalogue up to 1868! It is +almost impossible that they can be really absent; yet their place seems to +he, to some extent, supplied by an unusual development of the allied +Passalidæ, of which there are five South American and six peculiar genera. + +Cetoniidæ.--All the larger South American genera extend to Mexico, which +country possesses 3 peculiar forms, _Ischnoscelis_, _Psilocnemis_, and +_Dialithus_; while _Trigonopeltastes_ is characteristic, having 4 Mexican, +1 Brazilian, and 1 North American species. + +{57}Buprestidæ.--In this family there are no peculiar genera. All the large +South American groups are absent, the only important and characteristic +genus being _Stenogaster_. + +Longicorns.--This important group is largely developed, the country being +well adapted to them; and their distribution presents some features of +interest. + +In the Prionidæ there are 6 peculiar genera, the largest being _Holonotus_ +with 3 species; two others, _Derotrachus_ and _Mallaspis_, are +characteristic; 3 more are common to South America, and 1 to Cuba. The +Cerambycidæ are much more numerous, and there are 24 peculiar genera, the +most important being _Sphenothecus_, _Entomosterna_, and _Cyphosterna_; +while _Crioprosopus_ and _Metaleptus_ are characteristic of the sub-region, +although extending into South America; about 12 Neotropical genera extend +to Mexico or Guatemala, while 12 more stop short, as far as yet known, at +Nicaragua. Lamiidæ have a very similar distribution; 13 genera are +peculiar, the most important being _Monilema_, _Hamatoderus_, and +_Carneades_, while _Phæa_ and _Lagochirus_ are characteristic. About +sixteen typical Neotropical genera extend to Mexico, and 15 more only reach +Nicaragua, among which are such important genera as _Anisopus_, +_Lepturgus_, and _Callia_. + +The land-shells are not sufficiently known to furnish any corresponding +results. They are however mostly of South American genera, and have +comparatively little affinity for those of the Antilles. + +_Relations of the Mexican sub-region to the North and South American +Continents._--The sudden appearance of numerous South American forms of +Edentata in temperate North America, in Post-Tertiary times, as narrated in +Chapter VII., together with such facts as the occurrence of a considerable +number of identical species of sea fish on the two sides of the Central +American isthmus, render it almost certain that the union of North and +South America is comparatively a recent occurrance, and that during the +Miocene and Pliocene periods, they were separated by a wide arm of the sea. +The low country of Nicaragua was probably the part submerged, leaving the +highlands of Mexico and Guatemala still united with the North {58}American +continent, and forming part of the Tertiary "Nearctic region." This is +clearly indicated both by the many Nearctic forms which do not pass south +of Nicaragua, of which the turkeys (_Meleagris_) are a striking example, +and by the comparative poverty of this area in typical Neotropical groups. +During the Miocene period there was not that marked diversity of climate +between North and South America that now prevails; for when a luxuriant +vegetation covered what are now the shores of the Arctic Ocean, the country +south of the great lakes must have been almost or quite tropical. At an +early Tertiary period, the zoological differences of the Nearctic and +Neotropical regions were probably more radical than they are now, South +America being a huge island, or group of islands--a kind of Australia of +the New World, chiefly inhabited by the imperfectly organized Edentata; +while North America abounded in Ungulata and Carnivora, and perhaps formed +a part of the great Old World continent. There were also one or more very +ancient unions (in Eocene or Miocene times) of the two continents, +admitting of the entrance of the ancestral types of Quadrumana into South +America, and, somewhat later, of the Camelidæ; while the isthmus south of +Nicaragua was at one time united to the southern continent, at another made +insular by subsidence near Panama, and thus obtained that rich variety of +Neotropical types that still characterises it. When the final union of the +two continents took place, the tropical climate of the lower portions of +Guatemala and Mexico would invite rapid immigration from the south; while +some northern forms would extend their range into and beyond the newly +elevated territory. The Mexican sub-region has therefore a composite +character, and we must not endeavour too rigidly to determine its northern +limits, nor claim as exclusively Neotropical, forms which are perhaps +comparatively recent immigrants; and it would perhaps be a more accurate +representation of the facts, if we were to consider all the highlands of +Mexico and Guatemala above the limits of the tropical forests, as still +belonging to the Nearctic region, of which the whole country so recently +formed a part. + +The long-continued separation of North and South America {59}by one or more +arms of the sea, as above indicated, is further rendered necessary by the +character of the molluscan fauna of the Pacific shores of tropical America, +which is much more closely allied to that of the Caribbean sea, and even of +West Africa, than to that of the Pacific islands. The families and many of +the genera are the same, and a certain proportion of very closely allied or +identical species, shows that the union of the two oceans continued into +late Tertiary times. When the evidence of both land and sea animals support +each other as they do here, the conclusions arrived at are almost as +certain as if we had (as we no doubt some day shall have) geological proof +of these successive subsidences. + +_Islands of the Mexican Sub-region._--The only islands of interest +belonging to this sub-region, are Tres Marias and Socorro, recently +investigated by Col. Grayson for some of the American Natural History +societies. + +Tres Marias consist of four small islands lying off the coast of +north-western Mexico, about 70 miles from San Blas. The largest is about 15 +miles long by 10 wide. They are of horizontally stratified deposits, of +moderate height and flat-topped, and everywhere covered with luxuriant +virgin forests. They appear to lie within the 100 fathom line of soundings. +Fifty-two species of birds, of which 45 were land-birds, were collected on +these islands. They consisted of 19 Passeres; 11 Picariæ (7 being +humming-birds); 10 Accipitres; 2 parrots, and 3 pigeons. All were Mexican +species except 4, which were new, and presumably peculiar to the islands, +and one tolerably marked variety. The new species belong to the following +genera;--_Parula_ and _Granatellus_ (Mniotiltidæ); _Icterus_ (Icteridæ); +and _Amazilia_ (Trochilidæ). A small _Psittacula_ differs somewhat from the +same species on the mainland. + +There are a few mammalia on the islands; a rabbit (_Lepus_) supposed to be +new; a very small opossum (_Didelphys_), and a racoon (_Procyon_). There +are also several tree-snakes, a _Boa_, and many lizards. The occurrence of +so many mammalia and snakes is a proof that these islands have been once +joined to the mainland; but the fact that some of the species of both birds +and {60}mammals are peculiar, indicates that the separation is not a very +recent one. At the same time, as all the species are very closely allied to +those of the opposite coasts when not identical, we may be sure that the +subsidence which isolated them is not geologically remote. + +Socorro, the largest of the Revillagigedo Islands, is altogether different +from the Tres Marias. It is situated a little further south (19 S. +Latitude), and about 300 miles from the coast, in deep water. It is about +2,000 feet high, very rugged and bare, and wholly volcanic. No mammalia +were observed, and no reptiles but a small lizard, a new species of a genus +(_Uta_) characteristic of the deserts of N.-Western Mexico. The only +observed land-shell (_Orthalicus undatus_) also inhabits N.-W. Mexico. Only +14 species of birds were obtained, of which 9 were land-birds; but of these +4 were new species, one a peculiar variety, and another (_Parula +insularis_) a species first found in the Tres Marias. With the exception of +this bird and a _Buteo_, all the land-birds belonged to different _genera_ +from any found on the Tres Marias, though all were Mexican forms. The +peculiar species belonged to the genera _Harporhynchus_ (Turdidæ); +_Troglodytes_ (Troglodytidæ); _Pipilo_ (Fringillidæ); _Zenaidura_ +(Columbidæ); and a variety of _Conurus holochrous_ (Psittacidæ). + +The absence of mammals and snakes, the large proportion of peculiar +species, the wholly volcanic nature of these islands, and their situation +in deep water 300 miles from land,--all indicate that they have not formed +part of the continent, but have been raised in the ocean; and the close +relation of their peculiar species to those living in N.-Western Mexico, +renders it probable that their antiquity is not geologically great. + +The Cocos Islands, about 300 miles S.-W. of the Isthmus of Panama, are +known to possess one peculiar bird, a cuckoo of the _Coccyzus_ type, which +is considered by some ornithologists to constitute a peculiar genus, +_Nesococcyx_. + + +_IV. The West Indian Islands, or Antillean Sub-region._ + +The West Indian islands are, in many respects, one of the most interesting +of zoological sub-regions. In position they {61}form an unbroken chain +uniting North and South America, in a line parallel to the great Central +American isthmus; yet instead of exhibiting an intermixture of the +productions of Florida and Venezuela, they differ widely from both these +countries, possessing in some groups a degree of speciality only to be +found elsewhere in islands far removed from any continent. They consist of +two very large islands, Cuba and Hayti;[3] two of moderate size, Jamaica +and Portorico; and a chain of much smaller islands, St. Croix, Anguilla, +Barbuda, Antigua, Guadeloupe, Dominica, Martinique, St. Lucia, St. Vincent, +Barbadoes, and Grenada, with a host of intervening islets. Tobago, +Trinidad, Margarita, and Curaçao, are situated in shallow water near the +coast of South America, of which they form part zoologically. To the north +of Cuba and Hayti are the Bahamas, an extensive group of coral reefs and +islands, 700 miles long, and although very poor in animal life, belonging +zoologically to the Antilles. All the larger islands, and most of the +smaller ones (except those of coral formation) are very mountainous and +rocky, the chains rising to about 8,000 feet in Hayti and Jamaica, and to +nearly the same height in Cuba. All, except where they have been cleared by +man, are covered with a luxuriant forest vegetation; the temperature is +high and uniform; the rains ample; the soil, derived from granitic and +limestone rocks, exceedingly fertile; and as the four larger islands +together are larger than Great Britain, we might expect an ample and +luxuriant fauna. The reverse is however the case; and there are probably no +land areas on the globe, so highly favoured by nature in all the essentials +for supporting animal life, and at the same time so poor in all the more +highly organised groups of animals. Before entering upon our sketch of the +main features of this peculiar but limited fauna, it will be well to note a +few peculiarities in the physical structure of the islands, which have an +important bearing on their past {62}history, and will enable us to account +for much that is peculiar in the general character of their natural +productions. + +If we draw a line immediately south of St. Croix and St. Bartholomew, we +shall divide the Archipelago into two very different groups. The southern +range of islands, or the Lesser Antilles, are, almost without exception, +volcanic; beginning with the small detached volcanoes of Saba and St. +Eustatius, and ending with the old volcano of Grenada. Barbuda and Antigua +are low islands of Tertiary or recent formation, connected with the +volcanic islands by a submerged bank at no great depth. The islands to the +north and west are none of them volcanic; many are very large, and these +have all a central nucleus of ancient or granitic rocks. We must also note, +that the channels between these islands are not of excessive depth, and +that their outlines, as well as the direction of their mountain ranges, +point to a former union. Thus, the northern range of Hayti is continued +westward in Cuba, and eastward in Portorico; while the south-western +peninsula extends in a direct line towards Jamaica, the depth between them +being 600 fathoms. Between Portorico and Hayti there is only 250 fathoms; +while close to the south of all these islands the sea is enormously deep, +from more than 1,000 fathoms south of Cuba and Jamaica, to 2,000 south of +Hayti, and 2,600 fathoms near the south-east extremity of Portorico. The +importance of the division here pointed out will be seen, when we state, +that indigenous mammalia of peculiar genera are found on the western group +of islands only; and it is on these that all the chief peculiarities of +Antillian zoology are developed. + +_Mammalia._--The mammals of the West Indian Islands are exceedingly few, +but very interesting. Almost all the orders most characteristic of South +America are absent. There are no monkeys, no carnivora, no edentata. +Besides bats, which are abundant, only two orders are represented; rodents, +by peculiar forms of a South American family; and insectivora (an order +entirely wanting in South America) by a genus belonging to a family largely +developed in Madagascar and found nowhere else. The early voyagers mention +"Coatis" and "Agoutis" as being {63}found in Hayti and the other large +islands, and it is not improbable that species allied to _Nasua_ and +_Dasyprocta_ did exist, and have been destroyed by the dogs of the +invaders; though, on the other hand, these names may have been applied to +the existing species, which do bear some general resemblance to these two +forms. + +The Chiroptera, or bats, are represented by a large number of species and +by several peculiar genera. The American family of Phyllostomidæ or +vampires, has six genera in the Antilles, of which three, _Lonchorina_, +_Brachyphylla_, and _Phyllonycteris_, are peculiar, the latter being found +only in Cuba. The Vespertilionidæ have four genera, of which one, +_Nycticellus_, is confined to Cuba. There are six genera of Noctilionidæ, +of which one, _Phyllodia_, is confined to Jamaica. + +The Insectivora are represented by the genus _Solenodon_, of which two +species are known, one inhabiting Cuba the other Hayti. These are small +animals about the size of a cat, with long shrew-like snout, bare rat-like +tail, and long claws. Their peculiar dentition and other points of their +anatomy shows that they belong to the family Centetidæ, of which five +different genera inhabit Madagascar; while there is nothing closely allied +to them in any other part of the world but in these two islands. + +Seals are said to be found on the shores of some of the islands, but they +are very imperfectly known. + +The rodents belong to the family Octodontidæ, or, according to some +authors, to the Echimyidæ, both characteristic South American groups. They +consist of two genera, _Capromys_, containing three or four species +inhabiting Cuba and Jamaica; while _Plagiodontia_ (very closely allied) is +confined to Hayti. A peculiar mouse, a species of the American genus +_Hesperomys_, is said to inhabit Hayti and Martinique, and probably other +islands. A _Dasyprocta_ or agouti, closely allied to, if not identical +with, a South American species, inhabits St. Vincent, St. Lucia, and +Grenada, and perhaps St. Thomas, and is the only mammal of any size +indigenous to the Lesser Antilles. All the islands in which sugar is +cultivated are, however, overrun with European rats and mice, and it is not +improbable that these may have {64}starved out and exterminated some of the +smaller native rodents. + +_Birds._--The birds of the Antilles, although very inferior in number and +variety to those of the mainland, are yet sufficiently abundant and +remarkable, to offer us good materials for elucidating the past history of +the country, when aided by such indications as geology and physical +geography can afford. + +The total number of land-birds which are permanent residents in the West +India islands is, as nearly as can be ascertained from existing materials, +203. There are, in addition to this number, according to Prof. Baird, 88 +migrants from North America, which either spend the winter in some of the +islands or pass on to Central or South America. These migrants belong to 55 +genera, and it is an interesting fact that so many as 40 of these genera +have no resident representatives in the islands. This is important, as +showing that this northern migration is probably a recent and superficial +phenomenon, and has not produced any (or a very slight) permanent effect on +the fauna. The migratory genera which have permanent residents, and almost +always representative species, in the islands, are in most cases +characteristic rather of the Neotropical than of the Nearctic fauna, as the +following list will show; _Turdus_, _Dendroeca_, _Vireo_, _Polioptila_, +_Agelæus_, _Icterus_, _Contopus_, _Myiarchus_, _Tyrannus_, _Antrostomus_, +_Chordeiles_, _Coccyzus_, _Columba_. By far the larger part of these birds +visit Cuba only; 81 species being recorded as occurring in that island, +while only 31 have been found in Jamacia, 12 in Porto Rico and St. Croix, +and 2 in Tobago and Trinidad. Setting aside these migratory birds, as +having no bearing on the origin of the true Antillean fauna, we will +discuss the residents somewhat in detail. + +The resident land-birds (203 in number) belong to 95 genera and 26 +families. Of these families 15 are cosmopolitan or nearly so--Turdidæ, +Sylviidæ, Corvidæ, Hirundinidæ, Fringillidæ, Picidæ, Cuculidæ, +Caprimulgidæ, Cypselidæ, Trogonidæ, Psittacidæ, Columbidæ, Tetraonidæ, +Falconidæ, and Strigidæ; 5 are American only--Vireonidæ, Mniotiltidæ, +Icteridæ, Tyrannidæ, Trochilidæ; 4 are Netropical only or almost +{65}exclusively--Coerebidæ, Tanagridæ, Cotingidæ, Conuridæ; 1 is Antillean +only--Todidæ; while 1--Ampelidæ--is confined (in the western hemisphere) to +North America, and almost to the Nearctic region. Of the 95 genera, no less +than 31, or almost exactly one-third, are peculiar; while of the 203 +resident species, 177 are peculiar, the other 26 being all inhabitants of +South or Central America. Considering how closely the islands approach the +continent in several places--Florida, Yucatan, and Venezuela--this amount +of speciality in such locomotive creatures as birds, is probably unexampled +in any other part of the globe. The most interesting of these peculiar +genera are the following: 4 of Turdidæ, or thrushes--1 confined to the +large islands, 1 to the whole archipelago, while 2 are limited to the +Lesser Antilles; 2 genera of Tanagridæ, confined to the larger islands; 2 +of Trogonidæ, also confined to the larger islands; 5 of hummingbirds, 3 +confined to the Greater, 1 to the Lesser Antilles; 2 of cuckoos, one +represented in all the large islands, the other in Jamaica only; 2 of owls, +one peculiar to Jamaica, the other represented in St. Croix, St. Thomas, +Portorico, and Cuba; and lastly, _Todus_, constituting a peculiar family, +and having representative species in each of the larger islands, is +especially interesting because it belongs to a group of families which are +wholly Neotropical--the Momotidæ, Galbulidæ, and Todidæ. The presence of +this peculiar form, with 2 trogons; 10 species of parrots, all but one +peculiar; 16 peculiar humming-birds belonging to 8 genera; a genus of +Cotingidæ; 10 peculiar tanagers belonging to 3 genera; 9 Coerebidæ of 3 +genera; together with species of such exclusively Netropical genera as +_Coereba_, _Certhiola_, _Sycalis_, _Phonipara_, _Elainea_, _Pitangus_, +_Campephilus_, _Chloronerpes_, _Nyctibius_, _Stenopsis_, _Lampornis_, +_Calypte_, _Ara_, _Chrysotis_, _Zenaida_, _Leptoptila_, and _Geotrygon_, +sufficiently demonstrate the predominant affinities of this fauna; although +there are many cases in which it is difficult to say, whether the ancestors +of the peculiar genera or species may not have been derived from the +Nearctic rather than from the Neotropical region. + +The several islands differ considerably in their apparent +{66}productiveness, but this is, no doubt, partly due to our knowledge of +Cuba and Jamaica being much more complete than of Hayti. The species of +resident land-birds at present known are as follows:-- + + Cuba 68 species of which 40 are peculiar to it. + Hayti 40 " " 17 " " + Jamaica 67 " " 41 " " + Portorico 40 " " 15 " " + Lesser Antilles 45 " " 24 " " + +If we count the peculiar genera of each island, and reckon as (½) when a +genus is common to two islands only, the numbers are as follows:--Cuba 7½, +Hayti 3½, Jamaica 8½, Portorico 1, Lesser Antilles 3½. These figures show +us, that although Jamaica is one of the smaller and the most isolated of +the four chief islands, it yet stands in the first rank, both for the +number of its species and of its peculiar forms of birds,--and although +this superiority may be in part due to its having been more investigated, +it is probably not wholly so, since Cuba has also been well explored. This +fact indicates, that the West Indian islands have undergone great changes, +and that they were not peopled by immigration from surrounding countries +while in the condition we now see them; for in that case the smaller and +more remote islands would be very much poorer, while Cuba, which is not +only the largest, but nearest to the mainland in two directions, would be +immensely richer, just as it really is in migratory birds. + +The number of birds common to the four larger islands is very +small--probably not more than half a dozen; between 20 and 30 are common to +some two of the islands (counting the Lesser Antilles as one island) and a +few to three; but the great mass of the species (at least 140) are confined +each to some one of the five islands or groups we have indicated. This is +an amount of isolation and speciality, probably not to be equalled +elsewhere, and which must have required a remarkable series of physical +changes to bring about. What those changes probably were, we shall be in a +better position to consider when we have completed our survey of the +various classes of land animals. + + + +Plate XVII. + +[Illustration] + +A SCENE IN CUBA, WITH CHARACTERISTIC ANIMALS. + +{67}In the preceding enumeration the Bahamas have been included with Cuba, +as regards the birds they have in common; but they possess some half dozen +species not found elsewhere, and even one central American genus of +humming-birds (_Doricha_) not found in any other part of the Antilles. We +have thus given Cuba rather more peculiar species than it really possesses, +so that the proportionate richness of Jamaica is rather greater than shown +by our figures. + +The destruction of the forests and the increase of population, with, +perhaps, the use of firearms, seem to have led to the extermination of some +species of birds in the smaller islands. Professor Newton has called +attention to the work of M. Ledru, who, in 1796, described the birds of St. +Thomas. He mentions a parrot and a parroquet in the island, the latter only +being now known, and very scarce; also a green pigeon and a tody, both now +unknown. No less than six species of parrots are said to have been formerly +found in Guadeloupe and Martinique, which are now extinct. + +_Plate XVII. Illustrating the peculiar Mammalia and Birds of the +Antilles._--The scene of this illustration is Cuba, the largest of the West +Indian islands, and one in which all its peculiar zoological features are +well developed. In the foreground is the agouta (_Solenodon cubanus_), a +remarkable insectivorous animal which, with another species inhabiting +Hayti, has no allies on the American continent; nor anywhere in the world +but in Madagascar, where a group of animals are found constituting the +family Centetidæ, to which _Solenodon_ is said undoubtedly to belong. Above +it are a pair of hutias (_Capromys fournieri_), rat-like animals belonging +to the South American family Octodontidæ. They live in the forests, and +climb trees readily, eating all kinds of vegetable food. Three species of +the genus are known, which are found only in Cuba and Jamaica. Just above +these animals is a white-breasted trogon (_Prionoteles temnurus_), confined +to Cuba, and the only species of the genus. Near the top of the picture are +a pair of todies (_Todus multicolor_), singular little insectivorous birds +allied to the motmots, but forming a very distinct family which is confined +to the islands of the {68}Greater Antilles. They are beautifully-coloured +birds,--green above, red and white beneath, and are exceedingly active in +their movements. To the right are a pair of small humming-birds +(_Sporadinus ricordi_), not very remarkable in this beautiful family, but +introduced here because they belong to a genus which is confined to the +Greater Antilles. + + +_Table of distribution of West-Indian Birds._--As the birds of the West +Indian islands are particularly interesting and their peculiarities +comparatively little known, we give here a table of the genera of +land-birds, compiled from all available sources of information. Owing to +the numerous independent observations on which it is founded, the +discrepancies of nomenclature, and uncertainty in some cases as to the +locality of species, it can only be looked upon as an approximative summary +of the existing materials on Antillean ornithology. + + +_TABLE OF THE RESIDENT LAND-BIRDS OF THE ANTILLES._ + +NOTE.--Genera confined to the West Indies are in Italics. An (_a_) after +(1) indicates a species common to two islands: but where there are two or +more species in an island, or the localities are doubtful, this indication +cannot be given. All species not otherwise noted are peculiar to the +Antilles. + + Key to columns: + Column 1 Cuba. + " 2 Bahamas. + " 3 Hayti. + " 4 Jamaica. + " 5 Portorico & St. Croix. + " 6 Lesser Antilles. + " 7 Total resident species. + + ----------------+-----------------------------+----+--------------------- + | Number of Species in | | + Family and | each Island. | | + Genus. +----+----+----+----+----+----+ | Remarks. + | 1 | 2 | 3 | 4 | 5 | 6 | 7 | + ----------------+----+----+----+----+----+----+----+--------------------- + TURDIDÆ. | | | | | | | | + Turdus | -- | -- | -- | 1 | -- | -- | 1 | Five species migrate + | | | | | | | | to Cuba + _Mimocichla_ | 2 | 1 | 1 | 1 | -- | -- | 5 | + _Margarops_ | -- | -- | 1a | -- | 1a | 3 | 4 | Martinique, St. + | | | | | | | | Lucia, Guada. + _Rhamphocinclus_ -- | -- | -- | -- | -- | 1 | 1 | Martinique and St. + | | | | | | | | Lucia + _Cinclocerthia_| -- | -- | -- | -- | -- | 3 | 3 | Nevis to St. Lucia + Mimus | 1 | 1 | -- | 1 | (?)| -- | 3 | Another species + | | | | | | | | migrates to the + | | | | | | | | Antilles + | | | | | | | | + SYLVIIDÆ. | | | | | | | | + Myiadestes | 1 | -- | -- | 1 | -- | 1 | 3 | St. Lucia + Polioptila | 1 | -- | -- | -- | -- | -- | 1 | + | | | | | | | | + VIREONIDÆ. | | | | | | | | + Vireosylvia | 1 | -- | 1 | 1 | 1 | 1 | 2 | One S. American + | | | | | | | | species + Vireo | 1 | 1 | -- | 1 | 1 | -- | 4 | Five species migrate + | | | | | | | | to Cuba + _Laletes_ | -- | -- | -- | 1 | -- | -- | 1 | + _Phoenicomanes_| -- | -- | -- | 1 | -- | -- | 1 | + | | | | | | | | + CORVIDÆ. | | | | | | | | + Corvus | 1 | -- | 1a | 1 | 1a | -- | 3 | + Cyanocorax | -- | -- | -- | 1 | -- | -- | 1 | S. American species + | | | | | | | | + MNIOTILTIDÆ. | | | | | | | | + Perissoglossa | 1 | -- | 1 | -- | 1 | -- | 1 | N. American species + Dendroeca | 2 | 2 | 1 | 3 | 1 | 1 | 7 | Twelve sp. migrate + | | | | | | | | to W. I. + _Teretristis_ | 2 | -- | -- | -- | -- | -- | 2 | + | | | | | | | | + COEREBIDÆ. | | | | | | | | + Certhiola | -- | 1 | 1 | 1 | 2 | 2 | 7 | Dominica and + | | | | | | | | Martinique + _Glossiptila_ | -- | -- | -- | 1 | -- | -- | 1 | + Coereba | 1 | -- | -- | -- | -- | -- | 1 | S. American species + | | | | | | | | + AMPELIDÆ. | | | | | | | | + _Dulus_ | (?)| -- | 1 | (?)| (?)| (?)| 2 | One species locality + | | | | | | | | unknown + | | | | | | | | + HIRUNDINIDÆ. | | | | | | | | + Progne | -- | -- | 1 | 1 | 1 | -- | 1 | + Pterochelidon | 1 | -- | -- | 1 | 1 | -- | 1 | + Hirundo | 1 | -- | 1a | 1a | -- | -- | 2 | One S. American + | | | | | | | | species + | | | | | | | | + TANAGRIDÆ. | | | | | | | | + Euphonia | 1a | -- | 1a | 1 | 1 | 1 | 4 | St. Bartholom. & + | | | | | | | | Martinique + _Spindalis_ | 2 | 1 | 1 | 1 | 1 | -- | 5 | + _Phænicophilus_| -- | -- | 1 | -- | -- | -- | 1 | + Saltator | -- | -- | -- | -- | -- | 1 | 1 | Guadeloupe and St. + | | | | | | | | Lucia + | | | | | | | | + FRINGILLIDÆ. | | | | | | | | + _Loxigilla_ | -- | -- | 1 | 1 | -- | 1 | 3 | Martinique and + | | | | | | | | Dominica + _Melopyrrha_ | 1 | -- | -- | -- | -- | -- | 1 | + Sycalis | -- | -- | -- | 1 | -- | -- | 1 | S. American species + Phonipara | 3 | -- | 3 | 3 | 2 | -- | 4 | One S. American + | | | | | | | | species + Chrysomitris | -- | -- | 1 | -- | -- | -- | 1 | + | | | | | | | | + ICTERIDÆ. | | | | | | | | + Icterus | 1 | -- | 1 | 1 | 2 | 2 | 6 | + Agelæus | 2 | -- | -- | -- | 1 | -- | 3 | + Sturnella | 1 | -- | -- | -- | -- | -- | 1 | Mexican species + _Nesopsar_ | -- | -- | -- | 1 | -- | -- | 1 | + Scolecophagus | 1 | -- | -- | -- | -- | -- | 1 | + Quiscalus | -- | -- | 1 | 1 | 2 | 2 | 4 | St. Lucia, + | | | | | | | | Martinique and + | | | | | | | | Barbadoes + | | | | | | | | + TYRANNIDÆ. | | | | | | | | + Elainea | -- | -- | -- | 2 | -- | 1 | 3 | + Pitangus | 1a | -- | -- | 1a | 1 | -- | 2 | + Contopus | -- | -- | -- | 1 | -- | 1 | 2 | St. Lucia + Myiarchus | 2 | -- | 1 | 3 | 1 | 1b | 7 | One S. American + | | | | | | | | species (b) + _Blacicus_ | 1a | -- | 1a | 1 | -- | -- | 2 | + Tyrannus | 2 | -- | -- | 1b | 1b | 2b | 3 | One sp. in Cen. + | | | | | | | | America (b) + | | | | | | | | + COTINGIDÆ. | | | | | | | | + Hadrostomus | -- | -- | -- | 1 | -- | -- | 1 | + | | | | | | | | + PICIDÆ. | | | | | | | | + Campephilus | 1 | -- | -- | -- | -- | -- | 1 | + _Xiphidiopicus_| 1 | -- | -- | -- | -- | -- | 1 | + Melanerpes | -- | -- | -- | -- | 1 | -- | 1 | + Chloronerpes | -- | -- | 1 | -- | -- | -- | 1 | + Centurus | 1 | -- | 1 | 1 | -- | -- | 3 | + Colaptes | 2 | -- | -- | -- | -- | -- | 2 | + _Nesoceleus_ | 1 | -- | -- | -- | -- | -- | 1 | + _Picumnus_ | -- | -- | ?1 | -- | -- | -- | 1 | + | | | | | | | | + CUCULIDÆ. | | | | | | | | + _Saurothera_ | 1 | -- | 1 | 1 | 1 | -- | 4 | + _Hyetornis_ | -- | -- | 1 | 1 | -- | -- | 2 | + Coccygus | 1 | -- | 2 | 1 | 1 | 1 | 3 | Dominica, St. Lucia, + | | | | | | | | all Neotropical + | | | | | | | | species + Crotophaga | 1 | -- | -- | 1 | 1 | 1 | 2 | N. & Cen. American + | | | | | | | | species + | | | | | | | | + TODIDÆ. | | | | | | | | + _Todus_ | 1 | -- | 1 | 2 | 1 | -- | 5 | + | | | | | | | | + TROGONIDÆ. | | | | | | | | + _Prionoteles_ | 1 | -- | -- | -- | -- | -- | 1 | + _Temnotrogon_ | -- | -- | 1 | -- | -- | -- | 1 | + | | | | | | | | + CAPRIMULGIDÆ. | | | | | | | | + Nyctibius | -- | -- | -- | 1 | -- | -- | 1 | Neotropical species + Chordeiles | -- | -- | -- | 1 | -- | -- | 1 | + Antrostomus | 2 | -- | -- | 1 | -- | 1 | 2 | One Neotropical + | | | | | | | | species + _Siphonorhis_ | -- | -- | -- | 1 | -- | -- | 1 | + Stenopsis | -- | -- | -- | -- | -- | 1 | 1 | Martinique + | | | | | | | | (S. America sp.) + | | | | | | | | + CYPSELIDÆ. | | | | | | | | + Cypselus | 1 | -- | -- | 1 | -- | -- | 1 | + Panyptila | -- | -- | 1 | -- | -- | -- | 1 | S. American species + Hemiprocne | -- | -- | 1 | 1 | -- | -- | 1 | Mexican species + Cypseloides | -- | -- | -- | 1 | -- | -- | 1 | + | | | | | | | | + TROCHILIDÆ. | | | | | | | | + Lampornis | -- | -- | 1a | 1 | 2a | 1a | 3 | + Doricha | -- | 2 | -- | -- | -- | -- | 2 | + _Eulampis_ | -- | -- | -- | -- | 1 | 2 | 2 | St. Croix, Dominica, + | | | | | | | | St. Lucia, + | | | | | | | | Martinique + _Aithurus_ | -- | -- | -- | 1 | -- | -- | 1 | + _Mellisuga_ | -- | -- | 1 | 1 | -- | -- | 1 | + Calypte | 1 | -- | -- | -- | -- | -- | 1 | + _Orthorhynchus_| -- | -- | -- | -- | 1 | 2 | 3 | Domin., Martini., + | | | | | | | | St. Lucia + _Sporadinus_ | 1 | -- | 1 | -- | 1 | -- | 3 | + | | | | | | | | + CONURIDÆ. | | | | | | | | + Ara | 1 | -- | -- | -- | -- | -- | 1 | S. American species + Conurus | 1 | -- | -- | 1 | 1 | 1 | 1 | St. Thomas + | | | | | | | | + PSITTACIDÆ. | | | | | | | | + Chrysotis | 1 | -- | 1 | 2 | 1 | 3 | 8 | + | | | | | | | | + COLUMBIDÆ. | | | | | | | | + Columba | 1 | -- | 1 | 2 | 2 | 1 | 3 | One in Honduras + Chamæpelia | -- | -- | -- | 1 | 1 | 1 | 1 | + Zenaida | 1 | -- | 1 | 1 | 1 | 2 | 2 | + Leptoptila | -- | -- | -- | 1 | -- | -- | 1 | + Geotrygon | 2 | -- | 1 | 2 | 1 | 2 | 5 | St. Lucia, + | | | | | | | | Martinique, one + | | | | | | | | species Mexican + _Starnoenas_ | 1 | -- | -- | -- | -- | -- | 1 | + | | | | | | | | + TETRAONIDÆ. | | | | | | | | + Ortyx | 1 | -- | -- | -- | -- | -- | 1 | + | | | | | | | | + FALCONIDÆ. | | | | | | | | + Accipiter | 2 | -- | -- | -- | -- | -- | 2 | + Hypotriorchis | -- | -- | -- | 1 | -- | -- | 1 | Mexican species + Cerchneis | 2 | -- | 1 | -- | -- | 1 | 2 | + Cymindis | 1 | -- | -- | -- | -- | -- | 1 | + Polyborus | 1 | -- | -- | -- | -- | -- | 1 | Mexican species + | | | | | | | | + STRIGIDÆ. | | | | | | | | + Nyctalops | 1 | -- | -- | -- | -- | -- | 1 | S. American species + _Pseudoscops_ | -- | -- | -- | 1 | -- | -- | 1 | + _Gymnoglaux_ | 1 | -- | -- | -- | 1 | -- | 2 | St. Croix and St. + | | | | | | | | Thomas + Glaucidium | 1 | -- | -- | -- | -- | -- | 1 | + ----------------+----+----+----+----+----+----+----+--------------------- + + {Number of families of resident land-birds in the Antilles 26 + TOTALS { " " genera " " " 95 + { " " species " " " 203 + +{72}_Reptiles and Amphibia._--These classes not having been systematically +collected, and the numerous described genera not having undergone careful +revision, little trustworthy information can be derived from them. The +following enumeration of the chief groups hitherto noticed or described, +will, however, show very similar features to those presented by the +birds--a general relation to Neotropical forms, a more special relation to +those of Central America and Mexico, and a considerable number of peculiar +types. + +Snakes.--_Arrhyton_ (Calamariidæ) from Cuba, _Hypsirhynchus_ from +Barbadoes, _Cryptodacus_ from Cuba, _Ialtris_ from Hayti, and _Coloragia_ +from Cuba (all Colubridæ), have been described as genera peculiar to the +Antilles. _Phylodryas_ and _Dromicus_ (Colubridæ) are Antillean and +Neotropical; _Ahætulla_, (Dendrophidæ) has the same distribution but +extends to tropical Africa; _Epicrates_ and _Corallus_ (Pythonidæ) are +Neotropical and Antillean; while _Chilabothrus_ from Jamaica and _Ungalia_ +from Cuba and Jamaica (both Pythonidæ) are found elsewhere only in Central +America and Mexico. There appear to be no Crotalidæ except an introduced +species of _Craspedocephalus_ in St. Lucia. + +Lizards are more numerous. _Ameiva_ (Teidæ) is found all over America, +_Gerrhonotus_ (Zonuridæ) is Neotropical and occurs in Cuba; _Gymnopthalmus_ +is South American and Antillean. Of Scincidæ seven genera are noted. +_Celestus_ (with 9 species) is peculiar to the Antilles; _Camilia_ (1 +species) to Jamaica, _Panoplus_ (1 species) and _Embryopus_ (1 species) to +Hayti; _Diplogossus_ is Antillean and South American; while _Plestiodon_ +and _Mabouya_ are cosmopolite. Of Geckotidæ there are four genera; +_Phyllodactylus_ and _Hemidactylus_ which are cosmopolite; _Sphærodactylus_ +which is wholly American; and _Cubina_ found only in Martinique and Brazil. +Of Iguanidæ there are six genera; _Anolis_, which ranges all over America; +_Polychrus_, which is Neotropical; _Iguana_ and _Liocephalus_ which are +South American; _Tropedurus_ found in Cuba and Brazil; and _Cyclura_ only +known from Jamaica, Cuba, and Central America. + +_Amphibia._--The genus _Trachycephalus_, belonging to the {73}Hylidæ or +tropical tree-frogs, is almost peculiar to the Antilles; Cuba, Hayti, and +Jamaica possessing seven species, while only one is recorded from South +America. Other genera are, _Peltaphryne_ (Bufonidæ) from Portorico; +_Phyllobates_ (Polypedatidæ) from Cuba; _Leiuperus_ (Ranidæ) from +Hayti,--all Neotropical. Of the Urodela, or tailed batrachians, no +representative occurs, although they are so characteristic a feature of the +Nearctic region. + +_Fresh-water fish._--The same general remarks apply to these as to the +reptiles. Only one peculiar genus is noted--_Lebistes_, a form of +Cyprinodontidæ from Barbadoes; other genera of the same family being, +_Haplochilus_, _Rivulus_, and _Girardinus_, widely spread in the +Neotropical region; while _Gambusia_ is confined to Central America, +Mexico, and the Antilles. Four other families are represented; Siluridæ by +_Chætostomus_, found in Portorico and South America; Chromidæ by the South +American _Acara_; Mugillidæ by the Central American _Agonostoma_; and +Percidæ by the North American _Centrarchus_, of which a species is recorded +from Cuba. + +_Insects._--The various West Indian islands have not been well explored +entomologically; one reason no doubt being, that their comparative poverty +renders them little attractive to the professional collector, while the +abounding riches of Central and South America lie so near at hand. We can, +therefore, hardly tell whether the comparative poverty, or even total +absence of some families while others seem fairly represented, is a real +phenomenon of distribution, or only dependent on imperfect knowledge. +Bearing this in mind, we proceed to give a sketch of what is known of the +chief groups of Lepidoptera and Coleoptera. + +_Lepidoptera._--The Neotropical butterfly-fauna is but poorly represented, +the majority of the most remarkable types being entirely wanting; yet there +are a few peculiar and very characteristic forms which show great +isolation, while the majority of the species are peculiar. Four genera are +exclusively or characteristically Antillean,--_Calisto_ belonging to the +Satyridæ, with four species, of which one ranges to South Carolina; +_Clothilda_ {74}(Nymphalidæ) a fine genus which has 4 Antillean species and +2 in Central America; _Lucinia_ (Nymphalidæ) 2 species, confined to Jamaica +and Hayti; and _Kricogonia_ belonging to the Pieridæ, which has 2 West +Indian species, while 1 inhabits Mexico and Florida. Genera which show a +special relation to Central America are _Euptoieta_, _Eumæus_, and +_Nathalis_. Almost all the other genera are South American, the total +number recorded in each family as occurring in the West Indian islands, +being, 3 of Danaidæ; 1 of Heliconiidæ; 2 of Satyridæ; 18 of Nymphalidæ; 1 +of Erycinidæ; 4 of Lycænidæ; 6 of Pieridæ; 1 of Papilionidæ, and 10 of +Hesperidæ. The genus _Papilio_ is represented by about 20 species, 2 of +which are North American, 4 South American, while the rest form little +characteristic groups allied to those of Central America. The most marked +feature seems to be the scarcity of Satyridæ and the almost total absence +of Erycinidæ, with a great deficiency in characteristic Neotropical forms +of Danaidæ and Nymphalidæ. + +_Coleoptera._--Cicindelidæ and Carabidæ are very poorly represented, by a +few species of wide-spread groups, and hardly any peculiar genera. No +Lucanidæ are recorded. Of Cetoniidæ, _Gymnetis_ only appears to be +represented. Buprestidæ seem to be more numerous; 15 genera being recorded, +but almost all of wide distribution. One only is +peculiar--_Tetragonoschoma_, found in Hayti; _Halecia_ is the only +exclusively South American genus; _Chalcophora_ is widely scattered over +the tropical regions but is absent from South America, yet it occurs in the +Nearctic region and extends to Jamaica and Guadeloupe. We now come to the +Longicorns, the only group of Coleoptera which seems to be well +represented, or which has been carefully collected. No less than 40 genera +are known from the West Indian islands, and 15 of these are peculiar. +Prionidæ are proportionately very numerous, there being 10 genera, 2 of +which are widely distributed in both South and North America, 1 is North +American, and 1 South American, while the following are +peculiar,--_Stenodontes_ (Hayti and Cuba); _Dendroblaptus_ (Cuba); +_Monodesmus_ (Cuba and Jamaica); _Prosternodes_ (Cuba); _Solenoptera_ and +_Elateropsis_, the two largest genera found in most of the {75}islands. Of +Cerambycidæ there are 16 genera, 2 of which range all over America, 4 are +Neotropical, 1 South American only, while the following are confined to the +islands,--_Merostenus_, _Pentomacrus_, and _Eburiola_ (Jamaica); +_Bromiades_ (Cuba); _Trichrous_, _Heterops_, and _Pæciloderma_ (Antilles). +One genus, _Smodicum_, is widely spread, having a species in Carolina, 1 in +South America, 1 in Hayti, and 1 in West Africa. Of Lamiidæ there are 14 +genera, 8 of which are Neotropical, 1 common to Central America and Mexico, +1 to the United States and Cuba, while 2, _Proecha_ and _Phidola_, are +confined to Cuba. Several of the genera are curiously +distributed;--_Spalacopsis_ is South American, with 4 species in Cuba and +Tropical Africa; _Lagocheirus_ is Neotropical, with a species in Australia; +while _Leptostilus_ is characteristic of the Antilles and North America, +with a few species in South America, and one in New Zealand. These cases of +erratic distribution, so opposed to the general series of phenomena among +which they occur, must be held to be sufficiently explained by the great +antiquity of these groups and their former wide distribution. They may be +supposed to be the remnants of types, now dying out, which were once, like +_Callichroma_, _Clytus_, and many others, almost universally distributed. + +All the peculiar Antillean genera of Cerambycidæ and Lamiidæ are allied to +Neotropical forms. The peculiar Prionidæ, however, are mostly allied to +Mexican and North American groups, and one, _Monodesmus_, belongs to a +group all the other genera of which inhabit the East Indies and South +Africa. + +_Land-shells._--This subject has already been generally treated under the +Region, of which, in this class of animals, the Antilles form so important +a part. We must therefore now confine ourselves mainly to the internal +distribution of the genera, and to a few remarks on the general bearing of +the facts. + +The excessive and altogether unexampled productiveness of the West Indian +islands in land-shells, may be traced to two main sets of causes. The first +and least known, consist of the peculiar influences and conditions which +render islands always more productive than continents. Whatever these +conditions {76}are, they will be more effective where the islands have been +long separated from the mainland, as is here undoubtedly the case. It seems +most probable that the great development of land-shells in islands, is due +to the absence or deficiency of the vertebrata, which on continents supply +a variety of species adapted to prey upon these molluscs. This view is +supported by the fact, that in such islands as have been united to a +continent at no very distant epoch, and still maintain a continental +variety of vertebrata, no such special development of land-shells has taken +place. If we compare the Philippine islands with the Sunda group, we find +the development of vertebrata and land-molluscs in inverse ratio to each +other. The same thing occurs if we compare New Zealand and Tasmania; and we +have a still more striking example in the Antillean group itself, +continental Trinidad having only 20 genera and 38 species, while the highly +insular Jamaica has about 30 genera and more than 500 species. + +The other causes favourable to the increase and development of land-shells +are of a physical nature. A great extent of limestone-rock is one; and in +the larger West Indian islands we have a considerable proportion of the +surface consisting of this rock. But perhaps equally or more important, is +the character of the land surface, and the texture of the exposed rock +itself. A much broken surface, with numerous deep ravines, cutting up the +whole country into isolated valleys and ridges, seems very favourable to +the specialization of forms in this very sedentary class of animals. +Equally favourable is a honeycombed and highly-fissured rock-surface, +affording everywhere cracks and crannies for concealment. Now, taking +Jamaica as an example of the archipelago, we find all these conditions in a +wonderful degree. Over a large part of this island, a yard of level ground +can hardly be found; but ridges, precipices, ravines, and rock-bound +valleys, succeed each other over the whole country. At least five-sixths of +the entire surface is limestone, and under the influence of tropical rains +this rock is worn, fissured, and honeycombed, so as to afford ample shelter +and concealment for land-shells. + +{77}It is probable that the three chief islands, Cuba, Jamaica and Hayti, +are nearly equally rich in land-shells; but the last is very much less +known, and therefore, perhaps, appears to be much poorer. Cuba has rather +more species than Jamaica; but while the former has only 1 peculiar genus +(_Diplopoma_), the latter has 3 (_Geomelania_, _Chittya_, and _Jamaicea_), +as well as two others only represented in the other islands by single +species. From Hayti, only about one-third as many species are known as from +the two former islands. It has no peculiar genera, but it has some forms in +common with Cuba and others with Jamaica, which show that those islands +have more connection with it, than with each other; just as we found to be +the case in birds. Portorico and the Virgin islands have still fewer +species than Hayti; and, as many of the genera common to the other three +islands are wanting, there is, no doubt, here a real deficiency. In the +islands farther south (Barbuda to Martinique) more Antillean genera +disappear or become very rare, while some continental forms take their +place. The islands from St. Lucia to Trinidad have a still more continental +character; the genus _Bulimus_, so largely developed on the continent, only +reaching St. Lucia. The Bahamas contain about 80 species of land-shells, of +which 25 are Antillean, the rest peculiar; all the genera being Antillean. +The affinity is chiefly with Hayti and Cuba, but closest with the latter +island. + +In the West Indian islands as a whole, there are 11 peculiar genera; 9 +operculate (_Geomelania_, _Chittya_, _Jamaicea_, _Licina_, _Choanopoma_, +_Ctenopoma_, _Diplopoma_, _Stoastoma_, _Lucidella_); and 2 inoperculate +(_Sagda_ and _Stenopus_), besides _Cyclostomus_, which belongs to the Old +World and is not found on the American continent. Mr. Bland considers, that +many of the Antillean land-shells exhibit decided African and Asiatic, +rather than South American affinities. A species of the Asiatic genus +_Diplommatina_ has been found in Trinidad, and an Indian species of _Ennea_ +occurs in Grenada and St. Thomas; a clear indication that land-shells are +liable to be accidentally imported, and to become established in the less +productive islands. + +Although these islands are so wonderfully rich even now, {78}there is good +reason to believe that many species have become extinct since the European +occupation of them. When small islands are much cultivated, many of these +molluscs which can only live under the shade of forests, are soon +extirpated. In St. Croix many species have become extinct at a +comparatively recent period, from the burning of forests; and as we know +that in all the islands many of the species are excessively local, being +often confined to single valleys or ridges, we may be sure that wherever +the native forests have disappeared before the hand of man, numbers of land +shells have disappeared with them. As some of the smaller islands have been +almost denuded of their wood, and in the larger ones extensive tracts have +been cleared for sugar cultivation, a very considerable number of species +have almost certainly been exterminated. + +_General Conclusions as to the Past History of the West Indian +Islands._--The preceding sketch of the peculiarities of the animal life of +these islands, enables us to state, that it represents the remains of an +ancient fauna of decided Neotropical type, having on the whole most +resemblance to that which now inhabits the Mexican sub-region. The number +of peculiar genera in all classes of animals is so great in proportion to +those in common with the adjacent mainland, as to lead us to conclude that, +subsequent to the original separation from the Mexican area, a very large +tract of land existed, calculated to support a rich and varied fauna, and, +by the interaction of competing types, give rise to peculiar and specially +modified organisms. We have already shown that the outline of the present +islands and the depths of the surrounding seas, give indications of the +position and extent of this ancient land; which not improbably occupied the +space enclosed by uniting Western Cuba with Yucatan, and Jamaica with the +Mosquito Coast. This land must have stretched eastward to include Anguilla, +and probably northward to include the whole of the Bahamas. At one time it +perhaps extended southward so as to unite Hayti with northern Venezuela, +while Panama and Costa Pica were sunk beneath the Pacific. At this time the +Lesser Antilles had no existence. + +The only large island of whose geology we have any detailed {79}account, is +Jamaica; and taking this as a type of what will probably be found in Cuba +and Hayti, we must place the continental period as having occurred after +the close of the Miocene, or during some part of the Pliocene epoch, since +a large portion of the surface of the former island consists of beds of +marine limestone from 2,000 to 3,000 thick, believed to be of Pliocene age. +After some time, the land between Hayti and South America subsided, and +still later that between Central America and Cuba with Jamaica; but a large +tract of land remained insulated, and no doubt supported a very much richer +and more varied fauna than now. We have evidence of this in extinct +Mammalia of large size, belonging to the peculiar South American family of +the chinchillas, which have been found in caves in the small islands of +Anguilla, and which, from the character of the land-shells associated with +them, are believed to be of Pliocene or Post-pliocene age. This discovery +is most interesting, and gives promise of very valuable results from the +exploration of the numerous caverns that undoubtedly exist in the abundant +limestone strata of the larger islands. This extensive Antillean land, +after long continuing undivided, was at length broken up by subsidence into +several islands; but as this alone would not account for the almost +complete annihilation of the mammalian fauna, it seems probable that the +subsidence was continued much farther, so as greatly to reduce the size and +increase the number of the islands. This is indicated, by the extensive +alluvial plains in Cuba and Hayti, and to a less extent in Jamaica; and by +elevated beds of Post-pliocene marls in the latter island. + +The series of changes now suggested, will account for all the main features +of the Antillean fauna in its relations to that of the American continent. +There remains the affinity with Madagascar, indicated by _Solenodon_, and a +few cases of African and Asiatic affinity in insects and land-shells; but +these are far too scanty to call for any attempt at special explanation. +Such cases of remote affinity and discontinuous distribution, occur in all +the regions, and in almost every group of animals; and we look upon them +almost all, as cases of survival, under favourable {80}conditions, of once +wide-spread groups. If no wild species of the genus _Equus_ were now to be +found, except in South Africa (where they are still most abundant), and in +South Temperate America, where their fossil remains show us they did exist +not very long ago, what a strong fact it would have appeared for the +advocates of continental extensions! Yet it would have been due to no +former union of the great southern continents, but to the former extensive +range of the family or the genus to which the two isolated remnants +belonged. And if such an explanation will apply to the higher vertebrata, +it is still more likely to be applicable to similar cases occurring among +insects or mollusca, the genera of which we have every reason to believe to +be usually much older than those of vertebrates. It is in these classes +that examples of widely scattered allied species most frequently occur; and +the facility with which they are diffused under favourable conditions, +renders any other explanation than that here given altogether superfluous. + +The _Solenodon_ is a member of an order of Mammalia of low type +(Insectivora) once very extensive and wide-spread, but which has begun to +die out, and which has left a number of curious and isolated forms thinly +scattered over three-fourths of the globe. The occurrence, therefore, of an +isolated remnant of this order in the Antilles is not in itself remarkable; +and the fact that the remainder of the family to which the Antillean +species belong has found a refuge in Madagascar, where it has developed +into several distinct types, does not afford the least shred of argument on +which to found a supposed independent land connection between these two +sets of islands. + + +_Summary of the Past History of the Neotropical Region._ + +We have already discussed this subject, both in our account of extinct +animals, and in various parts of the present chapter. It is therefore only +necessary here, briefly to review and summarise the conclusions we have +arrived at. + +The whole character of Neotropical zoology, whether as regards its +deficiencies or its specialities, points to a long continuance of isolation +from the rest of the world, with a few very distant {81}periods of union +with the northern continent. The latest important separation took place by +the submergence of parts of Nicaragua and Honduras, and this separation +probably continued throughout much of the Miocene and Pliocene periods; but +some time previous to the coming on of the glacial epoch, the union between +the two continents took place which has continued to our day. Earlier +submergences of the isthmus of Panama probably occurred, isolating Costa +Rica and Veragua, which then may have had a greater extension, and have +thus been able to develope their rich and peculiar fauna. + +The isthmus of Tehuantepec, at the south of Mexico, may, probably, also +have been submerged; thus isolating Guatemala and Yucatan, and leading to +the specialization of some of the peculiar forms that now characterise +those countries and Mexico. + +The West Indian Islands have been long isolated and have varied much in +extent. Originally, they probably formed part of Central America, and may +have been united with Yucatan and Honduras in one extensive tropical land. +But their separation from the continent took place at a remote period, and +they have since been broken up into numerous islands, which have probably +undergone much submergence in recent times. This has led to that poverty of +the higher forms of life, combined with the remarkable speciality, which +now characterises them; while their fauna still preserves a sufficient +resemblance to that of Central America to indicate its origin. + +The great continent of South America, as far as we can judge from the +remarkable characteristics of its fauna and the vast depths of the oceans +east and west of it, has not during Tertiary, and probably not even during +Secondary times, been united with any other continent, except through the +intervention of North America. During some part of the Secondary epoch it +probably received the ancestral forms of its Edentates and Rodents, at a +time when these were among the highest types of Mammalia on the globe. It +appears to have remained long isolated, and to have already greatly +developed these groups of animals, before it received, in early Tertiary +times, the ancestors of its marmosets and monkeys, and, perhaps also, some +of its peculiar forms of {82}Carnivora. Later, it received its Camelidæ, +peccaries, mastodons, and large Carnivora; and later still, just before the +Glacial epoch, its deer, tapir, opossums, antelopes, and horses, the two +latter having since become extinct. All this time its surface was +undergoing important physical changes. What its earlier condition was we +cannot conjecture, but there are clear indications that it has been broken +up into at least three large masses, and probably a number of smaller ones; +and these have no doubt undergone successive elevations and subsidences, so +as at one time to reduce their area and separate them still more widely +from each other, and at another period to unite them into continental +masses. The richness and varied development of the old fauna of South +America, as still existing, proves, however, that the country has always +maintained an extensive area; and there is reason to believe that the last +great change has been a long continued and steady increase of its surface, +resulting in the formation of the vast alluvial plains of the Amazon, +Orinoko, and La Plata, and thus greatly favouring the production of that +wealth of specific forms, which distinguishes South America above all other +parts of our globe. + +The southern temperate portion of the continent, has probably had a +considerable southward extension in late Tertiary times; and this, as well +as the comparatively recent elevation of the Andes, has given rise to some +degree of intermixture of two distinct faunas, with that proper to South +Temperate America itself. The most important of these, is the considerable +Australian element that appears in the insects, and even in the reptiles +and fresh-water fishes, of South Temperate America. These may be traced to +several causes. Icebergs and icefloes, and even solid fields of ice, may, +during the Glacial epoch, have afforded many opportunities for the passage +of the more cold-enduring groups; while the greater extension of southern +lands and islands during the warm periods--which there is reason to believe +prevailed in the southern as well as in the northern regions in Miocene +times--would afford facilities for the passage of the reptiles and insects +of more temperate zones. That no actual land-connection occurred, is proved +by the total absence {83}of interchange of the mammals or land-birds of the +two countries, no less than by the very fragmentary nature of the +resemblances that do exist. The northern element consists almost wholly of +insects; and is evidently due to the migration of arctic and north +temperate forms along the ridges and plateaus of the Andes; and most likely +occurred when these organisms were driven southward at successive cold or +Glacial periods. + +A curious parallel exists between the past history and actual zoological +condition of South America and Africa. In both we see a very ancient +land-area extending into the South Temperate zone, isolated at a very early +period, and developing only a low grade of Mammalian life; chiefly +Edentates and Rodents on the one, Lemurs and Insectivora in the other. +Later we find an irruption into both of higher forms, including Quadrumana, +which soon acquired a large and special development in the tropical +portions of each country. Still later we have an irruption into both of +northern forms, which spread widely over the two regions, and having become +extinct in the land from whence they came, have been long held to be the +original denizens of their adopted country. Such are the various forms of +antelopes, the giraffe, the elephant, rhinoceros, and lion in Africa; while +in America we have deer and peccaries, the tapir, opossums, and the puma. + +On the whole, we cannot but consider that the broad outlines of the +zoological history of the Neotropical region can be traced with some degree +of certainty; but, owing to the absence of information as to the most +important of the geological periods--the Miocene and Eocene--we have no +clue to the character of its early fauna, or to the land connections with +other countries, which may possibly have occurred in early Tertiary times. + + + + +{84}TABLES OF DISTRIBUTION. + + +In drawing up these tables, showing the distribution of the various classes +of animals in the Neotropical region, the following sources of information +have been relied on, in addition to the general treatises, monographs, and +catalogues used in the compilation of the Fourth Part of this work. + +_Mammalia._--D'Orbigny, and Burmeister, for Brazil and La Plata; Darwin, +and Cunningham, for Temperate S. America; Tschudi, for Peru; Frazer, for +Ecuador; Salvin, for Guatemala; Frantzius, for Costa Rica; Sclater, for +Quadrumana N. of Panama; Gundlach, for Cuba; and papers by Dr. J. E. Gray, +and Mr. Tomes. + +_Birds._--Sclater and Salvin's Nomenclator; Notes by Darwin, and +Cunningham; Gundlach, March, Bryant, Baird, Elliot, Newton, Semper, and +Sundevall, for various islands of the Antilles; and papers by Hudson, +Lawrence, Grayson, Abbott, Sclater, and Salvin. + + +{85}TABLE I. + +_FAMILIES OF ANIMALS INHABITING THE NEOTROPICAL REGION._ + +EXPLANATION. + + Names in _italics_ show the families which are peculiar to the region. + + Names enclosed thus (......) indicate families which barely enter the + region, and are not considered properly to belong to it. + + Numbers correspond with those of the series of families in Part IV. + + ---------------------+-------------------+------------------------------- + | Sub-regions | + | 1=Chili. | + Order and Family | 2=Brazil. | Range beyond the Region. + | 3=Mexico. | + | 4=Antilles. | + ---------------------+----+----+----+----+------------------------------- + | 1. | 2. | 3. | 4. | + ---------------------+----+----+----+----+------------------------------- + | | | | | + MAMMALIA. | | | | | + PRIMATES. | | | | | + 4. _Cebidæ_ | | -- | -- | | + 5. _Hapalidæ_ | | -- | (?)| | + | | | | | + CHIROPTERA. | | | | | + 10. _Phyllostomidæ_ | -- | -- | -- | -- | California + 12. Vespertilionidæ | -- | -- | -- | -- | Cosmopolite + 13. Noctilionidæ | -- | -- | -- | -- | All tropical regions + | | | | | + INSECTIVORA. | | | | | + 18. Centetidæ | | | | -- | Madagascar + | | | | | + CARNIVORA. | | | | | + 23. Felidæ | -- | -- | -- | | All regions but Australian + 28. Canidæ | -- | -- | -- | | All regions but Australian + 29. Mustelidæ | -- | -- | -- | | All regions but Australian + 30. Procyonidæ | -- | -- | -- | | N. America + 32. Ursidæ | -- | | | | All regions but Ethiopian and + | | | | | Australian + 33. Otariidæ | -- | | | | S. temperate zone + 35. Phocidæ | -- | | | (?)| N. and S. temperate zones + | | | | | + CETACEA. | | | | | + 36 to 41 | -- | | | | Oceanic + | | | | | + SIRENIA. | | | | | + 42. Manatidæ | | -- | -- | -- | Tropical shores + | | | | | + UNGULATA. | | | | | + 44. Tapiridæ | | -- | -- | | Indo-Malaya + 47. Suidæ | | -- | -- | | Cosmopolite, excl. Australia + 48. Camelidæ | -- | | | | Palæarctic + 50. Cervidæ | -- | -- | -- | | All regions but Ethiopian and + | | | | | Australian + | | | | | + RODENTIA. | | | | | + 55. Muridæ | -- | -- | -- | -- | Cosmopolite + 59. Saccomyidæ | | | -- | | Nearctic + 61. Sciuridæ | | -- | -- | | All regions but Australian + 63. _Chinchillidæ_ | -- | | | | + 64. Octodontidæ | -- | -- | | -- | Africa + 65. Echimyidæ | -- | -- | | | Ethiopian + 66. Cercolabidæ | | -- | -- | | Nearctic + 68. _Caviidæ_ | -- | -- | -- | -- | + 70. Leporidæ | | -- | -- | | All regions but Australian + | | | | | + EDENTATA. | | | | | + 71. _Bradypodidæ_ | | -- | -- | | + 73. _Dasypodidæ_ | -- | -- | -- | | + 75. _Myrmecophagidæ_| | -- | -- | | + | | | | | + MARSUPIALIA. | | | | | + 76. Didelphyidæ | -- | -- | -- | | Temperate N. America + | | | | | + BIRDS. | | | | | + PASSERES. | | | | | + 1. Turdidæ | -- | -- | -- | -- | Almost cosmopolite + 2. Sylviidæ | | -- | -- | -- | Almost cosmopolite + 5. Cinclidæ | | -- | -- | | Nearctic, Palæarctic, Oriental + 6. Troglodytidæ | -- | -- | -- | -- | Nearctic, Palæarctic, Oriental + 8. Certhiidæ | | | -- | | Nearctic, Palæarctic, Oriental + 9. Sittidæ | | | -- | | All regions, excl. Africa + 10. Paridæ | | | -- | | Nearctic, Palæarctic, Oriental + 20. Corvidæ | -- | -- | -- | -- | Cosmopolite + 26. _Coerebidæ_ | | -- | -- | -- | + 27. Mniotiltidæ | | -- | -- | -- | Nearctic + 28. Vireonidæ | | -- | -- | -- | Nearctic + 29. Ampelidæ | | | -- | -- | Nearctic, Palæarctic + 30. Hirundinidæ | -- | -- | -- | -- | Cosmopolite + 31. Icteridæ | -- | -- | -- | -- | Nearctic + 32. Tanagridæ | -- | -- | -- | -- | Nearctic + 33. Fringillidæ | -- | -- | -- | -- | All regions but Australian + 38. Motacillidæ | -- | -- | -- | -- | Cosmopolite + 38a. _Oxyrhamphidæ_ | | -- | -- | | + 39. Tyrannidæ | -- | -- | -- |-- | Nearctic + 40. _Pipridæ_ | | -- | -- | | + 41. _Cotingidæ_ | -- | -- | -- | -- | + 42. _Phytotomidæ_ | -- | | | | + 44._Dendrocolaptidæ_| -- | -- | -- | | + 45. _Formicariidæ_ | | -- | -- | | + 46. _Pteroptochidæ_ | -- | -- | | | + | | | | | + PICARIÆ. | | | | | + 51. Picidæ | -- | -- | -- | -- | All regions but Australian + 54. Megalæmidæ | | -- | -- | | Ethiopian, Oriental + 55. _Rhamphastidæ_ | | -- | -- | | + 58. Cuculidæ | -- | -- | -- | -- | Cosmopolite + 60. _Bucconidæ_ | | -- | -- | | + 61. _Galbulidæ_ | | -- | -- | | + 64. _Todidæ_ | | | | -- | + 65. _Momotidæ_ | | -- | -- | | + 66. Trogonidæ | | -- | -- | -- | Ethiopian, Oriental + 67. Alcedinidæ | -- | -- | -- | -- | Cosmopolite + 72. _Steatornithidæ_| | -- | | | + 73. Caprimulgidæ | -- | -- | -- | -- | Cosmopolite + 74. Cypselidæ | -- | -- | -- | -- | Almost cosmopolite + 75. Trochilidæ | -- | -- | -- | -- | Nearctic + | | | | | + PSITTACI. | | | | | + 80. Conuridæ | -- | -- | -- | -- | S. United States + 81. Psittacidæ | | -- | -- | -- | Ethiopian + | | | | | + COLUMBÆ. | | | | | + 84. Columbidæ | -- | -- | -- | -- | Cosmopolite + | | | | | + GALLINÆ. | | | | | + 87. Tetraonidæ | | -- | -- | -- | Almost cosmopolite + 88. Phasianidæ | | | -- | | All regions but Australian + 91. _Cracidæ_ | | -- | -- | | + 92. _Tinamidæ_ | -- | -- | -- | | + | | | | | + OPISTHOCOMI. | | | | | + 93. _Opisthocomidæ_ | | -- | | | + | | | | | + ACCIPITRES. | | | | | + 94. Vulturidæ | -- | -- | -- | -- | All regions but Australian + 96. Falconidæ | -- | -- | -- | -- | Cosmopolite + 97. Pandionidæ | | -- | -- | -- | Cosmopolite + 98. Strigidæ | -- | -- | -- | -- | Cosmopolite + | | | | | + GRALLÆ. | | | | | + 99. Rallidæ | -- | -- | -- | -- | Cosmopolite + 100. Scolopacidæ | -- | -- | -- | -- | Cosmopolite + 101. _Chionididæ_ | -- | | | | + 102. _Thinocoridæ_ | -- | | | | + 103. Parridæ | | -- | -- | | Tropical regions + 105. Charadriidæ | -- | -- | -- | -- | Cosmopolite + 108. _Cariamidæ_ | -- | -- | | | + 109. _Aramidæ_ | | -- | -- | | + 110. _Psophiidæ_ | | -- | | | + 111. _Eurypygidæ_ | | -- | -- | | + 113. Ardeidæ | -- | -- | -- | -- | Cosmopolite + 114. Plataleidæ | -- | -- | -- | -- | Almost cosmopolite + 115. Ciconiidæ | -- | -- | -- | | Nearly cosmopolite + 116. _Palamedeidæ_ | -- | -- | | | + 117. Phoenicopteridæ | -- | -- | -- | | Ethiopian, Indian + | | | | | + ANSERES. | | | | | + 118. Anatidæ | -- | -- | -- | -- | Cosmopolite + 119. Laridæ | -- | -- | -- | -- | Cosmopolite + 120. Procellariidæ | -- | -- | -- | -- | Cosmopolite + 121. Pelecanidæ | -- | -- | -- | -- | Cosmopolite + 122. Spheniscidæ | -- | | | | S. temperate zone + 124. Podicipidæ | -- | -- | -- | -- | Cosmopolite + | | | | | + STRUTHIONES. | | | | | + 126. Struthionidæ | -- | | | | Ethiopian + | | | | | + REPTILIA. | | | | | + OPHIDIA. | | | | | + 1. Typhlopidæ | | -- | -- | -- | Tropical regions and + | | | | | S. Palæarctic + 2. Tortricidæ | | -- | | | Oriental, N.-W. America + 5. Calamariidæ | -- | -- | -- | -- | All warm countries + 6. Oligodontidæ | | -- | | | Oriental, Japan + 7. Colubridæ | -- | -- | -- | -- | Almost cosmopolite + 8. Homalopsidæ | -- | | | -- | All the regions + 11. Dendrophidæ | -- | -- | -- | -- | All tropical regions + 12. Dryiophidæ | | -- | -- | | Oriental, Ethiopian + 13. Dipsadidæ | | -- | -- | | All tropical regions + 14. Scytalidæ | | -- | -- | | Philippine Islands + 16. Amblycephalidæ | | -- | -- | | Oriental + 17. Pythonidæ | -- | -- | -- | -- | All tropical regions, + | | | | | California + 20. Elapidæ | -- | -- | -- | | Tropical regions, Japan, + | | | | | S. Carolina + 23. Hydrophidæ | | | -- | | Oriental, Australian, + | | | | | Madagascar + 24. Crotalidæ | -- | -- | -- | -- | Nearctic, Palæarctic, Oriental + | | | | | + LACERTILIA. | | | | | + 27. Chirotidæ | | | -- | | Missouri + 28. Amphisbænidæ | -- | -- | | -- | Ethiopian, S. Palæarctic + 29. Lepidosternidæ | -- | -- | | | Ethiopian + 31. _Helodermidæ_ | | | -- | | + 32. Teidæ | -- | -- | -- | -- | Nearctic + 34. Zonuridæ | | -- | -- | -- | Nearctic, Ethiopian, + | | | | | S. Europe, and N. India + 35. _Chalcidæ_ | -- | -- | -- | | Nearctic + 36. _Anadiadæ_ | | -- | | | + 37. _Chirocolidæ_ | | -- | | | + 38. _Iphisadæ_ | | -- | | | + 39. _Cercosauridæ_ | | -- | | | + 41. Gymnopthalmidæ | | -- | | -- | Australian, Ethiopian, + | | | | | Palæarctic + 45. Scincidæ | -- | -- | -- | -- | Almost cosmopolite + 49. Geckotidæ | -- | -- | -- | -- | Almost cosmopolite + 50. Iguanidæ | -- | -- | -- | -- | Nearctic + | | | | | + CROCODILIA. | | | | | + 55. Crocodilidæ | | -- | -- | -- | Ethiopian, Oriental, + | | | | | N. Australian + 56. Alligatoridæ | | -- | -- | | Nearctic + | | | | | + CHELONIA. | | | | | + 57. Testudinidæ | -- | -- | -- | -- | All continents but Australian + 58. Chelydidæ | | -- | | | Ethiopian, Australian + 60. Cheloniidæ | | | | | Marine + | | | | | + AMPHIBIA. | | | | | + PSEUDOPHIDIA. | | | | | + 1. Ceciliadæ | | -- | -- | | Oriental, Ethiopian + | | | | | + URODELA. | | | | | + 6. (Salamandridæ) | | -- | -- | | Nearctic, Palæarctic + | | | | | + ANOURA. | | | | | + 7. _Rhinophrynidæ_ | | | -- | | + 8. Phryniscidæ | -- | -- | -- | | Ethiopian, Australian, Java + 9. _Hylaplesidæ_ | -- | -- | | -- | + 10. Bufonidæ | -- | -- | -- | -- | All continents but Australia + 12. Engystomidæ | -- | -- | -- | | All regions but Palæarctic + 13. Bombinatoridæ | -- | -- | | | Palæarctic, New Zealand + 14. _Plectromantidæ_| -- | | | | + 15. Alytidæ | | -- | | | All regions but Oriental + 16. Pelodryadæ | -- | -- | | | Australia + 17. Hylidæ | -- | -- | -- | -- | All regions but Ethiopian + 18. Polypedatidæ | -- | -- | -- | -- | All the regions + 19. Ranidæ | -- | -- | -- | -- | Almost cosmopolite + 20. Discoglossidæ | -- | -- | | | All regions but Nearctic + 21. _Pipidæ_ | | -- | | | + | | | | | + FISHES. | | | | | + (FRESHWATER). | | | | | + ACANTHOPTERYGII. | | | | | + 3. Percidæ | -- | -- | | -- | All regions but Australian + 11. (Trachinidæ) | -- | | | | Australia + 12. Scienidæ | (?)| -- | -- | (?)| All regions but Australian + 33. Nandidæ | | -- | | | Oriental + 34. _Polycentridæ_ | | -- | -- | | + 38. Mugillidæ | | (?)| -- | -- | Australian, Ethiopian + 52. Chromidæ | -- | -- | -- | -- | Ethiopian, Oriental + | | | | | + PHYSOSTOMI. | | | | | + 59. Siluridæ | -- | -- | -- | -- | All warm regions + 60. Characinidæ | -- | -- | -- | -- | Ethiopian + 61. Haplochitonidæ | -- | | | | S. Australia + 67. Galaxidæ | -- | | | | Tasmania and New Zealand + 73. Cyprinodontidæ | -- | -- | -- | -- | Absent from Australia + 78. Osteoglossidæ | | -- | | | All tropical regions + 84. _Gymnotidæ_ | | -- | | | + 85. Symbranchidæ | | -- | | | Oriental, Australian, + | | | | | (? marine) + | | | | | + DIPNOI. | | | | | + 92. Sirenoidei | | -- | | | Ethiopian, Australian + | | | | | + PLAGIOSTOMATA. | | | | | + 112. _Trygonidæ_ | | -- | | | + | | | | | + INSECTS. | | | | | + LEPIDOPTERA (PART). | | | | | + DIURNI (BUTTERFLIES).| | | | | + 1. Danaidæ | -- | -- | -- | -- | All warm regions, and to + | | | | | Canada + 2. Satyridæ | -- | -- | -- | -- | Cosmopolite + 4. Morphidæ | | -- | -- | | Australian, Oriental + 5. _Brassolidæ_ | | -- | -- | | + 6. Acræidæ | | -- | -- | | All tropical regions + 7. _Heliconiidæ_ | | -- | -- | -- | + 8. Nymphalidæ | -- | -- | -- | -- | Cosmopolite + 9. Libytheidæ | | -- | | -- | Absent from Australia + 10. Nemeobiidæ | | -- | -- | | Not in Australia or Nearctic + | | | | | regions + 11. _Eurygonidæ_ | | -- | -- | | + 12. Erycinidæ | | -- | -- | -- | Nearctic + 13. Lycænidæ | -- | -- | -- | -- | Cosmopolite + 14. Pieridæ | -- | -- | -- | -- | Cosmopolite + 15. Papilionidæ | -- | -- | -- | -- | Cosmopolite + 16. Hesperidæ | -- | -- | -- | -- | Cosmopolite + | | | | | + SPHINGIDEA. | | | | | + 17. Zygænidæ | -- | -- | -- | -- | Cosmopolite + 18. Castniidæ | | -- | -- | -- | Australian + 20. Uraniidæ | | -- | -- | -- | All tropical regions + 21. Stygiidæ | | -- | | | Palæarctic + 22. Ægeriidæ | -- | -- | -- | -- | Not in Australia + 23. Sphingidæ | -- | -- | -- | -- | Cosmopolite + ---------------------+----+----+----+----+------------------------------- + +{91}TABLE II. + +_GENERA OF TERRESTRIAL MAMMALIA AND BIRDS INHABITING THE NEOTROPICAL +REGION._ + +EXPLANATION. + + Names in _italics_ show the genera peculiar to the region. + + Names enclosed thus (......) indicate genera which barely enter the + region, and are not considered properly to belong to it. + + Genera undoubtedly belonging to the region are numbered consecutively. + + + _MAMMALIA._ + + -------------------+-------+----------------------+---------------------- + Order, Family, and | No. of| Range within | Range beyond + Genus. |Species| the Region. | the Region. + -------------------+-------+----------------------+---------------------- + | | | + PRIMATES. | | | + CEBIDÆ. | | | + 1. _Cebus_ | 18 | Costa Rica to | + | | Paraguay | + 2. _Lagothrix_ | 5 | Upper Amazon and | + | | E. Andes | + 3. _Eriodes_ | 3 | East Brazil, S. of | + | | Equator | + 4. _Ateles_ | 14 | Almost all tropical | + | | America | + 5. _Mycetes_ | 10 | E. Guatemala to | + | | Paraguay | + 6. _Pithecia_ | 7 | Equatorial Forests | + 7. _Brachiurus_ | 5 | Equatorial Forests | + 8. _Nyctipithecus_ | 5 | Nicaragua to Amazonia| + 9. _Saimiris_ | 3 | Costa Rica to Brazil | + | | and Bolivia | + 10. _Callithrix_ | 11 | Panama to Paraguay | + | | | + HAPALIDÆ. | | | + 11. _Hapale_ | 9 | Brazil and Upper | + | | Amazon | + 12. _Midas_ | 24 | Equatorial America | + | | to Panama | + | | | + CHIROPTERA. | | | + PHYLLOSTOMIDÆ. | | | + 13. _Lonchorina_ | 1 | West Indian Islands | + 14. _Macrophyllum_ | 1 | Brazil | + 15. _Vampyrus_ }| | | + 16. _Lophostoma_ }| 25 | Tropical America and | + }| | Chili | + 17. _Phyllostoma_ }| | | + 18. Macrotus | 1 | Antilles and Mexico | California + 19. _Schizostoma_ | 5 | South America | + 20. _Brachyphylla_ | 1 | Antilles | + 21. _Glossophaga_ | 8 | Tropical America | + 22._Phyllonycteris_| 2 | Cuba | + 23. _Artibeus_ | 4 |S. America & Antilles,| + | | Costa Rica | + 24. _Stenoderma_ | 7 | The whole region | + 25. _Sturnira_ | 3 | Chili to Guatemala | + 26. _Desmodus_ | 3 | Chili to Mexico | + 27. _Saccopteryx_ | 1 | Ecuador | + 28. _Diphylla_ | 1 | Brazil | + 29. _Centurio_ | 3 | Brazil to Mexico | + | | | + VESPERTILIONIDÆ. | | | + 30. Lasiurus | 2 | Tropical America | Nearctic + 31. Scotophilus | 7 | Antilles, Mexico to | Nearc., Austral., + | | S. America | Orien. + 32. Vespertilio | 12 | The whole region | Cosmopolite + 33. Nycticejus | 3 | S. Temperate America | Nearctic, India, + | | | Tropical Africa + 34. _Natalus_ | 1 | S. America and | + | | Antilles | + 35. _Furipterus_ | 2 | S. America | + 36. _Thyroptera_ | 2 | S. America | + 37. _Nycticellus_ | 1 | Cuba | + 38. Taphozous | 5 | S. America | Ethiopian, Oriental, + | | | Austro-Malayan + 39. _Diclidurus_ | 1 | Brazil | + | | | + NOCTILIONIDÆ. | | | + 40. _Noctilio_ | 2 | Paraguay to W. Indies| + 41. _Mormops_ | 1 | Antilles and Mexico | + 42. _Phyllodia_ | 1 | Jamaica | + 43. _Chilonycteris_| 5 | Brazil and West | + | | Indies | + 44. _Pteronotus_ | 1 | Trinidad | + 45. Nyctinomus | 2 | La Plata to Antilles | S. Nearc., Orien., + | | & Costa Rica | Madag. + 46. Molossus | 16 | Paraguay and Chili | Ethiopian, S. + | | to Antilles | Palæarc., Australian + | | | + INSECTIVORA. | | | + CENTETIDÆ. | | | + 47. _Solenodon_ | 2 | Cuba and Hayti | + | | | + SORICIDÆ. | | | + (Sorex | 1 | Guatemala and Costa | All other reg. but + | | Rica) | Austrl. + | | | + CARNIVORA. | | | + FELIDÆ. | | | + 48. Felis | 13 | The whole region, | All regions but + | | excl. Antilles | Austral. + | | | + CANIDÆ. | | | + 49. _Icticyon_ | 1 | Brazil | + 50. _Chrysocyon_ | 1 | S. America | + (Lupus | 2 | Mexico to Costa Rica)| Northern genus + 51. _Lycalopex_ | 2 | S. America | + 52. _Pseudalopex_ | 5 | S. America, Falkland | + | | Islands, & Tierra | + | | del Fuego | + 53. _Thous_ | 2 | S. America to Chili | + | | | + MUSTELIDÆ. | | | + 54. Mustela | 2 | Andes of Peru | All other reg. but + | | | Austrl. + 55. _Galictis_ | 2 | S. America to Chili | + | | & Patagonia | + 56. _Lontra_ | 3 |Central and S. America| + | | to Chonos Archipelago| + 57. Nutria | 1 | W. coast of America | W. coast of N. + | | to Chiloe | America + 58. _Pteronura_ | 1 | Surinam and Brazil | + 59. Mephitis | 3 | Mexico to Sts. of | Nearctic to Canada + | | Magellan | + | | | + PROCYONIDÆ. | | | + 60. Procyon | 1 | Tropical America | Nearctic to Canada + 61. _Nasua_ | 5 | Mexico to Paraguay & | + | | La Plata | + 62. _Cercoleptes_ | 1 | Mexico to Peru and | + | | N. Brazil | + 63. Bassaris | 2 | Mexico and Guatemala | California and Texas + | | | + URSIDÆ. | | | + 64. _Tremarctos_ | 1 | Andes of Peru and | + | | Chili | + | | | + OTARIIDÆ. | | | + 65. _Otaria_ | 1 | Chili, La Plata, and | + | | Patagonia | + 66. Arctocephalus | 1 | Falkland Islands & | New Zealand + | | Cape Horn | + | | | + PHOCIDÆ. | | | + 67. Stenorhynchus | 1 | Falkland Islands | New Zealand + 68. Lobodon | 1 | Antarctic shores | + 69. Leptonyx | 1 | Antarctic shores, | S. Australia + | | E. Patagonia | + 70. Ommatophoca | 1 | Antarctic shores | + 71. Morunga | 1 | Falkland Islands | California, S. temp. + | | | zone + 72. Cystophora | 1 | Antilles | N. Atlantic + | | | + CETACEA. | | | + DELPHINIDÆ. | | | + 73. _Inia_ | 1 | Upper Amazon | + | | | + SIRENIA. | | | + MANATIDÆ. | | | + 74. Manatus | 1 | Gulf of Mexico to N. | W. Africa + | | Brazil, Amazon R. | + | | | + UNGULATA. | | | + TAPIRIDÆ. | | | + 75. Tapirus | 2 | Equatorial S. America| Indo-Malaya + 76. _Elasmognathus_| 1 | Panama to Guatemala | + | | | + SUIDÆ. | | | + 77. _Dicotyles_ | 2 | Mexico to Paraguay | Texas + | | | + CAMELIDÆ. | | | + 78. _Auchenia_ | 4 | Temp. S. America, | + | | from Cape Horn to | + | | Andes of Peru | + | | | + CERVIDÆ. | | | + 79. Cervus | 12 | Mexico to Patagonia | All regions but + | | and Tierra del Fuego| Ethiopian and + | | | Australian + | | | + RODENTIA. | | | + MURIDÆ. | | | + 80. Reithrodon | 4 | South Temp. America | United States + | | to Tierra del Fuego| + 81. _Acodon_ | 1 | Peru, 14,000 ft. | + | | elevation | + 82. _Myxomys_ | 1 | Guatemala | + 83. Hesperomys | 76 | The whole region | Nearctic + 84. _Holochilus_ | 4 | S. America | + 85. _Oxymycterus_ | 3 | Brazil and La Plata | + 86. _Drymomys_ | 1 | Peru | + 87. _Neotomys_ | 2 | S. America | + (Fiber | 1 | Mexico) | Nearctic genus + | | | + SACCOMYIDÆ. | | | + 88. _Heteromys_ | 6 | Mexico, Honduras, | + | | Costa Rica & Trinidad| + | | | + | | | + SCIURIDÆ. | | | + 89. Sciurus | 30 | Mexico to Paraguay | All reg. but + | | | Australian + | | | + CHINCHILLIDÆ. | | | + 90. _Chinchilla_ | 2 | Andes of Chili and | + | | Peru | + 91. _Lagidium_ | 3 | Chili to Ecuador | + | | (11,000 to | + | | 16,000 ft.) | + 92. _Lagostomus_ | 1 | Uruguay to Rio Negro | + | | of Patagonia | + | | | + OCTODONTIDÆ. | | | + 93. _Habrocomus_ | 2 | Chili | + 94. _Capromys_ | 3 | Cuba and Jamaica | + 95. _Plagiodontia_ | 1 | Hayti | + 96. _Spalacopus_ | 2 | Chili and E. of Andes| + 97. _Octodon_ | 3 | Chili, Peru, and | + | | Bolivia | + 98. _Ctenomys_ | 6 | S. Brazil to Tierra | + | | del Fuego | + | | | + ECHIMYIDÆ. | | | + 99. _Dactylomys_ | 2 | Guiana and Brazil | + 100. _Cercomys_ | 1 | Central Brazil | + 101. _Lasiuromys_ | 1 | St. Paulo, Brazil | + 102. _Myopotamus_ | 1 | S. half of tropical | + | | S. America | + 103. _Carterodon_ | 1 | Central Brazil | + 104. _Mesomys_ | 1 | Upper Amazon | + 105. _Echimys_ | 11 | Equatorial America | + | | to Paraguay | + 106. _Loncheres_ | 10 | New Granada to Brazil| + | | | + CERCOLABIDÆ. | | | + 107. _Cercolabes_ | 12 | Mexico to Paraguay | + 108. _Chætomys_ | 1 | N. Brazil | + | | | + CAVIIDÆ. | | | + 109. _Dasyprocta_ | 9 | Paraguay to Mexico | + | | and Lesser Antilles| + 110. _Coelogenys_ | 2 | Guatemala to Paraguay| + 111. _Hydrochoerus_| 1 | Guiana to La Plata | + 112. _Cavia_ | 9 | Brazil and Peru to | + | | Magellan Sts. | + 113. _Kerodon_ | 6 | Brazil and Peru to | + | | Magellan Sts. | + 114. _Dolichotis_ | 1 | The Pampas and | + | | Patagonia | + | | | + LEPORIDÆ. | | | + 115. Lepus | 1 | Central Brazil and | All regions but + | | Andes, Costa Rica | Austral. + | | to Mexico | + | | | + EDENTATA. | | | + BRADYPODIDÆ. | | | + 116. _Choloepus_ | 2 | Costa Rica to Brazil | + 117. _Bradypus_ | 2 | Amazon to Rio de | + | | Janeiro | + 118._Arctopithecus_| 8 | Costa Rica to Brazil | + | | and Bolivia | + | | | + DASYPODIDÆ. | | | + 119. _Tatusia_ | 5 | Rio Grande, Texas, | + | | to Patagonia | + 120. _Prionodontes_| 1 | Surinam to Paraguay | + 121. _Dasypus_ | 4 | Brazil to Chili and | + | | La Plata, Costa Rica?| + 122. _Xenurus_ | 3 | Guiana to Paraguay, | + | | Costa Rica? | + 123. _Tolypeutes_ | 2 | Bolivia and La Plata | + 124._Chlamydophorus_ 2 | La Plata and Bolivia | + | | | + MYRMECOPHAGIDÆ. | | | + 125. _Myrmecophaga_| 1 | Costa Rica?, & | + | | N. Braz., to Parag.| + 126. _Tamandua_ | 2 | Guatemala to Paraguay| + 127. _Cyclothurus_ | 2 | Honduras and Costa | + | | Rica to Paraguay | + | | | + MARSUPIALIA. | | | + DIDELPHYIDÆ. | | | + 128. Didelphys | 20 | Mexico to Uruguay | Temperate N. America + | | and S. Chili | + 129. _Chironectes_ | 1 | Guiana and Brazil, | + | | Costa Rica | + 130. _Hyracodon_ | 1 | Ecuador | + + _BIRDS._ + + PASSERES. | | | + TURDIDÆ. | | | + 1. Turdus | 32 | The whole reg. to | Almost cosmopolite + | | Tierra del Fuego | + 2._Rhodinocichla_| 1 | Mexico to Venezuela | + 3. _Melanoptila_ | 1 | Honduras | + 4. _Catharus_ | 10 | Mexico to Ecuador | + | | and Columbia | + 5. _Margarops_ | 4 | Hayti and Lesser | + | | Antilles | + 6. Mimus | 16 | Nearly the whole | Nearctic + | | region | + 7. _Melanotis_ | 2 | Mexico and Guatemala | + 8. Galeoscoptes | 1 | Mexico to Panama | Nearctic + 9. _Mimocichla_ | 4 | Cuba to Porto Rico | + (Harporhynchus | 3 | Mexico) | Nearctic genus + 10._Cinclocerthia_| 3 | Lesser Antilles | + 11._Ramphocinclus_| 1 | Martinique and | + | | St. Lucia | + | | | + SYLVIIDÆ. | | | + 12. _Myiadestes_ | 8 |Mexico and Antilles | N. & W. of N. America + | | to Peru and Bolivia | + 13. _Cichlopsis_ | 1 | Brazil | + (Sialia | 2 | Mexico and Guatemala)| United States & + | | | Canada + 14. Regulus | 2 | Mexico and Guatemala | Nearctic, Palæarctic + 15. Polioptila | 6 | Mexico and Cuba to | Cen. and S. U. States + | | Bolivia and La Plata| + | | | + CINCLIDÆ. | | | + 16. Cinclus | 4 | Mexico to Venezuela | Nearctic, Palæarctic + | | and Peru | + | | | + TROGLODYTIDÆ. | | | + 17. Troglodytes | 5 | Mexico to Straits of | Nearctic, Palæarctic + | | Magellan | + 18. Thryophilus | 13 | Mexico to Central | N.-W. America + | | Brazil | + 19. Thryothorus | 12 | Mexico to S. Brazil | N. America + 20. Cistothorus | 3 | Mexico to Chili and | N. America + | | Patagonia | + 21. _Donacobius_ | 2 | Columbia to Brazil | + | | and Bolivia | + 22._Campylorhynchus_ 18 | Mexico to Brazil and | New Mexico + | | Bolivia | + 23. _Cyphorhinus_ | 5 | Costa Rica to Peru | + 24._Microcerculus_| 5 | Mexico to Peru | + 25. _Henicorhina_ | 2 | Mexico to Peru | + (Salpinctes | 1 | Mexico and Guatemala)| Nearctic genus + (Catherpes | 1 | Mexico) | Gila and Colorado + 26. _Cinnicerthia_| 2 | Columbia and Ecuador | + 27. _Uropsila_ | 1 | Mexico | + | | | + CERTHIIDÆ. | | | + (Certhia | 1 | Mexico and Guatemala)| North temperate genus + | | | + SITTIDÆ. | | | + (Sitta | 2 | Mexico) | North temperate genus + | | | + PARIDÆ. | | | + (Parus | 1 | Mexico) | Nearc., Palæarc., + | | | Orient. + (Lophophanes | 2 | Mexico) | North temperate genus + (Psaltriparus | 1 | Mexico and Guatemala)| Nearctic + | | | + CORVIDÆ. | | | + 28. Cyanocitta | 16 | Mexico to Peru and |Nearctic + | | Bolivia | + 29. _Cyanocorax_ | 12 | Mexico to Paraguay, | + | | Jamaica | + 30. _Calocitta_ | 2 | Mexico to Guatemala | + 31. _Psilorhinus_ | 3 | Mexico to Costa Rica | + 32. Corvus | 4 | Mexico to Guatemala, | Cosmop., excl. + | | Cuba to Porto Rico | S. Amer. + | | | + COEREBIDÆ. | | | + 33. _Diglossa_ | 14 | Mexico to Guiana, | + | | Peru, and Bolivia | + 34. _Diglossopis_ | 1 | Venezuela to Ecuador | + 35. _Oreomanes_ | 1 | Ecuador | + 36. _Conirostrum_ | 6 | Columbia to Bolivia | + 37. _Hemidacnis_ | 1 | Columbia and Upper | + | | Amazon | + 38. _Dacnis_ | 13 | Costa Rica to Guiana | + | | & S. Brazil | + 39. _Certhidea_ | 2 | Galapagos Islands | + 40. _Chlorophanes_| 2 | Brazil to Central | + | | America, Cuba | + 41. _Coereba_ | 4 | Mexico and Cuba to | + | | Guiana and Brazil | + 42. _Certhiola_ | 10 | Antilles to Ecuador | Florida + | | and Brazil | + 43. _Glossiptila_ | 1 | Jamaica | + | | | + MNIOTILTIDÆ. | | | + 44. Siurus | 3 | Mexico to Columbia, | S. & E. States & + | | Antilles | Canada + 45. Mniotilta | 1 | Columbia to Mexico | Eastern United States + | | and Antilles | + 46. Parula | 5 | Brazil and Ecuador | Eastern U. S. & + | | to Mexico | Canada + 47. Protonotaria | 1 | Venezuela to Central | Florida to Ohio + | | America and W. India| + 48. Helminthophaga| 5 | Mexico to Columbia | North America + 49. Helmintherus | 1 | Mexico to Veragua | U. States to Canada + 50. Perissoglossa | 1 | Cuba, Hayti, and | E. United States + | | Porto Rico | + 51. Dendroeca | 25 | Mexico & W. Indies to| All N. America + | | Ecuador and Chili | + 52. _Oporornis_ | 1 | Guatemala to Panama | + 53. Geothlypis | 10 | Brazil to Mexico | All N. America + 54. Setophaga | 12 | Mexico to Brazil | E. U. States & Canada + 55. _Cardellina_ | 1 | Guatemala and Mexico | + 56. _Ergaticus_ | 2 | Guatemala and Mexico | + 57. Myiodioctes | 3 | Columbia to Mexico | U. States and Canada + 58. _Basileuterus_| 22 | Mexico to Brazil | + 59. Icteria | 1 | Costa Rica to Mexico | E. and Central United + | | | States to Canada + 60. _Granatellus_ | 3 | Amazon to Mexico | + 61. _Teretristis_ | 2 | Cuba | + | | | + VIREONIDÆ. | | | + 62. Vireosylvia | 9 | Venezuela to Mexico | All N. America + | | & Antilles | + 63. Vireo | 10 | Mexico to Costa Rica | All United States + | | & Antilles | + 64. _Neochloe_ | 1 | Mexico | + 65. _Hylophilus_ | 16 | Brazil to Mexico | + 66. _Laletes_ | 1 | Jamaica | + 67._Phoenicomanes_| 1 | Jamaica | + 68. _Vireolanius_ | 4 | Mexico to Amazon | + 69. _Cychloris_ | 9 | Mexico to Paraguay | + | | | + AMPELIDÆ. | | | + 70. _Dulus_ | 2 | Hayti | + (Ampelis | 1 | Mexico and Guatemala)| N. temperate genus + 71. _Ptilogonys_ | 2 | Mexico to Costa Rica | + (Phainopepla | 1 | Mexico) | Gila and Lower + | | | Colorado + | | | + HIRUNDINIDÆ. | | | + 72. Hirundo | 9 | Mexico and Antilles | Almost cosmopolite + | | to Chili and | + | | La Plata | + 73. Petrochelidon | 3 | Mexico and Antilles | Nearctic + | | to Paraguay | + 74. _Atticora_ | 6 | Guatemala to Peru | + | | and Brazil | + 75. Cotyle | 2 | Central America to | All regions but + | | La Plata | Austral. + 76. Stelgidopteryx| 4 | Mexico to Brazil | S. United States + 77. Progne | 4 | The whole region | Nearctic + | | | + ICTERIDÆ. | | | + 78. _Clypeicterus_| 1 | Upper Amazon | + 79. _Ostinops_ | 8 | Mexico to Guiana, | + | | Brazil, and Bolivia| + 80. _Cassiculus_ | 1 | Mexico | + 81. _Cassicus_ | 10 | Mexico to S. Brazil | + | | and Bolivia | + 82. Icterus | 33 | Mexico to Antilles | All U. States & + | | and La Plata | Canada + 83. Dolichonyx | 1 | Mexico to Paraguay, | E. U. States and + | | Galapagos | Canada + 84. Molothrus | 8 | Mexico to La Plata | All U. States & + | | and Bolivia | Canada + 85. Agelæus | 6 | Mexico to Paraguay, | All U. States & + | | Cuba, Porto Rico | Canada + (Xanthocephalus| 1 | Mexico) | Nearctic genus + 86. _Xanthosomus_ | 4 | Venezuela to La Plata| + 87._Amblyrhamphus_| 1 | Bolivia and La Plata | + 88._Gymnomystax_ | 1 | Guiana and Amazonia | + 89._Pseudoleistes_| 2 | Brazil and La Plata | + 90. _Leistes_ | 3 | Venezuela to Paraguay| + | | & Bolivia | + 91. Sturnella | 4 | Cuba and Mexico to | All U. States & + | | Chili, Falkland | Canada + | | Islands & Tierra | + | | del Fuego | + 92. _Curoeus_ | 1 | Chili to Magellan | + | | Straits | + 93. _Nesopsar_ | 1 | Jamaica | + (Scolecophagus| 1 | Mexico, Cuba ?) | Nearctic genus + 94. _Lampropsar_ | 4 | Guatemala to Peru | + | | and Guiana | + 95. Quiscalus | 9 | Mexico to Antilles & | S. and E. United + | | Venezuela | States to Labrador + 96. _Hypopyrrhus_ | 1 | Columbia | + 97. _Aphobus_ | 1 | Brazil Paraguay and | + | | Bolivia | + 98. _Cassidix_ | 1 | Mexico to Brazil and | + | | Guiana | + | | | + TANAGRIDÆ. | | | + 99. _Procnias_ | 2 | Brazil and Peru to | + | | Columbia | + 100. _Chlorophonia_| 7 | Brazil to Mexico | + 101. _Euphonia_ | 32 | Mexico and W. Indies | + | | to Brazil and Bolivia| + | | | + 102. _Tanagrella_ | 4 | Columbia to Guiana | + | | and Brazil | + 103. _Chlorochrysa_| 2 | Columbia to Peru | + 104. _Pipridea_ | 2 | Venezuela to Brazil | + | | and Bolivia | + 105. _Diva_ | 1 | Columbia and Ecuador | + 106. _Calliste_ | 56 | Guatemala to Bolivia | + | | & Paraguay | + 107. _Iridornis_ | 4 | Columbia to Peru | + 108._Poecilothraupis_ 4 |Columbia to Bolivia | + 109._Stephanophorus_ 1 | Brazil and La Plata | + 110. _Buthraupis_ | 5 | Veragua to Bolivia | + 111. _Compsocoma_ | 5 | Columbia to Bolivia | + 112. _Dubusia_ | 2 | Columbia and Ecuador | + 113. _Tanagra_ | 12 | Mexico to Bolivia and| + | | La Plata | + 114. _Spindalis_ | 5 | Porto Rico to Bahamas| + 115._Rhamphocoelus_| 11 | Guatemala to Brazil | + | | and Bolivia | + 116._Phlogothraupis_ 1 | Mexico to Costa Rica | + 117. _Euchætes_ | 1 | Eastern Ecuador | + 118. _Pyranga_ | 11 | Mexico to Bolivia and| U. States and Canada + | | Paraguay | + 119. _Orthogonys_ | 2 | Brazil and Guiana | + 120. _Lamprotes_ | 2 | Brazil and Columbia | + 121._Phænicothraupis_ 7 | Mexico to Paraguay | + | | and Bolivia | + 122. _Lanio_ | 4 | Mexico to Bolivia | + 123. _Eucometis_ | 5 | Costa Rica to Bolivia| + 124._Trichothraupis_ 1 | S. Brazil and Paraguay + 125. _Creurgops_ | 1 | West Ecuador | + 126. _Tachyphonus_ | 11 | Nicaragua to Paraguay| + 127. _Cypsnagra_ | 1 | S. Brazil and Bolivia| + 128. _Nemosia_ | 11 |Venezuela, W. Ecuador,| + | | to Brazil and | + | | Bolivia | + 129. _Pyrrhocoma_ | 1 | S. Brazil and Paraguay + 130._Chlorospingus_| 18 | Mexico to Peru and | + | | Bolivia | + 131. _Buarremon_ | 20 | Mexico to S. Brazil | + | | and Bolivia | + 132._Phænicophilus_| 1 | Hayti | + 133. _Arremon_ | 12 | Mexico to S. Brazil | + 134. _Oreothraupis_| 1 | East Ecuador | + 135. _Cissopis_ | 3 | Columbia to Peru and | + | | Bolivia | + 136. _Lamprospiza_ | 1 | Guiana | + 137. _Psittospiza_ | 2 | Columbia to Peru | + 138. _Saltator_ | 17 | Mexico to La Plata | + | | and Bolivia | + 139. _Diucopis_ | 2 | Upper Amazon and | + | | S. Brazil | + 140. _Orchesticus_ | 3 | Tropical S. America | + 141. _Pitylus_ | 8 | Mexico to Brazil and | + | | Ecuador | + | | | + FRINGILLIDÆ. | | | + 142. Chrysomitris | 12 | Mexico to Brazil, | Nearctic, Palæarctic + | | Chili and Patagonia| + 143. _Sycalis_ | 9 | Mexico to Chili and | + | | La Plata, Jamaica | + 144. Coccothraustes| 2 | Mexico and Guatemala | Nearctic, Palæarctic + 145. _Geospiza_ | 7 | Galapagos Islands | + 146. _Camarhynchus_| 5 | Galapagos Islands | + 147. _Cactornis_ | 4 | Galapagos Islands | + 148. _Phrygilus_ | 10 | Columbia to Fuegia and + | | Falkland Islands | + 149. _Xenospingus_ | 1 | Peru | + 150. _Diuca_ | 3 | Peru, Chili, and | + | | Patagonia | + 151. _Emberizoides_| 3 | Venezuela to Paraguay| + 152. _Donacospiza_ | 1 |S. Brazil and La Plata| + 153. _Chamæospiza_ | 1 | Mexico | + 154. Embernagra | 9 | Mexico to La Plata | Rocky Mountains + 155. _Hæmophila_ | 6 | Mexico to Costa Rica | + 156. Atlapetes | 1 | Mexico | Nearctic? + 157. _Pyrgisoma_ | 5 | Mexico to Costa Rica | + 158. Pipilo | 4 | Mexico to Guatemala | All Nearctic region + 159. Junco | 2 | Mexico and Guatemala | United States + 160. Zonotrichia | 5 | Mexico to Straits of | Nearctic + | | Magellan | + (Melospiza | 2 | Mexico and Guatemala)| Nearctic genus + (Spizella | 3 | Mexico and Guatemala)| Nearctic genus + (Passerculus | 1 | Mexico and Guatemala)| Nearctic genus + (Pooecetes | 1 | Mexico) | Nearctic genus + 161. Ammodramus | 1 | Guatemala | Nearctic + 162. Coturniculus | 4 | Mexico to Bolivia, | E. & N. of N. America + | | Jamaica | + 163. Peucæa | 4 | Mexico | S. E. States & + | | | California + 164. _Tiaris_ | 1 | Brazil | + 165. _Volatinia_ | 1 | Mexico to Brazil | + (Cyanospiza | 4 | Mexico and Central | Nearctic + | | America) | + 166. _Paroaria_ | 6 | Trop. S. America, | + | | E. of Andes | + 167._Coryphospingus_ 4 | Tropical S. America | + 168._Porphyrospiza_| 1 | Brazil | + 169. _Haplospiza_ | 2 | Mexico and Brazil | + 170. _Phonipara_ | 5 | Mexico to Columbia, | + | | Greater Antilles | + 171. Poospiza | 12 | Mexico to Bolivia and| W. & Central + | | La Plata | U. States + 172. _Spodiornis_ | 1 | Ecuador | + (Carpodacus | 2 | Mexico) | Nearctic, Palæarctic + 173. Cardinalis | 2 | Mexico to Venezuela | S. & S. Cent. + | | | U. States + 174. Guiraca | 6 | Mexico to Brazil and | Southern U. States + | | La Plata | + 175. _Amaurospiza_ | 2 | Costa Rica and Brazil| + 176. Hedymeles | 2 | Mexico to Columbia | Nearctic + 177. _Pheucticus_ | 5 | Mexico to Peru and | + | | Bolivia | + 178. _Oryzoborus_ | 6 | Mexico to Ecuador and| + | | S. Brazil | + 179. _Melopyrrha_ | 1 | Cuba | + 180. _Loxigilla_ | 4 | Antilles | + 181. _Spermophila_ | 44 | Mexico to Bolivia and| Texas + | | Uruguay | + 182. _Catamenia_ | 4 | Columbia to Bolivia | + 183. _Neorhynchus_ | 1 | W. Peru | + 184._Catamblyrhynchus_ 1 | Columbia | + (Loxia | 1 | Mexico) | North temperate genus + (Calamospiza | 1 | Mexico) | Arizona and Texas + (Chondestes | 1 | Mexico) | W. and Cent. + | | | U. States + (Euspiza | 1 | Mexico to Columbia) | S.-E. U. States, + | | | Palæarc. + 185. _Gubernatrix_ | 1 | Paraguay and La Plata| + (Plectrophanes| 1 | Mexico) | N. temp. & Arctic + | | | genus + | | | + ALAUDIDÆ. | | | + 186. Otocorys | 1 | Mexico, Andes of | Nearc. & Palæarc. + | | Columbia | genus + | | | + MOTACILLIDÆ. | | | + 187. Anthus | 4 | Mexico to Patagonia | Cosmopolite + | | and Falkland Islands| + | | | + OXYRHAMPHIDÆ. | | | + 187a. _Oxyrhamphus_| 2 | Brazil to Costa Rica | + | | | + TYRANNIDÆ. | | | + 188. _Conophaga_ | 11 | Columbia to Bolivia | + | | and Brazil | + 189. _Corythopis_ | 2 | Brazil and Guiana | + 190. _Agriornis_ | 5 | Ecuador, Peru, and | + | | Chili | + 191. _Myiotheretes_| 3 | Columbia to Ecuador, | + | | Patagonia | + 192. _Tænioptepa_ | 8 | S. Brazil and Bolivia| + | | to Patago. | + 193. _Ochthodioeta_| 1 | Columbian Andes | + 194. _Ochthæca_ | 17 | Andes, Bolivia to | + | | Columbia and | + | | Venezuela | + 195. Sayornis | 4 | Mexico to Ecuador | E. United Sts. to + | | | Canada + 196. _Fluvicola_ | 4 | Guiana & W. Ecuador | + | | to Brazil, and | + | | Bolivia | + 197. _Arundinicola_| 1 | Tropical S. America | + 198. _Alectorurus_ | 2 |S. Brazil and La Plata| + 199. _Cybernetes_ | 1 | Brazil | + 200. _Sysopygis_ | 1 |S. Brazil and La Plata| + 201. _Cnipolegus_ | 9 | Amazonia to Patagonia| + 202. _Lichenops_ | 1 | Brazil and La Plata | + 203. _Muscipipra_ | 1 | S. Brazil | + 204. _Copurus_ | 3 | Costa Rica to | + | | S. Brazil | + 205. _Machetornis_ | 1 | Venezuela to Brazil | + 206._Muscisaxicola_| 11 | Andes of Ecuador to | + | | Chili and Patagonia| + 207. _Centrites_ | 2 | Bolivia to Patagonia | + 208. _Muscigralla_ | 1 | W. Ecuador | + 209._Platyrhynchus_| 7 | Mexico to Brazil | + 210. _Todirostrum_ | 11 | Tropical N. and | + | | S. America | + 211. _Oncosotma_ | 2 | Tropical N. America | + 212. _Euscarthmus_ | 12 | Costa Rica to W. | + | | Ecuador, Brazil, | + | | and Bolivia | + 213. _Orchilus_ | 2 | Costa Rica to Brazil | + | | and Bolivia | + 214. _Colopterus_ | 2 | Veragua to Columbia | + | | and Guiana | + 215. _Hemitriccus_ | 1 | Brazil | + 216._Phylloscartes_| 1 | Columbia to Brazil | + 217. _Hapalocercus_| 3 | Brazil to Chili and | + | | La Plata | + 218. _Habrura_ | 1 | Uruguay | + 219._Pogonotriccus_| 2 | Brazil and Columbia | + 220. _Leptotriccus_| 2 | Brazil and Veragua | + 221. _Stigmatura_ | 2 | Upper Amazon to | + | | La Plata | + 222. _Serphophaga_ | 7 | Columbia to Chili and| + | | La Plata | + 223. _Anæretes_ | 4 | Columbia to Chili and| + | | La Plata, Magell. | + | | Sts. & Juan Fernand.| + 224. _Cyanotis_ | 1 | W. Peru to La Plata | + 225. _Mionectes_ | 4 | Mexico to Brazil and | + | | Bolivia | + 226. _Leptopogon_ | 6 | Mexico to Peru and | + | | Brazil | + 227. _Capsiempis_ | 1 | Chiriqui to Brazil | + 228. _Phyllomyias_ | 5 | Columbia to Brazil | + 229. _Ornithion_ | 4 | Mexico to Brazil | + 230. _Tyrannulus_ | 3 | Guatemala to Amazonia| + 231. _Tyranniscus_ | 9 | Guatemala to E. Peru | + 232. _Elainea_ | 18 | Mexico to Tierra del | + | | Fuego, Antilles | + 233. _Empidagra_ | 1 | Bolivia and La Plata | + 234. _Legatus_ | 2 | Mexico to Brazil | + 235. _Sublegatus_ | 2 | Venezuela and Lower | + | | Amazon | + 236. _Myiozetetes_ | 8 | Mexico to W. Peru and| + | | Brazil | + 237._Rhynchocyclus_| 10 | Mexico to W. Ecuador | + | | & Brazil | + 238. _Conopias_ | 3 | Venezuela to Peru and| + | | Brazil | + 239. _Pitangus_ | 7 | Mexico to La Plata, | + | | Antilles | + 240. _Sirystes_ | 2 | Panama to Brazil | + 241. _Myiodynastes_| 6 | Mexico to Bolivia and| + | | Paraguay | + 242. _Megarhynchus_| 1 | Mexico to Brazil | + 243. _Muscivora_ | 5 | Mexico to W. Ecuador | + | | & Brazil | + 244. _Hirundinea_ | 3 | Columbia & Guiana to | + | | Paraguay | + 245. _Cnipodectes_ | 1 | Panama to W. Ecuador | + | | & Amazon | + 246. _Myiobius_ | 13 | Mexico to W. Peru, | + | | Bolivia, and | + | | La Plata | + 247. _Pyrocephalus_| 3 | Tropical N. and S. | Gila and Rio Grande + | | America and | + | | Galapagos Islands | + 248. _Empidochanes_| 4 | Venezuela to | + | | S. Brazil. | + 249. _Mitrephorus_ | 2 | Mexico to Costa Rica | + 250. Empidonax | 12 | Mexico to Columbia & | All N. America + | | Ecuador | + 251. Contopus | 10 | Mexico to Amazonia, | N. & E. of Rocky + | | Antilles | Mtns. + 252. _Myiochanes_ | 1 | Amazonia and Brazil | + 253. Myiarchus | 12 | Mexico to W. Ecuador | East and West Coasts + | | & Brazil, Galapagos| to Canada + | | and Antilles | + 254. _Blacicus_ | 2 | Cuba, Hayti, Jamaica | + (Empidias | 1 | Mexico) | Eastern United States + 255. _Empidonomus_ | 1 | Guiana and Brazil | + 256. Tyrannus | 11 | All tropical | All U. States to + | | sub-regions | Canada + 257. _Milvulus_ | 2 | Tropical N. and S. | Texas + | | America | + | | | + PIPRIDÆ. | | | + 258. _Piprites_ | 4 | Costa Rica to Brazil | + 259. _Masius_ | 2 | Columbia and Ecuador | + 260. _Chloropipo_ | 1 | Columbia | + 261. _Xenopipo_ | 1 | Guiana and Columbia | + 262. _Pipra_ | 19 |Trop. N. and S. America + 263. _Neopipo_ | 1 | Upper Amazon | + 264._Machæropterus_| 4 | Columbia to Brazil | + 265. _Ilicura_ | 1 | Brazil | + 266. _Chiroxiphia_ | 5 | Guatemala to Brazil | + 267. _Metopia_ | 1 | Brazil | + 268. _Metopothrix_ | 1 | Upper Amazon | + 269._Chiromachæris_| 6 | Mexico to Ecuador and| + | | Brazil | + 270. _Hetoropelma_ | 10 | Mexico to Guiana and | + | | Brazil | + 271. _Heterocercus_| 2 | Guiana and Upper | + | | Amazon | + 272. _Schiffornis_ | 2 |Upper Amazon and Brazil + | | | + COTINGIDÆ. | | | + 273. _Tityra_ | 6 | Tropical N. and S. | + | | America | + 274. _Hadrostomus_ | 5 | Mexico to W. Ecuador | + | | & Brazil, Jamaica | + 275. _Pachyhamphus_| 11 | Mexico to W. Ecuador | + | | & Brazil | + 276. _Lathria_ | 5 | Mexico to Brazil | + 277. _Aulia_ | 3 | Veragua to Brazil | + 278. _Lipaugus_ | 3 | Guatemala to Brazil | + | | and Guiana | + 279. _Ptilochloris_| 2 | Brazil | + 280. _Attila_ | 8 | Costa Rica to Brazil | + | | and Guiana | + 281. _Casiornis_ | 2 | S. Brazil to Paraguay| + 282. _Rupicola_ | 3 | Guiana to W. Ecuador | + | | & Bolivia | + 283._Phoenicocercus_ 2 | Guiana and Amazonia | + 284. _Tijuca_ | 1 | Brazil | + 285. _Phibalura_ | 1 | Brazil | + 286. _Pipreola_ | 7 | Venezuela to Ecuador | + | | and Peru | + 287. _Ampelio_ | 4 | Columbia to Peru and | + | | Brazil | + 288. _Carpodectes_ | 1 | Nicaragua and Costa | + | | Rica | + 289. _Heliochæra_ | 2 | Columbia to Peru and | + | | Bolivia | + 290. _Cotinga_ | 6 | Guatemala to Peru and| + | | Brazil | + 291. _Xipholena_ | 3 | Guiana to Brazil | + 292. _Iodopleura_ | 3 | Guiana to Brazil | + 293. _Calyptura_ | 1 | Brazil | + 294. _Querula_ | 1 | Panama to Amazonia | + 295. _Hæmatoderus_ | 1 |Guiana and Lower Amazon + 296._Chasmorhynchus_ 4 | Costa Rica to Guiana | + | | and Brazil | + 297._Gymnocephalus_| 1 | Guiana and Rio Negro | + 298. _Gymnoderus_ | 1 |Guiana and Upper Amazon + 299. _Pyroderus_ | 3 | Venezuela to Brazil | + 300._Cephalopterus_| 3 | Costa Rica to W. | + | | Ecuador & Upr. Amazon| + | | | + PHYTOTOMIDÆ. | | | + 301. _Phytotoma_ | 3 | Bolivia, Chili, and | + | | La Plata | + | | | + DENDROCOLAPTIDÆ. | | | + 302. _Geobates_ | 1 | South Brazil | + 303. _Geositta_ | 6 | Peru to Chili and | + | | Patagonia | + 304. _Furnarius_ | 9 | Guiana & W. Ecuador | + | | to La Plata | + 305. _Clibanornis_ | 1 | S. Brazil | + 306. _Upucerthia_ | 4 | Andes of Ecuador to | + | | Chili and Patagonia| + 307. _Cinclodes_ | 5 | Ecuador to Chili, | + | | Patagonia and | + | | Tierra del Fuego | + 308. _Henicornis_ | 2 | Patagonia | + 309. _Lochmias_ | 2 | Venezuela and Brazil | + 310. _Sclerurus_ | 6 | Mexico to Brazil | + 311. _Oxyurus_ | 2 | Chili to Tierra del | + | | Fuego, and | + | | Masafuera Islands | + 312. _Sylviortho- | 1 | Chili | + rhynchus_ | 1 | Chili | + 313. _Phlæocryptes_| 1 | W. Peru to La Plata | + 314._Leptasthenura_| 5 | Andes of Ecuador to | + | | Brazil and Patagonia| + 315. _Synallaxis_ | 55 | The whole region | + | | (excl. Antilles) | + 316. _Coryphistera_| 1 | La Plata | + 317. _Anumbius_ | 1 | Paraguay and La Plata| + 318. _Limnornis_ | 1 | Uruguay and La Plata | + 319._Placellodomus_| 4 | Venezuela to Peru and| + | | La Plata | + 320. _Thripophaga_ | 3 | Brazil and Columbia | + 321._Pseudocolaptes_ 1 | Columbia to Peru | + 322. _Homorus_ | 3 | Brazil, Bolivia, and | + | | La Plata | + 323. _Thripadectes_| 1 | Columbia | + 324. _Ancistrops_ | 1 | Upper Amazon | + 325. _Automolus_ | 9 | Mexico to Amazonia | + 326. _Philydor_ | 14 |Tropical South America| + | | | + 327. _Heliobletus_ | 1 | Brazil | + 328. _Anabatoides_ | 1 | Brazil | + 329. _Anabazenops_ | 5 | Mexico to Brazil | + 330. _Xenops_ | 3 | Trop. North and South| + | | America | + 331. _Sittasomus_ | 3 | Mexico to Ecuador and| + | | Brazil | + 332. _Margarornis_ | 4 | Costa Rica to Peru | + | | and Bolivia | + 333._Glyphorhynchus_ 1 | Trop. North and South| + | | America | + 334. _Pygarrhicus_ | 1 | Chili | + 335. _Dendrocincla_| 10 | Mexico to Venezuela | + | | and Brazil | + 336._Dendrocolaptes_ 7 | Guatemala to Peru and| + | | Brazil | + 337. _Nasica_ | 1 | Guiana | + 338. _Drymornis_ | 1 | La Plata | + 339._Xiphocolaptes_| 5 | Mexico to Bolivia and| + | | Paraguay | + 340._Dendrexetastes_ 2 | Guiana | + 341. _Dendrornis_ | 14 | Mexico, W. Ecuador | + | | and Brazil | + 342. _Dendroplex_ | 2 | Columbia & Venezuela | + | | to Brazil | + 343. _Picolaptes_ | 14 | Mexico to Bolivia and| + | | La Plata | + 344._Xiphorhynchus_| 4 | Veragua to Brazil | + | | | + FORMICARIIDÆ. | | | + 345. _Cymbilanius_ | 1 | Amazonia and Guiana | + 346. _Batara_ | 1 | S. Brazil | + 347. _Thamnophilus_| 47 | Trop. North and South| + | | America | + 348. _Biatas_ | 1 | Brazil | + 349. _Thamnistes_ | 2 | Central America and | + | | Ecuador | + 350. _Pygoptila_ | 2 | Amazonia | + 351. _Neoctantes_ | 1 | Amazonia | + 352. _Clytoctantes_| 1 | Eastern Ecuador | + 353. _Dysithamnus_ | 12 | Mexico to Bolivia and| + | | Brazil | + 354. _Thamnomanes_ | 2 | Ecuador, Guiana, and | + | | Brazil | + 355._Herpsilochmus_| 4 | Venezuela to Brazil | + | | and Bolivia | + 356. _Myrmotherula_| 21 | Tropical S. America | + 357. _Formicivora_ | 14 | Trop. North and South| + | | America | + 358. _Terenura_ | 3 | Veragua to W. Ecuador| + | | & Brazil | + 359._Psilorhamphus_| 1 | Central Brazil | + 360. _Microbates_ | 1 | Cayenne | + 361. _Rhamphocænus_| 4 | Guatemala to Brazil | + 362. _Cercomacra_ | 9 | Cen. America to W. | + | | Equador & S. Brazil| + 363. _Pyriglena_ | 4 | Ecuador to Peru and | + | | Brazil | + 364. _Gymnocichla_ | 2 | Honduras to Panama | + 365. _Percnostola_ | 3 |Guiana and Upper Amazon + 366. _Heterocnemis_| 3 |Guiana and Upper Amazon + 367. _Myrmeciza_ | 11 |Veragua to W. Ecuador,| + | | Bolivia, and Brazil| + 368. _Hypocnemis_ | 15 | Costa Rica to W. | + | | Ecuador & Brazil | + 369. _Pithys_ | 5 | Nicaragua to Amazonia| + 370. _Rhopoterpe_ | 1 | Guiana | + 371. _Phlogopsis_ | 4 | Nicaragua to Guiana | + | | and Bolivia | + 372. _Formicarius_ | 9 | Mexico to Brazil and | + | | Bolivia | + 373. _Pittasoma_ | 1 | Panama and Veragua | + 374. _Chamæza_ | 4 | Columbia to Brazil | + 375. _Grallaria_ | 20 | Mexico to W. Ecuador | + | | & Brazil | + 376. _Grallaricula_| 5 | Costa Rica to Ecuador| + | | | + PTEROPTOCHIDÆ. | | | + 377. _Scytalopus_ | 8 | Columbia & Brazil to | + | | Chili and Tierra | + | | del Fuego | + 378. _Merulaxis_ | 1 | Central Brazil | + 379. _Rhinocrypta_ | 2 | La Plata and | + | | Patagonia | + 380. _Liosceles_ | 1 | Madeira Valley | + 381. _Pteroptochus_| 2 | Chili and Chiloe | + 382. _Hylactes_ | 3 | Chili | + 383. _Acropternis_ | 1 | Columbia and Ecuador | + 384. _Triptorhinus_| 1 | Chili | + | | | + PICARIÆ. | | | + PICIDÆ. | | | + 385. _Picumnus_ | 14 | Honduras to Brazil | + | | and Bolivia | + 386. Picus | 6 | Mexico, Chili, La | All reg. but Austral. + | | Plata, and S. | & Ethiopian + | | Patagonia | + (Sphyrapicus | 1 | Mexico and Guatemala)| Nearctic genus + 387. Campephilus | 12 | Mexico to Patagonia, | Nearctic + | | Cuba | + 388. Dryocopus | 4 | Mexico to S. Brazil | Palæarctic + 389. _Celeus_ | 15 | Mexico and S. Brazil | + 390. _Nesoceleus_ | 1 | Cuba | + 391. _Chrysoptilus_| 6 | Tropical S. America | + 392. Centurus | 10 | Mexico to Venezuela, | Nearctic + | | Antilles | + 393. _Chloronerpes_| 35 | Tropical America, | + | | Hayti | + 394._Xiphidiopicus_| 1 | Cuba | + 395. Melanerpes | 9 | Mexico to Brazil, | Nearctic + | | Porto Rico | + 396. _Leuconerpes_ | 1 | Brazil, Bolivia | + 397. Colaptes | 7 | Open country of trop.| Nearctic + | | America, Greater | + | | Antilles | + 398. _Hypoxanthus_ | 1 | Venezuela and Ecuador| + | | | + MEGALÆMIDÆ. | | | + 399. _Capito_ | 10 | Costa Rica to Peru | + | | and Guiana | + 400. _Tetragonops_ | 2 | Costa Rica and | + | | Ecuador | + | | | + RHAMPHASTIDÆ. | | | + 401. _Rhamphastos_ | 12 | All tropical America | + 402. _Pteroglossus_| 16 | Mexico to Guiana and | + | | Brazil | + 403. _Selenidera_ | 7 | Veragua to Brazil | + 404. _Andigena_ | 6 |Columbia to W. Ecuador, + | | Bolivia and Brazil | + 405._Aulacorhamphus_ 10 | Mexico to Venezuela | + | | and Bolivia | + | | | + CUCULIDÆ. | | | + 406. _Crotophaga_ | 3 | Tropical America and | Nearctic to + | | Antilles | Pennsylvania + 407. _Guira_ | 1 | Brazil and Paraguay | + 408. _Neomorphus_ | 4 | Nicaragua to Brazil | + | | and Upper Amazon | + 409. _Geococcyx_ | 1 | Guatemala | Texas to Calfornia + 410. _Dromococcyx_ | 2 | Mexico to Brazil | + 411. _Diplopterus_ | 1 | Mexico to Ecuador and| + | | Brazil | + 412. _Saurothera_ | 4 | Greater Antilles | + 413. _Hyetornis_ | 2 | Jamaica and Hayti | + 414. _Piaya_ | 3 | Mexico to W. Ecuador | + | | & Brazil | + 415. _Morococcyx_ | 1 | Mexico to Costa Rica | + 416. Coccygus | 10 | Tropical America and | Nearctic + | | Antilles, Cocos | + | | Islands | + | | | + BUCCONIDÆ. | | | + 417. _Bucco_ | 21 | Guatemala to Guiana, | + | | Paraguay and Bolivia| + 418. _Malacoptila_ | 10 | Guatemala to Guiana, | + | | W. Ecuador and | + | | Bolivia | + 419. _Nonnula_ | 5 | Columbia and Amazonia| + 420. _Monasa_ | 7 | Costa Rica to Brazil | + 421. _Chelidoptera_| 2 | Columbia to Guiana | + | | and Brazil | + GALBRILIDÆ. | | | + 422. _Galbula_ | 9 | Guatemala to Brazil | + | | and Bolivia | + 423. _Urogalba_ | 2 |Guiana to Lower Amazon| + | | | + 424. _Brachygalba_ | 4 | Columbia to Brazil | + | | and Bolivia | + 425._Jacamaralcyon_| 1 | Brazil | + 426. _Jacamerops_ | 2 | Columbia to Amazonia | + 427._Galbalcyrhynchus_ 1 | Upper Amazon | + | | | + TODIDÆ. | | | + 428. _Todus_ | 5 | Greater Antilles | + | | | + MOMOTIDÆ. | | | + 429. _Momotus_ | 10 | Mexico to W. Ecuador,| + | | Brazil and Bolivia | + 430. _Urospatha_ | 1 |Costa Rica to Columbia| + 431._Baryphthengus_| 1 | Brazil and Paraguay | + 432. _Hylomanes_ | 2 | Mexico and Guatemala | + 433._Prionirhynchus_ 2 | Guatemala to Upper | + | | Amazon | + 434. _Eumomota_ | 1 | Honduras to Chiriqui | + | | | + TROGONIDÆ. | | | + 435. _Prionoteles_ | 1 | Cuba | + 436. _Temnotrogon_ | 1 | Hayti | + 437. _Trogon_ | 22 | Mexico to W. Ecuador | + | | & Parag. | + 438. _Euptilotis_ | 1 | Mexico | + 439. _Pharomacrus_ | 5 | Guatemala to Upper | + | | Amazon and Bolivia | + | | | + ALCEDINIDÆ. | | | + 440. Ceryle | 8 | Mexico to Brazil, | Nearc., S. Palæarc., + | | Patagonia and Chili| Orien. + | | | + STEATORNITHIDÆ. | | | + 441. _Steatornis_ | 1 | Columb., Venezuela, &| + | | Trinidad | + | | | + CAPRIMULGIDÆ. | | | + 442. _Nyctibius_ | 6 | Brazil to Guatemala | + | | & Jamaica | + 443. _Hydropsalis_ | 8 | Columbia & Guiana to | + | | La Plata | + 444. Antrostomus | 10 | Mexico and Cuba to | All U. States to + | | Bolivia and La Plata| Canada + 445. _Stenopsis_ | 4 |Martinique to Columb.,| + | | W. Peru and Chili | + 446. _Siphonorhis_ | 1 | Jamaica | + 447._Heleothreptus_| 1 | Central Brazil | + 448. _Nyctidromus_ | 1 | Central America to | + | | S. Brazil | + 449. _Podager_ | 1 | Tropical S. America | + 450. _Lurocalis_ | 2 | Guiana to Brazil | + 451. Chordeiles | 7 | Mexico to W. Peru and| All U. States to + | | Brazil Jamaica and | Canada + | | Porto Rico | + 452. _Nyctiprogne_ | 1 | Amazonia | + | | | + CYPSELIDÆ. | | | + 453. Cypselus | 3 | Antilles to Guiana | The Eastern + | | and Bolivia | Hemisphere + 454. _Panyptila_ | 3 | Guatemala and Guiana | + 455. Chætura | 9 | Mexico to Ecuador and| Almost cosmopolite + | | Brazil | + 456. _Hemiprocne_ | 3 | Mexico to La Plata, | + | | Jamaica and Hayti | + 457. _Cypseloides_ | 2 | Brazil and Peru | + 458. _Nephoecetes_ | 1 | Jamaica | + | | | + TROCHILIDÆ. | | | + 459. _Grypus_ | 1 | Brazil | + 460. _Androdon_ | 1 | Ecuador | + 461. _Eutoxeres_ | 2 | Costa Rica to Ecuador| + 462. _Glaucis_ | 2 | Panama to Brazil | + 463. _Phaethornis_ | 14 | Tropical N. and | + | | S. America | + 464. _Pygmornis_ | 8 | Mexico to Guiana and | + | | Brazil | + 465. _Threnetes_ | 4 |Costa Rica to Amazonia| + | | and W. Ecuador | + 466. _Dolerisca_ | 1 | Venezuela | + 467. _Eupetomena_ | 1 | Guiana to Brazil | + 468._Sphenoproctus_ 2 | Mexico to Guatemala | + 469._Campylopterus_ 9 | Mexico to Amazonia | + 470. _Phæochroa_ | 2 | Guatemala to Columbia| + 471. _Aphantochroa_| 3 | Ecuador and Brazil | + 472. _Urochroa_ | 1 | Ecuador | + 473. _Sternoclyta_ | 1 | Venezuela | + 474. _Eugenes_ | 2 | Mexico to Costa Rica | + 475. _Coeligena_ | 1 | Mexico | + 476. _Lamprolæma_ | 1 | Mexico and Guatemala | + 477. _Delattria_ | 2 | Guatemala | + 478. _Oreopyra_ | 4 |Costa Rica to Chiriqui| + 479. _Heliopædica_ | 2 | Mexico and Guatemala | + 480. _Topaza_ | 2 | Guiana | + 481._Oreotrochilus_| 6 | Ecuador to Peru and | + | | Chili | + 482. _Lampornis_ | 7 | Mexico & W. India to | + | | Amazonia | + 483. _Eulampis_ | 2 | Lesser Antilles | + 484. _Avocettula_ | 1 | Guiana | + 485. _Lafresnaya_ | 2 |Venezuela and Columbia| + 486. _Doryphora_ | 5 | Costa Rica to Ecuador| + 487. _Chalybura_ | 5 |Costa Rica to Columbia| + 488. _Heliodoxa_ | 5 |Costa Rica to Venezue.| + | | & Boliv. | + 489. _Iolæma_ | 2 | Ecuador to Peru | + 490. _Phæolæma_ | 2 | Columbia and Ecuador | + 491. _Eugenia_ | 1 | Ecuador | + 492. _Aithurus_ | 1 | Jamaica | + 493. _Thalurania_ | 10 | Costa Rica to Guiana,| + | | Ecuador and Brazil | + 494. _Panoplites_ | 3 | Columbia and Ecuador | + 495. _Florisuga_ | 2 | Guatemala to Brazil | + 496. _Microchera_ | 2 | Nicaragua to Veragua | + 497. _Lophorius_ | 7 | Mexico to Brazil, | + | | Peru, & Bolivia | + 498. _Polemistria_ | 2 | Columbia to S. Brazil| + 499. _Discura_ | 2 | Brazil | + 500. _Gouldia_ | 4 | Costa Rica to Brazil | + | | & Bolivia | + 501. Trochilus | 2 | Mexico to Veragua | To Canada and Sitka + 502. _Mellisuga_ | 1 | Jamaica to Hayti | + 503. _Calypte_ | 3 | Mexico and Cuba | + 504. Selasphorus | 7 | Mexico to Veragua | W. & Cen. United + | | | States + 505. Atthis | 1 | Mexico and Guatemala | California and + | | | Colorado + 506. _Stellula_ | 1 | Mexico | + 507. _Calothorax_ | 2 | Mexico | + 508. _Acestrura_ | 3 | Venezuela to Ecuador | + | | & Bolivia | + 509. _Chætocercus_ | 3 | Venezuela and Ecuador| + 510. _Myrtis_ | 2 | Ecuador to Bolivia, | + | | W. of Andes | + 511. _Thaumastura_ | 1 | W. Peru | + 512. _Rhodopis_ | 2 | W. Peru and Chili | + 513. _Doricha_ | 5 | Mexico to Veragua, | + | | Bahamas | + 514. _Tilmatura_ | 1 | Guatemala | + 515. _Calliphlox_ | 2 | Ecuador and Brazil | + 516. _Loddigesia_ | 1 | Peruvian Andes | + 517. _Steganura_ | 6 | Venezuela to Ecuador | + | | & Bolivia | + 518. _Lesbia_ | 6 | Columbia to Peru | + 519. _Cynanthus_ | 2 | Venezuela to Ecuador | + 520. _Sparganura_ | 4 | Columbia to Bolivia &| + | | La Plata | + 521. _Pterophanes_ | 1 | Columbia to Peru | + 522. _Aglæactis_ | 4 | Columbia to Bolivia | + 523. _Oxypogon_ | 2 | Venezuela and Columbia | + 524. _Oreonympha_ | 1 | Peru | + 525._Rhamphomicron_| 6 | Columbia to Bolivia | + 526. _Urosticte_ | 2 | Ecuador | + 527. _Metallura_ | 6 | Columbia to Bolivia | + 528. _Adelomia_ | 4 | Venezuela to Peru & | + | | Bolivia | + 529. _Avocettinus_ | 1 | Columbia | + 530. _Anthocephala_| 1 | Columbia | + 531. _Chrysolampis_| 1 | Venezuela to Brazil | + 532._Orthorhynchus_| 2 | Lesser Antilles | + 533. _Cephalolepis_| 3 | Brazil | + 534. _Clais_ | 1 |Venezuela and Columbia| + 535. _Baucis_ | 1 | Mexico to Veragua | + 536. _Heliactin_ | 1 | Brazil | + 537. _Heliothrix_ | 3 | Guatemala to Ecuador | + | | & Brazil | + 538. _Schistes_ | 2 | Columbia and Ecuador | + 539. _Phlogophilus_| 1 | Ecuador | + 540. _Augastes_ | 2 | Brazil | + 541. _Petasophora_ | 5 | Mexico to Peru and | + | | Brazil | + 542._Chrysobronchus_ 3 | Venezuela to Brazil | + 543. _Patagona_ | 1 | Ecuador to Bolivia | + | | and Chili | + 544. _Docimastes_ | 1 | Columbia and Ecuador | + 545. _Helianthea_ | 7 | Columbia to Bolivia | + 546. _Heliotrypha_ | 2 | Columbia and Ecuador | + 547. _Heliangelus_ | 6 | Venezuela to Peru | + 548. _Diphlogæna_ | 3 | Bolivia | + 549. _Clytolæma_ | 2 | E. Ecuador and Brazil| + 550. _Bourcieria_ | 5 | Venezuela to Peru | + 551. _Lampropygia_ | 4 | Venezuela to Bolivia | + 552. _Heliomastes_ | 5 | Mexico to Ecuador & | + | | Venezuela | + 553. _Lepidolarynx_| 1 | Brazil | + 554. _Calliperidia_| 1 | Central Brazil and | + | | Paraguay | + 555. _Eustephanus_ | 3 | Chili, S. Patagonia, | + | | and Juan Fernandez | + | | Island | + 556. _Eriocnemis_ | 14 | Venezuela to Ecuador | + 557. _Cyanomyia_ | 6 | Mexico to Peru | + 558. _Hemistilbon_ | 1 | Mexico | + 559. _Leucippus_ | 2 | Peru and Bolivia | + 560. _Thaumatias_ | 15 | Mexico to Guiana, | + | | Upr. Amazon, and | + | | Brazil | + 561. _Amazilia_ | 14 | Mexico to W. Ecuador | + | | & Peru | + 562. _Saucerottia_ | 7 | Costa Rica to Columb.| + | | & Venezue. | + 563. _Eupherusa_ | 3 | Mexico to Veragua | + 564. _Chrysuronia_ | 5 | Guatemala to Ecuador | + | | & La Plata | + 565. _Eucephala_ | 7 | Venezuela to Guiana | + | | and Brazil | + 566. _Panterpe_ | 1 | Costa Rica and | + | | Chiriqui | + 567. _Juliamyia_ | 2 | Panama to Ecuador | + 568. _Circe_ | 3 | Mexico | + 569. _Phæoptila_ | 1 | Mexico | + 570. _Damophila_ | 1 | Costa Rica to Ecuador| + 571. _Hylocharis_ | 3 | Amazonia and Brazil | + 572. _Sapphironia_ | 2 | Columbia and Veragua | + 573. _Sporadinus_ | 3 | Cuba, Bahamas, Hayti,| + | | Porto Rico | + 574._Chlorostilbon_ 8 | Mexico to Brazil and | + | | La Plata | + 575. _Panychlora_ | 3 |Venezuela and Columbia| + 576._Smaragdochrysis_ 1 | Brazil | + | | | + PSITTACI. | | | + CONURIDÆ. | | | + 577. _Ara_ | 15 | Trop. North and South| + | | America, Cuba, | + | | Jamaica (extinct) | + 578. _Rhyncopsitta_| 1 | Mexico | + 579._Henicognathus_| 1 | Chili | + 580. Conurus | 30 | The whole region | S. & S.E. United + | | | States + 581. _Pyrrhura_ | 16 | Costa Rica to | + | | Paraguay & Bolivia | + 582._Bolborhynchus_| 7 | Mexico to Peru, | + | | Central Brazil, and| + | | La Plata | + 583. _Brotogerys_ | 9 | Trop. North and South| + | | America | + | | | + PSITTACIDÆ. | | | + 584. _Caica_ | 9 | Mexico to Amazonia | + 585. _Chrysotis_ | 32 | All the tropical | + | | sub-regions | + 586. _Triclaria_ | 1 | Brazil | + 587. _Deroptyus_ | 1 | Guiana and Rio Negro | + 588. _Pionus_ | 9 | Costa Rica to Bolivia| + | | and Brazil | + 589. _Urochroma_ | 7 | Venezuela to Brazil | + 590. _Psittacula_ | 6 | Mexico to W. Ecuador | + | | & Brazil | + | | | + COLUMBÆ. | | | + 591. Columba | 18 |Trop. sub-regions with| All regions but + | | Chili and La Plata | Austral. + 592. Zenaidura | 2 | Mexico to Veragua | Nearctic + 593. Chamæpelia | 6 | Mexico to Brazil and | S. Nearctic + | | Bolivia | + 594. _Columbula_ | 2 | Brazil and La Plata | + | | to Chili | + 595. _Scardafella_ | 2 | Guatemala and Brazil | + 596. _Zenaida_ | 10 | Antilles and S. | + | | America to Chili | + | | and La Plata | + 597. Melopelia | 2 | Mexico to Chili | South & West + | | | Nearctic + 598. _Peristera_ | 4 | Mexico to Brazil | + 599. _Metriopelia_ | 2 | W. America from | + | | Ecuador to Chili | + 600. _Gymnopelia_ | 1 | West Peru and Bolivia| + 601. _Leptoptila_ | 11 | Tropical sub-regions | + 602. _Geotrygon_ | 14 | Tropical sub-regions | + 603. _Starnoenas_ | 1 | Cuba | + | | | + GALLINÆ. | | | + TETRAONIDÆ. | | | + 604. _Odontophorus_| 17 | Trop. North and South| + | | America | + 605. _Dendrortyx_ | 3 | Mexico to Costa Rica | + 606. Cyrtonyx | 3 | Mexico to Guatemala | S. Central United + | | | States + 607. Ortyx | 5 | Mexico to Costa Rica,| Nearctic to Canada + | | Cuba | + 608. _Eupsychortyx_| 5 | Mexico to Columbia | + | | and Guiana | + (Callipepla | 2 | Mexico) | California + | | | + PHASIANIDÆ. | | | + 609. Meleagris | 2 | Mexico and Honduras | Nearctic + | | | + CRACIDÆ. | | | + 610. _Crax_ | 8 | Mexico to Venezuela &| + | | S. Brazil | + 611. _Nothocrax_ | 1 | Guiana and Upper | + | | Amazon | + 612. _Pauxi_ | 1 | Guiana and Venezuela | + 613. _Mitua_ | 2 | Guiana to Peru | + 614. _Stegnolæma_ | 1 | Columbia and Ecuador | + 615. _Penelope_ | 13 | Trop. North and South| + | | America | + 616. _Penelopina_ | 1 | Guatemala | + 617. _Pipile_ | 3 | Venezuela to Brazil | + | | and Peru | + 618. Aburria | 1 | Columbia | + 619. _Chamæpetes_ | 2 | Costa Rica to Peru | + 620. _Ortalida_ | 18 | Trop. North and South| New Mexico + | | America | + 621. _Oreophasis_ | 1 | Guatemala | + | | | + TINAMIDÆ. | | | + 622. _Tinamus_ | 7 | Trop. North and South| + | | America | + 623. _Nothocercus_ | 3 |Costa Rica to Venezue.| + | | & Ecuador | + 624. _Crypturus_ | 16 | Trop. North and South| + | | America | + 625. _Rhynchotus_ | 2 | Brazil to Bolivia and| + | | La Plata | + 626. _Nothoprocta_ | 4 | Ecuador to Bolivia | + | | and Chili | + 627. _Nothura_ | 4 | Brazil to Bolivia and| + | | La Plata | + 628. _Taoniscus_ | 1 | Brazil and Paraguay | + 629. _Calodromas_ | 1 | La Plata | + 630. _Tinamotis_ | 1 | Andes of Peru and | + | | Bolivia | + | | | + OPISTHOCOMI. | | | + OPISTHOCOMIDÆ | | | + 631. _Opisthocomus_| 1 | Guiana and Lower | + | | Amazon | + | | | + ACCIPITRES. | | | + VULTURIDÆ. | | | + (CATHARTINÆ.) | | | + 632._Sarcorhamphus_| 2 | The Andes and S. of | + | | 41° S. Lat. | + 633. _Cathartes_ | 1 | Mexico to 20° S. Lat.| + 634. Catharista | 1 | Mexico to 40° S. Lat.| S. United States + 635. Pseudogryphis | 3 | Mexico to Falkland | United States + | | Ids., Cuba, Jamaica| + | | | + FALCONIDÆ. | | | + 636. Polyborus | 2 | The whole region | California and + | | | Florida + 637. _Ibycter_ | 8 | Guatemala to Terra | + | | del Fuego | + 638. Circus | 3 | Nearly the whole | Almost cosmopolite + | | region | + 639. _Micrastur_ | 7 | Trop. North and South| + | | America | + 640. _Geranospiza_ | 2 | Trop. North and South| + | | America | + 641. Antenor | 1 | Mexico to Chili and | California and Texas + | | La Plata | + 642. Astur | 2 | Trop. N. and S. | Almost cosmopolite + | | America | + 643. Accipiter | 9 | The whole region | Almost cosmopolite + 644._Heterospizias_| 1 | Trop. S. America, | + | | E. of Andes | + 645. Tachytriorchis| 2 | Mexico to Paraguay | California + 646. Buteo | 9 | Mexico to Patagonia | Almost cosmopolite + 647. _Buteola_ | 1 | Veragua to Amazonia | + 648. Asturina | 7 | Mexico to Bolivia and| S.E. United States + | | La Plata | + 649. _Busarellus_ | 1 | Brazil and Guiana | + 650. _Buteogallus_ | 1 | Columbia and Guiana | + 651. _Urubutinga_ | 12 | Mexico to Brazil and | + | | Bolivia | + 652._Harpyhaliæetus_ 1 | Veragua to Chili & | + | | N. Patagonia | + 653. _Morphnus_ | 1 | Panama to Amazonia | + 654. _Thrasaëtus_ | 1 | Mexico to Bolivia and| + | | Paraguay | + 655. Lophotriorchis| 1 | Bogota | Indo-Malaya + 656. _Spiziastur_ | 1 | Guatemala to Brazil | + 657. Spizaëtus | 4 | Mexico to Paraguay | Africa, India, Malaya + 658._Herpetotheres_| 1 | S. Mexico to Bolivia | + | | & Paraguay | + 659. Nauclerus | 1 | Mexico to Brazil | S. United States + 660. _Rostrhamus_ | 3 | Antilles to Brazil | Florida + | | and Peru | + 661. _Leptodon_ | 4 | Central America to S.