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+<p><b>The Student&rsquo;s Elements of Geology</b></p>
+
+<hr>
+<p class="page"><a name="page 211">[ 211 ]</a></p>
+
+<p>&nbsp;</p>
+
+<center><b>Chapter XIV</b><br>
+<br>
+MIOCENE PERIOD&mdash;UPPER MIOCENE.</center>
+
+<p class="intro">Upper Miocene Strata of France&mdash;Faluns of
+Touraine. &mdash; Tropical Climate implied by Testacea. &mdash;
+Proportion of recent Species of Shells. &mdash; faluns more ancient
+than the Suffolk Crag. &mdash; Upper Miocene of Bordeaux and the
+South of France. &mdash; Upper Miocene of &OElig;ningen, in
+Switzerland. &mdash; Plants of the Upper Fresh-water Molasse.
+&mdash; Fossil Fruit and Flowers as well as Leaves. &mdash; Insects
+of the Upper Molasse. &mdash; Middle or Marine Molasse of
+Switzerland. &mdash; Upper Miocene Beds of the Bolderberg, in
+Belgium. &mdash; Vienna Basin. &mdash; Upper Miocene of Italy and
+Greece. &mdash; Upper Miocene of India; Siwalik Hills. &mdash;
+Older Pliocene and Miocene of the United States.</p>
+
+<p><b>Upper Miocene Strata of France&mdash;Faluns of
+Touraine.</b>&mdash;The strata which we meet with next in the
+descending order are those called by many geologists &ldquo;Middle
+Tertiary,&rdquo; for which in 1833 I proposed the name of Miocene,
+selecting the &ldquo;faluns&rdquo; of the valley of the Loire, in
+France, as my example or type. I shall now call these falunian
+deposits Upper Miocene, to distinguish them from others to which
+the name of Lower Miocene will be given.</p>
+
+<p>No British strata have a distinct claim to be regarded as Upper
+Miocene, and as the Lower Miocene are also but feebly represented
+in the British Isles, we must refer to foreign examples in
+illustration of this important period in the earth&rsquo;s history.
+The term &ldquo;faluns&rdquo; is given provincially by French
+agriculturists to shelly sand and marl spread over the land in
+Touraine, just as similar shelly deposits were formerly much used
+in Suffolk to fertilise the soil, before the coprolitic or
+phosphatic nodules came into use. Isolated masses of such faluns
+occur from near the mouth of the Loire, in the neighbourhood of
+Nantes, to as far inland as a district south of Tours. They are
+also found at Pontlevoy, on the Cher, about seventy miles above the
+junction of that river with the Loire, and thirty miles south-east
+of Tours. Deposits of the same age also appear under new mineral
+conditions near the towns of Dinan and Rennes, in Brittany. I have
+visited all the localities above enumerated, and found the beds on
+the Loire to consist principally of sand and marl, in which are
+shells and corals, some entire, some rolled, and others in minute
+fragments. In certain districts, as at Dou&eacute;, in the
+Department of Maine and Loire, ten miles south-west</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 212">[ 212 ]</a></p>
+
+<p>of Saumur, they form a soft building-stone, chiefly composed of
+an aggregate of broken shells, bryozoa, corals, and echinoderms,
+united by a calcareous cement; the whole mass being very like the
+Coralline Crag near Aldborough, and Sudbourn in Suffolk. The
+scattered patches of faluns are of slight thickness, rarely
+exceeding fifty feet; and between the district called Sologne and
+the sea they repose on a great variety of older rocks; being seen
+to rest successively upon gneiss, clay-slate, various secondary
+formations, including the chalk; and, lastly, upon the upper
+fresh-water limestone of the Parisian tertiary series, which, as
+before mentioned <a href="ch9.html#page 142">(p. 142)</a>,
+stretches continuously from the basin of the Seine to that of the
+Loire.</p>
+
+<img src="../images1/fig136.jpg" width="170" height="235" alt=
+"Fig. 136: Dinotherium giganteum." align="left">
+
+<p>At some points, as at Louans, south of Tours, the shells are
+stained of a ferruginous colour, not unlike that of the Red Crag of
+Suffolk. The species are, for the most part, marine, but a few of
+them belong to land and fluviatile genera. Among the former, <i>
+Helix turonensis)</i> (<a href="../images/fig38.jpg">Fig. 38</a>) is
+the most abundant. Remains of terrestrial quadrupeds are here and
+there intermixed, belonging to the genera Dinotherium (Fig. 136),
+Mastodon, Rhinoceros, Hippopotamus, Ch&aelig;ropotamus, Dichobune,
+Deer, and others, and these are accompanied by cetacea, such as the
+Lamantin, Morse, Sea-calf, and Dolphin, all of extinct species.</p>
+
+<p>The fossil testacea of the faluns of the Loire imply, according
+to the late Edward Forbes, that the beds were formed partly on the
+shore itself at the level of low water, and partly at very moderate
+depths, not exceeding ten fathoms below that level. The molluscan
+fauna is, on the whole, much more littoral than that of the
+Pliocene Red and Coralline Crag of Suffolk, and implies a shallower
+sea. It is, moreover, contrasted with the Suffolk Crag by the
+indications it affords of an extra-European climate. Thus it
+contains seven species of Cypr&aelig;a, some larger than any
+existing cowry of the Mediterranean, several species of <i>Oliva,
+Ancillaria, Mitra, Terebra, Pyrula, Fasciolaria,</i> and <i>
+Conus.</i> Of the cones there are no less than eight species, some
+very large, whereas the only European cone now living is of
+diminutive size. The genus <i>Nerita,</i> and many others, are also
+represented by individuals</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 213">[ 213 ]</a></p>
+
+<p>of a type now characteristic of equatorial seas, and wholly
+unlike any Mediterranean forms. These proofs of a more elevated
+temperature seem to imply the higher antiquity of the faluns as
+compared with the Suffolk Crag, and are in perfect accordance with
+the fact of the smaller proportion of testacea of recent species
+found in the faluns.</p>
+
+<p>Out of 290 species of shells, collected by myself in 1840 at
+Pontlevoy, Louans, Boss&eacute;e, and other villages twenty miles
+south of Tours, and at Savign&eacute;, about fifteen miles
+north-west of that place, seventy-two only could be identified with
+recent species, which is in the proportion of twenty-five per cent.
