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| author | nfenwick <nfenwick@pglaf.org> | 2025-03-08 19:12:04 -0800 |
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| committer | nfenwick <nfenwick@pglaf.org> | 2025-03-08 19:12:04 -0800 |
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diff --git a/40880-0.txt b/40880-0.txt new file mode 100644 index 0000000..65b24a7 --- /dev/null +++ b/40880-0.txt @@ -0,0 +1,1592 @@ +*** START OF THE PROJECT GUTENBERG EBOOK 40880 *** + + [Illustration: + + _Frontispiece._ + + Canon Horsley in his study examining a rare variety of whelk + (var. _Babylonica_) from a stall in the Walworth Road. It is now + in the South Kensington Museum.] + + + + + OUR BRITISH SNAILS + + BY THE + REV. CANON J. W. HORSLEY + + + AUTHOR OF "SOME FOLK-LORE AND LEGENDS OF BIRDS," ETC. + + LONDON + SOCIETY FOR PROMOTING CHRISTIAN KNOWLEDGE + NORTHUMBERLAND AVENUE, W.C. + 43, QUEEN VICTORIA STREET, E.C. + BRIGHTON: 129, NORTH STREET + 1915 + + + PRINTED BY + WILLIAM CLOWES AND SONS, LIMITED, + LONDON AND BECCLES. + + + + +LIST OF ILLUSTRATIONS + + + PAGE + + Canon Horsley in his study examining a rare variety of whelk + (var. _Babylonica_) from a stall in the Walworth Road. + _Frontispiece._ + + _H. pomatia_, half natural size 11 + + Dextral _H. aspersa_ and _H. pomatia_ 13 + + Love-darts of _H. pomatia_, much magnified 15 + + _H. nemoralis_ at rest on hawthorn 17 + + Names of parts of shell and of body. _Unio_, _Limnæa_, + _Vivipara_, and _Arion_ 22 + + Body of snail and of slug 23 + + Three specimens of _Arion ater_, showing tentacles, + breathing orifice, and slime gland 31 + + _Testacella haliotidea_ 35 + + _Helicella virgata_ at rest on thistle, natural size 45 + + Some of our smaller shells 47 + + _Paludina contecta_ (two) and _Limnæa stagnalis_ on + water-weeds 57 + + _Neritina_ and _Ancylus_ 59 + + Freshwater mussel breathing and eating 61 + + + + +OUR BRITISH SNAILS + + +It has been said that a child's education should begin thirty years +before its birth, since what he is, or becomes, or does, depends +largely upon what his parents were, and not solely on what he learns +at home or in school, or from his companions and surroundings. + +But the principle of what is called "atavism" shows us that the +appearance, tastes, and character of a child's grandparents may +reappear, even more than those of his parents; and that, therefore, +his education begins sixty years before his birth. + +My education, viewing me as a naturalist, began even earlier than +that, for nearly all my ancestors of whom I know anything more than +their names and abiding place were botanists or horticulturists, and I +cannot recollect the time when I was not an observer of nature and a +collector of the common objects of the field, the ditch, the seashore, +the wood, and the cliff. My father died before I was four, and I have +never had any remembrance of his words or looks, yet I remember his +cutting down a tree in the shrubbery of his Kentish vicarage garden +which forked curiously from the ground, and also of finding that +handsome fungus which is scarlet flecked with white. This shows that +the observation of the marvels and beauties of God's Green Bible, or +Book of Nature, began early in me. The habits of observation, of +comparison, and of method, are those which all naturalists and +collectors must have; habits which are of great value in other ways as +well. Firstly, one must have the seeing eye, and train it to notice +what many people do not. (Get and read the old book, much read when I +was young, called "Eyes and no Eyes.") Secondly, one must learn to +observe the difference (sometimes very small, although important) +between one object and others of the same family. Every one knows a +wild rose by sight; but nearly every one would be surprised to hear +that botanists make out twenty kinds of English wild roses, to say +nothing of varieties and hybrids. In all departments of natural +history a magnifying glass, for the dissection of inward parts, is +necessary in many cases to separate two kinds which look alike. And, +thirdly, if you want to make a collection, whether of dried plants, of +insects, of shells, or of anything else, you must cultivate ways of +order and method and neatness in the arrangement of your collection. +And then your increased powers of observation, of comparison, and of +method will stand you, and others, in good stead in higher matters of +thought and action, and the virtues of Prudence, Justice, Temperance, +and Fortitude will all increase in you as you learn more about what is +in man, what man should be, and how men should be treated. Let us take +Fortitude for example. I have known boys who collected one kind of +thing eagerly for a while, but soon got tired of it, and generally had +little power of "sticking" to anything. On the other hand, I was once +admiring the magnificent collection of shells owned by a middle-aged +doctor, and asked him, "When did you begin to collect?" "When I was +seven," was his answer. I should expect to find more Fortitude in that +doctor's character than in that of a boy who collected "all things in +turn and nothing long." + +Yet I myself was middle-aged before I felt disgusted with myself, when +gazing on a lad's collection of British land shells, that I should so +long have been groping in hedges and ditches, and yet never have +noticed the variety and the beauty of members of the snail family. +(That lad, by the bye, is now a Professor in an American University, +and a great authority on shells and other matters.) Since then I have +gathered a complete collection of the British land and fresh-water +shells, and a very large and valuable one of the _Helicidæ_--_i.e._ +the family to which the common or garden snail belongs--of every +country in the world; and have been President of the Conchological +Society of Great Britain and Ireland. + +I am now, therefore, writing about our British land shells, "slugs and +snails" in common speech, with the hope that it may add a new interest +to the country walks of lads and lasses. + +I could show you a wall-case I made for a school. It contains +specimens of all the British land shells with the exception of the +slugs, which (with the exception of one of which I shall speak in its +place) have no external or covering shell, although a small sort of +shell, or at any rate some chalky grains, is found inside most of +them. You would see that some are as small as a pin's head although +full grown, and they would require a magnifying glass to distinguish +one from the other. The largest is _Helix pomatia_ (figured on pp. 11 +and 12), which often goes by the name of "the edible snail." All +snails are edible and nutritious; but this is the one cultivated in +snail farms and sold as food abroad. Sometimes it is called "the Roman +snail," from an idea, probably wrong, that it was introduced by +Cæsar's soldiers, although as a matter of fact it is unknown in South +Italy. Sometimes also it is called "the apple snail," partly because +it is as large as a middle-sized apple, and partly because people +thought the name _pomatia_ came from the Latin _pomum_, "an apple," +whereas it really comes from the Greek [Greek: pôma]. This word means +a lid, or closing arrangement, and this mollusc makes a hard front +door for itself when it hibernates, _i.e._ suspends active life and +buries itself in the winter. + +[Illustration: _H. pomatia_, half natural size.] + +It is much to be regretted that in most cases scientific names fail to +give much information to the young student, and in some cases they +give none at all. The first or generic name is supposed to be formed +from Greek, the second, or specific, from the Latin, but there are +some hybrids and many mere "nonsense names" to puzzle beginners. Thus +the slug Limax gets its name from _limus_, "mud"; but a scientist, who +ought to have known better, when wanting a name for another kind of +slug, transposed the initial letters and made Milax! Vitrina is a +sensible and descriptive name, the Latin for glassy, given to a shell +like thin glass; but the Greek Arion recalls either a certain musician +or a certain swift steed, neither of whom naturally suggests a slug. +For Balea at least four derivations have been suggested--none of them +probable. Two facts concerning the life or appearance of a mollusc we +should learn from its two names, but this is not the case with +_Agriolimax agrestis_, which is by interpretation "the field slug +inhabiting fields." Nor are we helped by the specific name _virgata_ +or striped when so many land shells are striped or banded, and still +less by _terrestris_ for one land shell when all land shells are +terrestrial. + +You would note, however, in this wall-case that the species are not +many (a good many of the specimens are varieties, not separate +species), and that, therefore, one can collect with the hope of +speedily forming a complete collection without that inevitable absence +of finality found when one collects postage stamps, or, still more, +picture postcards, of which one might secure thousands, only to find +that fresh thousands were brought out next year. Here, however, is no +impossible ideal of perfection. There are but eighty-two land and +forty-five freshwater shells in Britain. + +[Illustration: Dextral _H. aspersa_ and _H. pomatia_. The right-hand +shell at the bottom shows the winter epiphragm of _H. pomatia_.] + +Let us imagine we are starting for an afternoon snailing near London. +Which way? To Oxshott? To Caterham? To the latter for choice, since it +is on the chalk, whereas the former is on the sand. Snails require +lime to make shells, and only on chalk or limestone will you find an +abundance. Here, too, as at Box Hill, we