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+<p><b>The Student&rsquo;s Elements of Geology</b></p>
+
+<hr>
+<p class="page"><a name="page 439">[ 439 ]</a></p>
+
+<p>&nbsp;</p>
+
+<center><b>Chapter XXV</b><br>
+<br>
+DEVONIAN OR OLD RED SANDSTONE GROUP.</center>
+
+<p class="intro">Classification of the Old Red Sandstone in
+Scotland and in Devonshire. &mdash; Upper Old Red Sandstone in
+Scotland, with Fish and Plants. &mdash; Middle Old Red Sandstone.
+&mdash; Classification of the Ichthyolites of the Old Red, and
+their Relation to Living Types. &mdash; Lower Old Red Sandstone,
+with Cephalaspis and Pterygotus. &mdash; Marine or Devonian Type of
+Old Red Sandstone. &mdash; Table of Devonian Series. &mdash; Upper
+Devonian Rocks and Fossils. &mdash; Middle. &mdash; Lower. &mdash;
+Eifel Limestone of Germany. &mdash; Devonian of Russia. &mdash;
+Devonian Strata of the United States and Canada. &mdash; Devonian
+Plants and Insects of Canada.</p>
+
+<p><b>Classification of the two Types of Old Red
+Sandstone.</b>&mdash;We have seen that the Carboniferous strata are
+surmounted by the Permian and Trias, both originally included in
+England under the name &ldquo;New Red Sandstone,&rdquo; from the
+prevailing red colour of the strata. Under the coal came other red
+sandstones and shales which were distinguished by the title of
+&ldquo;Old Red Sandstone.&rdquo; Afterwards the name of
+&ldquo;Devonian&rdquo; was given by Sir R. Murchison and Professor
+Sedgwick to marine fossiliferous strata which, in the south of
+England, occupy a similar position between the overlying coal and
+the underlying Silurian formations.</p>
+
+<p>It may be truly said that in the British Isles the rocks of this
+age present themselves in their mineral aspect, and even to some
+extent in their fossil contents, under two very different forms;
+the one as distinct from the other as are often lacustrine or
+fluviatile from marine strata. It has indeed been suggested that by
+far the greater part of the deposits belonging to what may be
+termed the Old Red Sandstone type are of fresh-water origin. The
+number of land-plants, the character of the fishes, and the fact
+that the only shell yet discovered belongs to the genus <i>
+Anodonta,</i> must be allowed to lend no small countenance to this
+opinion. In this case the difficulty of classification when the
+strata of this type are compared in different regions, even where
+they are contiguous, may arise partly from their having been formed
+in distinct hydrographical basins, or in the neighbourhood of the
+land in shallow parts of the sea into which large bodies of
+fresh-water entered, and where no marine mollusca or corals could
+flourish. Under such geographical conditions the limited extent of
+some kinds of sediment, as well as the</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 440">[ 440 ]</a></p>
+
+<p>absence of those marine forms by which we are able to identify
+or contrast marine formations, may be explained, while the great
+thickness of the rocks, which might seem at first sight to require
+a corresponding depth of water, can often be shown to have been due
+to the gradual sinking down of the bottom of the estuary or sea
+where the sediment was accumulated.</p>
+
+<p>Another active cause of local variation in Scotland was the
+frequency of contemporaneous volcanic eruptions; some of the rocks
+derived from this source, as between the Grampians and the Tay,
+having formed islands in the sea, and having been converted into
+shingle and conglomerate, before the upper portions of the red
+shales and sandstones were superimposed.</p>
+
+<p>The dearth of calcareous matter over wide areas is
+characteristic of the Old Red Sandstone. This is, no doubt, in
+great part due to the absence of shells and corals; but why should
+these be so generally wanting in all sedimentary rocks the colour
+of which is determined by the red oxide of iron? Some geologists
+are of opinion that the waters impregnated with this oxide were
+prejudicial to living beings, others that strata permeated with
+this oxide would not preserve such fossil remains.</p>
+
+<p>In regard to the two types, the Old Red Sandstone and the
+Devonian, I shall first treat of them separately, and then allude
+to the proofs of their having been to a great extent
+contemporaneous. That they constitute a series of rocks
+intermediate in date between the lowest Carboniferous and the
+uppermost Silurian is not disputed by the ablest geologists; and it
+can no longer be contended that the Upper, Middle, and Lower Old
+Red Sandstone preceded in date the three divisions to which, by aid
+of the marine shells, the Devonian rocks have been referred, while,
+on the other hand, we have not yet data for enabling us to affirm
+to what extent the subdivisions of the one series may be the
+equivalents in time of those of the other.</p>
+
+<p><b>Upper Old Red Sandstone.</b>&mdash;The highest beds of the
+series in Scotland, lying immediately below the coal in Fife, are
+composed of yellow sandstone well seen at Dura Den, near Coupar, in
+Fife, where, although the strata contain no mollusca, fish have
+been found abundantly, and have been referred to the genera <i>
+Holoptychius, Pamphractus, Glyptopomus,</i> and many others. In the
+county of Cork, in Ireland, a similar yellow sandstone occurs
+containing fish of genera characteristic of the Scotch Old Red
+Sandstone, as for example Coccosteus (a form represented by many
+species in the</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 441">[ 441 ]</a></p>
+
+<img src="../images4/fig494.jpg" width="212" height="150" alt=
