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+The Project Gutenberg EBook of Food Habits of the Thrushes of the United
+States, by F. E. L. Beal
+
+This eBook is for the use of anyone anywhere at no cost and with
+almost no restrictions whatsoever. You may copy it, give it away or
+re-use it under the terms of the Project Gutenberg License included
+with this eBook or online at www.gutenberg.org
+
+
+Title: Food Habits of the Thrushes of the United States
+ USDA Bulletin 280
+
+Author: F. E. L. Beal
+
+Release Date: October 11, 2010 [EBook #33935]
+
+Language: English
+
+Character set encoding: UTF-8
+
+*** START OF THIS PROJECT GUTENBERG EBOOK FOOD HABITS OF THE THRUSHES ***
+
+
+
+
+Produced by Chris Curnow, Tom Cosmas, Joseph Cooper and
+the Online Distributed Proofreading Team at
+http://www.pgdp.ne
+
+
+
+
+
+
+
+
+
+
+ UNITED STATES DEPARTMENT OF AGRICULTURE
+
+ BULLETIN No. 280
+
+ Contribution from the Bureau of Biological Survey
+
+ HENRY W. HENSHAW, Chief
+
+ -----------------------------------------------------------------
+ Washington, D. C. PROFESSIONAL PAPER September 27, 1915
+ -----------------------------------------------------------------
+
+
+ FOOD HABITS OF THE THRUSHES
+ OF THE UNITED STATES
+
+ By F. E. L. Beal, _Assistant Biologist_.
+
+
+
+
+
+ CONTENTS.
+
+
+
+ Page.
+
+ Introduction 1
+
+ Townsend's solitaire 3
+
+ Wood thrush 5
+
+ Veery and willow thrush 9
+
+ Gray-cheeked and Bicknell's thrushes 11
+
+ Olive-backed and russet-backed thrushes 13
+
+ Hermit thrushes 18
+
+
+ [Illustration: logo]
+
+
+ WASHINGTON
+ GOVERNMENT PRINTING OFFICE
+ 1915
+
+
+
+
+
+ UNITED STATES DEPARTMENT OF AGRICULTURE
+
+[Illustration: shield] BULLETIN No. 280 [Illustration: shield]
+
+ Contribution from the Bureau of Biological Survey
+
+ HENRY W. HENSHAW, Chief
+
+ ------------------------------------------------------------------
+ Washington, D. C. PROFESSIONAL PAPER September 27, 1915
+ ------------------------------------------------------------------
+
+
+
+
+ FOOD HABITS OF THE THRUSHES OF THE
+ UNITED STATES.
+
+ By F. E. L. Beal, _Assistant Biologist_.
+
+
+
+
+INTRODUCTION.
+
+North American thrushes (Turdidæ) constitute a small but interesting
+group of birds, most of which are of retiring habits but noted as
+songsters. They consist of the birds commonly known as thrushes,
+robins, bluebirds, Townsend's solitaire, and the wheatears. The
+red-winged thrush of Europe (_Turdus musicus_) is accidental in
+Greenland, and the wheatears (_Saxicola œnanthe_ subspp.) are rarely
+found in the Western Hemisphere except in Arctic America. Within the
+limits of the United States are 11 species of thrushes, of which the
+following 6 are discussed in this bulletin: Townsend's solitaire
+(_Myadestes townsendi_), the wood thrush (_Hylocichla mustelina_), the
+veery and willow thrush (_Hylocichla fuscescens_ subspp.), the
+gray-cheeked and Bicknell's thrushes (_Hylocichla aliciæ_ subspp.),
+the olive-backed and russet-backed thrushes (_Hylocichla ustulata_
+subspp.), and the hermit thrushes (_Hylocichla guttata_ subspp.). An
+account of the food habits of the 5 species of robins and bluebirds
+appeared in Department Bulletin No. 171.
+
+As a group thrushes are plainly colored and seem to be especially
+adapted to thickly settled rural districts, as the shyest of them, with
+the exception of the solitaire, do not require any greater seclusion
+than that afforded by an acre or two of woodland or swamp.
+
+The thrushes are largely insectivorous, and also are fond of spiders,
+myriapods, sowbugs, snails, and angleworms. The vegetable portion of
+their diet consists mostly of berries and other small fruits. As a
+family thrushes can not be called clean feeders, for the food eaten
+often contains a considerable proportion of such matter as dead
+leaves, stems, and other parts of more or less decayed vegetation. It
+might be supposed that this was gathered from the ground with insects
+and other food, but investigation shows that much of it has a
+different origin. It was noticed that the setæ or spines of earthworms
+were a very common accompaniment of this decayed vegetation.
+Earthworms themselves are rather rarely found in stomachs, although
+some birds, as the robin, eat them freely. It is well known that the
+food of earthworms consists largely of partially decayed vegetable
+matter found in the soil. Hence it is probable that decayed vegetation
+found in the stomachs of thrushes is the food contained in the
+earthworms when they were swallowed. The tissues of worms are quickly
+digested, leaving the contents of their alimentary canals mixed with
+the hard indigestible setæ or spines.
+
+Thrushes of the genus _Hylocichla_ show a very pronounced taste for
+ants, and the average consumption of these insects by the five species
+is 12.65 per cent. Few birds other than woodpeckers show so strong a
+liking for this highly flavored food. Hymenoptera in general,
+including ants, bees, and wasps, are the second largest item of insect
+food. Lepidoptera (caterpillars) stand next as an article of thrush
+diet, while Orthoptera (grasshoppers), which are a favorite food with
+most birds, do not seem to appeal much to the thrushes.
+
+The thrushes are pronounced ground feeders, and may often be seen
+picking small fruit that has fallen to the ground. The vegetable
+portion of their food (40.72 per cent) is largely composed of fruit,
+which constitutes over 34 per cent of the total food. Of this 30.88
+per cent is made up of wild berries, which outweigh the domestic
+varieties with every species. In all, 94 species of wild fruits or
+berries were identified in the stomachs of these birds, although it is
+not always practicable to identify such material unless seeds or some
+other characteristic parts are present. As this is not often the case,
+a considerable portion of the stomach contents must be pronounced
+"fruit pulp" without further identification; thus probably many more
+species are eaten than are recorded. Moreover, in the case of some
+fruits, it is not possible to distinguish species by the seeds, so
+that many species go unrecognized except as to genus. Domestic fruits
+are eaten so sparingly by the thrushes here considered as to be of no
+economic importance.
+
+ Note.--This bulletin treats of the economic relations
+ and value to agriculture of the thrushes of the United States
+ other than robins and bluebirds. These two forms were discussed
+ in Department Bulletin No. 171, issued February 5, 1915.
+
+
+
+
+
+TOWNSEND'S SOLITAIRE
+
+(_Myadestes townsendi._)
+
+
+Townsend's solitaire, a bird of the far West, is a resident of high
+mountains and lonely gorges. It is partial to running streams and
+often builds its nest just above a rushing mountain torrent. It ranges
+from Alaska through the Sierras south to San Bernardino, Cal., and
+through the Rockies to Arizona and New Mexico, and occasionally
+farther east. The species is not evenly distributed over this region,
+but is restricted to such high mountainous portions as afford its
+favorite surroundings. As long as it retains these habits the bird
+will have little or no effect upon the products of husbandry, and its
+food can have only a scientific interest. The song of this species is
+said to be at times the finest of any of the thrush family.
+
+As this bird is comparatively rare in settled regions only 41 stomachs
+are available for determining the character of its food. The most
+southerly and easterly one was taken in Texas, the most westerly in
+California, and the most northerly in Wyoming. They are distributed
+through all the months of the year, although April and May are
+represented by but one each and December by but two. Every other month
+has three or more. An investigation based upon such limited material
+can be considered only as preliminary, but will serve to show some of
+the more important elements of the food. This was made up of 35.90 per
+cent of animal matter to 64.10 of vegetable.
+
+_Animal food._--The animal food consists of insects and spiders, with
+a few hair worms (_Gordius_) found in one stomach. These last may have
+been contained in the insects eaten. Among insects, beetles constitute
+the second largest item (10.74 per cent), but 5.89 per cent of these
+were the useful predatory ground beetles (Carabidæ). This is not a
+good showing, but too few stomachs have been examined to allow
+sweeping conclusions. As evidence that this can not be taken as a fair
+sample of the bird's food habits it may be stated that all of these
+beetles were taken in January and October. The one stomach collected
+in January contained 95 per cent of Carabidæ--the only animal food in
+it--and 93 per cent of the contents of one October stomach was made up
+of the same material. Evidently in these cases the bird had found a
+colony of the beetles and filled up with them. Had they constituted
+the usual diet of the species they would have appeared in other months
+and in more stomachs, but in smaller quantities. Other families of
+beetles are eaten so sparingly as to be of little importance.
+Scarabæidæ stand the next highest, but they amount to less than 2 per
+cent of the food.
