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+ The Project Gutenberg eBook of Fishes of the Big Blue River Basin Kansas, by W. L. Minckley.
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+<pre>
+
+The Project Gutenberg EBook of Fishes of the Big Blue River Basin, Kansas, by
+W. L. Minckley
+
+This eBook is for the use of anyone anywhere at no cost and with
+almost no restrictions whatsoever. You may copy it, give it away or
+re-use it under the terms of the Project Gutenberg License included
+with this eBook or online at www.gutenberg.org
+
+
+Title: Fishes of the Big Blue River Basin, Kansas
+
+Author: W. L. Minckley
+
+Release Date: October 13, 2011 [EBook #37742]
+
+Language: English
+
+Character set encoding: ISO-8859-1
+
+*** START OF THIS PROJECT GUTENBERG EBOOK FISHES OF THE BIG BLUE RIVER ***
+
+
+
+
+Produced by Chris Curnow, Tom Cosmas, Joseph Cooper and
+the Online Distributed Proofreading Team at
+http://www.pgdp.net
+
+
+
+
+
+
+</pre>
+
+
+
+<div class="book">
+<p><span class="pagenum"><a name="Page_401" id="Page_401">[401]</a></span></p>
+
+<br />
+<img src="images/bar_double.png" width="100%" height="15" alt="double bar" title="double bar" />
+<div class="caption2"><div class="smcap">University of Kansas Publications<br />
+Museum of Natural History</div></div>
+<div class="center"><img src="images/bar_single.png" width="75%" height="15" title="bar" alt="bar" /></div>
+<br />
+<div class="caption2">Volume 11, No. 7, pp. 401-442, 2 plates, 4 figs. in text, 5 tabl.</div><br />
+<div class="center">
+ <img src="images/bar_single.png" width="28%" height="15" title="bar" alt="bar" />&nbsp;&nbsp;
+ <span class="caption2">May&nbsp;8,&nbsp;1959</span>&nbsp;&nbsp;
+ <img src="images/bar_single.png" width="28%" height="15" title="bar" alt="bar" />
+</div>
+<br />
+<br />
+<br />
+<div class="caption1">Fishes of the Big Blue River Basin, Kansas</div>
+<br />
+<br />
+<br />
+<div class="caption3">BY</div>
+<br />
+<div class="caption2">W. L. MINCKLEY</div>
+<br />
+<br />
+<br />
+<br />
+<br />
+<br />
+<div class="caption2">
+<span class="smcap">University of Kansas</span><br />
+<span class="smcap">Lawrence</span><br />
+1959
+</div>
+<br />
+<br />
+<br />
+
+<p><span class="pagenum"><a name="Page_402" id="Page_402">[402]</a></span></p>
+
+<div class="center">
+<div class="caption3">
+<span class="smcap">University of Kansas Publications, Museum of Natural History</span><br />
+<br />
+Editors: E. Raymond Hall, Chairman, Henry S. Fitch,<br />
+Robert W. Wilson<br />
+</div>
+<br />
+<br />
+<br />
+Volume 11, No. 7, pp. 401-442, 2 plates, 4 figs. in text, 5 tables<br />
+<br />
+Published May 8, 1959<br />
+<br />
+<br />
+<br />
+<span class="smcap">University of Kansas</span><br />
+Lawrence, Kansas<br />
+<br />
+<br />
+<span class="smcap">A Contribution From<br />
+The State Biological Survey of Kansas</span><br />
+<br />
+<br />
+<br />
+<div class="caption4">
+PRINTED IN<br />
+THE STATE PRINTING PLANT<br />
+TOPEKA, KANSAS<br />
+1959<br />
+<img src="images/union_label.png" width="71" height="26" alt="Look for the Union Label" title="Look for the Union Label" /><br />
+27-7080<br />
+</div>
+</div>
+<br />
+<br />
+<br />
+
+<p><span class="pagenum"><a name="Page_403" id="Page_403">[403]</a></span></p>
+
+<div class="caption1">Fishes of the Big Blue River Basin,<br />
+Kansas</div>
+<br />
+<div class="caption3">BY</div>
+
+<div class="caption2">W. L. MINCKLEY</div>
+
+<br />
+<div class="caption2">CONTENTS</div>
+
+<table width="100%" summary="ToC">
+<tr><td>&nbsp;</td><td class="text_rt"><small>PAGE</small></td></tr>
+<tr><td><a href="#INTRODUCTION">Introduction</a></td><td class="text_rt">403</td></tr>
+<tr><td><a href="#ACKNOWLEDGMENTS">Acknowledgments</a></td><td class="text_rt">404</td></tr>
+<tr><td><a href="#TUTTLE_CREEK_DAM_AND_RESERVOIR">Tuttle Creek Dam and Reservoir</a></td><td class="text_rt">404</td></tr>
+<tr><td><a href="#BIG_BLUE_RIVER_BASIN">Big Blue River Basin</a></td><td class="text_rt">404</td></tr>
+<tr><td><a href="#GEOLOGY_OF_THE_BASIN">Geology of the basin</a></td><td class="text_rt">405</td></tr>
+<tr><td><a href="#CLIMATE_POPULATION_AND_LAND-USE">Climate, population, and land-use</a></td><td class="text_rt">406</td></tr>
+<tr><td><a href="#PHYSICAL_FEATURES_OF_STREAMS">Physical features of streams</a></td><td class="text_rt">407</td></tr>
+<tr><td><a href="#PREVIOUS_RECORDS_OF_FISHES">Previous records of fishes</a></td><td class="text_rt">410</td></tr>
+<tr><td><a href="#METHODS_AND_MATERIALS">Methods and materials</a></td><td class="text_rt">410</td></tr>
+<tr><td><a href="#COLLECTING_STATIONS">Collecting stations</a></td><td class="text_rt">412</td></tr>
+<tr><td><a href="#ANNOTATED_LIST_OF_SPECIES">Annotated list of species</a></td><td class="text_rt">414</td></tr>
+<tr><td><a href="#HYBRID_COMBINATIONS">Hybrid combinations</a></td><td class="text_rt">431</td></tr>
+<tr><td><a href="#RELATIVE_ABUNDANCE_AND_DISCUSSION_OF_SPECIES">Relative abundance and discussion of species</a></td><td class="text_rt">431</td></tr>
+<tr><td><a href="#CREEL_CENSUS">Creel census</a></td><td class="text_rt">435</td></tr>
+<tr><td><a href="#RECOMMENDATIONS">Recommendations</a></td><td class="text_rt">437</td></tr>
+<tr><td><a href="#SUMMARY">Summary</a></td><td class="text_rt">438</td></tr>
+<tr><td><a href="#LITERATURE_CITED">Literature cited</a></td><td class="text_rt">438</td></tr>
+</table>
+<br />
+
+<a name="INTRODUCTION" id="INTRODUCTION"></a>
+<div class="caption2">INTRODUCTION</div>
+
+<p>The Big Blue River in northeastern Kansas will soon be impounded
+by the Tuttle Creek Dam, located about five miles north
+of Manhattan, Kansas. Since the inception of this project by the
+U. S. Army Corps of Engineers much argument has arisen as to
+the values of the dam and reservoir as opposed to the values of
+farmland and cultural establishments to be inundated (Schoewe,
+1953; Monfort, 1956; and Van Orman, 1956). Also, there has
+been some concern about the possible effects of impoundment on
+the fish-resources of the area, which supports "a catfish fishery
+that is notable throughout most of the State of Kansas and in some
+neighboring states (U. S. Fish and Wildlife Service, 1953:9)."
+The objectives of my study, conducted from March 30, 1957, to
+<span class="pagenum"><a name="Page_404" id="Page_404">[404]</a></span>
+August 9, 1958, were to record the species of fish present and their
+relative abundance in the stream system, and to obtain a measure
+of angler success prior to closure of the dam. These data may
+be used as a basis for future studies on the fish and fishing in the
+Big Blue River Basin, Kansas.</p>
+
+<a name="ACKNOWLEDGMENTS" id="ACKNOWLEDGMENTS"></a>
+<div class="caption2">ACKNOWLEDGMENTS</div>
+
+<p>I thank Messrs. J. E. Deacon, D. A. Distler, Wallace Ferrel, D. L. Hoyt,
+F. E. Maendele, C. O. Minckley, B. C. Nelson, and J. C. Tash for assistance
+in the field and for valuable suggestions. Dr. J. B. Elder, Kansas State
+College, arranged for loan of specimens, and Mr. B. C. Nelson supplied data
+on <i>Notropis deliciosus</i> (Girard) in Kansas, and on specimens in the University
+of Michigan Museum of Zoology.</p>
+
+<p>I thank the many landowners who allowed me access to streams in the
+Big Blue River Basin. The U. S. Army Corps of Engineers, Kansas City
+District, also allowed access in the reservoir area, and furnished information
+and some photographs. Mr. J. C. Tash did chemical determinations on my
+water samples.</p>
+
+<p>Dr. Frank B. Cross guided me in this study and in preparation of this report.
+Drs. E. Raymond Hall and K. B. Armitage offered valuable suggestions
+on the manuscript. Equipment and funds for my study were furnished by
+the State Biological Survey of Kansas, and the Kansas Forestry, Fish and
+Game Commission granted necessary permits.</p>
+
+<a name="TUTTLE_CREEK_DAM_AND_RESERVOIR" id="TUTTLE_CREEK_DAM_AND_RESERVOIR"></a>
+<div class="caption2">TUTTLE CREEK DAM AND RESERVOIR</div>
+
+<p>The data on Tuttle Creek Dam and Reservoir that follow were
+furnished by Mr. Donald D. Poole, U. S. Army Corps of Engineers,
+Kansas City District. The dam, an earth-fill structure, will be
+7,500 feet in length, with a maximum height of 157 feet
+above the valley floor. Release of water will be from beneath the
+west end of the dam, through two tunnels 20 feet in diameter that
+have a capacity of 45,000 cubic feet per second; however, releases
+exceeding 25,000 c. f. s. are not planned. The gated spillway is
+located at the east end of the dam. Freeboard will be 23 feet
+at the top of flood-control pool.</p>
+
+<p>The reservoir will have a maximum pool of 2,280,000 acre-feet capacity,
+a 53,500-acre surface area, and 368 miles of shoreline.
+The present operational plan provides for a conservation pool
+having a surface area of 15,700 acres, a shoreline of 112 miles,
+and a length of 20 miles.</p>
+
+<a name="BIG_BLUE_RIVER_BASIN" id="BIG_BLUE_RIVER_BASIN"></a>
+<div class="caption2">BIG BLUE RIVER BASIN</div>
+
+<p>Big Blue River and its tributaries, a sub-basin of the Kansas
+River System, drain approximately 9,600 square miles, of which
+2,484 miles are in Kansas (Colby, <i>et al.</i>, 1956:44). The headwaters
+<span class="pagenum"><a name="Page_405" id="Page_405">[405]</a></span>
+of the Big Blue River are in central Hamilton County,
+Nebraska, near the Platte River (<a href="#fig_1">Fig.&nbsp;1</a>). The stream flows generally
+south and east for 283 miles to its confluence with the Kansas
+River near Manhattan, Kansas. Little Blue River, the largest
+tributary to the Big Blue, rises in eastern Kearney and western
+Adams counties, Nebraska, and flows southeast for 208 miles to
+join the Big Blue near Blue Rapids, Kansas (Nebraska State Planning
+Board, 1936:628). The Big Blue River Basin varies in width
+from 129 miles in the northwest, to approximately ten miles near
+the mouth (Colby, <i>et al.</i>, 1956:44).</p>
+
+<a name="GEOLOGY_OF_THE_BASIN" id="GEOLOGY_OF_THE_BASIN"></a>
+<div class="caption2">GEOLOGY OF THE BASIN</div>
+<br />
+<a name="fig_1" id="fig_1"></a>
+<div class="figcenter">
+<img src="images/fig_1.png" width="594" height="486" alt="" title="" /><br />
+<span class="fig_caption"><span class="smcap">Fig. 1.</span> Big Blue River Basin, Kansas and Nebraska.</span>
+</div>
+<br />
+
+<p>In Kansas, outcrops of Pennsylvanian and Cretaceous age occur
+along the extreme eastern and western sides of the Big Blue River
+Basin, respectively, whereas Permian beds (overlain by Pleistocene
+deposits) occur throughout most of the remainder of the watershed
+(see Moore and Landes, 1937). The Big Blue and Little
+Blue rivers and their tributaries have deeply incised the Permian
+beds of the Flint Hills in Kansas, exposing limestones and shales
+<span class="pagenum"><a name="Page_406" id="Page_406">[406]</a></span>
+of the Admire, Council Grove, Chase, and Sumner groups (Wolfcampian
+and Leonardian series) (Walters, 1954:41-44). Pleistocene
+deposits in the Big Blue Basin in Kansas consist of alluvium,
+glacial till, and glacial outwash from the Kansan glacial stage,
+overlain by loess deposits of Wisconsin and Recent stages (Frye
+and Leonard, 1952: pl.&nbsp;1).</p>
+
+<p>The Big Blue River was formed "in part on the till plain surface
+and in part by integration of spillway channels," in the latter portion
+of the Kansan glaciation (Frye and Leonard, 1952:192). This
+stream, and the Republican River to the west, carried waters from
+the areas that are now the Platte, Niobrara, and upper Missouri
+River basins (Lugn, 1935:153). Drainage was southward, through
+Oklahoma, until establishment of the east-flowing Kansas River
+(Frye and Leonard, 1952:189-190). As Kansan ice receded the
+Blue and Republican rivers retained what is now the Platte River
+Basin. The lower Platte River developed and the surface drainage
+became distinct in the Iowan (Tazwellian) portion of the Wisconsin
+glacial stage (Lugn, 1935:152-153). However, according to Lugn
+(1935:203) the Platte River Basin contributes about 300,000 acre-feet
+of water per year to the Big Blue and Republican rivers by
+percolation through sands and gravels underlying the uplands that
+now separate the basins.</p>
+
+<a name="CLIMATE_POPULATION_AND_LAND-USE" id="CLIMATE_POPULATION_AND_LAND-USE"></a>
+<div class="caption2">CLIMATE, POPULATION, AND LAND-USE</div>
+
+<p>Climate of the Big Blue River Basin is of the subhumid continental
+type, with an average annual precipitation of 22 inches
+in the northwest and 30 inches in the southeast. The mean annual
+evaporation from water surfaces exceeds annual precipitation by
+approximately 30 inches (Colby, <i>et al.</i>, 1956:32-33).</p>
+
+<p>The average annual temperature for the basin is 53&deg;&nbsp;F. (Flora,
+1948:148). According to Kincer (1941:704-705) the average
+temperature in July, the warmest month, is 78&deg;&nbsp;F., and the
+coolest month, January, averages 28&deg;&nbsp;F. Periods of extreme cold
+and heat are sometimes of long duration. Length of the growing
+season varies from less than 160 days in the northwest to 180 days
+in the southeast (Kincer, <i>loc. cit.</i>).</p>
+
+<p>The human population of the Big Blue Basin varies from about
+90 persons per square mile in one Nebraska county in the northwest
+and one Kansas county in the southeast, to as few as six
+persons per square mile in some northeastern counties. The
+population is most dense along the eastern border of the basin,
+decreasing toward the west. This decrease in population is correlated
+<span class="pagenum"><a name="Page_407" id="Page_407">[407]</a></span>
+with the decrease in average annual precipitation from east
+to west (Colby, <i>et al.</i>, 1956:80).</p>
+
+<p>The principal land-use in the Big Blue Watershed is tilled
+crops, with wheat, sorghums, and corn being most important.
+Beef cattle are important in some portions of the basin. Colby,
+<i>et al.</i> (1956:24) reported that in 1954 as much as 55 per cent of
+the land in some counties near the mouth of the Big Blue River
+was in pasture. Only one Nebraska county had less than 15 per
+cent in pastureland.</p>
+
+<a name="PHYSICAL_FEATURES_OF_STREAMS" id="PHYSICAL_FEATURES_OF_STREAMS"></a>
+<div class="caption2">PHYSICAL FEATURES OF STREAMS</div>
+
+<p>Streams of the Big Blue River Basin are of three kinds: turbid,
+sandy-bottomed streams, usually 150 to 300 feet in width; relatively
+clear, mud-bottomed streams, ten to 60 feet in width; and clear,
+deeply incised, gravel-bottomed streams, usually five to 30 feet
+in width.</p>
+
+<p><span class="smcap">Sand-bottomed Streams.</span>&mdash;The Big Blue and Little Blue rivers
+represent this kind of stream. The bottoms of these rivers consist
+almost entirely of fine sand; nevertheless, their channels are primarily
+deep and fairly uniform in width, rather than broad, shallow,
+and braided as in the larger Kansas and Arkansas rivers in
+Kansas (<a href="#pl11fig1">Plate 11, Fig.&nbsp;1</a>). In the Big Blue River, gravel occurs
+rarely on riffles, and gravel-rubble bottoms are found below dams
+(<a href="#pl11fig2">Plate 11, Fig.&nbsp;2</a>). The Big Blue flows over a larger proportion
+of gravelly bottom than does the Little Blue.</p>
+
+<p>Big Blue River rises at about 1,800 feet above mean sea level
+and joins the Kansas River at an elevation of 1,000 feet above
+m. s. l. The average gradient is 2.8 feet per mile. Little Blue River,
+originating at 2,200 feet, has an average gradient of 5.3 feet per
+mile, entering the Big Blue at 1,100 feet above mean sea level
+(Nebraska State Planning Board, 1936:628, 637). The Little Blue
+is the shallower stream, possibly because of the greater amount
+of sandy glacial deposits in its watershed and the swift flow that
+may cause lateral cutting, increased movement, and "drifting"
+of the sandy bottom.</p>
+
+<p>For approximately a 50-year period, stream-flow in the Big Blue
+River at its point of entry into Kansas (Barnston, Nebraska)
+averaged 603 cubic feet per second, with maximum and minimum
+instantaneous flows of 57,700 c. f. s. and one c. f. s. The Little Blue
+River at Waterville, Kansas, averaged a daily discharge of 601
+c. f. s. (maximum 50,400, minimum 28). Below the confluence
+of the Big Blue and Little Blue rivers, at Randolph, Kansas, the
+<span class="pagenum"><a name="Page_408" id="Page_408">[408]</a></span>
+average daily discharge was 1,690 c.f.s. (maximum 98,000, minimum
+31) (Kansas Water Resources Fact-finding and Research
+Committee, 1955:27).</p>
+
+<p>The turbidity of the Big Blue River, as determined by use of
+a Jackson turbidimeter, varied from 27 parts per million in winter
+(January 10, 1958) to as high as 14,000 p.p.m. (July 12, 1958).
+The Little Blue River has similar turbidities, with high readings
+being frequent. In the summer of 1957, pH ranged from 7.2 to
+8.4 in the Big Blue River Basin&mdash;values that correspond closely
+with those of Canfield and Wiebe (1931:3) who made 25 determinations
+ranging from 7.3 to 8.3 in the streams of the Nebraskan
+portion of this basin in July, 1930. Surface temperatures at various
+stations varied from 38&deg;&nbsp;F. on January 10, 1958, to 90&deg;&nbsp;F.