+"Fig. 494: Anodonta Jukesii." align="right">
+
+<p>Old Red Sandstone and by one only in the Carboniferous group),
+and <i>Glytolepis</i> and <i>Asterolepis,</i> both exclusively
+confined to the &ldquo;Old Red.&rdquo; In the same Irish sandstone
+at Kiltorkan has been found an <i>Anodonta</i> or fresh-water
+mussel, the only shell hitherto discovered in the Old Red Sandstone
+of the British Isles (see Fig. 494).</p>
+
+<img src="../images4/fig495.jpg" width="124" height="271" alt=
+"Fig. 495: Bifurcating branch of Lepidendron Griffithsii." align=
+"left"> <img src="../images4/fig496.jpg" width="114" height="335" alt=
+"Fig. 496: Pal&aelig;opteris Hibernia." align="right">
+
+<p>In the same formation are found the fern (Fig. 496) and the <i>
+Lepidodendron</i> (Fig. 495), and other species of plants, some of
+which, Professor Heer remarks, agree specifically with species from
+the lower carboniferous beds. This induces him to lean to the
+opinion long ago advocated by Sir Richard Griffiths, that the
+yellow sandstone, in spite of its fish remains, should be classed
+as Lower Carboniferous, an opinion which I am not yet prepared to
+adopt. Between the Mountain Limestone and the yellow sandstone in
+the south-west of Ireland there intervenes a formation no less than
+5000 feet thick, called the &ldquo;Carboniferous slate,&rdquo; and
+at the base of this, in some places, are local deposits, such as
+the Glengariff Grits, which appear to be beds of passage between
+the Carboniferous and Old Red Sandstone groups.</p>
+
+<p>It is a remarkable result of the recent examination of the
+fossil flora of Bear Island, latitude 74&deg; 30' N., that
+Professor Heer has described as occurring in that part of the
+Arctic region (nearly twenty-six degrees to the north of the Irish
+locality) a flora agreeing in several of its species with that of
+the yellow sandstones of Ireland. This Bear Island flora is
+believed by Professor Heer to comprise species of plants some of
+which ascend even to the higher stages of the European
+Carboniferous formation, or as high as the Mountain Limestone and
+Millstone Grit. Pal&aelig;ontologists have long maintained that</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 442">[ 442 ]</a></p>
+
+<p>the same species which have a wide range in space are also the
+most persistent in time, which may prepare us to find that some
+plants having a vast geographical range may also have endured from
+the period of the Upper Devonian to that of the Millstone Grit.</p>
+
+<img src="../images4/fig497.jpg" width="141" height="165" alt=
+"Fig. 497: Scale of Holoptychius nobilissimus." align="left">
+
+<p>Outliers of the Upper &ldquo;Old Red&rdquo; occur unconformably
+on older members of the group, and the formation represented at
+Whiteness, near Arbroath, <i>a,</i> <a href="../images/fig55.jpg">Fig.
+55,</a> may probably be one of these outliers, though the want of
+organic remains renders this uncertain. It is not improbable that
+the beds given in this section as Nos. 1, 2, and 3, may all belong
+to the early part of the period of the Upper Old Red, as some
+scales of <i>Holoptychius nobilissimus</i> have been found
+scattered through these beds, No. 2, in Strathmore. Another nearly
+allied <i>Holoptychius</i> occurs in Dura Den, see Fig. 498 of this
+fish and also Fig. 497 of one of its scales, as these last are
+often the only parts met with; being scattered in Forfarshire
+through red-coloured shales and sandstones, as are scales of a
+large species of the same genus in a corresponding matrix in
+Herefordshire.* The number of fish obtained from the British Upper
+Old Red Sandstone amounts to fifteen species referred to eleven
+genera.</p>
+
+<center><img src="../images4/fig498.jpg" width="389" height="230" alt=
+"Fig. 498: Holoptychius, as restored by Professor Huxley.">
+</center>
+
+<p>Sir R. Murchison groups with this upper division of the Old Red
+of Scotland certain light-red and yellow sandstones and grits which
+occur in the northernmost part of the mainland, and extend also
+into the Orkney and Shetland Islands.</p>
+
+<p class="fnote">* Siluria, 4th ed., p. 265.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 443">[ 443 ]</a></p>
+
+<p>They contain Calamites and other plants which agree generically
+with Carboniferous forms.</p>
+
+<p><b>Middle Old Red Sandstone.</b>&mdash;In the northern part of
+Scotland there occur a great series of bituminous schists and
+flagstones, to the fossil fish of which attention was first called
+by the late Hugh Miller. They were afterwards described by Agassiz,
+and the rocks containing them were examined by Sir R. Murchison and
+Professor Sedgwick, in Caithness, Cromarty, Moray, Nairn, Gamrie in
+Banff, and the Orkneys and Shetlands, in which great numbers of
+fossil fish have been found. These were at first supposed to be the
+oldest known vertebrate animals, as in Cromarty the beds in which
+they occur seem to form the base of the Old Red system resting
+almost immediately on the crystalline or metamorphic rocks. But in
+fact these fish-bearing beds, when they are traced from north to
+south, or to the central parts of Scotland, thin out, so that their
+relative age to the Lower Old Red Sandstone, presently to be
+mentioned, was not at first detected, the two formations not
+appearing in superposition in the same district. In Caithness,
+however, many hundred feet below the fish-zone of the middle
+division, remains of <i>Pteraspis</i> were found by Mr. Peach in
+1861. This genus has never yet been found in either of the two
+higher divisions of the Old Red Sandstone, and confirms Sir R.