+A large number of the 290 species are common to all the localities,
+those peculiar to each not being more numerous than we might expect
+to find in different bays of the same sea.</p>
+
+<p>The total number of species of testaceous mollusca from the
+faluns in my possession is 302, of which forty-five only, or
+fourteen per cent, were found by Mr. Wood to be common to the
+Suffolk Crag. The number of corals, including bryozoa and
+zoantharia, obtained by me at Dou&eacute; and other localities
+before adverted to, amounts to forty-three, as determined by Mr.
+Lonsdale, of which seven (one of them a zoantharian) agree
+specifically with those of the Suffolk Crag. Some of the genera
+occurring fossil in Touraine, as the corals Astrea and <i>
+Dendrophyllia</i>, and the bryozoan <i>Lunulites</i>, have not been
+found in European seas north of the Mediterranean; nevertheless,
+the zoantharia of the faluns do not seem to indicate, on the whole,
+so warm a climate as would be inferred from the shells.</p>
+
+<p>It was stated that, on comparing about 300 species of Touraine
+shells with about 450 from the Suffolk Crag, forty-five only were
+found to be common to both, which is in the proportion of only
+fifteen per cent. The same small amount of agreement is found in
+the corals also. I formerly endeavoured to reconcile this marked
+difference in species with the supposed co-existence of the two
+faunas, by imagining them to have severally belonged to distinct
+zoological provinces or two seas, the one opening to the north and
+the other to the south, with a barrier of land between them, like
+the Isthmus of Suez, now separating the Red Sea and the
+Mediterranean. But I now abandon that idea for several reasons;
+among others, because I succeeded in 1841 in tracing the Crag fauna
+southward in Normandy to within seventy miles of the Falunian type,
+near Dinan, yet found that both assemblages of fossils retained
+their distinctive characters, showing no signs of any blending of
+species or transition of climate.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 214">[ 214 ]</a></p>
+
+<p>The principal grounds, however, for referring the English Crag
+to the older Pliocene and the French faluns to the Upper Miocene
+epochs, consist in the predominance of fossil shells in the British
+strata identifiable with species not only still living, but which
+are now inhabitants of neighbouring seas, while the accompanying
+extinct species are of genera such as characterise Europe. In the
+faluns, on the contrary, the recent species are in a decided
+minority; and most of them are now inhabitants of the
+Mediterranean, the coast of Africa, and the Indian Ocean; in a
+word, less northern in character, and pointing to the prevalence of
+a warmer climate. They indicate a state of things receding farther
+from the present condition of Central Europe in physical geography
+and climate, and doubtless, therefore, receding farther from our
+era in time.</p>
+
+<img src="../images1/fig137.jpg" width="134" height="300" alt=
+"Fig. 137: Voluta Lamberti." align="left">
+
+<p>Among the conspicuous fossils common to the faluns of the Loire
+and the Suffolk Crag is a variety of the <i>Voluta Lamberti</i>, a
+shell already alluded to <a href="../images1/fig123.jpg">(Fig.
+123).</a> The specimens of this shell which I have myself collected
+in Touraine, or have seen in museums, are thicker and heavier than
+British individuals of the same species, and shorter in proportion
+to their width, and have the folds on the columella less oblique,
+as represented in Fig. 137.</p>
+
+<p><b>Upper Miocene of Bordeaux and the South of
+France.</b>&mdash;A great extent of country between the Pyrenees
+and the Gironde is overspread by tertiary deposits of various ages,
+and chiefly of Miocene date. Some of these, near Bordeaux, coincide
+in age with the faluns of Touraine, already mentioned, but many of
+the species of shells are peculiar to the south. The succession of
+beds in the basin of the Gironde implies several oscillations of
+level by which the same wide area was alternately converted into
+sea and land and into brackish-water lagoons, and finally into
+fresh-water ponds and lakes.</p>
+
+<p>Among the fresh-water strata of this age near the base of the
+Pyrenees are marls, limestones and sands, in which the eminent
+comparative anatomist, M. Lartet, has obtained a great number of
+fossil mammalia common to the faluns of the Loire and the Upper
+Miocene beds of Switzerland, such as <i>Dinotherium giganteum</i>
+and <i>Mastodon angustidens</i>; also</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 215">[ 215 ]</a></p>
+
+<p>the bones of quadrumana, or of the ape and monkey tribe, which
+were discovered in 1837, the first of that order of quadrupeds
+detected in Europe. They were found near Auch, in the Department of
+Gers, in latitude 43&deg; 39' N. About forty miles west of
+Toulouse. They were referred by MM. Lartet and Blainville to a
+genus closely allied to the Gibbon, to which they gave the name of
+<i>Pliopithecus.</i> Subsequently, in 1856, M. Lartet described
+another species of the same family of long-armed apes
+(<i>Hylobates</i>), which he obtained from strata of the same age
+at Saint-Gaudens, in the Haute Garonne. The fossil remains of this
+animal consisted of a portion of a lower jaw with teeth and the
+shaft of a humerus. It is supposed to have been a tree-climbing
+frugivorous ape, equalling man in stature. As the trunks of oaks
+are common in the lignite beds in which it lay, it has received the
+generic name of <i>Dryopithecus.</i> The angle formed by the
+ascending ramus of the jaw and the alveolar border is less open,
+and therefore more like the human subject, than in the Chimpanzee,
+and what is still more remarkable, the fossil, a young but adult
+individual, had all its milk teeth replaced by the second set,
+while its last true molar (or wisdom-tooth) was still undeveloped,
+or only existed as a germ in the jaw-bone. In the mode, therefore,
+of the succession of its teeth (which, as in all the old-World
+apes, exactly agree in number with those in man) it differed from
+the Gorilla and Chimpanzee, and corresponded with the human
+species.</p>
+
+<p><b>Upper Miocene Beds of &OElig;ningen, in
+Switzerland.</b>&mdash;The faluns of the Loire first served, as
+already stated (p. 211), as the type of the Miocene formations in
+Europe. They yielded a plentiful harvest of marine fossil shells
+and corals, but were entirely barren of plants and insects. In
+Switzerland, on the other hand, deposits of the same age have been
+discovered, remarkable for their botanical and entomological
+treasures. We are indebted to Professor Heer, of Zurich, for the
+description, restoration, and classification of several hundred
+species and varieties of these fossil plants, the whole of which he
+has illustrated by excellent figures in his &ldquo;Flora Tertiaria
+Helveti&aelig;.&rdquo; This great work, and those of Adolphe
+Brongniart, Unger, Goppert and others, show that this class of
+fossils is beginning to play the same important part in the