shall find the big _Helix +pomatia_, only found in a few English counties, and very local there. +If we were very fortunate, we might find a sinistral, or "left-handed" +specimen. In the case of the _pomatia_ on the right hand there is +shown the thick epiphragm which the mantle secretes before the mollusc +hibernates. It hardens on exposure to the air like plaster-of-paris; +but is not a true operculum, for that is a constant possession of the +shells which have it. Opercula are mainly found in marine or +fluviatile shells, and may be either horny (like the winkle) or stony. +Amongst our British land shells _Cyclostoma elegans_ and _Acicula +lineata_ alone have true opercula, though others form some thin +epiphragm for the exclusion of cold air and enemies when they +hibernate. + +Most shells grow to the right, and a freak which does the contrary is +so rare that of the millions of the common _H. virgata_ that I have +seen and handled, only one delighted me with its left-handedness. If +it is early summer (nearly all snails hide, burrow, and sleep during +the winter), look about on the grass for some half-chalky, half-stony +shields, which are the winter front doors of _H. pomatia_, now +discarded; while sharper eyes might even descry the flinty little +darts with which they have been love-making. The illustration on p. 15 +shows three of these darts, much magnified. Only the most highly +developed Helices possess these courting weapons, not unlike bayonets +in form, sometimes rounded and smooth, and sometimes with two or even +four lateral blades, so that the section of the dart of _H. pomatia_ +is in the form of a Greek cross. Not many British shells have these +darts, but in one case their study is useful, since _H. nemoralis_ and +_H. hortensis_, though so closely allied that early conchologists +considered them to be of the same species, have darts remarkably +distinct one from the other, so that they become a court of final +appeal if from outward appearance it is difficult to distinguish, say, +a white-mouthed _nemoralis_ from a dark-mouthed _hortensis_. + +[Illustration: Love-darts of _H. pomatia_, much magnified.] + +Whenever you see a stone, a brick, a branch of dead wood, or even an +old boot or a piece of newspaper in the hedge or on the grass, turn it +over, for many of the smaller shells are thus found, and "leave no +stone unturned" is eminently a motto for the conchologist. Some of the +shells will be tiny, and must be studied under a magnifying +glass--which all naturalists should always have in their pockets--or +even under a microscope at home, in order to discover, not only their +beauty of marking or sculpture, but even to what species they belong. + +When you see a man sweeping herbage with a net, or beating hedges and +shrubs over an inverted umbrella, he is probably an entomologist in +search of caterpillars or beetles; but the same methods will often +reward the snail-hunter. + +Especially in the hedges will you find the two allied species _Helix_ +(_Cepea_) _nemoralis_ and _hortensis_, to which the attention of +beginners should first be directed, inasmuch as they are so common, so +beautiful, and so varying both in colour and the number of the +chocolate bands they usually bear. See the illustration of some of +these at rest on hawthorn, p. 17. Canary-yellow, flesh-colour, +chocolate, and almost white, are the prevailing ground-colours. Five +is the normal number of bands on the largest or body-whorl, although +sometimes all run into one, and often one, some, or all are wanting. +Where only one band is found--throughout the Helicidæ--it is usually +that on the periphery or middle of the whorl, and a shell in which +this band is wanting, while others are found, is a rarity. People are +usually astonished, on seeing a good series of the colour and +variations of these two shells, how they vie with those of warmer +regions. + +[Illustration: _H. nemoralis_ at rest on hawthorn.] + +Next search trunks of trees, and especially the smooth boles of the +beeches. The rough bark of the elm or oak is not congenial to slugs or +snails. Where trees are moss-covered at their foot, or walls at their +top, many of the smaller shells may be expected; while handfuls of +dead leaves may be shaken over something white, or taken home in a +large bag to be treated there. Hurdles leaning against a hedge are +often found to bear a good crop of snails. Damp places must be sought +in dry weather; but a rainy day, that troubles some kinds of +naturalists, sends the conchologist forth rejoicing, especially if a +warm evening follows a wet day. A night search with a lantern will +often be profitable. Where they will be undisturbed, traps may be set, +such as flat pieces of wood (the older the better), or cardboard, +lying on the grass; while most of those species that belong to the +group which seems to prefer the sun, _e.g._ _H. itala_, _virgata_, +etc., are fond of a newspaper for food rather than for shelter. + +During the hibernating season, which extends from November to April, +we turn rather to ditches than to hedges, and, armed with a perforated +scoop at the end of a long stick, we dredge among the water-weeds, or +sift, like gold-washers, the sand or mud in ditches, ponds, and +backwaters of rivers. Here we are introduced to the great bivalve +family which is unknown on land, and our trophies range from the +freshwater mussels, as large as our hand, to others hardly larger than +a pin's head. These must be sought at the bottom; but on the weeds, or +on the bottom, will be found not a few species of gasteropods or +univalves, some of which we may have noticed in a freshwater aquarium. +These, of course, are closely connected with the land shells, which +the bivalves are not. They can be brought home alive in a tin box with +a little moss, whereas for the land shells a calico bag with a little +foliage therein is best. In both cases some small glass tubes with +corks should be brought in a tin box in order to keep safely and +separately the tinier kinds. You can often discover what small shells +inhabit a particular ditch or pond by noticing the cases of +caddis-worms, some of which are formed almost entirely of shells +instead of vegetable fragments. + +Using the precious gift of observation, we have found our shells; at +home we exercise the other gifts of comparison and order, in the +preparation and arrangement of our collection. A dash of quite boiling +water kills instantaneously any molluscs whose shells we want to +preserve, and then the body is extracted after the fashion observed +with regard to winkles at tea. Be careful to get out all the body of +the animal, and then it is well to wash out any slime or particles by +directing a fine but strong jet of cold water into the shell. This can +be done by holding your thumb nearly over the mouth of a watertap, +while the shell is held in the left hand. Only adult shells should +usually be taken, and those which are weather-worn or bleached should +be neglected. In most the lip, or opening, of the shell will be hard +if adult, and membranous if young; but experience alone will enable +you to discriminate, especially where the young of one species is like +the adult of another. + +Get into the way of carrying a note-book with you to record not only +what shells, or varieties of a species, are found in any particular +spot, but also anything you observe as to the habits or peculiarities +of the objects of your search. Notes as to protective colouring or +mimicry; the influences of a wet or a dry season on the relative +thickness of shells; the difference in size caused by abundance or +scarcity of diet; what plants are preferred and what avoided as food +by particular helices,--are some of the points of interest, apart from +the earliest and latest dates at which certain species are abroad and +active. + +If you possess, or borrow, a microscope, many new wonders and fresh +lines of inquiry will open out. I know one professor who devotes +himself to the study of the teeth of molluscs. A snail may possess +over twenty thousand tiny flinty teeth set on a ribbon so as to make a +mowing-machine for the vegetable matter on which it feeds. With its +aid also you might study the life-history of a mollusc from the egg +onwards, and be able to determine by minute anatomical points whether +two molluscs were of the same species or not--a matter in which the +shape or appearance of the shell is not always a safe guide. + +Here, then, is a new hobby for some of my readers, or, at any rate, a +fresh source of interest when they are in the country. If any +collector lives near you, I am sure he or she would be delighted to +have your company during an expedition, and you would learn more by +sight and hearing than by reading. If, however, you must fall back +upon a book, get _The Collector's Manual_ by L. E. Adams, published by +Taylor Bros., Leeds. This is invaluable both to the beginner and to +the owner of a good collection. + +From this I borrow by leave the plate on p. 22, which will enable the +beginner to understand from the first certain names of parts of the +shell or the body of the bivalve, univalve, or slug which otherwise +might not be clear. The "muscular scars" are indents in the shell +which mark where the muscles were fixed whose function was to bring +close together the two valves of the shell when it has need to exclude +air or enemies. + +[Illustration: Names of parts of shell and of body. _Unio_, _Limnæa_, +_Vivipara_, and _Arion_.] + +The figures of the snail and the slug below are introduced to give +further knowledge of the soft parts. B is the body, soft and with a +surface generally wrinkled or covered with small tubercles. F is the +foot or muscular pad which forms