| + | | Brazil and Bolivia | + 662. Elanus | 1 | Mexico to Chili | Califor., Old World + | | | trop. + 663. _Gampsonyx_ | 1 | Trinidad to Brazil | + 664. _Harpagus_ | 3 | Central America to | + | | Brazil & Peru | + 665. _Ictinia_ | 2 | Mexico to Brazil | South United States + 666. _Spiziapteryx_| 1 | La Plata | + 667. Falco | 3 | The whole region | Almost cosmopolite + 668. Cerchneis | 3 | The whole region | Almost cosmopolite + | | | + PANDIONIDÆ. | | | + 669. Pandion | 1 | The whole region | Cosmopolite + | | | + STRIGIDÆ. | | | + 670. Glaucidium | 6 | The whole region | W. United Sts., + | | | Palæarc. + 671. Micrathene | 1 | Mexico | Arizona, New Mexico + 672. Pholeoptynx | 1 | The whole region | N. W. America & Texas + 673. Bubo | 1 | The whole region | All regions but + | | | Austral. + 674. Scops | 6 | Mexico to Brazil and | Almost cosmopolite + | | La Plata | + 675. _Gymnoglaux_ | 2 | West India Islands | + 676. _Lophostrix_ | 2 | Guatemala to Lower | + | | Amazon | + 677. Syrnium | 3 | Mexico to Patagonia | All regions but + | | | Austral. + 678. _Ciccaba_ | 10 | Mexico to Peru and | + | | Paraguay | + 679. _Nyctalatinus_| 1 | Columbia | + 680. _Pulsatrix_ | 2 | Guatemala to Brazil | + | | and Peru | + 681. Asio | 2 | The whole region | All regions but + | | | Austral. + 682. _Nyctalops_ | 1 | Cuba and Mexico to | + | | Brazil | + 683. _Pseudoscops_ | 1 | Jamaica | + (Nyctale | 1 | Mexico) | N. Temperate genus + 684. Strix | 2 | The whole region | Almost cosmopolite + + _Peculiar or very Characteristic Genera of Wading and Swimming Birds._ + + | | | + GRALLÆ. | | | + RALLIDÆ. | | | + Aramides | 23 | The whole region | Nearctic + _Heliornis_ | 1 | Tropical America | + | | | + SCOLOPACIDÆ. | | | + Eureunetes | 3 | The whole region | Nearctic + | | | + CHIONIDIDÆ. | | | + Chionis | 2 | Sts. of Magellan, | Kerguelen's Island + | | Falkland Ids. | + | | | + THINOCORIDÆ. | | | + _Attagis_ | 4 | Andes to Fuegia and | + | | Falkland Islands | + _Thinocoris_ | 2 | Peru, Chili, and | + | | La Plata | + | | | + CHARADRIIDÆ. | | | + _Phægornis_ | 1 | Temperate S. America | + _Oreophilus_ | 1 | Temperate S. America | + _Pluvianellus_ | 1 | Temperate S. America | + _Aphriza_ | 1 |W. coast of S. America| W. coast of + | | | N. America + | | | + CARIAMIDÆ. | | | + _Cariama_ | 2 |S. Brazil and La Plata| + | | | + ARAMIDÆ. | | | + _Aramus_ | 5 | Mexico and Cuba to | + | | Brazil | + | | | + PSOPHIIDÆ. | | | + _Psophia_ | 6 | Equatorial S. America| + | | | + EURYPYGIDÆ. | | | + _Eurypyga_ | 2 | Tropical America | + | | | + ARDEIDÆ. | | | + _Tigrisoma_ | 3 | The whole region | + _Cancroma_ | 1 | Tropical S. America | + | | | + PALAMEDEIDÆ. | | | + _Palamedea_ | 1 | Equatorial America | + _Chauna_ | 2 | Columbia, Brazil, and| + | | La Plata | + | | | + ANSERES. | | | + ANATIDÆ. | | | + _Cairina_ | 1 | Tropical S. America | + _Merganetta_ | 3 | Andes | + _Micropterus_ | 1 | Temperate S. America | + | | | + SPHENISCIDÆ. | | | + Eudyptes | 6 | Temperate S. America| Antarctic shores + Aptenodytes | 2 | Falkland Islands | Antarctic shores + | | | + STRUTHIONES. | | | + STRUTHIONIDÆ. | | | + 685. _Rhea_ | 3 | S. Temperate America | + +{114}CHAPTER XV. + +THE NEARCTIC REGION. + + +This region consists almost wholly of Temperate North America as defined by +physical geographers. In area it is about equal to the Neotropical region. +It possesses a vast mountain range traversing its entire length from north +to south, comparable with, and in fact a continuation of, the Andes,--and a +smaller range near the east coast, equally comparable with the mountains of +Brazil and Guiana. These mountains supply its great river-system of the +Mississippi, second only to that of the Amazon; and in its vast group of +fresh-water lakes or inland seas, it possesses a feature unmatched by any +other region, except perhaps by the Ethiopian. It possesses every variety +of climate between arctic and tropical; extensive forests and vast +prairies; a greatly varied surface and a rich and beautiful flora. But +these great advantages are somewhat neutralized by other physical features. +It extends far towards the north, and there it reaches its greatest width; +while in its southern and warmest portion it suddenly narrows. The northern +mass of land causes its isothermal lines to bend southwards; and its winter +temperature especially, is far lower than at corresponding latitudes in +Europe. This diminishes the available area for supporting animal life; the +amount and character of which must be, to a great extent, determined by the +nature of the least favourable part of the year. Again, owing to the +position of its mountain ranges and the direction of prevalent winds, a +large extent of its interior, east of the Rocky Mountains, is bare and +arid, and often almost desert; while the most favoured districts,--those +east of the Mississippi and west of the Sierra Nevada, bear but a small +proportion to its whole area. Again, we know that at a very recent period +geologically, it was subjected to a very severe Glacial epoch, which +wrapped a full half of it in a mantle of ice, and exterminated a large +number of animals which previously inhabited it. Taking all this into +account, we need not be surprised to find the Nearctic region somewhat less +rich and varied in its forms of life than the Palæarctic or the Australian +regions, with which alone it can fairly be compared. The wonder rather is +that it should be so little inferior to them in this respect, and that it +should possess such a variety of groups, and such a multitude of forms, in +every class of animals. + + +[Illustration: NEARCTIC REGION] + +{115}_Zoological characteristics of the Nearctic Region._--Temperate North +America possesses representatives of 26 families of Mammalia, 48 of Birds, +18 of Reptiles, 11 of Amphibia, and 18 of Fresh-water Fish. The first three +numbers are considerably less than the corresponding numbers for the +Palæarctic region, while the last two are greater--in the case of fishes +materially so, a circumstance readily explained by the wonderful group of +fresh-water lakes and the noble southward-flowing river system of the +Mississippi, to which the Palæarctic region has nothing comparable. But +although somewhat deficient in the total number of its families, this +region possesses its full proportion of peculiar and characteristic family +and generic forms. No less than 13 families or sub-families of Vertebrata +are confined to it, or just enter the adjacent Neotropical region. These +are,--three of mammalia, Antilocaprinæ, Saccomyidæ and Haploodontidæ; one +of birds, Chamæidæ; one of reptiles, Chirotidæ; two of amphibia, Sirenidæ +and Amphiumidæ; and the remaining six of fresh-water fishes. The number of +peculiar or characteristic genera is perhaps more important for our +purpose; and these are very considerable, as the following enumeration will +show. + +_Mammalia._--Of the family of moles (Talpidæ) we have 3 peculiar genera: +_Condylura_, _Scapanus_, and _Scalops_, as well as the remarkable +_Urotrichus_, found only in California and Japan. In the weasel family +(Mustelidæ) we have _Latax_, a peculiar kind of otter; _Taxidea_, allied to +the badgers; and one of the {116}remarkable and characteristic skunks is +separated by Dr. J. E. Gray as a genus--_Spilogale_. In the American family +Procyonidæ, a peculiar genus (_Bassaris_) is found in California and Texas, +extending south along the mountains of Mexico and Guatemala. _Eumetopias_, +and _Halicyon_, are seals confined to the west coast of North America. The +Bovidæ, or hollow-horned ruminants, contain three peculiar forms; +_Antilocapra_, the remarkable prong-buck of the Rocky Mountains; +_Aplocerus_, a goat-like antelope; and _Ovibos_, the musk-sheep, confined +to Arctic America and Greenland. Among the Rodents are many peculiar +genera: _Neotoma_, _Sigmodon_, and _Fiber_, belong to the Muridæ, or rats; +_Jaculus_ to the Dipodidæ, or jerboas. The very distinct family +_Saccomyidæ_, or pouched rats, which have peculiar cheek pouches, or a kind +of outer hairy mouth, consists of five genera all confined to this region, +with one of doubtful affinities in Trinidad and Central America. In the +squirrel family (Sciuridæ), _Cynomys_, the prairie-dogs, are peculiar; and +_Tamias_, the ground squirrel, is very characteristic, though found also in +North Asia. _Haploodon_, or sewellels, consisting of two species, forms a +distinct family; and _Erethizon_ is a peculiar form of tree porcupine +(Cercolabidæ). True mice and rats of the genus _Mus_ are not indigenous to +North America, their place being supplied by a distinct genus +(_Hesperomys_), confined to the American continent. + +_Birds._--The genera of birds absolutely peculiar to the Nearctic region +are not very numerous, because, there being no boundary but one of climate +between it and the Neotropical region, most of its characteristic forms +enter a short distance within the limits we are obliged to concede to the +latter. Owing also to the severe winter-climate of a large part of the +region (which we know is a comparatively recent phenomenon), a large +proportion of its birds migrate southwards, to pass the winter in the +West-Indian islands or Mexico, some going as far as Guatemala, and a few +even to Venezuela. + +In our chapter on extinct animals, we have shown, that there is good reason +for believing that the existing union of North and South America is a quite +recent occurrence; and that the {117}separation was effected by an arm of +the sea across what is now Nicaragua, with perhaps another at Panama. This +would leave Mexico and Guatemala joined to North America, and forming part +of the Nearctic region, although no doubt containing many Neotropical +forms, which they had received during earlier continental periods; and +these countries might at other times have been made insular by a strait at +the isthmus of Tehuantepec, and have then developed some peculiar species. +The latest climatal changes have tended to restrict these Neotropical forms +to those parts where the climate is really tropical; and thus Mexico has +attained its present strongly marked Neotropical character, although +deficient in many of the most important groups of that region. + +In view of these recent changes, it seems proper not to draw any decided +line between the Nearctic and Neotropical regions, but rather to apply, in +the case of each genus, a test which will show whether it was probably +derived at a comparatively recent date from one region or the other. The +test referred to, is the existence of peculiar species of the genus, in +what are undoubtedly portions of ancient North or South America. If, for +example, all the species of a genus occur in North America, some, or even +all, of them, migrating into the Neotropical region in winter, while there +are _no peculiar Neotropical species_, then we must class that genus as +strictly Nearctic; for if it were Neotropical it would certainly have +developed _some_ peculiar resident forms. Again, even if there should be +one or two resident species peculiar to that part of Central America north +of the ancient dividing strait, with an equal or greater number of species +ranging over a large part of Temperate North America, the genus must still +be considered Nearctic. Examples of the former case, are _Helminthophaga_ +and _Myiodioctes_, belonging to the Mniotiltidæ, or wood-warblers, which +range over _all_ Temperate North America to Canada, where _all_ the species +are found, but in each case one of the species is found in South America, +probably as a winter migrant. Of the latter, are _Ammodramus_ and _Junco_ +(genera of finches), which range over the whole United States, but each +have one peculiar species in Guatemala. These {118}may be claimed as +exclusively Nearctic genera, on the ground that Guatemala was recently +Nearctic; and is now really a transition territory, of which the lowlands +have been invaded and taken exclusive possession of by a Neotropical fauna, +while the highlands are still (in part at least) occupied by Nearctic +forms. + +In his article on "Birds," in the new edition of the "Encyclopædia +Britannica" (now publishing), Professor Newton points out, that the number +of _peculiar genera_ of Nearctic birds is much less than in each of the +various sub-divisions of the Neotropical region; and that the total number +of genera is also less, while the bulk of them are common either to the +Neotropical or Palæarctic regions. This is undoubtedly the case if any +fixed geographical boundary is taken; and it would thus seem that the +"Nearctic" should, in birds, form a sub-region only. But, if we define +"Nearctic genera" as above indicated, we find a considerable amount of +speciality, as the following list will show. The names not italicised are +those which are represented in Mexico or Guatemala by peculiar species:-- + + +LIST OF TYPICAL NEARCTIC GENERA OF LAND BIRDS. + + 1. _Oreoscoptes_ + 2. _Harporhynchus_ + 3. _Sialia_ + 4. _Chamæa_ + 5. _Catherpes_ + 6. _Salpinctus_ + 7. _Psaltriparus_ + 8. _Auriparus_ + 9. _Gymnokitta_ + 10. _Picicorvus_ + 11. _Mniotilta_ + 12. _Oporornis_ + 13. _Icteria_ + 14. _Helmintherus_ + 15. _Helminthophaga_ + 16. _Myiodioctes_ + 17. _Phænopepla_ + 18. _Xanthocephalus_ + 19. _Scolecophagus_ + 20. Pipilo + 21. Junco + 22. _Melospiza_ + 23. Spizella + 24. _Passerculus_ + 25. _Pooecetes_ + 26. Ammodromus + 27. _Cyanospiza_ + 28. _Pyrrhuloxia_ + 29. _Calamospiza_ + 30. Chondestes + 31. _Centronyx_ + 32. _Neocorys_ + 33. _Empidias_ + 34. _Sphyrapicus_ + 35. _Hylatomus_ + 36. _Trochilus_ + 37. _Atthis_ + 38. _Ectopistes_ + 39. _Centrocercus_ + 40. _Pediocætes_ + 41. _Cupidonia_ + ? Ortyx + 42. _Oreortyx_ + 43. _Lophortyx_ + 44. Callipepla + 45. Cyrtonyx + 46. Meleagris + 47. _Micrathene_ + +The above are all groups which are either wholly Nearctic or typically so, +but entering more or less into the debatable ground of the Neotropical +region; though none possess any peculiar species in the ancient Neotropical +land south of Nicaragua. But we have, besides these, a number of genera +which we are {119}accustomed to consider as typically European, or +Palæarctic, having representatives in North America; although in many cases +it would be more correct to say that they are Nearctic genera, represented +in Europe, since America possesses more species than Europe or North Asia. +The following is a list of genera which have as much right to be considered +typically Nearctic as Palæarctic:-- + + 1. Regulus + 2. Certhia + 3. Sitta + 4. Parus + 5. Lophophanes + 6. Lanius + 7. Perisoreus + 8. Pica + 9. Corvus + 10. _Ampelis_ + 11. Loxia + 12. Pinicola + 13. Linota + 14. _Passerelia_ + 15. _Leucosticte_ + 16. _Euspiza_ + 17. _Plectrophanes_ + 18. Tetrao + 19. Lagopus + 20. _Nyctala_ + 21. _Archibuteo_ + 22. Haliæetus + +The seven genera italicized have a decided preponderance of Nearctic +species, and have every right to be considered typically Nearctic; while +the remainder are so well represented by peculiar species, that it is quite +possible many of them may have originated here, rather than in the +Palæarctic region, all alike being quite foreign to the Neotropical. + +On the whole, then, we have 47 in the first and 7 in the second table, +making 54 genera which we may fairly class as typically Nearctic, out of a +total of 168 genera of land-birds, or nearly one-third of the whole. This +is an amount of peculiarity which is comparable with that of either of the +less isolated regions; and, combined with the more marked and more +exclusively peculiar forms in the other orders of vertebrates, fully +establishes Temperate North America as a region, distinct alike from the +Neotropical and the Palæarctic. + +_Reptiles._--Although temperate climates are always comparatively poor in +reptiles, a considerable number of genera are peculiar to the Nearctic +region. Of snakes, there are, _Conophis_, _Chilomeniscus_, _Pituophis_, and +_Ischnognathus_, belonging to the Colubridæ; _Farancia_, and _Dimodes_, +Homalopsidæ; _Lichanotus_, one of the Pythonidæ; _Cenchris_, +_Crotalophorus_, _Uropsophorus_, and _Crotalus_, belonging to the Crotalidæ +or rattlesnakes. + +Of Lizards, _Chirotes_, forming a peculiar family; _Ophisaurus_, {120}the +curious glass-snake, belonging to the Zonuridæ; with _Phrynosoma_ (commonly +called horned toads), _Callisaurus_, _Uta_, _Euphryne_, _Uma_, and +_Holbrookia_, genera of Iguanidæ. + +Testudinidæ, or Tortoises, show a great development of the genus _Emys_; +with _Aromochelys_ and _Chelydra_ as peculiar genera. + +_Amphibia._--In this class the Nearctic region is very rich, possessing +representatives of nine of the families, of which two are peculiar to the +region, and there are no less than fifteen peculiar genera. _Siren_ forms +the family Sirenidæ; _Menobranchus_ belongs to the Proteidæ; _Amphiuma_ is +the only representative of the Amphiumidæ; there are nine peculiar genera +of Salamandridæ. Among the tail-less batrachians (frogs and toads) we have +_Scaphiopus_, belonging to the Alytidæ; _Pseudacris_ to the Hylidæ; and +_Acris_ to the Polypedatidæ. + +_Fresh-water Fishes._--The Nearctic region possesses no less than five +peculiar family types, and twenty-four peculiar genera of this class. The +families are Aphredoderidæ, consisting of a single species found in the +Eastern States; Percopsidæ, founded on a species peculiar to Lake Superior; +Heteropygii, containing two genera peculiar to the Eastern States; +Hyodontidæ and Amiidæ, each consisting of a single species. The genera are +as follows: _Paralabrax_, found in California; _Huro_, peculiar to Lake +Huron; _Pileoma_, _Boleosoma_, _Bryttus_ and _Pomotis_ in the Eastern +States--all belonging to the perch family. _Hypodelus_ and _Noturus_, +belonging to the Siluridæ. _Thaleichthys_, one of the Salmonidæ peculiar to +the Columbia river. _Moxostoma_, _Pimephales_, _Hyborhynchus_, +_Rhinichthys_, in the Eastern States; _Ericymba_, _Exoglossum_, +_Leucosomus_, and _Carpiodes_, more widely distributed; _Cochlognathus_, in +Texas; _Mylaphorodon_ and _Orthodon_, in California; _Meda_, in the river +Gila; and _Acrochilus_, in the Columbia river--all belonging to the +Cyprinidæ. _Scaphirhynchus_, found only in the Mississippi and its +tributaries, belongs to the sturgeon family (Accipenseridæ). + +_Summary of Nearctic Vertebrata._--The Nearctic region possesses 24 +peculiar genera of mammalia, 49 of birds, 21 of reptiles, and 29 of +fresh-water fishes, making 123 in all. Of these 70 are mammals and +land-birds, out of a total of 242 {121}genera of these groups, a proportion +of about two-sevenths. This is the smallest proportion of peculiar genera +we have found in any of the regions; but many of the genera are of such +isolated and exceptional forms that they constitute separate families, so +that we have no less than 12 families of vertebrata confined to the region. +The Palæarctic region has only 3 peculiar families, and even the Oriental +region only 12; so that, judged by this test, the Nearctic region is +remarkably well characterized. We must also remember that, owing to the +migration of many of its peculiar forms during the Glacial period, it has +recently lost some of its speciality; and we should therefore give some +weight to the many characteristic groups it possesses, which, though not +quite peculiar to it, form important features in its fauna, and help to +separate it from the other regions with which it has been thought to be +closely allied. It is thus well distinguished from the Palæarctic region by +its Procyonidæ, or racoons, _Hesperomys_, or vesper mice, and _Didelphys_, +or opossums, among Mammalia; by its Vireonidæ, or greenlets, Mniotiltidæ, +or wood-warblers, Icteridæ, or hang-nests, Tyrannidæ, or tyrant shrikes, +and Trochilidæ, or humming-birds, among birds, families which, extending to +its extreme northern limits must be held to be as truly characteristic of +it as of the Neotropical region; by its Teidæ, Iguanidæ, and _Cinosternum_, +among reptiles; and by its Siluridæ, and Lepidosteidæ, among fishes. From +the Neotropical region it is still more clearly separated, by its numerous +insectivora; by its bears; its Old World forms of ruminants; its beaver; +its numerous _Arvicolæ_, or voles; its _Sciuropterus_, or flying squirrels; +_Tamias_, or ground-squirrels; and _Lagomys_, or marmots, among mammals; +its numerous Paridæ, or tits, and Tetraonidæ, or grouse, among birds; its +Trionychidæ among reptiles; its Proteidæ, and Salamandridæ, among Amphibia; +and its Gasterosteidæ, Atherinidæ, Esocidæ, Umbridæ, Accipenseridæ, and +Polydontidæ, among fishes. + +These characteristic features, taken in conjunction with the absolutely +peculiar groups before enumerated, demonstrate that the Nearctic region +cannot with propriety be combined with {122}any other. Though not very +rich, and having many disadvantages of climate and of physical condition, +it is yet sufficiently well characterized in its zoological features to +rank as one of the well-marked primary divisions of the earth's surface. + +There is one other consideration bearing on this question which should not +be lost sight of. In establishing our regions we have depended wholly upon +their _now_ possessing a sufficient number and variety of animal forms, and +a fair proportion of peculiar types; but when the validity of our +conclusion on these grounds is disputed, we may supplement the evidence by +an appeal to the past history of the region in question. In this case we +find a remarkable support to our views. During the whole Tertiary period, +North America was, zoologically, far more strongly contrasted with South +America than it is now; while, during the same long series of ages, it was +always clearly separated from the Eastern hemisphere or the Palæarctic +region by the exclusive possession of important families and numerous +genera of Mammalia, as shown by our summary of its extinct fauna in Chapter +VII. Not only may we claim North America as now forming one of the great +zoological regions, but as having continued to be one ever since the Eocene +period. + + +_Insects._ + +In describing the Palæarctic and Neotropical regions, many of the +peculiarities of the insect-fauna of this region have been incidentally +referred to; and as a tolerably full account of the distribution of the +several families is given in the Fourth Part of our work (Chapter XXI.), we +shall treat the subject very briefly here. + +_Lepidoptera._--The butterflies of the Nearctic region have lately been +studied with much assiduity, and we are now able to form some idea of their +nature and extent. Nearly 500 species belonging to about 100 genera have +been described; showing that the region, which a few years ago was thought +to be very poor in species of butterflies, is really much richer than +Europe, and probably about as rich as the more extensive Palæarctic region. +There is, however, very little speciality in the {123}forms. A considerable +number of Neotropical types enter the southern States; but there are hardly +any peculiar genera, except one of the Lycænidæ and perhaps a few among the +Hesperidæ, The most conspicuous feature of the region is its fine group of +Papilios, belonging to types (_P. turnus_ and _P. troilus_) which are +characteristically Nearctic. It is also as rich as the Palæarctic region in +some genera which we are accustomed to consider as pre-eminently European; +such as _Argynnis_, _Melitæa_, _Grapta_, _Chionabas_, and a few others. +Still, we must acknowledge, that if we formed our conclusions from the +butterflies alone, we could hardly separate the Nearctic from the +Palæarctic region. This identity probably dates from the Miocene period; +for when our existing arctic regions supported a luxuriant vegetation, +butterflies would have been plentiful; and as the cold came on, these would +move southwards both in America and Europe, and, owing to the long +continuance of the generic types of insects, would remain little modified +till now. + +_Coleoptera._--Only a few indications can be given of the peculiarities of +the Nearctic coleoptera. In Cicindelidæ the region possesses, besides the +cosmopolite _Cicindela_, four other genera, two of which--_Amblychile_ and +_Omus_--are peculiar to the West Coast and the Rocky Mountains. Of Carabidæ +it possesses _Dicælus_, _Pasimachus_, _Eurytrichus_, _Sphæroderus_, +_Pinacodera_, and a number of smaller genera, altogether peculiar to it; +_Helluomorpha_, _Galerita_, _Callida_, and _Tetragonoderus_, in common with +South America; and a large number of characteristic European forms. + +The Lucanidæ are all of European types. The region is poor in Cetoniidæ, +but has representatives of the South American _Euphoria_, as well as of +four European genera. Of Buprestidæ it has the South American _Actenodes_; +a single species of the Ethiopian and Eastern _Belionota_, in California; +and about a dozen other genera of European and wide distribution. + +Among Longicorns it possesses fifty-nine peculiar genera, representatives +of five Neotropical, and thirteen Palæarctic genera; as well as many of +wider distribution. _Prionus_ is the chief representative of the Prionidæ; +_Leptura_ and _Crossidius_ of the {124}Cerambycidæ; _Leptostylus_, +_Liopus_, _Graphidurus_, and _Tetraopes_, of the Lamiidæ, the latter genus +being confined to the region. + + +_Terrestrial and Fluviatile Mollusca._ + +The land-shells of temperate North America almost all belong to the +Inoperculate or Pulmoniferous division; the Operculata being represented +only by a few species of _Helicina_ and _Truncatella_, chiefly in the +Southern States. According to Mr. Binney's recent "Catalogue of the +Terrestrial Air-breathing Mollusks of North America," the fauna consists of +the following genera:--_Glandina_ (6 sp.); _Macrocyclis_ (5 sp.); _Zonites_ +(37 sp.); _Vitrina_ (4 sp.); _Limax_ (5 sp.); _Arion_ (3 sp.); _Ariolimax_ +(3 sp.); _Prophysaon_ (1 sp.); _Binneia_ (1 sp.); _Hemiphillia_ (1 sp.); +_Patula_ (16 sp.); _Helix_ (80); _Holospira_ (2 sp.); _Cylindrella_ (2 +sp.); _Macroceramus_ (2 sp.); _Bulimulus_ (8 sp.); _Cionella_ (2 sp.); +_Stenogyra_ (4 sp.); _Pupa_ (19 sp.); _Strophia_ (1 sp.); _Vertigo_ (6 +sp.); _Liguus_ (1 sp.); _Orthalicus_ (2 sp.); _Punctum_ (1 sp.); _Succinea_ +(26 sp.); _Tebennophorus_ (1 sp.); _Pallifera_ (1 sp.); _Veronicella_ (2 +sp.). + +All the larger genera range over the whole region, but the following have a +more restricted distribution; _Macrocyclis_ has only one species in the +East, the rest being Californian or Central; _Ariolimax_, _Prophysaon_, +_Binneia_, and _Hemiphillia_, are confined to the Western sub-region. Lower +California has affinities with Mexico, 18 species being peculiar to it, of +which two are true _Bulimi_, a genus unknown in other parts of the region. +The Central or Rocky Mountain sub-region is chiefly characterised by six +peculiar species of _Patula_. The Eastern sub-region is by far the richest, +nine-tenths of the whole number of species being found in it. The Alleghany +Mountains form the richest portion of this sub-region, possessing nearly +half the total number of species, and at least 24 species found nowhere +else. The southern States have also several peculiar species, but they are +not so productive as the Alleghanies. The Canadian sub-region possesses 32 +species, of which nearly half are northern forms more or less common to the +whole Arctic regions, and several of this character have spread southwards +all {125}over the United States. Species of _Vitrina_, _Zonites_, _Pupa_, +and _Succinea_, are found in Greenland; and Eastern Palæarctic species of +_Vitrina_, _Patula_, and _Pupa_ occur in Alaska. More than 30 species of +shells living in the Eastern States, are found fossil in the Post-Pliocene +deposits of the Ohio and Mississippi. + +_Fresh-water Shells._--North America surpasses every other part of the +globe in the number and variety of its fresh-water mollusca, both univalve +and bivalve. The numbers up to 1866 were as follows:--Melaniadæ, 380 +species; Paludinidæ, 58 species; Cycladidæ, 44 species; and Unionidæ, 552 +species. The last family had, however, increased to 832 species in 1874, +according to Dr. Isaac Lea, who has made them his special study; but it is +probable that many of these are such as would be considered varieties by +most conchologists. Many of the species of _Unio_ are very large, of varied +forms, and rich internal colouring, and the group forms a prominent feature +of the Nearctic fauna. By far the larger proportion of the fresh-water +shells inhabit the Eastern or Alleghany sub-region; and their great +development is a powerful argument against any recent extensive submergence +beneath the ocean of the lowlands of North America. + + +_The Nearctic Sub-regions._ + +The sub-divisions of the Nearctic region, although pretty clearly indicated +by physical features and peculiarities of climate and vegetation, are by no +means so strongly marked out in their zoology as we might expect. The same +genera, as a rule, extend over the whole region; while the species of the +several sub-regions are in most cases different. Even the vast range of the +Rocky Mountains has not been an effectual barrier against this wide +dispersal of the same forms of life; and although some important groups are +limited by it, these are exceptions to the rule. Even now, we find fertile +valleys and plateaus of moderate elevation, penetrating the range on either +side; and both to the north and south there are passes which can be freely +traversed by most animals during the summer. Previous to the glacial epoch +there was probably a warm period, when every part of the range supported an +abundant and varied {126}fauna, which, when the cold period arrived, would +descend to the lowlands, and people the country to the east, west, and +south, with similar forms of life. + +The first, and most important sub-division we can make, consists of the +Eastern United States, extending across the Mississippi and the more +fertile prairies, to about the 100°th. meridian of west longitude, where +the arid and almost desert country commences. Southwards, the boundary +bends towards the coast, near the line of the Brazos or Colorado rivers. To +the north the limits are undefined; but as a considerable number of species +and genera occur in the United States but not in Canada, it will be +convenient to draw the line somewhere near the boundary of the two +countries, except that the district between lakes Huron and Ontario, and +probably Nova Scotia, may be included in the present sub-region. As far +west as the Mississippi, this was originally a vast forest country; and it +is still well wooded, and clothed with a varied and luxuriant vegetation. + +The next, or Central sub-region, consists of the dry, elevated, and often +arid district of the Rocky Mountains, with its great plateaus, and the +barren plains of its eastern slope; extending northwards to near the +commencement of the great forests north of the Saskatchewan, and southward +to the Rio Grande del Norte, the Gulf of California, and to Cape St. Lucas, +as shown on our maps. This sub-region is of an essentially desert +character, although the higher valleys of the Rocky Mountains are often +well wooded, and in these are found some northern and some western types. + +The third, or Californian sub-region, is small, but very luxuriant, +occupying the comparatively narrow strip of country between the Sierra +Nevada and the Pacific. To the north it may include Vancouver's Island and +the southern part of British Columbia, while to the south it extends to the +head of the Gulf of California. + +The fourth division, comprises the remainder of North America; and is a +country of pine forests, and of barren wastes towards the Arctic Ocean. It +has fewer peculiar species to characterise it than any other, but it +possesses several characteristic arctic {127}forms, while many of those +peculiar to the south are absent; so that it is a very convenient, if it +should not be considered an altogether natural, sub-region. + +We will now give an outline of the most important zoological features of +each of these divisions, taking them in the order in which they are +arranged in the Fourth Part of this work. California comes first, as it has +some tropical forms not found elsewhere, and thus forms a transition from +the Neotropical region. + + +_I. The Western or Californian Sub-region._ + +This small district possesses a fruitful soil and a highly favourable +climate, and is, in proportion to its extent, perhaps the richest portion +of the continent, both zoologically and botanically. Its winters are far +milder than those of the Eastern States in corresponding latitudes; and +this, perhaps, has enabled it to support several tropical forms which give +a special character to its fauna. It is here only, in the whole region, +that bats of the families Phyllostomidæ and Noctilionidæ, and a serpent of +the tropical family, Pythonidæ, are found, as well as several Neotropical +forms of birds and reptiles. + +_Mammalia._--The following genera are not found in any other part of the +Nearctic region. _Macrotus_ (Phyllostomidæ), one species in California; +_Antrozous_ (Vespertilionidæ), one species on the West Coast; _Urotrichus_ +(Talpidæ) one species in British Columbia; sub-genus _Nesorex_ (Soricidæ), +one species in Oregon; _Bassaris_ (Procyonidæ), California; _Enhydra_ +(Mustelidæ), Pacific Coast; _Morunga_ (Phocidæ), California; _Haploodon_ +(Haploodontidæ) a rat-like animal, allied to the beavers and marmots, and +constituting a peculiar family found only in California and British +Columbia. The following characteristic Nearctic forms also extend into this +sub-region:--_Taxidea_, _Procyon_, _Didelphys_, _Sciuropterus_, _Tamias_, +_Spermophilus_, _Dipodomys_, _Perognathus_, _Jaculus_. + +_Birds._--Few genera of birds are quite peculiar to this sub-region, since +most of the Western forms extend into the central district, yet it has a +few. _Glaucidium_, a genus of Owls, is confined {128}(in the Nearctic +region) to California; _Chamæa_, a singular form allied to the wrens, and +forming a distinct family, is quite peculiar; _Geococcyx_, a Neotropical +form of cuckoo, extends to California and Southern Texas. The following +genera are very characteristic of the sub-region, and some of them almost +confined to it: _Myiadestes_ (Sylviidæ); _Psaltriparus_ (Paridæ); +_Cyanocitta_, _Picicorvus_ (Corvidæ); _Hesperiphona_, _Peucæa_, +_Chondestes_ (Fringillidæ); _Selasphorus_, _Atthis_ (Trochilidæ); +_Columba_, _Melopelia_ (Columbidæ); _Oreortyx_ (Tetraonidæ). + +_Reptiles._--The following genera are not found in any other part of the +Nearctic region: _Charina_ (Tortricidæ); _Lichanotus_ (Pythonidæ); +_Gerrhonotus_ (Zonuridæ); _Phyllodactylus_ (Geckotidæ); _Anolius_ and +_Tropidolepis_ (Iguanidæ). _Sceloporus_ (Iguanidæ) is only found elsewhere +in Florida. All the larger North American groups of lizards and snakes are +also represented here; but in tortoises it is deficient, owing to the +absence of lakes and large rivers. + +_Amphibia._--California possesses two genera of Salamandridæ, _Aneides_ and +_Heredia_, which do not extend to the other sub-regions. + +_Fresh-water Fish._--There are two or three peculiar genera of Cyprinidæ, +but the sub-region is comparatively poor in this group. + +_Plate XVIII. Illustrative of the Zoology of California and the Rocky +Mountains._--We have chosen for the subject of this illustration, the +peculiar Birds of the Western mountains. The two birds in the foreground +are a species of grouse (_Pediocætes Columbianus_), entirely confined to +this sub-region; while the only other species of the genus is found in the +prairies north and west of Wisconsin, so that the group is peculiar to +northern and western America. The crested birds in the middle of the +picture (_Oreortyx picta_), are partridges, belonging to the American +sub-family Odontophorinæ. This is the only species of the genus which is +confined to California and Oregon. The bird at the top is the blue crow +(_Gymnokitta cyanocephala_), confined to the Rocky Mountains and Sierra +Nevada from New Mexico and Arizona northwards, and more properly belonging +to the Central sub-region. It is allied to the European nutcracker; but +according to the American ornithologist, Dr. Coues, has also resemblances +to the jays, and certainly forms a distinct genus. The grizzly bear (_Ursus +ferox_) in the background, is one of the characteristic animals of the +Californian highlands. + + + +Plate XVIII. + +[Illustration] + +SCENE IN CALIFORNIA, WITH SOME CHARACTERISTIC BIRDS. + + +{129}_II. The Central, or Rocky Mountain Sub-region._ + +This extensive district is, for the greater part of its extent, from 2,000 +to 5,000 feet above the sea, and is excessively arid; and, except in the +immediate vicinity of streams and on some of the higher slopes of the +mountains, is almost wholly treeless. Its zoology is therefore peculiar. +Many of the most characteristic genera and families of the Eastern States +are absent; while a number of curious desert and alpine forms give it a +character of its own, and render it very interesting to the naturalist. + +_Mammalia._--The remarkable prong-horned antelope (_Antilocapra_), the +mountain goat (_Aplocerus_), the mountain sheep or bighorn (_Ovis +montana_), and the prairie-dog (_Cynomys_), one of the Rodentia, are +peculiar to this sub-region; while the family of the Saccomyidæ, or pouched +rats, is represented by many forms and is very characteristic. Here is also +the chief home of the bison. The glutton (_Gulo_) and marmot (_Lagomys_) +enter it from the north; while it has the racoon (_Procyon_), flying +squirrel (_Sciuropterus_), ground squirrel (_Tamias_), pouched marmot +(_Spermophilus_) and jumping mouse (_Jaculus_) in common with the countries +east or west of it. + +_Plate XIX. Illustrative of the Zoology of the Central Plains or +Prairies._--We here introduce four of the most characteristic mammalia of +the great American plains or prairies, three of them being types confined +to North America. The graceful animals on the left are the prong-horned +antelopes (_Antilocapra americana_), whose small horns, though hollow like +those of the antelopes, are shed annually like those of the deer. To the +right we have the prairie-dogs of the trappers (_Cynomys ludovicianus_) +which, as will be easily seen, are rodents, and allied to the marmots of +the European Alps. Their burrows are numerous on the prairies, and the +manner in which they perch {130}themselves on little mounds and gaze on +intruders, is noticed by all travellers. On the left, in the foreground, is +one of the extraordinary pouched rats of America (_Geomys bursarius_). +These are burrowing animals, feeding on roots; and the mouth is, as it +were, double, the outer portion very wide and hairy, behind which is the +small inner mouth. Its use may be to keep out the earth from the mouth +while the animal is gnawing roots. A mouth so constructed is found in no +other animals but in these North American rats. In the distance is a herd +of bisons (_Bison americanus_), the typical beast of the prairies. + +_Birds._--This sub-region has many peculiar forms of birds, both residents, +and migrants from the south or north. Among the peculiar resident species +we may probably reckon a dipper, (_Cinclus_); _Salpinctes_, one of the +wrens; _Poospiza_, _Calamospiza_, genera of finches; _Picicorvus_, +_Gymnokitta_, genera of the crow family; _Centrocercus_ and _Pediocætes_, +genera of grouse. As winter migrants from the north it has _Leucosticte_ +and _Plectrophanes_, genera of finches; _Perisoreus_, a genus of the crow +family; _Picoides_, the Arctic woodpecker; and _Lagopus_, ptarmigan. Its +summer migrants, many of which may be resident in the warmer districts, are +more numerous. Such are, _Oreoscoptes_, a genus of thrushes; +_Campylorhynchus_ and _Catherpes_, wrens; _Paroides_, one of the tits; +_Phænopepla_, allied to the waxwing; _Embernagra_ and _Spermophila_, genera +of finches; _Pyrocephalus_, one of the tyrant shrikes; _Callipepla_ and +_Cyrtonyx_, American partridges. Besides these, the more widely spread +genera, _Harporhynchus_, _Lophophanes_, _Carpodacus_, _Spizella_, and +_Cyanocitta_, are characteristic of the central district, and two genera of +humming-birds--_Atthis_ and _Selasphorus_--only occur here and in +California. Prof. Baird notes 40 genera of birds which are represented by +distinct allied species in the western, central, and eastern divisions of +the United States, corresponding to our sub-regions. + + + +Plate XIX. + +[Illustration] + +THE AMERICAN PRAIRIES, WITH CHARACTERISTIC MAMMALIA. + +{131}It is a curious fact that the birds of this sub-region should extend +across the Gulf of California, and that Cape St. Lucas, at the southern +extremity of the peninsula, should be decidedly more "Central" than +"Californian" in its ornithology. Prof. Baird says, that its fauna is +almost identical with that of the Gila River, and has hardly any relation +to that of Upper California. It possesses a considerable number (about +twenty) of peculiar species of birds, but all belong to genera +characteristic of the present sub-region; and there is no resemblance to +the birds of Mazatlan, just across the gulf in the Neotropical region. + +_Reptiles, Amphibia, and Fishes._--A large number of snakes and lizards +inhabit this sub-region, but they have not yet been classified with +sufficient precision to enable us to make much use of them. Among lizards, +Iguanidæ, Geckotidæ, Scincidæ, and Zonuridæ, appear to be numerous; and +many new genera of doubtful value have been described. Among snakes, +Calamariidæ, Colubridæ, and Crotalidæ are represented. Among Amphibia, +_Siredon_, one of the Proteidæ, is peculiar. The rivers and lakes of the +Great Central Basin, and the Colorado River, contain many peculiar forms of +Cyprinidæ. + + +_III. The Eastern or Alleghany Sub-region._ + +This sub-region contains examples of all that is most characteristic of +Nearctic zoology. It is for the most part an undulating or mountainous +forest-clad country, with a warm or temperate climate, but somewhat extreme +in character, and everywhere abounding in animal and vegetable life. To the +west, across the Mississippi, the country becomes more open, gradually +rises, becomes much drier, and at length merges into the arid plains of the +central sub-region. To the south, in Georgia, Florida, and Louisiana, a +sub-tropical climate prevails, and winter is almost unknown. To the north, +in Michigan and New England, the winters are very severe, and streams and +lakes are frozen for months together. These different climates, however, +produce little effect on the forms of animal life; the species to some +extent change as we go from north to south, but the same types everywhere +prevail. This portion of the United States, having been longest inhabited +by Europeans, has been more thoroughly explored than other parts of North +America; and to this more complete knowledge its superior zoological +richness {132}may be to some extent due; but there can be little doubt that +it is also positively, and not merely relatively, more productive in varied +forms of animal life than either of the other sub-regions. + +_Mammalia._--There seems to be only one genus absolutely peculiar to this +sub-region--the very remarkable _Condylura_, or star-nosed mole, only found +from Pennsylvania to Nova Scotia, and as far as about 94° west longitude. +It also has opossums (_Didelphys_) in common with California, and three out +of four species of _Scalops_, a genus of moles; as well as the skunk +(_Mephitis_), American badger (_Taxidea_), racoon (_Procyon_), pouched rat +(_Geomys_), beaver rat (_Fiber_), jumping mouse (_Jaculus_), tree porcupine +(_Erethizon_), and other characteristic Nearctic forms. + +_Birds._--The birds of this sub-region have been carefully studied by +American naturalists, and many interesting facts ascertained as to their +distribution and migrations. About 120 species of birds are peculiar to the +east coast of the United States, but only about 30 of these are residents +all the year round in any part of it; the bird population being essentially +a migratory one, coming from the north in winter and the south in summer. +The largest number of species seems to be congregated in the district of +the Alleghany mountains. A considerable proportion of the passerine birds +winter in Central America and the West Indian Islands, and go to the Middle +States or Canada to breed; so that even the luxuriant Southern States do +not possess many birds which may be called permanent residents. Thus, in +East Pennsylvania there are only 52, and in the district of Columbia 54 +species, found all the year round, out of about 130 which breed in these +localities; very much below the number which permanently reside in Great +Britain. + +This sub-region is well characterised by its almost exclusive possession of +_Ectopistes_, the celebrated passenger pigeon, whose enormous flocks and +breeding places have been so often described; and _Cupidonia_, a remarkable +genus of grouse. The only Nearctic parrot, _Conurus carolinensis_, is found +in the Southern States; as well as _Crotophaga_, a South American genus +usually associated with the cuckoos. _Helmintherus_ and {133}_Oporornis_, +genera of wood-warblers, may be considered to be peculiar to this +sub-region, since in each case only one of the two species migrates as far +as Central America; while two other genera of the same family, _Siurus_ and +_Setophaga_, as well as the finch genus, _Euspiza_, do not extend to either +of the western sub-regions. _Parus_, a genus of tits, comes into the +district from the north; _Otocorys_, an alpine lark, and _Coturniculus_, an +American finch, from the west; and such characteristic Nearctic genera as +_Antrostomus_ (the whip-poor-will goatsuckers); _Helminthophaga_, +_Dendræca_, and _Myiodioctes_ (wood-warblers); _Vireo_ (greenlets); +_Dolichonyx_ (rice-bird); _Quiscalus_ (troupial); _Meleagris_ (turkey); and +_Ortyx_ (American partridge), are wide-spread and abundant. In Mr. J. A. +Allen's elaborate and interesting paper on the birds of eastern North +America, he enumerates 32 species which breed only in the more temperate +portions of this province, and may therefore be considered to be especially +characteristic of it. These belong to the following genera:--_Turdus_, +_Galeoscoptes_, _Harporhynchus_, _Sialia_, _Dendræca_, _Wilsonia_, +_Pyranga_, _Vireo_, _Lanivireo_, _Lophophanes_, _Coturniculus_, +_Ammodromus_, _Spizella_, _Euspiza_, _Hedymeles_, _Cyanospiza_, _Pipilo_, +_Cardinalis_, _Icterus_, _Corvus_, _Centurus_, _Melanerpes_, _Antrostomus_, +_Coccyzus_, _Ortyx_, and _Cupidonia_. + +_Reptiles._--In this class the Eastern States are rich, possessing many +peculiar forms not found in other parts of the region. Among snakes it has +the genera _Farancia_ and _Dimodes_ belonging to the fresh-water snakes +(Homalopsidæ); the South American genus _Elaps_; and 3 genera of +rattlesnakes, _Cenchris_, _Crotalophorus_, and _Crotalus_. The following +genera of snakes are said to occur in the State of New York:--_Coluber_, +_Tropidonotus_, _Leptophis_, _Calamaria_, _Heterodon_, _Trigonocephalus_, +_Crotalus_, _Psammophis_, _Helicops_, _Rhinostoma_, _Pituophis_, and +_Elaps_. + +Among lizards, _Chirotes_, forming a peculiar family of Amphisbenians, +inhabits Missouri and Mexico; while the remarkable glass-snake, +_Ophisaurus_, belonging to the family Zonuridæ, is peculiar to the Southern +States; and the South American _Sphærodactylus_, one of the gecko family, +reaches Florida. Other genera which extend as far north as the State of New +{134}York are, _Scincus_, _Tropidolepis_, _Plestiodon_, _Lygosoma_, +_Ameiva_, and _Phrynosoma_. + +Tortoises, especially the fresh-water kind, are very abundant; and the +genera _Aromochelys_, _Chelydra_, _Terrapene_, and _Trionyx_, are nearly, +if not quite, confined to this division of the region. + +_Amphibia._--Almost all the remarkable forms of Urodela, or tailed +batrachians, peculiar to the region are found here only; such as _Siren_ +and _Pseudobranchus_, constituting the family Sirenidæ; _Menobranchus_, +allied to the _Proteus_ of Europe; _Amphiuma_, an eel-like creature with +four rudimentary feet, constituting the family Amphiumidæ; _Notopthalmus_, +_Desmognathus_, and _Menopoma_, belonging to the Salamandridæ; together +with several other genera of wider range. Of Anura, or tail-less +batrachians, there are no peculiar genera, but the Neotropical genus of +toads, _Engystoma_, extends as far as South Carolina. + +_Fishes._--Owing to its possession of the Mississippi and the great lakes, +almost all the peculiar forms of North American fishes are confined to this +sub-region. Such are _Perca_, _Pileoma_, _Huro_, _Bryttus_, and _Pomotis_ +(Percidæ); the families Aphredoderidæ and Percopsidæ; several genera of +Cyprinodontidæ and Cyprinidæ; and the family Polydontidæ. + + +_Islands of the Alleghany Sub-region._ + +_The Bermudas._--These islands, situated in the Atlantic, about 700 miles +from the coast of Carolina, are chiefly interesting for the proof they +afford of the power of a great variety of birds to cross so wide an extent +of ocean. There are only 6 or 8 species of birds which are permanent +residents on the islands, all common North American species; while no less +than 140 species have been recorded as visiting them. Most of these are +stragglers, many only noticed once; others appear frequently and in great +numbers, but very few, perhaps not a dozen, come every year, and can be +considered regular migrants. The permanent residents are, a greenlet +(_Vireo noveboracensis)_, the catbird (_Galeoscoptes carolinensis_), the +blue bird (_Sialia sialis_), the cardinal (_Cardinalis virginianus_), the +American crow (_Corvus {135}americanus_), and the ground dove (_Chamæpelia +passerina_). The most regular visitants are a kingfisher (_Ceryle alcyon_), +the wood-wagtail (_Siurus noveboracensis_), the rice-bird (_Dolichonyx +oryzivorus_), and a moorhen (_Gallinula galeata_). Besides the American +species, four European birds have been taken at the Bermudas: _Saxicola +oenanthe_, _Alauda arvensis_ (perhaps introduced), _Crex pratensis_, and +_Scolopax gallinago_. + +A common American lizard, _Plestiodon longirostris_, is the only land +reptile found on the islands. + + +_IV. The Sub-Arctic or Canadian Sub-region._ + +This sub-region serves to connect together the other three, since they all +merge gradually into it; while to the north it passes into the circumpolar +zone which is common to the Palæarctic and Nearctic regions. The greater +portion of it is an extensive forest-district, mostly of coniferæ; and +where these cease towards the north, barren wastes extend to the polar +ocean. It possesses several northern or arctic forms of Mammalia, such as +the glutton, lemming, reindeer, and elk, which barely enter the more +southern sub-regions; as well as the polar bear and arctic fox; but it also +has some peculiar forms, and many of the most characteristic Nearctic +types. The remarkable musk-sheep (_Ovibos_) is confined to this sub-region, +ranging over a considerable extent of country north of the forests, as well +as Greenland. It has been extinct in Europe and Asia since the +Post-pliocene epoch. Such purely Nearctic genera as _Procyon_, _Latax_, +_Erethizon_, _Jaculus_, _Fiber_, _Thomomys_, and _Hesperomys_, abound, many +of them ranging to the shores of Hudson's Bay and the barren wastes of +northern Labrador. Others, such as _Blarina_, _Condylura_, and _Mephitis_, +are found only in Nova Scotia and various parts of Canada. About 20 species +of Mammalia seem to be peculiar to this sub-region. + +_Plate XX. Illustrating the Zoology of Canada._--We have here a group of +Mammalia characteristic of Canada and the colder parts of the United +States. Conspicuous in the foreground is the skunk (_Mephitis mephitica_), +belonging to a genus of the weasel family found only in America. This +animal is {136}celebrated for its power of ejecting a terribly offensive +liquid, the odour of which is almost intolerable. The skunks are nocturnal +animals, and are generally marked, as in the species represented, with +conspicuous bands and patches of white. This enables them to be easily seen +at night, and thus serves to warn larger animals not to attack them. To the +left is the curious little jumping mouse (_Jaculus hudsonius_), the +American representative of the Palæarctic jerboa. Climbing up a tree on the +left is the tree porcupine (_Erethizon dorsatus_), belonging to the family +Cercolabidæ, which represents, on the American continent, the porcupines of +the Old World. In the background is the elk or moose (_Alces americanus_), +perhaps identical with the European elk, and the most striking inhabitant +of the northern forests of America, as the bison is of the prairies. + +_Birds._--Although the Canadian sub-region possesses very few resident +birds, the numbers which breed in it are perhaps greater than in the other +sub-regions, because a large number of circumpolar species are found here +exclusively. From a comparison of Mr. Allen's tables it appears, that more +than 200 species are regular migrants to Canada in the breeding season, and +nearly half of these are land-birds. Among them are to be found a +considerable number of genera of the American families Tyrannidæ and +Mniotiltidæ, as well as the American genera _Sialia_, _Progne_, _Vireo_, +_Cistothorus_, _Junco_, _Pipilo_, _Zonotrichia_, _Spizella_, _Melospiza_, +_Molothrus_, _Agelæus_, _Cyanura_, _Sphyrapicus_, and many others; so that +the ornithology of these northern regions is still mainly Nearctic in +character. Besides these, it has such specially northern forms as _Surnia_ +(Strigidæ); _Picoides_ (Picidæ); _Pinicola_ (Fringillidæ); as well as +_Leucosticte_, _Plectrophanes_, _Perisoreus_, and _Lagopus_, which extend +further south, especially in the middle sub-region. No less than 212 +species of birds have been collected in the new United States territory of +Alaska (formerly Russian America), where a humming-bird (_Selasphorus +rufus_) breeds. The great majority of these are typically American, +including such forms as _Colaptes_, _Helminthophaga_, _Siurus_, _Dendræca_, +_Myiodioctes_, _Passerculus_, _Zonotrichia_, _Junco_, _Spizella_, +_Melospizpa_, _Passerella_, _Scoleophagas_, _Pediocetes_, and _Bonasa_; +together with many northern birds common to both continents. Yet a few +Palæarctic forms, not known in other parts of the sub-region, appear here. +These are _Budytes flava_, _Phylloscopus kennicottii_, and _Pyrrhula +coccinea_, all belonging to genera not occurring elsewhere in North +America. Considering the proximity of the district to North-east Asia, and +the high probability that there was an actual land connection at, and south +of, Behring's Straits, in late Tertiary times, it is somewhat remarkable +that the admixture of Palæarctic and Nearctic groups is not greater than it +is. The Palæarctic element, however, forms so small apportion of the whole +fauna, that it may be satisfactorily accounted for by the establishment of +immigrants since the Glacial period. The great interest felt by +ornithologists in the discovery of the three genera above-named, with a +wren allied to a European species, is an indication that the faunas even of +the northern parts of the Nearctic and Palæarctic regions are, as regards +birds, radically distinct. It may be mentioned that the birds of the +Aleutian Isles are also, so far as known, almost wholly Nearctic. The +number of land-birds known from Alaska is 77; and from the Aleutian Isles +16 species, all of which, except one, are North American. + + + +Plate XX. + +[Illustration] + +A CANADIAN FOREST, WITH CHARACTERISTIC MAMMALIA. + +{137}_Reptiles._--These are comparatively few and unimportant. There are +however five snakes and three tortoises which are limited to Canada proper; +while further north there are only Amphibia, represented by frogs and +toads, and a salamander of the genus _Plethodon_. + +_Fishes._--Most of the groups of fresh-water fish of the Nearctic region +are represented here, especially those of the perch, salmon, and pike +families; but there seem to be few or no peculiar genera. + +_Insects._--These are far less numerous than in the more temperate +districts, but are still tolerably abundant. In Canada there are 53 species +of butterflies, viz., Papilionidæ, 4; Pieridæ, 2; Nymphalidæ, 21; Satyridæ, +3; Lycænidæ 16, and Hesperidæ 7. Most of these are, no doubt, found chiefly +in the southern parts of Canada. That Coleoptera are pretty numerous is +shown, by more than 800 species having been collected on the {138}shores of +Lake Superior; 177 being Geodephaga and 39 Longicorns. + +_Greenland._--This great arctic island must be considered as belonging to +the Nearctic region, since of its six land mammals, three are exclusively +American (_Myodes torquatus_, _Lepus glacialis_, and _Ovibos moschatus_), +while the other three (_Vulpes lagopus_, _Ursus maritimus_, and _Rangifer +tarandus_) are circumpolar. Only fourteen land-birds are either resident +in, or regular migrants to the country; and of these two are European +(_Haliæetus albicilla_, and _Falco peregrinus_), while three are American +(_Anthus ludovicianus_, _Zonotrichia leucophrys_, and _Lagopus rupestris_), +the rest being arctic species common to both continents. The waders and +aquatics (49 in number) are nearly equally divided between both continents; +but the land-birds which visit Greenland as stragglers are mostly American. +Yet although the Nearctic element somewhat preponderates, Greenland really +belongs to that circumpolar debateable land, which is common to the two +North Temperate regions. + + + +_Concluding remarks._--We have already discussed pretty fully, though +somewhat incidentally, the status and relations of the Nearctic region; +first in our chapter on Zoological regions, then in our review of extinct +faunas, and lastly in the earlier part of this chapter. It will not +therefore be necessary to go further into the question here; but we shall, +in our next chapter, give a brief summary of the general conclusions we +have reached as to the past history and mutual zoological relations of all +the great divisions of the earth. + + +{139}TABLES OF DISTRIBUTION. + +In drawing up these tables, showing the distribution of various classes of +animals in the Nearctic region, the following sources of information have +been chiefly relied on, in addition to the general treatises, monographs, +and catalogues used in the compilation of the 4th Part of this work. + +_Mammalia._--Professor Baird's Catalogue; Allen's List of the Bats; Mr. +Lord's List for British Columbia; Brown, for Greenland; Packard for +Labrador. + +_Birds._--Baird, Cassin, and Allen's Lists for United States; Richardson's +Fauna Boreali Americana; Jones, for Bermudas; and papers by Brown, Coues, +Lord, Packard, Dall, and Professor Newton. + + +{140}TABLE I. + +_FAMILIES OF ANIMALS INHABITING THE NEARCTIC REGION._ + +EXPLANATION. + + Names in _italics_ show the families which are peculiar to the region. + + Names inclosed thus (......) show families which barely enter the region, + and are not considered properly to belong to it. + + Numbers correspond to the series of numbers to the families in Part IV. + + ---------------------+-------------------+------------------------------- + | Sub-regions | + | 1=California. | + Order and Family | 2=Rocky Mntns. | Range beyond the Region. + | 3=Alleghanies. | + | 4=Canada. | + ---------------------+----+----+----+----+------------------------------- + | 1. | 2. | 3. | 4. | + ---------------------+----+----+----+----+------------------------------- + | | | | | + MAMMALIA. | | | | | + CHIROPTERA. | | | | | + 10. Phyllostomidæ | -- | | | | Neotropical + 12. Vespertilionidæ | -- | -- | -- | -- | Cosmopolite + 13. Noctilionidæ | -- | | | | Tropical regions + | | | | | + INSECTIVORA. | | | | | + 21. Talpidæ | -- | -- | -- | -- | Palæarctic + 22. Soricidæ | -- | -- | -- | -- | The Eastern Hemisphere, excl. + | | | | | Australia + | | | | | + CARNIVORA. | | | | | + 23. Felidæ | -- | -- | -- | -- | All regions but the Australian + 28. Canidæ | -- | -- | -- | -- | All regions but the Australian + 29. Mustelidæ | -- | -- | -- | -- | All regions but the Australian + 30. Procyonidæ | -- | -- | -- | -- | Neotropical + 32. Ursidæ | -- | -- | -- | -- | Palæarctic, Oriental + 33. Otariidæ | -- | | | -- | N. and S. temperate zones + 34. Trichechidæ | | | | -- | Arctic regions + 35. Phocidæ | -- | | | -- | N. and S. temperate zones + | | | | | + CETACEA. | | | | | + 36 to 41. | | | | | Oceanic + | | | | | + UNGULATA. | | | | | + 47. Suidæ | | | -- | | All other continents but + | | | | | Australia + 50. Cervidæ | -- | -- | -- | -- | All regions but Ethiopian and + | | | | | Australian + 52. Bovidæ | -- | -- | | -- |Palæarctic, Ethiopian, Oriental + | | | | | + RODENTIA. | | | | | + 55. Muridæ | -- | -- | -- | -- | Almost cosmopolite + 57. Dipodidæ | -- | -- | -- | -- | Palæarctic, Ethiopian + 59. _Saccomyidæ_ | -- | -- | -- | -- | Mexican sub-region + 60. Castoridæ | -- | -- | -- | -- | Palæarctic + 61. Sciuridæ | -- | -- | -- | -- | All regions but Australian + 62. _Haploodontidæ_ | -- | | | | + 66. Cercolabidæ | -- | -- | -- | -- | Neotropical + 69. Lagomyidæ | | -- | | -- | Palæarctic + 70. Leporidæ | -- | -- | -- | -- | All regions but Australian + | | | | | + MARSUPIALIA. | | | | | + 76. Didelphyidæ | -- | | -- | | Neotropical + | | | | | + BIRDS. | | | | | + PASSERES. | | | | | + 1. Turdidæ | -- | -- | -- | -- | Almost cosmopolite + 2. Sylviidæ | -- | -- | -- | -- | Almost cosmopolite + 5. Cinclidæ | | -- | | -- | Palæarctic, Oriental, Andes + 6. Troglodytidæ | -- | -- | -- | -- | All regions but Australian + 7. _Chamæidæ_ | -- | | | | + 8. Certhiidæ | -- | -- | -- | -- | Palæarctic, Oriental, + | | | | | Australian + 9. Sittidæ | -- | -- | -- | -- | Palæarctic, Oriental, + | | | | | Australian + 10. Paridæ | -- | -- | -- | -- | The Eastern Hemisphere + 19. Laniidæ | -- | -- | -- | -- | The Eastern Hemisphere + 20. Corvidæ | -- | -- | -- | -- | Cosmopolite + 26. (Coerebidæ) | | | -- | | Neotropical family + 27. Mniotiltidæ | -- | -- | -- | -- | Neotropical + 28. Vireonidæ | -- | -- | -- | -- | Neotropical + 29. Ampelidæ | -- | -- | -- | -- |Palæarctic, Antilles, Guatemala + 30. Hirundinidæ | -- | -- | -- | -- | Cosmopolite + 31. Icteridæ | -- | -- | -- | -- | Neotropical + 32. Tanagridæ | | -- | -- | | Neotropical + 33. Fringillidæ | -- | -- | -- | -- | All regions but Australian + 37. Alaudidæ | | -- | -- | -- | All regions but Neotropical + 38. Motacillidæ | -- | -- | -- | -- | Cosmopolite + 39. Tyrannidæ | -- | -- | -- | -- | Neotropical + | | | | | + PICARIÆ. | | | | | + 51. Picidæ | -- | -- | -- | -- | All regions but Australian + 58. Cuculidæ | -- | -- | -- | | Almost cosmopolite + 67. Alcedinidæ | -- | -- | -- | -- | Cosmopolite + 73. Caprimulgidæ | -- | -- | -- | -- | Cosmopolite + 74. Cypselidæ | -- | -- | -- | -- | Almost cosmopolite + 75. Trochilidæ | -- | -- | -- | -- | Neotropical + | | | | | + PSITTACI. | | | | | + 80. Conuridæ | | | -- | | Neotropical + | | | | | + COLUMBÆ. | | | | | + 84. Columbidæ | -- | -- | -- | -- | Cosmopolite + | | | | | + GALLINÆ. | | | | | + 87. Tetraonidæ | -- | -- | -- | -- | Almost cosmopolite + 88. Phasianidæ | | -- | -- | | Palæarctic, Oriental, + | | | | | Ethiopian, Honduras + 91. (Cracidæ) | | -- | | | Neotropical + | | | | | + ACCIPITRES. | | | | | + 94. Vulturidæ | -- | -- | -- | | All regions but Australian + 96. Falconidæ | -- | -- | -- | -- | Cosmopolite + 97. Pandionidæ | -- | -- | -- | -- | Cosmopolite + 98. Strigidæ | -- | -- | -- | -- | Cosmopolite + | | | | | + GRALLÆ. | | | | | + 99. Rallidæ | -- | -- | -- | -- | Cosmopolite + 100. Scolopacidæ | -- | -- | -- | -- | Cosmopolite + 105. Charadriidæ | -- | -- | -- | -- | Cosmopolite + 107. Gruidæ | -- | -- | -- | | All regions but Neotropical + 113. Ardeidæ | -- | -- | -- | -- | Cosmopolite + 114. Plataleidæ | -- | -- | -- | -- | Almost cosmopolite + 115. Ciconiidæ | | | -- | | All the regions + | | | | | + ANSERES. | | | | | + 118. Anatidæ | -- | -- | -- | -- | Cosmopolite + 119. Laridæ | -- | -- | -- | -- | Cosmopolite + 120. Procellariidæ | -- | -- | -- | -- | Cosmopolite + 121. Pelecanidæ | -- | -- | -- | -- | Cosmopolite + 123. Colymbidæ | | | | -- | North temperate and arctic + | | | | | zones + 124. Podicipidæ | -- | -- | -- | -- | Cosmopolite + 125. Alcidæ | -- | | | -- | North temperate and arctic + | | | | | zones + | | | | | + REPTILIA. | | | | | + OPHIDIA. | | | | | + 5. Calamariidæ | -- | -- | -- | | All the regions + 6. Oligodontidæ | | | -- | | Neotropical, Oriental, Japan + 7. Colubridæ | -- | -- | -- | -- | Almost cosmopolite + 8. Homalopsidæ | | | -- | | All the regions + 17. Pythonidæ | -- | | | | All tropical regions + 20. Elapidæ | | | -- | | All tropical regions, Japan + 24. Crotalidæ | -- | -- | -- | -- | Neotropical, Palæarctic, + | | | | | Oriental + | | | | | + LACERTILIA. | | | | | + 27. _Chirotidæ_ | | -- | -- | | Mexico + 32. Teidæ | -- | -- | -- | | Neotropical + 34. Zonuridæ | -- | -- | -- | -- | All regions but Australian + 35. Chalcidæ | | | -- | | Neotropical + 45. Scincidæ | -- | -- | -- | | Almost cosmopolite + 49. Geckotidæ | -- | -- | -- | | Almost cosmopolite + 50. Iguanidæ | -- | -- | -- | | Neotropical + | | | | | + CROCODILIA. | | | | | + 56. Alligatoridæ | | | -- | | Neotropical + | | | | | + CHELONIA. | | | | | + 57. Testudinidæ | -- | -- | -- | -- | All continents but Australian + 59. Trionychidæ | | | -- | | Ethiopian, Oriental, Japan + 60. Cheloniidæ | | | | | Marine + | | | | | + AMPHIBIA. | | | | | + URODELA. | | | | | + 2. _Sirenidæ_ | | | -- | | + 3. Proteidæ | | | -- | | Palæarctic + 4. _Amphiumidæ_ | | | -- | | + 5. Menopomidæ | | | -- | | Palæarctic + 6. Salamandridæ | -- | -- | -- | -- | Andes, Palæarctic + | | | | | + ANOURA. | | | | | + 10. Bufonidæ | -- | -- | -- | -- | All continents but Australia + 12. Engystomidæ | | | -- | | All regions but Nearctic + 15. Alytidæ | -- | -- | -- | | All regions but Oriental + 17. Hylidæ | -- | -- | -- | -- | All regions but Ethiopian + 18. Polypedatidæ | | | -- | | All the regions + 19. Ranidæ | -- | -- | -- | -- | Almost cosmopolite + | | | | | + FISHES (FRESHWATER). | | | | | + ACANTHOPTERYGII. | | | | | + 1. Gasterosteidæ | -- | -- | -- | -- | Palæarctic + 3. Percidæ | -- | -- | -- | -- | Cosmopolite + 4. _Aphredoderidæ_ | | | -- | | + 12. Scienidæ | -- | -- | -- | -- | All regions but Australian + 37. Atherinidæ | -- | -- | -- | -- | Palæarctic + | | | | | + PHYSOSTOMI. | | | | | + 59. Siluridæ | -- | -- | -- | -- | All warm regions + 65. Salmonidæ | -- | -- | -- | -- | Palæarctic, New Zealand + 66. _Percopsidæ_ | | | | -- | + 70. Esocidæ | -- | -- | -- | -- | Palæarctic + 71. Umbridæ | -- | -- | -- | -- | Palæarctic + 73. Cyprinodontidæ | -- | -- | -- | | All regions but Australian + 74. _Heteropygii_ | | | -- | | + 75. Cyprinidæ | -- | -- | -- | -- | Not in S. America or Australia + 77. _Hyodontidæ_ | -- | -- | -- | -- | + | | | | | + GANOIDEI. | | | | | + 93. _Amiidæ_ | -- | -- | -- | -- | + 95. _Lepidosteidæ_ | -- | -- | -- | -- | + 96. Accipenseridæ | -- | -- | -- | -- | Palæarctic + 97. Polydontidæ | | | -- | | Palæarctic + | | | | | + INSECTS. | | | | | + LEPIDOPTERA (PART) | | | | | + DIURNI (BUTTERFLIES).| | | | | + 1. Danaidæ | -- | -- | -- | -- | All warm regions + 2. Satyridæ | -- | -- | -- | -- | Cosmopolite + 7. (Heliconidæ) | | | -- | | Neotropical + 8. Nymphalidæ | -- | -- | -- | -- | Cosmopolite + 9. Libytheidæ | | -- | -- | | Not in Australia + 12. Erycinidæ | -- | -- | -- | | Neotropical + 13. Lycænidæ | -- | -- | -- | -- | Cosmopolite + 14. Pieridæ | -- | -- | -- | -- | Cosmopolite + 15. Papilionidæ | -- | -- | -- | -- | Cosmopolite + 16. Hesperidæ | -- | -- | -- | -- | Cosmopolite + | | | | | + SPHINGIDEA. | | | | | + 17. Zygænidæ | -- | -- | -- | -- | Cosmopolite + 18. Castniidæ | | | -- | | Neotropical, Australian + 22. Ægeriidæ | -- | -- | -- | -- | Not in Australia + 23. Sphingidæ | -- | -- | -- | -- | Cosmopolite + +{145}TABLE II. + +_LIST OF GENERA OF TERRESTRIAL MAMMALIA AND BIRDS INHABITING THE NEARCTIC +REGION._ + +EXPLANATION. + + Names in _italics_ show genera peculiar to the region. + + Names enclosed thus (...) indicate genera which barely enter the region, + and are not considered properly to belong to it. + + Genera properly belonging to the region are numbered consecutively. + + +_MAMMALIA._ + + -------------------+-------+----------------------+---------------------- + Order, Family, and | No. of| Range within | Range beyond + Genus. |Species| the Region. | the Region. + -------------------+-------+----------------------+---------------------- + | | | + CHIROPTERA. | | | + PHYLLOSTOMIDÆ. | | | + 1. Macrotus | 1 | California Mexico, | + | | Antilles | + | | | + VESPERTILIONIDÆ. | | | + 2. Scotophilus | 5 | Universal, to Hudson's| Neotr., Orient., + | | Bay | Austral. + 3. Vespertilio | 6 | Universal, to Hudson's| Cosmopolite. + | | Bay | + 4. Nycticejus | 1 | South and East | India, Tropical + | | | Africa, temperate + | | | S. America + 5. Lasiurus | 3 | Temp. N. Amer. to | Tropical America + | | Nova Scotia | + 6. _Synotus_ | 2 |S. E. and Central States + 7. _Autrozous_ | 1 | W. Coast | + | | | + NOCTILIONIDÆ. | | | + 8. Nyctinomus | 1 | Cal. and S. Central |Neotropical, Oriental, + | | Sub-region | S. Palæarctic + | | | + INSECTIVORA. | | | + TALPIDÆ. | | | + 9. _Condylura_ | 1 | Eastern N. America | + 10. _Scapanus_ | 2 | New York to San | + | | Francisco | + 11. _Scalops_ | 3 | S. of Great Lakes & | + | | Brit. Columb. | + 12. Urotrichus | 1 | British Columbia | Japan + | | | + SORICIDÆ. | | | + 13. Sorex | 16 | The whole region | Palæarc., Ethiop., + | | | Orien. + 14. Neosorex | 1 | Vancouver's Island | + | | (a sub-genus) | + 15. Blarina | 7 | Canada to Mexico | + | | (a sub-genus) | + | | | + CARNIVORA. | | | + FELIDÆ. | | | + 16. Felis | 5 | S. of 55° N. Latitude | All regs. but + | | | Australian + 17. Lynx | 3 | S. of 56° N. Latitude | Palæarctic + | | | + CANIDÆ. | | | + 18. Lupus | 6 | All N. America | Palæarctic, Oriental + 19. Vulpes | 6 | N. America to Arctic | Palæarc., Ethiop., + | | Ocean and Greenland | Orient. + | | | + MUSTELIDÆ. | | | + 20. Martes | 2 | Pennsylvania to | Palæarctic, Oriental + | | Paget's Sound | + 21. Mustela | 11 | All N. America | Peru, Palæarctic, + | | | Ethiopian, Oriental + 22. Gulo | 1 | Rocky Mountains and | N. Palæarctic + | | Canada | + 23. _Latax_ | 2 | United States and | + | | Canada | + 24. Enhydris | 1 | Pacific coast |W. coast of S. America + 25. _Taxidea_ | 2 |Arkansas to 58° N. Lat.| + 26. Mephitis | 6 | United States and | Neotropical + | | Canada | + | | | + PROCYONIDÆ. | | | + 27. Procyon | 2 | Texas to Canada, | Neotropical + | | California | + 28. Bassaris | 1 | California and Texas | Guatemala and Mexico + | | | + URSIDÆ. | | | + 29. Ursus | 3 | N. America and | Palæarctic, Oriental + | | Greenland | + | | | + OTARIIDÆ. | | | + 30. Callorhinus | 1 | Behring's Straits | Kamschatka + 31. Zalophus | 1 | S. California to | Japan + | | N. Pacific | + _Eumetopias_ | 1 | California to | + | | Behring's Straits | + | | | + TRICHECHIDÆ. | | | + 32. Trichechus | 1 | Arctic Ocean to 66° N.| Palæarctic + | | Lat. in N. America | + PHOCIDÆ. | | | + 33. Callocephalus | 1 | Greenland | Palæarctic + 34. Pagomys | 1 | N. Atlantic and | Japan + | | N. Pacific | + 35. Pagophilus | 1 | N. Atlantic and | Palæarctic + | | N. Pacific | + 36. _Halicyon_ | 1 | N. W. coast of America| + 37. Phoca | 1 | Northern Coast | Palæarctic + 38. Halichoerus | 1 | Greenland | Palæarctic + 39. Morunga | 1 | California | S. temperate shores + 40. Cystophora | | Greenland | N. Atlantic + | | | + UNGULATA. | | | + SUIDÆ. | | | + 41. Dicotyles | 1 | Texas to Red River, | Neotropical + | | Arkansas | + | | | + CERVIDÆ. | | | + 42. Alces | 1 | N. E. United States & | N. Palæarctic + | | Canada | + 43. Rangifer | 2 | Maine to Arctic Ocean | Arctic zone + | | & Greenl. | + 44. Cervus | 6 | N. America to 57° N. | Neotr., Palæarc., + | | Lat. | Orien. + | | | + BOVIDÆ. | | | + 45. Bison | 1 | Between Missouri & | E. Europe + | | Rocky Mtns. | + 46. _Antilocapra_ | 1 | Central plains from | + | | Rio Grande to | + | | British Columbia | + 47. _Aplocerus_ | 1 | Northern Rocky | + | | Mountains | + 48. Capra | 1 | Upper Missouri and | Palæarctic + | | Rocky Mountains | + | | northwards | + 49. _Ovibos_ | 1 | Arctic America and | + | | Greenland | + | | | + RODENTIA. | | | + MURIDÆ. | | | + 50. Reithrodon | 5 | N. America to Lat. | Neotropical + | | 39° N. | + 51. Hesperomys | 16 | Temperate N. America | Neotropical + 52. _Neotoma_ | 7 | Temperate N. America | + 53. _Sigmodon_ | 2 | S. and S. E. States | + 54. Arvicola | 27 | Texas and California | Palæarctic + | | to Hudson's Bay | + 55. Myodes | 3 | N. United States to | N. Palæarctic + | | Arctic Reg. and | + | | Greenland | + 56. _Fiber_ | 1 | All N. America | Mexico + | | | + DIPODIDÆ. | | | + 57. _Jaculus_ | 1 | Pennsylvania to Canada| + | | and California | + | | | + SACCOMYIDÆ. | | | + 58. _Dipodomys_ | 5 | New Mexico to Columbia| + | | River and Carolina | + 59. _Perognathus_ | 6 | New Mexico to British | + | | Columbia | + 60. _Thomomys_ | 2 | Upper Missouri to | + | | Hudson's Bay | + 61. _Geomys_ | 5 | New Mexico to Alabama | + | | and Nebraska | + 62. _Saccomys_ | 1 | N. America | + | | | + CASTORIDÆ. | | | + 63. Castor | 1 | N. Mexico to Labrador | Palæarctic + | | | + SCIURIDÆ | | | + 64. Sciurus | 18 | N. America to Labrador| All regs. but + | | | Australian + 65. Sciuropterus | 4 | California & E. States| Palæarctic, Oriental + | | northwds. | + 66. Tamias | 4 | Mexico and Virginia | Mexico, N. Asia + | | to Canada | + 67. Spermophilus | 15 | N., W., & Central | Palæarctic + | | N. America | + 68. _Cynomys_ | 2 | Rio Grande to Missouri| + | | (Central) | + 69. Arctomys | 4 | Virginia and Nebraska,| N. Palæarctic + | | northws. | + | | | + HAPLOODONTIDÆ. | | | + 70. _Haploodon_ | 2 | California and British| + | | Columbia | + | | | + CERCOLABIDÆ | | | + 71. _Erethizon_ | 2 |Pennsylvania to Canada,| + | | & Pacific coast | + LAGOMYIDÆ. | | | + 72. Lagomys | 1 | Rocky Mountains, 42° | Palæarctic + | | to 60° N. Lat. | + | | | + LEPORIDÆ. | | | + 73. Lepus | 15 | All N. America to | All regs. but + | | Greenland | Australian + | | | + MARSUPIALIA. | | | + DIDELPHYIDÆ. | | | + 74. Didelphys | 2 | From Hudson's River & | Neotropical + | | Lower California, | + | | southward | + + _BIRDS._ + + PASSERES. | | | + TURDIDÆ. | | | + 1. Turdus | 9 | The whole region | Almost cosmopolite + 2. Mimus | 2 | All U. States and to | Neotropical + | | Canada | + 3. Galeoscoptes | 1 | E. of N. America | To Panama + 4. _Oreoscoptes_ | 1 | California and Rocky | Mexico + | | Mountains | + 5. _Harporhynchus_| 7 | N. America, chiefly | Mexico + | | the west | + | | | + SYLVIIDÆ. | | | + 6. Myiadestes | 1 | W. of Rocky Mountains | Neotropical + | | and to Canada | + 7. _Sialia_ | 3 | All United States and | Mexico and Guatemala + | | to Canada | + 8. Regulus | 3 | All United States & | Palæarc., Cent. + | | to Labrador | America + 9. Polioptila | 3 | Central and Southern | Neotropical + | | U. States | + | | | + CINCLIDÆ. | | | + 10. Cinclus | 1 | Rocky Mountains and | Andes, Palæarctic + | | British America | + | | | + TROGLODYTIDÆ. | | | + 11. Troglodytes | 3 | N. America | Neotropical, + | | | Palæarctic + 12. Thryophilus | 1 | N. W. America | Neotropical + 13. Thryothorus | 3 | All N. America | Neotropical + 14. Cistothorus | 2 | N. America | Neotropical + (Campylorhynchus| 1 | Gila and Rio Grande) | Neotropical genus + 15. _Salpinctes_ | 1 | Rocky Mountains to | + | | Oregon | + 16. _Catherpes_ | 1 | Gila and Colorado | + | | | + CHAMÆIDÆ. | | | + 17. _Chamæa_ | 1 | California | + | | | + CERTHIIDÆ. | | | + 18. Certhia | 2 | All United States and | Palæarctic, Guatemala + | | Canada | + | | | + SITTIDÆ. | | | + 19. Sitta | 5 | All United States and | Palæarctic, Mexico + | | Canada | + | | | + PARIDÆ. | | | + 20. Parus | 8 | All United States and | Palæarc., Orien., + | | Canada | Mexico + 21. Lophophanes | 4 | All United States | Palæarctic, Mexico + 22. _Psaltriparus_ | 3 | Central & Western | Mexico and Guatemala + | | N. America | + 23. _Auriparus_ | 1 | Rio Grande Valley | + | | | + LANIIDÆ. | | | + 24. Lanius | 4 | All N. America | Palæarc., Ethio., + | | | Orient. + | | | + CORVIDÆ. | | | + 25. Perisoreus | 1 | Canada and Rocky | Palæarctic + | | Mountains | + 26. Cyanocitta | 9 | All United States and | Neotropical + | | to Canada | + 27. _Gymnokitta_ | 1 | Central and N. W. | + | | States | + 28. _Picicorvus_ | 1 | Central and Western | + | | States to Sitka | + 29. Pica | 2 | Central and Western | Palæarctic + | | States to Arctic | + | | Ocean | + 30. Corvus | 7 | All N. America | Cosmop., excl. + | | | S. Amer. + | | | + COEREBIDÆ. | | | + (Certhiola | 1 | Florida; summer | Neotropical genus + | | migrant) | + | | | + MNIOTILTIDÆ. | | | + 31. _Mniotilta_ | 1 | Eastern States | Antilles, Andes of + | | | Columbia (migrant) + 32. Parula | 1 | Eastern States and | Neotropical + | | Canada | + 33. Protonotaria | 1 | Ohio and southwards | Neotrop. to Venezuela + 34._Helminthophaga_| 8 | All N. America | Mexico to Columbia + 35. _Helmintherus_ | 2 | S. and E. States to | Mexico to Veragua + | | Canada | + 36. Perissoglossa | 1 | Eastern United States | Antilles + 37. Dendroeca | 22 | All N. America | Mex. to Ecuador & + | | | Chili + 38. _Oporornis_ | 2 | Eastern States | Guatemala and Panama + 39. Geothlypis | 4 | All N. America | Neotropical + 40. Setophaga | 2 | E. States & Canadian | Neotropical + | | sub-region | + 41. _Myiodioctes_ | 5 | United States and | Mex. to Columb. + | | Canada | (migr.) + 42. Siurus | 3 | S. and E. States to | Mexico to Columbia + | | Canada | + 43. _Icteria_ | 2 | E. and Central States | Mexico to Costa Rica + | | to Canada | + | | | + VIREONIDÆ. | | | + 44. Vireosylvia | 7 | All N. America |Antilles and Venezuela + 45. Vireo | 6 | All United States | Antilles and Costa + | | | Rica + | | | + AMPELIDÆ. | | | + 46. Ampelis | 2 | All N. America | Palæarctic, Guatemala + 47. _Phænopepla_ | 1 |Gila and Lower Colorado| Mexico + | | | + HIRUNDINIDÆ. | | | + 48. Hirundo | 3 | All N. America | Almost cosmopolite + 49. Petrochelidon | 1 | All N. America | Neotropical + 50. Cotyle | 1 | All N. America | All regs. but + | | | Australian + 51. Stelgidopteryx | 1 | Southern States | Neotropical + 52. Progne | 1 | All N. America | Neotropical + | | | + ICTERIDÆ. | | | + 53. Icterus | 7 | All United States and | Neotropical + | | Canada | + 54. Dolichonyx | 1 | Eastern States and | Neotropical + | | Canada | + 55. Molothrus | 1 | All United States and | Neotropical + | | Canada | + 56. Agelæus | 3 | All United Slates and | Neotropical + | | Canada | + 57._Xanthocephalus_| 1 | The whole region | Mexico + 58. Sturnella | 2 | All United States and | Neotropical + | | Canada | + 59. _Scolecophagus_| 2 | All United States and | Mexico + | | Canada | + 60. Quiscalus | 4 | S. and E. States to | Mexico to Venezuela + | | Labrador | + | | | + TANAGRIDÆ. | | | + 61. Pyranga | 4 | United Stales and | Neotropical + | | Canada | + | | | + FRINGILLIDÆ. | | | + 62. Chrysomitris | 7 | The whole region | Neotropical, + | | | Palæarctic + 63. Coccothraustes | 1 | W. and N. W. America | Palæarctic, Guatemala + 64. Embernagra | 1 |Rocky Mountain district| Neotropical + 65. _Pipilo_ | 9 | All N. America | Mexico and Guatemala + 66. _Junco_ | 5 | All United States | Mexico and Guatemala + 67. Zonotrichia | 5 | The whole region | Neotropical + 68. _Melospiza_ | 7 | All United States to | Mexico and Guatemala + | | Sitka | + 69. _Spizella_ | 6 | N. America | Mexico and Guatemala + 70. _Passerella_ | 3 | The whole region | Northern Asia + 71. _Passerculus_ | 6 | The whole region | Mexico and Guatemala + 72. _Pooecetes_ | 1 | All United States | Mexico + 73. _Ammodromus_ | 3 | All United States | Mexico and Guatemala + 74. Coturniculus | 3 |E. and N. of N. America| Neotropical + 75. Peucæa | 3 | S. Atlantic States and| Mexico + | | California | + 76. _Cyanospiza_ | 5 | All United States to | Central American + | | Canada | + 77. Poospiza | 2 | California and S. | Neotropical + | | Central States | + 78. Carpodacus | 5 | The whole region | Mexico, Palæarctic + 79. Cardinalis | 1 | S. and S. Central | Mexico to Venezuela + | | States | + 80. _Pyrrhuloxia_ | 1 | Texas and Rio Grande | + 81. Guiraca | 1 | Southern States | Neotropical + 82. Hedymeles | 2 | All United States | Mexico to Columbia + (Spermophila | 1 | Texas) | Neotropical genus + 83. Loxia | 2 | N. of Pennsylvania | Palæarctic + 84. Pinicola | 1 | Boreal America | Palæarctic + 85. Linota | 2 | E. and N. of N. | Palæarctic + | | America | + 86. Leucosticte | 4 | Alaska to Utah | Palæarctic + 87. _Calamospiza_ | 1 | Arizona and Texas to | Mexico + | | Mexico | + 88. _Chondestes_ | 1 | Western, Cen., & | Mexico + | | Southern States | + 89. Euspiza | 2 | S. Eastern States | Palæarc., Columb. + | | | (mig.) + 90. Plectrophanes | 6 | Boreal America and E. | Palæarctic + | | side of Rocky | + | | Mountains | + 91. _Centronyx_ | 1 | Mouth of Yellowstone | + | | River | + | | | + ALAUDIDÆ. | | | + 92. Otocorys | 1 | High central plains to| Palæarc., Mexico, + | | E. States and Canada| Andes of Columbia + | | | + MOTACILLIDÆ. | | | + 93. Anthus | 1 | The whole region | Cosmopolite + 94. _Neocorys_ | 1 | Nebraska | + | | | + TYRANNIDÆ. | | | + 95. Sayornis | 3 | E. States to Canada, | Mexico to Ecuador + | | California | + (Pyrocephalus | 1 | Gila and Rio Grande) | Neotropical + 96. Empidonax | 7 | The whole region | Mexico to Ecuador + 97. Contopus | 3 | N. and E. of Rocky | Mexico to Amazonia + | | Mountains | + 98. Myiarchus | 2 | E. and W. coasts and | Neotropical + | | Canada | + 99. _Empidias_ | 1 | Eastern States | Mexico + 100. Tyrannus | 4 | All United States to | Neotropical + | | Canada | + (Milvulus | 1 | Texas) | Neotropical genus + | | | + PICARIÆ. | | | + PICIDÆ. | | | + 101. Picoides | 3 | Arctic zone and Rocky | Palæarctic + | | Mounts. | + 102. Picus | 6 | All United States and | All regs. but Eth. + | | Canada | & Aus. + 103. _Sphyrapicus_ | 6 | Brit. Columbia and | Mexico and Guatemala + | | Pennsylvania | + | | southwards | + 104. Campephilus | 2 | United States and | Neotropical + | | Canada | + 105. _Hylatomus_ | 1 | E. and W. States and | + | | Canada | + 106. Centurus | 3 | The whole region | Mexico to Venezuela + 107. Melanerpes | 3 | United States and | Neotropical + | | S. Canada | + 108. Colaptes | 3 | United States and | Neotropical + | | Canada | + | | | + CUCULIDÆ. | | | + 109. Crotophaga | 2 | E. States from | Neotropical + | | Pennsylvania S. | + 110. Coccyzus | 3 | S. E. and Cen. States | Neotropical + | | to Canada | + 111. Geococcyx | 1 | California to New Mex.| Guatemala + | | & Texas | + | | | + ALCEDINIDÆ. | | | + 112. Ceryle | 2 | The whole region, | Neotropical S. + | | | Palæarctic, Oriental + | | | + CAPRIMULGIDÆ. | | | + 113. Chordeiles | 3 | All United States to | Neotropical + | | Canada | + 114. Antrostomus | 3 | All United States to | Neotropical + | | Canada | + | | | + CYPSELIDÆ. | | | + 115. Nephoecetes | 1 | N. W. America Jamaica | + 116. Chætura | 2 | All U. States & | Almost cosmopolite + | | British Columbia | + | | | + TROCHILIDÆ. | | | + 117. _Trochilus_ | 2 | The whole region | Mexico to Veragua + | | | (? mi.) + 118. Selasphorus | 2 | W. coast and Centre | Mexico to Veragua + 119. _Atthis_ | 2 | California and | Mexico to Guatemala + | | Colorado Valley | + | | | + PSITTACI. | | | + CONURIDÆ. | | | + 120. Conurus | 1 | S. and S. E. States | Neotropical + | | | + COLUMBÆ. | | | + COLUMBIDÆ. | | | + 121. Columba | 3 | W. and Central States | All regs. but + | | to Canada | Australian + 122. _Ectopistes_ | 1 | E. coast to Cen. | + | | plains, Canada and | + | | British Columbia | + 123. Melopelia | 1 | W. and S. Central | Neotropical + | | States | + 124. Zenaidura | 1 | All United States to | Mexico to Veragua + | | Canada | + 125. Chæmepelia | 1 | California and S. E. | Neotropical + | | States | + | | | + GALLINÆ. | | | + TETRAONIDÆ. | | | + 126. Cyrotonyx | 1 | S. Central States | Mexico and Guatemala + 127. Ortyx | 5 | All United States and | Mexico to Honduras + | | to Canada | and Costa Rica + 128. Callipepla | 1 | California | Mexico + 129. _Lophortyx_ | 2 | Arizona and California| + 130. _Oreortyx_ | 1 | California and Oregon | + 131. Tetrao | 3 | N. and N. W. America | Palæarctic + 132. _Centrocercus_ | 1 | Rocky Mountains | + 133. _Pediocætes_ | 2 | N. and N. W. America | + 134. _Cupidonia_ | 1 | E. & N. Cen. States | + | | and Canada | + 135. Bonasa | 1 | N. United States and | Palæarctic + | | Canada | + 136. Lagopus | 4 | Arctic zone and to 39°| Palæarctic + | | N. Lat. in Rocky | + | | Mountains | + | | | + PHASIANIDÆ. | | | + 137. _Meleagris_ | 2 | E. and Central States | Mexico, Honduras + | | to Canada | + CRACIDÆ. | | | + (Ortalida | 1 | New Mexico) | Neotropical genus + | | | + ACCIPITRES. | | | + VULTURIDÆ. | | | + _Sub-Family_ | | | + (CATHARTINÆ.) | | | + 138. Catharista | 1 | United States to | Neotropical + | | 40° N. Lat. | + 139. Psuedogryphis | 2 | United States to | Neotropical + | | 49° N. Lat. | + | | | + FALCONIDÆ. | | | + 140. Polyborus | 1 | S. States to Florida | Neotropical + | | & California | + 141. Circus | 1 | All N. America | Nearly cosmopolite + 142. Antenor | 2 | California and Texas | Neotropical + 143. Astur | 1 | All N. America | Almost cosmopolite + 144. Accipiter | 3 | All temperate | Almost cosmopolite + | | N. America | + 145. Tachytriorchis | 1 | New Mexico to | Neotropical + | | California | + 146. Buteo | 12 | All N. America | All regs. but + | | | Australian + 147. Archibuteo | 3 | All N. America N. | Palæarctic + 148. Asturina | 1 | S. E. States | Neotropical + 149. Aquila | 1 | The whole region | Palæarc., Ethiop., + | | | Indian + 150. Haliæetus | 2 | All N. America | All regs. but + | | | Neotropical + 151. Nauclerus | 1 | E. coast to | Neotropical + | | Pennsylvania and | + | | Wisconsin | + (Rostrhamus | 1 | Florida) | Neotropical + 152. Elanus | 1 | Southern and Western | Tropical regions + | | States | + 153. Ictinia | 1 | Southern States | Neotropical + 154. Falco | 7 | The whole region | Almost cosmopolite + 155. Hierofalco | 2 | N. of N. America | N. Palæarctic + 156. Cerchneis | 1 | All N. America | Almost cosmopolite + | | | + PANDIONIDÆ. | | | + 157. Pandion | 1 | Temperate N. | America Cosmopolite + | | | + STRIGIDÆ. | | | + 158. Surnia | 1 | Arctic & N. Temperate | N. Palæarctic + | | America | + 159. Nyctea | 1 | S. Carolina to | N. Palæarctic + | | Greenland | + 160. Glaucidium | 1 | Oregon and California | Neotropical, + | | | Palæarctic + 161. _Micrathene_ | 1 | Arizona and New Mexico| Mexico + 162. Pholeoptynx | 1 | N. W. America, Texas | Neotropical + 163. Bubo | 1 | All N. America | All regs. but + | | | Australian + 164. Scops | 2 | The whole region | Almost cosmopolite + 165. Syrnium | 2 | E. States, California,| All regs. but + | | Canada | Australian + 166. Asio | 2 | The whole region | All regs. but + | | | Australian + 167. Nyctale | 3 | All N. America | Palæarctic + 168. Strix | 1 | Temperate N. America | Almost cosmopolite + + _Peculiar or very Characteristic Genera of Wading and Swimming Birds._ + + GRALLÆ | | | + SCOLOPACIDÆ. | | | + Micropelma | 1 | N. America | Andes to Chili + _Philohela_ | 1 | Eastern States to | + | | Canada | + | | | + CHARADRIIDÆ. | | | + Aphriza | 1 | W. coast of America | West of S. America + | | | + ANSERES. | | | + ANATIDÆ. | | | + Aix | 1 | N. America | China + Bucephala | 4 | N. America | Europe + Oedemia | 3 | N. America | Europe + Harelda | 1 | Arctic | Arctic Seas + Somateria | 5 | Arctic | North Palæarctic + _Camptolæmus_ | 1 | N. E. America | + | | (? extinct) | + | | | + LARIDÆ. | | | + _Creagrus_ | 1 | California and | + | | N. Pacific coasts | + +{154}CHAPTER XVI. + +SUMMARY OF THE PAST CHANGES AND GENERAL RELATIONS OF THE SEVERAL REGIONS. + + +Having now closed our survey of the animal life of the whole earth--a +survey which has necessarily been encumbered with a multiplicity of +detail--we proceed to summarize the general conclusions at which we have +arrived, with regard to the past history and mutual relations of the great +regions into which we have divided the land surface of the globe. + +All the palæontological, no less than the geological and physical evidence, +at present available, points to the great land masses of the Northern +Hemisphere as being of immense antiquity, and as the area in which the +higher forms of life were developed. In going back through the long series +of the Tertiary formations, in Europe, Asia, and North America, we find a +continuous succession of vertebrate forms, including all the highest types +now existing or that have existed on the earth. These extinct animals +comprise ancestors or forerunners of all the chief forms now living in the +Northern Hemisphere; and as we go back farther and farther into the past, +we meet with ancestral forms of those types also, which are now either +confined to, or specially characteristic of, the land masses of the +Southern Hemisphere. Not only do we find that elephants, and rhinoceroses, +and hippopotami, were once far more abundant in Europe than they are now in +the tropics, but we also find that the apes of West Africa and Malaya, the +lemurs of Madagascar, the Edentata of Africa and South America, and the +{155}Marsupials of America and Australia, were all represented in Europe +(and probably also in North America) during the earlier part of the +Tertiary epoch. These facts, taken in their entirety, lead us to conclude +that, during the whole of the Tertiary and perhaps during much of the +Secondary periods, the great land masses of the earth were, as now, +situated in the Northern Hemisphere; and that here alone were developed the +successive types of vertebrata from the lowest to the highest. In the +Southern Hemisphere there appear to have been three considerable and very +ancient land masses, varying in extent from time to time, but always +keeping distinct from each other, and represented, more or less completely, +by Australia, South Africa, and South America of our time. Into these +flowed successive waves of life, as they each in turn became temporarily +united with some part of the northern land. Australia appears to have had +but one such union, perhaps during the middle or latter part of the +Secondary epoch, when it received the ancestors of its Monotremata and +Marsupials, which it has since developed into a great variety of forms. The +South African and South American lands, on the other hand, appear each to +have had several successive unions and separations, allowing first of the +influx of low forms only (Edentata, Insectivora and Lemurs); subsequently +of Rodents and small Carnivora, and, latest of all, of the higher types of +Primates, Carnivora and Ungulata. + +During the whole of the Tertiary period, at least, the Northern Hemisphere +appears to have been divided, as now, into an Eastern and a Western +continent; always approximating and sometimes united towards the north, and +then admitting of much interchange of their respective faunas; but on the +whole keeping distinct, and each developing its own special family and +generic types, of equally high grade, and generally belonging to the same +Orders. During the Eocene and Miocene periods, the distinction of the +Palæarctic and Nearctic regions was better marked than it is now; as is +shown by the floras no less than by the faunas of those epochs. Dr. +Newberry, in his Report on the Cretaceous and Tertiary floras of the +Yellowstone and Missouri Rivers, states, that although the Miocene flora of +Central North {156}America corresponds generally with that of the European +Miocene, yet many of the tropical, and especially the Australian types, +such as _Hakea_ and _Dryandra_, are absent. Owing to the recent discovery +of a rich Cretaceous flora in North America, probably of the same age as +that of Aix-la-Chapelle in Europe, we are able to continue the comparison; +and it appears, that at this early period the difference was still more +marked. The predominant feature of the European Cretaceous flora seems to +have been the abundance of Proteaceæ, of which seven genera now living in +Australia or the Cape of Good Hope have been recognised, besides others +which are extinct. There are also several species of _Pandanus_, or +screw-pine, now confined to the tropics of the Eastern Hemisphere, and +along with these, oaks, pines, and other more temperate forms. The North +American Cretaceous flora, although far richer than that of Europe, +contains no Proteaceæ or _Pandani_, but immense numbers of forest trees of +living and extinct genera. Among the former we have oaks, beeches, willows, +planes, alders, dog-wood, and cypress; together with such American forms as +magnolias, sassafras, and liriodendrons. There are also a few not now found +in America, as _Araucaria_ and _Cinnamomum_, the latter still living in +Japan. This remarkable flora has been found over a wide extent of +country--New Jersey, Alabama, Kansas, and near the sources of the Missouri +in the latitude of Quebec--so that we can hardly impute its peculiarly +temperate character to the great elevation of so large an area. The +intervening Eocene flora approximates closely, in North America, to that of +the Miocene period; while in Europe it seems to have been fully as tropical +in character as that of the preceding Cretaceous period; fruits of _Nipa_, +_Pandanus_, _Anona_, _Acacia_, and many Proteaceæ, occurring in the London +clay at the mouth of the Thames. + +These facts appear, at first sight, to be inconsistent, unless we suppose +the climates of Europe and North America to have been widely different in +these early times; but they may perhaps be harmonised, on the supposition +of a more uniform and a somewhat milder climate then prevailing over the +whole Northern Hemisphere; the contrast in the vegetation of these +countries {157}being due to a radical difference of type, and therefore not +indicative of climate. The early European flora seems to have been a +portion of that which now exists only in the tropical and sub-tropical +lands of the Eastern Hemisphere; and, as much of this flora still survives +in Australia, Tasmania, Japan, and the Cape of Good Hope, it does not +necessarily imply more than a warm and equable temperate climate. The early +North American flora, on the other hand, seems to have been essentially the +same in type as that which now exists there, and which, in the Miocene +period, was well represented in Europe; and it is such as now flourishes +best in the warmer parts of the United States. But whatever conclusion we +may arrive at on the question of climate, there can be no doubt as to the +distinctness of the floras of the ancient Nearctic and Palæarctic regions; +and the view derived from our study of their existing and extinct +faunas--that these two regions have, in past times, been more clearly +separated than they are now--receives strong support from the unexpected +evidence now obtained as to the character and mutations of their vegetable +forms, during so vast an epoch as is comprised in the whole duration of the +Tertiary period. + +The general phenomena of the distribution of living animals, combined with +the evidence of extinct forms, lead us to conclude that the Palæarctic +region of early Tertiary times was, for the most part, situated beyond the +tropics, although it probably had a greater southward extension than at the +present time. It certainly included much of North Africa, and perhaps +reached far into what is now the Sahara; while a southward extension of its +central mass may have included the Abyssinian highlands, where some truly +Palæarctic forms are still found. This is rendered probable by the fossils +of Perim Island a little further east, which show that the characteristic +Miocene fauna of South Europe and North India prevailed so far within the +tropics. There existed, however, at the extreme eastern and western limits +of the region, two extensive equatorial land-areas, our Indo-Malayan and +West African sub-regions--both of which must have been united for more or +less considerable periods with the northern continent. They would then have +received {158}from it such of the higher vertebrates as were best adapted +for the peculiar climatal and organic conditions which everywhere prevail +near the equator; and these would be preserved, under variously modified +forms, when they had ceased to exist in the less favourable and constantly +deteriorating climate of the north. At later epochs, both these equatorial +lands became united to some part of the great South African continent (then +including Madagascar), and we thus have explained many of the similarities +presented by the faunas of these distant, and generally very different +countries. + +During the Miocene period, when a subtropical climate prevailed over much +of Europe and Central Asia, there would be no such marked contrast as now +prevails between temperate and tropical zones; and at this time much of our +Oriental region, perhaps, formed a hardly separable portion of the great +Palæarctic land. But when, from unknown causes, the climate of Europe +became less genial, and when the elevation of the Himalayan chain and the +Mongolian plateau caused an abrupt difference of climate on the northern +and southern sides of that great mountain barrier, a tropical and a +temperate region were necessarily formed; and many of the animals which +once roamed over the greater part of the older and more extensive region, +now became restricted to its southern or northern divisions respectively. +Then came the great change we have already described (vol. i. p. 288), +opening the newly-formed plains of Central Africa to the incursions of the +higher forms of Europe; and following on this, a still further +deterioration of climate, resulting in that marked contrast between +temperate and tropical faunas, which is now one of the most prominent +features in the distribution of animal as well as of vegetable forms. + +It is not necessary to go into any further details here, as we have +already, in our discussion of the origin of the fauna of the several +regions, pointed out what changes most probably occurred in each case. +These details are, however, to a great extent speculative; and they must +remain so till we obtain as much knowledge of the extinct faunas and past +geological history of the southern lands, as we have of those of Europe and +North {159}America. But the broad conclusions at which we have now arrived +seem to rest on a sufficiently extensive basis of facts; and they lead us +to a clearer conception of the mutual relations and comparative importance +of the several regions than could be obtained at an earlier stage of our +inquiries. + +If our views of the origin of the several regions are correct, it is clear +that no mere binary division--into north and south, or into east and +west--can be altogether satisfactory, since at the dawn of the Tertiary +period we still find our six regions, or what may be termed the rudiments +of them, already established. The north and south division truly represents +the fact, that the great northern continents are the seat and birth-place +of all the higher forms of life, while the southern continents have derived +the greater part, if not the whole, of their vertebrate fauna from the +north; but it implies the erroneous conclusion, that the chief southern +lands--Australia and South America--are more closely related to each other +than to the northern continent. The fact, however, is that the fauna of +each has been derived, independently, and perhaps at very different times, +from the north, with which they therefore have a true genetic relation; +while any intercommunion between themselves has been comparatively recent +and superficial, and has in no way modified the great features of animal +life in each. The east and west division, represents--according to our +views--a more fundamental diversity; since we find the northern continent +itself so divided in the earliest Eocene, and even in Cretaceous times; +while we have the strongest proof that South America was peopled from the +Nearctic, and Australia and Africa from the Palæarctic region: hence, the +Eastern and Western Hemispheres are the two great branches of the tree of +life of our globe. But this division, taken by itself, would obscure the +facts--firstly, of the close relation and parallelism of the Nearctic and +Palæarctic regions, not only now but as far back as we can clearly trace +them in the past; and, secondly, of the existing radical diversity of the +Australian region from the rest of the Eastern Hemisphere. + +Owing to the much greater extent of the old Palæarctic region (including +our Oriental), and the greater diversity of {160}Mammalia it appears to +have produced, we can have little doubt that here was the earliest seat of +the development of the vertebrate type; and probably of the higher forms of +insects and land-molluscs. Whether the Nearctic region ever formed one mass +with it, or only received successive immigrations from it by northern +land-connections both in an easterly and westerly direction, we cannot +decide; but the latter seems the most probable supposition. In any case, we +must concede the first rank to the Palæarctic and Oriental regions, as +representing the most important part of what seems always to have been the +Great Continent of the earth, and the source from which all the other +regions were supplied with the higher forms of life. These once formed a +single great region, which has been since divided into a temperate and a +tropical portion, now sufficiently distinct; while the Nearctic region has, +by deterioration of climate, suffered a considerable diminution of +productive area, and has in consequence lost a number of its more +remarkable forms. The two temperate regions have thus come to resemble each +other more than they once did, while the Oriental retains more of the +zoological aspect of the great northern regions of Miocene times. The +Ethiopian, from having been once an insular region, where lower types of +vertebrates alone prevailed, has been so overrun with higher types from the +old Palæarctic and Oriental lands that it now rivals, or even surpasses, +the Oriental region in its representation of the ancient fauna of the great +northern continent. Both of our tropical regions of the Eastern Hemisphere +possess faunas which are, to some extent, composite, being made up in +different proportions of the productions of the northern and southern +continents,--the former prevailing largely in the Oriental, while the +latter constitutes an important feature in the Ethiopian fauna. The +Neotropical region has probably undergone great fluctuations in early +times; but it was, undoubtedly, for long periods completely isolated, and +then developed the Edentate type of Mammals and the Formicaroid type of +Passerine birds into a variety of forms, comparable with the diversified +Marsupials of Australia, and typical Passeres of the Eastern Hemisphere. +{161}It has, however, received successive infusions of higher types from +the north, which now mingle in various degrees with its lower forms. At an +early period it must have received a low form of Primates, which has been +developed into the two peculiar families of American monkeys; while its +llamas, tapirs, deer, and peccaries, came in at a later date, and its +opossums and extinct horses probably among the latest. The Australian +region alone, after having been united with the great northern continent at +a very early date (probably during the Secondary period) has ever since +remained more or less completely isolated; and thus exhibits the +development of a primeval type of mammal, almost wholly uninfluenced by any +incursions of a later and higher type. In this respect it is unique among +all the great regions of the earth. + +We see, then, that each of our six regions has had a history of its own, +the main outlines of which we have been able to trace with tolerable +certainty. Each of them is now characterised--as it seems to have been in +all past time of which we have any tolerably full record--by well-marked +zoological features; while all are connected and related in the complex +modes we have endeavoured to unravel. To combine any two or more of these +regions, on account of existing similarities which are, for the most part, +of recent origin, would obscure some of the most important and interesting +features of their past history and present condition. And it seems no less +impracticable to combine the whole into groups of higher rank; since it has +been shown that there are two opposing modes of doing this, and that each +of them represents but one aspect of a problem, which can only be solved by +giving equal attention to all its aspects. + + + +For reasons which have been already stated, and which are sufficiently +obvious, we have relied almost exclusively on the distribution of living +and extinct mammalia, in arriving at these conclusions. But we believe they +will apply equally to elucidate the phenomena presented by the distribution +of all terrestrial organisms, when combined with a careful consideration of +the {162}various means of dispersal of the different groups, and the +comparative longevity of their species and genera. Even insects, which are +perhaps of all animals the farthest removed from mammalia in this respect, +agree, in the great outlines of their distribution, with the vertebrate +orders. The Regions are admittedly the same, or nearly the same for both; +and the discrepancies that occur are of a nature which can be explained by +two undoubted facts--the greater antiquity, and the greater facilities for +dispersal, of insects. + +But this principle, if sound, must be carried farther, and be applied to +plants also. There are not wanting indications that this may be +successfully done; and it seems not improbable, that the reason why +botanists have hitherto failed to determine, with any unanimity, which are +the most natural phytological regions, and to work out any connected theory +of the migrations of plants, is, because they have not been furnished with +the clue to the past changes of the great land masses, which could only be +arrived at by such an examination of the past and present distribution of +the higher animals as has been here attempted. The difficulties in the way +of the study of the distribution of plants, from this point of view, will +be undoubtedly very great; owing to the unusual facilities for distribution +many of them possess, and the absence of any group which might take the +place of the mammalia among animals, and serve as a guide and standard for +the rest. We cannot expect the regions to be so well defined in the case of +plants as in that of animals; and there are sure to be many anomalies and +discrepancies, which will require long study to unravel. The Six Great +Regions here adopted, are however, as a whole, very well characterised by +their vegetable forms. The floras of tropical America, of Australia, of +South Africa, and of Indo-Malaya, stand out with as much individuality as +do the faunas; while the plants of the Palæarctic and Nearctic regions, +exhibit resemblances and diversities, of a character not unlike those found +among the animals. + +This is not a mere question of applying to the vegetable kingdom a series +of arbitrary divisions of the earth which have been {163}found useful to +zoologists; for it really involves a fundamental problem in the theory of +evolution. The question we have to answer, is, firstly--whether the +distribution of plants is, like that of animals, mainly and primarily +dependent on the past revolutions of the earth's surface; or, whether +other, and altogether distinct causes, have had a preponderating influence +in determining the range and limits of vegetable forms; and, +secondly--whether those revolutions have been, in their general outlines, +correctly interpreted by means of a study of the distribution and +affinities of the higher animals. The first question is one for botanists +alone to answer; but, on the second point, the author ventures to hope for +an affirmative reply, from such of his readers as will weigh carefully the +facts and arguments he has adduced. + + + +The remaining part of this volume, will consist, of a systematic review of +the distribution of each family of animals, and an application of the +principles already established to elucidate the chief phenomena they +present. The present chapter must, therefore, be considered as the +conclusion of the argumentative and theoretical part of the present work; +but it must be read in connection with the various discussions in Parts II. +and III., in which the conclusions to be drawn from the several groups of +facts have been successively given;--and especially in connection with the +general observations at the end of each of the six chapters on the +Zoological Regions. + +The hypothetical view, as to the more recent of the great Geographical +changes of the Earth's surface, here set forth, is not the result of any +preconceived theory, but has grown out of a careful study of the facts +accumulated, and has led to a considerable modification of the author's +previous views. It may be described, as an application of the general +theory of Evolution, to solve the problem of the distribution of animals; +but it also furnishes some independent support to that theory, both by +showing what a great variety of curious facts are explained by its means, +and by answering some of the objections, {164}which have been founded on +supposed difficulties in the distribution of animals in space and time. + +It also illustrates and supports the geological doctrine, of the general +permanence of our great continents and oceans, by showing how many facts in +the distribution of animals can only be explained and understood on such a +supposition; and it exhibits, in a striking manner, the enormous influence +of the Glacial epoch, in determining the existing zoological features of +the various continents. + +And, lastly, it furnishes a more consistent and intelligible idea than has +yet been reached by any other mode of investigation, of all the more +important changes of the earth's surface that have probably occurred during +the entire Tertiary period; and of the influence of these changes, in +bringing about the general features, as well as many of the more +interesting details and puzzling anomalies, of the Geographical +Distribution of Animals. + + + + + + +PART IV. + +_GEOGRAPHICAL ZOOLOGY:_ + +_A SYSTEMATIC SKETCH OF THE CHIEF FAMILIES OF LAND ANIMALS IN THEIR +GEOGRAPHICAL RELATIONS._ + + +{167}INTRODUCTION. + + +In the preceding part of our work, we have discussed the geographical +distribution of animals from the point of view of the geographer; taking +the different regions of the earth in succession, and giving as full an +account as our space would permit of their chief forms of animal life. Now, +we proceed from the standpoint of the systematic zoologist; taking in +succession each of the families with which we deal, and giving an account +of the distribution, both of the entire family and, as far as practicable, +of each of the genera of which it is composed. As in the former part, our +mode of treatment led us to speculate on the past changes of the earth's +surface; so here we shall endeavour to elucidate the past migrations of +animals, and thus, to some extent, account for their actual distribution. + +The tabular headings, showing the range of the family in each region, will +enable the reader to determine at a glance the general distribution of the +group, as soon as he has familiarised himself, by a study of our general +and regional maps, with the limits of the regions and sub-regions, and the +figures (1 to 4) by which the latter are indicated. Much pains have been +taken, to give the number of the known genera and species in each family, +correctly; but these numbers must, in most cases, only be looked upon as +approximations; because, owing to constant accessions of fresh material on +the one hand, and the discovery that many supposed species are only +varieties, on the other, such statistics are in a continual state of +fluctuation. In the number of genera there is the greatest uncertainty; as +will be seen by the two sets of numbers sometimes given, which denote the +genera according to different modern authorities. + +{168}There is also a considerable difference in the dependence to be placed +on the details given in the different classes of animals. In Mammalia and +Birds some degree of accuracy has, it is hoped, been attained; the +classification of these groups being much advanced, and the materials for +their study ample. In Reptiles this is not the case, as there is no +recently published work dealing with the whole subject, or with either of +the larger orders. An immense number of new species and new genera of +snakes and lizards, have been described in the last twenty years; and Dr. +Günther--our greatest authority on reptiles in this country--has kindly +assisted me in incorporating such of these as are most trustworthy, in a +general system; but until entire Orders have been described or catalogued +on a uniform plan, nothing more than a general approximation to the truth +can be arrived at. Still, so many of the groups are well defined, and have +a clearly limited distribution, that some interesting and valuable +comparisons may be made. + +For Fishes, the valuable "Catalogue" of Dr. Günther was available, and it +has rarely been attempted to go beyond it. A large number of new species +have since been described, in all parts of the world; but it is impossible +to say how many of these are really new, or what genera they actually +belong to. The part devoted to this Class is, therefore, practically a +summary of Dr. Günther's Catalogue; and it is believed that the discoveries +since made will not materially invalidate the conclusions to be drawn from +such a large number of species, which have been critically examined and +classified on a uniform system by one of our most able naturalists. When a +supplement to this catalogue is issued, it will be easier to make the +necessary alterations in distribution, than if a mass of untrustworthy +materials had been mixed up with it. + +For Insects, excellent materials are furnished, in the Catalogue of Mr. +Kirby for Butterflies and in that of Drs. Gemminger and Harold for +Coleoptera. I have also made use of some recently published memoirs on the +Insects of Japan and St. Helena, and a few other recent works; and have, I +believe, elaborated a more extensive series of facts to illustrate the +distribution of insects, {169}than has been made use of by any previous +writer. Several discussions on the bearing of the facts of insect +distribution, will also be found under the several Regions, in the +preceding part of this work. + +Terrestrial Mollusca form a group, as to the treatment of which I have most +misgivings; owing to my almost entire ignorance of Malacology, and the +great changes recently made in the classification of shells. There is also +much uncertainty as to genera and sub-genera, which is very puzzling to one +who merely wishes to get at general results. Finding it impossible to +incorporate the new matter with the old, or to harmonise the different +classifications of modern conchologists, I thought it better to confine +myself to the standard works of Martens and Pfeiffer, with such additions +of new species as I could make without fear of going far wrong. In some +cases I have made use of recent monographs--especially on the shells of +Europe, North America, the West Indian Islands, and the Sandwich Islands; +and have, I venture to hope, not fallen into much error in the general +conclusions at which I have arrived. + + + + +{170}CHAPTER XVII. + +THE DISTRIBUTION OF THE FAMILIES AND GENERA OF MAMMALIA. + + + +_Order I.--PRIMATES._ + +FAMILY 1.--SIMIIDÆ. (4 Genera, 12 Species). + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- |-- -- 3. 4 |-- -- -- -- + | | | | | + +The Simiidæ, or Anthropoid Apes, comprehend those forms of the monkey-tribe +which, in general organization, approach nearest to man. They inhabit the +tropics of the Old World, and are most abundant near the equator; but they +are limited to certain districts, being quite unknown in eastern and +southern Africa, and the whole peninsula of Hindostan. + +The genus _Troglodytes_ (or _Mimetes_, as it is sometimes named) +comprehends the chimpanzee and gorilla. It is confined to the West African +sub-region, being found on the coast about 12° North and South of the +equator, from the Gambia to Benguela, and as far inland as the great +equatorial forests extend. There are perhaps other species of chimpanzee; +since Livingstone met with what he supposed to be a new species in the +forest region west of Lake Tanganyika, while Dr. Schweinfurth found one in +the country beyond the western watershed of the Nile. The gorilla is +confined within narrower limits on and near the equator. + +{171}We have to pass over more than 70° of longitude before we again meet +with Anthropoid Apes, in the northern part of Sumatra--where a specimen of +the orang-utan (_Simia satyrus_) now in the Calcutta Museum, was obtained +by Dr. Abel, and described by him in the _Asiatic Researches_, vol. +xv.--and in Borneo, from which latter island almost all the specimens in +European museums have been derived. There are supposed to be two species of +_Simia_ in Borneo, a larger and a smaller; but their distinctness is not +admitted by all naturalists. Both appear to be confined to the swampy +forests near the north, west, and south coasts. + +The Gibbons, or long-armed apes, forming the genus _Hylobates_, (7 species) +are found in all the large islands of the Indo-Malayan sub-region, except +the Philippines; and also in Sylhet and Assam south of the Brahmaputra +river, eastward to Cambodja and South China to the west of Canton, and in +the island of Hainan. + +The Siamang (_Siamanga syndactyla_) presents some anatomical peculiarities, +and has the second and third toes united to the last joint, but in general +form and structure it does not differ from _Hylobates_. It is the largest +of the long-armed apes, and inhabits Sumatra and the Malay peninsula. + + +FAMILY 2.--SEMNOPITHECIDÆ. (2 Genera, 30 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- 4 |1. 2 -- -- |1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Semnopithecidæ, are long-tailed monkeys without cheek-pouches, and with +rather rounded faces, the muzzle not being prominent. They have nearly the +same distribution as the last family, but are more widely dispersed in both +Africa and Asia, one species just entering the Palæarctic region. + +The Eastern genus _Presbytes_ or _Semnopithecus_ (29 species), is spread +over almost the whole of the Oriental region wherever the forests are +extensive. They extend along the Himalayas to beyond Simla, where a species +has been observed at an altitude of 11,000 {172}feet, playing among +fir-trees laden with snow wreaths. On the west side of India they are not +found to the north of 14° N. latitude. On the east they extend into Arakan, +and to Borneo and Java, but not apparently into Siam or Cambodja. Along the +eastern extension of the Himalayas they again occur in East Thibet; a +remarkable species with a large upturned nose (_S. roxellana_) having been +discovered by Père David at Moupin (about Lat. 32° N.) in the highest +forests, where the winters are severe and last for several months, and +where the vegetation, and the other forms of animal life, are wholly those +of the Palæarctic region. It is very curious that this species should +somewhat resemble the young state of the proboscis monkey (_S. nasalis_), +which inhabits one of the most uniform, damp, and hot climates on the +globe--the river-swamps of Borneo. + +_Colobus_, the African genus (11 species), is very closely allied to the +preceding, differing chiefly in the thumb being absent or rudimentary. They +are confined to the tropical regions--Abyssinia on the east, and from the +Gambia to Angola and the island of Fernando Po, on the west. + + +FAMILY 3.--CYNOPITHECIDÆ. (7 Genera, 67 Species). + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --| -- 2 -- 4 |1. 2. 3 -- |1. 2. 3. 4 |1 -- -- -- + | | | | | + +This family comprehends all the monkeys with cheek pouches, and the +baboons. Some of these have very long tails, some none; some are dog-faced, +others tolerably round-faced; but there are so many transitions from one to +the other, and such a general agreement in structure, that they are now +considered to form a very natural family. Their range is more extensive +than any other family of Quadrumana, since they not only occur in every +part of the Ethiopian and Oriental regions, but enter the Palæarctic region +in the east and west, and the Australian region as far as the islands of +Timor and Batchian. The African genera {173}are _Myiopithecus_, +_Cercopithecus_, _Cercocebus_, _Theropithecus_, and _Cynocephalus_; the +Oriental genera, _Macacus_, and _Cynopithecus_. + +_Myiopithecus_ (1 species), consisting of the talapoin monkey of West +Africa, differs from the other African monkeys in the structure of the last +molar tooth; in the large ears, short face, and wide internasal septum; in +this respect, as well as in its grace and gentleness, resembling some of +the American monkeys. + +_Cercopithecus_ (24 species), contains all the more graceful and prettily +coloured monkeys of tropical Africa, and comprises the guenons, the +white-nosed, and the green monkeys. They range from the Gambia to the +Congo, and from Abyssinia to the Zambesi. + +_Cercocebus_ (5 species), the mangabeys, of West Africa, are very closely +allied to the eastern genus _Macacus_. + +_Theropithecus_ (2 species), including the gelada of Abyssinia and an +allied species, resemble in form the baboons, but have the nostrils placed +as in the last genus. + +_Cynocephalus_ (10 species), the baboons, are found in all parts of Africa. +They consist of animals which vary much in appearance, but which agree in +having an elongated dog-like muzzle with terminal nostrils, and being of +terrestrial habits. Some of the baboons are of very large size, the +mandrill (_C. maimon_) being only inferior to the orang and gorilla. + +_Macacus_ (25 species), is the commonest form of eastern monkey, and is +found in every part of the Oriental region, as well as in North Africa, +Gibraltar, Thibet, North China, and Japan; and one of the commonest +species, _M. cynomolgus_, has extended its range from Java eastward to the +extremity of Timor. The tail varies greatly in length, and in the Gibraltar +monkey (_M. innus_) is quite absent. A remarkable species clothed with very +thick fur, has lately been discovered in the snowy mountains of eastern +Thibet. + +_Cynopithecus_ (? 2 sp.).--This genus consists of a black baboon-like Ape, +inhabiting Celebes, Batchian, and the Philippine Islands; but perhaps +introduced by man into the latter islands and into Batchian. It is doubtful +if there is more than one species. The tail of this animal is a fleshy +tubercle, the nostrils as in _Macacus_, but the muzzle is very prominent; +and the {174}development of the maxillary bones into strong lateral ridges +corresponds to the structure of the most typical baboons. This species +extends further east than any other quadrumanous animal. + + +FAMILY 4.--CEBIDÆ. (10 Genera, 78 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Cebidæ, which comprehend all the larger American Monkeys, differ from +those of the Old World by having an additional molar tooth in each jaw, and +a broad nasal septum; while they have neither cheek-pouches nor ischial +callosities, and the thumb is never completely opposable. Some have +prehensile tails, especially adapting them for an arboreal life. They are +divided into four sub-families,--Cebinæ, Mycetinæ, Pitheciinæ, and +Nyctipithecinæ. The Cebidæ are strictly confined to the forest regions of +tropical America, from the southern part of Mexico to about the parallel of +30° South Latitude. The distribution of the genera is as follows:-- + +_Sub-family_, Cebinæ.--_Cebus_ (18 sp.), is the largest genus of American +monkeys, and ranges from Costa Rica to Paraguay. They are commonly called +sapajous. _Lagothrix_ (5 sp.), the woolly monkeys, are rather larger and +less active than the preceding; they are confined to the forests of the +Upper Amazon Valley, and along the slopes of the Andes to Venezuela and +Bolivia. _Ateles_ (14 sp.), the spider monkeys, have very long limbs and +tail. They range over the whole area of the family, and occur on the west +side of the Equatorial Andes and on the Pacific coast of Guatemala. +_Eriodes_ (3 sp.), are somewhat intermediate between the last two genera, +and are confined to the eastern parts of Brazil south of the equator. The +three last mentioned genera have very powerful prehensile tails, the end +being bare beneath; whereas the species of _Cebus_ have the tail +{175}completely covered with hair, although prehensile, and therefore not +so perfect a grasping organ. + +_Sub-family_, Mycetinæ, consists of but a single genus, _Mycetes_ (10 sp.), +the howling monkeys, characterized by having a hollow bony vessel in the +throat formed by an enlargement of the hyoid bone, which enables them to +produce a wonderful howling noise. They are large, heavy animals, with a +powerful and perfect prehensile tail. They range from East Guatemala to +Paraguay. (Plate XIV., vol. ii., p. 24.) + +_Sub-family_, Pitheciinæ, the sakis, have a non-prehensile bushy tail. +_Pithecia_ (7 sp.), has the tail of moderate length; while _Brachiurus_ (5 +sp.) has it very short. Both appear to be restricted to the great +equatorial forests of South America. + +_Sub-family_, Nyctipithecinæ, are small and elegant monkeys, with long, +hairy, non-prehensile tails. _Nyctipithecus_ (5 sp.), the night-monkeys or +douroucoulis, have large eyes, nocturnal habits, and are somewhat lemurine +in their appearance. They range from Nicaragua to the Amazon and eastern +Peru. _Saimiris_ or _Chrysothrix_ (3 sp.), the squirrel-monkeys, are +beautiful and active little creatures, found in most of the tropical +forests from Costa Rica to Brazil and Bolivia. _Callithrix_ (11 sp.), are +somewhat intermediate between the last two genera, and are found all over +South America from Panama to the southern limits of the great forests. + + +FAMILY 5.--HAPALIDÆ. (2 Genera, 32 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Hapalidæ, or marmosets, are very small monkeys, which differ from the +true Cebidæ in the absence of one premolar tooth, while they possess the +additional molar tooth; so that while they have the same number of teeth +(thirty-two) as the Old World monkeys, they differ from them even more than +do the {176}Cebidæ. The thumb is not at all opposable, and all the fingers +are armed with sharp claws. The hallux, or thumb-like great toe, is very +small; the tail is long and not prehensile. The two genera _Hapale_ (9 +sp.), and _Midas_ (24 sp.), are of doubtful value, though some naturalists +have still further sub-divided them. They are confined to the tropical +forests of South America, and are most abundant in the districts near the +equator. + + +_Sub-order--LEMUROIDEA._ + +FAMILY 6.--LEMURIDÆ. (11 Genera, 53 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3. 4 |-- 2. 3. 4 |-- -- -- --|-- -- -- -- + | | | | | + +The Lemuridæ, comprehending all the animals usually termed Lemurs and many +of their allies, are divided by Professor Mivart--who has carefully studied +the group--into four sub-families and eleven genera, as follows:-- + +_Sub-family_ Indrisinæ, consisting of the genus _Indris_ (5 sp.), is +confined to Madagascar. + +_Sub-family_ Lemurinæ, contains five genera, viz.:--_Lemur_, (15 sp.); +_Hapalemur_ (2 sp.); _Microcebus_ (4 sp.); _Chirogaleus_ (5 sp.); and +_Lepilemur_ (2 sp.);--all confined to Madagascar. + +_Sub-family_ Nycticebinæ, contains four genera, viz.:--_Nycticebus_ (3 +sp.)--small, short-tailed, nocturnal animals, called slow-lemurs,--range +from East Bengal to South China, and to Borneo and Java; _Loris_ (1 sp.)--a +very small, tail-less, nocturnal lemur, which inhabits Madras, Malabar, and +Ceylon; _Perodicticus_ (1 sp.)--the potto--a small lemur with almost +rudimentary forefinger, found at Sierra Leone (Plate V., vol. i., p. 264); +_Arctocebus_ (1 sp.)--the angwantibo,--another extraordinary form in which +the forefinger is quite absent and the first toe armed with a long +claw,--inhabits Old Calabar. + +{177}_Sub-family_ Galaginæ, contains only the genus _Galago_ (14 sp.), +which is confined to the African continent, ranging from Senegal and +Fernando Po to Zanzibar and Natal. + + +FAMILY 7.--TARSIIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 |-- -- -- -- + | | | | | + +The curious _Tarsius spectrum_, which constitutes this family, inhabits +Sumatra, Banca, and Borneo, and is also found in some parts of Celebes, +which would bring it into the Australian region; but this island is +altogether so anomalous that we can only consider its productions to have +somewhat more affinity with the Australian than the Oriental region, but +hardly to belong to either. The Tarsier is a small, long-tailed, nocturnal +animal, of curious structure and appearance; and it forms the only link of +connection with the next family, which it resembles in the extraordinary +development of the toes, one of which is much larger and more slender than +the rest. (Plate VIII., vol. i. p. 337.) + + +FAMILY 8.--CHIROMYIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 |-- -- -- --|-- -- -- -- + | | | | | + +The Aye-aye, (_Chiromys_), the sole representative of this family, is +confined to the island of Madagascar. It was for a long time very +imperfectly known, and was supposed to belong to the Rodentia; but it has +now been ascertained to be an exceedingly specialized form of the Lemuroid +type, and must be considered to be one of the most extraordinary of the +mammalia now inhabiting the globe. (Plate VI., vol. i., p. 278.) + + +{178}_Fossil Quadrumana._ + +Not much progress has yet been made in tracing back the various forms of +Apes and Monkeys to their earliest appearance on the globe; but there have +been some interesting recent discoveries, which lead us to hope that the +field is not yet exhausted. The following is a summary of what is known as +to the early forms of each family:-- + +_Simiidæ._--Two or three species of this family have been found in the +Upper Miocene deposits of France and Switzerland. _Pliopithecus_, of which +a species has been found at each locality, was allied to the gibbons +(_Hylobates_), and perhaps to _Semnopithecus_. A more remarkable form, +named _Dryopithecus_, as large as a man, and having peculiarities of +structure which are thought by Gervais and Lartet to indicate a nearer +approach to the human form than any existing Ape, has been found in strata +of the same age in France. + +_Semnopithecidæ._--Species of _Semnopithecus_ have been found in the Upper +Miocene of Greece, and others in the Siwalik Hills of N. W. India, also of +Upper Miocene age. An allied form also occurs in the Miocene of Wurtemburg. +_Mesopithecus_ from Greece is somewhat intermediate between _Semnopithecus_ +and _Macacus_. + +Remains supposed to be of _Semnopithecus_, have also occurred in the +Pliocene of Montpellier. + +_Cynopithecidæ._--_Macacus_ has occurred in Pliocene deposits at Grays, +Essex; and also in the South of France along with _Cercopithecus_. + +_Cebidæ._--In the caves of Brazil remains of the genera _Cebus_, _Mycetes_, +_Callithrix_, and _Hapale_, have been found; as well as an extinct form of +larger size--_Protopithecus_. + +_Lemuroidea._--A true lemur has recently been discovered in the Eocene of +France; and it is supposed to be most nearly allied to the peculiar West +African genera, _Perodicticus_ and _Arctocebus_. + +_Cænopithecus_, from the Swiss Jura, is supposed to have affinities both +for the Lemuridæ and the American Cebidæ. + +In the lower Eocene of North America remains have been {179}discovered, +which are believed to belong to this sub-order: but they form two distinct +families,--Lemuravidæ and Limnotheridæ. Other remains from the Miocene are +believed to be intermediate between these and the Cebidæ,--a most +interesting and suggestive affinity, if well founded. For the genera of +these American Lemuroidea, see vol. i., p. 133. + + +_General Remarks on the Distribution of Primates._ + +The most striking fact presented by this order, from our present point of +view, is the strict limitation of well-marked families to definite areas. +The Cebidæ and Hapalidæ would alone serve to mark out tropical America as +the nucleus of one of the great zoological divisions of the earth. In the +Eastern Hemisphere, the corresponding fact is the entire absence of the +order from the Australian region, with the exception of one or two outlying +forms, which have evidently transgressed the normal limits of their group. +The separation of the Ethiopian and Oriental regions is, in this order, +mainly indicated by the distribution of the genera, no one of which is +common to the two regions. The two highest families, the Simiidæ and the +Semnopithecidæ, are pretty equally distributed about two equatorial foci, +one situated in West Africa, the other in the Malay archipelago,--in Borneo +or the Peninsula of Malacca;--while the third family, Cynopithecidæ, ranges +over the whole of both regions, and somewhat overpasses their limits. The +Lemuroid group, on the other hand, offers us one of the most singular +phenomena in geographical distribution. It consists of three families, the +species of which are grouped into six sub-families and 13 genera. One of +these families and two of the sub-families, comprising 7 genera, and no +less than 30 out of the total of 50 species, are confined to the one island +of Madagascar. Of the remainder, 3 genera, comprising 15 species, are +spread over tropical Africa; while three other genera with 5 species, +inhabit certain restricted portions of India and the Malay islands. These +curious facts point unmistakably to the former existence of a large tract +of land in what is now the Indian Ocean, connecting Madagascar on the one +hand with Ceylon, and with the Malay countries on the {180}other. About +this same time (but perhaps not contemporaneously) Madagascar must have +been connected with some portion of Southern Africa, and the whole of the +country would possess no other Primates but Lemuroidea. After the +Madagascar territory (very much larger than the existing island) had been +separated, a connection appears to have been long maintained (probably by a +northerly route) between the more equatorial portions of Asia and Africa; +till those higher forms had become developed, which were afterwards +differentiated into _Simia_, _Presbytes_, and _Cynopithecus_, on the one +hand, and into _Troglodytes_, _Colobus_, and _Cynocephalus_, on the other. +In accordance with the principle of competition so well expounded by Mr. +Darwin, we can understand how, in the vast Asiatic and African area north +of the Equator, with a great variety of physical conditions and the +influence of a host of competing forms of life, higher types were developed +than in the less extensive and long-isolated countries south of the +Equator. In Madagascar, where these less complex conditions prevailed in a +considerable land-area, the lowly organized Lemuroids have diverged into +many specialized forms of their own peculiar type; while on the continents +they have, to a great extent, become exterminated, or have maintained their +existence in a few cases, in islands or in mountain ranges. In Africa the +nocturnal and arboreal _Galagos_ are adapted to a special mode of life, in +which they probably have few competitors. + +How and when the ancestors of the Cebidæ and Hapalidæ entered the South +American continent, it is less easy to conceive. The only rays of light we +yet have on the subject are, the supposed affinities of the fossil +_Cænopithecus_ of the Swiss, and the Lemuravidæ of the North American +Eocene, with both Cebidæ and Lemuroids, and the fact that in Miocene or +Eocene times a mild climate prevailed up to the Arctic circle. The +discovery of an undoubted Lemuroid in the Eocene of Europe, indicates that +the great Northern Continent was probably the birthplace of this low type +of mammal, and the source whence Africa and Southern Asia were peopled with +them, as it was, at a later period, with the higher forms of monkeys and +apes. + + +{181}_Order II.--CHIROPTERA._ + +FAMILY 9.--PTEROPIDÆ. (9 Genera, 65 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +The Pteropidæ, or fruit-eating Bats, sometimes called flying-foxes, are +pretty evenly distributed over the tropical regions of the Old World and +Australia. They range over all Africa and the whole of the Oriental Region, +and northward, to Amoy in China and to the South of Japan. They are also +found in the more fertile parts of Australia and Tasmania, and in the +Pacific Islands as far east as the Marianne and Samoa Islands; but not in +the Sandwich Islands or New Zealand. + +The genera of bats are exceedingly numerous, but they are in a very +unsettled state, and the synonymy is exceedingly confused. The details of +their distribution cannot therefore be usefully entered into here. The +Pteropidæ differ so much from all other bats, that they are considered to +form a distinct suborder of Chiroptera, and by some naturalists even a +distinct order of Mammalia. + +No fossil Pteropidæ have been discovered. + + +FAMILY 10.--PHYLLOSTOMIDÆ. (31 Genera, 60 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. --|1 -- -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Phyllostomidæ, or simple leaf-nosed Bats, are confined to the +Neotropical region, from Mexico and the Antilles to the {182}southern +limits of the forest region east of the Andes, and to about lat. 33° S. in +Chili. None are found in the Nearctic region, with the exception of one +species in California (_Macrotus Californicus_), closely allied to Mexican +and West Indian forms. The celebrated blood-sucking vampyre bats of South +America belong to this group. Two genera, _Desmodus_ and _Diphylla_, form +Dr. Peters' family Desmodidæ. Mr. Dobson, in his recently published +arrangement, divides the family into five groups:--Mormopes, Vampyri, +Glossophagæ, Stenodermata, and Desmodontes. + +Numerous remains of extinct species of this family have been found in the +bone-caves of Brazil. + + +FAMILY 11.--RHINOLOPHIDÆ. (7 Genera, 70 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. -- -- + | | | | | + +The Rhinolophidæ, or Horse-shoe Bats (so-called from a curiously-shaped +membranous appendance to the nose), range over all the Ethiopian and +Oriental regions, the southern part of the Palæarctic region, Australia and +Tasmania. They are most abundant and varied in the Oriental region, where +twelve genera are found; while only five inhabit the Australian and +Ethiopian regions respectively. Europe has only one genus and four species, +mostly found in the southern parts, and none going further north than the +latitude of England, where two species occur. Two others are found in +Japan, at the opposite extremity of the Palæarctic region. + +The genera _Nycteris_ and _Megaderma_, which range over the Ethiopian and +Oriental regions to the Moluccas, are considered by Dr. Peters to form a +distinct family, Megadermidæ; and Mr. Dobson in his recent arrangement +(published after our first {183}volume was printed) adopts the same family +under the name of Nycteridæ. The curious Indian genus _Rhinopoma_, which, +following Dr. J. E. Gray, we have classed in this family, is considered by +Mr. Dobson to belong to the Noctilionidæ. + +_Fossil Rhinolophidæ._--Remains of a species of _Rhinolophus_ still living +in England, have been found in Kent's Cavern, near Torquay. + + +FAMILY 12.--VESPERTILIONIDÆ (18 Genera, 200 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The small bats constituting the family Vespertilionidæ, have no +nose-membrane, but an internal earlet or _tragus_, and often very large +ears. They range over almost the whole globe, being apparently only limited +by the necessity of procuring insect food. In America they are found as far +north as Hudson's Bay and the Columbia river; and in Europe they approach, +if they do not pass the Arctic circle. Such remote islands as the Azores, +Bermudas, Fiji Islands, Sandwich Islands, and New Zealand, all possess +species of this group of bats, some of which probably inhabit every island +in warm or temperate parts of the globe. + +The genus _Taphozous_, which, in our Tables of Distribution in vol. i. we +have included in this family, is placed by Mr. Dobson in his family +Emballonuridæ, which is equivalent to our next family, Noctilionidæ. + +_Fossil Vespertilionidæ._--Several living European bats of this +family--_Scotophilus murinus_, _Plecotus auritus_, _Vespertilio noctula_, +and _V. pipestrellus_--have been found fossil in bone-caves in various +parts of Europe. + +Extinct species of _Vespertilio_ have occurred in the Lower Miocene at +Mayence, in the Upper Miocene of the South of France, and in the Upper +Eocene of the Paris basin. + + +{184}FAMILY 13.--NOCTILIONIDÆ. (14 Genera, 50 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1 -- -- -- |-- 2 -- -- |1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- 4 + | | | | | + +The Noctilionidæ, or short-headed Bats, are found in every region, but are +very unequally distributed. Their head-quarters is the Neotropical region, +where most of the genera occur, and where they range from Mexico to Buenos +Ayres and Chili, while in North America there is only one species in +California. They are unknown in Australia; but one species occurs in New +Zealand, and another in Norfolk Island. Several species of _Dysopes_ (or +_Molossus_) inhabit the Oriental region; one or two species being widely +distributed over the continent, while two others inhabit the Indo-Malayan +Islands. A species of this same genus occurs in South Africa, and another +in Madagascar and in the Island of Bourbon; while one inhabits Southern +Europe and North Africa, and another is found at Amoy in China. It will be +seen therefore, that these are really South American bats, which have a few +allies widely scattered over the various regions of the globe. Their +affinities are, according to Mr. Tomes, with the Phyllostomidæ, a purely +South American family. The species which forms the connecting link is the +_Mystacina tuberculata_, a New Zealand bat, which may, with almost equal +propriety be placed in either family, and which affords an interesting +illustration of the many points of resemblance between the Australian and +Neotropical regions. + +Dr. Peters has separated this family into three,--Mormopidæ, which is +wholly Neotropical, and is especially abundant in the West Indian Islands; +Molossidæ, chiefly consisting of the genus _Molossus_; and Noctilionidæ, +comprising the remainder of the family, and wholly Neotropical. Mr. Dobson, +however, classes the Mormopes with the Phyllostomidæ, and reduces the +{185}Molossi to the rank of a sub-family. In our first volume we have +classed _Rhinopoma_ with the Rhinolophidæ, and _Taphozous_ with the +Vespertilionidæ; but according to Mr. Dobson both these genera belong to +the present family. + + +_Remarks on the Distribution of the Order Chiroptera._ + +Although the bats, from their great powers of flight, are not amenable to +the limitations which determine the distribution of other terrestrial +mammals, yet certain great facts of distribution come out in a very +striking manner. The speciality of the Neotropical region is well shown, +not only by its exclusive possession of one large family (Phyllostomidæ), +but almost equally so by the total absence of two others (Pteropidæ and +Rhinolophidæ). The Nearctic region is also unusually well marked, by the +total absence of a family (Rhinolophidæ) which is tolerably well +represented in the Palæarctic. The Pteropidæ well characterize the tropical +regions of the Old World and Australia; while the Vespertilionidæ are more +characteristic of the Palæarctic and Nearctic regions, which together +possess about 60 species of this family. + +The bats are a very difficult study, and it is quite uncertain how many +distinct species are really known. Schinz, in his _Synopsis Mammalium_ +(1844) describes 330, while the list given by Mr. Andrew Murray in his +_Geographical Distribution of Mammalia_ (1866), contains 400 species. A +small number of new species have been since described, but others have been +sunk as synonyms, so that we can perhaps hardly obtain a nearer +approximation to the truth than the last number. In Europe there are 35 +species, and only 17 in North America. + +_Fossil Chiroptera._--The fossil remains of bats that have yet been +discovered, being chiefly allied to forms still existing in the same +countries, throw no light on the origin or affinities of this remarkable +and isolated order of Mammalia; but as species very similar to those now +living were in existence so far back as Miocene or even Eocene times, we +may be sure the group is one of immense antiquity, and that there has been +ample time for the amount of variation and extinction required to bring +about {186}the limitation of types, and the peculiarities of distribution +we now find to exist. + + +_Order III.--INSECTIVORA._ + +FAMILY 14.--GALEOPITHECIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 |-- -- -- -- + | | | | | + +The singular and isolated genus _Galeopithecus_, or flying lemur, has been +usually placed among the Lemuroidea, but it is now considered to come best +at the head of the Insectivora. Its food however, seems to be purely +vegetable, and the very small, blind, and naked young, closely attached to +the wrinkled skin of the mother's breast, perhaps indicates some affinity +with the Marsupials. This animal seems, in fact, to be a lateral offshoot +of some low form, which has survived during the process of development of +the Insectivora, the Lemuroidea, and the Marsupials, from an ancestral +type. Only two species are known, one found in Malacca, Sumatra, and +Borneo, but not in Java; the other in the Philippine islands (Plate VIII. +vol. i. p. 337). + + +FAMILY 15.--MACROSCELIDIDÆ. (3 Genera, 10 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- | 1 -- 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The Macroscelides, or elephant shrews, are extraordinary little animals, +with trunk-like snout and kangaroo-like hind-legs. They are almost confined +to South Africa, whence they extend up the east coast as far as the Zambezi +and Mozambique. A {187}single outlying species of _Macroscelides_ inhabits +Barbary and Algeria; while the two genera _Petrodromus_, and _Rhyncocyon_, +each represented by a single species, have only been found at Mozambique. + + +FAMILY 16.--TUPAIIDÆ. (3 Genera, 10 species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- 2. 3. 4 |-- -- -- -- + | | | | | + +The Tupaiidæ are squirrel-like shrews, having bushy tails, and often +climbing up trees, but also feeding on the ground and among low bushes. The +typical _Tupaia_ (7 species), are called ground squirrels by the Malays. +They are most abundant in the Malay islands and Indo-Chinese countries, but +one species is found in the Khasia Mountains, and one in the Eastern Ghauts +near Madras. The small shorter-tailed _Hylomys_ (2 species) is found from +Tenasserim to Java and Borneo; while the elegant little _Ptilocerus_ (1 +species) with its long pencilled tail, is confined to Borneo; (Plate VIII. +vol. i. p. 337). The family is therefore especially Malayan, with outlying +species in northern and continental India. + +_Extinct Species._--_Oxygomphus_, found in the Tertiary deposits of +Germany, is believed to belong to this family; as is _Omomys_, from the +Pliocene of the United States. + + +FAMILY 17.--ERINACEIDÆ. (2 Genera, 15 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3. 4 |-- -- 3 -- |1. 2. -- 4 |-- -- -- -- + | | | | | + +The Hedgehogs, comprised in the genus _Erinaceus_ (14 species), are widely +distributed over the Palæarctic, and a part of the {188}Oriental regions; +but they only occur in the Ethiopian region in South Africa and in the +Deserts of the north, which more properly belong to the Palæarctic region. +They are absent from the Malayan, and also from the Indo-Chinese +sub-regions; except that they extend from the north of China to Amoy and +Formosa and into the temperate highlands of the Western Himalayas. The +curious _Gymnura_ (1 species) is found in Borneo, Sumatra, and the Malay +peninsula. + +_Extinct Species._--The common hedgehog has been found fossil in several +Post-tertiary deposits, while extinct species occur in the lower Miocene of +Auvergne and in some other parts of Europe. Many of these remains are +classed in different genera from the living species;--(_Amphechinus_, +_Tetracus_, _Galerix_.) + + +FAMILY 18.--CENTETIDÆ. (6 Genera, 10 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- 4 |-- -- -- --|-- -- -- --|-- -- -- 4 |-- -- -- --|-- -- -- -- + | | | | | + +The Centetidæ are small animals, many of them having a spiny covering, +whence the species of _Centetes_ have been called Madagascar hedgehogs. The +genera _Centetes_ (2 species), _Hemicentetes_ (1 species), _Ericulus_ (1 +species), _Echinops_ (3 species), and the recently described _Oryzorictes_ +(1 species), are all exclusively inhabitants of Madagascar, and are almost +or quite tail-less. The remaining genus, _Solenodon_, is a more slender and +active animal, with a long, rat-like tail, shrew-like head, and coarse fur; +and the two known species are among the very few indigenous mammals of the +West Indian islands, one being found in Cuba (Plate XVII., vol. ii., p. +67), the other in Hayti. Although presenting many points of difference in +detail, the essential characters of this curious animal are, according to +Professors Peters and Mivart, identical with the rest of the Centetidæ. We +have thus a most remarkable and well-established case of discontinuous +distribution, two portions of the same family {189}being now separated from +each other by an extensive continent, as well as by a deep ocean. + +_Extinct Species._--Remains found in the Lower Miocene of the South of +France are believed to belong to the genus _Echinops_, or one closely +allied to it. + + +FAMILY 19.--POTAMOGALIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- |-- -- -- --|-- -- -- -- + | | | | | + +The genus _Potamogale_ was founded on a curious, small, otter-like animal +from West Africa, first found by M. Du Chaillu at the Gaboon, and +afterwards by the Portuguese at Angola. Its affinities are with several +groups of Insectivora, but it is sufficiently peculiar to require the +establishment of a distinct family for its reception. (Plate V., vol. i., +p. 264.) + + +FAMILY 20.--CHRYSOCHLORIDÆ. (2 Genera, 3 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- 3.-- |-- -- -- --|-- -- -- -- + | | | | | + +The Chrysochloridæ, or golden moles, of the Cape of Good Hope have been +separated by Professor Mivart into two genera, _Chrysochloris_ and +_Chalcochloris_. They are remarkable mole-like animals, having beautiful +silky fur, with a metallic lustre and changeable golden tints. They are +peculiar to the Cape district, but one species extends as far north as the +Mozambique territory. Their dentition is altogether peculiar, so as to +completely separate them from the true moles. + + +{190}FAMILY 21.--TALPIDÆ. (8 Genera, 19 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- --|-- -- 3 -- |-- -- -- -- + | | | | | + +The Moles comprise many extraordinary forms of small mammalia especially +characteristic of the temperate regions of the northern hemisphere, only +sending out a few species of _Talpa_ along the Himalayas as far as Assam, +and even to Tenasserim, if there is no mistake about this locality; while +one species is found in Formosa, the northern part of which is almost as +much Palæarctic as Oriental. The genus _Talpa_ (7 species), spreads over +the whole Palæarctic region from Great Britain to Japan; _Scaptochirus_ (1 +species) is a recent discovery in North China; _Condylura_ (1 species), the +star-nosed mole, inhabits Eastern North America from Nova Scotia to +Pennsylvania; _Scapanus_ (2 species) ranges across from New York to St. +Francisco; _Scalops_ (3 species), the shrew-moles, range from Mexico to the +great lakes on the east side of America, but on the west only to the north +of Oregon. An allied genus, _Myogale_ (2 species), has a curious +discontinuous distribution in Europe, one species being found in South-East +Russia, the other in the Pyrenees (Plate II., vol. i., p. 218). Another +allied genus, _Nectogale_ (1 species), has recently been described by +Professor Milne-Edwards from Thibet. _Urotrichus_ is a shrew-like mole +which inhabits Japan, and a second species has been discovered in the +mountains of British Columbia; an allied form, _Uropsilus_, inhabits East +Thibet. _Anurosorex_ and _Scaptonyx_, are new genera from North China. + +_Extinct Species._--The common mole has been found fossil in bone-caves and +diluvial deposits, and several extinct species of mole-like animals occur +in the Miocene deposits of the South of France and of Germany. These have +been described under the generic names _Dimylus_, _Geotrypus_, +_Hyporissus_, _Galeospalax_; while _Palæospalax_ has been found in the +Pliocene forest-beds of Norfolk {191}and Ostend. Species of _Myogale_ also +occur from the Miocene downwards. + + +FAMILY 22.--SORICIDÆ. (1 Genus, 11 Sub-genera, 65 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Shrews have a wide distribution, being found throughout every region +except the Australian and Neotropical; although, as a species is found in +Timor and in some of the Moluccas, they just enter this part of the former +region, while one found in Guatemala brings them into the latter. A number +of species have recently been described from India and the Malay Islands, +so that the Oriental region is now the richest in shrews, having 28 +species; the Nearctic comes next with 24; while the Ethiopian has 11, and +the Palæarctic 10 species. The sub-genera are _Crossopus_, _Amphisorex_, +_Neosorex_, _Crocidura_, _Diplomesodon_, _Pinulia_, _Pachyura_, _Blarina_, +_Feroculus_, _Anausorex_. + +_Extinct Species._--Several species of _Sorex_ have been found fossil in +the Miocene of the South of France, as well as the extinct genera +_Mysarachne_ and _Plesiosorex_; and some existing species have occurred in +Bone Caves and Diluvial deposits. + + +_General Remarks on the Distribution of the Insectivora._ + +The most prominent features in the distribution of the Insectivora +are,--their complete absence from South America and Australia; the presence +of _Solenodon_ in two of the West Indian islands while the five allied +genera are found only in Madagascar; and the absence of hedgehogs from +North America. If we consider that there are only 135 known species of the +order, 65 of which belong to the one genus _Sorex_; while the remaining 26 +genera contain only 70 species, which have to be classed in 8 distinct +families, and present such divergent and highly specialized forms as +_Galeopithecus_, _Erinaceus_, _Solenodon_, and _Condylura_, it becomes +evident that we have here the detached fragments of a much more +{192}extensive group of animals, now almost extinct. Many of the forms +continue to exist only in islands, removed from the severe competition of a +varied mammalian population, as in Madagascar and the Antilles; while +others appear to have escaped extermination either by their peculiar +habits--as the various forms of Moles; by special protection--as in the +Hedgehogs; or by a resemblance in form, coloration, and habits to dominant +groups in their own district--as the Tupaias of Malay which resemble +squirrels, and the Elephant-shrews of Africa which resemble the jerboas. +The numerous cases of isolated and discontinuous distribution among the +Insectivora, offer no difficulty from this point of view; since they are +the necessary results of an extensive and widely-spread group of animals +slowly becoming extinct, and continuing to exist only where special +conditions have enabled them to maintain themselves in the struggle with +more highly organized forms. + +The fossil Insectivora do not throw much light on the early history of the +order, since even as far back as the Miocene period they consist almost +wholly of forms which can be referred to existing families. In North +America they go back to the Eocene period, if certain doubtful remains have +been rightly placed. The occurrence of fossil Centetidæ in Europe, supports +the view we have maintained in preceding chapters, that the existing +distribution of this family between Madagascar and the Antilles, proves no +direct connection between those islands, but only shows us that the family +once had an extensive range. + + +_Order IV--CARNIVORA._ + +FAMILY 23.