+Murchison&rsquo;s previous suspicion that the rocks in which it
+occurs belong to the Lower &ldquo;Old Red,&rdquo; or agree in age
+with the Arbroath paving-stone.*</p>
+
+<p><i>Fossil Fish of the Middle Old Red Sandstone.</i>&mdash;The
+Devonian fish were referred by Agassiz to two of his great orders,
+namely, the Placoids and Ganoids. Of the first of these, which in
+the Recent period comprise the shark, the dog-fish, and the ray, no
+entire skeletons are preserved, but fin-spines, called
+ichthyodorulites, and teeth occur. On such remains the genera <i>
+Onchus, Odontacanthus,</i> and <i>Ctenodus,</i> a supposed
+cestraciont, and some others, have been established.</p>
+
+<p>By far the greater number of the Old Red Sandstone fishes belong
+to a sub-order of Ganoids instituted by Huxley in 1861, and for
+which he has proposed the name of <i>
+Crossopterygid&aelig;</i>,&dagger; or the fringe-finned, in
+consideration of the peculiar manner in which the fin-rays of the
+paired fins are arranged so as to form a fringe round a central
+lobe, as in the Polypterus (see <i>a,</i> Fig. 499), a genus of
+which there are several species now inhabiting the Nile and other
+African rivers. The reader will at once recognise in <i>
+Osteolepis</i> (Fig. 500), one of the common fishes of the Old Red
+Sandstone, many points of</p>
+
+<p class="fnote">* Siluria, 4th ed., p. 258.<br>
+&dagger; Abridged from <i>crossotos,</i> a fringe, and <i>
+pteryx,</i> a fin.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 444">[ 444 ]</a></p>
+
+<center><img src="../images4/fig499.jpg" width="408" height="159" alt=
+"Fig. 499: Polypterus. Living in the Nile and other rivers.">
+</center>
+
+<p>analogy with <i>Polypterus.</i> They not only agree in the
+structure of the fin, at first pointed out by Huxley, but also in
+the position of the pectoral, ventral, and anal fins, and in having
+an elongated body and rhomboidal scales. On the other hand, the
+tail is more symmetrical in the recent fish, which has also an
+apparatus of dorsal finlets of a very abnormal character, both as
+to number and structure. As to the dorsals of <i>Osteolepis,</i>
+they are regular in structure and position, having nothing
+remarkable about them, except that there are two of them, which is
+comparatively unusual in living fish.</p>
+
+<center><img src="../images4/fig500.jpg" width="373" height="144" alt=
+"Fig. 500: Restoration of Osteolepis."></center>
+
+<p>Among the &ldquo;fringe-finned&rdquo; Ganoids we find some with
+rhomboidal scales, such as <i>Osteolepis,</i> Fig. 500; others with
+cycloidal scales, as <i>Holoptychius,</i> before mentioned (see
+Fig. 498). In the genera <i>Dipterus</i> and <i>Diplopterus,</i> as
+Hugh Miller pointed out, and in several other of the fringe-finned
+genera, as in <i>Gyroptychius</i> and <i>Glyptolepis,</i> the two
+dorsals are placed far backward, or directly over the ventral and
+anal fins. The <i>Asterolepis</i> was a ganoid fish of gigantic
+dimensions. <i>A. Asmusii,</i> Eichwald, a species characteristic
+of the Old Red Sandstone of Russia, as well as that of Scotland,
+attained the length of between twenty and thirty feet. It was
+clothed with strong bony armour, embossed with star-like tubercles,
+but it had only a cartilaginous skeleton. The mouth was furnished
+with two rows of teeth, the outer ones small and fish-like, the
+inner larger and with a reptilian character. The <i>Asterolepis</i>
+occurs also in the Devonian rocks of North America.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 445">[ 445 ]</a></p>
+
+<p>If we except the Placoids already alluded to, and a few other
+families of doubtful affinities, all the Old Red Sandstone fishes
+are Ganoids, an order so named by Agassiz from the shining outer
+surface of their scales; but Professor Huxley has also called our
+attention to the fact that, while a few of the primary and the
+great majority of the secondary Ganoids resemble the living bony
+pike, <i>Lepidosteus,</i> or the <i>Amia,</i> genera now found in
+North American rivers, and one of them, <i>Lepidosteus,</i>
+extending as far south as Guatemala, the Crossopterygii, or
+fringe-finned Ichthyolites, of the Old Red are closely related to
+the African <i>Polypterus,</i> which is represented by five or six
+species now inhabiting the Nile and the rivers of Senegal. These
+North American and African Ganoids are quite exceptional in the
+living creation; they are entirely confined to the northern
+hemisphere, unless some species of <i>Polypterus</i> range to the
+south of the line in Africa; and, out of about 9000 living species
+of fish known to M. G&uuml;nther, and of which more than 6000 are
+now preserved in the British Museum, they probably constitute no
+more than nine.</p>
+
+<img src="../images4/fig501.jpg" width="222" height="282" alt=
+"Fig. 501: Pterichthys. Upper side, showing mouth." align="right">
+
+<p>If many circumstances favour the theory of the fresh-water
+origin of the Old Red Sandstone, this view of its nature is not a
+little confirmed by our finding that it is in Llake Superior and
+the other inland Canadian seas of fresh water, and in the
+Mississippi and African rivers, that we at present find those fish
+which have the nearest affinity to the fossil forms of this ancient
+formation.</p>
+
+<p>Among the anomalous forms of Old Red fishes not referable to
+Huxley&rsquo;s Crossopterygii is the <i>Pterichthys,</i> of which
+five species have been found in the middle division of the Old Red
+of Scotland. Some writers have compared their shelly covering to