+
+Lepidoptera (caterpillars) make the largest item in the food of
+_Myadestes_. Eaten much more regularly than beetles, they probably are
+a standard article of diet. They were found in the stomachs collected
+in every month of the year but four, and a greater number of stomachs
+would probably show them in every month. The one stomach taken in May
+contained the maximum (72 per cent). The total for the year is 12.95
+per cent. Ants are eaten to the extent of 4.71 per cent, while other
+Hymenoptera, as bees and wasps, make up less than half of 1 per cent.
+Diptera (flies) are represented by a mere trace in the stomachs.
+Observers who have seen this bird in its native haunts testify that it
+takes a considerable portion of its food on the wing. In view of this
+fact it seems curious that the two orders of insects most active on
+the wing (Hymenoptera and Diptera) should be so scantily represented
+in the food. Hymenoptera are a standard diet with flycatchers and
+would seem to be the natural food of any bird that feeds upon the
+wing.
+
+Hemiptera (bugs) were found to the extent of 3.51 per cent of the
+total food. All were contained in three stomachs taken in March, June,
+and July. In the July stomach four cicadas, or dog-day flies,
+constituted the whole contents. Grasshoppers amount to less than 1 per
+cent and all other insects to but a trifle. Spiders were eaten to the
+extent of 2.94 per cent of the food and were found in the stomachs
+taken in seven of the twelve months, and judging from their
+distribution they are eaten whenever available. A hair snake
+(_Gordius_) was found in one stomach. Following is a list of insects
+identified and the number of stomachs in which found:
+
+
+ COLEOPTERA.
+
+ _Amara erratica_ 1
+ _Aphodius_ sp 1
+ _Balaninus_ sp 1
+
+ HEMIPTERA.
+
+ _Platypedia putnami_ 1
+
+
+_Vegetable food._--The vegetable portion of the food of _Myadestes_ is
+64.10 per cent of the whole, and 58.70 per cent of this, or more than
+half the whole food, is classified as wild fruit or berries. These
+were found in stomachs collected in every month. From the even
+distribution of this food through the year and from the quantity eaten
+it is evidently a favorite article of diet. Nothing was found in any
+of the stomachs that could be identified as cultivated fruit, with the
+possible exception of a mass of fruit pulp found in one. A few seeds
+of poison ivy and sumac, with fragments of flowers and a few weed
+seeds, complete the vegetable food. Following is a list of fruits,
+seeds, etc., identified, and the number of stomachs in which found:
+
+
+ Rocky Mountain cedar (_Juniperus scopulorum_) 3
+ Western cedar (_Juniperus monospermum_) 1
+ Other cedars (_Juniperus_ sp.) 2
+ Hackberries (_Celtis occidentalis_) 1
+ Douglas hackberries (_Celtis douglasii_) 1
+ Service berries (_Amelanchier_ sp.) 1
+ Rose haws (_Rosa_ sp.) 2
+ Wild cherries (_Prunus_ sp.) 1
+ Sumac berries (_Rhus_ sp.) 1
+ Poison ivy (_Rhus toxicodendron_) 1
+ Waxwork (_Celastrus_ sp.) 1
+ Madrona berries (_Arbutus menziesii_) 5
+ Honeysuckle berries (_Lonicera_ sp.) 1
+ Elderberries (_Sambucus_ sp.) 1
+ Fruit not further identified 3
+
+
+_Summary._--With so small an amount of material it is not safe to draw
+general conclusions, but in the case of _Myadestes_ one point seems
+clear--the bird's favorite food is small wild fruit, and as long as
+this is abundant the bird will probably not attack cultivated
+varieties; but should any portion of the region occupied by the
+solitaire be cleared of its wild fruit and cultivated species be
+introduced these would likely be preyed upon. Under such conditions
+this bird, now perfectly harmless, might inflict considerable damage.
+
+
+
+
+WOOD THRUSH.
+
+(_Hylocichla mustelina._)
+
+
+The wood thrush is distributed over the eastern part of the United
+States wherever suitable conditions are found. It is a lover of open
+groves and bushy pastures, and may be found along little-traveled
+roads and near low bushy swamps. The bird is noted for its sweet song,
+and many country people who are well acquainted with its notes know
+little or nothing of the bird itself. Its favorite time for singing is
+in the early evening at the close of a sultry afternoon when a shower
+has cooled the air. As a rule, it does not nest in gardens or orchards
+and is seldom seen about farm buildings. It is strictly migratory, and
+the greater number pass out of the United States in winter, though a
+few remain in the Southern States. It usually migrates north in April
+or early May.
+
+For the investigation of the food habits of the wood thrush 171
+stomachs were available. One of these was collected in Florida in
+January and another in Alabama in February, and these two will be
+treated separately. The remaining 169 were collected from April to
+October, and are fairly well distributed over that time. The food
+consisted of 59.59 per cent of animal matter to 40.41 per cent of
+vegetable. The greatest quantity of animal food was eaten in April,
+the month of arrival from the south, and the least in October, the
+month of the return migration.
+
+_Animal food._--Beetles, collectively (20.40 per cent), constitute the
+largest item of animal food. Of these, 2.23 per cent are the
+predacious ground beetles (Carabidæ), generally considered useful. The
+remainder belong to several more or less harmful families, of which
+the May-beetle family (Scarabæidæ) amount to 10.17 per cent. Snout
+beetles, or weevils (Rhynchophora), are eaten to the extent of 2.16
+per cent only, and the wood-boring chick-beetles (Elateridæ) to 2.13
+per cent.
+
+Among the various species of these insects were noted the remains of
+the well-known Colorado potato beetle (_Leptinotarsa decemlineata_),
+in two stomachs, and _Coptocycla signifera_, also injurious to the
+potato, in one stomach. Remains of _Otiorhynchus ovatus_, a weevil
+destructive to strawberry plants, were found in two stomachs, and in
+one other a weevil, _Sphenophorus parvulus_, that injures the roots of
+grass. The well-known white grubs that attack grass roots and a host
+of other plants are the immature forms of many species of
+_Lachnosterna_, of several species of _Euphoria_ and of _Allorhina
+nitida_. Of these, remains of _Lachnosterna_ were found in 27 stomachs
+and of _Allorhina_ and _Euphoria_ in one each.
+
+
+ [Illustration:
+ B2084-73
+
+ Fig. 1.--Wood thrush (_Hylocichla mustelina_).]
+
+
+Lepidoptera (caterpillars) stand next to Coleoptera (beetles) in the
+animal diet of the wood thrush. Although eaten with a fair degree of
+regularity during every month of the bird's stay in the north, the
+most were taken in July (16.32 per cent). The average for the season
+is 9.42 per cent. Ants as an item of food are third in importance,
+though if other Hymenoptera were included the order would rank next to
+beetles. They seem to be a rather favorite food with all birds of the
+genus _Hylocichla_. With the wood thrush they begin with 18.12 per
+cent in April and gradually decrease through the summer and disappear
+in October. The total for the season is 8.89 per cent. Hymenoptera
+other than ants were eaten with great regularity (3.86 per cent)
+throughout the season, but not in large quantities. Diptera (flies)
+are eaten in small quantities and rather irregularly. Most of them
+were the long-legged crane flies (Tipulidæ), both in the adult and
+larval form. The total for the season is 2.70 per cent. Hemiptera
+(bugs) do not appear to be a favorite food, though a few were taken in
+all of the seven months except October. The average for the season is
+only 1.33 per cent. Orthoptera (grasshoppers) are eaten in small
+quantities until July, after which they form a fair percentage till
+September. The total consumption amounts to 2.28 per cent of the food.
+A few other insects make up a fraction of 1 per cent. Spiders and
+myriapods (thousand-legs) appear to be a favorite food with the wood
+thrush, constituting in April 20.94 per cent of the food, but
+gradually decreasing in quantity until September. The aggregate for
+the year is 8.49 per cent. A few sowbugs (isopods), snails, and
+earthworms (1.83 per cent) close the account of animal food.
+
+Following is a list of the insects identified in the stomachs of the
+wood thrush and the number of stomachs in which each was found:
+
+
+ HYMENOPTERA.
+
+ _Tiphia inornata_ 1
+
+ COLEOPTERA.