--FELIDÆ. (3 Genera, 14 Sub-genera, 66 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. --|1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Cats are very widely distributed over the earth--with the exception of +the Australian region and the island sub-region {193}of Madagascar and the +Antilles--universally; ranging from the torrid zone to the Arctic regions +and the Straits of Magellan. They are so uniform in their organization that +many naturalists group them all under one genus, _Felis_; but it is now +more usual to class at least the lynxes as a separate genus, while the +hunting leopard, or cheetah, forms another. Dr. J. E. Gray divides these +again, and makes 17 generic groups; but as this subdivision is not +generally adopted, and does not bring out any special features of +geographical distribution, I shall not further notice it. + +The genus _Felis_ (56 species) has the same general range as the whole +family, except that it does not go so far north; the Amoor river in Eastern +Asia, and 55° N. Lat. in America, marking its limits. _Lyncus_ (10 species) +is a more northern group, ranging to the polar regions in Europe and Asia, +and to Lat. 66° N. in America, but not going further south than Northern +Mexico and the European shores of the Mediterranean, except the caracal, +which may be another genus, and which extends to Central India, Persia, +North Africa and even the Cape of Good Hope. The lynxes are thus almost +wholly peculiar to the Nearctic and Palæarctic regions. _Cynælurus_ (1 +species) the hunting leopard, ranges from Southern and Western India +through Persia, Syria, Northern and Central Africa, to the Cape of Good +Hope. + +_Extinct Felidæ._--More than twenty extinct species of true Felidæ have +been described, ranging in time from the epoch of prehistoric man back to +the Miocene or even the Eocene period. They occur in the south of England, +in Central and South Europe, in North-West India, in Nebraska in North +America, and in the caves of Brazil. Most of them are referred to the genus +_Felis_, and closely resemble the existing lions, tigers, and other large +cats. Another group however forms the genus _Machairodus_, a highly +specialized form with serrated teeth. Five species have been described from +Europe, Northern India, and both North and South America; and it is +remarkable that they exhibit at least as wide a range, both in space and +time, as the more numerous species referred to _Felis_. One of them +undoubtedly coexisted {194}with man in England, while another, as well as +the allied _Dinictis_, has been found in the Mauvaises Terres of Nebraska, +associated with _Anchitherium_ and other extinct and equally remarkable +forms, which are certainly Miocene if not, as some geologists think, +belonging to the Eocene period. These facts clearly indicate that we have +as yet made little approach to discovering the epoch when Felidæ +originated, since the oldest forms yet discovered are typical and highly +specialized representatives of a group which is itself the most specialized +of the Carnivora. Another genus, _Pseudælurus_, is common to the Miocene +deposits of Europe and North America. + + +FAMILY 24.--CRYPTOPROCTIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 |-- -- -- --|-- -- -- -- + | | | | | + +The _Cryptoprocta ferox_, a small and graceful cat-like animal, peculiar to +Madagascar, was formerly classed among the Viverridæ, but is now considered +by Professor Flower to constitute a distinct family between the Cats and +the Civets. + + +FAMILY 25.--VIVERRIDÆ. (8-33 Genera, 100 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- |1. 2. 3. 4 |1. 2. 3. 4 |1 -- -- -- + | | | | | + +The Viverridæ comprise a number of small and moderate-sized carnivorous +animals, popularly known as civets, genets, and ichneumons, highly +characteristic of the Ethiopian and Oriental regions, several of the genera +being common to both. A species of _Genetta_, and one of _Herpestes_, +inhabit South Europe; while _Viverra_ extends to the Moluccas, but is +doubtfully indigenous. The extreme geographical limits of the family are +marked by {195}_Genetta_ in France and Spain, _Viverra_ in Shanghae and +Batchian Island, and _Herpestes_ in Java and the Cape of Good Hope. + +The following are the genera with their distribution as given by Dr. J. E. +Gray in his latest British Museum Catalogue: + +Sub-family VIVERRINÆ.--_Viverra_ (3 species), North and tropical Africa, +the whole Oriental region to the Moluccas; _Viverricula_ (1 species) India +to Java; _Genetta_ (5 species), South Europe, Palestine, Arabia, and all +Africa; _Fossa_ (1 species), Madagascar; _Linsang_ (2 species), Malacca to +Java; _Poiana_ (1 species), West Africa; _Galidia_ (3 species), Madagascar; +_Hemigalea_ (1 species), Malacca and Borneo; _Arctictis_ (1 species) Nepal +to Sumatra and Java; _Nandinia_ (1 species), West Africa; _Paradoxurus_ (9 +species), the whole Oriental region; _Paguma_ (3 species), Nepal to China, +Sumatra, and Borneo; _Arctogale_ (1 species), Tenasserim to Java. + +Sub-family HERPESTINÆ.--_Cynogale_ (1 species), Borneo; _Galidictis_ (2 +species), Madagascar; _Herpestes_ (22 species), South Palæarctic, +Ethiopian, and Oriental regions; _Athylax_ (3 species), Tropical and South +Africa; _Galogale_ (13 species), all Africa, North India, to Cambodja; +_Galerella_ (1 species), East Africa; _Calictis_ (1 species), Ceylon (?); +_Ariella_ (1 species), South Africa; _Ichneumia_ (4 species), Central, +East, and South Africa; _Bdeogale_ (3 species), West and East Africa; +_Urva_ (1 species), Himalayas to Aracan; _Tæniogale_ (1 species), Central +India; _Onychogale_ (1 species), Ceylon; _Helogale_ (2 species) East and +South Africa; _Cynictis_ (3 species), South Africa. + +Sub-family RHINOGALIDÆ.--_Rhinogale_ (1 species), East Africa; _Mungos_ (3 +species), all Africa; _Crossarchus_ (1 species), Tropical Africa; +_Eupleres_ (1 species), Madagascar; _Suricata_ (1 species), South Africa. + +_Fossil Viverridæ._--Several species of _Viverra_ and _Genetta_ have been +found in the Upper Miocene of France, and many extinct genera have also +been discovered. The most remarkable of these was _Ictitherium_, from the +Upper Miocene of Greece, which has also been found in Hungary, Bessarabia, +and France. Some of the species were larger than any living forms of +Viverridæ, and approached the hyænas. Other extinct genera are +_Thalassictis_ {196}and _Soricictis_ from the Upper Miocene, the former as +large as a panther; _Tylodon_, of small size, from the Upper Eocene; and +_Palæonyctis_ from the Lower Eocene, also small and showing a very great +antiquity for this family, if really belonging to it. + + +FAMILY 26.--PROTELIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- 3 -- |-- -- -- --|-- -- -- -- + | | | | | + + +The curious _Proteles_ or Aard-wolf, a highly-modified form of hyæna, +approaching the ichneumons, and feeding on white ants and carrion, is +peculiar to South Africa. + + +FAMILY 27.--HYÆNIDÆ. (1 Genus, 3 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- |1. 2. 3 -- |1 -- -- -- |-- -- -- -- + | | | | | + +The Hyænas are characteristically Ethiopian, to which region two of the +species are confined. The third, _Hyæna striata_, ranges over all the open +country of India to the foot of the Himalayas, and through Persia, Asia +Minor, and North Africa. Its fossil remains have been found in France. + +_Extinct Species._--The cave hyæna (_H. spelæa_) occurs abundantly in the +caverns of this country and of Central Europe, and is supposed to be most +nearly allied to the _H. crocuta_ of South Africa. Another species is found +in some parts of France. The earliest known true hyænas occur in the +Pliocene formation in France, in the Red Crag (Older Pliocene) of England, +and in the Upper Miocene of the Siwalik hills. In the Miocene period in +Europe, quite distinct genera are found, such as Hyænictis and _Lycæna_ +from the Upper Miocene of Greece; {197}_Ictitherium_, supposed to be +intermediate between Viverridæ and Hyænidæ; and _Thalassictis_, uniting the +weasels and hyænas. + + +FAMILY 28.--CANIDÆ. (3 Genera, 17 Sub-Genera, 54 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3. 4 |-- 2? -- -- + | | | | | + +The Canidæ, comprising the animals commonly known as dogs, wolves, and +foxes, have an almost universal range over the earth, being only absent +from the island sub-regions of Madagascar, the Antilles, Austro-Malaya, New +Zealand, and the Pacific Islands. With the exception of two remarkable +forms--the hyæna dog (_Lycaon picta_), and the great-eared fox (_Megalotis +Lalandei_), both from South Africa--all the species are usually placed in +the genus _Canis_, the distribution of which will be the same as that of +the family. Dr. J. E. Gray, in his arrangement of the family (Proc. Zool. +Soc., 1868), subdivides it into fifteen genera, the names and general +distribution of which are as follows:-- + +_Icticyon_ (1 species), Brazil; _Cuon_ (4 species), Siberia to Java; +_Lupus_ (5 species), North America, Europe, India to Ceylon; _Dieba_ (1 +species), North and West Africa; _Simenia_ (1 species), Abyssinia; +_Chrysocyon_ (2 species), North and South America; _Canis_ (4 species), +India, Australia (indigenous?) _Lycalopex_ (2 species), South America; +_Pseudalopex_ (5 species), South America and Falkland Islands; _Thous_ (2 +species), South America to Chili; _Vulpes_ (17 species), all the great +continents, except South America and Australia; _Fennecus_ (4 species), all +Africa; _Leucocyon_ (1 species), Arctic regions; _Urocyon_ (2 species), +North America; _Nyctereutes_ (1 species), Japan, Amoorland to Canton (Plate +III., vol. i. p. 226). These are all sub-genera according to Professor +Carus, except _Icticyon_. The same author makes Lycaon a sub-genus, while +Dr. Gray makes it a sub-family! + +_Extinct Species._--The dog, wolf, and fox, are found fossil in +{198}caverns in many parts of Europe, and several extinct species have been +found in Tertiary deposits in Europe, North India, and South America. Two +species have been found so far back as the Eocene of France, but the +fragments discovered are not sufficient to determine the characters with +any certainty. In North America, several species of _Canis_ occur in the +Pliocene of Nebraska and La Plata. The genus _Galecynus_, of the Pliocene +of Oeninghen, and _Palæocyon_, of the Brazilian caves, are supposed to +belong to the Canidæ. _Amphicyon_ abounded in the Miocene period, both in +Europe and North America; and some of the species were as large as a tiger. +Other extinct genera are, _Cynodictis_, _Cyotherium_, and _Galethylax_, +from the Eocene of France; _Pseudocyon_, _Simocyon_, and _Hemicyon_, from +the Miocene; but all these show transition characters to Viverridæ or +Ursidæ, and do not perhaps belong to the present family. + + +FAMILY 29.--MUSTELIDÆ. (21-28 Genera, 92 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 --|1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Mustelidæ constitute one of those groups which range over the whole of +the great continental areas. They may be divided into three +sub-families--one, the Mustelinæ, containing the weasels, gluttons, and +allied forms; a second, the Lutrinæ, containing the otters; and a third, +often considered a distinct family, the Melininæ, containing the badgers, +ratels, skunks, and their allies. + +In the first group (Mustelinæ) the genera _Martes_ and _Putorius_ (13 +species), range over all the Palæarctic region, and a considerable part of +the Oriental, extending through India to Ceylon, and to Java and Borneo. +Two species of _Martes_ (= _Mustela_ of Baird) occur in the United States. +The weasels, forming the genus _Mustela_ (20 species), have a still wider +range, extending into tropical Africa and the Cordilleras of Peru, but +{199}not going south of the Himalayas in India. The North American species +are placed in the genus _Putorius_ by Professor Baird. An allied genus, +_Gymnopus_ (4 species), is confined to the third and fourth Oriental +sub-regions. _Gulo_ (1 species), the glutton, is an arctic animal keeping +to the cold regions of Europe and Asia, and coming as far south as the +great lakes in North America. _Galictis_ (2 species), the grisons, are +confined to the Neotropical region. + +The Otters (Lutrinæ) range over the whole area occupied by the family. They +have been subdivided into a number of groups, such as _Barangia_ (1 +species), found only in Sumatra; _Lontra_, containing 3 South American +species; _Lutra_ (7 species), ranging over the whole of the Palæarctic and +Oriental regions; _Nutria_ (1 species), a sea-otter confined to the west +coast of America from California to Chiloe; _Lutronectes_ (1 species), from +Japan only; _Aonyx_ (5 species), found in West and South Africa, and the +third and fourth Oriental sub-regions. _Hydrogale_ (1 species), confined to +South Africa; _Latax_ (2 species), Florida and California to Canada and +British Columbia; _Pteronura_ (1 species), Brazil and Surinam; and +_Enhydris_ (1 species), the peculiar sea-otter of California, Kamschatka +and Japan. The last two are the only groups of otters, besides _Lutra_, +admitted by Professor Carus as genera. + +The Badgers and allies (Melininæ) have also a wide range, but with one +exception are absent from South America. They comprise the following +genera: _Arctonyx_ (1 species), Nepal to Aracan; _Meles_ (4 species), North +Europe to Japan, and China as far south as Hongkong (Plate I., vol. i., p. +195); _Taxidea_ (2 species), Central and Western North America to 58° N. +Lat.; _Mydaus_ (1 species), mountains of Java and Sumatra; _Melivora_ (3 +species), Tropical and South Africa and India to foot of Himalayas; +_Mephitis_ (12 species), America from Canada and British Columbia to the +Straits of Magellan (Plate XX., vol. ii., p. 136). _Ictonyx_ (2 species), +Tropical Africa to the Cape; _Helictis_ (4 species), Nepal to Java, Formosa +and Shanghai (Plate VII., vol. i. p. 331). + +_Fossil Mustelidæ._--Species of otter, weasel, badger, and glutton, occur +in European bone caves and other Post-tertiary deposits; and in North +America _Galictis_, now found only in the Neotropical region, and, with +_Mephitis_, occurring in Brazilian caves. + +{200}Species of _Mustela_ have been found in the Pliocene of France and of +South America; and _Lutra_ in the Pliocene of North America. + +In the Miocene deposits of Europe several species of _Mustela_ and _Lutra_ +have been found; with the extinct genera _Taxodon_, _Potamotherium_, and +_Palæomephitis_; as well as _Promephitis_ in Greece. + +In the Upper Miocene of the Siwalik Hills species of _Lutra_ and +_Mellivora_ are found, as well as the extinct genera _Enhydrion_ and +_Ursitaxus_. + +The family appears to have been unknown in North America during the Miocene +period. + + +FAMILY 30.--PROCYONIDÆ. (4 Genera, 8 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |1. 2. 3. 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Procyonidæ are a small, but very curious and interesting family of +bear-like quadrupeds, ranging from British Columbia and Canada on the +north, to Paraguay and the limits of the tropical forests on the south. + +The Racoons, forming the genus _Procyon_, are common all over North +America; a well-marked variety or distinct species inhabiting the west +coast, and another, most parts of South America. The genus _Nasua_, or the +coatis (5 species?), extends from Mexico and Guatemala to Paraguay. The +curious arboreal prehensile-tailed kinkagou (_Cercoleptes candivolvus_) is +also found in Mexico and Guatemala, and in all the great forests of Peru +and North Brazil. _Bassaris_ (2 species), a small weasel-like animal with a +banded tail, has been usually classed with the Viverridæ or Mustelidæ, but +is now found to agree closely in all important points of internal structure +with this family. It is found in California, Texas, and the highlands of +Mexico, and belongs therefore as much to the Nearctic as to the Neotropical +region. A second species has recently been described by Professor Peters +{201}from Coban in Guatemala, in which, country it has also been observed +by Mr. Salvin. + +_Fossil Procyonidæ._--A species of _Nasua_ has been found in the bone caves +of Brazil, and a _Procyon_ in the Pliocene or Post-pliocene deposits of +Illinois and Carolina. + + +FAMILY 31.--ÆLURIDÆ. (2 Genera, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- 4 |-- -- -- --|-- -- 3 -- |-- -- -- -- + | | | | | + +The Panda (_Ælurus fulgens_), of the forest regions of the Eastern +Himalayas and East Thibet, a small cat-like bear, has peculiarities of +organization which render it necessary to place it in a family by itself. +(Plate VII. vol. i. p. 331). An allied genus, _Æluropus_, a remarkable +animal of larger size and in colour nearly all white, has recently been +described by Professor Milne-Edwards, from the mountains of East Thibet; so +that the family may be said to inhabit the border lands of the Oriental and +Palæarctic regions. These animals have their nearest allies in the coatis +and bears. + + +FAMILY 32.--URSIDÆ. (5 Genera, or Sub-genera, 15 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1 -- -- -- |1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- --|1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Bears have a tolerably wide distribution, although they are entirely +absent from the Australian and Ethiopian, and almost so from the +Neotropical region, one species only being found in the Andes of Peru and +Chili. They comprise the following groups, some of which are doubtfully +ranked as genera. + +_Thalassarctos_, the polar bear (1 species) inhabiting the Arctic regions; +_Ursus_, the true bears (12 species), which range over {202}all the +Nearctic and Palæarctic regions as far as the Atlas Mountains, the +Indo-Chinese sub-region in the mountains, and to Hainan and Formosa; +_Helarctos_, the Malay or sun-bear (1 species) confined to the Indo-Malayan +sub-region; _Melursus_ or _Prochilus_, the honey-bear (1 species), confined +to the first and second Oriental sub-regions, over which it ranges from the +Ganges to Ceylon; and _Tremarctos_, the spectacled bear--commonly known as +_Ursus ornatus_--which is isolated in the Andes of Peru and Chili, and +forms a distinct group. + +_Fossil Ursidæ._--Two bears (_Ursus spelæus_ and _U. priscus_) closely +allied to living species, abound in the Post-tertiary deposits of Europe; +and others of the same age are found in North America, as well as an +extinct genus, _Arctodus_. + +_Ursus arvernensis_ is found in the Pliocene formation of France, and the +extinct genus _Leptarchus_ in that of North America. + +Several species of _Amphicyon_, which appears to be an ancestral form of +this family, are found in the Miocene deposits of Europe and N. India; +while _Ursus_ also occurs in the Siwalik Hills and Nerbudda deposits. + + +FAMILY 33.--OTARIIDÆ (4 Genera, 8 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1 -- -- -- | 1 -- -- 4 |-- -- -- --|-- -- 3 -- |-- -- -- --|-- 2. 3 -- + | | | | | + +The Otariidæ, or Eared Seals, comprehending the sea-bears and sea-lions, +are confined to the temperate and cold shores of the North Pacific, and to +similar climates in the Southern Hemisphere, where the larger proportion of +the species are found. They are entirely absent from the North Atlantic +shores. Mr. J. A. Allen, in his recent discussion of this family (Bull. +Harvard Museum) divides them into the following genera:-- + +_Otaria_ (1 species), Temperate South America, from Chili to La Plata; +_Callorhinus_ (1 species), Behring's Straits and Kamschatka; +_Arctocephalus_ (3 species), temperate regions of the {203}Southern +Hemisphere; _Zalophus_ (2 species), North Pacific, from California to +Japan, and the shores of Australia and New Zealand; _Eumetopias_ (1 +species), Behring's Straits and California. + +_Fossil Otariidæ._--Remains supposed to belong to this family have been +found in the Miocene of France. + + +FAMILY 34.--TRICHECHIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- 4 | 1 -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Morse, or Walrus (_Trichecus rosmarus_), which alone constitutes this +family, is a characteristic animal of the North Polar regions, hardly +passing south of the Arctic circle except on the east and west coasts of +North America, where it sometimes reaches Lat. 60°. It is most abundant on +the shores of Spitzbergen, but is not found on the northern shores of Asia +between Long. 80° and 160° E., or on the north shores of America from 100° +to 150° west. + +Its remains have been found fossil in Europe as far south as France, and in +America as far as Virginia; but the small fragments discovered may render +the identification uncertain. + + +FAMILY 35.--PHOCIDÆ. (13 Genera, 21 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1 -- -- 4? | 1 -- -- 4 |1. 2. 3. 4 |-- -- -- --|-- -- -- --|-- 2. 3 -- + | | | | | + +The earless or true Seals are pretty equally divided between the Northern +and Southern Hemispheres, frequenting almost exclusively the temperate and +cold regions, except two species said to occur among the West Indian +islands. The genus _Phoca_ and its close allies, as well as _Halichoerus_ +and _Pelagius_, are {204}northern; while _Stenorhynchus_ and _Morunga_, +with their allies, are mostly southern. The genera admitted by Dr. Gray in +his catalogue are as follows:-- + +_Callocephalus_ (3 species), Greenland, North Sea, also the Caspian Sea, +and Lakes Aral and Baikal; _Pagomys_ (2 species), North Sea, North Pacific, +and Japan; _Pagophilus_ (2 species), North Pacific and North Atlantic; +_Halicyon_ (1 species), North West coast of America; _Phoca_ (2 species), +North Atlantic and North Pacific, Japan; _Halichoerus_ (1 species), +Greenland, North Sea, and Baltic; _Pelagius_ (2 species), Madeira, +Mediterranean, Black Sea; _Stenorhynchus_ (1 species), Antarctic Ocean, +Falkland Islands, New Zealand; _Lobodon_ (1 species), Antarctic Ocean; +_Leptonyx_ (1 species), Antarctic Ocean, South Australia, East Patagonia; +_Ommatophoca_ (1 species), Antarctic Ocean; _Morunga_ (2 species), +California, Falkland Islands, Temperate regions of Southern Ocean; +_Cystophora_ (2 species), North Atlantic, Antilles. + +_Fossil Seals._--Remains of living species of seals have been found in +Post-tertiary deposits in many parts of Europe and in Algeria, as well as +in New Zealand. _Pristiphoca occitana_ is a fossil seal from the Pliocene +of Montpellier, while a species of _Phoca_ is said to have been found in +the Miocene deposits of the United States. + + +_General Remarks on the Distribution of the Carnivora._ + +_Terrestrial Carnivora._--For the purposes of geographical distribution, +the terrestrial and aquatic Carnivora differ too widely to be considered in +one view, their areas being limited by barriers of a very different nature. +The terrestrial Carnivora form a very extensive and considerably varied +group of animals, having, with the doubtful exception of Australia, a +world-wide distribution. Yet the range of modification of form is not very +great, and the occurrence of three families consisting of but one species +each, is an indication of a great amount of recent extinction. One of the +most marked features presented by this group is its comparative scarcity in +the Neotropical region, only four families being represented there (not +counting the Ursidæ, which has only one Andean species), and both genera +and species are few in number. Even the Procyonidæ, which are especially +South {205}American, have but two genera and six species in that vast area. +We might therefore, from these considerations alone, conclude that +Carnivora are a development of the northern hemisphere, and have been +introduced into the Neotropical region at a comparatively recent epoch. The +claim of the Nearctic region to be kept distinct from the Palæarctic (with +which some writers have wished to unite it) is well maintained by its +possession of at least six species of _Mephitis_, or skunk, a group having +no close allies in any other region,--and the genera _Procyon_ and +_Bassaris_,--for the latter, ranging from the high lands of Guatemala and +Mexico to Texas and California, may be considered a Nearctic rather than a +Neotropical form. In the other families, the most marked feature is the +total absence of Ursidæ from the Ethiopian region. The great mass of the +generic forms of Carnivora, however, are found in the Oriental and +Ethiopian regions, which possess all the extensive group of Viverridæ +(except a few species in the fourth Palæarctic sub-region) and a large +number of Felidæ and Mustelidæ. + +_Aquatic Carnivora._--The aquatic Carnivora present no very marked features +of distribution, except their preference for cold and temperate rather than +tropical seas. Their nearest approximation to the terrestrial group, is +supposed to be that of the Otariidæ to the Ursidæ; but this must be very +remote, and the occurrence of both seals and bears in the Miocene period, +shows, that until we find some late Secondary or early Tertiary formation +rich in Mammalian remains, we are not likely to get at the transition forms +indicating the steps by which the aquatic Carnivora were developed. The +most interesting special fact of distribution to be noticed, is the +occurrence of seals, closely allied to those inhabiting the northern seas, +in the Caspian, Lake Aral, and Lake Baikal. In the case of the two +first-named localities there is little difficulty, as they are connected +with the North Sea by extensive plains of low elevation, so that a +depression of less than 500 feet would open a free communication with the +ocean. At a comparatively recent epoch, a great gulf of the Arctic ocean +must have occupied the valley of the Irtish, and extended to the Caspian +Sea; till the elevation of the Kirghiz Steppes cut off the +{206}communication with the ocean, leaving an inland sea with its seals. +Lake Baikal, however, offers much greater difficulties; since it is not +only a fresh-water lake, but is situated in a mountain district nearly +2,000 feet above the sea level, and entirely separated from the plains by +several hundred miles of high land. It is true that such an amount of +submergence and elevation is known to have occurred in Europe so recently +as during the Glacial period; but Lake Baikal is so surrounded by +mountains, that it must at that time have been filled with ice, if at +anything like its present elevation. Its emergence from the sea must +therefore have taken place since the cold epoch, and this would imply that +an enormous extent of Northern Asia has been very recently under water. + +We are accustomed to look on Seals as animals which exclusively inhabit +salt water; but it is probably from other causes than its saltness that +they usually keep to the open sea, and there seems no reason why +fresh-water should not suit them quite as well, provided they find in it a +sufficiency of food, facilities for rearing their young, and freedom from +the attacks of enemies. As already remarked in vol. i. p. 218, Mr. Belt's +ingenious hypothesis (founded on personal examination of the Siberian +Steppes), that during the Glacial period the northern ice-cap dammed up the +waters of the northward flowing Asiatic rivers, and thus formed a vast +fresh-water lake which might have risen as high as Lake Baikal, seems to +offer the best solution of this curious problem of distribution. + +_Range of Carnivora in Time._--Carnivora have been found in all the +Tertiary deposits, and comprise a number of extinct genera and even +families. Several genera of Canidæ occur in the Upper Eocene of Europe; but +the most remarkable fact is, that even in the Lower Eocene are found two +well-marked forms, _Palæonyctis_, one of the Viverridæ, and _Arctocyon_, +forming a distinct family type of very generalized characters, but +unmistakably a carnivore. This last has been found at La Fère, in the +north-east of France, in a deposit which, according to M. Gaudry, is the +very lowest of the Lower Eocene formation in Europe. _Arctocyon_ is +therefore one of the oldest, if not the very oldest, of the higher forms of +mammal yet discovered. + + +{207}_Order V.--CETACEA._ + + +FAMILY 36.--BALÆNIDÆ. (6 Genera, 14 Species.) + +GENERAL DISTRIBUTION.--Temperate and Cold Seas of both Northern and +Southern Hemispheres. + +This family comprises the whalebone or "right" whales, the best known +species being the Greenland whale (_Balæna mysticetus_). Allied species are +found in all parts of the southern seas, as far north as the Cape of Good +Hope; while some of the northern species are found off the coast of Spain, +and even enter the Mediterranean. As most of the species indicated are +imperfectly known, and their classification by no means well settled, no +useful purpose will be served by enumerating the genera or sub-genera. + + +FAMILY 37.--BALÆNOPTERIDÆ. (9 Genera, 22 Species.) + +GENERAL DISTRIBUTION.--Cold and Temperate Seas of both Hemispheres. + +This family comprises the finner whales and rorquals, and are characterised +by possessing a dorsal fin and having the baleen or whalebone less +developed. They are abundant in all northern seas, less so in the southern +hemisphere, but they seem occasionally to enter the tropical seas. The best +known genera are _Megaptera_ (7 species); _Physalus_ (4 species); and +_Balænoptera_ (2 species); all of which have species in the North Sea. + + +FAMILY 38.--CATODONTIDÆ. (4 Genera, or Sub-Genera, 6 Species.) + +GENERAL DISTRIBUTION.--All the Tropical Oceans, extending north and south +into Temperate waters. + +This family, comprising the cachalots or sperm whales, and black-fish, are +separated from the true whales by having teeth in the lower jaw and no +whalebone. They are pre-eminently a tropical, as distinguished from the two +preceding which are {208}arctic and antarctic families. The spermaceti +whale (_Catodon macrocephalus_) abounds in the Pacific Ocean and in the +deep Moluccan Sea, and also in the Indian Ocean and the Mozambique Channel. +In the Atlantic it is scarce, although it occasionally comes north as far +as our shores. + +The genera of Catodontidæ as given by Dr. Gray are, _Catodon_ (2 species?), +Warm Eastern Oceans; _Physeter_ (1 species), "the black fish," North Sea; +_Cogia_ (2 species), South Temperate Oceans; _Euphysetes_ (1 species), +Coast of Australia. + + +FAMILY 39.--HYPEROODONTIDÆ. (9 Genera or Sub-Genera, 12 Species.) + +GENERAL DISTRIBUTION.--Atlantic, Mediterranean, Indian Ocean, and Southern +Ocean. + +This family consists of the beaked whales, which have no permanent teeth in +the upper jaw. The genera, according to Dr. Gray, are, _Hyperoodon_ (2 +species) "bottle-nosed whales," North Sea; _Lagenocetus_ (1 species), North +Sea; _Epiodon_ (2 species), North and South Atlantic; _Petrorhynchus_ (2 +species), Mediterranean Sea and Southern Ocean; _Berardius_ (1 species), +New Zealand; _Xiphius_ (1 species) North Atlantic; _Dolichodon_ (1 +species), Cape of Good Hope; _Neoziphius_ (1 species) Mediterranean; +_Dioplodon_ (1 species), Indian Ocean. + + +FAMILY 40.--MONODONTIDÆ. (1 Genus, 1 Species.) + +The "Narwhal" (_Monodon monoceros_) which constitutes this family, is +placed by Dr. Gray along with the "white whales," in his family Belugidæ. +It inhabits the North Sea. + + +FAMILY 41.--DELPHINIDÆ. (24 Genera or Sub-Genera, 100 Species.) + +GENERAL DISTRIBUTION.--All Oceans, Seas, and Great Rivers of the globe. + +This family, including the Porpoises, Dolphins, White Whales, &c., may be +described as small, fish-shaped whales, having teeth {209}in both jaws. +According to Dr. Gray they form seven families and 24 genera; according to +Professor Carus, four sub-families and 8 genera, but as these groups appear +to be established on quite different principles, and often differ widely +from each other, I shall simply enumerate Dr. Gray's genera with their +distribution as given in his British Museum Catalogue. + +_Platanista_ (2 species), long-snouted porpoises, inhabiting the Ganges and +Indus; _Inia_ (1 species), a somewhat similar form, inhabiting the upper +waters of the Amazonian rivers: _Steno_ (8 species), Indian Ocean, Cape of +Good Hope, and West Pacific; _Sotalia_ (1 species), Guiana; _Delphinus_ (10 +species), all the oceans; _Clymenia_ (14 species), all the oceans; +_Delphinapterus_ (1 species), South Atlantic; _Tursio_ (7 species), +Atlantic and Indian Oceans; _Eutropia_ (2 species), Chili, and Cape of Good +Hope; _Electra_, (8 species), all the oceans; _Leucopleurus_ (1 species), +North Sea; _Lagenorhynchus_ (1 species), North Sea; _Pseudorca_ (2 +species), North Sea, Tasmania; _Orcaella_ (2 species), Ganges; +_Acanthodelphis_ (1 species), Brazil; _Phocæna_ (2 species), North Sea; +_Neomeris_ (1 species), India; _Grampus_ (3 species), North Sea, +Mediterranean, Cape of Good Hope; _Globiocephalus_ (14 species), all the +oceans; _Sphærocephalus_ (1 species), North Atlantic; _Orca_ (9 species), +Northern and Southern Oceans; _Ophysia_ (1 species), North Pacific; +_Beluga_ (6 species), Arctic Seas, Australia; _Pontoporia_ (1 species), +Monte Video. + + +_Fossil Cetacea._ + +Remains of Cetacea are tolerably abundant in Tertiary deposits, both in +Europe and North America. In the Lower Pliocene of England, France, and +Germany, extinct species of five or six living genera of whales and +dolphins have been found; and most of these occur also in the Upper +Miocene, along with many others, referred to about a dozen extinct genera. + +In the Post-pliocene deposits of Vermont and South Carolina, several +extinct species have been found belonging to living genera; but in the +Miocene deposits of the Eastern United States cetacean remains are much +more abundant, more than 30 species of {210}extinct whales and dolphins +having been described, most of them belonging to extinct genera. + +The Zeuglodontidæ, an extinct family of carnivorous whales, with +double-fanged serrated molar teeth, whose affinities are somewhat doubtful, +are found in the older Pliocene of Europe, and in the Miocene and Eocene of +the Eastern United States. _Zeuglodon_ abounds in the United States, and +one species reached a length of seventy feet. A species of this genus is +said to have been found in Malta. _Squalodon_ occurs in Europe and North +America; and in the latter country four or five other genera have been +described, of which one, _Saurocetes_, has been found also at Buenos Ayres. + + +_Order VI.--SIRENIA._ + +FAMILY 42.--MANATIDÆ. (3 Genera, 5 Species?) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- 4 |-- -- -- --|1. -- 3 -- |1. 2 -- -- |1. 2. -- 4 |1 -- -- -- + | | | | | + +The Sea-cows are herbivorous aquatic animals living on the coasts or in the +great rivers of several parts of the globe. _Manatus_ (2 species) inhabits +both shores of the Atlantic, one species ranging from the Gulf of Mexico to +North Brazil, and ascending the Amazon far into the interior of the +continent; while the other is found on the west coast of Africa. _Halicore_ +(2 species?), the Dugong, is peculiar to the Indian Ocean, extending from +Mozambique to the Red Sea, thence to Western India and Ceylon, the Malay +Archipelago and the north coast of Australia. _Rytina_ (1 species), +supposed to be now extinct, inhabited recently the North Pacific, between +Kamschatka and Behring's Straits. + +_Fossil Sirenia._--Extinct species of _Manatus_ have been found in the +Post-pliocene deposits of Eastern North America from {211}Maryland to +Florida; and an extinct genus, _Prorastomus_, in some Tertiary deposits in +the Island of Jamaica. + +In Post-pliocene deposits in Siberia, remains of _Rytina_ have been found; +while several species of the extinct genus _Halitherium_, perhaps +intermediate between _Manatus_ and _Halicore_, have been found in the older +Pliocene and Upper Miocene of France and Germany. + + +_Order VII.--UNGULATA._ + +FAMILY 43.--EQUIDÆ. (1 Genus, 8 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + LIVING SPECIES. + | | | | | + -- -- -- --|-- -- -- --|-- 2. 3 -- |1. 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + EXTINCT SPECIES. + | | | | | + 1. 2 -- -- |1. 2. 3 -- |1. 2. 3. 4 |-- -- -- --| 1 -- 3 -- |-- -- -- -- + | | | | | + +The Horses, Asses, and Zebras form a highly specialized group now confined +to the Ethiopian and Palæarctic regions, but during the middle and later +tertiaries having a very extensive range. The zebras (3 species) inhabit +the greater part of the Ethiopian region, while the asses (4 species) are +characteristic of the deserts of the Palæarctic region from North Africa +and Syria to Western India, Mongolia, and Manchuria. The domestic horse is +not known in a wild state, but its remains are found in recent deposits +from Britain to the Altai Mountains, so that its disappearance is probably +due to human agency. + +_Extinct Equidæ._--Extinct forms of this family are very numerous. The +genus _Equus_ occurs in Post-pliocene and Pliocene deposits in Europe, +North America, and South America. In North America the species are most +numerous. An allied genus _Hipparion_, having rudimentary lateral toes, is +represented {212}by several species in the Pliocene of North America, while +in Europe it occurs both in the Older Pliocene and Upper Miocene. Various +other allied forms, in which the lateral toes are more and more developed, +and most of which are now classed in a distinct family, Anchitheridæ, range +back through the Miocene to the Eocene period. A sufficient account of +these has already been given in vol. i. chap. vi. p. 135, to which the +reader is referred for the supposed origin and migrations of the horse. + + +Family 44--TAPIRIDÆ. (2 Genera? 6 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 |-- -- -- -- + | | | | | + +The Tapirs form a small group of animals whose discontinuous distribution +plainly indicates their approaching extinction. For a long time only two +species were known, the black American, and the white-banded Malay tapir, +the former confined to the equatorial forests of South America, the latter +to the Malay peninsula, Sumatra, and Borneo (Plate VIII. vol. i. p. 337). +Lately however another, or perhaps two distinct species (or according to +Dr. J. E. Gray, four!) have been discovered in the Andes of New Granada and +Ecuador, at an elevation of from 8,000 to 12,000 feet; while one or perhaps +two more, forming the allied genus _Elasmognathus_, have been found to +inhabit Central America from Panama to Guatemala. + +_Extinct Tapirs._--True tapirs inhabited Western Europe, from the latest +Pliocene back to the earliest Miocene times; while they only occur in +either North or South America in the Post-pliocene deposits and caves. The +singular distribution of the living species is thus explained, since we see +that they are an Old World group which only entered the American continent +at a comparatively recent epoch. An ancestral form of this +group--_Lophiodon_--is found in Miocene and Eocene deposits of {213}Europe +and North America; while a still more ancient form of large size is found +in the Lower Eocene of France and England, indicating an immense antiquity +for this group of Mammalia. There are many other extinct forms connecting +these with the Palæotheridæ, already noticed in chapter vi. (vol. i. pp. +119-125). + + +FAMILY 45.--RHINOCEROTIDÆ. (1 Genus, 9 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + LIVING SPECIES. + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- |-- -- 3. 4 |-- -- -- -- + | | | | | + EXTINCT SPECIES. + | | | | | + -- -- -- --|1. 2 -- -- |1. 2. 3. 4 |-- -- -- --| 1 -- 3 -- |-- -- -- -- + | | | | | + +Living Rhinoceroses are especially characteristic of Africa, with Northern +and Malayan India. Four or perhaps five species, all two-horned, are found +in Africa, where they range over the whole country south of the desert to +the Cape of Good Hope. In the Oriental region there are also four or five +species, which range from the forests at the foot of the Himalayas +eastwards through Assam, Chittagong, and Siam, to Sumatra, Borneo and Java. +Three of these are one-horned, the others found in Sumatra, and northwards +to Pegu and Chittagong, two-horned. The Asiatic differ from the African +species in some dental characters, but they are in other respects so much +alike that they are not generally considered to form distinct genera. In +his latest catalogue however (1873), Dr. Gray has four genera, _Rhinoceros_ +(4 species), and _Ceratorhinus_ (2 species), Asiatic; _Rhinaster_ (2 +species), and _Ceratotherium_ (2 species), African. + +_Extinct Rhinocerotidæ._--Numerous species of _Rhinoceros_ ranged over +Europe and Asia from the Post-pliocene back to the Upper Miocene period, +and in North America during the Pliocene period {214}only. The hornless +_Acerotherium_ is Miocene only, in both countries. Other genera are +_Leptodon_ from Greece, and _Hyracodon_ from Nebraska, both of Miocene age. +More than 20 species of extinct rhinoceroses are known, and one has even +been found at an altitude of 16,000 feet in Thibet. + + +FAMILY 46.--HIPPOPOTAMIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + LIVING SPECIES. + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + EXTINCT SPECIES. + | | | | | + -- -- -- --|-- -- -- --|1. 2 -- -- |-- -- -- --| 1 -- 3 -- |-- -- -- -- + | | | | | + +The Hippopotamus inhabits all the great rivers of Africa; a distinct +species of a smaller size being found on the west coast, and on some of the +rivers flowing into Lake Tchad. + +_Fossil Hippopotami._--Eight extinct species of _Hippopotamus_ are known +from Europe and India, the former Post-pliocene or Pliocene, the latter of +Upper Miocene age. They ranged as far north as the Thames valley. An +extinct genus from the Siwalik Hills, _Merycopotamus_, according to Dr. +Falconer connects _Hippopotamus_ with _Anthracotherium_, an extinct form +from the Miocene of Europe, allied to the swine. + + +FAMILY 47.--SUIDÆ. (5 Genera, 22 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- 2. 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. -- -- -- + | | | | | + +The Swine may be divided into three well-marked groups, from peculiarities +in their dentition. 1. The Dicotylinæ, or {215}peccaries (1 genus, +_Dicotyles_). These offer so many structural differences that they are +often classed as a separate family. 2. The true swine (3 genera, _Sus_, +_Potamochoerus_, and _Babirusa_); and, 3. The Phacochoerinæ, or wart hogs +(1 genus, _Phacochoerus_). These last are also sometimes made into a +separate family, but they are hardly so distinct as the Dicotylinæ. + +The Peccaries (2 species), are peculiar to the Neotropical region, +extending from Mexico to Paraguay. They also spread northwards into Texas, +and as far as the Red River of Arkansas, thus just entering the Nearctic +region; but with this exception swine are wholly absent from this region, +forming an excellent feature by which to differentiate it from the +Palæarctic. + +_Sus_ (14 species), ranges over the Palæarctic and Oriental regions and +into the first Australian sub-region as far as New Guinea; but it is absent +from the Ethiopian region, or barely enters it on the north-east. +_Potamochoerus_ (3 species?), is wholly Ethiopian (Plate V. vol. i. p. +278). _Babirusa_ (1 species), is confined to two islands, Celebes and +Bouru, in the first Australian sub-region. + +_Phacochoerus_ (2 species), ranges over tropical Africa from Abyssinia to +Caffraria. + +Dr. J. E. Gray divides true swine (_Sus_) into 7 genera, but it seems far +better to keep them as one. + +_Fossil Suidæ._--These are very numerous. Many extinct species of wild hog +(_Sus_), are found in Europe and North India, ranging back from the +Post-pliocene to the Upper Miocene formations. In the Miocene of Europe are +numerous extinct genera, _Bothriodon_, _Anthracotherium_, _Palæochoerus_, +_Hyotherium_, and some others; while in the Upper Eocene occur +_Cebochoerus_, _Choeropotamus_, and _Acotherium_,--these early forms having +more resemblance to the peccaries. + +None of these genera are found in America, where we have the living genus +_Dicotyles_ in the Post-pliocene and Pliocene deposits, both of North and +South America; with a number of extinct genera in the Miocene. The chief of +these are, _Elotherium_, _Perchoerus_, _Leptochoerus_, and _Nanohyus_, all +from Dakota, and _Thinohyus_, from Oregon. One extinct genus, _Platygonus_, +closely allied to _Dicotyles_, is found in the Post-pliocene of Nebraska, +{216}Oregon, and Arkansas. _Elotherium_ is said to be allied to the peccary +and hippopotamus. _Hyopotamus_, from the Miocene of Dakota, is allied to +_Anthracotherium_, and forms with it (according to Dr. Leidy) a distinct +family of ancestral swine. + +It thus appears, that the swine were almost equally well represented in +North America and Europe, during Miocene and Pliocene times, but by +entirely distinct forms; and it is a remarkable fact that these hardy +omnivorous animals, should, like the horses, have entirely died out in +North America, except a few peccaries which have preserved themselves in +the sub-tropical parts and in the southern continent, to which they are +comparatively recent emigrants. We can hardly have a more convincing proof +of the vast physical changes that have occurred in the North American +continent during the Pliocene and Post-pliocene epochs, than the complete +extinction of these, along with so many other remarkable types of Mammalia. + +According to M. Gaudry, the ancestors of all the swine, with the +hippopotami and extinct _Anthracotherium_, _Merycopotamus_, and many allied +forms,--are the _Hyracotherium_ and _Pliolophus_, both found only in the +London clay belonging to the Lower Eocene formation. + + +FAMILY 48.--CAMELIDÆ. (2 Genera, 6 Species). + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + LIVING SPECIES. + | | | | | + 1. -- -- --|-- -- -- --|-- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + EXTINCT SPECIES. + | | | | | + 1. -- -- --|-- 2. 3. 4 |-- -- 3 -- |-- -- -- --|-- -- 3. --|-- -- -- -- + | | | | | + +The Camels are an exceedingly restricted group, the majority of the species +now existing only in a state of domestication. The genus _Camelus_ (2 +species), is a highly characteristic desert form {217}of the Palæarctic +region, from the Sahara to Mongolia as far as Lake Baikal. _Auchenia_ (4 +species), comprehending the Llamas and Alpacas, is equally characteristic +of the mountains and deserts of the southern part of South America. Two +species entirely domesticated inhabit the Peruvian and Bolivian Andes; and +two others are found in a wild state, the vicuna in the Andes of Peru and +Chili (Plate XVI. vol. ii. p. 40), and the guanaco over the plains of +Patagonia and Tierra del Fuego. + +_Extinct Camelidæ._--No fossil remains of camels have been found in Europe, +but one occurs in the deposits of the Siwalik Hills, usually classed as +Upper Miocene, but which some naturalists think are more likely of Older +Pliocene age. _Merycotherium_, teeth of which have been found in the +Siberian drift, is supposed to belong to this family. + +In North America, where no representative of the family now exists, the +camel-tribe were once abundant. In the Post-pliocene deposits of California +an _Auchenia_ has been found, and in those of Kansas one of the extinct +genus _Procamelus_. In the Pliocene period, this genus, which was closely +allied to the living camels, abounded, six or seven species having been +described from Nebraska and Texas, together with an allied form +_Homocamelus_. In the Miocene period different genera +appear,--_Poebrotherium_, and _Protomeryx_,--while a _Procamelus_ has been +found in deposits of this age in Virginia. + +In South America a species of _Auchenia_ has been found in the caves of +Brazil, and others in the Pliocene deposits of the pampas, together with +two extinct genera, _Palæolama_ and _Camelotherium_. + +We thus find the ancestors of the Camelidæ in a region where they do not +now exist, but which is situated so that the now widely separated living +forms could easily have been derived from it. This case offers a remarkable +example of the light thrown by palæontology on the distribution of living +animals; and it is a warning against the too common practice of assuming +the direct land connection of remote continents, in order to explain +similar instances of discontinuous distribution to that of the present +family. + + +{218}FAMILY 49.--TRAGULIDÆ. (2 Genera, 6 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- |1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Tragulidæ are a group of small, hornless, deer-like animals, with tusks +in the upper jaw, and having some structural affinities with the camels. +The musk-deer was formerly classed in this family, which it resembles +externally; but a minute examination of its structure by M. Milne-Edwards, +has shown it to be more nearly allied to the true deer. The Chevrotains, or +mouse-deer, _Tragulus_ (5 species), range over all India to the foot of the +Himalayas and Ceylon, and through Assam, Malacca, and Cambodja, to Sumatra, +Borneo, and Java (Plate VIII., vol. i. p. 337). _Hyomoschus_ (1 species), +is found in West Africa. + +_Extinct Tragulidæ._--A species of _Hyomoschus_ is said to have been found +in the Miocene of the South of France, as well as three extinct genera, +_Dremotherium_ (also found in Greece), with _Lophiomeryx_ from the Upper +Miocene, said to be allied to _Tragulus_; and _Amphitragulus_ from the +Lower Miocene, of more remote affinities, and sometimes placed among the +Deer. There seems to be no doubt, however, that this family existed in +Europe in Miocene times; and thus another case of discontinuous +distribution is satisfactorily accounted for. + + +FAMILY 50.--CERVIDÆ. (8 Genera, 52 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- --|1. 2. 3. 4 |1 -- -- -- + | | | | | + +The Cervidæ, or deer tribe, are an extensive group of animals equally +adapted for inhabiting forests or open plains, the Arctic {219}regions or +the Tropics. They range in fact over the whole of the great continents of +the globe, with the one striking exception of Africa, where they are only +found on the shores of the Mediterranean which form part of the Palæarctic +region. The following is the distribution of the genera. + +_Alces_ (1 species), the elk or moose, ranges all over Northern Europe and +Asia, as far south as East Prussia, the Caucasus, and North China; and over +Arctic America to Maine on the East, and British Columbia on the west. The +American species may however be distinct, although very closely allied to +that of Europe. _Tarandus_ (1 species), the reindeer, has a similar range +to the last, but keeps farther north in Europe, inhabiting Greenland and +Spitzbergen; and in America extends farther south, to New Brunswick and the +north shore of Lake Superior. There are several varieties or species of +this animal confined to special districts, but they are not yet well +determined. _Cervus_ (40 species), the true deer, have been sub-divided +into numerous sub-genera characteristic of separate districts. They range +over the whole area of the family, except that they do not go beyond 57° N. +in America and a little further in Europe and Asia. In South America they +extend over Patagonia and even to Tierra del Fuego. They are found in the +north of Africa, and over the whole of the Oriental region, and beyond it +as far as the Moluccas and Timor, where however they have probably been +introduced by man at an early period. _Dama_ (1 species), the fallow deer, +is a native of the shores of the Mediterranean, from Spain and Barbary to +Syria. _Capreolus_ (2 species), the roe-deer, inhabits all Temperate and +South Europe to Syria, with a distinct species in N. China. _Cervulus_ (4 +species), the muntjacs, are found in all the forest districts of the +Oriental region, from India and Ceylon to China as far north as Ningpo and +Formosa, also southward to the Philippines, Borneo, and Java. _Moschus_ (1 +species), the musk-deer, inhabits Central Asia from the Amoor and Pekin, to +the Himalayas and the Siamese mountains above 8000 ft. elevation. This is +usually classed as a distinct family, but M. Milne-Edwards remarks, that it +differs in no important points of organisation from the rest of the +Cervidæ. _Hydropotes_ {220}(1 species) inhabits China from the Yang-tse +Kiang northwards. This new genus has recently been discovered by Mr. +Swinhoe, who says its nearest affinities are with _Moschus_. Other new +forms are _Lophotragus_, and _Elaphodus_, both inhabiting North China; the +former is hornless, the latter has very small horns about an inch long. + +_Extinct Deer._--Numerous extinct species of the genus _Cervus_ are found +fossil in many parts of Europe, and in all formations between the +Post-pliocene and the Upper Miocene. The Elk and Reindeer are also found in +caves and Post-pliocene deposits, the latter as far south as the South of +France. Extinct genera only, occur in the Upper Miocene in various parts of +Europe:--_Micromeryx_, _Palæomeryx_, and _Dicrocercus_ have been described; +with others referred doubtfully to _Moschus_, and an allied genus +_Amphimoschus_. + +In N. America, remains of this family are very scarce, a _Cervus_ allied to +the existing wapiti deer, being found in Post-pliocene deposits, and an +extinct genus, _Leptomeryx_, in the Upper Miocene of Dakota and Oregon. +Another extinct genus, _Merycodus_, from the Pliocene of Oregon, is said to +be allied to camels and deer. + +In South America, several species of _Cervus_ have been found in the +Brazilian caves, and in the Pliocene deposits of La Plata. + +It thus appears, that there are not yet sufficient materials for +determining the origin and migrations of the Cervidæ. There can be little +doubt that they are an Old World group, and a comparatively recent +development; and that some time during the Miocene period they passed to +North America, and subsequently to the Southern continent. They do not +however appear to have developed much in North America, owing perhaps to +their finding the country already amply stocked with numerous forms of +indigenous Ungulates. + + +{221}FAMILY 51.--CAMELOPARDALIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + LIVING SPECIES. + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1 -- 3 -- |-- -- -- --|-- -- -- -- + | | | | | + EXTINCT SPECIES. + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- |1 -- -- -- |-- -- 3 -- |-- -- -- -- + | | | | | + +The Camelopardalidæ, or giraffes, now consist of but a single species which +ranges over all the open country of the Ethiopian region, and is therefore +almost absent from West Africa, which is more especially a forest district. +During the Middle Tertiary period, however, these animals had a wider +range, over Southern Europe and Western India as far as the slopes of the +Himalayas. + +_Extinct Species._--Species of _Camelopardalis_ have been found in Greece, +the Siwalik Hills, and Perim Island at the entrance to the Red Sea; and an +extinct genus, _Helladotherium_, more bulky but not so tall as the giraffe, +ranged from the south of France to Greece and North-west India. + + +FAMILY 52.--BOVIDÆ. (34 Genera, 149 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --| 1. 2 -- 4 |1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3. 4 |1 -- -- -- + | | | | | + +This large and important family, includes all the animals commonly known as +oxen, buffaloes, antelopes, sheep, and goats, which have been classed by +many naturalists in at least three, and sometimes four or five, distinct +families. Zoologically, they {222}are briefly and accurately defined as, +"hollow-horned ruminants;" and, although they present wide differences in +external form, they grade so insensibly into each other, that no +satisfactory definition of the smaller family groups can be found. As a +whole they are almost confined to the great Old World continent, only a few +forms extending along the highlands and prairies of the Nearctic region; +while one peculiar type is found in Celebes, an island which is almost +intermediate between the Oriental and Australian regions. In each of the +Old World regions there are found a characteristic set of types. Antelopes +prevail in the Ethiopian region; sheep and goats in the Palæarctic; while +the oxen are perhaps best developed in the Oriental region. + +Sir Victor Brooke, who has paid special attention to this family, divides +them into 13 sub-families, and I here adopt the arrangement of the genera +and species which he has been so good as to communicate to me in MSS. + +Sub-family I. BOVINÆ (6 genera, 13 species). This group is one of the best +marked in the family. It comprises the Oxen and Buffaloes with their +allies, and has a distribution very nearly the same as that of the entire +family. The genera are as follows: BOS (1 sp.), now represented by our +domestic cattle, the descendants of the _Bos primigenius_, which ranged +over a large part of Central Europe in the time of the Romans. The +Chillingham wild cattle are supposed to be the nearest approach to the +original species. _Bison_ (2 sp.), one still wild in Poland and the +Caucasus; the other in North America, ranging over the prairies west of the +Mississippi, and on the eastern slopes of the Rocky Mountains (Plate XIX., +vol. ii., p. 129). _Bibos_ (3 sp.), the Indian wild cattle, ranging over a +large part of the Oriental region, from Southern India to Assam, Burmah, +the Malay Peninsula, Borneo, and Java. _Poephagus_ (1 sp.), the yak, +confined to the high plains of Western Thibet. _Bubalus_ (5 sp.), the +buffaloes, of which three species are African, ranging over all the +continental parts of the Ethiopian region; one Northern and Central Indian; +and the domesticated animal in South Europe and North Africa. _Anoa_ (1 +sp.), the small wild cow of Celebes, {223}a very peculiar form more nearly +allied to the buffaloes than to any other type of oxen. + +Sub-family II. TRAGELAPHINÆ: (3 genera, 11 species). The Bovine Antelopes +are large and handsome animals, mostly Ethiopian, but extending into the +adjacent parts of the Palæarctic and Oriental regions. The genera are: +_Oreas_ (2 sp.), elands, inhabiting all Tropical and South Africa. +_Tragelaphus_ (8 sp.), including the bosch-bok, kudu, and other large +antelopes, ranges over all Tropical and South Africa (Plate IV., vol. i., +p. 261). _Portax_ (1 sp.) India, but rare in Madras and north of the +Ganges. + +Sub-family III. ORYGINÆ: (2 genera, 5 species). _Oryx_ (4 sp.) is a desert +genus, ranging over all the African deserts to South Arabia and Syria; +_Addax_ (1 sp.) inhabits North Africa, North Arabia, and Syria. + +Sub-family IV. HIPPOTRAGINÆ (1 genus, 3 species). The Sable Antelopes, +_Hippotragus_, form an isolated group inhabiting the open country of +Tropical Africa and south to the Cape. + +Sub-family V. GAZELLINÆ (6 genera, 23 species). This is a group of small or +moderate-sized animals, most abundant in the deserts on the borders of the +Palæarctic, Oriental, and Ethiopian regions. _Gazella_ (17 sp.) is +typically a Palæarctic desert group, ranging over the great desert plateaus +of North Africa, from Senegal and Abyssinia to Syria, Persia, Beloochistan, +and the plains of India, with one outlying species in South Africa. +_Procapra_ (2 sp.), Western Thibet and Mongolia to about 110° east +longitude. _Antilope_ (1 sp.) inhabits all the plains of India. _Æpyceros_ +(1 sp.) the pallah, inhabits the open country of South and South-east +Africa. _Saiga_ (1 sp.) a singular sheep-faced antelope, which inhabits the +steppes of Eastern Europe and Western Asia from Poland to the Irtish River, +south of 55° north latitude. (Plate II., vol. i., p. 218.) _Panthalops_ (1 +sp.) confined to the highlands of Western Thibet and perhaps Turkestan. + +Sub-family VI. ANTILOCAPRINÆ (1 genus, 1 species), _Antilocapra_, the +prong-horned antelope, inhabit both sides of the Rocky Mountains, extending +north to the Saskatchewan and {224}Columbia River, west to the coast range +of California, and east to the Missouri. Its remarkable deciduous horns +seem to indicate a transition to the Cervidæ. (Plate XIX., vol. ii., p. +129.) + +Sub-family VII. CERVICAPRINÆ (5 genera, 21 species). This group of +Antelopes is wholly confined to the continental portion of the Ethiopian +region. The genera are: _Cervicapra_ (4 sp.), Africa, south of the equator +and Abyssinia; _Kobus_ (6 sp.), grassy plains and marshes of Tropical +Africa; _Pelea_ (1 sp.), South Africa; _Nanotragus_ (9 species), Africa, +south of the Sahara; _Neotragus_ (1 sp.) Abyssinia and East Africa. + +Sub-family VIII. CEPHALOPHINÆ (2 genera, 24 species), Africa and India; +_Cephalophus_ (22 sp.), continental Ethiopian region; _Tetraceros_ (2 sp.) +hilly part of all India, but rare north of the Ganges. + +Sub-family IX. ALCEPHALINÆ (2 genera, 11 species), large African Antelopes, +one species just entering the Palæarctic region. The genera are: +_Alcephalus_ (9 sp.) all Africa and north-east to Syria; _Catoblepas_ (2 +sp.), gnus, Africa, south of the Equator. + +Sub-region X. BUDORCINÆ (1 genus, 2 species) _Budorcas_ inhabits the high +Himalayas from Nepal to East Thibet. + +Sub-family XI. RUPICAPRINÆ (1 genus, 2 species) the Chamois, _Rupicapra_, +inhabit the high European Alps from the Pyrenees to the Caucasus. (Plate +I., vol. i., p. 195.) + +Sub-family XII. NEMORHEDINÆ (2 genera, 10 species). These goat-like +Antelopes inhabit portions of the Palæarctic and Oriental regions, as well +as the Rocky Mountains in the Nearctic region. _Nemorhedus_ (9 sp.) ranges +from the Eastern Himalayas to N. China and Japan, and south to Formosa, the +Malay Peninsula and Sumatra. _Aplocerus_ (1 sp.), the mountain goat of the +trappers, inhabits the northern parts of California and the Rocky +Mountains. + +Sub-family XIII. CAPRINÆ (2 genera, 23 species). The Goats and Sheep form +an extensive series, highly characteristic of the Palæarctic region, but +with an outlying species on the Neilgherries in Southern India, and one in +the Rocky Mountains and California. The genera are _Capra_ (22 sp.) and +_Ovibos_ (1 sp.). {225}The genus _Capra_ consists of several sub-groups +which have been named as genera, but it is unnecessary here to do more than +divide them into "Goats and Ibexes" on the one hand and "Sheep" on the +other--each comprising 11 species. The former range over all the South +European Alps from Spain to the Caucasus; to Abyssinia, Persia, and Scinde; +over the high Himalayas to E. Thibet and N. China; with an outlying species +in the Neilgherries. The latter are only found in the mountains of Corsica, +Sardinia, and Crete, in Europe; in Asia Minor, Persia, and in Central and +North-Eastern Asia, with one somewhat isolated species in the Atlas +mountains; while in America a species is found in the Rocky Mountains and +the coast range of California. _Ovibos_ (1 sp.), the musk-sheep, inhabits +Arctic America north of lat. 60; but it occurs fossil in Post-glacial +gravels on the Yena and Obi in Siberia, in Germany and France along with +the Mammoth and with flint implements, and in caves of the Reindeer period; +also in the brick earth in the south of England, associated with +_Rhinoceros megarhinus_ and _Elephas antiquus_. + +_Extinct Bovidæ._--In the caverns and diluviums of Europe, of the +Post-Pliocene period, the remains are found of extinct species of _Bos_, +_Bison_, and _Capra_; and in the caverns of the south of France +_Rupicapra_, and an antelope near _Hippotragus_. _Bos_ and _Bison_ also +occur in Pliocene deposits. In the Miocene of Europe, the only remains are +antelopes closely allied to existing species, and these are especially +numerous in Greece, where remains referred to two living and four extinct +genera have been discovered. In the Miocene of India numerous extinct +species of _Bos_, and two extinct genera, _Hemibos_ and _Amphibos_, have +been found, one of them at a great elevation in Thibet. Antelopes, allied +to living Indian species, are chiefly found in the Nerbudda deposits. + +In North America, the only bovine remains are those of a _Bison_, and a +sheep or goat, in the Post-pliocene deposits; and of two species of +musk-sheep, sometimes classed in a distinct genus _Bootherium_, from beds +of the same age in Arkansas and Ohio. _Casoryx_, from the Pliocene of +Nebraska, is supposed to be allied to the antelopes and to deer. + +{226}In the caves of Brazil remains of two animals said to be antelopes, +have been discovered. They are classed by Gervais in the genera _Antilope_ +and _Leptotherium_, but the presence of true antelopes in S. America at +this period is so improbable, that there is probably some error of +identification. + +The extinct family Sivatheridæ, containing the extraordinary and gigantic +four-horned _Sivatherium_ and _Bramatherium_, of the Siwalik deposits, are +most nearly allied to the antelopes. + +From the preceding facts we may conclude, that the great existing +development of the Bovidæ is comparatively recent. The type may have +originated early in the Miocene period, the oxen being at first most +tropical, while the antelopes inhabited the desert zone a little further +north. The sheep and goats seem to be the most recent development of the +bovine type, which was probably long confined to the Eastern Hemisphere. + + +_General Remarks on the Distribution of the Ungulata._ + +With the exception of the Australian region, from which this order of +mammalia is almost entirely wanting, the Ungulata are almost universally +distributed over the continental parts of all the other regions. Of the ten +families, 7 are Ethiopian, 6 Oriental, 5 Palæarctic, 4 Neotropical, and 3 +Nearctic. The Ethiopian region owes its superiority to the exclusive +possession of the hippopotamus and giraffe, both of which inhabited the +Palæarctic and Oriental regions in Miocene times. The excessive poverty of +the Nearctic region in this order is remarkable; the swine being +represented only by _Dicotyles_ in its extreme southern portion, while the +Bovidæ are restricted to four isolated species. Deer alone are fairly well +represented. But, during the Eocene and Miocene periods, North America was +wonderfully rich in varied forms of Ungulates, of which there were at least +8 or 9 families; while we have reason to believe that during the same +periods the Ethiopian region was excessively poor, and that it probably +received the ancestors of all its existing families from Europe or Western +Asia in later Miocene or Pliocene times. Many types that once abounded in +both Europe and North America are now preserved only in South America and +Central or Tropical Asia,--as {227}the tapirs and camels; while others once +confined to Europe and Asia have found a refuge in Africa,--as the +hippopotamus and giraffe; so that in no other order do we find such +striking examples of those radical changes in the distribution of the +higher animals which were effected during the latter part of the Tertiary +period. The present distribution of this order is, in fact, utterly +unintelligible without reference to the numerous extinct forms of existing +and allied families; but as this subject has been sufficiently discussed in +the Second Part of this work (Chapters VI. and VII.) it is unnecessary to +give further details here. + + +_Order VIII.--PROBOSCIDEA._ + +FAMILY 53.--ELEPHANTIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + LIVING SPECIES. + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- |1. 2. 3. 4 |-- -- -- -- + | | | | | + EXTINCT SPECIES. + | | | | | + 1. 2 -- -- |1. 2. 3. 4 |1. 2. 3. 4 |1 -- -- -- |1 -- 3 -- |-- -- -- -- + | | | | | + +The elephants are now represented by two species, the African, which ranges +all over that continent south of the Sahara, and the Indian, which is found +over all the wooded parts of the Oriental region, from the slopes of the +Himalayas to Ceylon, and eastward, to the frontiers of China and to Sumatra +and Borneo. These, however, are but the feeble remnants of a host of +gigantic creatures, which roamed over all the great continents except +Australia during the Tertiary period, and several of which were +contemporary with man. + +_Extinct Elephants._--At least 14 extinct species of _Elephas_, and a +rather greater number of the allied genus _Mastodon_ (distinguished by +their less complex grinding teeth) have now been {228}discovered. Elephants +ranged over all the Palæarctic and Nearctic regions in Post-Pliocene times; +in Europe and Central India they go back to the Pliocene; and only in India +to the Upper Miocene period; the number of species increasing as we go back +to the older formations. + +In North America two or three species of _Mastodon_ are Post-pliocene and +Pliocene; and a species is found in the caves of Brazil, and in the +Pliocene deposits of the pampas of La Plata, of the Bolivian Andes, and of +Honduras and the Bahamas. In Europe the genus is Upper Miocene and +Pliocene, but is especially abundant in the former period. In the East, it +extends from Perim island to Burmah and over all India, and is mostly +Miocene, but with perhaps one species Pliocene in Central India. + +An account of the range of such animals as belong to extinct families of +Proboscidea, will be found in Chapters VI. and VII.; from which it will be +seen that, although the family Elephantidæ undoubtedly originated in the +Eastern Hemisphere, it is not improbable that the first traces of the order +Proboscidea are to be found in N. America. + + +_Order IX.--HYRACOIDEA._ + +FAMILY 54.--HYRACIDÆ. (1 Genus. 10-12 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2. -- --|1. 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The genus _Hyrax_, which alone constitutes this family, consists of small +animals having the appearance of hares or marmots, but which more resemble +the genus _Rhinoceros_ in their teeth and skeleton. They range all over the +Ethiopian region, except Madagascar; a peculiar species is found in +Fernando Po, and they just enter the Palæarctic as far as Syria. They may +therefore be considered as an exclusively Ethiopian group. In Dr. Gray's +{229}last Catalogue (1873) he divides the genus into three--_Hyrax_, +_Euhyrax_ and _Dendrohyrax_--the latter consisting of two species confined +apparently to West and South Africa. + +No extinct forms of this family have yet been discovered; the +_Hyracotherium_ of the London clay (Lower Eocene) which was supposed to +resemble _Hyrax_, is now believed to be an ancestral type of the Suidæ or +swine. + + +_Order X.--RODENTIA._ + +FAMILY 55.--MURIDÆ. (37 Genera, 330 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |-- 2 -- -- + | | | | | + +The Muridæ, comprising the rats and mice with their allies, are almost +universally distributed over the globe (even not reckoning the domestic +species which have been introduced almost everywhere by man), the +exceptions being the three insular groups belonging to the Australian +region, from none of which have any species yet been obtained. Before +enumerating the genera it will be as well to say a few words on the +peculiarities of distribution they present. The true mice, forming the +genus _Mus_, is distributed over the whole of the world except N. and S. +America where not a single indigenous species occurs, being replaced by the +genus _Hesperomys_; five other genera, comprehending all the remaining +species found in South America are peculiar to the Neotropical region. +Three genera are confined to the Palæarctic region, and three others to the +Nearctic. No less than twelve genera are exclusively Ethiopian, while only +three are exclusively Oriental and three Australian. + +_Mus_ (100-120 sp.) the Eastern Hemisphere, but absent from the Pacific and +Austro-Malayan Islands, except Celebes and Papua; _Lasiomys_ (1 sp.) +Guinea; _Acanthomys_ (5-6 sp.) Africa, India and {230}N. Australia; +_Cricetomys_ (1 sp.) Tropical Africa; _Saccostomus_ (2 sp.) Mozambique; +_Cricetus_ (9 sp.) Palæarctic region and Egypt; _Cricetulus_ (1 sp., +Milne-Edwards, 1870) Pekin; _Pseudomys_ (1 sp.) Australia; _Hapalotis_ (13 +sp.) Australia; _Phlæomys_ (1 sp.) Philippines; _Platacanthomys_ (1 sp., +Blyth, 1865) Malabar; _Dendromys_ (2 sp.) S. Africa; _Nesomys_ (1 sp. +Peters, 1870) Madagascar; _Steatomys_ (2 sp.) N. and S. Africa; _Pelomys_ +(1 sp.) Mozambique; _Reithrodon_ (9 sp.) N. America, Lat. 29° to Mexico, +and south to Tierra del Fuego; _Acodon_ (1 sp.) Peru; _Myxomys_ (1 sp.) +Guatemala; _Hesperomys_ (90 sp.) North and South America; _Holochilus_ (4 +sp.) South America; _Oxymycterus_ (4 sp.) Brazil and La Plata; _Neotoma_ (6 +sp.) U.S., East coast to California; _Sigmodon_ (2 sp.) Southern United +States; _Drymomys_ (1 sp.) Peru; _Neotomys_ (2 sp.) S. America; _Otomys_ (6 +sp.) S. and E. Africa; _Meriones_ = _Gerbillus_ (20-30 sp.) Egypt, Central +Asia, India, Africa; _Rhombomys_ (6 sp.) S. E. Europe, N. Africa, Central +Asia; _Malacothrix_ (2 sp.) South Africa; _Mystromys_ (1 sp.) South Africa; +_Psammomys_ (1 sp.) Egypt; _Spalacomys_ (1 sp.) India; _Sminthus_ (1-3 sp.) +East Europe, Tartary, Siberia; _Hydromys_ (5 sp.) Australia and Tasmania; +_Hypogeomys_ (1 sp., Grandidier, 1870) Madagascar; _Brachytarsomys_ (1 sp., +Günther, 1874) Madagascar; _Fiber_ (2 sp.) N. America to Mexico; _Arvicola_ +(50 sp.) Europe to Asia Minor, North Asia, Himalayas, Temp. N. America; +_Cuniculus_ (1 sp.) N. E. Europe, Siberia, Greenland, Arctic America; +_Myodes_ (4 sp.) Europe, Siberia, Arctic America, and Northern United +States; _Myospalax_ = _Siphneus_ (2 sp.) Altai Mountains and N. China[4]; +_Lophiomys_ (1 sp.) S. Arabia, and N. E. Africa; _Echiothrix_ (1 sp.) +Australia. + +_Extinct Muridæ._--Species of _Mus_, _Cricetus_, _Arvicola_, and _Myodes_, +occur in the Post-Pliocene deposits of Europe; _Arvicola_, _Meriones_, and +the extinct genus _Cricetodon_, with some others, in the Miocene. + +In North America, _Fiber_, _Arvicola_, and _Neotoma_, occur in caves; +{231}an extinct genus, _Eumys_, in the Upper Miocene of Dakota, and +another, _Mysops_, in the Eocene of Wyoming. + +In South America _Mus_, or more probably _Hesperomys_, is abundant in +Brazilian caverns, and _Oxymycterus_ in the Pliocene of La Plata; while +_Arvicola_ is said to have occurred both in the Pliocene and Eocene +deposits of the same country. + + +FAMILY 56.--SPALACIDÆ. (7 Genera, 17 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3 -- |1. 2. 3 -- | 1 -- 3. 4 |-- -- -- -- + | | | | | + +The Spalacidæ, or mole-rats, have a straggling distribution over the Old +World continents. They are found over nearly the whole of Africa, but only +in the South-east of Europe, and West of Temperate Asia, but appearing +again in North India, Malacca, and South China. _Ellobius_ (1 sp.), is +found in South Russia and South-west Siberia; _Spalax_ (1 sp.), Southern +Russia, West Asia, Hungary, Moldavia, and Greece (Plate II., vol. i. p. +218); _Rhizomys_ (6 sp.), Abyssinia, North India, Malacca, South China; +_Heterocephalus_ (1 sp.), Abyssinia; _Bathyerges_ (= _Orycterus_ 1 sp.), +South Africa; _Georychus_ (6 sp.), South, Central, and East Africa; +_Heliophobus_ (1 sp.), Mozambique. + + +FAMILY 57.--DIPODIDÆ. (3 Genera, 22 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|1. 2. 3. 4 |-- 2. 3. 4 |1. 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The Jerboas, or jumping mice, are especially characteristic of the regions +about the eastern extremity of the Mediterranean, being found in South +Russia, the Caspian district, Arabia, Egypt, {232}and Abyssinia; but they +also extend over a large part of Africa, and eastward to India; while +isolated forms occur in North America, and the Cape of Good Hope. _Dipus_ = +_Gerbillus_ (20 sp.), inhabits North and Central Africa, South-East Europe, +and across Temperate Asia to North China, also Afghanistan, India, and +Ceylon; _Pedetes_ (1 sp.), South Africa to Mozambique and Angola; _Jaculus_ += _Meriones_ (1 sp.), North America, from Nova Scotia and Canada, south to +Pennsylvania and west to California and British Columbia (Plate XX., vol. +ii. p. 135). + +_Extinct Dipodidæ._--_Dipus_ occurs fossil in the Miocene of the Alps; and +an extinct genus, _Issiodromys_, said to be allied to _Pedetes_ of the Cape +of Good Hope, is from the Pliocene formations of Auvergne in France. + + +FAMILY 58.--MYOXIDÆ. (1 Genus, 12 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The Dormice (_Myoxus_), are small rodents found over all the temperate +parts of the Palæarctic region, from Britain to Japan; and also over most +parts of Africa to the Cape, but wanting in India. Some of the African +species have been separated under the name of _Graphidurus_, while those of +Europe and Asia form the sub-genera _Glis_, _Muscardinus_, and _Eliomys_. + +_Extinct Myoxidæ._--_Myoxus_ ranges from the Post-pliocene of the Maltese +caverns to the Miocene of Switzerland and the Upper Eocene of France; and +an extinct genus _Brachymys_ is found in the Miocene of Central Europe. + + +{233}FAMILY 59.--SACCOMYIDÆ. (6 Genera, 33 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|1. 2. 3. 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Saccomyidæ, or pouched rats, are almost wholly confined to our second +Nearctic sub-region, comprising the Rocky Mountains and the elevated plains +of Central North America. A few species range from this district as far as +Hudson's Bay on the north, to South Carolina on the east, and to California +on the west, while one genus, doubtfully placed here, goes south as far as +Honduras and Trinidad. The group must therefore be considered to be +pre-eminently characteristic of the Nearctic region. + +The genera are,--_Dipodomys_ (5 sp.), North Mexico, California, the east +slope of the Rocky Mountains to the Columbia River, and one species in +South Carolina; _Perognathus_ (6 sp.), North Mexico, California, east slope +of the Rocky Mountains to British Columbia; _Thomomys_ (2 sp.), Upper +Missouri, and Upper Columbia Rivers to Hudson's Bay; _Geomys_ (5 sp.), +North Mexico, and east slope of Rocky Mountains to Nebraska (Plate XIX., +vol. ii. p. 129); _Saccomys_ (1 sp.), North America, locality unknown; +_Heteromys_ (6 sp.), Mexico, Honduras, and Trinidad. _Geomys_ and +_Thomomys_ constitute a separate family Geomyidæ, of Professor Carus; but I +follow Professor Lilljeborg, who has made a special study of the Order, in +keeping them with this family. + +In the Post-Pliocene deposits of Illinois and Nebraska, remains of an +existing species of _Geomys_ have been found. + + +{234}FAMILY 60.--CASTORIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|1. 2. 3. 4 | 1 -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Beavers, forming the genus _Castor_, consist of two species, the +American (_Castor canadensis_) ranging over the whole of North America from +Labrador to North Mexico; while the European (_Castor fiber_) appears to be +confined to the temperate regions of Europe and Asia, from France to the +River Amoor, over which extensive region it doubtless roamed in prehistoric +times, although now becoming rare in many districts. + +_Extinct Castoridæ._--Extinct species of _Castor_ range back from the +Post-pliocene to the Upper Miocene in Europe, and to the Newer Pliocene in +North America. Extinct genera in Europe are, _Trogontherium_, Post-Pliocene +and Pliocene; _Chalicomys_, Older Pliocene; and _Steneofiber_, Upper +Miocene. In North America _Castoroides_ is Post-Pliocene, and +_Palæocastor_, Upper Miocene. The family thus first appears on the same +geological horizon in both Europe and North America. + + +FAMILY 61.--SCIURIDÆ.--(8 Genera, 180-200 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Squirrel family, comprehending also the marmots and prairie-dogs, are +very widely spread over the earth. They are especially abundant in the +Nearctic, Palæarctic, and Oriental regions, and rather less frequent in the +Ethiopian and Neotropical, in which last region they do not extend south of +Paraguay. They are absent from the West Indian islands, Madagascar, and +Australia, only occurring in Celebes which doubtfully belongs to the +Australian region. The genera are as follows:-- + +{235}_Sciurus_ (100-120 sp., including the sub-genera Spermosciurus, Xerus, +Macroxus, Rheithrosciurus, and Rhinosciurus), comprises the true squirrels, +and occupies the area of the whole family wherever woods and forests occur. +The approximate number of species in each region is as follows: Nearctic +18, Palæarctic 6, Ethiopian 18, Oriental 50, Australian (Celebes) 5, +Neotropical 30. _Sciuropterus_ (16-19 sp.), comprises the flat-tailed +flying squirrels, which range from Lapland and Finland to North China and +Japan, and southward through India and Ceylon, to Malacca and Java, with a +species in Formosa; while in North America they occur from Labrador to +British Columbia, and south to Minnesota and Southern California. +_Pteromys_ (12 sp.), comprising the round-tailed flying squirrels, is a +more southern form, being confined to the wooded regions of India from the +Western Himalayas to Java and Borneo, with species in Formosa and Japan. +_Tamias_ (5 sp.), the ground squirrels, are chiefly North American, ranging +from Mexico to Puget's Sound on the west coast, and from Virginia to +Montreal on the Atlantic coast; while one species is found over all +northern Asia. _Spermophilus_ (26 sp.), the pouched marmots, are confined +to the Nearctic and Palæarctic regions; in the former extending from the +Arctic Ocean to Mexico and the west coast, but not passing east of Lake +Michigan and the lower Mississippi; in the latter from Silesia through +South Russia to the Amoor and Kamschatka, most abundant in the desert +plains of Tartary and Mongolia. _Arctomys_ (8 sp.), the marmots, are found +in the northern parts of North America as far down as Virginia and Nebraska +to the Rocky Mountains and British Columbia, but not in California; and +from the Swiss Alps eastward to Lake Baikal and Kamschatka, and south as +far as the Himalayas, above 8,000 feet elevation. _Cynomys_ (2 sp.), the +prairie-dogs, inhabit the plains east of the Rocky Mountains from the Upper +Missouri to the Red River and Rio Grande (Plate XIX., vol. ii. p. 129). +_Anomalurus_ (5 sp.), consists of animals which resemble flying-squirrels, +but differ from all other members of the family in some points of internal +structure. They form a very aberrant portion of the Sciuridæ, and, +according to some naturalists, a distinct family. They inhabit West Africa +and the island of Fernando Po. + +{236}_Extinct Sciuridæ._--These are tolerably abundant. The genus Sciurus +appears to be a remarkably ancient form, extinct species being found in the +Miocene, and even in the Upper Eocene formations of Europe. _Spermophilus_ +goes back to the Upper Miocene; _Arctomys_ to the Newer Pliocene. Extinct +genera are, _Brachymys_, _Lithomys_ and _Plesiarctomys_, from the European +Miocene, the latter said to be intermediate between marmots and squirrels. + +In North America, _Sciurus_, _Tamias_, and _Arctomys_ occur in the +Post-pliocene deposits only. The extinct genera are _Ischyromys_, from the +Upper Miocene of Nebraska; _Paramys_, allied to the marmots, and +_Sciuravus_, near the squirrels, from the Eocene of Wyoming. + +Here we have unmistakable evidence that the true squirrels (_Sciurus_) are +an Old World type, which has only recently entered North America; and this +is in accordance with the comparative scarcity of this group in South +America, a country so well adapted to them, and their great abundance in +the Oriental region, which, with the Palæarctic, was probably the country +of their origin and early development. The family, however, has been traced +equally far back in Europe and North America, so that we have as yet no +means of determining where it originated. + + +FAMILY 62--HAPLOODONTIDÆ.--(1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|1 -- -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The genus _Haploodon_ or _Aplodontia_, consists of two curious rat-like +animals, inhabiting the west coast of America, from the southern part of +British Columbia to the mountains of California. They seem to have +affinities both with the beavers and marmots, and Professor Lilljeborg +constitutes a separate family to receive them. + + +{237}FAMILY 63.--CHINCHILLIDÆ. (3 Genera, 6 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1 -- -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Chinchillidæ, including the chinchillas and viscachas, are confined to +the alpine zones of the Andes, from the boundary of Ecuador and Peru to the +southern parts of Chili; and over the Pampas, to the Rio Negro on the +south, and the River Uruguay on the east. _Chinchilla_ (2 sp.), the true +chinchillas, are found in the Andes of Chili and Peru, south of 9° S. lat., +and from 8,000 to 12,000 feet elevation (Plate XVI. vol. ii. p. 40); +_Lagidium_ (3 sp.), the alpine viscachas, inhabit the loftiest plateaus and +mountains from 11,000 to 16,000 feet, and extend furthest north of any of +the family; while _Lagostomus_ (1 sp.), the viscacha of the Pampas, has the +range above indicated. The family is thus confined within the limits of a +single sub-region. + +_Extinct Chinchillidæ._--_Lagostomus_ has been found fossil in the caves of +Brazil, and in the Pliocene deposits of La Plata. The only known extinct +forms of this family are _Amblyrhiza_ and _Loxomylus_, found in +cavern-deposits in the island of Anguilla, of Post-Pliocene age. These are +very interesting, as showing the greater range of this family so recently; +though its absence from North America and Europe indicates that it is a +peculiar development of the Neotropical region. + + +FAMILY 64.--OCTODONTIDÆ. (8 Genera, 19 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 -- 4 |-- -- -- --|-- 2 -- -- |1 -- -- -- |-- -- -- --|-- -- -- -- + | | | | | + +{238}The Octodontidæ include a number of curious and obscure rat-like +animals, mostly confined to the mountains and open plains of South America, +but having a few stragglers in other parts of the world, as will be seen by +our notes on the genera. The most remarkable point in their distribution +is, that two genera are peculiar to the West Indian islands, while no +species of the family inhabits the northern half of South America. The +distribution of the genera is as follows:--_Habrocomus_ (2 sp.), Chili; +_Capromys_ (3 sp.), two of which inhabit Cuba, the third Jamaica (Plate +XVII. vol. ii. p. 67); _Plagiodontia_ (1 sp.), only known from _Hayti_; +_Spalacopus_, including _Schizodon_ (2 sp.), Chili, and east side of +Southern Andes; _Octodon_ (3 sp.), Peru, Bolivia, and Chili; _Ctenomys_ (6 +sp.), the tuco-tuco of the Pampas, the Campos of Brazil to Bolivia and +Tierra del Fuego; _Ctenodactylus_ (1 sp.), Tripoli, North Africa; +_Pectinator_ (1 sp.), East Africa, Abyssinia, 4,000 to 5,000 feet. + +_Capromys_ and _Plagiodontia_, the two West Indian genera, were classed +among the Echimyidæ by Mr. Waterhouse, but Professor Lilljeborg removes +them to this family. + +_Extinct Octodontidæ._--Species of _Ctenomys_ have been found in the +Pliocene of La Plata, and an extinct genus _Megamys_, said to be allied to +_Capromys_, in the Eocene of the same country. In Europe, _Palæomys_ and +_Archæomys_ from the lower Miocene of Germany and France, are also said to +be allied to _Capromys_. + + +FAMILY 65.--ECHIMYIDÆ. (10 Genera, 30 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 -- -- |-- -- -- --|-- -- -- --| 1 -- 3 -- |-- -- -- --|-- -- -- -- + | | | | | + + +The Echimyidæ, or spiny rats, are a family, chiefly South American, of +which the Coypu, a large beaver-like water-rat from Peru and Chili is the +best known. Two of the genera are found in South Africa, but all the rest +inhabit the continent of South America, East of the Andes, none being yet +known north {239}of Panama. The genera are as follows:--_Dactylomys_ (2 +sp.), Guiana and Brazil; _Cercomys_ (1 sp.), Central Brazil; _Lasiuromys_ +(1 sp.), San Paulo, Brazil; _Petromys_ (1 sp.), South Africa; _Myopotamus_ +(1 sp.), the coypu, on the East side of the Andes from Peru to 42° S. lat., +on the West side from 33° to 48° S. lat.; _Carterodon_ (1 sp.), Minaes +Geraes, Brazil; _Aulacodes_ (1. sp.), West and South Africa; _Mesomys_ (1 +sp.), Borba on the Amazon; _Echimys_ (11 sp.), from Guiana and the +Ecuadorian Andes to Paraguay; _Loncheres_ (10 sp.), New Granada to Brazil. + +_Fossil and Extinct Echimyidæ._--The genus _Carterodon_ was established on +bones found in the Brazilian caves, and it was several years afterwards +that specimens were obtained showing the animal to be a living species. +Extinct species of _Myopotamus_ and _Loncheres_ have also been found in +these caves, with the extinct genera _Lonchophorus_ and _Phyllomys_. + +No remains of this family have been discovered in North America; but in the +Miocene and Upper Eocene deposits of France there are many species of an +extinct genus _Theridomys_, which is said to be allied to this group or to +the next (Cercolabidæ). _Aulacodon_, from the Upper Miocene of Germany, is +allied to the West African _Aulacodes_; and some other remains from the +lower Miocene of Auvergne, are supposed to belong to _Echimys_. + + +FAMILY 66.--CERCOLABIDÆ. (3 Genera, 13-15 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |1. 2. 3. 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Cercolabidæ, or arboreal porcupines, are a group of rodents entirely +confined to America, where they range from the northern limit of trees on +the Mackenzie River, to the southern limit of forests in Paraguay. There is +however an intervening district, the Southern United States, from which +they are absent. _Erethizon_ (3 sp.), the Canadian porcupine, is found +throughout {240}Canada and as far south as Northern Pennsylvania, and west +to the Mississippi (Plate XX., vol. ii. p. 135); an allied species +inhabiting the west coast from California to Alaska, and inland to the head +of the Missouri River; while a third is found in the north-western part of +South America; _Cercolabes_ (12 sp.), ranges from Mexico and Guatemala to +Paraguay, on the eastern side of the Andes; _Chætomys_ (1 sp.), North +Brazil. + +_Extinct Cercolabidæ._--A large species of _Cercolabes_ has been found in +the Brazilian caves, but none have been discovered in North America or +Europe. We may conclude therefore that this is probably a South American +type, which has thence spread into North America at a comparatively recent +epoch. The peculiar distribution of _Cercolabes_ may be explained by +supposing it to have migrated northwards along the west coast by means of +the wooded slopes of the Rocky Mountains. It could then only reach the +Eastern States by way of the forest region of the great lakes, and then +move southward. This it may be now doing, but it has not yet reached the +Southern States of Eastern North America. + + +_Family_ 67.--HYSTRICIDÆ. (3 Genera, 12 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- |1. 2. 3 -- |1. 2. 3. 4 |-- -- -- -- + | | | | | + +The true Porcupines have a very compact and well-marked distribution, over +the whole of the Oriental and Ethiopian regions (except Madagascar), and +the second Palæarctic sub-region. There is some confusion as to their +sub-division into genera, but the following are those most usually +admitted:--_Hystrix_ (5 sp.), South Europe to the Cape of Good Hope, all +India, Ceylon, and South China; _Atherura_ (5 sp.), "brush-tailed +porcupines," inhabit West Africa, India, to Siam, Sumatra, and Borneo; +_Acanthion_ (2 sp.), Nepal and Malacca, to Sumatra, Borneo, and Java. + +_Extinct Hystricidæ._--Several extinct species of _Hystrix_ have {241}been +found in the Pliocene and Miocene deposits of Europe, and one in the +Pliocene of Nebraska in North America. + +FAMILY 68.--CAVIIDÆ. (6 Genera, 28 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Cavies and Agoutis were placed in distinct families by Mr. Waterhouse, +in which he is followed by Professor Carus, but they have been united by +Professor Lilljeborg, and without pretending to decide which classification +is the more correct I follow the latter, because there is a striking +external resemblance between the two groups, and they have an identical +distribution in the Neotropical region, and with one exception are all +found east of the Andes. _Dasyprocta_ (9 sp.), the agouti, ranges from +Mexico to Paraguay, one species inhabiting the small West Indian islands of +St. Vincent, Lucia, and Grenada; _Cælogenys_ (2 sp.), the paca, is found +from Guatemala to Paraguay, and a second species (somewhat doubtful) in +Eastern Peru; _Hydrochoerus_ (1 sp.), the capybara inhabits the banks of +rivers from Guayana to La Plata; _Cavia_ (9 sp.), the guinea-pigs, Brazil +to the Straits of Magellan, and one species west of the Andes at Yça Peru; +_Kerodon_ (6 sp.), Brazil and Peru to Magellan; _Dolichotis_ (1 sp,), the +Patagonian cavy, from Mendoza to 48° 30' south latitude, on sterile plains. + +_Extinct Caviidæ._--_Hydrochoerus_, _Cælogenys_, _Dasyprocta_, and +_Kerodon_, have occurred abundantly in the caves of Brazil, and the +last-named genus in the Pliocene of La Plata. _Hydrochoerus_ has been found +in the Post-Pliocene deposits of South Carolina. _Cavia_ and _Dasyprocta_ +are said to have been found in the Miocene of Switzerland and France. No +well-marked extinct genera of this family have been recorded. + +If the determination of the above-mentioned fossil species of _Cavia_ and +_Dasyprocta_ are correct, it would show that this now {242}exclusively +South American family is really derived from Europe, where it has long been +extinct. + + +FAMILY 69.--LAGOMYIDÆ. (1 Genus, 11 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --| -- 2 -- 4 |-- -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Lagomyidæ, or pikas, are small alpine and desert animals which range +from the south of the Ural Mountains to Cashmere and the Himalayas, at +heights of 11,000 to 14,000 feet, and northward to the Polar regions and +the north-eastern extremity of Siberia. They just enter the eastern +extremity of Europe as far as the Volga, but with this exception, seem +strictly limited to the third Palæarctic sub-region. In America they are +confined to the Rocky Mountains from about 42° to 60° north latitude. + +_Extinct Lagomyidæ._--Extinct species of _Lagomys_ have occurred in the +southern parts of Europe, from the Post-Pliocene to the Miocene formations. +_Titanomys_, an extinct genus, is found in the Miocene of France and +Germany. + + +FAMILY 70.--LEPORIDÆ. (1 Genus, 35-40 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |1. 2. 3. 4 |1. 2. 3. 4 | 1 -- 3 -- |1. 2. 3 -- |-- -- -- -- + | | | | | + +The Hares and Rabbits are especially characteristic of the Nearctic and +Palæarctic, but are also thinly scattered over the Ethiopian and Oriental +regions. In the Neotropical region they are very scarce, only one species +being found in South America, in the mountains of Brazil and various parts +of the Andes, while one or two of the North American species extend into +Mexico {243}and Guatemala. In the Nearctic region, they are most abundant +in the central and western parts of the continent, and they extend to the +Arctic Ocean and to Greenland. They are found in every part of the +Palæarctic region, from Ireland to Japan; three species range over all +India to Ceylon, and others occur in Hainan, Formosa, South China, and the +mountains of Pegu; the Ethiopian region has only four or five species, +mostly in the southern extremity and along the East coast. An Indian +species is now wild in some parts of Java, but it has probably been +introduced. + +_Extinct Leporidæ._--Species of _Lepus_ occur in the Post-Pliocene and +Newer Pliocene of France; but only in the Post-Pliocene of North America, +and the caves of Brazil. + + +_General Remarks on the Distribution of the Rodentia._ + +With the exception of the Australian region and Madagascar, where Muridæ +alone have been found, this order is one of the most universally and evenly +distributed over the entire globe. Of the sixteen families which compose +it, the Palæarctic region has 10; the Ethiopian, Nearctic, and Neotropical, +each 9; and the Oriental only 5. These figures are very curious and +suggestive. We know that the rodentia are exceedingly ancient, since some +of the living genera date back to the Eocene period; and some ancestral +types might thus have reached the remote South American and South African +lands at the time of one of their earliest unions with the northern +continents. In both these countries the rodents diverged into many special +forms, and being small animals easily able to conceal themselves, have +largely survived the introduction of higher Mammalia. In the Palæarctic and +Nearctic regions, their small size and faculty of hibernation may have +enabled them to maintain themselves during those great physical changes +which resulted in the extermination or banishment of so many of the larger +and more highly organised Mammalia, to which, in these regions, they now +bear a somewhat inordinate proportion. The reasons why they are now less +numerous and varied in the Oriental region, may be of two kinds. The +comparatively small area of that region and its {244}uniformity of climate, +would naturally lead to less development of such a group as this, than in +the vastly more extensive and varied and almost equally luxuriant +Palæarctic region of Eocene and Miocene times; while on the other hand the +greater number of the smaller Carnivora in the tropics during the Pliocene +and Post-Pliocene epochs, would be a constant check upon the increase of +these defenceless animals, and no doubt exterminate a number of them. + +The Rodents thus offer a striking contrast to the Ungulates; and these two +great orders afford an admirable illustration of the different way in which +physical and organic changes may affect large and small herbivorous +Mammalia; often leading to the extinction of the former, while favouring +the comparative development of the latter. + + +_Order XI.--EDENTATA._ + +FAMILY 71.--BRADYPODIDÆ. (3 Genera, 12 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Sloths are a remarkable group of arboreal mammals, strictly confined to +the great forests of the Neotropical region, from Guatemala to Brazil and +Eastern Bolivia. None are found west of the Andes, nor do they appear to +extend into Paraguay, or beyond the Tropic of Capricorn on the east coast. +The genera as defined by Dr. Gray in 1871 are:--_Choloepus_ (2 sp.), +"Sloths with two toes on fore limbs, sexes alike," Costa Rica to Brazil; +_Bradypus_ (2 sp.), "Sloths with three toes on fore limbs, sexes alike," +Central Brazil, Amazon to Rio de Janeiro; _Arctopithecus_ (8 sp.), "Sloths +with three toes on fore limbs, males with a coloured patch on the back," +Costa Rica to Brazil and Eastern Bolivia (Plate XIV., vol ii. p. 24). + +{245}_Extinct Bradypodidæ._--In the caves of Brazil are found three extinct +genera of Sloths--_Cælodon_, _Sphenodon_, and _Ochotherium_. More distantly +allied, and probably forming distinct families, are _Scelidotherium_ and +_Megatherium_, from the caves of Brazil and the Pliocene deposits of La +Plata and Patagonia. + + +FAMILY 72.--MANIDIDÆ. (1 Genus, 8 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- |1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Manididæ, or scaly ant-eaters, are the only Edentate Mammalia found out +of America, They are spread over the Ethiopian and Oriental regions; in the +former from Sennaar to West Africa and the Cape; in the latter from the +Himalayas to Ceylon, and Eastward to Borneo and Java, as well as to South +China, as far as Amoy, Hainan, and Formosa. They have been sub-divided, +according to differences in the scaly covering, into five groups, _Manis_, +_Phatagin_, _Smutsia_, _Pholidotus_ and _Pangolin_, the three former being +confined to Africa, the last common to Africa and the East, while +_Pholidotus_ seems confined to Java. It is doubtful if these divisions are +more than sub-genera, and as such they are treated here. + +No extinct species referable to this family are yet known. + + +FAMILY 73.--DASYPODIDÆ. (6 Genera, 17 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Dasypodidæ, or armadillos, are a highly characteristic Neotropical +family, ranging from the northern extremity of the region {246}in south +Texas, to 50° south latitude on the plains of Patagonia. The distribution +of the genera is as follows:--_Tatusia_ (5 sp.), has the range of the whole +family from the lower Rio Grande of Texas to Patagonia; _Prionodontes_ (1 +sp.), the giant armadillo, Surinam to Paraguay; _Dasypus_ (4 sp.), Brazil +to Bolivia, Chili, and La Plata; _Xenurus_ (3 sp.), Guiana to Paraguay; +_Tolypeutes_ (2 sp.), the three-banded armadillos, Bolivia and La Plata; +_Chlamydophorus_ (2 sp.), near Mendoza in La Plata, and Santa Cruz de la +Sierra in Bolivia. + +_Extinct Armadillos._--Many species of _Dasypus_ and _Xenurus_ have been +found in the caves of Brazil, together with many extinct +genera--_Hoplophorus_, _Euryodon_, _Heterodon_, _Pachytherium_, and +_Chlamydotherium_, the latter as large as a rhinoceros. _Eutatus_, allied +to _Tolypeutes_, is from the Pliocene deposits of La Plata. + + +FAMILY 74.--ORYCTEROPODIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --| 1 -- 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The Aard-vark, or Cape ant-eater (_Orycteropus capensis_) is a curious form +of Edentate animal, with the general form of an ant-eater, but with the +bristly skin and long obtuse snout of a pig. A second species inhabits the +interior of North-East Africa and Senegal, that of the latter country +perhaps forming a third species (Plate IV. vol. i. p. 261). + +_Extinct Orycteropodidæ._--The genus _Macrotherium_, remains of which occur +in the Miocene deposits of France, Germany, and Greece, is allied to this +group, though perhaps forming a separate family. The same may be said of +the _Ancylotherium_, a huge animal found only in the Miocene deposits of +Greece. + + +{247}FAMILY 75.--MYRMECOPHAGIDÆ. (3 Genera, 5 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The true ant-eaters are strictly confined to the wooded portions of the +Neotropical region, ranging from Honduras to Paraguay on the East side of +the Andes. The three genera now generally admitted are: _Myrmecophaga_ (1 +sp.), the great ant-eater, Northern Brazil to Paraguay; _Tamandua_ (2 sp.), +4-toed ant-eaters, Guatemala, Ecuador to Paraguay (Plate XIV. vol. ii. p. +24); _Cyclothurus_ (2 sp.), 2-toed ant-eaters, Honduras and Costa Rica to +Brazil. + +_Extinct Ant-eaters._--The only extinct form of this family seems to be the +_Glossotherium_, found in the caves of Brazil, and the Tertiary deposits of +Uruguay. It is said to be allied to _Myrmecophaga_ and _Manis_. + + +_General Remarks on the Distribution of the Edentata._ + +These singular animals are almost confined to South America, where they +constitute an important part of the fauna. In Africa, two family types are +scantily represented, and one of these extends over all the Oriental +region. In Pliocene and Post-Pliocene times the Edentata were wonderfully +developed in South America, many of them being huge animals, rivalling in +bulk, the rhinoceros and hippopotamus. As none of these forms resemble +those of Africa, while the only European fossil Edentata are of African +type, it seems probable that South Africa, like South America, was a centre +of development for this group of mammalia; and it is in the highest degree +probable that, should extensive fluviatile deposits of Pliocene or Miocene +age be discovered in the former country, an extinct fauna, not less strange +and grotesque than that of South America, will be brought to {248}light. +From the fact that so few remains of this order occur in Europe, and those +of one family type, and in Miocene deposits only, it seems a fair +conclusion, that this represents an incursion of an ancient Ethiopian form +into Europe analogous to that which invaded North America from the south +during the Post-Pliocene epoch. The extension of the Manididæ, or scaly +ant-eaters, over tropical Asia may have occurred at the same, or a somewhat +later epoch. + +For a summary of the Numerous Edentata of North and South America which +belong to extinct families, see vol. i. p. 147. + + +_Order XII.--MARSUPIALIA._ + +FAMILY 76.--DIDELPHYIDÆ. (3 Genera, 22 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- | 1 -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Didelphyidæ, or true opossums, range throughout all the wooded +districts of the Neotropical region from the southern boundary of Texas to +the River La Plata, and on the west coast to 42° S. Lat., where a species +of _Didelphys_ was obtained by Professor Cunningham. One species only is +found in the Nearctic region, extending from Florida to the Hudson River, +and west to the Missouri. The species named _Didelphys californica_ +inhabits Mexico, and only extends into the southern extremity of +California. The species are most numerous in the great forest region of +Brazil, and they have been recently found to the west of the Andes near +Guayaquil, as well as in Chili. The exact number of species is very +doubtful, owing to the difficulty of determining them from dried skins. All +but two belong to the genus _Didelphys_, which has the range above given +for the family (Plate XIV., vol. ii. p. 24); _Chironectes_ (1 sp.), the +yapock or water opossum, inhabits Guiana and Brazil; _Hyracodon_ (1 sp.), +is a small {249}rat-like animal discovered by Mr. Fraser in Ecuador, and +which may perhaps belong to another family. + +_Extinct Didelphyidæ._--No less than seven species of _Didelphys_ have been +found in the caves of Brazil, but none in the older formations. In North +America the living species only, has been found in Post-Pliocene deposits. +In Europe, however, many species of small opossums, now classed as a +distinct genus, _Peratherium_, have been found in various Tertiary deposits +from the Upper Miocene to the Upper Eocene. + +We have here a sufficient proof that the American Marsupials have nothing +to do with those of Australia, but were derived from Europe, where their +ancestors lived during a long series of ages. + + +FAMILY 77.--DASYURIDÆ. (10 Genera, 30 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2 -- -- + | | | | | + +The Dasyuridæ, or native cats, are a group of carnivorous or insectivorous +marsupials, ranging from the size of a wolf to that of a mouse. They are +found all over Australia and Tasmania, as well as in New Guinea and the +adjacent Papuan islands. Several new genera and species have recently been +described by Mr. G. Krefft, of the Sydney Museum, and are included in the +following enumeration. _Phasgogale_ (3 sp.), New Guinea, West, East, and +South Australia; _Antechinomys_ (1 sp.), Interior of South Australia; +_Antechinus_ (12 sp.), Aru Islands, all Australia, and Tasmania; +_Chætocercus_ (1 sp.), South Australia; _Dactylopsila_ (1 sp.), Aru Islands +and North Australia; _Podabrus_ (5 sp.), West, East, and South Australia, +and Tasmania; _Myoictis_ (1 sp.), Aru Islands; _Sarcophilus_ (1 sp.), +Tasmania; _Dasyurus_ (4 sp.), North, East, and South, Australia, and +Tasmania; _Thylacinus_ (1 sp.), Tasmania (Plate XI., vol. i. p. 439). + +Extinct species of _Dasyurus_ and _Thylacinus_ have been found in the +Post-Pliocene deposits of Australia. + + +{250}FAMILY 78.--MYRMECOBIIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- + | | | | | + +The only representative of this family is the _Myrmecobius fasciatus_, or +native ant-eater, a small bushy-tailed squirrel-like animal, found in the +South and West of Australia. + + +FAMILY 79.--PERAMELIDÆ. (3 Genera, 10 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2 -- -- + | | | | | + +The Peramelidæ, or bandicoots, are small insectivorous Marsupials, having +something of the form of the kangaroos. They range over the whole of +Australia and Tasmania, as well as the Papuan Islands. The genus +_Perameles_ (8 sp.), has the range of the family, one species being found +in New Guinea and the Aru Islands (Plate XI., vol. i. p. 440); _Peragalea_ +(1 sp.), inhabits West Australia only; and _Choeropus_ (1 sp.), a beautiful +little animal with something of the appearance of a mouse-deer, is found in +both South, East, and West Australia. + + +FAMILY 80.--MACROPODIDÆ. (10 Genera, 56 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2 -- -- + | | | | | + +{251}The well-known Kangaroos are the most largely developed family of +Marsupials, and they appear to be the form best adapted for the present +conditions of life in Australia, over every part of which they range. One +genus of true terrestrial kangaroos (_Dorcopsis_), inhabits the Papuan +Islands, as do also the curious tree kangaroos (_Dendrolagus_) which, +without much apparent modification of form, are able to climb trees and +feed upon the foliage. The genera, as established by Mr. Waterhouse, are as +follows: _Macropus_ (4 sp.), West, South, and East Australia, and Tasmania +(Plate XII., vol. i. p. 441); _Osphranter_ (5 sp.), all Australia; +_Halmaturus_ (18 sp.), all Australia and Tasmania; _Petrogale_ (7 sp.), all +Australia; _Dendrolagus_ (2 sp.), New Guinea (Plate X., vol. i. p. 414); +_Dorcopsis_ (2 sp.) Aru and Mysol Islands, and New Guinea; _Onychogalea_ (3 +sp.), Central Australia; _Lagorchestes_ (5 sp.), North, West, and South +Australia; _Bettongia_ (6 sp.), West, South, and East, Australia, and +Tasmania; _Hypsiprymnus_ (4 sp.), West and East Australia, and Tasmania. + +_Extinct Macropodidæ._--Many species of the genera _Macropus_ and +_Hypsiprymnus_ have been found in the cave-deposits and other Post-Tertiary +strata of Australia. Among the extinct genera are _Protemnodon_ and +_Sthenurus_, which are more allied to the tree-kangaroos of New Guinea than +to living Australian species; the gigantic _Diprotodon_, a kangaroo nearly +as large as an elephant; and _Nototherium_, of smaller size. + + +FAMILY 81.--PHALANGISTIDÆ. (8 Genera, 27 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2 -- -- + | | | | | + +The Phalangistidæ, or phalangers, are one of the most varied and +interesting groups of Marsupials, being modified in a variety of ways for +an arboreal life. We have the clumsy-looking tail-less koala, or native +sloth; the prehensile-tailed opossum-like phalangers; the beautiful flying +oppossums, so closely resembling {252}in form the flying squirrels of North +America and India, but often no larger than a mouse; the beautiful +dormouse-like _Dromiciæ_, one species of which is only 2¼ inches long or +less than the harvest-mouse; and the little _Tarsipes_, a true honey-sucker +with an extensile tongue, and of the size of a mouse. These extreme +modifications and specializations within the range of a single family, are +sufficient to indicate the great antiquity of the Australian fauna; and +they render it almost certain that the region it occupied was once much +more extensive, so as to supply the variety of conditions and the struggle +between competing forms of life, which would be required to develop so many +curiously modified forms, of which we now probably see only a remnant. + +The Phalangistidæ not only range over all Australia and Tasmania, but over +the whole of the Austro-Malayan sub-region from New Guinea to the Moluccas +and Celebes. The distribution of the genera is as follows:--_Phascolarctos_ +(1 sp.), the koala, East Australia; _Phalangista_ (5 sp.), East, South, and +West Australia, and Tasmania; _Cuscus_ (8 sp.), woolly phalangers, New +Guinea, North Australia, Timor, Moluccas and Celebes; _Petaurista_ (1 sp.) +large flying phalanger, East Australia; _Belideus_ (5 sp.), flying +opossums, South, East, and North Australia, New Guiana and Moluccas; +_Acrobata_ (1 sp.), pigmy flying opossum, South and East Australia; +_Dromicia_ (5 sp.), dormouse-phalangers, West and East Australia, and +Tasmania; _Tarsipes_ (1 sp.), West Australia. + +_Thylacoleo_, a large extinct marsupial of doubtful affinities, seems to be +somewhat intermediate between this family and the kangaroos. Professor Owen +considered it to be carnivorous, and able to prey upon the huge +_Diprotodon_, while Professor Flower and Mr. Gerard Krefft, believe that it +was herbivorous. + + +FAMILY 82.--PHASCOLOMYIDÆ. (1 Genus, 3 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- + | | | | | + +{253}The Wombats are tail-less, terrestrial, burrowing animals, about the +size of a badger, but feeding on roots and grass. They inhabit South +Australia and Tasmania (Plate XI. vol. i. p. 439). + +An extinct wombat, as large as a tapir, has been found in the Australian +Pliocene deposits. + + +_General Remarks on the Distribution of Marsupialia._ + +We have here the most remarkable case, of an extensive and highly varied +order being confined to one very limited area on the earth's surface, the +only exception being the opossums in America. It has been already shown +that these are comparatively recent immigrants, which have survived in that +country long after they disappeared in Europe. As, however, no other form +but that of the Didelphyidæ occurs there during the Tertiary period, we +must suppose that it was at a far more remote epoch that the ancestral +forms of all the other Marsupials entered Australia; and the curious little +mammals of the Oolite and Trias, offer valuable indications as to the time +when this really took place. + +A notice of these extinct marsupials of the secondary period will be found +at vol. i. p. 159. + + +_Order XIII.--MONOTREMATA._ + +FAMILY 83.--ORNITHORHYNCHIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- + | | | | | + +The _Ornithorhynchus_, or duck-billed Platypus, one of the most remarkable +and isolated of existing mammalia, is found in East and South Australia, +and Tasmania. + + +{254}FAMILY 84.--ECHIDNIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- + | | | | | + +The _Echidna_, or Australian Hedgehog, although quite as remarkable in +internal structure as the Ornithorhynchus, is not so peculiar in external +appearance, having very much the aspect of a hedgehog or spiny armadillo. +The two species of this genus are very closely allied; one inhabits East +and South Australia, the other Tasmania. + +_Extinct Echidnidæ._--Remains of a very large fossil species of _Echidna_ +have lately (1868) been discovered at Darling Downs in Australia. + + +_Remark on the Distribution of the Monotremata._ + +This order is the lowest and most anomalous of the mammalia, and nothing +resembling it has been found among the very numerous extinct animals +discovered in any other part of the world than Australia. + + + + +{255}CHAPTER XVIII. + +THE DISTRIBUTION OF THE FAMILIES AND GENERA OF BIRDS. + + + +_Order I.--PASSERES._ + +FAMILY 1.--TURDIDÆ. (21 Genera, 205 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +The extensive and familiar group of Thrushes ranges over every region and +sub-region, except New Zealand. It abounds most in the North Temperate +regions, and has its least development in the Australian region. Thrushes +are among the most perfectly organized of birds, and it is to this cause, +perhaps, as well as to their omnivorous diet, that they have been enabled +to establish themselves on a number of remote islands. Peculiar species of +true thrush are found in Norfolk Island, and in the small Lord Howes' +Island nearer Australia; the Island of St. Thomas in the Gulf of Guinea has +a peculiar species; while the Mid-Atlantic island Tristan d'Acunha,--one of +the most remote and isolated spots on the globe,--has a peculiarly modified +form of thrush. Several of the smaller West Indian Islands have also +peculiar species or genera of thrushes. + +The family is of somewhat uncertain extent, blending insensibly with the +warblers (Sylviidæ) as well as with the Indian bulbuls {256}(Pycnonotidæ), +while one genus, usually placed in it (_Myiophonus_) seems to agree better +with _Enicurus_ among the Cinclidæ. The genera here admitted into the +thrush family are the following, the numbers prefixed to some of the genera +indicating their position in Gray's _Hand List of the Genera and Species of +Birds_:-- + +(1143) _Brachypteryx_ (8 sp.), Nepaul to Java and Ceylon (this may belong +to the Timaliidæ); _Turdus_ (100 sp.) has the range of the whole family, +abounding in the Palæarctic, Oriental and Neotropical regions, while it is +less plentiful in the Nearctic and Ethiopian, and very scarce in the +Australian; (934) _Oreocincla_ (11 sp.), Palæarctic and Oriental regions, +Australia and Tasmania; (942) _Rhodinocichla_ (1 sp.), Venezuela; (946) +_Melanoptila_ (1 sp.), Honduras; (947 948) _Catharus_ (10 sp.) Mexico to +Equador; (949 950) _Margarops_ (4 sp.), Hayti and Porto Rico to St. Lucia; +(951) _Nesocichla_ (1 sp.), Tristan d'Acunha; (952) _Geocichla_ (8 sp.), +India to Formosa and Celebes, Timor and North Australia; (954 955) +_Monticola_ (8 sp.), Central Europe to South Africa and to China, +Philippine Islands, Gilolo and Java; (956) _Orocætes_ (3 sp.), Himalayas +and N. China; _Zoothera_ (3 sp.) Himalayas, Aracan, Java, and Lombok; +_Mimus_ (20 sp.) Canada to Patagonia, West Indies and Galapagos; (962) +_Oreoscoptes_ (1 sp.), Rocky Mountains and Mexico; (963) _Melanotis_ (2 +sp.), South Mexico and Guatemala; (964) _Galeoscoptes_ (1 sp.), Canada and +Eastern United States to Cuba and Panama; (965 966) _Mimocichla_ (5 sp.), +Greater Antilles; (967 968) _Harporhynchus_ (7 sp.), North America, from +the great lakes to Mexico; _Cinclocerthia_ (3 sp.), Lesser Antilles; (970) +_Rhamphocinclus_ (1 sp.), Lesser Antilles; _Chætops_ (3 sp.), South Africa; +_Cossypha_ = _Bessonornis_ (15 sp.) Ethiopian region and Palestine. + + +FAMILY 2.--SYLVIIDÆ. (74 Genera, 640 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +{257}This immense family, comprising all the birds usually known as +"warblers," is, as here constituted, of almost universal distribution. Yet +it is so numerous and preponderant over the whole Eastern Hemisphere, that +it may be well termed an Old-World group; only two undoubted genera with +very few species belonging to the Nearctic region, while two or three +others whose position is somewhat doubtful, are found in California and the +Neotropical region. + +Canon Tristram, who has paid great attention to this difficult group, has +kindly communicated to me a MSS. arrangement of the genera and species, +which, with a very few additions and alterations, I implicitly follow. He +divides the Sylviidæ into seven sub-families, as follows: + +1. Drymoecinæ (15 genera, 194 sp.), confined to the Old World and +Australia, and especially abundant in the three Tropical regions. 2. +Calamoherpinæ (11 genera, 75 sp.), has the same general distribution as the +last, but is scarce in the Australian and abundant in the Palæarctic +region; 3. Phylloscopinæ (11 genera, 139 sp.), has the same distribution as +the entire family, but is most abundant in the Oriental and Palæarctic +regions. 4. Sylviinæ (6 genera, 33 sp.), most abundant in the Palæarctic +region, very scarce in the Australian and Oriental regions, absent from +America. 5. Ruticillinæ (10 genera, 50 sp.); entirely absent from America +and Australia; abounds in the Oriental and Palæarctic regions. 6. +Saxicolinæ (12 genera, 126 sp.), absent from America (except the extreme +north-west), abundant in the Oriental region and moderately so in the +Palæarctic, Ethiopian, and Australian. 7. Accentorinæ (6 genera, 21 sp.), +absent from the Ethiopian region and South America, most abundant in +Australia, one small genus (_Sialia_), in North America. + +The distribution of the several genera arranged under these sub-families, +is as follows: + +1. DRYMOECINÆ.--(736) Orthotomus (13 sp.), all the Oriental region; (737) +_Prinia_ (11 sp.), all the Oriental region; (738 740 742 746) _Drymoeca_ +(83 sp.), Ethiopian and Oriental regions, most abundant in the former; (743 +to 745 and 749 to 752) _Cisticola_ (32 sp.), Ethiopian and Oriental +regions, with South Europe, China {258}and Australia; (741) _Suya_ (5 sp.), +Nepal to South China and Formosa; (773) _Sphenæacus_ (7 sp.), Australia, +New Zealand, and Chatham Island, with one species (?) in South Africa; (770 +772) _Megalurus_ (4 sp.), Central India to Java and Timor; (774 775) +_Poodytes_ (2 sp.), Australia; (766) _Amytis_ (3 sp.), Australia; (768) +_Sphenura_ (4 sp.), Australia; (764) _Malurus_ (16 sp.), Australia and +Tasmania; (762 763) _Chthonicola_ (3 sp.), Australia; (761) _Calamanthus_ +(2 sp.), Australia and Tasmania; (759) _Camaroptera_ (5 sp.), Africa and +Fernando Po; (753) _Apalis_ (1 sp.), South Africa. + +2. CALAMOHERPINÆ.--(777 to 781 and sp. 2968) _Acrocephalus_ (35 sp.), +Palæarctic, Ethiopian, continental part of Oriental region, Moluccas, +Caroline Islands, and Australia; (782 818) _Dumeticola_ (4 sp.), Nepal to +East Thibet, Central Asia, high regions; (783 790) _Potamodus_ (3 sp.), +Central and South Europe, and East Thibet; (789 and sp. 2969) _Lusciniola_ +(1 sp.), South Europe; (791 792) _Locustella_ (8 sp.), Palæarctic region to +Central India and China; (739) _Horites_ (5 sp.), Nepal to North-west China +and Formosa; (784-786) _Bradyptetus_ = _Cettia_ (10 sp.), South Europe, +Palestine, and South Africa; (747 748) _Catriscus_ (3 sp.), Tropical and +South Africa; _Bernieria_ (2 sp.), and (756) _Ellisia_ (3 sp.), Madagascar; +(832 a) _Mystacornis_ (1 sp.), Madagascar; (787) _Calamodus_ (2 sp.), +Europe and Palestine; (734) _Tatare_ (2 sp.) Samoa to Marquesas Islands. + +3. PHYLLOSCOPINÆ.[5]--_Phylloscopus_ (18 sp.), all Palæarctic and Oriental +regions to Batchian; (757 758 820) _Eremomela_ (16 sp.), Tropical and South +Africa; (754) _Eroessa_ (1 sp.), Madagascar; [5]_Hypolais_ (12 sp.), +Palæarctic region, all India, Timor, North and South Africa; (815 816 819) +_Abrornis_ (26 sp.), Oriental region; (814) _Reguloides_ (4 sp.), +Palæarctic and continental Oriental regions; (822) _Sericornis_ (7 sp.), +Australia and Tasmania (823 824 1451) _Acanthiza_ (14 sp.), Australia and +New Caledonia; (821) _Regulus_ (7 sp.), all Palæarctic and Nearctic regions +and south to Guatemala; (890) _Polioptila_ (13 sp,); Paraguay to New +Mexico; (825) _Gerygone_ (22 sp.), Australia, Papuan and Timor groups, New +Zealand and Norfolk Island. + +{259}4. SYLVIINÆ.--(793) _Aedon_ (9 sp.), Spain and Palestine, to East and +South Africa; (858) _Drymodes_ (2 sp.), Australia; (800) _Pyrophthalma_ (2 +sp.), South Europe and Palestine; (801) _Melizophilus_ (3 sp.), South-west +Europe and North-east Africa; (802 804) _Sylvia_ = _Alsecus_ (8 sp.), +Palæarctic region to India and Ceylon, and North-east Africa; (806 809) +_Curruca_ (7 sp.), Central and South Europe, Madeira, Palestine, Central +India, North-east Africa, and South Africa. + +5. RUTICILLINÆ.--(827) _Luscinia_ (2 sp.), West Asia, Europe, North Africa; +(839) _Cyanecula_ (3 sp.), Europe, North-east Africa, India, Ceylon, and +China; (840) _Calliope_ (2 sp.), North Asia, Himalayas, Central India, and +China; (838) _Erithacus_ (3 sp.), Europe, North-east Africa, Japan, and +North China; (828 830 837) _Ruticilla_ (20 sp.), Palæarctic and Oriental +regions to Senegal and Abyssinia, and east to Timor; abounds in Himalayas; +(829) _Chæmarrhornis_ (1 sp.), Himalayas; (831 832 834) _Larvivora_ (10 +sp.), Oriental region and Japan; (833) _Notodela_ (3 sp.), Himalayas, Pegu, +Formosa, Java; (835) _Tarsiger_ (2 sp.), Nepal; (841) _Grandala_ (1 sp.), +High Himalayas of Nepal. + +6. SAXICOLINÆ.--(975) _Copsychus_ (7 sp.), all Oriental region and +Madagascar; (976) _Kittacincla_ (5 sp.), Oriental region to {260}Ceylon, +Andaman Islands, Formosa, and Borneo; (794-799) _Thamnobia_ (10 sp.), +Ethiopian region and India to foot of Himalayas; (977) _Gervasia_ (2 sp.), +Madagascar and Seychelle Islands; (845 847) _Dromolæa_ (18 sp.), Africa to +South Europe, Palestine, North-west India, and North China; (842 843 846) +_Saxicola_ (36 sp.), Africa, North-west India, whole Palæarctic region, +migrating to Alaska and Greenland; (848 849) _Oreicola_ (5 sp.), Timor, +Lombok, and Burmah; (844) _Cercomela_ (6 sp.), North-east Africa to +North-west India; (850) _Pratincola_ (15 sp.), Europe, Ethiopian, and +Oriental regions to Celebes and Timor; (917) _Ephthianura_ (3 sp.), +Australia; (851-856) _Petroeca_ (17 sp.), Australian region, Papua to New +Zealand, Chatham and Auckland Islands, and Samoa; (857) _Miro_ (2 sp.), New +Zealand (doubtfully placed here). + +7. ACCENTORINÆ.--(771) _Cinclorhamphus_ (2 sp.), Australia; (860) _Origma_ +(1 sp.), East Australia; (859) _Sialia_ (8 sp.), United States to +Guatemala; (861) _Accentor_ (12 sp.), Palæarctic region to Himalayas and +North-west China; (703) _Orthonyx_ (4 sp.), East Australia and New Zealand +(doubtfully placed here). + +The following two genera, which have been usually classed as Ampelidæ, are +arranged by Messrs. Sclater and Salvin in the Sylviidæ:-- + +(1362) _Myiadestes_ (8 sp.), Peru and Bolivia, along the Andes to Mexico +and California, also the Antilles; (1364) _Cichlopsis_ (1 sp.), Brazil. + + +FAMILY 3.--TIMALIIDÆ. (35 Genera, 240 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --| -- 2 -- 4 |1. 2. 3. 4 |1. 2. 3. 4 | 1. 2 -- 4 + | | | | | + +The Timaliidæ, or babbling thrushes, are a group of small strong-legged +active birds, mostly of dull colours, which are especially characteristic +of the Oriental region, in every part of which they abound, while they are +much less plentiful in {261}Australia and Africa. The Indo-Chinese +sub-region is the head quarters of the family, whence it diminishes rapidly +in all directions in variety of both generic and specific forms. Viscount +Walden has kindly assisted me in the determination of the limits of this +family, as to which there is still much difference of opinion. The +distribution of the genera here admitted is as follows; and as the genera +are widely scattered in the _Hand List_, reference numbers are prefixed in +every case. + +(1023-1026 1008) _Pomatorhinus_ (27 sp.), the whole Oriental region +(excluding Philippines), Australia and New Guinea; (1027) _Pterohinus_ (3 +sp.), North China, East Thibet; (1029 1030) _Malacocircus_ (9 sp.), +Continental India and Ceylon, Arabia, Nubia; (1031) _Chatarrhæa_ (5 sp.), +Abyssinia, Palestine, India, Nepal, Burmah, and Philippines; (1032) +_Layardia_ (3 sp.), India and Ceylon; (1033) _Acanthoptila_ (1 sp.), Nepal; +(1034) _Cinclosoma_ (4 sp.), Australia and Tasmania; (1035 1036) +_Crateropus_ (18 sp.), all Africa, Persia; (1037) _Hypergerus_ (1 sp.), +West Africa; (1038) _Cichladusa_ (3 sp.), Tropical Africa; (1039) +_Garrulax_ (23 sp.), the Oriental region (excluding Philippines); (1040) +_Janthocincla_ (10 sp.), Nepal, to East Thibet, Sumatra, Formosa; (1041 +1042) _Gampsorhynchus_ (2 sp.), Himalayas; (1049) _Grammatoptila_ (1 sp.), +North India; (1043-1045) _Trochalopteron_ (24 sp.), all India to China and +Formosa; (1046) _Actinodura_ (4 sp.), Nepal to Burmah, 3,000-10,000 feet; +(1047) _Pellorneum_ (4 sp.), Nepal to Ceylon, Tenasserim; (1158 1159) +_Timalia_ (12 sp.), Malaya;[6] (1160) _Dumetia_ (2 sp.), Central India and +Ceylon; (1162) _Stachyris_ (6 sp.), Nepal to Assam, Sumatra, Formosa; +(1164) _Pyctorhis_ (3 sp.), India to Ceylon and Burmah; (1165) _Mixornis_ +(8 sp.), Himalayas and Malaya; (1167) _Malacopteron_ (3 sp.), Malaya; (1168 +1169) _Alcippe_ (15 sp.), Ceylon and South India, Himalayas to Aracan, +Malaya, Formosa, New Guinea; (1170) _Macronus_ (2 sp.), Malaya; (1171) +_Cacopitta_ (5 sp.), Malaya; (1172) _Trichastoma_ (11 sp.), Nepal, Burmah, +Malaya, Celebes; (1173) _Napothera_ (6 sp.), Malaya; (1174) _Drymocataphus_ +(8 sp.), Burmah, Malaya, Ceylon, {262}Timor; (1175) _Turdinus_ (5 sp.), +Khasia Hills, Malacca, Tenasserim; (1176) _Trichixos_ (1 sp.), Borneo, +Malacca; (1004) _Sibia_ (6 sp.), Nepal to Assam, Tenasserim, Formosa; (1177 +1178) _Alethe_ (4 sp.), West Africa; (1178 a) _Oxylabes_ (1 sp.), +Madagascar; (1050) _Psophodes_ (2 sp.), South, East, and West Australia; +(1048) _Turnagra_ (3 sp.), New Zealand. + + +FAMILY 4.--PANURIDÆ. (4 Genera, 13 Species). + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --| 1. 2 -- 4 |-- -- -- --|-- -- 3 -- |-- -- -- -- + | | | | | + +This new family is adopted, at the suggestion of Professor Newton, to +include some peculiar groups of Himalayan birds whose position has usually +been among the Timaliidæ or the Paridæ, but which are now found to be +allied to our Bearded Reedling. The supposed affinity of this bird for the +Tits has been long known to be erroneous, and the family Panuridæ was +formed for its reception (Yarrell's _British Birds_, 4th edit. p. 512). The +genera having hitherto been widely scattered in systematic works, are +referred to by the numbers of Mr. G. E. Gray's _Hand List_. + +(1901) _Paradoxornis_ (3 sp.), Himalayas and East Thibet; (1904) +_Conostoma_ (1 sp.), Himalayas and East Thibet; (876) _Suthora_ (8 sp.), +Himalayas to North-west China, Formosa; (877) _Chlenasicus_ (1 sp.), +Darjeeling; (887) _Panurus_ (1 sp.), Central and Southern Europe; (1902) +_Heteromorpha_ (1 sp.), Nepal, 10,000 feet altitude; _Cholornis_ (1 sp.), +Moupin in East Thibet. + + +FAMILY 5.--CINCLIDÆ. (4 Genera, 27 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- | -- 2 -- 4 |1. 2. 3. 4 |-- -- -- ?4|1. 2. 3. 4 |1 -- -- -- + | | | | | + +{263}The Cinclidæ consist of a number of more or less thrush-like +ground-birds, of which the most remarkable are the Dippers, forming the +genus _Cinclus_. These are curiously distributed, from the Palæarctic +region as a centre, to the alpine districts of North and South America; +while the three genera which are here included as somewhat allied to +_Cinclus_, all inhabit the Oriental region. The genera which I class in +this family are the following:-- + +(978) _Cinclus_ (9 sp.), Palæarctic region to West China and Formosa, Rocky +Mountains, and Mexico in North America, and southward to the Andes of Peru; +(916) _Enicurus_ (9 sp.), Himalayas to Java and West China; (979) _Eupetes_ +(4 sp.), Indo-Malay sub-region and New Guinea; (971) _Myiophonus_ (5 sp.), +Himalayas to Ceylon, Java, South China, and Formosa. + +(981) _Mesites_ (1 sp.), Madagascar, is an anomalous bird placed with +_Eupetes_ by Mr. G. R. Gray, but of very uncertain affinities. + + +FAMILY 6.--TROGLODYTIDÆ. (17 Genera, 94 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- |-- -- 3. 4 |1 -- -- -- + | | | | | + +The Troglodytidæ, or Wrens, are small birds, rather abundant and varied in +the Neotropical region, with a few species scattered through the Nearctic, +Palæarctic, and parts of the Oriental regions, and one doubtful genus in +Africa. The constitution of the family is by no means well determined. The +South American genera are taken from Messrs. Sclater and Salvin's +_Nomenclator Avium Neotropicalium_. + +_Tesia_ (2 sp.), Eastern Himalayas; _Pnoepyga_ (6 sp.), Himalayas to East +Thibet, Java; (716 and 723) _Troglodytes_ (15 sp.), Neotropical, Nearctic, +and Palæarctic regions to the Higher Himalayas; (697) _Rimator_ (1 sp.), +Darjeeling; _Thryothorus_ (13 sp.), South Brazil to Mexico, Martinique, and +Nearctic region; _Thryophilus_ (13 sp.), Brazil to Mexico, and North-west +America; _Cistothorus_ {264}(5 sp.), Patagonia to Greenland; _Uropsila_ (1 +sp.), Mexico; _Donacobius_ (2 sp.), Tropical America; _Campylorhynchus_ (18 +sp.), Brazil, and Bolivia to Mexico and the Gila valley; _Cyphorhinus_ (5 +sp.), Equatorial South America to Costa Rica; _Microcerculus_ (5 sp.), +Brazil and Peru to Mexico; _Henicorhina_ (2 sp.), Peru and Guiana to Costa +Rica; _Salpinctes_ (1 sp.), High Plains of Rocky Mountains; _Catherpes_ (1 +sp.), Mexico and Rio Grande; _Cinnicerthia_ (2 sp.), Ecuador and Columbia. +(760) _Sylvietta_ (2 sp.), Tropical and South Africa,--is placed in this +family by Mr. Tristram. + + +FAMILY 7.--CHAMÆIDÆ. (1 Genus, 1 Species). + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|1 -- -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The bird which forms the genus _Chamæa_ inhabits California; and though +allied to the wrens it has certain peculiarities of structure which, in the +opinion of many ornithologists, require that it should be placed in a +distinct family. + + +FAMILY 8.--CERTHIIDÆ. (6 Genera, 18 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- 3 -- |1. 2. 3. 4 |-- -- -- --| 1 -- 3. 4 | 1.2 -- -- + | | | | | + +The Certhiidæ, or Creepers, form a small family whose species are thinly +scattered over North America from Mexico, the Palæarctic region, parts of +the Oriental region, and Australia, where they are somewhat more abundant. +The distribution of the genera is as follows: + +_Certhia_ (6 sp.), Nearctic and Palæarctic regions, Nepal, and Sikhim; +_Salpornis_ (1 sp.), Central India; _Tichodroma_, (1 sp.), South +{265}Europe to Abyssinia, Nepal, and North China; _Rhabdornis_ (1 sp.), +Philippine Islands; _Climacteris_ (8 sp.), Australia and New Guinea. + + +FAMILY 9.--SITTIDÆ. (6 Genera, 31 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- --|1. 2. 3. 4 | 1. 2 -- 4 + | | | | | + +The Sittidæ, or Nuthatches, are another small family of tree-creeping +birds, whose distribution is very similar to that of the Certhiidæ, but +with a more uniform range over the Oriental region, and extending to New +Zealand and Madagascar. The genera are as follows:-- + +_Sitta_ (17 sp.), Palæarctic and Nearctic regions to South India and +Mexico; _Dendrophila_ (2 sp.), Ceylon and India to Burmah and Malaya; +_Hypherpes_ (1 sp.), Madagascar; _Sittella_ (6 sp.), Australia and New +Guinea. _Acanthisitta_ (1 sp.) and _Xenicus_ (4 sp.), New Zealand, are +placed with some doubt in this family. + + +FAMILY 10.--PARIDÆ. (14 Genera, 92 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3. 4 | -- 2 -- 4 + | | | | | + +The Paridæ, or Tits, are very abundant in the Nearctic and Palæarctic +regions; many fine species are found in the Himalayas, but they are +sparingly scattered through the Ethiopian, Oriental, and Australian +regions. The genera usually admitted into this family are the following, +but the position of some of them, especially of the Australian forms, is +doubtful. + +(864-867 870) _Parus_ (46 sp.), North America, from Mexico, Palæarctic, and +Oriental regions, Tropical and South Africa; {266}(868 869) _Lophophanes_ +(10 sp.), Europe, the Higher Himalayas to Sikhim, North America to Mexico; +_Acredula_ = _Orites_ (6 sp.), Palæarctic region; _Melanochlora_ (2 sp.), +Nepal to Sumatra; _Psaltria_ (1 sp.), Java; _Psaltriparus_ (3 sp.), +Guatemala to California, and Rocky Mountains; _Auriparus_ (1 sp.), Rio +Grande; (881 882) _Parisoma_ (5 sp.), Tropical and South Africa; (883 884) +_Ægithalus_ (6 sp.), South-east Europe to South Africa; (885 889) +_Ægithaliscus_ (6 sp.), Afghanistan and Himalayas to Amoy; _Cephalopyrus_ +(1 sp.), North-west Himalayas; _Sylviparus_ (1 sp.), Himalayas and Central +India; _Certhiparus_ (2 sp.), New Zealand; (879 880) _Sphenostoma_ (2 sp.), +East and South Australia. + + +FAMILY 11.--LIOTRICHIDÆ. (11 Genera, 35 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- 3. 4 |-- -- -- -- + | | | | | + +The Liotrichidæ, or Hill-Tits, are small, active, delicately-coloured +birds, almost confined to the Himalayas and their extension eastward to +China. They are now generally admitted to form a distinct family. The +genera are distributed as follows: + +(1146) _Liothrix_ (3 sp.), Himalayas to China; _Siva_ (3 sp.), Himalayas; +_Minla_ (4 sp.), Himalayas and East Thibet; _Proparus_ (7 sp.), Nepal to +East Thibet and Aracan; (1153) _Pteruthius_ (6 sp.), Himalayas to Java and +West China; (1155) _Cutia_ (2 sp.), Nepal; (1019) _Yuhina_ (3 sp.), High +Himalayas and Moupin; (1020) _Ixulus_ (3 sp.), Himalayas to Tenasserim; +(1021) _Myzornis_ (1 sp.), Darjeeling. + + +FAMILY 12.--PHYLLORNITHIDÆ. (3 Genera, 14 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3. 4 |-- -- -- -- + | | | | | + +{267}The Phyllornithidæ, or "Green Bulbuls," are a small group of +fruit-eating birds, strictly confined to the Oriental region, and ranging +over the whole of it, with the one exception of the Philippine Islands. The +genera are:-- + +(1022) _Phyllornis_ (12 sp.), India to Java, Ceylon, and Hainan; (1166) +_Iora_ (4 sp.), the whole Oriental region; (1163) _Erpornis_ (2 sp.), +Borneo, Himalayas, Hainan, and Formosa. + + +FAMILY 13.--PYCNONOTIDÆ. (9 Genera, 139 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --| -- 2 -- 4 |1. 2. 3. 4 |1. 2. 3. 4 |1 -- -- -- + | | | | | + +The Pycnonotidæ, Bulbuls, or fruit-thrushes, are highly characteristic of +the Oriental region, in every part of which they abound; less plentiful in +the Ethiopian region, and extending to Palestine and Japan in the +Palæarctic, and to the Moluccas in the Australian region, but absent from +the intervening island of Celebes. The genera are:-- + +MICROSCELIS (6 sp.), Burmah, the Indo-Malay Islands, and Japan; +_Pycnonotus_ (52 sp., in many sub-genera), Palestine to South Africa, the +whole Oriental region, China and Japan; _Alcurus_ (1 sp.), Himalayas; +_Hemixus_ (2 sp.), Nepal, Bootan, Hainan; _Phyllastrephus_ (4 sp.), West +and South Africa; _Hypsipetes_ (20 sp.), the whole Oriental region, +Madagascar and the Mascarene Islands; _Tylas_ (1 sp.), Madagascar; +_Criniger_ (30 sp.), the whole Oriental region (excluding Philippines), +West and South Africa, Moluccas; _Ixonotus_ (7 sp.), West Africa; (1015 +1017) _Setornis_ (3 sp.), Malacca, Sumatra, and Borneo; _Iole_ (4 sp.), +Aracan and Malaya; _Andropadus_ (9 sp.), Tropical Africa; (1157) +_Lioptilus_ (1 sp.), South Africa. + + +{268}FAMILY 14.