+that of Crustaceans, with which, however, they have no real
+affinity. The wing-like appendages, whence the genus is named, were
+first supposed by Hugh Miller to be paddles, like those of the
+turtle; and there can now be no doubt that they do really
+correspond with the pectoral fins.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 446">[ 446 ]</a></p>
+
+<p>The number of species of fish already obtained from the middle
+division of the Old Red Sandstone in Great Britain is about 70, and
+the principal genera, besides <i>Osteolepis</i> and <i>
+Pterichthys,</i> already mentioned, are <i>Glyptolepis,
+Diplacanthus, Dendrodus, Coccosteus, Cheirancanthus,</i> and <i>
+Acanthoides.</i></p>
+
+<center><img src="../images4/fig502.jpg" width="360" height="268" alt=
+"Fig. 502: Cephalapsis Lyellii."></center>
+
+<p><b>Lower Old Red Sandstone.</b>&mdash;The third or lowest
+division south of the Grampians consists of grey paving-stone and
+roofing-slate, with associated red and grey shales; these strata
+underlie a dense mass of conglomerate. In these grey beds several
+remarkable fish have been found of the genus named by Agassiz <i>
+Cephalaspis,</i> or &ldquo;buckler-headed,&rdquo; from the
+extraordinary shield which covers the head (see Fig. 502), and
+which has often been mistaken for that of a trilobite, such as <i>
+Asaphus.</i> A species of <i>Pteraspis,</i> of the same family, has
+also been found by the Reverend Hugh Mitchell in beds of
+corresponding age in Perthshire; and Mr. Powrie enumerates no less
+than five genera of the family Acanthodid&aelig;, the spines,
+scales, and other remains of which have been detected in the grey
+flaggy sandstones.*</p>
+
+<img src="../images4/fig503.jpg" width="190" height="163" alt=
+"Fig. 503: Pteygotus anglicus." align="left">
+
+<p>In the same formation at Carmylie, in Forfarshire, commonly
+known as the Arbroath paving-stone, fragments of a huge crustacean
+have been met with from time to time. They are called by the Scotch
+quarrymen the &ldquo;Seraphim,&rdquo; from the</p>
+
+<p class="fnote">* Powrie, Geol. Quart. Journ., vol. xx, p.
+417.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 447">[ 447 ]</a></p>
+
+<table border="0" cellspacing="0" cellpadding="0" bgcolor="white"
+summary=
+"Fig. 504: Pterygotus anglicus. Ventral aspect. Restored by H. Woodward, F.G.S.">
+<tr>
+<td align="left" valign="top"><img src="../images4/fig504.jpg" width=
+"214" height="361" alt=
+"Fig. 504: Pterygotus anglicus. Ventral aspect."></td>
+<td align="left" valign="bottom">
+<ol class="tab">
+<li>Carapace, showing the large sessile eyes at the anterior
+angles.</li>
+
+<li>The <i>metastoma</i> or post-oral plate (serving the office of
+a lower lip).</li>
+
+<li>Chelate appendages (antennules).</li>
+
+<li>First pair of simple palpi (antenn&aelig;).</li>
+
+<li>Second pair of simple palpi (mandibles).</li>
+
+<li>Third pair of simple palpi (first maxill&aelig;).</li>
+
+<li>Pair of swimming feet with their broad basal joints, whose
+serrated edges serve the office of maxill&aelig;.</li>
+
+<li>Thoracic plate covering the first two thoracic segments, which
+are indicated by the figures 1, 2, and a dotted line. 1-6. Thoracic
+segments. 7-12. Abdominal segments. 13. Telson, or
+tail-plate.)</li>
+</ol>
+</td>
+</tr>
+</table>
+
+<p>wing-like form and feather-like ornament of the thoracic
+appendage, the part most usually met with. Agassiz, having
+previously referred some of these fragments to the class of fishes,
+was the first to recognise their crustacean character, and,
+although at the time unable correctly to determine the true
+relation of the several parts, he figured the portions on which he
+founded his opinion, in the first plate of his &ldquo;Poissons
+Fossiles du Vieux Gr&egrave;s Rouge.&rdquo;</p>
+
+<p>A restoration in correct proportion to the size of the fragments
+of <i>P. anglicus</i> (Fig. 504), from the Lower Old Red Sandstone
+of Perthshire and Forfarshire, would give us a creature measuring
+from five to six feet in length, and more than one foot across.</p>
+
+<p>The largest crustaceans living at the present day are the <i>
+Inachus Kaempferi,</i> of De Haan, from Japan (a brachyurous or
+short-tailed crab), chiefly remarkable for the extraordinary length
+of its limbs; the fore-arm measuring four feet in length, and the
+others in proportion, so that it covers about 25 square feet of
+ground; and the <i>Limulus Moluccanus,</i> the great King Crab of
+China and the Eastern seas, which, when adult, measures 1&frac12;
+foot across its carapace, and is three feet in length.</p>
+
+<p>Besides some species of <i>Pterygotus,</i> several of the allied
+genus <i>Eurypterus</i> occur in the Lower Old Red Sandstone, and
+with them the remains of grass-like plants so abundant in
+Forfarshire and Kincardineshire as to be useful to the geologist by
+enabling him to identify the inferior strata at distant points.
+Some botanists have suggested that these</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 448">[ 448 ]</a></p>
+
+<center><img src="../images4/fig505.jpg" width="342" height="184" alt=
+"Fig. 505: Parka decipiens. In sandstone of lower beds of Old Red, Ley's Mill, Forfarshire. Fig. 506: Parka decipiens. In shale of Lower Old Red, Park Hill, Fife.">
+</center>
+
+<p>plants may be of the family <i>Fluviales,</i> and of fresh-water
+genera. They are accompanied by fossils, called
+&ldquo;berries&rdquo; by the quarrymen, which they compared to a
+compressed blackberry (see Figs. 505, 506), and which were called
+&ldquo;Parka&rdquo; by Dr. Fleming. They are now considered by Mr.
