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| author | Roger Frank <rfrank@pglaf.org> | 2025-10-15 04:44:06 -0700 |
|---|---|---|
| committer | Roger Frank <rfrank@pglaf.org> | 2025-10-15 04:44:06 -0700 |
| commit | 3d8d97591af8f43392d3e1a88545250ae6979998 (patch) | |
| tree | 004d75f073c3923053bccac61bd582aac5060e95 /14279-h | |
Diffstat (limited to '14279-h')
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diff --git a/14279-h/14279-h.htm b/14279-h/14279-h.htm new file mode 100644 index 0000000..65e8c7a --- /dev/null +++ b/14279-h/14279-h.htm @@ -0,0 +1,24271 @@ +<!DOCTYPE html PUBLIC "-//W3C//DTD XHTML 1.0 Strict//EN" +"http://www.w3.org/TR/xhtml1/DTD/xhtml1-strict.dtd"> +<html xmlns="http://www.w3.org/1999/xhtml" xml:lang="en" lang="en"> +<head> +<meta http-equiv="Content-Type" content="text/html;charset=UTF-8" /> +<meta http-equiv="Content-Style-Type" content="text/css" /> +<title>The Project Gutenberg eBook of The Ancient Life History of the Earth, by Henry Alleyne Nicholson</title> + +<style type="text/css"> + + BODY { background: white; + margin-left: 15%; margin-right: 15%; } + P.indent { text-indent: 3mm; text-align: justify; } + P.contents { text-align: justify; font-size: smaller; + text-indent: -5mm; margin-left: 5mm; } + P.center { text-align: center; } + P.part { text-align: center; font-size: x-large; + font-weight: bold; } + P.subtitle { text-align: center; font-size: large; + font-style: italic; } + P.right { text-align: right; } + P.footnote { font-size: smaller; } + P.index { font-size: smaller; } + H1 { text-align: center; margin-top: 2em; + line-height: 150%; } + H2 { text-align: center; margin-top: 4em; } + H3 { text-align: center; margin-top: 4em; } + H4 { text-align: center; font-weight: normal; } + HR { color: #000000; + width: 10%; } + DIV.center { text-align: center; } + SPAN.image { font-size: smaller; } + TABLE.left { float: left; margin: 2mm; } + TABLE.right { float: right; margin: 2mm; } + TD.right { text-align: right; } + TD.center { text-align: center; } + TD.left { text-align: left; } + TD.genera { width: 33%; text-align: center; } + SPAN.page { position: absolute; left: 90%; right: auto; + text-align: right; text-indent: 0em; + color: gray; font-size: 9pt; + font-weight: normal; } + SPAN.big { font-size: xx-large; } + SPAN.smaller { font-size: smaller; } + +a:link {color:blue; text-decoration:none} +a:visited {color:blue; text-decoration:none} +a:hover {color:red} + + </style> +</head> + +<body> +<div>*** START OF THE PROJECT GUTENBERG EBOOK 14279 ***</div> + +<h1> +<span class="smaller">THE</span><br /> +ANCIENT LIFE-HISTORY<br /> +<span class="smaller">OF</span><br /> +THE EARTH +</h1> + +<p class="center"> +A COMPREHENSIVE OUTLINE OF THE PRINCIPLES AND LEADING FACTS OF +PALÆONTOLOGICAL SCIENCE +</p> + +<p class="center"><strong> +BY<br /> +H. ALLEYNE NICHOLSON +</strong></p> + +<p class="center"> +M.D., D.SC., M.A., PH. D. (GÖTT), F.R.S.E, F.L.S. +</p> + +<p class="center"> +PROFESSOR OF NATURAL HISTORY IN THE UNIVERSITY OF ST ANDREWS +</p> + +<h2> +<a name="page_v"><span class="page">Page v</span></a> +PREFACE.</h2> + +<hr /> + +<p class="indent"> +The study of Palæontology, or the science which is concerned +with the living beings which flourished upon the globe during +past periods of its history, may be pursued by two parallel but +essentially distinct paths. By the one method of inquiry, we may +study the anatomical characters and structure of the innumerable +extinct forms of life which lie buried in the rocks simply as so +many organisms, with but a slight and secondary reference to +the <i>time</i> at which they lived. By the other method, fossil +animals are regarded principally as so many landmarks in the +ancient records of the world, and are studied <i>historically</i> +and as regards their relations to the chronological succession +of the strata in which they are entombed. In so doing, it is of +course impossible to wholly ignore their structural characters, +and their relationships with animals now living upon the earth; +but these points are held to occupy a subordinate place, and to +require nothing more than a comparatively general attention. +</p> + +<p class="indent"> +In a former work, the Author has endeavoured to furnish a summary +of the more important facts of +<a name="page_vi"><span class="page">Page vi</span></a> +Palæontology regarded in its +strictly scientific aspect, as a mere department of the great +science of Biology. The present work, on the other hand, is an +attempt to treat Palæontology more especially from its +historical side, and in its more intimate relations with Geology. +In accordance with this object, the introductory portion of the +work is devoted to a consideration of the general principles of +Palæontology, and the bearings of this science upon various +geological problems—such as the mode of formation of the +sedimentary rocks, the reactions of living beings upon the crust +of the earth, and the sequence in time of the fossiliferous +formations. The second portion of the work deals exclusively with +Historical Palæontology, each formation being considered +separately, as regards its lithological nature and subdivisions, +its relations to other formations, its geographical distribution, +its mode of origin, and its characteristic life-forms. +</p> + +<p class="indent"> +In the consideration of the characteristic fossils of each successive +period, a general account is given of their more important zoological +characters and their relations to living forms; but the technical +language of Zoology has been avoided, and the aid of illustrations +has been freely called into use. It may therefore be hoped that +the work may be found to be available for the purposes of both the +Geological and the Zoological student; since it is essentially an +outline of Historical Palæontology, and the student of either +of the above-mentioned sciences must perforce possess some knowledge +of the last. Whilst primarily intended for students, it may be added +that the method of treatment adopted has been so far untechnical as +not to render the work useless to the general reader who may desire +<a name="page_vii"><span class="page">Page vii</span></a> +to acquire some knowledge of a subject of such vast and universal +interest. +</p> + +<p class="indent"> +In carrying out the object which he has held before him, the +Author can hardly expect, from the nature of the materials with +which he has had to deal, that he has kept himself absolutely +clear of errors, both of omission and commission. The subject, +however, is one to which he has devoted the labour of many years, +both in studying the researches of others and in personal +investigations of his own; and he can only trust that such errors +as may exist will be found to belong chiefly to the former class, +and to be neither serious nor numerous. It need only be added +that the work is necessarily very limited in its scope, and that +the necessity of not assuming a thorough previous acquaintance +with Natural History in the reader has inexorably restricted its +range still further. The Author does not, therefore, profess to +have given more than a merely general outline of the subject; and +those who desire to obtain a more minute and detailed knowledge +of Palæontology, must have recourse to other and more elaborate +treatises. +</p> + +<p> +UNITED COLLEGE, ST ANDREWS.<br/> + <i>October</i> 2, 1876. +</p> + +<h2> +<a name="page_ix"><span class="page">Page ix</span></a> +CONTENTS.</h2> + +<hr /> + +<p class="center"><strong> +PART I. +</strong></p> + +<p class="center"><strong> +PRINCIPLES OF PALÆONTOLOGY. +</strong></p> + +<p class="center"> +<a href="#page_1"> +INTRODUCTION.</a> +</p> + +<p class="contents"> +The general objects or geological science—The older theories +of catastrophistic and intermittent action—The more modern +doctrines of continuous and uniform action—Bearing of these +doctrines respectively on the origin or the existing terrestrial +order—Elements or truth in Catastrophism—General truth +of the doctrine of Continuity—Geological time. +</p> + +<p class="center"> +<a href="#page_10"> +CHAPTER I.</a> +</p> + +<p class="contents"> +Definition of Palæontology—Nature of +Fossils—Different processes of fossilisation. +</p> + +<p class="center"> +<a href="#page_14"> +CHAPTER II.</a> +</p> + +<p class="contents"> +Aqueous and igneous rocks—General characters of the +sedimentary rocks—Mode or formation of the sedimentary +rocks—Definition of the term "formation"—Chief divisions +of the aqueous rocks—Mechanically-formed rocks, their +characters and mode of origin—Chemically and organically +formed rocks—Calcareous rocks—Chalk, its microscopic +structure and mode of formation—Limestone, varieties, +structure, and origin—Phosphate of +lime—Concretions—Sulphate of lime—Silica and +siliceous deposits of various kinds—Greensands—Red +clays—Carbon and carbonaceous deposits. +</p> + +<p class="center"> +<a href="#page_37"> +CHAPTER III.</a> +</p> + +<p class="contents"> +Chronological succession of the fossiliferous rocks—Tests +or age of strata—Value of Palæontological evidence +in stratigraphical Geology—General sequence of the great +formations. +</p> + +<p class="center"> +<a name="page_x"><span class="page">Page x</span></a> +<a href="#page_44"> +CHAPTER IV.