+
+ _Harpalus herbivagus_ 1
+ _Necrophorus tomentosus_ 1
+ _Philonthus lomatus_ 1
+ _Hister abbreviatus_ 1
+ _Hister depurator_ 1
+ _Hister americanus_ 2
+ _Ips quadriguttatus_ 1
+ _Melanotus americanus_ 1
+ _Corymbites cylindriformis_ 1
+ _Agrilus bilineatus_ 1
+ _Telephorus carolinus_ 1
+ _Onthophagus striatulus_ 1
+ _Onthophagus tuberculifrons_ 1
+ _Onthophagus_ sp 3
+ _Atænius_ sp 2
+ _Aphodius granarius_ 1
+ _Aphodius_ sp 1
+ _Dichelonycha testacea_ 1
+ _Dichelonycha_ sp 1
+ _Lachnosterna_ sp 27
+ _Ligyrus_ sp 1
+ _Allorhina nitida_ 1
+ _Euphoria fulgida_ 1
+ _Euphoria_ sp 2
+ _Chrysomela pulchra_ 1
+ _Leptinotarsa decemlineata_ 2
+ _Odontota_ sp 1
+ _Coptocycla signifera_ 1
+ _Coptocycla_ sp 1
+ _Anametus griseus_ 1
+ _Phyxelis rigidus_ 1
+ _Otiorhynchus ovatus_ 2
+ _Tanymecus confertus_ 1
+ _Pandeletejus hilaris_ 1
+ _Barypithes pellucidus_ 1
+ _Listronotus latiusculus_ 1
+ _Macrops_ sp 1
+ _Conotrachelus posticatus_ 2
+ _Acalles carinatus_ 1
+ _Balaninus_ sp 2
+ _Eupsalis minuta_ 1
+ _Sphenophorus parvulus_ 1
+
+ HEMIPTERA.
+
+ _Nezara hilaris_ 2
+
+ ORTHOPTERA.
+
+ _Diapheromera femorata_ 1
+
+ ISOPTERA.
+
+ _Termes flavipes_ 1
+
+
+_Vegetable food._--More than nine-tenths of the vegetable food of the
+wood thrush can be included in a single item--fruit. Cultivated fruit,
+or what was thought to be such, was found in stomachs taken from June
+to September, inclusive. It was eaten regularly and moderately, and
+the total for the season was 3.74 per cent of the whole food. Wild
+fruits or berries of 22 species were found in 72 stomachs, distributed
+through every month of the bird's stay at the north. Beginning with
+1.18 per cent in April, the quantity gradually increases to 87.17 per
+cent in October, when it makes more than five-sixths of the whole
+food. The average for the season is 33.51 per cent. In this
+investigation _Rubus_ seeds (blackberries or raspberries) are always
+reckoned as cultivated fruit, though probably most often wild. Besides
+fruit, a few seeds and rose haws were found, which with a little
+rubbish complete the vegetable food (40.41 per cent).
+
+Following is a list of fruits, seeds, etc., identified and the number
+of stomachs in which found:
+
+
+ Yew berries (_Taxus minor_) 1
+ False Solomon's seal (_Smilacina racemosa_) 1
+ Bayberries (_Myrica carolinensis_) 1
+ Mulberries (_Morus_ sp.) 10
+ Spiceberries (_Benzoin æstivale_) 5
+ Currants (_Ribes_ sp.) 1
+ Mountain ash (_Pyrus americanus_) 2
+ Service berries (_Amelanchier canadensis_) 2
+ Blackberries or raspberries (_Rubus_ sp.) 17
+ Rose haws (_Rosa_ sp.) 1
+ Wild black cherries (_Prunus serotina_) 1
+ Chokecherries (_Prunus virginiana_) 7
+ Domestic cherries (_Prunus cerasus_) 4
+ Croton (_Croton_ sp.) 1
+ American holly (_Ilex opaca_) 2
+ Woodbine berries (_Psedera quinquefolia_) 1
+ Frost grapes (_Vitis cordifolia_) 4
+ Wild sarsaparilla (_Aralia nudicaulis_) 1
+ Flowering dogwood (_Cornus florida_) 3
+ Rough-leaved cornel (_Cornus asperifolia_) 4
+ Dogwood (_Cornus_ sp.) 1
+ Black gum (_Nyssa sylvatica_) 1
+ Huckleberries (_Gaylussacia_ sp.) 1
+ Blueberries (_Vaccinium_ sp.) 6
+ French mulberry (_Callicarpa americana_) 1
+ Black elderberries (_Sambucus canadensis_) 1
+ Other elderberries (_Sambucus_ sp.) 3
+ Fruit pulp not further identified 12
+
+
+Of the two stomachs not included in the foregoing discussion, the one
+taken in Florida in January contained 93 per cent of wild fruit and 7
+per cent of weevils, wasps, and spiders; the one collected in Alabama
+in February was entirely filled with animal food, of which 88 per cent
+was caterpillars, 5 per cent May beetles, 6 per cent bugs, and 1 per
+cent spiders.
+
+_Summary._--The animal food of the wood thrush includes remarkably few
+useful insects and contains some very harmful ones, as the Colorado
+potato beetle and many of the Scarabæidæ, the larval forms of which
+are the well-known white grubs which are a pest to agriculture in
+preying upon roots of plants. The vegetable portion of the food
+contains a small quantity of cultivated fruit, but observation shows
+that the thrush is in the habit of picking up fallen fruit instead of
+taking it fresh from the tree. The eating of wild fruit has no
+economic interest except that it serves to distribute the seeds of
+many shrubs and trees. There is no occasion to discriminate against
+this bird in any way. It should be rigidly protected.
+
+
+
+
+VEERY AND WILLOW THRUSH.
+
+(_Hylocichla fuscescens fuscescens_ and _Hylocichla fuscescens
+salicicola_.)
+
+
+The veery is distributed over the eastern portion of the United
+States during migration and breeds in the Northern States as far
+south as Pennsylvania, and in New England and Canada. In winter
+it disappears almost entirely from the country, only a few remaining
+in Florida and perhaps in other Southern States. Its
+western representative is the willow thrush. Like other thrushes,
+birds of this species are shy and retiring in disposition, keeping for
+the most part in the shade of woods or bushy swamps, or building
+nests in a damp ravine with a brook gurgling past. They have been
+known, however, to visit orchards and sometimes gardens which are
+not kept too trim. It is thus evident that the food has little direct
+economic interest, as this bird does not come in contact with the
+farmer's crops.
+
+For investigating the food of the species 176 stomachs were available.
+They were collected during the seven months from April to
+October, and represent 18 States, the District of Columbia, and
+Canada. The food separates into 57.27 per cent of animal matter
+and 42.73 per cent of vegetable. The former consists mostly of
+remains of insects, and the latter of fruit.
+
+_Animal food._--Predacious ground beetles (Carabidæ) amount to
+0.82 per cent. They are evidently not a preferred food. Beetles in
+general comprise 14.67 per cent of the food, but no family or other
+group appears to be distinguished except the Carabidæ, which are
+conspicuous by their absence. Weevils, or snout beetles, amount
+to 2.49 per cent, and one stomach contained a specimen of the notorious
+plum curculio (_Conotrachelus nenuphar_). A number of other
+harmful beetles were noted, but none are so well known as the plum
+destroyer. Ants make up 10.35 per cent and are eaten with great
+regularity. Hymenoptera other than ants amount to only 3.26 per
+cent, but are eaten regularly throughout the season. Hemiptera
+(bugs) were eaten to a small extent (1.30 per cent) in the first four
+months, but they are not seen after July. Exactly the same may be
+said of Diptera, which total only 0.85 per cent.
+
+Lepidoptera (caterpillars) are, next to Hymenoptera, the favorite
+insect food. They were eaten in goodly quantities in every month
+except October. The average for the season is 11.91 per cent.
+Grasshoppers appear to some extent in every month except April, the
+greatest consumption taking place in October (24 per cent), but as
+only small numbers are eaten in the earlier months the aggregate for
+the year is only 4.91 per cent. A few other insects of various orders
+amount to 0.98 per cent. Spiders (6.34 per cent) are eaten regularly
+and constantly through the season, except that none were taken in
+October. A few sowbugs, snails, etc. (2.70 per cent), complete the
+quota of animal food. Following is a list of insects identified and
+the number of stomachs in which found:
+
+
+ HYMENOPTERA.
+
+ _Tiphia inornata_ 1
+
+ COLEOPTERA
+
+ _Elaphrus ruscarius_ 1
+ _Anisodactylus harrisi_ 1
+ _Anisodactylus_ sp 1
+ _Pterostichus lucublandus_ 1
+ _Hydrobius fuscipes_ 1
+ _Ips fasciata_ 1
+ _Byrrhus murinus_ 1
+ _Dolopius lateralis_ 2
+ _Limonius æger_ 1
+ _Corymbites cylindriformis_ 1
+ _Corymbites spinosus_ 1
+ _Corymbites tarsalis_ 1
+ _Corymbites hieroglyphicus_ 1
+ _Podabrus flavicollis_ 1
+ _Telephorus bilineatus_ 2
+ _Telephorus_ sp 1
+ _Onthophagus_ sp 2
+ _Atænius cognatus_ 1
+ _Aphodius_ sp 3
+ _Dichelonycha_ sp 2
+ _Serica sericea_ 1
+ _Lachnosterna hirticula_ 1
+ _Lachnosterna_ sp 13
+ _Chrysomela pulchra_ 3
+ _Chlamys plicata_ 1
+ _Typophorus canellus_ 1
+ _Graphops simplex_ 1
+ _Graphops_ sp 1
+ _Calligrapha philadelphica_ 1
+ _Œdionychis quercata_ 1
+ _Microrhopala vittata_ 1
+ _Hormorus undulatus_ 1
+ _Phyxelis rigidus_ 1
+ _Otiorhynchus ovatus_ 1
+ _Neoptochus adspersus_ 1
+ _Cercopeus chrysorrhœus_ 2
+ _Barypithes pellucidus_ 2
+ _Sitones_ sp 2
+ _Phytonomus nigrirostris_ 2
+ _Conotrachelus nenuphar_ 1
+ _Conotrachelus posticatus_ 1
+ _Tyloderma_ sp 1
+ _Monarthrum mali_ 1
+ _Xyloteres politus_ 1
+
+ DIPTERA.