+Powrie to be the eggs of crustaceans, which is highly probable, for
+they have not only been found with <i>Pterygotus anglicus</i> in
+Forfarshire and Perthshire, but also in the Upper Silurian strata
+of England, in which species of the same genus, Pterygotus,
+occur.</p>
+
+<img src="../images4/fig507.jpg" width="231" height="251" alt=
+"Fig. 507: Shale of Old Red Sandstone. Forfarshire. With impression of plants and eggs of Crustaceans."
+ align="left">
+
+<p>The grandest exhibitions, says Sir R. Murchison, of the Old Red
+Sandstone in England and Wales appear in the escarpments of the
+Black Mountains and in the Fans of Brecon and Carmarthen, the one
+2862, and the other 2590 feet above the sea. The mass of red and
+brown sandstone in these mountains is estimated at not less than
+10,000 feet, clearly intercalated between the Carboniferous and
+Silurian strata. No shells or corals have ever been found in the
+whole series, not even where the beds are calcareous, forming
+irregular courses of concretionary lumps called
+&ldquo;corn-stones,&rdquo; which may be described as mottled red
+and green earthy limestones. The fishes of this lowest English Old
+Red are <i>Cephalaspis</i> and <i>Pteraspis,</i> specifically
+different from species of the same genera which occur in the
+uppermost Ludlow or Silurian tilestones. Crustaceans also of the
+genus <i>Eurypterus</i> are met with.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 449">[ 449 ]</a></p>
+
+<p><b>Marine or Devonian Type.</b>&mdash;We may now speak of the
+marine type of the British strata intermediate between the
+Carboniferous and Silurian, in treating of which we shall find it
+much more easy to identify the Upper, Middle, and Lower divisions
+with strata of the same age in other countries. It was not until
+the year 1836 that Sir R. Murchison and Professor Sedgwick
+discovered that the culmiferous or anthracitic shales and
+sandstones of North Devon, several thousand feet thick, belonged to
+the coal, and that the beds below them, which are of still greater
+thickness, and which, like the carboniferous strata, had been
+confounded under the general name &ldquo;graywacke,&rdquo; occupied
+a geological position corresponding to that of the Old Red
+Sandstone already described. In this reform they were aided by a
+suggestion of Mr. Lonsdale, who, after studying the Devonshire
+fossils, perceived that they belonged to a peculiar
+pal&aelig;ontological type of intermediate character between the
+Carboniferous and Silurian.</p>
+
+<p>It is in the north of Devon that these formations may best be
+studied, where they have been divided into an Upper, Middle, and
+Lower Group, and where, although much contorted and folded, they
+have for the most part escaped being altered by intrusive
+trap-rocks and by granite, which in Dartmoor and the more southern
+parts of the same county have often reduced them to a crystalline
+or metamorphic state.</p>
+
+<center><small>DEVONIAN &nbsp;SERIES&nbsp; IN &nbsp;NORTH
+&nbsp;DEVON.</small></center>
+
+<table border="1" cellspacing="0" cellpadding="10" summary=
+"Left column &mdash; Upper, Middle and Lower Devonian Groups; right column &mdash; Types of strata found in each group.">
+<tr>
+<td align="center" valign="middle">U<small>PPER</small>
+D<small>EVONIAN OR</small> P<small>ILTON</small>
+G<small>ROUP</small></td>
+<td align="left" valign="top">
+<b>(a)</b> &nbsp;Sandy slates and
+schists with fossils, 36 species out of 110 common to the
+Carboniferous group (Pilton, Barnstaple, etc.), resting on soft
+schists in which fossils are very abundant (Croyde, etc.), and
+which pass down into<br>
+<b>(b)</b> &nbsp;Yellow, brown, and red sandstone, with land
+plants (<i>Cyclopteris,</i> etc.) and marine shells. One zone,
+characterised by the abundance of cucull&aelig;a (Baggy Point,
+Marwood, Sloly, etc.) resting on hard grey and reddish sandstone
+and micaceous flags, no fossils yet found (Dulverton, Pickwell,
+Down, etc.)</td>
+</tr>
+
+<tr>
+<td align="center" valign="middle">M<small>IDDLE</small>
+D<small>EVONIAN OR</small> I<small>LFRACOMBE</small>
+G<small>ROUP.</small></td>
+<td align="left"><b>(a)</b> &nbsp;Green glossy slates of
+considerable thickness, no fossils yet recorded from these beds
+(Mortenoe, Lee Bay, etc.).<br>
+<b>(b)</b> &nbsp;Slates and schists, with several irregular
+courses of limestone containing shells and corals like those of the
+Plymouth Limestone (Combe Martin, Ilfracombe, etc.).</td>
+</tr>
+
+<tr>
+<td align="center" valign="middle">L<small>OWER</small>
+D<small>EVONIAN OR</small> L<small>YNTON</small>
+G<small>ROUP.</small></td>
+<td align="left"><b>(a)</b> &nbsp;Hard, greenish, red, and purple
+sandstone&mdash;no fossils yet found (Hangman Hill, etc.).<br>
+<b>(b)</b> &nbsp;Soft slates with subordinate
+sandstones&mdash;fossils numerous at various horizons&mdash;Orthis,
+Corals, Encrinites, etc. (Valley of Rocks, Lynmouth, etc.).</td>
+</tr>
+</table>
+
+<p>The above table exhibits the sequence of the strata or
+subdivisions as seen both on the sea-coast of the British Channel
+and in the interior of Devon. It will be seen that</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 450">[ 450 ]</a></p>
+
+<p>in all main points it agrees with the table drawn up in 1864 for
+the sixth edition of my &ldquo;Elements.&rdquo; Mr. Etheridge* has
+since published an excellent account of the different subdivisions
+of the rocks and their fossils, and has also pointed out their
+relation to the corresponding marine strata of the Continent. The
+slight modifications introduced in my table since 1864 are the
+result of a tour made in 1870 in company with Mr. T. Mck. Hughes,
+when we had the advantage of Mr. Etheridge&rsquo;s memoir as our
+guide.</p>
+
+<p>The place of the sandstones of the Foreland is not yet clearly
+made out, as they are cut off by a great fault and disturbance.</p>
+
+<img src="../images4/fig508.jpg" width="218" height="437" alt=
+"Fig. 508: Spirifera disjuncta. Fig. 509: Phacops latifrons."
