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+<p><b>The Student&rsquo;s Elements of Geology</b></p>
+
+<hr>
+<p class="page"><a name="page 366">[ 366 ]</a></p>
+
+<p>&nbsp;</p>
+
+<center><b>Chapter XXI</b><br>
+<br>
+TRIAS, OR NEW RED SANDSTONE GROUP.</center>
+
+<p class="intro">Beds of Passage between the Lias and Trias,
+Rh&aelig;tic Beds. &mdash; Triassic Mammifer. &mdash; Triple
+Division of the Trias. &mdash; Keuper, or Upper Trias of England.
+&mdash; Reptiles of the Upper Trias. &mdash; Foot-prints in the
+Bunter formation in England. &mdash; Dolomitic Conglomerate of
+Bristol. &mdash; Origin of Red Sandstone and Rock-salt. &mdash;
+Precipitation of Salt from inland Lakes and Lagoons. &mdash; Trias
+of Germany. &mdash; Keuper. &mdash; St. Cassian and Hallstadt Beds.
+&mdash; Peculiarity of their Fauna. &mdash; Muschelkalk and its
+Fossils. &mdash; Trias of the United States. &mdash; Fossil
+Foot-prints of Birds and Reptiles in the Valley of the Connecticut.
+&mdash; Triassic Mammifer of North Carolina. &mdash; Triassic
+Coal-field of Richmond, Virginia. &mdash; Low Grade of early
+Mammals favourable to the Theory of Progressive Development.</p>
+
+<p><b>Beds of Passage between the Lias and Trias&mdash;Rh&aelig;tic
+Beds.</b>&mdash;We have mentioned in the last chapter (<a href=
+"ch20.html#page 356">p. 356</a>) that the base of the Lower Lias is
+characterised, both in England and Germany, by beds containing
+distinct species of Ammonites, the lowest subdivision having been
+called the zone of <i>Ammonites planorbis.</i> Below this zone, on
+the boundary line between the Lias and the strata of which we are
+about to treat, called &ldquo;Trias,&rdquo; certain cream-coloured
+limestones devoid of fossils are usually found. These white beds
+were called by William Smith the White Lias, and they have been
+shown by Mr. Charles Moore to belong to a formation similar to one
+in the Rh&aelig;tian Alps of Bavaria, to which Mr. Gumbel has
+applied the name of Rh&aelig;tic. They have also long been known as
+the Koessen beds in Germany, and may be regarded as beds of passage
+between the Lias and Trias. They are named the Penarth beds by the
+Government surveyors of Great Britain, from Penarth, near Cardiff,
+in Glamorganshire, where they sometimes attain a thickness of fifty
+feet.</p>
+
+<p>The principal member of this group has been called by Dr. Wright
+the <i>Avicula contorta</i> bed,* as this shell is very abundant,
+and has a wide range in Europe. General Portlock first described
+the formation as it occurs at Portrush, in Antrim, where the <i>
+Avicula contorta</i> is accompanied by <i>Pecten Valoniensis,</i>
+as in Germany.</p>
+
+<p>The best known member of the group, a thin band or bone-breccia,
+is conspicuous among the black shales in the neigh-</p>
+
+<p class="fnote">* Dr. Wright, on Lias and Bone Bed, Quart. Geol.
+Journ., 1860, vol. xvi.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 367">[ 367 ]</a></p>
+
+<p>bourhood of Axmouth in Devonshire, and in the cliffs of
+Westbury-on-Severn, as well as at Aust and other places on the
+borders of the Bristol Channel. It abounds in the remains of
+saurians and fish, and was formerly classed as the lowest bed of
+the Lias; but Sir P. Egerton first pointed out, in 1841, that it
+should be referred to the Upper New Red Sandstone, because it
+contained an assemblage of fossil fish which are either peculiar to
+this stratum, or belong to species well-known in the Muschelkalk of
+Germany. These fish belong to the genera <i>Acrodus, Hybodus,
+Gyrolepis,</i> and <i>Saurichthys.</i></p>
+
+<center><img src="../images3/fig383.jpg" width="407" height="410" alt=
+"Fig. 383: Cardium rh&aelig;ticum. Fig. 384: Pecten Valoniensis. Fig. 385: Avicula contorta. Fig. 386: Hybodus plica ilis. Fig. 387: Saurichthys apicalis. Fig. 388: Gyrolepsis tenuistriatus.">
+</center>
+
+<p>Among those common to the English bone-bed and the Muschelkalk
+of Germany are <i>Hybodus plicatilis</i> (Fig. 386), <i>Saurychthys
+apicalis</i> (Fig. 387), <i>Gyrolepis tenuistriatus</i> (Fig. 388),
+and <i>G. Albertii.</i> Remains of saurians, <i>Plesiosaurus</i>
+among others, have also been found in the bone-bed, and plates of
+an <i>Encrinus.</i> It may be questioned whether some of those
+fossils which have the most Triassic character may</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 368">[ 368 ]</a></p>
+
+<p>not have been derived from the destruction of older strata,
+since in bone-beds, in general, many of the organic remains are
+undoubtedly derivative.</p>
+
+<img src="../images3/fig389.jpg" width="246" height="158" alt=
+"Fig. 389: Microlestes antiquus, molar tooth." align="left">
+
+<p><i>Triassic Mammifer.</i>&mdash;In North-western Germany, as in
+England, there occurs beneath the Lias a remarkable bone breccia.