the foot by the wavelike contractions +of which it moves. H is the head, bearing the tentacles T_{1} and +T_{2}, of which the upper pair have the eyes, E. The mantle, M, makes +the shell by secreting lime, etc. In it is the breathing orifice, BO, +obvious in the slug, but in the snail nearly hidden by the shell. L in +the snail is the spiral part, the liver, and it occupies a large part +of the shell. + +[Illustration: Body of snail and of slug.] + +Without going into details of classification and anatomy, which would +only deter or puzzle a beginner, let me take two typical molluscs of +those which we shall find in England, the common garden snail _Helix +aspersa_, and a freshwater mussel, _Unio margaritifer_, and see where +they come in the scale of creation and what are their powers and +peculiarities. + +Molluscs (_mollis esca_, soft food--boneless creatures) are below the +aristocracy of the vertebrates or backboned creatures, and so they +come just below the Fishes, but above the Insects. They are divided +into those possessing a head and those possessing no head (although +with some sort of a brain or organ of sense), the snail being of the +former class and the mussel of the latter. The former are univalves +and the latter bivalves having two shells for protection. The latter +also are restricted to life in water, whereas the former are found +both on land and in water, _e.g._ the snail and the whelk, although +for ages probably no molluscs were air-breathing land dwellers. In the +class of Cephala, to which our snail belongs, there is the sub-class +of Gasteropoda, or stomach-footed, because on the ventral side of the +body a sole-like disc or foot exists, by the wave-like expansions and +contractions of which the animal progresses. + +In this sub-class there is a division according to their having or not +having an operculum, or means of closing and protecting the orifice of +the shell. Most gasteropods which live in water have this; most which +live on land (only two exceptions in British molluscs) have not. Here +again we must trace our snail down to the sub-order of Pulmonata, or +lung or air-sac breathers as distinct from its sisters which inhabit +water and breathe by gills. This sub-order is again divided into +various families, Arion, Limax, Testacella, Vitrina, Zonites, Helix, +etc., and Helix again is divided into various genera, of which Helix +is one, and even this is subdivided into sub-genera, Patula, Punctum, +Acanthinula, Vallonia, Chilotrema, Gonostoma, Pomatia, Tachea, etc., +and to the sub-genus Pomatia our garden snail as well as the "Roman +snail" belongs. Looking backwards we, therefore, place our friend as +the species _aspersa_, of the sub-genus _Pomatia_, of the genus +_Helix_, of the family _Helicidæ_, of the sub-order _Pulmonata_, of +the order _Inoperculata_, of the sub-class _Gasteropoda_, of the class +_Cephala_, of the sub-kingdom of _Mollusca_, of the kingdom +_Invertebrata_ or backboneless animals. + +It belongs by origin not to the earliest form of snail, but to the +most highly organized group in the world, especially characteristic of +the European region, and possessing in their superiority the power to +colonize and dispossess the original native snails of other lands. The +shell is globular in form with five whorls (the Greek word "helix" +means a coil), each usually marked with five bands of pigment. It is +mainly a vegetarian, and by habit a lover of the twilight and of +moisture. With the exception of _H. pomatia_ it is the largest of our +native shells, and is too common to satisfy gardeners. A powerful +animal of its kind, it can travel a yard in twelve minutes, or at the +rate of a mile in a fortnight, can bear or draw on level ground a +weight fifty times its own. It breathes about four times a minute, and +its heart-beat varies from sixty to eighty per minute according to +temperature, or its activity. It takes its winter rest in clusters, +closing its mouth with a membranous film, while if the cold increases +it shrinks farther into its shell and makes more epiphragms or film +curtains to keep out the cold. Not only on the Continent, but in +several parts of England, notably about Bath and Bristol, it is +sought, sold, and used for food, and in Belgium it is said to be +preferred to the larger and more firm-fleshed _H. pomatia_. The eggs, +from forty to a hundred, are laid in the earth and hatched in from a +fortnight to a month, according to the weather. I had observed them as +a boy, and used to call tapioca pudding "snail's egg pudding." In the +year of their hatching they attain but half their proper size, but +after hibernation they eat voraciously and grow rapidly, so as to +attain full size in a little more than a year. Most die in their +second hibernation (if not destroyed by their many enemies, gardeners, +collectors, rats, rabbits, ducks, thrushes, and beetles); but when +kept and protected for observation they have achieved the great age of +even ten years. + +They have a great power of "homing" like pigeons, however far (for +them) is their journey after favourite food. The slime-marked journeys +or feeding tracks of this species (and still more of slugs) afford +matter of great interest. As to sight the two eyes are the dark specks +on the tip of the upper pair of "horns," but the range of vision is +very short indeed, and the difference between light and approaching +darkness is all that some seem able to perceive. The organs of hearing +are two small sacs filled with fluid in which are some calcareous +grains. They hear little which is audible to human ears, and if not +altogether deaf they are dumb as far as we can hear. The power of +taste they possess, as is shown by the preference of some foods to +others. The sense of touch is acute and resides in all parts of the +soft and moist external skin, and especially in the upper tentacles or +horns in the _Helicidæ_. Jaws they have with which to seize and to +bite off food, and in _H. aspersa_ and others these bear teeth, but +the chief work is done by a sort of toothed tongue, the radula, which +rasps off particles of food with a side to side motion of the head as +the animal advances. Our _aspersa_ has 12,615 teeth on this ribbon, +contained in 145 transverse rows. The organs of digestion are complex +and practically much the same as our own. Little vegetation would be +left in nature had not, on the one hand, snails been kept down by many +enemies as well as by their need of hibernation and their short life; +while on the other by numerous devices in the course of ages many +plants have protected themselves against the moving machine of a +snail's mouth. Cultivated plants, which generally lose their natural +protections, have to be guarded by human guards or gardeners. Some +plants defend themselves by prickles or hairs, some by hardening +themselves with lime or flint, some by bitter or acrid juices. A heart +of two chambers, veins, arteries, and blood our snail possesses, and, +like man, the old snail has a slower pulse than the young one, and in +both exercise increases the pulse rate and also warmth. Breathing is +accomplished by a single chamber or air-cell, but also through the +skin. As in the case of plants, some kinds are male and female +separately, and as some have both powers and products in the same +plant, so also is it with mollusca. _H. aspersa_ and most Gasteropoda +are of the latter kind. + +Having now taken _H. aspersa_ as the representative of our univalves, +let us take the "Pearl Mussel"--_Unio margaritifer_--as that of our +bivalves, all of which live in the water, whereas of univalves some +are "land snails" and some "water snails." It would say of itself, "I +am a species of the genus Unio (_unio_, a pearl), which belongs to the +family Unionidæ, which belongs to the sub-order Isomya (_i.e._ having +muscles of equal power to close the two valves of the shell), which +belongs to the order Lamellibranchiata (_i.e._ having gills arranged +in leaf-like fashion), which belongs to the sub-class Pelecypoda +(_i.e._ having a foot somewhat of an axe-shape), which belongs to the +class Acephala (headless), which is the second of the two chief +classes into which Mollusca are divided. + +"I differ from the Gasteropoda (whether they be terrestrial or +aquatic) in that I and my near relations are exclusively aquatic and +of a sedentary life, which makes the protection of two encompassing +shells necessary. These shells are secreted by my mantle lobes, and +are united by a ligament which tends to make the valves 'gape' for +water and food and by two contracting muscles which close them in +danger. I have a degenerate brain and no eyes. My mouth has neither +jaw nor teeth, but possesses nervous lips covered with cilia, the +vibration of which carries food-laden water to my mouth. My foot, when +protruded, is seen as a large muscular appendage, and, by alternately +expanding and contracting, it enables me to burrow or plough through +mud or even sand, and so disturb the minute organisms on which I +feed. I can thus travel fifteen feet a day, or about a mile in a year. + +"I have no eyes, but distinguish well between light and shade by means +of the surface of my body when exposed. I breathe, that is, get oxygen +from the water, by means of gill-plates. As regards other internal +organs, I differ not much from _H. aspersa_, but I am either male or +female. Outside I am black and uncomely; but within I am pearly-white, +and but for my power of forming pearls round an irritating grain of +sand the civilization of England would have come to pass later than it +did, for it was the report of my pearls which brought Cæsar to +Britain." + + * * * * * + +Now let us enumerate the species of land and freshwater shells to be +found, (all but two) in England, and most of them in Ireland or +Scotland. + +_Arion ater_ is a large (3 to 5 inches) and common slug, usually black +(whence its name _ater_), but also red, brown, or white. In some +varieties the foot-fringe is orange. When irritated it contracts into +a hemispherical lump. A few chalky granules under the mantle are the +representatives of a shell. See the illustration of three specimens on +p. 31. That hole in the mantle is the breathing orifice, and its +forward position is a characteristic of the group _Arion_. The body of +slugs is kept moist by a constant exuding of slime from a gland in the +tail. + +[Illustration: Three specimens of _Arion ater_, showing tentacles, +breathing orifice, and slime gland.] + +_Arion subfuscus_ (_i.e._ somewhat tawny). Smaller (2 to 3 inches) +than _A. ater_, grey or yellowish, with usually a dark stripe on each +side. Foot-sole white, and its fringe white with dark cross streaks. +Never very abundant. + +_Arion minimus._--The smallest Arion: not an inch long. Grey or +yellowish. Feeds on fungi. Body wrinkled with microscopic spikes. +Common. The _young_ of _A. ater_ might be mistaken for it. + +_Arion hortensis._--Grey with purple side bands. Foot-sole yellow. 