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+
+The Project Gutenberg EBook of A New Pocket Gopher (Genus Thomomys), from
+Eastern Colorado, by E. Raymond Hall
+
+This eBook is for the use of anyone anywhere at no cost and with
+almost no restrictions whatsoever. You may copy it, give it away or
+re-use it under the terms of the Project Gutenberg License included
+with this eBook or online at www.gutenberg.org
+
+
+Title: A New Pocket Gopher (Genus Thomomys), from Eastern Colorado
+
+Author: E. Raymond Hall
+
+Release Date: January 4, 2011 [EBook #34836]
+
+Language: English
+
+Character set encoding: UTF-8
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+*** START OF THIS PROJECT GUTENBERG EBOOK A NEW POCKET GOPHER (GENUS ***
+
+
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+Produced by Chris Curnow, Joseph Cooper and the Online
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+
+
+
+
+
+<p><span class="pagenum"><a name="Page_81" id="Page_81">[Pg 81]</a></span></p>
+
+<h1>A New Pocket Gopher (Genus Thomomys)<br />
+from Eastern Colorado</h1>
+
+<h5>BY</h5>
+
+<h3>E. RAYMOND HALL</h3>
+
+<h4>University of Kansas Publications<br />
+Museum of Natural History<br />
+<br />
+Volume 5, No. 8, pp. 81-85<br />
+October 1, 1951</h4>
+
+<h4>University of Kansas<br />
+LAWRENCE<br />
+1951</h4>
+
+
+
+<hr style="width: 65%;" />
+<p><span class="pagenum"><a name="Page_82" id="Page_82">[Pg 82]</a></span></p>
+
+<h4><span class="smcap">University of Kansas Publications, Museum of Natural History</span><br />
+<br />
+Editors: E. Raymond Hall, Chairman, A. Byron Leonard,<br />
+Edward H. Taylor, Robert W. Wilson<br />
+<br />
+Volume 5, No. 8, pp. 81-85<br />
+October 1, 1951<br />
+<br />
+<br />
+<big><span class="smcap">University of Kansas</span><br />
+Lawrence, Kansas</big></h4>
+
+<h5>PRINTED BY<br />
+FERD VOILAND, JR., STATE PRINTER<br />
+TOPEKA, KANSAS<br />
+1951<br />
+<br />
+23-7439<br />
+</h5>
+
+
+<hr style="width: 65%;" />
+<p><span class="pagenum"><a name="Page_83" id="Page_83">[Pg 83]</a></span></p>
+<h2>A New Pocket Gopher (Genus Thomomys)<br />
+from Eastern Colorado</h2>
+
+<h4>By</h4>
+
+<h3>E. RAYMOND HALL</h3>
+
+
+<p>The pocket gophers of the species <i>Thomomys talpoides</i> in east-central
+Colorado have long been referred to the subspecies
+<i>Thomomys talpoides clusius</i> Coues with type locality at Bridger
+Pass, Wyoming. Recently, two subspecies, <i>T. t. attenuatus</i> and
+<i>T. t. rostralis</i> (see Hall and Montague, Univ. Kansas Publ., Mus.
+Nat. Hist., 5(3):25-32, February 28, 1951) were named from along
+the Wyoming-Colorado boundary with the result that the populations
+of <i>Thomomys talpoides</i> in east-central Colorado are separated
+from <i>T. t. clusius</i> of Wyoming by the geographic ranges now
+ascribed to the recently named <i>T. t. attenuatus</i> and <i>T. t. rostralis</i>.