+It is filled with shells and with the remains of fishes and
+reptiles, almost all the genera of which, and some even of the
+species, agree with those of the subjacent Trias. This breccia has
+accordingly been considered by Professor Quenstedt, and other
+German geologists of high authority, as the newest or uppermost
+part of the Trias. Professor Plieninger found in it, in 1847, the
+molar tooth of a small Triassic mammifer, called by him <i>
+Microlestes antiquus.</i> He inferred its true nature from its
+double fangs, and from the form and number of the protuberances or
+cusps on the flat crown; and considering it as predaceous, probably
+insectivorous, he called it <i>Microlestes</i> from micros, little,
+and lestes, a beast of prey. Soon afterwards he found a second
+tooth, also at the same locality, Diegerloch, about two miles to
+the south-east of Stuttgart.</p>
+
+<p>No anatomist had been able to give any feasible conjecture as to
+the affinities of this minute quadruped until Dr. Falconer, in
+1857, recognised an unmistakable resemblance between its teeth and
+the two back molars of his new genus <i>Plagiaulax</i> (<a href=
+"../images2/fig306.jpg">Fig. 306</a>), from the Purbeck strata. This
+would lead us to the conclusion that Microlestes was marsupial and
+plant-eating.</p>
+
+<p>In W&uuml;rtemberg there are two bone-beds, namely, that
+containing the Microlestes, which has just been described, which
+constitutes, as we have seen, the uppermost member of the Trias,
+and another of still greater extent, and still more rich in the
+remains of fish and reptiles, which is of older date, intervening
+between the Keuper and Muschelkalk.</p>
+
+<p>The genera <i>Saurichthys, Hybodus,</i> and <i>Gyrolepis</i> are
+found in both these breccias, and one of the species, <i>
+Saurichthys Mongeoti,</i> is common to both bone-beds, as is also a
+remarkable reptile called <i>Nothosaurus mirabilis.</i> The saurian
+called <i>Belodon</i> by H. von Meyer, of the Thecodont family, is
+another Triassic form, associated at Diegerloch with
+Microlestes.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 369">[ 369 ]</a></p>
+
+<center><small>TRIAS OF ENGLAND.</small></center>
+
+<p>Between the Lias and the Coal (or Carboniferous group) there is
+interposed, in the midland and western counties of England, a great
+series of red loams, shales, and sandstones, to which the name of
+the &ldquo;New Red Sandstone formation&rdquo; was first given, to
+distinguish it from other shales and sandstones called the
+&ldquo;Old Red,&rdquo; often identical in mineral character, which
+lie immediately beneath the coal. The name of &ldquo;Red
+Marl&rdquo; has been incorrectly applied to the red clays of this
+formation, as before explained (<a href="ch2.html#page 38">p.
+38</a>), for they are remarkably free from calcareous matter. The
+absence, indeed, of carbonate of lime, as well as the scarcity of
+organic remains, together with the bright red colour of most of the
+rocks of this group, causes a strong contrast between it and the
+Jurassic formations before described.</p>
+
+<p>The group in question is more fully developed in Germany than in
+England or France. It has been called the Trias by German writers,
+or the Triple Group, because it is separable into three distinct
+formations, called the &ldquo;Keuper,&rdquo; the
+&ldquo;Muschelkalk,&rdquo; and the &ldquo;Bunter-sandstein.&rdquo;
+Of these the middle division, or the Muschelkalk, is wholly wanting
+in England, and the uppermost (Keuper) and lowest (Bunter) members
+of the series are not rich in fossils.</p>
+
+<p><b>Upper Trias or Keuper.</b>&mdash;In certain grey indurated
+marls below the bone-bed Mr. Boyd Dawkins has found at Watchet, on
+the coast of Somersetshire, a molar tooth of Microlestes, enabling
+him to refer to the Trias strata formerly supposed to be Liassic.
+Mr. Charles Moore had previously discovered many teeth of mammalia
+of the same family near Frome, in Somersetshire, in the contents of
+a vertical fissure traversing a mass of carboniferous limestone.
+The top of this fissure must have communicated with the bed of the
+Triassic sea, and probably at a point not far from the ancient
+shore on which the small marsupials of that era abounded.</p>
+
+<p>This upper division of the Trias called the Keuper is of great
+thickness in the central counties of England, attaining, according
+to Mr. Hull&rsquo;s estimate, no less than 3450 feet in Cheshire,
+and it covers a large extent of country between Lancashire and
+Devonshire.</p>
+
+<p>In Worcestershire and Warwickshire in sandstone belonging to the
+uppermost part of the Keuper the bivalve crustacean <i>Estheria
+minuta</i> occurs. The member of the English &ldquo;New Red&rdquo;
+containing this shell, in those parts of England, is, according to
+Sir Roderick Murchison and Mr. Strickland, 600 feet thick, and
+consists chiefly of red marl or slate, with</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 370">[ 370 ]</a></p>
+
+<img src="../images3/fig390.jpg" width="95" height="105" alt=
+"Fig. 390: Estheria minuta." align="right">
+
+<p>a band of sandstone. Ichthyodorulites, or spines of <i>
+Hybodus,</i> teeth of fishes, and footprints of reptiles were
+observed by the same geologists in these strata.</p>
+
+<img src="../images3/fig391.jpg" width="276" height="260" alt=
+"Fig. 391: Hyperodapedon Gordoni. Left Plate, Maxillary." align=
+"left">
+
+<p>In the Upper Trias or Keuper the remains of two saurians of the
+order Lacertilia have been found. The one called <i>
+Rhynchosaurus</i> occurred at Grinsell near Shrewsbury, and is
+characterised by having a small bird-like skull and jaws without
+teeth. The other <i>Hyperodapedon</i> (Fig. 391) was first noticed
+in 1858, near Elgin, in strata now recognised as Upper Triassic,
+and afterwards in beds of about the same age in the neighbourhood
+of Warwick. Remains of the same genus have been found both in
+Central India and Southern Africa in rocks believed to be of
+Triassic age. The Hyperodapedon has been shown by Professor Huxley
+to be a terrestrial reptile having numerous palatal teeth, and
+closely allied to the living Sphenodon of New Zealand.</p>
+
+<p>The recent discoveries of a living saurian in New Zealand so
+closely allied to this supposed extinct division of the Lacertilia
+seems to afford an illustration of a principle pointed out by Mr.