--ORIOLIDÆ. (5 Genera, 40 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --| 1. 2 -- 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Orioles, or Golden Thrushes, are a small group characteristic of the +Oriental and Ethiopian regions, migrating into the western Palæarctic +region, and with some of the less typical forms in Australia. The genera +are:-- + +_Oriolus_ (24 sp.), Central Europe, throughout Africa, and the whole +Oriental region, northward to Pekin, and eastward to Flores; (1073) +_Analcipus_ (3 sp.), Himalayas, Formosa, Java and Borneo; _Mimeta_ (9 sp.), +the Moluccas and Australia; _Sphecotheres_ (3 sp.), Timor and Australia. +_Artamia_ (1 sp.), Madagascar,--perhaps belongs to the next family or to +Laniidæ. + + +FAMILY 15.--CAMPEPHAGIDÆ (3 Genera, 100 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +The Campephagidæ, or Cuckoo Shrikes, (Campephaginæ of the _Hand List_, with +the addition of _Cochoa_) are most abundant in the Australian region +(especially in the Austro-Malay sub-region), less so in the Oriental, and +still less in the Ethiopian region. The genera, for the most part as +adopted by Dr. Hartlaub, are as follows:-- + +_Pericrocotus_ (22 sp.), the whole Oriental region, extending north to +Pekin, and east to Lombok; (1242-1244) _Lanicterus_ (4 sp.), West and South +Africa; (1245 1246) _Graucalus_ (25 sp.), the whole Oriental region, and +eastward to Austro-Malaya, the New {269}Hebrides, and Tasmania; _Artamides_ +(1 sp.), Celebes; _Pteropodocys_ (1 sp.), Australia; (1248 1250 1257 1258) +_Campephaga_ (16 sp.), Austro-Malaya, and New Caledonia, Philippines, the +Ethiopian region; _Volvocivora_ (8 sp.) the Oriental region (excluding +Philippines); _Lalage_ (18 sp.), the whole Malay Archipelago to New +Caledonia and Australia; _Symmorphus_ (1 sp.), Australia; _Oxynotus_ (2 +sp.), Mauritius and Bourbon; (1204) _Cochoa_ (3 sp.), Himalayas, Java. The +position of this last genus is doubtful. Jerdon puts it in the Liotrichidæ; +Sundeval in the Sturnidæ; Bonaparte in the Dicruridæ; Professor Newton +suggests the Pycnonotidæ; but it seems on the whole best placed here. + + +FAMILY 16.--DICRURIDÆ. (6 Genera, 58 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Dicruridæ, or Drongo Shrikes (Dicruridæ of the _Hand List_, omitting +the genus _Melænornis_), have nearly the same distribution as the last +family, with which they are sometimes united. They are, however, most +abundant and varied in the Oriental region, much less so both in the +Australian and Ethiopian regions. The distribution of the genera is as +follows:-- + +_Dicrurus_ (46 sp., in several sub-genera), has the range of the whole +family, extending east to New Ireland, and one species in Australia; +_Chætorhynchus_ (1 sp.), New Guinea; _Bhringa_ (2 sp.), Himalayas to Borneo +(Plate IX. vol. i. p. 339); _Chibia_ (2 sp.), Himalayas eastward to North +China; _Chaptia_ (3 sp.), all India to Malacca and Formosa; _Irena_ (4 +sp.), Central India, Assam, and Burmah to Borneo and the Philippine +Islands. This last genus is placed by Jerdon among the Pycnonotidæ, but +seems to come most naturally here or in the last family. + + +{270}FAMILY 17.--MUSCICAPIDÆ. (44 Genera, 283 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Muscicapidæ, or Flycatchers (Muscicapinæ and Myiagrinæ of the _Hand +List_, omitting _Cochoa_ and including _Pogonocichla_) form an extensive +family of usually small-sized and often bright-coloured birds, very +abundant in the warmer regions of the Old World and Australia, but becoming +scarce as we approach the temperate and colder regions. They are wholly +absent from North and South America. The genera, many of which are not well +defined, are distributed as follows:-- + +_Peltops_ (1 sp.), Papuan Islands; _Monarcha_ (28 sp.), Moluccas to the +Carolines and Marquesas Islands, Australia and Tasmania; _Leucophantes_ (1 +sp.), New Guinea; _Butalis_ (4 sp.), Ethiopian and Palæarctic regions, +Moluccas and Formosa; _Muscicapa_ (12 sp.), Europe and Africa; +_Muscicapula_ (6 sp.), India to Western China; _Alseonax_ (1 sp.), South +Africa; _Erythrosterna_ (7 sp.), Europe to China and Java; _Newtonia_ (1 +sp.), Madagascar; _Xanthopygia_ (2 sp.), Japan, China, Malacca; _Hemipus_ +(1 sp.), India and Ceylon; _Pycnophrys_ (1 sp.), Java; _Hyliota_ (2 sp.), +West Africa; _Erythrocercus_ (2 sp.), West Africa and Zambesi; _Micræca_ (6 +sp.), Australia, Timor, and Papuan Islands; _Artomyias_ (2 sp.), West +Africa; _Pseudobias_ (1 sp.), Madagascar; _Hemichelidon_ (3 sp.), the +Oriental region and North China; _Smithornis_ (2 sp.), West and South +Africa; _Megabias_ (1 sp.), West Africa; _Cassinia_ (2 sp.), West Africa; +_Bias_, (1 sp.), Tropical Africa; _Niltava_ (3 sp.), Himalayas to West +China; _Cyornis_ (16 sp.), the whole Oriental region; _Cyanoptila_ (1 sp.), +Japan, China, Hainan; _Eumyias_ (7 sp.), India to South China, Ceylon, and +Sumatra; (1213 and 1216) _Siphia_ (8 sp.), North India, Formosa, Timor; +_Anthipes_ (1 sp.), Nepal; _Seisura_ (5 sp.), Australia and +{271}Austro-Malaya (excluding Celebes); _Myiagra_ (16 sp.), Australia and +Moluccas to Caroline and Samoa Islands; _Hypothymis_ (2 sp.), Oriental +region and Celebes; _Elminia_ (2 sp.), Tropical Africa; _Muscitodus_ (2 +sp.), Fiji Islands; _Machærirhynchus_ (4 sp.), Papuan Islands and North +Australia; _Platystira_ (12 sp.), Tropical and South Africa; _Rhipidura_ +(45 sp.), the Oriental and Australian regions to the Samoa Islands and +Tasmania; _Chelidorynx_ (1 sp.), North India; _Myialestes_ (2 sp.), India +to Ceylon, China, Java and Celebes; _Tchitrea_ (26 sp.), the entire +Ethiopian and Oriental regions, and to North China and Japan; _Philentoma_ +(4 sp.), Malacca, Sumatra, Borneo, and Philippine Islands; _Todopsis_ (6 +sp.), Papuan Islands; (836) _Pogonocichla_ (1 sp.), South Africa; +(1061-1063) _Bradyornis_ (7 sp.), Tropical and South Africa; (1460) +_Chasiempis_ (2 sp.), Sandwich Islands. + + +FAMILY 18.--PACHYCEPHALIDÆ. (5 Genera, 62 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 |-- -- 3. 4 |1. 2. 3 -- + | | | | | + +The Pachycephalidæ, or Thick-headed Shrikes (Pachycephalinæ of the _Hand +List_ omitting _Colluricincla_, _Cracticus_, and _Pardalotus_) are almost +confined to the Australian region, a single species extending to Java and +Aracan, and another (?) to Madagascar. The family has generally been united +with the Laniidæ, but most modern ornithologists consider it to be +distinct. The distribution of the genera is as follows:-- + +_Oreoeca_ (1 sp.), Australia; _Falcunculus_ (2 sp.), Australia; +_Pachycephala_ (44 sp.), Sula Islands (east of Celebes) to the Fiji +Islands, and Australia; _Hylocharis_ (4 sp.), Timor, Celebes, Indo-Malaya, +and Aracan; _Calicalicus_ (1 sp.), Madagascar; _Eopsaltria_ (14 sp.), +Australia, New Caledonia, and the New Hebrides; _Artamia_ (4 sp.), +Madagascar,--may belong to this family, or to Laniidæ, Oriolidæ, or +Artamidæ, according to different authors. + + +{272}FAMILY 19.--LANIIDÆ. (19 Genera, 145 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +The Laniidæ, or Shrikes (Laniinæ and Malaconotinæ of the _Hand List_, and +including _Colluricincla_), are most abundant and varied in Africa, less +plentiful in the Oriental, Australian, and Palæarctic regions, with a few +species in the Nearctic region as far as Mexico. The constitution of the +family is, however, somewhat uncertain. The genera here admitted are:-- + +_Colluricincla_ (4 sp.), Australia and Tasmania; _Rectes_ (18 sp.), Papuan +Islands, North Australia, to Pelew and Fiji Islands; (1462-1464 1466 1470 +1471-1473) _Lanius_ (50 sp.), the whole Nearctic, Palæarctic, Ethiopian, +and Oriental regions, one species reaching Timor, none in Madagascar; +_Laniellus_ (1 sp.), Java; _Hypocolius_ (1 sp.), Abyssinia and Upper Nile; +_Corvinella_ (1 sp.), South and West Africa; _Urolestes_ (1 sp.), South and +East Africa; _Tephrodornis_ (4 sp.), Oriental region to Hainan and Java; +_Hypodes_ (1 sp.), West Africa; _Fraseria_ (2 sp.), West Africa; +_Cuphopterus_ (1 sp.), Princes' Island; _Nilaus_ (1 sp.), South and West +Africa; _Prionops_ (9 sp.), Tropical Africa; _Eurocephalus_ (2 sp.), North, +East, and South Africa, and Abyssinia; _Chaunonotus_ (1 sp.), West Africa; +_Vanga_ (4 sp.), Madagascar (Plate VI. vol. i. p. 278); _Laniarius_ (36 +sp.), the whole Ethiopian region; _Telephonus_ (10 sp.), all Africa and +South Europe; _Meristes_ (2 sp.), Tropical and South Africa; _Nicator_ (1 +sp.), East Africa. + + +FAMILY 20.--CORVIDÆ. (24 Genera, 190 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +{273}The Corvidæ, or Crows, Jays, &c., form an extensive and somewhat +heterogeneous group, some members of which inhabit almost every part of the +globe, although none of the genera are cosmopolitan. The true crows are +found everywhere but in South America; the magpies, choughs, and +nutcrackers are characteristic of the Palæarctic region; the jays are +Palæarctic, Oriental, and American; while the piping crows are peculiarly +Australian. The more detailed distribution of the genera is as follows:-- + +Sub-family I. Gymnorhininæ (Piping Crows).--_Strepera_ (4 sp.), and +_Gymnorhina_ (3 sp.), are Australian only; _Cracticus_ (9 sp.), ranges from +New Guinea to Tasmania (this is usually put with the Shrikes, but it has +more affinity with the preceding genera); _Pityriasis_ (1 sp.), Borneo (an +extraordinary bird of very doubtful affinities); _Grallina_ (1 sp.), +Australia, is put here by Sundevall,--among Motacillidæ, by Gould. + +Sub-family II. _Garrulinæ_ (Jays).--_Platylophus_ = _Lophocitta_ (4 sp.), +Malaya; _Garrulus_ (12 sp.), Palæarctic region, China and Himalayas; +_Perisoreus_ (2 sp.), North of Palæarctic and Nearctic regions; _Cyanurus_ +(22 sp.), American, from Bolivia to Canada, most abundant in Central +America, but absent from the Antilles; _Cyanocorax_ (15 sp.), La Plata to +Mexico; _Calocitta_ (2 sp.), Guatemala and Mexico; _Psilorhinus_ (3 sp.), +Costa Rica to Texas; _Urocissa_ (6 sp.), Western Himalayas to China and +Formosa; _Cissa_ (3 sp.), South-eastern Himalayas to Tenasserim, Ceylon, +Sumatra, and Java. + +Sub-family III. Dendrocittinæ (Tree Crows).--_Temnurus_ (3 sp.), Cochin +China, Malacca to Borneo (not Java); _Dendrocitta_ (9 sp.), the Oriental +region to Sumatra, Hainan, and Formosa; _Crypsirhina_ (3 sp.), Pegu, Siam, +and Java; _Ptilostomus_ (2 sp.), West, East, and South Africa. + +Sub-family IV. Corvinæ (Crows and Magpies).--_Nucifraga_ (4 sp.), +Palæarctic region to the Himalayas and North China; _Picicorvus_ (1 sp.), +the Rocky Mountains and California; _Gymnokitta_ (1 sp.), Rocky Mountains +and Arizona (Plate XVIII., Vol. II., p. 128); _Pica_ (9 sp.), Palæarctic +region, Arctic America, and California; _Cyanopica_ (3 sp.), Spain, +North-east Asia, Japan; {274}_Streptocitta_ (2 sp.), Celebes; _Charitornis_ +(1 sp.), Sula Islands; _Corvus_ (55 sp.), universally distributed except +South America and New Zealand, but found in Guatemala and the Antilles to +Porto Rico; reaches the extreme north of Europe and Asia; _Gymnocorvus_ (2 +sp.), Papuan Islands; _Picathartes_ (1 sp.), West Africa; _Corvultur_ (2 +sp.), Tropical and South Africa. + +Sub-family V. Fregilinæ (Choughs).--_Fregilus_ (3 sp.), mountains and +cliffs of Palæarctic region from West Europe to the Himalayas and North +China, Abyssinia (Plate I., Vol. I., p. 195); _Corcorax_ (1 sp.), +Australia. + + +FAMILY 21.--PARADISEIDÆ. (19 Genera, 34 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2 -- -- + | | | | | + +The Paradiseidæ, or "Birds of Paradise," form one of the most remarkable +families of birds, unsurpassed alike for the singularity and the beauty of +their plumage. Till recently the family was restricted to about eight +species of the more typical Paradise birds, but in his splendid monograph +of the group, Mr. Elliot has combined together a number of allied forms +which had been doubtfully placed in several adjacent families. The various +species of true Paradise birds, having ornamental plumes developed from +different parts of the body, are almost wholly confined to New Guinea and +the adjacent Papuan Islands, one species only being found in the Moluccas +and one in North Australia; while the less typical Bower-birds, having no +such developments of plumage, are most characteristic of the north and east +of Australia, with a few species in New Guinea. The distribution of the +genera according to Mr. Elliot's monograph is as follows:-- + +Sub-family I. Paradiseinæ.--_Paradisea_ (4 sp.), Papuan Islands; +_Manucodia_ (3 sp.), Papuan Islands and North Australia; _Astrapia_ (1 +sp.), New Guinea; _Parotia_ (1 sp.), New Guinea; _Lophorhina_ (1 sp.), New +Guinea; _Diphyllodes_ (3 sp.), Papuan {275}Islands; _Xanthomelus_ (1 sp.), +New Guinea; _Cicinnurus_ (1 sp.), Papuan Islands; _Paradigalla_ (1 sp.), +New Guinea; _Semioptera_ (1 sp.), Gilolo and Batchian. + +Sub-family II. Epimachinæ.--_Epimachus_ (1 sp.), New Guinea; _Drepanornis_ +(1 sp.), New Guinea; _Seleucides_ (1 sp.), New Guinea (Plate X., Vol. I., +p. 414); _Ptilorhis_ (4 sp.), New Guinea and North Australia. + +Sub-family III. Tectonarchinæ (Bower-birds).--_Sericulus_ (1 sp.), Eastern +Australia; _Ptilonorhynchus_ (1 sp.), Eastern Australia; _Chlamydodera_ (4 +sp.), North and East Australia; _Ælurædus_ (3 sp.), Papuan Islands and East +Australia; _Amblyornis_ (1 sp.), New Guinea. + + +FAMILY 22.--MELIPHAGIDÆ. (23 Genera, 190 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3. 4 + | | | | | + + (As in the _Hand List_, but omitting Zosterops, and slightly altering the + arrangement.) + +The extensive group of the Meliphagidæ, or Honey-suckers, is wholly +Australian, for the genus _Zosterops_, which extends into the Oriental and +Ethiopian regions, does not naturally belong to it. Several of the genera +are confined to Australia, others to New Zealand, while a few range over +the whole Australian region. The genera are distributed as follows:-- + +_Myzomela_ (18 sp.), has the widest range, extending from Celebes to the +Samoa Islands, and to Timor and Eastern Australia; _Entomophila_ (4 sp.), +Australia and New Guinea; _Gliciphila_ (10 sp.), Australia, Timor, New +Guinea, and New Caledonia; _Acanthorhynchus_ (2 sp.), Australia and +Tasmania; Meliphaga (1 sp.), Australia; _Ptilotis_ (40 sp.), Gilolo and +Lombok to Australia and Tasmania, and to the Samoa and Tonga Islands; +_Meliornis_ (5 sp.), Australia and Tasmania; _Prosthemadera_ (1 sp.), +_Pogonornis_ (1 sp.), New Zealand; _Anthornis_ (4 sp.), New Zealand and +Chatham Islands; _Anthochæra_ (4 sp.), Australia and Tasmania; +{276}_Xanthotis_ (4 sp.), Papuan Islands and Australia; _Leptornis_ (2 +sp.), Samoa Islands and New Caledonia; _Philemon_ = _Tropidorhyncus_ (18 +sp.), Moluccas and Lombok to New Guinea, Australia, Tasmania and New +Caledonia; _Entomiza_ (2 sp.), Australia; _Manorhina_ (5 sp.), Australia +and Tasmania; _Euthyrhynchus_ (3 sp.), New Guinea; _Melirrhophetes_ (2 +sp.), New Guinea; _Melidectes_ (1 sp.), New Guinea; _Melipotes_ (1 sp.), +New Guinea; _Melithreptus_ (8 sp.), New Guinea, Australia, and Tasmania; +(397) _Moho_ (3 sp.), Sandwich Islands; _Chætoptila_ (1 sp.), Sandwich +Islands. + + +FAMILY 23.--NECTARINIIDÆ. (11 Genera, 122 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Nectariniidæ, or Sun-birds, form a rather extensive group of +insectivorous honey-suckers, often adorned with brilliant metallic plumage, +and bearing a superficial resemblance to the American humming-birds, +although not in any way related to them. They abound in the Ethiopian, +Oriental, and Australian regions, as far east as New Ireland, and south to +Queensland, while one species inhabits the hot Jordan Valley in the +Palæarctic region. For the Eastern genera I follow Lord Walden's +classification (Ibis, 1870); the African species not having been so +carefully studied are mostly placed in one genus. The genera adopted are as +follows:-- + +_Promerops_ (1 sp.), South Africa; _Nectarinia_ (60 sp.), the whole +Ethiopian region; _Cinnyricinclus_ (5 sp.), West Africa; _Neodrepanis_ (1 +sp.), Madagascar; _Arachnecthra_ (13 sp.), Palestine, all India to Hainan, +the Papuan Islands, and North-east Australia; _Æthopyga_ (15 sp.), +Himalayas and Central India to West China, Hainan, Java, and Northern +Celebes; _Nectarophila_ (5 sp.), Central India and Ceylon, Assam and Aracan +to Java, Celebes and the Philippines; _Chalcostetha_ (6 sp.), Malay +Peninsula to New Guinea; _Anthreptes_ (1 sp.), Siam, Malay Peninsula to +{277}Sula Islands, and Flores; _Cosmeteira_ (1 sp.), Papuan Islands; +_Arachnothera_ (15 sp.), the Oriental region (excluding Philippines) +Celebes, Lombok, and Papuan Islands. + + +FAMILY 24.--DICÆIDÆ. (5 Genera, 107 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Dicæidæ, or Flower-peckers, consist of very small, gaily-coloured +birds, rather abundant over the whole Oriental and much of the Australian +regions, and one genus extending over the Ethiopian region. The genera here +adopted are the following:-- + +(622) _Zosterops_ (68 sp.), the whole Ethiopian, Oriental, and Australian +regions, as far east as the Fiji Islands, and north to Pekin and Japan; +(400-403) _Dicæum_ (25 sp.), the whole Oriental region, except China, with +the Australian region as far as the Solomon Islands; (404) _Pachyglossa_ (2 +sp. 1437 1442), Nepal and Northern Celebes; (405) _Piprisoma_ (2 sp.), +Himalayas to Ceylon and Timor; (1450) _Pardalotus_ (10 sp.), Australia and +Tasmania; (407-409) _Prionochilus_ (5 sp.), Indo-Malay sub-region and +Papuan Islands. + + +FAMILY 25.--DREPANIDIDÆ. (4 Genera, 8 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- 3 -- + | | | | | + +The Drepanididæ are confined to the Sandwich Islands, and I follow Mr. +Sclater's suggestion in bringing together the following genera to form this +family:-- + +_Drepanis_ (3 sp.); _Hemignathus_ (3 sp.); _Loxops_ (1 sp.); _Psittirostra_ +(1 sp.). If these are correctly associated, the great {278}differences in +the bill indicate that they are the remains of a larger and more varied +family, once inhabiting more extensive land surfaces in the Pacific. + + +FAMILY 26.--COEREBIDÆ. (11 Genera, 55 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |-- -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + + (According to the arrangement of Messrs. Sclater and Salvin.) + +The Coerebidæ, or Sugar-birds, are delicate little birds allied to the +preceding families, but with extensile honey-sucking tongues. They are +almost wholly confined to the tropical parts of America, only one species +of _Certhiola_ ranging so far north as Florida. The following is the +distribution of the genera:-- + +_Diglossa_ (14 sp.), Peru and Bolivia to Guiana and Mexico; _Diglossopis_ +(1 sp.), Ecuador to Venezuela; _Oreomanes_ (1 sp.), Ecuador; _Conirostrum_ +(6 sp.), Bolivia to Ecuador and Columbia; _Hemidacnis_ (1 sp.), Upper +Amazon and Columbia; _Dacnis_ (13 sp.), Brazil to Ecuador and Costa Rica; +_Certhidea_ (2 sp.), Galapagos Islands; _Chlorophanes_ (2 sp.), Brazil to +Central America and Cuba; _Coereba_ (4 sp.), Brazil to Mexico; _Certhiola_ +(10 sp.), Amazon to Mexico, West Indies, and Florida; _Glossoptila_ (1 +sp.), Jamaica. + + +FAMILY 27.--MNIOTILTIDÆ. (18 Genera, 115 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |1. 2. 3. 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + + (Messrs. Sclater and Salvin are followed for the Neotropical, Baird and + Allen for the Nearctic region.) + +The Mniotiltidæ, or Wood-warblers, are an interesting group of small and +elegant birds, allied to the preceding family and to the greenlets, and +perhaps also to the warblers and tits of Europe. {279}They range over all +North America from Panama to the Arctic regions, but do not extend far +beyond the tropic in Southern America. They are almost as abundant in the +Nearctic as in the Neotropical region; and considering the favourable +conditions of existence in Tropical America, this fact, in connection with +their absence from the South Temperate zone would lead us to suppose that +they originated in North Temperate America, and subsequently spread +southward into the tropics. This supposition is strengthened by the fact +that their metropolis, in the breeding season, is to the north of the +United States. The genera adopted by Messrs. Sclater and Salvin are as +follows:-- + +(918) _Siurus_ (4 sp.), Venezuela and West Indies to Eastern States and +Canada; _Mniotilta_ (1 sp.), Venezuela, Mexico, and Antilles to the Eastern +States; _Parula_ (5 sp.), Brazil to Mexico, and the Eastern States, and +Canada; _Protonotaria_ (1 sp.), Antilles to Ohio; _Helminthophaga_ (8 sp.), +Columbia to Arctic America; _Helmintherus_ (2 sp.), Central America to +Eastern States; _Perissoglossa_ (1 sp.), Antilles and Eastern States; +_Dendroeca_ (33 sp.), Amazon to Antilles, and Arctic America, and south to +Chili; _Oporornis_ (2 sp.), Guatemala to Eastern States; _Geothlypis_ (11 +sp.), all North America and Brazil; _Myiodioctes_ (5 sp.), all North +America and Columbia; _Basileuterus_ (22 sp.), Bolivia and Brazil to +Mexico; _Setophaga_ (15 sp.), Brazil to Canada; _Ergaticus_ (2 sp.), +Guatemala and Mexico; _Cardellina_ (1 sp.), Guatemala and Mexico; (1440) +_Granatellus_ (3 sp.), Amazon to Mexico; (1441) _Teretristis_ (2 sp.), +Cuba; (1439) _Icteria_ (2 sp.), Costa Rica and United States to Canada. + + +FAMILY 28.--VIREONIDÆ. (7 Genera, 63 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |1. 2. 3. 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +(Messrs. Sclater and Salvin are followed for the Neotropical genera; +Professor Baird and Mr. Allen for those of the Nearctic region.) + +{280}The Vireonidæ, or Greenlets, are a family of small fly-catching birds +wholly restricted to the American continent, where they range from Paraguay +to Canada. They are allied to the Mniotiltidæ and perhaps also to the +Australian Pachycephalidæ. Only two of the genera, with about a dozen +species, inhabit the Nearctic region. The distribution of the genera is as +follows:-- + +_Vireosylvia_ (13 sp.), Venezuela to Mexico, the Antilles, the Eastern +States and Canada; _Vireo_ (14 sp.), Central America and the Antilles to +Canada; _Neochloe_ (1 sp.), Mexico; _Hylophilus_ (20 sp.), Brazil to +Mexico; _Laletes_ (1 sp.), Jamaica; _Vireolanius_ (5 sp.), Amazonia to +Mexico; _Cychlorhis_ (9 sp.), Paraguay to Mexico. + + +FAMILY 29.--AMPELIDÆ. (4 Genera, 9 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Ampelidæ, represented in Europe by the waxwing, are a small family, +characteristic of the Nearctic and Palæarctic regions, but extending +southward to Costa Rica and the West Indian islands. The genera are +distributed as follows:-- + +(1539) _Ampelis_ (3 sp.), the Palæarctic and Nearctic regions, and +southward to Guatemala; (1360) _Ptilogonys_ (2 sp.), Central America; +(1442) _Dulus_ (2 sp.), West Indian Islands; (1361) _Phænopepla_ (1 sp.), +Mexico and the Gila Valley. + + +FAMILY 30.--HIRUNDINIDÆ. (9 Genera, 91 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +{281}The Hirundinidæ, or Swallows, are true cosmopolites. Although they do +not range quite so far north (except as stragglers) as a few of the extreme +polar birds, yet they pass beyond the Arctic Circle both in America and +Europe, _Cotyle riparia_ having been observed in the Parry Islands, while +_Hirundo rustica_ has been seen both in Spitzbergen and Nova Zembla. +_Cotyle riparia_ and _Chelidon urbica_ also breed in great numbers in +northern Lapland, latitude 67° to 70° north. Many of the species also, have +an enormous range, the common swallow (_Hirundo rustica_) inhabiting +Europe, Asia and Africa, from Lapland to the Cape of Good Hope and to the +Moluccas. The genera of swallows are not well determined, a number having +been established of which the value is uncertain. I admit the following, +referring by numbers to the _Hand List_:-- + +(215-221 226-228) _Hirundo_ (40 sp.), the range of the entire family; (222 +223) _Psalidoprogne_ (10 sp.), Tropical and South Africa; (224) _Phedina_ +(1 sp.), Madagascar and Mascarene Islands; (225) _Petrochelidon_ (5 sp.), +North and South America and Cape of Good Hope; (220-232 ?234) _Atticora_ (8 +sp.), the Neotropical region and ? Australia; (235 237) _Cotyle_ (11 sp.), +Europe, India, Africa, North America, Antilles and Ecuador; (236) +_Stelgidopteryx_ (5 sp.), La Plata to United States; (238 and 239) +_Chelidon_ (6 sp.), Palæarctic region, Nepal, Borneo; (240-242) _Progne_ (5 +sp.), all North and South America. + + +FAMILY 31.--ICTERIDÆ. (24 Genera, 110 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Icteridæ, or American hang-nests, range over the whole continent, from +Patagonia and the Falkland Islands to the Arctic Circle. Only about 20 +species inhabit the Nearctic region, while, as usual with exclusively +American families, the larger proportion of the genera and species are +found in the {282}tropical parts of South America. The genera adopted by +Messrs. Sclater and Salvin are the following:-- + +_Clypeicterus_ (1 sp.), Upper Amazon; _Ocycalus_ (2 sp.), Upper Amazon to +Mexico; _Ostinops_ (8 sp.), Brazil and Bolivia to Mexico; _Cassiculus_ (1 +sp.), Mexico; _Cassicus_ (10 sp.), South Brazil and Bolivia to Costa Rica; +_Icterus_ (34 sp.), La Plata to the Antilles and United States; +_Dolichonyx_ (1 sp.), Paraguay to Canada; _Molothrus_ (8 sp.), La Plata to +Northern United States; _Agelæus_ (7 sp.), La Plata and Chili to Northern +United States; _Xanthocephalus_ (1 sp.), Mexico to California and Canada; +_Xanthosomus_ (4 sp.), La Plata to Venezuela; _Amblyrhamphus_ (1 sp.), La +Plata and Bolivia; _Gymnomystax_ (1 sp.), Amazonia and Guiana; +_Pseudoleistes_ (2 sp.), La Plata and Brazil; _Leistes_ (3 sp.), La Plata +to Venezuela; _Sturnella_ (5 sp.), Patagonia and Falkland Islands to Middle +United States; _Curæus_ (1 sp.), Chili; _Nesopsar_ (1 sp.), Jamaica; +_Scolecophgaus_ (2 sp.), Mexico to Arctic Circle; _Lampropsar_ (4 sp.), +Amazonia and Ecuador to Mexico; _Quiscalus_ (10 sp.), Venezuela and +Columbia to South and Central United States; _Hypopyrrhus_ (1 sp.), +Columbia; _Aphobus_ (1 sp.), Brazil and Bolivia; _Cassidix_ (2 sp.), Brazil +to Mexico and Cuba. + + +FAMILY 32.--TANAGRIDÆ. (43 Genera, 304 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |-- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Tanagers are an extensive family of varied and beautiful fruit-eating +birds, almost peculiar to the Neotropical region, only four species of a +single genus (_Pyranga_) extending into the Eastern United States and Rocky +Mountains. Southward they range to La Plata. They are especially abundant +in the forest regions of South America east of the Andes, where no less +than 40 out of the 43 genera occur; 23 of the genera are peculiar to this +sub-region, while only 1 (_Phlogothraupis_) is {283}peculiar to Central +America and Mexico, and 2 (_Spindalis_ and _Phænicophilus_) to the West +Indian islands. The genera adopted by Messrs. Sclater and Salvin with their +distribution will be found at Vol. II., p. 99, in our account of +Neotropical Zoology. + + +FAMILY 33.--FRINGILLIDÆ. (74 Genera, 509 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- -- + | | | | | + +The great family of the Fringillidæ, or finches, is in a very unsettled +state as regards their division into genera, the most divergent views being +held by ornithologists as to the constitution and affinities of many of the +groups. All the Australian finch-like birds appear to belong to the +Ploceidæ, so that the finches, as here constituted, are found in every +region and sub-region, except the Australian region from which they are +entirely absent--a peculiar distribution hardly to be found in any other +family of birds. + +Many European ornithologists separate the Emberizidæ, or buntings, as a +distinct family, but as the American genera have not been so divided I am +obliged to keep them together; but the genera usually classed as "buntings" +are placed last, as a sub-family. In the following arrangement of the +genera, I have done what I could to harmonize the views of the best modern +writers. For convenience of reference the succession of the genera is that +of the _Hand List_, and the numbers of the sub-genera are given whenever +practicable:-- + +(1793 1795) _Fringilla_ (6 sp.), the whole Palæarctic region, including the +Atlantic Islands; (1794) _Acanthis_ (3 sp.), Europe to Siberia, Persia, and +North-West Himalayas; (1796) _Procarduelis_ (1 sp.), High Himalayas and +East Thibet; (1797-1803) _Chrysomitris_ (18 sp.), Neotropical and Nearctic +regions, Europe, and Siberia; (1804) _Metoponia_ (1 sp.), East Europe to +North West Himalayas; (1805 and 1809) _Chlorospiza_ (9 sp.), Palæarctic +region and Africa to the {284}Cape of Good Hope; (1806-1809) _Dryospiza_ +(14 sp.), South Europe, Palestine, Canaries, and all Africa; (1810) +_Sycalis_ (18 sp.), the whole Neotropical region; (1811-1813 1816-1819) +_Pyrgita_ (34 sp.), Palæarctic and Oriental regions, and all Africa; (1814) +_Montifringilla_ (4 sp.), Palæarctic region; (1815) _Fringillauda_ (2 sp.), +North-West Himalayas to East Thibet; (1820-1822) _Coccothraustes_ (6 sp.), +Palæarctic region and Nepal, Nearctic region to Mexico; (1823) _Eophona_ (2 +sp.), China and Japan; (1824) _Mycerobas_ (2 sp.), Central Asia to Persia, +High Himalayas, and East Thibet; (1825) _Chaunoproctus_ (1 sp.), Bonin +Islands, south-east of Japan, (probably Palæarctic); (1826) _Geospiza_ (7 +sp.), Galapagos Islands; (1827) _Camarhynchus_ (5 sp.), Galapagos Islands; +(1828) _Cactornis_ (4 sp.), Galapagos Islands; (1830-1832) _Phrygilus_ (10 +sp.), Columbia to Fuegia and the Falkland Islands; (1833) _Xenospingus_ (1 +sp.), Peru; (1834) _Diuca_ (3 sp.), Peru to Chili and Patagonia; (1835 and +1837) _Emberizoides_ (3 sp.), Venezuela to Paraguay; (1836) _Donacospiza_ +(1 sp.), South Brazil and La Plata; (1839) _Chamæospiza_ (1 sp.), Mexico; +(1838 and 1840) _Embernagra_ (9 sp.), Arizona to La Plata; (1841) +_Hæmophila_ (6 sp.), Mexico to Costa Rica; (1842) _Atlapetes_ (1 sp.), +Mexico; (1843) _Pyrgisoma_ (5 sp.). Mexico to Costa Rica; (1844 and 1845) +_Pipilo_ (12 sp.), all North America to Guatemala; (1846) _Junco_ (6 sp.), +all the United States to Guatemala; (1847) _Zonotrichia_ (9 sp.), the whole +Nearctic and Neotropical regions; (1848 1849) _Melospiza_ (7 sp.), Sitka +and United States to Guatemala; (1850) _Spizella_ (7 sp.), Canada to +Guatemala; (1851) _Passerella_ (4 sp.), the Nearctic region and Northern +Asia; (1852) _Passerculus_ (6 sp.), Nearctic region and to Guatemala; +(1853) _Pooecetes_ (1 sp.), all United States and Mexico; (1854) +_Ammodromus_ (4 sp.), all United States to Guatemala; (1855) _Coturniculus_ +(6 sp.), north and east of North America to Jamaica and Bolivia; (1856) +_Peucæa_ (6 sp.), South Atlantic States and California to Mexico; (1857) +_Tiaris_ (1 sp.), Brazil; (1858) _Volatinia_ (1 sp.), Mexico to Brazil and +Bolivia; (1859) _Cyanospiza_ (5 sp.), Canada to Guatemala; (1860 1861) +_Paroaria_ (6 sp.), Tropical South America, east of the Andes; (1862) +_Coryphospingus_ (4 sp.), Tropical South America; (1863) _Haplospiza_ (2 +sp.), Mexico and Brazil; (1864 1891) _Phonipara_ (8 sp.), Mexico to +Columbia, the greater Antilles; (1865) _Poospiza_ {285}(13 sp.), California +and South Central States to Bolivia and La Plata; (424) _Spodiornis_ (1 +sp.), Andes of Quito; (1866 1867) _Pyrrhula_ (9 sp.), the whole Palæarctic +region to the Azores and High Himalayas; (1868) _Crithagra_ (17 sp.), +Tropical and South Africa, Mauritius, Syria; (1869) _Ligurnus_ (2 sp.), +West Africa; (1870 1871) _Carpodacus_ (18 sp.), Nearctic and Palæarctic +regions to Mexico and Central India; (1872-1874) _Erythrospiza_ (6 sp.), +Southern parts of Palæarctic region; (1875) _Uragus_ (2 sp.), Siberia and +Japan; (1876) _Cardinalis_ (2 sp.), South and Central States to Venezuela; +(1877) _Pyrrhuloxia_ (1 sp.), Texas and Rio Grande; (1878 1879) _Guiraca_ +(6 sp.), Southern United States to La Plata; (1880) _Amaurospiza_ (2 sp.), +Costa Rica and Brazil; (1881) _Hedymeles_ (2 sp.), all United States to +Columbia; (1882) _Pheucticus_ (5 sp.), Mexico to Peru and Bolivia; (1883) +_Oryzoborus_ (6 sp.), Mexico to Ecuador and South Brazil; (1884) +_Melopyrrha_ (1 sp.), Cuba; (1885) _Loxigilla_ (4 sp.), Antilles; (1886 +1887) _Spermophila_ (44 sp.), Texas to Bolivia and Uruguay; (1888) +_Catamenia_ (4 sp.), Columbia to Bolivia; (1889) _Neorhynchus_ (3 sp.), +West Peru; (1892) _Catamblyrhyncus_ (1 sp.), Columbia; (1893) _Loxia_ (7 +sp.), Europe to North-west India and Japan, Arctic America to Pennsylvania, +Mexico; (1894) _Pinicola_ (3 sp.), Arctic America, North-east Europe to the +Amoor, Camaroons Mountains West Africa; (1895) _Propyrrhula_ (1 sp.), +Darjeeling in the winter,? Thibet; (1896) _Pyrrhospiza_ (1 sp.), Snowy +Himalayas; (1897) _Hæmatospiza_ (1 sp.), South-east Himalayas, 5,000-10,000 +feet; (1898 1899) _Linota_ (12 sp.), Europe to Central Asia, north and east +of North America; (1900) _Leucosticte_ (7 sp.), Siberia and Thibet to +Kamschatka, and from Alaska to Utah. + +Sub-family Emberizinæ.--(1995) _Calamospiza_ (1 sp.), Arizona and Texas to +Mexico; (1906) _Chondestes_ (2 sp.), Western, Central, and Southern States +to Mexico and Nicaragua; (1907-1910) _Euspiza_ (9 sp.), Palæarctic region, +India, Burmah, and South China, South-east United States to Columbia; +(1911-1920) _Emberiza_ (28 sp.), the whole Palæarctic region (continental), +to Central India in winter; (1921) _Gubernatrix_ (1 sp.), Paraguay and La +Plata, (according to Messrs. Sclater and Salvin this comes next to +_Pipilo_); (1922) _Fringillaria_ (8 sp.), Africa and South Europe; +{286}(1923-1925) _Plectrophanes_ (6 sp.), Arctic Zone to Northern Europe +and North China, Arctic America, and east side of Rocky Mountains; (1926) +_Centronyx_ (1 sp.), Mouth of Yellowstone River. + + +FAMILY 34.--PLOCEIDÆ. (29 Genera, 252 species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +The Ploceidæ, or Weaver-finches, are especially characteristic of the +Ethiopian region, where most of the genera and nearly four-fifths of the +species are found; the remainder being pretty equally divided between the +Oriental and Australian regions. Like the true finches these have never +been properly studied, and it is exceedingly difficult to ascertain what +genera are natural and how far those of Australia and Africa are distinct. +The following enumeration must therefore be taken as altogether tentative +and provisional. When the genera adopted differ from those of the _Hand +List_ they will be referred to by numbers. + +_Textor_ (5 sp.), Tropical and South Africa; (1650-1654 1657) +_Hyphantornis_ (32 sp.), Tropical and South Africa; (1655 1656) +_Symplectes_ (8 sp.), Tropical and South Africa; _Malimbus_ (9 sp.), West +Africa; (1659 1661) _Ploceus_ (6 sp.), West and East Africa, the Oriental +region (excluding Philippines); (1660) _Nelicurvius_ (1 sp.), Madagascar; +_Foudia_ (12 sp.), Madagascar and Mascarene Islands, Tropical Africa; (1663 +1664) _Sporopipes_ (2 sp.), Tropical and South Africa; (1665-1667) +_Pyromelana_ (14 sp.), Tropical and South Africa, Abyssinia to 10,500 feet; +_Philetærus_ (1 sp.), South Africa; _Nigrita_ (7 sp.), West Africa to Upper +Nile; _Plocepasser_ (4 sp.), East and South Africa; (1672-1674) _Vidua_ (7 +sp.), Tropical and South Africa (Plate V., Vol. I., p. 264); (1675-1677) +_Coliuspasser_ (9 sp.), Tropical and South Africa; _Chera_ (1 sp.), South +Africa; _Spermospiza_ (2 sp.), West Africa; _Pyrenestes_ (6 sp.), Tropical +and South Africa; (1682-1687 1689 1692 1693 1698) _Estrilda_ (26 sp.), +Tropical and South Africa, India, Burmah, and Java to Australia; (1688 1690 +1691 1695 1696) {287}_Pytelia_ (24 sp.), Tropical and South Africa; (1694) +_Hypargos_ (2 sp.), Mozambique and Madagascar; (1697) _Emblema_ (1 sp.), +North-west Australia (1699 1712-1717) _Amadina_ (15 sp.), Tropical and +South Africa, Moluccas to Australia and the Samoa Islands; (1700 1701 1710) +_Spermestes_ (8 sp.), Tropical Africa and Madagascar; (1702) _Amauresthes_ +(1 sp.), East and West Africa; (1703 1707-1709 1711) _Munia_ (30 sp.), +Oriental region to Timor and New Guinea; (1704) _Donacola_ (3 sp.), +Australia; (1705 1706) _Poephila_ (6 sp.), Australia; (1718-1721) +_Erythrura_ (7 sp.), Sumatra to Java, Moluccas, Timor, New Guinea, and Fiji +Islands; (1722) _Hypochera_ (3 sp.), Tropical and South Africa. + + +FAMILY 35.--STURNIDÆ. (29 Genera, 124 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 | 1 -- 3. 4 + | | | | | + +The Sturnidæ or Starlings, are a highly characteristic Old-World group, +extending to every part of the great Eastern continent and its islands, and +over the Pacific Ocean to the Samoa Islands and New Zealand, yet wholly +absent from the mainland of Australia. The family appears to be tolerably +well-defined, and the following genera are generally considered to belong +to it: (1558 1559 1562) _Eulabes_ (13 sp.), the Oriental region to +South-west China, Hainan, and Java,--and Flores, New Guinea and the Solomon +Islands in the Australian region; _Ampeliceps_ (1 sp.), _Tenasserim_, +Burmah, and Cochin China; _Gymnops_ (1 sp.), Philippine Islands; +_Basilornis_ (2 sp.), Celebes and Ceram; _Pastor_ (1 sp.), South-east +Europe to India, Ceylon, and Burmah; _Acridotheres_ (7 sp.), the whole +Oriental region and Celebes; (1568 1569) _Sturnia_ (12 sp.), the whole +Oriental region, North China, Japan, and Siberia, Celebes; _Dilophus_ (1 +sp.), South Africa; _Sturnus_ (6 sp.), Palæarctic region, to India and +South China in winter; _Sturnopastor_ (4 sp.), India to Burmah and East +Java; _Creadion_ (2 sp.), New Zealand; _Heterolocha_ (1 sp.), New Zealand; +(1520) _Callæas_ {288}(2 sp.), New Zealand; _Buphaga_ (2 sp.), Tropical and +South Africa; _Euryceros_ (1 sp.), Madagascar (see Plate VI., Vol. I., p. +278.) This genus and the last should perhaps form distinct families. (1577) +_Juida_ (5 sp.), Central, West, and South Africa; (1578) _Lamprocolius_ (20 +sp.), Tropical and South Africa; _Cinnyricinclus_ (2 sp.), Tropical and +South Africa; _Onychognathus_ (2 sp.), West Africa; (1581) _Spreo_ (4 sp.), +Tropical and South Africa; (1582-1585) _Amydrus_ (7 sp.), South and East +Africa, Palestine; _Aplonis_ (9 sp.), New Caledonia to the Tonga Islands; +(1587-1589) _Calornis_ (18 sp.), the whole Malay Archipelago and eastward +to the Ladrone and Samoa Islands; (1590) _Enodes_ (1 sp.), Celebes; +_Scissirostrum_ (1 sp.), Celebes; (1592) _Saroglossa_ (1 sp.), Himalayas; +(1593) _Hartlaubius_ (1 sp), Madagascar; _Fregilupus_ (1 sp.), Bourbon, but +it has recently become extinct; (363) _Falculia_ (1 sp)., Madagascar. + + +FAMILY 36.--ARTAMIDÆ. (1 Genus, 17 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\___________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS |SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+------------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --?|1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +The Artamidæ, or Swallow-shrikes, are a curious group of birds, ranging +over the greater part of the Oriental and Australian regions as far east as +the Fiji Islands and south to Tasmania. Only a single species inhabits +India, and they are more plentiful in Australia than in any other locality. +The only well-marked genus is _Artamus_. + +There are a few Madagascar birds belonging to the genus _Artamia_, which +some ornithologists place in this family, others with the Laniidæ, but +which are here classed with the Oriolidæ. + + +{289}FAMILY 37.--ALAUDIDÆ. (15 Genera, 110 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Alaudidæ, or Larks, may be considered as exclusively belonging to the +great Eastern continent, since the Nearctic, Neotropical, and Australian +regions have each only a single species. They abound most in the open +plains and deserts of Africa and Asia, and are especially numerous in South +Africa. The genera, including those recently established by Mr. Sharpe, are +as follows:-- + +_Otocorys_ (8 sp.); the Palæarctic region, North America and south to the +Andes of Columbia, North India; (1928 1929) _Alauda_ (17 sp.), Palæarctic +region, all Africa, the Peninsula of India, and Ceylon; (1931) _Galerita_ +(10 sp.), Central Europe to Senegal and Abyssinia, Persia, India and North +China; (1932) _Calendula_ (2 sp.), Abyssinia and South Africa; (1933 1934) +_Calandrella_ (6 sp.), Europe, North Africa, India, Burmah, North China, +and Mongolia; (1935-1937) _Melanocorypha_ (7 sp.), South Europe to Tartary, +Abyssinia, and North-west India; _Pallasia_ (sp. 7781), East Asia; (1938) +_Certhilauda_ (4 sp.), South Europe, South Africa; _Heterocorys_ (sp. 7792) +South Africa; (1939) _Alæmon_ (3 sp.), South-east Europe to Western India, +and South Africa; (1940) _Mirafra_ (25 sp.), the Oriental and Ethiopian +regions to Australia; (1941) _Ammomanes_ (10 sp.), South Europe to +Palestine and Central India, and to Cape Verd Islands and South Africa; +(1942 1943) _Megalophonus_ (6 sp.), Tropical and South Africa; +_Tephrocorys_ (1 sp.), South Africa; _Pyrrhulauda_ (9 sp.), all Africa, +Canary Islands, India and Ceylon. + + +{290}FAMILY 38.--MOTACILLIDÆ. (9 Genera, 80 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. -- 4 + | | | | | + +The Motacillidæ, or Wagtails and Pipits, are universally distributed, but +are most abundant in the Palæarctic, Ethiopian, and Oriental regions, to +which the true wagtails are almost confined. The following genera are +usually adopted, but some of them are not very well defined:-- + +_Motacilla_ (15 sp.), ranges over the greater part of Europe, Asia, and +Africa, and to Alaska in North-west America; _Budytes_ (10 sp.), Europe, +Africa, Asia to Philippines, Moluccas, Timor, and North Australia; +_Calobates_ (3 sp.), South Palæarctic and Oriental regions to Java; +_Nemoricola_ (1 sp.), Oriental region; _Anthus_ (30 sp.), all the great +continents; _Neocorys_ (1 sp.), Central North America; _Corydalla_ (14 +sp.), South Europe to India, China, the Malay Islands, Australia, New +Zealand and the Auckland Islands: _Macronyx_ (5 sp.), Tropical and South +Africa; _Heterura_ (1 sp.), Himalayas. + + +FAMILY 39.--TYRANNIDÆ. (71 Genera, 329 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Tyrannidæ, or Tyrant Shrikes, form one of the most extensive and truly +characteristic American families of birds; as they extend over the whole +continent from Patagonia to the Arctic regions, and are found also in all +the chief American islands--the Antilles, the Galapagos, the Falkland +Islands, and {291}Juan Fernandez. As the genera are all enumerated in the +table, at p. 101 of this volume, I shall here confine myself to the +distribution of the sub-families, only referring to such genera as are of +special geographical interest. + +Sub-family I. CONOPHAGINÆ (2 genera, 13 species). Confined to tropical +South America, from Brazil and Bolivia to Guiana and Columbia. + +Sub-family II. TÆNIOPTERINÆ (19 genera, 76 species). This group ranges from +Patagonia and the Falkland Islands to the northern United States; yet it is +almost wholly South American, only 2 genera and 4 species passing north of +Panama, and none inhabiting the West Indian islands. _Empidias_ has 3 +species in North America, while _Tænioptera_, _Cnipolegus_, +_Muscisaxicola_, and _Centrites_, range south to Patagonia. + +Sub-family III. PLATYRHYNICHINÆ (16 genera, 60 species). This sub-family is +wholly Neotropical and mostly South American, only 7 of the genera passing +Panama and but 3 reaching Mexico, while there are none in the West Indian +islands. Only 3 genera extend south to the temperate sub-region, and one of +these, _Anæretes_, has a species in Juan Fernandez. + +Sub-family IV. ELAINEINÆ (17 genera, 91 species). This sub-family is more +exclusively tropical, only two genera extending south as far as Chili and +La Plata, while none enter the Nearctic region. No less than 10 of the +genera pass north of Panama, and one of these, _Elainea_, which ranges from +Chili to Costa Rica has several species in the West Indian islands. About +one fourth of the species of this sub-family are found north of Panama. + +Sub-family V. TYRANNINÆ (17 genera, 89 species). This sub-family is that +which is best represented in the Nearctic region, where 6 genera and 24 +species occur. _Milvulus_ reaches Texas; _Tyrannus_ and _Myiarchus_ range +over all the United States; _Sayornis_, the Eastern States and California; +_Contopus_ extends to Canada; _Empidonax_ ranges all over North America; +and _Pyrocephalus_ reaches the Gila Valley as well as the Galapagos +Islands. No less than 5 genera of this sub-family occur in the West Indian +islands. + + +{292}FAMILY 39_a_.--OXYRHAMPHIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The genus _Oxyrhamphus_ (2 sp.) which ranges from Brazil to Costa Rica, has +usually been placed in the Dendrocolaptidæ; but Messrs Sclater and Salvin +consider it to be the type of a distinct family group, most allied to the +Tyrannidæ. + + +FAMILY 40.--PIPRIDÆ. (15 Genera, 60 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Pipridæ, or Manakins, have generally been associated with the next +family, and they have a very similar distribution. The great majority of +the genera and species are found in the equatorial regions of South +America, only 9 species belonging to 5 genera ranging north of Panama, +while 2 or 3 species extend to the southern limit of the tropical forests +in Paraguay and Brazil. The genera which go north of Panama are _Piprites_, +_Pipra_, _Chiroxiphia_, _Chiromachæris_, and _Hetoropelma_. _Pipra_ is the +largest genus, containing 19 species, and having representatives throughout +the whole range of the family. As in all the more extensive families +peculiar to the Neotropical region, the distribution of the genera will be +found in the tables appended to the chapter on the Neotropical region in +the Third Part of this work. (Vol. II. p. 103). + + +{293}FAMILY 41.--COTINGIDÆ. (28 Genera, 93 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Cotingidæ, or Chatterers, comprise some of the most beautiful and some +of the most remarkable of American birds, for such we must consider the +azure and purple Cotingas, the wine-coloured white-winged Pompadour, the +snowy carunculated Bell-birds, the orange-coloured Cocks-of-the-Rock, and +the marvellously-plumed Umbrella-birds, (Plate XV. Vol. II. p. 28). The +Cotingidæ are also one of the most pre-eminently Neotropical of all the +Neotropical families, the great mass of the genera and species being +concentrated in and around the vast equatorial forest region of the Amazon. +Only 13 species extend north of Panama, one to the Antilles, and not more +than 20 are found to the south of the Amazon Valley. Messrs. Sclater and +Salvin divide the family into six sub-families, the distribution of which +will be briefly indicated. + +Sub-family I. TITYRINÆ (3 genera, 22 species). Ranges from Brazil to +Mexico, one species of _Hadrostomus_ inhabiting Jamaica. + +Sub-family II. LIPAUGINÆ (4 genera, 14 species) also ranges from Brazil to +Mexico; one genus (_Ptilochloris_) is confined to Brazil. + +Sub-family III. ATTALINÆ (2 genera, 10 species). Ranges from Paraguay to +Costa Rica; one genus (_Casiornis_) is confined to South Brazil and +Paraguay. + +Sub-family IV. RUPICOLINÆ (2 genera, 5 species). This sub-family is +restricted to the Amazonian region and Guiana, with one species extending +along the Andean valleys to Bolivia. The genera are _Rupicola_ (3 species) +and _Phænicocercus_ (2 species). + +Sub-family V. COTINGINÆ (10 genera, 28 species). Ranges from Southern +Brazil and Bolivia to Nicaragua; only two species {294}(belonging to the +genera _Carpodectes_ and _Cotinga_) are found north of Panama, and there +are none in the West Indian islands. The great majority of these, the true +Chatterers, are from the regions about the Equator. + +Sub-family VI. GYMNODERINÆ (7 genera, 14 species). Ranges from Brazil to +Costa Rica; two species, of the genera _Chasmorhynchus_ and +_Cephalopterus_, are found north of Panama, while there are none in the +West Indian islands. Only 2 species are found south of the Amazon valley. + + +FAMILY 42.--PHYTOTOMIDÆ. (1 Genus, 3 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1 -- -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Phytotomidæ, or Plant-cutters, are singular thick-billed birds, +strictly confined to the temperate regions of South America. The single +genus, _Phytotoma_, is found in Chili, La Plata, and Bolivia. Their +affinities are uncertain, but they are believed to be allied to the series +of families with which they are here associated. (Plate XVI. Vol. II. p. +128). + + +FAMILY 43.--EURYLÆMIDÆ. (6 Genera, 9 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- 3. 4 |-- -- -- -- + | | | | | + +The Eurylæmidæ, or Broad-bills, form a very small family of birds, often +adorned with striking colours, and which have their nearest allies in the +South American Cotingidæ. They have a very limited distribution, from the +lower slopes of the Himalayas through Burmah and Siam, to Sumatra, Borneo, +and Java. They are evidently the remains of a once extensive group, and +from the small number of specific forms remaining, seem to be on {295}the +road to extinction. Thus we may understand their isolated geographical +position. The following are the names and distribution of the genera:-- + +_Eurylæmus_ (2 species), Malay Peninsula, Sumatra, Java, and Borneo; +_Corydon_ (1 species), Malacca, Sumatra and Borneo (Plate IX. Vol. I. p. +339); _Psarisomus_ (1 species), Himalayas to Burmah, up to 6,000 feet; +_Serilophus_ (2 species), Nepal to Tenasserim; _Cymbirhynchus_ (2 species), +Siam to Sumatra and Borneo; _Calyptomena_ (1 species), Penang to Sumatra +and Borneo. + + +FAMILY 44.--DENDROCOLAPTIDÆ. (43 Genera, 217 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Dendrocolaptidæ, or American Creepers, are curious brown-coloured birds +with more or less rigid tail feathers, strictly confined to the continental +Neotropical region, and very numerous in its south-temperate extremity. +They are divided by Messrs. Sclater and Salvin into five sub-families, to +which I shall confine my remarks on their distribution. The details of the +numerous genera, being only interesting to specialists, will be given in +the table of genera of the Neotropical region. No less than 13 of the +genera are confined to South-Temperate America and the High Andes; 14 are +restricted to Tropical South America, while not one is peculiar to Tropical +North America, and only 15 of the 43 genera extend into that sub-region, +showing that this is one of the pre-eminently South American groups. + +Sub-family I. FURNARIINÆ (8 genera, 30 species). Ranges over all South +America, 4 genera and 18 species being restricted to the temperate +sub-region; one species is found in the Falkland Islands. + +Sub-family II. SCLERURINÆ (1 genus, 6 species). Brazil to Guiana, Columbia, +and north to Mexico. + +Sub-family III. SYNALLAXINÆ (12 genera, 78 species). Ranges from Patagonia +to Mexico; 7 genera and 28 species are confined {296}to the temperate +sub-region; species occur in the islands of Mas-a-fuera, Trinidad, and +Tobago. + +Sub-family IV. PHILYDORINÆ (6 genera, 35 species). Confined to Tropical +America from Brazil to Mexico; 4 genera and 8 species occur in Tropical +North America. + +Sub-family V. DENDROCOLAPTINÆ (14 genera, 59 species). Ranges from Chili +and La Plata to Mexico; only 3 species occur in the South Temperate +sub-region, while 9 of the genera extend into Tropical North America. Two +of the continental species occur in the island of Tobago, which, together +with Trinidad, forms part of the South American rather than of the true +Antillean sub-region. + +FAMILY 45.--FORMICARIIDÆ. (32 Genera, 211 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Formicariidæ, comprising the Bush-Shrikes and Ant-thrushes, form one of +the most exclusively Neotropical families; and the numerous species are +rigidly confined to the warm and wooded districts, only a single species +extending to La Plata, and none to the Antilles or to the Nearctic region. +Less than 30 species are found north of Panama. Messrs. Sclater and Salvin +divide the group into three sub-families, whose distribution may be +conveniently treated, as in the Dendrocolaptidæ, without enumerating the +genera. + +Sub-family I. THAMNOPHILINÆ.--(10 genera, 70 species.) One species of +_Thamnophilus_ inhabits La Plata; only 3 genera and 12 species are found +north of Panama, the species of this sub-family being especially abundant +in the Equatorial forest districts. + +Sub-family II. FORMICIVORINÆ.--(14 genera, 95 species.) Only 8 species +occur north of Panama, and less than one-third of the species belong to the +districts south of the Equator. + +{297}Sub-family III. FORMICARIINÆ.--(8 genera, 46 species.) About 12 +species occur north of Panama, and only 5 south of the Equatorial district. + +It appears, therefore, that this extensive family is especially +characteristic of that part of South America from the Amazon valley +northwards. + + +FAMILY 46.--PTEROPTOCHIDÆ. (8 Genera, 19 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Pteroptochidæ are a group of curious Wren-like birds, almost confined +to the temperate regions of South America, extending along the Andes beyond +the Equator, and with a few species in South-east Brazil, and one in the +valley of the Madeira. The genera are as follows:-- + +_Scytalopus_ (8 sp.), Chili and West Patagonia to the Andes of Columbia; +_Merulaxis_ (1 sp.), South-east Brazil; _Rhinocrypta_ (2 sp.), Northern +Patagonia and La Plata; _Lioscelis_ (1 sp.), Madeira valley; _Pteroptochus_ +(2 sp.), Chili; _Hylactes_ (3 sp.), Western Patagonia and Chili; +_Acropternis_ (1 sp.), Andes of Ecuador and Columbia; _Triptorhinus_ (1 +sp.), Chili. + + +FAMILY 47.--PITTIDÆ. (4 Genera, 40 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- 4 |-- 2 -- -- |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Pittas comprise a number of beautifully-coloured Thrush-like birds, +which, although confined to the Old World, are more nearly allied to the +South American Pteroptochidæ than to any other family. They are most +abundant in the Malay Archipelago, {298}between the Oriental and Australian +divisions of which they are pretty equally divided. They seem, however, to +attain their maximum of beauty and variety in the large islands of Borneo +and Sumatra; from whence they diminish in numbers in every direction till +we find single species only in North China, West Africa, and Australia, The +genera here adopted are the following:-- + +(1087 1088 1090 1092 1093) _Pitta_ (33 sp.), has the range of the family; +(1089) _Hydrornis_ (3 sp.), Himalayas and Malaya; _Eucichla_ (3 sp.), +Malaya; _Melampitta_ (1 sp.), recently discovered in New Guinea. + + +FAMILY 48.--PAICTIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 |-- -- -- --|-- -- -- -- + | | | | | + +This family was established by Professor Sundevall, for an anomalous bird +of Madagascar, which he believes to have some affinity for the American +Formicariidæ, but which perhaps comes best near the Pittas. The only genus +is _Philepitta_, containing two species. + + +FAMILY 49.--MENURIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- + | | | | | + +The Menuridæ, or Lyre Birds, remarkable for the extreme elegance of the +lyre-shaped tail in the species first discovered, are birds of a very +anomalous structure, and have no near affinity to any other family. Two +species of _Menura_ are known, confined to South and East Australia (Plate +XII. Vol. I. p. 441). + + +{299}FAMILY 50.--ATRICHIIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- + | | | | | + +The genus _Atrichia_, or Scrub-birds of Australia, have been formed into a +separate family by Professor Newton, on account of peculiarities in the +skeleton which separate them from all other Passeres. Only two species are +known, inhabiting East and West Australia respectively. They are very +noisy, brown-coloured birds, and have been usually classed with the +warblers, near _Amytis_ and other Australian species. + + +_General remarks on the distribution of the Passeres._ + +The order Passeres, is the most extensive among birds, comprehending about +5,700 species grouped in 870 genera, and 51 families. The distribution of +the genera, and of the families considered individually, has been already +sufficiently given, and we now have to consider the peculiarities of +distribution of the families collectively, and in their relations to each +other, as representing well-marked types of bird-structure. The first thing +to be noted is, how very few of these families are truly cosmopolitan; for +although there are seven which are found in each of the great regions, yet +few of these are widely distributed throughout all the regions, and we can +only find three that inhabit every sub-region, and are distributed with +tolerable uniformity; these are the Hirundinidæ, or swallows, the +Motacillidæ or wagtails and pipits, and the Corvidæ or crows,--but the +latter is a family of so heterogeneous a nature, that it possibly contains +the materials of several natural families, and if so divided, the parts +would probably all cease to be cosmopolitan. The Sylviidæ, the +{300}Turdidæ, and the Paridæ, are the only other families that approach +universality of distribution, and all these are wanting in one or more +sub-regions. If, now, we divide the globe into the New and the Old World, +the former including the whole American continent, the latter all the rest +of the earth, we find that the Old World possesses exclusively 23 families, +the New World exclusively 14, of which 5 are common to North and South +America. But if we take the division proposed by Professor Huxley--a +northern world, comprising our first four regions (from Nearctic to +Oriental), and a southern world comprising our last two regions (the +Australian and Neotropical)--we find that the northern division possesses +only 5 families exclusively, and the southern division 13 exclusively, of +which not one is common to Australia and South America. This plainly +indicates that, as far as the Passeres are concerned, the latter bipartite +division is not so natural as the former. Again, if we compare temperate +with tropical families (not too rigidly, but as regards their general +character), we find in the northern hemisphere only two families that have +the character of being typically temperate--the Cinclidæ, and in a less +degree the Ampelidæ--both of small extent. In the southern hemisphere we +have also two, the Phytotomidæ, and in a less degree, the Pteroptochidæ; +making two wholly and two mainly temperate families. Of exclusively +tropical families on the other hand, we have about 12, and several others +that are mainly tropical. + +The several regions do not differ greatly in the number of families found +in each. The Nearctic has 19, the Palæarctic 21, the Ethiopian 23, the +Oriental 28, the Australian 29, and the Neotropical 23. But many of these +families are only represented by a few species, or in limited districts; +and if we count only those families which are tolerably well represented, +and help to form the ornithological character of the region, the richness +of the several tropical regions will appear to be (as it really is) +comparatively much greater. The families that are confined to single +regions are not very numerous, except in the case of the Neotropical +region, which has 5. The Australian has only {301}3, the Oriental 1, the +Ethiopian 1, and the other regions have no peculiar families. + +The distribution of the Passeres may be advantageously considered as +divided into the five series of Turdoid, Tanagroid, Sturnoid, Formicarioid, +and Anomalous Passeres. The Turdoid Passeres, consisting of the first 23 +families, are especially characteristic of the Old World, none being found +exclusively in America, and only two or three being at all abundant there. +The Tanagroid Passeres (Families 24-33) are very characteristic of the New +World, five being confined to it, and three others being quite as abundant +there as in the Old World; while there is not a single exclusively Old +World family in the series, except the Drepanididæ confined to the Sandwich +Islands. The Sturnoid Passeres (Families 34-38) are all exclusively Old +World, except that two larks inhabit parts of North America, and a few +pipits South America. The Formicarioid Passeres (Families 39-48) are +strikingly characteristic of the New World, to which seven of the families +exclusively belong; the two Old World groups being small, and with a very +restricted distribution. The Anomalous Passeres (Families 49-50) are +confined to Australia. + +The most remarkable feature in the geographical distribution of the +Passeres is the richness of the American continent, and the large +development of characteristic types that occurs there. The fact that +America possesses 14 altogether peculiar families, while no less than 23 +Old-World families are entirely absent from it, plainly indicates, that, if +this division does not represent the most ancient and radical separation of +the land surface of the globe, it must still be one of very great +antiquity, and have modified in a very marked way the distribution of all +living things. Not less remarkable is the richness in specific forms of the +13 peculiar American families. These contain no less than 1,570 species, +leaving only about 500 American species in the 13 other Passerine families +represented in the New World. If we make a deduction for those Nearctic +species which occur only north of Panama, we may estimate the truly +Neotropical species of Passerine birds at 1,900, which is almost exactly +{302}_one-third_ of the total number of Passeres; a wonderful illustration +of the Ornithological riches of South America. + + +_Order II.--PICARIÆ._ + +FAMILY 51.--PICIDÆ. (36 Genera, 320 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3. 4 |1 -- -- -- + | | | | | + +The Woodpeckers are very widely distributed, being only absent from the +Australian region beyond Celebes and Flores. They are most abundant in the +Neotropical and Oriental regions, both of which possess a number of +peculiar genera; while the other regions possess few or no peculiar forms, +even the Ethiopian region having only three genera not found elsewhere. The +soft-tailed Picumninæ inhabit the tropical regions only, _Picumnus_ being +Neotropical, _Vivia_ and _Sasia_ Oriental, and _Verreauxia_ Ethiopian. +_Picoides_, or _Apternus_, is an Arctic form peculiar to the Nearctic and +Palæarctic regions. _Celeus_, _Chrysoptilus_, _Chloronerpes_, and some +smaller genera, are Neotropical exclusively, and there are two peculiar +forms in Cuba. _Yungipicus_, _Chrysocolaptes_, _Hemicercus_, +_Mulleripicus_, _Brachypternus_, _Tiga_, and _Micropternus_, are the most +important of the peculiar Oriental genera. _Dendropicus_ and _Geocolaptes_ +are Ethiopian; but there are no woodpeckers in Madagascar. The Palæarctic +woodpeckers belong to the genera _Picus_--which is widely distributed, +_Gecinus_--which is an Oriental form, and _Dryocopus_--which is South +American. Except _Picoides_, the Nearctic woodpeckers are mostly of +Neotropical genera; but _Sphyrapicus_ and _Hylatomus_ are peculiar. The +geological record is, as yet, almost silent as to this family; but remains +doubtfully referred to it have been found in the Miocene of Europe and the +Eocene of the United States. Yet the group is evidently one of very high +antiquity, as is shown by {303}its extreme isolation, its great +specialization of structure, its abundant generic forms, and its wide +distribution. It originated, probably, in Central Asia, and passed through, +the Nearctic region to South America, in whose rich and varied forests it +found the conditions for rapid development, and for the specialization of +the many generic forms now found there. + +A large number of genera have been established by various authors, but +their limitations and affinities are not very well made out. Those which +seem best established are the following:-- + +(2107-2112) _Picumnus_ (22 sp.). Tropical South America to Honduras; (2113) +_Vivia_ (1 sp.), Himalayas to East Thibet; (2114) _Sasia_ (2 sp.), Nepal to +Java; (2115) _Verreauxia_ (1 sp.), West Africa; _Picoides_ (5 sp.), +northern parts of Nearctic and Palæarctic regions, and Mountains of East +Thibet; _Picus_ (42 sp.), the whole Palæarctic, Oriental, Nearctic, and +Neotropical regions; (2123) _Hyopicus_ (2 sp.), Himalayas and North China; +(2124) _Yungipicus_ (16 sp.), Oriental region, and to Flores, Celebes, +North China, and Japan; (2127-2129) _Sphyrapicus_ (7 sp.), Nearctic region, +Mexico, and Bolivia; (2130-2133 2139) _Campephilus_ (14 sp.), Neotropical +and Nearctic regions; _Hylatomus_ (1 sp.), Nearctic region; (2137 2140) +_Dryocopus_ (5 sp.), Mexico to South Brazil, Central and Northern Europe; +(2134) _Reinwardtipicus_ (1 sp.), Penang to Borneo; (2135 2136) _Venilia_ +(2 sp.), Nepal to Borneo; _Chrysocolaptes_ (8 sp.), India and Indo-Malaya; +_Dendropicus_ (16 sp.), Tropical and South Africa; _Hemicercus_ (5 sp.), +Malabar and Pegu to Malaya; _Gecinus_ (18 sp.), Palæarctic and Oriental +regions to Java; (2151-2156) _Dendromus_ (15 sp.), West and South Africa, +Zanzibar, and Abyssinia; (2157-2159) _Mulleripicus_ (6 sp.), Malabar, Pegu, +Indo-Malaya, and Celebes; _Celeus_ (17 sp.), Paraguay to Mexico; +_Nesoceleus_ (sp. 8833) Cuba; (2162) _Chrysoptilus_ (9 sp.), Chili and +South Brazil to Mexico; _Brachypternus_ (5 sp.), India, Ceylon, and China; +(2165 2166) _Tiga_ (5 sp.), all India to Malaya; (2167) _Gecinulus_ (2 +sp.), South-east Himalayas to Burmah; _Centurus_ (13 sp.), Nearctic Region +to Antilles and Venezuela; _Chloronerpes_ (35 sp.), Tropical America, +Hayti; (2171) _Xiphidiopicus_ (1 sp.), Cuba; _Melanerpes_ (11 sp.), Brazil +to {304}Canada, Porto Rico; _Leuconerpes_ (1 sp.), Bolivia to North Brazil; +_Colaptes_ (9 sp.), La Plata and Bolivia to Arctic America, Greater +Antilles; _Hypoxanthus_ (1 sp.), Venezuela and Ecuador; (2187) +_Geocolaptes_ (1 sp.), South Africa; _Miglyptes_ (3 sp.), Malaya; +_Micropternus_ (8 sp.), India and Ceylon to South China, Sumatra and +Borneo. + + +FAMILY 52.--YUNGIDÆ. (1 Genus, 5 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3. 4 | 1 -- 3 -- |1 -- -- -- |-- -- -- -- + | | | | | + +The Wrynecks (_Yunx_), which constitute this family, are small +tree-creeping birds characteristic of the Palæarctic region, but extending +into North and East Africa, over the greater part of the peninsula of India +(but not to Ceylon), and just reaching the lower ranges of the Himalayas. +There is also one species isolated in South Africa. + + +FAMILY 53.--INDICATORIDÆ. (1 Genus, 12 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3. --|-- -- 3. 4 |-- -- -- -- + | | | | | + +The Honey-guides (_Indicator_) constitute a small family of doubtful +affinities; perhaps most nearly allied to the woodpeckers and barbets. They +catch bees and sometimes kill small birds; and some of the species are +parasitical like the cuckoo. Their distribution is very interesting, as +they are found in every part of the Ethiopian region, except Madagascar, +and in the Oriental region only in Sikhim and Borneo, being absent from the +peninsula of India which is nearest, both geographically and zoologically, +to Africa. + + +{305}FAMILY 54.--MEGALÆMIDÆ. (13 Genera, 81 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|1. 2. 3 -- |1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Megalæmidæ, or Barbets, consist of rather small, fruit-eating birds, of +heavy ungraceful shape, but adorned with the most gaudy colours, especially +about the head and neck. They form a very isolated family; their nearest +allies being, perhaps, the still more isolated Toucans of South America. +Barbets are found in all the tropics except Australia, but are especially +characteristic of the great Equatorial forest-zone; all the most remarkable +forms being confined to Equatorial America, West Africa, and the Indo-Malay +Islands. They are most abundant in the Ethiopian and Oriental regions, and +in the latter are universally distributed. + +In the beautiful monograph of this family by the Messrs. Marshall, the +barbets are divided into three sub-families, as follows:-- + +Pogonorhynchinæ (3 genera, 15 sp.), which are Ethiopian except the 2 +species of _Tetragonops_, which are Neotropical; Megalæminæ (6 genera, 45 +sp.), which are Oriental and Ethiopian; and Capitoninæ (4 genera, 18 sp.), +common to the three regions. + +The genera are each confined to a single region. Africa possesses the +largest number of peculiar forms, while the Oriental region is richest in +species. + +This is probably a very ancient group, and its existing distribution may be +due to its former range over the Miocene South Palæarctic land, which we +know possessed Trogons, Parrots, Apes, and Tapirs, groups which are now +equally abundant in Equatorial countries. + +{306}The following is a tabular view of the genera with their +distribution:-- + + --------------------+------------------+----------------+---------------- + Genera | Ethiopian | Oriental | Neotropical + | Region. | Region. | Region. + --------------------+------------------+----------------+---------------- + | | | + POGONORHYNCHINÆ. | | | + Tricholæma 1 sp.|W. Africa | | + Pogonorhynchus 12 " |All Trop. & S. Af.| | + Tetragonops 2 " | | |Peru & Costa + | | | Rica + | | | + MEGALÆMINÆ. | | | + Megalæma 29 " | |The whole region| + Xantholæma 4 " | |The whole region| + Xylobucco 2 " |W. Africa | | + Barbatula 9 " |Trop. & S. Africa | | + Psilopogon 1 " | | |Sumatra + Gymnobucco 2 " |W. Africa | | + | | | + CAPITONINÆ. | | | + Trachyphonus 5 " |Trop. & S. Africa | | + Capito 10 " | | |Equatorial Amer. + | | | to Costa Rica + Calorhamphus 2 " | |Malay Pen., | + | | Sumatra, Borneo| + Stactolæma 1 " |W. Africa | | + | | | + --------------------+------------------+----------------+---------------- + + +FAMILY 55.--RHAMPHASTIDÆ. (5 Genera, 51 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Toucans form one of the most remarkable and characteristic families of +the Neotropical region, to which they are strictly confined. They differ +from all other birds by their long feathered tongues, their huge yet +elegant bills, and the peculiar texture and coloration of their plumage. +Being fruit-eaters, and strictly adapted for an arboreal life, they are not +found beyond the forest regions; but they nevertheless range from Mexico to +Paraguay, and from the Atlantic to the Pacific. One genus, {307}_Andigena_, +is confined to the forest slopes of the South American Andes. The genera +are:-- + +_Rhamphastos_ (12 sp.), Mexico to South Brazil; _Pteroglossus_ (16 sp.), +Nicaragua to South Brazil (Plate XV. Vol. II. p. 28); _Selenidera_ (7 sp.), +Veragua to Brazil, east of the Andes; _Andigena_ (6 sp.), the Andes, from +Columbia to Bolivia, and West Brazil; _Aulacorhamphus_ (10 sp.), Mexico to +Peru and Bolivia. + + +FAMILY 56.--MUSOPHAGIDÆ. (2 Genera, 18 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The Musophagidæ, or Plantain-eaters and Turacos, are handsome birds, +somewhat intermediate between Toucans and Cuckoos. They are confined to the +Ethiopian region and are most abundant in West Africa. The Plantain eaters +(_Musophaga_, 2 sp.), are confined to West Africa; the Turacos (_Turacus_, +16 sp., including the sub-genera _Corythaix_ and _Schizorhis_) range over +all Africa from Abyssinia to the Cape (Plate V. Vol. I. p. 264). + + +FAMILY 57.--COLIIDÆ. (1 Genus, 7 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The Colies, consisting of the single genus _Colius_, are an anomalous group +of small finch-like birds, occuping a position between the Picariæ and +Passeres, but of very doubtful affinities. Their range is nearly identical +with that of the Musophagidæ, but they are most abundant in South and East +Africa. + + +{308}FAMILY 58.--CUCULIDÆ. (35 Genera, 180 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Cuculidæ, of which our well-known Cuckoo is one of the most widely +distributed types, are essentially a tropical group of weak insectivorous +birds, abounding in varied forms in all the warmer parts of the globe, but +very scarce or only appearing as migrants in the temperate and colder +zones. Many of the smaller Eastern species are adorned with the most +intense golden or violet metallic lustre, while some of the larger forms +have gaily-coloured bills or bare patches of bright red on the cheeks. Many +of the cuckoos of the Eastern Hemisphere are parasitic, laying their eggs +in other birds' nests; and they are also remarkable for the manner in which +they resemble other birds, as hawks, pheasants, or drongo-shrikes. The +distribution of the Cuckoo family is rather remarkable. They abound most in +the Oriental region, which produces no less than 18 genera, of which 11 are +peculiar; the Australian has 8, most of which are also Oriental, but 3 are +peculiar, one of these being confined to Celebes and closely allied to an +Oriental group; the Ethiopian region has only 7 genera, all of which are +Oriental but three, 2 of these being peculiar to Madagascar, and the other +common to Madagascar and Africa. America has 11 genera, all quite distinct +from those of the Eastern Hemisphere, and only three enter the Nearctic +region, one species extending to Canada. + +Remembering our conclusions as to the early history of the several regions, +these facts enable us to indicate, with considerable probability, the +origin and mode of dispersal of the cuckoos. They were almost certainly +developed in the Oriental and Palæarctic regions, but reached the +Neotropical at a very early date, where they have since been completely +isolated. Africa must have long remained without cuckoos, the earliest +immigration {309}being to Madagascar at the time of the approximation of +that sub-region to Ceylon and Malaya. A later infusion of Oriental forms +took place probably by way of Arabia and Persia, when those countries were +more fertile and perhaps more extensive. Australia has also received its +cuckoos at a somewhat late date, a few having reached the Austro-Malay +Islands somewhat earlier. + +The classification of the family is somewhat unsettled. For the American +genera I follow Messrs. Sclater and Salvin; and, for those of the Old +World, Mr. Sharpe's suggestive paper in the _Proceedings of the Zoological +Society_, 1873, p. 600. The following is the distribution of the various +genera:-- + +(2195) _Phænicophaës_ (1 sp.), Ceylon; (2196) _Rhamphococcyx_ (1 sp.), +Celebes; (2196) _Rhinococcyx_ (1 sp.), Java; (2196 pt. and 2203) +_Rhopodytes_ (6 sp.), Himalayas to Ceylon, Hainan, and Malaya; (2203 pt) +_Poliococcyx_ (1 sp.), Malacca, Sumatra, and Borneo; (2197) _Dasylophus_ (1 +sp.), Philippine Islands; (2198) _Lepidogrammus_ (1 sp.), Philippine +Islands; (2200) _Zanclostomus_ (1 sp.), Malaya; (2201) _Ceuthmochares_ (2 +sp.), Tropical and South Africa and Madagascar; (2202) _Taccocua_ (4 sp.), +Himalayas to Ceylon and Malacca; (2204) _Rhinortha_ (1 sp.), Malacca, +Sumatra, Borneo; (2199) _Carpococcyx_ (1 sp.), Borneo and Sumatra; (2220) +_Neomorphus_ (4 sp.), Brazil to Mexico; (2205 2206) _Coua_ (10 sp.), +Madagascar; (2207) _Cochlothraustes_ (1 sp.), Madagascar; (2221) +_Centropus_ (35 sp.), Tropical and South Africa, the whole Oriental region, +Austro-Malaya and Australia; (2213) _Crotophaga_ (3 sp.), Brazil to +Antilles and Pennsylvania; (2212) _Guira_ (1 sp.), Brazil and Paraguay; +(2209) _Geococcyx_ (2 sp.), Guatemala to Texas and California; (2211) +_Dromococcyx_ (2 sp.), Brazil to Mexico; (2210) _Diplopterus_ (1 sp.), +Mexico to Ecuador and Brazil; (2208) _Saurothera_ (4 sp.), Greater +Antilles; (2219) _Hyetornis_ (2 sp.), Jamaica and Hayti; (2215) _Piaya_ (3 +sp.), Mexico to West Ecuador and Brazil; (2218) _Morococcyx_ (1 sp.), Costa +Rica to Mexico; (2214) _Coccygus_ (10 sp.), La Plata to Antilles, Mexico +and Pennsylvania, Cocos Island; (2227) _Cuculus_ (22 sp.), Palæarctic, +Ethiopian, and Oriental regions, to Moluccas and Australia; (2229) +_Caliecthrus_ (1 sp.), Papuan Islands; (2230-2232) _Cacomantis_ (15 sp.), +Oriental and Australian {310}regions to Fiji Islands and Tasmania; +(2233-2237) _Chrysococcyx_ (16 sp.), Tropical and South Africa, the +Oriental and Australian regions to New Zealand and Fiji Islands; (2238) +_Surniculus_ (2 sp.), India, Ceylon, and Malaya; (2239) _Hierococcyx_ (7 +sp.), the Oriental region to Amoorland and Celebes; (2240 2241) _Coccystes_ +(6 sp.), Tropical and South Africa, the Oriental region, excluding +Philippines; (2242) _Eudynamis_ (8 sp.), the Oriental and Australian +regions, excluding Sandwich Islands; (2243) _Scythrops_ (1 sp.), East +Australia to Moluccas and North Celebes. + + +FAMILY 59.--LEPTOSOMIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 |-- -- -- --|-- -- -- -- + | | | | | + +The _Leptosomus discolor_, which constitutes this family, is a bird of very +abnormal characters, having some affinities both with Cuckoos and Rollers. +It is confined to Madagascar (Plate VI. Vol. I. p. 278). + + +FAMILY 60.--BUCCONIDÆ. (5 Genera, 43 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Bucconidæ, or Puff-birds, are generally of small size and dull colours, +with rather thick bodies and dense plumage. They form one of the +characteristic Neotropical families, being most abundant in the great +Equatorial forest plains, but extending as far north as Guatemala, though +absent from the West Indian Islands. + +The genera are:--_Bucco_ (21 sp.), Guatemala to Paraguay, and West of the +Andes in Ecuador; _Malacoptila_ (10 sp.), Guatemala {311}to Bolivia and +Brazil; _Nonnula_ (3 sp.), Amazon and Columbia; _Monasa_ (7 sp.), Costa +Rica to Brazil; _Chelidoptera_ (2 sp.), Columbia and Guiana to Brazil. + + +FAMILY 61.--GALBULIDÆ. (6 Genera, 19 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Galbulidæ, or Jacamars, are small slender birds, of generally metallic +plumage; somewhat resembling in form the Bee-eaters of the Old World but +less active. They have the same general distribution as the last family, +but they do not occur west of the Equatorial Andes. The genera are:-- + +_Galbula_ (9 sp.), Guatemala to Brazil and Bolivia; _Urogalba_ (2 sp.), +Guiana and the lower Amazon; _Brachygalba_ (4 sp.), Venezuela to Brazil and +Bolivia; _Jacamaralcyon_ (1 sp.), Brazil; _Jacamerops_ (2 sp.), Panama to +the Amazon; _Galbalcyrhynchus_ (1 sp.), Upper Amazon. + + +FAMILY 62.--CORACIIDÆ. (3 Genera, 19 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Rollers are a family of insectivorous birds allied to the Bee-eaters, +and are very characteristic of the Ethiopian and Oriental regions; but one +species (_Coracias garrula_) spreads over the Palæarctic region as far +north as Sweden and the Altai mountains, while the genus _Eurystomus_ +reaches the Amoor valley, Australia, and the Solomon Islands. The +distribution of the genera is as follows:-- + +_Coracias_ (8 sp.), the whole Ethiopian region, the Oriental {312}region +except Indo-Malaya, the Palæarctic to the above-named limits, and the +island of Celebes on the confines of the Australian region; _Eurystomus_ (8 +sp.), West and East Africa and Madagascar, the whole Oriental region except +the Peninsula of India, and the Australian as far as Australia and the +Solomon Islands; _Brachypteracias_ (possibly allied to _Leptosomus_?) (4 +sp.), Madagascar only, but these abnormal birds form a distinct sub-family, +and according to Mr. Sharpe, three genera, _Brachypteracias_, _Atelornis_, +and _Geobiastes_. + +A most remarkable feature in the distribution of this family is the +occurrence of a true roller (_Coracias temminckii_) in the island of +Celebes, entirely cut off from the rest of the genus, which does not occur +again till we reach Siam and Burmah. + +The curious _Pseudochelidon_ from West Africa may perhaps belong to this +family or to the Cypselidæ. (Ibis. 1861, p. 321.) + + +FAMILY 63.--MEROPIDÆ. (5 Genera, 34 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2 -- -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Meropidæ, or Bee-eaters, have nearly the same distribution as the +Rollers, but they do not penetrate quite so far either into the Eastern +Palæarctic or the Australian regions. The distribution of the genera is as +follows:-- + +_Merops_ (21 sp.), has the range of the family extending on the north to +South Scandinavia, and east to Australia and New Guinea; _Nyctiornis_ (3 +sp.), the Oriental region, except Ceylon and Java; _Meropogon_ (1 sp.), +Celebes; _Meropiscus_ (3 sp.), West Africa; _Melittophagus_ (6 sp.), +Ethiopian region, except Madagascar. + + +{313}FAMILY 64.--TODIDÆ. (1 Genus, 5 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Todies are delicate, bright-coloured, insectivorous birds, of small +size, and allied to the Motmots, although externally more resembling +flycatchers. They are wholly confined to the greater Antilles, the islands +of Cuba, Hayti, Jamaica, and Porto Rico having each a peculiar species of +_Todus_, while another species, said to be from Jamaica, has been recently +described (Plate XVI. Vol. II. p. 67). + + +FAMILY 65.--MOMOTIDÆ. (6 Genera, 17 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Motmots range from Mexico to Paraguay and to the west coast of Ecuador, +but seem to have their head-quarters in Central America, five of the genera +and eleven species occurring from Panama northwards, two of the genera not +occurring in South America. The genera are as follows:-- + +_Momotus_ (10 sp.), Mexico to Brazil and Bolivia, one species extending to +Tobago, and one to Western Ecuador; _Urospatha_ (1 sp.), Costa Rica to the +Amazon; _Baryphthengus_ (1 sp.), Brazil and Paraguay; _Hylomanes_ (2 sp.), +Guatemala; _Prionirhynchus_ (2 sp.), Guatemala to Upper Amazon; _Eumomota_ +(1 sp.), Honduras to Chiriqui. + + +{314}FAMILY 66.--TROGONIDÆ. (7 Genera, 44 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |-- -- -- --|-- -- -- --|1. 2. 3 -- |1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Trogons form a well-marked family of insectivorous forest-haunting +birds, whose dense yet puffy plumage exhibits the most exquisite tints of +pink, crimson, orange, brown, or metallic green, often relieved by delicate +bands of pure white. In one Guatemalan species the tail coverts are +enormously lengthened into waving plumes of rich metallic green, as +graceful and marvellous as those of the Paradise-birds. Trogons are +tolerably abundant in the Neotropical and Oriental regions, and are +represented in Africa by a single species of a peculiar genus. The genera +now generally admitted are the following:-- + +_Trogon_ (24 sp.), Paraguay to Mexico, and west of the Andes in Ecuador; +_Temnotrogon_ (1 sp.), Hayti; _Prionoteles_ (1 sp.), Cuba (Plate XVII. Vol. +II. p. 67); _Apaloderma_ (2 sp.), Tropical and South Africa; _Harpactes_ +(10 sp.), the Oriental region, excluding China; _Pharomacrus_ (5 sp.), +Amazonia to Guatemala; _Euptilotis_ (1 sp.), Mexico. + +Remains of _Trogon_ have been found in the Miocene deposits of France; and +we are thus able to understand the existing distribution of the family. At +that exceptionally mild period in the northern hemisphere, these birds may +have ranged over all Europe and North America; but, as the climate became +more severe they gradually became restricted to the tropical regions, where +alone a sufficiency of fruit and insect-food is found all the year round. + + +{315}FAMILY 67.--ALCEDINIDÆ. (19 Genera, 125 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Kingfishers are distributed universally, but very unequally, over the +globe, and in this respect present some of the most curious anomalies to be +found among birds. They have their metropolis in the eastern half of the +Malay Archipelago (our first Australian sub-region), from Celebes to New +Guinea, in which district no less than 13 out of the 19 genera occur, 8 of +them being peculiar; and it is probable that in no other equally varied +group of universal distribution, is so large a proportion of the generic +forms confined to so limited a district. From this centre kingfishers +decrease rapidly in every direction. In Australia itself there are only 4 +genera with 13 species; the whole Oriental region has only 6 genera, 1 +being peculiar; the Ethiopian also 6 genera, but 3 peculiar; and each of +these have less than half the number of species possessed by the Australian +region. The Palæarctic region possesses only 3 genera, all derived from the +Oriental region; but the most extraordinary deficiency is shown by the +usually rich Neotropical region, which possesses but a single genus, common +to the larger part of the Eastern Hemisphere, and the same genus is alone +found in the Nearctic region, the only difference being that the former +possesses eight, while the latter has but a single species. These facts +almost inevitably lead to the conclusion that America long existed without +kingfishers; and that in comparatively recent times (perhaps during the +Miocene or Pliocene period), a species of the Old World genus, _Ceryle_, +found its way into North America, and spreading rapidly southward along the +great river-valleys has become differentiated in South America into the few +closely allied forms that alone inhabit that vast country--the richest in +the world in {316}fresh-water fish, and apparently the best fitted to +sustain a varied and numerous body of kingfishers. + +The names of the genera, with their distribution and the number of species +in each, as given by Mr. Sharpe in his excellent monograph of the family, +is as follows:-- + +_Alcedo_ (9 sp.), Palæarctic, Ethiopian, and Oriental regions (but absent +from Madagascar), and extending into the Austro-Malayan sub-region; +_Corythornis_ (3 sp.), the whole Ethiopian region; _Alcyone_ (7 sp.), +Australia and the Austro-Malayan sub-region, with one species in the +Philippine Islands; _Ceryle_ (13 sp.), absent only from Australia, the +northern half of the Palæarctic region, and Madagascar; _Pelargopsis_ (9 +sp.), the whole Oriental region; and extending to Celebes and Timor in the +Austro-Malayan sub-region; _Ceyx_ (11 sp.), the Oriental region and +Austro-Malayan sub-region, but absent from Celebes, and only one species in +continental India and Ceylon; _Ceycopsis_ (1 sp.), Celebes; _Myioceyx_ (2 +sp.), West Africa; _Ipsidina_ (4 sp.), Ethiopian region; _Syma_ (2 sp.), +Papua and North Australia; _Halcyon_ (36 sp.), Australian, Oriental, and +Ethiopian regions, and the southern part of the Palæarctic; _Dacelo_ (6 +sp.), Australia and New Guinea; _Todirhamphus_ (3 sp.), Eastern Pacific +Islands only; _Monachalcyon_ (1 sp.), Celebes; _Caridonax_ (1 sp.), Lombok +and Flores; _Carcineutes_ (2 sp.), Siam to Borneo and Java; _Tanysiptera_ +(14 sp.), Moluccas New Guinea, and North Australia (Plate X. Vol. I. p. +414); _Cittura_ (2 sp.), Celebes group; _Melidora_ (1 sp.), New Guinea. + + +FAMILY 68.--BUCEROTIDIÆ. (12 Genera, 50 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- |1. 2. 3. 4 |1 -- -- -- + | | | | | + +The Hornbills form an isolated group of generally large-sized birds, whose +huge bills form their most prominent feature. They are popularly associated +with the American Toucans, but have no close relationship to them, and are +now generally {317}considered to show most resemblance, though still a very +distant one, to the kingfishers. They are abundant in the Ethiopian and +Oriental regions, and extend eastward to the Solomon Islands. Their +classification is very unsettled, for though they have been divided into +more than twenty genera they have not yet been carefully studied. The +following grouping of the genera--referring to the numbers in the _Hand +List_--must therefore be considered as only provisional:-- + +(1957 1958 1963) _Buceros_ (6 sp.), all Indo-Malaya, Arakan, Nepal and the +Neilgherries (Plate IX. Vol. I. p. 339); (1959-1961) _Hydrocissa_ (7 sp.), +India and Ceylon to Malaya and Celebes; (1962) _Berenicornis_ (2 sp.), +Sumatra and West Africa; (1964) _Calao_ (3 sp.), Tennaserim, Malaya, +Moluccas to the Solomon Islands; (1965) _Aceros_ (1 sp.), South-east +Himalayas; (1966 1967) _Cranorrhinus_ (3 sp.), Malacca, Sumatra, Borneo, +Philippines, Celebes; (1968) _Penelopides_ (1 sp.), Celebes; (1969-1971) +_Tockus_ (15 sp.), Tropical and South Africa; (1972) _Rhinoplax_ (1 sp.), +Sumatra and Borneo; (1973-1975) _Bycanistes_ (6 sp.), West Africa with East +and South Africa; (1976 1977) _Meniceros_ (3 sp.), India and Ceylon to +Tenasserim; (1978) _Bucorvus_ (2 sp.), Tropical and South Africa. + + +FAMILY 69.--UPUPIDÆ. (1 Genus, 6 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --| -- 2 -- 4 |1. 2. 3. 4 |1. 2. 3 -- |-- -- -- -- + | | | | | + +The Hoopoes form a small and isolated group of semi-terrestrial +insectivorous birds, whose nearest affinities are with the Hornbills. They +are most characteristic of the Ethiopian region, but extend into the South +of Europe and into all the continental divisions of the Oriental region, as +well as to Ceylon, and northwards to Pekin and Mongolia. + + +{318}FAMILY 70.--IRRISORIDÆ. (1 Genus, 12 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The Irrisors are birds of generally metallic plumage, which have often been +placed with the Epunachidæ and near the Sun-birds, or Birds of Paradise, +but which are undoubtedly allied to the Hoopoes. They are strictly confined +to the continent of Africa, ranging from Abyssinia to the west coast, and +southward to the Cape Colony. They have been divided into several +sub-genera which it is not necessary here to notice (Plate IV. Vol. I. p. +261). + + +FAMILY 71.--PODARGIDÆ. (3 Genera, 20 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Podargidæ, or Frog-mouths, are a family of rather large-sized nocturnal +insectivorous birds, closely allied to the Goat-suckers, but distinguished +by their generally thicker bills, and especially by hunting for their food +on trees or on the ground, instead of seizing it on the wing. They abound +most in the Australian region, but one genus extends over a large part of +the Oriental region. The following are the genera with their +distribution:-- + +_Podargus_ (10 sp.), Australia, Tasmania, and the Papuan Islands (Plate +XII. Vol. I. p. 441); _Batrachostomus_ (6 sp.), the Oriental region +(excluding Philippine Islands and China) and the northern Moluccas; +_Ægotheles_ (4 sp.), Australia, Tasmania, and Papuan Islands. + + +{319}FAMILY 72.--STEATORNITHIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +This family contains a single bird--the Guacharo--forming the genus +_Steatornis_, first discovered by Humboldt in a cavern in Venezuela, and +since found in deep ravines near Bogota, and also in Trinidad. Although +apparently allied to the Goat-suckers it is a vegetable-feeder, and is +altogether a very anomalous bird whose position in the system is still +undetermined. + + +FAMILY 73.--CAPRIMULGIDÆ. (17 Genera, 91 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Goat-suckers, or Night-jars, are crepuscular insectivorous birds, which +take their prey on the wing, and are remarkable for their soft and +beautifully mottled plumage, swift and silent flight, and strange cries +often imitating the human voice. They are universally distributed, except +that they do not reach New Zealand or the remoter Pacific Islands. The +South American genus, _Nyctibius_, differs in structure and habits from the +other goat-suckers and should perhaps form a distinct family. More than +half the genera inhabit the Neotropical region. The genera are as +follows:-- + +_Nyctibius_ (6 sp.), Brazil to Guatemala, Jamaica; _Caprimulgus_ (35 sp.), +Palæarctic, Oriental, and Ethiopian regions, with the Austro-Malay Islands +and North Australia; _Hydropsalis_ (8 sp.), Tropical South America to La +Plata; _Antrostomus_ (10 {320}sp.), La Plata and Bolivia to Canada, Cuba; +_Stenopsis_ (4 sp.), Martinique to Columbia, West Peru and Chili; +_Siphonorhis_ (1 sp.), Jamaica; _Heleothreptus_ (1 sp.), Demerara; +_Nyctidromus_ (2 sp.), South Brazil to Central America; _Scortornis_ (3 +sp.), West and East Africa; _Macrodipteryx_ (2 sp.), West and Central +Africa; _Cosmetornis_ (1 sp.), all Tropical Africa; _Podager_ (1 sp.), +Tropical South America to La Plata; _Lurocalis_ (2 sp.), Brazil and Guiana; +_Chordeiles_ (8 sp.), Brazil and West Peru to Canada, Porto Rico, Jamaica; +_Nyctiprogne_ (1 sp.), Brazil and Amazonia; _Eurostopodus_ (2 sp.), +Australia and Papuan Islands; _Lyncornis_ (4 sp.), Burmah, Philippines, +Borneo, Celebes. + + +FAMILY 74.--CYPSELIDÆ. (7 Genera, 53 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +The Swifts can almost claim to be a cosmopolitan group, but for their +absence from New Zealand. They are most abundant both in genera and species +in the Neotropical and Oriental regions. The following is the distribution +of the genera:-- + +_Cypselus_ (1 sp.), absent only from the whole of North America and the +Pacific; _Panyptila_ (3 sp.), Guatemala and Guiana, and extending into +North-west America; _Collocalia_ (10 sp.), Madagascar, the whole Oriental +region and eastward through New Guinea to the Marquesas Islands; +_Dendrochelidon_ (5 sp.), Oriental region and eastward to New Guinea; +_Chætura_ (15 sp.), Continental America (excluding South Temperate), West +Africa and Madagascar, the Oriental region, North China and the Amoor, +Celebes, Australia; _Hemiprocne_ (3 sp.), Mexico to La Plata, Jamaica and +Hayti; _Cypseloides_ (2 sp.), Brazil and Peru; _Nephæcetes_ (2 sp.), Cuba, +Jamaica, North-west America. + + +{321}FAMILY 75.--TROCHILIDÆ. (118 Genera, 390 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The wonderfully varied and beautiful Humming-Birds are confined to the +American continent, where they range from Sitka to Cape Horn, while the +island of Juan Fernandez has two peculiar species. Only 6 species, +belonging to 3 genera, are found in the Nearctic region, and most of these +have extended their range from the south. They are excessively abundant in +the forest-clad Andes from Mexico to Chili, some species extending up to +the limits of perpetual snow; but they diminish in number and variety in +the plains, however luxuriant the vegetation. In place of giving here the +names and distribution of the numerous genera into which they are now +divided (which will be found in the tables of the genera of the Neotropical +region), it may be more useful to present a summary of their distribution +in the sub-divisions of the American continent, as follows:-- + + Sub-region I. = Patagonia & S. Andes. + Sub-region II. = Tropical S. America. + Sub-region III. = Tropical N. America. + Sub-region IV. = Antilles. + Nearctic region = Temperate N. America. + + -------- Sub-regions -------- Nearctic + I. II. III. IV. Regions. + Genera in each Sub-region 10 90 41 8 3 + Peculiar Genera 3 58 14 5 0 + Species in each Sub-region 15 275 100 15 6 + +The island of Juan Fernandez has two species, and Masafuera, an island +beyond it, one; the three forming a peculiar genus. The island of Tres +Marias, about 60 miles from the west coast of Mexico, possesses a peculiar +species of humming-bird, and the Bahamas two species; but none inhabit +either the Falkland Islands or the Galapagos. + +Like most groups which are very rich in species and in generic forms, the +humming-birds are generally very local, small {322}generic groups being +confined to limited districts; while single mountains, valleys, or small +islands, often possess species found nowhere else. It is now well +ascertained that the Trochilidæ are really insectivorous birds, although +they also feed largely, but probably never exclusively, on the nectar of +flowers. Their nearest allies are undoubtedly the Swifts; but the wide gap +that now separates them from these, as well as the wonderful variety of +form and of development of plumage, that is found among them, alike point +to their origin, at a very remote period, in the forests of the once +insular Andes. There is perhaps no more striking contrast of the like +nature, to be found, than that between the American kingfishers--confined +to a few closely allied forms of one Old World genus--and the American +humming-birds with more than a hundred diversified generic forms unlike +everything else upon the globe; and we can hardly imagine any other cause +for this difference, than a (comparatively) very recent introduction in the +one case, and a very high antiquity in the other. + + +_General Remarks on the Distribution of the Picariæ._ + +The very heterogeneous mass of birds forming the Order Picariæ, contains 25 +families, 307 genera and 1,604 species. This gives about 64 species to each +family, while in the Passeres the proportion is nearly double, or 111 +species per family. There are, in fact, only two very large families in the +Order, which happen to be the first and last in the series--Picidæ and +Trochilidæ. Two others--Cuculidæ and Alcedinidæ--are rather large; while +the rest are all small, seven of them consisting only of a single genus and +from one to a dozen species. Only one of the families--Alcedinidæ--is +absolutely cosmopolitan, but three others are nearly so, Caprimulgidæ and +Cypselidæ being only absent from New Zealand, and Cuculidæ from the +Canadian sub-region of North America. Eleven families inhabit the Old World +only, while seven are confined to the New World, only one of +these--Trochilidæ--being common to the Neotropical and Nearctic regions. + +The Picariæ are highly characteristic of tropical faunas, for {323}while no +less than 15 out of the 25 families are exclusively tropical, none are +confined to, or have their chief development in, the temperate regions. +They are best represented in the Ethiopian region, which possesses 17 +families, 4 of which are peculiar to it; while the Oriental region has only +14 families, none of which are peculiar. The Neotropical region has also 14 +families, but 6 of them are peculiar. The Australian region has 8, the +Palæarctic 9 and the Nearctic 6 families, but none of these are peculiar. +We may see a reason for the great specialization of this tropical +assemblage of birds in the Ethiopian and Neotropical regions, in the fact +of the large extent of land on both sides of the Equator which these two +regions alone possess, and their extreme isolation either by sea or deserts +from other regions,--an isolation which we know was in both cases much +greater in early Tertiary times. It is, perhaps, for a similar reason that +we here find hardly any trace of the connection between Australia and South +America which other groups exhibit; for that connection has most probably +been effected by a former communication between the temperate southern +extremities of those two continents. The most interesting and suggestive +fact, is that presented by the distribution of the Megalæmidæ and Trogonidæ +over the tropics of America, Africa, and Asia. In the absence of +palæontological evidence as to the former history of the Megalæmidæ, we are +unable to say positively, whether it owes its present distribution to a +former closer union between these continents in intertropical latitudes, or +to a much greater northern range of the group at the period when a +luxuriant sub-tropical vegetation extended far toward the Arctic regions; +but the discovery of _Trogon_ in the Miocene deposits of the South of +France renders it almost certain that the latter is the true explanation in +the case of both these families. + +The Neotropical region, owing to its enormous family of humming-birds, is +by far the richest in Picariæ, possessing nearly half the total number of +species, and a still larger proportion of genera. Three families, the +Bucerotidæ, Meropidæ and Coraciidæ are equally characteristic of the +Oriental and {324}Ethiopian regions, a few outlying species only entering +the Australian or the Palæarctic regions. One family (Todidæ) is confined +to the West Indian Islands; and another (Leptosomidæ) consisting of but a +single species, to Madagascar; parallel cases to the Drepanididæ among the +Passeres, peculiar to the Sandwich Islands, and the Apterygidæ among the +Struthiones, peculiar to New Zealand. + + +_Order III.--PSITTACI._ + +The Parrots have been the subject of much difference of opinion among +ornithologists, and no satisfactory arrangement of the order into families +and genera has yet been reached. Professor Garrod has lately examined +certain points in the anatomy of a large number of genera, and proposes to +revolutionize the ordinary classifications. Until, however, a general +examination of their whole anatomy, internal and external, has been made by +some competent authority, it will be unsafe to adopt the new system, as we +have as yet no guide to the comparative value of the characters made use +of. I therefore keep as much as possible to the old groups, founded on +external characters, only using the indications furnished by Professor +Garrod's paper, to determine the position of doubtful genera. + + +FAMILY 76.--CACATUIDÆ. (5 Genera, 35 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 |1. 2 -- -- + | | | | | + +The Cacatuidæ, Plyctolophidæ, or Camptolophidæ, as they have been variously +termed, comprise all those crested parrots usually termed Cockatoos, +together with one or two doubtful forms. They are very abundant in the +Australian region, more especially in the Austro-Malayan portion of it, one +species inhabiting {325}the Philippine Islands; but they do not pass +further east than the Solomon Islands and are not found in New Zealand. The +distribution of the genera is as follow:-- + +_Cacatua_ (18 sp.), ranges from the Philippine Islands, Celebes and Lombok, +to the Solomon Islands and to Tasmania; _Calopsitta_ (1 sp.), Australia; +_Calyptorhynchus_ (8 sp.), is confined to Australia and Tasmania; +_Microglossus_ (2 sp.), (perhaps a distinct family) to the Papuan district +and North Australia; _Licmetis_ (3 sp.), Australia, Solomon Islands, and +(?) New Guinea; _Nasiterna_ (3 sp.), a minute form, the smallest of the +whole order, and perhaps not belonging to this family, is only known from +the Papuan and Solomon Islands. + + +FAMILY 77.--PLATYCERCIDÆ. (11 Genera, 57 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3. 4 + | | | | | + +The Platycercidæ comprise a series of large-tailed Parrots, of weak +structure and gorgeous colours, with a few ground-feeding genera of more +sober protective tints; the whole family being confined to the Australian +region. The genera are:-- + +(1996 1999 2000) _Platycercus_ (14 sp.), Australia, Tasmania, and Norfolk +Island; _Psephotus_ (6 sp.), Australia; _Polytelis_ (3 sp.), Australia; +_Nymphicus_ (1 sp.), Australia and New Caledonia; (2002 2003) _Aprosmictus_ +(6 sp.), Australia, Papua, Timor, and Moluccas; _Pyrrhulopsis_ (3 sp.), +Tonga and Fiji Islands; _Cyanoramphus_ (14 sp.), New Zealand, Norfolk +Island, New Caledonia, and Society Islands; _Melopsittacus_ (1 sp.), +Australia; _Euphema_ (7 sp.), Australia; _Pezoporus_ (1 sp.), Australia and +Tasmania; _Geopsittacus_ (1 sp.), West Australia. The four last genera are +ground-feeders, and are believed by Professor Garrod to be allied to the +Owl-Parrot of New Zealand (_Stringops_). + + +{326}FAMILY 78.--PALÆORNITHIDÆ. (8 Genera, 65 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --| 1. 2 -- 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +I class here a group of birds brought together, for the most part, by +geographical distribution as well as by agreement in internal structure, +but which is nevertheless of a very uncertain and provisional character. + +_Palæornis_ (18 sp.), the Oriental region, Mauritius, Rodriguez, and +Seychelle Islands, and a species in Tropical Africa, apparently identical +with the Indian _P. torquatus_, and therefore--considering the very ancient +intercourse between the two countries, and the improbability of the +_species_ remaining unchanged if originating by natural causes--most likely +the progeny of domestic birds introduced from India. _Prioniturus_ (3 sp.), +Celebes and the Philippine Islands; (2061) _Geoffroyus_ (5 sp.), Bouru to +Timor and the Solomon Islands; _Tanygnathus_ (5 sp.), Philippines, Celebes, +and Moluccas to New Guinea; _Eclectus_ (8 sp.), Moluccas and Papuan +Islands; _Psittinus_ (1 sp.), Tenasserim to Sumatra and Borneo; +_Cyclopsitta_ (8 sp.), Papuan Islands, Philippines and North-east +Australia; _Loriculus_ (17 sp.), ranges over the whole Oriental region to +Flores, the Moluccas, and the Papuan island of Mysol; but most of the +species are concentrated in the district including the Philippines, +Celebes, Gilolo, and Flores, there being 1 in India, 1 in South China, 1 in +Ceylon, 1 in Java, 1 in Malacca, Sumatra, and Borneo, 3 in Celebes, 5 in +the Philippines, and the rest in the Moluccas, Mysol, and Flores. This +genus forms a transition to the next family. + + +{327}FAMILY 79.--TRICHOGLOSSIDÆ. (6 Genera, 57 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- + | | | | | + +The Trichoglossidæ, or Brush-tongued Paroquets, including the Lories, are +exclusively confined to the Australian region, where they extend from +Celebes to the Marquesas Islands, and south to Tasmania. The genus +_Nanodes_ (= _Lathamus_) has been shown by Professor Garrod to differ from +_Trichoglossus_ in the position of the carotid arteries. I therefore make +it a distinct genus but do not consider that it should be placed in another +family. The genera here admitted are as follows:-- + +_Trichoglossus_ (29 sp.), ranges over the whole Austro-Malay and Australian +sub-regions, and to the Society Islands; (2047) _Nanodes_ (1 sp.), +Australia and Tasmania; _Charmosyna_ (1 sp.), New Guinea (Plate X. Vol. I. +p. 414); _Eos_ (9 sp.), Bouru and Sanguir Island north of Celebes, to the +Solomon Islands, and in Puynipet Island to the north-east of New Ireland; +(2039 2040) _Lorius_ (13 sp.), Bouru and the Solomon Islands; (2041 2043) +_Coriphilus_ (4 sp.), Samoa, Tonga, Society and Marquesas Islands. + + +FAMILY 80.--CONURIDÆ. (7 Genera, 79 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |-- -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Conuridæ, which consist of the Macaws and their allies, are wholly +confined to America, ranging from the Straits of Magellan to South Carolina +and Nebraska, with Cuba and Jamaica. Professor Garrod places _Pyrrhura_ +(which has generally {328}been classed as a part of the genus _Conurus_) in +a separate family, on account of the absence of the ambiens muscle of the +knee, but as we are quite ignorant of the classificational value of this +character, it is better for the present to keep both as distinct genera of +the same family. The genera are:-- + +_Ara_ (15 sp.), Paraguay to Mexico and Cuba; _Rhyncopsitta_ (1 sp.), +Mexico; _Henicognathus_ (1 sp.), Chili; _Conurus_ (30 sp.), the range of +the family; _Pyrrhura_ (16 sp.), Paraguay and Bolivia to Costa Pica; +_Bolborhynchus_ (7 sp.), La Plata, Bolivia and West Peru, with one species +in Mexico and Guatemala; _Brotogerys_ (9 sp.), Brazil to Mexico. + + +FAMILY 81.--PSITTACIDÆ.--(12 Genera, 87 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |-- -- -- --|-- -- -- --|1. 2. 3. 4 |-- -- -- --|-- -- -- -- + | | | | | + +The Psittacidæ comprise a somewhat heterogeneous assemblage of Parrots and +Paroquets of the Neotropical and Ethiopian regions, which are combined here +more for convenience than because they are believed to form a natural +group. The genera _Chrysotis_ and _Pionus_ have no oil-gland, while +_Psittacula_ and _Agapornis_ have lost the furcula, but neither of these +characters are probably of more than generic value. The genera are:-- + +_Psittacus_ (2 sp.), West Africa; _Coracopsis_ (5 sp.), Madagascar, Comoro, +and Seychelle Islands; _Pæocephalus_ (9 sp.), all Tropical and South +Africa; (2063-2066) _Caica_ (9 sp.), Mexico to Amazonia; _Chrysotis_ (32 +sp.), Paraguay to Mexico and the West Indian Islands; _Triclaria_ (1 sp.), +Brazil; _Deroptyus_ (1 sp.), Amazonia; _Pionus_ (9 sp.), Paraguay to +Mexico; _Urochroma_ (7 sp.), Tropical South America; _Psittacula_ (6 sp.), +Brazil to Mexico; _Poliopsitta_ (2 sp.), Madagascar and West Africa; +_Agapornis_ (4 sp.), Tropical and South Africa. + + +{329}FAMILY 82.--NESTORIDÆ. (? 2 Genera, 6 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --| 1 -- -- 4 + | | | | | + +The present family is formed to receive the genus _Nestor_ (5 sp.), +confined to New Zealand and Norfolk Island. Its affinities are doubtful, +but it appears to have relations with the American Conuridæ and the +Australian Trichoglossidæ. With it is placed the rare and remarkable +_Dasyptilus_ (1 sp.), of New Guinea, of which however very little is known. + + +FAMILY 83.--STRINGOPIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 + | | | | | + +This family contains only the curious owl-like nocturnal Parrot of New +Zealand, _Stringops habroptilus_ (Plate XIII. Vol. I. p. 455). An allied +species is said to inhabit the Chatham Islands, if not now extinct. + + +_General Remarks on the Distribution of the Psittaci._ + +Although the Parrots are now generally divided into several distinct +families, yet they form so well marked and natural a group, and are so +widely separated from all other birds, that we may best discuss their +peculiarities of geographical distribution by treating them as a whole. By +the preceding enumeration we find that there are about 386 species of known +parrots, which are divided into 52 genera. They are pre-eminently a +tropical group, for although a few species extend a considerable distance +into the temperate zone, these are {330}marked exceptions to the rule which +limits the parrot tribe to the tropical and sub-tropical regions, roughly +defined as extending about 30° on each side of the equator. In America a +species of _Conurus_ reaches the straits of Magellan on the south, while +another inhabits the United States, and once extended to the great lakes, +although now confined to the south-eastern districts. In Africa parrots do +not reach the northern tropic, owing to the desert nature of the country; +and in the south they barely reach the Orange River. In India they extend +to about 35° N. in the western Himalayas; and in the Australian region, not +only to New Zealand but to Macquarie Islands in 54° S., the farthest point +from the equator reached by the group. But although found in all the +tropical regions they are most unequally distributed. Africa is poorest, +possessing only 6 genera and 25 species; the Oriental region is also very +poor, having but 6 genera and 29 species; the Neotropical region is much +richer, having 14 genera and 141 species; while the smallest in area and +the least tropical in climate--the Australian region, possesses 31 genera +and 176 species, and it also possesses exclusively 5 of the families, +Trichoglossidæ, Platycercidæ, Cacatuidæ, Nestoridæ, and Stringopidæ. The +portion of the earth's surface that contains the largest number of parrots +in proportion to its area is, undoubtedly, the Austro-Malayan sub-region, +including the islands from Celebes to the Solomon Islands. The area of +these islands is probably not one-fifteenth of that of the four tropical +regions, yet they contain from one-fifth to one-fourth of all the known +parrots. In this area too are found many of the most remarkable forms,--all +the crimson lories, the great black Cockatoos, the pigmy _Nasiterna_, the +raquet-tailed _Prioniturus_, and the bareheaded _Dasyptilus_. + +The almost universal distribution of Parrots wherever the climate is +sufficiently mild or uniform to furnish them with a perennial supply of +food, no less than their varied details of organization, combined with a +great uniformity of general type,--tell us, in unmistakable language, of a +very remote antiquity. The only early record of extinct parrots is, +however, in the Miocene of France, where remains apparently allied to the +West {331}African _Psittacus_, have been found. But the origin of so +widespread, isolated, and varied a group, must be far earlier than this, +and not improbably dates back beyond the dawn of the Tertiary period. Some +primeval forms may have entered the Australian region with the Marsupials, +or not long after them; while perhaps at a somewhat later epoch they were +introduced into South America. In these two regions they have greatly +flourished, while in the two other tropical regions only a few types have +been found, capable of maintaining themselves, among the higher forms of +mammalia, and in competition with a more varied series of birds. This seems +much more probable than the supposition that so highly organized a group +should have originated in the Australian region, and subsequently become so +widely spread over the globe. + + +_Order IV.--COLUMBÆ._ + +FAMILY 84.--COLUMBIDÆ. (44 Genera, 355 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Columbidæ, or Pigeons and Doves, are almost universally distributed, +but very unequally in the different regions. Being best adapted to live in +warm or temperate climates, they diminish rapidly northwards, reaching +about 62° N. Latitude in North America, but considerably farther in Europe. +Both the Nearctic and Palæarctic regions are very poor in genera and +species of pigeons, those of the former region being mostly allied to +Neotropical, and those of the latter to Oriental and Ethiopian types. The +Ethiopian region is, however, itself very poor, and several of its peculiar +forms are confined to the Madagascar sub-region. The Neotropical region is +very rich in peculiar genera, though but moderately so in number of +species. The Oriental {332}region closely approaches it in both respects; +but the Australian region is by far the richest, possessing nearly double +the genera and species of any other region, and abounding in remarkable +forms quite unlike those of any other part of the globe. The following +table gives the number of genera and species in each region, and enables us +readily to determine the comparative richness and isolation of each, as +regards this extensive family:-- + + Regions. No. of Genera. Peculiar Genera. No. of Species. + Neotropical 13 9 75 + Nearctic 5 1 7 + Palæarctic 3 0 9 + Ethiopian 6 1 37 + Oriental 12 1 66 + Australian 24 14 148 + +With the exception of _Columba_ and _Turtur_, which have a wide range, +_Treron_, common to the Oriental and Ethiopian regions, and _Carpophaga_, +to the Oriental and Australian, most of the genera of pigeons are either +restricted to or very characteristic of a single region. + +The distribution of the genera here admitted is as follows:-- + +_Treron_ (37 sp.), the whole Oriental region, and eastward to Celebes, +Amboyna and Flores, also the whole Ethiopian region to Madagascar; +_Ptilopus_ (52 sp.), the Australian region (excluding New Zealand) and the +Indo-Malay sub-region; _Alectroenas_ (4 sp.), Madagascar and the Mascarene +Islands; _Carpophaga_ (50 sp.), the whole Australian and Oriental regions, +but much the most abundant in the former; (2274) _Ianthoenas_ (11 sp.), +Japan, Andaman, Nicobar, and Philippine Islands, Timor and Gilolo to Samoa +Islands; (2278) _Leucomelæna_ (1 sp.), Australia; _Lopholaimus_ (1 sp.), +Australia; (2279 and 2283) _Alsæcomus_ (2 sp.), Himalayas to Ceylon and +Tenasserim; _Columba_ (46 sp.), generally distributed over all the regions +except the Australian, one species however in the Fiji Islands; +_Ectopistes_ (1 sp.), east of North America with British Columbia; +_Zenaidura_ (2 sp.), Veragua to Canada and British Columbia; _Oena_ (1 +sp.), Tropical and South Africa; _Geopelia_ (6 sp.), Philippine Islands and +Java to Australia; _Macropygia_ (14 sp.), Nepal, Hainan, Nicobar, Java, +{333}and Philippines to Australia and New Ireland; _Turacoena_ (3 sp.), +Celebes, Timor, and Solomon Islands; _Reinwardtoenas_ (1 sp.), Celebes to +New Guinea; _Turtur_ (24 sp.), Palæarctic, Ethiopian and Oriental regions +with Austro-Malaya; _Chæmepelia_ (7 sp.), Brazil and Bolivia to Jamaica, +California, and South-east United States; _Columbula_ (2 sp.), Brazil and +La Plata to Chili; _Scardafella_ (2 sp.), Brazil and Guatemala; _Zenaida_ +(10 sp.), Chili and La Plata to Columbia and the Antilles, Fernando +Noronha; _Melopelia_ (2 sp.), Chili to Mexico and California; _Peristera_ +(4 sp.), Brazil to Mexico; _Metriopelia_ (2 sp.), West America from Ecuador +to Chili; _Gymnopelia_ (1 sp.), West Peru and Bolivia; _Leptoptila_ (11 +sp.), Paraguay to Mexico and the Antilles; (2317 2318 and 2820) _Geotrygon_ +(14 sp.), Paraguay to Mexico and the Antilles; _Aplopelia_ (5 sp.), +Tropical and South Africa, St. Thomas and Princes Island; _Chalocopelia_ (4 +sp.), Tropical and South Africa; _Starnoenas_ (1 sp.), Cuba; Ocyphaps (1 +sp.), Australia (Plate XII. Vol. I. p. 441); _Petrophassa_ (1 sp.), +North-west Australia; _Chalocophaps_ (8 sp.), the Oriental region to New +Guinea and Australia; _Trugon_ (1 sp.), New Guinea; _Henicophaps_ (1 sp.), +Waigiou and New Guinea; _Phaps_ (3 sp.), Australia and Tasmania; +_Leucosarcia_ (1 sp.), East Australia; _Phapitreron_ (2 sp.), Philippine +Islands; _Geophaps_ (2 sp.), North and East Australia; _Lophophaps_ (3 +sp.), Australia; _Caloenas_ (1 sp.), scattered on the smaller islands from +the Nicobars and Philippines to New Guinea; _Otidiphaps_ (1 sp.), New +Guinea; _Phlogoenas_ (7 sp.), Philippine Islands and Celebes to the +Marquesas Islands; _Goura_ (2 sp.), New Guinea and the islands on the +north-east (Plate X. Vol. I. p. 414). + + +FAMILY 84_a_.--DIDUNCULIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- 3 -- + | | | | | + +{334}The _Didunculus stigirostris_, a hook-billed ground-pigeon, found only +in the Samoa Islands, is so peculiar in its structure that it is considered +to form a distinct family. + + +FAMILY 85.--DIDIDÆ.--(2 Genera, 3 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 |-- -- -- --|-- -- -- -- + | | | | | + +The birds which constitute this family are now all extinct; but as numerous +drawings are in existence, taken from living birds some of which were +exhibited in Europe, and a stuffed specimen, fragments of which still +remain, was in the Ashmolean Museum at Oxford down to 1755, they must be +classed among recent, as opposed to geologically extinct species. The Dodo +(_Didus ineptus_) a large, unwieldy, flightless bird, inhabited Mauritius +down to the latter part of the 17th century; and an allied form, the +Solitaire (_Pezophaps solitaria_), was found only in the island of +Rodriguez, where it survived about a century later. Old voyagers mention a +Dodo also in Bourbon, and a rude figure of it exists; but no remains of +this bird have been found. Almost complete skeletons of the Dodo and +Solitaire have, however, been recovered from the swamps of Mauritius and +the caves of Rodriguez, proving that they were both extremely modified +forms of pigeon. These large birds were formerly very abundant, and being +excellent eating and readily captured, the early voyagers to these islands +used them largely for food. As they could be caught by man, and very easily +by dogs, they were soon greatly diminished in numbers; and the introduction +of swine, which ran wild in the forests and fed on the eggs and young +birds, completed their extermination. + +The existence in the Mascarene Islands of a group of such remarkable +terrestrial birds, with aborted wings, is parallel to that of the _Apteryx_ +and _Dinornis_ in New Zealand, the Cassowaries of Austro-Malaya, and the +short-winged Rails of New {335}Zealand, Tristan d'Acunha, and other oceanic +islands; and the phenomenon is clearly dependent on the long-continued +absence of enemies, which allowed of great increase of bulk and the total +loss of the power of flight, without injury. In some few cases (the Ostrich +for example) birds incapable of flight co-exist with large carnivorous +mammalia; but these birds are large and powerful, as well as very swift, +and are thus able to escape from some enemies and defend themselves against +others. The entire absence of the smaller and more defenceless ground-birds +from the adjacent island of Madagascar, is quite in accordance with this +view, because that island has several small but destructive carnivorous +animals. + + +_General Remarks on the Distribution of the Columbæ._ + +The striking preponderance of Pigeons, both as to genera and species, in +the Australian region, would seem to indicate that at some former period it +possessed a more extensive land area in which this form of bird-life took +its rise. But there are other considerations which throw doubt upon this +view. The western half of the Malay Archipelago, belonging to the Oriental +region, is also rich in pigeons, since it has 43 species belonging to 11 +genera, rather more than are found in all the rest of the Oriental region. +Again, we find that the Mascarene Islands and the Antilles both possess +more pigeons than we should expect, in proportion to those of the regions +to which they belong, and to their total amount of bird-life. This looks as +if islands were more favourable to pigeon-development than continents; and +if we group together the Pacific and the Malayan Islands, the Mascarene +group and the Antilles, we find that they contain together about 170 +species of pigeons belonging to 24 out of the 47 genera here adopted; while +all the great continents united only produce about the same number of +species belonging (if we omit those peculiar to Australia) to only 20 +genera. The great development of the group in the Australian region may, +therefore, be due to its consisting mainly of islands, and not to the order +having originated there, and thus having had a longer period in which to +develop. I have elsewhere suggested (_Ibis_ 1865, p. 366) {336}a physical +cause for this peculiarity of distribution. Pigeons build rude, open nests, +and their young remain helpless for a considerable period. They are thus +exposed to the attacks of such arboreal quadrupeds or other animals as feed +on eggs or young birds. Monkeys are very destructive in this respect; and +it is a noteworthy fact that over the whole Australian region, the +Mascarene Islands and the Antilles, monkeys are unknown. In the Indo-Malay +sub-region, where monkeys are generally plentiful, the greatest variety of +pigeons occurs in the Philippines, where there is but a single species in +one island; and in Java, where monkeys are far less numerous than in +Sumatra or Borneo. If we add to this consideration the fact, that mammalia +and rapacious birds are, as a rule, far less abundant in islands than on +continents; and that the extreme development of pigeon-life is reached in +the Papuan group of islands, in which mammalia (except a few marsupials, +bats, and pigs) are wholly absent, we see further reason to adopt this +view. It is also to be noted that in America, comparatively few pigeons are +found in the rich forests (comparable to those of the Australian insular +region in which they abound), but are mostly confined to the open campos, +the high Andes, and the western coast districts, from which the +monkey-tribe are wholly absent. + +This view is further supported by the great development of colour that is +found in the pigeons of these insular regions, culminating in the +golden-yellow fruit-dove of the Fiji Islands, the metallic green +Nicobar-pigeon of Malaya, and the black and crimson _Alectroenas_ of +Mauritius. Here also, alone, we meet with crested pigeons, rendering the +possessors more conspicuous; such as the _Lopholaimus_ of Australia and the +crowned _Goura_ of New Guinea; and here too are more peculiar forms of +terrestrial pigeons than elsewhere, though none have completely lost the +power of flight but the now extinct Dididæ. + +The curious liking of pigeons for an insular habitat is well shown in the +genera _Ianthoenas_ and _Caloenas_. The former, containing 11 species, +ranges over a hundred degrees of longitude, and forty-five of latitude, +extending into three regions, yet nowhere inhabits a continent or even a +large island. It is {337}found in the Andaman and Nicobar Islands; in the +Philippines, Gilolo, and the smaller Papuan Islands, and in Japan; yet not +in any of the large Malay Islands or in Australia. The other genus, +_Caloenas_, consists of but a single species, yet this ranges from the +Nicobar Islands to New Guinea. It is not, however, as far as known, found +on any of the large islands, but seems to prefer the smaller islands which +surround them. We here have the general preference of pigeons for islands, +further developed in these two genera into a preference for small islands; +and it is probable that the same cause--the greater freedom from +danger--has produced both phenomena. + +Of the geological antiquity of the Columbæ we have no evidence; but their +wide distribution, their varied forms, and their great isolation, all point +to an origin, at least as far back as that we have assigned as probable in +the case of the Parrots. + + +_Order V.--GALLINÆ._ + + +FAMILY 86.--PTEROCLIDÆ. (2 Genera, 16 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2. 3. 4 | 1 -- 3. 4 |1 -- -- -- |-- -- -- -- + | | | | | + +The Pteroclidæ, or Sand-grouse, are elegantly formed birds with pointed +tails, and plumage of beautifully varied protective tints, characteristic +of the Ethiopian region and Central Asia, though extending into Southern +Europe and Hindostan. Being pre-eminently desert-birds, they avoid the +forest-districts of all these countries, but abound in the most arid +situations and on the most open and barren plains. The distribution of the +genera is as follows:-- + +_Pterocles_ (14 sp.), has the same range as the family; _Syrrhaptes_ (2 +sp.), normally inhabits Tartary, Thibet, and Mongolia to the country around +Pekin, and occasionally visits Eastern Europe. But a few years back (1863) +great numbers suddenly appeared in {338}Europe and extended westward to the +shores of the Atlantic, while some even reached Ireland and the Færoes. +(Plate III. Vol. I. p. 226.) + + +FAMILY 87.--TETRAONIDÆ. (29 Genera, 170 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- 4. + | | | | | + +The Tetraonidæ, including the Grouse, Partridges, Quails, and allied forms, +abound in all parts of the Eastern continents; they are less plentiful in +North America and comparatively scarce in South America, more than half the +Neotropical species being found north of Panama; and in the Australian +region there are only a few of small size. The Ethiopian region probably +contains most species; next comes the Oriental--India proper from the +Himalayas to Ceylon having twenty; while the Australian region, with 15 +species, is the poorest. These facts render it probable that the Tetraonidæ +are essentially denizens of the great northern continents, and that their +entrance into South America, Australia, and even South Africa, is, +comparatively speaking, recent. They have developed into forms equally +suited to the tropical plains and the arctic regions, some of them being +among the few denizens of the extreme north, as well as of the highest +alpine snows. The genera are somewhat unsettled, and there is even some +uncertainty as to the limits between this family and the next; but the +following are those now generally admitted:-- + +_Ptilopachus_ (1 sp.), West Africa; _Francolinus_ (34 sp.), all Africa, +South Europe, India to Ceylon, and South China; _Ortygornis_ (3 sp.), +Himalayas to Ceylon, Sumatra, and Borneo; _Peliperdix_ (1 sp.), West +Africa; _Perdix_ (3 sp.), the whole Continental Palæarctic region; +_Margaroperdix_ (1 sp.), Madagascar; _Oreoperdix_ (1 sp.), Formosa; +_Arborophila_ (8 sp.), the Oriental Continent and the Philippines; +_Peloperdix_ (4 sp.), Tenasserim and Malaya; _Coturnix_ (21 sp.), Temperate +Palæarctic, Ethiopian and {339}Oriental regions, and the Australian to New +Zealand; _Rollulus_ (2 sp.), Siam to Sumatra, Borneo, and Philippines; +_Caloperdix_ (1 sp.), Malacca and Sumatra; _Odontophorus_ (17 sp.), Brazil +and Peru to Mexico; _Dendrortyx_ (3 sp.), Guatemala and Mexico; _Cyrtonyx_ +(3 sp.), Guatemala to New Mexico; _Ortyx_ (8 sp.), Honduras and Cuba to +Canada; _Eupsychortyx_ (6 sp.), Brazil and Ecuador to Mexico; _Callipepla_ +(3 sp.), Mexico to California; _Lophortyx_ (2 sp.), Arizona and California; +_Oreortyx_ (1 sp.), California and Oregon (Plate XVIII., Vol. II. p. 128); +_Lerwa_ (1 sp.), Snowy Himalayas and East Thibet; _Caccabis_ (10 sp.), +Palæarctic region to Abyssinia, Arabia and the Punjaub; _Tetraogallus_ (4 +sp.), Caucasus and Himalayas to Altai Mountains; _Tetrao_ (7 sp.), northern +parts of Palæarctic and Nearctic regions; _Centrocercus_ (1 sp.), Rocky +Mountains; _Pediocætes_ (2 sp.), North and North-west America (Plate XVIII. +Vol. II. p. 128); _Cupidonia_ (1 sp.), East and North-Central United States +and Canada; _Bonasa_ (3 sp.), north of Nearctic and Palæarctic regions; +_Lagopus_ (6 sp.), Arctic Zone and northern parts of Nearctic and +Palæarctic regions. + + +FAMILY 88.--PHASIANIDÆ. (18 Genera, 75 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- 3 -- |-- 2. 3 -- |-- 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1 -- -- -- + | | | | | + +The Phasianidæ, including the Pea-fowl, Pheasants, and Jungle-fowl, the +Turkeys, and the Guinea-fowl, are very widely distributed, but are far more +abundant than elsewhere in the Eastern parts of Asia, both tropical and +temperate. Leaving out the African guinea-fowls and the American turkeys, +we have 13 genera and 63 species belonging to the Oriental and Palæarctic +regions. These are grouped by Mr. Elliot (whose arrangement we mainly +follow) in 5 sub-families, of which 3--Pavonniæ, Euplocaminæ, and +Gallinæ--are chiefly Oriental, while the Lophophorniæ and Phasianinæ are +mostly Palæarctic or from the highlands on the {340}borders of the two +regions. The genera adopted by Mr. Elliot in his _Monograph_ are the +following:-- + +PAVONINÆ, 4 genera.--_Pavo_ (2 sp.), Himalayas to Ceylon, Siam, to +South-west China and Java; _Argusianus_ (4 sp.), Siam, Malay Peninsula, and +Borneo (Plate IX. Vol. I. p. 339); _Polyplectron_ (5 sp.), Upper Assam to +South-west China and Sumatra; _Crossoptilon_ (4 sp.), Thibet and North +China. (Plate III. Vol. I. p. 226.) + +LOPHOPHORINÆ, 4 genera.--_Lophophorus_ (3 sp.), High woody region of +Himalayas from Cashmere to West China; _Tetraophasis_ (1 sp.), East Thibet; +_Ceriornis_ (5 sp.), Highest woody Himalayas from Cashmere to Bhotan and +Western China (Plate VII. Vol. I. p. 331); _Pucrasia_ (3 sp.), Lower and +High woody Himalayas from the Hindoo Koosh to North-west China. + +PHASIANINÆ, 2 genera.--_Phasianus_ (12 sp.), Western Asia to Japan and +Formosa, south to near Canton and Yunan, and the Western Himalayas, north +to the Altai Mountains; _Thaumalea_ (3 sp.), North-western China and +Mongolia. (Plate III. Vol. I. p. 226.) + +EUPLOCAMINÆ, 2 genera.--_Euplocamus_ (12 sp.), Cashmere, along Southern +Himalayas to Siam, South China and Formosa, and to Sumatra and Borneo; +_Ithaginis_ (2 sp.), High Himalayas from Nepal to North-west China. + +GALLINÆ, 1 genus.--_Gallus_ (4 sp.), Cashmere to Hainan, Ceylon, Borneo, +Java, and eastwards to Celebes and Timor. (Central India, Ceylon, and East +Java, have each a distinct species of Jungle-fowl.) + +MELEAGRINÆ, 1 genus.--_Meleagris_ (3 sp.), Eastern and Central United +States and south to Mexico, Guatemala and Yucatan. + +AGELASTINÆ, 2 genera.--_Phasidus_ (1 sp.), West Africa; _Agelastes_ (1 +sp.), West Africa. + +NUMIDINÆ, 2 genera.--_Acryllium_ (1 sp.), West Africa; _Numida_ (9 sp.), +Ethiopian region, east to Madagascar, south to Natal and Great Fish River. + + +{341}FAMILY 89.--TURNICIDÆ. (2 Genera, 24 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --| -- 2 -- 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Turnicidæ are small Quail-like birds, supposed to have remote +affinities with the American Tinamous, and with sufficient distinctive +peculiarities to constitute a separate family. They range over the Old +World, from Spain all through Africa and Madagascar, and over the whole +Oriental region to Formosa, and then north again to Pekin, as well as +south-eastward to Australia and Tasmania. The genus _Turnix_ (23 sp.), has +the range of the family; _Ortyxelos_ (1 sp.), inhabits Senegal; but the +latter genus may not belong to this family. + + +FAMILY 90.--MEGAPODIIDÆ. (4 Genera, 20 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- |-- -- -- 4 + | | | | | + +The Megapodiidæ, or Mound-makers and Brush-turkeys, are generally +dull-coloured birds of remarkable habits and economy, which have no near +allies, but are supposed to have a remote affinity with the South American +Curassows. They are highly characteristic of the Australian region, +extending into almost every part of it except New Zealand and the remotest +Pacific islands, and only sending two species beyond its limits,--a +_Megapodius_ in the Philippine Islands and North-west Borneo, and another +in the Nicobar Islands, separated by about 1,800 miles from its nearest +ally in Lombok. The Philippine species offers little difficulty, for these +birds are found on the smallest {342}islands and sand-banks, and can +evidently pass over a few miles of sea with ease; but the Nicobar bird is a +very different case, because none of the numerous intervening islands offer +a single example of the family. Instead of being a well-marked and clearly +differentiated form, as we should expect to find it if its remote and +isolated habitat were due to natural causes, it so nearly resembles some of +the closely-allied species of the Moluccas and New Guinea, that, had it +been found with them, it would hardly have been thought specifically +extinct. I therefore believe that it is probably an introduction by the +Malays, and that, owing to the absence of enemies and general suitability +of conditions, it has thriven in the islands and has become slightly +differentiated in colour from the parent stock. The following is the +distribution of the genera at present known:-- + +_Talegallus_ (2 sp.), New Guinea and East Australia; _Megacephalon_ (1 +sp.), East Celebes; _Lipoa_ (1 sp.), South Australia; _Megapodius_ (16 +sp.), Philippine Islands and Celebes, to Timor, North Australia, New +Caledonia, the Marian and Samoa Islands, and probably every intervening +island,--also a species (doubtfully indigenous) in the Nicobar Islands. + + +FAMILY 91.--CRACIDÆ, (12 Genera, 53 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +(Messrs. Sclater and Salvin's arrangement is here followed). + +The Cracidæ, or Curassows and Guans, comprise the largest and handsomest +game-birds of the Neotropical region, where they take the place of the +grouse and pheasants of the Old World. They are almost all forest-dwellers, +and are a strictly Neotropical family, only one species just entering the +Nearctic region as far as New Mexico. They extend southward to Paraguay and +the extreme south of Brazil, but none are found in the {343}Antilles, nor +west of the Andes south of the bay of Guayaquil. The sub-families and +genera are as follows:-- + +CRACINÆ, 4 genera.--_Crax_ (8 sp.), Mexico to Paraguay (Plate XV., Vol. II. +p. 28); _Nothocrax_ (1 sp.), Guiana, Upper Rio Negro, and Upper Amazon; +_Pauxi_ (1 sp.), Guiana to Venezuela; _Mitua_ (2 sp.), Guiana and Upper +Amazon. + +PENELOPINÆ, 7 genera.--_Stegnolæma_ (1 sp.), Columbia and Ecuador; +_Penelope_ (14 sp.), Mexico to Paraguay and to western slope of Ecuadorian +Andes; _Penelopina_ (1 sp.), Guatemala; _Pipile_ (3 sp.), Venezuela to +Eastern Brazil; _Aburria_ (1 sp), Columbia; _Chamæpetes_ (2 sp.), Costa +Rica to Peru; _Ortalida_ (18 sp.), New Mexico to Paraguay, also Tobago. + +OREOPHASINÆ, 1 genus.--_Oreophasis_ (1 sp.), Guatemala. + +It thus appears that the Cracinæ are confined to South America east of the +Andes, except one species in Central America; whereas nine Penelopinæ and +_Oreophasis_ are found north of Panama. The species of the larger genera +are strictly representative, each having its own distinct geographical +area, so that two species of the same genus are rarely or never found in +the same locality. + + +FAMILY 92.--TINAMIDÆ. (9 Genera, 39 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Tinamous are a very remarkable family of birds, with the general +appearance of partridges or hemipodes, but with the tail either very small +or entirely wanting. They differ greatly in their organization from any of +the Old World Gallinæ, and approach, in some respects, the Struthiones or +Ostrich tribe. They are very terrestrial in their habits, inhabiting the +forests, open plains, and mountains of the Neotropical region, from +Patagonia and Chili to Mexico; but, like the Cracidæ, they are absent from +the Antilles. Their colouring is very sober and protective, as is the case +with so many ground-birds, and they are seldom adorned {344}with crests or +other ornamental plumes, so prevalent in the order to which they belong. +The sub-families and genera, according to the arrangement of Messrs. +Sclater and Salvin, are as follows:-- + +TINAMINÆ, 7 genera.--_Tinamus_ (7 sp.), Mexico to Paraguay; _Nothocercus_ +(3 sp.), Costa Rica to Venezuela and Ecuador; _Crypturus_ (16 sp.), Mexico +to Paraguay and Bolivia; _Rhynchotus_ (2 sp.), Bolivia and South Brazil to +La Plata; _Nothoprocta_ (4 sp.), Ecuador to Bolivia and Chili; _Nothura_ (4 +sp.), Brazil and Bolivia to Patagonia; _Taoniscus_ (1 sp.), Brazil to +Paraguay. + +TINAMOTINÆ, 2 genera.-- _Calodromas_ (1 sp.), La Plata and Patagonia; +_Tinamotis_ (1 sp.), Andes of Peru and Bolivia. + + +_General Remarks on the Distribution of Gallinæ._ + +There are about 400 known species of Gallinaceous birds grouped into 76 +genera, of which no less than 65 are each restricted to a single region. +The Tetraonidæ are the only cosmopolitan family, and even these do not +extend into Temperate South America, and are very poorly represented in +Australia. The Cracidæ and Tinamidæ are strictly Neotropical, the +Megapodiidæ almost as strictly Australian. There remains the extensive +family of the Phasianidæ, which offers some interesting facts. We have +first the well-marked sub-families of the Numidinæ and Meleagrinæ, confined +to the Ethiopian and Nearctic regions respectively, and we find the +remaining five sub-families, comprising about 60 species, many of them the +most magnificent of known birds, spread over the Oriental and the +south-eastern portion of the Palæarctic regions. This restriction is +remarkable, since there is no apparent cause in climate or vegetation why +pheasants should not be found wild throughout southern Europe, as they were +during late Tertiary and Post-Tertiary times. We have also to notice the +remarkable absence of the Pheasant tribe from Hindostan and Ceylon, where +the peacock and jungle-fowl are their sole representatives. These two forms +also alone extend to Java, whereas in the adjacent islands of Borneo and +Sumatra we have _Argusianus_, _Polyplectron_, and _Euplocamus_. The common +jungle-fowl (the origin of our domestic poultry) is the only {345}species +which enters the Australian region as far as Celebes and Timor, and another +species (_Gallus æneus_) as far as Flores, and it is not improbable that +these may have been introduced by man and become wild. + +We have very little knowledge of the extinct forms of Gallinæ, but what we +have assures us of their high antiquity, since we find such distinct groups +as the jungle-fowl, partridges, and _Pterocles_, represented in Europe in +the Miocene period; while the Turkey, then as now, appears to have been a +special American type. + + +_Order VI.