+The subspecific identity of the animals from east-central Colorado
+thus is left in doubt. Examination of pertinent materials was made
+in the expectation that the names <i>Thomomys talpoides macrotis</i>
+F. W. Miller (Proc. Colorado Mus. Nat. Hist., 9:41, December 14,
+1930) and <i>Thomomys talpoides cheyennensis</i> Swenk (Missouri
+Valley Fauna, 4:5, March 1, 1941) would apply to the specimens,
+the identity of which is in doubt. This examination discloses instead,
+as set forth in more detail below, that neither of the two
+names mentioned immediately above does apply; the Coloradan
+specimens in question are referable to an heretofore unrecognized
+subspecies which may be named and described as follows:</p>
+
+<div class="blockquot">
+<p style="text-align: center"><b>Thomomys talpoides retrorsus</b> new subspecies</p>
+
+<div class="blockquot">
+<p><i>Thomomys clusius</i>, Warren, The Mammals of Colorado, G. P. Putnam's
+Sons, New York, p. 80, 1910; Cary, N. Amer. Fauna, 33:132,
+August 17, 1911.</p>
+
+<p><i>Thomomys talpoides clusius</i>, Bailey, N. Amer. Fauna, 39:100, November
+15, 1915; F. W. Miller, Proc. Colorado Mus. Nat. Hist., 9:41,
+December 14, 1930; Warren, The Mammals of Colorado, Univ. Oklahoma
+Press, Norman, p. 162, 1942.</p>
+</div>
+
+<p><i>Type.</i>&mdash;Male, subadult, skull and skin, No. 69840 Biological Surveys Collection,
+U. S. Nat. Hist.; from Flagler, Kit Carson County, Colorado; obtained
+on November 26, 1894, by Clark P. Streator; original No. 4460.</p>
+
+<p><i>Range.</i>&mdash;Western end of the Arkansas Divide in Colorado from eight miles
+south of Seibert westward to Colorado Springs<span class="pagenum"><a name="Page_84" id="Page_84">[Pg 84]</a></span></p>
+
+<p><i>Diagnosis.</i>&mdash;Size medium (see measurements); upper parts grayish brown;
+underparts lighter; skull small; tympanic bullae moderately inflated and
+angular anterolaterally; interpterygoid space narrowly U-shaped; pterygoid
+hamuli without transverse enlargement; nasals truncate posteriorly; premaxillary
+tongues projecting posteriorly behind nasals for distance of eight-tenths
+(0.5-1.1) of a millimeter.</p>
+
+<p><i>Comparisons.</i>&mdash;In comparison with <i>T. t. fossor</i> and <i>T. t. rostralis</i>, which
+occur farther west, selected differences of <i>T. t. retrorsus</i> are: lighter color;
+larger skull; more inflated tympanic bullae; greater relative (to length of skull)
+breadth across upper incisors, rostrum, and zygomata. The difference in
+color is greater in comparison with <i>fossor</i> than with <i>rostralis</i>.</p>
+
+<p>In comparison with <i>T. t. macrotis</i> (specimens from the type locality), <i>T. t.
+retrorsus</i> is indistinguishable in color, length of tail, and length of tooth-row,
+but averages smaller in all other measurements. There is no overlap in
+length of body, basilar length, zygomatic breadth, mastoidal breadth or
+length of nasals. The temporal ridges, which mark the limits of the temporal
+muscles, are straight as opposed to curved and are lower. The tympanic
+bullae are more angular anterolaterally in <i>T. t. retrorsus</i>.</p>
+
+<p>From <i>T. t. attenuatus</i> to the north, <i>T. t. retrorsus</i> differs in darker (more
+brown) color, consistently longer body, relatively (to length of skull) shorter
+rostrum and nasals. Linear measurements of the two latter structures and
+length of tail are approximately the same in the two subspecies but all other
+measurements average more in <i>T. t. retrorsus</i>. Also in the latter the temporal
+lines are approximately parallel instead of being bowed outward in their
+middle extent and instead of being more widely separated posteriorly than
+anteriorly.</p>
+
+<p>From <i>T. t. cheyennensis</i> to the northeast, <i>T. t. retrorsus</i> differs in slightly
+darker (more brownish) color, consistently shorter body and rostrum, usually
+a more narrowly V-shaped interpterygoid space, and smaller average dimensions
+of the skull, notably in mastoidal breadth and length of the rostrum.</p>
+</div>
+
+<p><i>Remarks.</i>&mdash;Miller's (Proc. Colorado Mus. Nat. Hist., 9:42, December
+14, 1930) mention of a specimen taken on November 9,
+1930, "near the head of Beaver Creek in extreme northeastern
+Elbert County" refers to the specimen, No. 2426 Colo. Mus. Nat.
+Hist., which is labeled as "8 mi. N. E. Agate, Elbert Co., Colo."
+Specimens from Colorado Springs, in the collection of the late E. R.