+Darwin of the survival in insulated tracts, after many changes in
+physical geography, of orders of which the congeners have become
+extinct on continents where they have been exposed to the severer
+competition of a larger progressive fauna.</p>
+
+<img src="../images3/fig392.jpg" width="85" height="138" alt=
+"Fig. 392: Tooth of Labyrinthodon." align="right">
+
+<p>Teeth of Labyrinthodon (Fig. 392) found in the Keuper in
+Warwickshire were examined microscopically by Professor Owen, and
+compared with other teeth from the German Keuper. He found after
+careful investigation that neither of them could be referred to
+true saurians, although they had been named <i>Mastodonsaurus</i>
+and <i>Phytosaurus</i> by J&auml;ger. It appeared that they were of
+the <i>Batrachian</i> order, and of gigantic</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 371">[ 371 ]</a></p>
+
+<p>dimensions in comparison with any representatives of that order
+now living. Both the Continental and English fossil teeth exhibited
+a most complicated texture, differing from that previously observed
+in any reptile, whether recent or extinct, but most nearly
+analogous to the <i>Ichthyosaurus.</i> A section of one of these
+teeth exhibits a series of irregular folds, resembling the
+labyrinthic windings of the surface of the brain; and from this
+character Professor Owen has proposed the name Labyrinthodon for
+the new genus. Fig. 393 of part of one is given from his
+&ldquo;Odontography,&rdquo; plate 64, A. The entire length of this
+tooth is supposed to have been about three inches and a half, and
+the breadth at the base one inch and a half.</p>
+
+<center><img src="../images3/fig393.jpg" width="354" height="336" alt=
+"Fig. 393: Transverse section of upper part of tooth of Labyrinthodon Jaegeri.">
+</center>
+
+<p><i>Rock-salt.</i>&mdash;In Cheshire and Lancashire there are red
+clays containing gypsum and salt of the age of the Trias which are
+between 1000 and 1500 feet thick. In some places lenticular masses
+of pure rock-salt nearly 100 feet thick are interpolated between
+the argillaceous beds. At the base of the formation beneath the
+rock-salt occur the Lower Sandstones and Marl, called provincially
+in Cheshire &ldquo;water-stones,&rdquo; which are largely quarried
+for building. They are often ripple-marked, and are impressed with
+numerous footprints of reptiles.</p>
+
+<p>The basement beds of the Keuper rest with a slight</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 372">[ 372 ]</a></p>
+
+<p>unconformability upon an eroded surface of the
+&ldquo;Bunter&rdquo; next to be described.</p>
+
+<img src="../images3/fig394.jpg" width="170" height="192" alt=
+"Fig. 394: Single footstep of Cheirotherium." align="left">
+
+<p><b>Lower Trias or Bunter.</b>&mdash;The lower division or
+English representative of the &ldquo;Bunter&rdquo; attains a
+thickness of 1500 feet in the counties last mentioned, according to
+Professor Ramsay. Besides red and green shales and red sandstones,
+it comprises much soft white quartzose sandstone, in which the
+trunks of silicified trees have been met with at Allesley Hill,
+near Coventry. Several of them were a foot and a half in diameter,
+and some yards in length, decidedly of coniferous wood, and showing
+rings of annual growth.* Impressions, also, of the footsteps of
+animals have been detected in Lancashire and Cheshire in this
+formation. Some of the most remarkable occur a few miles from
+Liverpool, in the whitish quartzose sandstone of Storton Hill, on
+the west side of the Mersey. They bear a close resemblance to
+tracks first observed in this member of the Upper New Red
+Sandstone, at the village of Hesseberg, near Hildburghausen, in
+Saxony. For many years these footprints have been referred to a
+large unknown quadruped, provisionally named <i>Cheirotherium</i>
+by Professor Kaup, because the marks both of the fore and hind feet
+resembled impressions made by a human hand. (See Fig. 394.) The
+foot-marks at Hesseberg are partly concave, and partly in relief,
+the former, or the depressions, are seen upon the upper surface of
+the sandstone slabs, but those in relief are only upon the lower
+surfaces, being, in fact, natural casts, formed in the subjacent
+footprints as in moulds. The larger impressions, which seem to be
+those of the hind foot, are generally eight inches in length, and
+five in width, and one was twelve inches long. Near each large
+footstep, and at a regular distance (about an inch and a half)
+before it, a smaller print of a fore foot, four inches long and
+three inches wide, occurs. The footsteps follow each other</p>
+
+<center><img src="../images3/fig395.jpg" width="367" height="82" alt=
+"Fig. 395: Line of footsteps on slab of sandstone."></center>
+
+<p class="fnote">* Buckland, Proc. Geol. Soc., vol. ii, p. 439; and
+Murchison and Strickland, Geol. Trans., Second Series., vol. v, p.
+347.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 373">[ 373 ]</a></p>
+
+<p>in pairs, each pair in the same line, at intervals of fourteen
+inches from pair to pair. The large as well as the small steps show
+the great toes alternately on the right and left side; each step
+makes the print of five toes, the first, or great toe, being bent
+inward like a thumb. Though the fore and hind foot differ so much
+in size, they are nearly similar in form.</p>
+
+<p>As neither in Germany nor in England had any bones or teeth been
+met with in the same identical strata as the footsteps, anatomists
+indulged, for several years, in various conjectures respecting the
+mysterious animals from which they might have been derived.