+in backwater-areas on July 19, 1957. The average surface temperature
+at mid-day in July and August, 1957, was approximately
+86.5&deg;&nbsp;F.</p>
+
+<p>Chemical determinations were made on water-samples from
+my Station 4-S on the Big Blue River, and Station 50-S on the
+Little Blue (<a href="#Table_1">Table&nbsp;1</a>). These samples were taken from the surface
+in strong current. Determinations were made by methods described
+in <i>Standard Methods for the Examination of Water and Sewage</i>,
+10th edition, 1955.</p>
+
+<a name="Table_1" id="Table_1"></a>
+<div class="smcap_tbl">Table 1.&mdash;Chemical Determinations in Milligrams Per Liter at Five
+Stations in the Big Blue River Basin, Kansas, 1958.</div>
+<br />
+<table width="100%" summary="Water Sample Chemical Data">
+<tr>
+<td class="bt2 bb smcap center">Station<br />and<br />Date</td>
+<td class="bt2 bb bl center"><ins title='Correction: was "Phenophthalein"'>Phenolphthalein</ins>&nbsp; alkalinity</td>
+<td class="bt2 bb bl center">Methyl-orange alkalinity</td>
+<td class="bt2 bb bl center">Chlorides</td>
+<td class="bt2 bb bl center">Sulphates</td>
+<td class="bt2 bb bl center">Nitrates</td>
+<td class="bt2 bb bl center">Nitrites</td>
+<td class="bt2 bb bl center">Ammonia</td>
+<td class="bt2 bb bl center">Phosphate</td>
+</tr>
+<tr>
+<td class="center">4-S<br />August&nbsp;9........</td>
+<td class="bl center">0.0</td>
+<td class="bl center">154</td>
+<td class="bl center">16</td>
+<td class="bl center">28</td>
+<td class="bl center">3.5</td>
+<td class="bl center">.083</td>
+<td class="bl center">.250</td>
+<td class="bl center">.225</td>
+</tr>
+<tr>
+<td class="center">50-S<br />August&nbsp;9........</td>
+<td class="bl center">0.0</td>
+<td class="bl center">125</td>
+<td class="bl center">24</td>
+<td class="bl center">20</td>
+<td class="bl center">2.5</td>
+<td class="bl center">.669</td>
+<td class="bl center">.427</td>
+<td class="bl center">.240</td>
+</tr>
+<tr>
+<td class="center">35-M<br />August&nbsp;9........</td>
+<td class="bl center">0.0</td>
+<td class="bl center">366</td>
+<td class="bl center">15</td>
+<td class="bl center">108</td>
+<td class="bl center">9.4</td>
+<td class="bl center">.220</td>
+<td class="bl center">.750</td>
+<td class="bl center">.080</td>
+</tr>
+<tr>
+<td class="center">11-G<br />July&nbsp;8........</td>
+<td class="bl center">0.0</td>
+<td class="bl center">272</td>
+<td class="bl center">15</td>
+<td class="bl center">60</td>
+<td class="bl center">4.5</td>
+<td class="bl center">.060</td>
+<td class="bl center">.625</td>
+<td class="bl center">.140</td>
+</tr>
+<tr>
+<td class="bb center">18-G<br />July&nbsp;22........</td>
+<td class="bb bl center">0.0</td>
+<td class="bb bl center">183</td>
+<td class="bb bl center">10</td>
+<td class="bb bl center">60</td>
+<td class="bb bl center">1.6</td>
+<td class="bb bl center">.938</td>
+<td class="bb bl center">.293</td>
+<td class="bb bl center">.240</td>
+</tr>
+</table>
+<br />
+
+<p><span class="pagenum"><a name="Page_409" id="Page_409">[409]</a></span></p>
+
+<p>The banks of both the Big Blue and Little Blue rivers support
+narrow riparian forests comprised primarily of elm, <i>Ulmus americanus</i>,
+cottonwood, <i>Populus deltoides</i>, sycamore, <i>Platanus occidentalis</i>,
+and willow, <i>Salix</i> spp. Maple, <i>Acer</i> sp., oak, <i>Quercus</i> spp.,
+and ash, <i>Fraxinus</i> sp. occur where the rivers flow near steep, rocky
+hillsides. Many of the hills are virgin bluestem prairies (<i>Andropogon</i>
+spp.), but the floodplains are heavily cultivated.</p>
+
+<p><span class="smcap">Mud-bottomed Streams.</span>&mdash;Streams of this kind are present in
+the watershed of the Black Vermillion River that enters Big Blue
+River from the east. The area east of the Big Blue River and
+north of the Black Vermillion River is till plains, where relief seldom
+exceeds 100 feet (Walters, 1954:12). Streams in this portion
+of the basin, and streams entering the Little Blue River from the
+west (Mill Creek and Horseshoe Creek systems), tend to have
+V-shaped channels, fewer riffles than the Little Blue and Big Blue
+rivers and in the gravelly streams (to be described later), and
+have bottoms of mud or clay, with few rocks (<a href="#pl12fig1">Plate 12, Fig.&nbsp;1</a>).
+However, in the extreme headwaters of most western tributaries
+of the Little Blue River (in Washington and Republic counties)
+sandy bottoms predominate. The Black Vermillion River flows
+on a broad floodplain and is a mud-bottomed, sluggish stream,
+with an average gradient of approximately one foot per mile.
+Fringe-forests of elm, cottonwood, sycamore, and willow persist
+along most of these stream-courses.</p>
+
+<p>Notwithstanding the mud bottoms, the water in this kind of
+stream in the Big Blue Basin remains clearer than that of the Big
+Blue and Little Blue rivers. Heavy algal blooms were noted in
+the Black Vermillion River and Mill Creek, Washington County,
+in 1957 and 1958. Temperatures at Stations 45-M and 46-M on
+Mill Creek, Washington County, averaged 85.5&deg;&nbsp;F. on July 31, 1957.
+Chemical characteristics of a water-sample from Station 35-M,
+Black Vermillion River, are in <a href="#Table_1">Table&nbsp;1</a>.</p>
+
+<p><span class="smcap">Gravel-bottomed Streams.</span>&mdash;Most streams of this kind are tributary
+to the Big Blue River; however, streams entering Black Vermillion
+River from the south are also of this type (<a href="#pl12fig2">Plate 12, Fig.&nbsp;2</a>).
+The streams are "characteristically a series of large pools (to 100
+feet in length and more than two feet in depth) connected by short
+riffles and smaller pools" (Minckley and Cross, in press). The
+average gradients are high: Carnahan Creek, 33 feet per mile;
+Mill Creek, Riley County, 21 feet; Clear Creek, 16 feet per mile.
+Stream-flow is usually less than five cubic feet per second. In summer,
+these streams may become intermittent, but springs and
+<span class="pagenum"><a name="Page_410" id="Page_410">[410]</a></span>
+subsurface percolation maintain pool-levels (Minckley and Cross,
+<i>loc. cit.</i>).</p>
+
+<p>The average temperatures of these small streams (79.5&deg;&nbsp;to
+81.0&deg;&nbsp;F. in July and August, 1957) were lower than temperatures
+in stream-types previously described. Turbidities were usually
+less than 25 p.p.m. The chemical properties of water-samples from
+two of these streams (Stations 11-G and 18-G) are listed in <a href="#Table_1">Table&nbsp;1</a>.</p>
+
+<a name="PREVIOUS_RECORDS_OF_FISHES" id="PREVIOUS_RECORDS_OF_FISHES"></a>
+<div class="caption2">PREVIOUS RECORDS OF FISHES</div>
+
+<p>The earliest records of fishes from the Big Blue River Basin are those of
+Cragin (1885) and Graham (1885) in independently published lists of the
+fishes of Kansas. Meek (1895) recorded fishes collected in 1891 "from both
+branches of the Blue River, a few miles west of Crete, Nebraska." Evermann
+and Cox (1896) reported five collections from the Nebraskan part of the
+basin. Their collections were made in October, 1892, and August, 1893, and
+the stations were: in 1892, Big Blue River at Crete; in 1893, Big Blue River
+at Seward, Lincoln Creek at Seward and York, and Beaver Creek at York.</p>
+
+<p>Canfield and Wiebe (1931) obtained fish from 18 localities in Nebraska
+in July, 1930; however, their major concern was determination of water quality.
+Their stations were: Big Blue River at Stromsburg, Polk Co.; Surprise and
+Ulysses, Butler Co.; Staplehurst, Seward, and Milford, Seward Co.; Crete and
+Wilber, Saline Co.; Beatrice, Blue Springs, and Barnston, Gage Co.; Little Blue
+River at Fairbury, Jefferson Co.; Hebron, Thayer Co.; Sandy Creek at Alexandria,
+Thayer Co.; West Fork of Big Blue River at Stockham, Hamilton Co.;
+McCool Junction, York Co.; Beaver Crossing, Seward Co.; and Beaver Creek
+at York, York Co.</p>
+
+<p>Breukelman (1940) and Jennings (1942) listed fishes from the University
+of Kansas Museum of Natural History and the Kansas State College Museum,
+respectively, including some specimens collected from the Big Blue River
+System in Kansas. Because records in these two papers pertain to collections
+that were widely spaced in the basin and in time, the specific localities are not
+given herein. One of Jennings' (<i>loc. cit.)</i> records, <i>Scaphirhynchus platorynchus</i>
+(Rafinesque), was cited by Bailey and Cross (1954:191). More recently,
+Minckley and Cross (in press) recorded several localities, and cited some
+papers mentioned above, in a publication dealing with <i>Notropis topeka</i>
+(Gilbert) in Kansas.</p>
+
+<p>Information on the fishes of the Nebraskan portion of the Big Blue River
+Basin was compiled, and additional localities were reported, in a doctoral
+thesis by Dr. Raymond E. Johnson, entitled The Distribution of Nebraska
+Fishes, 1942, at the University of Michigan.</p>
+
+<a name="METHODS_AND_MATERIALS" id="METHODS_AND_MATERIALS"></a>
+<div class="caption2">METHODS AND MATERIALS</div>
+
+<div class="caption3"><i>Collection of Fishes</i></div>
+
+<p>The gear and techniques used are listed below:</p>
+
+<p><span class="smcap">Entrapment Devices.</span>&mdash;Hoop and fyke nets and wire traps were used
+for 288 trap/net hours in 1957. The nets were not baited, and were set
+parallel to the current, with the mouths downstream. Hoop nets were 1&frac12; to
+three feet in diameter at the first hoop, with a pot-mesh of one inch; fyke nets
+<span class="pagenum"><a name="Page_411" id="Page_411">[411]</a></span>
+were three feet at the first hoop, pot-mesh of one inch; wire traps, with an
+opening at each end, were 2&frac12; feet in diameter and covered with one-inch-mesh,
+galvanized chicken wire.</p>
+
+<p><span class="smcap">Gill Nets.</span>&mdash;Experimental gill nets were set on three occasions in areas
+with little current. These nets were 125 feet in length, with &frac34; to two inch
+bar-mesh in 25-foot sections.</p>
+
+<p><span class="smcap">Seines.</span>&mdash;Seining was used more than other methods. An attempt was
+made to seine all habitats at each station. In swift water, seine-hauls were
+usually made downstream, but in quiet areas seining was done randomly.
+Haul-seines six to 60 feet in length, three to eight feet in depth, and with
+meshes of &#8539; to &frac12; inch were used. For collection of riffle-fishes, the seine was
+planted below a selected area and the bottom was kicked violently by one
+member of the party, while one or two persons held the seine, raising it when
+the area had been thoroughly disturbed. Seining on riffles was done with a
+four-foot by four-foot bobbinet seine.</p>
+
+<p><span class="smcap">Rotenone.</span>&mdash;Rotenone was used in pools of smaller streams, mouths of
+creeks, borrow-pits, and cut-off areas. Both powdered and emulsifiable
+rotenone were used. The rotenone was mixed with water and applied by
+hand, or into the backwash of an outboard motor.</p>
+
+<p><span class="smcap">Electric Shocker.</span>&mdash;The electrical unit used in this study generated 115
+volts and 600 to 700 watts, alternating current. The shocking unit consisted
+of two booms, each with two electrodes, mounted on and operated from a
+slowly moving boat. Fish were recovered in scape nets, or in many cases
+were identified as they lay stunned and were not collected.</p>
+
+<div class="caption3"><i>Estimation of Relative Abundance</i></div>
+
+<p>Data on relative abundance of fishes were obtained by counts of seine
+hauls at 29 of the 59 stations, counts of rotenoned fish at seven stations, and
+results with the electric shocker at nine stations. Counts were usually made
+in the field; however, in some collections all fish were preserved and counted
+in the laboratory. Some fish (or "swirls" presumed to be fish) observed
+while shocking were not identified and are not included in the calculations.
+However, all fish positively identified while shocking are included.</p>
+
+<div class="caption3"><i>Age and Growth of Fishes</i></div>
+
+<p>Fish from selected size-groups were aged in this study. Scales for age-determinations
+were removed from positions recommended by Lagler (1952:108).
+Scales were placed in water between glass slides and were read on a
+standard scale-projection device.</p>
+
+<p>Pectoral spines of catfish were removed from one or both sides, sectioned,
+and read by methods described by Marzolf (1955:243-244).</p>
+
+<p>Calculation of length at the last annulus for both scale-fish and catfish
+was made by direct proportion. All measurements are of total length to the
+nearest tenth of an inch unless specified otherwise.</p>
+
+<div class="caption3"><i>Creel Census</i></div>
+
+<p>From April 6 to May 28, 1957, a creel census was taken below Turtle Creek
+Dam. From June 16 to July 24, 1958, I periodically visited the main points of
+access to the Big Blue River, beginning approximately eight miles downstream
+from Tuttle Creek Dam and ending six miles upstream from the maximal extension
+<span class="pagenum"><a name="Page_412" id="Page_412">[412]</a></span>
+of the reservoir at capacity level. Access-points consisted of 11 bridges,
+two power dams, and three areas where county roads approached the river.
+Eleven eight-hour days were spent in the 1957 census and 22 checks in 15 days
+were made in 1958. An equal number of morning (6:00 a.m. to 12:00 noon)
+and afternoon (12:00 noon to 8:30 p.m.) checks were made.</p>
+
+<p>Fishermen contacted were asked the following questions: home address (or
+residence at the time of the fishing trip); time they started fishing; kind of fish
+sought; number and kinds of fish in possession; and baits used. Also, the
+number of poles and type of fishing (from the bank, from boat, <i>etc.</i>) were
+recorded. Fishes caught were examined to confirm identifications. About 80
+per cent of all fishermen seen were contacted.</p>
+
+<p>Fish per man-hour, as used in this report, refers to the average number of
+fish of all species caught by one fisherman in one hour. Fisherman-day is the
+average time spent fishing in one day by one person. Because some fishermen
+used more than one pole, the data are also expressed as catch per pole-hour.</p>
+
+<a name="COLLECTING_STATIONS" id="COLLECTING_STATIONS"></a>
+<div class="caption2">COLLECTING STATIONS</div>
+
+<p>In the list that follows, stations are numbered consecutively from the mouth
+of the Big Blue River, listing stations on each tributary as it is ascended. The
+letters following station-numbers indicate the general type of stream: S =
+sandy; M = muddy; and G = gravelly. The Big Blue River is the boundary
+between Riley and Pottawatomie counties, Kansas, along part of its length.
+Stations in this area have been designated Riley County. The legal description
+of each station is followed by the date(s) of collection, and each station is
+plotted in <a href="#fig2">Figure&nbsp;2</a>.</p>
+
+<a name="fig2" id="fig2"></a>
+<div class="figcenter">
+<img src="images/fig_2.png" width="601" height="429" alt="" title="" /><br />
+<span class="fig_caption"><span class="smcap">Fig. 2.</span> Collection stations in the Big Blue River Basin, Kansas, 1957 and 1958.</span>
+</div>
+<br />
+<br />
+
+<div class="caption1">PLATE 11</div>
+<br />
+<div class="figcenter" style="width:559px">
+<a name="pl11fig1" id="pl11fig1"></a>
+<img src="images/pl_11_fig_1.png" width="559" height="309" alt="" title="" /><br />
+<span class="fig_caption"><span class="smcap">Fig. 1.</span> Big Blue River at Station 3-S. U.S. Army Corps of Engineers photograph
+No. 563697.</span>
+</div>
+<br />
+<br />
+
+<div class="figcenter" style="width:560px">
+<a name="pl11fig2" id="pl11fig2"></a>
+<img src="images/pl_11_fig_2.png" width="560" height="372" alt="" title="" /><br />
+<span class="fig_caption"><span class="smcap">Fig. 2.</span> Big Blue River at Oketo, Marshall County, Kansas. U.S. Army Corps
+of Engineers, photograph No. 67516.</span>
+</div>
+<br />
+<br />
+
+<div class="caption1">PLATE 12</div>
+<br />
+<div class="figcenter" style="width:531px">
+<a name="pl12fig1" id="pl12fig1"></a>
+<img src="images/pl_12_fig_1.png" width="531" height="352" alt="" title="" /><br />
+<span class="fig_caption"><span class="smcap">Fig. 1.</span> Black Vermillion River, approximately one mile upstream from its
+mouth. Photograph by Robert G. Webb.</span>
+</div>
+<br />
+<br />
+
+<div class="figcenter" style="width:526px">
+<a name="pl12fig2" id="pl12fig2"></a>
+<img src="images/pl_12_fig_2.png" width="526" height="352" alt="" title="" /><br />
+<span class="fig_caption"><span class="smcap">Fig. 2.</span> Carnahan Creek at Station 11-G. Photograph by Robert G. Webb.</span>
+</div>
+<br />
+<br />
+
+<p><span class="pagenum"><a name="Page_413" id="Page_413">[413]</a></span></p>
+
+<div class="reference">1-S: Pottawatomie Co., mouth of Big Blue River, Sec. 16, T. 10S, R. 8E,
+June 20, 1958.</div>
+
+<div class="reference">2-S: Riley Co., Big Blue River, Sec. 4, T. 10S, R. 8E, June 6, 12, and 14,
+1957.</div>
+
+<div class="reference">3-S: Riley Co., Big Blue River, E ½, Sec. 30, T. 9S, R. 8E, Mar. 30, Apr. 6,
+July 15, 16, 17, Aug. 14, and Dec. 26, 1957; Apr. 26, June 20, and Aug.
+5, 1958.</div>
+
+<div class="reference">4-S: Riley Co., Big Blue River at Rocky Ford Dam, W ½, Sec. 30, T. 9S,
+R. 8E, Aug. 14, 1957; and Aug. 5, 1958.</div>
+
+<div class="reference">5-G: Pottawatomie Co., McIntire Creek, Sec. 12, T. 9S, R. 7E, July 14, 1958.</div>
+
+<div class="reference">6-S: Riley Co., Big Blue River and adjacent borrow-pit, Sec. 24, T. 9S, R.
+7E, July 18 and 19, 1957; and July 11, 1958.</div>
+
+<div class="reference">7-G: Riley Co., Tuttle Creek, Sec. 10, T. 9S, R. 7E, Aug. 5, 1958.</div>
+
+<div class="reference">8-S: Riley Co., Big Blue River, Sec. 10, T. 9S, R. 7E, Aug. 14, 1957.</div>
+
+<div class="reference">9-G: Riley Co., Mill Creek, Sec. 4, T. 9S, R. 7E, July 20 and 25, 1958.</div>
+
+<div class="reference">10-G: Riley Co., Mill Creek, Sec. 2, T. 9S, R. 6E, Aug. 13, 1957.</div>
+
+<div class="reference">11-G: Pottawatomie Co., Carnahan Creek, Sec. 22, 27, and 34, T. 8S, R. 7E,
+Aug. 1, 1957; and July 8, 1958.</div>
+
+<div class="reference">12-G: Pottawatomie Co., unnamed tributary to Carnahan Creek, Sec. 15, T.