+
+ _Bibio_ sp 1
+
+
+_Vegetable food._--The vegetable portion of the food of the species is
+made up of fruit, with a few seeds and a little miscellaneous matter
+more or less accidental. Fruit collectively amounts to 35.30 per cent,
+of which 12.14 per cent was thought to be of cultivated varieties and
+so recorded, while the remainder, 23.16 per cent, was quite certainly
+of wild species. This percentage of cultivated fruit is more than
+three times the record of the wood thrush, while the wild fruit eaten
+is correspondingly less, as the sum total of the fruit consumed is
+very nearly the same with both birds. From this percentage of domestic
+fruit one might infer that the veery was, or might be, a serious
+menace to fruit growing, but no such complaints have been heard, and
+it is probable that the species is not numerous enough to damage
+cultivated crops. A close inspection, however, of the fruit eating of
+the veery removes all doubts. The cultivated fruit, so called, was in
+every case either strawberries or _Rubus_ fruits, i. e., blackberries
+or raspberries, and as both of these grow wild and in abundance
+wherever the veery spends its summer, it is probable that all of the
+fruit eaten was taken from wild plants, though 12.14 per cent has been
+conventionally recorded as cultivated.
+
+Besides fruit, the veery eats a few seeds of grasses and weeds and a
+few of sumac, but none of the poisonous species were found in the
+stomachs. These seeds (7.25 per cent of the food) were eaten so
+irregularly as to suggest that they are merely a makeshift taken for
+want of something better. Rubbish (0.18 per cent), consisting of
+decayed wood, bits of leaves, plant stems, etc., completes the
+vegetable food.
+
+Following is a list of the items of vegetable food and the number of
+stomachs in which found:
+
+
+ Yew berries (_Taxus minor_) 1
+ Pigeon grass seed (_Chætochloa_ sp.) 1
+ Rush grass seed (_Sporobolus minor_) 1
+ False Solomon's seal (_Smilacina_ sp.) 1
+ Greenbrier berries (_Smilax_ sp.) 2
+ Hackberries (_Celtis occidentalis_) 1
+ Poke berries (_Phytolacca decandra_) 3
+ Spice berries (_Benzoin æstivale_) 2
+ Service berries (_Amelanchier canadensis_) 3
+ June berries (_Amelanchier_ sp.) 9
+ Mountain ash (_Pyrus americana_) 1
+ Crab apples (_Pyrus_ sp.) 1
+ Strawberries (_Fragaria_ sp.) 3
+ Blackberries or raspberries (_Rubus_ sp.) 8
+ Wild black cherries (_Prunus serotina_) 1
+ Bird cherries (_Prunus pennsylvanica_) 1
+ Chokecherries (_Prunus virginiana_) 1
+ Staghorn sumac (_Rhus hirta_) 2
+ Dwarf sumac (_Rhus copallina_) 1
+ Three-leaved sumac (_Rhus trilobata_) 1
+ Other sumac (_Rhus_ sp.) 1
+ American holly (_Ilex opaca_) 1
+ Woodbine berries (_Psedera quinquefolia_) 1
+ White cornel (_Cornus candidissima_) 2
+ Alternate-leaved cornel (_Cornus alternifolia_) 3
+ Rough-leaved cornel (_Cornus asperifolia_) 1
+ Dogwood berries (_Cornus_ sp.) 2
+ Sour gum berries (_Nyssa sylvatica_) 1
+ Huckleberries (_Gaylussacia_ sp.) 1
+ Blueberries (_Vaccinium_ sp.) 4
+ Snowberries (_Symphoricarpos racemosus_) 2
+ Black elderberries (_Sambucus canadensis_) 2
+ Red elderberries (_Sambucus pubens_) 4
+ Other elderberries (_Sambucus_ sp.) 3
+ Fruit pulp not further identified 4
+
+
+_Summary._--It is hardly necessary to make a summary of the
+food of this bird in order to bring out its good points, for it seems
+to have no others. The animal food includes less than 1 per cent of
+useful beetles, and the remainder is either harmful or neutral.
+In the matter of vegetable food there seems to be no chance for
+criticism, as nature evidently supplies all it needs. The bird has
+never been harmed, but has been held in high esteem for sentimental
+reasons; let it also be valued and protected for its economic worth.
+
+
+
+
+GRAY-CHEEKED AND BICKNELL'S THRUSHES.
+
+(_Hylocichla aliciæ aliciæ_ and _Hylocichla aliciæ bicknelli_.)
+
+
+The gray-cheeked thrush (_H. a. aliciæ_) is found in migration over
+all the Eastern States, but breeds farther north, beyond our limits.
+Bicknell's thrush (_H. a. bicknelli_), a closely related form, while
+having somewhat the same general range, breeds farther south and nests
+in the mountains of northern New York and New England. Both subspecies
+have the same general habits as other forms of the genus so far as
+haunts and choice of residence are concerned, but their far-northern
+range excludes them from coming into contact with cultivated crops.
+The species does not seem to be very abundant anywhere, and
+consequently only a few stomachs have been received for examination.
+In all they number but 111 and are very irregularly distributed in
+time. None were taken in August and only one in July and two in June.
+From so scanty and unevenly distributed material it is impossible to
+draw final conclusions, but we can get some idea as to the nature of
+the bird's food and its economic importance.
+
+The first analysis of the food gives 74.86 per cent of animal matter
+to 25.14 per cent of vegetable. This is the most animal food found in
+the stomachs of any bird of the genus _Hylocichla_ and the largest but
+two of any of the thrushes.
+
+_Animal food._--Beetles collectively amount to about one-third of all
+the food (33.32 per cent). Of these, 2.83 per cent are the useful
+Carabidæ. The rest belong to harmful families, such as the Scarabæidæ,
+Elateridæ, and the weevils, or snout beetles. Ants amount to 16.34 per
+cent and are eaten very regularly--the most in the early part of the
+season. Hymenoptera other than ants, as wasps and bees, were eaten to
+the extent of 5.60 per cent, and with the ants make 21.94 per cent,
+placing this food next in rank to beetles. As in the case of ants,
+most of the bees and wasps were eaten in the first three months of the
+season. No honey bees were found. Lepidoptera (caterpillars) were
+third in order of abundance (8.81 per cent). No special pest was
+discovered, but all caterpillars may be considered as harmful. A few
+grasshoppers were found in the stomachs taken in April and May, and
+more in those collected in September and October. They do not appear
+to be a favorite food and amount to only 1.72 per cent. Other insects,
+as flies, bugs, and a few others, collectively amount to 2.89 per
+cent. Among these, it is of interest to note in one stomach the
+remains of the famous seventeen-year locust (_Tibicen septemdecem_),
+rather large game for so small a bird. Spiders are freely eaten by the
+gray-cheeked thrush in spring, and sparingly in fall. For the season
+they constitute 5.77 per cent of the food. A few other animals, as
+crawfish, sowbugs, and angleworms (0.41 per cent), complete the animal
+food.
+
+Following is a list of the insects identified and the number of
+stomachs in which found:
+
+
+ HYMENOPTERA.
+
+ _Lophyrus_ sp 1
+ _Aphænogaster tennesseense_ 1
+
+ COLEOPTERA.