+Professor Kaup suggested that the unknown quadruped might have been
+allied to the <i>Marsupialia</i>; for in the kangaroo the first toe
+of the fore foot is in a similar manner set obliquely to the
+others, like a thumb, and the disproportion between the fore and
+hind feet is also very great. But M. Link conceived that some of
+the four species of animals of which the tracks had been found in
+Saxony might have been gigantic <i>Batrachians,</i> and when it was
+afterwards inferred that the Labyrinthodon was an air-breathing
+reptile, it was conjectured by Professor Owen that it might be one
+and the same as the Cheirotherium.</p>
+
+<p><b>Dolomitic Conglomerate of Bristol.</b>&mdash;Near Bristol, in
+Somersetshire, and in other counties bordering the Severn, the
+lowest strata belonging to the Triassic series consist of a
+conglomerate or breccia resting unconformably upon the Old Red
+Sandstone, and on different members of the Carboniferous rocks,
+such as the Coal Measures, Millstone Grit, and Mountain Limestone.
+This mode of superposition will be understood by reference to the
+section below Dundry Hill (<a href="../images/fig85.jpg">Fig. 85</a>),
+where No. 4 is the dolomitic conglomerate. Such breccias may have
+been partly the result of the sub&AElig;rial waste of an old
+land-surface which gradually sank down and suffered littoral
+denudation in proportion as it became submerged. The pebbles and
+fragments of older rocks which constitute the conglomerate are
+cemented together by a red or yellow base of dolomite, and in some
+places the encrinites and other fossils derived from the Mountain
+Limestone are so detached from the parent rocks that they have the
+deceptive appearance of belonging to a fauna contemporaneous with
+the dolomitic beds in which they occur. The imbedded fragments are
+both rounded and angular, some consisting of sandstone from the
+coal-measures, being of vast size, and weighing nearly a ton.
+Fractured bones and teeth of saurians which are truly of
+contemporaneous origin are dispersed through some parts of the
+breccia, and two of these reptiles called Thecodont saurians, named
+from the</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 374">[ 374 ]</a></p>
+
+<p>manner in which the teeth were implanted in the jawbone,
+obtained great celebrity because the patches of red conglomerate in
+which they were found, near Bristol, were originally supposed to be
+of Permian or Pal&aelig;ozoic age, and therefore the only
+representatives in England of vertebrate animals of so high a grade
+in rocks of such antiquity. The teeth of these saurians are
+conical, compressed, and with finely serrated edges (see Fig. 396);
+they are referred by Professor Huxley to the Dinosaurian order.</p>
+
+<img src="../images3/fig396.jpg" width="133" height="204" alt=
+"Fig. 396: Tooth of Thecodontosaurus." align="left">
+
+<p><b>Origin of Red Sandstone and Rock-salt.</b>&mdash;In various
+parts of the world, red and mottled clays and sandstones, of
+several distinct geological epochs, are found associated with salt,
+gypsum, and magnesian limestone, or with one or all of these
+substances. There is, therefore, in all likelihood, a general cause
+for such a coincidence. Nevertheless, we must not forget that there
+are dense masses of red and variegated sandstones and clays,
+thousands of feet in thickness, and of vast horizontal extent,
+wholly devoid of saliferous or gypseous matter. There are also
+deposits of gypsum and of common salt, as in the blue-clay
+formation of Sicily, without any accompanying red sandstone or red
+clay.</p>
+
+<p>These red deposits may be accounted for by the decomposition of
+gneiss and mica schist, which in the eastern Grampians of Scotland
+has produced a mass of detritus of precisely the same colour as the
+Old Red Sandstone.</p>
+
+<p>It is a general fact, and one not yet accounted for, that
+scarcely any fossil remains are ever preserved in stratified rocks
+in which this oxide of iron abounds; and when we find fossils in
+the New or Old Red Sandstone in England, it is in the grey, and
+usually calcareous beds, that they occur. The saline or gypseous
+interstratified beds may have been produced by submarine gaseous
+emanations, or hot mineral springs, which often continue to flow in
+the same spots for ages. Beds of rock-salt are, however, more
+generally attributed to the evaporation of lakes or lagoons
+communicating at intervals with the ocean. In Cheshire two beds of
+salt occur of the extraordinary thickness of 90 or even 100 feet,
+and extending over an area supposed to be 150 miles in diameter.
+The adjacent beds present ripple-marked sandstones and footprints
+of animals at so many levels as to imply that the whole area
+underwent a slow and gradual depression during the formation of the
+red sandstone.</p>
+
+<p>Major Harris, in his &ldquo;Highlands of Ethiopia,&rdquo;
+describes a</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 375">[ 375 ]</a></p>
+
+<p>salt lake, called the Bahr Assal, near the Abyssinian frontier,
+which once formed the prolongation of the Gulf of Tadjara, but was
+afterwards cut off from the gulf by a broad bar of lava or of land
+upraised by an earthquake. &ldquo;Fed by no rivers, and exposed in
+a burning climate to the unmitigated rays of the sun, it has shrunk
+into an elliptical basin, seven miles in its transverse axis, half
+filled with smooth water of the deepest c&aelig;rulean hue, and
+half with a solid sheet of glittering snow-white salt, the
+offspring of evaporation.&rdquo; &ldquo;If,&rdquo; says Mr. Hugh
+Miller, &ldquo;we suppose, instead of a barrier of lava, that
+sand-bars were raised by the surf on a flat arenaceous coast during
+a slow and equable sinking of the surface, the waters of the outer
+gulf might occasionally topple over the bar, and supply fresh brine
+when the first stock had been exhausted by evaporation.&rdquo;</p>
+
+<p>The Runn of Cutch, as I have shown elsewhere,* is a low region
+near the delta of the Indus, equal in extent to about a quarter of
+Ireland, which is neither land nor sea, being dry during part of