+8S, R. 7E, Mar. 19, 1956 (collection made before my formal study was
+begun).</div>
+
+<div class="reference">13-G: Pottawatomie Co., Carnahan Creek, Sec. 36, T. 7S, R. 7E, Aug. 13,
+1957.</div>
+
+<div class="reference">14-S: Riley Co., Big Blue River, Sec. 18, T. 8S, R. 7E, Mar. 22, 1958.</div>
+
+<div class="reference">15-S: Riley Co., Big Blue River, Sec. 7, T. 8S, R. 7E, Apr. 3, and June 12,
+1958.</div>
+
+<div class="reference">16-G: Riley Co., unnamed creek, Sec. 1, T. 8S, R. 6E, July 10, and Aug. 5,
+1958.</div>
+
+<div class="reference">17-G: Riley Co., unnamed creek, Sec. 10, T. 8S, R. 6E, June 26, 1958.</div>
+
+<div class="reference">18-G: Riley Co., Fancy Creek, Sec. 14, T. 7S, R. 6E, July 29, 1957.</div>
+
+<div class="reference">19-G: Riley Co., Walnut Creek, Sec. 20, T. 7S, R. 6E, June 26, 1958.</div>
+
+<div class="reference">20-G: Riley Co., Fancy Creek, Sec. 2, T. 7S, R. 5E, Mar. 13, 1957; and June
+26, 1958.</div>
+
+<div class="reference">21-G: Riley Co., Schoolhouse Branch, Sec. 35, T. 6S, R. 5E, July 22, 1958.</div>
+
+<div class="reference">22-G: Riley Co., Fancy Creek, Sec. 33, T. 6S, R. 5E, June 1, 1957.</div>
+
+<div class="reference">23-G: Riley Co., West Branch Fancy Creek, Sec. 32 and 33, T. 6S, R. 5E, June
+1 and 3, 1957.</div>
+
+<div class="reference">24-G: Clay Co., West Branch Fancy Creek, Sec. 32 and 33, T. 6S, R. 4E, July
+22, 1958.</div>
+
+<div class="reference">25-S: Riley Co., Big Blue River, Sec. 5, T. 7S, R. 7E, Aug. 7, 1958.</div>
+
+<div class="reference">26-G: Riley Co., Swede Creek, Sec. 21, T. 6S, R. 7E, Mar. 22, 1958.</div>
+
+<div class="reference">27-G: Pottawatomie Co., unnamed creek, Sec. 14, T. 6S, R. 7E, Sept. 10, 1957.</div>
+
+<div class="reference">28-G: Pottawatomie Co., Bluff Creek, Sec. 6, T. 6S, R. 8E, Oct. 6, 1957.</div>
+
+<div class="reference">29-G: Pottawatomie Co., Bluff Creek, Sec. 15, T. 6S, R. 8E, June 29, 1958.</div>
+
+<div class="reference">30-M: Marshall Co., Black Vermillion River, Sec. 9, T. 5S, R. 8E, Mar. 5,
+1958.</div>
+
+<div class="reference">31-G: Pottawatomie Co., Clear Creek, Sec. 3, T. 6S, R. 9E, July 14, 1958.</div>
+
+<div class="reference">32-G: Pottawatomie Co., unnamed creek, Sec. 14, T. 6S, R. 9E, July 14, 1958.</div>
+
+<div class="reference">33-M: Marshall Co., Robidoux Creek, Sec. 20, T. 2S, R. 9E, July 23, 1958.</div>
+
+<div class="reference">34-M: Marshall Co., Little Timber Creek, Sec. 10, T. 4S, R. 9E, Oct. 6, 1957.</div>
+
+<div class="reference">35-M: Marshall Co., Black Vermillion River, Sec. 15, T. 4S, R. 9E, Aug. 9,
+1958.</div>
+
+<div class="reference">36-M: Marshall Co., unnamed creek, Sec. 8, T. 4S, R. 9E, Oct. 6, 1957.</div>
+
+<div class="reference">37-M: Marshall Co., Black Vermillion River, Sec. 11, T. 4S, R. 10E, Oct. 6,
+1957.<span class="pagenum"><a name="Page_414" id="Page_414">[414]</a></span></div>
+
+<div class="reference">38-S: Marshall Co., Big Blue River, Sec. 18, T. 5S, R. 8E, Aug. 8, 1958.</div>
+
+<div class="reference">39-S: Marshall Co., Big Blue River, Sec. 20, T. 4S, R. 7E, May 29, 1958.</div>
+
+<div class="reference">40-M: Washington Co., Coon Creek, Sec. 27, T. 4S, R. 4E, July 22, 1958.</div>
+
+<div class="reference">41-S: Marshall Co., Little Blue River, Sec. 9, 16, and 17, T. 4S, R. 6E, June
+27, 1958.</div>
+
+<div class="reference">42-S: Washington Co., Little Blue River, Sec. 21, T. 3S, R. 5E, Aug. 8, 1958.</div>
+
+<div class="reference">43-S: Washington Co., Little Blue River, Sec. 5 and 8, T. 3S, R. 5E, July 30,
+1957.</div>
+
+<div class="reference">44-S: Washington Co., Little Blue River, Sec. 36, T. 1S, R. 4E, July 31, 1957.</div>
+
+<div class="reference">45-M: Washington Co., Mill Creek, Sec. 35 and 36, T. 1S, R. 4E, July 31,
+1957.</div>
+
+<div class="reference">46-M: Washington Co., Mill Creek, Sec. 4, T. 2S, R. 4E, July 31, 1957.</div>
+
+<div class="reference">47-M: Washington Co., Spring Creek, Sec. 11 and 12, T. 2S, R. 3E, June 19,
+1958.</div>
+
+<div class="reference">48-M: Washington Co., Mill Creek, Sec. 28, T. 2S, R. 2E, June 19, 1958.</div>
+
+<div class="reference">49-M: Republic Co., Mill Creek, Sec. 8 and 17, T. 2S, R. 1W, July 23, 1958.</div>
+
+<div class="reference">50-S: Washington Co., Little Blue River, Sec. 5, T. 1S, R. 4E, Aug. 9, 1958.</div>
+
+<div class="reference">51-M: Republic Co., Rose Creek, Sec. 20, T. 1S, R. 2W, July 23, 1958.</div>
+
+<div class="reference">52-S: Marshall Co., Big Blue River, Sec. 6, T. 4S, R. 7E, Aug. 6, 1958.</div>
+
+<div class="reference">53-S: Marshall Co., Big Blue River, Sec. 18, T. 3S, R. 7E, July 29 and 30,
+1957; May 28, and Aug. 6, 1958.</div>
+
+<div class="reference">54-G: Marshall Co., Hop Creek, Sec. 13 and 18, T. 3S, R. 7E, May 28, 1958.</div>
+
+<div class="reference">55-M: Marshall Co., Spring Creek, Sec. 29, T. 2S, R. 8E, July 9, 1958.</div>
+
+<div class="reference">56-S: Marshall Co., Big Blue River at Marysville Dam, Sec. 20, T. 2S, R. 7E,
+June 16, 1958.</div>
+
+<div class="reference">57-M: Marshall Co., Horseshoe Creek, Sec. 6, T. 2S, R. 7E, July 1, 1958.</div>
+
+<div class="reference">58-G: Marshall Co., unnamed creek, Sec. 2, T. 1S, R. 7E, July 1, 1958.</div>
+
+<div class="reference">59-G: Marshall Co., Mission Creek, Sec. 3, T. 1S, R. 8E, Nov. 30, 1957.</div>
+
+<a name="ANNOTATED_LIST_OF_SPECIES" id="ANNOTATED_LIST_OF_SPECIES"></a>
+<div class="caption2">ANNOTATED LIST OF SPECIES</div>
+
+<p>Forty-eight species were obtained in this survey and five others
+have been recorded in literature or are deposited in museums:
+KSC = Kansas State College Museum; and UMMZ = University
+of Michigan Museum of Zoology. Specimens, unless designated
+otherwise, are in the University of Kansas Museum of Natural
+History (KU).</p>
+
+<p>In this list, the scientific name of each species is followed by
+the common name, citations of previous records, and the stations
+where the species was obtained. I follow Bailey (1956:328-329)
+in treating <i>Lepisosteus osseus</i> (Linnaeus), <i>Catostomus commersonnii</i>
+(Lacépède), <i>Semotilus atromaculatus</i> (Mitchill), <i>Notropis
+lutrensis</i> (Baird and Girard), <i>Pimephales promelas</i> Rafinesque, <i>Ictalurus
+melas</i> (Rafinesque), <i>Ictalurus punctatus</i> (Rafinesque), and
+<i>Lepomis macrochirus</i> Rafinesque, in binomial form only.</p>
+
+<p><b><i>Scaphirhynchus platorynchus</i></b> (Rafinesque), shovelnose sturgeon:
+<span class="pagenum"><a name="Page_415" id="Page_415">[415]</a></span>
+Jennings (1942:364) as <i>Scaphirhynchus platorhynchus</i> (Rafinesque);
+Bailey and Cross (1954:191). Stations 3-S and 4-S.</p>
+
+<p>Shovelnose sturgeon were found only in the lower portion of the
+Big Blue River. On April 20, 1957, many were seen in fishermen's
+creels at Stations 3-S and 4-S. One male and two females that I
+examined on that date were ripe or nearly so; eggs seemed well
+developed and milt flowed freely from the male. After April, 1957,
+none was collected or observed until April 26, 1958, when one
+specimen was obtained while shocking. Forbes and Richardson
+(1920:27) reported that shovelnose sturgeon spawn in Illinois
+between April and June, and Eddy and Surber (1947:80) reported
+spawning in May and early June in Wisconsin and Minnesota.</p>
+
+<p><b><i>Lepisosteus platostomus</i></b> Rafinesque, shortnose gar: Jennings
+(1942:364). Stations 3-S and 4-S.</p>
+
+<p>I saw shortnose gar at various times in 1956 and 1957 at Rocky
+Ford Dam on the Big Blue River (Station 4-S). One was seen
+while shocking at Station 3-S on December 26, 1957.</p>
+
+<p><b><i>Lepisosteus osseus</i></b> (Linnaeus), longnose gar: Jennings (1942:364)
+as <i>Lepisosteus osseus oxyurus</i> Rafinesque. Stations 1-S, 2-S,
+3-S, 4-S, 6-S, 8-S, 9-G, 15-S, 18-G, 25-S, 41-S, 44-S, 52-S, and 53-S.</p>
+
+<p>Longnose gar were abundant in the mainstream of the Big Blue
+River but usually evaded capture. This species, and the shortnose
+gar, resided in the larger rivers, with <i>L. osseus</i> being taken in only
+two creeks near their mouths. In periods of high water, gar moved
+into the flooded creeks, but returned to the river as stream-levels
+subsided.</p>
+
+<p>Young-of-the-year <i>L. osseus</i>, averaging 21.5 mm. in total length
+(range 13 to 30 mm.), were taken on June 14, 1957, and larger young
+(estimated 60 to 70 mm. total length) were taken on June 27, 1958.</p>
+
+<p><b><i>Dorosoma cepedianum</i></b> (LeSueur), gizzard shad: Jennings
+(1942:364). Stations 1-S, 3-S, 4-S, 6-S, 8-S, 44-S, 45-M, and 53-S.</p>
+
+<p>Most gizzard shad were young-of-the-year, taken on July 16 and
+17, 1957, at Stations 3-S and 4-S. Twenty specimens from Station
+6-S that were in their second summer of life were from 3.8 to 5.9
+inches total length at the last annulus (average 4.3). This species
+was usually found in quiet water and was most abundant near the
+mouth of the Big Blue River.</p>
+
+<p><b><i>Hiodon alosoides</i></b> (Rafinesque), goldeye. Stations 3-S, 4-S, and
+53-S.</p>
+
+<p>I caught five specimens of <i>H. alosoides</i> from the Big Blue River,
+<span class="pagenum"><a name="Page_416" id="Page_416">[416]</a></span>
+and another specimen, obtained by Dr. R. B. Moorman in 1954, is
+at Kansas State College (KSC 4984).</p>
+
+<p>One goldeye that I caught on April 20, 1956, prior to the beginning
+of my study, was a ripe female measuring 15.5 inches total
+length. The fish was beginning its seventh summer of life.</p>
+
+<p><b><i>Cycleptus elongatus</i></b> LeSueur, blue sucker. The blue sucker is
+included on the basis of a single specimen (KSC 2917) collected
+by I. D. Graham and labeled "Blue River." No other data are with
+the specimen; however, most fishes deposited at Kansas State College
+by Graham are dated "1885" or "1886" and were caught near
+"Manhattan" (Riley County).</p>
+
+<p><b><i>Ictiobus cyprinella</i></b> (Valenciennes), bigmouth buffalo. Stations
+3-S, 6-S, and 30-M.</p>
+
+<p>Bigmouth buffalo were rare, and were taken only in quiet parts
+of larger streams, and in the borrow-pit at Station 6-S.</p>
+
+<p><b><i>Ictiobus niger</i></b> (Rafinesque), black buffalo. Stations 3-S, 41-S,
+and 53-S.</p>
+
+<p>Only four individuals of <i>I. niger</i> were taken. All were large
+adults (more than 20 inches in total length), and all were shocked
+in the deeper, swifter areas, where the channel narrowed.</p>
+
+<p><b><i>Ictiobus bubalus</i></b> (Rafinesque), smallmouth buffalo. Stations
+1-S, 3-S, 6-S, 7-G, 18-G, 38-S, 41-S, 43-S, 46-M, and 53-S.</p>
+
+<p>This species was found in relatively quiet waters in the main
+channel, in cut-off areas, and in creek-mouths. The ages and total
+lengths of 30 individuals obtained at Station 6-S were (average
+followed by number of fish in parentheses): I, 2.4 (11); II, 4.4
+(14); and III, 6.6 (5).</p>
+
+<p>Canfield and Wiebe (1931:6-7, 10) recorded "buffalo-fish" and
+"buffalo" from the Big Blue Basin in Nebraska; however, no specific
+designation was given.</p>
+
+<p><b><i>Carpiodes forbesi</i></b> Hubbs, plains carpsucker. Station 3-S.</p>
+
+<p>This represents the first record known to me of the plains carpsucker
+from Kansas. The specimen (KU 4180), 430 mm. in standard
+length, has the following characters: lower lip without a
+median, nipple-like projection; dorsal fin-rays, 25; lateral-line scales,
+38; diameter of orbit into distance from anterior nostril to tip of
+snout, 1.1; body-depth into standard length, 3.3; and head-length
+into standard length, 3.9. The specimen was taken while shocking
+a wide, shallow channel, over sand bottom.</p>
+
+<p><b><i>Carpiodes carpio carpio</i></b> (Rafinesque), river carpsucker: Jennings
+(1942:364). Stations 1-S, 2-S, 3-S, 4-S, 5-G, 6-S, 7-G, 8-S,
+<span class="pagenum"><a name="Page_417" id="Page_417">[417]</a></span>
+9-G, 11-G, 14-S, 15-S, 18-G, 19-G, 23-G, 25-S, 27-G, 28-G, 30-M,
+38-S, 39-S, 41-S, 42-S, 43-S, 44-S, 45-M, 50-S, 52-S, and 53-S.</p>
+
+<p>The river carpsucker occurred at most stations on the larger
+streams, and in many of the smaller tributaries. In smaller streams
+<i>C. c. carpio</i> frequented the largest pools, in or near the floodplains
+of larger streams. A marked preference for still water, soft, silty
+bottoms, and areas with drift or other cover was apparent; however,
+the species also occurred in open waters with moderate to swift
+currents.</p>
+
+<p>The sizes attained by the river carpsucker at different ages were
+(averages followed by number of fish in parentheses): I, 1.9 (10);
+II, 3.9 (5); III, 5.3 (8); IV, 7.7 (5); V, 11.9 (2); VI, 11.6 (7);
+VII, 12.8 (6); VIII, 13.1 (1); IX, 14.9 (2); X, 15.8 (8); and XI,
+17.6 (1). These averages are significantly less than those reported
+by Buchholz (1957:594) for the river carpsucker in the Des
+Moines River, Iowa.</p>
+
+<p>Examination of the gonads of river carpsucker in summer, 1957,
+indicated that spawning occurred in late July. Young-of-the-year,
+averaging 21 mm. in total length, first appeared in my collections
+on July 30, 1957.</p>
+
+<p><b><i>Carpiodes velifer</i></b> (Rafinesque), highfin carpsucker: Meek
+(1895:135); Evermann and Cox (1896:389).</p>
+
+<p>The highfin carpsucker was not taken in my survey. Meek
+(1895:135) reported "this small sucker [<i>C. velifer</i>] ... common
+in Blue River at Crete," characterizing the specimens as having
+"Dorsal rays, 24 to 30; scales in the lateral-line, 36 to 41; head
+3&frac34; to 4; and depth 2&frac12; to 3." The ranges in the number of dorsal
+rays and the number of scales in the lateral-line are higher than
+usual in <i>C. velifer</i>, or in <i>C. c. carpio</i>, which is now common in the
+Big Blue River Basin. Both species normally have 33 to 37 lateral-line
+scales and 27 or fewer dorsal rays (Bailey, 1956:352-353;
+Moore, 1957:79; and Trautman, 1957:81-82). The other characters
+listed by Meek would fit the young and some adults of either
+species, or possibly a composite including <i>C. forbesi</i>.</p>
+
+<p>Graham (1885:72) and Cragin (1885:107) reported <i>Ictiobus
+velifer</i> (= <i>Carpiodes velifer</i>) from "Eureka Lake," Riley County,
+Kansas. This lake, which no longer exists, was in the Kansas River
+Valley, about ten miles upstream from the mouth of the Big Blue
+River. Other, more recent records from the Kansas River Basin,
+in the vicinity of the Big Blue River, are: Maple Leaf Lake, Riley
+<span class="pagenum"><a name="Page_418" id="Page_418">[418]</a></span>
+Co., Oct. 4, 1925; Deep Creek, Riley Co., no date; Wildcat Creek,
+Riley Co., Sept. 7, 1923; and Wildcat Creek, Riley Co., Sept. 29,
+1925 (UMMZ 122187-90). Most of the collections were made by
+Minna E. Jewell (Nelson, personal communication).</p>
+
+<p><b><i>Moxostoma aureolum</i></b> (LeSueur), northern redhorse: Cragin
+(1885:108) as <i>Moxostoma macrolepidotum</i> LeSueur; Meek (1895:136)
+as <i>Moxostoma macrolepidotum duquesnei</i> (LeSueur); Evermann
+and Cox (1896:394-395); and Jennings (1942:364) as <i>Moxostoma
+erythrurum</i> (Rafinesque). Stations 41-S, 43-S, 44-S, and
+53-S.</p>
+
+<p>I collected three northern redhorse from the Big Blue River
+Basin, and another specimen was seined in the mouth of Mill Creek,
+Riley County (my present Station 9-G) by the Kansas State College
+class in fisheries management in 1954 (KSC 5068). I reidentify
+as <i>M. aureolum</i> the two specimens recorded by Jennings (<i>loc. cit.</i>)
+as <i>M. erythrurum</i>.</p>
+
+<p>The subspecific status of <i>M. aureolum</i> in the Kansas River Basin is
+to be the subject of another paper.</p>
+
+<p><b><i>Catostomus commersonnii</i></b> (Lacépède), white sucker: Canfield
+and Wiebe (1931:8) as "common suckers"; and Breukelman (1940:380).