+align="left">
+
+<p><b>Upper Devonian Rocks.</b>&mdash;The slates and sandstones of
+Barnstaple (<i>a</i> and <i>b</i> of the preceding section) contain
+the shell <i>Spirifera disjuncta,</i> Sowerby (S. Verneuilii,
+Murch.), (see Fig. 508), which has a very wide range in Europe,
+Asia Minor, and even China; also <i>Strophalosia caperata,</i>
+together with the large trilobite <i>Phacops latifrons,</i> Bronn.
+(See Fig. 509), which is all but world-wide in its distribution.
+The fossils are numerous, and comprise about 150 species of
+mollusca, a fifth of which pass up into the overlying Carboniferous
+rocks. To this Upper Devonian belong a series of limestones and
+slates well developed at Petherwyn, in Cornwall, where they have
+yielded 75 species of fossils. The genus of Cephalopoda called <i>
+Clymenia</i> (Fig. 510) is represented by no less than eleven
+species, and strata occupying the same position in Germany are
+called Clymenien-Kalk, or sometimes Cypridinen-Schiefer, on account
+of the number of minute bivalve shells of the crustacean called <i>
+Cypridina serrato-striata</i> (Fig. 511), which is found in these
+beds, in the Rhenish provinces, the Harz, Saxony, and Silesia, as
+well as in Cornwall and Belgium.</p>
+
+<p><b>Middle Devonian Rocks.</b>&mdash;We come next to the most
+typical portion of the Devonian system, including the great
+limestones of Plymouth and Torbay, replete with</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 451">[ 451 ]</a></p>
+
+<center><img src="../images4/fig510.jpg" width="390" height="194" alt=
+"Fig. 510: Clymenia linearis. Fig. 511: Cypridina serrato-striata.">
+</center>
+
+<img src="../images4/fig512.jpg" width="203" height="165" alt=
+"Fig. 512: Heliolites porosa." align="right">
+
+<p>shells, trilobites, and corals. Of the corals 51 species are
+enumerated by Mr. Etheridge, none of which pass into the
+Carboniferous formation. Among the genera we find <i>Favosites,
+Heliolites,</i> and <i>Cyathophyllum.</i> The two former genera are
+very frequent in Silurian rocks: some few even of the species are
+said to be common to the Devonian and Silurian groups, as, for
+example, <i>Favosites cervicornis</i> (Fig. 513), one of the
+commonest of all</p>
+
+<center><img src="../images4/fig513.jpg" width="382" height="316" alt=
+"Fig. 513: Favosites cervicornis. Fig. 514: Cyathophyllum c&aelig;spitosum.">
+</center>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 452">[ 452 ]</a></p>
+
+<p>the Devonshire fossils. The <i>Cyathophyllum
+c&aelig;spitosum</i> (Fig. 514) and <i>Heliolites pyriformis</i>
+(Fig. 512) are species peculiar to this formation.</p>
+
+<center><img src="../images4/fig515.jpg" width="359" height="207" alt=
+"Fig. 515: Stringocephalus Burtini. Fig. 516: Uncites Gryphus.">
+</center>
+
+<p>With the above are found no less than eleven genera of
+stone-lilies or crinoids, some of them, such as <i>
+Cupressocrinites,</i> distinct from any Carboniferous forms. The
+mollusks, also, are no less characteristic; of 68 species of
+Brachiopoda, ten only are common to the Carboniferous Limestone.
+The <i>Stringocephalus Burtini</i> (Fig. 515) and <i>Uncites
+Gryphus</i> (Fig. 516) may be mentioned as exclusively Middle
+Devonian genera, and extremely characteristic of the same division
+in Belgium. The <i>Stringocephalus</i> is also so abundant in the
+Middle Devonian of the banks of the Rhine as to have suggested the
+name of Stringocephalus Limestone.</p>
+
+<img src="../images4/fig517.jpg" width="262" height="213" alt=
+"Fig. 517: Megalodon cucullatus." align="left">
+
+<p>The only two species of Brachiopoda common to the Silurian and
+Devonian formations are <i>Atrypa reticularis</i> (Fig. 532), which
+seems to have been a cosmopolite species, and <i>Strophomena
+rhomboidalis.</i></p>
+
+<p>Among the peculiar lamellibranchiate bivalves common to the
+Plymouth limestone of Devonshire and the Continent, we find the <i>
+Megalodon</i> (Fig. 517). There are also twelve genera of
+Gasteropods which have yielded 36 species, four of which pass to
+the Carboniferous group, namely <i>Macrocheilus,</i></p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 453">[ 453 ]</a></p>
+
+<img src="../images4/fig518.jpg" width="132" height="280" alt=
+"Fig. 518: Conularia ornata." align="left"><img src=
+"../images4/fig519.jpg" width="132" height="217" alt=
+"Fig. 519: Bronteus flabellifer." align="right">
+
+<p><i>Acroculia, Euomphalus,</i> and <i>Murchisonia.</i> Pteropods
+occur, such as <i>Conularia</i> (Fig. 518), and Cephalopods, such
+as <i>Cyrtoceras, Gyroceras, Orthoceras,</i> and others, nearly all
+of genera distinct from those prevailing in the Upper Devonian
+Limestone, or Clymenien-kalk of the Germans already mentioned.