</a> +</p> + +<p class="contents"> +The breaks in the palæontological and geological +record—Use of the term "contemporaneous" as applied to +groups of strata—General sequence of strata and of +life-forms interfered with by more or less extensive +gaps—Unconformability—Phenomena implied by +this—Causes of the imperfection of the +palæontological record. +</p> + +<p class="center"> +<a href="#page_52"> +CHAPTER V.</a> +</p> + +<p class="contents"> +Conclusions to be drawn from fossils—Age of rocks—Mode +of origin of any fossiliferous bed—Fluviatile, lacustrine, +and marine deposits—Conclusions as to climate—Proofs +of elevation and subsidence of portions of the earth's crust +derived from fossils. +</p> + +<p class="center"> +<a href="#page_57"> +CHAPTER VI.</a> +</p> + +<p class="contents"> +The biological relations of fossils—Extinction of +life-forms—Geological range of different +species—Persistent types of life—Modern origin of +existing animals and plants—Reference of fossil forms to +the existing primary divisions of the animal +kingdom—Departure of the older types of life from those +now in existence—Resemblance of the fossils of a given +formation to those of the formation next above and next +below—Introduction of new life-forms. +</p> + +<hr /> + +<p class="center"><strong> +PART II. +</strong></p> + +<p class="center"><strong> +HISTORICAL PALÆONTOLOGY. +</strong></p> + +<p class="center"> +<a href="#page_65"> +CHAPTER VII.</a> +</p> + +<p class="contents"> +The Laurentian and Huronian periods—General nature, divisions, +and geographical distribution of the Laurentian deposits—Lower +and Upper Laurentian—Reasons for believing that the Laurentian +rocks are not azoic based upon their containing limestones, beds of +oxide of iron, and graphite—The characters, chemical +composition, and minute structure of <i>Eozoön +Canadense</i>—Comparison of <i>Eozoön</i> with existing +Foraminifera—<i>ArchĹ“osphĹ“rinĹ“</i>—Huronian +formation—Nature and distribution of Huronian +deposits—Organic remains of the Huronian—Literature. +</p> + +<p class="center"> +<a href="#page_77"> +CHAPTER VIII.</a> +</p> + +<p class="contents"> +The Cambrian period—General succession of Cambrian deposits in +Wales—Lower Cambrian and Upper Cambrian—Cambrian deposits of +the continent of Europe and North American—Life of the Cambrian +period — Fucoids — Eophyton — Oldhamia — Sponges +— Echinoderms — Annelides — Crustaceans — +Structure of Trilobites—Brachiopods—Pteropods, Gasteropods, +and Bivalves—Cephalopods—Literature. +</p> + +<p class="center"> +<a name="page_xi"><span class="page">Page xi</span></a> +<a href="#page_90"> +CHAPTER IX.</a> +</p> + +<p class="contents"> +The Lower Silurian period—The Silurian rocks generally—Limits +of Lower and Upper Silurian—General succession, subdivisions, and +characters of the Lower Silurian rocks of Wales—General succession, +subdivisions, and characters of the Lower Silurian rocks of the +North American continent—Life of the period — Fucoids — +Protozoa — Graptolites — Structure of Graptolites — +Corals — General structure of Corals — Crinoids — +Cystideans — General characters of Cystideans — Annelides +— Crustaceans — Polyzoa — Brachiopods — Bivalve +and Univalve Molluscs—Chambered Cephalopods—General +characters of the Cephalopoda—Conodonts. +</p> + +<p class="center"> +<a href="#page_115"> +CHAPTER X.</a> +</p> + +<p class="contents"> +The Upper Silurian period—General succession of the Upper Silurian +deposits of Wales—Upper Silurian deposits of North +America—Life of the Upper Silurian — Plants — Protozoa +— Graptolites — Corals — Crinoids — General +structure of Crinoids — Star-fishes — Annelides — +Crustaceans — Eurypterids — Polyzoa — Brachiopods +— Structure of Brachiopods — Bivalves and Univalves — +Pteropods — Cephalopods — Fishes — Silurian literature. +</p> + +<p class="center"> +<a href="#page_132"> +CHAPTER XI.</a> +</p> + +<p class="contents"> +The Devonian period—Relations between the Old Red Sandstone +and the marine Devonian deposits—The Old Red Sandstone of +Scotland—The Devonian strata of Devonshire—Sequence and +subdivisions of the Devonian deposits of North America—Life of +the period — Plants — Protozoa — Corals — +Crinoids — Pentremites — Annelides — Crustaceans +— Insects — Polyzoa — Brachiopods — Bivalves +— Univalves — Pteropods — Cephalopods — +Fishes — General divisions of the +Fishes—Palæontological evidence as to the +independent existence of the Devonian system as a distinct +formation—Literature. +</p> + +<p class="center"> +<a href="#page_157"> +CHAPTER XII.</a> +</p> + +<p class="contents"> +The Carboniferous period—Relations of Carboniferous rocks to +Devonian—The Carboniferous Limestone or Sub-Carboniferous +series—The Millstone-grit and the Coal-measures—Life of +the period—Structure and mode of formation of Coal—Plants +of the Coal. +</p> + +<p class="center"> +<a href="#page_170"> +CHAPTER XIII.</a> +</p> + +<p class="contents"> +Animal life of the Carboniferous period — Protozoa — +Corals — Crinoids — Pentremites — Structure of +Pentremites — Echinoids — Structure of Echinoidea +— Annelides — Crustacea — Insects — +Arachnids — Myriapods — Polyzoa — Brachiopods +— Bivalves and Univalves — Cephalopods — Fishes +— Labyrinthodont Amphibians—Literature. +</p> + +<p class="center"> +<a name="page_xii"><span class="page">Page xii</span></a> +<a href="#page_192"> +CHAPTER XIV.</a> +</p> + +<p class="contents"> +The Permian period — General succession, characters, and +mode of formation of the Permian deposits — Life of the +period — Plants — Protozoa — Corals — +Echinoderms — Annelides — Crustaceans — +Polyzoa — Brachiopods — Bivalves — Univalves +— Pteropods — Cephalopods — Fishes — +Amphibians — Reptiles — Literature. +</p> + +<p class="center"> +<a href="#page_203"> +CHAPTER XV.</a> +</p> + +<p class="contents"> +The Triassic period-—General characters and subdivisions of +the Trias of the Continent of Europe and Britain—Trias of +North America—Life of the period — Plants — +Echinoderms — Crustaceans — Polyzoa — +Brachiopods — Bivalves — Univalves — Cephalopods +— Intermixture of Palæozoic with Mesozoic types of +Molluscs — Fishes — Amphibians — Reptiles — +Supposed footprints of Birds — Mammals — Literature. +</p> + +<p class="center"> +<a href="#page_226"> +CHAPTER XVI.</a> +</p> + +<p class="contents"> +The Jurassic period—General sequence and subdivisions of the +Jurassic deposits in Britain—Jurassic rocks of North +America—Life of the period — Plants — Corals +— Echinoderms — Crustaceans — Insects — +Brachiopods — Bivalves — Univalves — Pteropods +— Tetrabranchiate Cephalopods — Dibranchiate +Cephalopods — Fishes — Reptiles — Birds — +Mammals — Literature. +</p> + +<p class="center"> +<a href="#page_256"> +CHAPTER XVII.</a> +</p> + +<p class="contents"> +The Cretaceous period—General succession and subdivisions +of the Cretaceous rocks in Britain—Cretaceous rocks of North +America—Life of the period — Plants — Protozoa +— Corals — Echinoderms — Crustaceans — +Polyzoa — Brachiopods — Bivalves — Univalves +— Tetrabranchiate and Dibranchiate Cephalopods — Fishes +— Reptiles — Birds — Literature. +</p> + +<p class="center"> +<a href="#page_284"> +CHAPTER XVIII.</a> +</p> + +<p class="contents"> +The Eocene period—Relations between the Kainozoic and Mesozoic +rocks in Europe and in North America—Classification of the +Tertiary deposits—The sequence and subdivisions of the Eocene +rocks of Britain and France—Eocene strata of the United +States—Life of the period — Plants — Foraminifera +— Corals — Echinoderms — Mollusca — Fishes +— Reptiles — Birds — Mammals. +</p> + +<p class="center"> +<a name="page_xiii"><span class="page">Page xiii</span></a> +<a href="#page_305"> +CHAPTER XIX.</a> +</p> + +<p class="contents"> +The Miocene period—Miocene strata of Britain—Of +France—Of Belgium—Of Austria—Of Switzerland—Of +Germany—Of Greece—Of India—Of North America—Of +the Arctic regions—Life of the period—Vegetation of the +Miocene period — Foraminifera — Corals — Echinoderms +— Articulates — Mollusca — Fishes — Amphibians +— Reptiles — Mammals. +</p> + +<p class="center"> +<a href="#page_323"> +CHAPTER XX.</a> +</p> + +<p class="contents"> +The Pliocene period—Pliocene deposits of Britain—Of +Europe—Of North America—Life of the period—Climate +of the period as indicated by the Invertebrate animals—The +Pliocene Mammalia—Literature relating to the Tertiary deposits +and their fossils. +</p> + +<p class="center"> +<a href="#page_334"> +CHAPTER XXI.</a> +</p> + +<p class="contents"> +The Post-Pliocene period—Division of the Quaternary deposits +into Post-Pliocene and Recent—Relations of the Post-Pliocene +deposits of the northern hemisphere to the "Glacial +period"—Pre-Glacial deposits—Glacial +deposits—Arctic Mollusca in Glacial beds—Post-Glacial +deposits—Nature and mode of formation of high-level and +low-level gravels—Nature and mode of formation of +cavern-deposits—Kent's Cavern-Post—Pliocene deposits of +the southern hemisphere. +</p> + +<p class="center"> +<a href="#page_344"> +CHAPTER XXII.</a> +</p> + +<p class="contents"> +Life of the Post-Pliocene period—Effect of the coming on and +departure of the Glacial period upon the animals inhabiting the +northern hemisphere—Birds of the Post-Pliocene—Mammalia +of the Post-Pliocene—Climate of the Post-Glacial period as +deduced from the Post-Glacial Mammals—Occurrence of the bones +and implements of Man in Post-Pliocene deposits in association with +the remains of extinct Mammalia—Literature relating to the +Post-Pliocene period. +</p> + +<p class="center"> +<a href="#page_367"> +CHAPTER XXIII.