+Stations 7-G, 11-G, 12-G, 13-G, 16-G, 18-G, 19-G, 23-G,
+29-G, 31-G, 53-S, 57-M, and 58-G.</p>
+
+<p>The white sucker occurred primarily in upland streams of the
+Flint Hills, with one occurrence in muddy habitat, and one in the
+main stream of the Big Blue River. Young <i>C. commersonnii</i> were
+often taken in riffles, but adults were in the larger, deeper pools.
+The ages and total lengths at the last annulus for 12 white suckers
+were: I, 2.8 (4); II, 3.9 (6); III, 8.2 (1); and IV, 9.2 (1).</p>
+
+<p><b><i>Cyprinus carpio</i></b> Linnaeus, carp: Canfield and Wiebe (1931:5-8,
+10) as "carp." Stations 1-S, 2-S, 3-S, 4-S, 6-S, 7-G, 8-S, 15-S,
+16-G, 18-G, 23-G, 24-G, 25-S, 27-G, 30-M, 35-M, 38-S, 41-S, 42-S,
+43-S, 44-S, 45-M, 52-S, 53-S, and 56-S.</p>
+
+<p>Carp occurred throughout the basin. The habitat of this species
+closely approximated that of the river carpsucker; however,
+carp were more often taken in moderate to swift water than were
+<i>C. c. carpio</i>.</p>
+
+<p>The ages and average lengths at the last annulus for 40 carp from
+the Big Blue River Basin were: I, 2.3 (4); II, 4.7 (10); III, 7.0 (10);
+IV, 9.0 (3); V, 11.3 (4); VI, 18.6 (1); VII, 18.9 (3); VIII, no fish;
+IX, 20.6 (3); X, 19.1 (2); XI, 21.1 (1); XII, 22.0 (1); and XIII,
+24.1 (2).</p>
+
+<p><b><i>Carassius auratus</i></b> (Linnaeus), goldfish. Station 4-S.
+<span class="pagenum"><a name="Page_419" id="Page_419">[419]</a></span></p>
+
+<p>I saw goldfish seined from Station 4-S by anglers obtaining bait
+on April 20, 1957. Goldfish were commonly used for bait at Stations
+4-S and 54-S.</p>
+
+<p><b><i>Semotilus atromaculatus</i></b> (Mitchill), creek chub: Evermann and
+Cox (1896:399); and Jennings (1942:364) as <i>Semotilus atromaculatus
+atromaculatus</i> (Mitchill). Stations 5-G, 7-G, 10-G, 11-G,
+12-G, 13-G, 16-G, 17-G, 18-G, 23-G, 24-G, 27-G, 28-G, 29-G, 31-G,
+32-G, 33-M, 34-M, 36-M, 37-M, 40-M, 46-M, 47-M, 48-M, 49-M,
+50-S, 53-S, 54-G, 55-M, 56-S, 57-M, 58-G, and 59-G.</p>
+
+<p>Creek chubs were found in all habitats in the Big Blue River
+Basin, but were abundant only in the headwaters of muddy streams
+and in clear upland creeks.</p>
+
+<p><b><i>Chrosomus erythrogaster</i></b> (Rafinesque), southern redbelly dace:
+Jennings (1942:365). Stations 11-G, 12-G, 13-G, 16-G, 27-G, 29-G,
+and 53-S.</p>
+
+<p>This colorful species occupied the headwaters of the clear,
+spring-fed creeks where it was abundant. Only one specimen was
+taken in muddy or sandy habitat (at the mouth of a small creek at
+Station 53-S), where it may have been washed by floods just prior
+to my collecting.</p>
+
+<p><b><i>Hybopsis storeriana</i></b> (Kirtland), silver chub. Station 3-S.</p>
+
+<p>One specimen of <i>H. storeriana</i> (KU 3810) was seined in swift
+water near a sandbar on April 6, 1957, and another was taken at
+the same locality on April 26, 1958.</p>
+
+<p><b><i>Hybopsis aestivalis</i></b> (Girard), speckled chub: Meek (1895:137);
+and Evermann and Cox (1896:409), both as <i>Hybopsis hyostomus</i>
+Gilbert. Stations 3-S, 4-S, 14-S, 25-S, 38-S, 39-S, 50-S, and 56-S.</p>
+
+<p>This species was restricted to wide, swift parts of the Big Blue
+and Little Blue rivers, and was found over clean, sometimes shifting,
+sand bottoms. On May 29, 1958, three males in breeding condition
+were collected and on June 16, 1958, a large series of both
+male and female <i>H. aestivalis</i>, all with well-developed gonads, was
+collected. The water temperature was 77.0&deg;&nbsp;F. Hubbs and Ortenburger
+(1929:25-26) reported that <i>Extrarius tetranemus</i> (Gilbert)
+(= <i>Hybopsis aestivalis tetranemus</i>) spawns in summer especially
+in early July. Cross (1950:135) reported a single pair of <i>H. a.
+tetranemus</i> that he considered in breeding condition on June 9, 1948.</p>
+
+<p>Breukelman (1940:380) recorded speckled chubs in the Kansas
+River Basin as <i>Extrarius</i> (= <i>Hybopsis</i>) <i>aestivalis</i>: <i>sesquialis</i> ×
+<i>tetranemus</i>; however, the name <i>sesquialis</i> is a <i>nomen nudum</i>, and
+<span class="pagenum"><a name="Page_420" id="Page_420">[420]</a></span>
+the status of this species in the Kansas River Basin is yet to be
+elucidated.</p>
+
+<p><b><i>Phenacobius mirabilis</i></b> (Girard), plains suckermouth minnow:
+Meek (1895:136); and Evermann and Cox (1896:408). Stations
+2-S, 3-S, 4-S, 5-G, 6-S, 7-G, 8-S, 9-G, 11-G, 16-G, 18-G, 25-S, 26-G,
+27-G, 35-M, 38-S, 39-S, 40-M, 42-S, 47-M, 50-S, 52-S, 53-S, 54-G,
+and 56-S.</p>
+
+<p><i>Phenacobius mirabilis</i> was widespread in the basin, occurring
+most frequently on riffles over bottoms of clean sand or gravel.
+Young-of-the-year were usually taken in backwaters.</p>
+
+<p><b><i>Notropis percobromus</i></b> (Cope), plains shiner. Stations 3-S and
+4-S.</p>
+
+<p>The plains shiner occurred only in the lower part of the main
+stream of the Big Blue River.</p>
+
+<p><b><i>Notropis rubellus</i></b> (Agassiz), rosyface shiner. Station 5-G.</p>
+
+<p>One rosyface shiner (KU 4195) was taken. This species was previously
+reported from only two localities in the Kansas River Basin:
+in the Mill Creek Watershed, Wabaunsee County, and Blacksmith
+Creek, Shawnee County as <i>Notropis rubrifrons</i> (Cope) (Gilbert,
+1886:208). Mill Creek and Blacksmith Creek are northward-flowing
+tributaries of the Kansas River that arise in the Flint Hills.
+Graham (1885:73) also recorded <i>N. rubellus</i> (as <i>N. rubrifrons</i>)
+from the "Kansas and Missouri Rivers"; however, I suspect that his
+specimens were <i>Notropis percobromus</i>, a species not generally
+recognized in Graham's time (see Hubbs, 1945:16-17). <i>Notropis
+rubellus</i> is now abundant in the Mill Creek Watershed (Wabaunsee
+County), but, except for my specimen No. 4195, has not been taken
+recently in other streams in the Kansas River Basin.</p>
+
+<p><b><i>Notropis umbratilis umbratilis</i></b> (Girard), redfin shiner. Station
+3-S.</p>
+
+<p>One specimen of <i>N. u. umbratilis</i> was captured near a sandbar
+on March 26, 1958. The absence of this species in Flint Hills
+streams of the Big Blue River Basin is unexplained; redfin shiners
+occur commonly in southern tributaries of the Kansas River both
+upstream and downstream from the mouth of the Big Blue River.
+In Kansas this species is usually associated with the larger pools
+of clear, upland streams.</p>
+
+<p>Canfield and Wiebe (1931:6-8) may have referred to this species
+in recording "black-fin minnows" from the Nebraskan portion of
+the Big Blue River Basin.</p>
+
+<p><b><i>Notropis cornutus frontalis</i></b> (Agassiz), common shiner. Stations
+<span class="pagenum"><a name="Page_421" id="Page_421">[421]</a></span>
+4-S, 5-G, 7-G, 10-G, 11-G, 12-G, 13-G, 18-G, 22-G, 26-G, 27-G, 28-G,
+29-G, 31-G, 32-G, and 59-G.</p>
+
+<p>Common shiners were most abundant in middle sections of the
+clear, gravelly creeks.</p>
+
+<p><b><i>Notropis lutrensis</i></b> (Baird and Girard), red shiner: Meek (1895:136);
+and Evermann and Cox (1896:404-405). All stations excepting
+1-S, 17-G, 30-M, and 51-M.</p>
+
+<p>Red shiners were the most widespread species taken in my survey,
+occurring in all habitats, and in all kinds of streams. On two
+occasions I observed what apparently was spawning behavior of
+this species. Both times the specimens collected were in the
+height of breeding condition, stripping in the hand easily, and
+often without pressure. At the first locality (Station 29-G) no attempt
+was made to obtain eggs, but by disturbing the bottom at
+the second (55-M) I found eggs that were thought to be those of
+red shiners. The eggs were slightly adhesive, clinging to the hand
+and to the bobbinet seine.</p>
+
+<p>On June 29, 1958, at Station 29-G, red shiners appeared to be
+spawning in an open-water area measuring about 15 by 15 feet,
+over nests of <i>Lepomis cyanellus</i> Rafinesque and <i>L. humilis</i> (Girard).
+No interspecific activity was noted between the sunfish and the red
+shiners. Water temperature at this station was 73.4&deg;&nbsp;F., and the
+bottom was gravel, sand, and mud. Observations were made from
+a high cut-bank, by naked eye and by use of 7-X binoculars.</p>
+
+<p>The red shiners moved rapidly at the surface of the water, with
+one male (rarely two or more) following one female. The male
+followed closely, passing the female and causing her to change
+direction. At the moment of the female's hesitation, prior to her
+turn, the male would erect his fins in display, at the side and a little
+in front of the female. After brief display, usually less than two
+seconds, the male resumed the chase, swimming behind and around
+the female in a spiral fashion. After a chase of two to three feet,
+the female would sometimes allow the male to approach closely on
+her left side. The male nudged the female on the caudal peduncle
+and in the anal region, moving alongside with his head near the
+lower edge of the left operculum of the female, thus placing his
+genital pore about a head-length behind and below that of the
+female. At this time spawning must have occurred; however,
+possibly because of the speed of the chase, I observed no vibration
+of the fish as described for other species of <i>Notropis</i> at the culmination
+<span class="pagenum"><a name="Page_422" id="Page_422">[422]</a></span>
+of spawning (Pfeiffer, 1955:98; Raney, 1947:106; and others).
+While the spawning act presumably occurred the pair was in
+forward motion in a straight course, for three to five feet, at the
+end of which the male moved rapidly away, gyrating to the side
+and down. The female then swam away at a slower rate. In
+instances when the female failed to allow the male to move alongside,
+the male sometimes increased his speed, striking the female,
+and often causing her to jump from the water.</p>
+
+<p>Some conflict between males was observed, usually when two
+or more followed one female. The males would leave the female,
+swerve to one side, and stop, facing each other or side by side. At
+this moment the fins were greatly elevated in display. There was
+usually a rush on the part of one male, resulting in the flight of
+the other, and the aggressive male would pursue for about two feet.
+Many times the pursued male jumped from the water.</p>
+
+<p>At Station 55-M, on July 9, 1958, activity similar to that described
+above was observed in a small pool near a mass of debris. At this
+station I watched from the bank, three feet from the spawning
+shiners. Water temperature was not recorded.</p>
+
+<p>The minnows performed the same types of chase and display,
+all in open water, as described for Station 29-G, However, at Station
+55-M, much activity of males occurred near the small deposit
+of debris. It seemed that conflict was taking place, with males behaving
+as described above, and milling violently about. Examination
+of the area revealed nests of <i>L. cyanellus</i> near the debris, and
+some of the activity by the shiners may have been raids on nests of
+the sunfish. However, females nearing the group of males were
+immediately chased by one to four individual males, with one
+usually continuing pursuit after a short chase by the group. The
+male again moved into position at the lower left edge of the operculum
+of the female as at Station 29-G.</p>
+
+<p>Another kind of behavior was observed also, in which the
+female sometimes stopped. The male approached, erecting his
+fins and arching his body to the left. The female also assumed this
+arch to the left, and the pair moved in a tight, counter-clockwise
+circle, with the male on the inside. After a short period in this
+position, the male moved aside in display, and gyrated to the side
+and down. Females at both stations moved about slowly, usually
+remaining in the immediate vicinity of activity by males, and returning
+to the area even when pursued and deserted some distance
+away.</p>
+
+<p><b><i>Notropis deliciosus</i></b> (Girard), sand shiner: Meek (1895:136);
+<span class="pagenum"><a name="Page_423" id="Page_423">[423]</a></span>
+Evermann and Cox (1896:402), both as <i>Notropis blennius</i> (Girard);
+and Jennings (1942:365) as <i>Notropis deliciosus missuriensis</i> (Cope).
+All stations excepting 1-S, 10-G, 12-G, 17-G, 20-G, 21-G, 22-G,
+24-G, 29-G, 30-M, 31-G, 32-G, 33-M, 35-M, 51-M, 55-M, 57-M, 58-G,
+and 59-G.</p>
+
+<p>Nelson (personal communication) has studied the sand shiner in
+Kansas, and has found that the Big Blue River is an area of intergradation
+between the southwestern subspecies (<i>deliciosus</i>) and the
+plains subspecies (<i>missuriensis</i>). <i>Notropis d. deliciosus</i> prefers
+cool, rocky habitat, and occurs in small streams of the Flint Hills,
+whereas <i>N. d. missuriensis</i> occupies the sandy, turbid Big Blue and
+Little Blue rivers. Intergrades occur most frequently in the Big
+Blue River, but are found in all habitats.</p>
+
+<p><b><i>Notropis topeka</i></b> (Gilbert), Topeka shiner: Meek (1895:136);
+Evermann and Cox (1896:403); and Minckley and Cross (in press).
+Stations 10-G, 11-G, 12-G, 19-G, 31-G, and 32-G.</p>
+
+<p>This species was common locally in the upland streams. Female
+Topeka shiners stripped easily at Station 11-G on July 8, 1958, and
+adult <i>N. topeka</i> in high breeding condition were collected at Station
+31-G on July 14, 1958. The water temperature at both stations
+was 77.5&deg;&nbsp;F. Evermann and Cox (1896:403-404) recorded female
+Topeka shiners "nearly ripe" on June 29, 1893.</p>
+
+<p><b><i>Notropis buchanani</i></b> Meek, ghost shiner. Stations 3-S and 4-S.
+Only two specimens of <i>N. buchanani</i> were taken, both on
+August 14, 1957. These specimens (KU 3833), a female with
+well-developed ova, and a tuberculate male, were near a sandbar
+in the main channel. To my knowledge, this is the first published
+record of the ghost shiner from the Kansas River Basin. Mr. James
+Booth, State Biological Survey, collected <i>N. buchanani</i> from two
+stations on Mill Creek, Wabaunsee County, Kansas, 1953.</p>
+
+<p><b><i>Hybognathus nuchalis</i></b> Agassiz, silvery minnow. Stations 2-S,
+3-S, 4-S, 7-G, 8-S, and 16-G.</p>
+
+<p>This species was taken sporadically, but sometimes abundantly,
+in the Big Blue River. At Stations 7-G and 16-G a few young-of-the-year
+were found.</p>
+
+<p>Bailey (1956:333) does not consider the southwestern <i>Hybognathus
+placita</i> (Girard) specifically distinct from the northeastern
+<i>H. nuchalis</i>, but little evidence of intergradation has been published.
+In <a href="#Table_2">Table&nbsp;2</a>, I have compared measurements and counts
+of 50 specimens of <i>Hybognathus</i> from the Big Blue River, 50 <i>H. n.
+placita</i> from the Walnut River, Kansas (Arkansas River Basin),
+and 50 <i>H. n. nuchalis</i> from Wisconsin. Measurements and counts
+were made by methods described by Hubbs and Lagler (1947:8-15)
+and measurements are expressed as thousandths of standard length.</p>
+
+<p><span class="pagenum"><a name="Page_424" id="Page_424">[424]</a></span></p>
+
+<a name="Table_2" id="Table_2"></a>
+<div class="sm_tbl"><span class="smcap">Table 2. Comparisons of Three Series of</span> <i>Hybognathus</i>, <span class="smcap">50 Specimens Each,
+from Different Stream Systems. Symbols:</span>&nbsp; X&#773;&nbsp;=&nbsp;<span class="smcap">Mean;</span>
+&#963;&nbsp;=&nbsp;<span class="smcap">One Standard Deviation; and</span> 2 &#963;<sub>m</sub>&nbsp;=&nbsp;<span class="smcap">Two Standard Errors.