+Warren, have not been examined but the fact that Cary, Warren
+1942, and Bailey (see under synonymy above) each referred the
+specimens to <i>clusius</i> instead of to the darker <i>fossor</i> gives basis for
+tentatively referring the specimens to <i>T. t. retrorsus</i>.</p>
+
+<p>Grateful acknowledgment is made to those persons in charge of
+the mammal collections of the Denver Museum of Natural History
+and the Biological Surveys collection of mammals in the United
+States National Museum for permission to examine and report upon
+the material listed below (see specimens examined). The study<span class="pagenum"><a name="Page_85" id="Page_85">[Pg 85]</a></span>
+here reported upon was aided also by a contract between the Office
+of Naval Research, Department of the Navy, and the University of
+Kansas (NR 161-791). Essential comparative materials were obtained
+with assistance from the Kansas University Endowment
+Association.</p>
+
+<div class="blockquot">
+<p><i>Measurements.</i>&mdash;Measurements of the type, a male, are followed by the
+measurements of three adult females (69835, 69839 and 69838) from the type
+locality. Total length, 216, 207, 210, 200; length of tail, 59, 58, 64, 56; length
+of hind foot, 28, 28, 28, 26; basilar length of skull, 32.8, 32.2, 32.3, 30.8;
+zygomatic breadth, 23.1, 22.5, &mdash;&mdash;, 20.5; least interorbital breadth, 6.0, 6.7,
+6.2, 6.1; mastoidal breadth, 18.2, 18.8, 17.7, 17.7; length of nasals, 13.0, 13.7,
+13.9, 14.0; breadth of rostrum, 7.6, 7.9, 7.4, 7.2; length of rostrum, 14.8, 15.6,
+15.7, 16.0; alveolar length of maxillary tooth-row, 7.6, 7.2, 7.7, 7.6.</p>
+
+<p><i>Specimens examined.</i>&mdash;Total number, 13, all from Colorado, as follows:
+<i>Elbert County</i> (Colorado Mus. Nat. Hist. [= Denver Mus. Nat. Hist.]): Bijou
+Creek, "near El Paso Co. line", 3; 8 mi. NE Elbert, 1. <i>Lincoln Co.</i> (U. S.
+Biol. Surv. Coll.): Limon, 1. <i>Kit Carson Co.</i> (U. S. Biol. Surv. Coll.):
+Flagler, 7; 8 mi. S Seibert, 1.</p></div>
+
+<p> &nbsp; &nbsp; <i>Transmitted, February 28, 1951.</i></p>
+
+<h5>23-7439</h5>
+
+
+
+
+
+
+
+
+<pre>
+
+
+
+
+
+End of the Project Gutenberg EBook of A New Pocket Gopher (Genus Thomomys),
+from Eastern Colorado, by E. Raymond Hall
+
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+++ b/34836.txt
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+The Project Gutenberg EBook of A New Pocket Gopher (Genus Thomomys), from
+Eastern Colorado, by E. Raymond Hall
+
+This eBook is for the use of anyone anywhere at no cost and with
+almost no restrictions whatsoever. You may copy it, give it away or
+re-use it under the terms of the Project Gutenberg License included
+with this eBook or online at www.gutenberg.org
+
+
+Title: A New Pocket Gopher (Genus Thomomys), from Eastern Colorado
+
+Author: E. Raymond Hall
+
+Release Date: January 4, 2011 [EBook #34836]
+
+Language: English
+
+Character set encoding: ASCII
+
+*** START OF THIS PROJECT GUTENBERG EBOOK A NEW POCKET GOPHER (GENUS ***
+
+
+
+
+Produced by Chris Curnow, Joseph Cooper and the Online
+Distributed Proofreading Team at http://www.pgdp.net
+
+
+
+
+
+
+
+
+
+ A New Pocket Gopher (Genus Thomomys)
+ from Eastern Colorado
+
+ BY
+
+ E. RAYMOND HALL
+
+
+ University of Kansas Publications
+ Museum of Natural History
+
+ Volume 5, No. 8, pp. 81-85
+ October 1, 1951
+
+
+ University of Kansas
+ LAWRENCE
+ 1951
+
+
+
+