</a> +</p> + +<p class="contents"> +The succession of life upon the globe—Gradual and successive +introduction of life-forms—What is meant by "lower" and +"higher" groups of animals and plants—Succession in time of +the great groups of animals in the main corresponding with their +zoological order—Identical phenomena in the vegetable +kingdom—Persistent types of life—High organisation of +many early forms—Bearings of Palæontology on the +general doctrine of Evolution. +</p> + +<p class="contents"> +<a href="#page_375"> +APPENDIX.</a>—Tabular view of the chief Divisions of the +Animal Kingdom. +</p> + +<p class="contents"> +<a href="#page_379"> +GLOSSARY.</a> +</p> + +<p class="contents"> +<a href="#page_396"> +INDEX.</a> +</p> + +<h2> +<a name="page_xv"><span class="page">Page xv</span></a> +LIST OF ILLUSTRATIONS.</h2> + +<hr /> + +<table summary="" style=""> +<tr><td valign="top" class="right">FIG.</td><td> + +</td></tr> + +<tr><td valign="top" class="right">1.</td><td> +Cast of <i>Trigonia longa</i>. +</td></tr> + +<tr><td valign="top" class="right">2.</td><td> +Microscopic section of the wood of a fossil Conifer. +</td></tr> + +<tr><td valign="top" class="right">3.</td><td> +Microscopic section of the wood of the Larch. +</td></tr> + +<tr><td valign="top" class="right">4.</td><td> +Section of Carboniferous strata, Kinghorn, Fife. +</td></tr> + +<tr><td valign="top" class="right">5.</td><td> +Diagram illustrating the formation of stratified deposits. +</td></tr> + +<tr><td valign="top" class="right">6.</td><td> +Microscopic section of a calcareous breccia. +</td></tr> + +<tr><td valign="top" class="right">7.</td><td> +Microscopic section of White Chalk. +</td></tr> + +<tr><td valign="top" class="right">8.</td><td> +Organisms in Atlantic Ooze. +</td></tr> + +<tr><td valign="top" class="right">9.</td><td> +Crinoidal marble. +</td></tr> + +<tr><td valign="top" class="right">10.</td><td> +Piece of Nummulitic limestone, Pyramids. +</td></tr> + +<tr><td valign="top" class="right">11.</td><td> +Microscopic section of Foraminiferal +limestone—Carboniferous, America. +</td></tr> + +<tr><td valign="top" class="right">12.</td><td> +Microscopic section of Lower Silurian limestone. +</td></tr> + +<tr><td valign="top" class="right">13.</td><td> +Microscopic section of oolitic limestone, Jurassic. +</td></tr> + +<tr><td valign="top" class="right">14.</td><td> +Microscopic section of oolitic limestone, Carboniferous. +</td></tr> + +<tr><td valign="top" class="right">15.</td><td> +Organisms in Barbadoes earth. +</td></tr> + +<tr><td valign="top" class="right">15.</td><td> +Organisms in Barbadoes earth. +</td></tr> + +<tr><td valign="top" class="right">16.</td><td> +Organisms in Richmond earth. +</td></tr> + +<tr><td valign="top" class="right">17.</td><td> +Ideal section of the crust of the earth. +</td></tr> + +<tr><td valign="top" class="right">18.</td><td> +Unconformable junction of Chalk and Eocene rocks. +</td></tr> + +<tr><td valign="top" class="right">19.</td><td> +Erect trunk of a <i>Sigillaria</i>. +</td></tr> + +<tr><td valign="top" class="right">20.</td><td> +Diagrammatic section of the Laurentian rocks +</td></tr> + +<tr><td valign="top" class="right">21.</td><td> +Microscopic section of Laurentian limestone. +</td></tr> + +<tr><td valign="top" class="right">22.</td><td> +Fragment of a mass of <i>Eozoön Canadense</i>. +</td></tr> + +<tr><td valign="top" class="right">23.</td><td> +Diagram illustrating the structure of <i>Eozoön</i>. +</td></tr> + +<tr><td valign="top" class="right">24.</td><td> +Microscopic section of <i>Eozoön Canadense</i>. +</td></tr> + +<tr><td valign="top" class="right">25.</td><td> +<i>Nonionina</i> and <i>Gromia</i>. +</td></tr> + +<tr><td valign="top" class="right">26.</td><td> +Group of shells of living <i>Foraminifera</i>. +</td></tr> + +<tr><td valign="top" class="right">27.</td><td> +Diagrammatic section of Cambrian strata. +</td></tr> + +<tr><td valign="top" class="right">28.</td><td> +<i>Eophyton Linneanum</i>. +</td></tr> + +<tr><td valign="top" class="right">29.</td><td> +<i>Oldhamia antiqua</i>. +</td></tr> + +<tr><td valign="top" class="right">30.</td><td> +<i>Scolithus Canadensis</i>. +</td></tr> + +<tr><td valign="top" class="right">31.</td><td> +Group of Cambrian Trilobites. +</td></tr> + +<tr><td valign="top" class="right">32.</td><td> +Group of characteristic Cambrian fossils. +</td></tr> + +<tr><td valign="top" class="right">33.</td><td> +Fragment of <i>Dictyonema sociale</i>. +</td></tr> + +<tr><td valign="top" class="right">34.</td><td> +Generalised section of the Lower Silurian rocks +of Wales. +</td></tr> + +<tr><td valign="top" class="right">35.</td><td> +Generalised section of the Lower Silurian rocks +of North America. +</td></tr> + +<tr><td valign="top" class="right">36.</td><td> +<i>Licrophycus Ottawaensis</i>. +</td></tr> + +<tr><td valign="top" class="right">37.</td><td> +<i>Astylospongia prĹ“morsa</i>. +</td></tr> + +<tr><td valign="top" class="right">38.</td><td> +<i>Stromatopora rugosa</i>. +</td></tr> + +<tr><td valign="top" class="right">39.</td><td> +<i>Dichograptus octobrachiatus</i>. +</td></tr> + +<tr><td valign="top" class="right">40.</td><td> +<i>Didymograptus divaricatus</i>. +<a name="page_xvi"><span class="page">Page xvi</span></a> +</td></tr> + +<tr><td valign="top" class="right">41.</td><td> +<i>Diplograptus pristis</i>. +</td></tr> + +<tr><td valign="top" class="right">42.</td><td> +<i>Phyllograptus typus</i>. +</td></tr> + +<tr><td valign="top" class="right">43.</td><td> +<i>Zaphrentis Stokesi</i>. +</td></tr> + +<tr><td valign="top" class="right">44.</td><td> +<i>Strombodes pentagonus</i>. +</td></tr> + +<tr><td valign="top" class="right">45.</td><td> +<i>Columnaria alveolata</i>. +</td></tr> + +<tr><td valign="top" class="right">46.</td><td> +Group of Cystideans. +</td></tr> + +<tr><td valign="top" class="right">47.</td><td> +Group of Lower Silurian Crustaceans. +</td></tr> + +<tr><td valign="top" class="right">48.</td><td> +<i>Ptilodictya falciformis</i>. +</td></tr> + +<tr><td valign="top" class="right">49.</td><td> +<i>Ptilodictya Schafferi</i>. +</td></tr> + +<tr><td valign="top" class="right">50.</td><td> +Group of Lower Silurian Brachiopods. +</td></tr> + +<tr><td valign="top" class="right">51.</td><td> +Group of Lower Silurian Brachiopods. +</td></tr> + +<tr><td valign="top" class="right">52.</td><td> +<i>Murchisonia gracilis</i>. +</td></tr> + +<tr><td valign="top" class="right">53.</td><td> +<i>Bellerophon argo</i>. +</td></tr> + +<tr><td valign="top" class="right">54.</td><td> +<i>Maclurea crenulata</i>. +</td></tr> + +<tr><td valign="top" class="right">55.</td><td> +<i>Orthoceras crebriseptum</i>. +</td></tr> + +<tr><td valign="top" class="right">56.</td><td> +Restoration of <i>Orthoceras</i>. +</td></tr> + +<tr><td valign="top" class="right">57.</td><td> +Generalised section of the Upper Silurian rocks. +</td></tr> + +<tr><td valign="top" class="right">58.</td><td> +<i>Monograptus priodon</i>. +</td></tr> + +<tr><td valign="top" class="right">59.</td><td> +<i>Halysites catenularia</i> and <i>H. agglomerata</i>. +</td></tr> + +<tr><td valign="top" class="right">60.</td><td> +Group of Upper Silurian Star-fishes. +</td></tr> + +<tr><td valign="top" class="right">61.</td><td> +<i>Protaster Sedgwickii</i>. +</td></tr> + +<tr><td valign="top" class="right">62.</td><td> +Group of Upper Silurian Crinoids. +</td></tr> + +<tr><td valign="top" class="right">63.</td><td> +<i>Planolites vulgaris</i>. +</td></tr> + +<tr><td valign="top" class="right">64.</td><td> +Group of Upper Silurian Trilobites. +</td></tr> + +<tr><td valign="top" class="right">65.</td><td> +<i>Pterygotus Anglicus</i>. +</td></tr> + +<tr><td valign="top" class="right">66.</td><td> +Group of Upper Silurian <i>Polyzoa</i>. +</td></tr> + +<tr><td valign="top" class="right">67.</td><td> +<i>Spirifera hysterica</i>. +</td></tr> + +<tr><td valign="top" class="right">68.</td><td> +Group of Upper Silurian Brachiopods. +</td></tr> + +<tr><td valign="top" class="right">69.</td><td> +Group of Upper Silurian Brachiopods. +</td></tr> + +<tr><td valign="top" class="right">70.</td><td> +<i>Pentamerus Knightii</i>. +</td></tr> + +<tr><td valign="top" class="right">71.</td><td> +<i>Cardiola interrupta, C. fibrosa</i>, and +<i>PterinĹ“a subfalcata</i>. +</td></tr> + +<tr><td valign="top" class="right">72.</td><td> +Group of Upper Silurian Univalves. +</td></tr> + +<tr><td valign="top" class="right">73.</td><td> +<i>Tentaculites ornatus</i>. +</td></tr> + +<tr><td valign="top" class="right">74.</td><td> +<i>Pteraspis Banksii</i>. +</td></tr> + +<tr><td valign="top" class="right">75.</td><td> +<i>Onchus tenuistriatus</i> and <i>Thelodus</i>. +</td></tr> + +<tr><td valign="top" class="right">76.</td><td> +Generalised section of the Devonian rocks of North America. +</td></tr> + +<tr><td valign="top" class="right">77.</td><td> +<i>Psilophyton princeps</i>. +</td></tr> + +<tr><td valign="top" class="right">78.</td><td> +<i>Prototaxites Logani</i>. +</td></tr> + +<tr><td valign="top" class="right">79.</td><td> +<i>Stromatopora tuberculata</i>. +</td></tr> + +<tr><td valign="top" class="right">80.</td><td> +<i>Cystiphyllum vesiculosum</i>. +</td></tr> + +<tr><td valign="top" class="right">81.</td><td> +<i>Zaphrentis cornicula</i>. +</td></tr> + +<tr><td valign="top" class="right">82.</td><td> +<i>Heliophyllum exiguum</i>. +</td></tr> + +<tr><td valign="top" class="right">83.</td><td> +<i>Crepidophyllum Archiaci</i>. +</td></tr> + +<tr><td valign="top" class="right">84.