+Although but few species of Trilobites occur, the characteristic
+<i>Bronteus flabellifer</i> (Fig. 519) is far from rare, and all
+collectors are familiar with its fan-like tail. In this same group,
+called, as before stated, the Stringocephalus, or Eifel Limestone,
+in Germany, several fish remains have been detected, and among
+others the remarkable genus Coccosteus, covered with its
+tuberculated bony armour; and these ichthyolites serve, as Sir R.
+Murchison observes (Siluria, p. 362), to identify this middle
+marine Devonian with the Old Red Sandstone of Britain and
+Russia.</p>
+
+<img src="../images4/fig520.jpg" width="252" height="145" alt=
+"Fig. 520: Calceola sandalina." align="right">
+
+<p>Beneath the Eifel Limestone (the great central and typical
+member of &ldquo;the Devonian&rdquo; on the Continent) lie certain
+schists called by German writers &ldquo;Calceola-schiefer,&rdquo;
+because they contain in abundance a fossil body of very curious
+structure, <i>Calceola sandalina</i> (Fig. 520), which has been
+usually considered a brachiopod, but which some naturalists have
+lately referred to a Goniophyllum, supposing it to be an abnormal
+form of the order <i>Zoantharia rugosa</i> (see <a href=
+"../images3/fig474.jpg">Fig. 474</a>), differing from all other corals
+in being furnished with a strong operculum. This is by no means a
+rare fossil in the slaty limestone of South Devon, and, like the
+Eifel form, is confined to the middle group of this country.</p>
+
+<p><b>Lower Devonian Rocks.</b>&mdash;A great series of sandstones
+and glossy slates, with Crinoids, Brachiopods, and some corals,</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 454">[ 454 ]</a></p>
+
+<img src="../images4/fig521.jpg" width="266" height="108" alt=
+"Fig. 521: Spirifora mucronata." align="left">
+
+<p>occurring on the coast at Lynmouth and the neighbourhood, and
+called the Lynton Group (see Table <a href="#page 449">p. 449</a>,
+form the lowest member of the Devonian in North Devon. Among the 18
+species of all classes enumerated by Mr. Etheridge, two-thirds are
+common to the Middle Devonian, but only one, the ubiquitous <i>
+Atrypa reticularis,</i> can with certainty be identified with
+Silurian species. Among the characteristic forms are <i>Alveolites
+suborbicularis,</i> also common to this formation in the Rhine, and
+<i>Orthis arcuata,</i> very widely spread in the North Devon
+localities. But we may expect a large addition to the number of
+fossils whenever these strata shall have been carefully searched.
+The Spirifer Sandstone of Sandberger, as exhibited in the rocks
+bordering the Rhine between Coblentz and Caub, belong to this Lower
+division, and the same broad-winged Spirifers distinguish the
+Devonian strata of North America.</p>
+
+<img src="../images4/fig522.jpg" width="132" height="370" alt=
+"Fig. 522: Homalonotus armatus." align="right">
+
+<p>Among the Trilobites of this era several large species of <i>
+Homalonotus</i> (Fig. 522) are conspicuous. The genus is still
+better known as a Silurian form, but the spinose species appear to
+belong exclusively to the &ldquo;Lower Devonian,&rdquo; and are
+found in Britain, Europe, and the Cape of Good Hope.</p>
+
+<p><b>Devonian of Russia.</b>&mdash;The Devonian strata of Russia
+extend, according to Sir R. Murchison, over a region more spacious
+than the British Isles; and it is remarkable that, where they
+consist of sandstone like the &ldquo;Old Red&rdquo; of Scotland and
+Central England, they are tenanted by fossil fishes often of the
+same species and still oftener of the same genera as the British,
+whereas when they consist of limestone they contain shells similar
+to those of Devonshire, thus confirming, as Sir Roderick has
+pointed out, the contemporaneous origin which had been previously
+assigned to formations exhibiting two very distinct mineral types
+in different parts of Britain.*</p>
+
+<p class="fnote">* Murchison&rsquo;s Siluria, p. 329.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 455">[ 455 ]</a></p>
+
+<p>The calcareous and the arenaceous rocks of Russia above alluded
+to alternate in such a manner as to leave no doubt of their having
+been deposited in different parts of the same great period.</p>
+
+<img src="../images4/fig523.jpg" width="238" height="615" alt=
+"Fig. 523: Psilophyton princeps." align="right">
+
+<p><b>Devonian Strata in the United States and
+Canada.</b>&mdash;Between the Carboniferous and Silurian strata
+there intervenes, in the United States and Canada, a great series
+of formations referable to the Devonian group, comprising some
+strata of marine origin abounding in shells and corals, and others
+of shallow-water and littoral origin in which terrestrial plants
+abound. The fossils, both of the deep and shallow water strata, are
+very analogous to those of Europe, the species being in some cases
+the same. In Eastern Canada Sir W. Logan has pointed out that in
+the peninsula of Gaspe, south of the estuary of St. Lawrence, a
+mass of sandstone, conglomerate, and shale referable to this period
+occurs, rich in vegetable remains, together with some fish-spines.