--OPISTHOCOMI._ + +FAMILY 93.--OPISTHOCOMIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Hoazin (_Opisthocomus cristatus_) is the sole representative of this +family and of the order Opisthocomi. It inhabits the eastern side of +Equatorial America in Guiana and the Lower Amazon; and at Pará is called +"Cigana" or gipsy. It is a large, brown, long-legged, weakly-formed and +loosely-crested bird, having such anomalies of structure that it is +impossible to class it along with any other family. It is one of those +survivors, which tell us of extinct groups, of whose past existence we +should otherwise, perhaps, remain for ever ignorant. + + +_Order VII.--ACCIPITRES._ + +FAMILY 94.--VULTURIDÆ. (10 Genera, 25 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3 -- |-- -- -- -- + | | | | | + +{346}Vultures range over all the great continents south of the Arctic +Circle, being only absent from the Australian region, the Malay Islands, +Ceylon, and Madagascar. The Old and New World forms are very distinct, +belonging to two well-marked divisions, often ranked as families. The +distribution of the genera is as follows:-- + +Sub-family I. VULTURINÆ (6 genera, 16 species), confined to the Old +World.--_Vultur_ (1 sp.), Spain and North Africa through Nepal to China +north of Ningpo; _Gyps_ (5 sp.), Europe south of 59°, Africa, except the +western sub-region, India, Siam, and Northern China; _Pseudogyps_ (2 sp.), +North-east Africa and Senegal, India and Burmah; _Otogyps_ (2 sp.), South +Europe, North-east and South Africa, India, and Siam; _Lophogyps_ (1 sp.), +North-east and South Africa and Senegal; _Neophron_ (4 sp.), South Europe, +India and the greater part of Africa. + +Sub-family II. SARCORHAMPHINÆ (4 genera, 9 species), confined to the New +World.--_Sarcorhamphus_ (2 sp.), "The Condor," Andes of South America, and +southern extremity below 41° south latitude; _Cathartes_ (1 sp.), America +from 20° south latitude to Trinidad and Mexico; _Catharistes_ (1 sp.), +America from 40° north to 40° south latitude, but not on Pacific coast of +United States; _Pseudogryphis_ (5 sp.), South America and Falkland Islands, +and to 49° north latitude in North America, also Cuba and Jamaica. + + +FAMILY 95.--SERPENTARIIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The singular Secretary Bird (_Serpentarius_) is found over a large part of +Africa. Its position is uncertain, as it has affinities both with the +Accipitres, through _Polyboroides_ (?) and with _Cariama_, which we place +near the Bustards. (Plate IV. Vol. I. p. 261.) + + +{347}FAMILY 96.--FALCONIDÆ. (69 Genera, 325 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Falconidæ, including the various groups of Hawks, Kites, Buzzards, +Eagles, and Falcons, are absolutely cosmopolitan, ranging far into the +arctic zone and visiting the most remote oceanic islands. They are abundant +in all the great continents and larger islands, preferring open to woody +regions. They are divided into several sub-families, the range of some of +which are restricted. For this family as well as the preceding I follow the +arrangement of Mr. Sharpe's _British Museum Catalogue_, and shall give the +approximate distribution of each sub-family, as well as of the several +genera. + +Sub-family I. POLYBORINÆ (2 genera, 10 species), the Neotropical region +with California and Florida, Tropical and South Africa.--_Polyborus_ (2 +sp.), South America, and to California and Florida; _Ibycter_ (8 sp.), +Tierra del Fuego to Honduras and Guatemala. + +_Cariama_ and _Serpentarius_, which Mr. Sharpe puts here, are so anomalous +that I think it better to class them in separate families--Serpentariidæ +among the Accipitres, and Cariamidæ near the Bustards. + +Sub-family II. ACCIPITRINÆ (10 genera, 87 +species).--Cosmopolitan.--_Polyboroides_ (2 sp.), Africa and Madagascar; +_Circus_ (15 sp.), Old and New Worlds, widely scattered, but absent from +Eastern Equatorial America, and the Malay Archipelago except Celebes; +_Micrastur_ (7 sp.), and _Geranospiza_ (2 sp.), Tropical parts of +Neotropical region; _Urotriorchis_ (1 sp.), West Africa; _Erythrocnema_ (1 +sp.), Chili and La Plata to California and Texas; _Melierax_ (5 sp.), +Africa except West African sub-region; _Astur_ (30 sp.), cosmopolitan, +except the Temperate South American sub-region; {348}_Nisoides_ (1 sp.), +Madagascar; _Eutriorchis_ (1 sp.), Madagascar; _Accipiter_ (23 sp.), +cosmopolitan, except Eastern Oceania. + +Sub-family III. BUTEONINÆ (13 genera, 51 sp.), cosmopolitan, except the +Malay and Pacific Islands.--_Urospizias_ (1 sp.), East and Central +Australia; _Heterospizias_ (1 sp.), Tropical South America east of the +Andes; _Tachytriorchis_ (2 sp.), Paraguay to California; _Buteo_ (18 sp.), +cosmopolitan, except the Australian region and the Indo-Malayan sub-region; +_Archibuteo_ (4 sp.), North America to Mexico and the cooler parts of the +Palæarctic region; _Buteola_ (1 sp.), Veragua to the Amazon Valley; +_Asturina_ (7 sp.), Paraguay and Bolivia to South-east United States; +_Busarellus_ (1 sp.), Brazil to Guiana; _Buteogallus_ (1 sp.), Guiana and +Columbia; _Urubutinga_ (12 sp.), South Brazil and Bolivia to Mexico; +_Harpyhaliæetus_ (1 sp.), Chili and North Patagonia to Veragua; _Morphnus_ +(1 sp.), Amazonia to Panama; _Thrasaëtus_ (1 sp.), Paraguay and Bolivia to +Mexico. + +Sub-family IV. AQUILINÆ (31 genera, 94 species), cosmopolitan.--_Gypaëtus_ +(2 sp.), south of Palæarctic region from Spain to North China, Abyssinia, +and South Africa; _Uroaëtus_ (1 sp.), Australia and Tasmania; _Aquila_ (9 +sp.), Nearctic, Palæarctic, and Ethiopian regions and India; _Nisaëtus_ (4 +sp.), Africa and South Europe, India, Ceylon, and Australia; +_Lophotriorchis_ (2 sp.), Indo-Malay sub-region, and Bogotá in South +America; _Neopus_ (1 sp.), India and Ceylon to Burmah, Java, Celebes and +Ternate; _Spiziastur_ (1 sp.), Guatemala to Brazil; _Spizaëtus_ (10 sp.), +Central and South America, Africa, India, and Ceylon, to Celebes and New +Guinea, Formosa, and Japan; _Lophoaëtus_ (1 sp.), all Africa; _Asturinula_ +(1 sp.), Africa, except extreme south; _Herpetotheres_ (1 sp.), Bolivia and +Paraguay to Southern Mexico; _Dryotriorchis_ (1 sp.), West Africa; +_Circaëtus_ (5 sp.) Africa to Central Europe, the Indian Peninsula, Timor; +_Spilornis_ (6 sp.), Oriental region and Celebes; _Butastur_ (4 sp.), +Oriental region to New Guinea and North-east Africa; _Helotarsus_ (2 sp.), +Africa south of the Sahara; _Haliæetus_ (7 sp.), cosmopolitan, except the +Neotropical region; _Gypohierax_ (1 sp.), West Africa and Zanzibar; +_Haliastur_ (2 sp.), Indian Peninsula to Ceylon, New {349}Caledonia, and +Australia; _Nauclerus_ (= _Elanoides_) (1 sp.), Brazil to Southern United +States; _Elanoides_ (= _Nauclerus_) (1 sp.), Western and North-eastern +Africa; _Milvus_ (6 sp.), the Old World and Australia; _Lophoictinia_ (1 +sp.), Australia; _Rostrhamus_ (3 sp.), Antilles and Florida to Brazil and +Peru; _Leptodon_ (4 sp.), Central America to South Brazil and Bolivia; +_Gypoictinia_ (1 sp.), South and West Australia; _Elanus_ (5 sp.), Africa, +India, and Malay Archipelago to Australia, South America to California; +_Gampsonyx_ (1 sp.), Trinidad to Brazil; _Henicopernis_ (1 sp.), Papuan +Islands; _Machærhamphus_ (2 sp.), South-west Africa, Madagascar, and +Malacca; _Pernis_ (3 sp.), Palæarctic, Oriental, and Ethiopian regions. + +Sub-family V. FALCONINÆ (11 genera, 80 species), cosmopolitan.--_Baza_ (10 +sp.), India and Ceylon to the Moluccas and North Australia, West Coast of +Africa, Natal, and Madagascar; _Harpagus_ (3 sp.), Central America to +Brazil and Peru; _Ictinia_ (2 sp.), Brazil to Southern United States; +_Hierax_ (= _Microhierax_, Sharpe), (4 sp.), Eastern Himalayas to Borneo +and Philippines; _Poliohierax_ (2 sp.), East Africa and Burmah; +_Spiziapteryx_ (1 sp.), La Plata; _Harpa_ (1 sp.), New Zealand and the +Auckland Islands; _Falco_ (27 sp.), cosmopolitan, except the Pacific +Islands; _Hierofalco_ (6 sp.), Nearctic and Palæarctic regions; +_Hieracidea_ (2 sp.), Australia; _Cerchneis_ (22 sp.), cosmopolitan, except +Oceania. + + +FAMILY 97.--PANDIONIDÆ. (2 Genera, 3 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Pandionidæ, or Fishing Hawks, are universally distributed, with the +exception of the Southern Temperate parts of South America. The genera +are:-- + +_Pandion_ (1 sp.), the range of the entire family; _Polioaëtus_ (2 sp.), +India through Malay Archipelago to Celebes and Sandwich Islands. + + +{350}FAMILY 98.--STRIGIDÆ. (23 Genera, 180 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Strigidæ, or Owls, form an extensive and well-known family of nocturnal +birds, which, although invariably placed next the Hawks, are now believed +to be not very closely allied to the other Accipitres. They range over the +whole globe, extending to the extreme polar regions and to the remotest +oceanic islands. Their classification is very unsettled, and we therefore +place the genera, for convenience, in the order in which they follow each +other in the _Hand List of Birds_. Those adopted by most ornithologists are +the following:-- + +_Surnia_ (1 sp.), the Arctic regions of both hemispheres; _Nyctea_ (1 sp.), +South Carolina to Greenland and Northern Europe; _Athene_ (40 sp.), the +Eastern hemisphere to New Zealand and the Solomon Islands; _Ninox_ (7 sp.), +the Oriental region, North China and Japan; _Glaucidium_ (7 sp.), +Neotropical region, California, and Oregon, Europe to North China; +_Micrathene_ (1 sp.), Mexico and Arizona; _Pholeoptynx_ (2 sp.), +Neotropical region, Texas, and North-west America; _Bubo_ (16 sp.), +universally distributed, excluding the Australian region; _Ketupa_ (3 sp.), +the Oriental region, Palestine; _Scotopelia_ (2 sp.), West and South +Africa; _Scops_ (30 sp.), universally distributed, excluding Australia and +Pacific Islands; _Gymnoglaux_ (2 sp.), Antilles; _Lophostrix_ (2 sp.), +Lower Amazon to Guatemala; _Syrnium_ (22 sp.), all regions but the +Australian; _Ciccaba_ (10 sp.), Paraguay to Mexico; _Nyctalatinus_ (1 sp.), +Columbia; _Pulsatrix_ (2 sp.), Brazil and Peru to Guatemala; _Asio_ (6 +sp.), all regions but the Australian, Sandwich Islands; _Nyctalops_ (1 +sp.), Cuba and Mexico to Brazil and Monte Video; _Pseudoscops_ (1 sp.), +Jamaica; _Nyctala_ (4 sp.), the North Temperate zone; _Strix_ (18 sp.), +universally distributed; _Phodilus_ (1 sp.), Himalayas and Malaya. + +{351}In Mr. Sharpe's Catalogue (published while this work was passing +through the press) the genera of Owls are reduced to 19, arranged in two +families--Strigidæ, containing our last two genera, and Bubonidæ, +comprising the remainder. The species are increased to 190; but some genera +are reduced, as _Strix_, which is said to contain only 5 species. + + +_General Remarks on the Distribution of the Accipitres._ + +The Birds of Prey are so widely distributed over the world's surface that +their general distribution calls for few remarks. Of the four families all +but one are cosmopolites, Vultures alone being absent from the Australian +region, as well as from Indo-Malaya and Madagascar. If we take the +sub-families, we find that each region has several which are confined to +it. The only parts of the world where there is a marked deficiency of +Accipitres is in the islands of the Pacific; and it may be noted, as a +rule, that these birds are more abundant in continents than in islands. +There is not so much difference between the number of Birds of Prey in +tropical and temperate regions, as is found in most other groups of +land-birds. North America and Europe have about 60 species each, while +India has about 80, and South America about 120. The total number of +Accipitres is 550 comprised in 104 genera, and 4 (or perhaps more properly +5) families. In this estimate I have not included the Serpentariidæ, +containing the Secretary Bird of Africa, as there is some doubt whether it +really belongs to the Order. + + +_Order VIII.--GRALLÆ._ + +FAMILY 99.--RALLIDÆ. (18 Genera, 153 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Rails are among the most widely distributed families of birds, many of +the genera being cosmopolitan, and several of the {352}species ranging over +half the globe. They are found in many remote islands; and in some of +these--as the _Gallinula_ of Tristan d'Acunha, and the _Notornis_ of Lord +Howe's Island and New Zealand,--they have lost the power of flight. The +classification of the Rallidæ is not satisfactory, and the following +enumeration of the genera must only be taken as affording a provisional +sketch of the distribution of the group:-- + +_Rallus_ (18 sp.), _Porzana_ (24 sp.), _Gallinula_ (17 sp.), and _Fulica_ +(10 sp.), have a world-wide range; _Ortygometra_ (1 sp.), ranges over the +whole North Temperate zone; _Porphyrio_ (14 sp.), is more especially +Oriental and Australian, but occurs also in South America, in Africa, and +in South Europe; _Eulabeornis_ (15 sp.), is Ethiopian, Malayan, and +Australian; _Himantornis_ (1 sp.), is West African only; _Aramides_ (24 +sp.), is North and South American; _Rallina_ (16 sp.), is Oriental, but +ranges eastward to Papua; _Habroptila_ (1 sp.), is confined to the +Moluccas; _Pareudiastes_ (1 sp.), the Samoa Islands; _Tribonyx_ (4 sp.), is +Australian, and has recently been found also in New Zealand; _Ocydromus_ (4 +sp.); _Notornis_ (2 sp.), (Plate XIII. Vol. I. p. 455); and _Cabalus_ (1 +sp.), are peculiar to the New Zealand group. + +The sub-family, Heliornithinæ (sometimes classed as a distinct family) +consists of 2 genera, _Heliornis_ (1 sp.), confined to the Neotropical +region; and _Podica_ (4 sp.), the Ethiopian region excluding Madagascar, +and with a species (perhaps forming another genus) in Borneo. + +_Extinct Rallidæ._--Remains of some species of this family have been found +in the Mascarene Islands, and historical evidence shows that they have +perhaps been extinct little more than a century. They belong to the genus +_Fulica_, and to two extinct genera, _Aphanapteryx_ and _Erythromachus_. +The _Aphanapteryx_ was a large bird of a reddish colour, with loose +plumage, and perhaps allied to _Ocydromus_. _Erythromachus_ was much +smaller, of a grey-and-white colour, and is said to have lived chiefly on +the eggs of the land-tortoises. (See _Ibis_, 1869, p. 256; and _Proc. Zool. +Soc._, 1875, p. 40.) + + +{353}FAMILY 100.--SCOLOPACIDÆ. (21 Genera, 121 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Scolopacidæ, comprehending the Snipes, Sandpipers, Curlews, and allied +genera, are perhaps as truly cosmopolitan as any family of birds, ranging +to the extreme north and visiting the remotest islands. The genera of +universal distribution are the following:-- + +_Numenius_ (16 sp.); _Limosa_ (6 sp.); _Totanus_ (12 sp.); _Tringoides_, (6 +sp.); _Himantopus_ (6 sp.); _Tringa_ (20 sp.); and _Gallinago_ (24 sp.). +Those which have a more or less restricted distribution are:-- + +_Ibidorhyncha_ (1 sp.), Central Asia and the Himalayas (Plate VII. Vol. I. +p. 331); _Helodromas_ (1 sp.), Palæarctic region and North India; _Terekia_ +(1 sp.), East Palæarctic, wandering to India and Australia; _Recurvirostra_ +(6 sp.), Nearctic region to the High Andes, South Palæarctic, East and +South Africa, Hindostan and Australia; _Micropelama_ (1 sp.), North America +to Chili; _Machetes_ (1 sp.), Palæarctic region and Hindostan (Plate I. +Vol. I. p. 195); _Ereunetes_ (3 sp.), Nearctic and Neotropical; +_Eurinorhynchus_ (1 sp.), North-east Asia and Bengal; _Calidris_ (1 sp.), +all regions but Australian; _Macrorhamphus_ (3 sp.), Palæarctic and +Nearctic, visits Brazil and India; _Scolopax_ (4 sp.), the whole Palæarctic +region, to India, Java, and Australia; _Philohela_ (1 sp.), East Nearctic; +_Rhynchæa_ (4 sp.), Ethiopian and Oriental, Australia, and Temperate South +America; _Phalaropus_ (3 sp.), North Temperate zone, and West Coast of +America to Chili. + + +{354}FAMILY 101.--CHIONIDIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1 -- -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Sheath-bills, _Chionis_ (2 sp.), are curious white birds, whose thick +bill has a horny sheath at the base. Their nearest ally is _Hæmatopus_, a +genus of Charadriidæ. These birds are confined to the Antarctic Islands, +especially the Falkland Islands, the Crozets and Kerguelen's Land. + + +FAMILY 102.--THINOCORIDÆ. (2 Genera, 6 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1 -- -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Thinocoridæ, or Quail-snipes, are small birds, confined to Temperate +South America. They have much the appearance of Quails but are more nearly +allied to Plovers. The two genera are:-- + +_Attagis_ (4 sp.), Falkland Islands, Straits of Magellan, Chili, Bolivia, +and the High Andes of Peru and Ecuador; _Thinocorus_ (2 sp.), La Plata, +Chili, and Peru. (Plate XVI. Vol. II. p. 40.) + + +FAMILY 103.--PARRIDÆ. (2 Genera, 11 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +{355}The Parridæ, or Jacanas, are remarkable long-toed birds, often of +elegant plumage, frequenting swamps and marshes, and walking on the +floating leaves of aquatic plants. They are found in all the tropics. +_Parra_ (10 sp.), has the distribution of the family; _Hydrophasianus_ (1 +sp.), is confined to the Oriental region. + + +FAMILY 104.--GLAREOLIDÆ. (3 Genera, 20 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +This family, comprising the Pratincoles and Coursers, is universally +distributed over the Old World and to Australia. + +_Glareola_ (9 sp.), has the distribution of the family; _Pluvianus_ (1 +sp.), is confined to North Africa; _Cursorius_ (10 sp.), ranges over +Africa, South Europe and India. + +The position of the genus _Glareola_ is uncertain, for though generally +classed here, Prof. Lilljeborg considers it to be an aberrant form of the +Caprimulgidæ! It differs, in its insectivorous habits and in many points of +external structure, from all its allies, and should probably form a +distinct family. + + +FAMILY 105.--CHARADIIDÆ. (19 Genera, 101 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The extensive family of the Plovers and their numerous allies, ranges over +the whole globe. The genera now usually admitted into this family are the +following:-- + +_Oedicnemus_ (9 sp.), is only absent from North America; _Æsacus_ (2 sp.), +India to Ceylon, Malay Islands and Australia; {356}_Vanellus_ (3 sp.), +Palæarctic and Neotropical regions; _Chætusia_ (15 sp.), the whole Eastern +Hemisphere; _Erythrogonys_ (1 sp.), Australia; _Hoplopterus_ (10 sp.), +widely scattered, but absent from North America; _Squatarola_ (1 sp), all +the regions; _Charadrius_ (14 sp.), cosmopolitan; _Eudromias_ (5 sp.), +Eastern Hemisphere and South Temperate America; _Ægialitis_ (22 sp.), +cosmopolitan; _Oreophilus_ (1 sp.), South Temperate America; _Thinornis_ (2 +sp.), New Zealand; _Anarhynchus_ (1 sp.), New Zealand (Plate XIII. Vol I. +p. 455); _Hæmatopus_ (9 sp.), cosmopolitan; _Strepsilas_ (2 sp.) almost +cosmopolitan; _Aphriza_ (1 sp.), West Coast of America; _Pluvianellus_ (1 +sp.), Straits of Magellan; _Dromas_ (1 sp.), India, Madagascar, and +North-east Africa; _Pedionomus_ (1 sp.), Australia. This last genus has +usually been placed with the Turnicidæ. + + +FAMILY 106.--OTIDIDÆ. (2 Genera, 26 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3 -- |-- 2 -- -- + | | | | | + +The Otididæ, or Bustards, occur in all parts of the Old World and Australia +where there are open tracts, being only absent from Madagascar and the +Malay Archipelago. + +_Otis_ (2 sp.), ranges over most of the Palæarctic region; while +_Eupodotis_ (24 sp.), has the range of the family, but is most abundant in +the Ethiopian region, which contains three-fourths of the whole number of +species. + + +FAMILY 107.--GRUIDÆ. (3 Genera, 16 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|1. 2. 3 -- |1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3 -- |-- 2 -- -- + | | | | | + +{357}The Gruidæ, or Cranes, are found in all the regions except the +Neotropical. + +_Grus_ (12 sp.) inhabits the southern and western United States, the whole +Palæarctic region, South-east Africa, India, and Australia; _Anthropoides_ +(2 sp.), Europe, North and South Africa and India; _Balearica_ (2 sp.), the +Ethiopian region (except Madagascar). + + +FAMILY 108.--CARIAMIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The genus _Cariama_ (2 sp.), consists of remarkable crested birds +inhabiting the mountains and open plains of Brazil and La Plata. In the +British Museum Catalogue of the Birds of Prey, they are classed as aberrant +Falconidæ, but their anomalous characters seem to require them to be placed +in a distinct family, which seems better placed among the Waders. + + +FAMILY 109.--ARAMIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Guaraünas are birds which have somewhat the appearance of Herons, but +which are usually classed with the Rails. They are now, however, considered +to form a distinct family. The only genus, _Aramus_ (2 sp.), inhabits the +Neotropical region, from Mexico and Cuba to Central Brazil. + + +{358}FAMILY 110.--PSOPHIIDÆ. (1 Genus, 6 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The remarkable and beautiful birds called Trumpeters, are confined to the +various parts of the Amazon valley; and it is an interesting fact, that the +range of each species appears to be bounded by some of the great rivers. +Thus, _Psophia crepitans_ inhabits the interior of Guiana as far as the +south bank of the Rio Negro; on the opposite or north bank of the Rio Negro +_Psophia ochroptera_ is found; beyond the next great rivers, Japura and +Iça, _Psophia napensis_ occurs; on the south bank of the Amazon, west of +the Madeira, we have the beautiful _Psophia leucoptera_; east of the +Madeira this is replaced by _Psophia viridis_, while near Pará, beyond the +Tapajoz, Xingu and Tocantins, there is another species, _Psophia obscura_. +Other species may exist in the intervening river districts; but we have +here, apparently, a case of a number of well-marked species of birds +capable of flight, yet with their range in certain directions accurately +defined by great rivers. (Plate XV. Vol. II. p. 28.) + + +FAMILY 111.--EURYPYGIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Eurypygidæ, or Sun-Bitterns, are small heron-like birds with +beautifully-coloured wings, which frequent the muddy and wooded river-banks +of tropical America. The only genus, _Eurypyga_ (2 sp.), ranges from +Central America to Brazil. + + +{359}FAMILY 112.--RHINOCHETIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The genus _Rhinochetus_ (1 sp.), consists of a singular bird called the +Kagu, which inhabits New Caledonia, an island which may be placed with +almost equal propriety in our 1st, 2nd, or 3rd Australian sub-regions. It +is a bird of a bluish ash-colour, with a loose plumage, partaking something +of the appearance of Rail, Plover, and Heron, but with peculiarities of +structure which require it to be placed in a distinct family. Its anatomy +shows that its nearest allies are the South American genera, _Eurypyga_ and +_Psophia_. + + +FAMILY 113.--ARDEIDÆ. (5 Genera, 80 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The well-known Herons and Bitterns are found in every part of the globe, +and everywhere closely resemble each other. Omitting the minuter +sub-divisions, the genera are as follows:-- + +_Ardea_ (60 sp.), cosmopolitan; _Botaurus_ (6 sp.), almost cosmopolitan; +_Tigrisoma_ (4 sp.), Tropical America and West Africa; _Nycticorax_ (9 +sp.), cosmopolitan; _Cancroma_ (1 sp.), Tropical America. + + +{360}FAMILY 114.--PLATALEIDÆ. (6 Genera, 30 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Plataleidæ, including the Spoonbills and Ibises, have been classed +either with the Herons or the Storks, but have most affinity with the +latter. Though not very numerous they are found over the greater part of +the globe, except the colder zones and the Pacific Islands. The following +is the distribution of the genera:-- + +_Platalea_ (6 sp.), all the warmer parts of the globe except the Moluccas +and Pacific Islands; _Ibis_ (2 sp.), Temperate North America and Tropical +South America; _Falcinellus_ (2 sp.), almost cosmopolitan; _Geronticus_ (19 +sp.), all Tropical countries and Temperate South America; _Scopus_ (1 sp.), +Tropical and South Africa; _Balæniceps_ (1 sp.), the Upper Nile. This last +genus the "Shoe-bird," or boat-billed heron, perhaps forms a distinct +family. + + +FAMILY 115.--CICONIIDÆ. (5 Genera, 20 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |-- -- 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Ciconiidæ, or Storks, are mostly an Old World family, only three +species inhabiting the Neotropical, and one, the Nearctic region. They are +also absent from the islands of the Pacific, the Antilles, and, with one +exception, from Madagascar. The genera are as follows:-- + +_Ciconia_ (6 sp.), ranges through the Palæarctic, Ethiopian and +{361}Oriental regions as far as Celebes, and in South America; _Mycteria_ +(4 sp.), inhabits Africa, India, Australia and the Neotropical region; +_Leptopiltus_ (3 sp.), the Ethiopian and Oriental regions to Java; +_Tantalus_ (5 sp.), the Ethiopian, Oriental and Neotropical regions, and +the South-east of North America; _Anastomus_ (2 sp.), the Ethiopian region, +and India to Ceylon. + + +FAMILY 116.--PALAMEDEIDÆ. (2 Genera, 3 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Palamedeidæ, or Screamers, are curious semi-aquatic birds of doubtful +affinities, perhaps intermediate between Gallinæ and Anseres. They are +peculiar to South America. The genera are:-- + +_Palamedea_ (1 sp.), which inhabits the Amazon valley; _Chauna_ (2 sp.), La +Plata, Brazil and Columbia, + + +FAMILY 117.--PHOENICOPTERIDÆ. (1 Genus, 8 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1 -- 3. 4 |-- -- -- --|-- 2 -- -- |1. 2. 3. 4 |1. 2 -- -- |-- -- -- -- + | | | | | + +The Flamingoes (_Phoenicopterus_) seem peculiar to the Ethiopian and +Neotropical regions, ranging from the former into India and South Europe. +America has four species, inhabiting Chili and La Plata, the Galapagos, +Mexico and the West Indian islands; the others range over all Africa, South +Europe, India and Ceylon. These singular birds are placed by some authors +near the Spoonbills and Ibises, by others with the Geese. Professor Huxley +considers them to be "completely {362}intermediate between the Anserine +birds on the one side and the Storks and Herons on the other." The +pterolysis according to Nitzsch is "completely stork-like." + + +_General Remarks on the Distribution of the Grallæ, or Wading and Running +Birds._ + +The Waders, as a rule, are birds of very wide distribution, the four +largest families Rallidæ, Scolopacidæ, Charadriidæ and Ardeidæ, being quite +cosmopolitan, as are many of the genera. But there are also a number of +small families of very restricted distribution, and these all occur in the +two most isolated regions, the Neotropical and the Australian. The +Neotropical region is by far the richest in varied forms of Waders, having +representatives of no less than 15 out of the 19 families, while 7 are +altogether peculiar to it. The Australian region has 11 families, with 1 +peculiar. The other two tropical regions each possess 11 families, but none +are peculiar. The Palæarctic region has 10, and the Nearctic 7 families. No +less than three families--Chionididæ, Thinocoridæ, and Cariamidæ--are +confined to the Temperate regions and highlands of South America; while +four others,--Aramidæ, Psophiidæ, Eurypygidæ and Palamedeidæ,--are found in +Tropical America only; and these present such an array of peculiar and +interesting forms as no other part of the globe can furnish. The +Phoenicopteridæ or Flamingoes, common to the Tropical regions of Asia, +Africa and America, but absent from Australia, is the only other feature of +general interest presented by the distribution of the Waders. + +The Order contains about 610 species, which gives about 32 species to each +family, a smaller average than in the Gallinæ or Accipitres, and only about +one-fourth of the average number in the Passeres. This is partly due to the +unusual number of very small families, and partly to the wide average range +of the species, which prevents that specialization of forms that occurs in +the more sedentary groups of birds. + + +{363}_Order IX.--ANSERES._ + +FAMILY 118.--ANATIDÆ. (40 Genera, 180 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Anatidæ, comprehending the Ducks, Geese, and Swans with their allies, +are of such universal distribution that there is probably no part of the +globe where some of them are not occasionally found. They are, however, +most abundant in temperate and cold regions; and, contrary to what occurs +in most other families, the most beautifully-coloured species are +extra-tropical, and some even arctic. The distribution of the genera is as +follows:-- + +_Anseranas_ (1 sp.), Australia; _Plectropterus_ (2 sp.), Tropical Africa; +_Sarkidiornis_ (1 sp.), South America, Africa, and India; _Chenalopex_ (1 +sp.), Amazonia; _Callochen_ (1 sp.), South Europe, North, East, and South +Africa; _Cereopsis_ (1 sp.), Australia; _Anser_ (13 sp.), Palæarctic and +Nearctic regions to Central America and the Antilles; _Bernicla_ (12 sp.), +Temperate regions of the Northern and Southern Hemispheres; _Chloephaga_ (5 +sp.), South Temperate America and Aleutian Islands; _Nettapus_ (4 sp.), +Tropical Africa and Madagascar, India and Ceylon to Malaya and Australia; +_Cygnus_ (10 sp.), Temperate regions of the Northern and Southern +Hemispheres; _Dendrocygna_ (10 sp.), Tropical and sub-tropical regions; +_Tadorna_ (3 sp.), Palæarctic and Australian regions; _Casarca_ (5 sp.), +Palæarctic, Oriental, Ethiopian, and Australian regions, to New Zealand; +_Aix_ (2 sp.), Temperate North America and Eastern Asia; _Mareca_ (4 sp.), +Palæarctic region, North America, Temperate South America, and Australia; +_Dafila_ (3 sp.), all America and the Palæarctic region; _Anas_ (16 sp.), +cosmopolitan; _Querquedula_ (17 sp.), {364}cosmopolitan; _Chaulelasmus_ (2 +sp.), Palæarctic region and North America; _Spatula_ (5 sp.), all Temperate +regions; _Malacorhynchus_ (1 sp.), Australia; _Cairina_ (1 sp.), Tropical +South America; _Branta_ (1 sp.), Palæarctic region and India; _Fuligula_ (5 +sp.), North Temperate regions and New Zealand; _Æthya_ (5 sp.), Palæarctic +and Nearctic regions, India, Australia, and South Africa; _Metopiana_ (1 +sp.), South Temperate America; _Bucephala_ (4 sp.), Nearctic and Palæarctic +regions; _Harelda_ (2 sp.), Northern Palæartic and Nearctic regions; +_Hymenolaimus_ (1 sp.), New Zealand; _Camptolaimus_ (1 sp.), North-east of +North America; _Micropterus_ (1 sp.), Temperate South America; _Somateria_ +(5 sp.), Arctic and sub-arctic regions; _Oedemia_ (5 sp.), Nearctic and +Palæarctic regions; _Biziura_ (1 sp.), Australia; _Thalassornis_ (1 sp.), +South Africa; _Erismatura_ (6 sp.), all America, South-east Europe and +South Africa; _Nesonetta_ (1 sp.), Auckland Islands; _Merganetta_ (3 sp.), +Andes of Columbia to Chili; _Mergus_ (6 sp.), Palæarctic and Nearctic +regions, Brazil, and the Auckland Islands. + + +FAMILY 119.--LARIDÆ. (13 Genera, 132 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Laridæ, or Gulls and Terns, are true cosmopolites, inhabiting the +shores and islands of every zone; and most of the genera have also a wide +range. They are therefore of little use in the study of geographical +distribution. The genera are as follows:-- + +_Stercorarius_ (6 sp.), cosmopolitan, most abundant in cold and temperate +zones; _Rhodostethia_ (1 sp), North America; _Larus_ (60 sp.), +cosmopolitan; _Xema_ (1 sp.), North Temperate zone; _Creagrus_ (1 sp.), +North Pacific; _Pagophila_ (1 sp.), Arctic seas; _Rissa_ (3 sp.), Arctic +and Northern seas; _Sterna_ (36 sp.), cosmopolitan; _Hydrochelidon_ (12 +sp.), Tropical and Temperate zones; {365}_Gygis_ (1 sp.), Indian Ocean and +Tropical Pacific Islands; _Anous_ (6 sp.), Tropical and Temperate zones; +_Nænia_ (1 sp.), South Temperate America; _Rhynchops_ (3 sp.), Tropical +America, Africa, and India. + + +FAMILY 120.--PROCELLARIIDÆ. (6 Genera, 96 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Procellariidæ, comprising the Shearwaters, Petrels, and Albatrosses, +are universally distributed, but some of the genera are local. + +_Puffinus_ (20 sp.), _Procellaria_ (18 sp.), and _Fulmarus_ (40 sp.), are +cosmopolitan; _Prion_ (5 sp.) and _Pelecanoides_ (3 sp.), belong to the +South Temperate and Antarctic regions; _Diomedia_ (10 sp.), comprises the +Albatrosses, which are tropical, occasionally wandering into temperate +seas. + + +FAMILY 121.--PELECANIDÆ. (6 Genera, 61 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Pelecanidæ, comprising the Gannets, Pelicans, Darters, and +Frigate-Birds, although universally distributed, are more abundant in +tropical and temperate regions. + +_Sula_ (8 sp.) and _Phalacrocorax_ (35 sp.), are cosmopolitan; _Pelecanus_ +(9 sp.) is tropical and temperate; _Fregetta_ (2 sp.) and _Phaeton_ (3 sp.) +are confined to Tropical seas; _Ptotus_ (4 sp.) to Tropical and warm +Temperate zones. + + +{366}FAMILY 122.--SPHENISCIDÆ. (3 Genera, 18 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 -- -- |-- -- -- --|-- -- -- --|-- -- 3 -- |-- -- -- --| -- 2 -- 4 + | | | | | + +The Penguins are entirely confined to the Antarctic and South Temperate +regions, except two species which are found on the coast of Peru and the +Galapagos. They are most plentiful in the southern parts of South America, +Australia, New Zealand, and most of the Antarctic islands, and one or two +species are found at the Cape of Good Hope. The genera as given in the +_Hand List_ are:-- + +_Spheniscus_ (1 sp.), South Africa and Cape Horn; _Eudyptes_ (15 sp.), with +the range of the family; _Aptenodytes_ (2 sp.), Antarctic Islands. + + +FAMILY 123.--COLYMBIDÆ. (1 Genus, 4 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- 4 | 1 -- 3. 4 |-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Northern Divers are confined to the Arctic and North Temperate Seas. +The only genus, _Colymbus_, has one species confined to the West Coast of +North America, the others being common to the two northern continents. + + +FAMILY 124.--PODICIPIDÆ. (2 Genera, 33 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +{367}The Grebes are universally distributed. The genera are _Podiceps_ (26 +sp.), cosmopolitan; and _Podilymbus_ (2 sp.), confined to North and South +America. Some ornithologists group these birds with the Colymbidæ. + + +FAMILY 125.--ALCIDÆ. (7 Genera, 28 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --| 1 -- -- 4 | 1 -- 3. 4 |-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Alcidæ, comprising the Auks, Guillemots, and Puffins, are confined to +the North Temperate and Arctic regions, where they represent the Penguins +of the Antarctic lands. One of the most remarkable of these birds, the +Great Auk, formerly abundant in the North Atlantic, is now extinct. The +genera are as follows:-- + +_Alca_ (2 sp.), North Atlantic and Arctic seas; _Fratercula_ (4 sp.), +Arctic and North Temperate zones; _Ceratorhina_ (2 sp.), North Pacific; +_Simorhynchus_ (8 sp.), North Pacific; _Brachyrhamphus_ (3 sp.), North +Pacific to Japan and Lower California; _Uria_ (8 sp.), Arctic and North +Temperate zones; _Mergulus_ (1 sp.), North Atlantic and Arctic Seas. The +last three genera constitute the family Uriidæ, of some ornithologists. + + +_General Remarks on the Distribution of the Anseres._ + +The Anseres, or Swimmers, being truly aquatic birds, possess, as might be +expected, a large number of cosmopolitan families and genera. No less than +5 out of the 8 families have a world-wide distribution, and the others are +characteristic either of the North or the South Temperate zones. Hence +arises a peculiarity of distribution to be found in no other order of +birds; the Temperate being richer than the Tropical regions. The Nearctic +and Palæarctic regions each have seven families of Anseres, two of which, +the Colymbidæ and Alcidæ, are peculiar to them. The Ethiopian, Australian, +and Neotropical regions, which all {368}extend into the South Temperate +zone, have six families, with one peculiar to them; while the Oriental +region, which is wholly tropical, possesses the five cosmopolitan families +only. + +There are about 78 genera and 552 species of Anseres, giving 69 species to +a family, a high number compared with the Waders, and due to there being +only one very small family, the Colymbidæ. The distribution of the Anseres, +being more determined by temperature than by barriers, the great regions +which are so well indicated by the genera and families of most other orders +of birds, hardly limit these, except in the case of the genera of Anatidæ. + + +_Order X.--STRUTHIONES._ + +FAMILY 126.--STRUTHIONIDÆ. (2 Genera, 4 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1 -- -- -- |-- -- -- --|-- 2 -- -- | 1 -- 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The Ostriches consist of two genera, sometimes formed into distinct +families. _Struthio_ (2 sp.) inhabits the desert regions of North, East, +and South Africa, as well as Arabia and Syria. It therefore just enters the +Palæarctic region. _Rhea_ (3 sp.) inhabits Temperate South America, from +Patagonia to the confines of Brazil. + + +FAMILY 127.--CASUARIIDÆ. (2 Genera, 11 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2 -- -- + | | | | | + +The Cassowaries and Emeus are confined to the Australian region. The Emeus, +_Dromæus_ (2 sp.), are found only on the {369}main-land of Australia (Plate +XII. Vol. I. p. 441). _Casuarius_ (9 sp.) inhabits the islands from Ceram +to New Britain, with one species in North Australia; it is most abundant in +the Papuan Islands. + + +FAMILY 128.--APTERYGIDÆ. (1 Genus, 4 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 + | | | | | + +The species of _Apteryx_ are entirely confined to the two larger islands of +New Zealand. They are supposed to have some remote affinity with +_Ocydromus_, a genus of Rails peculiar to Australia and New Zealand; but +they undoubtedly form one of the most remarkable groups of living birds +(Plate XIII. Vol. I. p. 445). + + +_Struthious Birds recently extinct._ + +A number of sub-fossil remains of birds, mostly large and some of gigantic +size, having affinities to the _Apteryx_ and, less closely, to the +Cassowaries, have been discovered in New Zealand. These are all classed by +Professor Owen in the genus _Dinornis_ and family _Dinornithidæ_; but Dr. +Haast, from the study of the rich collections in the Canterbury (New +Zealand) Museum, is convinced that they belong to two distinct families and +several genera. His arrangement is as follows. (See _Ibis_, 1874, p. 209). + + +FAMILY 129.--DINORNITHIDÆ. (2 Genera, 7 Species.) + +_Dinornis_ (5 sp.); _Meionornis_ (2 sp.). + +These had no hind toe, and include the largest species. Professor Newton +thinks that they were absolutely wingless, being the only birds in which +the fore limbs are entirely wanting. + + +{370}FAMILY 130.--PALAPTERYGIDÆ. (2 Genera, 4 Species.) + +_Palapteryx_ (2 sp.); _Euryapteryx_ (2 sp.). + +These had a well-developed hind toe, and rudimentary wings. + + +FAMILY 131.--ÆPYORNITHIDÆ. (1 Genus, 3 Species.) + +A gigantic Struthious bird (_Æpyornis_), belonging to a distinct family, +inhabited Madagascar. + +It was first made known by its enormous eggs, eight times the bulk of those +of the ostrich, which were found in a sub-fossil condition. Considerable +portions of skeletons have since been discovered, showing that these huge +birds formed an altogether peculiar family of the order. + + +_General Remarks on the Distribution of the Struthiones._ + +With the exception of the Ostrich, which has spread northward into the +Palæarctic region, the Struthious birds, living and extinct, are confined +to the Southern hemisphere, each continent having its peculiar forms. It is +a remarkable fact that the two most nearly allied genera, _Struthio_ and +_Rhea_, should be found in Africa and South Temperate America respectively. +Equally remarkable is the development of these large forms of wingless +birds in Australia and the adjacent islands, and especially in New Zealand, +where we have evidence which renders it probable that about 20 species +recently coexisted. This points to the conclusion that New Zealand must, +not long since, have formed a much more extensive land, and that the +diminution of its area by subsidence has been one of the causes--and +perhaps the main one--in bringing about the extinction of many of the +larger species of these wingless birds. + +The wide distribution of the Struthiones may, as we have already suggested +(Vol. I., p. 287.), be best explained, by supposing them to represent a +very ancient type of bird, developed at a time when the more specialized +carnivorous mammalia had {371}not come into existence, and preserved only +in those areas which were long free from the incursions of such dangerous +enemies. The discovery of Struthious remains in Europe in the Lower Eocene +only, supports this view; for at this time carnivora were few and of +generalized type, and had probably not acquired sufficient speed and +activity to enable them to exterminate powerful and quick-running +terrestrial birds. It is, however, at a much more remote epoch that we may +expect to find the remains of the earlier forms of this group; while these +Eocene birds may perhaps represent that ancestral wide-spread type which, +when isolated in remoter continents and islands, became modified into the +American and African ostriches, the Emeus and Cassowaries of Australia, the +_Dinornis_ and _Æpyornis_ of New Zealand. + + + + +{372}CHAPTER XIX. + +THE DISTRIBUTION OF THE FAMILIES AND GENERA OF REPTILES AND AMPHIBIA. + + + +REPTILIA. + +_Order I.--OPHIDIA._ + +FAMILY 1.--TYPHLOPIDÆ.--(4 Genera, 70 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |-- -- -- --| -- 2 -- 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Typhlopidæ, or Blind Burrowing Snakes, are widely scattered over the +warmer regions of the earth, but are most abundant in the Oriental and +Australian regions, and least so in the Neotropical. They are absent from +the Nearctic region; and in the Palæarctic are found only in South-eastern +Europe and Japan. + +The most extensive genus is _Typhlops_, comprising over 60 species, and +having a range almost as extensive as the entire family. The other well +characterised genera are:-- + +_Typhlina_ (1 sp.), ranging from Penang to Java and Hong Kong; _Typhline_ +(1 sp.), the Cape of Good Hope; _Dibamus_ (1 sp.), New Guinea. + + +{373}FAMILY 2.--TORTRICIDÆ. (3 Genera, 5 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |1 -- -- -- |-- -- -- --|-- -- -- --|1. 2. 3. 4 |1 -- -- -- + | | | | | + +The Tortricidæ, or Short-tailed Burrowing Snakes, are a small family, one +portion of which ranges from India to Cambodja, and through the Malay +islands as far as Celebes and Timor; these form the genus _Cylindrophis_. +Another portion inhabits America, and consists of:-- + +_Charina_ (1 sp.), found in California and British Columbia; and _Tortrix_ +(1 sp.), in Tropical America. + +We have here a case of discontinuous distribution, indicating, either very +imperfect knowledge of the group, or that it is the remnant of a once +extensive family, on the road to extinction. + + +FAMILY 3.--XENOPELTIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- 3. 4 |1 -- -- -- + | | | | | + +The curious nocturnal carnivorous Snake, forming the genus _Xenopeltis_, +and the sole representative of this family, ranges from Penang to Cambodja, +and through the Malay Islands to Celebes. + + +FAMILY 4.--UROPELTIDÆ. (5 Genera, 18 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- |-- -- -- -- + | | | | | + +{374}The Uropeltidæ, or Rough-tailed Burrowing Snakes, are strictly +confined to Ceylon and the adjacent parts of Southern India, and would +almost alone serve to mark out our second Oriental sub-region. The genera +are:-- + +_Rhinophis_ (7 sp.), Ceylon; _Uropeltis_ (1 sp.), Ceylon; _Silybura_ (8 +sp.), Anamally Hills and Neilgherries; _Plecturus_ (3 sp.), Neilgherries +and Madras; and _Melanophidium_ (1 sp.), the Wynand. + + +FAMILY 5.--CALAMARIIDÆ. (32 Genera, 75 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3 -- |-- 2 -- -- |1. 2. 3 -- |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Calamariidæ, or Dwarf Ground Snakes, are found in all warm parts of the +globe, extending north into the United States as far as British Columbia +and Lake Superior; but they are absent from the Palæarctic region, with the +exception of a species found in Persia. The species are in a very confused +state. The best characterised genera are the following:-- + +_Calamaria_ (20 sp.), Persia, India to Java and the Philippine Islands, +Celebes, and New Guinea; _Rhabdosoma_ (18 sp.), Mexico and South America, +and also the Malay Islands as far east as Amboyna, Timor, and New Guinea; +_Typhlocalamus_ (1 sp.), Borneo; _Macrocalamus_ (1 sp.), India; _Aspidura_ +(3 sp.), India and Ceylon; _Haplocerus_ (1 sp.), Ceylon; _Streptophorus_ (3 +sp.), Central and South America;--with a host of others of less importance +or ill-defined. + + +FAMILY 6.--OLIGODONTIDÆ. (4 Genera, 40 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- 3 -- |-- -- -- 4 |-- -- -- --|1. 2. 3. 4 |-- -- -- -- + | | | | | + +{375}The Oligodontidæ are a small family of Ground Snakes which have been +separated from the Calamariidæ, and, with the exception of a few species, +are confined to the Oriental region. The best characterised genera are:-- + +_Oligodon_ (12 sp.), India, Ceylon, and Philippines; and, _Simotes_ (24 +sp.), India to China and Borneo. In addition to these, _Achalinus_ is +founded on a single species from Japan; and _Teleolepis_ consists of three +species from North and South America. + + +FAMILY 7.--COLUBRIDÆ. (50 Genera, 270 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Colubrine Snakes are universally distributed over the globe, and they +reach the extreme northern limits of the order. They are, however, almost +absent from Australia, being there represented only by a few species of +_Tropidonotus_ and _Coronella_ in the northern and eastern districts. This +great family consists of four divisions or sub-families: the Coronellinæ +(20 genera, 100 species), the Colubrinæ (16 genera, 70 species), the +Dryadinæ (7 genera, 50 species), and the Natricinæ (7 genera, 50 species). +The more important genera of Colubridæ are the following:-- + +_Ablabes_, _Coronella_, _Ptyas_, _Coluber_, and _Tropidonotus_--all have a +very wide distribution, but the two last are absent from South America, +although _Tropidonotus_ reaches Guatemala; _Tomodon_, _Xenodon_, _Liopis_, +_Stenorhina_, _Erythrolampus_, _Elapochrus_, _Callirhinus_, _Enophrys_, and +_Dromicus_--are confined to the Neotropical region; _Hypsirhynchus_, +_Cryptodacus_, _Jaltris_, and _Coloragia_, are confined to the West Indian +Islands; _Chilomeniscus_, _Conophis_, _Pituophis_, and _Ischcognathus_, to +North America, the latter going as far south as Guatemala; _Compsosoma_, +_Zamenis_, _Zaocys_, _Atretium_, _Xenochrophys_, and _Herpetoreas_, are +peculiarly Oriental, but _Zamenis_ extends into South Europe; +{376}_Lytorhynchus_, _Rhamnophis_, _Herpetethiops_ and _Grayia_, are +Ethiopian; _Rhinechis_ is peculiar to Europe; _Megablabes_ to Celebes, and +_Styporhynchus_ to Gilolo; _Cyclophis_, is found in the Oriental region, +Japan, and North America; _Spilotes_, in the Nearctic and Neotropical +regions; _Xenelaphis_ in the Oriental, Ethiopian, and Palæarctic regions; +_Philodryas_, _Heterodon_ and _Herpetodryas_ in America and Madagascar, the +latter genus being also found in China. + + +FAMILY 8.--HOMALOPSIDÆ. (24 Genera, 50 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1 -- 3 -- |-- -- 3 -- |-- 2. 3. 4 |-- 2 -- -- |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Homalopsidæ, or Fresh-water Snakes, have been separated from the +Hydridæ by Dr. Günther, and they include some groups which have been +usually classed with the Natricinæ. They are especially characteristic of +the Oriental region, where considerably more than half the genera and +species are found; next comes the Neotropical region which has 6 species; +while none of the other regions have more than 4 or 5. It is to be observed +that the Ethiopian species occur in West Africa only, and mostly constitute +peculiar genera, so that in this family the separation of the Ethiopian and +Oriental regions is very well marked. The best characterised genera of the +family are the following:-- + +_Cantoria_ (10 sp.), ranging from Europe to Japan, the Philippines, and +Timor, with one species in Guinea; _Hypsirhina_ (6 sp.), Bengal, China, and +Borneo; _Fordonia_ (3 sp.), Rangoon to Borneo and Timor; _Homalopsis_ (2 +sp.), Cambodja to Java; _Cerberus_ (2 sp.), Ceylon and Siam, the Malay +Islands, New Guinea, and North Australia; _Herpeton_ (1 sp.), Siam; +_Ferania_ (1 sp.), Bengal to Penang; _Pythonopsis_ (1 sp.), Borneo; _Myron_ +(2 sp.), India and North Australia; _Homalophis_ (1 sp.), Borneo; +_Hipistes_ (1 sp.), Penang; _Xenodermus_ (1 sp.), Java; _Neusterophis_ and +_Limnophis_, with one species each, are peculiar to West {377}Africa; +_Helicops_ (2 sp.), North and South America; _Farancia_ and _Dimodes_, with +one species each, are from New Orleans; and a few others imperfectly known +from Tropical America. + + +FAMILY 9.--PSAMMOPHIDÆ. (5 Genera, 20 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- |1. 2. 3. 4 | 1 -- 3. 4 |-- -- -- -- + | | | | | + +The Psammophidæ, or Desert Snakes, are a small group characteristic of the +Ethiopian and Oriental regions, but more abundant in the former. The +distribution of the genera is as follows:-- + +_Psammophis_ (16 sp.), ranges from West Africa to Persia and Calcutta; +_Coelopeltis_ (1 sp.), North and West Africa; _Mimophis_ (1 sp.), +Madagascar; _Psammodynastes_ (2 sp.), Sikhim to Cochin China, Borneo and +the Philippine Islands; and _Dromophis_ (1 sp.), Tropical Africa. + + +FAMILY 10.--RACHIODONTIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The Rachiodontidæ are a small and very isolated group of snakes of doubtful +affinities. The only genus, _Dasypeltis_ (2 sp.), is confined to West and +South Africa. + + +{378}FAMILY 11.--DENDROPHIDÆ. (7 Genera, 35 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |-- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Dendrophidæ, or Tree Snakes, are found in all the Tropical regions, but +are most abundant in the Oriental. The genera are distributed as follows:-- + +_Dendrophis_ ranges from India and Ceylon to the Pelew Islands and North +Australia, and has one species in West Africa; _Ahætulla_ is almost equally +divided between Tropical Africa and Tropical America; _Gonyosoma_ ranges +from Persia to Java and the Philippines; _Chrysopelea_ is found in India, +Borneo, the Philippines, Amboyna, and Mysol; _Hapsidrophis_ and +_Bucephalus_ are confined to Tropical Africa; and _Ithycyphus_ (1 sp.), is +peculiar to Madagascar. + + +FAMILY 12.--DRYIOPHIDÆ. (5 Genera, 15 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --| -- 2 -- 4 |1. 2. 3. 4 |1 -- -- -- + | | | | | + +The Dryiophidæ, or Whip Snakes, are a very well characterised family of +slender, green-coloured, arboreal serpents, found in the three tropical +regions but absent from Australia, although they just enter the Australian +region in the island of Celebes. In Africa they are confined to the West +Coast and Madagascar. The genera are:-- + +_Dryiophis_ (4 sp.), Tropical America and West Africa; _Tropidococcyx_ (1 +sp.), Central India; _Tragops_ (4 sp.), Bengal to China, the Philippines, +Java, and Celebes; _Passerita_ (2 sp.), Ceylon {379}and the Indian +Peninsula; and _Langaha_ (2 sp.), confined to Madagascar. + + +FAMILY 13.--DIPSADIDÆ. (11 Genera, 45 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- 2 -- -- |1. 2. 3 -- |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Dipsadidæ, or Nocturnal Tree Snakes, are distinguished from the last +family by their dark colours and nocturnal habits. They are about equally +abundant in the Oriental and Neotropical regions, less so in the Ethiopian, +while only a single species extends to North Australia. The following are +the best known genera:-- + +_Dipsas_, comprising all the Oriental species with one in Asia-Minor, and a +few from the Moluccas, New Guinea, North Australia, West Africa, and +Tropical America; _Thamnodyastes_, _Tropidodipsas_, and several others, +from Tropical America; _Dipsadoboa_, from West Africa and Tropical America; +_Leptodeira_, from Tropical and South Africa, South America, and Mexico; +and _Pythonodipsas_, from Central Africa. + + +FAMILY 14.--SCYTALIDÆ. (3 Genera, 10 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 |-- -- -- -- + | | | | | + +It is doubtful how far the three genera which constitute this family form a +natural assemblage. We can therefore draw no safe conclusions from the +peculiarity of their distribution--_Scytale_ and _Oxyrhopus_ being confined +to Tropical America; while _Hologerrhum_ inhabits the Philippine Islands. + + +{380}FAMILY 15.--LYCODONTIDÆ. (11 Genera, 35 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- |1. 2. 3. 4 |1 -- -- -- + | | | | | + +The Lycodontidæ, or Fanged Ground Snakes, are confined to the Ethiopian and +Oriental regions, over the whole of which they range, except that they are +absent from Madagascar and extend eastward to New Guinea. The genera have +often a limited distribution:-- + +_Lycodon_ ranges from India and Ceylon to China, the Philippines, and New +Guinea; _Tetragonosoma_, the Malay Peninsula and Islands; _Leptorhytaon_ +and _Ophites_, India; _Cercaspis_, Ceylon; and _Cyclocorus_, the +Philippines. The African genera are _Boædon_, _Lycophidion_, +_Holuropholis_, _Simocephalus_, and _Lamprophis_, the latter being found +only in South Africa. The species are nearly equally abundant in both +regions, but no genus is common to the two. + + +FAMILY 16.--AMBLYCEPHALIDÆ. (5 Genera, 12 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- 3. 4 |-- -- 3? -- + | | | | | + +The Amblycephalidæ, or Blunt Heads, are very singularly distributed, being +nearly equally divided between Tropical America and the eastern half of the +Oriental region, as will be seen by the following statement of the +distribution of the genera:-- + +_Amblycephalus_ (1 sp.), Malay Peninsula to Borneo and the Philippines; +_Pareas_ (3 sp.), Assam, China, Java, and Borneo; {381}_Asthenodipsas_ (1 +sp.), Malacca; _Leptognathus_ (6 sp.), Central and South America; and +_Anoplodipsas_ (1 sp.), supposed to come from New Caledonia, and, if so, +furnishing a link, though a very imperfect one, between the disconnected +halves of the family. + + +FAMILY 17.--PYTHONIDÆ. (21 Genera, 46 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1 -- -- -- |-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +The Pythonidæ, comprising the Rock Snakes, Pythons, and Boas, are confined +to the tropics, with the exception of one species in California. They are +very abundant in the Neotropical region, where nearly half the known +species occur; the Australian region comes next, while the Oriental is the +least prolific in these large serpents. The genera which have been +described are very numerous, but they are by no means well defined. The +following are the most important:-- + +_Python_ is confined to the Oriental region; _Morelia_, _Liasis_, and +_Nardoa_ are Australian and Papuan; _Enygrus_ is found in the Moluccas, New +Guinea and the Fiji Islands; _Hortulia_ is African; _Sanzinia_ is peculiar +to Madagascar; _Boa_, _Epicrates_, _Corallus_, _Ungalia_, and _Eunectes_ +are Tropical American; _Chilabothrus_ is peculiar to Jamaica and Mexico; +and _Lichanotus_ to California. + +An extinct species belonging to this family has been found in the +Brown-coal formation of Germany, of Miocene age. + + +FAMILY 18.--ERYCIDÆ. (3 Genera, 6 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- |-- 2 -- -- | 1 -- 3 -- |-- -- -- -- + | | | | | + +{382}The Erycidæ, or Land Snakes, form a small but natural family, chiefly +found in the desert zone on the confines of the Palæarctic, Oriental, and +Ethiopian regions. They range from South Europe to West Africa and to +Sikhim. The three genera are distributed as follows:-- + +_Cursoria_ (1 sp.), Afghanistan; _Gongylophis_ (1 sp.), India and Sikhim; +_Eryx_ (4 sp.), has the range of the entire family. + + +FAMILY 19.--ACROCHORDIDÆ. (2 Genera, 3 Species) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --| -- 2 -- 4 |1 -- -- -- + | | | | | + +The Acrochordidæ, or Wart Snakes, form a small and isolated group, found +only in two sub-divisions of the Oriental region--the South Indian and the +Malayan, and in New Guinea. + +_Acrochordus_, inhabits Penang, Singapore, and Borneo; _Chersydrus_, +Southern India and the Malay Peninsula, with a species recently discovered +in New Guinea. + + +FAMILY 20.--ELAPIDÆ. (23 Genera, 100 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |-- -- 3 -- |-- -- -- 4 |1. 2. 3 -- |1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +The Elapidæ, or Terrestrial venomous Colubrine Snakes, are an extensive +group, spread over the tropics of the whole world, but especially abundant +in Australia, where half the known species occur, some of them being the +most deadly of venomous serpents. In the Oriental region they are also +abundant, containing amongst other forms, the well-known Cobras. The +American species are almost equally numerous, but they all belong to one +{383}genus, and they are annulated with rings of various colours in a +manner quite distinct from any other members of this family. The genera, +which are all very distinct, are distributed as follows:-- + +_Diemenia_, _Acanthophis_, _Hoplocephalus_, _Brachiurophis_, _Tropidechis_, +_Pseudechis_, _Cacophis_, _Pseudonaje_, _Denisonia_, and _Vermicella_, are +Australian, the first two ranging to the Moluccas and New Guinea; _Ogmodon_ +occurs in the Fiji Islands; _Naja_, _Bungarus_, _Ophiophagus_, +_Pseudonaje_, _Xenurelaps_, _Doliophis_, _Megærophis_, and _Callophis_ are +Oriental, one species of the latter genus being found in Japan, while an +_Ophiophagus_ has been discovered in New Guinea; _Cyrtophis_, _Elapsoidea_, +and _Poecilophis_ are African: _Elaps_ is American, ranging as far north as +South Carolina, but not to the West Indian Islands. + + +FAMILY 21.--DENDRASPIDIDÆ. (1 Genus, 5 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2 -- -- |-- -- -- --|-- -- -- -- + | | | | | + +The single genus _Dendraspis_, constituting the family, is confined to +Tropical Africa. + + +FAMILY 22.--ATRACTASPIDIDÆ. (1 Genus, 4 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +This small family, consisting of the genus _Atractaspis_, is also confined +to Africa, but has hitherto only been found in the West and South. + + +{384}FAMILY 23.--HYDROPHIDÆ. (8 Genera, 50 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Hydrophidæ, or Sea Snakes, are a group of small-sized marine serpents, +abundant in the Indian and Australian seas, and extending as far west as +Madagascar, and as far east as Panama. They are very poisonous, and it is +probable that many species remain to be discovered. The genera are +distributed as follows:-- + +_Hydrophis_ (37 sp.), ranging from India to Formosa and Australia; +_Platurus_ (2 sp.), from the Bay of Bengal to New Guinea and New Zealand; +_Aipysurus_ (3 sp.), Java to New Guinea and Australia; _Disteira_ (1 sp.), +unknown locality; _Acalyptus_ (1 sp.), South-west Pacific; _Enhydrina_ (1 +sp.), Bay of Bengal to New Guinea; _Pelamis_ (1 sp.), Madagascar to New +Guinea, New Zealand, and Panama; _Emydocephalus_ (1 sp.), Australian Seas. + + +FAMILY 24.--CROTALIDÆ. (11 Genera, 40 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 .3. 4 |1. 2. 3. 4 |-- -- 3. 4 |-- -- -- --|1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Crotalidæ, or Pit Vipers, including the deadly Rattlesnakes, form a +well-marked family of fanged serpents, whose distribution is very +interesting. They abound most in the Oriental region, at least 5 of the +genera and 20 species being found within its limits, yet they are quite +unknown in the Ethiopian region--a parallel case to that of the Bears and +Deer. A few species are peculiar to the eastern portion of the Palæarctic +region, while {385}the Nearctic is actually richer than the Neotropical +region both in genera and species. This would point to the conclusion, that +the group originated in the Indo-Chinese sub-region and spread thence +north-east to North America, and so onward to South America, which, having +been the last to receive the group, has not had time to develop it largely, +notwithstanding its extreme adaptability to Reptilian life. The genera are +divided among the several regions as follows:-- + +_Craspedocephalus_ (7 sp.), Tropical America and the West Indian Islands; +_Cenchris_, _Crotalophorus_, _Uropsophorus_, and _Crotalus_, inhabiting +North America from Canada and British Columbia to Texas, one species +(_Crotalus horridus_) extending into South America; _Trimeresurus_ (16 +sp.), all India from Ceylon to Assam, Formosa, the Philippines and Celebes; +_Peltopelor_ and _Hypnale_ (1 sp. each), peculiar to India; _Calloselasma_ +(1 sp.), Siam; _Atropos_ (1 sp.), Java and Borneo; _Halys_ (3 sp.), +peculiar to Tartary, Thibet, Japan, North China, and Formosa. + + +FAMILY 25.--VIPERIDÆ. (3 Genera, 22 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Viperidæ, or True Vipers, are especially characteristic of the +Palæarctic and Ethiopian regions, only one species being found over a large +part of the Oriental region, and another reaching Central India. They are +especially abundant in Africa, and the Palæarctic confines in South-western +Asia. The common Viper ranges across the whole Palæarctic region from +Portugal to Saghalien Island, reaching to 67° North Latitude, in +Scandinavia, and to 58° in Central Siberia. The genera, according to Dr. +Strauch's synopsis, are distributed as follows:-- + +_Vipera_ (17 sp.), which has the range of the family, extending over the +whole of the Palæarctic and Ethiopian regions, except Madagascar, and as +far as Ceylon, Siam, and Java, in the Oriental {386}region; _Echis_ (2 +sp.), inhabiting North Africa to Persia and to Continental India; and +_Atheris_ (3 sp.), confined to West Africa. + + +_Remarks on the General Distribution of Ophidia._ + +The Ophidia, being preeminently a Tropical order--rapidly diminishing in +numbers as we go north in the Temperate Zone, and wholly ceasing long +before we reach the Arctic Circle--we cannot expect the two Northern +regions to exhibit any great variety or peculiarity. Yet in their warmer +portions they are tolerably rich; for, of the 25 families of snakes, 6 are +found in the Nearctic region, 10 in the Palæarctic, 13 in the Australian, +16 in the Neotropical, 17 in the Ethiopian, and no less than 22 in the +Oriental, which last is thus seen to be by far the richest of the great +regions in the variety of its forms of Ophidian life. The only regions that +possess altogether peculiar families of this order, are the Ethiopian (3), +and the Oriental (2); the usually rich and peculiar Neotropical region not +possessing exclusively, any family of snakes; and what is still more +remarkable, the Neotropical and Australian regions together, do not possess +a family peculiar to them. Every family inhabiting these two regions is +found also in the Oriental; and this fact, taken in connection with the +superior richness of the latter region both in families and genera, would +indicate that the Ophidia had their origin in the northern hemisphere of +the Old World (the ancient Palæarctic region) whence they spread on all +sides, in successive waves of migration, to the other regions. The +distribution of the genera peculiar to, or highly characteristic of, the +several regions is as follows:-- + +The Nearctic possesses 9; four of these belong to the Colubridæ, one to the +Pythonidæ, and four to the Crotalidæ. The Palæarctic region has only 2 +peculiar genera, belonging to the Colubridæ and Crotalidæ. The Ethiopian +has 25, belonging to 11 families; four to Colubridæ, five to Lycodontidæ, +and three to Elapidæ. The Oriental has no less than 50, belonging to 15 +families; five are Colubridæ, five Uropeltidæ, twelve Homalopsidæ, six +Lycodontidæ, three Amblycephalidæ, eight Elapidæ, and four {387}Crotalidæ. +The Australian has 16, belonging to three families only; eleven being +Elapidæ, and four Pythonidæ. The Neotropical has about 24, belonging to +eight families; ten are Colubridæ, six Pythonidæ, and the rest Dipsadidæ, +Scytalidæ, Amblycephalidæ, Elapidæ, and Crotalidæ. + +We find then, that in the Ophidia, the regions adopted in this work are +remarkably distinct; and that, in the case of the Oriental and Ethiopian, +the difference is strongly marked, a very large number of the genera being +confined to each region. It is interesting to observe, that in many cases +the affinity seems to be rather between the West Coast of Africa and the +Oriental region, than between the East Coast and the plains of India; thus +the Homalopsidæ--a highly characteristic Oriental family--occur on the West +Coast of Africa only; the Dryiophidæ, which range over the whole Oriental +region, only occur in Madagascar and West Africa in the Ethiopian; the +genus _Dipsas_ is found over all the Oriental region and again in West +Africa. A cause for this peculiarity has been suggested in our sketch of +the past history of the Ethiopian region, Vol. I. p. 288. In the +Lycodontidæ, which are strictly confined to these two regions, the genera +are all distinct, and the same is the case with the more widely distributed +Elapidæ; and although a few desert forms, such as _Echis_ and the Erycidæ, +are common to Africa and the dry plains of India, this is evidently due to +favourable climatic conditions, and cannot neutralise the striking +differences in the great mass of the family and generic forms which inhabit +the two regions. The union of Madagascar with the South-western part of the +Oriental region under the appellation Lemuria, finds no support in the +distribution of Ophidia; which, however, strikingly accords with the views +developed in the Third Part of this work, as to the great importance and +high antiquity of the Euro-Asiatic continent, as the chief land-centre from +which the higher organisms have spread over the globe. + +_Fossil Ophidia._--The oldest known remains of Ophidia occur in the Eocene +formation in the Isle of Sheppey; others are found in the Miocene (Brown +Coal) of Germany, and in some Tertiary beds in the United States. Most of +these appear to have been {388}large species belonging to the Pythonidæ, so +that we are evidently still very far from knowing anything of the earliest +forms of this order. In some of the later Tertiary deposits the poison +fangs of venomous species have been found; also a Colubrine snake from the +Upper Miocene of the South of France. + + +_Order II.--LACERTILIA._ + +FAMILY 26.--TROGONOPHIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The single species of _Trogonophis_, forming this family, is found only in +North Africa. + + +FAMILY 27.--CHIROTIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- 3 -- |-- -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +_Chirotes_, the genus which constitutes this family, inhabits Mexico, and +has also been found in Missouri, one of the Southern United States. + + +FAMILY 28.--AMPHISBÆNIDÆ. (1 Genus, 13 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 -- 4 |-- -- -- --|-- 2 -- -- |1. 2 -- -- |-- -- -- --|-- -- -- -- + | | | | | + +{389}The Amphisbænidæ, which, in the opinion of Dr. Günther, are all +comprised in the genus _Amphisbæna_, inhabit Spain and Asia Minor, North +and Tropical Africa, South America as far as Buenos-Ayres and the West +Indian Islands. + + +FAMILY 29.--LEPIDOSTERNIDÆ. (3 Genera, 6 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 -- -- |-- -- -- --|-- -- -- --|-- 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The small family of Lepidosternidæ has nearly the same distribution as the +last, indicating a curious relationship between the Tropical parts of +Africa and America. _Lepidosternon_ and _Cephalopeltis_ are American +genera, while _Monotrophis_ is African. + + +FAMILY 30.--VARANIDÆ. (3 Genera, 30 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- |1. 2. 3 -- |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Varanidæ, or Water Lizards, are most abundant in the Oriental region, +whence they extend into the Austro-Malay Islands as far as New Guinea, and +into Australia. Several species are found in Africa. _Psammosaurus_ (1 +sp.), is found in North Africa and North-western India; _Monitor_ (18 sp.), +has the range of the family; while _Hydrosaurus_ (8 sp.) ranges from Siam +to the Philippines, New Guinea, and Australia. + + +{390}FAMILY 31.--HELODERMIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The genus _Heloderma_, which constitutes this family, is found in Mexico. + + +FAMILY 32.--TEIDÆ. (12 Genera, 74 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Teidæ, or Teguexins--a group of Lizards allied to the European +Lacertidæ, but with differently formed superciliary scales--are highly +characteristic of the Neotropical region, abounding almost everywhere from +Patagonia to the Antilles and Mexico, and extending northwards to +California on the west and to Pennsylvania on the east. The most extensive +genus is _Ameiva_, containing nearly 60 species and having the range of the +entire family; _Teius_ (3 sp.), inhabits Brazil and Mendoza; _Callopistes_ +(2 sp.), Chili; _Centropyx_ (3 sp.), Paraguay to Alabama; _Dicrodon_ +(Peru); _Monoplocus_ (Western Ecuador); with _Acrantus_, _Acanthopyga_, +_Emminia_, _Crocodilurus_, _Custa_, and _Ada_, which each consist of a +single species, and all inhabit Tropical America. + + +FAMILY 33.--LACERTIDÆ. (18 Genera, 80 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3. 4 |-- 2 -- -- + | | | | | + +{391}The Lacertidæ, or Land Lizards, are small-sized, terrestrial, +non-burrowing lizards, very characteristic of the Palæarctic region, which +contains more than half the known species, and of the adjacent parts of the +Oriental and Ethiopian regions, but extending also to South Africa, to +Java, and even to Australia. The best-defined genera are the following:-- + +_Lacerta_ (10 sp.), ranging over all Central and South Europe to Poland, +and farther north in Russia and Siberia, eastward to Persia, and southward +to North and West Africa; _Zootoca_ (8 sp.), has nearly the same range in +Europe as the last genus, but has representatives in Madeira, South Africa, +and Australia; _Tachydromus_ (7 sp.) is widely scattered in Chinese Asia, +Japan, Borneo, and West Africa; _Acanthodactylus_ (10 sp.) is most abundant +in North Africa, but has a species in South Africa, and two in Central +India; _Eremias_ (18 sp.) is found all over Africa, and also in the Crimea, +Persia, Tartary and China; _Psammodromus_ (2 sp.), is confined to Spain, +France, and Italy; _Ophiops_ (6 sp.), inhabits India, Persia, and Asia +Minor to South Russia. Less strongly marked and perhaps less natural genera +are the following:-- + +_Thetia_ (1 sp.), Algiers; _Teira_ (1 sp.), Madeira; _Nucras_ (4 sp.), +Caucasus and South Africa; _Notopholis_ (4 sp.), South Europe and South +Africa; _Algira_ (3 sp.), North and South Africa; _Scrapteira_ (1 sp.), +Nubia; _Aspidorhinus_ (1 sp.), Caspian district; _Messalina_ (4 sp.), North +Africa, Persia, and North-west India _Cabrita_ (1 sp.), Central India; +_Pachyrhynchus_ (1 sp.), Benguela. + + +FAMILY 34.--ZONURIDÆ. (15 Genera, 52 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |1. 2. 3. 4 |-- 2 -- -- |1. 2. 3. 4 |-- -- 3 -- |-- 2 -- -- + | | | | | + +The Zonuridæ, or Land Lizards, characterised by a longitudinal fold of skin +on each side of the body, have a very remarkable {392}distribution. Their +head-quarters is the Ethiopian region, which contains more than half the +known genera and species, most of which are found in South Africa and +several in Madagascar. Next to Africa the largest number of genera and +species are found in Mexico and Central America, with a few in the +Antilles, South America, and California, and even as far north as British +Columbia. Three of the genera form a distinct sub-group--the Glass +Snakes,--the four species composing it being located in North Africa, North +America, South-eastern Europe, and the Khasya Hills. + +The prominent fact in the distribution of this family is, that the mass of +the genera and species form two groups, one in South Africa, the other in +Mexico,--countries between which it would be difficult to imagine any means +of communication. We have here, probably, an example of a once much more +extensive group, widely distributed over the globe, and which has continued +to maintain itself only in those districts especially adapted to its +peculiar type of organization. This must undoubtedly have been the case +with the genus _Pseudopus_, whose two species now inhabit South-eastern +Europe and the Khasya Hills in Assam respectively. + +The genera are,--_Cordylus_, _Pseudocordylus_, _Platysaurus_, +_Cordylosaurus_, _Pleurostrichus_, and _Saurophis_, confined to South +Africa; _Zonurus_, South and East Africa and Madagascar; _Gerrhosaurus_, +ranges over the whole Ethiopian region; _Cicigna_ is confined to +Madagascar; _Gerrhonotus_ (22 sp.), ranges from British Columbia, +California, and Texas, to Cuba and South America, but is most abundant in +Mexico and Central America; _Abronia_ and _Barissia_, are two genera of +doubtful distinctness, peculiar to Mexico; _Ophisaurus_ (the Glass Snake) +is found in the Southern United States as far as Virginia; the allied genus +_Hyalosaurus_ in North Africa; and _Pseudopus_, as above stated, in +South-east Europe and the Khasya Hills. + + +{393}FAMILY 35.--CHALCIDÆ. (3 Genera, 8 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |-- -- ?3 --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Chalcidæ are a small group of Lizards characteristic of Tropical +America, one species extending into the United States. + +The genera are _Chalcis_ (6 sp.), ranging from Central America to Chili; +two other species, which have been placed in distinct genera, inhabit North +America and Peru. + + +FAMILY 36.--ANADIADÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The single species of _Anadia_, constituting this family, inhabits Tropical +America. + + +FAMILY 37.--CHIROCOLIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The genus _Heterodactylus_, which constitutes this family, inhabits Brazil. + + +{394}FAMILY 38.--IPHISADÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The single species of _Iphisa_, has been found only at Para in Equatorial +America. + + +FAMILY 39.--CERCOSAURIDÆ. (1 Genus, 5 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The genus _Cercosaura_, is known only from Brazil and Ecuador. + + +FAMILY 40.--CHAMÆSAURIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +This family, consisting of a single species of the genus _Chamæsaura_, is +confined to South Africa. + + +{395}FAMILY 41.--GYMNOPTHALMIDÆ . (5 Genera, 14 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- 4 |-- -- -- --|1. 2. 3 -- |-- 2 -- 4 |-- -- -- --|1. 2. 3 -- + | | | | | + +The Gymnopthalmidæ, or Gape-eyed Scinks, so called from their rudimentary +eyelids, form a small group, which is widely and somewhat erratically +distributed, as will be seen by the following account of the distribution +of the genera:-- + +_Lerista_ (1 sp.) and three other species for which Dr. Gray has +established the genera--_Morethria_ (1 sp.), and _Menetia_ (2 sp.), are +confined to Australia; _Cryptoblepharus_ (4 sp.), is found in West +Australia, Timor, New Guinea, the Fiji Islands, and Mauritius; _Ablepharus_ +(4 sp.), inhabits Eastern and South-eastern Europe, Persia, Siberia, West +Africa, and the Bonin Islands; and _Gymnopthalmus_ (3 sp.), is found in +Brazil and the West Indies. + + +FAMILY 42.--PYGOPODIDÆ. (2 Genera, 3 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- + | | | | | + +This small family of two-legged Lizards, comprising the genera _Pygopus_ +and _Delma_, is found only in Australia proper and Tasmania. + + +{396}FAMILY 43.--APRASIADÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- + | | | | | + +The genus _Aprasia_, constituting this family, is found in West and South +Australia. + + +FAMILY 44.--LIALIDÆ. (1 Genus, 3 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- + | | | | | + +This family is also confined to Australia, the single genus, _Lialis_, +inhabiting the Western and Northern districts. + + +FAMILY 45.--SCINCIDÆ. (60 Genera, 300 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Scincidæ, or Scinks, are an extensive family of smooth-scaled lizards, +frequenting dry and stony places, and almost universally distributed over +the globe, being only absent from the cold northern and southern zones. The +family itself is a very natural one, and it contains many natural genera; +but a large number have been established which probably require careful +revision. The following include the more important and the best established +groups:-- + +{397}_Scincus_ (2 sp.), North Africa and Arabia; _Hinulia_ (20 sp.), most +of the Australian and Oriental regions; _Cyclodina_ (1 sp.), _Hombronia_ (1 +sp.), and _Lygosomella_ (1 sp.), all from New Zealand; _Keneuxia_ (1 sp.), +Philippines, Moluccas, and Papuan Islands; _Elania_ (1 sp.) New Guinea; +_Carlia_ (2 sp.), North Australia and New Guinea; _Mocoa_ (16 sp.), +Australia and New Zealand, with species in Borneo, West Africa, and Central +America; _Lipinia_ (3 sp.), Philippine Islands and New Guinea; _Lygosoma_ +(12 sp.), Australia, New Caledonia, Pelew and Philippine Islands; +_Tetradactylus_ (1 sp.), _Hemierges_ (2 sp.), _Chelomeles_ (2 sp.), +_Omolepida_ (1 sp.), _Lissolepis_ (1 sp.), _Siaphos_ (1 sp.), _Rhodona_ (3 +sp.) _Anomalpus_ (1 sp.), _Soridia_ (2 sp.), and _Ophioscincus_ (1 sp.) all +confined to Australia; _Cophoscincus_ (3 sp.), Philippine Islands, Celebes, +and Queensland; _Plestiodon_ (18 sp.), China and Japan, Africa, and America +as far north as Pennsylvania and Nebraska; _Eumeces_ (30 sp.), South +Palæarctic, Oriental and Australian regions, to New Ireland and North +Australia; _Mabouya_ (20 sp.), Oriental region, Austro-Malaya, North +Australia, the Neotropical region, and to Lat. 42° 30' in North America; +_Amphixestus_ (1 sp.), Borneo; _Hagria_ (1 sp.), and _Chiamela_ (1 sp.), +India; _Senira_ (1 sp.), Philippine Islands; _Brachymeles_ (2 sp.), +Philippine Islands and Australia; _Ophiodes_ (1 sp.), Brazil; _Anguis_ (3 +sp.), West Palæarctic region and South Africa; _Tribolonotus_ (1 sp.), New +Guinea; _Tropidophorus_ (2 sp.), Cochin-China and Philippine Islands; +_Norbea_ (2 sp.), Borneo and Australia; _Trachydosaurus_ (1 sp.), +Australia; Cyclodus (8 sp.), Australia, Aru Islands, and Ceram; +_Silubosaurus_ (2 sp.), _Egerina_ (2 sp.), and _Tropidolepisma_ (6 sp.), +all peculiar to Australia; _Heteropus_ (7 sp.), Australia, Austro-Malaya, +and Bourbon; _Pygomeles_ (1 sp.), Madagascar; _Dasia_ (1 sp.), Malaya; +_Euprepes_ (70 sp.), Ethiopian and Oriental regions, Austro-Malaya, South +America (?); _Celestus_ (9 sp.), peculiar to the Antilles, except a species +in Costa Rica; _Diploglossus_ (7 sp.), the Neotropical region;--with a +number of other genera founded on single species from various parts of the +world. + + +{398}FAMILY 46.--OPHIOMORIDÆ. (2 Genera, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The snake-like Lizard constituting the genus _Ophiomorus_, is found in +Southern Russia, Greece, and Algeria; while _Zygnopsis_ having four weak +limbs, has been recently discovered by Mr. Blanford in South Persia. The +family is therefore confined to our Mediterranean sub-region. + + +FAMILY 47.--SEPIDÆ. (7 Genera, 22 species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- |1. 2. 3. 4 |-- -- -- --|-- -- -- -- + | | | | | + +The Sepidæ, or Sand-Lizards, are a very natural group, almost confined to +the Ethiopian region, but extending into the desert country on the borders +of the Oriental region, and into the south of the Palæarctic region as far +as Palestine, Madeira, Spain, Italy, and even the South of France. The +genera are:-- + +_Seps_ (10 sp.), South Europe, Madeira, Teneriffe, Palestine, North Africa, +South Africa and Madagascar; _Sphenops_ (2 sp.), North Africa, Syria, West +Africa; _Scelotes_ (3 sp.), Angola to South Africa, Madagascar; _Thyrus_ (1 +sp.), Bourbon and Mauritius; _Amphiglossus_ (1 sp.), Madagascar; +_Sphenocephalus_ (1 sp.), Afghanistan; and _Sepsina_ (4 sp.), South-west +Africa. + + +{399}FAMILY 48.--ACONTIADÆ. (3 Genera, 7 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- 2. 3. 4 |-- 2 -- -- |1 -- -- -- + | | | | | + +This small family of snake-like Lizards has a very curious distribution, +being found in South and West Africa, Madagascar, Ceylon, and Ternate in +the Moluccas. _Acontias_ (4 sp.), is found in the four first-named +localities; _Nessia_ (2 sp.), is confined to Ceylon; _Typhloscincus_ (1 +sp.), to Ternate. + + +FAMILY 49.--GECKOTIDÆ. (50 Genera, 200 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Geckoes, or Wall-Lizards, form an extensive family, of almost universal +distribution in the warmer parts of the globe; and they must have some +exceptional means of dispersal, since they are found in many of the most +remote islands of the great oceans,--as the Galapagos, the Sandwich +Islands, Tahiti, New Zealand, the Loo-Choo and the Seychelle Islands, the +Nicobar Islands, Mauritius, Ascension, Madeira, and many others. The +following are the larger and more important genera:-- + +_Oëdura_ (3 sp.), Australia; _Diplodactylus_ (8 sp)., Australia, South +Africa, and California; _Phyllodactylus_ (8 sp.), widely scattered in +Tropical America, California, Madagascar, and Queensland; _Hemidactylus_ +(40 sp.), all tropical and warm countries; _Peropus_ (12 sp.), the Oriental +region, Papuan Islands, Mauritius, and Brazil; _Pentadactylus_ (7 sp.), +Oriental region and Australia; _Gecko_ (12 sp.), Oriental region to New +Guinea and {400}North Australia; _Gehyra_ (5 sp.), Australia, New Guinea +and Fiji Islands; _Tarentola_ (7 sp.), North Africa, North America, +Madeira, Borneo, South Africa; _Phelsuma_ (6 sp.), Madagascar, Bourbon, and +Andaman Islands; _Pachydactylus_ (5 sp.), South and West Africa, and +Ascension Island; _Sphærodactylus_ (5 sp.), the Neotropical region; +_Naultinus_, (6 sp.), New Zealand; _Goniodactylus_ (5 sp.), Australia, +Timor, South America and Algiers; _Heteronota_ (4 sp.), Australia, Fiji +Islands, New Guinea and Borneo; _Cubina_ (4 sp.), the Neotropical region; +_Gymnodactylus_ (16 sp.), all warm countries except Australia; _Phyllurus_ +(3 sp.), Australia; _Stenodactylus_ (4 sp.), North and West Africa, and Rio +Grande in North America. + +The remaining genera mostly consist of single species, and are pretty +equally distributed over the various parts of the world indicated in the +preceding list. Madagascar, the Seychelle Islands, Chili, the Sandwich +Islands, South Africa, Tahiti, the Philippine Islands, New Caledonia, and +Australia--all have peculiar genera, while two new ones have recently been +described from Persia. + + +FAMILY 50.--IGUANIDÆ. (56 Genera, 236 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- 3 -- + | | | | | + +The extensive family of the Iguanas is highly characteristic of the +Neotropical region, in every part of which the species abound, even as far +as nearly 50° South Latitude in Patagonia. They also extend northwards into +the warmer parts of the Nearctic region, as far as California, British +Columbia, and Kansas on the west, and to 43° North Latitude in the Eastern +States. A distinct genus occurs in the Fiji Islands, and one has been +described as from Australia, and another from Madagascar, but there is some +doubt about these. The most extensive genera are:-- + +_Anolius_ (84 sp.), found in most parts of Tropical America and {401}north +to California; _Tropidolepis_ (15 sp.), which has nearly the same range; +_Leiocephalus_ (14 sp.), Antilles, Guayaquil, and Galapagos Islands; +_Leiolæmus_ (14 sp.), Peru to Patagonia; _Sceloporus_ (9 sp.), from Brazil +to California and British Columbia, and on the east to Florida; +_Proctotretus_ (6 sp.), Chili and Patagonia; _Phrynosoma_ (8 sp), New +Mexico, California, Oregon and British Columbia, Arkansas and Florida; +_Iguana_ (5 sp.), Antilles and South America; _Cyclusa_ (4 sp.), Antilles, +Honduras, and Mexico. + +Among the host of smaller genera may be noted:-- + +_Brachylophus_, found in the Fiji Islands; _Trachycephalus_ and +_Oreocephalus_, peculiar to the Galapagos; _Oreodeira_, said to be from +Australia; _Diplolæmus_ and _Phymaturus_, found only in Chili and +Patagonia; and _Callisaurus_, _Uta_, _Euphryne_, _Uma_, and _Holbrookia_, +from New Mexico and California. All the other genera are from various parts +of Tropical America. + + +FAMILY 51.--AGAMIDÆ. (42 Genera, 156 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +The extensive family Agamidæ--the Eastern representative of the Iguanas--is +highly characteristic of the Oriental region, which possesses about half +the known genera and species. Of the remainder, the greater part inhabit +the Australian region; others range over the deserts of Central and Western +Asia and Northern Africa, as far as Greece and South Russia. One genus +extends through Africa to the Cape of Good Hope, and there are three +peculiar genera in Madagascar, but the family is very poorly represented in +the Ethiopian region. Many of these creatures are adorned with beautifully +varied and vivid colours, and the little "dragons" or flying-lizards are +among the most interesting forms in the entire order. The larger genera are +distributed as follows:-- + +{402}_Draco_ (18 sp.), the Oriental region, excluding Ceylon; _Otocryptis_ +(4 sp.), Ceylon, North India, Malaya; _Ceratophora_ (3 sp.), Ceylon; +_Gonyocephalus_ (8 sp.), Papuan Islands, Java, Borneo, Pelew Islands; +_Dilophyrus_ (7 sp.), Indo-Malaya and Siam; _Japalura_ (6 sp.), Himalayas, +Borneo, Formosa, and Loo Choo Islands; _Sitana_ (2 sp.), Central and South +India and Ceylon; _Bronchocela_ (3 sp.), Indo-Malaya, Cambodja, and +Celebes; _Calotes_ (12 sp.), Continental India to China, Philippine +Islands; _Oriocalotes_ (2 sp.), Himalayas; _Acanthosaura_ (5 sp.), Malacca +and Siam; _Tiaris_ (3 sp.), Andaman Islands, Borneo, Philippine and Papuan +Islands; _Physignathus_ (3 sp.), Cochin-China and Australia; _Uromastix_ (5 +sp.), South Russia, North Africa, Central India; _Stellio_ (5 sp.), +Caucasus and Greece to Arabia, High Himalayas and Central India; _Trapelus_ +(5 sp.), Tartary, Egypt, and Afghanistan; _Phrynocephalus_ (10 sp.), +Tartary and Mongolia, Persia and Afghanistan; _Lophura_ (2 sp.), Amboyna +and Pelew Islands; _Grammatophorus_ (14 sp.), Australia and Tasmania; +_Agama_ (14 sp.), North Africa to the Punjaub, South Africa. The remaining +genera each consist of a single species. Eight are peculiar to Australia, +one to the Fiji Islands, one to the Aru Islands, three to Ceylon, five to +other parts of the Oriental region, one to Persia, and one to South Russia. + + +FAMILY 52.--CHAMÆLEONIDÆ. (1 Genus, 30 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- 2 -- -- |1. 2. 3. 4 |1. 2 -- -- |-- -- -- -- + | | | | | + +The Chamæleons are an almost exclusively Ethiopian group, only one species, +the common Chamæleon, inhabiting North Africa and Western Asia as far as +Central India and Ceylon. They abound all over Africa, and peculiar species +are found in Madagascar and Bourbon, as well as in the Island of Fernando +Po. + + +{403}_General Remarks on the Distribution of the Lacertilia._ + +The distribution of the Lacertilia is, in many particulars, strikingly +opposed to that of the Ophidia. The Oriental, instead of being the richest +is one of the poorest regions, both in the number of families and in the +number of peculiar genera it contains; while in both these respects the +Neotropical is by far the richest. The distribution of the families is as +follows:-- + +The Nearctic region has 7 families, none of which are peculiar to it; but +it has 3 peculiar genera--_Chirotes_, _Ophisaurus_, and _Phrynosoma_. + +The Palæarctic region has 12 families, with two (Ophiomoridæ and +Trogonophidæ, each consisting of a single species) peculiar; while it has 6 +peculiar or very characteristic genera, _Trogonophis_ in North Africa, +_Psammodromus_ in South Europe, _Hyalosaurus_ in North Africa, _Scincus_ in +North Africa and Arabia, _Ophiomorus_ in East Europe and North Africa, and +_Phrynocephalus_ in Siberia, Tartary, and Afghanistan. We have here a +striking amount of diversity between the Nearctic and Palæarctic regions +with hardly a single point of resemblance. + +The Ethiopian region has 13 families, only one of which (the Chamæsauridæ, +consisting of a single species) is altogether peculiar; but it possesses 21 +peculiar or characteristic genera, 9 belonging to the Zonuridæ, 2 to the +Sepidæ, 7 to the Geckotidæ, and 3 to the Agamidæ. + +The Oriental region has only 8 families, none of which are peculiar; but +there are 28 peculiar genera, 6 belonging to the Scincidæ, 1 to the +Acontiadæ, 5 to the Geckotidæ, and 16 to the Agamidæ. Many lizards being +sand and desert-haunters, it is not surprising that a number of forms are +common to the borderlands of the Oriental and Ethiopian regions; yet the +Sepidæ, so abundant in all Africa, do not range to the peninsula of India; +and the equally Ethiopian Zonuridæ have only one Oriental species, found, +not in the peninsula but in the Khasya Hills. The Acontiadæ alone offer +some analogy to the distribution of the Lemurs, being found in Africa, +Madagascar, Ceylon, and the Moluccas. + +The Australian region has 11 families, 3 of which are {404}peculiar; and it +has about 40 peculiar genera in ten families, about half of these genera +belonging to the Scincidæ. Only 3 families of almost universal distribution +are common to the Australian and Neotropical regions, with one species of +the American Iguanidæ in the Fiji Islands, so that, as far as this order is +concerned, these two regions have little resemblance. + +The Neotropical region has 15 families, 6 of which are peculiar to it, and +it possesses more than 50 peculiar genera. These are distributed among 12 +families, but more than half belong to the Iguanidæ, and half the remainder +to the Teidæ,--the two families especially characteristic of the +Neotropical region. All the Nearctic families which are not of almost +universal distribution are peculiarly Neotropical, showing that the +Lacertilia of the former region have probably been derived almost +exclusively from the latter. + +On the whole the distribution of the Lacertilia shows a remarkable amount +of specialization in each of the great tropical regions, whence we may +infer that Southern Asia, Tropical Africa, Australia, and South America, +each obtained their original stock of this order at very remote periods, +and that there has since been little intercommunication between them. The +peculiar affinities indicated by such cases as the Lepidosternidæ, found +only in the tropics of Africa and South America, and _Tachydromus_ in +Eastern Asia and West Africa, may be the results either of once widely +distributed families surviving only in isolated localities where the +conditions are favourable,--or of some partial and temporary geographical +connection, allowing of a limited degree of intermixture of faunas. The +former appears to be the more probable and generally efficient cause, but +the latter may have operated in exceptional cases. + + +_Fossil Lacertilia._ + +These date back to the Triassic period, and they are found in most +succeeding formations, but it is not till the Tertiary period that forms +allied to existing genera occur. These are at present too rare and too +ill-defined to throw much light on the geographical distribution of the +order. + + +{405}_Order III.--RHYNCOCEPHALINA._ + +FAMILY 53.--RHYNCOCEPHALIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- 4 + | | | | | + +The singular and isolated genus _Hatteria_--the "Tuatara" or fringed +lizard--which alone constitutes this family, has peculiarities of structure +which separate it from both lizards and crocodiles, and mark it out as an +ancestral type, as distinct from other living reptiles as the Marsupials +are from other Mammalia. It is confined to New Zealand, and is chiefly +found on small islands near the north-east coast, being very rare, if not +extinct, on the main land. A fossil reptile named _Hyperodapedon_, of +Triassic age, has been found in Scotland and India, and is supposed by +Professor Huxley to be more nearly allied to _Hatteria_ than to any other +living animal. + + +_Order IV.--CROCODILIA._ + +FAMILY 54.--GAVIALIDÆ. (2 Genera, 3 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --| 1 -- -- 4 |1 -- -- -- + | | | | | + +The Gavials are long-snouted Crocodiles with large front teeth, and canines +fitting in notches of the upper jaw. They consist of two genera, _Gavialis_ +(1 sp.), inhabiting the Ganges; _Tomistoma_ (2 sp.), found in the rivers of +Borneo and North Australia. + + +{406}FAMILY 55.--CROCODILIDÆ. (1 Genus, 12 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |-- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1 -- -- -- + | | | | | + +The true Crocodiles, which have the canines in notches, and the large front +teeth in pits in the upper jaw, are widely distributed over the tropical +regions of the globe, inhabiting all the rivers of Africa, the shores and +estuaries of India, Siam, and eastward to North Australia. Other forms +inhabit Cuba, Yucatan, and Guatemala, to Ecuador and the Orinooko. Four +species are Asiatic, one exclusively Australian, three African, and four +American. These have been placed in distinct groups, but Dr. Günther +considers them all to form one genus, _Crocodilus_. + + +FAMILY 56.--ALLIGATORIDÆ. (1 Genus, 10 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Alligators, which are distinguished by having both the large front +teeth and the canines fitting into pits of the upper jaw, are confined to +the Neotropical, and the southern part of the Nearctic regions, from the +lower Mississippi and Texas through all Tropical America, but they appear +to be absent from the Antilles. They are all placed by Dr. Günther in the +single genus, _Alligator_. + + +_General Remarks on the Distribution of Crocodilia._ + +These animals, being few in number and wholly confined to the tropical and +sub-tropical regions, are of comparatively {407}little interest as regards +geographical distribution. America possesses both Crocodiles and +Alligators; India, Crocodiles and Gavials; while Africa has Crocodiles +only. Both Crocodiles and Gavials are found in the northern part of the +Australian region, so that neither of the three families are restricted to +a single region. + + +_Fossil Crocodilia._ + +The existing families of the order date back to the Eocene period in +Europe, and the Cretaceous in North America. In the south of England, +Alligators, Gavials and Crocodiles, all occur in Eocene beds, indicating +that the present distribution of these families is the result of partial +extinction, and a gradual restriction of their range--a most instructive +fact, suggesting the true explanation of a large number of cases of +discontinuous distribution which are sometimes held to prove the former +union of lands now divided by the deepest oceans. In more ancient +formations, a number of Crocodilian remains have been discovered which +cannot be classed in any existing families, and which, therefore, throw no +light on the existing distribution of the group. + + +_Order V.--CHELONIA._ + +FAMILY 57.--TESTUDINIDÆ. (14 Genera, 126 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 | 1. 2 -- 4 |1. 2. 3. 4 |1. 2. 3. 4 |1 -- -- -- + | | | | | + +The Testudinidæ, including the land and many fresh-water tortoises, are +very widely distributed over the Old and New worlds, but are entirely +absent from Australia. They are especially abundant in the Nearctic region, +as far north as Canada and British Columbia, and almost equally so in the +{408}Neotropical and Oriental regions; in the Ethiopian there is a +considerable diminution in the number of species, and in the Palæarctic +they are still less numerous, being confined to the warmer parts of it, +except one species which extends as far north as Hungary and Prussia. The +genera are:-- + +_Testudo_ (25 sp.), most abundant in the Ethiopian region, but also +extending over the Oriental region, into South Europe, and the Eastern +States of North America; _Emys_ (64 sp.), abundant in North America and +over the whole Oriental region, less so in the Neotropical and the +Palæarctic regions; _Cinosternon_ (13 sp.), United States and California, +and Tropical America; _Aromochelys_ (4 sp.), confined to the Eastern States +of North America; _Staurotypus_ (2 sp.), Guatemala and Mexico; _Chelydra_ +(1 sp.), Canada to Louisiana; _Claudius_ (1 sp.), Mexico; _Dermatemys_ (3 +sp.), South America, Guatemala, and Yucatan; _Terrapene_ (4 sp.), Maine to +Mexico, Sumatra to New Guinea, Shanghae and Formosa--a doubtfully natural +group; _Cinyxis_ (3 sp.), _Pyxis_ (1 sp.), _Chersina_ (4 sp.), are all +Ethiopian; _Dumerilia_ (1 sp.), is from Madagascar only. + + +FAMILY 58.--CHELYDIDÆ. (10 Genera, 44 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- -- |-- -- -- --|-- -- -- --|1. 2. 3. 4 |-- -- -- --|-- 2 -- -- + | | | | | + +The Chelydidæ, or fresh-water tortoises with imperfectly retractile heads, +have a remarkable distribution in the three great southern continents of +Africa, Australia, and South America; the largest number of species being +found in the latter country. The genera are:-- + +_Peltocephalus_ (1 sp.), _Podocnemis_ (6 sp.), _Hydromedusa_ (4 sp.), +_Chelys_ (1 sp.), and _Platemys_ (16 sp.), inhabiting South America from +the Orinooko to the La Plata, the latter genus occurring also in Australia +and New Guinea; _Chelodina_ (5 sp.), _Chelemys_ (1 sp.), and _Elseya_ (2 +sp.) from Australia; while _Sternotheres_ {409}(6 sp.), and _Pelomedusa_ (3 +sp.), inhabit Tropical and South Africa and Madagascar. + + +FAMILY 59.--TRIONYCHIDÆ. (3 Genera, 25 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- 3 -- |-- -- -- 4 |1. 2. 3 -- |1. 2. 3. 4 |-- -- -- -- + | | | | | + +The distribution of the Trionychidæ, or Soft Tortoises, is very different +from that of the Chelydidæ, yet is equally interesting. They abound most in +the Oriental region, extending beyond it to Northern China and Japan. In +the Nearctic region they are only found in the Eastern States, +corresponding curiously to the distribution of plants, in which the +affinity of Japan to the Eastern States is greater than to California. The +Trionychidæ are also found over the Ethiopian region, but not in +Madagascar. + +The genera are,--_Trionyx_ (17 sp.), which extends over the whole area of +the family as above indicated; _Cycloderma_ (5 sp.), peculiar to Africa; +_Emyda_ (3 sp.), the peninsula of India, Ceylon, and Africa. + + +FAMILY 60.--CHELONIIDÆ. (2 Genera, 5 Species.) + +GENERAL DISTRIBUTION.--All the warm and tropical Seas. + +The Marine Turtles are almost universally distributed. _Dermatochelys_ (1 +sp.), is found in the temperate seas of both the Northern and Southern +Hemispheres; _Chelone_ (4 sp.), ranges over all the tropical seas--_C. +viridis_, the epicureans' species, inhabiting the Atlantic, while _C. +imbricata_ which produces the "tortoiseshell" of commerce is found in the +Indian and Pacific oceans. + + +{410}_Remarks on the Distribution of the Chelonia._ + +The four families into which the Chelonia are classed have all of them a +wide distribution, though none are universal. The Ethiopian region seems to +be the richest, as it possesses 3 of the four families, while no other +region has more than 2; and it also possesses 7 peculiar genera. Next comes +the Neotropical region with 2 families and 6 peculiar genera; the +Australian with 3, and the Nearctic with 2 peculiar genera; while the +Oriental and Palæarctic regions possess none that are peculiar. There are +about 30 genera and 200 species in the whole order. + +_Fossil Chelonia._--The earliest undoubted remains of this order occur in +the Upper Oolite. These belong to the Cheloniidæ and Emydidæ, which are +also found in the Chalk. In the Tertiary beds Chelonia are more abundant, +and the Trionychidæ now appear. The Testudinidæ are first met with in the +Miocene formation of Europe and the Eocene of North America, the most +remarkable being the gigantic _Colossochelys Atlas_ of the Siwalik Hills. +It appears, therefore, that the families of the order Chelonia were already +specialised in the Secondary period, a fact which, together with their more +or less aquatic habits, sufficiently accounts for their generally wide +distribution. Species of _Testudo_, _Emys_, and _Trionyx_, are found in the +Upper Miocene of the south of France. + + +{411}AMPHIBIA. + + +_Order I.--PSEUDOPHIDIA._ + +FAMILY 1.--CÆCILIADÆ. (4 Genera, 10 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- 2 -- -- |1. 2. 3 -- |-- -- -- -- + | | | | | + +The Cæciliadæ are a curious group of worm-like Amphibia sparingly scattered +over the three great tropical regions. The genera are,--_Cæcilia_, which +inhabits West Africa, Malabar and South America; _Siphonopsis_, peculiar to +Brazil and Mexico; _Ichthyopsis_, from Ceylon and the Khasya Mountains; and +_Rhinatrema_ from Cayenne. + + +_Order II.--URODELA._ + +FAMILY 2.--SIRENIDÆ. (1 Genus, 3 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The genus _Siren_, consisting of eel-like Batrachians with two anterior +feet and permanent branchiæ, inhabits the South-Eastern States of North +America from Texas to Carolina. + + +{412}FAMILY 3.--PROTEIDÆ. (2 Genera, 4 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- 3 -- |1 -- -- -- |-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Proteidæ have four feet and persistent external branchiæ. The two +genera are,--_Proteus_ (1 sp.), found only in caverns of Central Europe; +and _Menobranchus_, which are like newts in form, and inhabit the Eastern +States of North America. + + +FAMILY 4.--AMPHIUMIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The genus _Amphiuma_, or _Murænopsis_, consists of slender eel-like +creatures with four rudimentary feet, and no external branchiæ. The species +inhabit the Southern United States from New Orleans to Carolina. + + +FAMILY 5.--MENOPOMIDÆ. (2 Genera, 4 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- 3 -- |-- -- -- 4 |-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +There are large Salamanders of repulsive appearance, found only in Eastern +Asia and the Eastern United States. The genera are,--_Sieboldia_ (2 sp.), +Japan and north-west China; _Menopoma_ = _Protonopsis_ (2 sp.), Ohio and +Alleghany rivers. + + +{413}FAMILY 6.--SALAMANDRIDÆ. (20 Genera, 85 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |1. 2. 3. 4 |1. 2. 3. 4 |-- -- -- --|-- -- 3 -- |-- -- -- -- + | | | | | + +The Salamandridæ, of which our common Newts are characteristic examples, +form an extensive family highly characteristic of the North Temperate +regions, a few species only extending into the Neotropical region along the +Andes to near Bogota, and one into the Oriental region in Western China. +The genera, as arranged by Dr. Strauch, are as follows:-- + +_Salamandra_ (2 sp.), Central and South Europe and North Africa; +_Pleurodeles_ (1 sp.), Spain, Portugal, and Morocco; _Bradybates_ (1 sp.), +Spain; _Triton_ (16 sp.), all Europe except the extreme north, Algeria, +North China and Japan, Eastern States of North America, California and +Oregon; _Chioglossa_ (2 sp.), Portugal and South Europe; _Salamandrina_ (1 +sp.), Italy to Dalmatia; _Ellipsoglossa_ (2 sp.), Japan; _Isodactylium_ (2 +sp.), East Siberia; _Onychodactylus_ (1 sp.), Japan; _Amblystoma_ (21 sp.), +Nearctic region from Canada and Oregon to Mexico, most abundant in Eastern +States; _Ranodon_ (1 sp.), Tartary and North-east China; _Dicamptodon_ (1 +sp.), California; _Plethodon_ (5 sp.), Massachusetts to Louisiana, and +Vancouver's Island to California; _Desmognathus_ (4 sp.), Eastern United +States south of latitude 43°; _Anaides_ (1 sp.), Oregon and Northern +California; _Hemidactylium_ (2 sp.), South-eastern United States and +Southern California; _Heredia_ (1 sp.), Oregon and California; _Spelerpes_ +(18 sp.), Eastern United States from Massachusetts to Mexico, Guatemala, +Costa Rica and Andes of Bogota, with a species in South Europe; +_Batrachoseps_ (2 sp.), South-eastern United States and California; +_Tylotriton_ (1 sp.), Yunan in West China. + + +{414}_Order III.--ANURA._ + +FAMILY 7.--RHINOPHRYNIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Rhinophrynidæ are Toads with imperfect ears and a tongue which is free +in front. The single species of _Rhinophrynus_, is a native of Mexico. + + +FAMILY 8.--PHRYNISCIDÆ. (5 Genera, 13 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |-- -- -- --|-- -- -- --|1. 2 -- -- |-- -- -- 4 |-- 2 -- -- + | | | | | + +The Phryniscidæ, or Toads with imperfect ears and tongue fixed in front, +are widely distributed over the warmer regions of the earth, but are most +abundant in the Neotropical region and Australia, while only single species +occur in the Old World. The genera are:-- + +_Phryniscus_ (7 sp.), from Costa Rica to Chili and Monte Video; +_Brachycephalus_ (1 sp.), Brazil; _Pseudophryne_ (3 sp.), Australia and +Tasmania; _Hemisus_ (1 sp.), Tropical Africa; _Micrhyla_ (1 sp.), Java. + + +FAMILY 9.--HYLAPLESIDÆ. (1 Genus, 5 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 -- 4 |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +{415}The Hylaplesidæ are Toads with perfect ears, and they seem to be +confined to the Neotropical region. The only genus, _Hylaplesia_ (5 sp.), +inhabits Brazil, Chili, and the Island of Hayti. + + +FAMILY 10.--BUFONIDÆ. (6 Genera, 64 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The rather extensive family of the Bufonidæ, which includes our common +Toad, and is characterised by prominent neck glands and tongue fixed in +front, is almost universally distributed, but is very rare in the +Australian region; one species being found in Celebes and one in Australia. +The genera are:-- + +_Kalophrynus_ (2 sp.), Borneo; _Bufo_ (58 sp.), has the range of the entire +family, except Australia; _Otilophus_ (1 sp.), South America; _Peltaphryne_ +(1 sp.), Porto Rico; _Pseudobufo_ (1 sp.), Malay Peninsula; _Schismaderma_ +(1 sp.), Natal; _Notaden_ (1 sp.), East Central Australia. + + +FAMILY 11.--XENORHINIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1 -- -- -- + | | | | | + +The Xenorhinidæ may be characterised as Toads with perfect ears and tongue +free in front. The only species of _Xenorhina_ is a native of New Guinea. + + +{416}FAMILY 12.--ENGYSTOMIDÆ. (15 Genera, 31 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3 -- |-- -- 3 -- |-- -- -- --|-- 2. 3 -- |1. 2. 3. 4 |-- 2 -- -- + | | | | | + +The Engystomidæ are Toads without neck-glands and with the tongue tied in +front. They are most abundant in the Oriental and Neotropical regions, +especially in the latter, which contains about half the known species, with +isolated species in Australia, Africa, and the Southern States of North +America. They appear to be the remnant of a once extensive and universally +distributed group, which has maintained itself in two remote regions, but +is dying out everywhere else. The genera are:-- + +_Engystoma_ (9 sp.), Carolina to La Plata, with one species in South China; +_Diplopelma_ (3 sp.), South India to China and Java; _Cacopus_ (2 sp.), +Central India; _Glyphoglossus_ (1 sp.), Pegu; _Callula_ (4 sp.), Sikhim, +Ceylon, China, and Borneo; _Brachymerus_ (1 sp.), South Africa; _Adenomera_ +(1 sp.), Brazil; _Pachybatrachus_ (1 sp.), Australia; _Breviceps_ (2 sp.), +South and West Africa; _Chelydobatrachus_ (1 sp.), West Australia; +_Hypopachus_ (1 sp.), Costa Rica; _Rhinoderma_ (1 sp.), Chili; _Atelopus_ +(1 sp.), Cayenne and Peru; _Copea_ (1 sp.), South America; _Paludicola_ (1 +sp.), New Granada. + + +FAMILY 13.--BOMBINATORIDÆ. (8 Genera, 9 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 -- -- |-- -- -- --|1. 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- 4 + | | | | | + +The Bombinatoridæ are a family of Frogs which have imperfect ears and no +neck-glands, and they have a very peculiar and {417}interesting +distribution, being confined to Central and South Europe, the southern part +of South America, and New Zealand. They consist of many isolated groups +forming five separate sub-families. The genera are:-- + +_Bombinator_, Central Europe and Italy; _Pelobates_ and _Didocus_, Central +Europe and Spain; _Telmatobius_ (2 sp.), Peru and Brazil; _Alsodes_, Chonos +Archipelago; _Cacotus_, Chili; _Liopelma_, New Zealand; _Nannophryne_, +Straits of Magellan. + + +FAMILY 14.--PLECTROMANTIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1 -- -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Plectromantidæ, which are Frogs with neck-glands, and the toes but not +the fingers dilated, consists of a single species of the genus +_Plectromantis_. It inhabits the region west of the Andes, and south of the +Equator. + + +FAMILY 15.--ALYTIDÆ. (5 Genera, 37 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- -- |1. 2. 3 -- |1 -- -- -- |1. 2. 3 -- |-- -- -- --|1. 2 -- -- + | | | | | + +The Alytidæ are Frogs with neck-glands and undilated toes. They are most +abundant in the Ethiopian region, with a few species in the Nearctic and +Australian regions, and one in Europe and Brazil respectively. The genera +are:-- + +_Alytes_ (1 sp.), Central Europe; _Scaphiopus_ (5 sp.), California to +Mexico and the Eastern States; _Hyperolius_ (29 sp.), all Africa, and two +in New Guinea and North Australia; _Helioporus_ (1 sp.), in Australia; +_Nattereria_ (1 sp.), Brazil. + + +{418}FAMILY 16.--PELODRYADÆ. (3 Genera, 7 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|1. 2 -- -- + | | | | | + +The Pelodryadæ are Tree Frogs with neck-glands, and are confined to the +Australian and Neotropical regions. The genera are:-- + +_Phyllomedusa_ (3 sp.), South America to Paraguay; _Chirodryas_, Australia; +and _Pelodryas_ (3 sp.), Moluccas, New Guinea and Australia. + + +FAMILY 17.--HYLIDÆ. (11 Genera, 94 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- |-- -- -- --|-- -- 3 -- |1. 2 -- -- + | | | | | + +The Hylidæ are glandless Tree Frogs with a broadened sacrum. They are most +abundant in the Neotropical region, which contains more than two-thirds of +the species; about twenty species are Australian; six or seven are +Nearctic, reaching northward to Great Bear Lake; while one only is +European, and one Oriental. The genera are:-- + +_Hyla_ (62 sp.), having the range of the whole family; _Hylella_ (1 sp.), +_Ololygon_ (1 sp.), _Pohlia_ (2 sp.), _Triprion_ (1 sp.), _Opisthodelphys_ +(1 sp.), and _Nototrema_ (4 sp.), are South American; while +_Trachycephalus_ (8 sp.), is peculiar to the Antilles, except one South +American species; _Pseudacris_ (1 sp.), ranges from Georgia, United States, +to Great Bear Lake; _Litoria_ (7 sp.), is Australian and Papuan, except one +species in Paraguay; _Ceratohyla_ (4 sp.), is only known from Ecuador. + + +{419}FAMILY 18.--POLYPEDATIDÆ. (24 Genera, 124 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |-- -- 3 -- |-- -- 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +The Polypedatidæ, or glandless Tree Frogs with narrowed sacrum, are almost +equally numerous in the Oriental and Neotropical regions, more than forty +species inhabiting each, while in the Ethiopian there are about half this +number, and the remainder are scattered over the other three regions, as +shown in the enumeration of the genera:-- + +_Ixalus_ (16 sp.), Oriental, except one in Japan, and one in Western +Polynesia; _Rhacophorus_ (7 sp.), and _Theloderma_ (1 sp.), are Oriental; +_Hylarana_ (10 sp.), Oriental, to the Solomon Islands and Tartary, Nicobar +Islands, West Africa, and Madagascar; _Megalixalus_ (1 sp.), Seychelle +Islands; _Leptomantis_ (1 sp.), Philippines; _Platymantis_ (5 sp.), New +Guinea, Philippines, and Fiji Islands; _Cornufer_ (2 sp.), Java and New +Guinea; _Polypedates_ (19 sp.), mostly Oriental, but two species in West +Africa, one Madagascar, two Japan, one Loo-Choo Islands, and one Hong Kong; +_Hylambates_ (3 sp.), _Hemimantis_ (1 sp.), and _Chiromantis_ (1 sp.), are +Ethiopian; _Rappia_ (13 sp.), is Ethiopian, and extends to Madagascar and +the Seychelle Islands; _Acris_ (2 sp.), is North American; _Elosia_ (1 +sp.), _Epirhixis_ (1 sp.), _Phyllobates_ (9 sp.), _Hylodes_ (26 sp.), +_Hyloxalus_ (1 sp.), _Pristimantis_ (1 sp.), _Crossodactylus_ (1 sp.), +_Calostethus_ (1 sp.), _Strabomantis_ (1 sp.), and _Leiyla_ (1 sp.), are +Neotropical, the last two being Central American, while species of +_Hylodes_ and _Phyllobates_ are found in the West Indian Islands. + + +{420}FAMILY 19.--RANIDÆ. (26 Genera, 150 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The Ranidæ, or true Frogs, are characterised by having simple undilated +toes, but neither neck-glands nor dilated sacrum. They are almost +cosmopolitan, extending to the extreme north and south from the North Cape +to Patagonia, and they are equally at home in the tropics. They are perhaps +most abundant in South America, where a large number of the genera and +species are found; the Ethiopian region comes next, while they are rather +less abundant in the Oriental and Australian regions; the Nearctic region +has much less (about 12 species), while the Palæarctic has only five, and +these two northern regions only possess the single genus _Rana_. The genera +are distributed as follows:-- + +_Rana_ (60 sp.), ranges all over the world, except Australia and South +America, although it extends into New Guinea and into Mexico and Central +America; it is most abundant in Africa. _Pyxicephalus_ (7 sp.), extends +over the whole Ethiopian region, Hindostan, the Himalayas, and Japan; +_Cystignathus_ (22 sp.), is mainly Neotropical, but has three species +Ethiopian. All the other genera are confined to single regions. The +Neotropical genera are:--_Odontophrynus_ (1 sp.), _Pseudis_ (1 sp.), +_Pithecopsis_ (1 sp.), _Ensophleus_ (1 sp.), _Limnocharis_ (1 sp.), +_Hemiphractus_ (1 sp.), all Tropical South American east of Andes; +_Ceratophrys_ (5 sp.), Panama to La Plata; _Cycloramphus_ (1 sp.), West +Ecuador and Chili; _Pleurodema_ (6 sp.), Venezuela to Patagonia; +_Leiuperus_ (12 sp.), Mexico and St. Domingo to Patagonia; _Hylorhina_ (1 +sp.), Chiloe. The Australian genera are:--_Myxophyes_ (1 sp.), Queensland; +_Platyplectrum_ (2 sp.), Queensland and West Australia; _Neobatrachus_ (1 +sp.), South Australia; _Limnodynastes_ (7 sp.), and _Crinia_ (11 sp.), +Australia and Tasmania. The {421}Oriental genera are:--_Dicroglossus_ (1 +sp.), Western Himalayas; _Oxyglossus_ (2 sp.), Siam to Java, Philippines +and China; _Hoplobatrachus_ (1 sp.), Ceylon; _Phrynoglossus_ (1 sp.), Siam. +The Ethiopian genera are:--_Phrynobatrachus_ (1 sp.), _Stenorhynchus_ (1 +sp.), both from Natal. + + +FAMILY 20.--DISCOGLOSSIDÆ, (14 Genera, 18 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2 -- -- |-- -- -- --|1. 2. 3. 4 |-- 2. 3 -- |-- 2. 3. 4 |1. 2 -- -- + | | | | | + +The Discoglossidæ, or Frogs with a dilated sacrum, are remarkable for the +number of generic forms scattered over a large part of the globe, being +only absent from the Nearctic and the northern half of the Neotropical +regions, and also from Hindostan and East Africa. The genera are:-- + +_Chiroleptes_ (4 sp.), Australia; _Calyplocephalus_ (1 sp.), allied to the +preceding, from Chili; _Cryptotis_ (1 sp.), Australia; _Asterophys_ (2 +sp.), New Guinea and Aru Islands; _Xenophrys_ (1 sp.), Eastern Himalayas; +_Megalophrys_ (2 sp.), Ceylon and the Malay Islands; _Nannophrys_ (1 sp.), +Ceylon; _Pelodytes_ (1 sp.), France only; _Leptobrachium_ (1 sp.), Java; +_Discoglossus_ (1 sp.), Vienna to Algiers; _Laprissa_ (1 sp.), _Latonia_ (1 +sp.), Palæarctic region; _Arthroleptis_ (2 sp.), West Africa and the Cape; +_Grypiscus_ (1 sp.), South Brazil. + + +FAMILY 21.--PIPIDÆ. (1 Genus, 1 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +{422}The Pipidæ are toads without a tongue or maxillary teeth, and with +enormously dilated sacrum. The only species of _Pipa_ is a native of +Guiana. + + +FAMILY 22.--DACTYLETHRIDÆ. (1 Genus, 2 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|1. 2. 3 -- |-- -- -- --|-- -- -- -- + | | | | | + +The Dactylethridæ are Toads with maxillary teeth but no tongue, and with +enormously dilated sacrum. The species of _Dactylethra_ are natives of +West, East, and South Africa. + + +_General Remarks on the Distribution of the Amphibia._ + +The Amphibia, as here enumerated, consist of 22 families, 152 genera, and +nearly 700 species. Many of the families have a very limited range, only +two (Ranidæ and Polypedatidæ) being nearly universal; five more extend each +into five regions, while no less than thirteen of the families are confined +to one, two, or three regions each. By far the richest region is the +Neotropical, possessing 16 families (four of them peculiar) and about 50 +peculiar or very characteristic genera. Next comes the Australian, with 11 +families (one of which is peculiar) and 16 peculiar genera. The Nearctic +region has no less than 9 of the families (two of them peculiar to it) and +15 peculiar genera, 13 of which are tailed Batrachians which have here +their metropolis. The other three regions have 9 families each; the +Palæarctic has no peculiar family but no less than 15 peculiar genera; the +Ethiopian 1 family and 12 genera peculiar to it; and the Oriental, 19 +genera but no family confined to it. + +It is evident, therefore, that each of the regions is well characterised by +its peculiar forms of Amphibia, there being only a few genera, such as +_Hyla_, _Rana_, and _Bufo_ which have a wide range. The connection of the +Australian and Neotropical {423}regions is well shown in this group, by the +Phryniscidæ, Hylidæ, and Discoglossidæ, which present allied forms in both; +as well as by the genus _Liopelma_ of New Zealand, allied to the +Bombinatoridæ of South America, and the absence of the otherwise +cosmopolitan genus _Rana_ from both continents. The affinity of the +Nearctic and Palæarctic regions is shown by the Proteidæ, which are +confined to them, as well as by the genus _Triton_ and almost the whole of +the extensive family of the Salamandridæ. The other regions are also well +differentiated, and there is no sign of a special Ethiopian Amphibian fauna +extending over the peninsula of India, or of the Oriental and Palæarctic +regions merging into each other, except by means of genera of universal +distribution. + +_Fossil Amphibia._--The extinct Labyrinthodontia form a separate order, +which existed from the Carboniferous to the Triassic period. No other +remains of this class are found till we reach the Tertiary formation, when +Newts and Salamanders as well as Frogs and Toads occur, most frequently in +the Miocene deposits. The most remarkable is the _Andrias scheuchzeri_ from +the Miocene of Oeningen, which is allied to _Sieboldia maxima_ the great +salamander of Japan. + + + + +{424}CHAPTER XX. + +THE DISTRIBUTION OF THE FAMILIES OF FISHES, WITH THE RANGE OF SUCH GENERA +AS INHABIT FRESH WATER. + + + +SUB-CLASS I.--TELEOSTEI. + +_Order I.--ACANTHOPTERYGII._ + +FAMILY 1.--GASTEROSTEIDÆ. (1 Genus, 11 Species.) + +"Fresh-water or marine scaleless fishes, with elongate compressed bodies +and with isolated spines before the dorsal fin." + +DISTRIBUTION.--Palæarctic and Nearctic regions. + +The species of _Gasterosteus_, commonly called Sticklebacks, are found in +rivers, lakes, estuaries, and seas, as far south as Italy and Ohio. Four +species occur in Britain. + + +FAMILY 2.--BERYCIDÆ. (10 Genera, 55 Species.) + +"Marine fishes, with elevated compressed bodies covered with toothed +scales, and large eyes." + +DISTRIBUTION.--Tropical and temperate seas of both hemispheres. + +Their northern limit is the Mediterranean and Japan. Most abundant in the +Malayan seas. + + +{425}FAMILY 3.--PERCIDÆ. (61 Genera, 476 Species.) + +"Marine or fresh-water carnivorous fishes, with oblong bodies covered with +toothed scales." + +DISTRIBUTION.--Seas, rivers and lakes, of all regions. + +The genera which inhabit fresh-waters are the following:-- + +_Perca_ (3 sp.), inhabits the Nearctic and Palæarctic regions as far south +as Ohio and Switzerland; one species, the common perch, is British. +_Percichthys_ (5 sp.), Chili and Patagonia, with one species in Java; +_Paralabrax_ (2 sp.), California; _Labrax_ (8 sp.), six species are marine, +inhabiting the shores of Europe and North America, one being British, two +species inhabit the rivers of the northern United States; _Lates_ (2 sp.), +Nile and large rivers of India and China; _Acerina_ (3 sp.), Europe, from +England to Russia and Siberia; _Percarina_ (1 sp.), River Dniester; +_Lucioperca_ (6 sp.), North America and Europe; _Pileoma_ (2 sp.), North +America, Texas to Lake Erie; _Boleosoma_ (3 sp.), Texas to Lake Superior; +_Aspro_ (2 sp.), Central Europe; _Huro_ (1 sp.), Lake Huron; _Percilia_, (1 +sp.), Rio de Maypu in Chili; _Centrarchus_ (10 sp.), North America and +Cuba; _Bryttus_ (8 sp.), South Carolina to Texas; _Pomotis_ (8 sp.), North +America, Lake Erie to Texas. + +Of the exclusively marine genera a species of _Polyprion_ and one of +_Serranus_ are British. The latter genus has nearly 150 species spread over +the globe, but is most abundant in the Tropics. _Mesoprion_ is another +extensive genus confined to the Tropics. _Apogon_ abounds from the Red Sea +to the Pacific, but has one species in the Mediterranean and one in the +coast of Brazil. + + +FAMILY 4.--APHREDODERIDÆ. (1 Genus, 1 Species.) + +"Fresh-water fish, with oblong body covered with toothed scales, and wide +cleft mouth." + +DISTRIBUTION.--Atlantic States of North America. + + +{426}FAMILY 5.--PRISTIPOMATIDÆ. (25 Genera, 206 Species.) + +"Marine carnivorous fishes, with compressed oblong bodies, and without +molar or cutting teeth." + +DISTRIBUTION.--Seas of temperate and tropical regions, a few only entering +fresh water. + +Of the more extensive genera, nine, comprising more than half the species, +are confined to the Indian and Australian seas, while only one large genus +(_Hæmulon_) is found in the Atlantic on the coast of Tropical America. The +extensive Pacific genus, _Diagramma_, has one species in the Mediterranean. +One genus is confined to the Macquarie River in Australia. A species of +_Dentex_ has occurred on the English coast, and this seems to be the +extreme northern range of the family, which does not regularly extend +beyond the coast of Portugal, and in the East to Japan. Australia seems to +form the southern limit. + + +FAMILY 6.--MULLIDÆ. (5 Genera, 34 Species.) + +"Marine fishes, with elongate slightly compressed bodies covered with large +scales, and two dorsal fins at a distance from each other." + +DISTRIBUTION.--All tropical seas, except the West Coast of America, +extending into temperate regions as far as the Baltic, Japan, and New +Zealand. + +Two species of _Mullus_ (Mullets) are British, and these are the only +European fish belonging to the family. + + +FAMILY 7.--SPARIDÆ. (22 Genera, 117 Species.) + +"Herbivorous or carnivorous marine fishes, with oblong compressed bodies +covered with minutely serrated scales, and with one dorsal fin." + +DISTRIBUTION.--Seas of temperate and tropical regions, a few entering +rivers. + +{427}_Cantharus_, _Pagellus_, and _Chrysophrys_, have occurred on the +English Coast. _Haplodactylus_ is confined to the West Coast of South +America, and Australia; _Sargus_ to the temperate and warm parts of the +Atlantic and the shores of East Africa; _Pagellus_ to the western coasts of +Europe and Africa. + +The other large genera have a wider distribution. + + +FAMILY 8.--SQUAMIPENNES. (12 Genera, 124 Species.) + +"Carnivorous marine fishes, with compressed and elevated bodies, and scaly +vertical fins." + +DISTRIBUTION.--The seas between the tropics, most abundant in the Oriental +and Australian regions, a few entering rivers or extending beyond the +tropics. + +The extensive genus _Chætodon_ (67 sp.), ranges from the Red Sea to the +Sandwich Islands, and from Japan to Western Australia, while two species +are found in the West Indies. _Holacanthus_ (36 sp.), has a similar +distribution, one species only occurring in the West Indies and on the +coast of South America. Only one genus (_Pomacanthus_), with a single +species, is confined to the West Atlantic. + + +FAMILY 9.--CIRRHITIDÆ. (8 Genera, 34 Species.) + +"Carnivorous marine fishes, with a compressed oblong body, covered with +cycloid scales." + +DISTRIBUTION.--The tropical and south temperate waters of the Indian and +Pacific oceans, from Eastern Africa to Western America. Absent from the +Atlantic. + + +FAMILY 10.--TRIGLIDÆ. (50 Genera, 259 Species.) + +"Carnivorous, mostly marine fishes, with oblong compressed or +subcylindrical bodies, and wide cleft mouths. They live at the bottom of +the water." + +DISTRIBUTION.--All seas, some entering fresh water, and a few inhabiting +exclusively the fresh waters of the Arctic regions. + +{428}They are divided by Dr. Günther into four groups. The Heterolepidina +(comprising 4 genera and 12 species) are confined to the North Pacific. The +Scorpænina (23 genera, 113 species) have an almost universal distribution, +but the genera are each restricted to one or other of the great oceans. +_Sebastes_ has occurred on the English coast. The Cottina (28 genera, 110 +species) have also a universal distribution; the numerous species of +_Cottus_ are found either in the seas or fresh waters of Europe and North +America; four species are British, as well as seven species of the +wide-spread genus _Trigla_. _Ptyonotus_ (1 sp.) is confined to Lake +Ontario. The Cataphracti (5 genera, 23 species) have also a wide range; one +genus, _Agonus_, is found in the British seas, and also in Kamschatka and +on the coast of Chili. _Peristethus_ is also British. + + +FAMILY 11.--TRACHINIDÆ (24 Genera, 90 Species.) + +"Carnivorous marine fishes, with elongate bodies, living at the bottom, +near the shore." + +DISTRIBUTION.--Almost or quite universal. + +_Trachinus_ is a British genus. A species of _Aphritis_ inhabits the fresh +waters of Tasmania, while its two allies are found on the coasts of +Patagonia. + + +FAMILY 12. SCIÆNIDÆ. (13 Genera, 102 Species.) + +"Marine or fresh-water fishes, with compressed and rather elongate bodies, +covered with toothed scales." + +DISTRIBUTION.--Temperate and tropical regions, but absent from Australia. + +_Larimus_ is found in the Atlantic, and in African and American rivers. +_Corvina_, _Sciæna_, and _Otilothus_ are also marine and fresh-water, both +in the Atlantic and Pacific. The other genera are of small extent and more +restricted range. _Umbrina_ and _Sciæna_ have occurred in British seas. + + +{429}FAMILY 13.--POLYNEMIDÆ. (3 Genera, 23 Species.) + +"Marine or fresh-water fishes, with compressed oblong bodies and entire or +ciliated scales." + +DISTRIBUTION.--Tropical seas and rivers of both the great oceans, but most +abundant in the Pacific. + + +FAMILY 14.--SPHYRENIDÆ. (1 Genus, 15 Species.) + +"Carnivorous marine fishes, with elongate sub-cylindrical bodies covered +with small cycloid scales." + +DISTRIBUTION.--The warm and tropical seas of the globe. + + +FAMILY 15.--TRICHIURIDÆ. (7 Genera, 18 Species.) + +"Marine fishes, with elongate compressed bodies covered with minute scales +or naked." + +DISTRIBUTION.--All the tropical and sub-tropical seas. + + +FAMILY 16.--SCOMBRIDÆ. (20 Genera, 108 Species.) + +"Marine fishes, with elongate compressed bodies, scaled or naked." + +DISTRIBUTION.--All the temperate and tropical oceans. Mostly inhabiting the +open seas. + +_Scomber_, (the Mackerel) _Thynnus_, _Naucrates_, _Zeus_, _Centrolophus_, +_Brama_, and _Lampris_, are genera which have occurred in the British seas. + + +FAMILY 17.--CARANGIDÆ. (27 Genera, 171 Species.) + +"Marine fishes, with compressed oblong or elevated bodies covered with +small scales or naked." + +DISTRIBUTION.--All temperate and tropical seas; some species occur in both +the great oceans, ranging from New York to Australia. + +_Trachurus_ and _Capros_ are genera which occur in British seas. + + +{430}FAMILY 18.--XIPHIIDÆ. (2 Genera, 8 Species.) + +"Marine fishes, with elongate compressed body and a produced sword-shaped +upper jaw." + +DISTRIBUTION.--Mediterranean, and open seas between or near the Tropics. + +_Xiphias_ (the Sword-fish) has occurred on the English coast. + + +FAMILY 19.--GOBIIDÆ. (24 Genera, 294 Species.) + +"Carnivorous fishes, with elongate low, naked, or scaly bodies, living at +the bottom of the shallow seas or fresh waters of temperate or tropical +regions. Individuals of the same species often differ in inhabiting +exclusively fresh or salt water." + +DISTRIBUTION.--All temperate and tropical regions, from Scotland and Japan +to New Zealand. Species of _Gobius_, _Latrunculus_, and _Callionymus_ occur +in Britain. Several genera are confined to the East Indian seas and rivers, +but none seem peculiar to America. The genus _Periopthalmus_ consists of +the curious, large-headed, projecting-eyed fishes, so abundant on the muddy +shores of African and Eastern tidal rivers, and which seem to spend most of +their time out of water, hunting after insects, &c. + + +FAMILY 20.--DISCOBOLI. (2 Genera, 11 Species.) + +"Carnivorous fishes, with oblong naked or tubercular bodies, living at the +bottom of shallow seas, and attaching themselves to rocks by means of a +ventral disc." + +DISTRIBUTION.--All northern seas, as far south as Belgium, England, and San +Francisco. + +Species of both genera (_Cyclopterus_ and _Liparis_) occur in British seas. + + +{431}FAMILY 21.--OXUDERCIDÆ. (1 Genus, 1 Species.) + +"A marine fish, with an elongate sub-cylindrical body and no ventral fins." + +DISTRIBUTION.--Macao, China. + + +FAMILY 22.--BATRACHIDÆ. (3 Genera, 12 Species.) + +"Marine fishes, with sub-cylindrical body and broad depressed head." + +DISTRIBUTION.--The coasts of nearly all tropical and south temperate +regions, ranging from New York and Portugal to Chili and Tasmania. + + +FAMILY 23.--PEDICULATI. (8 Genera, 40 Species.) + +"Marine carnivorous fishes, with very large heads and without scales." + +DISTRIBUTION.--Seas of all temperate and tropical regions, extending south +to New Zealand and north to Greenland. + +A species of _Lophius_ (the Fishing-frog or Sea-Devil) is found in British +seas. The genus _Antennarius_, comprising two-thirds of the species, is +wholly tropical. + + +FAMILY 24.--BLENNIDÆ. (33 Genera, 201 Species.) + +"Carnivorous fishes, with long sub-cylindrical naked bodies, living at the +bottom of shallow water in seas, or tidal rivers." + +DISTRIBUTION.--All seas from the Arctic regions to New Zealand, Chili, and +the Cape of Good Hope. + +Species of _Anarrhichas_, _Blennius_, _Blenniops_, _Centronotus_ and +_Zoarces_ occur in British seas. _Chasmodes_ (3 sp.) is confined to the +Atlantic coasts of Temperate North America; _Petroscirtes_ (26 sp.) to the +tropical parts of the Indian and Pacific Oceans; and _Stichæus_ (9 sp.) to +the Arctic Seas. + + +{432}FAMILY 25.--ACANTHOCLINIDÆ. (1 Genus, 1 Species.) + +"A carnivorous marine fish, with long flat body and very long dorsal fin." + +DISTRIBUTION.--Coasts of New Zealand. + + +FAMILY 26.--COMEPHORIDÆ. (1 Genus, 1 Species.) + +"An elongate, naked, large-headed fish, with two dorsal fins." + +DISTRIBUTION.--Lake Baikal. + +Dr. Günther remarks, that this fish approaches the Scombrina (Mackerel) in +several characters. These are exclusively marine fishes, while Lake Baikal +is fresh-water, and is situated among mountains, at an elevation of nearly +2000 feet, and more than a thousand miles from the ocean! + + +FAMILY 27.--TRACHYPTERIDÆ. (3 Genera, 16 Species.) + +"Deep sea fishes, with elongate, much compressed, naked bodies." + +DISTRIBUTION.--Europe, East Indies, West Coast of South America, New +Zealand. Dr. Günther remarks, that little is known of these fishes, from +their being so seldom thrown on shore, and then rapidly decomposing. The +Ribbon-fish (_Regalecus banksii_) has occurred frequently on our shores. +They have soft bones and muscles, small mouths, and weak dentition. + + +FAMILY 28.--LOPHOTIDÆ. (1 Genus, 1 Species.) + +"A marine fish, with elongate compressed naked body, and high crested +head." + +DISTRIBUTION.--Mediterranean Sea and Japan. + + +{433}FAMILY 29.--TEUTHIDIDÆ. (1 Genus, 29 Species.) + +"Marine, herbivorous fishes, with compressed, oblong, small-scaled bodies." + +DISTRIBUTION.--Eastern tropical seas, from Bourbon and the Red Sea to the +Marianne and Fiji Islands. + + +FAMILY 30.--ACRONURIDÆ. (5 Genera, 64 Species.) + +"Marine, herbivorous fishes, with compressed, minutely-scaled bodies." + +DISTRIBUTION.--All tropical seas, but most abundant in the Malay region, +and extending to Japan and New Zealand. + + +FAMILY 31.--HOPLEGNATHIDÆ. (1 Genus, 3 Species.) + +"Marine fishes, with compressed elevated bodies, covered with very small +toothed scales." + +DISTRIBUTION.--Seas of Australia, China, and Japan. + + +FAMILY 32.--MALACANTHIDÆ. (1 Genus, 3 Species.) + +"Marine fishes, with elongate bodies covered with very small scales, and +with very long dorsal and anal fins." + +DISTRIBUTION.--Atlantic coasts of Tropical America, Mauritius, and New +Guinea. + + +FAMILY 33.--NANDIDÆ. (6 Genera, 14 Species.) + +"Marine or fresh-water carnivorous fishes, with oblong, compressed, scaly +bodies." + +DISTRIBUTION.--From the Red Sea to the coasts of China and Australia; and +the fresh waters of the Neotropical and Oriental regions. _Badis_, +_Nandus_, and _Catopra_ inhabit the {434}rivers of India and the Malay +Islands; _Acharnes_ the rivers of British Guiana. + + +FAMILY 34.--POLYCENTRIDÆ. (2 Genera, 3 Species.) + +"Fresh-water carnivorous fishes, with compressed elevated scaly bodies, and +many-spined dorsal and anal fins." + +DISTRIBUTION.--Rivers of Tropical America. + + +FAMILY 35.--LABYRINTHICI. (9 Genera, 25 Species.) + +"Fresh-water fishes, with compressed oblong bodies, and capable of living +for some time out of water or in dried mud." + +DISTRIBUTION.--Fresh waters of South Africa and the East Indies from the +Mauritius to China, the Philippines, Celebes, and Amboyna. + + +FAMILY 36.--LUCIOCEPHALIDÆ. (1 Genus, 1 Species.) + +"Fresh-water fish, with elongate scaled body, and a dilated branchial +membrane." + +DISTRIBUTION.--Rivers of Borneo, Biliton, and Banca. + + +FAMILY 37.--ATHERINIDÆ. (3 Genera, 39 Species.) + +"Marine or fresh-water carnivorous fishes, with subcylindrical scaled +bodies, and feeble dentition." + +DISTRIBUTION.--All temperate and tropical seas, from Scotland and New York +to the Straits of Magellan and Tasmania. + +_Atherina presbyter_ occurs in British seas. Species of _Atherina_ and +_Atherinichthis_ are found in fresh-water lakes and rivers in Europe, +America, and Australia. + + +{435}FAMILY 38.--MUGILIDÆ. (3 Genera, 78 Species.) + +"Fresh-water and marine fishes, with oblong compressed bodies, cycloid +scales, and small mouths, often without teeth." + +DISTRIBUTION.--Coasts and fresh waters of all temperate and tropical +regions. + +_Mugil_ (66 sp.) is mostly marine, and is very widely distributed; several +species (Grey Mullets) occur on the British coasts. _Agonostoma_ (9 sp.) is +confined to the fresh waters of the West Indies, Central America, New +Zealand, Australia, Celebes, and the Comoro Islands. _Myxus_ (3 sp.) is +marine, and occurs both in the Atlantic and Pacific. + + +FAMILY 39.--OPHIOCEPHALIDÆ. (2 Genera, 26 Species.) + +"Fresh-water fishes, with elongate subcylindrical scaled bodies; often +leaving the water for a considerable time." + +DISTRIBUTION.--Rivers of the Oriental region:--India, Ceylon, China, Malay +Islands to Philippines and Borneo. + + +FAMILY 40.--TRICHONOTIDÆ. (2 Genera, 2 Species.) + +"Marine carnivorous fishes, with elongate subcylindrical bodies, cycloid +scales, and eyes directed upwards." + +DISTRIBUTION.--Coasts of Celebes, Ceram, and New Zealand. + + +FAMILY 41.--CEPOLIDÆ. (1 Genus, 7 Species.) + +"Marine fishes, with very long, compressed, band-like bodies, covered with +small cycloid scales." + +DISTRIBUTION.--Temperate seas of Western Europe and Eastern Asia, and one +species in the Malayan Seas. + +_Cepola rubescens_ (the Band fish) ranges from Scotland to the +Mediterranean. All the other species but one are from Japan. + + +{436}FAMILY 42.--GOBIESOCIDÆ. (9 Genera, 21 Species.) + +"Carnivorous marine fishes, elongate, anteriorly depressed and scaleless, +with dorsal fin on the tail." + +DISTRIBUTION.--Temperate and tropical seas; Scandinavia to the Cape, +California to Chili, West Indies, Red Sea, Australia, New Zealand, and Fiji +Islands. + +Three species of _Lepadogaster_ have occurred in the English Channel. + + +FAMILY 43.--PSYCHROLUTIDÆ. (1 Genus, 1 Species.) + +"A large-headed, elongate, naked marine fish, with small teeth, and dorsal +fin on the tail." + +DISTRIBUTION.--West Coast of North America (Vancouver's Island.) + +FAMILY 44.--CENTRISCIDÆ. (2 Genera, 7 Species.) + +"Marine fishes, with compressed, oblong or elevated bodies, elongate +tubular mouth and no teeth." + +DISTRIBUTION.--West Coast of Europe and Africa, Mediterranean, Indian Ocean +to Java, Philippines, and Japan. + +A species of _Centriscus_ has occurred on the South Coast of England, and +another species is found both at Madeira and Japan. + + +FAMILY 45.--FISTULARIDÆ. (2 Genera, 4 Species.) + +"Marine fishes, very elongate, with long tubular mouth and small teeth." + +DISTRIBUTION.--Tropical seas, both in the Atlantic and Indian Ocean, and as +far east as the New Hebrides. + + +{437}FAMILY 46.--MASTACEMBELIDÆ. (2 Genera, 9 Species.) + +"Fresh-water fishes, with eel-like bodies and very long dorsal fin." + +DISTRIBUTION.--Rivers of the Oriental region, one species from Ceram (?). + + +FAMILY 47.--NOTACANTHI. (1 Genus, 5 Species.) + +"Marine fishes, with elongate bodies covered with very small scales, and +snout protruding beyond the mouth." + +DISTRIBUTION.