</td><td> +<i>Favosites Gothlandica</i>. +</td></tr> + +<tr><td valign="top" class="right">85.</td><td> +<i>Favosites hemisphĹ“rica</i>. +</td></tr> + +<tr><td valign="top" class="right">86.</td><td> +<i>Spirorbis omphalodes</i> and <i>S. Arkonensis</i>. +</td></tr> + +<tr><td valign="top" class="right">87.</td><td> +<i>Spirorbis laxus</i> and <i>S. Spinulifera</i>. +</td></tr> + +<tr><td valign="top" class="right">88.</td><td> +Group of Devonian Trilobites. +</td></tr> + +<tr><td valign="top" class="right">89.</td><td> +Wing of <i>Platephemera antiqua</i>. +</td></tr> + +<tr><td valign="top" class="right">90.</td><td> +<i>Clathropora intertexta</i>. +</td></tr> + +<tr><td valign="top" class="right">91.</td><td> +<i>Ceriopora Hamiltonensis</i>. +</td></tr> + +<tr><td valign="top" class="right">92.</td><td> +<i>Fenestella magnifica</i>. +</td></tr> + +<tr><td valign="top" class="right">93.</td><td> +<i>Retepora Phillipsi</i>. +</td></tr> + +<tr><td valign="top" class="right">94.</td><td> +<i>Fenestella cribrosa</i>. +</td></tr> + +<tr><td valign="top" class="right">95.</td><td> +<i>Spirifera sculptilis</i>. +</td></tr> + +<tr><td valign="top" class="right">96.</td><td> +<i>Spirifera mucronata</i>. +</td></tr> + +<tr><td valign="top" class="right">97.</td><td> +<i>Atrypa reticularis</i>. +</td></tr> + +<tr><td valign="top" class="right">98.</td><td> +<i>Strophomena rhomboidalis</i>. +</td></tr> + +<tr><td valign="top" class="right">99.</td><td> +<i>Platyceras dumosum</i>. +</td></tr> + +<tr><td valign="top" class="right">100.</td><td> +<i>Conularia ornata</i>. +</td></tr> + +<tr><td valign="top" class="right">101.</td><td> +<i>Clymenia Sedgwickii</i>. +</td></tr> + +<tr><td valign="top" class="right">102.</td><td> +Group of Fishes from the Devonian rocks of North America. +</td></tr> + +<tr><td valign="top" class="right">103.</td><td> +<i>Cephalaspis Lyellii</i>. +</td></tr> + +<tr><td valign="top" class="right">104.</td><td> +<i>Pterichthys cornutus</i>. +</td></tr> + +<tr><td valign="top" class="right">105.</td><td> +<i>Polypterus</i> and <i>Osteolepis</i>. +</td></tr> + +<tr><td valign="top" class="right">106.</td><td> +<i>Holoptychius nobilissimus</i>. +</td></tr> + +<tr><td valign="top" class="right">107.</td><td> +Generalised section of the Carboniferous rocks of the +North of England. +</td></tr> + +<tr><td valign="top" class="right">108.</td><td> +<i>Odontopteris Schlotheimii</i>. +</td></tr> + +<tr><td valign="top" class="right">109.</td><td> +<i>Calamites cannĹ“formis</i>. +</td></tr> + +<tr><td valign="top" class="right">110.</td><td> +<i>Lepidodendron Sternbergii</i>. +</td></tr> + +<tr><td valign="top" class="right">111.</td><td> +<i>Sigillaria GrĹ“seri</i>. +</td></tr> + +<tr><td valign="top" class="right">112.</td><td> +<i>Stigmaria ficoides</i>. +</td></tr> + +<tr><td valign="top" class="right">113.</td><td> +<i>Trigonocarpum ovatum</i>. +</td></tr> + +<tr><td valign="top" class="right">114.</td><td> +Microscopic section of Foraminiferal +limestone—Carboniferous, North America. +</td></tr> + +<tr><td valign="top" class="right">115.</td><td> +<i>Fusulina cylindrica</i>. +</td></tr> + +<tr><td valign="top" class="right">116.</td><td> +Group of Carboniferous Corals. +</td></tr> + +<tr><td valign="top" class="right">117.</td><td> +<i>Platycrinus tricontadactylus</i>. +</td></tr> + +<tr><td valign="top" class="right">118.</td><td> +<i>Pentremites pyriformis</i> and <i>P. conoideus</i>. +<a name="page_xvii"><span class="page">Page xvii</span></a> +</td></tr> + +<tr><td valign="top" class="right">119.</td><td> +<i>ArchĹ“ocidaris ellipticus</i>. +</td></tr> + +<tr><td valign="top" class="right">120.</td><td> +<i>Spirorbis Carbonarius</i>. +</td></tr> + +<tr><td valign="top" class="right">121.</td><td> +<i>Prestwichia rotundata</i>. +</td></tr> + +<tr><td valign="top" class="right">122.</td><td> +Group of Carboniferous Crustaceans. +</td></tr> + +<tr><td valign="top" class="right">123.</td><td> +<i>Cyclophthalmus senior</i>. +</td></tr> + +<tr><td valign="top" class="right">124.</td><td> +<i>Xylobius SigillariĹ“</i>. +</td></tr> + +<tr><td valign="top" class="right">125.</td><td> +<i>Haplophlebium Barnesi</i>. +</td></tr> + +<tr><td valign="top" class="right">126.</td><td> +Group of Carboniferous <i>Polyzoa</i>. +</td></tr> + +<tr><td valign="top" class="right">127.</td><td> +Group of Carboniferous <i>Brachiopoda</i>. +</td></tr> + +<tr><td valign="top" class="right">128.</td><td> +<i>Pupa vetusta</i>. +</td></tr> + +<tr><td valign="top" class="right">129.</td><td> +<i>Goniatites FossĹ“</i>. +</td></tr> + +<tr><td valign="top" class="right">130.</td><td> +<i>Amblypterus macropterus</i>. +</td></tr> + +<tr><td valign="top" class="right">131.</td><td> +<i>Cochliodus contortus</i>. +</td></tr> + +<tr><td valign="top" class="right">132.</td><td> +<i>Anthracosaurus Russelli</i>. +</td></tr> + +<tr><td valign="top" class="right">133.</td><td> +Generalised section of the Permian rocks. +</td></tr> + +<tr><td valign="top" class="right">134.</td><td> +<i>Walchia piniformis</i>. +</td></tr> + +<tr><td valign="top" class="right">135.</td><td> +Group of Permian <i>Brachiopods</i>. +</td></tr> + +<tr><td valign="top" class="right">136.</td><td> +<i>Arca antiqua</i>. +</td></tr> + +<tr><td valign="top" class="right">137.</td><td> +<i>Platysomus gibbosus</i>. +</td></tr> + +<tr><td valign="top" class="right">138.</td><td> +<i>Protorosaurus Speneri</i>. +</td></tr> + +<tr><td valign="top" class="right">139.</td><td> +Generalised section of the Triassic rocks. +</td></tr> + +<tr><td valign="top" class="right">140.</td><td> +<i>Zamia spiralis</i>. +</td></tr> + +<tr><td valign="top" class="right">141.</td><td> +Triassic Conifers and Cycads. +</td></tr> + +<tr><td valign="top" class="right">142.</td><td> +<i>Encrinus liliiformis</i>. +</td></tr> + +<tr><td valign="top" class="right">143.</td><td> +<i>Aspidura loricata</i>. +</td></tr> + +<tr><td valign="top" class="right">144.</td><td> +Group of Triassic Bivalves. +</td></tr> + +<tr><td valign="top" class="right">145.</td><td> +<i>Ceratites nodosus</i>. +</td></tr> + +<tr><td valign="top" class="right">146.</td><td> +Tooth of <i>Ceratodus serratus</i> and <i>C. Altus</i>. +</td></tr> + +<tr><td valign="top" class="right">147.</td><td> +<i>Ceratodus Fosteri</i>. +</td></tr> + +<tr><td valign="top" class="right">148.</td><td> +Footprints of <i>Cheirotherium</i>. +</td></tr> + +<tr><td valign="top" class="right">149.</td><td> +Section of tooth of <i>Labyrinthodont</i>. +</td></tr> + +<tr><td valign="top" class="right">150.</td><td> +Skull of <i>Mastodonsaurus</i>. +</td></tr> + +<tr><td valign="top" class="right">151.</td><td> +Skull of <i>Rhynchosaurus</i>. +</td></tr> + +<tr><td valign="top" class="right">152.</td><td> +<i>Belodon</i>, <i>Nothosaurus</i>, +<i>PalĹ“osaurus</i>, &c. +</td></tr> + +<tr><td valign="top" class="right">153.</td><td> +<i>Placodus gigas</i>. +</td></tr> + +<tr><td valign="top" class="right">154.</td><td> +Skulls of <i>Dicynodon</i> and <i>Oudenodon</i>. +</td></tr> + +<tr><td valign="top" class="right">155.</td><td> +Supposed footprint of Bird, from the Trias of Connecticut. +</td></tr> + +<tr><td valign="top" class="right">156.</td><td> +Lower jaw of <i>Dromatherium sylvestre</i>. +</td></tr> + +<tr><td valign="top" class="right">157.</td><td> +Molar tooth of <i>Microlestes antiquus</i>. +</td></tr> + +<tr><td valign="top" class="right">158.</td><td> +<i>Myrmecobius fasciatus</i>. +</td></tr> + +<tr><td valign="top" class="right">159.</td><td> +Generalised section of the Jurassic rocks. +</td></tr> + +<tr><td valign="top" class="right">160.</td><td> +<i>Mantellia megalophylla</i>. +</td></tr> + +<tr><td valign="top" class="right">161.</td><td> +<i>Thecosmilia annularis</i>. +</td></tr> + +<tr><td valign="top" class="right">162.</td><td> +<i>Pentacrinus fasciculosus</i>. +</td></tr> + +<tr><td valign="top" class="right">163.</td><td> +<i>Hemicidaris crenularis</i>. +</td></tr> + +<tr><td valign="top" class="right">164.</td><td> +<i>Eryon arctiformis</i>. +</td></tr> + +<tr><td valign="top" class="right">165.</td><td> +Group of Jurassic Brachiopods. +</td></tr> + +<tr><td valign="top" class="right">166.</td><td> +<i>Ostrea Marshii</i>. +</td></tr> + +<tr><td valign="top" class="right">167.</td><td> +<i>GryphĹ“a incurva</i> +</td></tr> + +<tr><td valign="top" class="right">168.</td><td> +<i>Diceras arietina</i>. +</td></tr> + +<tr><td valign="top" class="right">169.</td><td> +<i>NerinĹ“a Goodhallii</i>. +</td></tr> + +<tr><td valign="top" class="right">170.</td><td> +<i>Ammonites Humphresianus</i>. +</td></tr> + +<tr><td valign="top" class="right">171.</td><td> +<i>Ammonites bifrons</i>. +</td></tr> + +<tr><td valign="top" class="right">172.</td><td> +<i>Beloteuthis subcostata</i>. +</td></tr> + +<tr><td valign="top" class="right">173.</td><td> +Belemnite restored; diagram of Belemnite; <i>Belemnites +canaliculata</i>. +</td></tr> + +<tr><td valign="top" class="right">174.</td><td> +<i>Tetragonolepis</i>. +</td></tr> + +<tr><td valign="top" class="right">175.</td><td> +<i>Acrodus nobilis</i>. +</td></tr> + +<tr><td valign="top" class="right">176.</td><td> +<i>Ichthyosaurus communis</i>. +</td></tr> + +<tr><td valign="top" class="right">177.</td><td> +<i>Plesiosaurus dolichodeirus</i>. +</td></tr> + +<tr><td valign="top" class="right">178.</td><td> +<i>Pterodactylus crassirostris</i>. +</td></tr> + +<tr><td valign="top" class="right">179.</td><td> +<i>Ramphorhynchus Bucklandi</i>, restored. +</td></tr> + +<tr><td valign="top" class="right">180.