1 +to 1-1/2 inch in length. Generally found in gardens, as its name +indicates. + +_Arion circumscriptus._--Very common in fields. A dark band down the +back, foot-sole white. Very "sluggish." + +_Geomalacus maculosus_ (_i.e._ the spotted earth-mollusc).--Only found +in south-west Ireland. Probably a relic of the prehistoric time when +Ireland was joined to Portugal and Spain. Has a solid chalky shell +beneath the shield. Blackish with oval yellow spots. Feeds on lichens. + +_Amalia gagates_ (_gagates_ is Greek for "jet").--Dark lead colour. +Foot-sole white. Length 2-1/2 inches. Local, and mainly near sea. + +_Amalia Sowerbyi._--Brown, speckled with black. Foot-sole yellowish. +Length 2-1/2 inches. Local. Shell often very thick. + +_Limax maximus._--Length 4 to 6 inches. Grey with two dark lateral +bands. Often found in cellars. + +_Limax cinereo-niger._--Ashy-black. Very like _L. maximus_, but with a +sharp keel, and the sole paler in the middle than at the sides. Less +nocturnal and less fond of houses; chiefly found in forests on hills. +Local, and not common. + +_Limax flavus._--Yellow, with a faint dark network of markings. +Tentacles blue. Sole cream. Length 4 inches. Only found in cellars and +near houses. + +_Limax marginatus._--Semi-transparent. Grey, with two dark bands on +each side. Foot-sole with a dark line down the middle. Shell solid, +often a cube. Length 3 inches. Fond of tree climbing. + +_Limax tenellus._--Yellow. Tentacles black. Mucus yellow. Found in +woods. Lives on fungi. Rare. + +_Agriolimax agrestis._--The common field slug. Swarms everywhere. Its +milk-white slime is characteristic. Very variable in colour and +markings. + +_Agriolimax lævis._--Slender. All chocolate brown. Length 3/4 inch. +Shell may be seen through the mantle. Active. Our smallest slug. +Usually found near ditches. + + * * * * * + +It may be useful here to give the chief differences between the genera +Arion, Amalia, Limax, and Agriolimax. The shield in the first two is +granulated, in the other concentrically striated. The breathing +orifice in Arion is in front of the centre of the mantle margin; in +the others behind. The shell is distinctly formed in all but Arion, +in which it is absent or represented by a few granules. Arion has no +dorsal keel. Amalia has one all down the back. In Limax and Agriolimax +it is confined to the caudal part. Other differences are only +discovered by dissection. + +One may also here note that to preserve slugs is difficult, and the +best plan is to have a coloured drawing made of them when extended. +Otherwise they may be drowned in cold water, cleaned of slime with a +soft brush, and then preserved in glass tubes with diluted formalin or +alcohol. Or, after drowning, they may be skinned and the skins dried +on a card and varnished. Note also that most slugs have many +variations in colour and markings. + +_Testacella haliotidea._--This genus of slugs forms a link between the +naked slugs with rudimentary shells within, and the snails which live +within their shells. The name _Testacella_, or little shell, was given +by Cuvier in 1800, because this slug has a small shell at the end of +the tail. Haliotidea means having a shell in the form of the marine +shell _Haliotis_, the meaning of which again is the ear-shaped +seashell, often called "Venus' Ear." It is subterranean in habit, and +lives on worms. It should be looked for on the surface on damp nights, +or is found when digging. Its length is 3 inches at most. Pale yellow +in colour. See the illustration on page 35. + +_Testacella scutulum_ (a little shield).--Not so common as the former +species, and differing chiefly in anatomy. + +_Testacella Maugei._--First found at Tenerife by M. Mauge. Reaches 4 +inches in length. Deep brown in colour. Shell larger. Rarer and more +western in habitat than the other species. + +[Illustration: _Testacella haliotidea._] + +_Vitrina pellucida._--The Vitrinas in several ways afford a connecting +link between the slugs and snails, having the same tooth-formation and +mantle as the former, while the shell cannot contain the whole body. +As the name indicates, the shell is like a bubble of clear greenish +glass and very delicate. It is small, and found in damp places, coming +out mostly at night. Omnivorous, it is often found feeding on dead +worms, and, unlike nearly all our earth molluscs, can be found abroad +in winter. + +_Vitrea (Polita) lucida._--This is the largest of our British +Hyaliniæ, which are difficult to distinguish. The body of this species +is cobalt blue, the apex of the shell is flat, its colour opaque, and +the last whorl more expanded than in others. All belong to the +sub-genus _Polita_, and have polished or glossy shells. All love shade +and moisture, and should be sought under stones or wood or in moss. +They only come out by day when it is wet, a habit they may have +acquired from their being a favourite food of birds, 416 having been +found in the crop of one nestling Stockdove; while various flies are +very destructive to them. This species prefers animal food, and is +more gregarious than others. Not common. + +_Vitrea (Polita) cellaria._--The next largest species is the most +common of all. It is fond of cellars (whence its name), and I found it +under the stone lid of a manhole in the drain of S. Peter's Rectory, +Walworth--the only shell left in that part of London. It resembles the +previous species, but is smaller and has a broader and deeper suture +between the whorls, while the foot-sole is paler than the body. + +_Vitrea (Polita) Rogersi._--Local. Found in dense woods. It is much +like both _H. cellaria_ and _H. alliaria_, and all three smell of +garlic, but the last is much smaller than the others. The tentacles +in the first are long, and in the third short; while in _Rogersi_ the +upper pair are long and the lower very short. It is also the most +glossy of all. If put in a box with other small shells it will clean +them by cannibalism. + +_Vitrea (Polita) alliaria_, _i.e._ smelling of garlic.--Often confused +by quite good conchologists with the preceding species, but the body +is much darker, and the shell smaller and less white below than either +_cellaria_ or _helvetica_. The always present smell is said to protect +it from ants. Common, but local, and often a pest in greenhouses and +ferneries. + +_Vitrea (Polita) nitidula._--Common. Less glossy. Marked expansion of +the last whorl as it nears the mouth. + +_Vitrea (Polita) pura._--Like _nitidula_ but smaller, and edge of +mantle white instead of dark. More common in the north. Shell thin and +dull white. + +_Vitrea (Polita) radiatula._--Never abundant. Striations on shell give +it a radiated appearance when magnified. Animal nearly black. + +_Vitrea crystallina._--The smallest of the genus. Shell transparent, +pearly white. Umbilicus (_i.e._ the opening in the centre of the +underside showing the whorls) very narrow. Subterranean in habit. +Whorls, four; whereas _H. pura_ has five; also more compressed. + +_Euconulus fulvus._--Distinctively pyramidal in shape. Small. Brown. +Common under rotten branches and moss in woods. Hardly hibernates. + +_Zonitoides nitidus._--Chocolate-brown, with no white round the +umbilicus (as has _H. nitudula_). Larger than, but not unlike, _H. +radiatula_. Gregarious. Chiefly found by water; also in damp +hothouses. Amphibious. + +_Zonitoides excavatus._--Its broad and deep umbilicus is quite +distinctive. Mainly British. Dislikes lime, and is most plentiful on +the coal measures. + +We come now to the Helicidæ family and its genus Helix, in which there +are various sub-genera of which the name is given in brackets. The +shell in this genus can wholly contain the body; the tentacles are +always four; the shell conical, and rarely with a depressed spire. The +word "helix" is Greek, and means a coil. + +_Helix (Gonyodiscus) rotundata._--Very common under stones, moss, etc. +Circular, flat, with a large open umbilicus. Horn colour with brown +markings. + +_H. (Pyramidula) rupestris_, _i.e._ inhabiting rocks.