+classification of the tertiary strata containing lignite or brown
+coal as an older flora has long played in enabling us to understand
+the ancient coal or carboniferous formation. No small skepticism
+has always prevailed among botanists as to whether the leaves alone
+and the wood of plants could</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 216">[ 216 ]</a></p>
+
+<p>ever afford sufficient data for determining even genera and
+families in the vegetable kingdom. In truth, before such remains
+could be rendered available a new science had to be created. It was
+necessary to study the outlines, nervation, and microscopic
+structure of the leaves, with a degree of care which had never been
+called for in the classification of living plants, where the flower
+and fruit afforded characters so much more definite and
+satisfactory. As geologists, we can not be too grateful to those
+who, instead of despairing when so difficult a task was presented
+to them, or being discouraged when men of the highest scientific
+attainments treated the fossil leaves as worthless, entered with
+full faith and enthusiasm into this new and unexplored field. That
+they should frequently have fallen into errors was unavoidable, but
+it is remarkable, especially if we inquire into the history of
+Professor Heer&rsquo;s researches, how often early conjectures as
+to the genus and family founded on the leaves alone were afterwards
+confirmed when fuller information was obtained. As examples to be
+found on comparing Heer&rsquo;s earlier and later works, I may
+instance the chestnut, elm, maple, cinnamon, magnolia, buckbean or
+Menyanthes, vine, buckthorn (<i>Rhamnus</i>), <i>Andromeda</i> and
+<i>Myrica,</i> and among the conifers <i>Sequoia</i> and <i>
+Taxodium.</i> In all these cases the plants were first recognised
+by their leaves, and the accuracy of the determination was
+afterwards confirmed when the fruit, and in some instances both
+fruit and flower, were found attached to the same stem as the
+leaves.</p>
+
+<p>But let us suppose that no fruit, seed, or flower had ever been
+met with in a fossil state, we should still have been indebted to
+the persevering labours of botanical pal&aelig;ontologists for one
+of the grandest scientific discoveries for which the present
+century is remarkable&mdash;namely, the proofs now established of
+the prevalence of a mild climate and a rich arborescent flora in
+the arctic regions in that Miocene epoch on the history of which we
+are now entering. It may be useful if I endeavour to give the
+reader in a few words some idea of the nature of the evidence of
+these important conclusions, to show how far they may be safely
+based on fossil leaves alone. When we begin by studying the fossils
+of the Newer Pliocene deposits, such as those of the Upper Val
+d&rsquo;Arno, before alluded to, we perceive that the fossil
+foliage agrees almost entirely with the trees and shrubs of a
+modern European forest. In the plants of the Older Pliocene strata
+of the same region we observe a larger proportion of species and
+genera which, although they may agree with well-known Asiatic or
+other foreign types, are at present</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 217">[ 217 ]</a></p>
+
+<p>wanting in Italy. If we then examine the Miocene formations of
+the same country, exotic forms become more abundant, especially the
+palms, whether they belong to the European or American fan-palms,
+<i>Cham&aelig;rops</i> and <i>Sabal</i>, or to the more tropical
+family of the date-palms or <i>Ph&oelig;nicites</i>, which last are
+conspicuous in the Lower Miocene beds of Central Europe. Although
+we have not found the fruit or flower of these palms in a fossil
+state, the leaves are so characteristic that no one doubts the
+family to which they belong, or hesitates to accept them as
+indications of a warm and sub-tropical climate.</p>
+
+<p>When the Miocene formations are traced to the northward of the
+50th degree of latitude, the fossil palms fail us, but the greater
+proportion of the leaves, whether identical with those of existing
+European trees or of forms now unknown in Europe, which had
+accompanied the Miocene palms, still continue to characterise rocks
+of the same age, until we meet with them not only in Iceland, but
+in Greenland, in latitude 70&deg; N., and in Spitzbergen, latitude
+78&deg; 56', or within about 11 degrees of the pole, and under
+circumstances which clearly show them to have been indigenous in
+those regions, and not to have been drifted from the south <a href=
+"ch15.html#page 240">(see p. 240).</a> Not only, therefore, has the
+botanist afforded the geologist much pal&aelig;ontological
+assistance in identifying distinct tertiary formations in distant
+places by his power of accurately discriminating the forms,
+veining, and microscopic structure of leaves or wood, but,
+independently of that exact knowledge derivable from the organs of
+fructification, we are indebted to him for one of the most novel,
+unexpected results of modern scientific inquiry.</p>
+
+<p>The Miocene formations of Switzerland have been called <i>
+Molasse</i>, a term derived from the French <i>mol</i>, and applied
+to a <i>soft</i>, incoherent, greenish sandstone, occupying the
+country between the Alps and the Jura. This molasse comprises three
+divisions, of which the middle one is marine, and being closely
+related by its shells to the faluns of Touraine, may be classed as
+Upper Miocene. The two others are fresh-water, the upper of which
+may be also grouped with the faluns, while the lower must be
+referred to the Lower Miocene, as defined in the next chapter.</p>
+
+<p><b>Upper Fresh-water Molasse.</b>&mdash;This formation is best
+seen at &OElig;ningen, in the valley of the Rhine, between Constance
+and Schaffhausen, a locality celebrated for having produced in the
+year 1700 the supposed human skeleton called by Scheuchzer
+&ldquo;homo diluvii testis,&rdquo; a fossil afterwards demonstrated
+by Cuvier to be a reptile, or aquatic salamander,</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 218">[ 218 ]</a></p>
+
+<p>of larger dimensions than even its great living representative,
+the salamander of Japan.</p>
+
+<p>The &OElig;ningen strata consist of a series of marls and
+limestones, many of them thinly laminated, and which appear to have
+slowly accumulated in a lake probably fed by springs holding
+carbonate of lime in solution. The elliptical area over which this
+fresh-water formation has been traced extends, according to Sir
+Roderick Murchison, for a distance of ten miles east and west from
+Berlingen, on the right bank of the river to Wangen, and to
+&OElig;ningen, near Stein, on the left bank. The organic remains
+have been chiefly derived from two quarries, the lower of which is
+about 550 feet above the level of the Lake of Constance, while the
+upper quarry is 150 feet higher. In this last, a section thirty
+feet deep displays a great succession of beds, most of them
+splitting into slabs and some into very thin lamin&aelig;.