--Greenland, Mediterranean, and West Australia. + + +_Order II.--ACANTHOPTERYGII PHARYNGOGNATHI._ + + +FAMILY 48.--POMACENTRIDÆ. (8 Genera, 143 Species.) + +"Marine fishes, with short compressed bodies covered with toothed scales, +and with feeble dentition." + +DISTRIBUTION.--Tropical parts of Pacific and Indian Ocean, less numerous in +Tropical Atlantic, a few reaching the Mediterranean, Japan, and South +Australia. _Pomacentrus_, _Glyphidodon_, and _Heliastes_ are Atlantic +genera. + + +FAMILY 49.--LABRIDÆ. (46 Genera, 396 Species.) + +"Herbivorous or carnivorous marine fishes, with elongate bodies covered +with cycloid scales, and teeth adapted for crushing the shells of +mollusca." + +DISTRIBUTION.--Temperate and tropical regions of all parts of the globe. + +The genera _Labrus_, _Crenilabrus_, _Ctenolabrus_, _Acantholabrus_, +_Centrolabrus_, and _Coris_, have occurred in British seas, and all of +{438}these, except the last, are confined to the Mediterranean and the +Atlantic as far as Madeira. Eight other genera are characteristic of the +Atlantic, most of them being West Indian, but one from the coasts of North +America. Seven genera are common to all the great oceans; the remainder +being confined to the Indian and Pacific Oceans, ranging from Japan to New +Zealand, but being far more abundant between the Tropics. + + +FAMILY 50.--EMBROTOCIDÆ. (2 Genera, 17 Species.) + +"Marine viviparous fishes, with compressed elevated bodies covered with +cycloid scales, and with small teeth." + +DISTRIBUTION.--Pacific Ocean from Japan and California northwards. One +species enters the fresh waters of California. + + +FAMILY 51.--GERRIDÆ. (1 Genus, 28 Species.) + +"Marine fishes, with compressed oblong bodies covered with minutely +serrated scales, and with small teeth." + +DISTRIBUTION.--Tropical seas; ranging south as far as the Cape of Good Hope +and Australia, and north to Japan and (one species) to New Jersey, U.S. + + +FAMILY 52.--CHROMIDÆ. (19 Genera, 100 Species.) + +"Fresh-water herbivorous or carnivorous fishes, with elevated or elongate +scaly bodies, and small teeth." + +DISTRIBUTION.--The Oriental, Ethiopian, and Neotropical regions. + +_Eutroplus_ (2 sp.) is from the rivers of Southern India and Ceylon; +_Chromis_ (15 sp.), _Sarotherodon_ (2 sp.), and _Hemichromis_ (4 sp.), are +from the rivers and lakes of Africa, extending to the Sahara and Palestine. +The remaining 15 genera are American, and several of them have a restricted +distribution. _Acara_ (17 sp.) inhabits Tropical South America and the +Antilies; _Theraps_ (1 sp.), Guatemala; _Heros_ (26 sp.), Texas and +{439}Mexico to La Plata; _Mesonauta_ (1 sp.), Brazil; _Petenia_ (1 sp.), +Lake Peten, Guatemala; _Uaru_ (2 sp.), Brazil; _Hygrogonus_ (1 sp.), +Brazil; _Cichla_ (4 sp.), Equatorial America; _Crenicichla_ (9 sp.), Brazil +and Guiana; _Chætobranchus_ (3 sp.), Brazil and Guiana; _Mesops_ (2 sp.), +Brazil; _Satanoperca_ (7 sp.), Amazon Valley and Guiana; _Geophagus_ (1 +sp.), North Brazil and Guiana; _Symphysodon_ (1 sp.), Lower Amazon; +_Pterophyllum_ (1 sp.), Lower Amazon. + + +_Order III.--ANACANTHINI._ + + +FAMILY 53.--GADOPSIDÆ. (1 Genus, 1 Species.) + +"Fresh-water fish, with rather elongate body covered with very small +scales, the upper jaw overhanging the lower, forming an obtuse snout." + +DISTRIBUTION.--Rivers of Australia and Tasmania. + + +FAMILY 53_a_.--LYCODIDÆ. (3 Genera, 14 Species.) + +"Marine fishes, with elongate bodies, and the dorsal united with the anal +fin." + +DISTRIBUTION.--Arctic seas of America and Greenland, and Antarctic seas +about the Falkland Islands and Chiloe Island. + + +FAMILY 54.--GADIDÆ. (21 Genera, 58 Species.) + +"Marine fishes, with more or less elongate bodies covered with small smooth +scales." + +DISTRIBUTION.--Cold and temperate regions of both hemispheres; in the North +extending as far south as the Mediterranean, Canary Islands, New York and +Japan (and one species to the Philippines and Bay of Bengal), and in the +South to Chili and New Zealand. + +_Gadus_ (Cod), _Merluccius_ (Hake), _Phycis_, _Lota_, _Molva_, _Couchia_, +_Motella_, and _Raniceps_, are British. _Lota_ inhabits fresh waters. + + +{440}FAMILY 55.--OPHIDIIDÆ. (16 Genera, 43 Species.) + +"Marine fishes, with more or less elongate bodies, the dorsal and anal fins +united, and the ventral fins rudimentary or absent." + +DISTRIBUTION.--Almost universal; from Greenland to New Zealand, but most +abundant in the Tropics. + +_Ophidium_ and _Ammodytes_ occur in British seas; _Lucifuga_ inhabits +subterranean fresh waters in Cuba. + + +FAMILY 56.--MACROURIDÆ. (3 Genera, 21 Species.) + +"Marine fishes, with the body terminating in a long, compressed tapering +tail, and covered with spiny, keeled or striated scales." + +DISTRIBUTION.--North Atlantic from Greenland to Madeira and the Canary +Islands, Mediterranean, Japanese and Australian seas. + +None of these fishes have occurred in the British seas. + + +FAMILY 57.--ATELEOPODIDÆ. (1 Genus, 1 Species.) + +"Marine fishes, with the naked body terminating in a long compressed, +tapering tail." + +DISTRIBUTION.--Japan. + + +FAMILY 58.--PLEURONECTIDÆ. (34 Genera, 185 Species.) + +"Marine carnivorous fishes, with strongly compressed flat bodies, one side +of which is colourless, and eyes unsymmetrically placed, both on the +coloured side. They inhabit the sandy bottoms of shallow seas, and often +ascend rivers." + +DISTRIBUTION.--Universal, on Arctic, Temperate, and Tropical coasts. + +{441}Seven genera occur in British seas, viz.: _Hippoglossus_, +_Hippoglossoides_, _Rhombus_, _Phrynorhombus_, _Arnoglossus_, +_Pleuronectes_ (Turbot), and _Solea_ (Sole). There are 13 genera in the +Atlantic and 23 in the Pacific, 4 being common to both; and 2 found only in +the Mediterranean. A Pacific genus, _Synaptura_, has one species in the +Mediterranean. + + +_Order IV.--PHYSOSTOMI._ + + +FAMILY 59.--SILURIDÆ. (114 Genera, 547 Species.) + +"Fresh-water or marine, scaleless fishes, often with bony shields, and the +head always furnished with barbels." + +DISTRIBUTION.--The fresh waters of all the temperate and tropical regions, +those which enter the salt water keeping near the coast. + +This extensive family is divided by Dr. Günther into eight sub-families and +seventeen groups, the distribution of which is as follows:-- + +Sub-family 1 (SILURIDÆ HOMALOPTERÆ) is confined to the Old World. It +consists of three groups: Clarina (2 genera, _Clarias_ and +_Heterobranchus_) ranges over the whole area of the Ethiopian and Oriental +regions, to which it appears to be strictly confined; Plotosina (3 genera, +_Plotosus_, _Copidoglanis_, and _Cnidoglanis_) ranges from the eastern +coasts of Africa to Japan, Polynesia, and Australia, in seas and rivers; +Chacina (1 genus, _Chaca_) ranges from India to Borneo. + +Sub-family 2 (SILURIDÆ HETEROPTERÆ) is also confined to the Old World; it +consists of one group,--Silurina, containing 19 genera, +viz.:--_Saccobranchus_ (4 sp.), India to Cochin China and Ceylon; _Silurus_ +(5 sp.), Palæarctic region from Central Europe to Japan, China, and +Afghanistan, and a species in Cochin China; _Silurichthys_ (3 sp.), +Cashmere, Java, and Borneo; _Wallago_ (2 sp.), Hindostan, Sumatra, and +Borneo; _Belodontichthys_ (1 sp.), Sumatra and Borneo; _Eutropiichthys_ (1 +sp.), Bengal; _Cryptopterus_ {442}(15 sp.), Java, Sumatra, and Borneo, with +a species in the Ganges, in Siam, and (?) in Amboyna; _Callichrous_ (10 +sp.), Afghanistan to Borneo and Java; _Schilbe_ (5 sp.), Tropical Africa; +_Eutropius_ (6 sp.), Tropical Africa and Central India; _Hemisilurus_ (2 +sp.), Java and Sumatra; _Siluranodon_ (1 sp.), Nile; _Ailia_ (2 sp.), +Bengal; _Schilbichthys_ (1 sp.), Bengal; _Laïs_ (1 sp.), Java, Sumatra, +Borneo; _Pseudeutropius_ (6 sp.), India and Sumatra; _Pangasius_ (7 sp.), +Ganges, Sumatra, Java, Borneo; _Helicophagus_ (2 sp.), Sumatra; _Silondia_ +(1 sp.), Ganges. + +Sub-family 3 (SILURIDÆ ANOMALOPTERÆ) is confined to Equatorial America; it +consists of the group Hypopthalmina, containing 2 genera: _Helogenes_ (1 +sp.), _Hypopthalmus_ (4 sp.), from the country north of the Amazon, +Surinam, and the Rio Negro. + +Sub-family 4 (SILURIDÆ PROTEROPTERÆ) ranges over all the tropical and most +of the temperate parts of the globe, except Europe and Australia. It +consists of four groups: Bagrina (16 genera), ranging over most of the Old +World and North America; Pimelodina (15 genera), confined to Tropical +America, except one genus which is African; Ariina (10 genera), all +Tropical regions; and Bagarina (3 genera), Oriental region. The +distribution of the genera is as follows:-- + +_Bagrus_ (2 sp.), Nile; _Chrysichthys_ (5 sp.), Tropical Africa; _Clarotes_ +(1 sp.), Upper Nile; _Macrones_ (19 sp.), India, Ceylon to Borneo, and one +species in Asia Minor; _Pseudobagrus_ (4 sp.), Japan, China, and Cochin +China; _Liocassis_ (5 sp.), Japan, China, Java, Sumatra, and Borneo; +_Bagroides_ (3 sp.), Sumatra and Borneo; _Bagrichthys_ (1 sp.), Sumatra and +Borneo; _Rita_ (5 sp.), Continental India and Manilla; _Acrochordonichthys_ +(6 sp.), Java and Sumatra; _Akysis_ (3 sp.), Java and Sumatra; _Olyra_ (1 +sp.), Khasya; _Branchiosteus_ (1 sp.), Khasya; _Amiurus_ (13 sp.), Nearctic +region to Guatemala and China; _Hopladelus_ (1 sp.), North America; +_Noturus_ (4 sp.), North America; _Sorubim_ (1 sp.), Amazon; _Platystoma_ +(11 sp.), Tropical South America; _Hemisorubim_ (1 sp.) Rio Negro, Brazil; +_Platistomatichthys_ (1 sp.), Rio Branco, Brazil; _Phractocephalus_ (1 +sp.), Amazon; _Piramutana_ (2 sp.), Equatorial America; _Platynematichthys_ +{443}(1 sp.), northern and southern tributaries of Amazon; _Piratinga_ (3 +sp.), Amazon Valley; _Sciades_ (2 sp.), Amazon; _Pimelodus_ (42 sp.), +Mexico to La Plata, single aberrant species from West Africa, Java and the +Sandwich Islands; _Pirinampus_ (1 sp.), Brazil; _Conorhynchus_ (1 sp.), +Brazil; _Notoglanis_ (1 sp.), Madeira, Amazon Valley; _Callophysus_ (3 +sp.), Tropical South America; _Auchenaspis_ (1 sp.), Tropical Africa; +_Arius_ (68 sp.), all Tropical regions; _Galeichthys_ (1 sp.), Cape of Good +Hope; _Genidens_ (1 sp.), Brazil; _Hemipimelodus_ (3 sp.), India, Sumatra, +and Borneo; _Ketingus_ (1 sp.), Sunda Islands; _Ælurichthys_ (4 sp.), +Eastern United States to Guiana; _Paradiplomystax_ (1 sp.), Brazil; +_Diplomystax_ (1 sp.), Chili; _Osteogeniosus_ (3 sp.), India to Java; +_Batrachocephalus_ (1 sp.), Java and Sumatra; _Bagarius_ (1 sp.), India to +Java; _Euclyptosternum_ (1 sp.), India; _Glyptosternum_ (8 sp.), Himalayas, +Central India, Java, and Sumatra; _Hara_ (3 sp.), Continental India; +_Amblyceps_ (3 sp.), Continental India. + +Sub-family 5 (SILURIDÆ STENOBRANCHIÆ) is confined to South America and +Africa, with one genus and species in the Ganges. It consists of three +groups: Doradina (12 genera), South America and Africa; Rhinoglanina (3 +genera), Central Africa and the Ganges; Malapterurina (1 genus), Tropical +Africa. The distribution of the genera is as follows:-- + +_Ageniosus_ (4 sp.), Surinam to La Plata; _Tetranematichthys_ (1 sp.), +Central Brazil, Rio Guaporé; _Euanemus_ (1 sp.), Surinam and Brazil; +_Auchenipterus_ (9 sp.), Equatorial America; _Centromochlus_ (2 sp.), +Equatorial America; _Trachelyopterus_ (2 sp.), Equatorial America; +_Cetopsis_ (3 sp.), Brazil; _Asterophysus_ (1 sp.), Rio Negro, North +Brazil; _Doras_ (13 sp.), Tropical South America east of Andes; _Oxydoras_ +(7 sp.), Amazon Valley and Guiana; _Rhinodoras_ (3 sp.), Tropical South +America east of Andes; _Synodontis_ (12 sp.), Tropical Africa; +_Rhinoglanis_ (1 sp.), Upper Nile; _Mochocus_ (1 sp.), Nile; _Callomystax_ +(1 sp.), Nile; _Malapterurus_ (3 sp.), Tropical Africa. + +Sub-family 6 (SILURIDÆ PROTEROPODES) inhabits Tropical America and Northern +India as far as Tenasserim. It consists of two groups: the Hypostomatina +(17 genera), with the same distribution as the sub-family, and the +Aspredinina (3 genera), {444}confined to Equatorial America. The +distribution of the genera is as follows:-- + +_Arges_ (2 sp.), Andes of Peru and Ecuador; _Stygogenes_ (2 sp.), Andes; +_Brontes_ (1 sp.), Andes; _Astroblepus_ (1 sp.), Popayan; _Callichthys_ (11 +sp.), Tropical South America east of Andes, and Trinidad; _Plecostomus_ (15 +sp.), Tropical South America east of Andes, and Trinidad; _Liposarcus_ (3 +sp.), Surinam and Brazil; _Chætostomus_ (25 sp.), Tropical America, +Trinidad, and Porto Rico; _Pterygoplichthys_ (4 sp.), Brazil; _Rhinelepis_ +(1 sp.), Brazil; _Acanthicus_ (2 sp.), Equatorial America; _Loricaria_ (17 +sp.), Tropical South America east of Andes; _Acestra_ (4 sp.), Brazil and +Guiana; _Sisor_ (1 sp.), Northern Bengal; _Erethistes_ (1 sp.), Assam; +_Pseudecheneis_ (1 sp.), Khasya Hills; _Exostoma_ (2 sp.), Assam and +Tenasserim; _Bunocephalus_ (2 sp.), Guiana; _Bunocephalichthys_ (1 sp.), +Rio Branco, North Brazil; _Aspredo_ (6 sp.), Guiana. + +Sub-family 7 (SILURIDÆ OPISTHOPTERÆ) consists of two groups: Nematogenyina +(2 genera), and Trichomycterina (3 genera), and is confined to South +America. The distribution of the genera is as follows:-- + +_Heptapterus_ (2 sp.), South America; _Nematogenys_ (1 sp.), Chili; +_Trichomycterus_ (7 sp.), South America to 15,000 feet elevation; +_Eremophilus_ (1 sp.), Andes of Bogota; _Pariodon_ (1 sp.), Amazon. + +Sub-family 8 (SILURIDÆ BRANCHICOLÆ) is confined to Tropical South America. +It consists of one group, Stegophilina, and 2 genera: _Stegophilus_ (1 +sp.), Brazil; and _Vandellia_ (2 sp.), Amazon Valley. + + +FAMILY 60. CHARACINIDÆ. (47 Genera, 230 Species.) + +"Fresh-water fishes, with scaly bodies and without barbels." + +DISTRIBUTION.--The Neotropical and Ethiopian regions. + +This extensive family is divided by Dr. Günther into 10 groups, viz.: +Erythrinina (5 genera), South America; Curumatina {445}(6 genera), South +America; Citharinina (1 genus), Tropical Africa; Anostomatina (3 genera), +South America; Tetragonopterina (16 genera), South America and Tropical +Africa; Hydrocyonina (9 genera), Tropical America and Tropical Africa; +Distichodontina (1 genus), Tropical Africa; Icthyborina (1 genus), Africa; +Crenuchina (1 genus), Equatorial America; Serrasalmonina (4 genera), South +America. + +The following is the distribution of the genera:-- + +_Macrodon_ (4 sp.), Tropical America; _Erythrinus_ (5 sp.), Brazil and +Guiana; _Lebiasina_ (1 sp.), West Equatorial America; _Pyrrhulina_ (1 sp.), +Guiana; _Corynopoma_ (4 sp.), Trinidad only; _Curimatus_ (15 sp.), Tropical +South America and Trinidad; _Prochilodus_ (12 sp.), South America to the La +Plata; _Cæntropus_ (2 sp.), East Equatorial America; _Hemiodus_ (8 sp.), +Equatorial America east of Andes; _Saccodon_ (1 sp.), Ecuador; _Parodon_ (1 +sp.), Brazil; _Citharinus_ (2 sp.), Tropical Africa; _Anostomus_ (8 sp.), +Tropical America; _Rhytiodus_ (2 sp.), Equatorial America; _Leporinus_ (14 +sp.), South America East of Andes; _Piabucina_ (2 sp.), Guiana; _Alestes_ +(4 sp.), Tropical Africa; _Brachyalestes_ (5 sp.), Tropical Africa; +_Tetragonopterus_ (32 sp.), Tropical America; _Scissor_ (1 sp.), South +America; _Pseudochalceus_ (1 sp.), West Ecuador; _Chirodon_ (2 sp.), Chili; +_Chalceus_ (1 sp.), Guiana; _Brycon_ (10 sp.), South America east of Andes; +_Chalcinopsis_ (4 sp.), Central America and Ecuador; _Bryconops_ (2 sp.), +Tropical America; _Creagrutus_ (1 sp.), Western Ecuador; _Chalcinus_ (4 +sp.), Tropical South America; _Gastropelecus_ (8 sp.), Tropical South +America; _Piabuca_ (2 sp.), Equatorial America; _Agoniates_ (1 sp.), +Guiana; _Anacyrtus_ (7 sp.), Central and South America; _Hystricodon_ (1 +sp.), Equatorial America; _Salminus_ (3 sp.), South America; _Hydrocyon_ (3 +sp.), Tropical Africa; _Sarcodaces_ (1 sp.), West Africa; _Oligosarcus_ (1 +sp.), Brazil; _Xiphoramphus_ (7 sp.), South America east of Andes; +_Xiphostoma_ (5 sp.), Equatorial America east of Andes; _Cynodon_ (3 sp.), +Tropical America East of Andes; _Distichodus_ (7 sp.), Tropical Africa; +_Icthyborus_ (3 sp.), Nile; _Crenuchus_ (1 sp.), Guiana; _Mylesinus_ (1 +sp.), Equatorial America; _Serrasalmo_ (13 sp.), Tropical South America +east of Andes; _Myletes_ (18 sp.), {446}Tropical South America east of +Andes; _Catoprion_ (1. sp.), Brazil and Guiana. + + +FAMILY 61.--HAPLOCHITONIDÆ. (2 Genera, 3 Species.) + +"Fresh-water fishes, with naked or scaly bodies and without barbels." + +DISTRIBUTION.--Temperate South America and South Australia. + +The genera are, _Haplochiton_ (2 sp.), Tierra del Fuego and the Falkland +Islands; _Prototroctes_ (2 sp.), Southern Australia and New Zealand. + + +FAMILY 62.--STERNOPTYCHIDÆ. (6 Genera, 12 Species.) + +"Marine fishes, with very thin deciduous scales or none, and with a row of +phosphorescent spots or organs on the under surface of the body." + +DISTRIBUTION.--Mediterranean and Atlantic. + +These are deep-sea fishes found in the Mediterranean sea, and in the deep +Atlantic from the coasts of Norway to the Azores and the Tropics. + + +FAMILY 63.--SCOPELIDÆ. (11 Genera, 47 Species.) + +"Marine fishes, somewhat resembling the fresh-water Siluridæ." + +DISTRIBUTION.--Almost universal, but most abundant in warm and tropical +seas. + +These are deep-sea fishes, abounding in the Mediterranean and the great +oceans, a few extending north to near Greenland and south to Tasmania. + + +{447}FAMILY 64.--STOMIATIDÆ. (4 Genera, 8 Species.) + +"Small marine fishes, naked or with very fine scales." + +DISTRIBUTION.--The Mediterranean and Atlantic. + +These are deep-sea fishes, ranging from Greenland to beyond the Equator. + + +FAMILY 65.--SALMONIDÆ (15 Genera, 157 Species.) + +"Fresh-water fishes, many species periodically descending to the sea and a +few altogether marine:--Salmon and Trout." + +DISTRIBUTION.--The Palæarctic and Nearctic Regions, and one genus and +species in New Zealand. A considerable number of species are confined to +single lakes or rivers, others have a wide distribution. + +The genera are distributed as follows:-- + +_Salmo_ (83 sp.), rivers and lakes of the Palæarctic and Nearctic Regions, +as far south as Algeria, Asia Minor, the Hindoo-Koosh and Kamschatka, and +to about 38° North Latitude in North America, many of the species +migratory; _Onchorhynchus_ (8 sp.), American and Asiatic rivers entering +the Pacific, as far south as San Francisco and the Amur; _Brachymystax_ (1 +sp.), Siberian rivers, from Lake Baikal and the Atlai Mountains northwards; +_Luciotrutta_ (2 sp.), Caspian Sea and Volga; _Plecoglossus_ (1 sp.), Japan +and Formosa; _Osmerus_ (3 sp.), rivers of temperate Europe and North +America entering the Atlantic, and one species in California; +_Thaleichthys_ (1 sp.), Columbia River, Vancouver's Island; _Hypomesus_ (1 +sp.), coasts of California, Vancouver's Island, and North-eastern Asia; +_Mallotus_ (1 sp.), coasts of Arctic America from Greenland to Kamschatka; +_Retropinna_ (1 sp.), fresh waters of New Zealand; _Coregonus_ (41 sp.), +fresh waters of northern parts of temperate Europe, Asia and North America, +many of the species migratory: _Thymallus_ (6 sp.), fresh waters of +temperate parts of {448}Europe, Asia, and North America; _Argentina_ (4 +sp.), Mediterranean and deep seas of Western Europe; _Microstoma_ (2 sp.), +Mediterranean, and seas of Greenland; _Salarix_ (2 sp.), China and Japan, +in seas and rivers. _Salmo_, _Osmerus_, _Coregonus_, and _Thymallus_, are +British genera. + + +FAMILY 66.--PERCOPSIDÆ. (1 Genus, 1 Species.) + +"A fresh-water fish covered with toothed scales." + +DISTRIBUTION.--Lake Superior, North America. + + +FAMILY 67.--GALAXIDÆ. (1 Genus, 12 Species.) + +"Fresh-water fishes, with neither scales nor barbels." + +DISTRIBUTION.--The temperate zone of the Southern Hemisphere. + +The only genus, _Galaxias_, is found in New Zealand, Tasmania, and Tierra +del Fuego, ranging north as far as Queensland and Chili; and one of the +species is absolutely identical in the two regions. + + +FAMILY 68.--MORMYRIDÆ. (3 Genera, 25 Species.) + +"Fresh-water fishes, with scales on the body and tail but not on the head, +and no barbels." + +DISTRIBUTION.--The Ethiopian Region. + +Most abundant in the Nile, a few from the Gambia, the Congo, and Rovuma. +The genera are:-- + +_Mormyrus_ (1 sp.), Nile, Gambia, West Africa, Mozambique, Rovuma; +_Hyperopsius_ (2 sp.), Nile and West Africa; _Mormyrops_ (4 sp.), Nile, +West Africa and Mozambique. + + +{449}FAMILY 69.--GYMNARCHIDÆ. (1 Genus, 1 Species.) + +"Fresh-water fishes, resembling the Mormyridæ, but with tapering finless +tail, and neither anal nor ventral fins." + +DISTRIBUTION.--Ethiopian region. + +The only genus, _Gymnarchus_, inhabits the Nile and the rivers of West +Africa. + + +FAMILY 70.--ESOCIDÆ. (1 Genus, 7 Species.) + +"Fresh-water fishes, with scaly bodies, no barbels, and dorsal fins +situated towards the tail." + +DISTRIBUTION.--The Nearctic and Palæarctic regions. + +One species, the Pike (_Esox lucius_) ranges from Lapland to Turkey, and in +America from the Arctic regions to the Albany river; the remainder are +American species extending South as far as New Orleans. + + +FAMILY 71.--UMBRIDÆ. (1 Genus, 2 Species.) + +"Small fresh-water scaly fishes, without barbels or adipose fin." + +DISTRIBUTION.--Central Europe and Temperate North America. + + +FAMILY 72.--SCOMBRESOCIDÆ. (5 Genera, 136 Species.) + +"Marine or fresh-water fishes, with scaly bodies and a series of keeled +scales along each side of the belly." + +DISTRIBUTION.--Temperate and tropical regions. + +All the genera have a wide distribution. A species of _Belone_ and one of +_Scombresox_ are found on the British coast. The Flying-fishes (_Exocetus_, +44 sp.), belong to this family. They abound in all tropical seas and extend +as far as the Mediterranean and Australia. None of the genera are +exclusively fresh-water, {450}but a few species of _Belone_, and +_Hemiramphus_ are found in rivers in various parts of the world. + + +FAMILY 73.--CYPRINODONTIDÆ. (20 Genera, 106 Species.) + +"Fresh-water fishes, covered with scales, the sexes frequently differing, +mostly viviparous." + +DISTRIBUTION.--Southern Europe, Asia, Africa and North America, but most +abundant in Tropical America. + +The distribution of the genera is as follows:-- + +_Cyprinodon_ (11 sp.), Italy, North Africa and Western Asia to Persia, also +North America from Texas to New York; _Fitzroya_ (1 sp.), Montevideo; +_Characodon_ (1 sp.), Central America; _Tellia_ (1 sp.), Alpine pools of +the Atlas: _Limnurgus_ (1 sp.), Mexican plateau; _Lucania_ (1 sp.), Texas; +_Haplochilus_ (18 sp.), India, Java, Japan, Tropical Africa, Madagascar, +and the Seychelle Islands, Carolina to Brazil, Jamaica; _Fundulus_ (17 +sp.), North and Central America and Ecuador, Spain and East Africa; +_Rivulus_ (3 sp.), Tropical America, Cuba and Trinidad; _Orestias_ (6 sp.), +Lake Titacaca, Andes; _Jenynsia_ (1 sp.), Rio Plata; _Pseudoxiphophorus_ (2 +sp.), Central America; _Belonesox_ (1 sp.), Central America; _Gambusia_ (8 +sp.), Antilles, Central America and Texas; _Anableps_ (3 sp.), Central and +Equatorial America; _Poecilia_ (16 sp.), Antilles, Central and South +America; _Mollienesia_ (4 sp.), Louisiana to Mexico; _Platypoecilus_ (1 +sp.), Mexico; _Girardinus_ (10 sp.), Antilles and South Carolina to +Uruguay; _Lepistes_ (1 sp.), Barbadoes. + + +FAMILY 74.--HETEROPYGII. (2 Genera, 2 Species.) + +"Fresh-water fishes, with posterior dorsal fin, and very small scales." + +DISTRIBUTION.--Fresh waters of the United States. + +_Amblyopsis_ (1 sp.) is a blind fish found in the caverns of Kentucky; +while _Chologastes_ (1 sp.), which only differs from it in having perfect +eyes, is found in ditches in South Carolina. + + +{451}FAMILY 75.--CYPRINIDÆ. (109 Genera, 790 Species.) + +"Fresh-water fishes, generally scaly, with no adipose fin, and pharyngeal +teeth only, the mouth being toothless." + +DISTRIBUTION.--Fresh waters of the Old World and North America, but absent +from Australia and South America. + +This enormous family is divided by Dr. Günther into fourteen groups, the +distribution of which is as follows:-- + +Catostomina (4 genera), North America and North-east Asia; Cyprinina (39 +genera), same range as the family; Rohteichthyina (1 genus), Malay +Archipelago; Leptobarbina (1 genus), Malay Archipelago; Rasborina (5 +genera), East Africa to China and Borneo; Semiplotina (2 genera), Western +Asia; Xenocypridina (3 genera), Eastern Asia; Leuciscina (10 genera), +Palæarctic and Nearctic regions; Rhodeina (3 genera), Palæarctic region; +Danionina (9 genera), India to China and Japan; Hypophthalmichthyina (1 +genus), China; Abramidina (16 genera), same range as the family; +Homalopterina (2 genera), India to Java; Cobitidina (10 genera), Palæarctic +and Oriental regions. + +The following is the distribution of the genera:-- + +_Catostomus_ (16 sp.), Nearctic region and Eastern Siberia; _Moxostoma_ (2 +sp.), Eastern United States; _Sclerognathus_ (5 sp.), Temperate North +America to Guatemala, also Northern China; _Carpiodes_ (1 sp.), United +States; _Cyprinus_ (2 sp.), Temperate parts of Palæarctic region (1 sp. +British); _Carassius_ (3 sp.), Temperate Palæarctic region (1 sp. British); +_Catla_ (1 sp.), Continental India; _Cirrhina_ (5 sp.), Continental India +to China; _Dangila_ (6 sp.), Java, Sumatra, Borneo; _Osteochilus_ (14 sp.), +Siam to Java and Sumatra; _Labeo_ (27 sp.), Tropical Africa and Oriental +region; _Tylognathus_ (10 sp.), Syria, India to Java; _Abrostomus_ (2 sp.), +South Africa; _Discognathus_ (4 sp.), Syria to India and Java, mostly in +mountain streams; _Crossochilus_ (9 sp.), India to Sumatra and Java; +_Gymnostomus_ (7 sp.), Continental India; _Epalzeorhynchus_ (1 sp.), +Sumatra and Borneo; _Capoeta_ (13 sp.), Western Asia; _Barbus_ (163 sp.), +Temperate or Tropical {452}parts of Europe, Asia, and Africa (1 sp. +British); _Thynnichthys_ (2 sp.), Pegu, Borneo, and Sumatra; _Barbichthys_ +(1 sp.), Java, Sumatra, and Borneo; _Amblyrhynchichthys_ (1 sp.), Sumatra +and Borneo; _Albulichthys_ (1 sp.), Sumatra and Borneo; _Oreinus_ (3 sp.), +Himalayan region; _Schizothorax_ (13 sp.), Himalayan region and west to +Afghanistan and Persia; _Ptychobarbus_ (1 sp.), Thibet; _Gymnocypris_ (1 +sp.), loc. unknown; _Schizopygopsis_ (1 sp.), Thibet; _Diptychus_ (1 sp.), +Himalayas and Thibet; _Aulopyge_ (1 sp.), Western Asia; _Gobio_ (2 sp.), +Temperate Europe (1 sp. British); _Pseudogobio_ (4 sp.), China, Japan, and +Formosa; _Ceratichthys_ (9 sp.), Temperate North America; _Bungia_ (1 sp.), +Western Asia, Herat; _Pimephales_ (2 sp.), Eastern United States; +_Hyborhynchus_ (3 sp.), Eastern United States; _Ericymba_ (1 sp.), United +States; _Pseudorasbora_ (1 sp.), Japan, China; _Cochlognathus_ (1 sp.), +Texas; _Exoglossum_ (2 sp.), United States; _Rhinichthys_ (6 sp.), Eastern +United States; _Rohteichthys_ (1 sp.), Borneo and Sumatra; _Leptobarbina_ +(1 sp.), Sumatra and Borneo; _Rasbora_ (12 sp.), East Coast of Africa, +India, to Java and Borneo; _Luciosma_ (3 sp.), Java, Sumatra, and Borneo; +_Nuria_ (2 sp.), India, Tenasserim, and Ceylon; _Aphyocypris_ (1 sp.), +North China; _Amblypharyngodon_ (3 sp.), India to Tenasserim; _Cyprinion_ +(3 sp.), Syria and Persia; _Semiplotus_ (1 sp.), Assam; _Xenocypris_ (1 +sp.), China; _Paracanthobrama_ (1 sp.), China; _Mystacoleucus_ (1 sp.), +Sumatra; _Leuciscus_ (84 sp.), Nearctic and Palæarctic regions (5 sp. are +British); _Ctenopharyngodon_ (1 sp.), China; _Mylopharodon_ (1 sp.), +California; _Paraphoxinus_ (2 sp.), South-eastern Europe; _Meda_ (1 sp.), +River Gila; _Tinca_ (1 sp.), Europe (Britain to Constantinople); +_Leucosomus_ (8 sp.), Nearctic region; _Chondrostoma_ (7 sp.), Europe and +Western Asia; _Orthodon_ (1 sp.), California; _Acrochilus_ (1 sp.), +Columbia River; _Achilognathus_ (6 sp.), China, Japan, and Formosa; +_Rhodeus_ (3 sp.), Central Europe and China; _Pseudoperilampus_ (1 sp.), +Japan; _Danio_ (8 sp.), India and Ceylon; _Pterosarion_ (2 sp.), Central +India and Assam; _Aspidoparia_ (3 sp.), Continental India; _Barilius_ (15 +sp.), East Africa and Continental India; _Bola_ (1 sp.), Ganges to +Bramahputra; _Schacra_ (1 sp.), Bengal; _Opsariichthys_ (5 sp.), Japan and +Formosa; _Squaliobarbus_ (1 sp.), China; _Ochetobius_ (1 sp.), North China; +{453}_Hypophthalmichthys_ (2 sp.), China; _Abramis_ (16 sp.), North +America, Central Europe, and Western Asia (1 sp. is British); _Aspius_ (3 +sp.), East Europe, Western Asia, China; _Alburnus_ (15 sp.), Europe and +Western Asia (1 British sp.); _Rasborichthys_ (1 sp.), Borneo; +_Elopichthys_ (1 sp.), China; _Pelotrophus_ (2 sp.), East Africa; +_Acanthobrama_ (3 sp.), Western Asia; _Osteobrama_ (5 sp.), Continental +India; _Chanodichthys_ (6 sp.), China and Formosa; _Smiliogaster_ (1 sp.), +Bengal; _Culter_ (2 sp.), China; _Pelecus_ (1 sp.), Eastern Europe; +_Eustira_ (1 sp.), Ceylon; _Chela_ (16 sp.), India to Siam, Java and +Borneo; _Pseudolabuca_ (1 sp.), China; _Cachius_ (1 sp.), Continental +India; _Homaloptera_ (8 sp.), India to Cochin China, Java, and Sumatra; +_Psilorhynchus_ (2 sp.), North-eastern India; _Misgurnus_ (5 sp.), Europe +to India, China, and Japan; _Nemachilus_ (37 sp.), Europe and Asia; +_Cobitis_ (3 sp.), Europe, India, Japan; _Lepidocepalichthys_ (3 sp.), +India, Ceylon, and Java; _Acanthopsis_ (2 sp.), Tenasserim, Sumatra, Java, +and Borneo; _Botia_ (7 sp.), India to Japan and Sunda Isles; _Oreonectes_ +(1 sp.), China; _Lepidocephalus_ (1 sp.), Java and Sumatra; +_Acanthopthalmus_ (2 sp.), Java and Sumatra; _Apua_ (1 sp.), Tenasserim; +_Kneria_ (2 sp.), Tropical Africa. + + +FAMILY 76.--GONORHYNCHIDÆ. (1 Genus, 1 Species.) + +"A marine fish with spiny scales, mouth with barbels, and with short dorsal +fin opposite the ventrals." + +DISTRIBUTION.--Temperate parts of Southern Oceans, and Japan. + + +FAMILY 77.--HYODONTIDÆ. (1 Genus, 1 Species.) + +"A fresh-water fish with cycloid scales and posterior dorsal fin." + +DISTRIBUTION.--Fresh waters of North America. + + +{454}FAMILY 78.--OSTEOGLOSSIDÆ. (3 Genera, 5 Species.) + +"Fresh-water fishes, with large hard scales, and dorsal fin opposite and +equal to the anal fin." + +DISTRIBUTION.--Tropical rivers. + +The genera are:--_Osteoglossum_ (3 sp.), Eastern South America, Sunda +Islands, and Queensland; _Arapaima_ (1 sp.), Eastern South America--the +"Pirarucú" of the Amazon; _Heterotis_ (1 sp.), Tropical Africa. + + +FAMILY 79.--CLUPEIDÆ. (18 Genera, 161 Species.) + +"Marine scaly fishes, without barbels, and with the abdomen often +compressed and serrated." + +DISTRIBUTION.--Seas of the whole globe, many species entering rivers. They +are most abundant in the Indian seas, less so in America, scarce in Africa, +while they are almost absent from Australia. The Herring, Sprat, Shad, and +Pilchard, are British species of _Clupea_, a genus which contains 61 +species and ranges all over the world. + + +FAMILY 80.--CHIROCENTRIDÆ. (1 Genus, 1 Species.) + +"A marine fish, with thin deciduous scales, no barbels, and posterior +dorsal fin." + +DISTRIBUTION.--The Eastern seas from Africa to China. + + +FAMILY 81.--ALEPOCEPHALIDÆ. (1 Genus, 1 Species.) + +"A marine fish, covered with thin cycloid scales, no barbels, and posterior +dorsal fin." + +DISTRIBUTION.--Deep waters of the Mediterranean. + + +{455}FAMILY 82.--NOTOPTERIDÆ. (1 Genus, 5 Species.) + +"Fresh-water fishes, without barbels, head and body scaly, long tapering +tail, and short posterior dorsal fin." + +DISTRIBUTION.--Rivers of India, Siam, the Sunda Islands, and West Africa. + + +FAMILY 83.--HALOSAURIDÆ. (1 Genus, 1 Species.) + +"Marine fishes, with cycloid scales, a short median dorsal fin, and no +barbels." + +DISTRIBUTION.--Deep waters of the Atlantic, Madeira. + + +FAMILY 84.--GYMNOTIDÆ. (5 Genera, 20 Species.) + +"Fresh-water fishes, with elongate bodies, pointed tail, and no dorsal +fin." + +DISTRIBUTION.--Tropical America from Trinidad to the River Parana. + +The genera are distributed as follows:-- + +_Sternarchus_ (8 sp.), Guiana and Brazil; _Rhamphichthys_ (6 sp.), Guiana +and Brazil; _Sternophygus_ (4 sp.), Tropical America; _Carapus_ (1 sp.), +Trinidad to Brazil; _Gymnotus_, (1 sp.--the Electric eel), Tropical South +America. + + +FAMILY 85.--SYMBRANCHIDÆ. (4 Genera, 6 Species.) + +"Marine and fresh-water fishes, having elongate bodies without fins, and +very minute scales or none." + +DISTRIBUTION.--Fresh waters and coasts of Western Australia and Tasmania. + +The genera are:-- + +_Amphipnous_ (1 sp.), Bengal; _Monopterus_ (1 sp.), Siam to Northern China +and Sunda Islands; _Symbranchus_ (3 sp.), Tropical {456}America, and India +to Australia; _Chilobranchus_ (1 sp.), Australia and Tasmania. + + +FAMILY 86.--MURÆNIDÆ. (26 Genera, 230 Species.) + +"Marine or fresh-water fishes, with cylindrical or band-like bodies and no +ventral fins." + +DISTRIBUTION.--The seas and fresh waters of temperate and tropical regions. +This family is divided by Dr. Günther into two sub-families and nine +sections. The genus _Anguilla_, comprising our common Eel and a number of +species from all parts of the world, is the only one which is found in +fresh water, though even here most of the species are marine. _Anguilla_ +and _Conger_ are the only British genera. + + +FAMILY 87.--PEGASIDÆ. (1 Genus, 4 Species.) + +"Small marine fishes, covered with bony plates, and short opposite dorsal +and anal fins." + +DISTRIBUTION.--Indian Ocean and seas of China and Australia. + + +_Order V.--LOPHOBRANCHII._ + +"Fish with a segmented bony covering, long snout, and small toothless +mouth." + + +FAMILY 88.--SOLENOSTOMIDÆ. (1 Genus, 3 Species.) + +"Marine Lophobranchii, with wide gill openings and two dorsal fins." + +DISTRIBUTION.--Indian Ocean, from Zanzibar to China and the Moluccas. + + +{457}FAMILY 89.--SYNGNATHIDÆ. (15 Genera, 112 Species.) + +"Marine Lophobranchii, with very small gill opening and one soft dorsal +fin." + +DISTRIBUTION.--All the tropical and temperate seas. Some species of +_Syngnathus_, _Doryichthys_, and _Coelonotus_ enter fresh water, and a few +live in it exclusively. _Siphonostoma_, _Syngnathus_, _Nerophis_, and +_Hippocampus_ are British genera. The _Hippocampina_ (5 genera, 25 sp.), or +Sea-horses, are peculiar to the Indian and Pacific Oceans, except three or +four species of _Hippocampus_ in the Atlantic and Mediterranean. + + +_Order VI.--PLECTOGNATHI._ + +"Fishes covered with rough scales or shields, having a narrow mouth, and +soft posterior dorsal fin." + + +FAMILY 90.--SCLERODERMI. (7 Genera, 95 Species.) + +"Marine Plectognathi, with toothed jaws." + +DISTRIBUTION.--Temperate and Tropical seas, but much more abundant in the +Tropics. + + +FAMILY 91.--GYMNODONTES. (10 Genera, 82 Species.) + +"Marine or fresh-water Plectognathi, with jaws modified into a beak." + +DISTRIBUTION.--Temperate and tropical regions. + +Some species of _Tetrodon_ are found in the rivers of Tropical America, +Africa, and Asia. Species of _Tetrodon_ and _Orthagoriscus_ have been found +on the British coasts. + + +{458}SUB-CLASS II.--DIPNOI. + + +FAMILY 92.--SIRENOIDEI. (3 Genera, 3 Species.) + +"Eel-shaped fresh-water fishes, covered with cycloid scales; the vertical +fins forming a continuous border to the compressed tapering tail." + +DISTRIBUTION.--Rivers of Tropical Africa, South America, and Australia. + +The genera are:--_Protopterus_ (1 sp.), Tropical Africa; _Lepidosiren_ (1 +sp.), Amazon Valley; _Ceratodus_ (1 sp.), Queensland. + + +SUB-CLASS III.--GANOIDEI. + + +_Order I.--HOLOSTEI._ + +"Body covered with scales." + + +FAMILY 93.--AMIIDÆ. (1 Genus, 1 Species.) + +"A fresh-water fish, with cycloid scales and a long soft dorsal fin." + +DISTRIBUTION.--United States. + + +FAMILY 94.--POLYPTERIDÆ. (2 Genera, 2 Species.) + +"Fresh-water fishes, with ganoid scales and dorsal spines." + +DISTRIBUTION.--Central and Western Africa. + +The genera are:-- + +_Polypterus_ (1 sp.), the Nile and rivers of West Africa; _Calamoichthys_ +(1 sp.), Old Calabar. + + +{459}FAMILY 95.--LEPIDOSTEIDÆ. (1 Genus, 3 Species.) + +"Fresh-water fishes, with ganoid scales, and dorsal and anal fins composed +of articulated rays." + +DISTRIBUTION.--The genus Lepidosteus, the Garfishes or Bony Pikes, inhabits +North America to Mexico and Cuba. + + +_Order II.--CHONDROSTEI._ + +"Sub-cartilaginous scaleless fishes with heterocercal tail, the skin with +osseous bucklers or naked." + + +FAMILY 96.--ACCIPENSERIDÆ. (2 Genera, 20 Species.) + +"Marine or fresh-water fishes with osseous bucklers and inferior mouth." + +DISTRIBUTION.--Temperate and Arctic regions of the northern hemisphere. +ACCIPENSER (19 sp.), comprising the Sturgeons, has the distribution of the +family; most of the species are marine, but some are confined to the +Caspian and Black Seas and the great American lakes with the rivers flowing +into them, while the Danube, Mississippi, and Columbia River have peculiar +species. The other genus, SCAPHIRHYNCHUS (1 sp.), is confined to the +Mississippi and its tributaries. + + +FAMILY 97.--POLYDONTIDÆ. (1 Genus, 2 Species.) + +"Fresh-water fishes, with wide lateral mouth and naked skin." + +DISTRIBUTION.--The Mississippi and Yang-tse-kiang rivers. + + +{460}SUB CLASS IV.--CHONDROPTERYGII. (SHARKS AND RAYS.) + + +_Order I.--HOLOCEPHALA. (Chimæras.)_ + + +FAMILY 98.--CHIMÆRIDÆ. (2 Genera, 4 Species.) + +"Shark-like marine fishes, snout of the male with a prehensile organ." + +DISTRIBUTION.--Northern and Southern temperate seas. _Chimæra_ is British. + + +_Order II.--PLAGIOSTOMATA._ + +Sub-order.--SELACHOIDEA. (Sharks.) + + +FAMILY 99.--CARCHARIIDÆ. (11 Genera, 59 Species.) + +"Sharks with two dorsals and a nictitating membrane." + +DISTRIBUTION.--Seas of the Arctic, temperate, and tropical regions. Species +of _Galeus_ and _Mustelus_ have occurred on our coasts. + + +FAMILY 100.--LAMNIDÆ. (5 Genera, 7 Species.) + +"Sharks with two dorsals and no nictitating membrane." + +DISTRIBUTION.--Temperate and tropical seas. Species of _Lamna_, +_Alopecias_, and _Selache_ have occurred in British seas. + + +{461}FAMILY 101.--RHINODONTIDÆ. (1 Genus, 1 Species.) + +"Sharks with two dorsal fins, the second small, and no nictitating +membrane." + +DISTRIBUTION.--South and East Africa. + + +FAMILY 102.--NOTIDANIDÆ. (1 Genus, 4 Species.) + +"Sharks with one dorsal fin and no nictitating membrane." + +DISTRIBUTION.--Temperate and tropical seas, from the North Atlantic to the +Cape of Good Hope and California. One species has occurred on our southern +coasts. + + +FAMILY 103.--SCYLLIIDÆ. (7 Genera, 25 Species.) + +"Sharks with one dorsal fin and no nictitating membrane." + +DISTRIBUTION.--All temperate and tropical seas. Species of _Scyllium_ and +_Pristiurus_ are British. + + +FAMILY 104.--CESTRACIONTIDÆ. (1 Genus, 4 Species.) + +"Sharks with two dorsal fins and no nictitating membrane." + +DISTRIBUTION.--Pacific Ocean from Japan to New Zealand, Moluccan Sea. + + +FAMILY 105.--SPINACIDÆ. (10 Genera, 21 Species.) + +"Sharks with two dorsal fins and no nictitating membrane, no anal fin." + +DISTRIBUTION.--Arctic, temperate, and tropical seas. Species of +_Acanthias_, _Læmargus_, and _Echinorhinus_ have occurred on our coasts. + + +{462}FAMILY 106.--RHINIDÆ (1 Genus, 1 Species.) + +"Sharks with depressed flat body and large expanded pectoral fins." + +DISTRIBUTION.--Temperate and tropical seas, from Britain to California and +Australia. + + +FAMILY 107.--PRISTIOPHORIDÆ. (1 Genus, 4 Species.) + +"Sharks with produced flat snout, armed with teeth on each edge." + +DISTRIBUTION.--Seas of Japan and Australia. + + +Sub-order BATOIDEI. (Rays.) + + +FAMILY 108.--PRISTIDÆ. (1 Genus, 5 Species.) + +"Rays with produced snout and lateral saw-like teeth." + +DISTRIBUTION.--Seas of tropical and sub-tropical regions. + + +FAMILY 109.--RHINOBATIDÆ. (3 Genera, 15 Species.) + +"Rays with long and strong tail, having a caudal and two dorsal fins." + +DISTRIBUTION.--Tropical and sub-tropical seas. + + +FAMILY 110.--TORPEDINIDÆ. (6 Genera, 15 Species.) + +"Rays with broad smooth disc, and an electric organ." + +DISTRIBUTION.--Tropical and temperate seas, from Britain to Tasmania. + + +FAMILY 111.--RAIIDÆ. (4 Genera, 29 Species.) + +"Rays with broad rhombic disc and no serrated caudal spine." + +DISTRIBUTION.--All temperate and tropical seas. Several species of _Raia_ +are found on our coasts. + + +{463}FAMILY 112.--TRYGONIDÆ (6 Genera, 43 Species.) + +"Rays with the pectoral fins extending to end of snout." + +DISTRIBUTION.--Seas of all temperate and tropical regions, and rivers of +Tropical America. A species of _Trygon_ has occurred on our Southern coast. +_Ellipesurus_ and _Tæniura_ are found in the fresh waters of the interior +of South America, while the latter genus occurs also in the Indian seas, +but not in the Atlantic. + + +FAMILY 113.--MYLOBATIDÆ. (5 Genera, 22 Species.) + +"Rays with very broad pectoral fins not extending to end of snout." + +DISTRIBUTION.--Temperate and tropical seas. A species of MYLIOBATIS is +British, but most of the species and genera are confined to tropical seas. +_Dicerobatis_ and _Ceratoptera_ are very large Rays, commonly called +Sea-devils. + + +SUB-CLASS V.--CYCLOSTOMATA. + +"Cartilaginous fishes, with suctorial mouths and without lateral fins." + + +FAMILY 114.--PETROMYZONTIDÆ. (4 Genera, 12 Species.) + +"Marine or fresh-water eel-like fishes, with suctorial mouths and without +barbels." + +DISTRIBUTION.--Coasts and fresh waters of temperate regions of both +hemispheres. Three species of _Petromyzon_ (Lampreys), are British. + + +{464}FAMILY 115.--MYXINIDÆ. (2 Genera, 5 Species.) + +"Marine eel-like fishes, with four pairs of barbels." + +DISTRIBUTION.--Seas of the temperate regions of both hemispheres. + + +SUB-CLASS VI.--LEPTOCARDII. + + +FAMILY 116.--CIRRHOSTOMI. (1 Genus, 1 Species.) + +"A small marine fish with no jaws or fins, and with rudimentary eyes." + +DISTRIBUTION.--The only species, the Lancelet (_Amphioxus_), is the lowest +form of living vertebrate. It is found in the temperate regions of both +hemispheres, and has occurred on our southern coast. + + +_Remarks on the Distribution of Fishes._ + +_Marine Fish._--There are about 80 families of marine fishes, and of these +no less than 50 are universally, or almost universally, distributed over +the seas and oceans of the globe. Of the remainder many are widely +distributed, some species even ranging from the North Atlantic to +Australia. Six families are confined to the Northern Seas, but four of +these consist of single species only, the other two being the Discoboli (2 +genera, 11 sp.), and the Accipenseridæ (2 genera and 20 sp.). Only one +family (Acanthoclinidæ) is confined to the Southern oceans, and that +consists of but a single species. Four families (Sternoptychidæ, +Stomiatidæ, Alepocephalidæ and Halosauridæ) are confined to the Atlantic +Ocean, while 13 are found only in the Pacific; and of the remainder several +are more abundant in the Pacific than the Atlantic. Two families (Lycodidæ +and Gadidæ) are found in the Arctic and Antarctic seas only, though the +{465}latter family has a single species in the Indian seas. Among the +curiosities of distribution are,--the extensive genus _Diagramma_, confined +to the Pacific with the exception of one species in the Mediterranean; the +single species constituting the family Lophotidæ, found only in the +Mediterranean and Japan; the small family of Notacanthi, confined to +Greenland, the Mediterranean, and West Australia; and the four families, +Sternoptychidæ, Stomiatidæ, Alepocephalidæ, and Halosauridæ, which are +believed to inhabit exclusively the depths of the ocean, and are therefore +very rarely obtained. + +_Fresh-water Fish._--There are 36 families of fishes which inhabit fresh +water exclusively, and 5 others, which are both marine and fresh-water. +These present many interesting peculiarities of distribution. The +Neotropical region is the richest in families, and probably also in genera +and species. No less than 22 families inhabit it, and of these 6 are +altogether peculiar. The Ethiopian and Nearctic regions each have 18 +families, the former with 3, and the latter with 5 peculiar. Several +isolated forms, requiring to be placed in distinct families, inhabit the +great American lakes; and, no doubt, when the African lakes are equally +well known, they will be found also to possess many peculiar forms. The +Oriental region comes next, with 17 families, of which 3 are peculiar. The +Palæarctic has 12, and the Australian 11 families, each with only 1 +altogether peculiar to it. + +If we take those regions which are sometimes supposed to be so nearly +related that they should be combined, we shall find the fresh-water fishes +in most cases markedly distinct. The Nearctic and Palæarctic regions, for +example, together contain 20 families, but only 11 of these occur in both, +and only 5 are exclusive inhabitants of these two regions. This shows an +amount of diversity that would not, perhaps, be exhibited by any other +class of animals. The Ethiopian and Oriental regions together possess 24 +families, only 11 of which are found in both, and only 1 exclusively +characteristic of the two. The Australian and Neotropical regions possess +together 27 families, of which 7 are found in both, and 3 are exclusively +characteristic of the two. This last fact is very interesting: the marine +family of {466}Trachinidæ possesses a fresh-water genus, _Aphritis_, one +species of which inhabits Tasmania, and two others Patagonia; the +Haplochitonidæ (2 genera, 3 sp.) are found only in Tierra del Fuego, the +Falkland Islands, and South Australia; and the Galaxidæ (1 genus, 12 sp.) +inhabit the same regions, but extend to Chili, to New Zealand and to +Queensland. We have here an illustration of that connection between South +America and Australia which is so strongly manifested in plants, but of +which there are only scattered indications in most classes of animals. The +dividing line across the Malay Archipelago, separating the Oriental from +the Australian regions, and which is so strikingly marked in mammalia and +birds, is equally so in fresh-water fishes. No less than six families have +their eastern limits in Java and Borneo; while the extensive family of +Cyprinidæ has no less than 23 genera in Java and Borneo, but not a single +species has been found in Celebes or the Moluccas. + +The distribution of fresh-water fishes lends no support to the view that +the peninsula of India belongs to the Ethiopian region. A large proportion +of the Oriental families are common to the whole region; while there is +hardly a single example, of a characteristic Ethiopian family or genus +extending into the peninsula of India and no further. + +Among the special peculiarities of distribution, is the curious fish, +forming the family Comephoridæ, which is confined to Lake Baikal, among the +mountains of Central Asia, 2,000 feet above the sea, and a thousand miles +distant from the ocean; yet having its nearest allies in the exclusively +oceanic family of the mackerels (Scomberidæ). The Characinidæ are confined +to Africa and South America, distinct genera inhabiting each region. The +Salmonidæ are confined to the two northern regions, except a single species +of a peculiar genus in New Zealand. The genus _Osteoglossum_ has a species +in South America, another in the Sunda Islands, and a third in Queensland; +while the curious Sirenoidei are represented by single species of peculiar +genera in Tropical America, Tropical Africa, and Tropical Australia. + +_Fossil Fishes._--Fishes have existed from a very remote era, and it is +remarkable that the first whose remains have been {467}discovered belong to +the Ganoidei, a highly developed group which has continued to exist down to +our times, and of which the sturgeon is the best known example. We may +therefore be sure that the Upper Silurian rocks in which these are found, +although so very far back in geological history, do not by any means lead +us to the time when the primitive fish-type appeared upon the earth. In the +Carboniferous and Permian formations numerous remains of fishes are found, +allied to the _Lepidosteus_ or Gar-pike of North America. The next group in +order of appearance, are the Plagiostomata, containing the existing Sharks +and Rays. Traces of these are found in the highest Silurian beds, and +become plentiful in the Devonian and Carboniferous formations and in all +succeeding ages, being especially abundant in Cretaceous and Eocene strata. +The Holocephali appear first in the Oolitic period, and are represented by +the living Chimæridæ. The Dipnoi, to which belong the _Lepidosiren_ and +_Ceratodus_, are believed to have existed in the Triassic period, from the +evidence of teeth almost identical with those of the existing Australian +fish. All the ancient fossil fishes belong to the above-mentioned groups, +and many of them have little resemblance to existing forms. The Teleostean +fishes, which form the great bulk of those now living, cannot be traced +back further than the Cretaceous period, while by far the larger number +first appear in the Tertiary beds. The Salmonidæ, Scopelidæ, Percidæ, +Clupeidæ, Scombresocidæ, Mugilidæ, and Siluridæ, or forms closely allied to +them, are found in the Cretaceous formation. In the Eocene beds we first +meet with Squammipennes, Cyprinidæ, Pleuronectidæ, Characinidæ, Murænidæ, +Gadidæ, Pediculati, Syngnathidæ, and Hippocampidæ. + +Most of these fossils represent marine fishes, those of fresh-water origin +being rare, and of little importance as an aid in determining the causes of +the distribution of living forms. To understand this we must look to the +various changes of the land surface which have led to the existing +distribution of all the higher vertebrates, and to those special means of +dispersal which Mr. Darwin has shown to be possessed by all fresh-water +productions. + + + +{468}CHAPTER XXI. + +THE DISTRIBUTION OF SOME OF THE MORE IMPORTANT FAMILIES AND GENERA OF +INSECTS. + + +Although insects are, for the most part, truly terrestrial animals, and +illustrate in a very striking manner the characteristic phenomena of +distribution, it is impossible here to treat of them in much detail. This +arises chiefly from their excessive numbers, but also from the minuteness +and obscurity of many of the groups, and our imperfect knowledge of all but +the European species. The number of described species of insects is +uncertain, as no complete enumeration of them has ever been made; but it +probably exceeds 100,000, and these may belong to somewhere about 10,000 +genera--many times more than all vertebrate animals together. Of the eight +Orders into which Insects are usually divided, only two--the Coleoptera and +Lepidoptera--have been so thoroughly collected in all parts of the globe +that they can be used, with any safety, to compare their distribution with +that of vertebrate animals; and even of these it is only certain favourite +groups which have been so collected. Among Lepidoptera, for example, +although the extensive group of Butterflies may be said, in a general +sense, to be thoroughly well known--every spot visited by civilized man +having furnished its quota to our collections--yet the minute Tineidæ, or +even the larger but obscure Noctuidæ, have scarcely been collected at all +in tropical countries, and any attempt to study their geographical +distribution would certainly lead to erroneous results. The same thing +occurs, though perhaps in a less degree, among the Coleoptera. While the +Carabidæ, Buprestidæ, and {469}Longicorns of the Tropics, are almost as +well known as those of the Temperate Zones, the Staphylinidæ, the smaller +Elateridæ, and many other obscure and minute groups, are very imperfectly +represented from extra-European countries. I therefore propose to examine +with some care the distribution of the Butterflies, and the Sphingina among +Lepidoptera, and the following large and well-known families of +Coleoptera:--Cicindelidæ, Carabidæ, Lucanidæ, Cetoniidæ, Buprestidæ, and +the three families of Longicorns. These families together contain over +30,000 species, classed in nearly 3,000 genera, and comprise a large +proportion of the best known and most carefully studied groups. We may +therefore consider, that a detailed examination of their distribution will +lead us to results which cannot be invalidated by any number of isolated +facts drawn from the less known members of the class. + + + +_Range of Insects in Time._--In considering how much weight is to be given +to facts in insect distribution, and what interpretation is to be put upon +the anomalies or exceptional cases that may be met with, it is important to +have some idea of the antiquity of the existing groups, and of the rate at +which the forms of insect life have undergone modification. The geological +record, if imperfect in the case of the higher animals, is fragmentary in +the extreme as regards indications of former insect life; yet the positive +facts that it does disclose are of great interest, and have an important +bearing on our subject. These facts and the conclusions they lead to have +been discussed in our first volume (p. 166), and they must be carefully +weighed in all cases of apparent conflict or incongruity between the +distribution of insects and that of the higher animals. + + +{470}_Order--LEPIDOPTERA._ + +Sub-order--_Lepidoptera_ RHOPALOCERA, or BUTTERFLIES. + +FAMILY 1.--DANAIDÆ. (24 Genera, 530 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |-- 2 -- -- |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Danaidæ are now held to comprehend, not only the whole of the group so +named by Doubleday, but a large portion of the Heliconidæ of that author. +Their range is thus extended over the whole of the tropical regions. A few +species spread northwards into the Palæarctic and Nearctic regions, but +these are only stragglers, and hardly diminish the exclusively tropical +character of the group. The more remarkable genera are,--_Hestia_ (10 sp.), +and _Ideopsis_ (6 sp.), confined to the Malayan and Moluccan districts; +_Danais_ (50 sp.), which has the range of the whole family; _Euploea_ (140 +sp.), confined to the Oriental and Australian regions, but especially +abundant in the Malayan and Moluccan districts; _Hamadryas_ (4 sp.), +Australian region only. The remaining genera constitute the Danaioid +Heliconidæ, and are strictly confined to Tropical America, except a few +species which extend into the southern parts of the Nearctic region. The +chief of these genera are:-- + +_Ithomia_ (160 sp.), _Melinæa_ (18 sp.), _Napeogenes_ (20 sp.), +_Mechanitis_ (4 sp.), _Ceratina_ (32 sp.), _Dircenna_ (10 sp.), and +_Lycorea_ (4 sp.). Florida, Louisiana, and Southern California, mark the +northern extent of these insects. + + +{471}FAMILY 2.--SATYRIDÆ. (60 Genera, 835 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +This family has an absolutely universal distribution, extending even into +the Arctic and Antarctic regions. Many of the genera are, however, +restricted in their range. + +_Hætera_, _Lymanopoda_, _Calisto_, _Corades_, _Taygetis_, _Pronophila_, +_Euptychia_, and some allied forms (25 genera in all) are Neotropical, the +last named extending north to Canada; _Debis_, _Melanitis_, _Mycalesis_ and +_Ypthima_, are mostly Oriental, but extending also into the Australian and +the Ethiopian regions; _Gnaphodes_, _Leptoneura_, and a few other small +genera, are exclusively Ethiopian; _Xenica_, _Hypocista_, and +_Heteronympha_, are Australian; _Erebia_, _Satyrus_, _Hipparchia_, +_Coenonympha_, and allies, are mostly Palæarctic, but some species are +Ethiopian, and others Nearctic; _Chionabas_, is characteristic of the whole +Arctic regions, but is also found in Chili and the Western Himalayas. The +peculiar genera in each region are,--Neotropical, 25; Australian, 7; +Oriental, 11; Ethiopian, 5; Palæarctic, 3; Nearctic, 0. + + +FAMILY 3.--ELYMNIIDÆ. (1 Genus, 28 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|-- -- -- --|-- -- -- --|-- 2 -- -- |-- -- 3. 4 |1 -- -- -- + | | | | | + +The genus _Elymnias_, which constitutes this family, is characteristic of +the Malayan and Moluccan districts, with some species in Northern India and +one in Ashanti. It thus agrees with several groups of Vertebrata, in +showing the resemblance {472}of Malaya with West Africa independently of +the Peninsula of India. + + +FAMILY 4. MORPHIDÆ. (10 Genera, 106 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- 3. 4 | 1 -- 3 -- + | | | | | + +The Morphidæ are a group of generally large-sized butterflies, especially +characteristic of the Malayan and Moluccan districts, and of Tropical +America; with a few species extending to the Himalayas on the west, and to +Polynesia on the east. The genera are:-- + +_Amathusia_ (6 sp.), Northern India to Java; _Zeuxidia_ (9 sp.), the Malay +district; _Discophora_ (7 sp.), Northern India to Philippines, Java and +Timor; _Enispe_ (3 sp.), Northern India; _Hyades_ (15 sp.), Moluccan and +Polynesian districts, except one species in Java; _Clerome_ (11 sp.), +Northern India to Philippines and Celebes; _Æmona_ (1 sp.), Sikhim; +_Hyantis_ (1 sp.), Waigiou; _Thaumantis_ (10 sp.), Indo-Chinese and Malayan +districts; _Morpho_ (40 sp.), Neotropical region, Brazilian and Central +American sub-regions. + + +FAMILY 5. BRASSOLIDÆ. (7 Genera, 62 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The Brassolidæ have the same distribution as the genus _Morpho_. The genera +are:-- + +_Brassolis_ (5 sp.); _Opsiphanes_ (17 sp.); _Dynastor_ (2 sp.); _Penetes_ +(1 sp.); _Caligo_ (21 sp.); _Narope_ (5 sp.); and _Dasyopthalma_ (3 sp.) + + +{473}FAMILY 6.--ACRÆIDÆ. (1 Genus, 90 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |1. 2 -- -- + | | | | | + +The genus _Acræa_ is especially abundant in the Ethiopian region, which +contains two-thirds of all the known species; 3 or 4 species only, range +over the whole Oriental, and most of the Australian regions; while all the +rest inhabit the same districts of the Neotropical region as the +Brassolidæ. + + +FAMILY 7.--HELICONIDÆ. (2 Genera, 114 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |-- -- 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +The true Heliconidæ are very characteristic of the Neotropical region; one +species only extending into the Southern States of North America as far as +Florida. The genus _Heliconius_ (83 sp.), has the range of the family; +while _Eueides_ (19 sp.), is confined to the Brazilian and Central American +sub-regions. + + +FAMILY 8.--NYMPHALIDÆ. (113 Genera, 1490 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +This is the largest and most universally distributed family of butterflies, +and is well illustrated by our common Fritillaries, {474}Tortoise-shell, +Peacock, Painted Lady, and Purple Emperor butterflies. They are found +wherever butterfly-life can exist, and some single species--like the +Painted Lady (_Pyrameis cardui_)--range almost over the globe. A few of the +more extensive and remarkable genera only, can be here noticed:-- + +_Colænis_, _Agraulis_, _Eresia_, _Synchloe_, _Epicalia_, _Eunica_, +_Eubagis_, _Catagramma_, _Callithea_, _Ageronia_, _Timetes_, _Heterochroa_, +_Prepona_, _Hypna_, _Paphia_, and _Siderone_, are wholly Neotropical, as +well as many others which have a smaller number of species. _Euryphene_, +_Romaleosoma_, _Aterica_, and _Harma_, are exclusively Ethiopian. +_Terinos_, _Athyma_, _Adolias_, and _Tanæcia_, are Oriental, but they +mostly extend into the Moluccan region; the last however is strictly +Malayan, and _Adolias_ only reaches Celebes. _Mynes_ alone, is exclusively +Australian, but _Prothoe_ is almost so, having only one outlying species in +Java. _Eurytela_ and _Ergolis_ are confined to the Oriental and Ethiopian +regions, but the latter reaches the Moluccas. _Cethosia_, _Cirrhochroa_, +_Messaras_, and _Symphædra_, are both Oriental and Australian; while +_Junonia_, _Cyrestis_, _Diadema_, _Neptis_, and _Nymphalis_, are common to +the three tropical regions of the Eastern Hemisphere, the latter extending +into the Mediterranean district, while _Junonia_ occurs also in South +America and the Southern United States. + +The most cosmopolitan genus is _Pyrameis_, which has representatives in +every region and every district. _Apatura_ is found in all but the +Ethiopian and the Australian, although it just enters the confines of the +latter region in Celebes; _Limenitis_ is abundant in the Oriental region, +but extends eastward to Celebes and westward into Europe, North America, +and even into South America. _Argynnis_, _Melitæa_, and _Vanessa_, are +almost confined to the Palæarctic and Nearctic regions; the former however +occurs in the Himalayas and in the mountains of Java, and also in Chili and +in Jamaica. Two genera--_Dicrorrhagia_ and _Helcyra_--have both one species +in North India and another in the island of Ceram. The number of genera +peculiar to each region is as follows:--Neotropical, 50; Australian, 2; +Oriental 15; Ethiopian, 14; Palæarctic, 1; Nearctic, 0. + + +{475}FAMILY 9.--LIBYTHEIDÆ. (1 Genus, 10 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- 4 | -- 2.3 -- |1. 2 -- -- | -- 2 -- 4 |1. 2. 3. 4 |1 -- -- -- + | | | | | + +The genus _Libythea_, which constitutes this family, appears to have its +head-quarters in the Oriental region, but extends on all sides in an +erratic manner, into various remote and disconnected portions of the globe, +as indicated above. + + +FAMILY 10.--NEMEOBIIDÆ. (12 Genera, 145 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|1 -- -- -- | -- 2 -- 4 |-- -- 3. 4 |1 -- -- -- + | | | | | + +This group has been separated from the Erycinidæ of the older authors, and +contains all the non-American genera and species. Half the genera and +nearly four-fifths of the species of this group are, however, Neotropical; +one is European; two or three African; and twenty-six Oriental and +Australian. The genera are:-- + +_Nemeobius_ (1 sp.), Europe; _Dodona_ (6 sp.), North India; _Zemeros_ (2 +sp.), North India and Malaya; _Abisara_ (11 sp.), North India, Malayan and +Moluccan districts, Madagascar and West Africa; _Taxila_ (8 sp.), North +India and Malaya; _Dicallaneura_ (2 sp.), Moluccan district; _Alesa_ (6 +sp.), _Eunogyra_ (2 sp.), _Cremna_ (7 sp.), _Bæotis_ (3 sp.), are all from +the Brazilian sub-region; _Eurybia_ (10 sp.), _Mesosemia_ (80 sp.), inhabit +both the Brazilian and Mexican sub-regions. + + +{476}FAMILY 11.--EURYGONIDÆ. (2 Genera, 78 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --| -- --- -- |-- -- -- -- + | | | | | + +This small family, separated from the true Erycinidæ by Mr. Bates, is +confined to the tropical forest-districts of continental America. The +genera are:-- + +_Eurygona_ (71 sp.); _Methonella_ (1 sp.); the latter found in Equatorial +South America. + + +FAMILY 12.--ERYCINIDÆ. (59 Genera, 560 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |1. 2. 3 -- |-- -- -- --|-- -- -- --|-- -- -- --|-- -- -- -- + | | | | | + +This extensive family of small, but exquisitely beautiful butterflies, is +especially characteristic of the virgin forests of the Neotropical region, +only a few species of three genera extending into the Nearctic region. The +more important genera, and those which have an exceptional distribution, +can alone be here noticed. _Charis_ extends from Brazil to New York; +_Apodemia_ from Brazil to California, Utah, and Oregon; _Amarynthis_ +inhabits the Brazilian and Antillean sub-regions; _Lepricornis_ and +_Metapheles_ are small genera found only in the Mexican sub-region; +_Lymnas_, _Necyria_, _Ancyluris_, _Diorhina_, _Esthemopsis_, _Anteros_, +_Emesis_, _Symmachia_, _Cricosoma_, _Calydna_, _Lemonias_, _Nymphidium_, +_Theope_, and _Aricoris_ are common to the Brazilian and Mexican +sub-regions. All the other genera (40 in number) are only known from the +Brazilian sub-region, and of these a considerable proportion are confined +to the damp equatorial forests of the Amazon Valley. + + +{477}FAMILY 13.--LYCÆNIDÆ. (39 Genera, 1,220 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Lycænidæ--of the variety and beauty of which in tropical regions our +own "Blues" and "Coppers" give but a faint idea--are a group of universal +distribution. We shall therefore indicate those genera which are restricted +to one or more regions, or are nearly cosmopolitan. The large genus +_Polyommatus_ (containing 325 species) has the same universal distribution +as the entire family. Our common "Blues" well represent this genus. +_Lycæna_ (comprising the "Coppers") is more especially characteristic of +the Palæarctic and Nearctic regions, but straggling species occur also in +North India, South Africa, Chili, and New Zealand. _Thecla_ is especially +characteristic of the Neotropical region, where there are about 370 +species; in the Nearctic region, 36; in the Palæarctic 13; and in the +Ethiopian 3. _Miletus_, _Lucia_, _Hypolycæna_, _Myrina_, and _Deudorix_ are +common to the three tropical regions of the Eastern Hemisphere--the +Ethiopian, Oriental, and Australian. _Aphneus_ and _Iolaus_ are common to +the Ethiopian and Oriental regions, the latter extending to Celebes. +_Ialmenus_, _Pseudodipsas_, _Curetis_, and _Amblypodia_ are common to the +Oriental and Australian regions, but the first-named is found also in +Madagascar. _Zephyrus_ is found only in the Nearctic and Palæarctic, +_Eumæus_ in the Nearctic and Neotropical regions. The Nearctic region has +one peculiar genus (_Feniseca_); the Palæarctic has two--_Thestor_ and +_Læosopis_; the Ethiopian has nine--_Pentila_, _Liptana_, _D'Urbania_, +_Axiocerces_, _Capys_, _Phytala_, _Epitola_, _Hewitsonia_, and _Deloneura_; +the Oriental has five--_Allotinus_, _Ilerda_, _Poritia_, _Camena_, and +_Liphyra_; the Australian has three--_Hypochrysops_, _Utica_, and _Ogyris_; +and the Neotropical also three--_Lamprospilus_, _Theorema_, and +_Trichonis_. + + +{478}FAMILY 14.--PIERIDÆ. (35 Genera, 817 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3 -- + | | | | | + +The Pieridæ are distributed almost, if not quite, as widely over the globe +as the last family, and we shall group the genera in the same manner. +_Pieris_ (130 sp.) is cosmopolitan; _Terias_ and _Callidryas_ are found in +all the four tropical regions, and as far north as Pennsylvania in the +Nearctic region; _Pontia_, _Tachyris_, _Eronia_, and _Thestias_ are common +to the Ethiopian, Oriental, and Australian regions, the last-named, +however, only extending as far as Timor; _Colias_ is pre-eminently +Palæarctic and Nearctic, with a few Ethiopian species, one Indian, two in +Chili, and one in the Sandwich Islands; _Anthocharis_ is wholly Palæarctic +and Nearctic; _Midea_ has two species Nearctic, and one in Japan; +_Gonepteryx_ is Palæarctic and Neotropical, extending into Texas; _Idmais_ +and _Callosune_ are Ethiopian and Oriental; _Thyca_ and _Iphias_ are +Oriental and Australian; _Meganostoma_ is Nearctic and Neotropical; +_Nathalis_ and _Kricogonia_ are Neotropical, ranging into Florida, Texas, +and Colorado. + +The peculiar genera are pretty equally distributed. The Neotropical region +has ten, two being confined to Chili; _Euterpe_ and _Leptalis_ are the most +remarkable, the latter containing a number of forms mimicking the +Heliconidæ and Danaidæ. The Oriental region has two, _Prioneris_ and +_Dercas_, the Australian one, _Elodina_; the Ethiopian two, _Teracolus_ and +_Pseudopontia_; the Palæarctic two, _Leucophasia_ and _Zegris_; the +Nearctic one, _Neophasia_. + + +{479}FAMILY 15.--PAPILIONIDÆ. (13 Genera, 455 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Papilionidæ, comprising many of the noblest and richest-coloured +butterflies, and long placed at the head of the group, are almost as +universally distributed as the Pieridæ, but they do not extend to so many +remote islands nor so far into the Arctic and Antarctic regions. +Nine-tenths of the species belong to the genus _Papilio_, and these are +especially abundant in tropical regions, although species occur in every +region and every sub-region. Well-marked sub-divisions of this large genus +are characteristic of each great region--as the "Æneas" group in the +Neotropical, the "Paris" group in the Oriental, the "Ægeus" group in the +Australian, the "Zenobius" group in the Ethiopian, and many others. The few +species of the Palæarctic region belong, on the other hand, to a group of +universal distribution, and the Nearctic has a good number of species +allied to Neotropical forms. + +The other genera have mostly a very restricted range. _Parnassius_ is an +Alpine genus, confined to the Palæarctic and Nearctic regions. The +Palæarctic region further possesses 5 peculiar genera--_Mesapia_, +_Hypermnestra_, _Doritis_, _Sericinus_, and _Thais_; the Oriental has 4, +_Calinaga_, _Teinopalpus_, _Bhutanitis_, and _Leptocircus_, the latter +going as far as Celebes; the Australian has 1, _Eurycus_; and the +Neotropical 1, _Euryades_, confined to the Chilian sub-region. The +Ethiopian and the Nearctic regions have no peculiar genera. + + +{480}FAMILY 16.--HESPERIDÆ. (52 Genera (?), 1,200 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Hesperidæ, or Skippers, are an immense group of mostly small obscurely +coloured butterflies, universally distributed, and of which hosts of +species still remain to be discovered and described. As the grouping of +these into genera is not yet satisfactorily accomplished, only the more +extensive and best known groups will be here noticed. _Pamphila_ and +_Hesperia_ are universally distributed; _Nisoniades_ seems to be only +absent from the Australian region. The Neotropical region is pre-eminently +rich in Hesperidæ, 33 genera being found there, of which 20 are peculiar to +it; the Australian region has 12 genera, only 1 (_Euschemon_) being +peculiar; the Oriental has 18, with 3 peculiar; the Ethiopian, 13, with 3 +peculiar; the Palæarctic 6, with 1 (_Erynnis_) almost peculiar, a species +occurring in Mexico; the Nearctic 9, with none peculiar, 4 being found also +in the Neotropical region, 2 in the Palæarctic, and the rest being of wide +distribution. Many new genera have, however, been recently described in the +United States, but it is impossible yet to determine how many, if any, of +these are peculiar. More than 100 species of the family are included in Mr. +Edwards' "Synopsis of North American Butterflies,"--a very large number +considering that Europe possesses only about 30. + + +{481}Sub-order--LEPIDOPTERA HETEROCERA, or MOTHS. + +The Lepidoptera Heterocera, or Moths, are of such immense extent, and are, +besides, so imperfectly known compared with the Butterflies, that it would +serve no purpose to go into the details of their distribution; especially +as most of the families and a considerable number of the genera are +cosmopolitan. We propose therefore to notice only the Sphingina, which, +being generally of large size and finely marked or coloured, and many of +them day-fliers, have been extensively collected; and whose numbers are +more manageable than the succeeding groups. + + +_Group I.--SPHINGINA._ + + +FAMILY 17.--ZYGÆNIDÆ (46 Genera, about 530 Species). + +The Zygænidæ are universally distributed, but many of the genera are +restricted in their range. _Zygæna_ (85 sp.) is mainly Palæarctic, but 2 +species are South African, and 1 North American; _Procris_ (22 sp.) has a +scattered distribution, from the Palæarctic region to South America, South +Africa and North India; _Heterogynis_ (3 sp.) and _Dysauxis_ (3 sp.) are +European; _Pollanisus_ (3 sp.) is Australian; _Glaucopis_ (120 sp.) is +mainly Neotropical, with a few Oriental; _Syntomis_ (94 sp.) is found in +all the Old-World regions; and _Euchromia_ (150 sp.) is found in all warm +countries, though especially abundant in South America. + + +FAMILY 18.--CASTNIIDÆ (7 Genera, 63 Species). + +The Castniidæ have an interesting distribution, being mainly Neotropical, +with four genera in Australia and New Guinea. _Castnia_, _Coronis_, and +_Gazera_, with 51 species, are Neotropical; _Synemon_, _Euschemon_, +_Damias_ and _Cocytia_, with 12 species, are Australian, the latter being +found only in the Papuan Islands. + + +{482}FAMILY 19.--AGARISTIDÆ (13 Genera, 76 Species). + +The Agaristidæ are beautiful diurnal moths, allied to the Castniidæ, but +almost confined to the Australian and Oriental regions, with a few in the +Ethiopian. The most important genera are,--_Agarista_ (21 sp.), Australia +and New Guinea; _Eusemia_ (31 sp.), _Ægocera_ (7 sp.), Oriental and +Ethiopian regions; the other genera being confined to the islands from Java +to New Guinea. + + +FAMILY 20.--URANIIDÆ (2 Genera, 12 Species). + +These magnificent insects have a singular distribution. The gold-spangled +_Urania_ (6 sp.) is characteristic of Tropical America, but a single +species of great magnificence occurs in Madagascar. The large but +sober-tinted _Nyctalemon_ (6 sp.) is found in the Neotropical, Oriental, +and Australian regions. + + +FAMILY 21.--STYGIIDÆ. (3 Genera, 14 Species.) + +These insects are confined to the Palæarctic and Neotropical regions, 2 +genera in the former, 1 in the latter. + + +FAMILY 22.--ÆGERIIDÆ. (24 Genera, 215 Species.) + +This family is found in all parts of the world except Australia. _Ægeria_ +is most abundant in Europe, but is found also in North and South America. + + +FAMILY 23.--SPHINGIDÆ. (40 Genera, 345 Species.) + +The Sphinx Moths are cosmopolitan. The most important genera +are,--_Macroglossa_ (26 sp.), _Chærocampa_ (46 sp.), and _Macrosila_ (21 +sp.), all cosmopolitan; _Sesia_ (12 sp.), Europe, Asia, and North America; +_Deilephila_ (19 sp.), Palæarctic and Oriental regions, Nearctic region, +and Chili; _Sphinx_ (21 sp.), Europe, {483}North and South America; +_Smerinthus_ (29 sp.), all regions except Australia. Our Death's Head Moth +(_Acherontia atropos_) ranges to Sierra Leone and the Philippine Islands. + + +_General Remarks on the Distribution of the Diurnal Lepidoptera and +Sphingidea._ + +The Diurnal Lepidoptera or Butterflies, comprehend 431 genera and 7,740 +species, arranged in 16 families, according to Mr. Kirby's Catalogue +published in 1871. The Sphingidea consist of 135 genera and 1,255 species, +arranged in 7 families, according to the British Museum Catalogue dated +1864; and as this includes all Mr. Bates' collections in America and my own +in the East, it is probable that no very large additions have since been +made. + +The distribution of the families and genera of Butterflies corresponds +generally with that of Birds--and more especially with that of the +Passerine birds--in showing a primary division of the earth into Eastern +and Western, rather than into Northern and Southern lands. The Neotropical +region is by far the richest and most peculiar. It possesses 15 families of +butterflies, whereas the other regions have only from 8, in the Palæarctic, +to 12 in the Ethiopian and Oriental regions; and as none of the Old World +regions possess any peculiar families, the New World has a very clear +superiority. In genera the preponderance is still greater, since the +Neotropical region possesses about 200 altogether peculiar to it, out of a +total of 431 genera, many of which are cosmopolitan. Comparing, now, the +Eastern regions with the Western, we have two peculiar families in the +former to 4 in the latter; while the Southern regions (Australian and +Neotropical) possess not a single peculiar family in common. + +In the Sphingidea the same general features recur in a less marked degree, +the Neotropical being the richest region; but here we have one family +(Castniidæ) which appears to be confined to the two southern regions,--the +Australian and Neotropical. + +The distribution of the genera affords us some facts of special interest, +which must be briefly noticed. There are several {484}genera typically +characteristic of the North Temperate regions which have a few species +widely scattered on mountains, or in the temperate parts of the Southern +Hemisphere. Chili possesses representatives of four of these +genera--_Argynnis_, _Lycæna_, _Colias_, and _Deilephila_; and this has been +thought by some naturalists to be of such importance as to outweigh the +purely Neotropical character of a large portion of the Chilian fauna, and +to render it advisable to join it on, as an outlying portion of a great +North Temperate zoological region. But when we remember that _Argynnis_ +occurs also in Java, and _Lycæna_ in New Zealand, while _Colias_ ranges to +Southern Africa, Malabar, and the Sandwich Islands, we can hardly admit the +argument to be a sound one. For a fuller discussion of this question see +Vol. II., pp. 43-47. The remarkable fact of the existence of the otherwise +purely Neotropical genus, _Urania_, in Madagascar is even more striking, +supported as it is by the Antillean, _Solenedon_, belonging to a family of +Mammalia otherwise confined to Madagascar, and by one or two Coleopterous +genera, to be noticed farther on as common to the two countries. Our view +as to the true explanation of this and analogous phenomena will be found at +Vol. I., p. 284. + +The division of the Castniidæ (a family almost confined to the Tropics), +between the Neotropical and Australian regions, is also a very curious and +important phenomenon, because it seems to point to a more remote connection +between the two countries than that indicated by the resemblance between +the productions of South Temperate America with those of Australia and New +Zealand; but we have already shown that the facts may be explained in +another way. (See Vol. I., pp. 398 and 404). + +The division of the Malay Archipelago between the Oriental and Australian +regions is clearly marked in the Lepidoptera, and it is very curious that +it should be so, for in this, if in any group of animals, we should expect +an almost complete fusion to have been effected. Lepidoptera fly readily +across wide tracts of sea, and there is absolutely no climatal difference +to interfere with their free migration from island to island. Yet we find +no less than 10 genera abundant in the Indo-Malayan {485}sub-region which +never cross the narrow seas to the east of them; 6 others which only pass +to Celebes; and 2 more which have extended from Java along the closely +connected line of islands eastwards to Timor. On the other side, we find 5 +strictly Austro-Malayan genera, and 2 others which have a single +representative in Java. The following is a list of these genera:-- + +INDO-MALAYAN GENERA:--_Amathusia_, _Thaumantis_, _Tanæcia_, _Eurytela_, +_Ilerda_, _Zemeros_, _Taxila_, _Aphneus_, _Prioneris_, _Dercas_, _Clerome_, +_Adolias_, _Apatura_, _Limenitis_, _Iolaus_, _Leptocircus_, (the last six +reach Celebes); _Discophora_, _Thestias_; (the last two reach Timor.) + +AUSTRO-MALAYAN GENERA:--_Hamadryas_, _Hypocista_, _Mynes_, _Dicallaneura_, +_Elodina_, _Hyades_, _Prothoë_ (the last two reach Java). + +The most characteristic groups, which range over the whole Archipelago and +give it a homogeneous character, are the various genera of Danaidæ, the +genus _Elymnias_, and _Amblypodia_ with a few other Lycænidæ. These are all +abundant and conspicuous groups, but they are nevertheless exceptions to +the general rule of limitation to one or other of the regions. The cause of +this phenomenon is probably to be found in the limitation of the larvæ of +many Lepidoptera to definite species, genera, and families of plants; and +we shall perhaps find, when the subject is carefully investigated, that the +groups which range over the whole Archipelago feed on genera of plants +which have an equally wide range, while those which are limited to one +region or the other, have food-plants belonging to genera which are +similarly limited. It is known that the vegetation of the two regions +differs largely in a botanical sense, although its general aspect is almost +identical; and this may be the reason why the proportion of wide-ranging +genera is greater among such insects as feed upon dead wood, than among +those which derive their support from the juices of the living foliage. +This subject will be again discussed under the various families of +Coleoptera, and it will be well to bear in mind the striking facts of +generic limitation which have been here brought forward. + +{486}Fossil Butterflies, apparently of existing genera, occur in the +Miocene and Eocene formations, and an extinct form in the Lower Oolite; but +these cannot be held to give any adequate idea of the antiquity of so +highly specialised a group, which, in all probability, dates back to +Palæozoic times, since one of the Bombycidæ,--a group almost as +highly-organised--has been discovered in the coal formation of Belgium. +(See Vol. I. p. 168.) + + +_Order--COLEOPTERA._ + +GEODEPHAGA, or CARNIVOROUS GROUND BEETLES. + +The Geodephaga consist of two families, Cicindelidæ and Carabidæ, differing +in their form and habits no less than in their numbers and distribution. +The former, comprising about 800 species, are far more abundant and varied +in Tropical regions; the latter, more than ten times as numerous, are +highly characteristic of the North Temperate zone, where fully half of all +the known species occur. + + +CICINDELIDÆ. (35 Genera, 803 Species.) + +The Cicindelidæ, or Tiger Beetles, are a moderately extensive group, spread +over the whole globe, but much more abundant in tropical than in temperate +or cold countries. More than half of the species (418) belong to the single +genus _Cicindela_, the only one which is cosmopolitan. The other large +genera are,--_Collyris_ (81 sp.), wholly Oriental; _Odontochila_ (57 sp.), +South American, with species in Java and Celebes; _Tetracha_ (46 sp.), +mostly South American, but with species in South Europe, North America, and +Australia; _Tricondyla_ (31 sp.), characteristic of the Oriental region, +but extending eastward to New Guinea; _Ctenostoma_ (26 sp.), wholly +Neotropical; _Dromica_ (24 sp.), wholly African, south of Lake Ngami and +Mozambique; _Therates_ (18 sp.), wholly Malayan, from Singapore to New +Guinea. + +The genera are distributed in the several regions as follows:--the Nearctic +region has 5 genera, 3 of which are peculiar to it; the {487}Palæarctic has +2, but none peculiar; the Ethiopian 13, with 11 peculiar; the Oriental 8, +with 3 peculiar; the Australian 9, with 2 peculiar; and the Neotropical 15, +with 10 peculiar. The connection between South America and Australia is +shown by the latter country possessing 9 species of the characteristic +South American genus _Tetracha_, as well as one of _Megacephala_. The small +number of peculiar genera in the Oriental and Australian regions is partly +owing to the circumstance that two otherwise peculiar Oriental genera have +spread eastward to the Moluccas and New Guinea, a fact to be easily +explained by the great facilities such creatures have for passing narrow +straits, and by the almost identical physical conditions in the Malayan +portion of the two regions. The insects of Indo-Malaya were better adapted +to live in the Austro-Malay Islands than those of Australia itself, and the +latter group of islands have thus acquired an Oriental aspect in their +entomology, though not without indications of the presence of an aboriginal +insect-fauna of a strictly Australian type. The relation of the Australian +and Neotropical regions is exhibited by this family in an unusually +distinct manner. _Tetracha_, a genus which ranges from Mexico to La Plata, +has 9 species in Australia; while _Megacephala_ has 2 American and 1 +Australian species. Another curious, and more obscure relation, is that +between the faunas of Tropical America and Tropical Africa. This is also +illustrated by the genus _Megacephala_, which has 4 African species as well +as 2 South American; and we have also the genus _Peridexia_, which has 2 +species in South America and 2 in Madagascar. + +Several of the sub-regions are also well characterised by peculiar genera; +as _Amblychila_ and _Omus_ confined to California and the Rocky Mountains; +_Manticora_, _Ophryodera_, _Platychile_ and _Dromica_, characteristic of +South Africa; _Megalomma_ and _Pogonostoma_ peculiar to the Mascarene +Islands; and _Caledonica_ to the islands east of New Guinea. The extensive +and elegant genus _Collyris_ is highly characteristic of the Oriental +region, over the whole of which it extends, only just passing the limits +into Celebes and Timor. + +The Cicindelidæ, therefore, fully conform to those divisions of {488}the +earth which have been found best to represent the facts of distribution in +the higher animals. + + +CARABIDÆ. (620 Genera, 8500 Species.) + +The enormous extent of this family, necessitates a somewhat general +treatment. It has been very extensively collected, while its classification +has been most carefully worked out, and a detailed exposition of its +geographical distribution by a competent entomologist would be of the +greatest interest. A careful study of Gemminger and Harold's Catalogue, +however, enables me to sketch out the main features of its distribution, +and to detail many of its peculiarities with considerable accuracy. + +The Carabidæ are remarkable among insects, and perhaps among all +terrestrial animals, as being a wonderfully numerous, varied, conspicuous, +and beautiful group, which is pre-eminently characteristic of the +Palæarctic region. So strikingly and unmistakably is this the case, that it +must be held completely to justify the keeping that region distinct from +those to which it has at various times been proposed to join it. Although +the Carabidæ are thoroughly well represented by hosts of peculiar genera +and abundant species in every part of the world without exception, yet the +Palæarctic region alone contains fully one-third, or perhaps nearer +two-fifths, of the whole. It may also be said, that the group is a +temperate as compared with a tropical one; so that probably half the +species are to be found in the temperate and cold regions of the globe, +leaving about an equal number in the much more extensive tropical and warm +regions. But, among the cold regions, the Palæarctic is pre-eminent. North +America is also rich, but it contains, by far, fewer genera and fewer +species. + +The magnificent genus _Carabus_, with its allies _Procerus_ and +_Procrustes_, containing about 300 species, all of large size, is almost +wholly confined to the Palæarctic region, only 10 species inhabiting North +America, and 11 Temperate South America, with one on the African mountain +of Kilimandjaro. Twelve large genera, containing together more than 2000 +species, are truly cosmopolitan, inhabiting both temperate and tropical +{489}countries all over the globe; but many of these are more abundant in +the Palæarctic region than elsewhere. Such are _Scarites_, _Calosoma_, +_Brachinus_, _Cymindis_, _Lebia_, _Chlænius_, _Platynus_, _Harpalus_, +_Bembecidium_, _Pæcilus_, and _Argutor_. Of tropical cosmopolites, or +genera found in all the tropical regions, but not in the temperate zones, +there seem to be only four,--_Catascopus_, _Coptodera_, _Colopodes_, and +_Caasnonia_. _Pheropsophus_ is confined to the tropics of the Old World; +while _Drimostoma_, though widely scattered, is characteristic of the +Southern Hemisphere. + +The Palæarctic region has about 50 genera of Carabidæ which are strictly +confined to it, the most important being,--_Leistus_ (30 sp.), _Procerus_ +(5 sp.), _Procrustes_, (17 sp.), _Zabrus_ (60 sp.), _Pristonychus_ (42 +sp.), and _Ophonus_ (60 sp.); but it possesses a large number in common +with the Nearctic region. The more remarkable of these are,--_Carabus_, +_Nebria_, _Amara_, _Cyrtonotus_, _Bradycellus_, _Anopthalmus_, _Celia_, +_Cychrus_, _Patrobus_, _Elaphrus_, _Notiophilus_, _Bradytus_, +_Callisthenus_, _Blethisa_, and several others. Many too, though not +strictly confined to the North Temperate regions, are very abundant there, +with a few species isolated in remote countries, or widely scattered, often +in an eccentric manner. Among these may be mentioned, _Trechus_ (120 sp.), +all North Temperate but 8, which are scattered in Java, New Caledonia and +South America; _Dyschirus_ (127 sp.), North Temperate, with 3 or 4 species +in Australia, China and La Plata; _Omaseus_, (88 sp.), _Steropus_ (90 sp.), +_Platysoma_ (114 sp.), and _Pterostichus_ (138 sp.), are mostly North +Temperate, but each has a few species in the South Temperate zone, New +Zealand, Australia, Chili, and the Cape of Good Hope. _Dromius_ (54 sp.), +is about two-thirds Palæarctic, the rest of the species being scattered +over the world, in Chili, North and South America, South Africa, Burmah, +Ceylon, and New Zealand. The North Temperate genera _Calathus_ and +_Olisthopus_, have each one species in New Zealand; _Percus_ has most of +its species in South Europe, but 3 in Australia; _Abax_ is confined to the +north temperate zone, but with one species in Madagascar; while +_Læmosthenes_ is said to have a species identically the same in South +Europe and Chili. Some of these apparent anomalies may be due to wrong +{490}determination of the genera, but there can be little doubt that most +of them represent important facts in distribution. + +The Nearctic region is comparatively poor in Carabidæ. Its more important +peculiar genera are,--_Dicælus_ (22 sp.), _Pasimachus_ (17 sp.), +_Eurytrichus_ (9 sp.), _Sphæroderus_ (7 sp.), _Pinacodera_ (6 sp.), and +others of smaller extent, about 30 in all. It also possesses +representatives of a considerable number of Palæarctic genera, as already +indicated; and a few of South American genera, of which _Helluomorpha_ and +_Galerita_ are the most important. + +The Neotropical region is very rich in peculiar forms of Carabidæ, as in +almost all other great groups. It possesses more than 100 peculiar genera, +but about 30 of these are confined to the South Temperate sub-region. The +more important peculiar genera of Tropical America are,--_Agra_ (144 sp.), +_Ardistomus_ (44 sp.), _Schizogenius_ (25 sp.), _Pelecium_, (24 sp.), +_Calophena_ (22 sp.), _Ctenodactyla_ (7 sp.). Among the Chilian and South +Temperate peculiar forms are,--_Antarctia_ (29 sp.), _Scelodontis_ (10 +sp.), _Tropidopterus_ (4 sp.). Among the Neotropical genera with outlying +species are,--_Pachyteles_ (50 sp.), one of which is West African; +_Selenophorus_ (70 sp.), with 4 African, 4 Oriental, and 1 from New +Caledonia; _Ega_ (11 sp), with one in the East Indies, and one in New +Caledonia; _Galerita_, with 36 American species, 8 African, and 3 Indian; +_Callida_ and _Tetragonoderus_, mostly American, but with a few African, +Oriental and Australian species; and _Pseudomorpha_, common to America and +Oceania. + +The Australian region is almost equally rich, possessing about 95 peculiar +genera of Carabidæ, no less than 20 of which are confined to New Zealand. +The most important are, _Carenum_, _Promecoderus_, _Scaraphites_, +_Notonomus_, _Ænigma_, _Sphallomorpha_, _Silphomorpha_, and _Adelotopus_. +The gigantic _Catadromus_ has 4 Australian species and 1 in Java; +_Homalosoma_ has 31 species in Australia and New Zealand, and 1 in +Madagascar. Celebes and New Guinea have each peculiar genera, and one is +common to Australia and the Cape of Good Hope. + +The Oriental region possesses 80 peculiar genera, 10 of which are confined +to Ceylon. The more important are,--_Pericallus_, _Planetes_, and +_Mormolyce_. _Distrigus_ is also characteristic of this {491}region, with +one species in Madagascar; while it has _Orthogonius_, _Hexagonia_, +_Macrochilus_, and _Thyreopterus_ in common with the Ethiopian region, and +is rich in the fine tropical genus, _Catascopus_. + +The Ethiopian region has 75 peculiar genera, 8 of which are confined to +Madagascar. The more important are,--_Polyhirma_, _Graphipterus_, and +_Piezia_. _Anthia_ is chiefly African, with a few species in India; +_Abacetus_ is wholly African, except a species in Java, and another in +South Europe; and _Hypolithus_ is typically African, but with 7 species in +South America and 1 in Java. + +The facts of distribution presented by this important family, looked at +broadly, do not support any other division of the earth into primary +regions than that deduced from a study of the higher animals. The amount of +speciality in each of these regions is so great, that no two of them can be +properly united; and in this respect the Carabidæ accord wonderfully with +the Vertebrates. In the details of distribution there occur many singular +anomalies; but these are not to be wondered at, if we take into +consideration the immense antiquity of Coleopterous insects--which existed +under specialised forms so far back as the Carboniferous epoch,--the ease +with which they may be dispersed as compared with larger animals, and the +facilities afforded by their small size, habits of concealment, and often +nocturnal habits, for adaptation to the most varied conditions, and for +surviving great changes of surface and of the surrounding organic forms. +The wonder rather is, not that there are so many, but so few cases of +exceptional and anomalous distribution; and the fact that these creatures, +so widely different from Vertebrates in organisation and mode of life, are +yet on the whole subject to the same limitations of range as were found to +occur among the higher animals, affords a satisfactory proof that the +principles on which our six primary regions are founded, are sound; and +that they are well adapted to exhibit the most interesting facts of +geographical distribution, among all classes of animals. + +Much stress has been laid on the fact of a few species of such typical +European genera as _Carabus_, _Dromius_, and others, being {492}found in +Chili and Temperate South America; and it has been thought, that in a +system of Entomological regions this part of the world must be united to +the Northern Hemisphere. But these writers omit to take into account, +either the large numbers of isolated and peculiar forms characteristic of +South Temperate America, or the indications of affinity with Tropical +America and Australia, both of which are really more important than the +connection with Europe. The three important Chilian genera, _Cascelius_, +_Barypus_, and _Cardiopthalmus_, are closely allied to the Australian +Promecoderus; others, as _Omostenus_ and _Plagiotelium_, are quite +isolated; while _Antarctia_ and _Metius_, according to Lacordaire, form a +distinct division of the family. Chili, too, has many species of +_Pachyteles_, _Coptodera_, and other South American genera; and this +affinity is far stronger in many other families than in the Carabidæ. The +existence of representatives of typical northern forms in Chili, is a fact +of great interest, and may be accounted for in a variety of ways; (see Vol. +II. p. 44) but it is not of such a magnitude as to be of primary importance +in geographical distribution, and it can only be estimated at its fair +value, by taking into account the affinities of all the groups inhabiting +that part of the world. + + +LUCANIDÆ. (45 Genera, 529 Species.) + +Passing over a number of obscure families, we come to the remarkable group +of the Lucanidæ, or Stag-beetles, which, being almost all of large size, +and many of them of the most striking forms, have been very thoroughly +collected and assiduously studied. + +The most curious feature of their general distribution, is their scarcity +in Tropical South America, and their complete absence from Tropical North +America and the West Indian Islands, though they appear again in Temperate +North America. In the New World they may, in fact, be looked upon as a +temperate group characteristic of the extra-tropical regions and the +highlands; while in the Old World, where they are far more abundant, they +are distinctly tropical, being especially numerous {493}in the Oriental and +Australian regions. No genus has the range of the whole family, _Dorcus_ +and _Lucanus_ being absent from Africa, while _Cladognathus_ is unknown in +the New World and on the continent of Australia. The Oriental region is the +richest in peculiar forms, possessing 16 genera, 7 of which are wholly +confined to it, while 3 others only just range beyond it to North China on +the one side, or to the Austro-Malayan islands on the other. The Australian +region comes next, with 15 genera, of which 7 are wholly peculiar. South +America has 12 genera, 10 of which are peculiar. The Ethiopian region has +10 genera, 7 of which are peculiar, and 2 of these are confined to the +island of Bourbon. The Palæarctic region has 8 genera, and the Nearctic 5; +one genus being peculiar to Europe, and two confined to Europe and North +America. The Ethiopian and Oriental regions have 3 genera in common and +peculiar to them; the Oriental and Australian 3; while the Australian and +Neotropical have 1 in common, to which may be added _Streptocerus_, which +represents in Chili the Australian _Lamprima_. + +Among the special features presented by the distribution of the Lucanidæ, +may be mentioned--the remarkable group of genera, _Pholidotus_, +_Chiasognathus_, and _Sphenognathus_, confined to Temperate South America, +the Andes, and mountains of Brazil; _Lucanus_ (19 sp.), almost confined to +the Oriental and Palæarctic regions, three species only inhabiting North +America; _Odontolabris_ (29 sp.), wholly Oriental, with 2 sp. in Celebes; +_Nigidius_ (11 sp.), Ethiopian, but with species in Formosa, the +Philippines, and Malacca; _Syndesus_ (11 sp.), common to Australia, New +Caledonia, and South America; _Figulus_ (20 sp.), divided between Africa +and Madagascar on the one hand, and Australia, with the Malay and Pacific +Islands, on the other. + +The facts of distribution here sketched out are in perfect accordance with +those of many groups of Vertebrates. The regions are sharply contrasted by +their peculiar and characteristic genera; the several relations of those +regions are truly indicated; while there is a comparatively small +proportion of cases of anomalous or eccentric distribution. + + +{494}CETONIIDÆ. (120 Genera, 970 Species.) + +As representative of the enormous group of the Lamellicorns, which, +according to continental entomologists, forms a single family numbering +nearly 7,000 species, we take the Cetoniidæ or Rose-Chafers. These comprise +a number of the most brilliant and beautifully-coloured insects, including +the gigantic _Goliathi_, which are among the largest of known beetles. They +have been assiduously collected in every part of the world, and their +classification has been elaborated by many of our most eminent +entomologists. + +The Cetoniidæ are especially abundant in tropical and warm countries, yet +far more so in the Old World than in the New; and in the Old World, the +Ethiopian region exhibits a marvellous richness in this family, no less +than 76 genera being found there, while 64, or more than half the total +number, are peculiar to it. Next in richness, though still very far behind, +comes the Oriental region, with 29 genera, 17 of which are peculiar. The +Neotropical has only 14 genera, but all except two are peculiar to it, and +one of these is not found out of the New World. The Australian region has +11 genera, three only being peculiar. The Palæarctic region has 13, with 4 +peculiar; the Nearctic 7, with 2 peculiar. The affinities of the regions +for each other, as indicated by the genera confined to two adjacent +regions, are in this family somewhat peculiar. The Ethiopian and Oriental +show the most resemblance, 6 genera being common and peculiar to the two; +the Oriental and the Australian are unusually well contrasted, having only +one genus exclusively in common, while 8 genera are found in the Indo-Malay +Islands which do not cross the boundary to the Austro-Malayan division, and +several others only pass to the nearest adjacent islands; on the other +hand, the only large Australian genus, _Schizorhina_, is found in many +parts of the Moluccas, but not further west. The Australian and Neotropical +regions exhibit no direct affinity, the nearest ally to the South American +Gymnetidæ being _Clinteria_, an African and Asiatic genus; while not a +single genus is common {495}to Australia and South America. The Nearctic +and Palæarctic regions have 3 genera in common, which are found in no other +part of the world. + +Among the special features of interest connected with the distribution of +this family, we must first notice the exceptional richness of Madagascar, +which alone possesses 21 peculiar genera. South Africa is also very rich, +having 8 peculiar genera. _Stethodesma_ is very peculiar, being divided +between South America and Mexico on the one hand, and West and South Africa +on the other. _Stalagmosoma_ is a desert genus, ranging from Persia to +Dongola. No genus is cosmopolitan, or even makes any approach to being so, +except _Valgus_, which occurs in all the regions except the Neotropical; +and even the family seems to be not universally distributed, since no +species are recorded either from New Zealand, the Pacific Islands, or the +Antilles. + +The facts here brought forward, lead us to the conclusion that the +Cetoniidæ are an Old-World tropical family, which had been well developed +in Africa and Asia before it spread to Australia and America; and that it +is only capable of being freely dispersed in the warmer regions of the +earth. This view will explain the absence of affinity between the +Australian and Neotropical regions, the only closer connection between +which, has almost certainly occurred in the colder portions of the +Temperate zone. + + +BUPRESTIDÆ. (109 Genera, 2,686 Species.) + +The next family suited to our purpose is that of the Buprestidæ, consisting +as it does of many large and some gigantic species, generally adorned with +brilliant metallic colours, and attracting attention in all warm countries. +Although these insects attain their full development of size and beauty +only in the Tropics, they are not much less abundant in the warmer parts of +the Temperate zone. In the Catalogue of the Coleoptera of Europe and the +Mediterranean Basin, by M. de Marseul (1863), we find 317 species of +Buprestidæ enumerated, although {496}the district in question only forms a +part of the Palæarctic region, which would thus seem to possess its full +proportion of the species of this family. Confining ourselves to the +generic forms, we find far less difference than usual between the numbers +possessed by the tropical and the temperate regions; the richest being the +Australian, with 47 genera, 20 of which are peculiar; and the poorest the +Nearctic, with 24 genera, of which 7 are peculiar. The Oriental has 41 +genera, 14 of which are peculiar; the Neotropical 39, of which the large +proportion of 18 are peculiar; the Ethiopian 27, of which 6 are peculiar; +and the Palæarctic also 27, but with 9 peculiar. + +A most interesting feature in the distribution of this family, is the +strong affinity shown to exist between the Australian and Neotropical +regions, which have 4 genera common to both and found nowhere else; but +besides this, the extensive and highly characteristic Australian genus, +_Stigmodera_, is closely related to a number of peculiar South American +genera, such as _Conognatha_, _Hyperantha_, _Dactylozodes_,--the last +altogether confined to Chili and Temperate South America. Here we have a +striking contrast to the Cetoniidæ, and we can hardly help concluding, +that, as the latter is typically a tropical group, so the present family, +although now so largely tropical, had an early and perhaps original +development in the temperate regions of Australia, spreading thence to +Temperate South America as well as to the tropical regions of Asia and +Africa. The Australian and Oriental regions have 4 genera exclusively in +common, but they also each possess a number of peculiar or characteristic +genera, such as the Indo-Malayan _Catoxantha_ (which has only a single +species in the Moluccas) and nine others of less importance; and the +exclusively Austro-Malayan genus, _Sambus_, with five smaller groups, and +_Cyphogastra_, with only 2 Indo-Malay species. The Oriental and Ethiopian +regions are very distinct, only possessing the single genus, _Sternocera_, +exclusively in common. The Nearctic and Palæarctic are also distinct, only +one genus, _Dicerca_, being confined to America (North and South) and +Europe, a fact which again points to a southern origin for this family, and +its comparatively recent extension into the {497}North Temperate zone. It +must be remembered, however, that in view of the immense geological +antiquity of the existing families of Beetles, dating back certainly to the +Secondary and probably to the Palæozoic epoch, "comparatively recent" may +still be of considerable antiquity. + +It is somewhat singular that North and South America have no genera +exclusively in common. The connection between South America and Africa +seems to be shown,--by the genus _Psiloptera_, the mass of the species +being divided between these regions, with a few widely scattered over the +globe; and the American genus _Actenodes_, which has one species in West +Africa. Somewhat allied, is the extensive genus _Polybothris_, strictly +confined to Madagascar. The genus _Agrilus_ is perhaps cosmopolitan, +although no species of the family is recorded from New Zealand. Among the +peculiarities of distribution we may notice,--the genus _Sponsor_, with 8 +species in the island of Mauritius, 1 in Celebes, and 1 in New Guinea; +_Ptosima_, scattered between the United States, Mendoza in South Temperate +America, South Europe, the Philippine Islands, and North China; +_Polycesta_, which besides inhabiting South America, North America, and +Europe, has a single species in Madagascar; and _Belionota_, which has 8 +species African, 8 Indo-Malayan, 2 Austro-Malayan, and 1 in California. The +extensive genus _Acmæodera_, is most abundant in the warm and dry portions +of the Palæarctic, Ethiopian, and Nearctic regions, with some in the Andes +and South Temperate America, a few in Brazil and the West Indies, and 1 +said to be from the Philippines. About one-third of the genera (containing +more than half the species) have a tolerably extensive range, while the +genera confined to single regions contain only about one-fourth of the +total number of species. + +It will, I think, be admitted, after a careful study of the preceding +facts, that the regions and sub-regions here adopted, serve to exhibit, +with great clearness, the chief phenomena of distribution presented by this +interesting family. + + +{498}LONGICORNIA. (1,488 Genera, 7,576 Species). + +The elegant and admired group of the Longicorn Beetles, is treated by +continental authors as a single family, consisting of three +sub-divisions--the Prionidæ, Cerambycidæ, and Lamiidæ of English +entomologists. These are so closely related, and are so similar in form, +habits, and general distribution, that it will be best to consider the +whole as one group, noticing whatever peculiarities occur in the separate +divisions. The endless structural differences among these insects, have led +to their being classed in an unusual number of genera, which average little +more than 5 species each; a number far below that in any of the other +families we have been considering, and probably below that which obtains in +any of the more extensive groups of animals or plants. This excessive +subdivision of the genera, a large number of which consist of only one or +two species, renders it difficult to determine with precision the relations +of the several regions, since the affinities of these genera for each other +are in many cases undetermined. A group of such enormous extent as this, +can only be properly understood after years of laborious study; we must +therefore content ourselves with such results as may be obtained from a +general survey of the group, and from a comparison of the range of the +several genera, by means of a careful tabulation of the mass of details +given in the recent Catalogue of Messrs. Gemminger and Harold and the noble +work of Lacordaire. + +The proportionate extent of the three families of Longicorns is very +unequal; the Prionidæ comprising about 7 per cent., the Cerambycidæ 44 per +cent., and the Lamiidæ 49 per cent. of the total number of species; and the +genera are nearly in the same proportions, being almost exactly 10, 40, and +50 per cent. of the whole, respectively; or, 135 Prionidæ, 609 Cerambycidæ, +and 746 Lamiidæ. The several regions, however, present marked differences +in their proportions of these families. In the two North Temperate regions, +the Cerambycidæ are considerably more numerous than the Lamiidæ, in the +proportion of about 12 to {499}9; and in this respect the Neotropical +region agrees with them, though the superiority in the proportion of +Cerambycidæ is somewhat less. In the Old World tropical regions, however, +and in Australia, the Lamiidæ greatly preponderate--being nearly double in +the Oriental and Ethiopian regions (or as 11 to 6), while in the Australian +it is as 6 to 5. The Prionidæ show a similar difference, though in a less +degree; being proportionately more numerous in the North Temperate and +Neotropical regions. Now, as regards the North Temperate regions, this +difference can be, to some extent explained, by a difference in the habits +of the insects. The Lamiidæ, which both in the larva and perfect state have +exceedingly powerful jaws, exclusively frequent timber trees, and almost +always such as are dead; while the Cerambycidæ, are generally more delicate +and have weaker mandibles, and many of the species live on shrubs, dead +twigs, foliage, and even on flowers. The immense superiority of the Tropics +in the number and variety of their timber trees, and the extent of their +forests, sufficiently accounts for their superiority to the Temperate +regions in the development of Lamiidæ; but the great excess of Cerambycidæ +in South America as compared with the rest of the Tropics, is not to be so +readily explained. + +Bearing in mind the different proportions of the families, as above noted, +we may now consider the distribution of the Longicorns as a whole. In +number of generic forms, the Neotropical region, as in so many other +groups, has a marked superiority. It possesses 516 genera, 489 of which (or +about 19/20 of the whole) are peculiar to it. The Australian and Oriental +regions come next, and are exactly equal, both possessing 360 genera, and +having almost exactly the same proportion (in each case a little less than +¾) peculiar. The Ethiopian region has 262 genera, with about 5/6 peculiar; +the Palæarctic 196, with 51 (rather more than ¼) peculiar; and the Nearctic +111, with 59 (a little more than half) peculiar. The more isolated of the +sub-regions are also well characterised by peculiar genera. Thus, Chili +with Temperate South America possesses 37, a large proportion being +Cerambycidæ; the Malagasi group 26, {500}with a preponderance of Lamiidæ; +and New Zealand 12, of which the Cerambycidæ are only slightly in excess. + +The relations between the Longicorn fauna of the several regions, are such +as are in accordance with the dependence of the group on a warm climate and +abundant vegetation; and indicate the efficiency of deserts and oceans as +barriers to their migration. The Neotropical and Australian regions have +only 4 genera in common, but these are sufficient to show, that there must +probably once have been some means of communication between the two +regions, better adapted to these insects than any they now possess. The +Nearctic and Neotropical regions have 5, and the Nearctic and Palæarctic 13 +genera in common and peculiar to them, the latter fact being the most +remarkable, because no means of inter-communication now exists, except in +high latitudes where the species of the Longicorns are very few. The +Oriental and Australian regions, on the other hand, are closely connected, +by having no less than 52 genera of Longicorns in common and peculiar to +them. Most of these are specially characteristic of the Malay Archipelago, +often extending over all the islands from Sumatra to New Guinea. This large +number of wide-spread genera of course gives a character of uniformity to +the entire area over which they extend; and, with analogous facts occurring +in other families, has led many entomologists to reject that division of +the Archipelago between the Australian and Oriental regions, which has been +so overwhelmingly demonstrated to be the natural one in the case of the +higher animals. The general considerations already advanced in Chapter II. +enable us, however, to explain such anomalies as this, by the great +facilities that exist for the transfer from island to island of such small +animals, so closely connected with woody vegetation in every stage of their +existence. That this is the true and sufficient explanation, is rendered +clear by certain additional facts, which those who object to the sharp +division of the Indo-Malay and Austro-Malay sub-regions have overlooked. + +An analysis of all the Malay Longicorns proves, that besides the 52 genera +characteristic of the Archipelago as a whole, there are 100 genera which +are confined to one or other of its component {501}sub-regions. Many of +these, it is true, consist of single species confined to a single island, +and we will not lay any stress on these; but there are also several +important groups, which extend over the Indo-Malay or the Austro-Malay +islands only, stopping abruptly at the dividing-line between them. For +example, on the Indo-Malay side we have _Euryarthrum_, _Leprodera_, +_Aristobia_, _Coelosterna_, and _Entelopes_, and what is perhaps even more +satisfactory, the large genera _Agelasta_ and _Astathes_, abundant in all +the Indo-Malay islands, but having only one or two species just passing the +boundary into Celebes. On the other side we have _Tethionea_, +_Sphingnotus_, _Arrhenotus_, _Tmesisternus_ (the last three genera +abounding from New Guinea to Celebes, but totally unknown further west), +_Hestima_, _Trigonoptera_, _Amblymora_, _Stesilea_, _Enes_, and the large +genus _Micracantha_, with but a single species beyond the boundary,--30 +Austro-Malayan genera in all, each found in more than one island, but none +of them extending west of Celebes. Here we have clear proof that the +boundary line between the two great regions exists for Longicorns, as well +as for all other animals; but in this case an unusually large number have +been able to get across it. This, however, does not abolish the barrier, +but only proves that it is not absolutely effectual in all cases. Those who +maintain that the Malay Archipelago forms a single Coleopterous region, +must disprove or explain the instances of limited range here adduced. + +Out of nearly 1500 known genera of these insects, only one genus, _Clytus_, +appears to be cosmopolitan. _Saperda_ and _Callichroma_ are the only others +that perhaps occur in every region; but these are both wanting over wide +tracts of the earth's surface, _Saperda_ being absent from Tropical Africa +and the Malay Archipelago; and _Callichroma_ from the Australian region, +except one species in Polynesia. Many of the genera of Longicorns have a +somewhat wide and scattered distribution, indicative of decadence or great +antiquity. _Mallodon_ and _Parandra_ are mostly South American, but have +species in Australia and Africa; _Oeme_ is found in Brazil and the United +States, with one species in West Africa; _Ceratophorus_ has 2 species in +West Africa and 1 in New Zealand. _Xystrocera_ is mostly African, but has +single species in {502}Borneo, Java, Amboyna and South Australia; _Phyton_ +has one species in North America and the other in Ceylon; _Philagetes_ has +2 in South Africa, and 1 in Malacca; _Toxotus_ abounds in North America and +Europe, with one species away in Madagascar. _Leptura_ is also North +Temperate, but has a species at the Cape, one at Singapore and a third in +Celebes. _Necydalis_ has species in North and South America, Europe and +Australia. _Hylotrupes_ has 1 species in North America and Europe, and 1 in +Australia; _Leptocera_ prefers islands, being found only in Ceylon, +Madagascar, Bourbon, Batchian, the New Hebrides, New Caledonia and North +Australia; _Hathliodes_ is Australian, with 1 species in Ceylon; +_Schoenionta_ has 3 Malayan species, and 1 in Natal. Many other cases +equally curious could be quoted, but these are sufficient. They cannot be +held to indicate any close relation between the distant countries in which +species of the same genus are now found, but perhaps serve to remind us +that groups of great antiquity, and probably of great extent, have dwindled +away, leaving a few surviving relics scattered far and wide, the sole +proofs of their former predominance. + + +_General Observations on the Distribution of Coleoptera._ + +We have now passed in review six of the most important and best known +groups of the Coleoptera or Beetles, comprising about 2,400 genera, and +more than 21,000 species. Although presenting certain peculiarities and +anomalies, we have found that, on the whole, their distribution is in very +close accordance with that of the higher animals. We have seen reason to +believe that these great and well-marked groups have a high geological +antiquity, and by constantly bearing this fact in mind, we can account for +many of the eccentricities of their distribution. They have probably +survived changes of physical geography which have altogether extinguished +many of the more highly organised animals, and we may perhaps gain some +insight into the bearing of those changes, by considering the cross +relations between the several regions indicated by them. On carefully +tabulating the indications given by each of the groups here discussed, I +arrive at the following approximate result. The {503}best marked affinities +between the regions are those between the Nearctic and Palæarctic,--the +Oriental and Australian,--the Australian and Neotropical,--which appear to +be about equal in each case. Next comes that between the Ethiopian and +Oriental on the one side, and the Ethiopian and Neotropical on the other, +which also appear about equal. Then follows that between the Nearctic and +Neotropical regions; and lastly, and far the least marked, that between the +North Temperate and South Temperate regions. That the relation between the +Ethiopian and Neotropical region should be so comparatively well marked, is +unexpected; but we must consider that in such a comparison as the present, +we probably get the result, not of any recent changes or intermigrations, +but of all the long series of changes and opportunities of migration that +have occurred during many geological epochs,--probably during the whole of +the Tertiary period, perhaps extending far back into the Secondary age. + +It appears evident that Insects exhibit in a very marked degree in their +actual distribution, the influence both of very ancient and very modern +conditions of the earth's surface. The effects of the ancient geographical +features of the earth, are to be traced, in the large number of cases of +discontinuous and widely scattered groups which we meet with in almost +every family, and which, to some extent, obscure the broader features of +distribution due to the period during which the barriers which divide the +several primary regions have continued to exist. And this, which we may +consider as the normal distribution, is still further obscured in those +cases where the barriers between existing regions are of such a nature as +to admit of the free passage of insects or their larva in a variety of +ways, and (what is perhaps of more importance) in which the physical +features on both sides of the barrier are so nearly identical, as to admit +of the ready establishment of such immigrants as may occasionally arrive. +These conditions concur, for some families of insects, in the case of the +Oriental and Australian portions of the Malay Archipelago; and it is there +that the normal distribution has been sometimes greatly obscured, but +never, as we have sufficiently shown, by any means obliterated. + + + + +{504}CHAPTER XXII. + +AN OUTLINE OF THE GEOGRAPHICAL DISTRIBUTION OF MOLLUSCA. + + +The Mollusca being for the most part marine, it does not enter into the +plan of this work to go into much detail as to their distribution. The +orders and families will, however, be passed briefly in review, and all +terrestrial and fresh-water groups discussed in somewhat more detail; with +the object of showing how far their distribution accords with that of the +higher animals, and to what extent the anomalies they present can be +explained by peculiarities of organisation and habits. If the views +advocated in our fifth chapter are correct, the regions there marked out +must apply to all classes of animals; and it will be the task of the +students of each group, to work out in detail the causes which have led to +any special features of distribution. All I can hope to do here, is to +show, generally and tentatively, that such a mode of treatment is possible; +and that it is not necessary, as it is certainly not convenient or +instructive, to have a distinct set of "Regions" established for each class +or order in the Animal and Vegetable Kingdoms. + +For all the Marine groups I have merely summarised the information +contained in Mr. Woodward's _Manual of the Mollusca_, but in the case of +the Land Shells I have consulted the most recent general works, and +endeavoured to give an accurate, though doubtless a very incomplete, +account of the most interesting facts in their distribution. As their +classification is very unsettled, I have followed that of the two latest +great works, by Martens and Pfeiffer. + + +{505}CLASS.--CEPHALOPODA. + + +_Order I.--DIBRANCHIATA._ + + +FAMILY 1.--ARGONAUTIDÆ. "Paper Nautilus." (1 Genus, 4 Species). + +DISTRIBUTION.--Open seas of all warm regions. Two species fossil in +Tertiary deposits. + + +FAMILY 2.--OCTOPODIDÆ. "Polypi." (7 Genera, 60 Species). + +DISTRIBUTION.--Norway to New Zealand, all tropical and temperate seas and +coasts. + + +FAMILY 3.--TEUTHIDÆ. "Squids or Sea-pens." (16 Genera, 102 Species.) + +DISTRIBUTION.--Universal, to Greenland; 2 other genera are fossil, in the +Lias and Oolite. + + +FAMILY 4.--SEPIADÆ. "Cuttle Fish." (1 Genus, 30 Species). + +DISTRIBUTION.--All seas: 4 other genera are fossil, in Eocene and Miocene +deposits. + + +FAMILY 5.--SPIRULIDÆ. (1 Genus, 3 Species). + +DISTRIBUTION.--All the warmer seas. + + +{506}FAMILY 6.--BELEMNITIDÆ. Fossil. (6 Genera, 100 Species). + +DISTRIBUTION.--Lias to Chalk in Europe, India and North America. + + +_Order II.--TETRABRANCHIATA._ + + +FAMILY 7.--NAUTILIDÆ. (1 Genus, 3 Species, Living; 4 Genera, 300 Species, +Fossil). + +DISTRIBUTION.--Indian and Pacific Oceans; and the fossil species from the +Silurian Period to the Tertiary, in all parts of the world. + + +FAMILY 8.--ORTHOCERATIDÆ. Fossil. (8 Genera, 400 Species). + +DISTRIBUTION.--Lower Silurian to Lias. + + +FAMILY 9.--AMMONITIDÆ. Fossil. (14 Genera, 1100 Species). + +DISTRIBUTION.--Upper Silurian to Chalk. Found at 16,000 feet elevation in +the Himalayas. + + +{507}Class.--GASTEROPODA. + +_Order I.--PROSOBRANCHIATA._ + + +FAMILY 1.--STROMBIDÆ. (4 Genera, 86 Species.) + +DISTRIBUTION.--The Strombidæ, or Wing-shells, inhabit tropical and warm +seas from the Mediterranean to New Zealand; most abundant in the Indian and +Pacific Oceans. There are nearly 200 fossil species, from the Lias to +Miocene and recent deposits. + + +FAMILY 2.--MURICIDÆ. (12 Genera, 1000 Species.) + +DISTRIBUTION.--All seas, most abundant in the Tropics. _Trichotropis_ is +confined to Northern seas; _Murex_ and _Fusus_ are cosmopolitan. There are +about 700 fossil species, ranging from the Oolite to the Miocene and recent +formations. + + +FAMILY 3.--BUCCINIDÆ. (24 Genera, 1100 Species.) + +DISTRIBUTION.--The Buccinidæ, or "Whelks," range over the whole world, but +some of the genera are restricted. _Buccinum_ inhabits the north and south +temperate seas; _Monoceros_ the West Coast of America; _Cassidaria_ the +Mediterranean; _Phos_, _Harpa_, _Eburna_, and _Ricinula_, are confined to +the Pacific; _Dolium_ inhabits the Mediterranean as well as the Pacific. +There are about 350 fossil species, mostly from the Eocene and Miocene +beds. + + +{508}FAMILY 4.--CONIDÆ. (3 Genera, 850 Species.) + +DISTRIBUTION.--The Cones are universally distributed, but this applies only +to the genus _Pleurotoma_. _Conus_ is tropical and sub-tropical, and +_Cithara_ is confined to the Philippine Islands. There are about 460 fossil +species, from the Chalk formation to the most recent deposits. + + +FAMILY 5.--VOLUTIDÆ, (5 Genera, 670 Species.) + +DISTRIBUTION.--The Volutes are mostly tropical; but a small species of +_Mitra_ is found at Greenland, and a _Marginella_ in the Mediterranean. +_Cymba_ is confined to the West Coast of Africa and Portugal. _Voluta_ +extends south to Cape Horn. There are about 200 fossil species, from the +Chalk and Eocene to recent formations. + + +FAMILY 6.--CYPRÆIDÆ. (3 Genera, 200 Species.) + +DISTRIBUTION.--The well-known Cowries are found all over the world, but +they are much more abundant in warm regions. One small species extends to +Greenland. There are nearly 100 fossil species, from the Chalk to the +Miocene and recent formations. + + +FAMILY 7.--NATICIDÆ. (5 Genera, 270 species.) + +DISTRIBUTION.--The Naticidæ, or Sea-snails, though most abundant in the +Tropics, are found also in temperate seas, and far into the Arctic regions. +Two other genera are fossil; and there are about 300 extinct species, +ranging from the Devonian to the Pliocene formations. + + +{509}FAMILY 8.--PYRAMIDELLIDÆ. (10 Genera, 220 Species.) + +DISTRIBUTION.--These turreted shells are very widely distributed both in +temperate and tropical seas; and most of the genera have also a wide range. +There are about 400 extinct species, from so far back as the Lower Silurian +to the Pliocene formations. + + +FAMILY 9.--CERITHIADÆ. (5 Genera, 190 Species.) + +DISTRIBUTION.--These are marine, estuary, or fresh-water shells, of an +elongated spiral form; they have a world-wide distribution, but are most +abundant in the Tropics. _Potamides_ (41 sp.), is the only fresh-water +genus, and is found in the rivers of Africa, India and China, to North +Australia and California. Another genus is exclusively fossil, and there +are about 800 extinct species, ranging from the Trias to the Eocene and +recent formations. + + +FAMILY 10.--MELANIADÆ. (3 Genera, 410 Species.) + +DISTRIBUTION.--Fresh-water only: lakes and rivers in warm countries, widely +scattered. South Palæarctic and Australian regions, from Spain to New +Zealand; South Africa, West Africa, and Madagascar; United States. There +are about 50 fossil species, from the Wealden and Eocene to recent +formations. + + +FAMILY 11.--TURRITELLIDÆ. (5 Genera, 230 Species.) + +DISTRIBUTION.--Universal. _Cæcum_ is found in north temperate seas only. +The other genera are mostly tropical, but some species reach Iceland and +Greenland. There are near 300 species fossil, ranging from the Neocomian to +the Pliocene formations. + + +{510}FAMILY 12.--LITTORINIDÆ. (9 Genera, 310 Species.) + +DISTRIBUTION.--The Littorinidæ are mostly found on the coasts in shallow +water; as the common Periwinkle (_Littorina littorea_). They are of +world-wide distribution; but _Solarium_ and _Phorus_ are tropical; while +_Lacuna_, _Skenea_, and most species of _Rissoa_ are Northern. About 180 +species are fossil, ranging from the Permian to the Pliocene formations. + + +FAMILY 13.--PALUDINIDÆ. (4 Genera, 217 Species.) + +DISTRIBUTION.--The Paludinidæ, or River-snails, are all fresh-water, and +range over the whole world. _Paludina_ (60 sp.), is confined to the +Northern Hemisphere; _Ampullaria_ (136 sp.), is tropical; _Amphibola_ (3 +sp.), inhabits New Zealand and the Pacific Islands; _Valvata_ (18 sp.), +North America and Britain. There are 72 fossil species of _Paludina_ and +_Valvata_, in the Wealden formation and more recent fresh-water deposits. + + +FAMILY 14.--NERITIDÆ. (10 Genera, 320 Species.) + +DISTRIBUTION.--All warm seas, ranging north to Norway and the Caspian Sea. +_Neritina_ and _Navicella_ inhabit fresh or brackish waters, the latter +confined to the countries bordering the Indian Ocean and the islands of the +Pacific. There are 80 fossil species, from the Trias, Lias, and Eocene +formations down to recent deposits. + + +FAMILY 15.--TURBINIDÆ. (10 Genera, 425 Species). + +DISTRIBUTION.--The genus TROCHUS (200 sp.) has a world-wide range, but the +other genera are mostly tropical, and are most abundant in the Indian and +Pacific Oceans. There are more than 900 fossil species, found in all parts +of the world, from the Lower Silurian to the Tertiary formations. + + +{511}FAMILY 16.--HALIOTIDÆ. (6 Genera, 106 Species). + +DISTRIBUTION.--The Ear-shells are most abundant in the Indian and Pacific +Oceans; some are found on the east coasts of the Atlantic, but there are +very few in the West Indies. _Ianthina_ (10 sp.) consists of floating +oceanic snails found in the warm parts of the Atlantic. Three other genera +are fossil, and there are near 500 fossil species of this family ranging +from the Lower Silurian to the Pliocene formations. + + +FAMILY 17.--FISSURELLIDÆ. (5 Genera, 200 Species). + +DISTRIBUTION.--All seas. _Puncturella_ (6 sp.) is confined to Northern and +Antarctic seas; _Rimula_ to the Philippines; and _Parmophorus_ (15 sp.) +from the Cape of Good Hope to the Philippines and New Zealand. There are +about 80 fossil species, ranging from the Carboniferous formation to the +deposits of the Glacial epoch. + + +FAMILY 18.--CALYPTRÆIDÆ. (4 Genera, 125 Species). + +DISTRIBUTION.--The Calptræidæ, or Bonnet-Limpets, are found on the coasts +of all seas from Norway to Chili and Australia; but are most abundant +within the Tropics. The genera are all widely scattered. There are 75 +fossil species, ranging from the Devonian to recent formations. + + +FAMILY 19.--PATELLIDÆ. (4 Genera, 254 Species). + +DISTRIBUTION.--The Patellidæ, or Limpets, are universally distributed, and +are as abundant in the temperate as in tropical seas. There are about 100 +fossil species, ranging from the Silurian to the Tertiary formations. + + +{512}FAMILY 20.--DENTALIADÆ. (1 Genus, 50 Species). + +DISTRIBUTION.--The genus _Dentalium_ is found in the North Atlantic, +Mediterranean, West Indies and India. There are 125 fossil species, found +in various formations as far back as the Devonian in Europe and in Chili. + + +FAMILY 21.--CHITONIDÆ. (1 Genus, 250 Species). + +DISTRIBUTION.--On rocky shores in all parts of the world. There are 37 +fossil species ranging back to the Silurian period. + + +_Order II.--PULMONIFERA. ("Terrestrial Molluscs.")_ + +The Land and Fresh-water snails are so important and extensive a group, and +their classification has been so carefully studied, that their geographical +distribution is a subject of much interest. The range of the genera will +therefore be given in some detail. For the Helicidæ I follow the classical +work of Albers--_Die Helicien_, Von Martens' Edition (1860); and for the +Operculate families, Pfeiffer's _Monographia Pneumonopomorum Viventium_, +2nd Supplement, 1865. The number of species is, of course, very +considerably increased since these works were published (and the probable +amount of the increase I have in most cases indicated), but this does not +materially affect the great features of their geographical distribution. + + +FAMILY 22.--HELICIDÆ. (33 Genera, 3,332 Species) (1860). + +GENERAL DISTRIBUTION.--Universal. + +The Helicidæ, or Snails, are a group of immense extent and absolutely +cosmopolitan in their range, being found in the most barren deserts and on +the smallest islands, all over the globe. They reach to near the line of +perpetual snow on mountains, and {513}to the limit of trees or even +considerably beyond it, in the Arctic regions; but they are comparatively +very scarce in all cold countries. The Antilles, the Philippine Islands, +Equatorial America, and the Mediterranean sub-region are especially rich in +this family. Comparatively few of the genera, and those generally small +ones, are restricted to single regions; but on the other hand very few are +generally distributed, only two--_Helix_ and _Pupa_--occurring in all the +six regions, while _Helix_ alone is truly cosmopolitan, occurring in every +sub-region, in every country, and perhaps in every island on the globe. + +The Neotropical region is, on the whole, the richest in this family, the +continental Equatorial districts producing an abundance of large and +handsome species, while the Antilles are pre-eminent for the number of +their peculiar forms. This region possesses 22 of the genera, and 6 of them +are peculiar. + +The Palæarctic region seems to come next in productiveness, but this may be +partly owing to its having been so thoroughly explored. It possesses 16 of +the genera, and 3 of them are confined to it. The great mass of the species +are found in the warm and fertile countries surrounding the Mediterranean +Sea. + +The Ethiopian region has 13 genera, only one of which is peculiar. + +The Australian region has 14 genera, 2 of which are confined to the Pacific +Islands. + +The Oriental has 15 genera and the Nearctic 12, but in neither case are +there any peculiar generic types. + +The following is the distribution of the several genera taken in the order +of their magnitude:-- + +_Helix_ (1,115 sp.), cosmopolitan. This genus is divided into 88 +sub-genera, a number of which have a limited distribution. An immense +quantity of species have been recently described, so that the number now +exceeds 2,000. + +_Nanina_ (290 sp.) is characteristic of the Oriental and Australian +regions, over the whole of which it extends, just entering the Palæarctic +region as far as North China and Japan. Isolated from this area is a small +group of 4 species occurring {514}in West Africa. The number of species in +this genus have now been increased to about 400. + +_Clausilia_ (272 sp.) is most abundant in Europe, with a few species widely +scattered in India, Malaya, China, Japan, Equatorial America, and one in +Porto Rico. The described species have been increased to nearly 500. + +_Bulimulus_ (210 sp.) is American, and almost exclusively Neotropical, +ranging from Montevideo and Chili, to the West Indian Islands, California +and Texas; with two sub-genera confined to the Galapagos Islands. About 100 +new species have been described since the issue of the second edition of +Dr. Woodward's Manual. + +_Pupa_ (210 sp.) abounds most in Europe and the Arctic regions, but has a +very wide range, being scattered throughout Africa, continental India, +Australia, the Pacific Islands, North America to Greenland, and the +Antilles; but it is absent from South America, the Himalayan and Malayan +sub-regions, China and Japan. An extinct species has occurred abundantly in +the carboniferous strata of North America. About 160 additional species +have been described. + +_Bulimus_ (172 sp.) abounds most in Tropical South America; it is also +found from Burmah eastward through Malaya to the Solomon and Fiji Islands; +there are also scattered species in Patagonia, St. Vincents, Texas, St. +Helena, and New Zealand. More than 100 additional species have been +described. + +_Buliminus_ (132 sp.) ranges from Central and South Europe over the whole +Ethiopian and Oriental regions to North China, and through the Australian +to New Zealand; there is also a single outlying species in the Galapagos +Islands. About 50 more species have been described. + +_Cochlostyla_ (127 sp.) is almost peculiar to the Philippine Islands, +beyond which, are a species in Borneo, one in Java, and two in Australia. +Very few new species have been added to this genus. + +_Achatinella_ (95 sp.) is absolutely confined to the Sandwich Island group. +Recent researches have more than tripled the number of described species. + +{515}_Achatina_ (87 sp.) is most abundant and finest in the Ethiopian +region, over the whole of which it ranges; but there are also species in +Florida, the Antilles, the Sandwich Islands, Ceylon and India. The +described species are now more than doubled. + +_Hyalina_ (84 sp.) inhabits all Tropical America and the Antilles, North +America to Greenland, and Europe to the Arctic regions. Comparatively few +new species have been described. + +_Cylindrella_ (83 sp.) inhabits the West Indian islands and Guatemala to +Texas, with a sub-genus in the Philippine Islands. Species since described +have more than trebled the number in this genus. + +_Cionella_ (67 sp.) is widely scattered; in India from Ceylon to the Khasia +Mountains, Brazil, New Granada, the West Indian islands, Palæarctic, and +northern part of Nearctic regions, Pacific Islands, New Zealand, and Juan +Fernandez. About 20 new species have since been described. + +_Glandina_ (66 sp.), Peru to South Carolina and the Antilles, with three +species in Central Africa and one in South Europe. About 40 species have +been added to this genus. + +_Stenogyra_ (49 sp.), widely distributed: Tropical America and West Indies +to Florida, South and West Africa, the Mediterranean region, India and the +Philippines. About a dozen new species have been described. + +_Succinea_ (41 sp.), widely scattered in all the regions, and in St. +Helena, Juan Fernandez, Tahiti, Chiloe, Greenland, West Africa, Himalayas +and Australia. The described species are now more than 100. + +_Partula_ (39 sp.), Solomon Islands to Tahiti and Sandwich Islands. This +genus has also been increased to near 100 species. + +_Streptaxis_ (34 sp.), most abundant in Tropical South America, but occurs +in West Africa, the Seychelles and Rodriguez Islands, Ceylon and Burmah. It +now contains over 100 described species. + +_Spiraxis_ (33 sp.), Yucatan to Mexico, and less abundant in the West +Indian Islands. About 20 species have been added. + +{516}_Macroceramus_ (27 sp.), Antilles, Florida, and Peru. The species have +been more than doubled. + +_Vitrina_ (26 sp.), widely scattered through North and Central Europe, +North-west America and Greenland, Abyssinia, Madagascar and South Africa, +Himalayas to Burmah and Australia. Species since described have more than +doubled the number in this genus. + +_Orthalicus_ (23 sp.), Bolivia to Mexico and Antilles. This genus has been +increased to about 40 species. + +_Sagda_ (19 sp.), Antilles only. Very few new species, if any, have been +described. + +_Zonites_ (12 sp.), South Europe, with one species of a distinct type in +Guatemala. The number of species in this genus has been since about +tripled. + +_Leucochroa_ (11 sp.), Mediterranean region to Syria and Arabia Petrea. + +_Simpulopsis_ (7 sp.), Bahia, Antilles, and far away in the Solomon +Islands. Two or three have been added. + +_Balea_ (6 sp.), Middle and North Europe, Brazil, and the Island of Tristan +d'Acunha. + +_Daudebardia_ (6 sp.), Central and South Europe; and a species has since +been discovered in New Zealand. + +_Macrocycles_ (4 sp.), Chili, California, Oregon and Central North America. + +_Columna_ (3 sp.), West Africa, Princes Islands and Madagascar. + +_Stenopus_ (2 sp.), Island of St. Vincent (West Indies.) + +_Pfeifferia_ (2 sp.), Philippines and Moluccas. + +_Testacella_ (2 sp.), West Europe and Teneriffe. About 8 species have been +since described, including one from New Zealand. + +Fossil species of _Helix_, _Bulimus_, _Achatina_, _Balea_, and _Clausilia_, +are found in all the Tertiary formations; while a species of _Pupa_ (as +already stated) occurs in the carboniferous formation. For interesting +details of the distribution of the sub-genera and species of _Achatinella_ +in the Sandwich Islands, see a paper by Rev. J. T. Gulick in the _Journal +of the Linnean Society_. (Zoology, vol. xi. p. 496.) + + +{517}FAMILY 23.