+Ranges are in Parentheses. Standard Lengths of Specimens are: Walnut River, Kansas, 60.0 to 72.7 mm.</span>, <span style="text-decoration:overline">X</span>&nbsp;=&nbsp;67.1;
+<span class="smcap">Big Blue River, Kansas, 43.3 to 63.3 mm.</span>, <span style="text-decoration:overline">X</span>&nbsp;=&nbsp;52.0; <span class="smcap">and
+Chippewa River, Wisconsin, 57.6 to 74.4 mm.</span>, <span style="text-decoration:overline">X</span>&nbsp;=&nbsp;65.9.</div>
+<br />
+<table width="100%" summary="Hybognathus Comparisons">
+<tr>
+ <td rowspan="2" class="bt2 bb center"><span class="smcap">Count or Proportional Measurement</span></td>
+ <td colspan="3" class="bt2 bb bl center">Walnut River, Kansas <br /><i>H. n. placita</i>, KU 3869</td>
+ <td colspan="3" class="bt2 bb bl center">Big Blue River,<br />Kansas KU 3812</td>
+ <td colspan="3" class="bt2 bb bl center">Chippewa River,<br /><i>H. n. nuchalis</i>, KU 2012</td>
+</tr>
+<tr>
+ <td class="bl bb center"><span style="text-decoration:overline">X</span></td>
+ <td class="bb bl center">&#963;</td>
+ <td class="bb bl center">2 &#963;<sub>m</sub></td>
+ <td class="bb bl center"><span style="text-decoration:overline">X</span></td>
+ <td class="bb bl center">&#963;</td>
+ <td class="bb bl center">2 &#963;<sub>m</sub></td>
+ <td class="bb bl center"><span style="text-decoration:overline">X</span></td>
+ <td class="bb bl center">&#963;</td>
+ <td class="bb bl center">2 &#963;<sub>m</sub></td>
+</tr>
+<tr>
+ <td>Lateral-line scales </td>
+ <td class="bl center">38.9<br />(37-41)</td>
+ <td class="bl center">1.1</td>
+ <td class="bl center">0.4</td>
+ <td class="bl center">37.2<br />(35-39)</td>
+ <td class="bl center">1.1</td>
+ <td class="bl center">0.4</td>
+ <td class="bl center">37.3<br />(35-39)</td>
+ <td class="bl center">1.0</td>
+ <td class="bl center">0.2</td>
+</tr>
+<tr>
+ <td>Predorsal scale-rows</td>
+ <td class="bl center">16.8<br />(15-19)</td>
+ <td class="bl center">0.9</td>
+ <td class="bl center">0.7</td>
+ <td class="bl center">15.9<br />(14-17)</td>
+ <td class="bl center">0.8</td>
+ <td class="bl center">0.2</td>
+ <td class="bl center">15.1<br />(14-17)</td>
+ <td class="bl center">0.5</td>
+ <td class="bl center">0.1</td>
+</tr>
+<tr>
+ <td>Scale-rows below lateral-line</td>
+ <td class="bl center">15.6<br />(13-18)</td>
+ <td class="bl center">1.2</td>
+ <td class="bl center">0.3</td>
+ <td class="bl center">14.9<br />(12-16)</td>
+ <td class="bl center">1.0</td>
+ <td class="bl center">0.3</td>
+ <td class="bl center">12.9<br />(12-15)</td>
+ <td class="bl center">0.7</td>
+ <td class="bl center">0.2</td>
+</tr>
+<tr>
+ <td>Scale-rows around caudal peduncle</td>
+ <td class="bl center">16.2<br />(15-19)</td>
+ <td class="bl center">1.1</td>
+ <td class="bl center">0.3</td>
+ <td class="bl center">15.8<br />(14-18)</td>
+ <td class="bl center">0.8</td>
+ <td class="bl center">0.2</td>
+ <td class="bl center">13.8<br />(12-15)</td>
+ <td class="bl center">0.6</td>
+ <td class="bl center">0.2</td>
+</tr>
+<tr>
+ <td>Orbit ÷ standard length</td>
+ <td class="bl center">.051<br />(044-61)</td>
+ <td class="bl center">.0035</td>
+ <td class="bl center">.0010</td>
+ <td class="bl center">.059<br />(047-71)</td>
+ <td class="bl center">.0047</td>
+ <td class="bl center">.0013</td>
+ <td class="bl center">.068<br />(059-77)</td>
+ <td class="bl center">.0044</td>
+ <td class="bl center">.0013
+ <span class="pagenum"><a name="Page_425" id="Page_425">[425]</a></span></td>
+</tr>
+<tr>
+ <td>Gape-width ÷ standard length</td>
+ <td class="bl center">.066<br />(055-75)</td>
+ <td class="bl center">.0046</td>
+ <td class="bl center">.0013</td>
+ <td class="bl center">.064<br />(055-74)</td>
+ <td class="bl center">.0044</td>
+ <td class="bl center">.0013</td>
+ <td class="bl center">.056<br />(046-64)</td>
+ <td class="bl center">.0038</td>
+ <td class="bl center">.0011</td>
+</tr>
+<tr>
+ <td class="bb">Orbit ÷ gape-width</td>
+ <td class="bb bl center">.776<br />(647-945)</td>
+ <td class="bb bl center">.0083</td>
+ <td class="bb bl center">.0024</td>
+ <td class="bb bl center">.907<br />(712-1.067)</td>
+ <td class="bb bl center">.0080</td>
+ <td class="bb bl center">.0023</td>
+ <td class="bb bl center">1.223<br />(953-1.566)</td>
+ <td class="bb bl center">.0119</td>
+ <td class="bb bl center">.0034</td>
+</tr>
+</table>
+<br />
+
+<p><span class="pagenum"><a name="Page_426" id="Page_426">[426]</a></span></p>
+
+<p><i>Hybognathus</i> from the Big Blue River tend to have fewer, larger
+scales than <i>H. n. placita</i> from the Walnut River, Kansas, but more
+and smaller scales than <i>H. n. nuchalis</i> from Wisconsin. In specimens
+from the Blue River, the size of the orbit divided by standard
+length, and the width of gape divided by standard length and
+width of orbit, are also intermediate between the Walnut River and
+Wisconsin specimens, but tend toward the former. Specimens from
+the Big Blue River resemble <i>H. n. placita</i> from the Walnut River
+in body shape, robustness, and in the embedding of scales on the
+nape.</p>
+
+<p><b><i>Pimephales notatus</i></b> (Rafinesque), bluntnose minnow: Meek
+(1895:136); and Evermann and Cox (1896:399). Stations 2-S, 3-S,
+5-G, 6-S, 8-S, 9-G, 10-G, 11-G, 12-G, 13-G, 16-G, 19-G, 27-G, 29-G,
+53-S, 54-G, and 58-G.</p>
+
+<p>The bluntnose minnow preferred the clearer creeks, with gravel
+or gravel-silt bottoms, but occurred rarely in the mainstream of the
+Big Blue River. Males and females in high breeding condition
+were taken on July 14, 1958. The temperature of the water was
+75.5&deg;&nbsp;F.</p>
+
+<p><b><i>Pimephales promelas</i></b> Rafinesque, fathead minnow: Meek (1895:
+136); and Evermann and Cox (1896:397-398). All stations excepting
+1-S, 4-S, 12-G, 30-M, 43-S, 44-S, and 56-S.</p>
+
+<p>Small muddy streams were preferred by <i>P. promelas</i>; however,
+the fathead minnow was taken in all habitats, and in association
+with most other species.</p>
+
+<p>Canfield and Wiebe (1931:6-7) may have recorded <i>P. promelas</i>
+from the Big Blue River Basin, Nebraska, as "blackhead minnows."</p>
+
+<p><b><i>Campostoma anomalum plumbeum</i></b> (Girard), stoneroller. All
+stations excepting 1-S, 2-S, 3-S, 14-S, 15-S, 21-G, 22-G, 28-G, 30-M,
+33-M, 34-M, 35-M, 36-M, 37-M, 38-S, 41-S, 44-S, 45-M, 51-M,
+52-S, and 55-M.</p>
+
+<p>Stonerollers were usually taken in riffles with gravel-rubble bottoms.
+Those individuals collected in areas with mud or sand
+bottoms were almost invariably in the current, or in the edge of
+currents.</p>
+
+<p>Specimens from the Big Blue River Basin have an average of
+47.4 scale-rows around the body (range 42-54).</p>
+
+<p><b><i>Ictalurus melas</i></b> (Rafinesque), black bullhead: Evermann and
+<span class="pagenum"><a name="Page_427" id="Page_427">[427]</a></span>
+Cox (1896:387) as <i>Ameiurus melas</i> (Rafinesque); and Canfield
+and Wiebe (1931:5-7, 10) as "bullheads." Stations 2-S, 6-S, 7-G,
+11-G, 16-G, 20-G, 22-G, 23-G, 24-G, 28-G, 35-M, 40-M, 51-M, 53-S,
+55-M, 56-S, 57-M, and 58-G.</p>
+
+<p>Black bullhead occurred in all habitats, but were less commonly
+taken in the Big Blue and Little Blue rivers than in other streams.</p>
+
+<p><b><i>Ictalurus natalis</i></b> (LeSueur), yellow bullhead. Stations 7-G, 9-G,
+10-G, 11-G, 17-G, 18-G, 19-G, 34-M, 35-M, 36-M, 37-M, 40-M, 47-M,
+48-M, 53-S, and 55-M.</p>
+
+<p>The yellow bullhead inhabited the muddy-bottomed streams and
+the upland, gravelly creeks, usually occurring in the headwaters.
+I obtained only one <i>I. natalis</i> in the sandy Big Blue River.</p>
+
+<p><b><i>Ictalurus punctatus</i></b> (Rafinesque), channel catfish: Cragin (1885:107);
+Meek (1895:135); Evermann and Cox (1896:386); and Canfield
+and Wiebe (1931:6-7, 10) as "channel catfish." Stations 1-S,
+2-S, 3-S, 4-S, 5-G, 6-S, 7-G, 8-S, 9-G, 11-G, 14-S, 15-S, 16-G, 18-G,
+25-S, 27-G, 30-M, 35-M, 38-S, 39-S, 41-S, 42-S, 43-S, 44-S, 46-M,
+50-S, 51-M, 52-S, 53-S, and 56-S.</p>
+
+<p>Channel catfish were most common in the larger, sandy streams,
+but occurred in other kinds of streams. The ages and calculated
+total lengths at the last annulus for 40 channel catfish were: I, no
+fish; II, 7.3 (16); III, 10.6 (5); IV, 12.3 (5); V, 13.3 (6); VI, 15.5
+(4); VII, 18.0 (3); and VIII, 21.9 (1). These lengths are slightly
+lower than averages reported by Finnell and Jenkins (1954:5) in
+Oklahoma impoundments.</p>
+
+<p>The length-frequency distribution of 438 channel catfish, collected
+by rotenone on August 5 and 7, 1958, indicated that two
+age-groups were represented. Without examination of spines, I
+assigned 265 fish to age-group O (1.3 to 2.9 inches, average 2.5)
+and 173 fish to age-group I (3.1 to 5.8 inches, average 4.5). The
+average total length of age group I (4.5 inches) is only slightly
+higher than the total length at the first annulus reported as average
+for Oklahoma (4.0 inches, Finnell and Jenkins, <i>loc. cit.</i>). It seems
+unlikely that my yearling fish taken in August, 1958, would have
+reached the length at the second annulus recorded in my study of
+spines (7.3 inches) by the end of the 1958 growing season.</p>
+
+<p>From 1952 to 1956, severe drought was prevalent in Kansas,
+probably causing streams to flow less than at any previously recorded
+time (Minckley and Cross, in press). This drought must
+have resulted in reduced populations of fishes in the streams. The
+channel catfish hatched in 1956 were therefore subjected to low
+<span class="pagenum"><a name="Page_428" id="Page_428">[428]</a></span>
+competition for food and space when normal flow was resumed in
+1957, and grew rapidly, reaching an average total length of 7.3
+inches at the second annulus, while channel catfish that were members
+of the large 1957 and 1958 hatches suffered more competition
+and grew more slowly.</p>
+
+<p><b><i>Noturus flavus</i></b> Rafinesque, stonecat: Jennings (1942:365). Stations
+3-S, 4-S, 6-S, 16-G, 25-S, 28-G, 38-S, 41-S, 42-S, 43-S, 52-S, 53-S,
+and 56-S.</p>
+
+<p><i>Noturus flavus</i> frequented riffles and swift currents along sandbars
+in the Big Blue and Little Blue rivers. Cross (1954:311) reported
+that "the shale-strewn riffles of the South Fork [of the
+Cottonwood River, Kansas] provide ideal habitat for the stonecat."
+In my study-area, this species was found not only on rubble-bottomed
+riffles, but occurred along both stationary and shifting
+sandbars where no cover was apparent.</p>
+
+<p><b><i>Pylodictis olivaris</i></b> (Rafinesque), flathead catfish: Canfield and
+Wiebe (1931:7) as "yellow catfish." Stations 3-S, 4-S, 6-S, 8-S,
+15-S, 25-S, 38-S, 41-S, 43-S, 44-S, 53-S, and 56-S.</p>
+
+<p>Flathead catfish were found only in the larger rivers. The species
+was taken rarely by seine, but was readily obtained by electric
+shocker. Data on the age and growth and food-habits of this
+species are to be the subject of another paper.</p>
+
+<p><b><i>Anguilla bostoniensis</i></b> (LeSueur), American eel: Jennings (1942:365).</p>
+
+<p>American eels are now rare in Kansas, and none was taken in my
+survey. The specimen reported by Jennings (<i>loc. cit.</i>) is at Kansas
+State College (KSC 2916), and was taken by I. D. Graham from
+the Big Blue River, Riley County, 1885.</p>
+
+<p><b><i>Fundulus kansae</i></b> Garman, plains killifish. Station 42-S.</p>
+
+<p>The plains killifish was collected by me only at Station 42-S.
+Specimens were collected from my Station 4-S by the Kansas State
+College class in fisheries management in 1954 (KSC 4985). My
+specimens were 11 to 13 mm. in total length.</p>
+
+<p><b><i>Roccus chrysops</i></b> (Rafinesque), white bass. Station 3-S.</p>
+
+<p>That the white bass is indigenous to Kansas is evidenced by
+records of Graham (1885:77) and Cragin (1885:111); however,
+since that time, and prior to the introduction of this species into
+reservoirs in the State, <i>R. chrysops</i> has rarely been recorded in
+Kansas. I collected young white bass at Station 3-S in both 1957
+and 1958, and I collected them also in an oxbow of the Kansas River
+four miles west of Manhattan, Riley County, Kansas, in the mouth
+<span class="pagenum"><a name="Page_429" id="Page_429">[429]</a></span>
+of McDowell's Creek, Riley County, and in Deep Creek, Wabaunsee
+County, and I saw other specimens from an oxbow of the
+Kansas River on the Fort Riley Military Reservation, Riley County,
+Kansas. The apparent increase in abundance of white bass in the
+Kansas River Basin must be attributable to introductions in reservoirs,
+with subsequent escape and establishment in the streams.</p>
+
+<p><b><i>Micropterus salmoides salmoides</i></b> (Lacépède), largemouth bass.
+Stations 6-S, 11-G, 43-S, and 45-M.</p>
+
+<p>Four largemouth bass were taken. This species has been widely
+stocked in farm-ponds and other impoundments in Kansas.</p>
+
+<p><b><i>Lepomis cyanellus</i></b> Rafinesque, green sunfish: Breukelman (1940:382);
+and Canfield and Wiebe (1931:5, 7-8, 10) as "green sunfish."
+All stations excepting 1-S, 2-S, 4-S, 8-S, 9-G, 15-S, 22-G, 25-S,
+30-M, 32-G, 34-M, 38-S, 39-S, 41-S, 42-S, 43-S, 44-S, 45-M, 46-M,
+47-M, 50-S, and 52-S.</p>
+
+<p>Green sunfish occurred primarily in the muddy streams. The ages
+and total lengths at the last annulus for 25 specimens are as follows:
+I, 1.1 (9); II, 2.2 (4); III, 3.1 (7); IV, 5.4 (4); and V, 6.0 (1).
+Male green sunfish were seen on nests on June 29, July 1, and
+July 9, 1958.</p>
+
+<p><b><i>Lepomis humilis</i></b> (Girard), orangespotted sunfish: Meek (1895:137);
+Evermann and Cox (1896:418); Canfield and Wiebe (1931:6)
+as "orange spots"; and Breukelman (1940:382). All stations excepting
+1-S, 9-G, 13-G, 15-G, 17-G, 21-G, 26-G, 34-M, 36-M, 38-M, 43-M,
+44-S, 47-M, 50-S, and 52-S.</p>
+
+<p><i>Lepomis humilis</i> was most common over sand-silt bottoms. Only
+two age-groups were found; their calculated total lengths were
+I, 1.7 (15); and II, 2.4 (10). Orangespotted sunfish were seen nesting
+on the same dates as <i>Lepomis cyanellus</i>.</p>
+
+<p><b><i>Lepomis macrochirus</i></b> Rafinesque, bluegill. Stations 7-G, 13-G,
+16-G, 24-G, and 59-G.</p>
+
+<p>This species has been widely stocked in Kansas. Only young-of-the-year
+and sub-adults were taken, and these were rare.</p>
+
+<p><b><i>Pomoxis annularis</i></b> Rafinesque, white crappie: Canfield and
+Wiebe (1931:5-8, 10) as "white crappie." Stations 3-S, 6-S, 8-S,
+12-G, 42-S, and 53-S.</p>
+
+<p>White crappie were rare, except in a borrow-pit at Station 6-S.
+Ages and calculated total lengths at the last annulus for 50 specimens
+from 6-S are as follows: I, 3.6 (22); II, 5.0 (14); III, 7.1 (5);
+IV, 8.3 (7); and V, 10.7 (2).</p>
+
+<p><b><i>Pomoxis nigromaculatus</i></b> (LeSueur), black crappie. Station 6-S.</p>
+
+<p>One black crappie (KU 4174) was taken. Canfield and Wiebe
+<span class="pagenum"><a name="Page_430" id="Page_430">[430]</a></span>
+(1931:10) noted: "The Black Crappie has been planted here [Big
+Blue River Basin in Nebraska] by the State, but, apparently, is not
+propagating itself."</p>
+
+<p><b><i>Stizostedion canadense</i></b> (Smith), sauger. Station 56-S.</p>
+
+<p>Mr. Larry Stallbaumer, of Marysville, Kansas, obtained a sauger
+(KU 4179) while angling on May 25, 1958.</p>
+
+<p><b><i>Stizostedion vitreum</i></b> (Mitchill), walleye.</p>
+
+<p>Though I failed to obtain the walleye in my survey, Dr. Raymond
+E. Johnson (personal communication) reported that the
+species occurred in the Nebraskan portion of the Big Blue River
+in recent years. Canfield and Wiebe (1931:6, 10) reported that
+"yellow pike are taken at Crete [Nebraska]," but may have referred
+to either the walleye or the sauger.</p>
+
+<p><b><i>Perca flavescens</i></b> (Mitchill), yellow perch: Canfield and Wiebe
+(1931:5-6, 10) as "ring perch" and "yellow perch."</p>
+
+<p>This fish was not taken in my survey. Canfield and Wiebe (<i>loc.