+
+ _Cychrus andrewsi_ 2
+ _Cychrus_ sp 2
+ _Dyschirius hispidus_ 1
+ _Hister sedecimstriatus_ 1
+ _Phelister vernus_ 1
+ _Epuræa rufa_ 3
+ _Stelidota 8-maculata_ 1
+ _Byrrhus murinus_ 1
+ _Eucinetus morio_ 1
+ _Monocrepidius vespertinus_ 1
+ _Agriotes limosus_ 1
+ _Corymbites signaticollis_ 1
+ _Podabrus flavicollis_ 1
+ _Telephorus bilineatus_ 1
+ _Onthophagus_ sp 1
+ _Atænius strigatus_ 1
+ _Atænius ovatulus_ 1
+ _Atænius_ sp 3
+ _Aphodius ruricola_ 1
+ _Aphodius inquinatus_ 3
+ _Aphodius_ sp 1
+ _Serica_ sp 1
+ _Lachnosterna_ sp 10
+ _Anomala_ sp 1
+ _Leptura sphæricollis_ 1
+ _Leptura mutabilis_ 1
+ _Chrysomela pulchra_ 4
+ _Blapstinus metallicus_ 1
+ _Helops micans_ 1
+ _Hormorus undulatus_ 1
+ _Otiorhynchus ovatus_ 1
+ _Cercopeus chrysorrhœus_ 2
+ _Pandeletejus hilaris_ 1
+ _Sitones_ sp 1
+ _Hylobius pales_ 1
+ _Desmoris constrictus_ 1
+ _Bagous sellatus_ 1
+ _Anthonomus sycophanta_ 1
+ _Conotrachelus posticatus_ 2
+ _Acalles clavatus_ 1
+ _Acalles_ sp 1
+ _Cryptorhynchus ferratus_ 1
+ _Sphenophorus melanocephalus_ 1
+
+ HEMIPTERA.
+
+ _Tibicen septendecem_ 1
+ _Nezara hilaris_ 1
+
+
+_Vegetable food._--A few _Rubus_ seeds were recorded as cultivated
+fruit, but they were found in only two stomachs and probably were
+wild, as the gray-cheeked thrush does not live where it is likely to
+come in contact with cultivated blackberries or raspberries. In any
+case they amount to only 0.15 per cent. Wild fruits of 18 different
+species (23.98 per cent) make up nearly one-fourth of the whole
+food--in fact, the vegetable food, other than wild fruit, is
+insignificant. Wild berries supplement the regular food, which
+consists of insects and spiders.
+
+The following list shows the fruits and seeds identified and the
+number of stomachs in which found:
+
+
+ False spikenard (_Smilacina racemosa_) 1
+ Greenbrier berries (_Smilax_ sp.) 2
+ Bayberries (_Myrica carolinensis_) 1
+ Poke berries (_Phytolacca decandra_) 2
+ Crab apples (_Pyrus_ sp.) 1
+ Wild black cherries (_Prunus serotina_) 5
+ Blackberries or raspberries (_Rubus_ sp.) 2
+ Sumac berries (_Rhus_ sp.) 1
+ Black-alder berries (_Ilex verticillata_) 1
+ Wild grapes (_Vitis_ sp.) 5
+ Wild sarsaparilla (_Aralia_ sp.) 1
+ Flowering dogwood (_Cornus florida_) 5
+ Rough-leaved dogwood (_Cornus asperifolia_) 2
+ White cornel (_Cornus candidissima_) 1
+ Dogwood (_Cornus_ sp.) 1
+ Sour gum (_Nyssa sylvatica_) 2
+ Black nightshade (_Solanum nigrum_) 1
+ Dockmackie (_Viburnum acerifolium_) 1
+ Arrowwood (_Viburnum_ sp.) 1
+ Elderberries (_Sambucus canadensis_) 3
+ Fruit not further identified 6
+
+
+_Summary._--In the food of the gray-cheeked thrush the only useful
+element is a small percentage (2.83) of useful beetles. The remainder
+of the animal food is composed of either harmful or neutral elements.
+The vegetable food, drawn entirely from nature's great storehouse,
+contains no product of human industry, either of grain or fruit.
+Whatever the sentimental reasons for protecting this bird, the
+economic ones are equally valid.
+
+
+
+
+OLIVE-BACKED AND RUSSET-BACKED THRUSHES.
+
+(_Hylocichla ustulata swainsoni_ and _Hylocichla ustulata ustulata_.)
+
+
+The olive-backed thrush and its relative, the russet-backed, occupy
+the whole of the United States at some time during the year. The
+olive-back breeds north of our northern border, except in the higher
+mountains, and the russet-back on the Pacific coast nests as far south
+as southern California. The habits of birds of this species resemble
+those of others of the genus in living in swamps and woodlands rather
+than in gardens and orchards. The russet-back on the Pacific coast,
+however, seems to have become quite domestic, and wherever a stream
+runs through or past an orchard or garden, or the orchard is near
+thick chaparral, this bird is sure to be found taking its toll of the
+fruit and rearing its young in the thicket beside the stream. During
+the cherry season it takes a liberal share of the fruit, but its
+young, then in the nest, are fed almost entirely on insects. The
+eastern subspecies, on the contrary, does not come in contact with
+domestic fruit or any other product of husbandry. A great number of
+the subspecies nest far north of the region of fruit raising.
+
+For this investigation 403 stomachs of the olive-backed thrush were
+available, collected in 25 States, the District of Columbia, and
+Canada. Florida, Louisiana, and Texas represent the most southern
+collections and New Brunswick, Ontario, and Northwest Territory the
+most northern. In California 157 stomachs were obtained, which, with
+those taken in Oregon and Washington, fairly represent the Pacific
+coast region. The whole collection was fairly well distributed over
+the nine months from March to November. The food consisted of 63.52
+per cent of animal matter to 36.48 per cent of vegetable.
+
+_Animal food._--Beetles of all kinds amount to 16.29 per cent. Of
+these 3.14 per cent are the useful Carabidæ. The others belong to
+harmful or neutral families. Weevils or snout-beetles (Rhynchophora)
+amount to 5.29 per cent, a high percentage for such insects. One
+Colorado potato beetle (_Leptinotarsa decemlineata_) was found in a
+stomach taken on Long Island. Hymenoptera collectively aggregate 21.50
+per cent. Of these, 15.20 per cent are ants--a favorite food of
+_Hylocichla_. The remainder (6.30 per cent) were wild bees and wasps.
+No honeybees were found. Caterpillars, which rank next in importance
+in the food of the olive-back, form a good percentage of the food of
+every month represented and aggregate 10.30 per cent for the season.
+
+Grasshoppers are not an important element in the food of thrushes, as
+they chiefly inhabit open areas, while _Hylocichla_ prefers thick damp
+cover, where grasshoppers are not found. An inspection of the record
+shows that most of the orthopterous food taken by the olive-back
+consists of crickets, whose habits are widely different from those of
+grasshoppers, and which are found under stones, old logs, or dead
+herbage. The greatest quantity is taken in August and September. The
+average for the season is 2.42 per cent.
+
+Diptera (flies) reach the rather surprisingly large figure of 6.23 per
+cent. These insects are usually not eaten to any great extent except
+by flycatchers and swallows, which take their food upon the wing. The
+flies eaten by the olive-back are mostly crane flies (Tipulidæ) or
+March flies (_Bibio_), both in the adult and larval state. Crane flies
+are slow of wing and frequent shady places. The larvæ of both groups
+are developed in moist ground, and often in colonies of several
+hundred. With these habits it is not surprising that they fall an easy
+prey to the thrushes.
+
+Hemiptera (bugs), a small but rather constant element of the food,
+were found in the stomachs collected every month, and in July reach
+11.11 per cent. They were of the families of stinkbugs (Pentatomidæ),
+shield bugs (Scutelleridæ), tree hoppers (Membracidæ), leaf hoppers
+(Jassidæ), and cicadas. Some scales were found in one stomach. The
+total for the season is 3.76 per cent. A few insects not included in
+any of the foregoing categories make up 0.48 per cent of the food.
+Spiders, with a few millipeds, amount to 2.22 per cent, the lowest
+figure for this item of any bird of the genus _Hylocichla_. Snails,
+sowbugs, angleworms, etc. (0.32 per cent), complete the animal food.
+
+Following is a list of insects identified and the number of stomachs
+in which found:
+
+
+ HYMENOPTERA.
+
+ _Camponotus pennsylvanicus_ 1
+ _Tiphia inornata_ 1
+
+ COLEOPTERA.