</td><td> +Skull of <i>Megalosaurus</i>. +</td></tr> + +<tr><td valign="top" class="right">181.</td><td> +<i>ArchĹ“opteryx macrura</i>. +</td></tr> + +<tr><td valign="top" class="right">182.</td><td> +<i>ArchĹ“opteryx, restored</i>. +</td></tr> + +<tr><td valign="top" class="right">183.</td><td> +Jaw of <i>Amphitherium Prevostii</i>. +</td></tr> + +<tr><td valign="top" class="right">184.</td><td> +Jaws of Oolitic Mammals. +</td></tr> + +<tr><td valign="top" class="right">185.</td><td> +Generalised section of the Cretaceous rocks. +</td></tr> + +<tr><td valign="top" class="right">186.</td><td> +Cretaceous Angiosperms. +</td></tr> + +<tr><td valign="top" class="right">187.</td><td> +<i>Rotalia Boueana</i>. +</td></tr> + +<tr><td valign="top" class="right">188.</td><td> +<i>Siphonia ficus</i>. +</td></tr> + +<tr><td valign="top" class="right">189.</td><td> +<i>Ventriculites simplex</i>. +</td></tr> + +<tr><td valign="top" class="right">190.</td><td> +<i>Synhelia Sharpeana</i>. +</td></tr> + +<tr><td valign="top" class="right">191.</td><td> +<i>Galerites albogalerus</i>. +</td></tr> + +<tr><td valign="top" class="right">192.</td><td> +<i>Discoidea cylindrica</i>. +</td></tr> + +<tr><td valign="top" class="right">193.</td><td> +<i>Escharina Oceani</i>. +</td></tr> + +<tr><td valign="top" class="right">194.</td><td> +<i>Terebratella Astieriana</i>. +</td></tr> + +<tr><td valign="top" class="right">195.</td><td> +<i>Crania Ignabergensis</i>. +</td></tr> + +<tr><td valign="top" class="right">196.</td><td> +<i>Ostrea Couloni</i>. +</td></tr> + +<tr><td valign="top" class="right">197.</td><td> +<i>Spondylus spinosus</i>. +</td></tr> + +<tr><td valign="top" class="right">198.</td><td> +<i>Inoceramus sulcatus</i>. +</td></tr> + +<tr><td valign="top" class="right">199.</td><td> +<i>Hippurites Toucasiana</i>. +</td></tr> + +<tr><td valign="top" class="right">200.</td><td> +<i>Voluta elongata</i>. +</td></tr> + +<tr><td valign="top" class="right">201.</td><td> +<i>Nautilus Danicus</i>. +</td></tr> + +<tr><td valign="top" class="right">202.</td><td> +<i>Ancyloceras Matheronianus</i>. +<a name="page_xviii"><span class="page">Page xviii</span></a> +</td></tr> + +<tr><td valign="top" class="right">203.</td><td> +<i>Turrilites catenatus</i> +</td></tr> + +<tr><td valign="top" class="right">204.</td><td> +Forms of Cretaceous <i>AmmonitidĹ“</i>. +</td></tr> + +<tr><td valign="top" class="right">205.</td><td> +<i>Belemnitella mucronata</i>. +</td></tr> + +<tr><td valign="top" class="right">206.</td><td> +Tooth of <i>Hybodus</i>. +</td></tr> + +<tr><td valign="top" class="right">207.</td><td> +Fin-spine of <i>Hybodus</i>. +</td></tr> + +<tr><td valign="top" class="right">208.</td><td> +<i>Beryx Lewesiensis</i> and <i>Osmeroides Mantelli</i>. +</td></tr> + +<tr><td valign="top" class="right">209.</td><td> +Teeth of <i>Iguanodon</i>. +</td></tr> + +<tr><td valign="top" class="right">210.</td><td> +Skull of <i>Mosasaurus Camperi</i>. +</td></tr> + +<tr><td valign="top" class="right">211.</td><td> +<i>Chelone Benstedi</i>. +</td></tr> + +<tr><td valign="top" class="right">212.</td><td> +Jaws and vertebræ of <i>Odontornithes</i>. +</td></tr> + +<tr><td valign="top" class="right">213.</td><td> +Fruit of <i>Nipadites</i>. +</td></tr> + +<tr><td valign="top" class="right">214.</td><td> +<i>Nummulina lĹ“vigata</i>. +</td></tr> + +<tr><td valign="top" class="right">215.</td><td> +<i>Turbinolia sulcata</i>. +</td></tr> + +<tr><td valign="top" class="right">216.</td><td> +<i>Cardita planicosta</i>. +</td></tr> + +<tr><td valign="top" class="right">217.</td><td> +<i>Typhis tubifer</i>. +</td></tr> + +<tr><td valign="top" class="right">218.</td><td> +<i>CyprĹ“a elegans</i>. +</td></tr> + +<tr><td valign="top" class="right">219.</td><td> +<i>Cerithium hexagonum</i>. +</td></tr> + +<tr><td valign="top" class="right">220.</td><td> +<i>LimnĹ“a pyramidalis</i>. +</td></tr> + +<tr><td valign="top" class="right">221.</td><td> +<i>Physa columnaris</i>. +</td></tr> + +<tr><td valign="top" class="right">222.</td><td> +<i>Cyclostoma Arnoudii</i>. +</td></tr> + +<tr><td valign="top" class="right">223.</td><td> +<i>Rhombus minimus</i>. +</td></tr> + +<tr><td valign="top" class="right">224.</td><td> +<i>Otodus obliquus</i>. +</td></tr> + +<tr><td valign="top" class="right">225.</td><td> +<i>Myliobatis Edwardsii</i>. +</td></tr> + +<tr><td valign="top" class="right">226.</td><td> +Upper jaw of Alligator. +</td></tr> + +<tr><td valign="top" class="right">227.</td><td> +Skull of <i>Odontopteryx toliapicus</i>. +</td></tr> + +<tr><td valign="top" class="right">228.</td><td> +<i>Zeuglodon cetoides</i>. +</td></tr> + +<tr><td valign="top" class="right">229.</td><td> +<i>PalĹ“otherium magnum</i>, restored. +</td></tr> + +<tr><td valign="top" class="right">230.</td><td> +Feet of <i>EquidĹ“</i>. +</td></tr> + +<tr><td valign="top" class="right">231.</td><td> +<i>Anoplothelium commune</i>. +</td></tr> + +<tr><td valign="top" class="right">232.</td><td> +Skull of <i>Dinoceras mirabilis</i>. +</td></tr> + +<tr><td valign="top" class="right">233.</td><td> +<i>Vespertilio Parisiensis</i>. +</td></tr> + +<tr><td valign="top" class="right">234.</td><td> +Miocene Palms. +</td></tr> + +<tr><td valign="top" class="right">235.</td><td> +<i>Platanus aceroides</i>. +</td></tr> + +<tr><td valign="top" class="right">236.</td><td> +<i>Cinnamomum polymorphum</i>. +</td></tr> + +<tr><td valign="top" class="right">237.</td><td> +<i>Textularia Meyeriana</i>. +</td></tr> + +<tr><td valign="top" class="right">238.</td><td> +<i>Scutella subrotunda</i>. +</td></tr> + +<tr><td valign="top" class="right">239.</td><td> +<i>Hyalea Orbignyana</i>. +</td></tr> + +<tr><td valign="top" class="right">240.</td><td> +Tooth of <i>Oxyrhina</i>. +</td></tr> + +<tr><td valign="top" class="right">241.</td><td> +Tooth of <i>Carcharodon</i>. +</td></tr> + +<tr><td valign="top" class="right">242.</td><td> +<i>Andrias Scheuchzeri</i>. +</td></tr> + +<tr><td valign="top" class="right">243.</td><td> +Skull of <i>Brontotherium ingens</i>. +</td></tr> + +<tr><td valign="top" class="right">244.</td><td> +<i>Hippopotamus Sivalensis</i>. +</td></tr> + +<tr><td valign="top" class="right">245.</td><td> +Skull of <i>Sivatherium</i>. +</td></tr> + +<tr><td valign="top" class="right">246.</td><td> +Skull of <i>Deinotherium</i>. +</td></tr> + +<tr><td valign="top" class="right">247.</td><td> +Tooth of <i>Elephas planfrons</i> and of <i>Mastodon +Sivalensis</i>. +</td></tr> + +<tr><td valign="top" class="right">248.</td><td> +Jaw of <i>Pliopithecus</i>. +</td></tr> + +<tr><td valign="top" class="right">249.</td><td> +<i>Rhinoceros Etruscus</i> and <i>R. megarhinus</i>. +</td></tr> + +<tr><td valign="top" class="right">250.</td><td> +Molar tooth of <i>Mastodon Arvernensis</i>. +</td></tr> + +<tr><td valign="top" class="right">251.</td><td> +Molar tooth of <i>Etephas meridionalis</i>. +</td></tr> + +<tr><td valign="top" class="right">252.</td><td> +Molar tooth of <i>Elephas antiquus</i>. +</td></tr> + +<tr><td valign="top" class="right">253.</td><td> +Skull and tooth of <i>Machairodus cultridens</i>. +</td></tr> + +<tr><td valign="top" class="right">254.</td><td> +<i>Pecten Islandicus</i>. +</td></tr> + +<tr><td valign="top" class="right">255.</td><td> +Diagram of high-level and low-level gravels. +</td></tr> + +<tr><td valign="top" class="right">256.</td><td> +Diagrammatic section of Cave. +</td></tr> + +<tr><td valign="top" class="right">257.</td><td> +<i>Dinornis elephantopus</i>. +</td></tr> + +<tr><td valign="top" class="right">258.</td><td> +Skull of <i>Diprotodon</i>. +</td></tr> + +<tr><td valign="top" class="right">259.</td><td> +Skull of <i>Thylacoleo</i>. +</td></tr> + +<tr><td valign="top" class="right">260.</td><td> +Skeleton of <i>Megatherium</i>. +</td></tr> + +<tr><td valign="top" class="right">261.</td><td> +Skeleton of <i>Mylodon</i>. +</td></tr> + +<tr><td valign="top" class="right">262.</td><td> +<i>Glyptodon clavipes</i>. +</td></tr> + +<tr><td valign="top" class="right">263.</td><td> +Skull of <i>Rhinoceros tichorhinus</i>. +</td></tr> + +<tr><td valign="top" class="right">264.</td><td> +Skeleton of <i>Cervus megaceros</i>. +</td></tr> + +<tr><td valign="top" class="right">265.</td><td> +Skull of <i>Bos primigenius</i>. +</td></tr> + +<tr><td valign="top" class="right">266.</td><td> +Skeleton of Mammoth. +</td></tr> + +<tr><td valign="top" class="right">267.</td><td> +Molar tooth of Mammoth. +</td></tr> + +<tr><td valign="top" class="right">268.</td><td> +Skull of <i>Ursus spelĹ“us</i>. +</td></tr> + +<tr><td valign="top" class="right">269.</td><td> +Skull of <i>HyĹ“na spelĹ“a</i>. +</td></tr> + +<tr><td valign="top" class="right">270.</td><td> +Lower jaw of <i>Trogontherium Cuvieri</i>. +</td></tr> + +</table> + +<h2> +<a name="page_xix"><span class="page">Page xix</span></a> +PART I. +</h2> + +<hr /> + +<p class="part"> +PRINCIPLES OF PALÆONTOLOGY. +</p> + +<h2> +<a name="page_1"><span class="page">Page 1</span></a> +<span class="smaller">THE</span><br/> +ANCIENT LIFE-HISTORY<br/> +<span class="smaller">OF</span><br/> +THE EARTH +</h2> + +<hr /> + +<h3>INTRODUCTION.