+ UNIVERSITY OF KANSAS PUBLICATIONS, MUSEUM OF NATURAL HISTORY
+
+ Editors: E. Raymond Hall, Chairman, A. Byron Leonard,
+ Edward H. Taylor, Robert W. Wilson
+
+ Volume 5, No. 8, pp. 81-85
+ October 1, 1951
+
+
+ UNIVERSITY OF KANSAS
+ Lawrence, Kansas
+
+
+ PRINTED BY
+ FERD VOILAND, JR., STATE PRINTER
+ TOPEKA, KANSAS
+ 1951
+
+ 23-7439
+
+
+
+
+ A New Pocket Gopher (Genus Thomomys)
+ from Eastern Colorado
+
+ By
+
+ E. RAYMOND HALL
+
+
+The pocket gophers of the species _Thomomys talpoides_ in east-central
+Colorado have long been referred to the subspecies _Thomomys talpoides
+clusius_ Coues with type locality at Bridger Pass, Wyoming. Recently,
+two subspecies, _T. t. attenuatus_ and _T. t. rostralis_ (see Hall and
+Montague, Univ. Kansas Publ., Mus. Nat. Hist., 5(3):25-32, February 28,
+1951) were named from along the Wyoming-Colorado boundary with the
+result that the populations of _Thomomys talpoides_ in east-central
+Colorado are separated from _T. t. clusius_ of Wyoming by the geographic
+ranges now ascribed to the recently named _T. t. attenuatus_ and _T. t.
+rostralis_. The subspecific identity of the animals from east-central
+Colorado thus is left in doubt. Examination of pertinent materials was
+made in the expectation that the names _Thomomys talpoides macrotis_ F.
+W. Miller (Proc. Colorado Mus. Nat. Hist., 9:41, December 14, 1930) and
+_Thomomys talpoides cheyennensis_ Swenk (Missouri Valley Fauna, 4:5,
+March 1, 1941) would apply to the specimens, the identity of which is in
+doubt. This examination discloses instead, as set forth in more detail
+below, that neither of the two names mentioned immediately above does
+apply; the Coloradan specimens in question are referable to an
+heretofore unrecognized subspecies which may be named and described as
+follows:
+
+ #Thomomys talpoides retrorsus# new subspecies
+
+ _Thomomys clusius_, Warren, The Mammals of Colorado,
+ G. P. Putnam's Sons, New York, p. 80, 1910;
+ Cary, N. Amer. Fauna, 33:132, August 17, 1911.
+
+ _Thomomys talpoides clusius_, Bailey, N. Amer. Fauna,
+ 39:100, November 15, 1915; F. W. Miller, Proc.
+ Colorado Mus. Nat. Hist., 9:41, December 14, 1930;
+ Warren, The Mammals of Colorado, Univ. Oklahoma
+ Press, Norman, p. 162, 1942.
+
+ _Type._--Male, subadult, skull and skin, No. 69840 Biological
+ Surveys Collection, U. S. Nat. Hist.; from Flagler, Kit Carson
+ County, Colorado; obtained on November 26, 1894, by Clark P.
+ Streator; original No. 4460.
+
+ _Range._--Western end of the Arkansas Divide in Colorado from
+ eight miles south of Seibert westward to Colorado Springs
+
+ _Diagnosis._--Size medium (see measurements); upper parts
+ grayish brown; underparts lighter; skull small; tympanic bullae
+ moderately inflated and angular anterolaterally; interpterygoid
+ space narrowly U-shaped; pterygoid hamuli without transverse
+ enlargement; nasals truncate posteriorly; premaxillary tongues
+ projecting posteriorly behind nasals for distance of
+ eight-tenths (0.5-1.1) of a millimeter.
+
+ _Comparisons._--In comparison with _T. t. fossor_ and _T. t.
+ rostralis_, which occur farther west, selected differences of
+ _T. t. retrorsus_ are: lighter color; larger skull; more
+ inflated tympanic bullae; greater relative (to length of skull)
+ breadth across upper incisors, rostrum, and zygomata. The
+ difference in color is greater in comparison with _fossor_ than
+ with _rostralis_.