+cit</i>.) reported that the yellow perch "had been planted by the
+State [Nebraska]."</p>
+
+<p><b><i>Etheostoma nigrum nigrum</i></b> Rafinesque, johnny darter: Jennings
+(1942:365) as <i>Boleosoma nigrum nigrum</i> (Rafinesque). Stations
+10-G, 11-G, 12-G, 13-G, 16-G, 29-G, 40-M, 53-S, and 54-G.</p>
+
+<p>The larger pools of gravelly streams were preferred by johnny
+darters, but one specimen was taken from the main stream of the
+Big Blue River, and the species was abundant in one stream over
+hard, sand-silt bottom.</p>
+
+<p><b><i>Etheostoma spectabile pulchellum</i></b> (Girard), orangethroat
+darter: Jennings (1942:365) as <i>Poecilichthys spectabilis pulchellus</i>
+(Girard). Stations 5-G, 7-G, 10-G, 11-G, 12-G, 13-G, 16-G, 17-G,
+18-G, 21-G, 23-G, 27-G, 28-G, 29-G, 33-M, 40-M, 49-M, 53-S,
+54-G, and 59-G.</p>
+
+<p>The orangethroat darter was less restricted in habitat than the
+johnny darter, occurring in all stream-types, but most often in the
+riffles of gravelly streams. Most specimens from muddy or sandy
+streams were small.</p>
+
+<p><b><i>Aplodinotus grunniens</i></b> Rafinesque, freshwater drum. Stations
+3-S, 4-S, 6-S, 7-G, 8-S, 15-S, 38-S, 39-S, 53-S, and 56-S.</p>
+
+<p>The ages and calculated total lengths at the last annulus for 42
+freshwater drum from the Big Blue River were: I, 3.0 (10); II,
+5.7 (6); III, 9.4 (7); IV, 12.1 (13); V, 14.0 (3); VI, 15.1 (2); and
+VII, 16.3 (1).</p>
+
+<p><span class="pagenum"><a name="Page_431" id="Page_431">[431]</a></span></p>
+
+<a name="HYBRID_COMBINATIONS" id="HYBRID_COMBINATIONS"></a>
+<div class="caption2">HYBRID COMBINATIONS</div>
+
+<p>I obtained two hybrid fishes in my study-area. One specimen of
+<i>Notropis cornutus frontalis</i> × <i>Chrosomus erythrogaster</i> was taken
+at Station 29-G. This combination was recorded by Trautman
+(1957:114) in Ohio. The other hybrid was <i>Lepomis cyanellus</i> ×
+<i>Lepomis humilis</i>, captured at Station 24-G. This combination was
+first recorded by Hubbs and Ortenburger (1929:42).</p>
+
+<p>Hubbs and Bailey (1952:144) recorded another hybrid combination
+from my area of study: <i>Campostoma anomalum plumbeum</i> ×
+<i>Chrosomus erythrogaster</i>, UMMZ 103132, from a "spring-fed creek
+on 'Doc' Wagner's farm, Riley County, Kansas; September 21,
+1927; L. O. Nolf [collector]."</p>
+
+<a name="RELATIVE_ABUNDANCE_AND_DISCUSSION_OF_SPECIES" id="RELATIVE_ABUNDANCE_AND_DISCUSSION_OF_SPECIES"></a>
+<div class="caption2">RELATIVE ABUNDANCE AND DISCUSSION OF SPECIES</div>
+
+<p>The relative abundance of different species was estimated by
+combining counts of individual fishes taken in 290 seine-hauls, 26
+hours and 15 minutes of shocking, and seven samples obtained with
+rotenone. At some stations all seine-hauls were counted. At other
+stations the seine-hauls in which complete counts were recorded
+had been selected randomly in advance; that is to say, prior to
+collecting at each station. I selected those hauls to be counted
+from a table of random numbers (Snedecor, 1956:10-13). I did
+not use the frequency-of-occurrence method as proposed by Starrett
+(1950:114), in which the species taken and not the total number
+of individuals are recorded for all seine-hauls. However, the frequency
+of occurrence of each species is indicated by the number
+of stations at which it was found, and those stations are listed in
+the previous accounts. <a href="#Table_3">Table 3</a> shows the percentage of the total
+number of fish that each species comprised in three kinds of streams:
+sandy (Big Blue and Little Blue rivers), muddy, and gravelly
+streams.</p>
+
+<p>The habitat preferences of some species affect their abundance
+in different stream-types. <i>Notropis lutrensis</i> and <i>P. mirabilis</i> seemed
+almost ubiquitous. <i>Notropis deliciosus</i> also occurred in all kinds
+of streams (rarely in muddy streams); however, this species was
+represented by the sand-loving <i>N. d. missuriensis</i> in the Big Blue
+and Little Blue rivers, and <i>N. d. deliciosus</i> in the clear, gravelly,
+upland creeks (Nelson, personal communication). Because of its
+widespread occurrence, and for purposes of later discussion, I refer
+to this minnow also as an ubiquitous species in the Big Blue River
+Basin.</p>
+
+<p><span class="pagenum"><a name="Page_432" id="Page_432">[432]</a></span></p>
+
+<a name="Table_3" id="Table_3"></a>
+<div class="sm_tbl justify"><span class="smcap">Table 3. Relative Abundance of Fishes in Per Cent of the Total
+Number Taken, Big Blue River Basin, Kansas. Trace (Tr.) Is Used
+for Values Less Than .05 Per Cent, and Dashes Signify that the
+Species Did Not Occur in the Counted Collections Although it May
+Have Occurred in Uncounted Collections from the Same Stream-type.
+Three Species,</span> <i>C. auratus</i>, <i>N. buchanani</i>, <span class="smcap">and</span> <i>S. canadense</i>, <span class="smcap">Were Not
+Taken in Counted Collections.</span></div>
+<br />
+<table width="100%" summary="Relative Abundance">
+<tr>
+ <td rowspan="2" class="bt2 bb center smcap">Species</td>
+ <td colspan="2" class="bt2 bl center">Sandy streams</td>
+ <td rowspan="2" class="bt2 bb bl center">Muddy streams</td>
+ <td rowspan="2" class="bt2 bb bl center">Gravelly streams</td>
+</tr>
+<tr>
+ <td class="bt bb bl center">Big Blue River</td>
+ <td class="bt bb bl center">Little Blue River</td>
+</tr>
+<tr>
+ <td style="padding-top: 0.5em;"><i>N. lutrensis</i></td><td class="bl center">43.5</td>
+ <td class="bl center">55.9</td>
+ <td class="bl center">27.6</td>
+ <td class="bl center">56.0</td>
+</tr>
+<tr>
+ <td><i>I. punctatus</i></td>
+ <td class="bl center">14.0</td>
+ <td class="bl center">7.0</td>
+ <td class="bl center">1.2</td>
+ <td class="bl center">4.2</td>
+</tr>
+<tr>
+ <td><i>Carpiodes carpio</i></td>
+ <td class="bl center">11.9</td>
+ <td class="bl center">2.0</td>
+ <td class="bl center">5.0</td>
+ <td class="bl center">0.5</td>
+</tr>
+<tr>
+ <td><i>N. deliciosus</i></td>
+ <td class="bl center">8.2</td>
+ <td class="bl center">28.2</td>
+ <td class="bl center">3.1</td>
+ <td class="bl center">11.1</td>
+</tr>
+<tr>
+ <td><i>I. melas</i></td>
+ <td class="bl center">2.5</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">1.3</td>
+ <td class="bl center">0.5</td>
+</tr>
+<tr>
+ <td><i>Cyprinus carpio</i></td>
+ <td class="bl center">2.3</td>
+ <td class="bl center">1.9</td>
+ <td class="bl center">2.7</td>
+ <td class="bl center">0.2</td>
+</tr>
+<tr>
+ <td><i>P. olivaris</i></td>
+ <td class="bl center">1.8</td>
+ <td class="bl center">0.8</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>L. humilis</i></td>
+ <td class="bl center">1.7</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">9.0</td>
+ <td class="bl center">5.1</td>
+</tr>
+<tr>
+ <td><i>I. bubalus</i></td>
+ <td class="bl center">1.4</td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">Tr.</td>
+</tr>
+<tr>
+ <td><i>P. mirabilis</i></td>
+ <td class="bl center">1.3</td>
+ <td class="bl center">0.7</td>
+ <td class="bl center">0.3</td>
+ <td class="bl center">1.3</td>
+ </tr>
+<tr>
+ <td><i>H. nuchalis</i></td>
+ <td class="bl center">1.2</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">Tr.</td>
+ </tr>
+<tr>
+ <td><i>P. promelas</i></td>
+ <td class="bl center">0.8</td>
+ <td class="bl center">1.0</td>
+ <td class="bl center">28.7</td>
+ <td class="bl center">4.0</td>
+ </tr>
+<tr>
+ <td><i>H. aestivalis</i></td>
+ <td class="bl center">0.7</td>
+ <td class="bl center">0.2</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ </tr>
+<tr>
+ <td><i>A. grunniens</i></td>
+ <td class="bl center">0.5</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">0.2</td>
+ </tr>
+<tr>
+ <td><i>L. osseus</i></td>
+ <td class="bl center">0.5</td>
+ <td class="bl center">1.0</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ </tr>
+<tr>
+ <td><i>C. anomalum</i></td>
+ <td class="bl center">0.4</td>
+ <td class="bl center">0.2</td>
+ <td class="bl center">2.7</td>
+ <td class="bl center">4.6</td>
+ </tr>
+<tr>
+ <td><i>C. commersonnii</i></td>
+ <td class="bl center">0.4</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">0.7</td>
+ </tr>
+<tr>
+ <td><i>D. cepedianum</i></td>
+ <td class="bl center">0.4</td>
+ <td class="bl center">Tr.</td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">&mdash;</td>
+ </tr>
+<tr>
+ <td><i>N. percobromus</i></td>
+ <td class="bl center">0.3</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ </tr>
+<tr>
+ <td><i>P. annularis</i></td>
+ <td class="bl center">0.3</td>
+ <td class="bl center">Tr.</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ </tr>
+<tr>
+ <td><i>N. flavus</i></td>
+ <td class="bl center">0.2</td>
+ <td class="bl center">0.4</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">Tr.</td>
+ </tr>
+<tr>
+ <td><i>S. atromaculatus</i></td>
+ <td class="bl center">0.2</td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">12.2</td>
+ <td class="bl center">1.7</td>
+ </tr>
+<tr>
+ <td><i>M. aureolum</i></td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">0.2</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>I. cyprinella</i></td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>P. notatus</i></td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">2.2</td>
+</tr>
+<tr>
+ <td><i>I. niger</i></td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>H. alosoides</i></td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>E. spectabile</i></td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">1.4</td>
+ <td class="bl center">1.6</td>
+</tr>
+<tr>
+ <td><i>R. chrysops</i></td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>L. cyanellus</i></td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">3.5</td>
+ <td class="bl center">Tr.</td>
+</tr>
+<tr>
+ <td><i>H. storeriana</i></td>
+ <td class="bl center">Tr.</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>L. platostomus</i></td>
+ <td class="bl center">Tr.</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>M. salmoides</i></td>
+ <td class="bl center">Tr.</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>P. nigromaculatus</i></td>
+ <td class="bl center">Tr.</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>I. natalis</i></td>
+ <td class="bl center">Tr.</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">1.0</td>
+ <td class="bl center">Tr.</td>
+</tr>
+<tr>
+ <td><i>N. umbratilis</i></td>
+ <td class="bl center">Tr.</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>C. forbesi</i></td>
+ <td class="bl center">Tr.</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>S. platorynchus</i></td>
+ <td class="bl center">Tr.</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>F. kansae</i></td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">Tr.</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+</tr>
+<tr>
+ <td><i>E. nigrum</i></td>
+ <td class="bl center">Tr.</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">0.1</td>
+ <td class="bl center">0.2</td>
+</tr>
+<tr>
+ <td><i>N. rubellus</i></td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">Tr.</td>
+</tr>
+<tr>
+ <td><i>N. topeka</i></td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">1.0</td>
+</tr>
+<tr>
+ <td><i>N. cornutus</i></td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">1.0</td>
+</tr>
+<tr>
+ <td><i>C. erythrogaster</i></td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">&mdash;</td>
+ <td class="bl center">1.0</td>
+</tr>
+<tr><td style="padding-bottom: 0.5em;" class="bb"><i>L. macrochirus</i></td>
+ <td class="bb bl center">&mdash;</td>
+ <td class="bb bl center">&mdash;</td>
+ <td class="bb bl center">&mdash;</td>
+ <td class="bb bl center">1.0</td>
+</tr>
+</table>
+<br />
+
+<p><span class="pagenum"><a name="Page_433" id="Page_433">[433]</a></span></p>
+
+<p><i>Carpiodes carpio</i>, <i>Cyprinus carpio</i>, <i>I. punctatus</i>, <i>I. melas</i>, and <i>L.
+humilis</i> were widespread, but each was absent or rare in one of
+the kinds of streams (<a href="#Table_3">Table 3</a>). <i>Carpiodes carpio</i>, <i>Cyprinus carpio</i>,
+and <i>I. punctatus</i> occurred most frequently in the sandy streams,
+whereas <i>L. humilis</i> was most common in muddy streams. The high
+per cent of <i>I. melas</i> in collections from the Big Blue River is a direct
+result of one large population that was taken with rotenone in a
+borrow-pit at Station 6-S. In my opinion, this species actually
+was most abundant in the muddy streams.</p>
+
+<p>Some fish were almost restricted to the sandy streams, apparently
+because of preference for larger waters, or sandy stream-bottoms:
+<i>P. olivaris</i>, <i>I. bubalus</i>, <i>H. nuchalis</i>, <i>H. aestivalis</i>, <i>A. grunniens</i>, <i>L.
+osseus</i>, <i>D. cepedianum</i>, <i>N. percobromus</i>, <i>P. annularis</i>, <i>N. flavus</i>, <i>M.
+aureolum</i>, <i>I. niger</i>, <i>H. alosiodes</i>, and <i>R. chrysops</i>. Other species
+that were taken only in the larger rivers, and that are sometimes
+associated with streams even larger (or more sandy) than the Big
+Blue River are <i>H. storeriana</i>, <i>L. platostomus</i>, <i>M. salmoides</i>, <i>P. nigromaculatus</i>,
+<i>C. forbesi</i>, <i>S. platorynchus</i>, <i>F. kansae</i>, <i>N. buchanani</i>,
+<i>S. canadense</i>, and <i>C. auratus</i>. <i>Ictiobus cyprinella</i> also occurred
+more frequently in the larger streams.</p>
+
+<p>The muddy-bottomed streams supported populations composed
+primarily of <i>P. promelas</i>, <i>N. lutrensis</i>, and <i>S. atromaculatus</i>. No
+species was restricted to this habitat, but the following were
+characteristic there: <i>P. promelas</i>, <i>S. atromaculatus</i>, <i>L. humilis</i>,
+<i>L. cyanellus</i>, and <i>I. natalis</i>. <i>Carpiodes carpio</i>, <i>Cyprinus carpio</i>,
+<i>C. anomalum</i>, <i>E. spectabile</i>, and <i>E. nigrum</i> were locally common in
+muddy streams, but the first two were most frequent in larger,
+sandy streams, and the last three in gravelly streams.</p>
+
+<p>In gravel-bottomed, upland streams, <i>N. cornutus</i>, <i>N. rubellus</i>,
+<i>N. topeka</i>, and <i>C. erythrogaster</i> characteristically occurred; with
+the exception of <i>N. rubellus</i> (only one specimen taken), all were
+common at some stations. Other species in gravelly creeks were
+<i>N. lutrensis</i>, <i>C. anomalum</i>, <i>C. commersonnii</i>, <i>P. notatus</i>, <i>L. macrochirus</i>,
+<i>E. spectabile</i>, and <i>E. nigrum</i>. Although the one specimen of
+<i>N. umbratilis</i> taken in this survey was from the Big Blue River, this
+species is more characteristic of the clearer creeks in Kansas.</p>
+
+<p>In order to illustrate the composition of the fauna in some specific
+streams in the Big Blue River Basin, I segregated the fishes into
+ecological groups, as in the above discussion: ubiquitous types;
+species of larger, sandy streams; fishes of muddy streams; and
+fishes of clear, gravelly creeks.</p>
+
+<p>The total number of species taken in each of the streams was
+<span class="pagenum"><a name="Page_434" id="Page_434">[434]</a></span>
+divided into the number of species from that stream that were in
+each of these units, to give a percentage. The resultant data are
+presented graphically in <a href="#fig3">Figure&nbsp;3</a>.</p>
+<br />
+
+<div class="figcenter" style="width:600px">
+<a name="fig3" id="fig3"></a>
+<img src="images/fig_3.png" width="600" height="329" alt="" title="" /><br />
+<span class="fig_caption"><span class="smcap">Fig. 3.</span> Composition of the fauna of the entire Big Blue River Basin, and of
+seven streams or stream systems in that basin. &quot;Mill Creek, Wash. Co.&quot; refers
+to all streams in the Mill Creek System, Washington and Republic counties.