--LIMACIDÆ.--(12 Genera, 116 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- -- -- --|1. 2. 3. 4 |1. 2. 3. 4 |-- -- 3 -- |1. 2. 3. 4 |1. 2. 3. 4 + | | | | | + +The Limacidæ, or Slugs, are widely distributed, but they are absent from +South America, where they are represented by the next family. They also +seem to be absent from the greater part of Africa. The genera are +distributed as follows:-- + +_Limax_ (51 sp.), Palæarctic region, Australia and the Sandwich Islands; +_Anadenus_ (2 sp.), Himalayas; _Philomychus_ (9 sp.), North America, China +and Java; _Arion_ (25 sp.), Norway to Spain and South Africa; _Parmacella_ +(7 sp.), South Europe, Canary Islands and North India; _Janella_ (1 sp.), +New Zealand; _Aneitea_ (1 sp.), New Hebrides and New Caledonia; _Parmarion_ +(4 sp.), India; _Triboniophorus_ (3 sp.), Australia; _Testacella_ (3 sp.), +South Europe, Canary Islands, and New Zealand; _Hyalimax_ (2 sp.), Bourbon +and Mauritius; _Krynickia_ (8 sp.), Eastern Europe and North America. A few +species of _Limax_, _Arion_, and _Testacella_ have been found fossil in +Tertiary deposits. + + +FAMILY 24.--ONCIDIADÆ. (2 Genera, 36 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |-- -- -- --|1. 2 -- -- |-- -- -- 4 |1. 2. 3. 4 | -- 2 -- 4 + | | | | | + +The Oncidiadæ, or Slugs with a coriaceous mantle, inhabit the Oriental +region, Mauritius, Australia, the Pacific Islands, South America and South +Europe. The genera are:-- + +{518}_Oncidium_ (16 sp.), South Europe (1 sp. British), Mauritius, +Australia and Pacific Islands; Vaginulus (20 sp.), Neotropical and Oriental +regions. + + +FAMILY 25.--LIMNÆIDÆ. (7 Genera, 332 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1. 2. 3. 4 |1. 2. 3. 4 |1. 2. 3. 4 |-- -- 3 -- |1. 2. 3. 4 |-- -- -- -- + | | | | | + +The Limnæidæ, or Fresh-water Snails, inhabit ponds and rivers in most parts +of the world, but appear to be absent from the Australian region. The +genera are distributed as follows:-- + +_Limnæa_ (95 sp.), Nearctic, Palæarctic, and Oriental regions; +_Choanomphalos_ (2 sp.), Lake Baikal; _Pompholyx_ (2 sp.), Western America; +_Chilinia_ (18 sp.), South America; _Physa_ (20 sp.), Nearctic, Palæarctic, +Ethiopian and Oriental regions, and extends to above 73° North Latitude in +Siberia, being the most Arctic of land or fresh-water shells; _Ancylus_ (49 +sp.), Nearctic and Neotropical regions, Europe and New Zealand; _Planorbis_ +(145 sp.), Nearctic, Palæarctic and Oriental regions. Several genera are +found fossil, chiefly in the Wealden, Eocene, and Miocene formations. + + +FAMILY 26.--AURICULIDÆ. (3 Genera, 210 Species.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + 1 -- -- 4 |1. 2. 3. 4 |1. 2 -- -- |1. 2. 3 -- |1. 2. 3. 4 | 1. 2 -- 4 + | | | | | + +The Auriculidæ are chiefly found near the sea in hot countries, and are +most abundant in the Eastern tropics. They are absent {519}from the East +coast of South America. The genera have a somewhat restricted distribution +as follows:-- + +_Auricula_ (128 sp.), India, Pacific Islands, Peru and West Indies; +_Melampus_ (56 sp.), West Indies and Europe; _Carychium_ (9 sp.), Europe +and North America; _Plectrotrema_ (14 sp.), Australia, Malay Islands, +China, Cuba; _Blauneria_ (2 sp.), West Indian and Sandwich Islands. There +are many fossil species ranging back to the Eocene formation. + + +FAMILY 27.--ACICULIDÆ. (4 Genera, 65 Species.) (1865.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |1. 2 -- -- | 1. 2 -- 4 |-- -- -- 4 | -- 2 -- 4 |1. 2. 3 -- + | | | | | + +The Aciculidæ are small cylindrical shells chiefly found in the West Indian +Islands, but with representatives widely scattered over the globe. + +_Acicula_ (5 sp.) is European only; _Geomelania_ (21 sp.), and _Chittya_ (1 +sp.), are confined to the Island of Jamaica; _Truncatella_ (38 sp.), is +most abundant in the Antilles, but is also found in some part of each of +the six regions, as indicated by the diagram of the family. But few new +species have been added to this group. + + +FAMILY 28.--DIPLOMMATINIDÆ. (3 Genera, 23 Species.) (1865.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2 -- -- |-- -- -- --|-- -- -- --|-- -- -- --| 1 -- 3. 4 | 1. 2. 3. 4 + | | | | | + +The Diplommatinidæ are minute shells of the Oriental and Australian +regions. + +{520}_Diplommatina_ (18 sp.) inhabits India to Burmah, and the greater part +of the Australian region; the number of species has now been doubled, and +one has been discovered in the island of Trinidad; _Clostophis_ (1 sp.), +Moulmein; _Paxillus_ (3 sp.), Borneo, Hong Kong, and Loo Choo Islands. + + +FAMILY 29.--CYCLOSTOMIDÆ. (41 Genera, 1009 Species.) (1865.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |-- -- 3 -- |-- -- -- 4 |-- -- 3. 4 |1. 2. 3. 4 |1 -- -- -- + | | | | | + +This extensive group, comprising the largest of the operculated +land-shells, is especially characteristic of the Oriental region, which +possesses 25 genera, no less than 12 of them being wholly confined to it. +The Neotropical region comes next, with 15 genera, 9 of which are peculiar; +but a large number of these are confined to the West Indian Islands, South +America itself being very poor in this group. The Palæarctic region has 3 +peculiar genera; the Ethiopian and Australian 1 each. The Nearctic region +has but a single West Indian species in Florida. The distribution of the +genera is as follows:-- + +Peculiar to or characteristic of the Oriental region are, _Opisthoporus_ +(11 sp.), _Rhiostoma_ (6 sp.), _Alycaeus_ (39 sp.), _Opisthostoma_ (1 sp.), +_Hybocistis_ (3 sp.), _Pterocyclos_ (19 sp.), extending to the Moluccas; +_Aulopoma_ (4 sp.), _Dermatocera_ (4 sp.), _Leptopoma_ (54 sp.), extending +west to the Seychelles and east to the Moluccas and New Guinea; +_Cyclophorus_ (163 sp.), most abundant in the Oriental region, but ranges +to Japan, to Chili, and all Tropical America, over the whole Australian +region, and to Natal and Madagascar; _Cataulus_ (15 sp.), confined to +Ceylon, the Neilgherries and Nicobar Islands; _Rhaphaulus_ (4 sp.), Penang +to Ceram; _Streptaulus_ (1 sp.), _Arinia_ (3 sp.), _Pupinella_ (2 sp.), +_Pupina_ (24 sp.), half in North India to Philippines and {521}Japan, the +other half in Moluccas, New Guinea and Australia; _Cyclotopsis_ (2 sp.), +India and Malaya; _Registoma_ (9 sp.), Philippines and Moluccas, New +Caledonia and Pacific. + +Characteristic of the Neotropical region are:--_Cyclotus_ (111 sp.), half +in the Antilles and Tropical America, the rest in the Moluccas, China, +Malaya, India, Natal, and the Seychelle Islands; _Megalomastoma_ (27 sp.), +abundant in Cuba, West Indies and South America, others in India, Malaya, +and Mauritius; _Jamaicia_ (2 sp.), Jamaica; _Licina_ (5 sp.), Antilles; +_Choanopoma_ (49 sp.), Antilles; _Ctenopoma_ (25 sp.), Antilles; +_Diplopoma_ (1 sp.), Cuba; _Adamsiella_ (15 sp.), Jamaica, Cuba, Guatemala; +_Cyclostomus_ (113 sp.), abundant in Antilles, also occurs in Madagascar, +Arabia, Syria, Hungary and New Zealand; _Tudora_ (34 sp.), Antilles, and +one species in Algeria; _Cistula_ (40 sp.), _Chondropoma_ (94 sp.), +_Bourcieria_ (2 sp.), Tropical America. + +Peculiar to or characteristic of the Palæarctic region are:--_Craspedopoma_ +(5 sp.), confined to Madeira, the Azores and Canaries; _Leonia_ (1 sp.), +Spain and Algeria; _Pomatias_ (22 sp.), Europe and Canaries with a species +in the Himalayas; _Cecina_ (1 sp.), Manchuria. + +The Ethiopian region has the peculiar genus _Lithodion_ (5 sp.), +Madagascar, Socotra and Arabia; and _Otopoma_ (19 sp.), Mascarene Islands +and Socotra, with a species in Western India and another in New Ireland. + +The Australian region is characterised by _Callia_ (3 sp.), in Ceram, +Australia, and the Philippines respectively; _Realia_ (7 sp.), New Zealand +and the Marquesas Islands; _Omphalotropis_ (38 sp.), the Australian region, +with some species in India, Malaya, and the Mauritius. + +The remaining genus, _Hydrocena_ (27 sp.), has a very widely scattered +distribution, being found in South Europe, Japan, the Cape, China, Malaya, +New Zealand, the Pacific Islands and Chili. From 10 to 20 per cent. of new +species have been since described in most of the genera of this family. + + +{522}FAMILY 30.--HELICINIDÆ. (7 Genera, 433 Species.) (1868.) + + GENERAL DISTRIBUTION. + __________________________________/\__________________________________ + / \ + NEOTROPICAL| NEARCTIC |PALÆARCTIC | ETHIOPIAN | ORIENTAL | AUSTRALIAN + SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS|SUB-REGIONS. + -----------+-----------+-----------+-----------+-----------+------------ + | | | | | + -- 2. 3. 4 |-- -- 3 -- |-- -- -- --|-- -- -- --|-- -- 3. 4 |1. 2. 3 -- + | | | | | + +The Helicinidæ are very characteristic of the Antilles, comparatively few +being found in any other part of the world except the Islands of the +Pacific. The genera are:-- + +_Trochatella_ (33 sp.), Antilles with a species in Venezuela, and another +in Cambodja; _Lucidella_ (5 sp.), Antilles; _Helicina_ (274 sp.), Antilles, +Pacific Islands, Tropical America, Southern United States, Moluccas, +Australia, Philippines, Java, Andaman Islands and North China; +_Schasicheila_ (5 sp.), Mexico, Guatemala and Bahamas; _Alcadia_ (28 sp.), +Antilles; _Georissa_ (5 sp.) Moulmein to Burmah. About 10 per cent. of new +species appear to have been since described in the larger genera of this +family. + + +_General Observations on the Distribution of the Land Mollusca._ + +A consideration of the distribution of the families and genera of +land-shells shows us, that although they possess some special features, yet +they agree in many respects with the higher animals in their limitation by +great natural barriers, such as oceans, deserts, mountain ranges, and +climatal zones. A remarkable point in the distribution of these animals, is +the number of genera which have a very limited range, and also the +prevalence of genera having species scattered, as it were at random, all +over the earth. No less than 14 genera (or about one-sixth of the whole +number) are confined to the Antilles, while the greater part of the +sub-genera of modern authors are restricted to limited areas. + +If we first compare the New World with the Old, we find the difference as +regards genera quite as great as in most of the {523}vertebrates. In the +Helicidæ, 10 genera are confined to the New, and 7 to the Old World, 16 +being common to both. In the Operculata the number of genera of restricted +range is greater,--the New World having 15, the Old World 32 genera, only 8 +being common to both. Of the New World genera 12 out of the 15 do not occur +at all in South America; and of those of the Old World, 22 out of the 32 +occur in a single region only. If we take the northern and southern +division proposed by Professor Huxley (the latter comprising the Australian +and Neotropical regions), we find a much less well-marked diversity. Among +the Helicidæ only 4 are exclusively northern, 8 southern; while among the +Operculata 22 are northern, 16 southern. The best way to compare these two +kinds of primary division will be to leave out all those genera confined to +a single region each, and to take account only of those characteristic of +two or more of the combined regions; which will evidently show which +division is the most natural one for this group. The result is as +follows:-- + +GENERA COMMON TO TWO OR MORE REGIONS IN, AND CONFINED TO, EACH PRIMARY +DIVISION OF THE EARTH. + + Helicidæ Operculata. Totals. + { Northern 0 0 0 } + { Southern 0 0 0 } + + { Old World 1 12 13 } + { New World 4 0 4 } + +We find then that the northern and southern division of the globe is not at +all supported by the distribution of the terrestrial molluscs. It is indeed +very remarkable, that the connection so apparent in many groups between +Australia and South America is so scantily indicated here. The only facts +supporting it seem to be, the occurrence of _Geotrochus_ (a sub-genus of +Helix) in Brazil, as well as in the Austro-Malayan and West Pacific Islands +and North Australia; and of _Bulimus_ in the same two parts of the globe, +but peculiar sub-genera in each. But in neither case is there any affinity +shown between the temperate portions of the two regions, so that we must +probably trace this resemblance to some more ancient diffusion of types +than that which led to the similarity of plants and insects. Still more +curious is the entire {524}absence of genera confined to, and +characteristic of Africa and India. One small sub-genus of _Helix_, +(_Rachis_), and one of _Achatina_, (_Homorus_), appear to have this +distribution,--a fact of but little significance when we find another +sub-genus of _Helix_, (_Hapalus_), common and confined to Guinea and the +Philippine Islands; and when we consider the many other cases of scattered +distribution which cannot be held to indicate any real connection between +the countries implicated. No genus is confined to the Palæarctic and +Nearctic regions as a whole. A large number of sub-genera, many of them of +considerable extent, are peculiar to one or other of these regions, but +only 3 sub-genera of _Helix_ and 2 of _Pupa_ are common and peculiar to the +two combined, and these are always such as have an Arctic range and whose +distribution therefore offers no difficulty. + +We find, then, that each of our six regions and almost all of our +sub-regions are distinctly confirmed by the distribution of the terrestrial +mollusca; while the different combinations of them which have at various +times been suggested, receive little or no support whatever. Even those +remarkably isolated sub-regions, New Zealand and Madagascar, have no +strictly peculiar genera of land-shells, although they both possess several +peculiar sub-genera; being thus inferior in isolation to some single West +Indian Islands, to the Sandwich Islands, and even to the North Atlantic +Islands (Canaries, Madeira, and Azores), each of which have peculiar +genera. This of course, only indicates that the means by which land +mollusca have been dispersed are somewhat special and peculiar. To +determine in what this speciality consists we must consider some of the +features of the specific distribution of this group. + +The range of genera, and even of sub-genera is, as we have seen, often wide +and erratic, but as a general rule the species have a very restricted area. + +Hardly a small island on the globe but has some land-shells peculiar to it. +Juan Fernandez has 20 species, all peculiar. Madeira and Porto Santo have +109 peculiar species out of a total of 134. Every little valley, plain, or +hill-top, in the Sandwich Islands, though only a few square miles in +extent, has its {525}peculiar species of _Achatinella_. Another striking +feature of the distribution of land molluscs, is the richness of islands as +compared with continents. The Philippines contain more species than all +India; and those of the Antilles according to Mr. Bland almost exactly +equal the numbers found in the entire American continent from Greenland to +Patagonia. Taking the whole world, it appears that many more species of +land-shells are found in the islands than on the continents of the globe, a +peculiarity that obtains in no other extensive group of animals. + +Looking at these facts it seems probable, that the air-breathing molluscs +have been chiefly distributed by air- or water-carriage, rather than by +voluntary dispersal on the land. Even seas and oceans have not formed +impassable barriers to their diffusion; whereas they only spread on dry +land with excessive slowness and difficulty. The exact mode in which their +diffusion is effected is not known, and it may depend on rare and +exceptional circumstances; but it seems likely to occur in two ways. Snails +frequently conceal themselves in crevices of trees or under bark, or attach +themselves to stems or foliage, and either by their operculum or mucous +diaphragm, are able to protect themselves from the injurious effects of +salt water for long periods. They might therefore, under favourable +conditions, be drifted across arms of the sea or from island to island; +while wherever there are large rivers and occasional floods, they would by +similar means be widely scattered over land areas. Another possible mode of +distribution is by means of storms and hurricanes, which would carry the +smaller species for long distances, and might occasionally transport the +eggs of the larger forms. Aquatic birds might occasionally get both shells +and eggs attached to their feet or their plumage, and convey them across a +wide extent of sea. But whether these, or some other unknown agency has +acted, the facts of distribution clearly imply that some means of transport +over water is, and has been, the chief agent in the distribution of these +animals; but that its action is very rare or intermittent, so that its +effects are hardly perceptible in the distribution of single species. + +Another important factor in enabling us to account for the +{526}distribution of these animals is the geological antiquity of the +group, and the amount of change exhibited in time, by species and genera. +Now we find that most of the genera of land-shells range back to the Eocene +period, while those inhabiting fresh water are found almost unchanged in +the Wealden. In North America a species of _Pupa_ and one of _Zonites_, +have been discovered in the coal measures, along with Labyrinthodonts; and +this fact seems to imply, that many more terrestrial molluscs would be +discovered, if fresh-water deposits, made under favourable conditions, were +more frequently met with in the older rocks. If then the existing groups of +land-molluscs are of such vast antiquity, and possess some means, however +rarely occurring, of crossing seas and oceans, we need not wonder at the +wide and erratic distribution now presented by so many of the groups; and +we must not expect them to conform very closely to those regions which +limit the range of animals of higher organization and less antiquity. + +The total number of species of pulmoniferous mollusca is about 7,000, +according to the estimate of Mr. Woodward, brought down to 1868 by Mr. +Tate. But this number would be largely increased if the estimates of +specialists were taken. Mr. Woodward for example, gives 760 as the number +of species in the West Indian Islands; whereas Mr. Thomas Bland, who has +made the shells of these islands a special study, considers that there were +1,340 species in 1866. So, the land-shells of the Sandwich Islands are +given at 267; but Mr. Gulick has added 120 species of Achatinellidæ, +bringing the numbers up to nearly 400,--but no doubt several of these are +so closely related that many conchologists would class them as varieties. +The land-shell fauna of the Antilles is undoubtedly the most remarkable in +the world, and it has been made the subject of much interesting discussion +by Mr. Bland and others. This fauna differs from that of all other parts of +the globe in the proportions of the operculate to the inoperculate shells. +The Operculata of the globe are about one-seventh, the Inoperculata about +six-sevenths of the whole; and some general approximation to this +proportion (or a much smaller one) exists in almost all the continents, +islands, and {527}archipelagoes. In the Philippines, for example, the +proportion of the Operculata is a little more than one-seventh; in the +Mauritius, between one-third and one-fourth; in Madeira, one-fourteenth; in +the whole American continent about one-eighth; but when we come to the +Antilles we find them to amount to nearly five-sixths, about half the +Operculata of the globe being found there! + +Mr. Bland endeavours to ascertain the source of some of the chief genera +found in the West Indian Islands, on the principle that "each genus has had +its origin where the greatest number of species is found;" and then +proceeds to determine that some have had an African, some an Asiatic, and +some an American origin, while others are truly indigenous. But we fear +there is no such simple way of arriving at so important a result; and in +the case of groups of extreme antiquity like the genera of mollusca, it +would seem quite as possible that the origin of a genus is generally _not_ +where the greatest number of species are now found. For during the repeated +changes of physical conditions that have everywhere occurred since the +Eocene period (to go no further back) every genus must have made extensive +migrations, and have often become largely developed in some other district +than that in which it first appeared. As a proof of this, we not +unfrequently find fossil shells where the species and even the genus now no +longer exists; as _Auricula_, found fossil in Europe, but only living in +the Malay and Pacific Islands; _Anastoma_ and _Megaspira_, now peculiar to +Brazil, but fossil in the Eocene of France; and _Proserpina_ of the West +Indies, found in the Eocene formation of the Isle of Wight. The only means +by which the origin of a genus can satisfactorily be arrived at, is by +tracing back its fossil remains step by step to an earlier form; and this +we have at present no means of doing in the case of the land-shells. Taking +existing species as our guide we should certainly have imagined that the +genus _Equus_ originated in Africa or Central Asia; but recent discoveries +of numerous extinct species and of less specialized forms of the same type, +seem to indicate that it originated in North America, and that the whole +tribe of "horses" may be, for anything we yet know {528}to the contrary, +recent immigrants into the Old World! This example alone must convince us, +that it is impossible to form any conclusion as to the origin of a genus, +from the distribution of existing species only. + +The general conclusion we arrive at, therefore, is, that the causes that +have led to the existing distribution of the genera and higher groups of +the terrestrial mollusca are so complex, and have acted through such long +periods, that most of the barriers which limit the range of other +terrestrial animals do not apply to them, although the species are, in most +cases, strictly limited by them. Some means of diffusion--which, though +probably acting very slowly and at long intervals, and more powerfully on +continents than between islands, is yet highly efficient when we consider +the long duration of genera--has, to a considerable extent, dispersed them +across continents, seas, and oceans. On the other hand, those mountain +barriers which separate many groups of the higher vertebrates, are +generally less ancient than the genera of land-shells, which are thus often +distributed independently of them. In order to compare the distribution of +the terrestrial mollusca on equal terms with those of land animals +generally, we must take genera of the former as equivalent to family groups +of the latter; and we shall, I believe, then find that the distribution of +the sub-genera and smaller groups of species do accord mainly with those +divisions of the earth into regions and sub-regions which we have here +indicated. Mr. Harper Pease, in a communication on Polynesian Land Shells +in the _Proceedings of the Zoological Society_ for 1871 (p. 449), marks out +the limits of the Polynesian sub-region, so as exactly to agree with that +arrived at here from a consideration of the distribution of vertebrata; and +he says that this sub-region, (or region, as he terms it) is distinctly +characterised by its land-shells from all the surrounding regions. The +genera (or sub-genera) _Partula_, _Pitys_, _Achatinella_, _Palaina_, +_Omphalotropis_, and many others, are either wholly confined to this +sub-region or highly characteristic of it. Mr. Binney, in his _Catalogue of +the Air-breathing Molluscs of North America_, marks out our Nearctic region +(with almost identical limits) as most clearly {529}characterised. He also +arrives at a series of sub-divisions, which generally (though not exactly) +agree with the sub-regions which I have here adopted. The Palæarctic, the +Ethiopian, and the Oriental regions, are also generally admitted to be well +characterised by their terrestrial molluscs. There only remain the +Australian and the Neotropical regions, in which some want of homogeneity +is apparent, owing to the vast development and specialisation of certain +groups in the islands which belong to these regions. The Antilles, on the +one hand, and the Polynesian Islands, on the other, are so rich in +land-shells and possess so many peculiar forms, that, judged by these +alone, they must form primary instead of secondary divisions. We have, +however, already pointed out the inconvenience of any such partial systems +of zoological geography, and the causes have been sufficiently indicated +which have, in the case of land-shells as of insects, produced certain +special features of distribution. + +We therefore venture to hope, that conchologists will give us the advantage +of their more full and accurate knowledge both of the classification and +distribution of this interesting group of animals, not to map out new sets +of regions for themselves, but to show what kind of barriers have been most +efficient in limiting the range of species, and how their distribution is +actually effected, so as to be able to explain whatever discrepancies exist +between the actual distribution of land-shells and that of the higher +animals. + + +_Order III.--OPISTHO-BRANCHIATA._ + +There are ten families in this order, all of which, as far as known, are +widely or universally distributed. Some of them are found fossil, ranging +back to the Carboniferous epoch. They are commonly termed Sea-slugs, and +have either a thin small shell or none. We shall therefore simply enumerate +the families, with the number of genera and species as given by Mr. +Woodward. + +{530}FAMILY 31.--TORNATELLIDÆ. (7 Genera, 62 Species living, 166 fossil.) + +FAMILY 32.--BULLIDÆ. (12 Genera, 168 Species living, 88 fossil.) + +FAMILY 33.--APHYSIADÆ. (8 Genera, 84 Species living, 4 fossil.) + +FAMILY 34.--PLEUROBRANCHIDÆ. (7 Genera, 28 Species living, 5 fossil.) + +FAMILY 35.--PHYLLIDIADÆ. (4 Genera, 14 Species living, 0 fossil.) + +FAMILY 36.--DORIDÆ. (23 Genera, 160 Species living, 0 fossil.) + +FAMILY 37.--TRITONIADÆ. (9 Genera, 38 Species living, 0 fossil.) + +FAMILY 38.--ÆOLIDÆ. (14 Genera, 101 Species living, 0 fossil.) + +FAMILY 39.--PHYLLYRHOIDÆ. (1 Genus, 6 Species living, 0 fossil.) + +FAMILY 40.--ELYSIADÆ. (5 Genera, 13 Species living, 0 fossil.) + + +{531}_Order IV.--NUCLEO-BRANCHIATA._ + +These are oceanic, swimming molluscs, of a delicate texture. They are found +in all warm seas, and range back to the Lower Silurian epoch. There are +only two families. + +FAMILY 41.--FIROLIDÆ. (2 Genera, 33 Species living, 1 fossil.) + +FAMILY 42.--ATLANTIDÆ. (5 Genera, 22 Species living, 159 fossil.) + + +CLASS.--PTEROPODA. + +These are swimming, oceanic mollusca, inhabiting both Arctic, Temperate, +and Tropical seas. The three families have each a wide distribution in all +the great oceans. They range back to the Silurian period. + +FAMILY 1.--HYALEIDÆ. (9 Genera, 52 Species living, 95 fossil.) + +FAMILY 2.--LIMACINIDÆ. (4 Genera, 19 Species living, 0 fossil.) + +FAMILY 3.--CLIONIDÆ. (4 Genera, 14 Species living, 0 fossil.) + + +{532}CLASS.--BRACHIOPODA. + +These are sedentary, bivalve, marine mollusca, having laterally symmetrical +shells, but with unequal valves. Both in space and time they are the most +widely distributed molluscs. They are found in all seas, and at all depths; +and when any of the families or genera have a restricted range, it seems to +be due to our imperfect knowledge, rather than to any real geographical +limitations. In time they range back to the Cambrian formation, and seem to +have had their maximum development in the Silurian period. It is not, +therefore, necessary for our purpose, to do more than give the names of the +families with the numbers of the genera and species, as before. + +FAMILY 1.--TEREBRATULIDÆ. (5 Genera, 67 Species living, 340 fossil.) + +FAMILY 2.--SPIRIFERIDÆ. (4 Genera, 0 Species living, 380 fossil.) + +FAMILY 3.--RHYNCHONELLIDÆ. (3 Genera, 4 Species living, 422 fossil.) + +FAMILY 4.--ORTHIDÆ. (4 Genera, 0 Species living, 328 fossil.) + +FAMILY 5.--PRODUCTIDÆ. (3 Genera, 0 Species living, 146 fossil.) + +FAMILY 6.--CRANIADÆ. (1 Genus, 5 Species living, 37 fossil.) + +FAMILY 7.--DISCINIDÆ. (2 Genera, 10 Species living, 90 fossil.) + +FAMILY 8.--LINGULIDÆ. (2 Genera, 16 Species living, 99 fossil.) + + +{533}CLASS.--CONCHIFERA. + +The Conchifera, or ordinary Bivalve Molluscs, may be distinguished from the +Brachiopoda by having their shells laterally unsymmetrical, while the +valves are generally (but not always) equal. They are mostly marine, but a +few inhabit fresh water. As the distribution of some of the families +presents points of interest, we shall treat them in the same manner as the +marine Gasteropoda. + + +FAMILY 1.--OSTREIDÆ. (5 Genera, 426 Species.) + +DISTRIBUTION.--The Ostreidæ, including the Oysters and Scallops, are found +in all seas, Arctic as well as Tropical. There are nearly 1,400 species +fossil, ranging back to the Carboniferous period. + + +FAMILY 2.--AVICULIDÆ. (3 Genera, 94 Species.) + +DISTRIBUTION.--The Aviculidæ, or Wing-shells and Pearl Oysters, are +characteristic of Tropical and warm seas, a few only ranging into temperate +regions. Nearly 700 fossil species are known from various formations +ranging back to the Devonian, and Lower Silurian. + + +FAMILY 3.--MYTILIDÆ. (3 Genera, 217 Species.) + +DISTRIBUTION.--The Mytilidæ, or Mussels, have a world-wide distribution. +There is one fresh-water species, which inhabits the Volga. There are about +350 fossil species, ranging back to the Carboniferous epoch. + + +{534}FAMILY 4.--ARCADÆ. (6 Genera, 360 Species.) + +DISTRIBUTION.--The Arcadæ are universally distributed, and are most +abundant in warm seas. The genus _Leda_ is, however, abundant in Arctic and +Temperate regions, and _Solenella_ is confined to the South Temperate zone. +There are near 1,200 fossil species, found in all strata as low as the +Lower Silurian. + + +FAMILY 5.--TRIGONIADÆ. (1 Genus, 3 Species.) + +DISTRIBUTION.--The living _Trigoniæ_ are confined to Australia, but there +are 5 other genera fossil, containing about 150 species, and found in +various formations from the Chalk to the Lower Silurian. + + +FAMILY 6.--UNIONIDÆ. (7 Genera, 549 Species.) + +DISTRIBUTION.--The Unionidæ, or Fresh-water Mussels, are found in all the +fresh waters of the globe, but some of the genera are restricted. +_Castalia_, _Mycetopus_ and _Mulleria_ are confined to the rivers of South +America; _Anodon_, to the Nearctic and Palæarctic regions; _Iridina_, and +_Etheria_, to the rivers of Africa; _Unio_ has a universal distribution, +but is especially abundant in North America. About 60 fossil species are +found in the Tertiary and Wealden formations. + + +FAMILY 7.--CHAMIDÆ. (1 Genus, 50 Species.) + +DISTRIBUTION.--The Chamidæ, or Giant Clams, are confined to Tropical seas, +chiefly among coral reefs. There are two other genera and 62 species +fossil, ranging from the Chalk to the Oolite formations. + + +FAMILY 8.--HIPPURITIDÆ. (5 Genera, 103 Species.) + +Fossils of doubtful affinity, from the Chalk formation. + + +{535}FAMILY 9.--TRIDACNIDÆ. (1 Genus, 8 Species.) + +DISTRIBUTION.--The Tridacnidæ, or Clam-shells, are of very large size, and +are confined to the Tropical regions of the Indian and Pacific Oceans. A +few species have been found fossil in the Miocene formation. + + +FAMILY 10.--CARDIADÆ. (1 Genus, 200 Species.) + +DISTRIBUTION.--The Cardiadæ, or Cockles, are of world-wide distribution. +Another genus is fossil, and nearly 400 fossil species are known, ranging +back to the Upper Silurian formation. + + +FAMILY 11.--LUCINIDÆ (8 Genera, 178 Species.) + +DISTRIBUTION.--The Lucinidæ inhabit the Tropical and Temperate seas of all +parts of the world; but the genus _Corbis_ is confined to the Indian and +Pacific Oceans, _Montacuta_ and _Lepton_, to the Atlantic. There are nearly +500 extinct species, ranging from the Tertiary back to the Silurian +formation. + + +FAMILY 12.--CYCLADIDÆ. (3 Genera, 176 Species.) + +DISTRIBUTION.--The Cycladidæ are small fresh- or brackish-water shells +found all over the globe. The genus _Cyclas_ is most abundant in the North +Temperate zone, while _Cyrena_ inhabits the warmer shores of the Atlantic +and Pacific, but is absent from the West Coast of America. There are about +150 species fossil, ranging back from the Pliocene to the Wealden +formations. + + +FAMILY 13.--CYPRINIDÆ. (10 Genera, 176 Species). + +DISTRIBUTION.--Universal. _Cyprina_ and _Astarte_ are Arctic and North +Temperate; _Cardita_ is Tropical and South Temperate. There are several +extinct genera and about 1,000 species found in all formations as far back +as the Lower Silurian. + + +{536}FAMILY 14.--VENERIDÆ. (10 Genera, 600 Species.) + +DISTRIBUTION.--Universal. _Lucinopsis_ is confined to the North Atlantic; +_Glauconeza_ to the months of rivers in the Oriental region; _Meroe_ and +_Trigona_ to warm seas. There are about 350 fossil species, ranging back to +the Oolitic period. + + +FAMILY 15.--MACTRIDÆ. (5 Genera, 147 Species.) + +DISTRIBUTION.--All seas, but more abundant in the Tropics. _Gnathodon_ is +found in the Gulf of Mexico; _Anatinella_ in the Oriental region. There are +about 60 fossil species, ranging back to the Carboniferous period. + + +FAMILY 16.--TELLINIDÆ. (11 Genera, 560 Species.) + +DISTRIBUTION.--All seas; most abundant in the Tropics. _Galatea_ is +confined to African rivers. There are about 60 fossil species, mostly +Tertiary, but ranging back to the Carboniferous period. + + +FAMILY 17.--SOLENIDÆ. (3 Genera, 63 Species.) + +DISTRIBUTION.--All Temperate and Tropical seas. There are 80 fossil species +which range back to the Carboniferous epoch. + + +FAMILY 18.--MYACIDÆ. (6 Genera, 121 Species.) + +DISTRIBUTION.--All seas. _Panopæa_ inhabits both North and South Temperate +seas; _Glycimeris_, Arctic seas. There are near 350 fossil species, ranging +back to the Lower Oolite formation. + + +FAMILY 19.--ANATINIDÆ. (8 Genera, 246 Species.) + +DISTRIBUTION.--All seas. _Pholadomya_ is from Tropical Africa; _Myadora_ +from the Western Pacific; _Myochama_ and _Chamostræa_ are Australian. There +are about 400 fossil species, ranging back to the Lower Silurian formation. + + +{537}FAMILY 20.--GASTROCHÆNIDÆ. (5 Genera, 40 Species.) + +DISTRIBUTION.--Temperate and warm seas. _Aspergillum_ ranges from the Red +Sea to New Zealand. There are 35 fossil species, ranging back to the Lower +Oolite. + + +FAMILY 21.--PHOLADIDÆ (4 Genera, 81 Species.) + +DISTRIBUTION.--These burrowing molluscs inhabit all Temperate and warm seas +from Norway to New Zealand. There are about 50 fossil species, ranging back +to the epoch of the Lias. + + +_General Remarks on the Distribution of the Marine Mollusca._ + +The marine Mollusca are remarkable for their usually wide distribution. +About 48 of the families are cosmopolitan, ranging over both hemispheres, +and in cold as well as warm seas. About 15 are restricted to the warmer +seas of the globe; but several of these extend from Norway to New Zealand, +a distribution which may be called universal, and only 2 or 3 are +absolutely confined to Tropical seas. Two small families only, are confined +to the Pacific and Indian Oceans. Marine fishes, on the other hand, have a +much less cosmopolitan character, no less than 30 families having a limited +distribution, while 50 are universal. Some of these 30 families are +confined to the Northern seas, some to the Atlantic and Mediterranean, and +a considerable number to the Indian Ocean and Western Pacific. Many of +these families, it is true, are much smaller than those of the Mollusca, +which seem to possess very few of those small isolated families of two or +three species only, which abound in all the Vertebrate classes. These +differences are no doubt connected with the higher organisation of fishes, +which renders them more susceptible to changed conditions of life; and this +is indicated by the much less antiquity of existing families of fishes, the +greater part of which do not date back beyond the Cretaceous epoch, and +many of them only to the Eocene. In striking contrast we have the vast +antiquity of most of the families of {538}Mollusca, as shown in the +following table of their range taken from Mr. Woodward's work, but +re-arranged, and somewhat modified. + + 1 = Lower Silurian. 6 = Trias. 11 = Eocene. + 2 = Upper Silurian. 7 = Lower Oolite. 12 = Miocene. + 3 = Devonian. 8 = Upper Oolite. 13 = Pliocene. + 4 = Carboniferous. 9 = Lower Cretaceous. 14 = Recent. + 5 = Permian. 10 = Upper Cretaceous. + + +-----------------------------+--+--+--+--+--+--+--+--+--+--+--+--+--+--+ + | Range of Families of | | | | | | | | | | | | | | | + | Mollusca in Time; arranged | | | | | | | | | | | | | | | + | in their order of appearance| | | | | | | | | | | | | | | + | and disappearance. | 1| 2| 3| 4| 5| 6| 7| 8| 9|10|11|12|13|14| + +-----------------------------+--+--+--+--+--+--+--+--+--+--+--+--+--+--+ + | Productidæ |--|--|--|--|--| | | | | | | | | | + | Orthoceratidæ |--|--|--|--|--|--| | | | | | | | | + | Spiriferidæ, Orthidæ |--|--|--|--|--|--|--| | | | | | | | + | Atlantidæ, Hyaleidæ |--|--|--|--|--|--|--|--|--|--|--|--|--|--| + | Pyramidellidæ, Turbinidæ |--|--|--|--|--|--|--|--|--|--|--|--|--|--| + | Ianthidæ, Chitonidæ |--|--|--|--|--|--|--|--|--|--|--|--|--|--| + | Lingulidæ |--|--|--|--|--|--|--|--|--|--|--|--|--|--| + | Aviculidæ, Mytilidæ |--|--|--|--|--|--|--|--|--|--|--|--|--|--| + | Arcadæ, Trigoniadæ |--|--|--|--|--|--|--|--|--|--|--|--|--|--| + | Cyprinidæ, Anatinidæ |--|--|--|--|--|--|--|--|--|--|--|--|--|--| + | Nautilidæ | |--|--|--|--|--|--|--|--|--|--|--|--|--| + | Rhynchonellidæ, Craniadæ, } | | | | | | | | | | | | | | | + | Discinidæ } | |--|--|--|--|--|--|--|--|--|--|--|--|--| + | Cardiadæ, Lucinidæ | |--|--|--|--|--|--|--|--|--|--|--|--|--| + | Ammonitidæ | | |--|--|--|--|--|--|--|--| | | | | + | Naticidæ, Calyptræidæ | | |--|--|--|--|--|--|--|--|--|--|--|--| + | Dentalidæ, Terebratulidæ | | |--|--|--|--|--|--|--|--|--|--|--|--| + | Helicidæ | | | |--| | | | | | |--|--|--|--| + | Fissurellidæ, Tornatellidæ | | | |--|--|--|--|--|--|--|--|--|--|--| + | Pectinidæ, Solenidæ | | | |--|--|--|--|--|--|--|--|--|--|--| + | Cerithiadæ, Littorinidæ, } | | | | | | | | | | | | | | | + | Astartidæ } | | | | | |--|--|--|--|--|--|--|--|--| + | Belemnitidæ | | | | | | |--|--|--|--| | | | | + | Teuthidæ, Sepiadæ | | | | | | |--|--|--|--|--|--|--|--| + | Neritidæ, Patellidæ, } | | | | | | | | | | | | | | | + | Bullidæ } | | | | | | |--|--|--|--|--|--|--|--| + | Gastrochænidæ, Pholadidæ | | | | | | |--|--|--|--|--|--|--|--| + | Limnæidæ, Melaniadæ | | | | | | | |--|--|--|--|--|--|--| + | Chamidæ, Myadæ | | | | | | | |--|--|--|--|--|--|--| + | Cycladidæ, Veneridæ, } | | | | | | | | | | | | | | | + | Tellinidæ } | | | | | | | |--|--|--|--|--|--|--| + | Hippuritidæ | | | | | | | | |--|--| | | | | + | Unionidæ | | | | | | | | |--|--|--|--|--|--| + | Strombidæ, Buccinidæ | | | | | | | | |--|--|--|--|--|--| + | Conidæ, Volutidæ | | | | | | | | |--|--|--|--|--|--| + | Auriculidæ, Cyclostomidæ | | | | | | | | |--|--|--|--|--|--| + | Mactridæ | | | | | | | | |--|--|--|--|--|--| + | Limacidæ | | | | | | | | | | |--|--|--|--| + | Argonautidæ | | | | | | | | | | | | |--|--| + | Tridacnidæ | | | | | | | | | | | | |--|--| + +-----------------------------+--+--+--+--+--+--+--+--+--+--+--+--+--+--+ + +Nor is this enormous antiquity confined to family types alone. Many genera +are equally ancient. The genus _Lingula_ has {539}existed from the earliest +Palæozoic times down to the present day; while _Terebratula_, +_Rhynchonella_, _Discina_, _Nautilus_, _Natica_, _Pleurotomaria_, +_Patella_, _Dentalium_, _Mytilus_ and many other living forms, range back +to the Palæozoic epoch. That groups of such immense antiquity, and having +power to resist such vast changes of external conditions as they must have +been subject to, should now be widely distributed, is no more than might +reasonably be expected. It is only in the case of sub-genera and species, +that we can expect the influence of recent geological or climatal changes +to be manifest; and it must be left to special students to work out the +details of their distribution, with reference to the general principles +found to obtain among the more highly organised animals. + + + + +{540}CHAPTER XXIII. + +SUMMARY OF THE DISTRIBUTION, AND LINES OF MIGRATION, OF THE SEVERAL CLASSES +OF ANIMALS. + +Having already given summaries of the distribution of the several orders, +and of some of the classes of land animals, we propose here to make a few +general remarks on the special phenomena presented by the more important +groups, and to indicate where possible, the general lines of migration by +which they have become dispersed over wide areas. + + +MAMMALIA. + +This class is very important, and its past history is much better known +than that of most others. We shall therefore briefly summarise the results +we have arrived at from our examination of the distribution of extinct and +living forms of each order. + +_Primates._--This order, being pre-eminently a tropical one, became +separated into two portions, inhabiting the Eastern and Western Hemispheres +respectively, at a very early epoch. In consequence of this separation it +has diverged more radically than most other orders, so that the two +American families, Cebidæ and Hapalidæ, are widely differentiated from the +Apes, Monkeys, and Lemurs of the Old World. The Lemurs were probably still +more ancient, but being much lower in organisation, they became extinct in +most of the areas where the higher forms of Primates became developed. +Remains found in the Eocene formation indicate, that the North American and +European {541}Primates had, even at that early epoch, diverged into +distinct series, so that we must probably look back to the secondary period +for the ancestral form from which the entire order was developed. + +_Chiroptera._--These are also undoubtedly very ancient. The most +generalised forms--the Vespertilionidæ and Noctilionidæ--are the most +widely distributed; while special types have arisen in America, and in the +Eastern Hemisphere. Remains found in the Upper Eocene formation of Europe +differ little from species still living in the same countries; so that we +can form no conjecture as to the origin or migration, of the group. Their +power of flight would, however, enable them rapidly to spread over all the +great continents of the globe. + +_Insectivora._--This very ancient group, now probably verging towards +extinction, appears to have originated in the Northern continent, and never +to have reached Australia or South America. It may, however, have become +extinct in the latter country owing to the competition of the numerous +Edentata. The Insectivora now often maintain themselves amidst more highly +developed forms, by means of some special protection. Some burrow in the +earth,--like the moles; others have a spiny covering,--as the hedgehog's +and several of the Centetidæ; others are aquatic,---as the _Potamogale_ and +the desman; others have a nauseous odour,--as the shrews; while there are +several which seem to be preserved by their resemblance to higher +forms,--as the elephant-shrews to jerboas, and the tupaias to squirrels. +The same need of protection is shown by the numerous Insectivora inhabiting +Madagascar, where the competing forms are few; and by one lingering in the +Antilles, where there are hardly any other mammalia. + +_Carnivora._--Although perhaps less ancient than the preceding, this form +of mammal is far more highly organised, and from its earliest appearance +appears to have become dominant in the world. It would therefore soon +spread widely, and diverge into the various specialised types represented +by existing families. Most of these appear to have originated in the +Eastern Hemisphere, the only Carnivora occurring in North {542}American +Miocene deposits being ancestral forms of Canidæ and Felidæ. It seems +probable, therefore, that the order had attained a considerable development +before it reached the Western Hemisphere. The Procyonidæ, now confined to +America, are not very ancient; and the occurrence of a few allied forms in +the Himalayas (_Ælurus_ and _Æluropus_) render it probable that their +common ancestors entered North America from the Palæarctic region during +the Miocene period, but being a rather low type they have succumbed under +the competition of higher forms in most parts of the Eastern Hemisphere. +Bears and Weasels are probably still more recent emigrants to America. The +aquatic carnivora (Seals, &c.) are, as might be expected, more widely and +uniformly distributed, but there is little evidence to show at what period +the type was first developed. + +_Ungulata._--These are the dominant vegetable-feeders of the great +continents, and they have steadily increased in numbers and in +specialisation from the oldest Tertiary times to the present day. Being +generally of larger size and less active than the Carnivora, they have +somewhat more restricted powers of dispersal. We have good evidence that +their wide range over the globe is a comparatively recent phenomenon. +Tapirs and Llamas have probably not long inhabited South America, while +Rhinoceroses and Antelopes were once, perhaps, unknown in Africa, although +abounding in Europe and Asia. Swine are one of the most ancient types in +both hemispheres; and their great hardiness, their omnivorous diet, and +their powers of swimming, have led to their wide distribution. The sheep +and goats, on the other hand, are perhaps the most recent development of +the Ungulata, and they seem to have arisen in the Palæarctic region at a +time when its climate already approximated to that which now prevails. +Hence they are pre-eminently a Temperate group, never found within the +Tropics except upon a few mountain ranges. + +_Proboscidea._--These huge animals (the Elephants and Mastodons) appear to +have originated in the warmer parts of the Palæarctic region, but they soon +spread over all the great {543}continents, even reaching the southern +extremity of America. Their extinction has probably depended more on +physical than on organic changes, and we can clearly trace their almost +total disappearance to the effects of the Glacial epoch. + +_Rodentia._--Rodents are a very dominant group, and a very ancient one. +Owing to their small size and rapid powers of increase, they soon spread +over almost every part of the globe, whence has resulted a great +specialisation of family types in the South American continent which +remained so long isolated. They are capable of living wherever there is any +kind of vegetable food, hence their range will be determined rather by +organic than by physical conditions; and the occupation of a country by +enemies or by competing forms, is probably the chief cause which has +prevented many of the families from acquiring a wide range. The occurrence +of isolated species of the South American families, Octodontidæ and +Echimyidæ in the Ethiopian and Palæarctic regions, is an indication that +the range of many of the families has recently become less extensive. + +_Edentata._--These singular and lowly-organised animals appear to have +become almost restricted to the two great Southern lands--South Africa and +South America--at an early period; and, being there free from the +competition of higher forms, developed a number of remarkable types often +of huge size, of which the Megatherium is one of the best known. The +incursion of the highly-organised Ungulates and Carnivora into Africa +during the Miocene epoch, probably exterminated most of them in that +continent; but in America they continued in full force down to the +Post-Pliocene period; and even now, the comparatively diminutive Sloths, +Ant-eaters, and Armadillos, form a large and important portion of the +fauna. + +_Marsupialia and Monotremata._--These are probably the representatives of +the most ancient and lowly-organised types of mammal. They once existed in +the northern continents, whence they spread into Australia; and being +isolated, and preserved from the competition of the higher forms which soon +arose in other parts of the world, they have developed into a variety of +types, which, however, still preserve a general {544}uniformity of +organisation. One family, which continued to exist in Europe till the +latter part of the Miocene period, reached America, and has there been +preserved to our day. + +_Lines of Migration of the Mammalia._--The whole series of phenomena +presented by the distribution of the Mammalia, looked at broadly, are in +harmony with the view that the great continents and oceans of our own epoch +have been in existence, with comparatively small changes, during all +Tertiary times. Each one of them has, no doubt, undergone considerable +modifications in its area, its altitude, and in its connection with other +lands. Yet some considerable portion of each continent has, probably, long +existed in its present position, while the great oceans seem to have +occupied the same depressions of the earth's crust (varied, perhaps, by +local elevations and subsidences) during all this vast period of time. +Hence, allowing for the changes of which we have more or less satisfactory +evidence, the migrations of the chief mammalian types can be pretty clearly +traced. Some, owing to their small size and great vitality, have spread to +almost all the chief land masses; but the majority of the orders have a +more restricted range. All the evidence at our command points to the +Northern Hemisphere as the birth-place of the class, and probably of all +the orders. At a very early period the land communication with Australia +was cut off, and has never been renewed; so that we have here preserved for +us a sample of one or more of the most ancient forms of mammal. Somewhat +later the union with South America and South Africa was severed; and in +both these countries we have samples of a somewhat more advanced stage of +mammalian development. Later still, the union by a northern route between +the Eastern and Western Hemispheres appears to have been broken, partly by +a physical separation, but almost as effectually by a lowering of +temperature. About the same period the separation of the Palæarctic region +from the Oriental was effected, by the rise of the Himalayas and the +increasing contrast of climate; while the formation of the great +desert-belts of the Sahara, Arabia, Persia, and Central Asia, helped to +complete the separation of {545}the Temperate and Tropical zones, and to +render further intermigration almost impossible. + +In a few cases--of which the Rodents in Australia and the pigs in +Austro-Malaya are perhaps the most striking examples--the distribution of +land-mammals has been effected by a sea-passage either by swimming or on +floating vegetation; but, as a rule, we may be sure that the migrations of +mammalia have taken place over the land; and their presence on islands is, +therefore, a clear indication that these have been once connected with a +continent. The present class of animals thus affords the best evidence of +the past history of the land surface of our globe; and we have chiefly +relied upon it in sketching out (in Part III.) the probable changes which +each of our great regions has undergone. + + +_Birds._ + +Although birds are, of all land-vertebrates, the best able to cross seas +and oceans, it is remarkable how closely the main features of their +distribution correspond with those of the Mammalia. South America possesses +the low Formicaroid type of Passeres,--which, compared with the more highly +developed forms of the Eastern Hemisphere, is analogous to the Cebidæ and +Hapalidæ as compared with the Old World Apes and Monkeys; while its Cracidæ +as compared with the Pheasants and Grouse, may be considered parallel to +the Edentata as compared with the Ungulates of the Old World. The +Marsupials of America and Australia, are paralleled among birds, in the +Struthionidæ and Megapodiidæ; the Lemurs and Insectivora preserved in +Madagascar are represented by the Mascarene Dididæ; the absence of Deer and +Bears from Africa is analogous to the absence of Wrens, Creepers, and +Pheasants; while the African Hyracidæ and Chrysochloridæ among mammals, may +well be compared with the equally peculiar Coliidæ and Musophagidæ among +birds. + +From these and many other similarities of distribution, it is clear that +birds have, as a rule, followed the same great lines of migration as +mammalia; and that oceans, seas, and deserts, have {546}always to a great +extent limited their range. Yet these barriers have not been absolute; and +in the course of ages birds have been able to reach almost every habitable +land upon the globe. Hence have arisen some of the most curious and +interesting phenomena of distribution; and many islands, which are entirely +destitute of mammalia, or possess a very few species, abound in birds, +often of peculiar types and remarkable for some unusual character or habit. +Striking examples of such interesting bird-faunas are those of New Zealand, +the Sandwich Islands, the Galapagos, the Mascarene Islands, the Moluccas, +and the Antilles; while even small and remote islets,--such as Juan +Fernandez and Norfolk Island, have more light thrown upon their past +history by means of their birds, than by any other portion of their scanty +fauna. + +Another peculiar feature in the distribution of this class is the +extraordinary manner in which certain groups and certain external +characteristics, have become developed in islands, where the smaller and +less powerful birds have been protected from the incursions of mammalian +enemies, and where rapacious birds--which seem to some degree dependent on +the abundance of mammalia--are also scarce. Thus, we have the Pigeons and +the Parrots most wonderfully developed in the Australian region, which is +pre-eminently insular; and both these groups here acquire conspicuous +colours very unusual, or altogether absent, elsewhere. Similar colours +(black and red) appear, in the same two groups, in the distant Mascarene +islands; while in the Antilles the parrots have often white heads, a +character not found in the allied species on the South American continent. +Crests, too, are largely developed, in both these groups, in the Australian +region only; and a crested parrot formerly lived in Mauritius,--a +coincidence too much like that of the colours as above noted, to be +considered accidental. + +Again, birds exhibit to us a remarkable contrast as regards the oceanic +islands of tropical and temperate latitudes; for while most of the former +present hardly any cases of specific identity with the birds of adjacent +continents, the latter often show hardly any differences. The Galapagos and +Madagascar {547}are examples of the first-named peculiarity; the Azores and +the Bermudas of the last; and the difference can be clearly traced to the +frequency and violence of storms in the one case and to the calms or steady +breezes in the other. + +It appears then, that although birds do not afford us the same convincing +proof of the former union of now disjoined lands as we obtain from mammals, +yet they give us much curious and suggestive information as to the various +and complex modes in which the existing peculiarities of the distribution +of animals have been brought about. They also throw much light on the +relation between distribution and the external characters of animals; and, +as they are often found where mammalia are quite absent, we must rank them +as of equal value for the purposes of our present study. + + +_Reptiles._ + +These hold a somewhat intermediate place, as regards their distribution, +between mammals and birds, having on the whole rather a wider range than +the former, and a more restricted one than the latter. + +Snakes appear to have hardly more facilities for crossing the ocean than +mammals; hence they are generally absent from oceanic islands. They are +more especially a tropical group, and have thus never been able to pass +from one continent to another by those high northern and southern routes, +which we have seen reason to believe were very effectual in the case of +mammalia and some other animals. Hence we find no resemblance between the +Australian and Neotropical regions, or between the Palæarctic and Nearctic; +while the Western Hemisphere is comparatively poor as regards variety of +types, although rich in genera and species. Deserts and high mountains are +also very effectual barriers for this group, and their lines of migration +have probably been along river valleys, and occasionally across narrow seas +by means of floating vegetation. + +Lizards, being somewhat less tropical than snakes, may have passed by the +northern route during warm epochs. They are also more suited to traverse +deserts, and they possess some unknown {548}means of crossing the ocean, as +they are not unfrequently found in remote oceanic islands. These various +causes have modified their distribution. The Western Hemisphere is much +richer in lizards than it is in snakes; and it is also very distinct from +the Eastern Hemisphere. The lines of migration of lizards appear to have +been along the mountains and deserts of tropical countries, and, under +special conditions, across tropical seas from island to island. + +Crocodiles are a declining group. They were once more generally +distributed, all the three families being found in British Eocene deposits. +Being aquatic and capable of living in the sea, they can readily pass along +all the coasts and islands of the warmer parts of the globe. Tortoises are +equally ancient, and the restriction of certain groups to definite areas +seems to be also a recent phenomenon. + + +_Amphibia._ + +The Amphibia differ widely from Reptiles in their power of enduring cold; +one of their chief divisions, the Urodela or Tailed-Batrachia, being +confined to the temperate parts of the Northern Hemisphere. To this class +of animals the northern and southern routes of migration were open; and we +accordingly find a considerable amount of resemblance between South America +and Australia, and a still stronger affinity between North America and the +Palæarctic continent. The other tropical regions are more distinct from +each other; clearly indicating that, in this group, it is tropical deserts +and tropical oceans which are the barriers to migration. The class however +is very fragmentary, and probably very ancient; so that descendants of once +widespread types are now found isolated in various parts of the globe, +between which we may feel sure there has been no direct transmission of +Batrachia. Remembering that their chief lines of migration have been by +northern and southern land-routes, by floating ice, by fresh-water +channels, and perhaps at rare intervals by ova being carried by aquatic +birds or by violent storms,--we shall be able to comprehend most of the +features of their actual distribution. + + +{549}_Fresh-water Fishes._ + +Although it would appear, at first sight, that the means of dispersal of +these animals are very limited, yet they share to some extent the wide +range of other fresh-water organisms. They are found in all climates; but +the tropical regions are by far the most productive, and of these South +America is perhaps the richest and most peculiar. There is a certain amount +of identity between the two northern continents, and also between those of +the South Temperate zone; yet all are radically distinct, even North +America and Europe having but a small proportion of their forms in common. +The occurrence of allied fresh-water species in remote lands--as the +_Aphritis_ of Tasmania and Patagonia, and the _Comephorus_ of Lake Baikal, +distantly allied to the mackerels of Northern seas--would imply that marine +fishes are often modified for a life in fresh waters; while other facts no +less plainly show that permanent fresh-water species are sometimes +dispersed in various ways across the oceans, more especially by the +northern and southern routes. + +The families of fresh-water fishes are often of restricted range, although +cases of very wide and scattered distribution also occur. The great +zoological regions are, on the whole, very well characterized; showing that +the same barriers are effectual here, as with most other vertebrates. We +conclude, therefore, that the chief lines of migration of fresh-water +fishes have been across the Arctic and Antarctic seas, probably by means of +floating ice as well as by the help of the vast flocks of migratory aquatic +birds that frequent those regions. On continents they are, usually, widely +dispersed; but tropical seas, even when of small extent, appear to have +offered an effectual barrier to their dispersal. The cases of affinity +between Tropical America, Africa, Asia, and Australia, must therefore be +imputed either to the survival of once widespread groups, or to analogous +adaptation to a fresh-water life of wide-spread marine types; and these +cases cannot be taken as evidence of any former land connection between +such remote continents. + + +{550}_Insects._ + +It has already been shown (Vol. I. pp. 209-213 and Vol. II. pp. 44-48) that +the peculiarities of distribution of the various groups of insects depend +very much on their habits and general economy. Their antiquity is so vast, +and their more important modifications of structure have probably occurred +so slowly, that modes of dispersal depending on such a combination of +favourable conditions as to be of excessive rarity, may yet have had time +to produce large cumulative effects. Their small specific gravity and their +habits of flight render them liable to dispersal by winds to an extent +unknown in other classes of animals; and thus, what are usually very +effectual barriers have been overstepped, and sometimes almost obliterated, +in the case of insects. A careful examination will, however, almost always +show traces of an ancient fauna, agreeing in character with other classes +of animals, intermixed with the more prominent and often more numerous +forms whose presence is due to this unusual facility of dispersal. + +The effectual migration of insects is, perhaps more than in any other class +of animals, limited by organic and physical conditions. The vegetation, the +soil, the temperature, and the supply of moisture, must all be suited to +their habits and economy; while they require an immunity from enemies of +various kinds, which immigrants to a new country seldom obtain. Few +organisms have, in so many complex ways, become adapted to their special +environment, as have insects. They are in each country more or less adapted +to the plants which belong to it; while their colours, their habits, and +the very nature of the juices of their system, are all modified so as to +protect them from the special dangers which surround them in their native +land. It follows, that while no animals are so well adapted to show us the +various modes by which dispersal may be effected, none can so effectually +teach us the true nature and vast influence of the organic barrier in +limiting dispersal. + +It is probable that insects have at one time or another taken advantage of +every line of migration by which any terrestrial {551}organisms have spread +over the earth, but owing to their small size and rapid multiplication, +they have made use of some which are exclusively their own. Such are the +passage along mountain ranges from the Arctic to the Antarctic regions, and +the dispersal of certain types over all temperate lands. It will perhaps be +found that insects have spread over the land surface in directions +dependent on our surface zones--forests, pastures, and deserts;--and a +study of these, with a due consideration of the fact that narrow seas are +scarcely a barrier to most of the groups, may assist us to understand many +of the details of insect-distribution. + + +_Terrestrial Mollusca._ + +The distribution of land-shells agrees, in some features, with that of +insects, while in others the two are strongly contrasted. In both we see +the effects of great antiquity, with some special means of dispersal; but +while in insects the general powers of motion, both voluntary and +involuntary, are at a maximum, in land-molluscs they are almost at a +minimum. Although to some extent dependent on vegetation and climate, the +latter are more dependent on inorganic conditions, and also to a large +extent on the general organic environment. The result of these various +causes, acting through countless ages, has been to spread the main types of +structure with considerable uniformity over the globe; while generic and +sub-generic forms are often wonderfully localized. + +Land-shells, even more than insects, seem, at first sight, to require +regions of their own; but we have already pointed out the disadvantages of +such a method of study. It will be far more instructive to refer them to +those regions and sub-regions which are found to accord best with the +distribution of the higher animals, and to consider the various anomalies +they present as so many problems, to be solved by a careful study of their +habits and economy, and especially by a search after the hidden causes +which have enabled them to spread so widely over land and ocean. + +The lines of migration which land-shells have followed, can {552}hardly be +determined with any definiteness. On continents they seem to spread +steadily, but slowly, in every direction, checked probably by organic and +physical conditions rather than by the barriers which limit the higher +groups. Over the ocean they are also slowly dispersed, by some means which +act perhaps at very long intervals, but which, within the period of the +duration of genera and families, are tolerably effective. It thus happens +that, although the powers of dispersal of land-shells and insects are so +very unequal, the resulting geographical distribution is almost the +opposite of what might have been expected,--the former being, on the whole, +less distinctly localized than the latter. + + +CONCLUSION. + +The preceding remarks are all I now venture to offer, on the distinguishing +features of the various groups of land-animals as regards their +distribution and migrations. They are at best but indications of the +various lines of research opened up to us by the study of animals from the +geographical point of view, and by looking upon their range in space and +time as an important portion of the earth's history. Much work has yet to +be done before the materials will exist for a complete treatment of the +subject in all its branches; and it is the author's hope that his volumes +may lead to a more systematic collection and arrangement of the necessary +facts. At present all public museums and private collections are arranged +zoologically. All treatises, monographs, and catalogues, also follow, more +or less completely, the zoological arrangement; and the greatest difficulty +the student of geographical distribution has to contend against, is the +total absence of geographical collections, and the almost total want of +complete and comparable local catalogues. Till every well-marked +district,--every archipelago, and every important island, has all its known +species of the more important groups of animals catalogued on a uniform +plan, and with a uniform nomenclature, a thoroughly satisfactory account of +the Geographical Distribution of Animals will not be possible. But more +than this is wanted. Many of the most curious relations between animal +{553}forms and their habitats, are entirely unnoticed, owing to the +productions of the same locality _never_ being associated in our museums +and collections. A few such relations have been brought to light by modern +scientific travellers, but many more remain to be discovered; and there is +probably no fresher and more productive field still unexplored in Natural +History. Most of these curious and suggestive relations are to be found in +the productions of islands, as compared with each other, or with the +continents of which they form appendages; but these can never be properly +studied, or even discovered, unless they are visibly grouped together. When +the birds, the more conspicuous families of insects, and the land-shells of +islands, are kept together so as to be readily compared with similar +associations from the adjacent continents or other islands, it is believed +that in almost every case there will be found to be peculiarities of form +or colour running through widely different groups, and strictly indicative +of local or geographical influences. Some of these coincident variations +have been alluded to in various parts of this work, but they have never +been systematically investigated. They constitute an unworked mine of +wealth for the enterprising explorer; and they may not improbably lead to +the discovery of some of the hidden laws (supplementary to Natural +Selection), which seem to be required, in order to account for many of the +external characteristics of animals. + +In concluding his task, the author ventures to suggest, that naturalists +who are disposed to turn aside from the beaten track of research, may find +in the line of study here suggested a new and interesting pursuit, not +inferior in attractions to the lofty heights of transcendental anatomy, or +the bewildering mazes of modern classification. And it is a study which +will surely lead them to an increased appreciation of the beauty and the +harmony of nature, and to a fuller comprehension of the complex relations +and mutual interdependence, which link together every animal and vegetable +form, with the ever-changing earth which supports them, into one grand +organic whole. + + +{557}GENERAL INDEX. + + +All names in Italics refer, either to the genera and other groups of +Extinct Animals in Part II. of the First Volume;--or to the genera whose +distribution is given under Geographical Zoology (Part IV.) in the Second +Volume; the Families and higher groups being in small capitals. All other +references are in ordinary type. + +The various matters discussed under Zoological Geography (Part III.), are +indexed as much as possible by subjects and localities. None of the genera +mentioned in this Part are indexed, as this would have more than doubled +the extent of the Index, and would have served no useful purpose, because +the general distribution of each genus is given in Part IV., and the +separate details can always be found by referring to the region, +sub-region, and class. + + + A. + + Aard-vark of East Africa, figure of, i. 261 + + Aard-vark, ii. 246 + + Aard-wolf, ii. 196 + + _Abacetus_, ii. 491 + + _Abax_, ii. 489 + + _Abisara_, ii. 475 + + _Ablabes_, ii. 375 + + _Ablepharus_, ii. 395 + + _Abramis_, ii. 453 + + _Abronia_, ii. 392 + + _Abrornis_, ii. 258 + + _Abrostomus_, ii. 451 + + _Aburria_, ii. 343 + + _Acalyptus_, ii. 384 + + _Acanthias_, ii. 401 + + _Acanthicus_, ii. 444 + + _Acanthion_, ii. 240 + + _Acanthis_, ii. 283 + + _Acanthisitta_, ii. 265 + + _Acanthiza_, ii. 258 + + _Acanthobrama_, ii. 453 + + ACANTHOCLINIDÆ, ii. 432 + + _Acanthodactylus_, ii. 391 + + _Acanthodelphis_, ii. 209 + + _Acantholabrus_, ii. 437 + + _Acanthomys_, ii. 229 + + _Acanthophis_, ii. 383 + + _Acanthopsis_, ii. 453 + + ACANTHOPTERYGII, ii. 424 + + ACANTHOPTERYGII PHARYNCOGNATHI, ii. 437 + + _Acanthopthalmus_, ii. 453 + + _Acanthoptila_, ii. 261 + + _Acanthopyga_, ii. 390 + + _Acanthorhynchus_, ii. 275 + + _Acanthosaura_, ii. 402 + + _Acara_, ii. 438 + + Accentor, ii. 260 + + Accentorinæ, ii. 257 + + _Accipenser_, ii. 459 + + ACCIPENSERIDÆ, ii. 459 + + _Accipiter_, ii. 348 + + _Accipitres_, European Eocene, i. 163 + + Accipitres, classification of, i. 97 + range of Palæarctic genera of, i. 248 + range of Ethiopian genera of, i. 312 + range of Oriental genera of, i. 385 + range of Australian genera of, i. 484 + + ACCIPITRES, ii. 345 + general remarks on the distribution of, ii. 351 + + ACCIPITRINÆ, ii. 347 + + _Acerina_, ii. 425 + + _Aceros_, ii. 317 + + _Acerotherium_, ii. 214 + + _Acerotherium_, European Miocene, i. 119 + N. American Tertiary, i. 136 + + _Acestra_, ii. 444 + + _Acestura_, ii. 108 + + _Achalinus_, ii. 375 + + _Acharnes_, ii. 434 + + _Achatina_, ii. 515 + + _Achatinella_, ii. 514 + + _Acherontia_, ii. 483 + + _Achilognathus_, ii. 452 + + _Achænodon_, N. American Tertiary, i. 138 + + _Acicula_, ii. 519 + + ACICULIDÆ, ii. 519 + + _Acmæodera_, ii. 497 + + _Acodon_, ii. 230 + + ACONTIADÆ, ii. 399 + + _Acontias_, ii. 399 + + _Acotherium_, European Eocene, i. 126 + + _Acotherium_, ii. 215 + + _Acræa_, ii. 473 + + ACRÆIDÆ, ii. 473 + + _Acrantus_, ii. 390 + + _Acredula_, ii. 266 + + _Acridotheres_, ii. 287 + + _Acris_, ii. 419 + + _Acrobata_, ii. 252 + + _Acrocephalus_, ii. 258 + + _Acrochilus_, ii. 452 + + ACROCHORDIDÆ, ii. 382 + + _Acrochordonichthys_, ii. 442 + + _Acrochordus_, ii. 382 + + ACRONURIDÆ, ii. 433 + + _Acropternis_, ii. 297 + + _Acryllium_, ii. 340 + + _Actenodes_, ii. 497 + + _Actinodura_, ii. 261 + + _Ada_, ii. 390 + + _Adamsiella_, ii. 521 + + _Adapis_, European Eocene, i. 125 + + _Addax_, ii. 223 + + _Adelomia_, ii. 108 + + _Adelotopus_, ii. 490 + + _Adenomera_, ii. 416 + + _Adolias_, ii. 474 + + _Aedon_, ii. 259 + + _Ægeria_, ii. 482 + + ÆGERIIDÆ, ii. 482 + + _Ægialitis_, ii. 356 + + _Ægithaliscus_, ii. 266 + + _Ægithalus_, ii. 266 + + _Ægocera_, ii. 482 + + _Ægotheles_, ii. 318 + + _Æluredus_, ii. 275 + + _Ælurichthys_, ii. 443 + + ÆLURIDÆ, ii. 201 + + _Ælurogale_, European Eocene, i. 125 + + _Æluropus_, ii. 201 + + _Ælurus_, ii. 201 + + _Æmona_, ii. 472 + + _Ænigma_, ii. 490 + + _Æolidæ_, ii. 530 + + _Æpyceros_, ii. 223 + + _Æpyornis_, of Madagascar, i. 164 + + _Æpyornis_, ii. 370 + + ÆPYORNITHIDÆ, ii. 370 + + _Æsacus_, ii. 355 + + _Æshna_, from the Lias, i. 167 + + _Æthopyga_, ii. 276 + + _Æthya_, ii. 364 + + _Agama_, ii. 402 + + AGAMIDÆ, ii. 401 + + _Agapornis_, ii. 328 + + _Agarista_, ii. 482 + + _Agaristidæ_, ii. 482 + + _Agelasta_, ii. 501 + + AGELASTINÆ, ii. 340 + + _Agelæus_, ii. 282 + + _Ageniosus_, ii. 443 + + _Ageronia_, ii. 474 + + _Aglæactis_, ii. 108 + + _Agnopterus_, European Eocene, i. 163 + + _Agoniates_, ii. 445 + + _Agonostoma_, ii. 435 + + _Agonus_, ii. 428 + + Agouti, ii. 241 + + _Agra_, ii. 490 + + _Agraulis_, ii. 474 + + _Agrilus_, ii. 497 + + _Agriochoerus_, N. American Tertiary, i. 138 + + _Agrion_, from the Lias, i. 167 + + _Agriornis_, ii. 100 + + _Ahætulla_, ii. 379 + + _Ailia_, ii. 442 + + _Aipysurus_, ii. 384 + + _Aithurus_, ii. 107 + + _Aix_, ii. 363 + + _Akysis_, ii. 442 + + _Alæmon_, ii. 289 + + Alaska, birds of, ii. 136 + + _Alauda_, ii. 289 + + ALAUDIDÆ, ii. 289 + + Albatrosses, ii. 365 + + _Albulichthys_, ii. 452 + + _Alburnus_, ii. 453 + + _Alca_, ii. 367 + + _Alcadia_, ii. 522 + + ALCEDINIDÆ, ii. 315 + + _Alcedo_, ii. 316 + + ALCEPHALINÆ, ii. 224 + + _Alcephalus_, Indian Miocene, i. 122 + + _Alcephalus_, ii. 224 + + _Alces_, ii. 219 + + ALCIDÆ, ii. 367 + + _Alcippe_, ii. 261 + + _Alcurus_, ii. 267 + + _Alcyone_, ii. 316 + + Aldabra Islands, land-tortoises of, i. 289 + + _Alectorurus_, ii. 100 + + _Alectroenas_, ii. 332 + + ALEPOCEPHALIDÆ, ii. 454 + + _Alesa_, ii. 475 + + _Alestes_, ii. 445 + + _Alethe_, ii. 262 + + _Aletornis_, N. American Eocene, i. 163 + + Aleutian Islands, birds of, ii. 137 + + Algeria, Post-Pliocene deposits and caves of, i. 111 + + _Algira_, ii. 391 + + Alleghany sub-region, ii. 130 + mammalia of, ii. 132 + birds of, ii. 132 + reptiles of, ii. 133 + amphibia of, ii. 134 + fishes of, ii. 134 + islands of, ii. 134 + + Allen, Mr. J. A., on Zoological regions, i. 61 + objections to his system of circumpolar zones, i. 67 + objections to his zoo-geographical nomenclature, i. 68 + on birds of N. America, ii. 133, 136 + + _Alligator_, ii. 406 + + ALLIGATORIDÆ, ii. 406 + + Alligators, ii. 406 + + _Allotinus_, ii. 477 + + _Alopecias_, ii. 460 + + _Alsæcomus_, ii. 332 + + _Alsecus_, ii. 259 + + _Alseonax_, ii. 270 + + _Alsodes_, ii. 417 + + Altai mountains, fossils in caves, i. 111 + + _Alytes_, ii. 417 + + ALYTIDÆ, ii. 417 + + _Amadina_, ii. 287 + + _Amara_, ii. 489 + + _Amarynthis_, ii. 476 + + _Amathusia_, ii. 472 + + _Amauresthes_, ii. 287 + + _Amaurospiza_, ii. 285 + + _Amazilia_, ii. 109 + + _Amblyrhiza_, Pliocene of Antilles, i. 148 + + AMBLYCEPHALIDÆ, ii. 380 + + _Amblycephalus_, ii. 380 + + _Amblyceps_, ii. 443 + + _Amblychila_, ii. 487 + + _Amblymora_, ii. 501 + + _Amblyopsis_, ii. 450 + + _Amblyornis_, ii. 275 + + _Amblypharyngodon_, ii. 452 + + _Amblypodia_, ii. 477 + + _Amblyrhamphus_, ii. 282 + + _Amblyrhiza_, ii. 237 + + _Amblyrhynchichthys_, ii. 452 + + _Amblystoma_, ii. 413 + + _Ameiva_, ii. 390 + + America, recent separation of North and South, i. 40 + extinct mammalia of, i. 129 + N., Post-Pliocene fauna of, i. 129 + + American Creepers, ii. 295 + + AMIIDÆ, ii. 458 + + _Amiurus_, ii. 442 + + _Ammodromus_, ii. 284 + + _Ammodytes_, ii. 440 + + _Ammomanes_, ii. 289 + + AMMONITIDÆ, ii. 506 + + _Amomys_, N. American Tertiary, i. 134 + + _Ampeliceps_, ii. 287 + + AMPELIDÆ, ii. 280 + + _Ampelio_, ii. 102 + + _Ampelis_, ii. 280 + + _Amphechinus_, European Miocene, i. 117 + ii. 188 + + Amphibia, means of dispersal of, i. 28 + classification of, i. 100 + peculiar to Palæarctic region, i. 186 + of Central Europe, i. 196 + of the Mediterranean sub-region, i. 205 + of the Siberian sub-region, i. 220 + of the Manchurian sub-region, i. 226 + table of Palæarctic families of, i. 237 + of the Ethiopian region, i. 255 + of West Africa, i. 264 + S. African, i. 268 + of Madagascar, i. 280 + table of Ethiopian families of, i. 298 + of the Oriental region, i. 317 + of the Indian sub-region, i. 326 + of Ceylon, i. 327 + of the Indo-Chinese sub-region, i. 331 + of the Indo-Malay sub-region, i. 340 + table of Oriental families of, i. 369 + of the Australian region, i. 397 + resemblances of Australian and South-American, i. 400 + of New Guinea, i. 416 + of New Zealand, i. 457 + Neotropical, ii. 11 + of South Temperate America, ii. 41 + of the Mexican sub-region, ii. 54 + of the Antilles, ii. 72 + table of Neotropical families of, ii. 89 + of the Nearctic region, ii. 120 + of California, ii. 128 + of Central North America, ii. 131 + of Eastern United States, ii. 134 + table of Nearctic families of, ii. 143 + + AMPHIBIA, ii. 411 + general remarks on the distribution of, ii. 422 + fossil, ii. 423 + summary and conclusion, ii. 548 + lines of migration of, ii. 548 + + _Amphibola_, ii. 510 + + _Amphibos_, Indian Miocene, i. 122 + ii. 225 + + _Amphicyon_, European Miocene, i. 118 + Indian Miocene, i. 121 + N. American Tertiary, i. 134 + ii. 198 + ii. 202 + + _Amphiglossus_, ii 398 + + _Amphimericidæ_, European Miocene, i. 119 + + _Amphimoschus_, European Miocene, i. 120 + ii. 220 + + _Amphioxus_, ii. 464 + + _Amphipnous_, ii. 455 + + _Amphisbæna_, ii. 389 + + AMPHISBÆNIDÆ, ii. 388 + + _Amphisorex_, European Miocene, i. 118 + ii. 191 + + _Amphitragulus_, European Miocene, i. 120 + ii. 218 + + _Amphiuma_, ii. 412 + + AMPHIUMIDÆ, ii. 412 + + _Amphixestus_, ii. 397 + + _Ampullaria_, ii. 510 + + _Amydrus_, ii. 288 + + _Amytis_, ii. 258 + + _Anabatoides_, ii. 103 + + _Anabazenops_, ii. 103 + + _Anableps_, ii. 450 + + ANACANTHINI, ii. 439 + + _Anacyrtus_, ii. 445 + + _Anadenus_, ii. 517 + + _Anadia_, ii. 393 + + ANADIADÆ, ii. 393 + + _Anæretes_, ii. 101, 291 + + _Anaides_, ii. 413 + + _Analcipus_, ii. 268 + + _Anarhynchus_, ii. 356 + + _Anarrhichas_, ii. 431 + + _Anas_, ii. 363 + + _Anastoma_, European Tertiary, i. 169 + ii. 527 + + _Anastomus_, ii. 361 + + ANATIDÆ, ii. 363 + + ANATINIDÆ, ii. 536 + + _Anatinella_, ii. 536 + + _Anausorex_, ii. 191 + + _Anchilophus_, European Eocene, i. 125 + + _Anchippodus_, N. American Eocene, i. 139 + + _Anchippus_, N. American Tertiary, i. 135 + + _Anchitheridæ_, N. American Tertiary, i. 135 + ii. 212 + + _Anchitherium_, European Miocene, i. 119 + European Eocene, i. 125 + N. American Tertiary, i. 135 + + Ancient fauna of New Zealand, i. 459 + + _Ancistrops_, ii. 103 + + _Ancylotherium_, Miocene of Greece, i. 116 + European Miocene, i. 121 + + _Ancylotherium_, ii. 246 + + _Ancyluris_, ii. 476 + + _Ancylus_, ii. 518 + + Andaman Islands, zoology of, i. 333 + probable past history of, i. 334 + + _Andigena_, ii. 307 + + _Andrias_, European Miocene, i. 165 + + _Androdon_, ii. 107 + + _Andropadus_, ii. 267 + + _Aneitea_, ii. 517 + + _Anguilla_, ii. 456 + + _Anguis_, ii. 397 + + Angwantibo, ii. 176 + + Animal kingdom, primary divisions of, i. 85 + + Animals, development of, affecting distribution, i. 7 + dispersal and migration of, i. 10 + rapid multiplication of, i. 10 + + _Anisacodon_, N. American Tertiary, i. 137 + + Anoa of Celebes, peculiarities of, i. 428 + + _Anoa_, ii. 222 + + _Anodon_, ii. 534 + + _Anolius_, ii. 400 + + _Anomalurus_, ii. 235 + + _Anomalpus_, ii. 397 + + _Anoplodipsas_, ii. 381 + + _Anoplotheriidæ_, European Miocene, i. 119 + + _Anoplotherium_, European Miocene, i. 119 + European Eocene, i. 126 + S. American Eocene, i. 1 + + _Anopthalmus_, ii. 489 + + _Anostomus_, ii. 445 + + _Anous_, ii. 365 + + _Anser_, ii. 363 + + _Anseranas_, ii. 363 + + Anseres, arrangement of, i. 98 + peculiar Palæarctic genera of, i. 250 + peculiar Ethiopian genera of, i. 313 + peculiar Australian genera of, i. 485 + + ANSERES, general remarks on the distribution of, ii. 367 + + _Antarctia_, ii. 490 + ii. 492 + + Ant-eaters, ii. 247 + + _Antechinomys_, ii. 249 + + _Antechinus_, ii. 249 + + Antelopes in the Indian Miocene deposits, i. 122 + birthplace and migrations of, i. 155 + Palæarctic, i. 182 + ii. 221 + + _Antelotherium_, Indian Miocene, i. 122 + + _Antennarius_, ii. 431 + + _Anteros_, ii. 476 + + _Anthia_, ii. 491 + + _Anthipes_, ii. 270 + + _Anthocharis_, ii. 478 + + _Anthochæra_, ii. 275 + + _Anthornis_, ii. 275 + + _Anthracotheridæ_, N. American Tertiary, i. 137 + + _Anthracotherium_, European Miocene, i. 119 + ii. 215 + + _Anthreptes_, ii. 276 + + Anthropoid apes, ii. 170 + + _Anthropoides_, ii. 357 + + _Anthus_, ii. 290 + + _Antiacodon_, N. American Tertiary, i. 133 + + Antillean sub-region, ii. 61 + mammalia of, ii. 62 + birds of, ii. 64 + illustration of zoology of, ii. 67 + table of distribution of resident land-birds of, ii. 68 + reptiles and amphibia of, ii. 72 + fresh-water fish of, ii. 73 + insects of, ii. 73 + land-shells of, ii. 75 + past history of, ii. 78 + + Antilles, Pliocene Mammalia of, i. 148 + land-shells of, ii. 526 + + _Antilocapra_, ii. 223 + + ANTILOCAPRINÆ, ii. 223 + + _Antilope_, Post-Pliocene, i. 112 + in Brazilian caves, i. 144 + ii. 223 + ii. 226 + + Antiquity of the genera of insects, i. 166 + of the genera of land and fresh-water shells, i. 168 + + _Antrostomus_, ii. 319 + + Ant-thrushes, ii. 297 + + _Anumbius_, ii. 103 + + ANURA, ii. 414 + + _Anurosorex_, ii. 190 + + _Aonyx_, ii. 199 + + _Apalis_, ii. 258 + + _Apaloderma_, ii. 314 + + _Apatura_, ii. 474 + + _Aphanapteryx_ of Mauritius, i. 164 + ii. 352 + + _Aphantocera_, ii. 107 + + _Aphelotherium_, European Eocene, i. 125 + + _Aphneus_, ii. 477 + + _Aphobus_, ii. 283 + + APHREDODERIDÆ, ii. 425 + + _Aphritis_, ii. 428 + ii. 549 + + _Aphriza_, ii. 356 + + _Aphysiadæ_, ii. 530 + + _Aphyocypris_, ii. 452 + + _Aplocerus_, ii. 224 + + _Aplodontia_, ii. 236 + + _Aplonis_, ii. 288 + + _Aplopelia_, ii. 383 + + _Apodemia_, ii. 476 + + _Apogon_, ii. 425 + + _Aprasia_, ii. 396 + + APRASIADÆ, ii. 396 + + _Aprosmictus_, ii. 325 + + _Aptenodytes_, ii. 366 + + APTERYGIDÆ, ii. 369 + + _Apteryx_, ii. 369 + + _Apua_, ii. 453 + + _Aquila_, European Miocene, i. 161 + ii. 348 + + AQUILINÆ, ii. 348 + + _Ara_, ii. 328 + + _Arachnechthra_, ii. 276 + + _Arachnothera_, ii. 277 + + ARAMIDÆ, ii. 357 + + _Aramides_, ii. 352 + + _Aramus_, ii. 357 + + _Arapaima_, ii. 454 + + _Arborophila_, ii. 338 + + ARCADÆ, ii. 534 + + _Archæomys_, ii. 238 + + _Archæopteryx_, Bavarian Oolite, i. 163 + + _Archibuteo_, ii. 348 + + Arctic shells, ii. 518 + zone not a separate region, i. 68 + + _Arctitis_, ii. 195 + + _Arctocebus_, ii. 176 + + _Arctocephalus_, ii. 202 + + _Arctocyon_, European Eocene, i. 125 + ii. 206 + + _Arctodus_, N. American Post-Pliocene, i. 130 + ii. 202 + + _Arctogale_, ii. 195 + + _Arctomys_, European Pliocene, i. 113 + ii. 235, 236 + + _Arctonyx_, ii. 199 + + _Arctopithecus_, ii. 244 + + _Arctotherium_ in Brazilian caves, i. 144 + S. American Pliocene, i. 146 + + _Ardea_, ii. 359 + + ARDEIDÆ, ii. 359 + + _Ardistomus_, ii. 490 + + _Argentina_, ii. 488 + + _Arges_, ii. 444 + + ARGONAUTIDÆ. ii. 505 + + _Argus pheasant_, figure of, i. 339 + peculiarity in display of plumage, and confirmation of Mr. Darwin's + views, i. 340 + + _Argusianus_, ii. 340 + + _Argutor_, ii. 489 + + _Argynnis_, ii. 474 + + _Aricoris_, ii. 476 + + _Ariella_, ii. 195 + + _Arinia_, ii. 520 + + _Arion_, ii. 517 + + _Aristobia_, ii. 501 + + _Arius_, ii. 443 + + Armadillos, ii. 245 + + _Arnoglossus_, ii. 441 + + _Aromochelys_, ii. 408 + + _Arremon_, ii. 99 + + _Arrhenotus_, ii. 501 + + _Artamia_, ii. 268 + ii. 271 + ii. 288 + + ARTAMIDÆ, ii. 288 + + _Artamides_, ii. 269 + + _Artamus_, ii. 288 + + _Arthroleptis_, ii. 421 + + _Artiodactyla_, European Eocene, i. 126 + N. American Tertiary, i. 137 + S. American Pliocene, i. 146 + + _Artomyias_, ii. 270 + + _Arundinicola_, ii. 100 + + _Arvicola_, European Pliocene, i. 113 + in Brazilian caves, i. 145 + S. American Pliocene, i. 147 + S. American Eocene, i. 148 + + _Arvicola_, ii. 230, 231 + + _Asio_, ii. 350 + + _Aspergillum_, ii. 537 + + _Aspidoparia_, ii. 452 + + _Aspidorhinus_, ii. 391 + + _Aspidura_, ii. 374 + + _Aspius_, ii. 453 + + _Aspredo_, ii. 444 + + _Aspro_, ii. 425 + + _Astarte_, ii. 535 + + _Astathes_, ii. 501 + + _Asterophys_, ii. 421 + + _Asterophysus_, ii. 443 + + _Asthenodipsas_, ii. 381 + + _Astrapia_, ii. 274 + + _Astroblepus_, ii. 444 + + _Astur_, ii. 348 + + _Asturina_, ii. 348 + + _Asturinula_, ii. 348 + + ATELEOPODIDÆ, ii. 440 + + _Ateles_, ii. 174 + + _Atelopus_, ii. 416 + + _Atelornis_, ii. 312 + + _Aterica_, ii. 474 + + _Athene_, ii. 350 + + _Atherina_, ii. 434 + + _Atherinichthys_, ii. 434 + + ATHERINIDÆ, ii. 434 + + _Atheris_, ii. 386 + + _Atherura_, ii. 240 + + _Athylax_, ii. 195 + + _Athyma_, ii. 474 + + ATLANTIDÆ, ii. 531 + + _Atlapetes_, ii. 284 + + ATRACTASPIDIDÆ, ii. 383 + + _Atractaspis_, ii. 383 + + _Atretium_, ii. 375 + + _Atrichia_, ii. 299 + + ATRICHIIDÆ, ii. 299 + + _Atropos_, ii. 385 + + _Attagis_, ii. 354 + + ATTALINÆ, ii. 293 + + _Atthis_, ii. 108 + + _Atticora_, ii. 281 + + _Attila_, ii. 102 + + _Auchenaspis_, ii. 443 + + _Auchenia_, N. American Post-Pliocene, i. 130 + ii. 217 + + _Auchenipterus_, ii. 443 + + Auckland Islands, birds of, i. 455 + + _Augastes_, ii. 108 + + _Auks_, ii. 367 + + _Aulia_, ii. 102 + + _Aulacodes_, ii. 239 + + _Aulacodon_, ii. 239 + + _Aulacorhamphus_, ii. 307 + + _Aulopoma_, ii. 520 + + _Aulopyge_, ii. 452 + + _Auricula_, ii. 519, 527 + + AURICULIDÆ, ii. 518 + + _Auriparus_, ii. 266 + + Australia, physical features of, i. 387 + + Australia and S. America, supposed land connection between, i. 398 + + Australian region, description of, i. 387 + zoological characteristics of, i. 390 + mammalia of, i. 390 + birds of, i. 391 + reptiles of, i. 396 + amphibia of, i. 397 + fresh-water fish of, i. 397 + summary of vertebrata of, i. 397 + supposed land-connection of with S. America, i. 398 + insects of, i. 403 + lepidoptera of, i. 404 + coleoptera of, i. 405 + land-shells of, i. 407 + sub-regions of, i. 408 + early history of, i. 465 + + Australian sub-region, mammalia of, i. 438 + illustration of mammalia of, i. 439 + birds of, i. 440 + illustration of fauna of, i. 441 + + Australian hedgehog, ii. 254 + + Austro-Malayan sub-region, physical features of, i. 388 + zoology of, i. 409 + + _Automolus_, ii. 103 + + AVICULIDÆ, ii. 533 + + _Avocettula_, ii. 107 + + _Avocettinus_, ii. 108 + + Aye-aye, figure of, i. 278 + ii. 177 + + _Axiocerces_, ii. 477 + + Azores, visited by European birds, i. 17 + birds of, i. 207 + butterflies of, i. 207 + beetles of, i. 207, 209 + peculiarly modified birds of, i. 207 + stragglers to, i. 208 + how stocked with animal life, i. 208 + + B. + + Babirusa of Celebes, peculiarities of, i. 428 + + _Babirusa_, ii. 215 + + Badger, figure of, i. 195 + + _Badis_, ii. 433 + + _Bæotis_, ii. 475 + + _Bagarius_, ii. 443 + + _Bagrichthys_, ii. 442 + + _Bagroides_, ii. 442 + + _Bagrus_, ii. 442 + + Baird, Professor, on fauna of Cape St. Lucas ii. 130 + on representative birds of United States, ii. 180 + + _Balæna_, European Pliocene, i. 112 + ii. 207 + + _Balæniceps_, ii. 360 + + BALÆNIDÆ, ii. 207 + + _Balænodon_, European Pliocene, i. 112 + + _Balænoptera_, ii. 207 + + _Balænopteridæ_, ii. 207 + + _Balea_, ii. 516 + + _Balearica_, ii. 357 + + Baly, Mr., on Phytophaga of Japan, i. 230 + + Banca, its peculiar species and solution of a problem in distribution, i. + 356 + + Band-fish, ii. 435 + + Bandicoots, ii. 250 + + _Barangia_, ii. 199 + + _Barbatula_, ii. 306 + + Barbets, ii. 305 + + _Barbichthys_, ii. 452 + + _Barbus_, ii. 451 + + _Barilius_, ii. 452 + + _Barissia_, ii. 392 + + Barriers, as affecting distribution, i. 6 + permanence of, as affecting distribution, i. 7 + to the dispersal of birds, i. 17 + + _Baryphthengus_, ii. 313 + + _Barypus_, ii. 492 + + _Basileuterus_, ii. 279 + + _Basilornis_, ii. 287 + + _Bassaris_, ii. 200 + + _Batara_, ii. 104 + + Bates, Mr., on Carabidæ of Japan, i. 228 + on Longicorns of Japan, i. 230 + + _Bathmodon_, N. American Tertiary, i. 136 + + _Bathrodon_, N. American Tertiary, i. 133 + + _Bathyerges_, ii. 231 + + BATOIDEI, ii. 462 + + BATRACHIDÆ, ii. 431 + + _Batrachocephalus_, ii. 443 + + _Batrachoseps_, ii. 413 + + _Batrachostomus_, ii. 318 + + Bats, powers of flight of, i. 15 + classification of, i. 87 + of New Zealand, i. 450 + + _Baucis_, ii. 108 + + _Baza_, ii. 349 + + _Bdeogale_, ii. 195 + + Bearded Reedling, ii. 262 + + Bears, probable cause of absence of, from tropical Africa, i. 291 + ii. 201 + + Beaver, N. American Tertiary, i. 140 + + Beavers, ii. 234 + + Bee-eaters, ii. 312 + + Beetles, families selected for study, i. 103 + from the Lias, i. 167 + of Azores, i. 207 + of Japan, i. 228 + of S. Temperate America, ii. 44 + + BELEMNITIDÆ, ii. 506 + + _Belemnoziphius_, European Pliocene, i. 112 + + _Belideus_, ii. 252 + + _Belionota_, ii. 497 + + _Belodontichthys_, ii. 441 + + _Belone_, ii. 450 + + _Belonesox_, ii. 450 + + Belt, Mr., his theory of a great Siberian lake during the glacial epoch, + i. 218; ii. 206 + on change of climate caused by diminution of obliquity of ecliptic, i. + 466 + + _Beluga_, ii. 209 + + _Bembecidium_, ii. 489 + + _Berardius_, ii. 208 + + _Berenicornis_, ii. 317 + + Bermudas, zoology of, ii. 134 + + _Bernicla_, ii. 363 + + _Bernieria_, ii. 258 + + BERYCIDÆ, ii. 424 + + _Bessonornis_, ii. 256 + + _Bettongia_, ii. 251 + + _Bhringa_, ii. 269 + + _Bhutanitis_, ii. 479 + + _Bias_, ii. 270 + + _Biatas_, ii. 104 + + _Bibos_, ii. 222 + + _Bison_, ii. 222, 225 + + Binney, Mr., on Air-breathing Molluscs of N. America, ii. 528 + + Birds, means of dispersal of, i. 15 + dispersal of by winds, i. 16 + American, found in Europe, i. 16 + reaching the Azores, i. 17 + barriers to dispersal of, i. 17 + limited by forests, i. 17 + classification of, i. 93 + Miocene of Greece, i. 116 + extinct, i. 160 + fossil of Palæarctic region, i. 161 + European of Miocene period, i. 161 + Eocene of Europe, i. 162 + relations of, i. 162 + extinct of North America, i. 163 + recently extinct in New Zealand, i. 164 + Cretaceous of N. America, i. 164 + remains of in Brazilian caves, i. 164 + recently extinct in Madagascar and the Mascarene Islands, i. 164 + cosmopolitan groups of, i. 176 + numerous Palæarctic genera, i. 183 + of the European sub-region, i. 193 + northern range of in Europe, i. 193 + of the zone of pine forests, i. 194 + of Iceland, i. 198 + of the Mediterranean sub-region, i. 203 + of Malta, i. 206 (_note_) + of the Azores, i. 207 + of the Cape Verd Islands, i. 215 + of the Siberian sub-region, i. 219 + Oriental found in Siberia, i. 219 + extreme northern Asiatic, i. 219 + of northern Asiatic forests, i. 220 + of the Manchurian sub-region, i. 223 + Palæarctic genera of, in the Manchurian + sub-region, i. 224 + Oriental genera of, in the Manchurian + sub-region, i. 224 + characteristic of N.W. China and Mongolia, i. 226 + table of Palæarctic families of, i. 235 + of West Africa, i. 243 + list of Palæarctic genera of, i. 243 + of the Ethiopian region, i. 253 + of the East African sub-region, i. 260 + S. African, i. 267 + genera of, peculiar to Madagascar, i. 275 + common to Madagascar and Oriental or Ethiopian regions, i. 276 + species common to Madagascar and Africa or Asia, i. 277 + table of Ethiopian families of, i. 295 + table of Ethiopian genera of, i. 306 + of the Oriental region, i. 316 + of the Indian sub-region, i. 323 + Oriental genera of in Central India, i. 324 + Palæarctic and Ethiopian genera in Central India, i. 325 + of Ceylon, i. 327 + of the Indo-Chinese sub region, i. 330 + of the Indo-Malayan sub-region, i. 337 + illustration of peculiar Malayan, i. 339 + of the Philippine Islands, i. 346 + table of Oriental families of, i. 366 + table of Oriental genera of, i. 375 + of Australian region, i. 391 + specially organized Australian families of, i. 392 + of the Papuan Islands, i. 410 + peculiarities of, i. 413 + brilliant colours of, i. 413 + remarkable forms of, i. 414 + of the Moluccas, i. 418 + peculiarities of, i. 421 + of the Timor group, i. 423 + of Celebes, i. 428 + of Australia, i. 440 + of New Zealand, i. 451 + peculiar to New Zealand, i. 452 + of Norfolk Island, i. 453 + of Lord Howe's Island, i. 453 + of the Chatham Islands, i. 454 + of the Auckland Islands, i. 455 + table of families of Australian, i. 471 + table of genera of Australian, i. 478 + of the Neotropical region, ii. 6 + distinctive characters of Neotropical, ii. 7 + of the Mexican sub region, ii. 52 + of the Antilles, ii. 64 + table of distribution of, ii. 68 + table of families of Neotropical, ii. 86 + table of genera of Neotropical, ii. 86 + of the Nearctic region, ii. 116 + list of typical genera of, ii. 118 + of California, ii. 127 + of Central N. America, ii. 130 + of Eastern United States, ii. 132 + of Canada, ii. 136 + table of Nearctic families of, ii. 141 + table of Nearctic genera of, ii. 148 + and Mammals, parallelism of distribution of, ii. 545 + lines of migration of, ii. 545 + peculiar development of, in islands, ii. 546 + contrast of distribution in tropical and + temperate oceanic islands, ii. 546 + + _Biziura_ , ii. 364 + + _Blacicus_, ii. 102 + + Black ape of Celebes, i. 427 + + Bland, Mr. Thomas, on Antillean land-shells, ii. 19 + ii. 526 + + Blanford, Mr. W. T., on the "Indian" region, i. 60 + on relations of Indian sub-region with Africa, i. 321 + + _Blapsidium_, Oolitic insect, i. 167 + + _Blarina_, ii. 191 + + _Blauneria_, ii. 519 + + BLENNIDÆ, ii. 431 + + _Blenniops_, ii. 431 + + _Blennius_, ii. 431 + + _Blethisa_, ii. 489 + + Blind burrowing snakes, ii. 372 + + Blunt-heads, ii. 380 + + Blyth, Mr., on zoological regions, i. 60 + on the relations of Indian sub-region with Africa, i. 321 + + _Boa_, ii. 381 + + _Boædon_, ii. 380 + + Boas, ii. 381 + + _Bola_, ii. 452 + + _Bolborhynchus_, ii. 328 + + _Boleosoma_, ii. 425 + + BOMBINATORIDÆ, ii. 416 + + _Bombinator_, ii. 417 + + _Bonasa_, ii. 339 + + Bonnet-limpets, ii. 511 + + Bony Pikes, ii. 459 + + _Bootherium_, ii. 225 + + Borneo, probable recent changes in, i. 357 + + _Bos_, Post-Pliocene, i. 112 + Indian Miocene, i. 122 + ii. 222, 225 + + _Botaurus_, ii. 359 + + _Bothriodon_, ii. 215 + + _Botia_, ii. 453 + + Bourbon, zoology of, i. 280 + reptiles of, i. 281 + + _Bourcieria_, ii. 108 + ii. 521 + + _Bovidæ_, European Miocene, i. 120 + + BOVIDÆ, ii. 221 + + BOVINÆ, ii. 222 + + _Brachinus_, ii. 489 + + BRACHIOPODA, ii. 532 + + _Brachiurophis_, ii. 383 + + _Brachiurus_, ii. 175 + + _Brachyalestes_, ii. 445 + + _Brachycephalus_, ii. 414 + + _Brachygalba_, ii. 311 + + _Brachylophus_, ii. 401 + + _Brachymeles_, ii. 397 + + _Brachymerus_, ii. 416 + + _Brachymys_, European Miocene, i. 120 + ii. 232 + ii. 236 + + _Brachymystax_, ii. 447 + + _Brachypteryx_, ii. 256 + + _Brachypternus_, ii. 303 + + _Brachytarsomys_, ii. 230 + + _Brachypteracias_, ii. 312 + + _Brachyrhamphus_, ii. 367 + + _Bradybates_, ii. 413 + + _Bradycellus_, ii. 489 + + _Bradyornis_, ii. 271 + + BRADYPODIDÆ, ii. 244 + + _Bradyptetus_, ii. 258 + + _Bradypus_, ii. 244 + + _Bradytus_, ii. 489 + + _Brama_, ii. 429 + + _Bramatherium_, Miocene of Perim Island, i. 122 + ii. 226 + + _Branchiosteus_, ii. 442 + + _Branta_, ii. 364 + + BRASSOLIDÆ, ii. 472 + + _Brassolis_, ii. 472 + + Brazilian cave-fauna, i. 143 + remarks on, i. 145 + + Brazilian sub-region, description of, ii. 21 + mammalia of, ii. 23 + birds of, ii. 24 + illustration of mammalia of, ii. 23 + illustration of birds of, ii. 28 + islands of, ii. 29 + + _Breviceps_, ii. 416 + + _Breyeria borinensis_, Carboniferous insect, i. 168 + + Britain, peculiar species in, i. 197 + + British Isles, zoology of, i. 197 + + Broad-bill, Malayan, figure of, i. 340 + + Broad-bills, ii. 294 + + _Bronchocela_, ii. 402 + + _Brontes_, ii. 444 + + _Brontotheridæ_, N. American Tertiary, i. 137 + + _Brontotherium_, N. American Tertiary, i. 137 + + _Brotogerys_, ii. 328 + + Brush-turkeys, ii. 341 + + _Brycon_, ii. 445 + + _Bryconops_, ii. 445 + + _Bryttus_, ii. 425 + + _Buarremon_, ii. 99 + + _Bubalus_, ii. 222 + + _Bubo_, European Miocene, i. 162 + ii. 350 + + BUCCINIDÆ, ii. 507 + + _Buccinum_, ii. 507 + + _Bucco_, ii. 310 + + BUCCONIIDÆ, ii. 310 + + _Bucephala_, ii. 364 + + _Bucephalus_, ii. 379 + + _Buceros_, ii. 317 + + BUCEROTIDÆ, ii. 316 + + _Bucorvus_, ii. 317 + + _Budorcas_, ii. 224 + + BUDORCINÆ, ii. 224 + + _Budytes_, ii. 290 + + Buffaloes, ii. 221 + + _Bufo_, ii. 415 + + BUFONIDÆ, ii. 415 + + _Bulbuls_, ii. 267 + + BULLIDÆ, ii. 530 + + _Buliminus_, ii. 514 + + _Bulimulus_, ii. 514 + + _Bulimus_, Eocene, i. 169 + ii. 514, 523 + + _Bunælurus_, N American Tertiary, i. 134 + + _Bungarus_, ii. 383 + + _Bungia_, ii. 452 + + _Bunocephalichthys_, ii. 444 + + _Bunocephalus_, ii. 444 + + _Buphaga_, ii. 288 + + BUPRESTIDÆ, ii. 495 + + _Buprestidium_, Oolitic insect, i. 167 + + _Busarellus_, ii. 348 + + Bush-shrikes, ii. 297 + + Bustards, ii. 356 + + _Butalis_, ii. 270 + + _Butastur_, ii. 348 + + _Buteo_, ii. 348 + + _Buteogallus_, ii. 348 + + BUTEONINÆ, ii. 348 + + _Buteola_, ii. 348 + + _Buthraupis_, ii. 98 + + Butterflies, arrangement of, i. 103 + Palæarctic, i. 187 + of Central Europe, i. 196 + of the Mediterranean sub-region, i. 205 + of Azores, i. 207 + peculiar to Siberian sub-region, i. 220 + of Japan and North China, i. 227 + of the Ethiopian region, i. 255 + number of Ethiopian species, i. 256 + of Indo-Malay sub-region, i. 342 + of the Australian region, i. 404 + of the Austro-Malay sub-region, i. 404 + of the Moluccas, i. 419 + of Celebes, peculiarities of, i. 434 + of New Zealand, i. 457 + ii. 470 + general remarks on the distribution of, ii. 483 + fossil, ii. 486 + of S. Temperate America, ii. 43 + + _Bycanistes_, ii. 317 + + C. + + _Cabalus_, ii. 352 + + _Cabrita_, ii. 391 + + _Cacatua_, ii. 325 + + CACATUIDÆ, ii. 324 + + _Caccabis_, ii. 339 + + _Cachius_, ii. 453 + + _Cacomantis_, ii. 309 + + _Cacophis_, ii. 383 + + _Cacopitta_, ii. 261 + + _Cacopus_, ii. 416 + + _Cacotus_, ii. 417 + + _Cactornis_, ii. 284 + + _Cadurcotherium_, European Eocene, i. 125 + + _Cæcilia_, ii. 411 + + CÆCILIADÆ, ii. 411 + + _Cæcum_, ii. 509 + + _Cælodon_, in Brazilian caves, i. 145 + + _Cælogenys_, in Brazilian caves, i. 144 + ii. 241 + + _Cænopithecus_, European Eocene, i. 124 + ii. 178 + + _Cæntropus_, ii. 445 + + _Cainotherium_, European Miocene, i. 120 + European Eocene, i. 126 + + _Cairina_, ii. 364 + + _Caica_, ii. 328 + + _Calamanthus_, ii. 258 + + _Calamaria_, ii. 374 + + CALAMARIIDÆ, ii. 374 + + _Calamodon_, N. American Eocene, i. 139 + + _Calamodus_, ii. 258 + + CALAMOHERPINÆ, ii. 287 + + _Calamoichthys_, ii. 458 + + _Calamospiza_, ii. 285 + + _Calandrella_, ii. 289 + + _Calao_, ii. 317 + + _Calathus_, ii. 489 + + _Caledonica_, ii. 487 + + _Calendula_, ii. 289 + + _Calicalicus_, ii. 271 + + _Calictis_, ii. 195 + + _Calidris_, ii. 353 + + _Caliecthrus_, ii. 309 + + California, illustration of zoology of, ii. 128 + + Californian sub-region, ii. 127 + mammalia of, ii. 127 + birds of, ii. 127 + reptiles of, ii. 128 + amphibia of, ii. 128 + fresh-water fishes of, ii. 128 + + _Caligo_, ii. 472 + + _Calinaga_, ii. 479 + + _Calisto_, ii. 471 + + _Callæas_, ii. 287 + + _Callia_, ii. 521 + + _Callichroma_, ii. 501 + + _Callichrous_, ii. 442 + + _Callichthys_, ii. 444 + + _Callida_, ii. 490 + + _Callidryas_, ii. 478 + + _Callionymus_, ii. 430 + + _Calliope_, ii. 259 + + _Callipepla_, ii. 339 + + _Calliperidia_, ii. 108 + + _Calliphlox_, ii. 198 + + _Callirhynus_, ii. 375 + + _Callisaurus_, ii. 401 + + _Calliste_, ii. 98 + + _Callisthenus_, ii. 489 + + _Callithea_, ii. 474 + + _Callithrix_, in Brazilian caves, i. 184 + ii. 175 + ii. 178 + + _Callocephalus_, ii. 204 + + _Callochen_, ii. 363 + + _Callomystax_, ii. 443 + + _Callophis_, ii. 383 + + _Callophysus_, ii. 443 + + _Callopistes_, ii. 390 + + _Callorhinus_, ii. 202 + + _Calloselasma_, ii. 385 + + _Callosune_, ii. 478 + + _Callula_, ii. 416 + + _Calobates_, ii. 290 + + _Calocitta_, ii. 273 + + _Calodromas_, ii. 344 + + _Caloenas_, ii. 333 + + _Caloperdix_, ii. 339 + + _Calophena_, ii. 490 + + _Calopsitta_, ii. 325 + + _Caloragia_, ii. 375 + + _Calorhamphus_, ii. 306 + + _Calornis_, ii. 288 + + _Calosoma_, ii. 489 + + _Calostethus_, ii. 419 + + _Calotes_, ii. 402 + + _Calothorax_, ii. 108 + + _Calydna_, ii. 476 + + _Calypte_, ii. 108 + + _Calyptocephalus_, ii. 421 + + _Calyptomena_, ii. 295 + + _Calyptorhynchus_, ii. 325 + + CALYPTRÆIDÆ, ii. 511 + + _Calyptura_, ii. 102 + + _Camarhynchus_, ii. 284 + + _Camaroptera_, ii. 258 + + Camel, fossil in Indian Miocene, i. 122 + birth-place and migrations of, i. 155 + Palæarctic, i. 182 + + _Camelidæ_, essentially extra-tropical, i. 112 + N. American Tertiary, i. 138 + + CAMELIDÆ, ii. 216 + + CAMELOPARDALIDÆ, ii. 221 + + _Camelopardalis_, Miocene of Greece, i. 116 + Indian Miocene, i. 122 + ii. 221 + + _Camelotherium_, S. American Pliocene, i. 147 + ii. 217 + + Camels, ii. 216 + + _Camelus_, ii. 216 + + _Camena_, ii. 477 + + _Campephaga_, ii. 269 + + CAMPEPHAGIDÆ, ii. 268 + + _Campephilus_, ii. 303 + + _Campsiempis_, ii. 101 + + _Camptolaimus_, ii. 364 + + _Campylopterus_, ii. 107 + + _Campylorhynchus_, ii. 264 + + Canadian sub-region, mammalia of, ii. 135 + birds of, ii. 136 + reptiles and fishes of, ii. 137 + insects of, ii. 137 + + Canaries, birds of, i. 208 + beetles of, i. 209 + + _Cancroma_, ii. 359 + + _Canidæ_, European Miocene, i. 118 + European Eocene, i. 125 + N. American Tertiary, i. 134 + remarkable S. African, i. 267 + + CANIDÆ, ii. 197 + + _Canis_, European Pliocene, i. 112 + Post-Pliocene, i. 112 + European Miocene, i. 118 + Indian Miocene, i. 121 + European Eocene, i. 125 + N. American Post-Pliocene, i. 129 + N. American Tertiary, i. 134, 135 + in Brazilian caves, i. 144 + S. American Pliocene, i. 146 + ii. 197 + + _Cantharus_, ii. 427 + + _Cantoria_, ii. 376 + + Cape Ant-eater, ii. 246 + + Cape of Good Hope, peculiar flora and fauna of, i. 266 + + Cape Verd Islands, zoology of, i. 214 + + Cape-hare, S. African, i. 267 + + _Capito_, ii. 306 + + CAPITONINÆ, ii. 306 + + _Capoeta_, ii. 451 + + _Capra_, ii. 224, 225 + + _Capreolus_, ii. 219 + + CAPRIMULGIDÆ, ii. 319 + + _Caprimulgus_, ii. 319 + + CAPRINÆ, ii. 224 + + _Capromys_, ii. 238 + + _Capys_, ii. 477 + + CARABIDÆ, ii. 488 + + _Carabus_, ii. 488 + ii. 489 + + CARANGIDÆ, ii. 429 + + _Carassius_, ii. 451 + + CARCHARIIDÆ, ii. 460 + + _Carcineutes_, ii. 316 + + _Cardellina_, ii. 279 + + CARDIADÆ, ii. 535 + + _Cardinalis_, ii. 285 + + _Cardiodus_, S. American Pliocene, i. 147 + + _Cardiopthalmus_, ii. 492 + + _Cardita_, ii. 535 + + _Carenum_, ii. 490 + + _Cariama_, Brazilian caves, i. 164 + ii. 357 + + CARIAMIDÆ, ii. 357 + + _Caridonax_, ii. 316 + + _Carlia_, ii. 397 + + _Carnivora_ of European Pliocene, i. 112 + Miocene of Greece, i. 115 + European Miocene, i. 118 + Indian Miocene, i. 121 + European Eocene, i. 125 + N. American Post-Pliocene, i. 129 + N. American Tertiary, i. 134 + of Brazilian caves, i. 144 + S. American Pliocene, i. 146 + + Carnivora, classification of, i. 88 + antiquity of, i. 153 + of the Palæarctic region, i. 182 + list of Palæarctic genera of, i. 240 + list of Ethiopian genera of, i. 302 + range of Oriental genera of, i. 373 + list of Australian genera of, i. 476 + + CARNIVORA, ii. 192 + general remarks on the distribution of, ii. 204 + range of, in time, ii. 206 + summary and conclusion, ii. 541 + + Caroline Islands, birds of, i. 444 + + Carpenter, Dr. Philip, on Panama shells, ii. 20 + + _Carpiodes_, ii. 451 + + _Carpococcyx_, ii. 309 + + _Carpodacus_, ii. 285 + + _Carpodectes_, ii. 102, 294 + + _Carpophaga_, ii. 332 + + _Carterodon_ in Brazilian caves, i. 145 + ii. 239 + + Carus, and Gerstaeker on classification of animals, i. 85 + Professor, on classification of the Cetacea, i. 88 + + _Carychium_, ii. 519 + + _Casarca_, ii. 363 + + _Cascelius_, ii. 492 + + _Casiornis_, ii. 102, 293 + + _Casoryx_, N. American Tertiary, i. 138 + ii. 225 + + _Casnonia_, ii. 489 + + _Cassiculus_, ii. 282 + + _Cassicus_, ii. 282 + + _Cassidaria_, ii. 507 + + _Cassidix_, ii. 283 + + _Cassinia_, ii. 270 + + Cassowaries, ii. 368 + + _Castalia_, ii. 534 + + _Castnia_, ii. 481 + + CASTNIIDÆ, ii. 481 + + _Castor_, European Pliocene, i. 113 + European Miocene, i. 120 + ii. 234 + + CASTORIDÆ, ii. 234 + + _Castoroides_, ii. 234 + + _Casuarius_, ii. 369 + + CASUARIIDÆ, ii. 368 + + _Catadromus_, ii. 490 + + _Catagramma_, ii. 474 + + _Catamblyrhynchus_, ii. 285 + + _Catamenia_, ii. 285 + + _Catascopus_, ii. 489 + ii. 491 + + _Cataulus_, ii. 520 + + _Catharistes_, ii. 346 + + _Cathartes_, Brazilian caves, i. 124 + ii. 346 + + _Catharus_, ii. 256 + + _Catherpes_, ii. 264 + + _Catla_, ii. 451 + + _Catoblepas_, ii. 224 + + _Catodon_, ii. 208 + + _Catodontidæ_, ii. 207 + + _Catopra_, ii. 433 + + _Catoprion_, ii. 446 + + _Catostomus_, ii. 451 + + _Catoxantha_, ii. 496 + + _Catriscus_, ii. 258 + + Cats, ii. 192 + + Cave-fauna of Brazil, i. 143 + + _Cavia_, European Miocene, i. 121 + in Brazilian caves, i. 144 + S. American Pliocene, i. 147 + ii. 241 + + Cavies, ii. 241 + + CAVIIDÆ, ii. 241 + + CEBIDÆ, ii. 174 + + _Cebochoerus_, European Eocene, i. 126 + ii. 215 + + _Cebus_ in Brazilian caves, i. 144 + ii. 174 + ii. 178 + + _Cecina_, ii. 521 + + Celebes, physical features of, i. 389 + mammalia of, i. 426 + birds of, i. 428 + insects of, i. 434 + origin of fauna of, i. 436 + + _Celestus_, ii. 327 + + _Celeus_, ii. 303 + + _Celia_, ii. 489 + + _Cenchris_, ii. 385 + + _Centetes_, ii. 188 + + _Centetidæ_, European Miocene, i. 118 + + CENTETIDÆ, ii. 188 + + _Centrarchus_, ii. 425 + + CENTRISCIDÆ, ii. 436 + + _Centriscus_, ii. 436 + + _Centrites_, ii. 101, 291 + + _Centrocercus_, ii. 339 + + _Centrolabrus_, ii. 437 + + _Centrolophus_, ii. 429 + + _Centromochlus_, ii. 443 + + _Centronotus_, ii. 431 + + _Centropus_, ii. 309 + + _Centronyx_, ii. 286 + + _Centropyx_, ii. 390 + + _Centurus_, ii. 303 + + _Cephalepis_, ii. 108 + + _Cephalopeltis_, ii. 389 + + CEPHALOPHINÆ, ii. 224 + + _Cephalophus_, ii. 224 + + CEPHALOPODA, ii. 505 + + _Cephalopterus_, ii. 103, 294 + + _Cephalopyrus_, ii. 266 + + _Cepola_, ii. 435 + + CEPOLIDÆ, ii. 435 + + CERAMBYCIDÆ, ii. 493 + + _Ceratichthys_, ii. 452 + + _Ceratina_, ii. 470 + + Ceratodus, remarkable Australian fish, i. 397 + + _Ceratodus_, ii. 458 + + _Ceratohyla_, ii. 418 + + _Ceratophora_, ii. 402 + + _Ceratophorus_, ii. 501 + + _Ceratophrys_, ii. 420 + + _Ceratoptera_, ii. 463 + + _Ceratorhina_, ii. 367 + + _Ceratorhinus_, ii. 213 + + _Ceratotherium_, ii. 213 + + _Cerberus_, ii. 376 + + _Cercaspis_, ii. 380 + + _Cerchneis_, ii. 349 + + _Cercocebus_, ii. 173 + + _Cercolabes_ in Brazilian caves, i. 145 + ii. 240 + + CERCOLABIDÆ, ii. 239 + + _Cercoleptes_, ii. 200 + + _Cercomacra_, ii. 104 + + _Cercomela_, ii. 260 + + _Cercomys_, ii. 239 + + _Cercopithecus_ in European Pliocene, i. 112 + ii. 173 + + _Cercosaura_, ii. 394 + + CERCOSAURIDÆ, ii. 394 + + _Cereopsis_, ii. 363 + + _Ceriornis_, ii. 340 + + CERITHIADÆ, ii. 509 + + _Certhia_, ii. 264 + + _Certhidea_, ii. 278 + + CERTHIADÆ, ii. 264 + + _Certhilauda_, ii. 289 + + _Certhiola_, ii. 278 + + _Certhiparus_, ii. 266 + + _Cervicapra_, ii. 224 + + CERVICAPRINÆ, ii. 224 + + _Cervidæ_, European Miocene, i. 120 + birth-place and migrations of, i. 155 + + CERVIDÆ, ii. 218 + + _Cervulus_, ii. 219 + + _Cervus_, European Pliocene, i. 113 + Indian Pliocene and Miocene, i. 122 + N. American Post-Pliocene, i. 130 + N. American Tertiary, i. 138 + in Brazilian caves, i. 144 + S. American Pliocene, i. 147 + ii. 219 + + _Ceryle_, ii. 316 + + CESTRACIONTIDÆ, ii. 461 + + _Cetacea_, European Pliocene, i. 112 + European Miocene, i. 119 + N. American Post-Pliocene, i. 130 + N. American Tertiary, i. 140 + + Cetacea, classification of, i. 89 + range of Oriental genus, i. 374 + + CETACEA, ii. 207 + + _Cethosia_, ii. 474 + + CETONIIDÆ, ii. 494 + + _Cetopsis_, ii. 443 + + _Cettia_, ii. 258 + + _Ceuthmochares_, ii. 309 + + _Ceycopsis_, ii. 316 + + Ceylon and Malaya, resemblance of insects of, i. 327 + + Ceylonese sub-region, i. 326 + mammalia of, i. 327 + birds of, i. 327 + reptiles of, i. 327 + amphibia of, i. 327 + insects of, i. 327 + past history of, as indicated by its fauna. i. 328 + + _Ceyx_, ii. 316 + + _Chaca_, ii. 441 + + _Chæmarrhornis_, ii. 259 + + _Chæmepelia_, ii. 333 + + _Chærocampa_, ii. 482 + + _Chætobranchus_, ii. 439 + + _Chætocercus_, ii. 108 + ii. 249 + + _Chætodon_, ii. 427 + + _Chætomys_, ii. 240 + + _Chætops_, ii. 256 + + _Chætoptila_, ii. 276 + + _Chætorhynchus_, ii. 269 + + _Chætostomus_, ii. 444 + + _Chætura_, ii. 320 + + _Chætusia_, ii. 356 + + _Chalceus_, ii. 445 + + CHALCIDÆ, ii. 393 + + _Chalcinopsis_, ii. 445 + + _Chalcinus_, ii. 445 + + _Chalcis_, ii. 393 + + _Chalcochloris_, ii. 189 + + _Chalcopelia_, ii. 333 + + _Chalcophaps_, ii. 333 + + _Chalcostetha_, ii. 276 + + _Chalicomys_, European Pliocene, i. 113 + + _Chalicotherium_, European Miocene, i. 119 + Indian Miocene, i. 122 + fossil in N. China, i. 123 + + _Chamæleo_, N. American Eocene, i. 165 + + _Chalybura_, ii. 107 + + _Chamæa_, ii. 264 + + CHAMÆIDÆ, ii. 264 + + CHAMÆLEONIDÆ, ii. 402 + + Chamæleons, ii. 402 + + _Chamæpetes_, ii. 343 + + _Chamæospiza_, ii. 284 + + _Chamæsaura_, ii. 394 + + CHAMÆSAURIDÆ, ii. 394 + + _Chamæza_, ii. 104 + + CHAMIDÆ, ii. 534 + + Chamois, figure of, i. 195 + ii. 224 + + _Chamostrea_, ii. 536 + + _Chanodichthys_, ii. 453 + + CHARACINIDÆ, ii. 444 + + _Characodon_, ii. 450 + + CHARADRIIDÆ, ii. 355 + + _Charadrius_, ii. 356 + + _Charina_, ii. 373 + + _Charis_, ii. 476 + + _Charitornis_, ii. 274 + + _Charmosyna_, ii. 327 + + _Chasiempis_, ii. 271 + + _Chasmodes_, ii. 431 + + _Chasmorhynchus_, ii. 103, 294 + + _Chatarrhæa_, ii. 261 + + Chatham Islands, birds of, i. 454 + + Chatterers, ii. 293 + + _Chaulelasmus_, ii. 364 + + _Chauna_, ii. 361 + + _Chaunonotus_, ii. 272 + + _Chaunoproctus_, ii. 284 + + _Chela_, ii. 453 + + _Chelemys_, ii. 408 + + _Chelidon_, ii. 281 + + _Chelidoptera_, ii. 311 + + _Chelidorynx_, ii. 271 + + _Chelodina_, ii. 408 + + _Chelomeles_ ii. 397 + + _Chelone_, ii. 409 + + Chelonia, classification of, i. 100 + + CHELONIA, ii. 407 + remarks on the distribution of, ii. 410 + fossil, ii. 410 + + CHELONIIDÆ, ii. 409 + + CHELYDIDÆ, ii. 408 + + _Chelydobatrachus_, ii. 416 + + _Chelydra_, European Pliocene, i. 165 + ii. 408 + + _Chelys_, ii. 408 + + _Chenalopex_, ii. 363 + + _Chera_, ii. 286 + + _Chersina_, ii. 408 + + _Chersydrus_, ii. 382 + + Chevrotain of Malaya, figure of, i. 336 + + Chevrotains, ii. 218 + + _Chiamela_, ii. 397 + + _Chiasognathus_, ii. 493 + + _Chibia_, ii. 269 + + _Chilabothrus_, ii. 381 + + Chili should not be placed in the Palæarctic or Nearctic regions, i. 63 + + Chili and Temperate S. America, distribution of Carabidæ in, ii. 492 + + Chilian Andes, illustration of zoology of, ii. 40 + + Chilian sub-region, ii. 36 + mammalia of, ii. 36 + birds of, ii. 37 + illustration of zoology of, ii. 40 + reptiles and amphibia of, ii. 40 + fresh-water fishes of, ii. 42 + insects of, ii. 42 + origin and migrations of insects of, ii. 47 + + Chili, islands of, ii. 49 + + _Chilinia_, ii. 518 + + _Chilobranchus_, ii. 456 + + _Chilomeniscus_, ii. 375 + + _Chimæra_, ii. 460 + + CHIMÆRIDÆ, ii. 460 + + China, fossil mammals in, resembling those of Indian and European + Miocene, i. 362 + North, mammalia of, i. 222 + + _Chinchilla_, ii. 237 + + _Chinchillidæ_ in Brazilian caves, i. 145 + S. American Pliocene, i. 147 + Pliocene of Antilles, i. 148 + + CHINCHILLIDÆ, ii. 237 + + _Chioglossa_, ii. 413 + + _Chionabas_, ii. 471 + + CHIONIDIDÆ, ii. 354 + + _Chionis_, ii. 354 + + CHIROCENTRIDÆ, ii. 454 + + CHIROCOLIDÆ, ii. 393 + + _Chirodon_, ii. 445 + + _Chirodryas_, ii. 418 + + _Chirogaleus_, ii. 176 + + _Chiroleptes_, ii. 421 + + _Chiromachæris_, ii. 102, 292 + + _Chiromantis_, ii. 419 + + CHIROMYIDÆ, ii. 177 + + _Chironectes_, ii. 248 + + Chiroptera, classification of, i. 87 + list of Palæarctic genera of, i. 239 + list of Ethiopian genera of, i. 300 + range of Oriental genera of, i. 371 + list of Australian genera of, i. 475 + + _Chiroptera_, European Eocene, i. 125 + in Brazilian caves, i. 144 + + CHIROPTERA, ii. 181 + remarks on the distribution of, ii. 185 + fossil, ii. 185 + summary and conclusion, ii. 441 + + _Chirotes_, ii. 388 + + CHIROTIDÆ, ii. 388 + + _Chiroxiphia_, ii. 102, 292 + + CHITONIDÆ, ii. 512 + + _Chittya_, ii. 519 + + _Chlænius_, ii. 489 + + _Chlamydodera_, ii. 275 + + _Chlamydophorus_, ii. 246 + + _Chlamydotherium_, ii. 246 + in Brazilian caves, i. 145 + + _Chlenasicus_, ii. 262 + + _Chloëphaga_, ii. 363 + + _Chlorochrysa_, ii. 98 + ii. 99 + + _Chloronerpes_, ii. 303 + + _Chlorophanes_, ii. 278 + + _Chlorophonia_, ii. 98 + ii. 99 + + _Chloropipo_, ii. 102 + + _Chlorospiza_, ii. 283 + + _Chlorospingus_, ii. 99 + ii. 100 + + _Chlorostilbon_, ii. 109 + + _Choanomphalus_, ii. 518 + + _Choanopoma_, ii. 521 + + _Choeromorus_, European Miocene, i. 119 + + _Choeropotamus_, European Eocene, i. 126 + ii. 215 + + _Choeropus_, ii. 250 + + _Choerotherium_, Indian Miocene, i. 122 + + _Cholæpus_, ii. 244 + + _Chologastes_, ii. 450 + + _Cholornis_, ii. 262 + + _Chondestes_, ii. 285 + + _Chondropoma_, ii. 521 + + CHONDROPTERYGII, ii. 46 + + CHONDROSTEI, ii. 459 + + _Chondrostoma_, ii. 452 + + _Choneziphius_, European Pliocene, i. 112 + + _Chordeiles_, ii. 320 + + Chough, Alpine, figure of, i. 195 + + Choughs, ii. 274 + + CHROMIDÆ, ii. 438 + + _Chromis_, ii. 438 + + _Chrysichthys_, ii. 442 + + _Chrysobronchus_, ii. 108 + + _Chrysococcyx_, ii. 310 + + CHRYSOCHLORIDÆ, ii. 189 + + _Chrysochloris_, ii. 189 + + _Chrysocolaptes_, ii. 303 + + _Chrysocyon_, ii. 197 + + _Chrysolampis_, ii. 108 + + _Chrysomitris_, ii. 283 + + _Chrysopelea_, ii. 379 + + _Chrysophrys_, ii. 427 + + _Chrysoptilus_, ii. 303 + + _Chrysothrix_, ii. 175 + + _Chrysotis_, ii. 328 + + _Chrysuronia_, ii. 109 + + _Chthonicola_, ii. 258 + + _Ciccaba_, ii. 350 + + _Cichla_, ii. 439 + + _Cichladusa_, ii. 261 + + _Cichlopsis_, ii. 260 + + _Cicigna_, ii. 392 + + _Cicindela_, ii. 486 + + CICINDELIDÆ, ii. 486 + + _Cicinnurus_, ii. 275 + + _Ciconia_, ii. 360 + + _Ciconiidæ_, ii. 360 + + CINCLIDÆ, ii. 262 + + _Cinclocerthia_, ii. 256 + + _Cinclodes_, ii. 103 + + _Cinclorhamphus_, ii. 260 + + _Cinclosoma,_ ii. 261 + + _Cinclus_, ii. 263 + + _Cinnicerthia_, ii. 264 + + _Cinnyricinclus_, ii. 276 + ii. 288 + + _Cinosternon_, ii. 408 + + _Cinyxis_, ii. 408 + + _Cionella_, ii. 515 + + _Circaëtus_, ii. 348 + + _Circe_, ii. 109 + + Circumpolar zones, objections to system of, i. 67 + + _Circus_, ii. 347 + + _Cirrhina_, ii. 451 + + CIRRHITIDÆ, ii. 427 + + _Cirrhochroa_, ii. 474 + + CIRRHOSTOMI, ii. 464 + + _Cissa_, ii. 273 + + _Cissopis_, ii. 99 + + _Cisticola_, ii. 257 + + _Cistothorus_, ii. 263 + + _Cistula_, ii. 521 + + _Cithara_, ii. 508 + + _Citharinus_, ii. 445 + + _Cittura_, ii. 316 + + _Cladognathus_, ii. 493 + + _Clais_, ii. 108 + + Clam-shells, ii. 535 + + _Clarias_, ii. 441 + + _Clarotes_, ii. 442 + + Classification as affecting the study of distribution, i. 83 + + _Claudius_, ii. 408 + + _Clausilia_, Eocene, i. 169 + ii. 514 + + _Clerome_, ii. 472 + + _Clibanornis_, ii. 103 + + _Climacteris_, ii. 265 + + Climate, as a limit to the range of mammalia, i. 11 + gradual change of, before the glacial epoch, i. 41 + + _Clinteria_, ii. 494 + + CLIONIDÆ, ii. 531 + + _Clostophis_, ii. 520 + + _Clupea_, ii. 454 + + CLUPEIDÆ, ii. 454 + + _Clymenia_, ii. 209 + + _Clypeicterus_, ii. 282 + + _Clytoctantes_, ii. 104 + + _Clytolæna_, ii. 108 + + _Clytus_, ii. 501 + + _Cnidoglanis_, ii. 441 + + _Cnipodectes_, ii. 101 + + _Cnipolegus_, ii. 101, 291 + + _Cobitis_, ii. 453 + + Cobras, ii. 382 + + _Coccothraustes_, ii. 284 + + _Coccygus_, ii. 309 + + _Coccystes_, ii. 310 + + _Cochlognathus_, ii. 452 + + _Cochlostyla_, ii. 514 + + _Cochlothraustes_, ii. 309 + + _Cochoa_, ii. 269 + + Cockatoos, ii. 324 + + Cockles, ii. 535 + + Cocos Islands, bird of, ii. 60 + + _Cocytia_, ii. 481 + + _Coeligena_, ii. 107 + + _Coelodon_, ii. 245 + + _Coelonotus_, ii. 457 + + _Coelopeltis_, ii. 377 + + _Coelosterna_, ii. 501 + + _Coenonympha_, ii. 471 + + _Coereba_, ii. 278 + + COEREBIDÆ, ii. 278 + + _Cogia_, ii. 208 + + _Colaptes_, ii. 304 + + Coleoptera, families selected for study, i. 103 + Palæarctic, i. 188 + number of Palæarctic species, i. 189 + of Central Europe, i. 196 + of the Mediterranean sub-region, i. 205 + of the Cape Verd Islands, i. 215 + of the Ethiopian region, i. 256 + S. African, i. 268 + of Madagascar, i. 282, 283 + of the Oriental region, i. 319 + of Indo-Malay sub-region, i. 342 + of the Australian region, i. 405 + affinity of Australian and South American, i. 406, 407 + of Celebes, i. 435 + of New Zealand, i. 457 + of the Neotropical region, ii. 15 + of S. Temperate America, ii. 44 + of the Mexican sub-region, ii. 56 + of the Antilles, ii. 74 + of the Nearctic region, ii. 123 + + COLEOPTERA, ii. 486 + general observations on the distribution of, ii. 502 (_see also_ + Beetles) + + _Colias_, ii. 478 + + Colies, ii. 307 + + COLIIDÆ, ii. 307 + + _Colius_, ii. 307 + + _Coliuspasser_, ii. 286 + + _Collocalia_, European Miocene, i. 161 + ii. 320 + + _Colluricincla_, ii. 272 + + _Collyris_, ii. 486, 487 + + _Colobus_, European Miocene, i. 117 + ii. 172 + + _Colænis_, ii. 474 + + _Colonoceras_, N. American Tertiary, i. 136 + + _Colopterus_, ii. 101 + + _Colossochelys_ of Indian Miocene, i. 123, 165 + + _Colpodes_, ii. 489 + + _Coluber_, ii. 375 + + COLUBRIDÆ, ii. 375 + + COLUBRINÆ, ii. 375 + + Columbæ, classification of, i. 96 + range of Palæarctic genera of, i. 248 + range of Ethiopian genera of, i. 311 + range of Oriental genera of, i. 384 + range of Australian genera of, i. 485 + + _Columba_, ii. 332 + + COLUMBÆ, ii. 331 + general remarks on the distribution of, ii. 335 + + COLUMBIDÆ, ii. 331 + + _Columbula_, ii. 333 + + _Columna_, ii. 516 + + COLYMBIDÆ, ii. 366 + + _Colymbus_, ii. 366 + + COMEPHORIDÆ, ii. 432 + + _Comephorus_, ii. 549 + + Comoro islands, zoology of, i. 281 + + _Compsosoma_, ii. 98, 375 + + CONCHIFERA, ii. 533 + + _Condylura_, ii. 190 + + Cones, ii. 508 + + _Conger_, ii. 456 + + CONIDÆ, ii. 508 + + _Conirostrum_, ii. 278 + + _Conognatha_, ii. 496 + + _Conophaga_, ii. 100 + + CONOPHAGINÆ, ii. 291 + + _Conophis_, ii. 375 + + _Conopias_, ii. 101 + + _Conorhynchus_, ii. 443 + + _Conostoma_, ii. 262 + + Continents, distribution of, i. 37 + recent changes of, i. 38 + + Continental extension in Mesozoic times, i. 156 + + _Contopus_, ii. 102, 291 + + CONURIDÆ, ii. 327 + + _Conurus_, ii. 328 + + _Conus_, ii. 508 + + _Copea_, ii. 416 + + _Cophoscincus_, ii. 397 + + _Copidoglanis_, ii. 441 + + _Coptodera_, ii. 489, 492 + + _Copurus_, ii. 101 + + _Copsychus_, ii. 259 + + _Coracias_, ii. 311 + + CORACIIDÆ, ii. 311 + + _Coracopsis_, ii. 328 + + _Corades_, ii. 471 + + _Corallus_, ii. 381 + + _Corbis_, ii. 535 + + _Cordylosaurus_, ii. 392 + + _Cordylus_, ii. 392 + + _Coregonus_, ii. 447 + + _Coriphilus_, ii. 327 + + _Coris_, ii. 437 + + _Coronella_, ii. 375 + + CORONELLINÆ, ii. 375 + + _Coronis_, ii. 481 + + _Cornufer_, ii. 419 + + CORVIDÆ, ii. 272 + + _Corvina_, ii. 428 + + _Corvinella_, ii. 272 + + _Corvultur_, ii. 274 + + _Corvus_, European Miocene, i. 161 + ii. 274 + + _Corydalla_, ii. 290 + + _Corydon_, ii. 295 + + _Corynopoma_, ii. 445 + + _Coryphistera_, ii. 103 + + _Coryphodon_, European Eocene, i. 126 + + _Coryphospingus_, ii. 284 + + _Corythaix_, ii. 307 + + _Corythopis_, ii. 100 + + _Corythornis_, ii. 316 + + _Cosmeteira_, ii. 277 + + _Cosmetornis_, ii. 320 + + Cosmopolitan groups enumerated, i. 175 + + _Cossypha_, ii. 256 + + _Cotinga_, ii. 102, 294 + + COTINGIDÆ, ii. 293 + + _Cottus_, ii. 428 + + _Coturniculus_, ii. 284 + + _Coturnix_, ii. 338 + + _Cotyle_, ii. 281 + + _Coua_, ii. 309 + + _Couchia_, ii. 439 + + Coues, Dr., on the blue crow of the Rocky Mountains, ii. 128 + + Coursers, ii. 355 + + Cowries, ii. 508 + + Coypu, ii. 238 + + CRACIDÆ, ii. 342 + + CRACINÆ, ii. 343 + + _Cracticus_, ii. 273 + + Cranes, ii. 357 + + CRANIADÆ, ii. 532 + + _Cranorrhinus_, ii. 317 + + _Craspedocephalus_, ii. 385 + + _Craspedopoma_, ii. 521 + + _Crateropus_, ii. 261 + + _Crax_, ii. 343 + + _Creadion_, ii. 287 + + _Creagrus_, ii. 364 + + _Creagrutus_, ii. 445 + + Creepers, ii. 264 + + _Cremna_, ii. 475 + + _Crenicichla_, ii. 439 + + _Crenilabrus_, ii. 437 + + _Crenuchus_, ii. 445 + + _Creurgops_, ii. 99 + + _Cricetodon_, European Miocene, i. 120 + ii. 230 + + _Cricetomys_, ii. 230 + + _Cricetulus_, ii. 230 + + _Cricetus_, European Pliocene, i. 113 + ii. 230 + + _Cricosoma_, ii. 476 + + _Crinia_, ii. 420 + + _Criniger_, ii. 267 + + _Crithagra_, ii. 285 + + _Crocidura_, ii. 191 + + Crocodiles, Eocene, i. 165 + ii. 406 + lines of migration of, ii. 548 + + Crocodilia, classification of, i. 100 + + CROCODILIA, ii. 405 + general remarks on the distribution of, ii. 406 + fossil, ii. 407 + + CROCODILIDÆ, ii. 406 + + _Crocodilurus_ ii. 390 + + _Crocodilus_, ii. 406 + + Crook-billed plovers of New Zealand, i. 456 + + _Crossarchus_, ii. 195 + + _Crossochilus_, ii. 451 + + _Crossodactylus_, ii. 419 + + _Crossoptilon_, ii. 340 + + _Crossopus_, ii. 191 + + CROTALIDÆ, ii. 384 + + _Crotalophorus_, ii. 385 + + _Crotalus_, ii. 385 + + Crotch, Mr., on beetles of the Azores, i. 209 + + _Crotophaga_, ii. 309 + + Crowned-pigeon, figure of, i. 415 + + Crows, ii. 273 + + _Crypsirhina_, ii. 273 + + _Cryptoblepharus_, ii. 395 + + _Cryptodacus_, ii. 375 + + _Cryptoprocta_, ii. 194 + + CRYPTOPROCTIDÆ, ii. 194 + + _Cryptopterus_, ii. 441 + + _Cryptornis_, European Eocene, i. 163 + + _Cryptotis_, ii. 421 + + _Crypturus_, ii. 344 + + _Ctenodactyla_, ii. 490 + + _Ctenodactylus_, ii. 238 + + _Ctenolabrus_, ii. 437 + + _Ctenomys_, S. American Pliocene, i. 147 + ii. 238 + + _Ctenopharyngodon_, ii. 452 + + _Ctenopoma_, ii. 521 + + _Ctenostoma_, ii. 486 + + Cuba, extinct mammalia of, i. 148 + + _Curculionidium_, Oolitic insect, i. 167 + + _Cubina_, ii. 400 + + CUCULIDÆ, ii. 308 + + _Cuculus_, ii. 309 + + Cuckoo-shrikes, ii. 268 + + Cuckoos, ii. 308 + + _Culter_, ii. 453 + + Cunningham, Professor, lizard discovered by, in Tierra-del-Fuego, ii. 41 + + _Cuniculus_, ii. 230 + + _Cuphopterus_, ii. 272 + + _Cupidonia_, ii. 339 + + Curassows, ii. 342 + + _Curæus_, ii. 282 + + _Curetis_, ii. 477 + + _Curimatus_, ii. 445 + + _Curruca_, ii. 259 + + _Cursoria_, ii. 382 + + _Cursorius_, ii. 355 + + _Cuscus_, ii. 252 + + _Custa_, ii. 390 + + _Cutia_, ii. 266 + + Cuttle fish, ii. 505 + + _Cyanecula_, ii. 259 + + _Cyanocorax_, ii. 273 + + _Cyanomyia_, ii. 109 + + _Cyanopica_, ii. 273 + + _Cyanoptila_, ii. 270 + + _Cyanorhamphus_, ii. 325 + + _Cyanospiza_, ii. 284 + + _Cyanotis_, ii. 101 + + _Cyanurus_, ii. 273 + + _Cybernetes_, ii. 101 + + _Cychloris_, ii. 280 + + _Cychrus_, ii. 489 + + CYCLADIDÆ, ii. 535 + + _Cyclas_, ii. 535 + + _Cyclocorus_, ii. 380 + + _Cycloderma_, ii. 409 + + _Cyclodina_, ii. 397 + + _Cyclodus_, ii. 397 + + _Cyclophis_, ii. 376 + + _Cyclophorus_, ii. 520 + + _Cyclopterus_, ii. 430 + + _Cyclopsitta_, ii. 326 + + _Cyclorhamphus_, ii. 420 + + _Cyclostoma_, Eocene, i. 169 + + CYCLOSTOMATA, ii. 463 + + CYCLOSTOMIDÆ, ii. 520 + + _Cyclostomus_, ii. 521 + + _Cyclothorus_, ii. 247 + + _Cyclotopsis_, ii. 521 + + _Cyclotus_, ii. 521 + + _Cyclusa_, ii. 401 + + _Cygnus_, ii. 363 + + _Cylindrella_, ii. 515 + + _Cylindrophis_, ii. 373 + + _Cyllo sepulta_, European Cretaceous, i. 167 + + _Cymba_, ii. 508 + + _Cymbilanius_, ii. 104 + + _Cymbirhynchus_, ii. 295 + + _Cymindis_, ii. 489 + + _Cynælurus_, in Brazilian caves, i. 144 + ii. 193 + + _Cynanthus_, ii. 108 + + _Cynictis_, ii. 195 + + _Cynocephalus_, ii. 173 + + _Cynodictis_, ii. 198 + + _Cynodon_, ii. 445 + + _Cynogale_, ii. 195 + + CYNOPITHECIDÆ, ii. 172 + + Cynopithecus of Celebes, affinities of, i. 427 + + _Cynopithecus_, ii. 173 + + _Cynomys_, ii. 235 + + _Cyornis_, ii. 270 + + _Cyotherium_, European Eocene, i. 125 + ii. 198 + + _Cyphogastra_, ii. 496 + + _Cyphorhinus_, ii. 264 + + CYPRÆIDÆ, ii. 508 + + _Cyprina_, ii. 535 + + CYPRINIDÆ, ii. 451 + ii. 535 + + _Cyprinus_, ii. 451 + + _Cyprinion_, ii. 452 + + _Cyprinodon_, ii. 450 + + CYPRINODONTIDÆ, ii. 450 + + CYPSELIDÆ, ii. 320 + + _Cypseloides_, ii. 320 + + _Cypselus_, ii. 320 + + _Cypsnagra_, ii. 99 + + _Cyrena_, ii. 535 + + _Cyrestis_, ii. 474 + + _Cyrtonotus_, ii. 489 + + _Cyrtonyx_, ii. 339 + + _Cyrtophis_, ii. 383 + + _Cystignathus_, ii. 420 + + _Cystophora_, ii. 204 + + D. + + _Dacelo_, ii. 316 + + _Dacnis_, ii. 278 + + _Dactylethra_, ii. 422 + + DACTYLETHRIDÆ, ii. 422 + + _Dactylomys_, ii. 239 + + _Dactylopsila_, ii. 249 + + _Dactylozodes_, ii. 496 + + _Dafila_, ii. 363 + + _Dama_, ii. 219 + + _Damias_, ii. 481 + + _Damophila_, ii. 109 + + DANAIDÆ, ii. 470 + + _Danais_, ii. 470 + + _Dangila_, ii. 451 + + _Danio_, ii. 452 + + _Daptophilus_, N. American Tertiary, i. 134 + + Darters, ii. 365 + + Darwin, Mr., his explanation of the cause of the abundance of apterous + insects in Madeira, i. 211 + on the relation of flowers and insects, i. 463 + amphibia collected by, in S. Temperate America, ii. 41 + mice collected by, in S. Temperate America, ii. 37 + on physical geography of the Galapagos, ii. 33 + + _Dasia_, ii. 397 + + _Dasylophus_, ii. 309 + + _Dasyopthalma_, ii. 472 + + _Dasypeltis_, ii. 377 + + DASYPODIDÆ, ii. 245 + + _Dasyprocta_, European Miocene, i. 121 + in Brazilian caves, i. 144 + ii. 241 + + _Dasyptilus_, ii. 329 + + _Dasypus_, in Brazilian caves, i. 145 + S. American Pliocene, i. 147 + ii. 246 + + DASYURIDÆ, ii. 249 + + _Dasyurus_, Australian Post-Tertiary, i. 157 + ii. 249 + + _Daudebardia_, ii. 516 + + David, Père, his researches in China and Thibet, i. 221, 222 + on birds of N. China, i. 226 + + _Debis_, ii. 471 + + Deer, fossil in N. American Tertiary formations, i. 138 + Palæarctic, i. 182 + probable cause of absence from tropical Africa, i. 291 + ii. 218 + + _Deilephila_, ii. 482 + + _Deltatria_, ii. 107 + + _Delma_, ii. 395 + + _Deloneura_, ii. 477 + + _Delphinapterus_, ii. 209 + + DELPHINIDÆ, ii. 208 + + _Delphinus_, European Pliocene, i. 112 + ii. 209 + + DENDRASPIDIDÆ, ii. 383 + + _Dendraspis_, ii. 383 + + _Dendrexetastes_, ii. 104 + + _Dendrochelidon_, ii. 320 + + _Dendrocitta_, ii. 273 + + _Dendrocolaptes_, ii. 103 + + DENDROCOLAPTIDÆ, ii. 295 + + DENDROCOLAPTINÆ, ii. 295 + + _Dendrocincla_, ii. 103 + + _Dendrocygna_, European Miocene, i. 162 + ii. 363 + + _Dendroeca_, ii. 279 + + _Dendrohyrax_, ii. 229 + + _Dendrolagus_, ii. 251 + + _Dendromus_, ii. 303 + + _Dendromys_, ii. 230 + + _Dendrophila_, ii. 265 + + DENDROPHIDÆ, ii. 378 + + _Dendrophis_, ii. 378 + + _Dendropicus_, ii. 303 + + _Dendroplex_, ii. 103 + + _Dendrornis_, ii. 103 + + _Dendrortyx_, ii. 339 + + _Denisonia_, ii. 383 + + DENTALIADÆ, ii. 512 + + _Dentalium_, ii. 512 + ii. 539 + + _Dentex_, ii. 426 + + _Dercas_, ii. 478 + + _Dermatemys_, ii. 408 + + _Dermatocera_, ii. 520 + + _Dermatochelys_, ii. 409 + + _Deroptyus_, ii. 328 + + _Dermognathus_, ii. 413 + + Desert-snakes, ii. 377 + + Desman of S. Russia, figure of, i. 219 + + _Desmodus_, ii. 182 + + _Deudorix_, ii. 477 + + _Diadema_, ii. 474 + + _Diagramma_, ii. 426 + + _Dibamus_, ii. 372 + + DIBRANCHIATA, ii. 505 + + DICÆIDÆ, ii. 277 + + _Dicælus_, ii. 490 + + _Dicæum_, ii. 277 + + _Dicallaneura_, ii. 475 + + _Dicamptodon_, ii. 413 + + _Diceratherium_, N. American Tertiary, i. 137 + + _Dicerca_, ii. 496 + + _Dicerobatis_, ii. 463 + + _Dichobune_, European Eocene, i. 126 + + _Dicotyles_, N. American Post-Pliocene, i. 130 + N. American Tertiary, i. 137 + in Brazilian caves, i. 144 + S. American Pliocene, i. 146 + birthplace and migrations of, i. 155 + ii. 215 + + _Dicotylinæ_, ii. 214 + + _Dicrocerus_, European Miocene, i. 120 + ii. 220 + + _Dicrodon_, ii. 390 + + _Dicroglossus_, ii. 421 + + _Dicrorrhagia_, ii. 474 + + DICRURIDÆ, ii. 269 + + _Dicrurus_, ii. 269 + + DIDELPHYIDÆ, ii. 248 + + _Didelphys_, European Eocene, i. 126 + N. American Post-Pliocene, i. 130 + in Brazilian caves, i. 145 + ii. 248 + + _Dididæ_, i. 164 + + DIDIDÆ, ii. 334 + + _Didocus_, ii. 417 + + DIDUNCULIDÆ, ii. 333 + + _Didunculus_, ii. 334 + + _Didus_, ii. 334 + + _Dieba_, ii. 197 + + _Diemenia_, ii. 383 + + _Diglossa_, ii. 278 + + _Diglossopis_, ii. 278 + + _Dilophus_, ii. 287 + + _Dilophyrus_, ii. 402 + + _Dimodes_, ii. 377 + + _Dimylus_, ii. 190 + + _Dinictis_, ii. 194 + + _Dinoceras_, N. American Eocene, i. 139 + + _Dinocerata_, N. American Tertiary, i. 139 + + _Dinornis_, allied form in European Eocene, i. 163 + of New Zealand and Australia, i. 164 + ii. 369 + + _Dinornithidæ_ of New Zealand, i. 164 + + DINORNITHIDÆ, ii. 269 + + _Dinotherium_, Miocene of Greece, i. 116 + European Miocene, i. 120 + Miocene of Perim Island, i. 123 + + _Dinyctis_, N. American Tertiary, i. 134 + + _Dinylus_, European Miocene, i. 117 + + _Diomedia_, ii. 365 + + _Dioplodon_, ii. 208 + + _Diorhina_, ii. 476 + + _Diphlogæna_, ii. 108 + + _Diphylla_, ii. 182 + + _Diphyllodes_, ii. 274 + + _Diplacodon_, N. American Tertiary, i. 136 + + _Diplodactylus_, ii. 399 + + _Diploglossus_, ii. 397 + + _Diplolæmus_, ii. 401 + + _Diplomesodon_, ii. 191 + + DIPLOMMATINIDÆ, ii. 519 + + _Diplommatina_, ii. 520 + + _Diplomystax_, ii. 443 + + _Diplopelma_, ii. 416 + + _Diplopoma_, ii. 521 + + _Diplopterus_, ii. 309 + + DIPNOI, ii. 458 + + DIPODIDÆ, ii. 231 + + _Dipodomys_, ii. 233 + + Dippers, ii. 263 + + _Diprotodon_, Australian Post-Tertiary, i. 157 + ii. 251 + + DIPSADIDÆ, ii. 379 + + _Dipsadoboa_, ii. 379 + + _Dipsas_, ii. 379 + + _Diptychus_, ii. 452 + + _Dipus_, ii. 232 + + _Dircenna_, ii. 470 + + _Discina_, ii. 539 + + DISCINIDÆ, ii. 532 + + _Discoboli_, ii. 430 + + DISCOGLOSSIDÆ, ii. 421 + + _Discoglossus_, ii. 421 + + _Discognathus_, ii. 451 + + _Discophora_, ii. 472 + + _Discura_, ii. 107 + + Dispersal of animals, i. 10 + of mammalia, i. 10 + of reptiles and amphibia, i. 28 + + _Disteira_, ii. 384 + + _Distichodus_, ii. 445 + + Distribution, affected by climate, i. 5 + affected by physical features, i. 5 + contrasts of, in similar climates, i. 5 + similarities of, in diverse climates, i. 6 + barriers as affecting, i. 6 + study of, dependent on a good classification, i. 83 + of animals an adjunct to geology, i. 8 + of animals requires certain preliminary studies, i. 8 + of animals dependent on physical geography, i. 35 + of animals, as affected by the glacial epoch, i. 40 + of animals, as affected by changes of vegetation, i. 43 + of animals, as affected by organic changes, i. 44 + of animals, hypothetical illustration of, i. 46 + of animals, complexity of the causes affecting the, i. 49 + of animals, problems in, i. 51 + of plants, as affected by the glacial epoch, i. 42 + + _Distrigus_, ii. 490 + + _Diuca_, ii. 284 + + _Diucopis_, ii. 99 + + _Diva_, ii. 98 + + Divers, ii. 366 + + _Docimastes_, ii. 108 + + Dodo of Mauritius, i. 282 + ii. 334 + + _Dodona_, ii. 475 + + _Dolerisca_, ii. 107 + + _Dolichodon_, ii. 208 + + _Dolichonyx_, ii. 282 + + _Dolichopterus_, European Miocene, i. 162 + + _Dolichotis_, ii. 241 + + _Doliophis_, ii. 383 + + _Dolium_, ii. 507 + + _Dommina_, N. American Tertiary, i. 134 + + _Donacobius_, ii. 264 + + _Donacola_, ii. 287 + + _Donacospiza_, ii. 284 + + _Doras_, ii. 443 + + _Dorcatherium_, European Miocene, i. 120 + + _Dorcopsis_, ii. 251 + + _Dorcus_, ii. 493 + + _Doricha_, ii. 108 + + DORIDÆ, ii. 530 + + _Doritis_, ii. 479 + + Dormice, ii. 232 + + _Doryichthys_, ii. 457 + + _Doryphora_, ii. 107 + + Douroucoulis, ii. 175 + + _Draco_, ii. 402 + + _Dremotherium_, Miocene of Greece, i. 116 + European Miocene, i. 120 + ii. 218 + + DREPANIDIDÆ, ii. 277 + + _Drepanis_, ii. 277 + + _Drepanornis_, ii. 275 + ii. 276 + + Dresser, Mr. H. E., on northern range of European birds, i. 193 + + _Drimostoma_, ii. 489 + + _Dromæus_, ii. 368 + + _Dromas_, ii. 356 + + _Dromatherium_, N. American Triassic, i. 134 + oldest American mammal, i. 160 + + _Dromica_, ii. 486, 487 + + _Dromicia_, ii. 252 + + _Dromicus_, ii. 375 + + _Dromius_, ii. 489 + + _Dromococcyx_, ii. 309 + + _Dromolæa_, ii. 260 + + _Dromophis_, ii. 377 + + Drongo-shrike, Malayan, figure of, i. 340 + ii. 269 + + DRYADINÆ, ii. 375 + + DRYIOPHIDÆ, ii. 379 + + _Dryiophis_, ii. 379 + + _Drymocataphus_, ii. 261 + + _Drymodes_, ii. 259 + + _Drymoeca_, ii. 257 + + DRYMOECINÆ, ii. 257 + + _Drymomys_, ii. 230 + + _Drymornis_, ii. 103 + + _Dryocopus_, ii. 303 + + _Dryopithecus_, European Miocene, i. 117 + ii. 178 + + _Dryospiza_, ii. 284 + + _Dryotriorchis_, ii. 348 + + _Dubusia_, ii. 98 + + Ducks, ii. 363 + + _Dulus_, ii. 280 + + _Dumerilia_, ii. 408 + + _Dumetia_, ii. 261 + + _Dumeticola_, ii. 258 + + Duncan, Dr., on fossil corals of the Antilles, ii. 21 + + _D'Urbania_, ii. 477 + + Dwarf-ground snakes, ii. 374 + + _Dynastor_, ii. 472 + + _Dyschirus_, ii. 489 + + _Dysauxis_, ii. 481 + + _Dysithamnus_, ii. 104 + + _Dysopes_, ii. 184 + + E. + + Eagles, ii. 347 + + Eared Seals, ii. 202 + + Ear-shells, ii. 511 + + East Africa, geographical features of, i. 258 + wide range of genera and species over, i. 259 + few special types in, i. 260 + + East African sub-region, description of, i. 258 + genera and species ranging over the whole of, i. 259 + mammalia of, i. 260 + birds of, i. 260 + reptiles of, i. 260 + amphibia and fishes of, i. 260 + insects of, i. 260 + few peculiar types in, i. 260 + illustration of zoology of, i. 261 + + East Australia, peculiar birds of, i. 440 + + East Thibet, mammalia of, i. 222 + + Eaton, Rev., A. E., on insects of Kerguelen Island, i. 211 + + _Eburna_, ii. 507 + + _Echidna_, ii. 254 + + ECHIDNIDÆ, ii. 254 + + _Echimyidæ_, in Brazilian caves, i. 145 + + ECHIMYIDÆ, ii. 238 + + _Echimys_, ii. 239 + + _Echinogale_, European Miocene, i. 118 + + _Echinops_, ii. 188, 189 + + _Echinorhinus_, ii. 461 + + _Echiothrix_, ii. 230 + + _Echis_, ii. 386 + + _Eclectus_, ii. 326 + + _Ectognathus_, N. American Eocene, i. 139 + + _Ectopistes_, ii. 332 + + _Edentata_, Miocene of Greece, i. 116 + European Miocene, i. 121 + N. American Post-Pliocene, i. 130 + N. American Pliocene, i. 140 + of Brazilian caves, i. 145 + S. American Pliocene, i. 147 + + Edentata, classification of, i. 90 + probable birthplace of, i. 155 + range of Ethiopian genera of, i. 305 + range of Oriental genus of, i. 375 + + EDENTATA, ii. 244 + general remarks on the distribution of, ii. 247 + summary and conclusion, ii. 543 + + _Ega_, ii. 490 + + _Egerina_, ii. 397 + + _Elainea_, ii. 101, 291 + + ELAINEINÆ, ii. 291 + + _Elania_, ii. 397 + + _Elanoides_, ii. 349 + + _Elanus_, ii. 349 + + _Elaphodus_, ii. 220 + + _Elaphrus_, ii. 489 + + ELAPIDÆ, ii. 382 + + _Elapochrus_, ii. 375 + + _Elaps_, ii. 383 + + _Elapsoidea_, ii. 383 + + _Elasmognathus_, ii. 212 + + _Electra_, ii. 209 + + ELEPHANTIDÆ, ii. 227 + + Elephants, fossil, of Indian Miocene, i. 123 + fossil in N. American Post-Pliocene formations, i. 130 + birthplace and migrations of, i. 155 + ii. 227 + + Elephant shrews, S. African, i. 267 + ii. 186 + + _Elephas_, Post-Pliocene, i. 112 + fossil in N. China, i. 123 + N. American Tertiary, i. 138 + ii. 227 + + _Eliomys_, ii. 232 + + Elliot, Mr., his great work on the birds of paradise, i. 415 + on classification of the birds of paradise, ii. 274 + + _Ellipesurus_, ii. 463 + + _Ellipsoglossa_, ii. 413 + + _Ellisia_, ii. 258 + + _Ellobius_, ii. 231 + + _Elminia_, ii. 271 + + _Elodina_, ii. 478 + + _Elopichthys_, ii. 453 + + _Elornis_, European Miocene, i. 162 + + _Elosia_, ii. 419 + + _Elotherium_, N. American Tertiary, i. 137, 139 + ii. 215, 216 + + _Elseya_, ii. 408 + + Elwes, Mr., on birds of Persia, i. 204 + on true relations of the birds of Central India, i. 323 + + _Elymnias_, ii. 471 + + ELYMNIIDÆ, ii. 471 + + ELSIADÆ, ii. 530 + + _Embasis_, N. American Tertiary, i. 134 + + _Emberiza_, ii. 285 + + EMBERIZINÆ, ii. 285 + + _Emberizoides_, ii. 284 + + _Emblema_, ii. 287 + + _Embernagra_, ii. 284 + + EMBROTOCIDÆ, ii. 438 + + _Emesis_, ii. 476 + + Emeu, figure of, i. 441 + + Emeus, ii. 368 + + _Emminia_, ii. 390 + + _Empidagra_, ii. 101 + + _Empidias_, ii. 102, 291 + + _Empidochanes_, ii. 102 + + _Empidonax_, ii. 102, 291 + + _Empidonomus_, ii. 102 + + _Emyda_, ii. 409 + + _Emydida_, Indian Miocene, i. 123 + + _Emydocephalus_, ii. 384 + + _Emys_, Indian Miocene, i. 123 + Miocene and Eocene, i. 165 + ii. 408 + + _Enes_, ii. 501 + + _Engystoma_, ii. 416 + + ENGYSTOMIDÆ, ii. 416 + + _Enhydrina_, ii. 384 + + _Enhydrion_, Indian Miocene, i. 121 + ii. 200 + + _Enhydris_, ii. 199 + + _Enicurus_, ii. 263 + + _Enispe_, ii. 472 + + _Enodes_, ii. 288 + + _Enophrys_, ii. 375 + + _Ensophleus_, ii. 420 + + _Entelopes_, ii. 501 + + _Entomiza_, ii. 276 + + _Entomophila_, ii. 275 + + _Enygrus_, ii. 381 + + _Eobasileus_, N. American Eocene, i. 139 + + Eocene period, i. 124 + fauna of S. America, i. 148 + + _Eophona_, ii. 284 + + _Eopsaltria_, ii. 271 + + _Eos_, ii. 327 + + _Epalzeorhynchus_, ii. 451 + + _Ephemera_, from the Lias, i. 167 + + _Ephthianura_, ii. 260 + + _Epicalia_, ii. 474 + + _Epicrates_, ii. 381 + + EPIMACHINÆ, ii. 275 + + _Epimachus_, ii. 275 + + _Epiodon_, ii. 208 + + _Epirhixis_, ii. 419 + + _Epitola_, ii. 477 + + _Eporeodon_, N. American Tertiary, i. 138 + + EQUIDÆ, ii. 211 + + _Equidæ_, European Pliocene, i. 112 + Miocene of Greece, i. 115 + European Eocene, i. 125 + + _Equus_, European Pliocene, i. 112 + Post-Pliocene, i. 112 + Indian Miocene, i. 121 + N. American Post-Pliocene, i. 130 + N. American Tertiary, i. 135 + Brazilian caves, i. 144 + S. American Pliocene, i. 146 + ii. 211 + + _Erebia_, ii. 471 + + _Eremias_, ii. 391 + + _Eremomela_, ii. 258 + + _Eremophilus_, ii. 444 + + _Ereptodon_, N. American Post-Pliocene, i. 130 + + _Eresia_, ii. 474 + + _Erethistes_, ii. 444 + + _Erethizon_, ii. 239 + + _Ereunetes_, ii. 353 + + _Ergaticus_, ii. 279 + + _Ergolis_, ii. 474 + + _Ericulus_, ii. 188 + + _Ericymba_, ii. 452 + + ERINACEIDÆ, ii. 187 + + _Erinaceus_, European Miocene, i. 117 + ii. 187 + + _Eriocnemis_, ii. 109 + + _Eriodes_, ii. 174 + + _Erismatura_, ii. 364 + + _Erithacus_, ii. 259 + + _Eroessa_, ii. 258 + + _Eronia_, ii. 478 + + _Erpornis_, ii. 267 + + ERYCIDÆ, ii. 381 + + ERYCINIDÆ, ii. 476 + + _Erynnis_, ii. 480 + + _Erythrinus_, ii. 445 + + _Erythrocercus_, ii. 270 + + _Erythrocnema_, ii. 347 + + _Erythrogonys_, ii. 356 + + _Erythrolampus_, ii. 375 + + _Erythromachus_ of Rodriguez, i. 164 + ii. 352 + + _Erythrospiza_, ii. 285 + + _Erythrosterna_, ii. 270 + + _Erythrura_, ii. 387 + + _Eryx_, ii. 382 + + ESOCIDÆ, ii. 449 + + _Esox_, ii. 449 + + _Esthemopsis_, ii. 476 + + _Esthonyx_, N. American Eocene, i. 139 + + _Estrilda_, ii. 286 + + _Etheria_, ii. 534 + + Ethiopian region should not include any part of India, i. 63 + defined, i. 73 + subdivisions of, i. 73 + general features of, i. 251 + zoological characteristics of, i. 252 + mammalia of, i. 253 + great speciality of, i. 253 + birds of, i. 253 + reptiles of, i. 254 + amphibia of, i. 255 + fresh-water fish of, i. 255 + summary of vertebrates of, i. 255 + insects of, i. 255 + coleoptera of, i. 256 + terrestrial mollusca of, i. 257 + sub-regions of, i. 258 + Atlantic islands of, i. 269 + the probable past history of, i. 285 + tables of distribution of animals of, i. 293 + + _Euanemus_, ii. 443 + + _Eubagis_, ii. 474 + + _Eucephala_, ii. 109 + + _Euchætes_, ii. 98 + + _Eucometis_, ii. 99 + + _Euchromia_, ii. 481 + + _Eucichla_, ii. 298 + + _Euclyptosternum_, ii. 443 + + _Eudromias_, ii. 356 + + _Eudynamis_, ii. 310 + + _Eudyptes_, ii. 366 + + _Eueides_, ii. 473 + + _Eugenes_, ii. 107 + + _Eugenia_, ii. 107 + + _Euhyrax_, ii. 229 + + _Eulabeornis_, ii. 352 + + _Eulabes_, ii. 287 + + _Eulampis_, ii. 107 + + _Eumæus_, ii. 477 + + _Eumeces_, ii. 397 + + _Eumetopias_, ii. 203 + + _Eumomota_, ii. 313 + + _Eumys_, N. American Tertiary, i. 140 + ii. 231 + + _Eumyias_, ii. 270 + + _Eunectes_, ii. 381 + + _Eunica_, ii. 474 + + _Eunogyra_, ii. 475 + + _Eupetes_, ii. 263 + + _Eupetomena_, ii. 107 + + _Euphema_, ii. 325 + + _Eupherusa_, ii. 109 + + _Euphonia_, ii. 98 + + _Euphractus_, S. American Pliocene, i. 147 + + _Euphryne_, ii. 401 + + _Euphysetes_, ii. 208 + + _Eupleres_, ii. 195 + + _Euploea_, ii. 470 + + EUPLOCAMINÆ, ii. 340 + + _Euplocamus_, ii. 340 + + _Eupodotis_, ii. 356 + + _Euprepes_, ii. 397 + + _Eupsychortyx_, ii. 339 + + _Euptilotis_, ii. 314 + + _Euptychia_, ii. 471 + + _Eurinorhynchus_, ii. 353 + + _Eurocephalus_, ii. 272 + + Europe, recent changes in physical geography of, i. 39 + Miocene fauna of Central, i. 117 + Miocene fauna of, allied to existing fauna of tropical Asia and Africa, + i. 124 + + European sub-region, description of, i. 191 + forests of, i. 192 + mammalia of, i. 192 + birds of, i. 193 + reptiles and amphibia of, i. 195 + fresh-water fish of, i. 196 + insects of, i. 196 + islands of, i. 197 + + _Eurostopodus_, ii. 320 + + _Euryades_, ii. 479 + + _Euryapteryx_, ii. 370 + + _Euryarthrium_, ii. 501 + + _Eurybia_, ii. 475 + + Euryceros of Madagascar, figure of, i. 278 + + _Euryceros_, ii. 288 + + _Eurycus_, ii. 479 + + _Eurygona_, ii. 476 + + EURYGONIDÆ, ii. 476 + + EURYLÆMIDÆ, ii. 294 + + _Eurylæmus_, ii. 295 + + _Euryodon_, in Brazilian caves, i. 145 + ii. 246 + + _Euryphene_, ii. 474 + + _Eurypyga_, ii. 358 + + EURYPYGIDÆ, ii. 358 + + _Eurystomus_, ii. 312 + + _Eurytela_, ii. 474 + + _Eurytherium_, European Eocene, i. 126 + + _Eurytrichus_, ii. 490 + + _Euscarthmus_, ii. 101 + + _Euschemon_, ii. 480 + ii. 481 + + _Eusemia_, ii. 482 + + _Euspiza_, ii. 285 + + _Eustephanus_, ii. 108 + + _Eustira_, ii. 453 + + _Eutatus_, S. American Pliocene, i. 147 + ii. 246 + + _Eutelodon_, European Eocene, i. 126 + + _Eutemnodus_, S. American Eocene, i. 148 + + _Euterpe_, ii. 285 + + _Euthyrhynchus_, ii. 276 + + _Eutoxeres_, ii. 107 + + _Eutriorchis_, ii. 348 + + _Eutropia_, ii. 209 + + _Eutropiichthys_, ii. 441 + + _Eutropius_, ii. 442 + + _Eutroplus_, ii. 438 + + _Exocetus_, ii. 449 + + _Exoglossum_, ii. 452 + + _Exostoma_, ii. 444 + + Extinct mammalian fauna of Europe, general considerations on, i. 126 + mammalia of N. America and Europe, comparison of, i. 140 + mammalia of the Antilles, i. 148 + mammalia of the Old and New Worlds, general remarks on, i. 148 + fauna of New Zealand, i. 459 + Amphibia, ii. 423 + Ant-eaters, ii. 247 + Armadillos, ii. 246 + Bovidæ, ii. 225 + Bradypodidæ, ii. 245 + Camelidæ, ii. 217 + Camelopardalidæ, ii. 221 + Canidæ, ii. 197 + Castoridæ, ii. 234 + Caviidæ, ii. 241 + Centetidæ, ii. 189 + Cercolabidæ, ii. 240 + Cetacea, ii. 209 + Chinchillidæ, ii. 237 + Chiroptera, ii. 185 + Crocodiles, ii. 407 + Deer, ii. 220 + Didelphyidæ, ii. 249 + Dipodidæ, ii. 232 + Echidnidæ, ii. 254 + Echimyidæ, ii. 239 + Elephants, ii. 227 + Equidæ, ii. 211 + Erinaceidæ, ii. 188 + Felidæ, ii. 193 + Hippopotami, ii. 214 + Hyænas, ii. 196 + Hystricidæ, ii. 240 + Insectivora, ii. 192 + Lacertilia, ii. 404 + Lagomyidæ, ii. 242 + Leporidæ, ii. 243 + Macropodidæ, ii. 251 + Muridæ, ii. 230 + Mustelidæ, ii. 199 + Myoxidæ, ii. 232 + Octodontidæ, ii. 238 + Ophidia, ii. 387 + Orycteropodidæ, ii. 246 + Otariidæ, ii. 203 + Procyonidæ, ii. 201 + Quadrumana, ii. 178 + Rallidæ, ii. 252 + Rhinocerotidæ, ii. 213 + Sciuridæ, ii. 236 + Seals, ii. 204 + Sirenia, ii. 210 + Struthionidæ, ii. 369 + Suidæ, ii. 215 + Talpidæ, ii. 190 + Tapirs, ii. 212 + Tortoises, ii. 410 + Tragulidæ, ii. 218 + Tupaiidæ, ii. 186 + Ursidæ, ii. 202 + Viverridæ, ii. 195 + + Extinction of large animals, causes of, i. 158 + + F. + + _Falcinellus_, ii. 360 + + _Falco_, ii. 349 + + FALCONIDÆ, ii. 347 + + FALCONINÆ, ii. 349 + + Falcons, ii. 347 + + _Falculia_, ii. 228 + + Falkland Islands, zoology of, ii. 49 + + Fanged ground-snakes, ii. 380 + + _Farancia_, ii. 377 + + Fauna of Japan, general character and affinities of, i. 230 + of Palæarctic region, general conclusions as to, i. 231 + extinct, of Madagascar and Mascarene Islands, i. 282 + Malayan, probable origin of, i. 359 + Moluccan, peculiarities of, i. 419 + Timorese, origin of, i. 422 + of Celebes, origin of, i. 436 + of New Zealand, origin of, i. 460 + of Galapagos, origin of, ii. 33 + of Central America, origin of, ii. 57 + of Antilles, origin of, ii. 78 + of Neotropical region, origin of, ii. 80 + + FELIDÆ, ii. 192 + + _Felis_, Miocene of Greece, i. 115 + European Miocene, i. 118 + Indian Miocene, i. 121 + N. American Post-Pliocene, i. 129 + in Brazilian caves, i. 144 + ii. 193 + + _Felis spelæa_, i. 110 + + _Feniseca_, ii. 477 + + _Fennecus_, ii. 197 + + _Ferania_, ii. 376 + + Fernando Po, zoological features of, i. 265 + + _Feroculus_, ii. 191 + + _Fiber_, ii. 230 + + _Figulus_, ii. 493 + + Fiji, Tonga, and Samoa Islands, birds of, i. 443 + + Finches, ii. 283 + + FIROLIDÆ, ii. 531 + + Fishes, means of dispersal of, i. 29 + classification of, i. 101 + cosmopolitan groups of, i. 176 + of the Palæarctic region, i. 186 + of the European sub-region, i. 196 + of the Mediterranean sub-region, i. 205 + of the Manchurian sub-region, i. 227 + fresh-water, table of Palæarctic families of, i. 227 + of the Ethiopian region, i. 255 + of South Africa, i. 268 + fresh-water, table of Ethiopian families of, i. 298 + fresh-water, of the Oriental region, i. 318 + of the Indo-Malay sub-region, i. 341 + fresh-water, table of Oriental families of i. 369 + fresh-water, of the Australian region, i. 397 + fresh-water, resemblance of Australian and S. American, i. 400 + how the transmission may have taken place, i. 401 + fresh-water, of New Zealand, i. 457 + fresh-water, table of Neotropical families of, ii. 89 + of Central N. America, ii. 131 + of Eastern United States, ii. 134 + of Canada, ii. 137 + fresh-water, table of Nearctic families of, ii. 143 + remarks on the distribution of, ii. 464 + fossil, ii. 466 + + Fishing-hawks, ii. 349 + + FISSURELLIDÆ, ii. 511 + + FISTULARIDÆ, ii. 436 + + _Fitzroya_, ii. 450 + + Flamingoes, European Miocene, i. 162 + ii. 361 + + Flora, of New Zealand, as influenced by scarcity of insects, i. 462 + fossil of Australia, i. 467 + + Floras, cretaceous and tertiary, of North America, ii. 155 + + _Florisuga_, ii. 107 + + Flower-peckers, ii. 277 + + Flower, Professor, on classification of mammalia, i. 85 + classification of carnivora, i. 87 + + _Fluvicola_, ii. 100 + + Flycatchers, ii. 270 + + Flying Lemur, Malayan, figure of, i. 337 + ii. 186 + + Flying Lizards, ii. 401 + + Flying Opossum, figure of, i. 442 + + _Fordonia_, ii. 376 + + Forests, essential to existence of many European animals, i. 192 + Siberian, greatest extent of, i. 216 + + FORMICARIIDÆ, ii. 297 + + FORMICARIINÆ, ii. 298 + + _Formicarius_, ii. 104 + + _Formicivora_, ii. 104 + + FORMICIVORINÆ, ii. 297 + + Formosa, zoology of, i. 332 + + _Fossa_, ii. 195 + + _Foudia_, ii. 286 + + _Francolinus_, ii. 338 + + _Fraseria_, ii. 272 + + _Fratercula_, ii. 367 + + _Fregetta_, ii. 365 + + _Fregilupus_, ii. 288 + + _Fregilus_, ii. 274 + + Fresh-water fishes, Neotropical, ii. 12 + of S. Temperate America, ii. 42 + of the Mexican sub-region, ii. 54 + of the Antilles, ii. 73 + of the Nearctic region, ii. 120 + of California, ii. 128 + summary and conclusion, ii. 549 + + Fresh-water mussels, ii. 534 + shell, the most Arctic, ii. 518 + snakes, ii. 376 + snails, ii. 518 + + _Fringilla_, ii. 283 + + _Fringillaria_, ii. 285 + + _Fringillauda_, ii. 282 + + FRINGILLIDÆ, ii. 284 + + Frog-mouths, ii. 318 + + Frogs, ii. 420 + + _Fulica_, ii. 352 + + _Fuligula_, ii. 364 + + _Fulmarus_, ii. 365 + + _Fundulus_, ii. 450 + + FURNARIINÆ, ii. 295 + + _Furnarius_, ii. 103 + + _Fusus_, ii. 507 + + G. + + GADIDÆ, ii. 439 + + GADOPSIDÆ, ii. 439 + + _Gadus_, ii. 439 + + _Galago_, ii. 177 + + Galapagos, scarcity of insects in, i. 463 + + Galapagos islands, ii. 29 + mammalia of, ii. 29 + birds of, ii. 30 + reptiles of, ii. 32 + insects of, ii. 33 + land-shells of, ii. 33 + conclusions as to the origin of their fauna, ii. 33 + + _Galatea_, ii. 536 + + _Galaxias_, ii. 448 + + GALAXIDÆ, ii. 448 + + _Galbalcyrhynchus_, ii. 311 + + _Galbula_, ii. 311 + + GALBULIDÆ, ii. 311 + + _Galecynus_, in European Pliocene, i. 112 + ii. 198 + + _Galeichthys_, ii. 443 + + GALEOPITHECIDÆ, ii. 186 + + _Galeoscoptes_, ii. 256 + + _Galeospalax_, European Miocene, i. 118 + ii. 190 + + _Galeotherium_, Post-Pliocene, i. 111 + + _Galera_, N. American Post-Pliocene, i. 130 + + _Galerella_, ii. 195 + + _Galerita_, ii. 289 + ii. 490 + + _Galerix_, ii. 188 + + _Galethylax_, European Eocene, i. 125 + ii. 198 + + _Galeus_, ii. 460 + + _Galictis_, in Brazilian caves, i. 144 + ii. 199 + + _Galidia_, ii. 195 + + _Galidictis_, ii. 195 + + Gallinæ, classification of, i. 96 + range of Palæarctic genera of, i. 248 + range of Ethiopian genera of, i. 311 + range of Oriental genera of, i. 384 + range of Australian genera of, i. 485 + + GALLINÆ, ii. 337 + ii. 340 + general remarks on the distribution of, ii. 344 + + _Gallinago_, ii. 353 + + _Gallinula_, ii. 352 + + _Gallus_, Miocene of Greece, i. 116 + ii. 340 + + _Gallus bravardi_, European Pliocene, i. 161 + + _Galogale_, ii. 195 + + _Gambusia_, ii. 450 + + _Gampsonyx_, ii. 349 + + _Gampsorhynchus_, ii. 261 + + Gannets, ii. 365 + + GANOIDEI, ii. 458 + + Gape-eyed Scinks, ii. 395 + + Gar-fish, ii. 459 + + Garrod, Professor, on the Classification of Parrots, ii. 324 + + _Garrulax_, ii. 261 + + _Garrulus_, ii. 273 + + GASTEROPODA, ii. 507 + + GASTEROSTEIDÆ, ii. 424 + + _Gasterosteus_, ii. 424 + + _Gastornis_, European Eocene, i. 163 + + GASTROCHÆNIDÆ, ii. 537 + + _Gastropelecus_, ii. 445 + + GAVIALIDÆ, ii. 405 + + _Gavialis_, ii. 405 + + Gavials, ii. 405 + + _Gazella_, ii. 223 + + GAZELLINÆ, ii. 223 + + _Gazera_, ii. 481 + + _Gecinulus_, ii. 303 + + _Gecinus_, ii. 303 + + _Gecko_, ii. 399 + + GECKOTIDÆ, ii. 399 + + Geese, ii. 363 + + _Gehyra_, ii. 400 + + Genera common to Post-Pliocene and Pliocene faunas of N. America, i. 132 + + _Genetta_, ii. 195 + + _Genidens_, ii. 443 + + _Geobates_, ii. 103 + + _Geobiastes_, ii. 312 + + _Geocichla_, ii. 256 + + _Geococcyx_, ii. 309 + + _Geocolaptes_, ii. 304 + + GEODEPHAGA, ii. 486 + + _Geoffroyus_, ii. 326 + + Geographical zoology, introduction, ii. 167 + materials for, ii. 168 + + Geological history of Oriental region, i. 362 + + Geology and Physical Geography of the Antilles, ii. 62, 79 + + _Geomelania_, ii. 519 + + _Geomys_, ii. 233 + + _Geopelia_, ii. 332 + + _Geophaps_, ii. 333 + + _Geophagus_, ii. 439 + + _Geopsittacus_, ii. 325 + + _Georissa_, ii. 522 + + _Georychus_, ii. 231 + + _Geositta_, ii. 103 + + _Geospiza_, ii. 284 + + _Geothlypis_, ii. 279 + + _Geotrochus_, ii. 523 + + _Geotryon_, ii. 333 + + _Geotrypus_, ii. 190 + + _Geranospiza_, ii. 347 + + _Gerbillus_, ii. 230 + ii. 232 + + _Geronticus_, ii. 360 + + _Gerrhonotus_, ii. 392 + + _Gerrhosaurus_, ii. 392 + + GERRIDÆ, ii. 438 + + _Gervasia_, ii. 260 + + _Gerygone_, ii. 258 + + Giant-Clams, ii. 534 + + Gibbon, ii. 171 + + Gibraltar, cave fauna of, i. 114 + + Giraffes, ii. 221 + + _Girardinus_, ii. 450 + + Glacial epoch, as affecting the distribution of animals, i. 40 + as a cause of the great change in the fauna of the temperate zones, + since Pliocene times, i. 151 + probably simultaneous in both hemispheres, i. 151 + causing a general subsidence of the ocean, i. 152 + + _Glandina_, Eocene, i. 169 + ii. 515 + + _Glareola_, ii. 355 + + GLAREOLIDÆ, ii. 355 + + _Glaucis_, ii. 107 + + _Glaucidium_, ii. 350 + + _Glauconeza_, ii. 536 + + _Glaucopis_, ii. 481 + + _Gliciphila_, ii. 275 + + _Glis_, ii. 232 + + _Globiocephalus_, ii. 209 + + _Glossoptila_, ii. 278 + + _Glossotherium_, in Brazilian caves, i. 145 + S. American Pliocene, i. 147 + ii. 247 + + _Glycimeris_, ii. 536 + + _Glyphidodon_, ii. 437 + + _Glyphoglossus_, ii. 416 + + _Glyphorhynchus_, ii. 103 + + _Glyptodon_, S. American Pliocene, i. 147 + + _Glyptosternum_, ii. 443 + + _Gnaphodes_, ii. 471 + + _Gnathodon_, ii. 536 + + _Gnathopsis_, S. American Pliocene, i. 147 + + Goats, Palæarctic, i. 182 + ii. 221 + + Goat-suckers, ii. 519 + + GOBIESOCIDÆ, ii. 436 + + GOBIIDÆ, ii. 430 + + _Gobio_, ii. 452 + + _Gobius_, ii. 430 + + Godman, Mr., on Natural History of the Azores, i. 207 + + Golden Moles, S. African, i. 267 + + _Goliathi_, ii. 494 + + _Gonepteryx_, ii. 478 + + _Goniodactylus_, ii. 400 + + _Gongylophis_, ii. 382 + + _Gonorhynchidæ_, ii. 453 + + _Gonyocephalus_, ii. 402 + + _Gonyosoma_, ii. 379 + + _Gouldia_, ii. 107 + + _Goura_, ii. 333 + + _Graculavus_, N. American Cretaceous, i. 164 + + Grallæ, arrangement of, i. 97 + peculiar or characteristic Palæarctic genera, i. 249 + peculiar Ethiopian genera of, i. 31 + peculiar Oriental genera of, i. 386 + peculiar Australian genera of, i. 484 + + GRALLÆ, ii. 351 + general remarks on the distribution of, ii. 362 + + _Grallaria_, ii. 104 + + _Grallaricula_, ii. 104 + + _Grallina_, ii. 273 + + _Grammatophorus_, ii. 402 + + _Grammatoptila_, ii. 261 + + _Grampus_, ii. 209 + + _Granatellus_, ii. 279 + + _Grandala_, ii. 259 + + _Graphidurus_, ii. 232 + + _Graphipterus_, ii. 491 + + _Graucalus_, ii. 268 + + Gray, Dr. J. E., on classification of Cetacea, i. 88 + + _Grayia_, ii. 376 + + Grayson, Col, on birds of Tres Marias, ii. 59 + + Grebes, ii. 367 + + Greece, Upper Miocene deposits of, i. 115 + summary of Miocene fauna of, i. 116 + + Green Bulbuls, ii. 267 + + Greenland, zoology of, ii. 138 + + Greenlets, ii. 280 + + Groups peculiar to a region, how defined, ii. 184 + + Grouse, ii. 328 + + GRUIDÆ, ii. 356 + + _Grus_, ii. 357 + + _Grypsicus_, ii. 421 + + _Grypus_, ii. 107 + + Guacharo, ii. 107 + + Guans, ii. 342 + + Guaraunas, ii. 357 + + _Gubernatrix_, ii. 285 + + _Guillemots_, ii. 267 + + _Guira_, ii. 309 + + _Guiraca_, ii. 285 + + Gulick, Rev. J. T., on Achatinellidæ of the Sandwich Islands, i. 446 + + Gulls, ii. 364 + + _Gulo_, ii. 199 + + Günther, Dr., his classification of reptiles, i. 98 + his classification of fishes, i. 101 + on gigantic tortoises of Galapagos and the Mascarene Islands, i. 289 + on range of Indian reptiles in the Himalayas, i. 329 + on identical Atlantiic and Pacific fishes, ii. 21 + on fresh-water fishes of Central America, ii. 54 + + _Gygis_, ii. 365 + + GYMNARCHIDÆ, ii. 449 + + _Gymnarchus_, ii. 449 + + GYMNETINÆ, ii. 494 + + _Gymnobucco_, ii. 306 + + _Gymnocephalus_, ii. 103 + + _Gymnocichla_, ii. 104 + + _Gymnocorvus_, ii. 274 + + _Gymnocypris_, ii. 452 + + _Gymnodactylus_, ii. 400 + + GYMNODERINÆ, ii. 293 + + _Gymnoderus_, ii. 103 + + GYMNODONTES, ii. 457 + + _Gymnoglaux_, ii. 350 + + _Gymnokitta_, ii. 273 + + _Gymnomystax_, ii. 282 + + _Gymnopelia_, ii. 333 + + _Gymnops_, ii. 287 + + GYMNOPHTHALMIDÆ, ii. 395 + + _Gymnophthalmus_, ii. 395 + + _Gymnopus_, ii. 199 + + _Gymnorhina_, ii. 273 + + _Gymnostomus_, ii. 451 + + GYMNOTIDÆ, ii. 455 + + _Gymnotus_, ii. 455 + + _Gymnura_, ii. 188 + + _Gypaëtus_, ii. 348 + + _Gypohierax_, ii. 348 + + _Gypoictinia_, ii. 349 + + _Gyps_, ii. 346 + + H. + + Haast, Dr., on extinct birds of New Zealand, i. 460 + + Habitat, definition of, i. 4 + + _Habrocomus_, ii. 238 + + _Habroptila_, ii. 352 + + _Habrura_, ii. 101 + + _Hadrostomus_, ii. 102, 293 + + _Hæmatoderus_, ii. 103 + + _Hæmatopus_, ii. 356 + + _Hæmatospiza_, ii. 285 + + _Hæmophila_, ii. 284 + + _Hæmulon_, ii. 426 + + _Hætera_, ii. 471 + + _Hagria_, ii. 397 + + Hainan, zoology of, i. 334 + + _Halcyon_, ii. 316 + + _Halcyornis_, European Eocene, i. 103 + + _Halicyon_, ii. 204 + + _Haliæetus_, ii. 348 + + _Haliastur_, ii. 348 + + _Halichærus_, ii. 204 + + _Halicore_, ii. 210 + + HALIOTIDÆ, ii. 511 + + _Halitherium_, European Pliocene, i. 112 + European Miocene, i. 119 + ii. 211 + + _Halmaturus_, ii. 251 + + HALOSAURIDÆ, ii. 455 + + _Halys_, ii. 385 + + _Hamadryas_, ii. 470 + + Hang-nests, ii. 281 + + _Hapale_, ii. 176 + ii. 178 + + _Hapalemur_, ii. 176 + + HAPALIDÆ, ii. 175 + + _Hapalotis_, ii. 230 + + _Hapalus_, ii. 524 + + _Haplocerus_, ii. 374 + + _Haplochilus_, ii. 450 + + _Haplochiton_, ii. 446 + + HAPLOCHITONIDÆ, ii. 446 + + _Haplodactylus_, ii. 427 + + _Haploodon_, ii. 236 + + HAPLOODONTIDÆ, ii. 236 + + _Haplospiza_, ii. 284 + + _Hapsidrophis_, ii. 379 + + _Harelda_, ii. 364 + + Hares, ii. 242 + + _Harma_, ii. 474 + + _Harpa_, ii. 349 + ii. 507 + + _Harpactes_, ii. 314 + + _Harpagus_, ii. 349 + + _Harpalus_, ii. 489 + + _Harporhynchus_, ii. 256 + + _Harpyhaliæetus_, ii. 348 + + _Hartlaubius_, ii. 288 + + _Hathliodes_, ii. 502 + + Hatteria of New Zealand, i. 456 + + _Hatteria_, ii. 405 + + Hawks, ii. 347 + + Hedgehogs, ii. 187 + + _Hedymeles_, ii. 285 + + _Helarctos_, ii. 202 + + _Helcyra_, ii. 474 + + _Heleothreptus_, ii. 320 + + _Heliactin_, ii. 108 + + _Heliangelus_, ii. 108 + + _Helianthea_, ii. 108 + + _Heliastes_, ii. 437 + + HELICIDÆ, ii. 512 + + _Helicina_, ii. 522 + + HELICONIDÆ, ii. 473 + ii. 522 + + _Heliconius_, ii. 473 + + _Helicophagus_, ii. 442 + + _Helicops_, ii. 377 + + Helictis, Himalayan, figure of, i. 331 + + _Helictis_, ii. 199 + + _Heliobletus_, ii. 103 + + _Heliochæra_, ii. 102 + + _Heliodoxa_, ii. 107 + + _Heliomastes_, ii. 108 + + _Heliopædica_, ii. 107 + + _Heliophobus_, ii. 231 + + _Helioporus_, ii. 417 + + _Heliornis_, ii. 352 + + _Heliothrix_, ii. 108 + + _Heliotrypha_, ii. 108 + + _Helix_, Eocene, i. 169 + ii. 513 + + _Helladotherium_, Miocene of Greece, i. 116 + European Miocene, i. 120 + ii. 221 + + _Helluomorpha_, ii. 490 + + _Helmintherus_, ii. 279 + + _Helminthophaga_, ii. 279 + + _Heloderma_, ii. 390 + + HELODERMIDÆ, ii. 390 + + _Helodromas_, ii. 353 + + _Helogale_, ii. 195 + + _Helogenes_, ii. 442 + + HELORNITHINÆ, ii. 352 + + _Helotarsus_, ii. 348 + + _Hemibos_, Indian Miocene, i. 122 + ii. 225 + + _Hemicentetes_, ii. 188 + + _Hemicercus_, ii. 303 + + _Hemichelidon_, ii. 290 + + _Hemichromis_, ii. 438 + + _Hemicyon_, European Miocene, i. 118 + ii. 198 + + _Hemidacnis_, ii. 278 + + _Hemidactylium_, ii. 413 + + _Hemidactylus_, ii. 399 + + _Hemierges_, ii. 397 + + _Hemigalea_, ii. 195 + + _Hemignathus_, ii. 277 + + _Hemimantis_, ii. 419 + + _Hemiodus_, ii. 445 + + _Hemiphractus_, ii. 420 + + _Hemipimelodus_, ii. 443 + + _Hemiprocne_, ii. 320 + + _Hemipus_, ii. 270 + + _Hemirhamphus_, ii. 450 + + _Hemisilurus_, ii. 442 + + _Hemisorubim_, ii. 442 + + _Hemistilbon_, ii. 109 + + _Hemisus_, ii. 414 + + _Hemitriccus_, ii. 101 + + _Hemixus_, ii. 267 + + _Henicognathus_, ii. 328 + + _Henicopernis_, ii. 349 + + _Henicophaps_, ii. 333 + + _Henicorhina_, ii. 264 + + _Henicornis_, ii. 103 + + _Heptapterus_, ii. 444 + + _Heredia_, ii. 413 + + Herons, ii. 359 + + _Heros_, ii. 438 + + _Herpestes_, ii. 195 + + _Herpetethiops_, ii. 376 + + _Herpetodryas_, ii. 376 + + _Herpeton_, ii. 376 + + _Herpetoreas_, ii. 375 + + _Herpetotheres_, ii. 348 + + _Herpetotherium_, N. American Tertiary, i. 134 + + _Herpsilochmus_, ii. 104 + + Herring, ii. 454 + + _Hesperia_, ii. 480 + + HESPERIDÆ, ii. 480 + + _Hesperomys_, N. American Tertiary, i. 140 + in Brazilian caves, i. 145 + S. American Pliocene, i. 147 + ii. 230, 231 + + _Hesperornis_, N. American Cretaceous, i. 164 + + _Hestia_, ii. 470 + + _Hestima_, ii. 501 + + _Heterobranchus_, ii. 441 + + _Heterocephalus_, ii. 231 + + _Heterocercus_, ii. 102 + + _Heterochroa_, ii. 474 + + _Heterocnemis_, ii. 104 + + _Heterocorys_, ii. 289 + + _Heterodactylus_, ii. 393 + + _Heterodon_, in Brazilian caves, i. 145 + ii. 246 + ii. 376 + + _Heterogynis_, ii. 481 + + _Heterolocha_, ii. 287 + + _Heteromorpha_, ii. 262 + + _Heteromys_, ii. 233 + + _Heteronota_, ii. 400 + + _Heteronympha_, ii. 471 + + _Heteropelma_, ii. 102, 292 + + _Heteropus_, ii. 397 + + HETEROPYGII, ii. 450 + + _Heterospizias_, ii. 348 + + _Heterotis_, ii. 454 + + _Heterura_, ii. 290 + + _Hewitsonia_, ii. 477 + + _Hexagonia_, ii. 491 + + _Hexaprotodon_, Indian Miocene, i. 122 + + Hickman, Mr. John, on a cause of the extinction of large animals, i. 158 + + _Hieracidea_, ii. 349 + + _Hierax_, ii. 349 + + _Hierococcyx_, ii. 310 + + _Hierofalco_, ii. 349 + + Hill-Tits, ii. 266 + + Himalayas, altitude reached by various groups in the, i. 329, 333 + + _Himantornis_, ii. 352 + + _Himantopus_, ii. 353 + + _Hinulia_, ii. 397 + + _Hipistes_, ii. 376 + + _Hipparchia_, ii. 471 + + _Hipparion_, European Pliocene, i. 112 + Miocene of Greece, i. 115 + European Miocene, i. 119 + N. American Post-Pliocene, i. 130 + N. American Tertiary, i. 135 + ii. 211 + + _Hippocampus_, ii. 457 + + _Hippoglossoides_, ii. 441 + + _Hippoglossus_, ii. 441 + + HIPPOPOTAMIDÆ, ii. 214 + + _Hippopotamus_, Post-Pliocene, i. 112 + European Pliocene, i. 113 + Indian Pliocene, i. 122 + ii. 214 + + _Hipposyus_, N. American Tertiary, i. 133 + + _Hippotherium_, European Miocene, i. 119 + Indian Miocene, i. 122 + + HIPPOTRAGINÆ, ii. 223 + + _Hippotragus_, European Miocene, i. 120 + ii. 223 + + HIPPURITIDÆ, ii. 534 + + _Hirundinea_, ii. 101 + + HIRUNDINIDÆ, ii. 280 + + _Hirundo_, ii. 281 + + _Hoazin_, ii. 345 + + _Holocanthus_, ii. 427 + + _Holbrookia_, ii. 401 + + _Holochilus_, ii. 230 + + _Hologerrhum_, ii. 379 + + HOLOSTEI, ii. 458 + + _Holurophis_, ii. 380 + + _Homalodontotherium_, S. American Pliocene, i. 146 + + _Homalophis_, ii. 376 + + _Homalophus_, European Miocene, i. 161 + + HOMALOPSIDÆ, ii. 376 + + _Homalopsis_, ii. 376 + + _Homaloptera_, ii. 453 + + _Homalosoma_, ii. 490 + + _Hombronia_, ii. 397 + + _Homocamelus_, N. American Tertiary, i. 138 + ii. 217 + + _Homorus_, ii. 103 + ii. 524 + + Honey-guides, ii. 304 + + Honeysuckers, birds specially adapted to Australia, i. 392 + ii. 275 + + Hooker, Dr., on deficiency of odours in New Zealand plants, i. 464 + + Hoopoes, ii. 317 + + _Hopladelus_, ii. 442 + + HOPLEGNATHIDÆ, ii. 433 + + _Hoplobatrachus_, ii. 421 + + HOPLOCEPHALA, ii. 460 + + _Hoplocephalus_, ii. 383 + + _Hoplocetus_, European Pliocene, i. 112 + + _Hoplophoneus_, N. American Tertiary, i. 134 + + _Hoplophorus_, ii. 246 + + _Hoplopterus_, ii. 356 + + _Horites_, ii. 258 + + Hornbills, ii. 316 + + Horses, fossil, in Indian Miocene, i. 121 + perfect series of ancestral, in N. America, i. 136 + probable birthplace of, i. 154 + ii. 211 + + Horse-shoe bats, ii. 182 + + _Hortulia_, ii. 381 + + Howling monkeys, ii. 175 + + Hudson, Mr., on land-birds of Patagonia, ii. 39 + + Humming-birds, ii. 321 + + _Huro_, ii. 425 + + Hutton, Capt. F. W., on origin of New Zealand fauna, i. 461 + + Huxley, Professor, on zoological regions, i. 59 + division of animal kingdom by, i. 85 + + _Hyades_, ii. 472 + + _Hyæna_, Post-Pliocene, i. 112 + Miocene of Greece, i. 115 + European Miocene, i. 118 + Indian Miocene, i. 121 + fossil in N. China, i. 123 + ii. 196 + + _Hyænarctos_ in European Pliocene, i. 112 + European Miocene, i. 118 + Indian Miocene, i. 121 + S. American Pliocene, i. 146 + + _Hyænictis_, Miocene of Greece, i. 115 + European Miocene, i. 118 + ii. 196 + + _Hyænidæ_, European Miocene, i. 118 + + HYÆNIDÆ, ii. 196 + + _Hyænodon_, European Miocene, i. 118 + European Eocene, i. 125 + N. American Tertiary, i. 134 + + _Hyænodontidæ_, European Miocene, i. 118 + + HYALEIDÆ, ii. 531 + + _Hyalimax_, ii. 517 + + _Hyalina_, ii. 515 + + _Hyalosaurus_, ii. 392 + + _Hyantis_, ii. 472 + + _Hybocystis_, ii. 520 + + _Hyborhynchus_, ii. 452 + + _Hydrocena_, ii. 521 + + _Hydrochelidon_, ii. 364 + + _Hydrochoerus_, N. American Post-Pliocene, i. 130 + + _Hydrochoerus_, ii. 241 + + _Hydrocissa_, ii. 317 + + _Hydrocyon_, ii. 445 + + _Hydrogale_, ii. 199 + + _Hydromedusa_, ii. 408 + + _Hydromys_, ii. 230 + + _Hydrophasianus_, ii. 355 + + HYDROPHIDÆ, ii. 384 + + _Hydrophis_, ii. 384 + + _Hydropotes_, ii. 219 + + _Hydropsalis_, ii. 319 + + _Hydrornis_, European Miocene, i. 162 + ii. 298 + + _Hydrosaurus_, ii. 389 + + _Hyetornis_, ii. 309 + + _Hygrogonus_, ii. 439 + + _Hyla_, ii. 418 + + _Hylactes_, ii. 297 + + _Hylambates_, ii. 419 + + _Hylaplesia_, ii. 415 + + HYLAPLESIDÆ, ii. 414 + + _Hylarana_, ii. 419 + + _Hylatomus_, ii. 303 + + _Hylella_, ii. 418 + + HYLIDÆ, ii. 418 + + _Hyliota_, ii. 270 + + _Hylobates_, ii. 171 + + _Hylocharis_, ii. 109 + ii. 271 + + _Hylodes_, ii. 419 + + _Hylomanes_, ii. 313 + + _Hylomys_, ii. 186 + + _Hylophilus_, ii. 280 + + _Hylorhina_, ii. 420 + + _Hylotrupes_, ii. 502 + + _Hyloxalus_, ii. 419 + + _Hymenolaimus_, ii. 364 + + HYODONTIDÆ, ii. 453 + + _Hyohippus_, N. American Tertiary, i. 135 + + _Hyomoschus_, European Miocene, i. 120 + ii. 218 + + _Hyopicus_, ii. 303 + + _Hyopotamus_, European Miocene, i. 119 + N. American Tertiary, i. 137 + ii. 216 + + _Hyopsodus_, N. American Tertiary, i. 133 + + _Hyotherium_, European Miocene, i. 119 + ii. 215 + + _Hypargos_, ii. 287 + + _Hyperantha_, ii. 496 + + _Hypergerus_, ii. 261 + + _Hypermnestra_, ii. 479 + + _Hyperodapedon_, ii. 405 + + _Hyperolius_, ii. 417 + + _Hyperoodon_, ii. 208 + + HYPEROODONTIDÆ, ii. 208 + + _Hyperopsius_, ii. 448 + + _Hypertragulus_, N. American Tertiary, i. 138 + + _Hyphantornis_, ii. 286 + + _Hypherpes_, ii. 265 + + _Hypisodus_, N. American Tertiary, i. 138 + + _Hypna_, ii. 474 + + _Hypnale_, ii. 385 + + _Hypochera_, ii. 287 + + _Hypochrysops_, ii. 477 + + _Hypocista_, ii. 471 + + _Hypocnemis_, ii. 104 + + _Hypocolius_, ii. 272 + + _Hypodes_, ii. 272 + + _Hypogeomys_, ii. 230 + + _Hypolais_, ii. 258 + + _Hypolithus_, ii. 491 + + _Hypolycæna_, ii. 477 + + _Hypomesus_, ii. 477 + + _Hypopachus_, ii. 416 + + _Hypophthalmichthys_, ii. 453 + + _Hypophthalmus_, ii. 442 + + _Hypopyrrhus_, ii. 282 + + _Hyporissus_, ii. 190 + + _Hypothymis_, ii. 271 + + _Hypoxanthus_, ii. 304 + + _Hypsipetes_, ii. 267 + + _Hypsiprymnus_, Australian Post-Tertiary, i. 157 + ii. 251 + + _Hypsirhina_, ii. 376 + + _Hypsirhynchus_, ii. 375 + + _Hyrachyus_, N. American Tertiary, i. 136 + + HYRACIDÆ, ii. 228 + + _Hyracodon_, N. American Tertiary, i. 136 + ii. 214 + ii. 248 + + _Hyracoidea_, classification of, i. 90 + Palæarctic, i. 242 + Ethiopian, i. 304 + + HYRACOIDEA, ii. 228 + + _Hyracotherium_, supposed, in European Eocene, i. 125 + European Eocene, i. 126 + ii. 216 + ii. 229 + + _Hyrax_, ii. 228 + + HYSTRICIDÆ, ii. 240 + + _Hystricodon_, ii. 445 + + _Hystrix_, European Pliocene, i. 113 + Miocene of Greece, i. 116 + N. American Tertiary, i. 140 + ii. 240 + + I. + + _Ialmenus_, ii. 477 + + _Ianthoenas_, ii. 332 + + _Ianthina_, ii. 511 + + _Ibidipodia_, European Miocene, i. 162 + + _Ibidorhynchus_, ii. 353 + + Ibidorhynchus, figure of, i. 331 + + _Ibis_, ii. 360 + + Ibises, ii. 360 + + _Ibycter_, ii. 347 + + Iceland, zoology of, i. 198 + + _Ichneumia_, ii. 195 + + _Ichthyoborus_, ii. 445 + + _Ichthyopsis_, ii. 411 + + _Icteria_, ii. 279 + + ICTERIDÆ, ii. 281 + + _Icterus_, ii. 282 + + _Icthyornis_, N. American Cretaceous, i. 164 + + _Icticyon_ in Brazilian caves, i. 144 + ii. 197 + + _Ictinia_, ii. 349 + + _Ictitherium_, Miocene of Greece, i. 115 + European Miocene, i. 118 + ii. 195 + ii. 197 + + _Ictonyx_, ii. 199 + + _Ictops_, N. American Tertiary, i. 133 + + _Ideopsis_, ii. 470 + + _Idmais_, ii. 478 + + _Iguana_, ii. 401 + + Iguanas, ii. 400 + + IGUANIDÆ, ii. 400 + + _Ilerda_, ii. 477 + + _Ilicura_, ii. 102 + + India, Miocene fauna of, allied to that of Europe, i. 123 + geological features of, i. 328 + + Indian sub-region, description of, i. 321 + supposed relation to Ethiopian region, i. 321 + mammalia of, i. 322 + birds of, i. 323 + reptiles and amphibia of, i. 326 + + _Indicator_, ii. 304 + + INDICATORIDÆ, ii. 304 + + Indo-Chinese sub-region, description of, i. 329 + zoological characteristics of, i. 330 + illustration of, i. 331 + reptiles of, i. 331 + amphibia of, i. 331 + insects of, i. 332 + islands belonging to, i. 333 + + Indo-Malayan sub-region, description of, i. 334 + mammalia of, i. 336 + illustrations of, i. 336, 339 + birds of, i. 337 + remote geographical relations of, i. 339 + reptiles and amphibia of, i. 340 + fishes of, i. 341 + insects of, i. 341 + coleoptera of, i. 342 + terrestrial mollusca of, i. 343 + zoological relations of islands of, i. 345 + recent geographical changes in, i. 357 + probable origin of fauna of, i. 359 + + _Inia_, ii. 209 + + _Insectivora_, European Miocene, i. 117 + N. American Post-Pliocene, i. 129 + N. American Tertiary, i. 133 + + Insectivora, classification of, i. 87 + of the Palæarctic region, i. 181 + of N. China and E. Thibet, i. 222 + range of Palæarctic genera of, i. 239 + of Madagascar, i. 273 + range of Ethiopian genera of, i. 301 + of the Oriental region, i. 315 + range of Oriental genera of, i. 372 + range of Australian genera of, i. 476 + + INSECTIVORA, ii. 186 + general remarks on the distribution of, ii. 191 + summary and conclusion, ii. 541 + + Insects, means of dispersal of, i. 32 + tenacity of life of, i. 33 + adapted to special conditions, i. 33 + groups selected for the study of their geographical distribution, i. + 102 + antiquity of the genera of, i. 166 + fossil of European Miocene, i. 166 + European Cretaceous, i. 167 + European Wealden, i. 167 + Palæozoic, i. 168 + Palæarctic, i. 187 + of Central Europe, i. 196 + of the Mediterranean sub-region, i. 205 + of the Siberian sub-region, i. 220 + of the Manchurian sub-region, i. 227 + of the Ethiopian region, i. 255 + of the E. African sub-region, i. 260 + of W. African, i. 265 + S. African, i. 268 + of Madagascar, i. 282 + general remarks on, i. 284 + of tropical Africa and America, probable cause of similarities in, i. + 291 + of Indo-Chinese sub-region, i. 332 + of the Oriental region, i. 318 + of Ceylon, i. 327 + of Indo-Malay sub-region, i. 341 + statistics of collecting in the various islands of the Malay + Archipelago, i. 343 + of the Australian region, i. 403 + of New Guinea, i. 417 + of the Moluccas, i. 420 + of the Timor group, i. 426 + of Celebes, i. 454 + of New Zealand, i. 458 + scarcity of, in New Zealand, i. 462 + influence of, on the flora, i. 463 + of the Neotropical region, ii. 13 + of S. Temperate America, ii. 42 + of S. Temperate America, Palæarctic affinity of, ii. 45 + of the Mexican sub-region, ii. 55 + of the Antilles, ii. 73 + of the Nearctic region, ii. 122 + of Canada, ii. 137 + distribution of, ii. 468 + range of, in time, ii. 469 + summary and conclusion, ii. 550 + lines of migration of, ii. 551 + + _Iodopleura_, ii. 102 + + _Iolæma_, ii. 107 + + _Iolaus_, ii. 477 + + _Iole_, ii. 267 + + _Iora_, ii. 267 + + _Iphias_, ii. 478 + ii. 394 + + IPHISADÆ, ii. 394 + + _Irena_, ii. 269 + + _Iridina_, ii. 534 + + _Iridornis_, ii. 98 + + _Irrisor_, ii. 318 + + IRRISORIDÆ, ii. 318 + + _Isacis_, N. American Tertiary, i. 133 + + _Ischcognathus_, ii. 375 + + _Ischyromys_, N. American Tertiary, i. 140 + ii. 236 + + Islands, N. European, zoology of, i. 197 + of the Mediterranean sub-region, i. 206 + of the W. African sub-region, i. 265 + of the Ethiopian region, i. 269 + Mascarene, i. 280 + of the Indo-Chinese sub-region, i. 333 + of the Indo-Malay sub-region, i. 345 + Fiji, Tonga, and Samoa, i. 443 + Society and Marquesas, i. 444 + New Caledonia and New Hebrides, i. 445 + Sandwich, i. 446 + of New Zealand sub-region, i. 453 + Norfolk, i. 453 + Lord Howe's, i. 454 + Chatham, i. 454 + Auckland, i. 455 + of Tropical S. America, ii. 29 + of the Mexican sub-region, ii. 59 + of Eastern United States, ii. 134 + peculiar colours of pigeons in, ii. 336 + abundance of land-shells in, ii. 525 + + _Isodactylium_, ii. 413 + + _Ispidina_, ii. 316 + + _Issiodromys_, European Pliocene, i. 113 + ii. 232 + + _Ithaginis_, ii. 340 + + _Ithomia_, ii. 470 + + _Ithycyphus_, ii. 379 + + _Ixalus_, ii. 419 + + _Ixonotus_, ii. 267 + + _Ixulus_, ii. 266 + + J. + + _Jacamaralcyon_, ii. 311 + + Jacamars, ii. 311 + + _Jacamerops_, ii. 311 + + Jacanas, ii. 255 + + _Jacchus_, in Brazilian caves, i. 144 + + _Jaculus_, ii. 232 + + _Jaltris_, ii. 375 + + _Jamaicia_, ii. 521 + + _Janella_, ii. 517 + + _Janthocincla_, ii. 261 + + _Japalura_, ii. 402 + + Japan and North China, physical features of, i. 221 + southern extremity of perhaps belongs to the Oriental region, i. 226 + + Japan, general character of the fauna of, i. 230 + former land-connexions of, i. 231 + + Java, mammalia of, i. 349 + productions of, well known, i. 350 + birds of, i. 351 + representative species of birds in, i. 352 + origin of the anomalous features of its fauna, i. 352 + Sumatra and Borneo, their geographical contrasts and zoological + peculiarities explained, i. 357 + + Jays, ii. 273 + + _Jenynsia_, ii. 450 + + Jerboas, ii. 231 + + Juan Fernandez, Carabidæ of, ii. 44 + birds of, ii. 49 + beetles and land-shells of, ii. 51 + + _Juida_, ii. 288 + + _Juliamyia_, ii. 109 + + _Junco_, ii. 284 + + _Junonia_, European Miocene, i. 167 + ii. 474 + + K. + + Kagu, ii. 359 + + Kakapoe, of New Zealand, i. 455 + + _Kalophrynus_, ii. 415 + + Kangaroos, extinct in Australia, i. 157 + ii. 251 + + _Keneuxia_, ii. 397 + + Kerguelen Island, apterous insects of, i. 211 (_note_) + + _Kerodon_, in Brazilian caves, i. 144 + S. American Pliocene, i. 147 + ii. 241 + + _Ketingus_, ii. 443 + + _Ketupa_, ii. 350 + + King-fisher, racquet-tailed, of New Guinea, figure of, i. 415 + + King-fishers, ii. 315 + + _Kittacincla_, ii. 259 + + Kiwi of New Zealand, i. 455 + + _Kneria_, ii. 453 + + _Kobus_, ii. 224 + + Koodoo antelope, figure of, i. 261 + + _Kricogonia_, ii. 478 + + _Krynickia_, ii. 517 + + L. + + _Labeo_, ii. 451 + + _Labrax_, ii. 425 + + LABRIDÆ, ii. 437 + + _Labrus_, ii. 437 + + LABYRINTHICI, ii. 434 + + _Lacerta_, ii. 391 + + LACERTIDÆ, ii. 390 + + Lacertilia, classification of, i. 99 + + LACERTILIA, ii. 388 + general remarks on the distribution of, ii. 403 + fossil, ii. 404 + + _Lacuna_, ii. 510 + + Ladrone Islands, birds of, i. 444 + + _Læmargus_, ii. 461 + + _Læmosthenes_, ii. 489 + + _Læosopis_, ii. 477 + + _Lafresnaya_, ii. 107 + + _Lagenocetus_, ii. 208 + + _Lagenorhynchus_, ii. 209 + + _Lagidium_, ii. 237 + + LAGOMYIDÆ, ii. 242 + + _Lagomys_, European Pliocene, i. 113 + European Miocene, i. 120 + ii. 242 + + _Lagopus_, ii. 339 + + _Lagorchestes_, ii. 251 + + _Lagostomus_, in Brazilian caves, i. 145 + S. American Pliocene, i. 147 + ii. 237 + + _Lagothrix_, ii. 174 + + _Lais_, ii. 442 + + Lake Baikal, seals of, i. 218 + ii. 206 + + _Lalage_, ii. 269 + + _Laletes_, ii. 280 + + LAMIIDÆ, ii. 498 + + _Lamna_, ii. 460 + + LAMNIDÆ, ii. 460 + + _Lampornis_, ii. 107 + + Lampreys, ii. 463 + + _Lamprima_, ii. 493 + + _Lampris_, ii. 429 + + _Lamprocolius_, ii. 288 + + _Lamprolæma_, ii. 107 + + _Lamrophis_, ii. 380 + + _Lampropsar_, ii. 282 + + _Lampropygia_, ii. 108 + + _Lamprospilus_, ii. 477 + + _Lamprospiza_, ii. 99 + + _Lamprotes_, ii. 98 + + Lancelet, ii. 464 + + Land-lizards, ii. 391 + + Land and water, proportions of, i. 35 + + Land and fresh-water shells, antiquity of the genera of, i. 168 + + Land-shells, Palæozoic, i. 169 + Palæarctic, i. 190 + of Madeira, i. 209 + of the Cape Verd Islands, i. 215 + of the Ethiopian region, i. 257 + of W. Africa, i. 265 + of Madagascar and the Mascarene Islands, i. 285 + of the Indo-Malay sub-region, i. 344 + of the Australian region, i. 407 + of the Sandwich Islands, i. 466 + of New Zealand, i. 459 + of the Neotropical region, ii. 19 + of the Antilles, ii. 75 + conditions favouring development of, ii. 75 + of N. America, ii. 124 + general observations on the distribution of, ii. 522 + richness of islands in, ii. 525 + their mode of diffusion, ii. 525, 528 + comparative distribution of Operculate and In-operculate, ii. 526 + estimated numbers of, ii. 526 + + Land-snakes, ii. 382 + + _Langaha_, ii. 379 + + _Laniarius_, ii. 272 + + _Lanicterus_, ii. 268 + + _Laniellus_, ii. 272 + + LANIIDÆ, ii. 272 + + _Lanio_, ii. 99 + + _Lanius_, European Miocene, i. 161 + ii. 272 + + _Laopithecus_, N. American Tertiary, i. 133 + + _Laornis_, N. American Cretaceous, i. 164 + + _Laprissa_, ii. 421 + + LARIDÆ, ii. 364 + + _Larimus_, ii. 428 + + Larks, ii. 289 + + _Larus_, ii. 364 + + _Larvivora_, ii. 259 + + _Lasiomys_, ii. 229 + + _Lasiuromys_, ii. 239 + + _Latax_, ii. 199 + + _Lates_, ii. 425 + + _Lathria_, ii. 102 + + _Latonia_, ii. 421 + + _Latrunculus_, ii. 430 + + _Layardia_, ii. 261 + + Lea, Dr. Isaac, on N. American Unionidæ, ii. 125 + + _Lebia_, ii. 489 + + _Lebiasina_, ii. 445 + + _Legatus_, ii. 101 + + _Leiocephalus_, ii. 401 + + _Leiolæmus_, ii. 401 + + _Leistes_, ii. 282 + + _Leistus_, ii. 489 + + _Leiuperus_, ii. 420 + + _Leiyla_, ii. 419 + + _Lemonias_, ii. 476 + + _Lemur_, ii. 176 + + Lemur, fossil, ii. 178 + + _Lemuravidæ_, N. American Tertiary, i. 133 + + _Lemuravus_, N. American Tertiary, i. 133 + + Lemuria, a hypothetical land, i. 76 + + _Lemuridæ_, European Eocene, i. 124 + + LEMURIDÆ, ii. 176 + + Lemuroidea, range of Ethiopian genera of, i. 300 + range of Oriental genera of, i. 371 + + LEMUROIDEA, ii. 176 + + Lemurs, ii. 176 + + _Leonia_, ii. 521 + + _Lepadogaster_, ii. 436 + + _Lepictis_, N. American Tertiary, i. 133 + + _Lepidocepalichthys_, ii. 453 + + _Lepidocephalus_, ii. 453 + + _Lepidogrammas_, ii. 309 + + _Lepidolarynx_, ii. 108 + + Lepidoptera, cosmopolitan families of, i. 177 + table of Palæarctic families of, i. 238 + S. African, i. 268 + table of Ethiopian families of, i. 299 + of the Oriental region, i. 318 + table of Oriental families of, i. 369 + of the Australian region, i. 404 + table of Australian families of, i. 472 + of the Neotropical region, ii. 13 + of the Antilles, ii. 73 + table of Neotropical families of, ii. 90 + of the Nearctic region, ii. 122 + Nearctic families of, ii. 143 + + LEPIDOPTERA, ii. 470 + + _Lepidosiren_, ii. 458 + + LEPIDOSTEIDÆ, ii. 459 + + _Lepidosteus_, ii. 459 + + LEPIDOSTERNIDÆ, ii. 389 + + _Lepidosternon_, ii. 389 + + _Lepilemur_, ii. 176 + + _Lepistes_, ii. 450 + + LEPORIDÆ, ii. 242 + + _Leporinus_, ii. 445 + + _Lepricornis_, ii. 476 + + _Leprodera_, ii. 501 + + _Leptalis_, ii. 478 + + _Leptarchus_, N. American Tertiary, i. 135 + ii. 202 + + _Leptasthenura_, ii. 103 + + _Leptauchenia_, N. American Tertiary, i. 138 + + _Leptobarbina_, ii. 452 + + _Leptobrachium_, ii. 421 + + LEPTOCARDII, ii. 464 + + _Leptocera_, ii. 502 + + _Leptochoerus_, N. American Tertiary, i. 137 + ii. 215 + + _Leptocircus_, ii. 479 + + _Leptodeira_, ii. 379 + + _Leptodon_, Miocene of Greece, i. 116 + ii. 214 + ii. 349 + + _Leptognathus_, ii. 381 + + _Leptomantis_, ii. 419 + + _Leptomeryx_, N. American Tertiary, i. 138 + ii. 220 + + _Lepton_, ii. 535 + + _Leptoneura_, ii. 471 + + _Leptonyx_, ii. 204 + + _Leptopogon_, ii. 101 + + _Leptoma_, ii. 520 + + _Leptoptila_, ii. 333 + + _Leptoptilus_, European Miocene, i. 162 + ii. 361 + + _Leptorhytaon_, ii. 380 + + _Leptornis_, ii. 276 + + LEPTOSOMIDÆ, ii. 310 + + _Leptosomus_, allied form in European Eocene, + i. 168 + ii. 310 + + Leptosomus of Madagascar, i. 278 + figure of, i. 279 + + _Leptotherium_, in Brazilian caves, i. 144 + ii. 226 + + _Leptotriccus_, ii. 101 + + _Leptura_, ii. 502 + + _Lepus_, in Brazilian caves, i. 145 + S. American Pliocene, i. 147 + + _Lerista_, ii. 395 + + _Lerwa_, ii. 339 + + _Lesbia_, ii. 108 + + _Lestodon_, S. American Pliocene, i. 147 + + _Leucippus_, ii. 109 + + _Leuciscus_, ii. 452 + + _Leucochroa_, ii. 516 + + _Leucocyon_, ii. 197 + + _Leucomelæna_, ii. 332 + + _Leuconerpes_, ii. 304 + + _Leucophantes_, ii. 270 + + _Leucophasia_, ii. 478 + + _Leucopleurus_, ii. 209 + + _Leucosarcia_, ii. 333 + + _Leucosomus_, ii. 452 + + _Leucosticte_, ii. 285 + + Lewis, Mr. George, his collection of Japan insects, i. 228 + + LIALIDÆ, ii. 396 + + _Lialis_, ii. 396 + + _Liasis_, ii. 381 + + _Libellula_, from the Lias, i. 167 + + _Libythea_, ii. 475 + + LIBYTHEIDÆ, ii. 475 + + _Lichanotus_, ii. 381 + + _Lichenops_, ii. 101 + + _Licina_, ii. 521 + + _Licmetis_, ii. 325 + + Lilljeborg, Professor, on classification of the Rodentia, i. 90 + + LIMACIDÆ, ii. 517 + + LIMACINIDÆ, ii. 531 + + _Limax_, ii. 517 + + _Limenitis_, ii. 474 + + _Limnæa_, Eocene, i. 169 + European Secondary, i. 169 + ii. 518 + + LIMNÆIDÆ, ii. 518 + + _Limnatornis_, European Miocene, i. 161 + + _Limnocharis_, ii. 420 + + _Limnocyon_, N. American Tertiary, i. 134 + + _Limnodynastes_, ii. 420 + + _Limnohyus_, N. American Tertiary, i. 136 + + _Limnophis_, ii. 376 + + _Limnornis_, ii. 103 + + _Limnotheridæ_, N. American Tertiary, i. 133 + + _Limnotherium_, N. American Tertiary, i. 133 + + _Limnurgus_, ii. 450 + + _Limosa_, ii. 353 + + Limpets, ii. 511 + + _Lingula_, ii. 538 + + LINGULIDÆ, ii. 532 + + _Linota_, ii. 285 + + _Linsang_, ii. 195 + + _Liocassis_, ii. 442 + + _Liopelma_, ii. 417 + + _Liopis_, ii. 375 + + _Lioptilus_, ii. 267 + + _Lioscelis_, ii. 297 + + LIOTRICHIDÆ, ii. 266 + + _Liothrix_, ii. 266 + + _Lipaugus_, ii. 102 + + _Liparis_, ii. 430 + + _Liphyra_, ii. 477 + + _Lipinia_, ii. 397 + + _Lipoa_, ii. 342 + + _Liposarcus_, ii. 444 + + _Liptala_, ii. 477 + + _Lissolepis_, ii. 397 + + _Listriodon_, European Miocene, i. 119 + + _Lithiodon_, ii. 521 + + _Lithomys_, European Miocene, i. 120 + ii. 236 + + _Lithornis_, European Eocene, i. 163 + + _Litoria_, ii. 418 + + _Littorina_, ii. 510 + + LITTORINIDÆ, ii. 510 + + Lizards, classification of, i. 90 + Tertiary, i. 165 + wide range of a species in Polynesia, i. 448 + distribution and lines of migration of, ii. 547 + + _Lobodon_, ii. 204 + + _Lochmias_, ii. 103 + + _Locustella_, ii. 258 + + _Loddigesia_, ii. 108 + + _Loncheres_, in Brazilian caves, i. 145 + ii. 239 + + _Lonchophorus_, in Brazilian caves, i. 145 + ii. 239 + + LONGICORNIA, ii. 498 + + Longicornia, Palæarctic, i. 188 + Ethiopian, i. 257 + Oriental, i. 320 + Australian, i. 407 + Neotropical, ii. 17 + of Chili, ii. 46 + Nearctic, ii. 123 + + _Lontra_, ii. 199 + + _Lophiodon_, European Eocene, i. 125 + N. American Tertiary, i. 136 + ii. 212 + + _Lophiomeryx_, ii. 218 + + _Lophiotherium_, N. American Tertiary, i. 136 + + _Lophius_, ii. 431 + + _Lophoaëtus_, ii. 348 + + LOPHOBRANCHII, ii. 456 + + _Lophocitta_, ii. 273 + + _Lophogyps_, ii. 346 + + _Lophoictinia_, ii. 349 + + _Lopholaimus_, ii. 362 + + _Lophiomys_, ii. 230 + + _Lophophaps_, ii. 333 + + _Lophophanes_, ii. 266 + + LOPHOPHORINÆ, ii. 340 + + _Lophophorus_, ii. 340 + + _Lophorhina_, ii. 274 + + _Lophornis_, ii. 107 + + _Lophortix_, ii. 339 + + _Lophostrix_, ii. 350 + + LOPHOTIDÆ, ii. 432 + + Lophotragus, ii. 220 + + _Lophotriorchis_, ii. 348 + + _Lophura_, ii. 402 + + Lord Howe's Island, birds of, i. 453 + + _Loricaria_, ii. 444 + + _Loriculus_, ii. 326 + + _Loris_, ii. 176 + + _Lorius_, ii. 327 + + _Lota_, ii. 439 + + _Loxia_, ii. 285 + + _Loxigilla_, ii. 285 + + _Loxomylus_, Pliocene of Antilles, i. 148 + ii. 237 + + _Loxops_, ii. 277 + + _Lucania_, ii. 450 + + LUCANIDÆ, ii. 492 + + _Lucanus_, ii. 493 + + _Lucia_, ii. 477 + + _Lucidella_, ii. 522 + + _Lucifuga_, ii. 440 + + LUCINIDÆ, ii. 535 + + _Lucinopsis_, ii. 536 + + LUCIOCEPHALIDÆ, ii. 434 + + _Lucioperca_, ii. 425 + + _Luciotrutta_, ii. 447 + + _Lucisoma_, ii. 452 + + Lund, Dr., his researches in caves of Brazil, i. 143 + + _Lupus_, ii. 197 + + _Lurocalis_, ii. 320 + + _Luscinia_, ii. 259 + + _Lusciniola_, ii. 258 + + _Lutra_, European Miocene, i. 118 + Indian Miocene, i. 121 + ii. 199 + + _Lutronectes_, ii. 199 + + _Lycæna_, Miocene of Greece, i. 115 + ii. 196 + + LYCÆNIDÆ, ii. 477 + + _Lycalopex_, i. 197 + + LYCODIDÆ, ii. 439 + + _Lycodon_, ii. 380 + + LYCODONTIDÆ, ii. 380 + + _Lycophidion_, ii. 380 + + _Lycorea_, ii. 470 + + _Lygosoma_, ii. 397 + + _Lygosomella_, ii. 397 + + _Lymanopoda_, ii. 471 + + _Lymnas_, ii. 476 + + _Lyncornis_, ii. 320 + + _Lyncus_, ii. 193 + + _Lytorhynchus_, ii. 376 + + Lyre-bird, figure of, i. 441 + ii. 298 + + M. + + _Mabouya_, ii. 397 + + _Macacus_, European Pliocene, i. 112 + Miocene of Greece, i. 115 + Indian Miocene, i. 121 + supposed in European Eocene, i. 125 + ii. 173 + ii. 178 + + Macaws, ii. 327 + + _Machairodus_, i. 110, 111 + Miocene of Greece, i. 115 + European Miocene, i. 118 + Indian Miocene, i. 121 + N. American Tertiary, i. 134 + in Brazilian caves, i. 144 + S. American Pliocene, i. 146 + ii. 193 + + _Machetornis_, ii. 101 + + _Machærhamphus_, ii. 349 + + _Machærirhynchus_, ii. 271 + + _Machæropterus_, ii. 102 + + _Machetes_, ii. 353 + + _Macrauchenia_, S. American Pliocene, i. 146 + + _Macrocalamus_, ii. 374 + + _Macroceramus_, ii. 516 + + _Macrochilus_, ii. 491 + + _Macrocyclis_, ii. 516 + + _Macrodipteryx_, ii. 320 + + _Macrodon_, ii. 445 + + _Macroglossa_, ii. 482 + + _Macrones_, ii. 442 + + _Macronus_, ii. 261 + + _Macronyx_, ii. 290 + + MACROPODIDÆ, ii. 250 + + _Macropus_, ii. 251 + + _Macropygia_, ii. 332 + + _Macrorhamphus_, ii. 353 + + MACROSCELIDIDÆ, ii. 186 + + _Macroscelides_, ii. 186 + + _Macrosila_, ii. 482 + + _Macrotherium_, Miocene of Greece, i. 116 + European Miocene, i. 121 + ii. 246 + + _Macrotus californicus_, ii. 182 + + MACROURIDÆ, ii. 440 + + MACTRIDÆ, ii. 443 + + Madagascar, extinct birds of, i. 164 + description of, i. 272 + mammalia of, i. 272 + birds of, i. 274 + reptiles of, i. 279 + amphibia of, i. 280 + extinct fauna of, i. 282 + general remarks on insect fauna of, i. 284 + + Madeira, birds of, i. 208 + land-shells of, i. 208 + beetles of, i. 210 + wingless insects numerous in, i. 211 + how stocked with animals, i. 213 + + MALACANTHIDÆ, ii. 433 + + Malacca, Sumatra, and Borneo, zoological unity of, i. 353 + comparison of mammalia, i. 354 + of birds, i. 355 + + _Malacocircus_, ii. 261 + + _Malacopteron_, ii. 261 + + _Malacoptila_, ii. 310 + + _Malacorhynchus_, ii. 364 + + _Malacothrix_, ii. 230 + + Malagasy sub-region, description of, i. 272 + mammalia of, i. 272 + birds of, i. 274 + illustration of zoology of, i. 278 + reptiles of, i. 279 + amphibia of, i. 280 + extinct fauna of, i. 282, 289 + insects of, i. 282 + early history of, i. 286 + + _Malapterurus_, ii. 443 + + Malaya and Indo-Malaya, terms defined, i. 345 (_note_) + + Malaya, meaning of term, ii. 261 + + Malay Archipelago, distribution of butterflies in, ii. 484 + distribution of Cicindelidæ in, ii. 487 + distribution of Longicorns in, ii. 500 + + Malayan forms of life reappearing in West Africa, i. 263 + fauna, probable origin of, i. 359 + resemblances to that of Madagascar and Ceylon explained, i. 361 + + _Malimbus_, ii. 286 + + _Mallodon_, ii. 501 + + _Mallotus_, ii. 447 + + Malta, Post-Pliocene fauna of, i. 114 + formerly joined to Africa, i. 201 + fossil elephants of, i. 201 + birds of, i. 206 (_note_) + + _Malurus_, ii. 258 + + Mammal, the most ancient American, i. 134 + + Mammalia, means of dispersal of, i. 10 + as limited by climate, i. 11 + as limited by rivers, i. 12 + how far limited by the sea, i. 13 + dispersed by ice-floes and drift-wood, i. 14 + means of dispersal of aquatic, i. 15 + of most importance in determining zoological regions, i. 57 + classification of, i. 85 + birthplace and migrations of some families of, i. 142, 153 + cosmopolitan groups of, i. 176 + of the Palæarctic region, i. 181 + of the European sub-region, i. 192 + of the Mediterranean sub-region, i. 202 + of the Siberian sub-region, i. 217 + characteristic of Western Tartary, i. 218 + of the Manchurian sub-region, i. 222 + Palæarctic genera of, in the Manchurian sub-region, i. 222 + Oriental genera of, on borders of same sub-region, i. 223 + peculiar to Japan, i. 223 + characteristic of N. W. China and Mongolia, i. 226 + table of Palæarctic families of, i. 234 + range of Palæarctic genera of, i. 239 + of the Ethiopian region, i. 253 + absence of certain important groups, i. 253 + of the E. African sub-region, i. 260 + of W. Africa, i. 262 + of S. Africa, i. 267 + of Madagascar, i. 272 + table of Ethiopian families of, i. 294 + table of Ethiopian genera of, i. 300 + of the Oriental region, i. 315 + range of the genera inhabiting the Indian sub-region, i. 322 + of Ceylon, i. 327 + of the Indo-Chinese sub-region, i. 330 + of the Indo-Malayan sub-region, i. 336 + illustration of characteristic Malayan, i. 336 + of the Philippine Islands, i. 345 + table of Oriental families of, i. 365 + table of Oriental genera of, i. 371 + of Australian region, i. 390 + of the Papuan Islands, i. 410 + of the Moluccas, i. 417 + of the Timor group, i. 422 + of Celebes, i. 427 + of Australia, i. 439 + illustration of, i. 439 + of New Zealand, i. 450 + table of families of Australian, i. 470 + table of genera of Australian, i. 475 + destinctive characters of Neotropical, ii. 6 + of S. Temperate America, ii. 36 + of Straits of Magellan, ii. 37 + of the Mexican sub-region, ii. 52 + of the Antilles, ii. 62 + table of Neotropical families of, ii. 85 + table of Neotropical genera of, ii. 91 + of the Nearctic region, ii. 115 + of California, ii. 127 + of N. American central plains, ii. 129 + of E. United States, ii. 132 + of Canada, ii. 135 + table of Nearctic families of, ii. 140 + table of Nearctic genera of, ii. 145 + + _Mammalia_, extinct, of Old World, i. 107 + extinct, of historic period, i. 110 + extinct, comparative age of in Europe, i. 127 + extinct, of the New World, i. 129 + extinct, of N. America and Europe, compared, i. 141 + original birthplace of some families and genera, i. 142, 153 + of the secondary period, i. 160 + + MAMMALIA, summary and conclusion, ii. 540 + lines of migration of, ii. 544 + + Manakins, ii. 102 + + MANATIDÆ, ii. 210 + + _Manatus_, N. American Post-Pliocene, i. 130 + ii. 210 + + Manchurian sub-region, description of, i. 220 + mammalia of, i. 222 + birds of, i. 223 + reptiles and amphibia of, i. 227 + fresh-water fish of, i. 227 + insects of, i. 227 + coleoptera of, i. 228 + + MANIDIDÆ, ii. 245 + + _Manis_, ii. 245 + + _Manorhina_, ii. 276 + + _Manticora_, ii. 487 + + _Manucodia_, ii. 274 + + _Mareca_, ii. 363 + + _Margaroperdix_, ii. 338 + + _Margarops_, ii. 256 + + _Margarornis_, ii. 103 + + _Marginella_, ii. 508 + + Marine Mollusca, general remarks on the distribution of, ii. 537 + + Marine shells of the Neotropical region, ii. 20 + + Marmosets, ii. 175 + + Marquesas Islands, birds of, i. 443 + + Marsh, Mr., on improvability of Asiatic and African deserts, i. 200 + on camels and goats as destructive to vegetation, i. 200 + + MARSUPIALIA, ii. 248 + general remarks on the distribution of, ii. 253 + + Marsupials, classification of, i. 91 + N. American Post-Pliocene, i. 130 + European Miocene, i. 121 + first migration to America, i. 155 + diversified forms of, i. 391 + of America prove no connection with Australia, i. 399 + list of Australian genera of, i. 476 + + MARSUPIALIA and MONOTREMATA, summary and conclusion, ii. 543 + + _Martes_, N. American Tertiary, i. 135 + ii. 198 + + Mascarene Islands, zoology of, i. 280 + extinct fauna of, i. 282 + gigantic land-tortoises of, i. 289 + + _Masius_, ii. 102 + + MASTACEMBELIDÆ, ii. 437 + + _Mastodon_, European Pliocene, i. 113 + Miocene of Greece, i. 116 + European Miocene, i. 120 + in Brazilian caves, i. 144 + S. American Pliocene, i. 147 + Indian Miocene, i. 123 + N. American Post-Pliocene, i. 130 + N. American Tertiary, i. 138 + ii. 227, 228 + + Mauritius, zoology of, i. 280 + reptiles of, i. 281 + + McCoy, Professor, on Palæontology of Victoria, i. 466 + + _Mechanitis_, ii. 470 + + _Meda_, ii. 452 + + Mediterranean, recent changes in, i. 39 + sub-region, description of, i. 199 + mammalia of, i. 202 + birds of, i. 203 + reptiles and amphibia of, i. 204 + fresh-water fish of, i. 205 + insects of, i. 205 + islands of, i. 206 + sea not separating distinct faunas, i. 201 + + _Megabias_, ii. 270 + + _Megablabes_, ii. 376 + + _Magacephala_, ii. 478 + + _Megacephalon_, ii. 342 + + _Megacerops_, N. American Tertiary, i. 137 + + _Megaderma_, ii. 182 + + _Megærophis_, ii. 383 + + _Megalæma_, ii. 306 + + MEGALÆMIDÆ, ii. 305 + + MEGALÆMINÆ, ii. 306 + + _Megalixalus_, ii. 419 + + _Megalocnus_, fossil in Cuba, i. 148 + + _Megalomastoma_, ii. 521 + + _Megalomeryx_, N. American Tertiary, i. 138 + + _Megalomma_, ii. 487 + + _Megalonyx_, N. American Post-Pliocene, i. 130 + in Brazilian caves, i. 145 + S. American Pliocene, i. 147 + + _Megalophrys_, ii. 421 + + _Megalostoma_, Eocene, i. 169 + + _Megalurus_, ii. 258 + + _Megalophonus_, ii. 289 + + _Megamys_, S. American Eocene, i. 148 + ii. 238 + + _Meganostoma_, ii. 478 + + MEGAPODIIDÆ, ii. 341 + + _Megapodius_, ii. 342 + + _Megaptera_, ii. 207 + + _Megarhynchus_, ii. 101 + + _Megaspira_, European Tertiary, i. 169 + ii. 527 + + _Megatheridæ_, in Brazilian caves, i. 145 + + _Megatherium_, N. American Post-Pliocene, i. 130 + in Brazilian caves, i. 145 + S. American Pliocene, i. 147 + ii. 245 + + _Meiornis_, ii. 369 + + _Melampitta_, ii. 298 + + _Melampus_, ii. 519 + + _Melanerpes_, ii. 303 + + _Melania_, European Secondary, i. 169 + + MELANIADÆ, ii. 509 + + _Melanitis_, ii. 471 + + _Melanochlora_, ii. 266 + + _Melanocorypha_, ii. 289 + + _Melanophidium_, ii. 374 + + _Melanoptila_, ii. 256 + + _Melanotis_, ii. 256 + + MELEAGRINÆ, ii. 340 + + _Meleagris_, N. American Miocene, i. 163 + ii. 340 + + _Meles_, ii. 199 + + _Melidectes_, ii. 276 + + _Melidora_, ii. 316 + + _Melierax_, ii. 348 + + _Melinæa_, ii. 470 + + _Meliornis_, ii. 275 + + _Meliphaga_, ii. 275 + + MELIPHAGIDÆ, ii. 275 + + _Melipotes_, ii. 276 + + _Melirrhophetes_, ii. 276 + + _Melitæa_, ii. 474 + + _Melithreptus_, ii. 276 + + _Melittophagus_, ii. 312 + + _Melizophilus_, ii. 259 + + _Mellisuga_, ii. 108 + + _Mellivora_, Indian Miocene, i. 121 + ii. 199 + + _Melolonthidium_, Oolitic insect, i. 167 + + _Melopelia_, ii. 333 + + _Melopsittacus_, ii. 325 + + _Melopyrrha_, ii. 285 + + _Melospiza_, ii. 284 + + _Melursus_, ii. 202 + + _Menetia_, ii. 395 + + _Meniceros_, ii. 317 + + _Meniscotherium_, N. American Tertiary, i. 138 + + _Menobranchus_, ii. 412 + + _Menopoma_, ii. 412 + + MENOPOMIDÆ, ii. 412 + + _Menotherium_, N. American Tertiary, i. 133 + + _Menura_, ii. 298 + + MENURIDÆ, ii. 298 + + _Mephitis_, in Brazilian caves, i. 144 + ii. 199 + + _Merganetta_, ii. 364 + + _Mergulus_, ii. 367 + + _Mergus_, ii. 364 + + _Meriones_, ii. 232 + ii. 230 + + _Meristes_, ii. 272 + + _Merluccius_, ii. 439 + + _Meroe_, ii. 536 + + MEROPIDÆ, ii. 312 + + _Meropiscus_, ii. 312 + + _Meropogon_, ii. 312 + + _Merops_, ii. 312 + + _Merulaxis_, ii. 297 + + _Merychus_, N. American Tertiary, i. 138 + + _Merychippus_, N. American Tertiary, i. 135 + + _Merychochoerus_, N. American Tertiary, i. 138 + + _Merycodus_, N. American Tertiary, i. 138 + ii. 220 + + _Merycopotamus_, Indian Miocene, i. 122 + ii. 214 + + _Merycotherium_, of Siberian drift, i. 112 + ii. 217 + + _Mesacodon_, N. American Tertiary, i. 133 + + _Mesapia_, ii. 479 + + _Mesites_, ii. 263 + + _Mesohippus_, N. American Tertiary, i. 135 + + _Mesomys_, ii. 239 + + _Mesonauta_, ii. 438 + + _Mesonyx_, N. American Tertiary, i. 134 + + _Mesopithecus_, Miocene of Greece, i. 115 + ii. 178 + + _Mesoprion_, ii. 425 + + _Mesops_, ii. 439 + + _Mesosemia_, ii. 475 + + _Messalina_, ii. 391 + + _Messaras_, ii. 474 + + _Metallura_, ii. 108 + + _Metapheles_, ii. 476 + + _Methonella_, ii. 476 + + _Metius_, ii. 492 + + _Metopia_, ii. 102 + + _Metopiana_, ii. 364 + + _Metoponia_, ii. 283 + + _Metopothrix_, ii. 102 + + _Metriopelia_, ii. 333 + + Mexican sub-region, ii. 51 + mammalia of, ii. 52 + birds of, ii. 52 + reptiles of, ii. 54 + amphibia of, ii. 54 + fresh-water fish of, ii. 54 + insects of, ii. 55 + land-shells of, ii. 57 + its relations to the N. and S. American continents, ii. 57 + islands of, ii. 59 + + Meyer, Dr. A.B., on reptiles and amphibia of New Guinea, i. 415¨ + + _Micræca_, ii. 270 + + _Micracantha_, ii. 501 + + _Micrastur_, ii. 347 + + _Micrathene_, ii. 350 + + _Micrhyla_, ii. 414 + + _Microbates_, ii. 104 + + _Microcebus_, ii. 176 + + _Microcerculus_, ii. 264 + + _Microchæra_, ii. 107 + + _Microglossus_, ii. 325 + + _Microhierax_, ii. 349 + + _Microlestes_, oldest European mammal, i. 160 + + _Micromeryx_, European Miocene, i. 120 + ii. 220 + + _Micropelama_, ii. 353 + + _Micropternus_, ii. 304 + + _Micropterus_, ii. 364 + + _Microscelis_, ii. 267 + + _Microstoma_, ii. 448 + + _Microsyops_, N. American Tertiary, i. 133 + + _Microtherium_, European Miocene, i. 120 + + _Midas_, ii. 176 + + Middendorf, on extreme northern birds, i. 219 + + _Midea_, ii. 478 + + _Miglyptes_, ii. 304 + + Migrating birds, in which region to be placed, i. 185 + + Migration of animals, i. 10 + general phenomena of, i. 18 + of birds, i. 19 + of birds in Europe, i. 19 + probable origin of, i. 22 + of birds in India and China, i. 23 + of birds in N. America, i. 23 + changes in extent of, i. 24 + of birds in S. Temperate America, i. 25 + general remarks on, i. 25 + + _Miletus_, ii. 477 + + _Milvulus_, ii. 102, 291 + + _Milvus_, European Miocene, i. 162 + ii. 349 + + _Mimeta_, ii. 268 + + _Mimetes_, ii. 170 + + _Mimocichla_, ii. 256 + + _Mimus_, ii. 256 + + _Minla_, ii. 266 + + Miocene fauna of the Old World, i. 114 + fauna of Geece, i. 115 + fauna of Greece, summary of, i. 116 + fauna of Central Europe, i. 117 + deposits of Siwalik Hills, i. 121 + faunas of Europe and Asia, general observations on, i. 123 + + _Miohippus_, N. American Tertiary, i. 135 + + _Mionectes_, ii. 101 + + _Mirafra_, ii. 289 + + _Miro_, ii. 260 + + _Misgurnus_, ii. 453 + + _Mitra_, ii. 508 + + _Mitrephorus_, ii. 102 + + _Mitua_, ii. 343 + + Mivart, Professor, on classification of primates, i. 86 + on classification of insectivora, i. 87 + on classification of amphibia, i. 101 + of classification of lemurs, ii. 176 + + _Mixornis_, ii. 261 + + _Mniotilta_, ii. 279 + + MNIOTILTIDÆ, ii. 278 + + _Mochocus_, ii. 443 + + _Mocoa_, ii. 397 + + _Moho_, ii. 276 + + Mole-rat of W. Tartary, i. 218 + + Mole-rats, ii. 231 + + Moles, almost wholly Palæarctic, i. 181 + ii. 190 + + _Mollienesia_, ii. 450 + + Mollusca, means of dispersal of, i. 30 + classification of, i. 104 + groups selected for study, i. 104 + + MOLLUSCA, distribution of, ii. 504 + range of families of, in time, ii. 538 + + Moluccas, zoology of, i. 417 + birds of, i. 419 + reptiles of, i. 420 + insects of, i. 420 + peculiarities of fauna of, i. 421 + + _Molossus_, ii. 184 + + _Molothrus_, ii. 282 + + _Molva_, ii. 439 + + MOMOTIDÆ, ii. 313 + + _Momotus_, ii. 313 + + _Monachalcyon_, ii. 316 + + _Monarcha_, ii. 270 + + _Monasa_, ii. 311 + + _Monitor_, ii. 389 + + Monkeys on the high Himalayas, i. 12 + fossil in N. American Miocene, i. 133 + in E. Thibet, i. 222 + abundance of in the Oriental region, i. 315 + + _Monoceros_, ii. 507 + + _Monodon_, ii. 208 + + MONODONTIDÆ, ii. 208 + + _Monoplocus_, ii. 390 + + _Monopterus_, ii. 455 + + Monotremata, classification of, i. 91 + list of Australian genera of, i. 477 + + MONOTREMATA, ii. 253 + remarks on the distribution of, ii. 254 + + _Monotrophis_, ii. 289 + + _Montacuta_, ii. 535 + + _Monticola_, ii. 256 + + _Montifringilla_, ii. 284 + + Mörch, Dr., on Panama shells, ii. 20 + + _Morelia_, ii. 381 + + "More-pork" of Australia, figure of, i. 442 + + _Morethria_, ii. 395 + + _Mormolyce_, ii. 490 + + MORMYRIDÆ, ii. 448 + + _Mormyrops_, ii. 448 + + _Mormyrus_, ii. 488 + + _Morococcyx_, ii. 309 + + _Morotherium_, N. American Pliocene, i. 140 + + MORPHIDÆ, ii. 472 + + _Morphnus_, ii. 348 + + _Morpho_, ii. 472 + + _Morunga_, ii. 204 + + _Moschus_, ii. 219 + + _Motacilla_, European Miocene, i. 161 + ii. 290 + + MOTACILLIDÆ, ii. 290 + + _Motella_, ii. 439 + + Moths, ii. 481 + + Motmots, ii. 313 + + Mound-builders, peculiar Australian birds, i. 393 + + Moupin, position and zoology of, i. 221 + + Mouse-deer, ii. 218 + + _Moxostoma_, ii. 451 + + _Mugil_, ii. 435 + + MUGILLIDÆ, ii. 435 + + _Mulleria_, ii. 534 + + _Mulleripicus_, ii. 303 + + MULLIDÆ, ii. 426 + + _Mullus_, ii. 426 + + _Mungos_, ii. 195 + + _Munia_, ii. 287 + + MURÆNIDÆ, ii. 456 + + _Murænopsis_, ii. 412 + + _Murex_, ii. 507 + + MURICIDÆ, ii. 507 + + _Muridæ_, S. American Pliocene, i. 147 + + MURIDÆ, ii. 229 + + Murray, Mr. Andrew, on zoological region, i. 60 + + _Mus_, ii. 229 + + _Muscardinus_, ii. 232 + + _Muscicapa_, ii. 270 + + MUSCICAPIDÆ, ii. 270 + + _Muscicapula_, ii. 270 + + _Muscigralla_, ii. 101 + + _Muscipipra_, ii. 101 + + _Muscisaxicola_, ii. 101, 291 + + _Muscitodus_, ii. 271 + + _Muscivora_, ii. 101 + + _Musophaga_, ii. 307 + + MUSOPHAGIDÆ, ii. 307 + + Mussels, ii. 533 + + _Mustela_, Miocene of Greece, i. 115 + European Miocene, i. 118 + S. American Pliocene, i. 146 + ii. 198 + + _Mustelidæ_, in Brazilian caves, i. 144 + + MUSTELIDÆ, ii. 198 + + _Mustelus_, ii. 460 + + MYACIDÆ, ii. 536 + + _Myadora_, ii. 536 + + _Mycalesis_, ii. 471 + + _Mycerobas_, ii. 284 + + _Mycetes_, ii. 175 + ii. 178 + + _Mycetopus_, ii. 534 + + _Mydaus_, ii. 199 + + _Myiadestes_, ii. 260 + + _Myiagra_, ii. 271 + + _Myialestes_, ii. 271 + + _Myiarchus_, ii. 102, 291 + + _Myiobius_, ii. 101 + + _Myioceyx_, ii. 316 + + _Myiochanes_, ii. 102 + + _Myiodioctes_, ii. 279 + + _Myiodynastes_, ii. 101 + + _Myiophonus_, ii. 263 + + _Myiopithecus_, ii. 173 + + _Myiotheretes_, ii. 100 + + _Myiozetetes_, ii. 101 + + _Mylesinus_, ii. 445 + + _Myletes_, ii. 445 + + MYLIOBATIDÆ, ii. 463 + + _Myliobatis_, ii. 463 + + _Mylodon_, N. American Post-Pliocene, i. 130 + S. American Pliocene, i. 147 + + _Mylopharadon_, ii. 452 + + _Mynes_, ii. 474 + + _Myochama_, ii. 536 + + _Myodes_, ii. 230 + + _Myogale_, European Miocene, i. 118 + ii. 190, 191 + + _Myoictis_, ii. 249 + + _Myomorphus_, fossil in Cuba, i. 148 + + _Myopotamus_, in Brazilian caves, i. 145 + ii. 239 + + _Myospalax_, ii. 230 + + MYOXIDÆ, ii. 232 + + _Myoxus_, European Miocene, i. 120 + European Eocene, i. 126 + ii. 232 + + _Myxomys_, ii. 230 + + _Myrina_, ii. 477 + + _Myrmeciza_, ii. 104 + + MYRMECOBIIDÆ, ii. 250 + + _Myrmecobius_, ii. 250 + + _Myrmecophaga_, ii. 247 + + MYRMECOPHAGIDÆ, ii. 247 + + _Myrmotherula_, ii. 104 + + _Myron_, ii. 376 + + _Myrtis_, ii. 108 + + _Mysarachne_, European Miocene, i. 118 + + _Mysops_, N. American Eocene, i. 140 + ii. 231 + + _Mystacina tuberculata_, ii. 184 + + _Mystacoleucus_, ii. 452 + + _Mystacornis_, ii. 258 + + _Mystromys_, ii. 230 + + MYTILIDÆ, ii. 533 + + _Mytilus_, ii. 539 + + MYXINIDÆ, ii. 464 + + _Myxophagus_, N. American Post-Pliocene, i. 130 + + _Myxophyes_, ii. 420 + + _Myxus_, ii. 435 + + _Myzomela_, ii. 275 + + _Myzornis_, ii. 266 + + N. + + _Nænia_, ii. 365 + + _Naja_, ii. 383 + + NANDIDÆ, ii. 433 + + _Nandinia_, ii. 195 + + _Nandus_, ii. 433 + + _Nanina_, ii. 513 + + _Nannophryne_, ii. 417 + + _Nannophrys_, ii. 421 + + _Nanodes_, ii. 327 + + _Nanohyus_, N. American Tertiary, i. 137 + ii. 215 + + _Nanotragus_, ii. 224 + + _Napeogenes_, ii. 470 + + _Napothera_, ii. 261 + + _Nardoa_, ii. 381 + + Narwhal, ii. 208 + + _Narope_, ii. 472 + + _Nasica_, ii. 103 + + _Nasiterna_, ii. 325 + + _Nasua_, in Brazilian caves, i. 144 + ii. 200 + + _Nathalis_, ii. 478 + + _Natica_, ii. 539 + + NATICIDÆ, ii. 508 + + _Nautilus_, ii. 539 + + NATRICINÆ, ii. 375 + + _Nattereria_, ii. 417 + + _Nauclerus_, ii. 349 + + _Naucrates_, ii. 429 + + _Naultinus_ ii. 400 + + NAUTILIDÆ, ii. 506 + + _Navicella_, ii. 510 + + Nearctic region, defined, i. 79 + subdivisions of, i. 80 + distinct from Palæarctic, i. 79 + ii. 114 + zoological characteristics of, ii. 115 + mammalia of, ii. 115 + birds of, ii. 116 + reptiles of, ii. 119 + amphibia of, ii. 120 + fresh-water fishes of, ii. 120 + summary of vertebrata of, ii. 120 + insects of, ii. 122 + land and fresh-water shells of, ii. 124 + sub-regions of, ii. 125 + concluding remarks on, ii. 138 + tables of distribution of animals of, ii. 139 + + Nearctic and Neotropical regions, no decided boundary between, ii. 117 + + _Nebria_, ii. 489 + + _Necrornis_, European Miocene, i. 161 + + NECTARINIIDÆ, ii. 276 + + _Nectarinia_, ii. 276 + + _Nectarophila_, ii. 276 + + _Nectogale_, ii. 190 + + _Necydalis_, ii. 502 + + _Necyria_, ii. 476 + + _Nelicurvius_, ii. 286 + + _Nemachilus_, ii. 453 + + _Nematogenys_, ii. 444 + + NEMEOBIIDÆ, ii. 475 + + _Nemeobius_, ii. 475 + + NEMORHEDINÆ, ii. 224 + + _Nemorhedus_, ii. 224 + + _Nemoricola_, ii. 290 + + _Nemosia_, ii. 99 + + _Neobatrachus_, ii. 420 + + _Neochloe_, ii. 280 + + _Neocorys_, ii. 290 + + _Neoctantes_, ii. 104 + + _Neomeris_, ii. 209 + + _Neomorphus_, ii. 309 + + _Neophasia_, ii. 478 + + _Neophron_, ii. 346 + + _Neopipo_, ii. 102 + + _Neopus_, ii. 348 + + _Neorhynchus_, ii. 285 + + _Neosorex_, ii. 191 + + _Neotoma_, ii. 230 + + _Neotomys_, ii. 230 + + _Neotragus_, ii. 224 + + Neotropical region, defined, i. 78 + subdivisions of, i. 78 + relations of W. African sub-region with, i. 265 + description of, ii. 1 + zoological features of, ii. 5 + birds of, ii. 6, 7 + distinctive features of mammalia of, ii. 6 + reptiles of, ii. 9 + amphibia of, ii. 11 + fresh-water fishes of, ii. 12 + summary of vertebrates of, ii. 13 + insects of, ii. 13 + land-shells of, ii. 19 + marine shells of, ii. 20 + summary of past history of, ii. 80 + tables of distribution of animals of, ii. 84 + + Neotropical sub-regions, ii. 21 + + _Neoziphius_, ii. 208 + + _Nephæcetes_, ii. 320 + + _Neptis_, ii. 474 + + NERITIDÆ, ii. 510 + + _Neritina_, ii. 510 + + _Nerophis_, ii. 457 + + _Nesoceleus_, ii. 303 + + _Nesocichla_, ii. 256 + + _Nesodon_, S. American Pliocene, i. 147 + + _Nesomys_, ii. 230 + + _Nesonetta_, ii. 364 + + _Nesopsar_, ii. 282 + + _Nessia_, ii. 399 + + _Nestor_, ii. 329 + + NESTORIDÆ, ii. 329 + + _Nettapus_, ii. 363 + + _Neusterophis_, ii. 376 + + Newberry, Dr., on Cretaceous and Tertiary floras of N. America, ii. 155 + + Newton, Professor, on position of _Menuridæ_ and _Atrichiidæ_, i. 95 + on birds of Iceland, i. 198 + on Neotropical sub-regions, ii. 25 + on genus _Camptolæmus_, ii. 39 + on peculiar genera of Nearctic and Neotropical birds, ii. 118 + on family _Panuridæ_, ii. 262 + + _Newtonia_, ii. 270 + + Newts, ii. 413 + + New Caledonia, birds of, i. 444 + + New Guinea, zoology of, i. 409 + mammalia of, i. 410 + birds of, i. 411 + peculiarities of its ornithology, i. 413 + illustration of ornithology of, i. 414 + reptiles and amphibia of, i. 415 + insects of, i. 416 + + New Zealand, objections to making a primary zoological region, i. 62 + extinct birds of, i. 164 + sub region, description of, i. 449 + compared with British Isles, i. 449 + mammalia of, i. 451 + islets of, i. 453 + illustration of ornithology of, i. 455 + reptiles of, i. 456 + amphibia of, i. 457 + fresh-water fish of, i. 457 + insects of, i. 458 + Longicorns of, i. 458 + Myriapoda of, i. 458 + land-shells of, i. 459 + ancient fauna of, i. 460 + origin of fauna of, i. 460 + poverty of insects in, i. 462 + relations of insect-fauna and flora of, i. 472 + + _Nicator_, ii. 272 + + Nicobar Islands, their zoological relations, i. 332 + + Nightingale, migration of the, i. 21 + + Night-jars, ii. 319 + + _Nigidius_, ii. 493 + + _Nigrita_, ii. 286 + + _Nilaus_, ii. 272 + + _Niltava_, ii. 270 + + _Ninox_, ii. 350 + + _Nisaëtus_, ii. 348 + + _Nisoides_, ii. 348 + + _Nisoniades_, ii. 480 + + NOCTILIONIDÆ, ii. 184 + + Nocturnal tree-snakes, ii. 379 + + _Nonnula_, ii. 311 + + _Norbea_, ii. 397 + + Norfolk Island, birds of, i. 453 + + North Africa, zoological relations of, i. 202 + + North America, remarks on Post-Pliocene, fauna of, i. 130 + Post-Pliocene fauna of, partly derived from S. America, i. 131 + extinct birds of, i. 163 + + Northern Hemisphere, zoological importance of, ii. 155 + + NOTACANTHI, ii. 437 + + _Notaden_, ii. 415 + + _Notharctos_, N. American Tertiary, i. 133 + + _Nothocerus_, ii. 344 + + _Nothocrax_, ii. 343 + + _Nothoprocta_, ii. 344 + + _Nothura_, ii. 344 + + NOTIDANIDÆ, ii. 461 + + _Notiophilus_, ii. 489 + + _Notodela_, ii. 259 + + _Notoglanis_, ii. 443 + + _Notonomus_, ii. 490 + + NOTOPTERIDÆ, ii. 455 + + _Notopholis_, ii. 391 + + Notornis of New Zealand, i. 455 + + _Notornis_, ii. 352 + + _Nototherium_, Australian Post-Tertiary, i. 157 + ii. 251 + + _Nototrema_, ii. 418 + + _Noturus_, ii. 442 + + _Nucifraga_, ii. 273 + + NUCLEO-BRANCHIATA. ii. 531 + + _Nucras_, ii. 391 + + _Numenius_, ii. 353 + + NUMIDINÆ, ii. 340 + + _Numida_, ii. 340 + + _Nuria_, ii. 452 + + Nuthatches, ii. 265 + + _Nutria_, ii. 199 + + _Nyctala_, ii. 350 + + _Nyctalatinus_, ii. 350 + + _Nyctalemon_, ii. 482 + + _Nyctalops_, ii. 350 + + _Nyctea_, ii. 350 + + _Nyctereutes_, ii. 197 + + _Nycteris_, ii. 182 + + _Nyctibius_, ii. 319 + + _Nycticorax_, ii. 359 + + _Nyctidromius_, ii. 320 + + _Nyctiornis_, ii. 312 + + _Nyctipithecus_, ii. 175 + + _Nyctiprogne_, ii. 320 + + _Nymphalis_, ii. 474 + + NYMPHALIDÆ, ii. 473 + + _Nymphicus_, ii. 325 + + _Nymphidium_, ii. 476 + + O. + + _Ochetobius_, ii. 452 + + _Ochotherium_, in Brazilian caves, i. 145 + ii. 245 + + _Ochthæca_, ii. 100 + + _Ochthodiæta_, ii. 100 + + _Octodontidæ_, S. American Pliocene, i. 147 + + OCTODONTIDÆ, ii. 237 + + _Octodon_, ii. 238 + + OCTOPODIDÆ, ii. 505 + + _Ocyalus_, ii. 282 + + _Ocydromus_, ii. 352 + + _Ocyphaps_, ii. 333 + + _Odontochila_, ii. 486 + + _Odontolabris_, ii. 493 + + _Odontophorus_, ii. 339 + + _Odontophrynus_, ii. 420 + + _Oedemia_, ii. 364 + + _Oedicnemis_, ii. 355 + + _Oëdura_, ii. 399 + + _Oena_, ii. 332 + + _Ogmodon_, ii. 383 + + _Ogyris_, ii. 477 + + _Oligdon_, ii. 375 + + OLIGODONTIDÆ, ii. 374 + + _Oligosarcus_, ii. 445 + + _Olisthopus_, ii. 489 + + _Olylogon_, ii. 418 + + _Olyra_, ii. 442 + + _Omaseus_, ii. 489 + + _Ommatophoca_, ii. 204 + + _Omolepida_, ii. 397 + + _Omostenus_, ii. 492 + + _Omphalotropis_, ii. 521 + + _Omus_, ii. 487 + + ONCIDIADÆ, ii. 517 + + _Oncidium_, ii. 518 + + _Onychodactylus_, ii. 413 + + _Onychogale_, ii. 195 + + _Onychogalea_, ii. 251 + + _Onychognathus_, ii. 288 + + _Onchorhynchus_, ii. 447 + + _Oncostoma_, ii. 101 + + Ophidia, classification of, i. 99 + + OPHIDIA, ii. 372 + remarks on the general distribution of, ii. 386 + fossil, ii. 387 + + OPHIDIIDÆ, ii. 440 + + _Ophidium_, ii. 440 + + OPHIOCEPHALIDÆ, ii. 435 + + _Ophiodes_, ii. 397 + + OPHIOMORIDÆ, ii. 398 + + _Ophiomorus_, ii. 398 + + _Ophiophagus_, ii. 383 + + _Ophiops_, ii. 391 + + _Ophioscincus_, ii. 397 + + _Ophisaurus_, ii. 392 + + _Ophites_, ii. 380 + + _Ophonus_, ii. 489 + + _Ophryodera_, ii. 487 + + _Ophysia_, ii. 209 + + OPISTHO-BRANCHIATA, ii. 529 + + OPISTHOCOMI, ii. 345 + + _Opisthocomus_, Brazilian caves, i. 164 + ii. 345 + + _Opisthodelphys_, ii. 418 + + _Opisthostoma_, ii. 520 + + _Opisthoporus_, ii. 520 + + _Oporornis_, ii. 279 + + _Opossum_, extinct in European Miocene, i. 121 + + Opossums, ii. 248 + + _Opsariichthys_, ii. 452 + + _Opsiphanes_, ii. 472 + + Orang-utan, ii. 171 + + _Orca_, ii. 209 + + _Orcaella_, ii. 209 + + _Orchesticus_, ii. 99 + + _Orchilus_, ii. 101 + + _Oreas_, ii. 223 + + _Oreicola_, ii. 260 + + _Oreinus_, ii. 452 + + _Oreocephalus_, ii. 401 + + _Oreocincla_, ii. 256 + + _Oreodeira_, ii. 401 + + _Oreodon_, N. American Tertiary, i. 138 + + _Oreodontidæ_, N. American Tertiary, i. 138 + + _Oreoeca_, ii. 271 + + _Oreomanes_, ii. 278 + + _Oreonectes_, ii. 453 + + _Oreonympha_, ii. 108 + + _Oreoperdix_, ii. 338 + + OREOPHASINÆ, ii. 343 + + _Oreophasis_, ii. 343 + + _Oreophilus_, ii. 356 + + _Oreopyra_, ii. 107 + + _Oreortyx_, ii. 339 + + _Oreoscoptes_, ii. 256 + + _Oreothraupis_, ii. 99 + + _Oreotrochilus_, ii. 107 + + _Orestias_, ii. 450 + + Oriental region, defined, i. 75 + subdivisions of, i. 75 + description of, i. 314 + zoological features of, i. 315 + mammalia of, i. 315 + birds of, i. 316 + reptiles of, i. 317 + amphibia of, i. 317 + fresh-water fishes of, i. 318 + summary of vertebrata of, i. 318 + insects of, i. 318 + sub-regions of, i. 321 + concluding remarks on, i. 362 + tables of distribution of animals of, i. 364 + + Oriental relations of W. African sub-region, i. 265 + + Oriental and Palæarctic faunas once identical, i. 362 + + Oriental and Ethiopian faunas, cause of their resemblances, i. 363 + + _Origma_, ii. 260 + + _Oriocalotes_, ii. 402 + + Orioles, ii. 268 + + ORIOLIDÆ, ii. 268 + + _Oriolus_, ii. 268 + + _Orites_, ii. 266 + + _Ornithion_, ii. 101 + + ORNITHORHYNCHIDÆ, ii. 253 + + _Ornithorhynchus_, ii. 253 + + _Orocætes_, ii. 256 + + _Orohippus_, N. American Tertiary, i. 136 + + _Ortalida_, ii. 343 + + _Orthagoriscus_, ii. 457 + + _Orthalicus_, ii. 516 + + ORTHIDÆ, ii. 532 + + ORTHOCERATIDÆ, ii. 506 + + _Orthodon_, ii. 452 + + _Orthogonius_, ii. 491 + + _Orthogonys_, ii. 98 + + _Orthonyx_, ii. 260 + + _Orthorhynchus_, ii. 108 + + _Orthotomus_, ii. 257 + + _Ortygometra_, ii. 352 + + _Ortygornis_, ii. 338 + + _Ortyx_, ii. 339 + + _Ortyxelos_, ii. 341 + + ORYCTEROPODIDÆ, ii. 246 + + _Orycteropus_, ii. 246 + + _Orycterus_, ii. 231 + + ORYGINÆ, ii. 223 + + _Oryx_, ii. 223 + + _Oryzoborus_, ii. 285 + + _Oryzorictes_, ii. 188 + + _Osmerus_, ii. 447 + + _Osphranter_, ii. 251 + + _Osteobrama_, ii. 453 + + _Osteochilus_, ii. 451 + + _Osteogeniosus_, ii. 443 + + OSTEOGLOSSIDÆ, ii. 454 + + _Osteoglossum_, ii. 454 + + _Ostinops_, ii. 282 + + OSTREIDÆ, ii. 533 + + _Ostrich_, Miocene of N. India, i. 162 + + Ostriches, ii. 368 + + _Otaria_, European Miocene, i. 118 + ii. 202 + + OTARIIDÆ, ii. 202 + + OTIDIDÆ, ii. 356 + + _Otidiphaps_, ii. 333 + + _Otilophus_, ii. 415, 428 + + _Otis_, ii. 356 + + _Otocorys_, ii. 289 + + _Otocryptis_, ii. 402 + + _Otogyps_, ii. 346 + + _Otomys_, ii. 230 + + _Otopoma_, ii. 521 + + _Ovibos_, N. American Post-Pliocene, i. 130 + ii. 224, 225 + + Owl-parrot, ii. 329 + + Owls, ii. 350 + + Oxen, birth-place and migrations of, i. 155 + Palæarctic, i. 182 + ii. 221 + + OXUDERCIDÆ, ii. 431 + + _Oxyæna_, N. American Tertiary, i. 134 + + _Oxydoras_, ii. 443 + + _Oxyglossus_, ii. 421 + + _Oxygomphus_, European Miocene, i. 118 + ii. 186 + + _Oxylabes_, ii. 262 + + _Oxymycterus_, in Brazilian caves, i. 145 + S. American Pliocene, i. 147 + ii. 230, 231 + + _Oxynotus_, ii. 269 + + _Oxypogon_, ii. 108 + + OXYRHAMPHIDÆ, ii. 292 + + _Oxyrhamphus_, ii. 292 + + _Oxyrhopus_, ii. 379 + + _Oxyurus_, ii. 103 + + Oysters, ii. 533 + + P. + + _Pachybatrachus_, ii. 416 + + _Pachycephala_, ii. 271 + + PACHYCEPHALIDÆ, ii. 271 + + _Pachydactylus_, ii. 400 + + _Pachyæna_, N. American Tertiary, i. 134 + + _Pachyglossa_, ii. 277 + + _Pachynolophus_, European Eocene, i. 126 + + _Pachyrhamphus_, ii. 102 + + _Pachyrhynchus_, ii. 391 + + _Pachyteles_, ii. 490, 492 + + _Pachytherium_, in Brazilian caves, i. 145 + ii. 246 + + _Pachyura_, ii. 191 + + _Pæocephalus_, ii. 328 + + _Pæcilus_, ii. 489 + + _Pagellus_, ii. 427 + + _Pagomys_, ii. 204 + + _Pagophila_, ii. 364 + + _Pagophilus_, ii. 204 + + _Paguma_, ii. 195 + + PAICTIDÆ, ii. 298 + + Palæarctic region, ancient limits of, ii. 157 + defined, i. 171 + subdivisions of, i. 71 + general features of. i. 180 + zoological charcteristics of, i. 181 + has few peculiar families, i. 181 + mammalia of, i. 181 + birds of, i. 182 + high degree of speciality of, i. 184 + reptiles and amphibia of, i. 186 + fresh-water fish of, i. 186 + summary of vertebrata of, i. 186 + insects of, i. 186 + coleoptera of, i. 187 + number of coleoptera of, i. 189 + land-shells of, i. 190 + sub-regions of, i. 190 + general conclusions on the fauna of, i. 231 + tables of distribution of animals of, i. 233 + + _Palæacodon_, N. American Tertiary, i. 133 + + _Palæetus_, European Miocene, i. 162 + + _Palægithalus_, European Eocene, i. 162 + + _Palælodus_, European Miocene, i. 162 + + _Palæocastor_, N. American Tertiary, i. 140 + ii. 234 + + _Palæocercus_, European Miocene, i. 162 + + _Palæochoerus_, European Miocene, i. 119 + ii. 215 + + _Palæocyon_, ii. 198 + + _Palæohierax_, European Miocene, i. 162 + + _Palæolagus_, N. American Tertiary, i. 140 + + _Palæolama_, S. American Pliocene, i. 147 + ii. 217 + + _Palæomephitis_, European Miocene, i. 118 + ii. 200 + + _Palæomeryx_, European Miocene, i. 120 + ii. 220 + + _Palæomys_, European Miocene, i. 121 + ii. 238 + + _Palæontina oolitica_, Oolitic insect, i. 167 + + Palæontology, i. 107 + how best studied in its bearing on geographical distribution, i. 168 + as an introduction to the study of geographical distribution, + concluding remarks on, i. 169 + + _Palæonyctis_, European Eocene, i. 125 + + _Palæoperdix_, European Miocene, i. 161 + + _Palæophrynus_, European Miocene, i. 166 + + _Palæoreas_, Miocene of Greece, i. 116 + + _Palæornis_, ii. 326 + + PALÆORNITHIDÆ, ii. 326 + + _Palæonyctis_, ii. 196, 206 + + _Palæortyx_, European Miocene, i. 161 + + _Palæoryx_, Miocene of Greece, i. 116 + + _Palæospalax_, i. 111 + European Miocene, i. 117 + ii. 190 + + _Palæosyops_, N. American Tertiary, i. 136 + + _Palæotheridæ_, European Eocene, i. 125 + + _Palæotherium_, Enropean Eocene, i. 125 + S. American Eocene, i. 148 + + _Palæotragus_, Miocene of Greece, i. 116 + + _Palæotringa_, N. American Cretaceous, i. 164 + + _Palamedea_, ii. 361 + + PALAMEDEIDÆ, ii. 361 + + _Palapterygidæ_ of New Zealand, i. 164 + + PALAPTERYGIDÆ, ii. 370 + + _Palapteryx_, ii. 370 + + Palestine, birds of, i. 203 + + _Pallasia_, ii. 289 + + _Paloplotherium_, European Miocene, i. 119 + European Eocene, i. 125 + + _Paludicola_, ii. 416 + + _Paludina_, Eocene, i. 169 + European Secondary, i. 169 + ii. 510 + + PALUDINIDÆ, ii. 510 + + Pampas, Pliocene deposits of, i. 146 + + _Pamphila_, ii. 480 + + Panda, of Nepaul and E. Thibet, i. 222 + Himalayan, figure of, i. 331 + ii. 201 + + _Pandion_, ii. 349 + + PANDIONIDÆ, ii. 349 + + _Pangasius_, ii. 442 + + _Pangolin_, ii. 245 + + _Panolax_, N. American Tertiary, i. 140 + + _Panopoea_, ii. 536 + + _Panoplites_, ii. 107 + + _Panterpe_, ii. 109 + + _Panthalops_, ii. 223 + + PANURIDÆ, ii. 262 + + _Panurus_, ii. 262 + + _Panychlora_, ii. 109 + + _Panyptila_, ii. 320 + + Paper-Nautilus, ii. 505 + + _Paphia_, ii. 474 + + _Papilio_, ii. 479 + + PAPILIONIDÆ, ii. 479 + + Papuan Islands, zoology of, i. 409 + + _Paracanthobrama_, ii. 452 + + _Paradigalla_, ii. 275 + + _Paradiplomystax_, ii. 443 + + _Paradisea_, ii. 274 + + Paradise-bird, twelve-wired, figure of, i. 414 + + Paradise-birds, ii. 274 + + PARADISEIDÆ, ii. 274 + + PARADISEINÆ, ii. 274 + + _Paradoxornis_, ii. 262 + + _Paradoxurus_, ii. 195 + + _Parahippus_, N. American Tertiary, i. 136 + + _Paralabraz_, ii. 425 + + _Paramys_, N. American Eocene, i. 140 + ii. 236 + + _Parandra_, ii. 501 + + _Paraphoxinus_, ii. 452 + + _Pardalotus_, ii. 277 + + _Pareas_, ii. 380 + + _Parodon_, ii. 445 + + _Pareudiastes_, ii. 352 + + PARIDÆ, ii. 265 + + _Pariodon_, ii. 444 + + _Parisoma_, ii. 266 + + _Parmacella_, ii. 517 + + _Parmarion_, ii. 517 + + _Parmophorus_, ii. 511 + + _Parnassius_, ii. 479 + + _Paroaria_, ii. 284 + + _Parotia_, ii. 274 + + _Parra_, ii. 355 + + PARRIDÆ, ii. 354 + + Parroquet, Papuan, figure of, i. 415 + + Parrots, classification of, i. 96 + ii. 324, 329 + + Partridges, ii. 338 + + _Partula_, ii. 515 + + _Parula_, ii. 279 + + _Parus_, ii. 265 + + _Pasimachus_, ii. 490 + + _Passerculus_, ii. 284 + + _Passerella_, ii. 284 + + Passeres, arrangement of, i. 94 + range of Palæarctic genera of, i. 243 + range of Ethiopian genera of, i. 306 + range of Oriental genera of, i. 375 + range of Australian genera of, i. 478 + + PASSERES, ii. 255 + general remarks on the distribution of, ii. 299 + + _Passerita_, ii. 379 + + _Pastor_, ii. 287 + + _Patagona_, ii. 108 + + _Patella_, ii. 539 + + PATELLIDÆ, ii. 511 + + _Patriofelis_, N. American Tertiary, i. 134 + + _Patrobus_, ii. 489 + + _Pauxi_, ii. 343 + + _Pavo_, ii. 340 + + PAVONINÆ, ii. 340 + + _Paxillus_, ii. 520 + + Pearl-oysters, ii. 533 + + Pease, Mr. Harper, on Polynesian region of Land-shells, ii. 528 + + Peccaries, ii. 215 + + _Pectinator_, ii. 238 + + Peculiar groups, geographically, how defined, ii. 184 + + _Pedetes_, ii. 232 + + PEDICULATI, ii. 431 + + _Pediocætes_, ii. 339 + + _Pedionomus_, ii. 356 + + PEGASIDÆ, ii. 456 + + _Pelagius_, ii. 204 + + _Pelagornis_, European Miocene, i. 162 + + _Pelamis_, ii. 384 + + _Pelargopsis_, ii. 316 + + _Pelea_, ii. 224 + + PELECANIDÆ, ii. 365 + + _Pelecanoides_, ii. 365 + + _Pelecanus_, ii. 365 + + _Pelecium_, ii. 490 + + _Pelecus_, ii. 453 + + Pelicans, ii. 365 + + _Peliperdix_, ii. 338 + + _Pellorneum_, ii. 261 + + _Pelobates_, ii. 417 + + PELODRYADÆ, ii. 418 + + _Pelodryas_, ii. 418 + + _Pelodytes_, ii. 421 + + _Pelomedusa_, ii. 409 + + _Pelomys_, ii. 230 + + _Pelonax_, N. American Tertiary, i. 138 + + _Peloperdix_, ii. 338 + + _Pelotrophus_, ii. 453 + + _Peltaphryne_, ii. 415 + + _Peltocephalus_, ii. 408 + + _Peltopelor_, ii. 385 + + _Peltops_, ii. 270 + + _Penelope_, ii. 343 + + _Penelopides_, ii. 317 + + _Penelopina_, ii. 343 + + PENELOPINÆ, ii. 343 + + _Penetes_, ii. 472 + + Penguins, ii. 366 + + _Pentadactylus_, ii. 399 + + _Pentila_, ii. 477 + + _Peragalea_, ii. 250 + + _Perameles_, ii. 250 + + PERAMELIDÆ, ii. 250 + + _Peratherium_, European Miocene, i. 121 + European Eocene, i. 126 + ii. 249 + + _Perca_, ii. 425 + + _Percarina_, i. 425 + + _Perchoerus_, N. American Tertiary, i. 137 + ii. 215 + + _Percilia_, ii. 425 + + _Percichthys_, ii. 425 + + PERCIDÆ, ii. 425 + + _Percnostola_, ii. 104 + + PERCOPSIDÆ, ii. 448 + + _Percus_, ii. 489 + + _Perdix_, ii. 338 + + _Pericallus_, ii. 490 + + _Pericrocotus_, ii. 268 + + _Peridexia_, ii. 487 + + Perim Island, extinct mammalia of, i. 122 + probable southern limit of old Palæarctic land, i. 362 + character of fossils of, ii. 157 + + _Periopthalmus_, ii. 430 + + _Perisoreus_, ii. 273 + + _Perissodactyla_, N. American Tertiary, i. 135 + + _Perissoglossa_, ii. 279 + + _Peristera_, ii. 333 + + _Peristethus_, ii. 428 + + Periwinkle, ii. 510 + + _Pernis_, ii. 349 + + _Perodicticus_, ii. 176 + + _Perognathus_, ii. 233 + + _Peropus_, ii. 399 + + Persia, birds of, i. 204 + + _Petasophora_, ii. 108 + + _Petaurista_, ii. 252 + + _Petenia_, ii. 438 + + _Petrochelidon_, ii. 281 + + _Petrodromus_, ii. 186 + + Petrels, ii. 365 + + _Petroeca_, ii. 260 + + _Petrogale_, ii. 251 + + _Petromys_, ii. 239 + + _Petrophassa_, ii. 333 + + _Petrorhynchus_, ii. 208 + + _Petroscirtes_, ii. 431 + + _Peucæa_, ii. 284 + + _Pezophaps_, ii. 334 + + _Pezoporus_, ii. 325 + + _Pfeifferia_, ii. 516 + + _Phacellodomus_, ii. 103 + + _Phacochoerus_, ii. 215 + + _Phænicophaës_, ii. 309 + + _Phænicophilus_, ii. 99 + + _Phænicothraupis_, ii. 98 + + _Phænopepla_, ii. 280 + + _Phæochroa_, ii. 107 + + _Phæolæma_, ii. 107 + + _Phæoptila_, ii. 109 + + _Phaëthornis_, ii. 107 + + _Phaeton_, ii. 365 + + _Phalacrocorax_, ii. 365 + + Phalangers, ii. 251 + + _Phalangista_, ii. 252 + + Phalangistidæ, ii. 251 + + _Phalaropus_, ii. 353 + + _Phapitreron_, ii. 333 + + _Phaps_, ii. 333 + + _Pharomacrus_, ii. 314 + + _Phascogale_, ii. 249 + + _Phascolarctos_, ii. 252 + + PHASCOLOMYIDÆ, ii. 252 + + _Phascolomys_, Australian Post-Tertiary, i. 157 + + PHASIANIDÆ, ii. 339 + + PHASIANINÆ, ii. 340 + + _Phasianus_, Miocene of Greece, i. 116 + European Post-Pliocene, i. 161 + ii. 340 + + _Phasidus_, ii. 340 + + _Phatagin_, ii. 245 + + Pheasants, in European Miocene, i. 161 + golden, of N. China, i. 226 + eared, of Mongolia, i. 226 + ii. 339 + + _Phedina_, ii. 281 + + _Phelsuma_, ii. 400 + + _Phenacodus_, N. American Tertiary, i. 138 + + _Pheropsophus_, ii. 489 + + _Pheucticus_, ii. 285 + + _Phibalura_, ii. 102 + + _Philagetes_, ii. 502 + + _Philemon_, ii. 276 + + _Philentoma_, ii. 271 + + _Philepitta_, ii. 298 + + _Philetærus_, ii. 286 + + _Philodryas_, ii. 376 + + Philippine Islands, mammals of, i. 345 + birds of, i. 346 + origin of peculiar fauna of, i. 448 + + _Philohela_, ii. 353 + + _Philomycus_, ii. 517 + + _Philydor_, ii. 103 + + PHILYDORINÆ, ii. 295 + + _Phlæomys_, ii. 230 + + _Phlæocryptes_, ii. 103 + + _Phlogoenas_, ii. 333 + + _Phlogophilus_, ii. 108 + + _Phlogopsis_, ii. 104 + + _Phlogothraupis_, ii. 98, 283 + + _Phoca_, ii. 204 + + _Phocæna_, ii. 209 + + _Phocidæ_, N. American Tertiary, i. 140 + + PHOCIDÆ, ii. 203 + + _Phodilus_, ii. 350 + + _Phoenicocercus_, ii. 102, 293 + + _Phoenicophaës_, ii. 309 + + PHOENICOPTERIDÆ, ii. 361 + + _Phoenicopterus_, ii. 361 + + PHOLADIDÆ, ii. 537 + + _Pholadomya_, ii. 536 + + _Pholeoptynx_, ii. 350 + + _Pholidotus_, ii. 245 + + _Pholidotus_, ii. 493 + + _Phonipara_, ii. 284 + + _Phorus_, ii. 510 + + _Phos_, ii. 507 + + _Phractocephalus_, ii. 442 + + _Phrygilus_, ii. 284 + + PHRYNISCIDÆ, ii. 414 + + _Phryniscus_, ii. 414 + + _Phrynobatrachus_, ii. 421 + + _Phrynocephalus_, ii. 402 + + _Phrynoglossus_, ii. 421 + + _Phrynorhombus_, ii. 441 + + _Phrynosoma_, ii. 401 + + _Phycis_, ii. 439 + + _Phyllastrephus_, ii. 267 + + PHYLLIDIADÆ, ii. 530 + + _Phyllobates_, ii. 419 + + _Phyllodactylus_, ii. 399 + + _Phyllomedusa_, ii. 418 + + _Phyllomyias_, ii. 101 + + _Phyllomys_, in Brazilian caves, i. 145 + ii. 239 + + _Phyllornis_, ii. 267 + + PHYLLORNITHIDÆ, ii. 267 + + _Phylloscartes_, ii. 101 + + PHYLLOSCOPINÆ, ii. 257 + + _Phylloscopus_, ii. 258 + + _Phyllostomidæ_, in Brazilian caves, i. 144 + + PHYLLOSTOMIDÆ, ii. 181 + + _Phyllurus_, ii. 400 + + PHYLLYRHOIDÆ, ii. 530 + + _Phymaturus_, ii. 401 + + _Physa_, ii. 518 + + _Physalus_, ii. 207 + + _Physeter_, European Pliocene, i. 112 + ii. 208 + + Physical changes affecting distribution, i. 7 + + _Physignathus_, ii. 402 + + PHYSOSTOMI, ii. 441 + + _Phytala_, ii. 477 + + _Phytotoma_, ii. 294 + + PHYTOTOMIDÆ, ii. 294 + + _Phyton_, ii. 502 + + _Piabuca_, ii. 445 + + _Piabucina_, ii. 445 + + _Piaya_, ii. 309 + + _Pica_, ii. 273 + + Picariæ, arrangement of, i. 95 + range of Palæarctic genera of, i. 247 + range of Ethiopian genera of, i. 309 + range of Oriental genera of, i. 381 + range of Australian genera of, i. 482 + + PICARIÆ, ii. 302 + general remarks on the distribution of, ii. 322 + + _Picathartes_, ii. 274 + + _Picicorvus_, ii. 273 + + PICIDÆ, ii. 302 + + _Picoides_, ii. 303 + + _Picolaptes_, ii. 103 + + _Picumnus_, ii. 303 + + _Picus_, European Miocene, i. 161 + ii. 303 + + PIERIDÆ, ii. 478 + + _Pieris_, ii. 478 + + _Piezia_, ii. 491 + + Pigeons, classification of, i. 96 + remarkable development of, in the Australian region, i. 395 + crested, of Australia, figure of, i. 441 + ii. 331 + abundant in islands, ii. 335 + + Pigs, power of swimming, i. 13 + + Pikas, ii. 242 + + Pike, ii. 449 + + Pikermi, Miocene fauna of, i. 115 + + Pilchard, ii. 454 + + _Pileoma_, ii. 425 + + _Pimelodus_, ii. 443 + + _Pimephales_, ii. 452 + + _Pinacodera_, ii. 490 + + _Pinicola_, ii. 285 + + _Pinulia_, ii. 191 + + _Pionus_, ii. 328 + + _Pipa_, ii. 422 + + PIPIDÆ, ii. 421 + + _Pipile_, i. 343 + + _Pipilo_, ii. 284 + + Piping crows, ii. 273 + + _Pipra_, ii. 102, 292 + + _Pipreola_, ii. 102 + + PIPRIDÆ, ii. 102 + + _Pipridea_, ii. 98 + + _Piprisoma_, ii. 277 + + _Piprites_, ii. 102, 292 + + _Piramutana_, ii. 442 + + _Piratinga_, ii. 443 + + _Pirinampus_, ii. 443 + + _Pitangus_, ii. 101 + + _Pithecia_, ii. 175 + + _Pithecopsis_, ii. 420 + + _Pithys_, ii. 104 + + _Pitta_, ii. 298 + + Pittas, ii. 297 + + _Pittasoma_, ii. 104 + + Pittidæ, abundant in Borneo, i. 355 + + PITTIDÆ, ii. 297 + + _Pituophis_, ii. 375 + + Pit-vipers, ii. 384 + + _Pitylus_, ii. 99 + + _Pityriasis_, ii. 273 + + _Plagiodontia_, ii. 238 + + _Plagiolophus_, European Eocene, i. 126 + + _Plagiotelium_, ii. 492 + + PLAGIOSTOMATA, ii. 460 + + _Planetes_, ii. 490 + + _Planorbis_, European Secondary, i. 169 + Eocene, i. 169 + ii. 518 + + Plantain-eaters, ii. 307 + + Plant-cutters, ii. 294 + + Plants, distribution of, probably the same fundamentally as that of + animals, ii. 162 + + _Platacanthomys_, ii. 230 + + _Platalea_, ii. 360 + + PLATALEIDÆ, ii. 360 + + _Platanista_, ii. 209 + + _Platemys_, ii. 408 + + _Platurus_, ii. 384 + + _Platycercidæ_, gorgeously-coloured Australian parrots, i. 394 + + PLATYCERCIDÆ, ii. 325 + + _Platycercus_, ii. 325 + + _Platychile_, ii. 487 + + _Platygonus_, N. American Post-Pliocene, i. 130 + ii. 215 + + _Platylophus_, ii. 273 + + _Platymantis_, ii. 419 + + _Platynematichthys_, ii. 442 + + _Platynus_, ii. 489 + + _Platypoecilus_, ii. 450 + + PLATYRHYNCHINÆ, ii. 291 + + _Platyrhynchus_, ii. 101 + + _Platysaurus_, ii. 392 + + _Platysoma_, ii. 489 + + _Platystira_, ii. 271 + + _Platystoma_, ii. 442 + + _Platystomatichthys_, ii. 442 + + _Plecoglossus_, ii. 447 + + _Plecostomus_, ii. 444 + + _Plecotus_, ii. 183 + + PLECTOGNATHI, ii. 457 + + PLECTROMANTIDÆ, ii. 417 + + _Plectromantis_, ii. 417 + + _Plectrophanes_, ii. 286 + + _Plectropterus_, ii. 363 + + _Plectrotrema_, ii. 519 + + _Plecturus_, ii. 374 + + _Plesiarctomys_, European Eocene, i. 126 + ii. 236 + + _Plesiomeryx_, European Eocene, i. 126 + + _Plesiosorex_, European Miocene, i. 118 + + _Plestiodon_, ii. 397 + + _Plethodon_, ii. 413 + + PLEUROBRANCHIDÆ, ii. 530 + + _Pleurodeles_, ii. 413 + + _Pleurodema_, ii. 420 + + _Pleuronectes_, ii. 441 + + PLEURONECTIDÆ, ii. 440 + + _Pleurostrichus_, ii. 392 + + _Pleurotoma_, ii. 508 + + _Pleurotomaria_, ii. 539 + + Pliocene period, Old World, mammalia of, i. 112 + + Pliocene and Post-Pliocene faunas of Europe, general conclusions from, i. + 113 + of N. America, i. 132 + of S. America, i. 146 + of Australia, i. 157 + + _Pliohippus_, N. American Tertiary, i. 135 + + _Pliolophus_, European Eocene, i. 126 + ii. 216 + + _Pliopithecus_, European Miocene, i. 117 + ii. 178 + + PLOCEIDÆ, ii. 286 + + _Plocepasser_, ii. 286 + + _Ploceus_, ii. 286 + + _Plotosus_, ii. 441 + + _Plotus_, ii. 365 + + Plovers, ii. 355 + + _Pluvianellus_, ii. 356 + + _Pluvianus_, ii. 355 + + PLYCTOLOPHIDÆ, ii. 324 + + _Pnoepyga_, ii. 263 + + _Podabrus_, ii. 249 + + _Podager_, ii. 320 + + PODARGIDÆ, ii. 318 + + _Podargus_, ii. 318 + + _Podica_, ii. 352 + + _Podiceps_, ii. 367 + + PODICIPIDÆ, ii. 366 + + _Podilymbus_, ii. 367 + + _Podocnemis_, ii. 408 + + _Poebrotherium_, N. American Tertiary, i. 138 + ii. 217 + + _Poecilia_, ii. 450 + + _Poecilophis_, ii. 383 + + _Poecilothraupis_, ii. 98 + + _Poephagus_, ii. 222 + + _Poephila_, ii. 287 + + _Pogonocichla_, ii. 271 + + POGONORHYNCHINÆ, ii. 306 + + _Pogonorhynchus_, ii. 306 + + _Pogonornis_, ii. 275 + + _Pogonostoma_, ii. 487 + + _Pogonotriccus_, ii. 101 + + _Pohlia_, ii. 418 + + _Poiana_, ii. 195 + + _Polemistria_, ii. 107 + + _Polioaëtus_, ii. 349 + + _Poliococcyx_, ii. 309 + + _Poliohierax_, ii. 349 + + _Poliopsitta_, ii. 328 + + _Polioptila_, ii. 258 + + _Pollanisus_, ii. 481 + + POLYBORINÆ, ii. 347 + + _Polyboroides_, ii. 347 + + _Polyborus_, ii. 347 + + _Polybothris_, ii. 497 + + POLYCENTRIDÆ, ii. 434 + + _Polycesta_, ii. 479 + + POLYDONTIDÆ, ii. 459 + + _Polyhirma_, ii. 491 + + POLYNEMIDÆ, ii. 429 + + _Polyommatus_, ii. 477 + + Polynesian sub-region, description of, i. 442 + birds of, i. 443 + reptiles of, i. 447 + + _Polypedates_, ii. 419 + + POLYPEDATIDÆ, ii. 419 + + _Polypi_, ii. 505 + + _Polyplectron_, ii. 340 + + _Polyprion_, ii. 425 + + POLYPTERIDÆ, ii. 458 + + _Polypterus_, ii. 458 + + _Polytelis_, ii. 325 + + _Pomacanthus_, ii. 427 + + POMACENTRIDÆ, ii. 437 + + _Pomacentrus_, ii. 437 + + _Pomatias_, ii. 521 + + _Pomatorhinus_, ii. 261 + + _Pomotis_, ii. 425 + + _Pompholyx_, ii. 518 + + _Pontia_, ii. 478 + + _Pontoporia_, ii. 209 + + _Pooecetes_, ii. 284 + + _Poodytes_, ii. 258 + + _Poospiza_, ii. 284 + + Porcupines, ii. 240 + + _Poritia_, ii. 477 + + _Porphyrio_, ii. 352 + + Porpoises, ii. 208 + + _Portax_, ii. 223 + + _Porzana_, ii. 352 + + Post-Pliocene, mammalia of Europe, i. 110 + remains imply changes of physical geography in Europe, i. 111 + fauna of N. America, i. 129 + fauna of N. America, remarks on, i. 130 + + _Potamides_, ii. 509 + + _Potamochoerus_, ii. 215 + + _Potamodus_, ii. 258 + + Potamogale of W. Africa, figure of, i. 264 + + _Potamogale_, ii. 189 + + POTAMOGALIDÆ, ii. 189 + + _Potamotherium_, European Miocene, i. 118 + ii. 200 + + Potto of W. Africa, figure of, i. 264 + ii. 176 + + Pouched Rats, ii. 233 + + _Praotherium_, N. American Post-Pliocene, i. 130 + + _Pratincola_, ii. 260 + + Pratincoles, ii. 355 + + _Presbytes_, ii. 171 + + _Prepona_, ii. 474 + + Primates, classification of, i. 86 + probable birthplace of, i. 153 + range of Palæarctic genera of, i. 239 + range of Ethiopian genera of, i. 300 + range of Oriental genera of, i. 371 + range of Australian genera of, i. 475 + + _Primates_, European Pliocene, i. 112 + Miocene of Greece, i. 115 + European Miocene, i. 117 + Indian Miocene, i. 121 + European Eocene, i. 124 + N. American Tertiary, i. 132 + of Brazilian caves, i. 144 + + PRIMATES, distribution of, ii. 170-180 + general remarks on the distribution of, ii. 179 + summary and conclusion, ii. 540 + + Prince's Island, birds of, i. 206 + + _Prinia_, ii. 257 + + _Prion_, ii. 365 + + _Prioneris_, ii. 478 + + PRIONIDÆ, ii. 498 + + _Prionidium_, Oolitic insects, i. 167 + + _Prionirhynchus_, ii. 313 + + _Prioniturus_, ii. 326 + + _Prionochilus_, ii. 277 + + _Prionodontes_, ii. 246 + + _Prionops_, ii. 272 + + _Prionoteles_, ii. 314 + + PRISTIDÆ, ii. 462 + + _Pristimantis_, ii. 419 + + PRISTIOPHORIDÆ, ii. 462 + + _Pristiphoca_, in European Pliocene, i. 112 + ii. 204 + + PRISTIPOMATIDÆ, ii. 426 + + _Pristiurus_, ii. 461 + + _Pristonychus_, ii. 489 + + _Proboscidea_, classification of, i. 90 + range of Ethiopian genus, i. 303 + range of Oriental genus, i. 374 + + _Proboscidea_, European Pliocene, i. 113 + Miocene of Greece, i. 116 + European Miocene, i. 120 + Indian Miocene, i. 122 + N. American Post-Pliocene, i. 130 + N. American Tertiary i. 138 + of Brazilian caves, i. 144 + S. American Pliocene, i. 147 + + PROBOSCIDEA, ii. 227 + summary and conclusion, ii. 542 + + _Procamelus_, N. American Post-Pliocene, i. 130 + N. American Tertiary, i. 138 + ii. 217 + + _Procapra_, ii. 223 + + _Procarduelis_, ii. 283 + + _Procellaria_, ii. 365 + + PROCELLARIIDÆ, ii. 365 + + _Procerus_, ii. 488 + ii. 489 + + _Prochilodus_, ii. 445 + + _Prochilus_, ii. 202 + + _Procnias_, ii. 98 + + _Procris_, ii. 481 + + _Procrustes_, ii. 488 + ii. 489 + + _Proctotretus_, ii. 401 + + _Procyon_, N. American Post-Pliocene, i. 130 + ii. 200 + + _Procyonidæ_, in Brazilian caves, i. 144 + + PROCYONIDÆ, ii. 200 + + PRODUCTIDÆ, ii. 532 + + _Progne_, ii. 281 + + _Promecoderus_, ii. 490 + + _Promephitis_, Miocene of Greece, i. 115 + European Miocene, i. 118 + ii. 200 + + Promerops of East Africa, figure of, i. 261 + + _Promerops_, ii. 276 + + _Pronophilia_, ii. 471 + + _Propalæotherium_, European Eocene, i. 126 + + _Proparus_, ii. 266 + + _Propyrrhula_, ii. 285 + + _Prorastomus_, ii. 211 + + _Proserpina_, ii. 527 + + PROSOBRANCHIATA, ii. 507 + + _Prosthemadera_, ii. 275 + + PROTEIDÆ, ii. 412 + + _Proteles_, ii. 196 + + PROTELIDÆ, ii. 196 + + _Protemnodon_, Australian Post-Tertiary, i. 157 + ii. 251 + + _Proteus_, ii. 412 + + _Prothoe_, ii. 474 + + _Protohippus_, N. American Tertiary, i. 135 + + _Protomeryx_, N. American Tertiary, i. 138 + ii. 217 + + _Protonopsis_, ii. 412 + + _Protonotaria_, ii. 279 + + _Protopithecus_, in Brazilian caves, i. 144 + ii. 178 + + _Protopterus_, ii. 458 + + _Protornis_, European Eocene, i. 162 + + _Prototomus_, N. American Tertiary, i. 134 + + _Prototroctes_, ii. 446 + + _Psalidoprogne_, ii. 281 + + _Psaltria_, ii. 266 + + _Psaltriparus_, ii. 266 + + _Psammodromus_, ii. 391 + + _Psammodynastes_, ii. 377 + + _Psammomys_, ii. 230 + + PSAMMOPHIDÆ, ii. 377 + + _Psammophis_, ii. 377 + + _Psammosaurus_, ii. 389 + + _Psarisomus_, ii. 295 + + _Psephotus_, ii. 325 + + _Pseudacris_, ii. 418 + + _Pseudælurus_, European Miocene, i. 118 + ii. 194 + + _Pseudalopex_, ii. 197 + + _Pseudecheneis_, ii. 444 + + _Pseudechis_, ii. 383 + + _Pseudeutropius_, ii. 442 + + _Pseudis_, ii. 420 + + _Pseudobagrus_, ii. 442 + + _Pseudobias_, ii. 270 + + _Pseudobufo_, ii. 415 + + _Pseudochalceus_, ii. 445 + + _Pseudochelidon_, ii. 312 + + _Pseudocolaptes_, ii. 103 + + _Pseudocordylus_, ii. 392 + + _Pseudocyon_, European Miocene, i. 118 + ii. 198 + + _Pseudodipsas_, ii. 477 + + _Pseudogobio_, ii. 452 + + _Pseudogryphis_, ii. 346 + + _Pseudogyps_, ii. 346 + + _Pseudohage_, ii. 383 + + _Pseudolabuca_, ii. 453 + + _Pseudoleistes_, ii. 282 + + _Pseudomorpha_, ii. 490 + + _Pseudomys_, ii. 230 + + _Pseudonaje_, ii. 383 + + _Pseudoperilampus_, ii. 452 + + PSEUDOPHIDIA, ii. 411 + + _Pseudophryne_, ii. 414 + + _Pseudopontia_, ii. 478 + + _Pseudopus_, ii. 392 + + _Pseudorasbora_, ii. 452 + + _Pseudorca_, ii. 209 + + _Pseudoscops_, ii. 350 + + _Pseudoxiphophorus_, ii. 450 + + _Psilopogon_, ii. 306 + + _Psiloptera_, ii. 497 + + _Psilorhamphus_, ii. 104 + + _Psilorhinus_, ii. 273 + + _Psilorhynchus_, ii. 453 + + Psittaci, classification of, i. 96 + range of Ethiopian genera of, i. 311 + range of Oriental genera of, i. 383 + range of Australian genera of, i. 484 + + PSITTACI, ii. 324 + general remarks on the distribution of, ii. 329 + + PSITTACIDÆ, ii. 328 + + _Psittacula_, ii. 328 + + _Psittacus_, European Miocene, i. 161 + ii. 328 + + _Psittinus_, ii. 326 + + _Psittirostra_, ii. 277 + + _Psittospiza_, ii. 99 + + _Psophia_, ii. 358 + + PSOPHIIDÆ, ii. 358 + + _Psophodes_, ii. 262 + + PSYCHROLUTIDÆ, ii. 436 + + _Pterocles_, European Miocene, i. 161 + ii. 337 + + PTEROCLIDÆ, ii. 337 + + _Pterocyclos_, ii. 520 + + _Pterodon_, European Miocene, i. 125 + + _Pteroglossus_, ii. 307 + + _Pteromys_, ii. 235 + + _Pteromyzon_, ii. 463 + + PTEROMYZONTIDÆ, ii. 463 + + _Pteronura_, ii. 199 + + _Pterophanes_, ii. 108 + + _Pterophyllum_, ii. 439 + + PTEROPIDÆ, ii. 181 + + PTEROPODA, ii. 531 + + _Pteropodocys_, ii. 269 + + PTEROPTOCHIDÆ, ii. 297 + + _Pteroptochus_, ii. 297 + + _Pterorhinus_, ii. 261 + + _Pterosarion_, ii. 452 + + _Pterostichus_, ii. 489 + + _Pteruthius_, ii. 266 + + _Pterygophlichthys_, ii. 444 + + _Ptilocerus_, ii. 186 + + _Ptilochloris_, ii. 102, 293 + + _Ptilogonys_, ii. 280 + + _Ptilonorhynchus_, ii. 275 + + _Ptilopachus_, ii. 338 + + _Ptilopus_, ii. 332 + + _Ptilorhis_, ii. 275 + + _Ptilostomus_, ii. 273 + + _Ptilotis_, ii. 275 + + _Ptosima_, ii. 497 + + _Ptyas_, ii. 375 + + _Ptychobarbus_, ii. 452 + + _Ptyonotus_, ii. 428 + + _Pucrasia_, ii. 340 + + Puff-birds, ii. 310 + + Puffins, ii. 367 + + _Puffinus_, ii. 365 + + PULMONIFERA, ii. 512 + + _Pulsatrix_, ii. 350 + + _Puncturella_, ii. 511 + + _Pupa_, Eocene, i. 169 + + _Pupa vetusta_, Palæozoic, i. 169 + + _Pupa_, ii. 514 + + _Pupina_, ii. 520 + + _Pupinella_, ii. 520 + + _Putorius_, ii. 198 + + PYCNONOTIDÆ, ii. 267 + + _Pycnonotus_, ii. 267 + + _Pycnophrys_, ii. 270 + + _Pyctorhis_, ii. 261 + + _Pygarrhicus_, ii. 103 + + _Pygmornis_, ii. 107 + + _Pygomeles_, ii. 397 + + PYGOPODIDÆ, ii. 395 + + _Pygoptila_, ii. 104 + + _Pygopus_, ii. 395 + + PYRAMIDELLIDÆ, ii. 509 + + _Pyrameis_, ii. 474 + + _Pyranga_, ii. 98 + + _Pyrenestes_, ii. 286 + + _Pyrgisoma_, ii. 284 + + _Pyrgita_, ii. 284 + + _Pyriglena_, ii. 104 + + _Pyrocephalus_, ii. 101, 291 + + _Pyroderus_, ii. 103 + + _Pyromelana_, ii. 286 + + _Pyrophthalma_, ii. 259 + + _Pyrrhocoma_, ii. 99 + + _Pyrrhospiza_, ii. 285 + + _Pyrrhula_, ii. 285 + + _Pyrrhulauda_, ii. 289 + + _Pyrrhulina_, ii. 445 + + _Pyrrhulopsis_, ii. 325 + + _Pyrrhuloxia_, ii. 285 + + _Pyrrhura_, ii. 328 + + _Pytelia_, ii. 287 + + _Python_, ii. 381 + + _Pythonidæ_, European Miocene, i. 165 + + PYTHONIDÆ, ii. 381 + + _Pythonodipsas_, ii. 379 + + _Pythonopsis_, ii. 376 + + Pythons, ii. 381 + + _Pyxicephalus_, ii. 420 + + _Pyxis_, ii. 408 + + Q. + + Quadrumana, fossil, ii. 178 + + Quail-snipes, ii. 354 + + _Querquedula_, ii. 363 + + _Querula_, ii. 102 + + _Quiscalus_, ii. 282 + + R. + + _Rachis_, ii. 524 + + Racoon-dog of N. China, i. 226 + + Racoons, ii. 200 + + _Raia_, ii. 462 + + RAIIDÆ, ii. 462 + + Rails, ii. 351 + + RALLIDÆ, ii. 351 + + _Rallina_, ii. 352 + + _Rallus_, ii. 352 + + _Rana_, European Miocene, i. 166 + ii. 420 + + _Raniceps_, ii. 439 + + RANIDÆ, ii. 420 + + _Ranodon_, ii. 413 + + _Rappia_, ii. 419 + + _Rasbora_, ii. 452 + + _Rasborichthys_, ii. 453 + + Rattle-snakes, ii. 384 + + Rays, ii. 462 + + _Realia_, ii. 521 + + _Rectes_, ii. 272 + + _Recurvirostra_, ii. 353 + + _Regalecus_, ii. 432 + + Region, the best term for the primary zoological divisions, i. 68 + Arctic, why not adopted, i. 69 + Palæarctic, defined, i. 71 + Palæarctic, subdivisions of, i. 71 + Ethiopian, defined, i. 73 + Ethiopian, subdivisions of, i. 73 + Oriental, defined, i. 75 + Oriental, subdivisions of, i. 75 + Australian, defined, i. 77 + Australian, subdivisions of, i. 77 + Neotropical, defined, i. 78 + Neotropical, subdivisions of, i. 78 + Nearctic, defined, i. 79 + Nearctic, distinct from Palæarctic, i. 79 + Nearctic, subdivisions of, i. 80 + + Regions, zoological, i. 50 + zoological, how they should be formed, i. 53 + zoological, may be defined by negative or positive characters, i. 54 + zoological, by what class of animals best determined, i. 56 + for each class of animals, not advisable, i. 58 + zoological, proposed since 1857, i. 58 + zoological, Mr. Sclater's, i. 59 + zoological, discussion of those proposed by various authors, i. 61 + zoological, proportionate richness of, i. 64 + temperate and tropical, well marked in northern hemisphere, i. 65 + and zones, table of, i. 66 + comparative richness of, i. 81 + and sub-regions, table of, i. 81 + order of succession of the, i. 173 + + _Registoma_, ii. 521 + + _Reguloides_, ii. 258 + + _Regulus_, ii. 258 + + _Reinwardtænas_, ii. 333 + + _Reinwardtipicus_, ii. 303 + + _Reithrodon_, ii. 230 + + Representative species, i. 4 + + Reptiles, means of dispersal of, i. 28 + classification of, i. 98 + Miocene of Greece, i. 116 + of Indian Miocene deposits, i. 123 + extinct Tertiary, i. 165 + cosmopolitan groups of, i. 176 + peculiar to Palæarctic region, i. 186 + of Central Europe, i. 195 + of the Mediterranean sub-region, i. 204 + of the Siberian sub-region, i. 220 + of the Manchurian sub-region, i. 227 + table of Palæarctic families of, i. 236 + of the Ethiopian region, i. 254 + of the E. African sub-region, i. 260 + of W. Africa, i. 264 + S. African, i. 268 + of Madagascar, i. 279 + table of Ethiopian families of, i. 297 + of the Oriental region, i. 317 + of the Indian sub-region, i. 326 + of Ceylon, i. 327 + of the Indo-Chinese sub-region, i. 331 + of the Indo-Malay sub-region, i. 340 + table of Oriental families of, i. 368 + of the Australian region, i. 396 + of New Guinea, i. 415 + of the Moluccas, i. 420 + of the Polynesian sub-region, i. 447 + of New Zealand, i. 456 + table of Australian families of, i. 472 + Neotropical, ii. 9 + of S. Temperate America, ii. 40 + of the Mexican sub-region, ii. 54 + of the Antilles, ii. 72 + table of Neotropical families of, ii. 88 + of the Nearctic region, ii. 119 + of California, ii. 128 + of Central N. America, ii. 131 + of Eastern United States, ii. 133 + of Canada, ii. 137 + table of Nearctic families of, ii. 142 + summary and conclusion, ii. 547 + + REPTILIA, ii. 372 + + _Retropinna_, ii. 447 + + Revillagigedo Islands, zoology of, ii. 60 + + _Rhabdornis_, ii. 265 + + _Rhabdosoma_, ii. 374 + + RACHIODONTIDÆ, ii. 377 + + _Rhacophorus_, ii. 419 + + _Rhamnophis_, ii. 376 + + RHAMPHASTIDÆ, ii. 306 + + _Rhamphastos_, ii. 307 + + _Rhamphichthys_, ii. 455 + + _Rhamphocænus_, ii. 104 + + _Rhamphococcyx_, ii. 309 + + _Rhamphocinclus_, ii. 256 + + _Rhamphocoelus_, ii. 98 + + _Rhamphomicron_, ii. 108 + + _Rhaphaulus_, ii. 520 + + _Rhea_, in Brazilian caves, i. 164 + ii. 368 + + _Rhinaster_, ii. 213 + + _Rhinatrema_, ii. 411 + + _Rhinechis_, ii. 376 + + _Rhinelepis_, ii. 444 + + _Rhinichthys_, ii. 452 + + RHINIDÆ, ii. 462 + + RHINOBATIDÆ, ii. 462 + + _Rhinoceros_, Post-Pliocene, i. 112 + European Pliocene, i. 113 + Miocene of Greece, i. 116 + Indian Miocene, i. 122 + fossil remains of, at 16,000 feet elevation in Thibet, i. 122 + fossil in N. China, i. 123 + N. American Tertiary, i. 136 + ii. 213 + + Rhinoceros-hornbill, figure of, i. 339 + + _Rhinocerotidæ_, N. American Tertiary, i. 136 + + RHINOCEROTIDÆ, ii. 213 + + RHINOCHETIDÆ, ii. 359 + + _Rhinochetus_, ii. 359 + + _Rhinococcyx_, ii. 309 + + _Rhinocrypta_, ii. 297 + + _Rhinoderma_, ii. 416 + + RHINODONTIDÆ, ii. 461 + + _Rhinodoras_, ii. 443 + + _Rhinogale_, ii. 195 + + _Rhinoglanis_, ii. 443 + + RHINOLOPHIDÆ, ii. 182 + + _Rhinolophus_, ii. 183 + + _Rhinophis_, ii. 374 + + RHINOPHRYNIDÆ, ii. 414 + + _Rhinophrynus_, ii. 414 + + _Rhinoplax_, ii. 317 + + _Rhinopoma_, ii. 183 + + _Rhinortha_, ii. 309 + + _Rhipidura_, ii. 271 + + _Rhizomys_, ii. 231 + + _Rhodeus_, ii. 452 + + _Rhodinocincla_, ii. 256 + + _Rhodona_, ii. 397 + + _Rhodopis_, ii. 108 + + _Rhodosttehia_, ii. 364 + + _Rhombomys_, ii. 230 + + _Rhombus_, ii. 441 + + _Rhopodytes_, ii. 309 + + _Rhopoterpe_, ii. 104 + + _Rhynchæa_, ii. 353 + + RHYNCHOCEPHALIDÆ, ii. 405 + + RHYNCHOCEPHALINA, ii. 405 + + _Rhynchocyon_, ii. 186 + + _Rhynchocyclus_, ii. 101 + + _Rhynchonella_, ii. 539 + + RHYNCHONELLIDÆ, ii. 532 + + _Rhynchops_, ii. 365 + + _Rhynchopsitta_, ii. 328 + + _Rhynchotus_, ii. 344 + + _Rhytina_, ii. 210, 211 + + _Rhytiodus_, ii. 445 + + _Ricinula_, ii. 507 + + _Rimator_, ii. 263 + + _Rimula_, ii. 511 + + _Rissa_, ii. 364 + + _Rissoa_, ii. 510 + + _Rita_, ii. 442 + + River-hog, of West Africa, figure of, i. 264 + of Madagascar, figure of, i. 278 + + Rivers, limiting the range of mammalia, i. 12 + limiting the range of birds, i. 17 + + River-scene in West Africa, i. 264 + + River-snails, ii. 510 + + _Rivulus_, ii. 450 + + Rock-snakes, ii. 381 + + Rocky mountain sub-region, ii. 129 + mammalia of, ii. 129 + birds of, ii. 130 + reptiles, amphibia, and fishes of, ii. 130 + + Rodentia, classification of, i. 90 + range of Palæarctic genera of, i. 242 + range of Ethiopian genera of, i. 304 + range of Oriental genera of, i. 374 + range of Australian genera of, i. 476 + + _Rodentia_, European Pliocene, i. 113 + Miocene of Greece, i. 116 + European Miocene, i. 120 + European Eocene, i. 126 + N. American Post-Pliocene, i. 130 + N. American Tertiary, i. 139 + of Brazilian caves, i. 144 + S. American Pliocene, i. 147 + of S. American Eocene, i. 148 + + RODENTIA, ii. 229 + + Rodentia, general remarks on the distribution of, ii. 243 + + _Rodentia_, summary and conclusion, ii. 543 + + _Rohteichthys_, ii. 452 + + Rollers, ii. 311 + + _Rollulus_, ii. 339 + + _Romaleosoma_, ii. 474 + + Rose-chafers, ii. 494 + + _Rostrhamus_, ii. 349 + + Rough-tailed burrowing snakes, ii. 374 + + Ruff, figure of, i. 195 + + _Rupicapra_, ii. 224, 225 + + RUPICAPRINÆ, ii. 224 + + _Rupicola_, ii. 102, 293 + + RUPICOLINÆ, ii. 293 + + _Ruticilla_, ii. 259 + + RUTICILLINÆ, ii. 257 + + S. + + _Saccobranchus_, ii. 441 + + _Saccodon_, ii. 445 + + SACCOMYIDÆ, ii. 233 + + _Saccomys_, ii. 233 + + _Saccostomus_, ii. 230 + + _Sagda_, ii. 516 + + Sahara, a debatable land, i. 251 + + Saiga, antelope of W. Tartary, i. 218 + + _Saiga_, ii. 223 + + _Saimiris_, ii. 175 + + Sakis, ii. 175 + + _Salamandra_, ii. 413 + + SALAMANDRIDÆ, ii. 413 + + _Salamandrina_, ii. 413 + + _Salarix_, ii. 448 + + _Salminus_, ii. 445 + + _Salmo_, ii. 447 + + SALMONIDÆ, ii. 447 + + _Salpinctes_, ii. 264 + + _Salpornis_, ii. 264 + + _Saltator_, ii. 99 + + Salvin, Mr., on birds of Galapagos, ii. 30 + + _Sambus_, ii. 496 + + Samoa Islands, birds of, i. 443 + + Sand-grouse, Pallas', of Mongolia, i. 226 + ii. 337 + + Sand-lizards, ii. 398 + + Sandpipers, ii. 353 + + Sandwich Islands, birds of, i. 445 + probable past history of, i. 446 + mountain plants of, i. 446 + depth of ocean around, i. 447 + + _Sanzinia_, ii. 381 + + _Saperda_, ii. 501 + + _Sapphironia_, ii. 109 + + _Sarcodaces_, ii. 445 + + _Sarcophilus_, ii. 249 + + SARCORHAMPHINÆ, ii. 346 + + _Sarcorhamphus_, ii. 346 + + _Sargus_, ii. 427 + + _Sarkidiornis_, ii. 363 + + _Saroglossa_, ii. 288 + + _Sarotherodon_, ii. 438 + + _Sasia_, ii. 303 + + _Satanoperca_, ii. 439 + + SATYRIDÆ, ii. 471 + + _Satyrites Reynesii_, European Cretaceous insect, i. 167 + + _Satyrus_, ii. 471 + + _Saucerottia_, ii. 109 + + Saunders, Mr. Edward, on the Buprestidæ of Japan, i. 229 + + _Saurocetes_, ii. 210 + + _Saurophis_, ii. 392 + + _Saurothera_, ii. 309 + + _Saxicola_, ii. 260 + + _Saxicolinæ_, ii. 257 + + _Sayornis_, ii. 100, 291 + + Scallops, ii. 533 + + _Scalops_, ii. 190 + + _Scapanus_, ii. 190 + + _Scaphiopus_, ii. 417 + + _Scaphirhynchus_, ii. 459 + + _Scaptochirus_, ii. 190 + + _Scaptonyx_, ii. 190 + + _Scaraphites_, ii. 490 + + _Scardafella_, ii. 333 + + _Scarites_, ii. 489 + + _Scelidotherium_, in Brazilian caves, i. 145 + S. American Pliocene, i. 147 + ii. 245 + + _Scelodontis_, ii. 490 + + _Sceloporus_, ii. 401 + + _Scelotes_, ii. 398 + + _Schacra_, ii. 452 + + _Schasicheila_, ii. 522 + + _Schiffornis_, ii. 102 + + _Schilbe_, ii. 442 + + _Schilbichthys_, ii. 442 + + _Schismaderma_, ii. 415 + + _Schistes_, ii. 108 + + _Schistopleurum_, S. American Pliocene, i. 147 + + _Schizodon_, ii. 238 + + _Schizogenius_, ii. 490 + + _Schizopygopsis_, ii. 452 + + _Schizorhina_, ii. 494 + + _Schizorhis_, ii. 307 + + _Schizothorax_, ii. 452 + + _Schoenionta_, ii. 502 + + Schweinfurth, Dr., on natural history of Central Africa, i. 252 + on limits of W. African sub-region, i. 262 (_note_) + + _Sciades_, ii. 443 + + _Sciæna_, ii. 428 + + SCIÆNIDÆ, ii. 428 + + SCINCIDÆ, ii. 396 + + _Scincus_, ii. 397 + + Scinks, ii. 396 + + _Scissirostrum_, ii. 288 + + _Scissor_, ii. 445 + + _Sciuravus_, N. American Eocene, i. 140 + + _Sciuravus_, ii. 236 + + SCIURIDÆ, ii. 234 + + _Sciuropterus_, ii. 235 + + _Sciurus_, European Miocene, i. 120 + European Eocene, i. 126 + ii. 235, 236 + + Sclater, Mr., on zoological regions, i. 59 + why his six regions are adopted, i. 63 + on birds of Sandwich Islands, i. 445 + on systematic position of _Certhidea_, ii. 31 + + Sclater and Salvin, Messrs., on Neotropical sub-regions, ii. 25 + + SCLERODERMI, ii. 457 + + _Sclerognathus_, ii. 451 + + SCLERURINÆ, ii. 295 + + _Sclerurus_, ii. 103 + + _Scolecophagus_, ii. 282 + + SCOLOPACIDÆ, ii. 353 + + _Scolopax_, ii. 353 + + _Scomber_, ii. 429 + + SCOMBRESOCIDÆ, ii. 449 + + _Scombresox_, ii. 449 + + SCOMBRIDÆ, ii. 429 + + SCOPELIDÆ, ii. 446 + + _Scops_, ii. 350 + + _Scopus_, ii. 360 + + _Scortornis_, ii. 320 + + _Scotopelia_, ii. 350 + + _Scotophilus_, ii. 183 + + _Scrapteira_, ii. 391 + + Screamers, ii. 361 + + Scrub-birds, ii. 299 + + SCYLLIDÆ, ii. 461 + + _Scyllium_, ii. 461 + + _Scytale_, ii. 379 + + SCYTALIDÆ, ii. 379 + + _Scytalopus_, ii. 297 + + _Scythrops_, ii. 310 + + Sea, as a barrier to mammalia, i. 13 + + Sea-devils, ii. 463 + + Seals, fossil in European Miocene, i. 118 + of Lake Baikal, i. 218 + ii. 203 + + Sea-pens, ii. 505 + + Sea-snails, ii. 508 + + Sea-snakes, ii. 384 + + _Sebastes_, ii. 428 + + Secondary formations, mammalian remains in, i. 169 + + Secretary bird, of Africa, figure of, i. 261 + ii. 346 + + Seemann, Dr., on protective resemblance of sloths, ii. 24 + + _Seisura_, ii. 270 + + _Selache_, ii. 460 + + _Selasphorus_, ii. 108 + + _Selenidera_, ii. 307 + + _Selenophorus_, ii. 490 + + _Seleucides_, ii. 275 + + _Semioptera_, ii. 275 + + _Semiplotus_, ii. 452 + + SEMNOPITHECIDÆ, ii. 171 + + _Semnopithecus_, European Pliocene, i. 112 + Miocene of Greece, i. 115 + European Miocene, i. 117 + Indian Miocene, i. 121 + ii. 171 + ii. 178 + + Semper, Dr., on Philippine mammalia, i. 345 + + _Senira_, ii. 397 + + SEPIADÆ, ii. 505 + + SEPIDÆ, ii. 398 + + _Seps_, ii. 398 + + _Sepsina_, ii. 398 + + _Sericinus_, ii. 479 + + _Sericornis_, ii. 258 + + _Sericulus_, ii. 275 + + _Serilophus_, ii. 295 + + SERPENTARIIDÆ, ii. 346 + + _Serpentarius_, European Miocene, i. 162 + ii. 346 + + _Serphophaga_, ii. 101 + + _Serranus_, ii. 425 + + _Serrasalmo_, ii. 445 + + _Sesia_, ii. 482 + + _Setophaga_, ii. 279 + + _Setornis_, ii. 267 + + Seychelle Islands, zoology of, i. 281 + amphibia of, i. 281 + + Shad, ii. 454 + + Sharks, ii. 460 + + Sharp, Dr., on Japan beetles, i. 229 + + Sharpe, Mr. R. B., his arrangement of Accipitres, i. 97 + on birds of Cape Verd Islands, i. 215 + on classification of Cuckoos, ii. 309 + + Sheath-bills, ii. 354 + + Sheep, Palæarctic, i. 182 + ii. 221 + + Short-tailed burrowing snakes, ii. 373 + + Shrikes, ii. 272 + + _Sialia_, ii. 260 + + _Siamanga_, ii. 171 + + _Siaphos_, ii. 397 + + Siberia, climate of, i. 217 + + Siberian sub-region, description of, i. 216 + mammalia of, i. 217 + birds of, i. 219 + reptiles and amphibia of, i. 220 + insects of, i. 220 + + _Sibia_, ii. 262 + + _Siderone_, ii. 474 + + _Sieboldia_, ii. 412 + + _Sigmodon_, ii. 230 + + _Silondia_, ii. 442 + + _Silphomorpha_, ii. 490 + + _Silubosaurus_, ii. 397 + + _Siluranodon_, ii. 442 + + _Silurichthys_, ii. 441 + + SILURIDÆ, ii. 441 + + _Silurus_, ii. 441 + + _Silybura_, ii. 374 + + _Simenia_, ii. 197 + + _Simia_, ii. 171 + + SIMIIDÆ, ii. 170 + + _Simocephalus_, ii. 380 + + _Simocyon_, Miocene of Greece, i. 115 + ii. 198 + + _Simorhynchus_, ii. 367 + + _Simotes_, ii. 375 + + _Simpulopsis_, ii. 516 + + _Sinopa_, N. American Tertiary, i. 134 + + _Siphia_, ii. 270 + + _Siphneus_, ii. 230 + + _Siphonopsis_, ii. 411 + + _Siphonorhis_, ii. 320 + + _Siphonostoma_, ii. 457 + + _Siren_, ii. 411 + + Sirenia, classification of, i. 89 + range of Ethiopian genera of, i. 303 + range of Oriental genus, i. 374 + range of Australian genus of, i. 476 + + _Sirenia_, European Pliocene, i. 112 + European Miocene, i. 119 + + SIRENIA, ii. 210 + + SIRENIDÆ, ii. 411 + + SIRENOIDEI, ii. 458 + + _Sirystes_, ii. 101 + + _Sisor_, ii. 444 + + _Sitana_, ii. 402 + + _Sitta_, ii. 265 + + _Sittasomus_, ii. 103 + + _Sittella_, ii. 265 + + SITTIDÆ, ii. 265 + + _Siurus_, ii. 279 + + _Siva_, ii. 266 + + _Sivatherium_, Indian Miocene, i. 122 + ii. 226 + + Siwalik Hills, Miocene deposits of, i. 121 + + _Skenea_, ii. 510 + + Sloths, ii. 244 + + Slugs, ii. 517 + + _Smaragdochrysis_, ii. 109 + + _Smerinthus_, ii. 483 + + _Smiliogaster_, ii. 453 + + _Sminthus_, ii. 230 + + Smith, Mr. Frederick, on Hymenoptera of Japan, i. 230 + + _Smithornis_, ii. 270 + + _Smutsia_, ii. 245 + + Snails, ii. 512 + + Snake, at great elevation in Himalayas, i. 220 + + Snakes, classification of, i. 99 + Eocene, i. 165 + large proportion of venomous species in Australia, i. 396 + of New Zealand, i. 457 + distribution and lines of migration of, ii. 547 + + Snipes, ii. 353 + + Society Islands, birds of, i. 443 + + Socorro, zoology of, ii. 60 + + Soft-tortoises, ii. 409 + + _Solarium_, ii. 510 + + _Solea_, ii. 441 + + SOLENIDÆ, ii. 536 + + _Solenodon_, ii. 188 + + SOLENOSTOMIDÆ, ii. 456 + + Solitaire, ii. 334 + + _Somateria_, ii. 364 + + _Soricictis_, European Miocene, i. 118 + ii. 196 + + _Soricidæ_, European Miocene, i. 118 + + SORICIDÆ, ii. 191 + + _Soridia_, ii. 397 + + _Sorubim_, ii. 442 + + _Sotalia_, ii. 209 + + South African sub-region, description of, i. 266 + mammalia of, i. 267 + birds of, i. 267 + reptiles of, i. 268 + amphibia of, i. 268 + fresh-water fish of, i. 268 + butterflies of, i. 268 + coleoptera of, i. 268 + summary of its zoology, i. 269 + + South America, fossil fauna of, i. 143 + Pliocene deposits of, i. 146 + supposed land connection with Australia, i. 398 + + South America and Africa, parallelism of their past zoological history, + ii. 83 + + South Australia, peculiar birds of, i. 441 + + SPALACIDÆ, ii. 231 + + _Spalacomys_, ii. 230 + + _Spalacopus_, ii. 238 + + _Spalax_, ii. 231 + + _Sparganura_, ii. 108 + + SPARIDÆ, ii. 426 + + _Spatula_, ii. 364 + + Species, representative, i. 4 + + _Spelerpes_, ii. 413 + + _Speothos_, in Brazilian caves, i. 145 + + _Spermestes_, ii. 287 + + _Spermophila_, ii. 285 + + _Spermophilus_, European Miocene, i. 120 + ii. 235, 236 + + _Spermospiza_, ii. 286 + + Sperm Whales, ii. 207 + + _Sphærocephalus_, ii. 209 + + _Sphærodactylus_, ii. 400 + + _Sphæroderus_, ii. 490 + + _Sphallomorpha_, ii. 490 + + _Sphecotheres_, ii. 268 + + _Sphenæacus_, ii. 258 + + SPHENISCIDÆ, ii. 366 + + _Spheniscus_, ii. 366 + + _Sphenocephalus_, ii. 398 + + _Sphenodon_, in Brazilian caves, i. 115 + ii. 245 + + _Sphenognathus_, ii. 493 + + _Sphenoproctus_, ii. 107 + + _Sphenops_, ii. 398 + + _Sphenostoma_, ii. 266 + + _Sphenura_, ii. 258 + + SPHINGIDÆ, ii. 482 + + Sphingidea, distribution of, ii. 483 + + SPHINGINA, ii. 481 + + _Sphingnotus_, ii. 501 + + _Sphinx_, in European Oolite, i. 167 + ii. 482 + + Sphinx Moths, ii. 482 + + _Sphyrapicus_, ii. 303 + + SPHYRENIDÆ, ii. 429 + + Spider monkeys, ii. 174 + + _Spilornis_, ii. 348 + + _Spilotes_, ii. 376 + + SPINACIDÆ, ii. 461 + + _Spindalis_, ii. 98, 284 + + _Spiraxis_, ii. 515 + + SPIRIFERIDÆ, ii. 532 + + SPIRULIDÆ, ii. 505 + + _Spizaëtus_, ii. 348 + + _Spizella_, ii. 284 + + _Spiziapteryx_, ii. 349 + + _Spiziastur_, ii. 348 + + _Spodiornis_, ii. 285 + + _Sponsor_, ii. 497 + + Spoonbills, ii. 360 + + _Sporadinus_, ii. 109 + + _Sporopipes_, ii. 286 + + Sprat, ii. 454 + + _Spreo_, ii. 288 + + _Squalodon_, ii. 210 + + _Squaliobarbus_, ii. 452 + + SQUAMIPENNES, ii. 427 + + _Squatarola_, ii. 356 + + Squirrel monkeys, ii. 175 + + Squirrels, ii. 234 + + St. Helena, zoological features of, i. 269 + coleoptera of, i. 270 + landshells of, i. 271 + + St. Thomas's Island, birds of, i. 266 + + _Stachyris_, ii. 261 + + _Stactolæma_, ii. 306 + + Stag-beetles, ii. 492 + + _Stalagmosoma_, ii. 495 + + Starlings, ii. 287 + + _Starnoenas_, ii. 33 + + Stations, definition of, i. 4 + + _Staurotypus_, ii. 408 + + _Steatomys_, ii. 230 + + _Steatornis_, ii. 319 + + STEATORNITHIDÆ, ii. 319 + + _Steganura_, ii. 108 + + _Stegnolæma_, ii. 343 + + _Stegophilus_, ii. 444 + + _Stelgidopteryx_, ii. 281 + + _Stellio_, ii. 402 + + _Stellula_, ii. 108 + + _Steneofiber_, European Miocene, i. 120 + ii. 234 + + _Steno_, ii. 209 + + _Stenodactylus_, ii. 400 + + _Stenogyra_, ii. 515 + + _Stenopsis_, ii. 320 + + _Stenopus_, ii. 516 + + _Stenorhina_, ii. 375 + + _Stenorhynchus_, ii. 204 + ii. 421 + + _Stephanophorus_, ii. 98 + + _Stercorarius_, ii. 304 + + _Sterna_, ii. 364 + + _Sternarchus_, ii. 455 + + _Sternocera_, ii. 496 + + _Sternoclyta_, ii. 107 + + STERNOPTYCHIDÆ, ii. 446 + + _Sternopygus_, ii. 455 + + _Sternotheres_, ii. 408 + + _Steropus_, ii. 489 + + _Stesilea_, ii. 501 + + _Stethodesma_, ii. 495 + + _Sthenurus_, Australian Post-Tertiary, i. 157 + ii. 251 + + _Stichæus_, ii. 431 + + Sticklebacks, ii. 424 + + _Stigmatura_, ii. 101 + + _Stigmodera_, ii. 496 + + STOMIATIDÆ, ii. 447 + + Storks, ii. 360 + + _Stabomantis_, ii. 419 + + Straits of Magellan, mammalia of, ii. 37 + birds of, ii. 39 + + _Strepera_, ii. 273 + + _Strepsilas_, ii. 356 + + _Streptaulus_, ii. 520 + + _Streptaxis_, ii. 515 + + _Streptocerus_, ii. 493 + + _Streptocitta_, ii. 274 + + _Streptophorus_, ii. 374 + + STRIGIDÆ, ii. 350 + + _Stringops_, ii. 329 + + STRINGOPIDÆ, ii. 329 + + _Strix_, European Miocene, i. 162 + ii. 350 + + STROMBIDÆ, ii. 507 + + _Struthio_, ii. 368 + + Struthiones, arrangement of, i. 98 + range of Ethiopian genera of, i. 313 + range of Australian genera of, i. 485 + + STRUTHIONES, ii. 368 + general remarks on the distribution of, ii. 370 + + STRUTHIONIDÆ, ii. 368 + + Struthious birds, probable origin of, i. 287 + + Sturgeons, ii. 459 + + _Sturnella_, ii. 282 + + _Sturnia_, ii. 287 + + STURNIDÆ, ii. 287 + + _Sturnopastor_, ii. 287 + + _Sturnus_, ii. 287 + + STYGIIDÆ, ii. 482 + + _Stygogenes_, ii. 444 + + _Stylinodontia_, N. American Eocene, i. 139 + + _Stylinodontidæ_, N. American Eocene, i. 139 + + _Styporhynchus_, ii. 376 + + _Sublegatus_, ii. 101 + + Sub-regions, on what principle formed, i. 180 + Palæarctic, i. 191 + Ethiopian, i. 258 + Oriental, i. 321 + Australian, i. 408 + Neotropical, ii. 21 + Nearctic, ii. 125 + + _Succinea_, ii. 515 + + Sugar-birds, ii. 278 + + _Suidæ_, European Miocene, i. 119 + + SUIDÆ, ii. 214 + + Sula Islands, fauna of, i. 433 + + _Sula_, ii. 365 + + Summary of relations of regions, ii. 155 + + Sun-birds, ii. 276 + + Sun-bitterns, ii. 358 + + _Suricata_, ii. 195 + + _Surnia_, ii. 350 + + _Surniculus_, ii. 310 + + _Sus_, European Pliocene, i. 113 + Miocene of Greece, i. 116 + European Miocene, i. 119 + Indian Miocene, i. 122 + ii. 215 + + _Suthora_, ii. 262 + + _Suya_, ii. 258 + + Swallows, ii. 281 + + Swallow-shrikes, ii. 288 + + Swifts, ii. 320 + + Swine, ii. 214 + + Swinhoe, Mr., on zoology of Formosa and Hainan, i. 332 + + _Sycalis_, ii. 284 + + _Sylvia_, ii. 259 + + _Sylvietta_, ii. 264 + + SYLVIIDÆ, ii. 256 + + SYLVIINÆ, ii. 257 + + _Sylviorthorhynchus_, ii. 103 + + _Sylviparus_, ii. 266 + + _Syma_, ii. 316 + + _Symborodon_, N. American Tertiary, i. 137 + + SYMBRANCHIDÆ, ii. 455 + + _Symbranchus_, ii. 455 + + _Symmachia_, ii. 476 + + _Symmorphus_, ii. 269 + + _Symphædra_, ii. 474 + + _Symphysodon_, ii. 439 + + _Symplectes_, ii. 286 + + SYNALLAXINÆ, ii. 295 + + _Synallaxis_, ii. 103 + + _Synaphodus_, European Miocene, i. 119 + + _Synaptura_, ii. 441 + + _Synchloe_, ii. 474 + + _Syndesus_, ii. 493 + + _Synemon_, ii. 481 + + _Syngnathus_, ii. 457 + + SYNGNATHIDÆ, ii. 457 + + _Synodontis_, ii. 443 + + _Synoplotherium_, N. American Tertiary, i. 134 + + _Syntomis_, ii. 481 + + _Syrnium_, ii. 350 + + _Syrrhaptes_, ii. 337 + + _Sysopygis_, ii. 101 + + T. + + Tables of distribution of families and genera explained, i. 177 + + _Taccocoua_, ii. 309 + + _Tachydromus_, ii. 391 + + _Tachyphonus_, ii. 99 + + _Tachyris_, ii. 478 + + _Tachytriorchis_, ii. 348 + + _Tadorna_, ii. 363 + + _Tæniogale_, ii. 195 + + _Tænioptera_, ii. 100, 291 + + TÆNIOPTERINÆ, ii. 291 + + _Tæniura_, ii. 463 + + _Talegallus_, ii. 342 + + _Talpa_, European Miocene, i. 117 + ii. 190 + + TALPIDÆ, ii. 190 + + _Tamandua_, ii. 247 + + _Tamias_, ii. 235, 236 + + _Tanæcia_, ii. 474 + + Tanagers, ii. 283 + + _Tanagra_, ii. 98 + + _Tanagrella_, ii. 98 + + TANAGRIDÆ, ii. 283 + + _Tantalus_, ii. 361 + + _Tanygnathus_, ii. 326 + + _Tanysiptera_, ii. 316 + + _Taoniscus_, ii. 344 + + _Taphozous_, ii. 183 + + _Tapir_, fossil in N. China, i. 123 + + Tapir, Malayan figure of, i. 337 + + _Tapiridæ_, European Eocene, i. 125 + + TAPIRIDÆ, ii. 212 + + Tapirs, birthplace and migrations of, i. 154 + ii. 212 + + _Tapirus_, European Pliocene, i. 113 + Indian Miocene, i. 122 + in Brazilian caves, i. 144 + + _Tarandus_, ii. 219 + + _Tarentola_, ii. 400 + + Tarsier, Malayan, figure of, i. 337 + + _Tarsiger_, ii. 259 + + TARSIIDÆ, ii. 177 + + _Tarsipes_, ii. 252 + + _Tarsius_, ii. 177 + + Tasmania, comparative zoological poverty of, i. 441 + + _Tatare_, ii. 258 + + _Tatusia_, ii. 246 + + _Taxidea_, ii. 199 + + _Taxila_, ii. 475 + + _Taxodon_, European Miocene, i. 118 + ii. 200 + + _Taygetis_, ii. 471 + + _Tchitrea_, ii. 271 + + TECTONARCHINÆ, ii. 275 + + Teguexius, ii. 390 + + TEIDÆ, ii. 390 + + _Teinopalpus_, ii. 479 + + _Teira_, ii. 391 + + _Teius_, ii. 390 + + _Teleopis_, ii. 375 + + TELEOSTEI, ii. 424 + + _Telephonus_, ii. 272 + + _Tellia_, ii. 450 + + TELLINIDÆ, ii. 506 + + _Telmatobius_, N. American Cretaceous, i. 164 + ii. 417 + + _Telmatolestes_, N. American Tertiary, i. 133 + + _Temnotrogon_, ii. 314 + + _Temnurus_, ii. 273 + + _Tephrocorys_, ii. 289 + + _Tephrodornis_, ii. 272 + + _Teracolus_, ii. 478 + + _Terebratula_, ii. 539 + + TEREBRATULIDÆ, ii. 532 + + _Terekia_, ii. 353 + + _Terenura_, ii. 104 + + _Teretristis_, ii. 279 + + _Terias_, ii. 478 + + _Terinos_, ii. 474 + + Terns, ii. 364 + + _Terrapene_, ii. 408 + + Terrestrial Molluscs, ii. 512 + + Terrestrial Mollusca, summary and conclusion, ii. 551 + lines of migration of, ii. 552 + + _Tesia_, ii. 263 + + _Testacella_, ii. 516 + ii. 517 + + TESTUDINIDÆ, ii. 407 + + _Testudo_, Miocene of Greece, i. 116 + Indian Miocene, i. 123 + great antiquity of the genus, i. 289 + + _Testudo_, ii. 408 + + _Tethionea_, ii. 501 + + TETRABRANCHIATA, ii. 506 + + _Tetracha_, ii. 486, 487 + + _Tetrachus_, European Miocene, i. 117 + + _Tetraceros_, ii. 224 + + _Tetracus_, ii. 188 + + _Tetradactylus_, ii. 397 + + _Tetragonoderus_, ii. 490 + + _Tetragonops_, ii. 306 + + _Tetragonopterus_, ii. 445 + + _Tetragonosoma, ii. 380_ + + _Tetranematichthys_, ii. 443 + + _Tetrao albus_, in Italian caverns, i. 161 + + _Tetrao_, ii. 339 + + _Tetraogallus_, ii. 339 + + TETRAONIDÆ, ii. 338 + + _Tetraophasis_, ii. 340 + + _Tetrodon_, ii. 457 + + TEUTHIDÆ, ii. 505 + + TEUTHIDIDÆ, ii. 433 + + _Textor_, ii. 286 + + _Thais_, ii. 479 + + _Thalassarctos_, ii. 201 + + _Thalassictis_, Miocene of Greece, i. 115 + European Miocene, i. 118 + ii. 195 + ii. 197 + + _Thalassornis_, ii. 364 + + _Thaleichthys_, ii. 447 + + _Thalurania_, ii. 107 + + _Thamnistes_, ii. 104 + + _Thamnobia_, ii. 260 + + _Thamnodyastes_, ii. 379 + + _Thamnomanes_, ii. 104 + + THAMNOPHILINÆ, ii. 297 + + _Thamnophilus_, ii. 104 + + _Thaumalea_, ii. 340 + + _Thaumantis_, ii. 472 + + _Thaumastura_, ii. 108 + + _Thaumatias_, ii. 109 + + _Thecla_, ii. 477 + + _Theloderma_, ii. 419 + + _Theope_, ii. 476 + + _Theorema_, ii. 477 + + _Theraps_, ii. 438 + + _Therates_, ii. 486 + + _Theridomys_, European Miocene, i. 126 + European Eocene, i. 126 + S. American Eocene, i. 148 + ii. 239 + + _Theropithecus_, ii. 173 + + _Thestias_, ii. 478 + + _Thestor_, ii. 477 + + _Thetia_, ii. 391 + + THINOCORIDÆ, ii. 354 + + _Thinocorus_, ii. 354 + + _Thinohyus_, N. American Tertiary, i. 137 + ii. 215 + + _Thinolestes_, N. American Tertiary, i. 133 + + _Thinornis_, ii. 356 + + _Thomomys_, ii. 233 + + _Thous_, ii. 197 + + _Thrasaëtus_, ii. 348 + + _Threnetes_, ii. 107 + + _Thripadectes_, ii. 103 + + _Thripophaga_, ii. 103 + + _Thryophilus_, ii. 263 + + _Thryothorus_, ii. 263 + + Thrushes, ii. 255 + + _Thyca_, ii. 471 + + _Thylacinus_, Australian Post-Tertiary, i. 157 + ii. 249 + + _Thylacoleo_, Australian Post-Tertiary, i. 157 + ii. 252 + + _Thymallus_, ii. 447 + + _Thynnichthys_, ii. 452 + + _Thynnus_, ii. 429 + + _Thyreopterus_, ii. 491 + + _Thyrus_, ii. 398 + + _Tiaris_, ii. 284 + ii. 402 + + _Tichodroma_, ii. 264 + + _Tiga_, ii. 303 + + Tiger-beetles, ii. 486 + + _Tigrisoma_, ii. 359 + + _Tijuca_, ii. 102 + + _Tillodontia_, N. American Eocene, i. 139 + + _Tillotheridæ_, N. American Eocene, i. 139 + + _Tillotherium_, N. American Eocene, i. 139 + + _Tilmatura_, ii. 108 + + _Timalia_, ii. 261 + + TIMALIIDÆ, ii. 260 + + _Timetes_, ii. 474 + + Timor, physical features of, i. 389 + group, mammalia of, i. 422 + birds of, i. 422 + origin of fauna of, i. 424 + insects of, i. 426 + + TINAMIDÆ, ii. 343 + + TINAMINÆ, ii. 344 + + Tinamous, ii. 343 + + TINAMOTINÆ, ii. 344 + + _Tinamotis_, ii. 344 + + _Tinamus_, ii. 344 + + _Tinca_, ii. 452 + + _Tinoceras_, N. American Eocene, i. 139 + + _Titanomys_, European Miocene, i. 121 + ii. 242 + + _Titanotherium_, N. American Tertiary, i. 137 + + Tits, ii. 265 + + _Tityra_, ii. 102 + + TITYRINÆ, ii. 293 + + _Tmesisternus_, ii. 501 + + Toads, ii. 415 + + _Tockus_, ii. 317 + + TODIDÆ, ii. 313 + + Todies, ii. 313 + + _Todirhamphus_, ii. 316 + + _Todirostrum_, ii. 101 + + _Todopsis_, ii. 271 + + _Todus_, ii. 313 + + _Tolypeutes_, ii. 246 + + _Tomarctos_, N. American Tertiary, i. 135 + + _Tomistoma_, ii. 405 + + _Tomodon_, ii. 175 + + Tonga Islands, birds of, i. 443 + + _Topaza_, ii. 107 + + TORNATELLIDÆ, ii. 530 + + TORPEDINIDÆ, ii. 462 + + Tortoises, classification of, i. 100 + of Mascarene Islands and Galapagos, i. 289 + ii. 407 + + TORTRICIDÆ, ii. 373 + + _Tortrix_, ii. 373 + + _Totanus_, ii. 353 + + Toucans, ii. 306 + + Touraco of W. Africa, figure of, i. 264 + + _Toxodon_, S. American Pliocene, i. 137 + + _Toxodontidæ_, S. American Pliocene, i. 147 + + _Toxotus_, ii. 502 + + _Trachelyopterus_, ii. 443 + + TRACHINIDÆ, ii. 428 + + _Trachinus_, ii. 428 + + _Trachurus_, ii. 429 + + _Trachycephalus_, ii. 401 + ii. 418 + + _Trachydosaurus_, ii. 397 + + _Trachyphonus_, ii. 306 + + TRACHYPTERIDÆ, ii. 432 + + _Trachytherium_, European Miocene, i. 119 + + TRAGELAPHINÆ, ii. 223 + + _Tragelaphus_, ii. 223 + + _Tragocerus_, Miocene of Greece, i. 116 + European Miocene, i. 120 + + Tragopan, Himalayan, figure of, i. 331 + + _Tragops_, ii. 379 + + TRAGULIDÆ, ii. 218 + + _Tragulus_, ii. 218 + + _Trapelus_, ii. 402 + + _Trechus_, ii. 489 + + Tree-crows, ii. 273 + + Tree-kangaroo, figure of, i. 415 + + Tree-shrew of Borneo, figure of, i. 337 + + Tree-snakes, ii. 378 + + _Tremarctos_, ii. 202 + + _Treron_, ii. 332 + + Tres Marias, zoology of, ii. 59 + + _Tribolonotus_, ii. 397 + + _Triboniophorus_, ii. 517 + + _Tribonyx_, ii. 352 + + _Trichastoma_, ii. 261 + + TRICHECHIDÆ, ii. 203 + + _Trichechus_, N. American Post-Pliocene, i. 130 + ii. 203 + + TRICHIURIDÆ, ii. 429 + + _Trichixos_, ii. 262 + + TRICHOGLOSSIDÆ, ii. 327 + + Trichoglossidæ, birds specially adapted to Australia, i. 393 + + _Trichoglossus_, ii. 327 + + _Tricholæma_, ii. 306 + + _Trichomycterus_, ii. 444 + + _Trycondyla_, ii. 486 + + _Trichonis_, ii. 477 + + TRICHONOTIDÆ, ii. 435 + + _Trichothraupis_, ii. 99 + + _Trichotropis_, ii. 507 + + _Triclaria_, ii. 328 + + TRIDACNIDÆ, ii. 535 + + _Trigla_, ii. 428 + + TRIGLIDÆ, ii. 427 + + _Trigona_, ii. 536 + + TRIGONIADÆ, ii. 534 + + _Trigonoptera_, ii. 501 + + _Trimeresurus_, ii. 385 + + _Tringa_, ii. 353 + + _Tringoides_, ii. 353 + + TRIONYCHIDÆ, ii. 409 + + _Trionyx_, Indian Miocene, i. 123 + Miocene and Eocene, i. 165 + ii. 409 + + _Triprion_, ii. 418 + + _Triptorhinus_, ii. 297 + + Tristan d'Acunha, zoology of, i. 271 + + Tristram, Canon, summary of the birds of Palestine, i. 203 + on the arrangement of the Sylviidæ, ii. 257 + + _Triton_, ii. 413 + + TRITONIADÆ, ii. 530 + + _Trochalopteron_, ii. 261 + + _Trochatella_, ii. 522 + + TROCHILIDÆ, ii. 321 + + _Trochilus_, ii. 108 + + _Trochus_, ii. 510 + + _Troglodytes_, ii. 170 + ii. 263 + + TROGLODYTIDÆ, ii. 263 + + Trogon, European Miocene, i. 161 + ii. 314 + + _Trogon_, ii. 314 + + TROGONIDÆ, ii. 314 + + TROGONOPHIDÆ, ii. 388 + + Trogonophis, ii. 388 + + _Trogontherium_, Post-Pliocene of Europe, i. 111 + ii. 234 + + _Tropidechis_, ii. 383 + + _Tropidococcyx_, ii. 379 + + _Tropidodipsas_, ii. 379 + + _Tropidolepis_, ii. 401 + + _Tropidolepisma_, ii. 397 + + _Tropidonotus_, ii. 375 + + _Tropidophorus_, ii. 397 + + _Tropidopterus_, ii. 490 + + _Tropidorhynchus_, ii. 276 + + _Trucifelis_, N. American Post-Pliocene, i. 129 + + _Trugon_, ii. 333 + + Trumpeters, ii. 358 + + _Truncatella_, ii. 519 + + _Trgyon_, ii. 463 + + TRYGONIDÆ, ii. 463 + + Tuatara, ii. 405 + + _Tudora_, ii. 521 + + Tundras of Siberia, greatest extent of, i. 216 + + _Tupaia_, ii. 186 + + TUPAIIDÆ, ii. 186 + + _Tupaiidæ_, European Miocene, i. 118 + + _Turacoena_, ii. 333 + + Turacos, ii. 307 + + _Turacus_, ii. 307 + + TURBINIDÆ, ii. 510 + + TURDIDÆ, ii. 255 + + _Turdinus_, ii. 262 + + _Turdus_, ii. 256 + + _Turnagra_, ii. 262 + + Turner, Mr., on classification of Edentata, i. 90 + + TURNICIDÆ, ii. 341 + + _Turnix_, ii. 341 + + TURRITELLIDÆ, ii. 509 + + _Tursio_, ii. 209 + + Turtles, ii. 409 + + _Turtur_, ii. 333 + + _Tylas_, ii. 267 + + _Tylodon_, European Eocene, i. 125 + ii. 196 + + _Tylognathus_, ii. 451 + + _Tylotriton_, ii. 413 + + _Typhlina_, ii. 372 + + _Typhline_, ii. 372 + + _Typhlocalamus_, ii. 374 + + TYPHLOPIDÆ, ii. 372 + + _Typhlops_, ii. 372 + + _Typhloscincus_, ii. 399 + + _Typotherium_, S. American Pliocene, i. 147 + + TYRANNIDÆ, ii. 290 + + TYRANNINÆ, ii. 291 + + _Tyranniscus_, ii. 101 + + _Tyrannulus_, ii. 101 + + _Tyrannus_, ii. 102, 291 + + Tyrant-Shrikes, ii. 290 + + U. + + _Uaru_, ii. 439 + + _Uintacyon_, N. American Tertiary, i. 134 + + _Uintatherium_, N. American Eocene, i. 139 + + _Uintornis_, N. American Eocene, i. 163 + + _Uma_, ii. 401 + + UMBRIDÆ, ii. 449 + + _Umbrina_, ii. 428 + + _Ungalia_, ii. 381 + + Ungulata, classification of, i. 89 + antiquity of, i. 154 + of the Palæarctic region, i. 182 + range of Palæarctic genera of, i. 241 + range of Ethiopian genera of, i. 303 + range of Oriental genera of, i. 374 + range of Australian genera of, i. 476 + + _Ungulata_, European Pliocene, i. 112 + Miocene of Greece, i. 115 + European Miocene, i. 119 + Indian Miocene, i. 121 + European Eocene, i. 125 + N. American Post-Pliocene, i. 130 + N. American Tertiary, i. 135 + of Brazilian caves, i. 144 + S. American Pliocene, i. 146 + + UNGULATA, ii. 211 + general remarks on the distribution of, ii. 226 + summary and conclusion, ii. 542 + + _Unio_, European Secondary, i. 169 + ii. 534 + + UNIONIDÆ, ii. 534 + + _Upucerthia_, ii. 103 + + UPUPIDÆ, ii. 317 + + _Uragus_, ii. 285 + + Urania of Madagascar, i. 282 + + _Urania_, ii. 482 + + URANIIDÆ, ii. 482 + + _Uria_, ii. 367 + + _Uroaëtus_, ii. 348 + + _Urocissa_, ii. 273 + + _Urochroa_, ii. 107 + + _Urochroma_, ii. 328 + + _Urocyon_, ii. 197 + + URODELA, ii. 411 + + _Urogalba_, ii. 311 + + _Urolestes_, ii. 272 + + _Uromastix_, ii. 402 + + UROPELTIDÆ, ii. 373 + + _Uuropeltis_, ii. 374 + + _Uropsila_, ii. 264 + + _Uropsilus_, ii. 190 + + _Uropsophorus_, ii. 385 + + _Urospatha_, ii. 313 + + _Urospizias_, ii. 348 + + _Urosticte_, ii. 108 + + _Urotrichus_, ii. 190 + + _Urotriorchis_, ii. 347 + + _Ursidæ_, N. American Tertiary, i. 135 + in Brazilian caves, i. 144 + + URSIDÆ, ii. 201 + + _Ursitaxus_, Indian Miocene, i. 121 + ii. 200 + + _Ursus_, Post-Pliocene, i. 112 + Indian Miocene, i. 121 + ii. 201 + + _Urubutinga_, ii. 348 + + _Urva_, ii. 195 + + _Uta_, ii. 401 + + _Utica_, ii. 477 + + V. + + _Vaginulus_, ii. 518 + + _Valgus_, ii. 495 + + _Valvata_, ii. 510 + + Vanga of Madagascar, figure of, i. 278 + + _Vanga_, ii. 272 + + _Vandellia_, ii. 444 + + _Vanellus_, ii. 356 + + _Vanessa_, ii. 474 + + VARANIDÆ, ii. 389 + + _Varanus_, Miocene of Greece, i. 116 + Indian Miocene, i. 123 + + VENERIDÆ, ii. 536 + + _Venilia_, ii. 303 + + _Vermicella_, ii. 383 + + _Verreauxia_, ii. 303 + + Vertebrata, summary of Palæarctic, i. 186 + summary of Ethiopian, i. 255 + summary of Oriental, i. 318 + summary of Australian, i. 397 + summary of Neotropical, ii. 13 + summary of Nearctic, ii. 120 + + _Vespertilio_, European Eocene, i. 125 + ii. 183 + + VESPERTILIONIDÆ, ii. 183 + + _Vidua_, ii. 286 + + _Vipera_, ii. 385 + + VIPERIDÆ, ii. 385 + + _Viperus_, European Miocene, i. 165 + + Vipers, ii. 385 + + _Vireo_, ii. 280 + + VIREONIDÆ, ii. 279 + + _Vireolanius_, ii. 280 + + _Vireosylvia_, ii. 280 + + Viscacha, ii. 237 + + _Vitrina_, ii. 516 + + _Viverra_, European Pliocene, i. 112 + European Miocene, i. 118 + ii. 195 + + _Viverricula_, ii. 195 + + _Viverridæ_, European Miocene, i. 118 + European Eocene, i. 125 + + VIVERRIDÆ, ii. 194 + + _Vivia_, ii. 303 + + _Volatinia_, ii. 284 + + _Voluta_, ii. 508 + + Volutes, ii. 508 + + VOLUTIDÆ, ii. 508 + + _Volvocivora_, ii. 269 + + _Vulpes_, ii. 197 + + _Vultur_, ii. 346 + + VULTURIDÆ, ii. 345 + + VULTURINÆ, ii. 346 + + W. + + Wagtails, ii. 290 + + Walden, Viscount, on birds of Philippine islands, i. 346 + on birds of Celebes, i. 428 + on arrangement of the Timaliidæ, ii. 261 + + _Wallago_, ii. 441 + + Wall-lizards, ii. 399 + + Walrus, ii. 203 + + Wart-snakes, ii. 382 + + _Washakius_, N. American Tertiary, i. 134 + + Waterhouse, Mr. G. R., on classification of rodentia, i. 90 + on classification of marsupials, i. 91 + + Water-lizards, ii. 389 + + Weaver-finches, ii. 286 + + West African sub-region, description of, i. 262 + mammalia of, i. 262 + birds of, i. 262 + Oriental or Malayan element in, i. 263 + river scene with characteristic animals, i. 264 + reptiles of, i. 264 + amphibia of, i. 264 + Oriental and Neotropical relations of, i. 265 + insects of, i. 265 + land-shells of, i. 265 + islands of, i. 265 + + West Australia, peculiar birds of, i. 441 + + Whelks, ii. 507 + + Whip-snakes, ii. 379 + + Whydah finch of W. Africa, figure of, i. 264 + + Wing-shells, ii. 507 + ii. 533 + + Wollaston, Mr. T. V., on the Coleoptera of the Atlantic Islands, i. 209 + on the wings of the Madeiran beetles, i. 211 + on the origin of the insect fauna of the Atlantic Islands, i. 214 + on the Coleoptera of the Cape Verd Islands, i. 215 + on the beetles of St. Helena, i. 270 + + Wombats, ii. 253 + + Woodpeckers, ii. 302 + + Wood-warblers, ii. 278 + + Woolly monkeys, ii. 174 + + Wrens, ii. 263 + + Wrynecks, ii. 304 + + X. + + _Xanthocephalus_, ii. 282 + + _Xantholæma_, ii. 306 + + _Xanthomelus_, ii. 275 + + _Xanthopygia_, ii. 270 + + _Xanthosomus_, ii. 282 + + _Xanthotis_, ii. 275 + + _Xema_, ii. 364 + + _Xenelaphis_, ii. 376 + + _Xenica_, ii. 471 + + _Xenicus_, ii. 265 + + _Xenochrophys_, ii. 375 + + _Xenocypris_, ii. 452 + + _Xenodermus_, ii. 376 + + _Xenodon_, ii. 375 + + XENOPELTIDÆ, ii. 373 + + _Xenopeltis_, ii. 373 + + _Xenophrys_, ii. 421 + + _Xenopipo_, ii. 102 + + _Xenops_, ii. 103 + + _Xenorhina_, ii. 415 + + XENORHINIDÆ, ii. 415 + + _Xenospingus_, ii. 284 + + _Xenurelaps_, ii. 383 + + _Xenurus_, in Brazilian caves, i. 145 + ii. 246 + + _Xiphias_, ii. 430 + + _Xiphidiopicus_, ii. 303 + + XIPHIIDÆ, ii. 430 + + _Xiphius_, ii. 208 + + _Xiphocolaptes_, ii. 103 + + _Xiphodontidæ_, European Miocene, i. 119 + + _Xipholena_, ii. 102 + + _Xiphorhampus_, ii. 445 + + _Xiphorhynchus_, ii. 103 + + _Xiphostoma_, ii. 445 + + _Xystrocera_, ii. 501 + + Y. + + _Ypthima_, ii. 471 + + _Yuhina_, ii. 266 + + YUNGIDÆ, ii. 304 + + _Yungipicus_, ii. 303 + + _Yunx_, ii. 304 + + Z. + + _Zabrus_, ii. 489 + + _Zalophus_, ii. 203 + + _Zamenis_, ii. 375 + + _Zanclostomus_, ii. 309 + + _Zaocys_, ii. 375 + + Zebras, ii. 211 + + _Zegris_, ii. 478 + + _Zemeros_, ii. 475 + + _Zenaida_, ii. 333 + + _Zenaidura_, ii. 332 + + _Zephyrus_, ii. 477 + + _Zeuglodon_, ii. 210 + + _Zeuglodontidæ_, N. American Tertiary, i. 140 + ii. 210 + + _Zeus_, ii. 429 + + _Zeuxidia_, ii. 472 + + _Zoarces_, ii. 431 + + _Zonites_, ii. 516 + + _Zonites priscus_, Palæozoic, i. 169 + + _Zonotrichia_, ii. 284 + + ZONURIDÆ, ii. 391 + + _Zonurus_, ii. 392 + + Zoological characteristics of Palæarctic region, i. 181 + Ethiopian region, i. 252 + Oriental region, i. 315 + Australian region, i. 390 + of Neotropical region, ii. 5 + of Nearctic region, ii. 115 + + Zoological regions, discussion on, i. 50 + their origin and relations, ii. 155-161 + + _Zoothera_, ii. 256 + + _Zootoca_, ii. 391 + + _Zosterops_, ii. 277 + + _Zygæna_, ii. 481 + + ZYGÆNIDÆ, ii. 481 + + _Zygnopsis_, ii. 398 + + +THE END. + + +LONDON: R. CLAY, SONS, AND TAYLOR, PRINTERS. + + + +Notes + +[1] Messrs. Sclater and Salvin, and Professor Newton, divide the + Neotropical Region into six sub-regions, of which our "Brazilian + sub-region" comprises three--the "Brazilian," the "Amazonian," and the + "Columbian;" but, after due consideration, it does not seem advisable + to adopt this subdivision in a general work which treats of all the + classes of terrestrial animals. (See p. 27.) + +[2] Mr. Salvin, who has critically examined the ornithological fauna of + these islands, has kindly corrected my MS. List of the Birds, his + valuable paper in the _Transactions of the Zoological Society_ not + having been published in time for me to make use of it. + +[3] This name will be used for the whole island of St. Domingo, as being + both shorter and more euphonious, and avoiding all confusion with + Dominica, one of the Lesser Antilles. It is also better known than + "Hispaniola," which is perhaps the most correct name. + +[4] Myospalax has hitherto formed part of the next family, Spalacidæ; but a + recent examination of its anatomy by M. Milne-Edwards shows that it + belongs to the Muridæ, and comes near Arvicola. + +[5] The species of the genera _Phylloscopus_ and _Hypolais_ are so mixed up + in the _Hand List_, that Mr. Tristram has furnished me with the + following enumeration of the species which in his view properly belong + to them, by the numbers in that work:-- + + _Phylloscopus._ + 3032 + 3033 + 3048 = 3038 + 3039 + 3063 = 3047 = 3054 = 3061 + 3048 + 3049 + 3050 + 3051 + 3052 + 3053 + 3056 = 3081 + 3057 + 3059 + 3060 + + _Hypolais._ + 3026 + 3028 + 3029 + 3054 = 3031 = 3036 + 3042 + 3043 + 304 + 3062 = 3047 + 3046 = 2932 + 3035 + 2976 + +[6] The term "Malaya" is used here to include the Malay Peninsula, Sumatra, + Borneo, and Java, a district to which many species and genera are + confined. "Malay Archipelago" will be used to include both Indo-Malaya + and Austro-Malaya. + + + + + + + +End of the Project Gutenberg EBook of The Geographical Distribution of +Animals, Volume II, by Alfred Russel Wallace + +*** END OF THE PROJECT GUTENBERG EBOOK 56507 *** |