--Small. +Gregarious. Dark brown. Mainly on exposed dry walls and cliffs. + +_H. (Punctum) pygmæum._--Very small. Yellowish brown and glossy shell. +Mainly on moist dead leaves. Not unlike _H. rupestris_ except as to +habitat. + +_H. (Acanthinula) lamellata._--Small. Horn-colour. Epidermis raised +into lamellæ or ridges in the line of growth. Mainly northern. +Frequents dead leaves, especially beech and holly. + +_H. (Acanthinula) aculeata._--More common than the former; which it +resembles in habitat. Differs chiefly by the ridges being produced +into spines. + +_H. (Vallonia) pulchella._--Tiny. White. Mouth trumpet-shaped. +Umbilicus wide. Under stones and at the roots of grass. Its variety +_costata_ (which some make a separate species) is strongly ribbed. + +_H. (Helicigona) lapicida._--Circular, flattened, dark brown, strong +white reflected rim to mouth. Large umbilicus. Marked keel, which +distinguishes it from all other British land shells. Chiefly on chalk +soils. Often on beech tree trunks. + +_H. (Gonostoma) obvoluta._--Common abroad, but confined in England to +a few spots in Sussex and Hants. Circular, flat above, mouth +triangular, with a strong pinkish-white rim with three denticles. + +_H. (Pomatia) pomatia._--Described earlier. Found in Hants, Sussex, +Kent, Surrey, Oxford, Gloucester, and Bedfordshire; but very local. +Elsewhere it may well be an escape from captivity, or the remains of +an attempt (always unsuccessful) to establish a colony. Box Hill and +Caterham are two good localities for Londoners. In Kent it has two +centres, Charing and Shoreham with their contiguous parishes, but +there is a great gap between them, and it is absent from places on the +same chalk ridge which are identical in soil and vegetation. + +_H. (Cryptomphalus) aspersa._--The sub-generic name means that the +umbilicus is hidden in adult shells by a fold of the pillar lip; the +specific name means sprinkled (with brown blotches); but it may be a +slip of the pen, for _aspera_, or rough, from the rough shagreening of +its surface. Five banded, like so many of the Helicidæ, but usually +the second and third band unite. No umbilicus. The variety _exalbida_ +(chiefly found in Kent and the West) is straw colour and somewhat +transparent. Commonly sold for food on the Continent as well as +_pomatia_, which is cultivated in "snail-farms," but not native in +Germany or Switzerland, and in France chiefly found in the coast +departments. Insipid; but as nourishing as calf's-foot jelly. Fond of +gardens (whence its common name), but not of gardeners. As most +animals are marvellously gifted with a knowledge of what food to eat +and what to avoid, it is curious that _aspersa_ will eat voraciously +the leaves of the spindle-tree, though this soon poisons them. It is +said also that they share with cows and horses the ignorance that the +leaves of the yew should be avoided on pain of sickness or even of +death. + +_H. (Cepæa) nemoralis._--As already stated, this is the most +brilliantly and variously coloured and diversely banded of all our +English land shells with the exception of its very close connection +_H. (Tachea) hortensis_. It is happily very common, and so the +attention of beginners should first be directed to this. Thrushes and +mice are its great enemies, the former smashing it on some stone which +may be found surrounded by the broken shells. The "mouth" or peristome +is normally black, the shell larger and stouter than _hortensis_, in +which the mouth is white. When a white-mouthed _nemoralis_ or +dark-mouthed _hortensis_ (both rare) is found, the shape of the +internal flinty dart at once distinguishes them. In some places both +live together: in most one is found and not the other. _Nemoralis_ is +fond of sand-hills by the coast, but is chiefly a hedge-snail, and the +edges of main roads are preferred because of the greater variety of +food, because the traffic scares away their bird enemies, and because +the dust gives them abundance of already prepared material for their +shells. When, however, the collector comes to a wayside cottage where +fowls are kept he need not waste his time in looking for snails in the +neighbouring hedge. The more the chicken industry extends and the more +the Bird Protection Acts operates, the worse it is for collectors of +snails. The banding is probably protective, as in the case of the +tiger and the zebra, and renders the shell less visible. + +_Helix (Cepæa) hortensis._--Rarely found in gardens in spite of its +specific name. A hedge-snail. White forms not uncommon, though almost +unknown in _nemoralis_. Though the weaker form, the coalescence of the +five bands into one broad one is more common here than in _nemoralis_. +Also the variety with only one band, and that on the periphery, is +very common in _nemoralis_ and rare in _hortensis_. It is more +dependent on shade and moisture than its congener. Smells of garlic +when immersed in boiling water to be killed. _Hortensis_ is a more +northern, and _nemoralis_ a more southern, shell by origin and +distribution. There are 89 possible band variations in any normally +five-banded shell, and all have been noted in the case of _nemoralis_, +but in _hortensis_ only 61. They are distinguished, for purposes of +record and exchange, by numbers. Thus the type is 12345, the usual +one-banded variety 00300, the common coalescence of the second and +third band is 1(23)45, and when all bands unite (12345). The +unicolourous or bandless variations would be 00000. + +_H. (Arianta) arbustorum._--Local. Usually found in hedges and by +ditches on chalk and limestone. Shell globose, brown or yellow, with a +check or willow leaf pattern, and a single dark band on the periphery. +Lip strong and white. Animal usually nearly black. Very fond of +moisture. Anatomically related to _A. lapicida_, but no external +resemblance. + +_Helix (Theba) cantiana._--First observed in Kent (where it is +especially fine and abundant), whence its specific name, but generally +dispersed in South and East England. A dull, creamy white shell with a +pink tinge, sometimes becoming partially or wholly reddish. + +_Helix (Theba) cartusiana_ (first noticed near a Carthusian +monastery). Much resembles _cantiana_, but is much smaller and more +smooth. Chiefly found on the downs of Kent and Sussex. Used to be +common on Deal sand-hills--now devastated by golf! The tint in this is +brown, in the former red. + +_H. (Hygromia) rufescens._--A flattish, dark brown shell, abundant in +the south of England, and not rare elsewhere. Has a semi-lunar mouth +with a white internal rib. In gardens seems to prefer violet beds. + +_H. (Hygromia) hispida_, _i.e._ hairy.--These hairs are deciduous, and +the hairless variety used to be considered a separate species under +the name of _concinna_ (_i.e._ neat), but would now be the variety +_depilata_, or bald. Broad and deep umbilicus. Common, except in +Ireland. Usually associates with _H. rufescens_ in moist places. + +_H. (Hygromia) granulata_ is also hairy with white silky bristles. +Yellowish in colour. Shell thin. Local, but abundant where found. Its +umbilicus is very small. It falls from its food plants at the least +shake. + +_H. (Hygromia) revelata._--Scantily haired. Globular thin shell. Pale +green. Mainly found in Cornwall and South Devon. In cold or dry +weather it buries itself rather deeply. + +_H. (Hygromia) fusca._--Very thin, glossy, brown shell. Local. Hardy, +and even active in frost. Chiefly found on nettles, which many shells +like as food, though avoiding the commonly associated horehound. + +_H. (Euparypha) pisana._--First noticed at Pisa. Somewhat like +_Helicella virgata_, but larger, sub-globular, and solid shell, +yellowish-white with dark lines or bands. Aperture or mouth yellowish +or rosy. Most common in Portugal and Morocco, and all round the +Mediterranean, dry places, especially near the sea. In England chiefly +confined to Tenby and other parts of Pembrokeshire; also in the +Channel Islands. Varies much in tint and markings. Swarms where found; +it loves sun and heat. Seems to lend itself better to colonization +than most species. + +_H. (Helicella) itala._--So named by Linnæus, who probably received it +first from Italy. Shell almost circular, flat. Umbilicus very large +and open. Common on heaths and downs, especially near the sea. + +_H. (Candidula) caperata._--(The specific name means wrinkled, like a +goat's horn.) Careless of heat or cold. Distinguished from the young +of _H. virgata_ by being more depressed, having a larger umbilicus, +regular and strong striation, and round mouth with white internal rib. +Found under stones and on grass. Common. + +[Illustration: _Helicella virgata_ at rest on thistle, natural size.] + +_H. (Heliomanes) virgata_ (_i.e._ striped).--A very variable shell. +See the illustration above of some at rest on thistles. Local, but +very abundant where found. Whitish shell with dark bands, but a +yellowish and a white variety usually is found with the type. The most +beautiful variety, _radiata_, is chiefly found in Romney Marsh, and +from Hythe to Rye. + +_H. (Turricola) terrestris._--A Mediterranean species, well +established since 1890, in one spot near Dover. A pyramidal shell, +greyish, with one dark band on each whorl. + +_H. (Cochlicella) barbara_ (_i.e._ foreign).