+Twenty-one beds are enumerated by Professor Heer, the uppermost a
+bluish-grey marl seven feet thick, with organic remains, resting on
+a limestone with fossil plants, including leaves of poplar,
+cinnamon, and pond-weed (<i>Potamogeton</i>), together with some
+insects; while in the bed No. 4, below, is a bituminous rock, in
+which the <i>Mastodon tapiroides</i>, a characteristic Upper
+Miocene quadruped, has been met with. The 5th bed, two or three
+inches thick, contains fossil fish, e.g., <i>Leuciscus</i> (roach),
+and the larv&aelig; of dragon-flies, with plants such as the elm
+(<i>Ulmus</i>), and the aquatic Chara. Below this are other
+plant-beds; and then, in No. 9, the stone in which the great
+salamander (Andrias Scheuchzeri) and some fish were found. Below
+this other strata occur with fish, tortoises, the great salamander
+before alluded to, fresh-water mussels, and plants. In No. 16 the
+fossil fox of &OElig;ningen, <i>Galecynus &OElig;ningensis,</i> Owen,
+was obtained by Sir R. Murchison. To this succeed other beds with
+mammalia (<i>Lagomys</i>), reptiles, (<i>Emys</i>), fish, and
+plants, such as walnut, maple, and poplar. In the 19th bed are
+numerous fish, insects, and plants, below which are marls of a blue
+indigo colour.</p>
+
+<p>In the lower quarry eleven beds are mentioned, in which, as in
+the upper, both land and fresh-water plants and many insects occur.
+In the 6th, reckoning from the top, many plants have been obtained,
+such as <i>Liquidambar, Daphnogene, Podogonium,</i> and <i>
+Ulmus</i>, together with tortoises, besides the bones and teeth of
+a ruminant quadruped, named by H. von Meyer <i>Pal&aelig;omeryx
+eminens.</i> No. 9 is called the insect-bed, a layer only a few
+inches thick, which, when exposed to the frost, splits into leaves
+as thin as paper. In these thin lamin&aelig; plants such as <i>
+Liquidambar, Daphnogene,</i> and <i>Glyptostrobus</i>,</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 219">[ 219 ]</a></p>
+
+<p>occur, with innumerable insects in a wonderful state of
+preservation, usually found singly. Below this is an indigo-blue
+marl, like that at the bottom of the higher quarry, resting on
+yellow marl ascertained to be at least thirty feet thick.</p>
+
+<img src="../images1/fig138.jpg" width="216" height="299" alt=
+"Fig. 138: Cinnamomum polymorphum." align="left">
+
+<p>All the above fossil-bearing strata were evidently formed with
+extreme slowness. Although the fossiliferous beds are, in the
+aggregate, no more than a few yards in thickness, and have only
+been examined in the small area comprised in the two quarries just
+alluded to, they give us an insight into the state of animal and
+vegetable life in part of the Upper Miocene period, such as no
+other region in the world has elsewhere supplied. In the year 1859,
+Professor Heer had already determined no less than 475 species of
+plants and more than 800 insects from these &OElig;ningen beds. He
+supposes that a river entering a lake floated into it some of the
+leaves and land insects, together with the carcasses of quadrupeds,
+among others a great Mastodon. Occasionally, during tempests, twigs
+and even boughs of trees with their leaves were torn off and
+carried for some distance so as to reach the lake. Springs,
+containing carbonate of lime, seem at some points to have supplied
+calcareous matter in solution, giving origin locally to a kind of
+travertin, in which organic bodies sinking to the bottom became
+hermetically sealed up. The lamin&aelig;, says Heer, which
+immediately succeed each other were not all formed at the same
+season, for it can be shown that, when some of them originated,
+certain plants were in flower, whereas, when the next of these
+layers was produced, the same plants had ripened their fruit. This
+inference is confirmed by independent proofs derived from insects.