+
+ _Cychrus nitidicollis_ 1
+ _Cychrus stenostomus_ 1
+ _Notiophilus æneus_ 1
+ _Pterostichus sayi_ 1
+ _Pterotichus lustrans_ 1
+ _Amara interstitialis_ 1
+ _Triæna longula_ 1
+ _Agonoderus pallipes_ 1
+ _Silpha ramosa_ 1
+ _Staphylinus cinnamopterus_ 1
+ _Tachyporus californicus_ 1
+ _Chilocorus orbus_ 1
+ _Scymnus_ sp 1
+ _Hister americanus_ 1
+ _Ips quadriguttatus_ 4
+ _Cytilus sericeus_ 1
+ _Agriotes stabilis_ 1
+ _Podabrus flavicollis_ 2
+ _Podabrus modestus_ 2
+ _Silis lutea_ 1
+ _Telephorus carolinus_ 1
+ _Telephorus bilineatus_ 5
+ _Telephorus divisus_ 2
+ _Onthophagus hecate_ 1
+ _Onthophagus striatulus_ 1
+ _Onthophagus tuberculifrons_ 2
+ _Onthophagus_ sp 4
+ _Atænius abditus_ 1
+ _Aphodius hamatus_ 1
+ _Aphodius fimetarius_ 6
+ _Aphodius inquinatus_ 7
+ _Aphodius_ sp 6
+ _Geotrupes_ sp 1
+ _Dichelonycha elongata_ 2
+ _Lachnosterna hirticula_ 1
+ _Lachnosterna_ sp 12
+ _Anomala undulata_ 1
+ _Anomala_ sp 1
+ _Euphoria fulgida_ 1
+ _Donacia emarginata_ 1
+ _Hæmonia nigricornis_ 1
+ _Syneta pallida_ 1
+ _Leptinotarsa decemlineata_ 1
+ _Gastroidea_ sp 1
+ _Galerucella decora_ 1
+ _Diabrotica soror_ 1
+ _Diabrotica_ sp 1
+ _Gonioctena pallida_ 1
+ _Luperodes bivittatus_ 1
+ _Opatrinus notus_ 1
+ _Blapstinus metallicus_ 1
+ _Blapstinus mæstus_ 1
+ _Blapstinus_ sp 1
+ _Otiorhynchus ovatus_ 1
+ _Thinoxenus_ sp 1
+ _Cercopeus chrysorrhæus_ 1
+ _Barypithes pellucidus_ 1
+ _Sitones flavescens_ 1
+ _Sitones_ sp 1
+ _Phytonomus punctatus_ 2
+ _Pachylobius picivorus_ 1
+ _Conotrachelus posticatus_ 1
+ _Micromastus elegans_ 1
+ _Acalles clavatus_ 1
+ _Cryptorhynchus bisignatus_ 1
+ _Rhinoncus pyrrhopus_ 1
+ _Balaninus_ sp 3
+ _Sphenophorus parvulus_ 1
+ _Sphenophorus_ sp 1
+ _Scolytus muticus_ 1
+
+ LEPIDOPTERA.
+
+ _Edema albifrons_ 1
+
+ TRICHOPTERA.
+
+ _Phryganea californica_ 1
+
+ HEMIPTERA.
+
+ _Myodocha serripes_ 1
+ _Sinea diadema_ 1
+
+
+This list of insects contains a considerable number of injurious
+species and some that at various times and places have become decided
+pests. Such are the Colorado potato beetle (_Leptinotarsa
+decemlineata_), the spotted squash beetle (_Diabrotica soror_), the
+cloverleaf weevil (_Phytonomus punctatus_), and the various species of
+_Lachnosterna_, the parent of the destructive white grubs. Many others
+are plant feeders and may increase to such an extent as to inflict
+great damage upon agriculture.
+
+_Vegetable food._--The vegetable food of the olive-backed thrush
+consists of small fruit. The bird has a weak bill and can not break
+through the tough skin of the larger kinds. In the cherry orchards of
+California the writer many times observed the western subspecies of
+this bird, the russet-back, on the ground pecking at cherries that had
+been bitten open and dropped by linnets and grosbeaks. Blackberries
+and raspberries have a very delicate skin and are successfully managed
+by weak-billed birds, so that all the records of domestic fruit eaten
+by the eastern form relate to these berries, and it is probable that
+in most cases the fruit was not cultivated. The total of cultivated
+fruit for the season is 12.63 per cent of the whole food, but if we
+consider the eastern subspecies alone this item would practically
+disappear. Wild fruit (19.73 per cent) is eaten regularly and in a
+goodly quantity in every month after April. Weed seeds and a few
+miscellaneous items of vegetable food (4.04 per cent) close the
+account.
+
+Following is a list of vegetable foods so far as identified and the
+number of stomachs in which found.
+
+
+ White cedar seeds (_Thuja occidentalis_) 1
+ Red cedar berries (_Juniperus communis_) 2
+ False Solomon's seal (_Smilacina trifolia_) 3
+ Greenbrier (_Smilax tamnifolia_) 1
+ Cat brier (_Smilax_ sp.) 1
+ Hackberry (_Celtis occidentalis_) 3
+ Mulberry (_Morus_ sp.) 2
+ Fig (_Ficus_ sp.) 3
+ Pale persicaria (_Polygonum lapathifolium_) 1
+ Poke berries (_Phytolacca decandra_) 9
+ Mountain ash (_Pyrus americana_) 1
+ Service berries (_Amelanchier_ sp.) 1
+ Blackberries or raspberries (_Rubus_ sp.) 67
+ Rose haws (_Rosa_ sp.) 1
+ Wild black cherries (_Prunus serotina_) 15
+ Bird cherries (_Prunus Pennsylvanica_) 2
+ Domestic cherries (_Prunus cerasus_) 29
+ Domestic plum (_Prunus domestica_) 2
+ Apricot (_Prunus armeniaca_) 3
+ Filaree (_Erodium_ sp.) 1
+ Poison oak (_Rhus diversiloba_) 4
+ Staghorn sumac (_Rhus hirta_) 2
+ Dwarf sumac (_Rhus copallina_) 3
+ Other sumac (_Rhus_ sp.) 4
+ Pepper tree (_Schinus molle_) 1
+ American holly (_Ilex opaca_) 1
+ Black alder (_Ilex verticillata_) 1
+ Coffee berries (_Rhamnus californicus_) 3
+ Woodbine (_Psedera quinquefolia_) 10
+ Frost grape (_Vitis cordifolia_) 6
+ Spikenard (_Aralia racemosa_) 2
+ Flowering dogwood (_Cornus florida_) 7
+ Kinnikinnik (_Cornus amomum_) 2
+ Red osier (_Cornus stolonifera_) 1
+ Panicled cornel (_Cornus paniculata_) 3
+ Dogwood unidentified (_Cornus_ sp.) 6
+ Huckleberries (_Gaylussacia_ sp.) 1
+ Three-flowered nightshade (_Solanum triflorum_) 1
+ Nightshade unidentified (_Solanum_ sp.) 8
+ Black twinberries (_Lonicera involucrata_) 2
+ Honeysuckle berries (_Lonicera_ sp.) 2
+ Snowberries (_Symphoricarpos racemosus_) 2
+ Dockmackie (_Viburnum acerifolium_) 1
+ Arrowwood (_Viburnum_ sp.) 1
+ Black elderberries (_Sambucus canadensis_) 6
+ Red elderberries (_Sambucus pubens_) 5
+ Blue elderberries (_Sambucus glauca_) 15
+ Tarweed (_Madia_ sp.) 1
+ Fruit pulp not further identified 17
+
+
+_Food of young of russet-backed thrush._--Before concluding the
+discussion of this species it will be of interest to note the results
+obtained from an investigation of stomachs of 25 nestlings of the
+russet-back taken in June and July when the birds were from two to
+eleven days old. These were from eight broods, ranging from three to
+five nestlings to the brood. The percentage of animal food of the
+young (92.60 per cent) is considerably higher than that of the parent
+birds.
+
+The distribution of the animal food is as follows: Caterpillars were
+found in every stomach but seven and aggregated nearly 27 per cent;
+beetles, including the useful Carabidæ (7.7 per cent), are irregularly
+distributed to the extent of 22 per cent; other more or less harmful
+species included five families of (Hemiptera) bugs, 13.8 per cent,
+viz, stinkbugs, leaf hoppers, tree hoppers, shield bugs, and cicadas;
+ants and a few other Hymenoptera amount to 12 per cent, and spiders
+the same. These latter were mostly harvestmen or daddy longlegs
+(Phalangidæ). The remainder (6 per cent) included a few miscellaneous
+insects. Only three stomachs contained remains of grasshoppers.
+Carabid beetles were eaten by the young birds to the extent of 7.7 per
+cent, which is more than three times the amount eaten by the adults, a
+remarkable fact when is considered that these insects are very hard
+shelled, thus seemingly unsuited for young birds.
+
+The vegetable food consisted of fruit (6.8 per cent), mainly
+blackberries or raspberries, found in 11 stomachs, and twinberries in
+1, and two or three other items, including a seed of filaree and some
+rubbish. From the irregular variety of food in the different stomachs,
+it would seem that the parents make little selection, but fill the
+gaping mouths of their young with the nearest obtainable supply.
+
+In addition to the examination of stomach contents of nestlings two
+nests were carefully and regularly watched, and from these it was
+determined that the parent birds fed each nestling 48 times in 14
+hours of daylight. This means 144 feedings as a day's work for the
+parents for a brood of three nestlings, and that each stomach was
+filled to its full capacity several times daily, an illustration that
+the digestion and assimilation of birds, especially the young, are
+constant and very rapid. Experiments in raising young birds have
+proved that they thrive best when fed small quantities at short
+intervals rather than greater quantities at longer periods. Aside from
+the insects consumed by the parents, a brood of three young birds will
+thus each require the destruction of at least 144 insects in a day and
+probably a very much greater number.