+Far down in the sandstones of Gaspe, Dr. Dawson found, in 1869, an
+entire specimen of the genus <i>Cephalaspis,</i> a form so
+characteristic, as we have already seen, of the Scotch Lower Old
+Red Sandstone. Some of the sandstones are ripple-marked, and
+towards the upper part of the whole series a thin seam of coal has
+been observed, measuring, together with some associated</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 456">[ 456 ]</a></p>
+
+<p>carbonaceous shale, about three inches in thickness. It rests on
+an under-clay in which are the roots of Psilophyton (see Fig. 523).
+At many other levels rootlets of this same plant have been shown by
+Principal Dawson to penetrate the clays, and to play the same part
+as do the rootlets of Stigmaria in the coal formation.</p>
+
+<p>We had already learnt from the works of G&ouml;ppert, Unger, and
+Bronn that the European plants of the Devonian epoch resemble
+generically, with few exceptions, those already known as
+Carboniferous; and Dr. Dawson, in 1859, enumerated 32 genera and 69
+species which he had then obtained from the State of New York and
+Canada. A perusal of his catalogue,* comprising <i>Conifer&aelig;,
+Sigillari&aelig;, Calamites, Asterophyllites, Lepidodendra,</i> and
+ferns of the genera <i>Cyclopteris, Neuropteris, Sphenopteris,</i>
+and others, together with fruits, such as <i>Cardiocarpum</i> and
+<i>Trigonocarpum,</i> might dispose geologists to believe that they
+were presented with a list of Carboniferous fossils, the difference
+of the species from those of the coal-measures, and even a slight
+admixture of genera unknown in Europe, being naturally ascribed to
+geographical distribution and the distance of the New from the Old
+World. But fortunately the coal formation is fully developed on the
+other side of the Atlantic, and is singularly like that of Europe,
+both lithologically and in the species of its fossil plants. There
+is also the most unequivocal evidence of relative age afforded by
+superposition, for the Devonian strata in the United States are
+seen to crop out from beneath the Carboniferous on the borders of
+Pennsylvania and New York, where both formations are of great
+thickness.</p>
+
+<p>The number of American Devonian plants has now been raised by
+Dr. Dawson to 120, to which we may add about 80 from the European
+flora of the same age, so that already the vegetation of this
+period is beginning to be nearly half as rich as that of the
+coal-measures which have been studied for so much longer a time and
+over so much wider an area. The Psilophyton above alluded to is
+believed by Dr. Dawson to be a lycopodiaceous plant, branching
+dichotomously (see <i>P. princeps,</i> Fig. 523), with stems
+springing from a rhizome, which last has circular areoles, much
+resembling those of Stigmaria, and like it sending forth
+cylindrical rootlets. The extreme points of some of the branchlets
+are rolled up so as to resemble the croziers or circinate vernation
+of ferns; the leaves or bracts, <i>a,</i> supposed to belong to the
+same plant, are described by Dawson as having inclosed the
+fructification. The remains of <i>Psilophyton princeps</i> have
+been traced through</p>
+
+<p class="fnote">* Quart. Geol. Journ., vol. xv, p. 477, 1859; also
+vol. xviii, p. 296, 1862.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 457">[ 457 ]</a></p>
+
+<p>all the members of the Devonian series in America, and Dr.
+Dawson has lately recognised it in specimens of Old Red Sandstone
+from the north of Scotland.</p>
+
+<p>The monotonous character of the Carboniferous flora might be
+explained by imagining that we have only the vegetation handed down
+to us of one set of stations, consisting of wide swampy flats. But
+Dr. Dawson supposes that the geographical conditions under which
+the Devonian plants grew were more varied, and had more of an
+upland character. If so, the limitation of this more ancient flora,
+represented by so many genera and species, to the gymnospermous and
+cryptogamous orders, and the absence or extreme rarity of plants of
+higher grade, lead us naturally to speculate on the theory of
+progressive development, however difficult it may be to avail
+ourselves of this explanation, so long as we meet with even a few
+exceptional cases of what may seem to be monocotyledonous or
+dicotyledonous exogens.</p>
+
+<p><b>Devonian Insects of Canada.</b>&mdash;The earliest known
+insects were brought to light in 1865 in the Devonian strata of St.
+John&rsquo;s, New Brunswick, and are referred by Mr. Scudder to
+four species of <i>Neuroptera.</i> One of them is a gigantic
+Ephemera, and measured five inches in expanse of wing.</p>
+
+<p>Like many other ancient animals, says Dr. Dawson, they show a
+remarkable union of characters now found in distinct orders of
+insects, or constitute what have been named &ldquo;synthetic
+types.&rdquo; Of this kind is a stridulating or musical apparatus
+like that of the cricket in an insect otherwise allied to the <i>
+Neuroptera.</i> This structure, as Dr. Dawson observes, if rightly
+interpreted by Mr. Scudder, introduces us to the sounds of the
+Devonian woods, bringing before our imagination the trill and hum
+of insect life that enlivened the solitudes of these strange old
+forests.</p>
+
+<br>
+<hr>
+<small><a href="contents.html">Contents</a> / <a href="ch24.html">
+Chapter XXIV</a> / <a href="ch26.html">Chapter XXVI</a></small>
+</body>
+</html>
+