+The principal insect-bed is rarely two inches thick, and is
+composed, says Heer, of about 250 leaf-like lamin&aelig;, some of
+which were deposited in the spring, when the <i>Cinnamomum
+polymorphum</i> (Fig. 138) was in flower, others in summer, when
+winged ants were numerous, and when the poplar and willow had
+matured their seed; others, again, in autumn, when the same <i>
+Cinnamomum polymorphum</i> (Fig. 138) was in fruit, as well as the
+liquidambar, oak, clematis,</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 220">[ 220 ]</a></p>
+
+<p>and many other plants. The ancient lake seems to have had a belt
+of poplars and willows round its borders, countless leaves of which
+were imbedded in mud, and together with them, at some points, a
+species of reed, <i>Arundo</i>, which was very common.</p>
+
+<p>One of the most characteristic shrubs is a papilionaceous and
+leguminous plant of an extinct genus, called by Heer <i>
+Podogonium</i>, of which two species are known. Entire twigs have
+been found with flowers, and always without leaves, as the flowers
+evidently came out, as in the poplar and willow tribe, before any
+leaves made their appearance. Other specimens have been obtained
+with ripe fruits accompanied by leaves, which resemble those of the
+tamarind, to which it was evidently allied, being of the family
+C&aelig;salpine&aelig;, now proper to warmer regions.</p>
+
+<img src="../images1/fig139.jpg" width="251" height="337" alt=
+"Fig. 138: Acer trilobatum." align="left">
+
+<p>The Upper Miocene flora of &OElig;ningen is peculiarly important,
+in consequence of the number of genera of which not merely the
+leaves, but, as in the case of the <i>Podogonium</i> just
+mentioned, the fruit also and even the flower are known. Thus there
+are nineteen species of maple, ten of which have already been found
+with fruit. Although in no one region of the globe do so many
+maples now flourish, we need not suspect Professor Heer of having
+made too many species in this genus when we consider the manner in
+which he has dealt with one of them, <i>Acer trilobatum</i>, Figs.
+139 and 140. Of this plant the number of marked varieties figured
+and named is very great, and no less than three of them had been
+considered as distinct species by other botanists, while six of the
+others might have laid claim, with nearly equal propriety, to a
+like distinction. The common form, called <i>Acer trilobatum</i>,
+Fig. 139, may be taken as a normal representative of the
+&OElig;ningen fossil, and Fig. 140, as one of the most divergent
+varieties, having almost four lobes in the leaf instead of
+three.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 221">[ 221 ]</a></p>
+
+<center><img src="../images1/fig140.jpg" width="285" height="233" alt=
+"Fig. 140: Acer trilobatum."></center>
+
+<p>Among the conspicuous genera which abounded in the Miocene
+period in Europe is the plane-tree, <i>Platanus,</i> the fossil
+species being considered by Heer to come nearer to the American <i>
+P. occidentalis</i> than to <i>P. orientalis</i> of Greece and Asia
+Minor. In some of the fossil specimens the male flowers are
+preserved. Among other points of resemblance with the living
+plane-trees, as we see them in the parks and squares of London,
+fossil fragments of the trunk are met with, having pieces of their
+bark peeling off.</p>
+
+<img src="../images1/fig141.jpg" width="166" height="271" alt=
+"Platanus aceroides." align="right">
+
+<p>The vine of &OElig;ningen, <i>Vitis teutonica</i>, Ad. Brong, is
+of a North American type. Both the leaves and seeds have been found
+at &OElig;ningen, and bunches of compressed grapes of the same
+species have been met with in the brown coal of Wetteravia in
+Germany. No less than eight species of smilax, a monocotyledonous
+genus, occur at &OElig;ningen and in other Upper Miocene localities,
+the flowers of some of them, as well as the leaves, being
+preserved; as in the case of the very common fossil, <i>S.
+sagittifera</i>, Fig. 142, <i>a.</i></p>
+
+<p>Leaves of plants supposed to belong to the order Proteace&aelig;
+have been obtained partly from &OElig;ningen and partly from the
+lacustrine formation of the same age at Locle in the Jura. They
+have been referred to the genera <i>Banksia, Grevillea, Hakea,</i>
+and <i>Persoonia.</i> Of Hakea there is the impression of a
+supposed seed-vessel, with its characteristic thick stalk and
+seeds, but as the fruit is without structure, and has not yet</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 222">[ 222 ]</a></p>
+
+<p>been found attached to the same stem as the leaf, the proof is
+incomplete.</p>
+
+<img src="../images1/fig142.jpg" width="208" height="291" alt=
+"Fig. 142: Smilax sagittifera." align="left">
+
+<p>To whatever family the foliage hitherto regarded as proteaceous
+by many able pal&aelig;ontologists may eventually be shown to
+belong, we must be careful not to question their affinity to that
+order of plants on those geographical considerations which have
+influenced some botanists. The nearest living Proteace&aelig; now
+feel the in Abyssinia in lat. 20&deg; N., but the greatest number
+are confined to the Cape and Australia. The ancestors, however, of
+the &OElig;ningen fossils ought not to be looked for in such distant
+regions, but from that European land which in Lower Miocene times
+bore trees with similar foliage, and these had doubtless an Eocene
+source, for cones admitted by all botanists to be proteaceous have
+been met with in one division of that older Tertiary group <a href=
+"../images1/fig206.jpg">(see Fig. 206</a>). The source of these last,
+again, must not be sought in the antipodes, for in the white chalk
+of Aix-la-Chapelle leaves like those of Grevillea and other
+proteaceous genera have been found in abundance, and, as we shall
+see <a href="ch17.html#page 304">(p. 304)</a> in a most perfect
+state of preservation. All geologists agree that the distribution
+of the Cretaceous land and sea had scarcely any connection with the
+present geography of the globe.</p>
+
+<center><img src="../images1/fig143.jpg" width="490" height="157" alt=
+"Fig. 143: Fruit of the fossil and recent species of Hakea, a genus of Proteace&aelig;.">
+</center>
+
+<p>In the same beds with the supposed Proteace&aelig; there occurs
+at Locle a fan-palm of the American type Sabal (for genus see Fig.