+
+_Summary._--In a résumé of the food of the olive-backed and
+russet-backed thrushes one is impressed with the fact that they come
+in contact with the products of industry but rarely. The olive-back's
+food habits infringe upon the dominion of man but little. The bird
+lives among men, but not with them. The western form, the russet-back,
+comes more into relations with the cultivated products because it
+visits orchards and partakes freely of the fruit. Even then the damage
+is slight, as much of the fruit eaten is that fallen to the ground.
+Moreover, while the adult bird is feeding upon fruit a nestful of
+young are being reared upon insects which must be largely taken from
+the orchard, thus not only squaring the account but probably
+overbalancing it in favor of the farmer.
+
+
+
+
+HERMIT THRUSHES.
+
+(_Hylocichla guttata_ subspp.)
+
+
+The hermit thrush of the subspecies _H. g. pallasi_ inhabits the
+Eastern States in winter as far north as Massachusetts and breeds from
+the mountains of Maryland and Pennsylvania and from northern Michigan
+and central Minnesota northward to Alaska. Several other subspecies
+occupy the Pacific coast region in suitable localities--that is, in
+the higher and more wooded sections, as this bird, like all of the
+genus _Hylocichla_, does not live in treeless or arid regions. In the
+East the bird is a late fall migrant and may often be seen sitting
+silent and alone on a branch in the forest in late October or even in
+November, when the great army of migrants have passed on to the South.
+While a beautiful songster, the species is so quiet and unobtrusive
+that by sight it is entirely unknown to many.
+
+Inquiry into the food habits of this bird covered 551 stomachs,
+collected in 29 States, the District of Columbia, and Canada, and
+representing every month of the year, though all the stomachs taken in
+winter were collected in the Southern States, the District of
+Columbia, and California. In the primary analysis the food was found
+to consist of 64.51 per cent of animal matter to 35.49 per cent of
+vegetable. The former is mostly composed of insects with some spiders,
+while the latter is largely fruit, chiefly wild species.
+
+_Animal food._--Beetles constitute 15.13 per cent of the food. Of
+these 2.98 per cent are of the useful family, Carabidæ. The remainder
+are mostly harmful. Scarabæidæ, the larvæ of which are the white grubs
+that destroy the roots of so many plants, were eaten to the extent of
+3.44 per cent. Snout beetles, among the most harmful of insects, were
+taken to the extent of 3.13 per cent. Among these was the notorious
+plum curculio (_Conotrachelus nenuphar_) found in two stomachs taken
+in the District of Columbia in April of different years. Two other
+species of the same genus also were found, as well as the clover
+weevil (_Epicærus imbricatus_). The Colorado potato beetle
+(_Leptinotarsa decemlineata_) and the striped squash beetle
+(_Diabrotica vittata_), with a number of other species of less
+notoriety, were found in several stomachs. Thus, in spite of the
+bird's retiring habits, it comes in contact with some of the pests of
+cultivation.
+
+
+ [Illustration:
+ B2085-73
+
+ Fig. 2.--Hermit thrush (_Hylocichla guttata_).]
+
+
+The ants destroyed--12.46 per cent of the food--keep up the reputation
+of thrushes as ant eaters. They were taken constantly every month,
+with the greatest number from May to September; a falling off in July
+is partly accounted for by the fact that more fruit is taken in that
+month. Other Hymenoptera (bees and wasps) were eaten to the extent of
+5.41 per cent, a surprising amount for a bird that feeds so largely
+upon the ground, as these insects are usually of fleet wing and live
+in sunshine and open air.
+
+Caterpillars, eaten in every month and mostly in goodly quantities,
+appear to be a favorite food of the hermit thrush. December is the
+month of least consumption (2.75 per cent), while the most were eaten
+in June (17.08 per cent). The average for the year is 9.54 per cent.
+Hemiptera (bugs) seem to be eaten whenever found, as they appear in
+the food of every month, but rather irregularly and not in large
+quantities. The greatest consumption was in June (9.17 per cent), but
+July, September, and December show the least (less than 1 per cent).
+The total for the year is 3.63 per cent. Of the six families
+represented, the Pentatomidæ, or stink bugs, predominate. These highly
+flavored insects are eaten by most insectivorous birds often, but
+usually in small quantities.
+
+Diptera (flies) comprise 3.02 per cent of the food of the hermit
+thrush. The record shows, however, that nearly all of them are either
+crane flies (Tipulidæ) and their eggs and larvæ, or March flies
+(_Bibio_) and their larvæ. Over 150 of the latter were found in one
+stomach. Both of these families of flies lay their eggs in the ground,
+which accounts for their consumption by ground-feeding birds.
+Orthoptera (grasshoppers and crickets) are eaten by the hermit thrush
+to the extent of 6.32 per cent of its food. While this figure is not
+remarkable, it is the highest for any of the genus. These birds are
+fond of dark moist nooks among trees and bushes and do not feed
+extensively in those dry sunshiny places so much frequented by
+grasshoppers. A close inspection of the food record shows that the
+Orthoptera eaten by the thrushes are mostly crickets, which live in
+shadier and moister places than those where grasshoppers abound. A few
+miscellaneous insects (0.27 per cent) close the insect account.
+Spiders and myriapods (7.47 per cent) seem to constitute a very
+acceptable article of diet, as they amount to a considerable
+percentage in nearly every month, and in May rise to 20.79 per cent. A
+few miscellaneous animals, as sowbugs, snails, and angleworms, make up
+the balance of the animal food (1.26 per cent).
+
+Following is a list of insects so far as identified and the number of
+stomachs in which found:
+
+
+ HYMENOPTERA.
+
+ _Tiphia inornata_ 2
+
+ COLEOPTERA.
+
+ _Elaphrus_ sp 1
+ _Notiophilus semistriatus_ 1
+ _Scarites subterraneus_ 1
+ _Dyschirius pumilis_ 1
+ _Pterostichus patruelis_ 1
+ _Pterostichus_ sp 1
+ _Amara_ sp 1
+ _Chlænius pennsylvanicus_ 2
+ _Stenolophus_ sp 1
+ _Anisodactylus agilis_ 1
+ _Tropisternus limbalis_ 2
+ _Hydrocharis obtusatus_ 1
+ _Sphæridium lecontei_ 1
+ _Ptomaphagus consobrinus_ 1
+ _Anisotoma valida_ 1
+ _Megilla maculata_ 1
+ _Anatis 15-maculata_ 1
+ _Psyllobora tædata_ 1
+ _Brachycantha ursina_ 1
+ _Endomychus biguttatus_ 1
+ _Cryptophagus_ sp 1
+ _Hister marginicollis_ 1
+ _Hister americanus_ 1
+ _Saprinus fimbriatus_ 1
+ _Carpophilus hemipterus_ 1
+ _Perthalycra murrayi_ 1
+ _Ips quadriguttatus_ 3
+ _Cytilus sericeus_ 2
+ _Cytilus_ sp 1
+ _Byrrhus kirbyi_ 1
+ _Byrrhus cyclophorus_ 1
+ _Cryptohypnus bicolor_ 2
+ _Drasterius dorsalis_ 1
+ _Dolopius lateralis_ 1
+ _Melanotus_ sp 2
+ _Podabrus tomentosus_ 1
+ _Canthon_ sp 1
+ _Onthophagus tuberculifrons_ 1
+ _Onthophagus_ sp 3
+ _Ægialia lacustris_ 1
+ _Rhyssemus scaber_ 1
+ _Atænius abditus_ 1
+ _Atænius cognatus_ 1
+ _Atænius_ sp 1
+ _Aphodius fimetarius_ 11
+ _Aphodius granarius_ 1
+ _Aphodius rugifrons_ 1
+ _Aphodius inquinatus_ 9
+ _Aphodius pardalis_ 1
+ _Aphodius prodromus_ 4
+ _Aphodius crassiusculus_ 1
+ _Aphodius_ sp 11
+ _Geotrupes semipunctata_ 1
+ _Dichelonycha_ sp 1
+ _Lachnosterna_ sp 17
+ _Chrysomela pulchra_ 3
+ _Lema nigrovittata_ 1
+ _Chlamys plicata_ 1
+ _Myochrous denticollis_ 2
+ _Xanthonia 10-notata_ 1
+ _Calligrapha scalaris_ 1
+ _Leptinotarsa decemlineata_ 1
+ _Phædon viridis_ 1
+ _Diabrotica vittata_ 1
+ _Odontota rubra_ 1
+ _Odontota_ sp 1
+ _Haltica torquata_ 1
+ _Crepidodera helxines_ 1
+ _Syneta ferruginea_ 1
+ _Systena elongata_ 1
+ _Chætocnema pulicaria_ 1
+ _Psylliodes punctulata_ 1
+ _Chelymorpha cribraria_ 1
+ _Opatrinus notus_ 1
+ _Opatrinus aciculatus_ 1
+ _Blapstinus metallicus_ 1
+ _Blapstinus rufipes_ 1
+ _Salpingus virescens_ 1
+ _Anthicus pubescens_ 1
+ _Notoxus monodon_ 1
+ _Notoxus denudatum_ 1
+ _Notoxus_ sp 1
+ _Attelabus rhois_ 1
+ _Rhigopsis effracta_ 1
+ _Cercopeus chrysorrhœus_ 4
+ _Pandetetejus hilaris_ 1
+ _Barypithes pellucidus_ 1
+ _Sitones hispidulus_ 4
+ _Sitones flavescens_ 1
+ _Trichalophus alternatus_ 1
+ _Apion_ sp 1
+ _Listronotus latiusculus_ 1
+ _Listronotus inæqualipennis_ 1
+ _Listronotus_ sp 1
+ _Macrops_ sp 2
+ _Smicronyx corniculatus_ 1
+ _Trachodes ptinoides_ 1
+ _Conotrachelus nenuphar_ 2
+ _Conotrachelus posticatus_ 5
+ _Conotrachelus erinaceus_ 1
+ _Rhinoncus pyrrhopus_ 1
+ _Onychobaris insidiosus_ 1
+ _Balaninus nasicus_ 1
+ _Balaninus_ sp 1
+ _Sphenophorus parvulus_ 1
+ _Sphenophorus_ sp 1
+ _Dendroctonus terebrans_ 1
+
+ HEMIPTERA.