+every year, and covered by salt water during the monsoons. Here and
+there its surface is incrusted over with a layer of salt caused by
+the evaporation of sea-water. A subsiding movement has been
+witnessed in this country during earthquakes, so that a great
+thickness of pure salt might result from a continuation of such
+sinking.</p>
+
+<br>
+
+
+<center><small>TRIAS OF GERMANY.</small></center>
+
+<p>In Germany, as before hinted, chapter 21, the Trias first
+received its name as a Triple Group, consisting of two sandstones
+with an intermediate marine calcareous formation, which last is
+wanting in England.</p>
+
+<center><small>NOMENCLATURE OF TRIAS.</small></center>
+
+<center>
+<table border="1" cellspacing="0" cellpadding="4" width="80%"
+summary="German, French and English nomenclature.">
+<tr>
+<td align="center">German</td>
+<td align="center">French</td>
+<td align="center">English</td>
+</tr>
+
+<tr>
+<td align="left" valign="top">Keuper</td>
+<td align="left" valign="top">Marnes iris&eacute;es</td>
+<td align="left">Saliferous and gypseous<br>
+shales and sandstone.</td>
+</tr>
+
+<tr>
+<td align="left" valign="top">Muschelkalk</td>
+<td align="left">Muschelkalk, on calcaire coquillier</td>
+<td align="left" valign="top">Wanting in England.</td>
+</tr>
+
+<tr>
+<td align="left" valign="top">Bunter-sandstein</td>
+<td align="left" valign="top">Gr&egrave;s bigarr&eacute;</td>
+<td align="left">Sandtone and quartzose conglomerate.</td>
+</tr>
+</table>
+</center>
+
+<p><b>Keuper.</b>&mdash;The first of these, or the Keuper,
+underlying the beds before described as Rh&aelig;tic, attains in
+W&uuml;rtemberg a thickness of about 1000 feet. It is divided by
+Alberti into sandstone, gypsum, and carbonaceous
+clay-slate.&dagger; Remains of reptiles called <i>Nothosaurus</i>
+and <i>Phytosaurus,</i> have been found in it with Labyrinthodon;
+the detached teeth, also, of</p>
+
+<p class="fnote">* Principles of Geology, chap. xxvii.<br>
+&dagger; Monog. des Bunter-Sandsteins.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 376">[ 376 ]</a></p>
+
+<img src="../images3/fig397.jpg" width="179" height="207" alt=
+"Fig. 397: Equisetites columnaris." align="left">
+
+<p>placoid fish and of Rays, and of the genera <i>Saurichthys</i>
+and <i>Gyrolepis</i> (<a href="../images3/fig383.jpg">Figs. 387,
+388</a>). The plants of the Keuper are generically very analogous
+to those of the oolite and lias, consisting of ferns, equisetaceous
+plants, cycads, and conifers, with a few doubtful monocotyledons. A
+few species such as <i>Equisetites columnaris,</i> are common to
+this group and the oolite.</p>
+
+<p><i>St. Cassian and Hallstadt Beds</i> (see Map, Fig.
+398).&mdash; The sandstones and clay of the Keuper resemble the
+deposits of estuaries and a shallow sea near the land, and afford,
+in the N.W. of Germany, as in France and England, but a scanty
+representation of the marine life of that period. We might,
+however, have anticipated, from its rich reptilian fauna, that the
+contemporaneous inhabitants of the sea of the Keuper period would
+be very numerous, should we ever have an opportunity of bringing
+their remains to light. This, it is believed, has at length been
+accomplished, by the position now assigned to certain Alpine rocks
+called the &ldquo;St. Cassian beds,&rdquo; the true place of which
+in the series was until lately a subject of much doubt and
+discussion. It has been proved that the Hallstadt beds on the
+northern flanks of the Austrian Alps correspond in age with the St.
+Cassian beds on their southern declivity, and the Austrian
+geologists, M. Suess of Vienna and others, have satisfied
+themselves that the Hallstadt formation is referable to the period
+of the Upper Trias.</p>
+
+<center><img src="../images3/fig398.jpg" width="389" height="237" alt=
+"Fig. 398: Map of Tyrol and Styria showing St. Cassian and Hallstadt Beds.">
+</center>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 377">[ 377 ]</a></p>
+
+<img src="../images3/fig399.jpg" width="137" height="274" alt=
+"Fig. 399: Scoliotoma. Fig. 400: Koninckia Leonhardi." align=
+"left">
+
+<p>Assuming this conclusion to be correct, we become acquainted
+suddenly and unexpectedly with a rich marine fauna belonging to a
+period previously believed to be very barren of organic remains,
+because in England, France, and Northern Germany the upper Trias is
+chiefly represented by beds of fresh or brackish water origin.</p>
+
+<p>About 600 species of invertebrate fossils occur in the Hallstadt
+and St. Cassian beds, many of which are still undescribed; some of
+the Mollusca are of new and peculiar genera, as <i>Scoliostoma,</i>
+Fig. 399, and <i>Platystoma,</i> Fig. 400, among the Gasteropoda;
+and <i>Koninckia,</i> Fig. 401, among the Brachiopoda.</p>
+
+<img src="../images3/fig401.jpg" width="246" height="238" alt=
+"Fig. 401: Koninckia Leonhardi." align="right">
+
+<p>The following table of genera of marine shells from the
+Hallstadt and St. Cassian beds, drawn up first on the joint
+authority of M. Suess and the late Dr. Woodward, and since
+corrected by Messrs. Etheridge and Tate, shows how many connecting
+links between the fauna of primary and secondary Pal&aelig;ozoic
+and Mesozoic rocks are supplied by the St. Cassian and Hallstadt
+beds.</p>
+
+<br>
+
+
+<center><small>GENERA OF FOSSIL MOLLUSCA IN THE ST. CASSIAN AND
+HALLSTADT BEDS.</small></center>
+
+<center>
+<table border="0" cellpadding="4" cellspacing="0" width="100%"
+summary=
+"Column 1: Common to Older Rocks. Column 2: Characteristic Triassic Genera. Column 3: Common to Newer Rocks.">
+<tr>
+<td colspan="2" align="center"><small>Common to Older
+Rocks</small></td>
+<td colspan="2" align="center"><small>Characteristic Triassic
+Genera</small></td>
+<td colspan="2" align="center"><small>Common to Newer
+Rocks</small></td>
+</tr>
+
+<tr>
+<td width="20"></td>
+<td align="left" valign="top">Orthoceras<br>
+Bactrites<br>
+Macrocheilus<br>
+Loxonema<br>
+Holopella<br>
+Murchisonia<br>
+Porcellia<br>
+Athyris<br>
+Retzia<br>
+Cyrtina<br>
+Euomphalus</td>
+<td width="24"></td>
+<td align="left" valign="top">Ceratites<br>
+Cochloceras<br>
+Choristoceras<br>