</h3> + +<p class="subtitle"> +THE LAWS OF GEOLOGICAL ACTION. +</p> + +<p class="indent"> +Under the general title of "Geology" are usually included at +least two distinct branches of inquiry, allied to one another in +the closest manner, and yet so distinct as to be largely capable +of separate study. <i>Geology</i>,[1] in its strict sense, is +the science which is concerned with the investigation of the +materials which compose the earth, the methods in which those +materials have been arranged, and the causes and modes of origin +of these arrangements. In this limited aspect, Geology is nothing +more than the Physical Geography of the past, just as Physical +Geography is the Geology of to-day; and though it has to call in +the aid of Physics, Astronomy, Mineralogy, Chemistry, and other +allies more remote, it is in itself a perfectly distinct and +individual study. One has, however, only to cross the threshold +of Geology to discover that the field and scope of the science +cannot be thus rigidly limited to purely physical problems. The +study of the physical development of the earth throughout past +ages brings us at once in contact with the forms of animal and +vegetable life which peopled its surface in bygone epochs, and it +is found impossible adequately to comprehend +<a name="page_2"><span class="page">Page 2</span></a> +the former, unless we possess some knowledge of the latter. +However great its physical advances may be, Geology remains +imperfect till it is wedded with Palæontology,[2] a study +which essentially belongs to the vast complex of the Biological +Sciences, but at the same time has its strictly geological side. +Dealing, as it does, wholly with the consideration of such living +beings as do not belong exclusively to the present order of +things, Palæontology is, in reality, a branch of Natural +History, and may be regarded as substantially the Zoology and +Botany of the past. It is the ancient life-history of the earth, +as revealed to us by the labours of palæontologists, with +which we have mainly to do here; but before entering upon this, +there are some general questions, affecting Geology and +Palæontology alike, which may be very briefly discussed. +</p> + +<p class="footnote"> +[Footnote 1: Gr. <i>ge</i>, the earth; <i>logos</i>, a discourse.] +</p> + +<p class="footnote"> +[Footnote 2: Gr. <i>palaios</i>, ancient; <i>onta</i>, beings; +<i>logos</i>, discourse.] +</p> + +<p class="indent"> +The working geologist, dealing in the main with purely physical +problems, has for his object to determine the material structure +of the earth, and to investigate, as far as may be, the long chain +of causes of which that structure is the ultimate result. No wider +or more extended field of inquiry could be found; but philosophical +geology is not content with this. At all the confines of his +science, the transcendental geologist finds himself confronted +with some of the most stupendous problems which have ever engaged +the restless intellect of humanity. The origin and primæval +constitution of the terrestrial globe, the laws of geologic action +through long ages of vicissitude and development, the origin of +life, the nature and source of the myriad complexities of living +beings, the advent of man, possibly even the future history of +the earth, are amongst the questions with which the geologist +has to grapple in his higher capacity. +</p> + +<p class="indent"> +These are problems which have occupied the attention of philosophers +in every age of the world, and in periods long antecedent to +the existence of a science of geology. The mere existence of +cosmogonies in the religion of almost every nation, both ancient +and modern, is a sufficient proof of the eager desire of the +human mind to know something of the origin of the earth on which +we tread. Every human being who has gazed on the vast panorama +of the universe, though it may have been but with the eyes of +a child, has felt the longing to solve, however imperfectly, +"the riddle of the painful earth," and has, consciously or +unconsciously, elaborated some sort of a theory as to the why and +wherefore of what he sees. Apart from the profound and perhaps +inscrutable problems which lie at the bottom of human existence, +men have in all ages invented +<a name="page_3"><span class="page">Page 3</span></a> +theories to explain the common phenomena of the material universe; +and most of these theories, however varied in their details, turn +out on examination to have a common root, and to be based on the +same elements. Modern geology has its own theories on the same +subject, and it will be well to glance for a moment at the +principles underlying the old and the new views. +</p> + +<p class="indent"> +It has been maintained, as a metaphysical hypothesis, that there +exists in the mind of man an inherent principle, in virtue of +which he believes and expects that what has been, will be; and +that the course of nature will be a continuous and uninterrupted +one. So far, however, from any such belief existing as a necessary +consequence of the constitution of the human mind, the real fact +seems to be that the contrary belief has been almost universally +prevalent. In all old religions, and in the philosophical systems +of almost all ancient nations, the order of the universe has +been regarded as distinctly unstable, mutable, and temporary. +A beginning and an end have always been assumed, and the course +of terrestrial events between these two indefinite points has +been regarded as liable to constant interruption by revolutions +and catastrophes of different kinds, in many cases emanating from +supernatural sources. Few of the more ancient theological creeds, +and still fewer of the ancient philosophies, attained body and +shape without containing, in some form or another, the belief +in the existence of periodical convulsions, and of alternating +cycles of destruction and repair. +</p> + +<p class="indent"> +That geology, in its early infancy, should have become imbued +with the spirit of this belief, is no more than might have been +expected; and hence arose the at one time powerful and +generally-accepted doctrine of "Catastrophism." That the succession +of phenomena upon the globe, whereby the earth's crust had assumed +the configuration and composition which we find it to possess, +had been a discontinuous and broken succession, was the almost +inevitable conclusion of the older geologists. Everywhere in +their study of the rocks they met with apparently impassable +gaps, and breaches of continuity that could not be bridged over. +Everywhere they found themselves conducted abruptly from one system +of deposits to others totally different in mineral character or +in stratigraphical position. Everywhere they discovered that +well-marked and easily recognisable groups of animals and plants +were succeeded, without the intermediation of any obvious lapse +of time, by other assemblages of organic beings of a different +character. Everywhere they found evidence that the earth's crust +<a name="page_4"><span class="page">Page 4</span></a> +had undergone changes of such magnitude as to +render it seemingly irrational to suppose that they could have +been produced by any process now in existence. If we add to the +above the prevalent belief of the time as to the comparative +brevity of the period which had elapsed since the birth of the +globe, we can readily understand the general acceptance of some +form of catastrophism amongst the earlier geologists. +</p> + +<p class="indent"> +As regards its general sense and substance, the doctrine of +catastrophism held that the history of the earth, since first +it emerged from the primitive chaos, had been one of periods +of repose, alternating with catastrophes and cataclysms of a +more or less violent character. The periods of tranquillity were +supposed to have been long and protracted; and during each of them +it was thought that one of the great geological "formations" was +deposited. In each of these periods, therefore, the condition of the +earth was supposed to be much the same as it is now—sediment +was quietly accumulated at the bottom of the sea, and animals and +plants flourished uninterruptedly in successive generations. +Each period of tranquillity, however, was believed to have been, +sooner or later, put an end to by a sudden and awful convulsion +of nature, ushering in a brief and paroxysmal period, in which +the great physical forces were unchained and permitted to spring +into a portentous activity. The forces of subterranean fire, +with their concomitant phenomena of earthquake and volcano, were +chiefly relied upon as the efficient causes of these periods of +spasm and revolution. Enormous elevations of portions of the +earth's crust were thus believed to be produced, accompanied by +corresponding and equally gigantic depressions of other portions. +In this way new ranges of mountains were produced, and previously +existing ranges levelled with the ground, seas were converted into +dry land, and continents buried beneath the ocean—catastrophe +following catastrophe, till the earth was rendered uninhabitable, +and its races of animals and plants were extinguished, never to +reappear in the same form. Finally, it was believed that this +feverish activity ultimately died out, and that the ancient peace +once more came to reign upon the earth. As the abnormal throes +and convulsions began to be relieved, the dry land and sea once +more resumed their relations of stability, the conditions of +life were once more established, and new races of animals and +plants sprang into existence, to last until the supervention +of another fever-fit. +</p> + +<p class="indent"> +Such is the past history of the globe, as sketched for us, in +alternating scenes of fruitful peace and revolutionary destruction, +by the earlier