--Long, conical, whitish, +with one dark band. By the sea-coast. In shape somewhat like a +Buliminus. + +We come now to the Pupa family and its genus Buliminus and its +sub-genus Ena. It is represented by:-- + +_Ena montana._--A local and southern shell, conical, slightly glossy, +brown. Lip white and deflected. Commonly found on the holes of +smooth-barked trees, and it closely resembles the small knobs on beech +trunks. + +_Ena obscura._--Like the former, but much smaller, and found nearly +everywhere in England and Wales. Found in hedgebanks, or on beech +trunks. Its specific name is derived from its habit of covering itself +with a coating of earth, and so becoming inconspicuous. + +The plate on p. 47, gives figures of some of our smaller shells, +enlarged in most cases so that their distinguishing marks can be seen. +The upright line by the side of each figure gives its actual height. +The shells as numbered are _Helix rupestris_, _H. pygmæa_, _H. +pulchella_, _H. lapicida_, _H. obvoluta_, _H. terrestris_, _H. +barbara_, _Ena montana_, _Ena obscura_, _Pupa secale_, _P. anglica_, +_P. cylindracea_, _P. muscorum_, _Vertigo antivertigo_, _V. +moulinsiana_, _V. pygmæa_, _V. alpestris_, _V. substriata_, _V. +pusilla_, _V. angustior_, _V. edentula_, and _V. minutissima_. Without +a magnifying glass it will be seen that it would be very hard to +distinguish some of the minute shells, but this enlargement enables us +to see the characteristic denticles in the mouth, and the presence or +absence of striations on the shell. + +[Illustration: Some of our smaller shells. Actual size indicated by +the upright line.] + +_Pupa (Abida) secale_ is named from the Latin for rye, a grain of +which the shell more or less resembles. Conical, brown, mouth +horseshoe-shaped with eight white denticles. Our largest Pupa. Local, +but abundant where found. Prefers calcareous rocks or woods. + +_Pupa (Lauria) anglica._--Small, ovate, purplish in colour; mouth like +that of _secale_. Lives in moss, mainly in the north of Britain. + +_Pupa (Lauria) cylindracea._--Small, cylindrical, paler than the last; +thick and reflected white lip with one denticle. Abundant. On stones, +in moss, under leaves and bark. + +_Pupa (Jaminia) muscorum._--Common, especially on sandy soils near the +sea. Mouth nearly circular, whereas in the two former species it is +horseshoe-shaped. The lip is thin and not reflected. + +The genus Vertigo (_i.e._ twisted, the Latin equivalent of the Greek +Helix) contains shells even smaller than the Pupæ, about the size of a +pin's head. + +_Vertigo (Alæa) antivertigo_ (_i.e._ not reversed or sinistral, as are +_V. pusilla_ and _V. angustior_). Semi-transparent, glossy, +horn-colour, with denticles (as have all except _V. edentula_ and _V. +minutissima_). Found in nearly all counties in moist places. + +_Vertigo (Alæa) moulinsiana._--Our largest species, though only 2-1/3 +millimetres in height. Mainly in marshy places. Not common. + +_Vertigo (Alæa) alpestris._--Rare and local, chiefly northern. Nearly +transparent shell. + +_Vertigo (Alæa) pygmæa._--Common, and often in colonies at roots of +grass and under stones and logs. Not confined to moist places. + +_Vertigo (Alæa) substriata._--Local. Strongly striated. + +_Vertigo (Vertilla) pusilla._--Sinistral, as is also + +_Vertigo (Vertilla) angustior._--Both species rare and local. The +former is the larger and broader. In the former the last whorl is +broadest, in the latter the penultimate. In the former the mouth is +semi-oval, in the latter triangular. In the former the outer lip is +very slightly, in the latter very deeply contracted. The former has 6 +to 7 teeth, the latter 4 to 5. + +_Vertigo (Sphyradium) edentula_ is dextral and without denticles. +Perhaps the most common _Vertigo_. Partial to bracken. + +_Vertigo (Isthmia) minutissima._--Dextral and without denticles. +Smaller, narrower, and more strongly striated than edentula, but +rarer. All the Pupæ should be examined with a magnifier. + +_Balea perversa_ (_i.e._ sinistral) is a much larger shell belonging +to the Clausilia family. Thin, dark horn-colour, semi-transparent, +glossy, 7 to 8 whorls, local, but abundant where found. Chiefly found +on trees. + +_Clausilia (Pirostoma) bidentata._--All our British clausilias are +sinistral. The clausilium (little door) is an internal contrivance +fastened to the pillar of the shell (whereas an operculum is attached +to the body of a mollusc) by an elastic ligament to protect it against +insect enemies when the animal withdraws. _Bidentata_ has two +denticles, fusiform and reddish-brown, as are all. Very common on +walls and trees. + +_Clausilia (Pirostoma) rolphii._--Rare and local. Almost subterranean +in habit. More coarsely striated than the last. The upper whorls +nearly of the same breadth, forming a short cylinder. + +_Clausilia (Alinda) biplicata._--Very local. Chiefly on Thames +willows. Larger than the two former, and streaked with white. + +_Clausilia (Marpessa) laminata._--Much like the former, but widely +distributed. Usually found on beech and ash trees, and on limestone +rocks. Smooth and glossy. + +In the family Stenogyra we have three genera, Azeca, Cochlicopa, and +Cæcilioides (with also the imported _Stenogyra Goodallii_, found only +in pine-houses). + +_Stenogyra (Azeca) tridens_ is a small chrysalis-shaped, solid but +semi-transparent shell, horn-coloured, with 3 denticles. Not rare in +moist places. + +_Stenogyra (Cochlicopa) lubrica_ (_i.e._ slippery).--Very common in +moss and under stones or logs. Much like the previous species, but no +denticles and fewer whorls, and broader mouth. + +_Stenogyra (Cæcilioides) acicula._--If this word is supposed to be +Latin it would mean either "like to a blind worm" or "like to a +lettuce"! _Cæcus_, however, being Latin for blind, the allusion is no +doubt to the fact that this wholly subterranean species is eyeless. +The only British representative of a large family of carnivorous +molluscs. I have found it on Saxon bones when unearthed, and in +crevices of limestone underground, but it is generally found dead +amongst the rejectamenta on the banks of rivers. It is a pretty, +glossy white shell, 5 millimetres in height by 1 in breadth. + +I may notice here two other land shells, although they scientifically +are grouped amongst the fluviatile Gasteropoda. + +_Cyclostoma (Pomatias) elegans._--Common on calcareous soils, +especially chalk. A spiral shell of 4-1/2 whorls, suture very deep. +Mouth circular (whence its name) and provided with a thick shelly +operculum which closes the orifice when the animal retires by means of +an elastic ligament. This and the next species are our only land +shells provided with an operculum, and this shows their derivation +from the marine Gasteropoda (_e.g._ whelk and winkle). Perhaps all +shells were originally marine, but some became first amphibious and +then terrestrial. It is quite unlike any other of our land shells. + +_Acicula lineata_ is a very small shell, the size of the Pupæ; mainly +northern in distribution. Feeds on liverworts and fungi. Very local; 6 +or 7 whorls. Mouth pear-shaped, with a horny operculum. + +The Family Succinea really ranks with the land shells, as belonging to +the sub-order Pulmonata or lung-breathing molluscs. It is, however, +amphibious, and hibernates in the mud at the bottom of a ditch. + +_Succinea putris_ (it is the mud, not the animal, which is putrid!) is +called the Amber Snail from the colour of its shell, which is unlike +any other. Common on flags, etc., at the edges of ditches and ponds. + +_Succinea elegans._--Difficult to distinguish from the former, but the +animal is darker and the shell more slender, with a deeper suture and +a narrower mouth. + +_Succinea oblonga_ is local and rare. Generally found near the sea. +Much smaller than the other Succineas, and easily mistaken for the +young of other species. Colour dull greenish. + +The family Auriculidæ is represented in Britain only by _Carychium +minimum_; a very small, semi-transparent, white and glossy shell found +under mossy stones and other moist places. Common, but sharp eyes are +needed to find it. + + * * * * * + +We now come to the freshwater shells, which we capture best by means +of a perforated scoop, whether they are on the waterweeds or hidden in +the sand or mud of the bottom. + +It may be noted that all freshwater shells are greenish-brown which is +an excellent protective colouring as rendering them less visible among +water weeds to the fish, which devour them greedily. + +The family of Limnæidæ (or lake dwellers) has the sub-families, +Planorbis, Physa, Limnæa, and Ancylus. In the Planorbinæ (_i.e._ +flat-coiled) the only representative of the genus Segmentina is +_Segmentina nitida_, a small, quoit-shaped, keeled, semi-transparent, +light brown shell, with internal divisions like those of a nautilus +which are visible from the outside of the shell. Local. Found in +stagnant or sluggish water. The genus Planorbis contains the +sub-genera Hippeutis, Gyraulus, Gyrorbis, Coretus, and Bathyomphalus. + +_Planorbis (Hippeutis) fontanus_ is much like Segmentina but has no +septa, and is flatter. Common, especially on watercress. Often +encrusted with mud. + +_Planorbis (Gyraulus) nautileus_ is very small; quoit-shaped, with the +upper side flat. Grey and striated. The variety crista has the ridges +of the epidermis drawn into points, and is beautiful when seen by a +magnifying glass. Common in ponds and ditches. + +_Planorbis (Gyraulus) dilatatus_ is a very small shell imported in +cotton bales from America, and naturalized in canals in Lancashire. No +other of its kind is so small. + +_Planorbis (Gyraulus) albus_ is dull white and striated. Flattish +above, with spire depressed. Frequently encrusted and black with mud. +Common. + +_Planorbis (Gyraulus) parvus_ (but not so small as +_dilatatus_).