+Rhabdoceras<br>
+Aulacoceras<br>
+Scoliostoma *<br>
+Naticella<br>
+Platystoma<br>
+Ptychostoma<br>
+Euchrysalis<br>
+Halobia<br>
+Hornesia<br>
+Amphiclina<br>
+Koninckia<br>
+Cassianella &dagger;<br>
+Myophoria &dagger;</td>
+<td width="20"></td>
+<td align="left" valign="top">Ammonites<br>
+Chemnitzia<br>
+Cerithium<br>
+Monodonta<br>
+Opis<br>
+Sphoera<br>
+Cardita<br>
+Myoconcha<br>
+Hinnites<br>
+Monotis<br>
+Plicatula<br>
+Pachyrisma<br>
+Thecidium</td>
+</tr>
+</table>
+</center>
+
+<p class="fnote">* Reaches its maximum in the Trias, but passes
+down to older rocks.<br>
+&dagger; Reach their maximum in the Trias, but pass up to newer
+rocks.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 378">[ 378 ]</a></p>
+
+<p>The first column marks the last appearance of several genera
+which are characteristic of Pal&aelig;ozoic strata. The second
+shows those genera which are characteristic of the Upper Trias,
+either as peculiar to it, or, as in the three cases marked by
+asterisks, reaching their maximum of development at this era. The
+third column marks the first appearance in Triassic rocks of genera
+destined to become more abundant in later ages.</p>
+
+<p>It is only, however, when we contemplate the number of species
+by which each of the above-mentioned genera are represented that we
+comprehend the peculiarities of what is commonly called the St.
+Cassian fauna. Thus, for example, the Ammonite, which is not common
+to older rocks, is represented by no less than seventy-three
+species; whereas Loxonema, which is only known as common to older
+rocks, furnishes fifteen Triassic species. Cerithium, so abundant
+in tertiary strata, and which still lives, is represented by no
+less than fourteen species. As the Orthoceras had never been met
+with in the marine Muschelkalk, much surprise was naturally felt
+that seven or eight species of the genus should appear in the
+Hallstadt beds, assuming these last to belong to the Upper Trias.
+Among these species are some of large dimensions, associated with
+large Ammonites with foliated lobes, a form never seen before so
+low in the series, while the Orthoceras had never been seen so
+high.</p>
+
+<p>On the whole, the rich marine fauna of Hallstadt and St.
+Cassian, now generally assigned to the lowest members of the Upper
+Trias or Keuper, leads us to suspect that when the strata of the
+Triassic age are better known, especially those belonging to the
+period of the Bunter sandstone, the break between the
+Pal&aelig;ozoic and Mesozoic Periods may be almost effaced. Indeed
+some geologists are not yet satisfied that the true position of the
+St. Cassian beds (containing so great an admixture of types, having
+at once both Mesozoic and Pal&aelig;ozoic affinities) is made out,
+and doubt whether they have yet been clearly proved to be newer
+than the Muschelkalk.</p>
+
+<p><b>Muschelkalk.</b>&mdash;The next member of the Trias in
+Germany, the <i>Muschelkalk,</i> which underlies the <i>Keuper</i>
+before described, consists chiefly of a compact greyish limestone,
+but includes beds of dolomite in many places, together with gypsum
+and rock-salt. This limestone, a formation wholly unrepresented in
+England, abounds in fossil shells, as the name implies. Among the
+Cephalopoda there are no belemnites, and no ammonites with foliated
+sutures, as in the Lias, and Oolite, and the Hallstadt beds; but we
+find instead a genus allied to</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 379">[ 379 ]</a></p>
+
+<center><img src="../images3/fig402.jpg" width="399" height="562" alt=
+"Fig. 402: Ceratites nodosus. Fig. 403: Gervillia (Avicula) socialis. Fig. 404: Enerinus liliiformis. Fig. 405: Aspidura loricata.">
+</center>
+
+<p>the Ammonite, called <i>Ceratites</i> by de Haan, in which the
+descending lobes (Fig. 402) terminate in a few small denticulations
+pointing inward. Among the bivalve crustacea, the <i>Estheria
+minuta,</i> Bronn (see <a href="../images3/fig390.jpg">Fig. 390</a>),
+is abundant, ranging through the Keuper, Muschelkalk, and
+Bunter-sandstein; and <i>Gervillia socialis</i> (Fig. 403), having
+a similar range, is found in great numbers in the Muschelkalk of
+Germany, France, and Poland.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 380">[ 380 ]</a></p>
+
+<img src="../images3/fig406.jpg" width="161" height="226" alt=
+"Fig. 406: Palatal teeth of Placodus gigas." align="left">
+
+<p>The abundance of the heads and stems of lily encrinites, <i>
+Encrinus liliiformis</i> (Fig. 404), (or <i>Encrinites
+moniliformis</i>), shows the slow manner in which some beds of this
+limestone have been formed in clear sea-water. The star-fish called
+<i>Aspidura loricata</i> (Fig. 405) is as yet peculiar to the
+Muschelkalk. In the same formation are found the skull and teeth of
+a reptile of the genus <i>Placodus</i> (see Fig. 406), which was
+referred originally by Munster, and afterwards by Agassiz, to the
+class of fishes. But more perfect specimens enabled Professor Owen,
+in 1858, to show that this fossil animal was a Saurian reptile,
+which probably fed on shell-bearing mollusks, and used its short
+and flat teeth, so thickly coated with enamel, for pounding and
+crushing the shells.</p>
+
+<img src="../images3/fig407.jpg" width="180" height="211" alt=
+"Fig. 407: Voltzia heterophylla." align="right">
+
+<p><b>Bunter-sandstein.</b>&mdash;The <i>Bunter-sandstein</i>
+consists of various-coloured sandstones, dolomites, and red clays,
+with some beds, especially in the Hartz, of calcareous pisolite or
+roe-stone, the whole sometimes attaining a thickness of more than
+1000 feet. The sandstone of the Vosges is proved, by its fossils,
+to belong to this lowest member of the Triassic group. At Sulzbad
+(or Soultz-les-bains), near Strasburg, on the flanks of the Vosges,
+many plants have been obtained from the &ldquo;bunter,&rdquo;
+especially conifers of the extinct genus <i>Voltzia,</i> of which
+the fructification has been preserved. (See Fig. 407.) Out of
+thirty species of ferns, cycads, conifers, and other plants,
+enumerated by M. Ad. Brongniart, in 1849, as coming from the