+
+ _Podops cinctipes_ 1
+ _Nezara hilaris_ 6
+ _Arhaphe cicindeloides_ 1
+ _Corimelæna denudata_ 1
+ _Myodocha serripes_ 2
+
+ ORTHOPTERA.
+
+ _Amblycorypha rotundifolia_ 1
+ _Œcanthus niveus_ 1
+
+
+_Vegetable food._--The vegetable diet of the hermit thrush consists
+largely of fruit, as with most birds of this group. As might be
+expected of a bird of such retiring habits, but little of the fruit
+eaten can be classed as cultivated. In September 5.45 per cent was so
+considered, but in most months the quantity was small, and in March,
+April, and May was completely wanting. The total for the year as found
+in 17 stomachs is 1.20 per cent. One stomach contained strawberries,
+one grapes, one figs, one currants, two apples, and the rest _Rubus_
+fruit, i. e., blackberries or raspberries. These last as well as the
+strawberries were probably wild. Of the wild fruit (26.19 per cent) 46
+species were identified with a reasonable degree of certainty in 243
+stomachs. A few seeds, ground-up vegetable matter not further
+identified, and rubbish make up the rest of the vegetable food (8.10
+per cent). Among the seeds were some of the various species of
+poisonous _Rhus_. These were found in 18 stomachs, mostly from
+California. The dissemination of these seeds is unfortunate from the
+standpoint of husbandry, but many birds engage in it, as the waxy
+coating of the seeds is nutritious, especially in winter, when fruit
+and insects are not easily obtainable.
+
+Following is a list of the components of the vegetable food so far as
+identified, and the number of stomachs in which found:
+
+
+ Cedar berries (_Juniperus virginiana_) 2
+ False Solomon's seal (_Smilacina racemosa_) 4
+ False spikenard (_Smilacina_ sp.) 1
+ Greenbrier (_Smilax walteri_) 2
+ Cat brier (_Smilax bona-nox_) 2
+ Laurel-leaved greenbrier (_Smilax laurifolia_) 1
+ Other greenbriers (_Smilax_ sp.) 11
+ Wax myrtle (_Myrica cerifera_) 1
+ Bayberries (_Myrica carolinensis_) 7
+ Chinquapin (_Castanea pumila_) 1
+ Western hackberries (_Celtis occidentalis_) 5
+ Other hackberries (_Celtis_ sp.) 3
+ Figs (_Ficus_ sp.) 1
+ Mulberries (_Morus_ sp.) 1
+ Mistletoe berries (_Phoradendron villosum_) 2
+ Poke berries (_Phytolacca decandra_) 16
+ Miner's lettuce (_Montia perfoliata_) 1
+ Sassafras berries (_Sassafras varifolium_) 2
+ Spice berries (_Benzoin æstivale_) 1
+ Currants (_Ribes_ sp.) 3
+ Sweet gum (_Liquidambar styraciflua_) 2
+ Chokeberries (_Pyrus arbutifolia_) 1
+ Service berries (_Amelanchier canadensis_) 9
+ Hawthorn (_Cratægus_ sp.) 1
+ Strawberries (_Fragaria_ sp.) 1
+ Blackberries or raspberries (_Rubus_ sp.) 5
+ Rose haws (_Rosa_ sp.) 1
+ Wild black cherries (_Prunus scrotina_) 3
+ Three-seeded mercury (_Acalypha virginica_) 1
+ Staghorn sumach (_Rhus typhina_) 5
+ Smooth sumach (_Rhus glabra_) 5
+ Dwarf sumach (_Rhus copallina_) 7
+ Poison ivy (_Rhus radicans_) 3
+ Poison oak (_Rhus diversiloba_) 15
+ Laurel-leaved sumach (_Rhus laurina_) 2
+ Other sumachs (_Rhus_ sp.) 12
+ Pepper berries (_Schinus molle_) 15
+ American holly (_Ilex opaca_) 9
+ Black alder (_Ilex verticillata_) 12
+ Ink berries (_Ilex glabra_) 9
+ Other hollies (_Ilex_ sp.) 7
+ Strawberry bush (_Euonymus americanus_) 1
+ Roxbury waxwork (_Celastrus scandens_) 1
+ Supple-Jack (_Berchemia volubilis_) 2
+ Coffee berries (_Rhamnus californicus_) 1
+ Woodbine (_Psedera quinquefolia_) 10
+ Frost grapes (_Vitis cordifolia_) 2
+ Wild grapes (_Vitis_ sp.) 1
+ Wild sarsaparilla (_Aralia nudicaulis_) 1
+ Flowering dogwood (_Cornus florida_) 32
+ Rough-leaved dogwood (_Cornus asperifolia_) 2
+ Black gum (_Nyssa sylvatica_) 2
+ Checkerberry (_Gaultheria procumbens_) 1
+ Huckleberries (_Gaylussacia_ sp.) 1
+ Blueberries (_Vaccinium_ sp.) 12
+ Black nightshade (_Solanum nigrum_) 4
+ Bittersweet (_Solanum_ sp.) 4
+ Goose grass (_Galium aparine_) 1
+ Honeysuckle (_Lonicera_ sp.) 2
+ Indian currant (_Symphoricarpos orbiculatus_) 1
+ Downy arrowwood (_Viburnum pubescens_) 1
+ Nanny berries (_Viburnum lentago_) 2
+ Black elderberries (_Sambucus canadensis_) 4
+ Red elderberries (_Sambucus pubens_) 3
+ Fruit not further identified 60
+
+
+In looking over this list one is impressed with the fact that the
+taste of human beings for fruit differs markedly from that of birds.
+For example, _Rhus_ seeds are hard and have little pulp to render them
+palatable or nutritious. They are usually passed through the
+alimentary canal of birds or regurgitated unharmed, and the slight
+outer coating alone is digested. In the case of the poisonous species,
+this outer coating is a white wax or tallow which appears to be very
+nutritious, for these species are eaten much more extensively than the
+nonpoisonous ones. The seed itself is rarely broken in the stomach to
+get any nutriment it may contain. But in spite of these facts _Rhus_
+seeds were found in 49 stomachs, while fruits of huckleberries and
+blueberries, which are delicious to the human taste, were found in
+only 13 stomachs; and blackberries and raspberries, highly esteemed by
+man, were found in only 5 stomachs. Next to _Rhus_ the fruit most
+eaten was the dogwood berry, found in 34 stomachs, yet from a human
+estimate these berries are distasteful and contain such large seeds
+that they afford but very little actual food.
+
+_Summary._--The hermit thrush, as it name indicates, is of solitary
+habits and neither seeks human companionship nor molests cultivated
+products. It destroys nothing indirectly helpful to man, as beneficial
+insects, but aids in the destruction of the myriad hosts of insect
+life which at all times threaten vegetation. While it is not easy to
+point out any especially useful function of the hermit thrush, it
+fills its place in the economy of nature, from which it should not be
+removed.
+
+
+
+
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+
+ * * * * * * *
+
+
+ Transcriber's Notes
+
+ The text presented is essentially that in the original printed
+ document with the exception of some minor punctuation changes and
+ the three typographical corrections detailed below. The original
+ version also had two copies of the Table of Contents. The second
+ copy which appeared on Page 1 was removed. Many of the tables
+ which were presented in a two-column format and sometimes split
+ between two pages were reformatted into one long table.
+
+
+ Typographical Corrections
+
+ Page 1 : thrust => thrush
+ Page 10 : COLEOFTERA => COLEOPTERA
+ " : Cormybites => Corymbites
+
+
+ * * * * * * *
+
+
+
+
+
+
+
+
+End of the Project Gutenberg EBook of Food Habits of the Thrushes of the
+United States, by F. E. L. Beal
+
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