+&quot;Bl. Vermillion R. System&quot; includes all streams in that watershed excepting
+Clear Creek and one of its tributaries (Stations 31-G and 32-G).</span>
+</div>
+<br />
+<br />
+
+<div class="figcenter" style="width:605px">
+<a name="fig4" id="fig4"></a>
+<img src="images/fig_4.png" width="605" height="271" alt="" title="" /><br />
+<span class="fig_caption"><span class="smcap">Fig. 4.</span> Composition of the fauna of the Big Blue River, and of five collecting-sites
+on Carnahan Creek, Pottawatomie County. Lowermost sites are at the
+left of the figure.</span>
+</div>
+<br />
+
+<p><a href="#fig3">Figure 3</a> gives a generalized picture of the faunal composition in
+different kinds of streams. However, the fauna of a small tributary
+becomes more distinct from the fauna of the larger stream into
+which the small stream flows as one moves toward the headwaters
+(Metcalf, 1957:92, 95-100). <a href="#fig4">Figure 4</a> illustrates this in Carnahan
+<span class="pagenum"><a name="Page_435" id="Page_435">[435]</a></span>
+Creek. Station 11-G included four sampling-sites, which were approximately
+one, two, three, and four miles upstream from the
+mouth of Carnahan Creek. Station 13-G (one collection) was
+about four miles upstream from the closest sampling-site of Station
+11-G. Applying the same methods as for <a href="#fig3">Figure 3</a>, my findings
+show a gradual decline in the per cent of the fauna represented
+by the "large-river-fishes," and an increase in the segment classified
+as "upland-fishes," from downstream to upstream.</p>
+
+<a name="CREEL_CENSUS" id="CREEL_CENSUS"></a>
+<div class="caption2">CREEL CENSUS</div>
+
+<p>Fifty-three fishermen were interviewed in the 1957 creel census
+period, and 152 in 1958. Only those fishermen using pole and line
+were interviewed. In the area censused, much additional fishing is
+done with set-lines, that are checked periodically by the owners.</p>
+
+<p>In the 1958 census, 22 checks along approximately 80 miles of
+river were made, and seven of these trips were made without seeing
+one fisherman. The average fishing pressure for the entire area
+was estimated at one fisherman per 7.9 miles of stream, or one
+fisherman per 15.7 miles of shoreline.</p>
+
+<p>Seven species of fish were identified from fishermen's creels in
+1957 and 1958. These, in order of abundance were: channel catfish;
+carp; freshwater drum; flathead catfish; shovelnose sturgeon;
+smallmouth buffalo; and river carpsucker. Shovelnose sturgeon
+occurred in fishermen's creels only in April, 1957, and freshwater
+drum occurred more frequently in the spring-census of 1957 than
+in the summer of 1958.</p>
+
+<p>Sixty-two of the fishermen interviewed in 1958 were fishing for
+"anything they could catch," 68 were fishing specifically for catfish,
+and 22 sought species other than catfish. The order of preference
+was as follows: channel catfish, 21.1 per cent; flathead catfish, 15.1
+per cent; unspecified catfish, 12.5 per cent; carp, 9.2 per cent; freshwater
+drum, 1.3 per cent; and unspecified, 40.8 per cent. The kinds
+of fish desired by those fishermen checked in 1957 were not
+ascertained.</p>
+
+<p>Of all fishermen checked in 1957 and 1958, 165 were men, 17 were
+women, and 24 were children. Ninety-three per cent were fishing
+from the bank, five per cent were fishing from bridges, and two
+per cent were wading. All but two per cent of those checked were
+fishing "tightline"; the remainder fished with a cork.</p>
+
+<p>The ten baits most commonly used, in order of frequency, were
+worms, doughballs, minnows, liver, beef-spleen, chicken-entrails,
+coagulated blood, crayfish, shrimp, and corn.</p>
+
+<p>For purposes of later comparison the data on angler success
+<span class="pagenum"><a name="Page_436" id="Page_436">[436]</a></span>
+(<a href="#Table_4">Table 4</a>) have been divided according to areas: Area I, below
+Tuttle Creek Dam; Area II, in the Tuttle Creek Reservoir area; and
+Area III, above the reservoir. Areas I and III received the most
+fishing pressure, especially Station 4-S (in Area I), and Station
+56-S (in Area III).</p>
+
+<p>In Area I, the success ranged from 0.91 fish per fisherman-day
+in 1957 to 0.26 fish per fisherman-day in 1958. The 1957 census
+was made in April and May, when fishing in warm-water streams
+is considered better than in July (Harrison, 1956:203). The 1958
+census was from late June through July, and stream-flow in this
+period was continuously above normal. Therefore, fewer people
+fished the river, and catches were irregular. Catches in 1958
+ranged from 0.26 fish per fisherman-day in Area I to 0.44 fish per
+fisherman-day in Area III. In 1951, in the Republican River of
+Kansas and Nebraska, the average fisherman-day yielded 0.36 fish,
+0.09 fish per man-hour, and 0.06 fish per pole-hour (U. S. Fish and
+Wildlife Service, 1952:13-14). The average fisherman-day in the
+Republican River study was 3.0 hours, whereas the average on the
+Big Blue River was 2.2 hours for all areas in 1958 (<a href="#Table_4">Table 4</a>).</p>
+
+<a name="Table_4" id="Table_4"></a>
+<div class="smcap_tbl">Table 4. Angling Success in the Big Blue River, Kansas, 1957 and 1958.</div>
+<br />
+<table width="100%" summary="Angling Success">
+<tr>
+ <td class="bt2 bb smcap center">Area, Year, and Number<br />of Fishermen</td>
+ <td class="bt2 bb bl center">Average length of fisherman-day</td>
+ <td class="bt2 bb bl center">Number fish per fisherman-day</td>
+ <td class="bt2 bb bl center">Number fish per man-hour</td>
+ <td class="bt2 bb bl center">Number fish per pole-hour
+ <a name="FNanchor_A_1" id="FNanchor_A_1"></a><a href="#Footnote_A_1" class="fnanchor">[A]</a></td>
+</tr>
+<tr>
+ <td style="padding-top: 0.5em;">Area I, 1957<br />&nbsp;&nbsp;&nbsp;&nbsp;53 fishermen</td>
+ <td class="bl center">2.7 hours</td>
+ <td class="bl center">0.91</td>
+ <td class="bl center">0.33</td>
+ <td class="bl center">0.23</td>
+</tr>
+<tr>
+ <td style="padding-top: 0.5em;">Area I, 1958<br />&nbsp;&nbsp;&nbsp;&nbsp;84 fishermen</td>
+ <td class="bl center">2.5 hours</td>
+ <td class="bl center">0.26</td>
+ <td class="bl center">0.10</td>
+ <td class="bl center">0.07</td>
+</tr>
+<tr>
+ <td style="padding-top: 0.5em;">Area II, 1958<br />&nbsp;&nbsp;&nbsp;&nbsp;27 fishermen</td>
+ <td class="bl center">1.7 hours</td>
+ <td class="bl center">0.37</td>
+ <td class="bl center">0.22</td>
+ <td class="bl center">0.14</td>
+</tr>
+<tr>
+ <td style="padding-top: 0.5em;">Area III, 1958<br />&nbsp;&nbsp;&nbsp;&nbsp;41 fishermen</td>
+ <td class="bl center">2.4 hours</td>
+ <td class="bl center">0.44</td>
+ <td class="bl center">0.16</td>
+ <td class="bl center">0.11</td>
+</tr>
+<tr>
+ <td class="bb" style="padding-top: 0.5em; padding-bottom: 0.5em;">All areas, 1958<br />&nbsp;&nbsp;&nbsp;&nbsp;152 fishermen</td>
+ <td class="bb bl center">2.2 hours</td>
+ <td class="bb bl center">0.33</td>
+ <td class="bb bl center">0.14</td>
+ <td class="bb bl center">0.09</td>
+</tr>
+</table>
+
+<div class="footnote"><p><a name="Footnote_A_1" id="Footnote_A_1"></a><a href="#FNanchor_A_1"><span class="label">[A]</span></a> Fishermen used an average of 1.44 poles.</p></div>
+<br />
+
+<p>In the Big Blue River 47.7 per cent of all fishermen were successful
+in Area I in 1957, while only 13.1 per cent were successful
+in the same area in 1958 (<a href="#Table_5">Table 5</a>). In the Republican River, 24
+per cent of the fishing parties were successful (1.64 persons per
+party) (U. S. Fish and Wildlife Service, <i>loc. cit.</i>). The average
+<span class="pagenum"><a name="Page_437" id="Page_437">[437]</a></span>
+distance that each fisherman had traveled to fish in the Big Blue
+River was 15.7 miles. Seventy-nine per cent of the persons contacted
+lived within 25 miles of the spots where they fished. In
+the study on the Republican River, 77 per cent of the parties interviewed
+came less than 25 miles to fish.</p>
+
+<a name="Table_5" id="Table_5"></a>
+<div class="smcap_tbl">Table 5. Per Cent of Total Fishermen Successful, and Distances
+Travelled to Fish, Big Blue River Basin, Kansas, 1957 and 1958. All
+Distances Were Measured in Airline Miles.</div>
+<br />
+<table width="100%" summary="Fishermaen Success Rates">
+<tr>
+ <td class="bt2 bb">&nbsp;</td>
+ <td class="bt2 bl bb center">1957 Area I</td>
+ <td class="bt2 bl bb center">1958 Area I</td>
+ <td class="bt2 bl bb center">1958 Area II</td>
+ <td class="bt2 bl bb center">1958 Area III</td>
+ <td class="bt2 bl bb center">1958 All areas</td>
+</tr>
+<tr>
+ <td style="padding-top: 0.5em;">Per cent of fishermen successful</td>
+ <td class="bl center">47.1</td>
+ <td class="bl center">13.1</td>
+ <td class="bl center">18.5</td>
+ <td class="bl center">19.5</td>
+ <td class="bl center">15.8</td>
+</tr>
+<tr>
+ <td style="padding-top: 0.5em; padding-bottom: 0.5em;" class="bb">Distances traveled to fish<br />(averages in parentheses)</td>
+ <td class="bb bl center">0-121<br />(15.6)</td>
+ <td class="bb bl center">1-197<br />(20.5)</td>
+ <td class="bb bl center">0-124<br />(13.5)</td>
+ <td class="bb bl center">0-60<br />(7.4)</td>
+ <td class="bb bl center">0-197<br />(15.7)</td>
+</tr>
+</table>
+<br />
+
+<a name="RECOMMENDATIONS" id="RECOMMENDATIONS"></a>
+<div class="caption2">RECOMMENDATIONS</div>
+
+<p>My primary recommendation is for continued study of the
+Tuttle Creek Reservoir, and the Big Blue River above and below
+the reservoir, to trace changes in the fish population that result
+from impoundment.</p>
+
+<p>Probably the fishes that inhabit the backwaters, creek-mouths,
+and borrow-pits in the Big Blue River Basin (gars, shad, carpsucker,
+buffalo, carp, sunfishes, and white bass) will increase in abundance
+as soon as Tuttle Creek Reservoir is formed. Also, as in eastern
+Oklahoma reservoirs (see Finnell, <i>et al</i>., 1956:61-73), populations
+of channel and flathead catfish should increase. Because of the
+presence of brood-stock of the major sport-fishes of Kansas (channel
+and flathead catfish, bullhead, bluegill, crappie, largemouth bass,
+and white bass), stocking of these species would be an economic
+waste: exception might be made for the white bass. It may be
+above Tuttle Creek Dam, but was not found there.</p>
+
+<p>I do recommend immediate introduction of walleye, and possibly
+northern pike (<i>Esox lucius</i> Linnaeus), the latter species having
+been successfully stocked in Harlan County Reservoir, Nebraska,
+in recent years (Mr. Donald D. Poole, personal communication).
+These two species probably are native to Kansas, but may have
+been extirpated as agricultural development progressed. Reservoirs
+may again provide habitats suitable for these species in the State.</p>
+
+<p>If Tuttle Creek Reservoir follows the pattern found in most
+Oklahoma reservoirs, large populations of "coarse fish"&mdash;fishes that
+<span class="pagenum"><a name="Page_438" id="Page_438">[438]</a></span>
+are, however, commercially desirable&mdash;will develop (Finnell, <i>et al.</i>,
+<i>loc. cit.</i>). To utilize this resource, and possibly to help control
+"coarse fish" populations for the betterment of sport-fishing, some
+provision for commercial harvest should be made in the reservoir.</p>
+
+<a name="SUMMARY" id="SUMMARY"></a>
+<div class="caption2">SUMMARY</div>
+
+<p>1. The Big Blue River Basin in northeastern Kansas was studied
+between March 30, 1957, and August 9, 1958. The objectives were
+to record the species of fish present and their relative abundance
+in the stream, and to obtain a measure of angling success prior
+to closure of Tuttle Creek Dam.</p>
+
+<p>2. Fifty-nine stations were sampled one or more times, using
+seines, hoop and fyke nets, wire traps, experimental gill nets,
+rotenone, and an electric fish shocker.</p>
+
+<p>3. Forty-eight species of fish were obtained, and five others have
+been recorded in literature or found in museums. One species,
+<i>Carpiodes forbesi</i>, is recorded from Kansas for the first time.</p>
+
+<p>4. <i>Notropis lutrensis</i> was the most abundant fish in the Big Blue
+River Basin, followed by <i>Notropis deliciosus</i> and <i>Ictalurus
+punctatus</i>. The most abundant sport-fishes were <i>I. punctatus</i>, <i>I.
+melas</i>, and <i>Pylodictis olivaris</i>, respectively.</p>
+
+<p>5. The spawning behavior of <i>Notropis lutrensis</i> is described.</p>
+
+<p>6. A creel census at major points of access to the Big Blue River,
+was taken in 1957 (below Tuttle Creek Dam) and in 1958 (above,
+in, and below the dam-site). Fishing pressure averaged one fisherman
+per 15.7 miles of shoreline. The average length of the fisherman-day
+averaged 2.2 hours, with an average of 0.33 fish per fisherman-day
+being caught in 1958. The average number of fish per
+man-hour in 1958 was 0.14 and 15.8 per cent of the fishermen were
+successful. Distances traveled in order to fish ranged from 0 to
+197 miles (airline) and averaged 15.7 miles.</p>
+
+<p>7. The primary recommendation is that studies be continued, to
+document changes that result from impoundment. Because brood-stock
+of the major sport-fishes is already present, stocking is unnecessary,
+except for walleye and northern pike. Also, I recommend
+commercial harvest of non-game food-fishes.</p>
+
+<p><span class="pagenum"><a name="Page_439" id="Page_439">[439]</a></span></p>
+
+<a name="LITERATURE_CITED" id="LITERATURE_CITED"></a>
+<div class="caption2">LITERATURE CITED</div>
+
+<div class="refauthor smcap">Bailey, R. M.</div>
+
+<div class="reference">1956. A revised list of fishes of Iowa, with keys for identification. <i>In</i>
+Iowa Fish and Fishing, by J. R. Harlan and E. B. Speaker. Iowa
+State Cons. Comm., Des Moines, pp. 325-377.</div>
+
+<div class="refauthor">&mdash;&mdash;, and <span class="smcap">Cross, F. B.</span></div>
+
+<div class="reference">1954. River sturgeons of the American genus <i>Scaphirhynchus</i>: characters,
+distribution, and synonymy. Pap. Michigan Acad. Sci., Arts,
+and Letters, 39 (1953): 169-208.</div>
+
+<div class="refauthor smcap">Breukelman, J.</div>
+
+<div class="reference">1940. A collection of Kansas fish in the State University Museum. Trans.
+Kansas Acad. Sci., 43: 377-384.</div>
+
+<div class="refauthor smcap">Buchholz, M.</div>
+
+<div class="reference">1957. Age and growth of river carpsucker in Des Moines River, Iowa.
+Proc. Iowa Acad. Sci., 64: 589-600.</div>
+
+<div class="refauthor"><span class="smcap">Canfield, H. L.</span>, and <span class="smcap">Wiebe, A. H.</span></div>
+
+<div class="reference">1931. A cursory survey of the Blue River System of Nebraska. U. S.
+Dept. Comm., Bur. of Fisheries, Econ. Circ. 73: 1-10.</div>
+
+<div class="refauthor">
+<span class="smcap">Colby, C.</span>, <span class="smcap">Dillingham, H.</span>,
+<span class="smcap">Erickson, E.</span>, <span class="smcap">Jenks, G.</span>,
+<span class="smcap">Jones, J</span>., and <span class="smcap">Sinclair, R.</span>
+</div>
+
+<div class="reference">1956. The Kansas Basin, Pilot Study of a Watershed. Univ. of Kansas
+Press, Lawrence, ix + 103 pp.</div>
+
+<div class="refauthor smcap">Cragin, F. W.</div>
+
+<div class="reference">1885. Preliminary list of Kansas fishes. Bull. Washburn Lab. of Nat.
+Hist., 1 (3):105-111.</div>
+
+<div class="refauthor smcap">Cross, F. B.</div>
+
+<div class="reference">1950. Effects of sewage and of a headwaters impoundment on the
+fishes of Stillwater Creek in Payne County, Oklahoma. Amer.
+Midl. Nat., 43 (1):128-145.</div>
+
+<div class="reference">1954. Fishes of Cedar Creek and the South Fork of the Cottonwood
+River, Chase County, Kansas. Trans. Kansas Acad. Sci., 57 (3): 303-314.</div>
+
+<div class="refauthor"><span class="smcap">Eddy, S.</span>, and <span class="smcap">Surber, T.</span></div>
+
+<div class="reference">1947. Northern Fishes, with Special Reference to the Upper Mississippi
+Valley. Univ. of Minnesota Press, Minneapolis, xii + 276 pp.</div>
+
+<div class="refauthor"><span class="smcap">Evermann, B. W.</span>, and <span class="smcap">Cox, U. O.</span></div>
+
+<div class="reference">1896. Report upon the fishes of the Missouri River Basin. Appendix 5.
+Rept. U. S. Commissioner of Fish and Fisheries for 1894. pp. 325-429.</div>
+
+<div class="refauthor"><span class="smcap">Finnell, J. C.</span>, and <span class="smcap">Jenkins, R. M.</span></div>
+
+<div class="reference">1954. Growth of channel catfish in Oklahoma waters: 1954 revision.
+Oklahoma Fish Res. Lab. Rept. 41: ii + 1-37.</div>
+
+<div class="refauthor">&mdash;&mdash;, <span class="smcap">Jenkins, R. M.</span> and <span class="smcap">Hall, G. E.</span></div>
+
+<div class="reference">1956. The fishery resources of the Little River System, McCurtain County,
+Oklahoma. Oklahoma Fish. Res. Lab. Rept. 55: ii + 1-82.</div>
+
+<div class="refauthor smcap">Flora, S. D.</div>
+
+<div class="reference">1948. Climate of Kansas. Rept. Kansas State Board of Agri., 67 (285):
+xii + 1-320.</div>
+
+<div class="refauthor"><span class="smcap">Forbes, S. A.</span>, and <span class="smcap">Richardson, R. E.</span>
+<span class="pagenum"><a name="Page_440" id="Page_440">[440]</a></span></div>
+
+<div class="reference">1920. The Fishes of Illinois. Nat. Hist. Survey of Illinois. Illinois Printing
+Co., Danville, cxxxi + 357 pp.</div>
+
+<div class="refauthor"><span class="smcap">Frye, J. C.</span>, and <span class="smcap">Leonard, A. B.</span></div>
+
+<div class="reference">1952. Pleistocene geology of Kansas. Kansas Geol. Survey, Bull. 99: 1-230.</div>
+
+<div class="refauthor smcap">Gilbert, C. H.</div>
+
+<div class="reference">1886. Third series of notes on the fishes of Kansas. Bull. Washburn Lab.
+of Nat. Hist., 1 (6): 207-211.</div>
+
+<div class="refauthor smcap">Graham, I. D.</div>
+
+<div class="reference">1885. Preliminary list of Kansas fishes. Trans. Kansas Acad. Sci., 9: 69-78.</div>
+
+<div class="refauthor smcap">Harrison, H. M.</div>
+
+<div class="reference">1956. Angling for channel catfish. <i>In</i> Iowa Fish and Fishing, by J. R.
+Harlan and E. B. Speaker. Iowa State Cons. Comm., Des Moines.
+Pp. 202-212.</div>
+
+<div class="refauthor smcap">Hubbs, C. L.</div>
+
+<div class="reference">1945. Corrected distributional records for Minnesota fishes. Copeia, 1945
+(1):13-22.</div>
+
+<div class="refauthor">&mdash;&mdash;, and <span class="smcap">Ortenburger, A. I.</span></div>
+
+<div class="reference">1929. Further notes on the fishes of Oklahoma with descriptions of new
+species of Cyprinidae. Pub. Univ. Oklahoma Biol. Survey, 1 (2):
+17-43.</div>
+
+<div class="refauthor">&mdash;&mdash;, and <span class="smcap">Lagler, K. F.</span></div>
+
+<div class="reference">1947. Fishes of the Great Lakes Region. Cranbrook Inst. of Sci., Bull.
+26: xi + 1-186.</div>
+
+<div class="refauthor">&mdash;&mdash;, and <span class="smcap">Bailey, R. M.</span></div>
+
+<div class="reference">1952. Identification of <i>Oxygeneum pulverulentum</i> Forbes, from Illinois,
+as a hybrid cyprinid fish. Pap. Michigan Acad. Sci., Arts, and
+Letters, 37 (1951): 143-152.</div>
+
+<div class="refauthor smcap">Jennings, D.</div>
+
+<div class="reference">1942. Kansas fish in the Kansas State College Museum at Manhattan.
+Trans. Kansas Acad. Sci., 45: 363-366.</div>
+
+<div class="refauthor smcap">Kansas Water Resources Fact-finding and Research Committee</div>
+
+<div class="reference">1955. Water in Kansas. A Report to the Kansas State Legislature ...
+Univ. of Kansas. 1-216 pp.</div>
+
+<div class="refauthor smcap">Kincer, J. B.</div>
+
+<div class="reference">1941. Climate and weather data for the United States. <i>In</i> Climate and
+Man, Yearbook of Agri. for 1941. House Doc. 27. pp. 685-699.</div>
+
+<div class="refauthor smcap">Lagler, K. F.</div>
+
+<div class="reference">1952. Freshwater Fishery Biology. Wm. C. Brown Co., Dubuque, Iowa.
+x + 360 pp.</div>
+
+<div class="refauthor smcap">Lugn, A. L.</div>
+
+<div class="reference">1935. The Pleistocene geology of Nebraska. Nebraska Geol. Survey,
+Bull. 10, 2nd series: 1-223.</div>
+
+<div class="refauthor smcap">Marzolf, R. C.</div>
+
+<div class="reference">1955. Use of pectoral spines and vertebrae for determining age and rate
+of growth of the channel catfish. Jour. Wildl. Mgmt., 19 (2): 243-249.</div>
+
+<p><span class="pagenum"><a name="Page_441" id="Page_441">[441]</a></span></p>
+
+<div class="refauthor smcap">Meek, S. E.</div>
+
+<div class="reference">1895. Notes on the fishes of western Iowa and eastern Nebraska. Bull.
+U. S. Fish Comm., 14 (1894): 133-138.</div>
+
+<div class="refauthor smcap">Metcalf, A. L.</div>
+
+<div class="reference">1957. Fishes of Chautauqua, Cowley and Elk counties, Kansas. Univ.