+&ldquo;Gr&egrave;s bigarr&eacute;,&rdquo; or Bunter, not one is
+common to the Keuper.</p>
+
+<p>The footprints of Labyrinthodon observed in the clays of this
+formation at Hildburghausen, in Saxony, have already been
+mentioned. Some idea of the variety and importance of the
+terrestrial vertebrate fauna of the three members of the Trias in
+Northern Germany may be derived from the fact that in the great
+monograph by the late Hermann von Meyer on the reptiles</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 381">[ 381 ]</a></p>
+
+<p>of the Trias, the remains of no less than eighty distinct
+species are described and figured.</p>
+
+<center><small>TRIAS OF THE UNITED STATES.</small></center>
+
+<p><b>New Red Sandstone of the Valley of the Connecticut
+River.</b>&mdash;In a depression of the granitic or hypogene rocks
+in the States of Massachusetts and Connecticut strata of red
+sandstone, shale, and conglomerate are found, occupying an area
+more than 150 miles in length from north to south, and about five
+to ten miles in breadth, the beds dipping to the eastward at angles
+varying from 5 to 50 degrees. The extreme inclination of 50 degrees
+is rare, and only observed in the neighbourhood of masses of trap
+which have been intruded into the red sandstone while it was
+forming, or before the newer parts of the deposit had been
+completed. Having examined this series of rocks in many places, I
+feel satisfied that they were formed in shallow water, and for the
+most part near the shore, and that some of the beds were from time
+to time raised above the level of the water, and laid dry, while a
+newer series, composed of similar sediment, was forming.</p>
+
+<img src="../images3/fig408.jpg" width="108" height="437" alt=
+"Fig. 408: Foot-prints of a bird, Turner's Falls, Valley of the Connecticut."
+ align="right">
+
+<p>According to Professor Hitchcock, the footprints of no less than
+thirty-two species of bipeds, and twelve of quadrupeds, have been
+already detected in these rocks. Thirty of these are believed to be
+those of birds, four of lizards, two of chelonians, and six of
+batrachians. The tracks have been found in more than twenty places,
+scattered through an extent of nearly 80 miles from north to south,
+and they are repeated through a succession of beds attaining at
+some points a thickness of more than 1000 feet.*</p>
+
+<p>The bipedal impressions are, for the most part, trifid, and show
+the same number of joints as exist in the feet of living
+tridactylous birds. Now, such birds have three phalangeal bones for
+the inner toe, four for the middle, and five for the outer one (see
+Fig. 408); but the impression of the terminal joint is that of the
+nail only. The fossil footprints exhibit regularly, where the
+joints are seen, the same number; and we see in each continuous
+line of tracks</p>
+
+<p class="fnote">* Hitchcock, Mem. of Amer. Acad., New Series, vol.
+iii, p. 129, 1848.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 382">[ 382 ]</a></p>
+
+<p>the three-jointed and five-jointed toes placed alternately
+outward, first on the one side, and then on the other. In some
+specimens, besides impressions of the three toes in front, the
+rudiment is seen of the fourth toe behind. It is not often that the
+matrix has been fine enough to retain impressions of the integument
+or skin of the foot; but in one fine specimen found at
+Turner&rsquo;s Falls, on the Connecticut, by Dr. Deane, these
+markings are well preserved, and have been recognised by Professor
+Owen as resembling the skin of the ostrich, and not that of
+reptiles.</p>
+
+<p>The casts of the footprints show that some of the fossil bipeds
+of the red sandstone of Connecticut had feet four times as large as
+the living ostrich, but scarcely, perhaps, larger than the Dinornis
+of New Zealand, a lost genus of feathered giants related to the
+Apteryx, of which there were many species which have left their
+bones and almost entire skeletons in the superficial alluvium of
+that island. By referring to what was said of the Iguanodon of the
+Wealden, the reader will perceive that the Dinosaur was somewhat
+intermediate between reptiles and birds, and left a series of
+tridactylous impressions on the sand.</p>
+
+<p>To determine the exact age of the red sandstone and shale
+containing these ancient footprints, in the United States, is not
+possible at present. No fossil shells have yet been found in the
+deposit, nor plants in a determinable state. The fossil fish are
+numerous and very perfect; but they are of a peculiar type, called
+<i>Ischypterus,</i> by Sir Philip Egerton, from the great size and
+strength of the fulcral rays of the dorsal fin, from ischus,
+strength, and pteron, a fin.</p>
+
+<p>The age of the Connecticut beds can not be proved by direct
+superposition, but may be presumed from the general structure of
+the country. That structure proves them to be newer than the
+movements to which the Appalachian or Allegheny chain owes its
+flexures, and this chain includes the ancient or pal&aelig;ozoic
+coal-formation among its contorted rocks.</p>
+
+<p><b>Coal-field of Richmond, Virginia.</b>&mdash;In the State of
+Virginia, at the distance of about 13 miles eastward of Richmond,
+the capital of that State, there is a coal-field occurring in a
+depression of the granite rocks, and occupying a geological
+position analogous to that of the New Red Sandstone,
+above-mentioned, of the Connecticut valley. It extends 26 miles
+from north to south, and from four to twelve from east to west.</p>
+
+<p>The plants consist chiefly of zamites, calamites, equiseta, and
+ferns, and, upon the whole, are considered by Professor</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 383">[ 383 ]</a></p>
+
+<p>Heer to have the nearest affinity to those of the European
+Keuper.</p>
+
+<p>The equiseta are very commonly met with in a vertical position
+more or less compressed perpendicularly. It is clear that they grew
+in the places where they are now buried in strata of hardened sand
+and mud. I found them maintaining their erect attitude, at points
+many miles apart, in beds both above and between the seams of coal.