geologists. As before said, we cannot +<a name="page_5"><span class="page">Page 5</span></a> +wonder at the former general acceptance of Catastrophistic +doctrines. Even in the light of our present widely-increased +knowledge, the series of geological monuments remains a broken +and imperfect one; nor can we ever hope to fill up completely +the numerous gaps with which the geological record is defaced. +Catastrophism was the natural method of accounting for these gaps, +and, as we shall see, it possesses a basis of truth. At present, +however, catastrophism may be said to be nearly extinct, and its +place is taken by the modern doctrine of "Continuity" or +"Uniformity"—a doctrine with which the name of Lyell must +ever remain imperishably associated. +</p> + +<p class="indent"> +The fundamental thesis of the doctrine of Uniformity is, that, +in spite of all apparent violations of continuity, the sequence +of geological phenomena has in reality been a regular and +uninterrupted one; and that the vast changes which can be shown +to have passed over the earth in former periods have been the +result of the slow and ceaseless working of the ordinary physical +forces—acting with no greater intensity than they do now, but +acting through enormously prolonged periods. The essential element +in the theory of Continuity is to be found in the allotment of +indefinite time for the accomplishment of the known series of +geological changes. It is obviously the case, namely, that there +are two possible explanations of all phenomena which lie so far +concealed in "the dark backward and abysm of time," that we can +have no direct knowledge of the manner in which they were produced. +We may, on the one hand, suppose them to be the result of some +very powerful cause, acting through a short period of time. That +is Catastrophism. Or, we may suppose them to be caused by a much +weaker force operating through a proportionately prolonged period. +This is the view of the Uniformitarians. It is a question of +<i>energy</i> versus <i>time</i> and it is <i>time</i> which is +the true element of the case. An earthquake may remove a mountain +in the course of a few seconds; but the dropping of the gentle rain +will do the same, if we extend its operations over a millennium. +And this is true of all agencies which are now at work, or ever +have been at work, upon our planet. The Catastrophists, believing +that the globe is but, as it were, the birth of yesterday, were +driven of necessity to the conclusion that its history had been +checkered by the intermittent action of paroxysmal and almost +inconceivably potent forces. The Uniformitarians, on the other +hand, maintaining the "adequacy of existing causes," and denying +that the known physical forces ever acted in past time with greater +intensity than they do at present, are, equally of necessity, +driven to the conclusion that +<a name="page_6"><span class="page">Page 6</span></a> +the world is +truly in its "hoary eld," and that its present state is really +the result of the tranquil and regulated action of known forces +through unnumbered and innumerable centuries. +</p> + +<p class="indent"> +The most important point for us, in the present connection, is +the bearing of these opposing doctrines upon the question, as +to the origin of the existing terrestrial order. On any doctrine +of uniformity that order has been evolved slowly, and, according +to law, from a pre-existing order. Any doctrine of catastrophism, +on the other hand, carries with it, by implication, the belief +that the present order of things was brought about suddenly and +irrespective of any pre-existent order; and it is important to +hold clear ideas as to which of these beliefs is the true one. In +the first place, we may postulate that the world had a beginning, +and, equally, that the existing terrestrial order had a beginning. +However far back we may go, geology does not, and cannot, reach the +actual beginning of the world; and we are, therefore, left simply +to our own speculations on this point. With regard, however, to +the existing terrestrial order, a great deal can be discovered, +and to do so is one of the principal tasks of geological science. +The first steps in the production of that order lie buried in +the profound and unsearchable depths of a past so prolonged as +to present itself to our finite minds as almost in eternity. +The last steps are in the prophetic future, and can be but dimly +guessed at. Between the remote past and the distant future, we +have, however, a long period which is fairly open to inspection; +and in saying a "long" period, it is to be borne in mind that +this term is used in its <i>geological</i> sense. Within this +period, enormously long as it is when measured by human standards, +we can trace with reasonable certainty the progressive march +of events, and can determine the laws of geological action, by +which the present order of things has been brought about. +</p> + +<p class="indent"> +The natural belief on this subject doubtless is, that the world, +such as we now see it, possessed its present form and configuration +from the beginning. Nothing can be more natural than the belief +that the present continents and oceans have always been where +they are now; that we have always had the same mountains and +plains; that our rivers have always had their present courses, +and our lakes their present positions; that our climate has always +been the same; and that our animals and plants have always been +identical with those now familiar to us. Nothing could be more +natural than such a belief, and nothing could be further removed +from the actual truth. On the contrary, a very slight acquaintance +with geology shows us, in the words of Sir John Herschel, that +<a name="page_7"><span class="page">Page 7</span></a> +"the actual configuration of our continents +and islands, the coast-lines of our maps, the direction and +elevation of our mountain-chains, the courses of our rivers, and +the soundings of our oceans, are not things primordially arranged +in the construction of our globe, but results of successive and +complex actions on a former state of things; <i>that</i>, again, +of similar actions on another still more remote; and so on, till +the original and really permanent state is pushed altogether out +of sight and beyond the reach even of imagination; while on the +other hand, a similar, and, as far as we can see, interminable +vista is opened out for the future, by which the habitability +of our planet is secured amid the total abolition on it of the +present theatres of terrestrial life." +</p> + +<p class="indent"> +Geology, then, teaches us that the physical features which now +distinguish the earth's surface have been produced as the ultimate +result of an almost endless succession of precedent changes. +Palæontology teaches us, though not yet in such assured +accents, the same lesson. Our present animals and plants have not +been produced, in their innumerable forms, each as we now know it, +as the sudden, collective, and simultaneous birth of a renovated +world. On the contrary, we have the clearest evidence that some +of our existing animals and plants made their appearance upon the +earth at a much earlier period than others. In the confederation +of animated nature some races can boast of an immemorial antiquity, +whilst others are comparative <i>parvenus</i>. We have also the +clearest evidence that the animals and plants which now inhabit the +globe have been preceded, over and over again, by other different +assemblages of animals and plants, which have flourished in +successive periods of the earth's history, have reached their +culmination, and then have given way to a fresh series of living +beings. We have, finally, the clearest evidence that these successive +groups of animals and plants (faunæ and floræ) are to +a greater or less extent directly connected with one another. Each +group is, to a greater or less extent, the lineal descendant of +the group which immediately preceded it in point of time, and is +more or less fully concerned with giving origin to the group +which immediately follows it. That this law of "evolution" has +prevailed to a great extent is quite certain; but it does not +meet all the exigencies of the case, and it is probable that +its action has been supplemented by some still unknown law of +a different character. +</p> + +<p class="indent"> +We shall have to consider the question of geological "continuity" +again. In the meanwhile, it is sufficient to state that this +doctrine is now almost universally accepted as the basis +<a name="page_8"><span class="page">Page 8</span></a> +of all inquiries, both in the domain of geology and +that of palæontology. The advocates of continuity possess +one immense advantage over those who believe in violent and +revolutionary convulsions, that they call into play only agencies +of which we have actual knowledge. We <i>know</i> that certain +forces are now at work, producing certain modifications in the +present condition of the globe; and we <i>know</i> that these +forces are capable of producing the vastest of the changes which +geology brings under our consideration, provided we assign a time +proportionately vast for their operation. On the other hand, the +advocates of catastrophism, to make good their views, are +compelled to invoke forces and actions, both destructive and +restorative, of which we have, and can have, no direct knowledge. +They