+
+ In comparison with _T. t. macrotis_ (specimens from the type
+ locality), _T. t. retrorsus_ is indistinguishable in color,
+ length of tail, and length of tooth-row, but averages smaller in
+ all other measurements. There is no overlap in length of body,
+ basilar length, zygomatic breadth, mastoidal breadth or length
+ of nasals. The temporal ridges, which mark the limits of the
+ temporal muscles, are straight as opposed to curved and are
+ lower. The tympanic bullae are more angular anterolaterally in
+ _T. t. retrorsus_.
+
+ From _T. t. attenuatus_ to the north, _T. t. retrorsus_ differs
+ in darker (more brown) color, consistently longer body,
+ relatively (to length of skull) shorter rostrum and nasals.
+ Linear measurements of the two latter structures and length of
+ tail are approximately the same in the two subspecies but all
+ other measurements average more in _T. t. retrorsus_. Also in
+ the latter the temporal lines are approximately parallel instead
+ of being bowed outward in their middle extent and instead of
+ being more widely separated posteriorly than anteriorly.
+
+ From _T. t. cheyennensis_ to the northeast, _T. t. retrorsus_
+ differs in slightly darker (more brownish) color, consistently
+ shorter body and rostrum, usually a more narrowly V-shaped
+ interpterygoid space, and smaller average dimensions of the
+ skull, notably in mastoidal breadth and length of the rostrum.
+
+_Remarks._--Miller's (Proc. Colorado Mus. Nat. Hist., 9:42, December 14,
+1930) mention of a specimen taken on November 9, 1930, "near the head
+of Beaver Creek in extreme northeastern Elbert County" refers to the
+specimen, No. 2426 Colo. Mus. Nat. Hist., which is labeled as "8 mi. N.
+E. Agate, Elbert Co., Colo." Specimens from Colorado Springs, in the
+collection of the late E. R. Warren, have not been examined but the fact
+that Cary, Warren 1942, and Bailey (see under synonymy above) each
+referred the specimens to _clusius_ instead of to the darker _fossor_
+gives basis for tentatively referring the specimens to _T. t.
+retrorsus_.
+
+Grateful acknowledgment is made to those persons in charge of the mammal
+collections of the Denver Museum of Natural History and the Biological
+Surveys collection of mammals in the United States National Museum for
+permission to examine and report upon the material listed below (see
+specimens examined). The study here reported upon was aided also by a
+contract between the Office of Naval Research, Department of the Navy,
+and the University of Kansas (NR 161-791). Essential comparative
+materials were obtained with assistance from the Kansas University
+Endowment Association.
+
+ _Measurements._--Measurements of the type, a male, are followed
+ by the measurements of three adult females (69835, 69839 and
+ 69838) from the type locality. Total length, 216, 207, 210, 200;
+ length of tail, 59, 58, 64, 56; length of hind foot, 28, 28, 28,
+ 26; basilar length of skull, 32.8, 32.2, 32.3, 30.8; zygomatic
+ breadth, 23.1, 22.5, ----, 20.5; least interorbital breadth,
+ 6.0, 6.7, 6.2, 6.1; mastoidal breadth, 18.2, 18.8, 17.7, 17.7;
+ length of nasals, 13.0, 13.7, 13.9, 14.0; breadth of rostrum,
+ 7.6, 7.9, 7.4, 7.2; length of rostrum, 14.8, 15.6, 15.7, 16.0;
+ alveolar length of maxillary tooth-row, 7.6, 7.2, 7.7, 7.6.
+
+ _Specimens examined._--Total number, 13, all from Colorado, as
+ follows: _Elbert County_ (Colorado Mus. Nat. Hist. [= Denver
+ Mus. Nat. Hist.]): Bijou Creek, "near El Paso Co. line", 3;
+ 8 mi. NE Elbert, 1. _Lincoln Co._ (U. S. Biol. Surv. Coll.):
+ Limon, 1. _Kit Carson Co._ (U. S. Biol. Surv. Coll.): Flagler,
+ 7; 8 mi. S Seibert, 1.
+
+ _Transmitted, February 28, 1951._
+
+
+ 23-7439
+
+
+
+
+
+End of the Project Gutenberg EBook of A New Pocket Gopher (Genus Thomomys),
+from Eastern Colorado, by E. Raymond Hall
+
+*** END OF THIS PROJECT GUTENBERG EBOOK A NEW POCKET GOPHER (GENUS ***
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