+In order to explain this fact, we must suppose such shales and
+sandstones to have been gradually accumulated during the slow and
+repeated subsidence of the whole region.</p>
+
+<img src="../images3/fig409.jpg" width="230" height="164" alt=
+"Fig. 409: Triassic coal-shale, Richmond, Virginia." align="right">
+
+
+<p>The fossil fish are Ganoids, some of them of the genus <i>
+Catopterus,</i> others belonging to the liassic genus <i>
+Tetragonolepis (&AElig;chmodus),</i> see <a href=
+"../images3/fig375.jpg">Fig. 376.</a> Two species of <i>
+Entomostraca</i> called <i>Estheria</i> are in such profusion in
+some shaly beds as to divide them like the plates of mica in
+micaceous shales (see Fig. 409).</p>
+
+<p>These Virginian coal-measures are composed of grits, sandstones,
+and shales, exactly resembling those of older or primary date in
+America and Europe, and they rival, or even surpass, the latter in
+the richness and thickness of the coal-seams. One of these, the
+main seam, is in some places from 30 to 40 feet thick, composed of
+pure bituminous coal. The coal is like the finest kinds shipped at
+Newcastle, and when analysed yields the same proportions of carbon
+and hydrogen&mdash;a fact worthy of notice, when we consider that
+this fuel has been derived from an assemblage of plants very
+distinct specifically, and in part generically, from those which
+have contributed to the formation of the ancient or pal&aelig;ozoic
+coal.</p>
+
+<p><b>Triassic Mammifer.</b>&mdash;In North Carolina, the late
+Professor Emmons has described the strata of the Chatham
+coal-field, which correspond in age to those near Richmond, in
+Virginia. In beds underlying them he has met with three jaws of a
+small insectivorous mammal which he has called <i>Dromatherium
+sylvestre,</i> closely allied to <i>Spalacotherium.</i> Its nearest
+living analogue, says Professor Owen, &ldquo;is found in
+Myrmecobius; for each ramus of the lower jaw contained ten small
+molars in a continuous series, one canine, and three</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 384">[ 384 ]</a></p>
+
+<p>conical incisors&mdash;the latter being divided by short
+intervals.&rdquo;</p>
+
+<p><b>Low Grade of Early Mammals favourable to the Theory of
+Progressive Development.</b>&mdash;There is every reason to believe
+that this fossil quadruped is at least as ancient as the
+Microlestes of the European Trias described in <a href=
+"ch21.html#page 368">p. 368</a>; and the fact is highly important,
+as proving that a certain low grade of marsupials had not only a
+wide range in time, from the Trias to the Purbeck, or uppermost
+oolitic strata of Europe, but had also a wide range in space,
+namely, from Europe to North America, in an east and west
+direction, and, in regard to latitude, from Stonesfield, in 52&deg;
+N., to that of North Carolina, 35&deg; N.</p>
+
+<p>If the three localities in Europe where the most ancient
+mammalia have been found&mdash;Purbeck, Stonesfield, and
+Stuttgart&mdash;had belonged all of them to formations of the same
+age, we might well have imagined so limited an area to have been
+peopled exclusively with pouched quadrupeds, just as Australia now
+is, while other parts of the globe were inhabited by placentals;
+for Australia now supports one hundred and sixty species of
+marsupials, while the rest of the continents and islands are
+tenanted by about seventeen hundred species of mammalia, of which
+only forty-six are marsupial, namely, the opossums of North and
+South America. But the great difference of age of the strata in
+each of these three localities seems to indicate the predominance
+throughout a vast lapse of time (from the era of the Upper Trias to
+that of the Purbeck beds) of a low grade of quadrupeds; and this
+persistency of similar generic and ordinal types in Europe while
+the species were changing, and while the fish, reptiles, and
+mollusca were undergoing great modifications, would naturally lead
+us to suspect that there must also have been a vast extension in
+space of the same marsupial forms during that portion of the
+Secondary or Mesozoic epoch which has been termed &ldquo;the age of
+reptiles.&rdquo; Such an inference as to the wide geographical
+range of the ancient marsupials has been confirmed by the discovery
+in the Trias of North America of the above-mentioned Dromatherium.
+The predominance in earlier ages of these mammalia of a low grade,
+and the absence, so far as our investigations have yet gone, of
+species of higher organisation, whether aquatic or terrestrial, is
+certainly in favour of the theory of progressive development.</p>
+
+<br>
+<hr>
+<small><a href="contents.html">Contents</a> / <a href="ch20.html">
+Chapter XX</a> / <a href="ch22.html">Chapter XXII</a></small>
+</body>
+</html>
+