+Kansas Publ., Mus. Nat. Hist., 11:345-400.</div>
+
+<div class="refauthor"><span class="smcap">Minckley, W. L.</span>, and <span class="smcap">Cross, F. B.</span></div>
+
+<div class="reference">In press. Habitat, distribution, and abundance of <i>Notropis topeka</i> (Gilbert)
+in Kansas. Amer. Midl. Nat.</div>
+
+<div class="refauthor smcap">Monfort, E.</div>
+
+<div class="reference">1956. A layman looks at water. Trans. Kansas Acad. Sci., 59 (1):118-123.</div>
+
+<div class="refauthor smcap">Moore, G. A.</div>
+
+<div class="reference">1957. Fishes. <i>In</i> Vertebrates of the United States, by W. Blair, A. Blair,
+P. Brodkorb, F. Cagle, and G. Moore. McGraw-Hill Book Co.,
+New York, N. Y. pp. 31-210.</div>
+
+<div class="refauthor"><span class="smcap">Moore, R. C.</span>, and <span class="smcap">Landes, K. K.</span></div>
+
+<div class="reference">1937. Geologic map of Kansas. Scale 1:500,000. Kansas Geol. Survey.</div>
+
+<div class="refauthor smcap">Nebraska State Planning Board.</div>
+
+<div class="reference">1936. Water resources of Nebraska. Mimeo. by Nebraska State Planning
+Board. Lincoln, xxviii + 695 pp.</div>
+
+<div class="refauthor smcap">Pfeiffer, R. A.</div>
+
+<div class="reference">1955. Studies on the life history of the rosyface shiner, <i>Notropis rubellus</i>.
+Copeia, 1955 (2):95-104.</div>
+
+<div class="refauthor smcap">Raney, E. C.</div>
+
+<div class="reference">1947. Subspecies and breeding behavior of the cyprinid fish <i>Notropis
+procne</i> (Cope). Copeia, 1947 (2):103-109.</div>
+
+<div class="refauthor smcap">Schoewe, W. H.</div>
+
+<div class="reference">1953. The geography of Kansas, Part III&mdash;concluded, hydrogeography.
+Trans. Kansas Acad. Sci., 56 (2):131-190.</div>
+
+<div class="refauthor smcap">Snedecor, G. W.</div>
+
+<div class="reference">1956. Statistical Methods. Iowa State College Press, Ames. xiii + 534 pp.</div>
+
+<div class="refauthor smcap">Starrett, W. C.</div>
+
+<div class="reference">1950. Distribution of the fishes of Boone County, Iowa, with special reference
+to the minnows and darters. Amer. Midl. Nat., 43 (1):
+112-127.</div>
+
+<div class="refauthor smcap">Trautman, M. B.</div>
+
+<div class="reference">1957. The Fishes of Ohio. Waverly Press, Inc., Baltimore, Md. xvii +
+683 pp.</div>
+
+<div class="refauthor smcap">U. S. Fish and Wildlife Service.</div>
+
+<div class="reference">1952. A one-year creel census and evaluation of the Republican River,
+Nebraska and Kansas, 1951. Mimeo. by the Staff, Missouri River
+Basin Studies, Billings, Mont. 29 pp., Appendix.</div>
+
+<div class="reference">1953. A preliminary report on fish and wildlife resources in relation to
+the water development plan for the Tuttle Creek Dam and Reservoir,
+Big Blue River, Missouri River Basin, Kansas. Mimeo. by the
+Staff, Missouri River Basin Studies, Billings, Mont. 25 pp.</div>
+
+<p><span class="pagenum"><a name="Page_442" id="Page_442">[442]</a></span></p>
+
+<div class="refauthor smcap">Van Orman, C. R.</div>
+
+<div class="reference">1956. Surface water&mdash;its control and retention for use. Trans. Kansas
+Acad. Sci., 59 (1):105-110.</div>
+
+<div class="refauthor smcap">Walters, K. L.</div>
+
+<div class="reference">1954. Geology and ground-water resources of Marshall County, Kansas.
+Kansas Geol. Survey, Bull. 106:1-116.</div>
+<br />
+<br />
+<p><i>Transmitted December 19, 1958.</i></p>
+<br />
+<br />
+
+<div class="center">
+<img src="images/square.png" width="16" height="17" alt="" title="" /><br />
+27-7080
+</div>
+<br />
+<br />
+
+<p><span class="pagenum"><a name="Page_i" id="Page_i">[i]</a></span></p>
+
+<div class="caption2">UNIVERSITY OF KANSAS PUBLICATIONS<br />
+MUSEUM OF NATURAL HISTORY</div>
+
+<p>Institutional libraries interested in publications exchange may obtain this
+series by addressing the Exchange Librarian, University of Kansas Library,
+Lawrence, Kansas. Copies for individuals, persons working in a particular
+field of study, may be obtained by addressing instead the Museum of Natural
+History, University of Kansas, Lawrence, Kansas. There is no provision for
+sale of this series by the University Library, which meets institutional requests,
+or by the Museum of Natural History, which meets the requests of individuals.
+Nevertheless, when individuals request copies from the Museum, 25 cents should
+be included, for each separate number that is 100 pages or more in length, for
+the purpose of defraying the costs of wrapping and mailing.</p>
+
+<p>* An asterisk designates those numbers of which the Museum's supply (not the Library's
+supply) is exhausted. Numbers published to date, in this series, are as follows:</p>
+
+<table class="pub_list" summary="pub_list">
+<tr><td class="text_rt">&nbsp;Vol.&nbsp;&nbsp;1.</td><td colspan="2" class="justify">Nos. 1-26 and index. Pp. 1-638, 1946-1950.</td></tr>
+
+<tr><td class="text_rt">*Vol.&nbsp;&nbsp;2.</td><td colspan="2" class="justify">(Complete) Mammals of Washington. By Walter W. Dalquest. Pp. 1-444, 140
+figures in text. April 9, 1948.</td></tr>
+
+<tr><td class="text_rt">Vol.&nbsp;&nbsp;3.</td><td class="text_rt">*1.</td><td class="justify">The avifauna of Micronesia, its origin, evolution, and distribution. By Rollin
+H. Baker. Pp. 1-359, 16 figures in text. June 12, 1951.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">*2.</td><td class="justify">A quantitative study of the nocturnal migration of birds. By George H.
+Lowery, Jr. Pp. 361-472, 47 figures in text. June 29, 1951.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">3.</td><td class="justify">Phylogeny of the waxwings and allied birds. By M. Dale Arvey. Pp. 473-530,
+49 figures in text, 13 tables. October 10, 1951.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">4.</td><td class="justify">Birds from the state of Veracruz, Mexico. By George H. Lowery, Jr., and
+Walter W. Dalquest. Pp. 531-649, 7 figures in text, 2 tables. October 10, 1951.</td></tr>
+
+<tr><td>&nbsp;</td><td colspan="2" class="justify">Index. Pp. 651-681.</td></tr>
+
+<tr><td class="text_rt">*Vol.&nbsp;&nbsp;4.</td><td colspan="2" class="justify">(Complete) American weasels. By E. Raymond Hall. Pp. 1-466, 41 plates, 31
+figures in text. December 27, 1951.</td></tr>
+
+<tr><td class="text_rt">Vol.&nbsp;&nbsp;5.</td><td colspan="2" class="justify">Nos. 1-37 and index. Pp. 1-676, 1951-1953.</td></tr>
+
+<tr><td class="text_rt">*Vol.&nbsp;&nbsp;6.</td><td colspan="2" class="justify">(Complete) Mammals of Utah, <i>taxonomy and distribution</i>. By Stephen D.
+Durrant. Pp. 1-549, 91 figures in text, 30 tables. August 10, 1952.</td></tr>
+
+<tr><td class="text_rt vtop">Vol.&nbsp;&nbsp;7.</td><td class="text_rt vtop">*1.</td><td class="justify">Mammals of Kansas.&nbsp; By E. Lendell Cockrum. Pp. 1-303, 73 figures in
+text, 37 tables. August 25, 1952.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">2.</td><td class="justify">Ecology of the opossum on a natural area in northeastern Kansas. By Henry
+S. Fitch and Lewis L. Sandidge. Pp. 305-338, 5 figures in text. August 24, 1953.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">3.</td><td class="justify">The silky pocket mice (Perognathus flavus) of Mexico. By Rollin H. Baker.
+Pp. 339-347, 1 figure in text. February 15, 1954.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">4.</td><td class="justify">North American jumping mice (Genus Zapus). By Philip H. Krutzsch. Pp.
+349-472, 47 figures in text, 4 tables. April 21, 1954.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">5.</td><td class="justify">Mammals from Southeastern Alaska. By Rollin H. Baker and James S.
+Findley. Pp. 473-477. April 21, 1954.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">6.</td><td class="justify">Distribution of Some Nebraskan Mammals. By J. Knox Jones, Jr. Pp. 479-487.
+April 21, 1954.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">7.</td><td class="justify">Subspeciation in the montane meadow mouse, Microtus montanus, in Wyoming
+and Colorado. By Sydney Anderson. Pp. 489-506, 2 figures in text. July 23, 1954.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">8.</td><td class="justify">A new subspecies of bat (Myotis velifer) from southeastern California and
+Arizona. By Terry A. Vaughn. Pp. 507-512. July 23, 1954.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">9.</td><td class="justify">Mammals of the San Gabriel mountains of California. By Terry A. Vaughn.
+Pp. 513-582, 1 figure in text, 12 tables. November 15, 1954.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">10.</td><td class="justify">A new bat (Genus Pipistrellus) from northeastern Mexico. By Rollin H.
+Baker. Pp. 583-586. November 15, 1954.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">11.</td><td class="justify">A new subspecies of pocket mouse from Kansas. By E. Raymond Hall. Pp.
+587-590. November 15, 1954.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">12.</td><td class="justify">Geographic variation in the pocket gopher, Cratogeomys castanops, in Coahuila,
+Mexico. By Robert J. Russell and Rollin H. Baker. Pp. 591-608. March
+15, 1955.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">13.</td><td class="justify">A new cottontail (Sylvilagus floridanus) from northeastern Mexico. By Rollin
+H. Baker. Pp. 609-612. April 8, 1955.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">14.</td><td class="justify">Taxonomy and distribution of some American shrews. By James S. Findley.
+Pp. 613-618. June 10, 1955.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">15.</td><td class="justify">The pigmy woodrat, Neotoma goldmani, its distribution and systematic position.
+By Dennis G. Rainey and Rollin H. Baker. Pp. 619-624, 2 figs. in
+text. June 10, 1955.</td></tr>
+
+<tr><td>&nbsp;</td><td colspan="2" class="justify">Index. Pp. 625-651.</td></tr>
+
+<tr><td class="text_rt vtop">Vol.&nbsp;&nbsp;8.<span class="pagenum"><a name="Page_ii" id="Page_ii">[ii]</a></span></td>
+<td class="text_rt vtop">1.</td><td class="justify">Life history and ecology of the five-lined skink, Eumeces fasciatus. By Henry
+S. Fitch. Pp. 1-156, 26 figs. in text. September 1, 1954.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">2.</td><td class="justify">Myology and serology of the Avian Family Fringillidae, a taxonomic study.
+By William B. Stallcup. Pp. 157-211, 23 figures in text, 4 tables. November 15, 1954.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">3.</td><td class="justify">An ecological study of the collared lizard (Crotaphytus collaris). By Henry
+S. Fitch. Pp. 213-274, 10 figures in text. February 10, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">4.</td><td class="justify">A field study of the Kansas ant-eating frog, Gastrophryne olivacea. By Henry
+S. Fitch. Pp. 275-306, 9 figures in text. February 10, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">5.</td><td class="justify">Check-list of the birds of Kansas. By Harrison B. Tordoff. Pp. 307-359, 1
+figure in text. March 10, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">6.</td><td class="justify">A population study of the prairie vole (Microtus ochrogaster) in northeastern
+Kansas. By Edwin P. Martin. Pp. 361-416, 19 figures in text. April 2, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">7.</td><td class="justify">Temperature responses in free-living amphibians and reptiles of northeastern
+Kansas. By Henry S. Fitch. Pp. 417-476, 10 figures in text, 6 tables. June 1, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">8.</td><td class="justify">Food of the crow, Corvus brachyrhynchos Brehm, in south-central Kansas. By
+Dwight Platt. Pp. 477-498, 4 tables. June 8, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">9.</td><td class="justify">Ecological observations on the woodrat, Neotoma floridana. By Henry S.
+Fitch and Dennis G. Rainey. Pp. 499-533, 3 figures in text. June 12, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt vtop">10.</td><td class="justify">Eastern woodrat, Neotoma floridana: Life history and ecology. By Dennis G.
+Rainey. Pp. 535-646, 12 plates, 13 figures in text August 15, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td colspan="2" class="justify">Index. Pp. 647-675.</td></tr>
+
+<tr><td class="text_rt">Vol.&nbsp;&nbsp;9.</td><td class="text_rt">1.</td><td class="justify">Speciation of the wandering shrew. By James S. Findley. Pp. 1-68, 18
+figures in text. December 10, 1955.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">2.</td><td class="justify">Additional records and extension of ranges of mammals from Utah. By
+Stephen D. Durrant, M. Raymond Lee, and Richard M. Hansen. Pp. 69-80.
+December 10, 1955.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">3.</td><td class="justify">A new long-eared myotis (Myotis evotis) from northeastern Mexico. By Rollin
+H. Baker and Howard J. Stains. Pp. 81-84. December 10, 1955.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">4.</td><td class="justify">Subspeciation in the meadow mouse, Microtus pennsylvanicus, in Wyoming.
+By Sydney Anderson. Pp. 85-104, 2 figures in text. May 10, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">5.</td><td class="justify">The condylarth genus Ellipsodon. By Robert W. Wilson. Pp. 105-116, 6
+figures in text. May 19, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">6.</td><td class="justify">Additional remains of the multituberculate genus Eucosmodon. By Robert
+W. Wilson. Pp. 117-123, 10 figures in text. May 19, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">7.</td><td class="justify">Mammals of Coahulia, Mexico. By Rollin H. Baker. Pp. 125-335, 75 figures
+in text. June 15, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">8.</td><td class="justify">Comments on the taxonomic status of Apodemus peninsulae, with description
+of a new subspecies from North China. By J. Knox Jones, Jr. Pp. 337-346,
+1 figure in text, 1 table. August 15, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">9.</td><td class="justify">Extensions of known ranges of Mexican bats. By Sydney Anderson. Pp.
+347-351. August 15, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">10.</td><td class="justify">A new bat (Genus Leptonycteris) from Coahulia. By Howard J. Stains.
+Pp. 353-356. January 21, 1957.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">11.</td><td class="justify">A new species of pocket gopher (Genus Pappogeomys) from Jalisco, Mexico.
+By Robert J. Russell. Pp. 357-361. January 21, 1957.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">12.</td><td class="justify">Geographic variation in the pocket gopher, Thomomys bottae, in Colorado.
+By Phillip M. Youngman. Pp. 363-387, 7 figures in text. February 21, 1958.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">13.</td><td class="justify">New bog lemming (genus Synaptomys) from Nebraska. By J. Knox Jones,
+Jr. Pp. 385-388. May 12, 1958.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">14.</td><td class="justify">Pleistocene bats from San Josecito Cave, Nuevo León, México. By J. Knox
+Jones, Jr. Pp. 389-396. December 19, 1958.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">15.</td><td class="justify">New subspecies of the rodent Baiomys from Central America. By Robert
+L. Packard. Pp. 397-404. December 19, 1958.</td></tr>
+
+<tr><td>&nbsp;</td><td colspan="2" class="justify">More numbers will appear in volume 9.</td></tr>
+
+<tr><td class="text_rt">Vol.&nbsp;10.</td><td class="text_rt">1.</td><td class="justify">Studies of birds killed in nocturnal migration. By Harrison B. Tordoff and
+Robert M. Mengel. Pp. 1-44, 6 figures in text, 2 tables. September 12, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">2.</td><td class="justify">Comparative breeding behavior of Ammospiza caudacuta and A. maritima.
+By Glen E. Woolfenden. Pp. 45-75, 6 plates, 1 figure. December 20, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">3.</td><td class="justify">The forest habitat of the University of Kansas Natural History Reservation.
+By Henry S. Fitch and Ronald R. McGregor. Pp. 77-127, 2 plates, 7 figures
+in text, 4 tables. December 31, 1956.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">4.</td><td class="justify">Aspects of reproduction and development in the prairie vole (Microtus ochrogaster).
+By Henry S. Fitch. Pp. 129-161, 8 figures in text, 4 tables. December
+19, 1957.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">5.</td><td class="justify">Birds found on the Arctic slope of northern Alaska. By James W. Bee.
+Pp. 163-211, plates 9-10, 1 figure in text. March 12, 1958.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">6.</td><td class="justify">The wood rats of Colorado: distribution and ecology. By Robert B. Finley,
+Jr. Pp. 213-552, 34 plates, 8 figures in text, 35 tables. November 7, 1958.</td></tr>
+
+<tr><td>&nbsp;</td><td colspan="2" class="justify">More numbers will appear in volume 10.</td></tr>
+
+<tr><td class="text_rt">Vol.&nbsp;11.<span class="pagenum"><a name="Page_iii" id="Page_iii">[iii]</a></span></td>
+<td class="text_rt">1.</td><td class="justify">The systematic status of the colubrid snake, Leptodeira discolor Günther.
+By William E. Duellman. Pp. 1-9, 4 figs. July 14, 1958.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">2.</td><td class="justify">Natural history of the six-lined racerunner, Cnemidophorus sexlineatus. By
+Henry S. Fitch. Pp. 11-62, 9 figs., 9 tables. September 19, 1958.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">3.</td><td class="justify">Home ranges, territories, and seasonal movements of vertebrates of the
+Natural History Reservation. By Henry S. Fitch. Pp. 63-326, 6 plates, 24
+figures in text, 3 tables. December 12, 1958.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">4.</td><td class="justify">A new snake of the genus Geophis from Chihuahua, Mexico. By John M.
+Legler. Pp. 327-334, January 28, 1959.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">5.</td><td class="justify">A new tortoise, genus Gopherus, from north-central Mexico. By John M.
+Legler. Pp. 335-343, April 24, 1959.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">6.</td><td class="justify">Fishes of Chautauqua, Cowley and Elk counties, Kansas. By Artie L.
+Metcalf. Pp. 345-400, 2 plates, 2 figures in text, 10 tables. May 6, 1959.</td></tr>
+
+<tr><td>&nbsp;</td><td class="text_rt">7.</td><td class="justify">Fishes of the Big Blue River Basin, Kansas. By W. L. Minckley. Pp. 401-442,
+2 plates, 4 figures in text, 5 tables. May 8, 1959.</td></tr>
+
+<tr><td>&nbsp;</td><td colspan="2" class="justify">More numbers will appear in volume 11.</td></tr>
+</table>
+<br />
+<br />
+
+<div class="trans_notes">
+<div class="caption2">Transcriber's Notes</div>
+
+<p>Except as noted below, the text presented herein is that contained in
+the original printed version. Minor corrections (such as missing
+punctuation) may have been corrected. The original version had a
+list of publications printed inside the cover and inside and on the
+back cover. The cover page was not retained as it is a copy of the
+first page and the list inside the cover was moved past the end of the
+article.</p>
+
+<div class="caption3">Typographical Corrections</div>
+<br />
+<table summary="Correction List">
+<tr><td>Page</td><td>&nbsp;&nbsp;&nbsp;</td><td>Correction</td></tr>
+<tr><td class="bt2">&nbsp;&nbsp;408</td><td>&nbsp;</td><td class="bt2">Phenophthalein &#8594; Phenolphthalein</td></tr>
+</table>
+<br />
+</div>
+</div>
+
+
+
+
+
+
+
+
+
+<pre>
+
+
+
+
+
+End of the Project Gutenberg EBook of Fishes of the Big Blue River Basin,
+Kansas, by W. L. Minckley
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+</body>
+</html>
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