+151), a genus which ranges throughout the low country near the sea
+from the Carolinas to Florida and</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 223">[ 223 ]</a></p>
+
+<p>Louisiana. Among the Conifer&aelig; of Upper Miocene age is
+found a deciduous cypress nearly allied to the <i>Taxodium
+distichum</i> of North America, and a <i>Glyptostrobus</i> (Fig.
+144), very like the Japanese <i>G. heterophyllus,</i> now common in
+our shrubberies.</p>
+
+<img src="../images1/fig144.jpg" width="120" height="214" alt=
+"Fig. 144: Glyptostrobus Europ&aelig;us." align="right">
+
+<p>Before the appearance of Heer&rsquo;s work on the Miocene Flora
+of Switzerland, Unger and Goppert had already pointed out the large
+proportion of living North American genera which distinguished the
+vegetation of the Miocene period in Central Europe. Next in number,
+says Heer, to these American forms at &OElig;ningen the European
+genera preponderate, the Asiatic ranking in the third, the African
+in the fourth, and the Australian in the fifth degree. The American
+forms are more numerous than in the Italian Pliocene flora, and the
+whole vegetation indicates a warmer climate than the Pliocene,
+though not so high a temperature as that of the older or Lower
+Miocene period.</p>
+
+<p>The conclusions drawn from the insects are for the most part in
+perfect harmony with those derived from the plants, but they have a
+somewhat less tropical and less American aspect, the South European
+types being more numerous. On the whole, the insect fauna is richer
+than that now inhabiting any part of Europe. No less than 844
+species are reckoned by Heer from the &OElig;ningen beds alone, the
+number of specimens which he has examined being 5080. The entire
+list of Swiss species from the Upper and Lower Miocene together
+amount to 1322. Almost all the living families of Coleoptera are
+represented, but, as we might have anticipated from the
+preponderance of arborescent and ligneous plants, the wood-eating
+beetles play the most conspicuous part, the Buprestid&aelig; and
+other long-horned beetles being particularly abundant.</p>
+
+<p>The patterns and some remains of the colours both of <i>
+Coleoptera</i> and <i>Hemiptera</i> are preserved at &OElig;ningen,
+as, for example in <i>Harpactor</i> (Fig. 145), in which the
+antenn&aelig;, one of the eyes, and the legs and wings are
+retained. The characters, indeed, of many of the insects are so
+well defined as to incline us to believe that if this class of the
+invertebrata were not so rare and local, they might be more useful
+than even the plants and shells in settling chronological points in
+geology.</p>
+
+<p><b>Middle or Marine Molasse (Upper Miocene) of
+Switzerland.</b>&mdash;It was before stated that the Miocene
+formation of Switzerland</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 224">[ 224 ]</a></p>
+
+<img src="../images1/fig145.jpg" width="172" height="284" alt=
+"Fig. 145: Harpactor maculipes." align="left">
+
+<p>consisted of, first, the upper fresh-water molasse, comprising
+the lacustrine marls of &OElig;ningen; secondly, the marine molasse,
+corresponding in age to the faluns of Touraine; and thirdly, the
+lower fresh-water molasse. Some of the beds of the marine or middle
+series reach a height of 2470 feet above the sea. A large number of
+the shells are common to the faluns of Touraine, the Vienna basin,
+and other Upper Miocene localities. The terrestrial plants play a
+subordinate part in the fossiliferous beds, yet more than ninety of
+them are enumerated by Heer as belonging to this falunian division,
+and of these more than half are common to subjacent Lower Miocene
+beds, while a proportion of about forty-five in one hundred are
+common to the overlying &OElig;ningen flora. Twenty-six of the
+ninety-two species are peculiar.</p>
+
+<img src="../images1/fig146.jpg" width="119" height="174" alt=
+"Fig. 146: Olica Dufresnii." align="right">
+
+<p><b>Upper Miocene of the Bolderberg, in Belgium.</b>&mdash;In a
+small hill or ridge called the Bolderberg, which I visited in 1851,
+situated near Hasselt, about forty miles E.N.E. of Brussels, strata
+of sand and gravel occur, to which M. Dumont first called attention
+as appearing to constitute a northern representative of the faluns
+of Touraine. On the whole, they are very distinct in their fossils
+from the two upper divisions of the Antwerp Crag before mentioned
+<a href="ch13.html#page 204">(p. 204)</a>, and contain shells of
+the genera <i>Oliva, Conus, Ancillaria, Pleurotoma,</i> and <i>
+Cancellaria</i> in abundance. The most common shell is an Olive
+(Fig. 146), called by Nyst <i>Oliva Dufresnii</i>; and
+constituting, as M. Bosquet observes, a smaller and shorter variety
+of the Bordeaux species.</p>
+
+<p>So far as the shells of the Bolderberg are known, the proportion
+of recent species agrees with that in the faluns of Touraine, and
+the climate must have been warmer than that of the Coralline Crag
+of England.</p>
+
+<p><b>Upper Miocene Beds of the Vienna Basin.</b>&mdash;In South
+Germany the general resemblance of the shells of the Vienna
+tertiary basin with those of the faluns of Touraine has long been
+acknowledged. In the late Dr. H&ouml;rnes&rsquo;s excellent
+work</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 225">[ 225 ]</a></p>
+
+<p>on the fossil mollusca of that formation, we see accurate
+figures of many shells, clearly of the same species as those found
+in the falunian sands of Touraine.</p>
+
+<p>According to Professor Suess, the most ancient and purely marine
+of the Miocene strata in this basin consist of sands,
+conglomerates, limestones, and clays, and they are inclined inward,
+or from the borders of the trough towards the centre, their
+outcropping edges rising much higher than the newer beds, whether
+Miocene or Pliocene, which overlie them, and which occupy a smaller
+area at an inferior elevation above the sea. M. Hornes has
+described no less than 500 species of gasteropods, of which he
+identifies one-fifth with living species of the Mediterranean,
+Indian, or African seas, but the proportion of existing species
+among the lamellibranchiate bivalves exceeds this average. Among
+many univalves agreeing with those of Africa on the eastern side of
+the Atlantic are <i>Cypr&aelig;a sanguinolenta, Buccinum
+lyratum,</i> and <i>Oliva flammulata.</i> In the lowest marine beds
+of the Vienna basin the remains of several mammalia have been
+found, and among them a species of <i>Dinotherium</i>, a Mastodon
+of the <i>Trilophodon</i> family, a Rhinoceros (allied to <i>R.