--Convex above with a central depression, concave +beneath. Suture deep, and umbilicus large. Smooth and glossy. Local. + +_Planorbis (Gyrorbis) spirorbis._--Very flat, glossy, brown, whorls 5 +to 6. Common in ponds and ditches. + +_Planorbis (Gyrorbis) vertex._--Very like the last, but flatter and +thinner, and with a prominent keel. More local than _spirorbis_, but +sometimes found with it. Whorls 6 to 8. + +_Planorbis (Gyrorbis) carinatus._--Larger than _spirorbis_ and +_vertex_. Sharply keeled in the centre of the outer margin. Mouth +angulated above and below. Local, mainly in the south and east of +England. + +_Planorbis (Gyrorbis) umbilicatus._--Like the last, but the keel is +below and not on the centre. Mouth rhomboidal. More common than +_carinatus_. + +_Planorbis (Coretus) corneus._--Far the largest species. Dark brown, +lighter below. Mouth nearly circular. Spire sunk. In boiling water +often exudes a crimson fluid. Common. + +_Planorbis (Bathyomphalus) contortus._--Small, 8-whorled, flat above, +very convex below. Fairly common in still water. Very compact in +appearance. + +The sub-family Physa has two genera, Aplecta and Physa. + +_Physa (Aplecta) hypnorum_ is a spindle-shaped, very glossy, +semi-transparent, dark reddish brown, shell, with 6 to 7 whorls. Not +common. Found in still water. + +_Physa (Physa) fontinalis._--More common, and found in running as well +as in still water. Shorter and more rounded than the last. Shell very +thin, greenish horn-colour. Lobes of the mantle expand over the shell. +Seen in an aquarium are its perpendicular threads of mucus, up and +down which the animals climb. + +_Limnæa (Amphipeplea) glutinosa._--Very local. Somewhat like _Ph. +fontinalis_, but larger and more thin. In young specimens the mantle +covers the shell, and in adults the animal is not wholly contained in +the shell. + +_Limnæa (sub-genus Radix) involuta._--Only found in one Irish tarn. +Whorls envelop the spire. Very thin, pale amber. + +_Limnæa (Radix) peregra._--The most common and variable of all our +freshwater shells. Spire pointed. Somewhat amphibious. Found +practically over the whole of the Eastern Hemisphere. + +_Limnæa (Radix) auricularia._--Mouth very large, with outer lip widely +reflected. Very common and fine in the Thames. Spire very short, apex +sharp. + +_Limnæa (sub-genus Limnophysa) stagnalis._--The largest of the genus. +Common, except in Wales. Shell greyish, spire long and tapering to a +point; 12210 teeth on its lingual ribbon. See the illustration on p. +57, which also shows above two specimens of _Paludina contecta_, one +being covered (as freshwater shells often are) by a vegetable growth, +which obscures the marking. + +_Limnæa (Limnophysa) palustris._--Shell tapering, somewhat solid, +brown, much smaller than stagnalis. Common in slow or stagnant water. +Some varieties much darker than the type. + +_Limnæa (Limnophysa) truncatula._--Like the last in shape, but much +smaller, and with a deeper suture. Common, and fond of being out of +the water. A parasite of this mollusc causes "fluke" in sheep which +have taken it in by drinking or by eating grass by the side of ponds +and ditches. + +[Illustration: _Paludina contecta_ (two) and _Limnæa stagnalis_ on +water-weeds.] + +_Limnæa (Omphiscola) glabra._--Also amphibious. About the same size as +_truncatula_. Local. Inner lip rather thick and reflected on the base +of the penultimate whorl. + +_Limnæa (Ancylus) fluviatilis._--"Freshwater limpet." Shell, rather +limpet-like, with a hooked apex (whence its generic name), adheres to +stones or piles in running water. Common. I once dredged a large +water-beetle with three of these shells adhering to its wing-cases; +thus it would be transported to fresh habitats. + +_Limnæa (Acroloxus) lacustris._--Like the former but more local, and +preferring sluggish or still waters. Shell more oblong, thinner, and +apex twisted to the left instead of to the right as in _fluviatilis_. + +The sub-order Pectinibranchiata (comb-like gill) contains the genera +Neritina, Paludina, and Valvata, in all of which there are two +tentacles with eye at the base, and an operculum to the shell. + +_Neritina fluviatilis._--Solid, glossy, chequered brown, white, and +purple (but also a lemon-coloured variety). Operculum semi-lunar, +orange, with a projection which serves as a lock to keep the operculum +in position. Not rare in England; on stones in running water. See +illustration below, which also shows above _L. (Ancylus) +fluviatilis_. + +_Paludina (Vivipara) contecta._--Shell dark green with darker bands. +Conical. Suture very deep. Operculum horny. Viviparous. Local. + +_Paludina (Vivipara) vivipara._--More common than contecta. Shell more +oval, not so glossy, light greenish yellow, suture not so deep, no +umbilicus, apex blunt. + +[Illustration: _Neritina_ and _Ancylus_.] + +_Paludina (Bythinia) tentaculata._--(The eyes in this genus are not on +foot-stalks; the operculum is shelly instead of horny). Common in slow +water and ditches. Shell semi-transparent, yellowish, mouth oval, +angulated above. Operculum made of plates rising one above another +formed at different stages of growth. + +_Paludina (Bythinia) leachii._--Much smaller and less common than the +last. Distinct umbilicus; mouth almost circular. + +_Paludina (Paludestrina) ventrosa._--A brackish-water shell, swarming +where found, _e.g._ from Erith to Gravesend, and in East Anglia. Shell +small, thin, semi-transparent. + +_Paludina (Paludestrina) jenkinsi._--A larger shell, not confined to +brackish water and spreading very rapidly. Swarms where found. A +variety has a marked keel which sometimes bears bunches of spines at +equal distances. + +_Paludina (Paludestrina) stagnalis._--Larger and with more whorls. Not +so common. + +_Paludina (Pseudamnicola) anatina._--Small, sub-conical, deep suture. +Found in brackish water, and apparently identical with _Hydrobia_ or +_Paludestrina similis_, which I used to find by the Thames, where it +is now apparently extinct. + +_Valvata piscinalis._--Globular, suture very deep, circular mouth, +operculum concentrically spiral. In ponds and slow water. Shell +yellowish, but commonly covered with conferva. + +_Valvata cristata._--Much smaller; shell disk-shaped. Frequents the +roots of flags. Shell striated and more or less ridged, but the name +_cristata_ refers to the plume-like appearance of its breathing +apparatus. + + * * * * * + +We now come to the bivalve shells with leaf-like gills. The Unionidæ +contain two genera, Unio and Anodonta, commonly called freshwater +mussels. + +_Unio tumidus._--Shell ovate, very solid, dark brown; common. See +accompanying illustration, which shows the fringed branchial siphon +which draws in food-bearing water, and the smaller anal siphon by +which it gets rid of undigested matter. + +[Illustration: Freshwater mussel breathing and eating.] + +_Unio pictorum._--More oblong and thinner shell, yellowish, girdled +with brown in the lines of growth. Common. The specific name recalls +that gold and silver paint used to be sold in these shells (or marine +mussels) for illuminating work. It is said to produce 220,000 eggs in +the three summer months. + +_Unio (margaritana) margaritifer._--Shell solid and black, beaks +always eroded. Mainly found in mountain streams. Its pearls are few +and poor compared with those of marine shells; but they attracted the +notice of Cæsar and so hastened the conquest (and development) of +Britain. + +_Anodonta cygnea._--(In this genus the hinge is toothless, whence its +generic name. The specific names _cygnea_ and _anatina_ mean "swan" +and "duck," in reference to their comparative size). This is the +largest of our freshwater shells, reaching even 9 inches in breadth by +4-1/2 in length. Common in ponds and slow water. Sometimes the shells +are yellowish green with rays of the same colour. + +_Anodonta anatina._--Doubtful if this is a separate species or only a +smaller form. The hinge line is raised instead of being straight, and +the posterior side slopes abruptly instead of gradually. + +In the next family are two genera, Sphærium and Pisidium. + +_Sphærium corneum._--Very common. Shell somewhat globular, glossy, +opaque, horn-coloured, marked with lighter bands in the line of +growth. Usually on the bottom, but can suspend itself by threads of +mucus. + +_Sphærium rivicola._--Much larger. Also flatter and more striated. +Yellowish brown or greenish. A whole series of young of different +sizes will be found in the animal. + +_Sphærium pallidum._--Local in canals and ponds. Oblong. +Distinguished also from the previous species by the body being +milk-white, and the shell is ashy-grey. + +_Sphærium lacustre._--Local. On the beaks is a calcareous nucleus +which distinguishes it. It is thinner than _corneum_, and rounder than +_pallidum_. + +_Pisidium amnicum._--(Our five _pisidia_ resemble _Sphærium_, but are +much smaller, all but _amnicus_ being minute. Very abundant where +found. _P. amnicum_ and _fortinale_ are triangular in shape, _P. +pusillum_ oval, _P. nitidum_ round, and _P. roseum_ or _milium_ +oblong; but they are difficult to distinguish on account of their +similarity and variation). _P. amnicum_ is nearly twice the size of +the others, and this and _fontinale_ may be found in slow rivers, +whereas the others prefer stagnant waters. + +_Pisidium fontinale._--Smaller and thinner, and with more prominent +beaks than _P. amnicum_. + +_Pisidium pusillum._--The most common species. Distinguished from the +last by being oval and by its beaks being blunter and more central. + +_Pisidium nitidum._--Rare. Very glossy and striated. + +_Pisidium roseum_ (from the colour of part of its body).