+megarhinus</i>, Christol), also an animal of the hog tribe, <i>
+Listriodon</i>, von Meyer, and a carnivorous animal of the canine
+family. The <i>Helix turonensis</i> <a href="../images/fig38.jpg">
+(Fig. 38)</a>, the most common land shell of the French faluns,
+accompanies the above land animals. In a higher member of the
+Vienna Miocene series are found <i>Dinotherium giganteum</i> <a
+href="../images1/fig136.jpg">(Fig. 136)</a>, <i>Mastodon longirostris,
+Rhinoceros Schleiermacheri, Acerotherium incisivum,</i> and <i>
+Hippotherium gracile,</i> all of them equally characteristic of an
+Upper Miocene deposit occurring at Eppelsheim, in Hesse Darmstadt;
+a locality also remarkable as having furnished in latitude 49&deg;
+50' N. the bone of a large ape of the Gibbon kind, the most
+northerly example yet discovered of a quadrumanous animal.</p>
+
+<img src="../images1/fig147.jpg" width="130" height="164" alt=
+"Fig. 147: Amphistegina Hauerina." align="right">
+
+<p>M. Alcide d&rsquo;Orbigny has shown that the foraminifera of the
+Vienna basin differ alike from the Eocene and Pliocene species, and
+agree with those of the faluns, so far as the latter are known.
+Among the Vienna foraminifera, the genus <i>Amphistegina</i> (Fig.
+147) is very characteristic, and is supposed by d&rsquo;Archiac to
+take the same place among the Rhizopods of the Upper Miocene era
+which the Nummulites occupy in the Eocene period.</p>
+
+<p>The flora of the Vienna basin exhibits some species which</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 226">[ 226 ]</a></p>
+
+<p>have a general range through the whole Miocene period, such as
+<i>Cinnamomum polymorphum</i> <a href="../images1/fig138.jpg">(Fig.
+138)</a>, and <i>C. Scheuchzeri,</i> also Planera Richardi, Mich.,
+<i>Liquidambar europ&aelig;um</i> <a href="../images1/fig134.jpg">
+(Fig. 135)</a> <i>Juglans bilinica, Cassia ambigua,</i> and <i>C.
+lignitum.</i> Among the plants common to the Upper Miocene beds of
+&OElig;ningen, in Switzerland, are <i>Platanus aceroides</i> <a
+href="../images1/fig141.jpg">(Fig. 141)</a>, <i>Myrica
+vindobonensis,</i> and others.</p>
+
+<p><b>Upper Miocene Strata of Italy.</b>&mdash;We are indebted to
+Signor Michelotti for a valuable work on the Miocene shells of
+Northern Italy. Those found in the hill called the Superga, near
+Turin, have long been known to correspond in age with the faluns of
+Touraine, and they contain so many species common to the Upper
+Miocene strata of Bordeaux as to lead to the conclusion that there
+was a free communication between the northern part of the
+Mediterranean and the Bay of Biscay in the Upper Miocene
+period.</p>
+
+<p><b>Upper Miocene Formations of Greece.</b>&mdash;At
+Pikerm&eacute;, near Athens, MM. Wagner and Roth have described a
+deposit in which they found the remains of the genera <i>Mastodon,
+Dinotherium, Hipparion,</i> two species of <i>Giraffe,
+Antelope,</i> and others, some living and some extinct. With them
+were also associated fossil bones of the <i>Semnopithecus,</i>
+showing that here, as in the south of France, the quadrumana were
+characteristic of this period. The whole fauna attests the former
+extension of a vast expanse of grassy plains where we have now the
+broken and mountainous country of Greece; plains, which were
+probably united with Asia Minor, spreading over the area where the
+deep &AElig;gean Sea and its numerous islands are now situated. We
+are indebted to M. Gaudry, who visited Pikerm&eacute;, for a
+treatise on these fossil bones, showing how many data they
+contribute to the theory of a transition from the mammalia of the
+Upper Miocene through the Pliocene and Post-pliocene forms to those
+of living genera and species.</p>
+
+<p><b>Upper Miocene of India. Siw&acirc;lik Hills.</b>&mdash;The
+Siw&acirc;lik Hills lie at the southern foot of the Himalayan
+chain, rising to the height of 2000 and 3000 feet. Between the
+Jumna and the Ganges they consist of inclined strata of sandstone,
+shingle, clay, and marl. We are indebted to the indefatigable
+researches of Dr. Falconer and Sir Proby Cautley, continued for
+fifteen years, for the discovery in these marls and sandstones of a
+great variety of fossil mammalia and reptiles, together with many
+fresh-water shells. Out of fifteen species of shells of the genera
+<i>Paludina, Melania, Ampullaria,</i> and <i>Unio,</i> all are
+extinct or unknown species with the exception of four, which are
+still inhabitants of Indian rivers. Such a</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 227">[ 227 ]</a></p>
+
+<p>proportion of living to extinct mollusca agrees well with the
+usual character of an Upper Miocene or Falunian fauna, as observed
+in Touraine, or in the basin of Vienna and elsewhere.</p>
+
+<p>The genera of mammalia point in the same direction. One of them,
+of the genus <i>Chalicotherium</i> (or <i>Anisodon</i> of Lartet),
+is a pachyderm intermediate between the <i>Rhinoceros</i> and <i>
+Anoplothere,</i> and characteristic of the Upper Miocene strata of
+Eppelsheim, and of the south of France. With it occurs also an
+extinct form of Hippopotamus, called Hexaprotodon, and a species of
+Hippotherium and pig, also two species of <i>Mastodon</i>, two of
+elephant, and three other elephantine proboscidians; none of them
+agreeing with any fossil forms of Europe, and being intermediate
+between the genera Elephas and Mastodon, constituting the sub-genus
+<i>Stegodon</i> of Falconer. With these are associated a monkey,
+allied to the <i>Semnopithecus entellus</i>, now living in the
+Himalaya, and many ruminants. Among these last, besides the
+giraffe, camel, antelope, stag, and others, we find a remarkable
+new type, the <i>Sivatherium,</i> like a gigantic four-horned deer.