--Like +_nitidum_, but oblong, with a straight lower margin, and with beaks +placed away from the centre. + +The last shell to be mentioned could not be mistaken for any other. It +belongs to the sub-order _Heteromya_ (_i.e._ with adductor or closing +muscles not equal); to the family of _Mytilidæ_ (or mussels) and the +genus _Dreissensia_ (named after a Dutch conchologist). + +_Dreissensia polymorpha_ is a triangular, boat-shaped, bivalve, +supposed to have been introduced with Russian timber (as was also +probably _Hydrobia Jenkinsi_). It is gregarious, and attaches itself +to objects by a byssus like our marine mussels. Shell yellowish-brown +with wavy purplish lines, wrinkled in the line of growth. Common in +the New River, and has been found in iron water-pipes in Oxford +Street. + + * * * * * + +All our shells have varieties (many an albino or white form), and the +collection and distinguishing of these varieties, which in some +species are numerous, adds much to the interest of the collector. In +addition there are also the variations in size or markings which can +hardly rank as varieties. Inasmuch as none of our shells are peculiar +to our country (which is from the natural history and the geological +point of view only a detached portion of the Continent), it may be +well to warn young collectors that if they receive shells from the +Continent, mere varieties are there often named as separate species +and variations considered as definite varieties. This is especially +the case with _Helicella virgata_. + +As to the arrangement of shells in a collection before a regular +cabinet is obtained, the tinier shells may be kept in small glass +tubes with corks (such as used for homoeopathic medicines), and the +medium sized ones in the trays of common matchboxes, these being +arranged in large shallow glass-covered trays which can be obtained +from any cardboard boxmaker at a small cost, and several of these, +stored one above the other, form an excellent substitute for a more +costly cabinet. In all cases the name, and the place where the shells +were found, should be written on a small slip of card placed in the +tube or tray. It is not well in most cases to fasten the shells on +card, but if this is done gum tragacanth is best. The collection +should be kept free from damp and from dust. + + + HINTS FOR COLLECTING AND PRESERVING SHELLS OF MOLLUSCS. + +The following notes supply a few general rules as to finding and +preserving shells:-- + +Of Shell-bearing Molluscs there are three classes--Marine, Freshwater, +and Land. The first two include Univalves and Bivalves, the last only +Univalves. + +1. MARINE SHELLS may be obtained, 1st, by searching on and under rocks +at low water, or on coral reefs, among seaweed attached to them, or +floating on the sea, or on a sandy beach. Bivalves may be found by +digging in the sand, or mud, on a beach, or at the mouth of a river: +their presence is generally indicated by a circular breathing hole in +the sand. 2nd. By dredging, by which means only deep-sea shells can be +obtained; but after a storm these may often be found upon the shore, +before they have lost their lustre. + +Limpets, etc., should be detached with a thin blade passed quickly +under the shell, taking care not to break the edges. Small shells on +and in seaweed, and limpets, etc., adhering to stones will drop off +and sink to the bottom in a vessel of cold fresh water. + +2. FRESH-WATER SHELLS may be obtained in any river, lake, pond, marsh +or reservoir. Univalves, chiefly on the banks, on reeds and plants +growing near the hedges, and on the under surface, leaves, and stems +of aquatic plants. Bivalves generally at the bottom, among stones, or +buried in the sand, or among the roots of aquatic plants. + +3. LAND SHELLS.--These resemble, more or less, in their habits the +garden snail, though varying greatly in character, size, and colour. +They mostly abound in a chalk or limestone district, and in moist and +wooded situations. Some species inhabit low and damp spots, roots of +trees, hollows and crevices of rocks and walls; some lie under stones +or pieces of wood, or in the earth; others climb shrubs, and in +tropical climates even lofty trees. Their haunts vary according to the +weather and the season. They come out early in the morning, and after +rain. Some bury themselves in moist places during the dry season, or +burrow under leaves, grass, or stones, often closing the mouths of +their shells with a white secretion to prevent evaporation during the +period of hibernation. + +The smallest shells, especially of land species, and young imperfect +shells should be collected. + +In all cases "live shells," _i.e._ shells in which the animal is +alive, are to be chosen; but, when these cannot be procured, "dead +shells," which have not lost their lustre, or their colour, especially +those of rare species, should be preserved. + + + _With regard to the mode of Preserving Shells._ + +1. No attempt should be made to clean them, or to remove the furry +skin, more or less thick, with which they are often covered, beyond +removing with a soft brush any mud or sand adhering to them. + +2. The animals of Land and Freshwater shells may be killed by +immersing them for a few minutes in _boiling_ water, after which the +bodies may be easily extracted whole with any suitable instrument, +_e.g._, a fork or a pin, according to size. Hot water should not be +used with marine shells: it often destroys their lustre. They should +be buried, if time permits, in sand, or other dry material, until the +animal dries up (in small shells) or rots (in large specimens); or +they may be drowned in cold fresh water, and hung up in the air to dry +or rot away. In the former case, if an operculum (with which some +species, both marine and land, close their mouths, more or less +partially) exists, it will, generally in the case of land shells, +remain in its place, adhering to the shell. In the latter, the decayed +matter should be washed out, and the operculum, if any, replaced and +fixed, say, on cotton filling the shell. This applies equally to land +shells. + +3. Care should be taken not to injure the edge or lip of the mouth of +univalves, or the ligament of the hinge of bivalves. When bivalves +gape on dying in water, or if the ligament be broken, the valves +should be closed and tied together. If the ligament of a gaping +bivalve should become dry and stiff, it can be softened by putting it +in water. + +4. The localities in which each species is found should be noted, and, +in the case of dredging, the depth of water. + + + _With regard to the mode of packing Shells for Transport._ + +All solid shells may be wrapped in one or two folds of paper of any +kind. Fragile and minute shells should be put, generally separately, +into a box or bottle--with or without cotton, as required. Such +packets may be heaped up in any box, heavy shells at the bottom, +without pressure, and any blank filled at the top with paper or other +elastic material. Sawdust injures the lustre of many species. + +Two books on shells should be procured at an early stage of the +collector's career, which will give not only minute descriptions of +all our land and freshwater shells and their varieties, but also +plates of illustrations. These are the _Collector's Manual_, by L. E. +Adams, 2nd ed., published by Taylor Brothers of Leeds; and Rimmers' +_Land and Fresh Water Shells_, published by George Grant of Edinburgh. + + +THE END. + + + PRINTED BY + + WILLIAM CLOWES AND SONS, LIMITED, + + LONDON AND BECCLES. + + + * * * * * + + +PUBLICATIONS + +OF THE + +Society for Promoting Christian Knowledge + + +BIRDS, SOME FOLK-LORE AND LEGENDS OF. + + Written mainly for Children, by Canon J. W. HORSLEY. Small post + 8vo. Cloth, 6_d._ net. + + +BIRDS (BRITISH) IN THEIR HAUNTS. + + Being a Popular Account of the Birds which have been observed + in the British Isles, their Haunts and Habits, their + systematic, common, and provincial Names. By the late C. A. + JOHNS. With 16 coloured Plates and many Woodcuts. Post 8vo, + cloth, 5_s._ net. + + +BIRDS (SKETCH BOOK OF BRITISH). + + By R. BOWDLER SHARPE, LL.D., F.L.S. With coloured Illustrations + by A. F. and C. LYDON. Crown 4to, cloth boards, 10_s._ 6_d._ + net. + + +BIRDS' EGGS (BRITISH). + + Described and Illustrated with 20 coloured Plates, by A. F. + LYDON. 4to, cloth boards, 5_s._ net. + + +BUTTERFLIES AND MOTHS. In Romance and Reality. + + By W. F. KIRBY, F.L.S., F.E.S. With 28 coloured Plates and + other Illustrations. Small 4to, cloth, 5_s._ net. + + +EDIBLE AND POISONOUS MUSHROOMS: What to eat and what to avoid. + + By the late M. C. COOKE, M.A., LL.D. With 18 coloured Plates of + 48 species. Crown 8vo, cloth boards, 2_s._ 6_d._ net. + + +PUBLICATIONS OF THE S.P.C.K. + + +FLORAL RAMBLES IN HIGHWAYS AND BYWAYS. + + By the Rev. Professor G. HENSLOW, M.A., F.L.S., F.G.S. With 32 + coloured plates and 100 illustrations in black-and-white. Large + crown 8vo, cloth boards, 6_s._ net. + + +FLOWERS IN THEIR NATURAL COLOURS AND FORM (BRITISH WILD). + + Text by the Rev. Professor HENSLOW, M.A., F.L.S., F.G.S. With + over 200 coloured Illustrations drawn from, and of the size of, + the natural plant. Large crown 8vo, cloth boards, 8_s._ net. + + +FLOWERS OF THE FIELD. + + By the late Rev. C. A. JOHNS, B.A., F.L.S. (33rd Edition). + Revised by Professor G. S. BOULGER, F.L.S., F.G.S. With + Portrait and Memoir of Author. Numerous Woodcuts. 64 coloured + Plates by GRACE LAYTON. Large crown 8vo, cloth boards, 6_s._ + net. + + +FOREST TREES OF BRITAIN (THE). + + By the Rev. C. A. JOHNS, B.A., F.L.S. Post 8vo. With 16 Plates + taken direct from Nature by Photography, giving the Natural + Colours, and 150 Woodcuts, cloth boards, 6_s._ net. + + +SELBORNE (THE NATURAL HISTORY OF). + + By the late Rev. GILBERT WHITE, M.A. With Map and numerous + Woodcuts. Post 8vo, cloth boards, 2_s._ net. + + +THE ANIMAL KINGDOM. + + Illustrated in 27 coloured Plates containing several hundreds + of species. The letterpress by Dr. ZWANZIGER, of Fuerth, + translated by GERARD K. GUDE, F.Z.S. Large 4to, half cloth, + 8_s._ 6_d._ net. + + +THE PROCESS OF THE YEAR. Notes on the Succession of Plant and Animal +Life. + + By HENRY HILTON BROWN, F.E.S. Numerous Illustrations. Crown + 8vo, cloth boards, 2_s._ 6_d._ net. + +SOCIETY FOR PROMOTING CHRISTIAN KNOWLEDGE + + +LONDON: NORTHUMBERLAND AVENUE, W.C. + + + + +Transcriber's Notes: + + +Archaic and inconsistent punctuation and spelling retained. + + + + + +End of Project Gutenberg's Our British Snails, by John William Horsley + +*** END OF THE PROJECT GUTENBERG EBOOK 40880 *** |