+There are also new forms of carnivora, both feline and canine, the
+<i>Machairodus</i> among the former, also hy&aelig;nas, and a
+subursine form called the Hy&aelig;narctos, and a genus allied to
+the otter (<i>Enhydriodon</i>), of formidable size.</p>
+
+<p>The giraffe, camel, and a large ostrich may be cited as proofs
+that there were formerly extensive plains where now a steep chain
+of hills, with deep ravines, runs for many hundred miles east and
+west. Among the accompanying reptiles are several crocodiles, some
+of huge dimensions, and one not distinguishable, says Dr. Falconer,
+from a species now living in the Ganges (<i>C. Gangeticus</i>); and
+there is still another saurian which the same anatomist has
+identified with a species now inhabiting India. There was also an
+extinct species of tortoise of gigantic proportions
+(<i>Colossochelys Atlas</i>), the curved shell of which was twelve
+feet three inches long and eight feet in diameter, the entire
+length of the animal being estimated at eighteen feet, and its
+probable height seven feet.</p>
+
+<p>Numerous fossils of the Siw&acirc;lik type have also been found
+in Perim Island, in the Gulf of Cambay, and among these a species
+of <i>Dinotherium,</i> a genus so characteristic of the Upper
+Miocene period in Europe.</p>
+
+<p><b>Older Pliocene and Miocene Formations in the United
+States.</b>&mdash;Between the Alleghany Mountains, formed of older
+rocks, and the Atlantic, there intervenes, in the United States, a
+low region occupied principally by beds of marl, clay, and sand,
+consisting of the cretaceous and tertiary formations,</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 228">[ 228 ]</a></p>
+
+<p>and chiefly of the latter. The general elevation of this plain
+bordering the Atlantic does not exceed 100 feet, although it is
+sometimes several hundred feet high. Its width in the middle and
+southern states is very commonly from 100 to 150 miles. It
+consists, in the South, as in Georgia, Alabama, and South Carolina,
+almost exclusively of Eocene deposits; but in North Carolina,
+Maryland, Virginia, Delaware, more modern strata predominate, of
+the age of the English Crag and faluns of Touraine.*</p>
+
+<center><img src="../images1/fig148.jpg" width="399" height="221" alt=
+"Fig. 148: Fulgur canaliculatus. Fig. 149: Fusus quadricostatus.">
+</center>
+
+<p>In the Virginian sands, we find in great abundance a species of
+Astarte (<i>A. undulata,</i> Conrad), which resembles closely, and
+may possibly be a variety of, one of the commonest fossils of the
+Suffolk Crag (<i>A. Omalii</i>); the other shells also, of the
+genera <i>Natica, Fissurella, Artemis, Lucina, Chama,
+Pectunculus,</i> and <i>Pecten,</i> are analagous to shells both of
+the English Crag and French faluns, although the species are almost
+all distinct. Out of 147 of these American fossils I could only
+find thirteen species common to Europe, and these occur partly in
+the Suffolk Crag, and partly in the faluns of Touraine; but it is
+an important characteristic of the American group, that it not only
+contains many peculiar extinct forms, such as <i>Fusus
+quadricostatus,</i> Say (see Fig. 149), and <i>Venus
+tridacnoides,</i> abundant in these same formations, but also some
+shells which, like <i>Fulgur carica</i> of Say and <i>F.
+canaliculatus</i> (see Fig. 148), <i>Calyptr&aelig;a costata, Venus
+mercenaria,</i> Lam., <i>Modiola glandula,</i> Totten, and <i>
+Pecten magellanicus,</i> Lam., are recent species, yet of forms now
+confined to the western side of the Atlantic&mdash;a fact implying
+that some traces of the beginning of the present geographical
+distribution of mollusca</p>
+
+<p class="fnote">* Proceedings of the Geol. Soc., vol. iv, pt. iii,
+1845, p. 547.</p>
+
+<p>&nbsp;</p>
+
+<hr>
+<p class="page"><a name="page 229">[ 229 ]</a></p>
+
+<p>date back to a period as remote as that of the Miocene
+strata.</p>
+
+<img src="../images1/fig150.jpg" width="152" height="184" alt=
+"Fig. 150: Astrangia lineata." align="right">
+
+<p>Of ten species of corals which I procured on the banks of the
+James River, one agrees generically with a coral now living on the
+coast of the United States. Mr. Lonsdale regarded these corals as
+indicating a temperature exceeding that of the Mediterranean, and
+the shells would lead to similar conclusions. Those occurring on
+the James River are in the 37th degree of N. latitude, while the
+French faluns are in the 47th; yet the forms of the American
+fossils would scarcely imply so warm a climate as must have
+prevailed in France when the Miocene strata of Touraine
+originated.</p>
+
+<p>Among the remains of fish in these post-eocene strata of the
+United States are several large teeth of the shark family, not
+distinguishable specifically from fossils of the faluns of
+Touraine.</p>
+
+<br>
+<hr>
+<small><a href="contents.html">Contents</a> / <a href="ch13.html">
+Chapter XIII</a> / <a href="ch15.html">Chapter XV</a></small>
+</body>
+</html>
+