endow the whirlwind and the earthquake, the central fire and +the rain from heaven, with powers as mighty as ever imagined in +fable, and they build up the fragments of a repeatedly shattered +world by the intervention of an intermittently active creative +power. +</p> + +<p class="indent"> +It should not be forgotten, however, that from one point of view +there is a truth in catastrophism which is sometimes overlooked +by the advocates of continuity and uniformity. Catastrophism +has, as its essential feature, the proposition that the known +and existing forces of the earth at one time acted with much +greater intensity and violence than they do at present, and they +carry down the period of this excessive action to the commencement +of the present terrestrial order. The Uniformitarians, in effect, +deny this proposition, at any rate as regards any period of the +earth's history of which we have actual cognisance. If, however, +the "nebular hypothesis" of the origin of the universe be well +founded—as is generally admitted—then, beyond question, +the earth is a gradually cooling body, which has at one time been +very much hotter than it is at present. There has been a time, +therefore, in which the igneous forces of the earth, to which we +owe the phenomena of earthquakes and volcanoes, must have been +far more intensely active than we can conceive of from anything +that we can see at the present day. By the same hypothesis, the +sun is a cooling body, and must at one time have possessed a +much higher temperature than it has at present. But increased +heat of the sun would seriously alter the existing conditions +affecting the evaporation and precipitation of moisture on our +earth; and hence the aqueous forces may also have acted at one +time more powerfully than they do now. The fundamental principle +of catastrophism is, therefore, not wholly vicious; and we have +reason to think that there must have been periods—very +<a name="page_9"><span class="page">Page 9</span></a> +remote, it is true, and perhaps unrecorded +in the history of the earth—in which the known physical +forces may have acted with an intensity much greater than direct +observation would lead us to imagine. And this may be believed, +altogether irrespective of those great secular changes by which +hot or cold epochs are produced, and which can hardly be called +"catastrophistic," as they are produced gradually, and are +liable to recur at definite intervals. +</p> + +<p class="indent"> +Admitting, then, that there <i>is</i> a truth at the bottom of +the once current doctrines of catastrophism, still it remains +certain that the history of the earth has been one of <i>law</i> +in all past time, as it is now. Nor need we shrink back affrighted +at the vastness of the conception—the vaster for its very +vagueness—that we are thus compelled to form as to the +duration of <i>geological time</i>. As we grope our way backward +through the dark labyrinth of the ages, epoch succeeds to epoch, +and period to period, each looming more gigantic in its outlines +and more shadowy in its features, as it rises, dimly revealed, from +the mist and vapour of an older and ever-older past. It is useless +to add century to century or millennium to millennium. When we +pass a certain boundary-line, which, after all, is reached very +soon, figures cease to convey to our finite faculties any real +notion of the periods with which we have to deal. The astronomer +can employ material illustrations to give form and substance +to our conceptions of celestial space; but such a resource is +unavailable to the geologist. The few thousand years of which +we have historical evidence sink into absolute insignificance +beside the unnumbered æons which unroll themselves one by +one as we penetrate the dim recesses of the past, and decipher +with feeble vision the ponderous volumes in which the record of +the earth is written. Vainly does the strained intellect seek to +overtake an ever-receding commencement, and toil to gain some +adequate grasp of an apparently endless succession. A beginning +there must have been, though we can never hope to fix its point. +Even speculation droops her wings in the attenuated atmosphere +of a past so remote, and the light of imagination is quenched +in the darkness of a history so ancient. In <i>time</i>, as in +<i>space</i>, the confines of the universe must ever remain +concealed from us, and of the end we know no more than of the +beginning. Inconceivable as is to us the lapse of "geological +time," it is no more than "a mere moment of the past, a mere +infinitesimal portion of eternity." Well may "the human heart, +that weeps and trembles," say, with Richter's pilgrim through +celestial space, "I will go no farther; for the spirit of man +acheth with +<a name="page_10"><span class="page">Page 10</span></a> +this infinity. Insufferable is the glory of God. Let me lie +down in the grave, and hide me from the persecution of the +Infinite, for end, I see, there is none." +</p> + +<h3>CHAPTER I.</h3> + +<p class="subtitle"> +THE SCOPE AND MATERIALS OF PALÆONTOLOGY. +</p> + +<p class="indent"> +The study of the rock-masses which constitute the crust of the +earth, if carried out in the methodical and scientific manner of +the geologist, at once brings us, as has been before remarked, in +contact with the remains or traces of living beings which formerly +dwelt upon the globe. Such remains are found, in greater or less +abundance, in the great majority of rocks; and they are not only of +great interest in themselves, but they have proved of the greatest +importance as throwing light upon various difficult problems in +geology, in natural history, in botany, and in philosophy. Their +study constitutes the science of palæontology; and though it +is possible to proceed to a certain length in geology and zoology +without much palæontological knowledge, it is hardly possible +to attain to a satisfactory general acquaintance with either of +these subjects without having mastered the leading facts of the +first. Similarly, it is not possible to study palæontology +without some acquaintance with both geology and natural history. +</p> + +<p class="indent"> +Palæontology, then, is the science which treats of the living +beings, whether animal or vegetable, which have inhabited the earth +during past periods of its history. Its object is to elucidate, +as far as may be, the structure, mode of existence, and habits +of all such ancient forms of life; to determine their position +in the scale of organised beings; to lay down the geographical +limits within which they flourished; and to fix the period of +their advent and disappearance. It is the ancient life-history +of the earth; and were its record complete, it would furnish +us with a detailed knowledge of the form and relations of all +the animals and plants which have at any period flourished upon +the land-surfaces of the globe or inhabited its waters; it would +enable us to determine precisely their succession in time; and +it would place in our hands an unfailing key to the problems of +evolution. Unfortunately, from causes which will be subsequently +discussed, the palæontological record is extremely imperfect, +and our knowledge is interrupted +<a name="page_11"><span class="page">Page 11</span></a> +by gaps, which not only bear a large proportion to our solid +information, but which in many cases are of such a nature that +we can never hope to fill them up. +</p> + +<p class="indent"> +FOSSILS.—The remains of animals or vegetables which we now +find entombed in the solid rock, and which constitute the working +material of the palæontologist, are termed "fossils,"[3] or +"petrifactions." In most cases, as can be readily understood, +fossils are the actual hard parts of animals and plants which +were in existence when the rock in which they are now found was +being deposited. Most fossils, therefore, are of the nature of +the shells of shell-fish, the skeletons of coral-zoophytes, the +bones of vertebrate animals, or the wood, bark, or leaves of +plants. All such bodies are more or less of a hard consistence +to begin with, and are capable of resisting decay for a longer +or shorter time—hence the frequency with which they occur +in the fossil condition. Strictly speaking, however, by the term +"fossil" must be understood "any body, <i>or the traces of the +existence of any body</i>, whether animal or vegetable, which +has been buried in the earth by natural causes" (Lyell). We shall +find, in fact, that many of the objects which we have to study +as "fossils" have never themselves actually formed parts of any +animal or vegetable, though they are due to the former existence +of such organisms, and indicate what was the nature of these. +Thus the footprints left by birds, or reptiles, or quadrupeds +upon sand or mud, are just as much proofs of the former existence +of these animals as would be bones, feathers, or scales, though +in themselves they are inorganic. Under the head of fossils, +therefore, come the footprints of air-breathing vertebrate animals; +the tracks, trails, and burrows of sea-worms, crustaceans, or +molluscs; the impressions left on the sand by stranded jelly-fishes; +the burrows in stone or wood of certain shell-fish; the "moulds" +or "casts" of shells, corals, and other organic remains; and +various other bodies of a more or less similar nature. +</p> + +<p class="footnote"> +[Footnote 3: Lat. <i>fossus</i>, dug up.] +</p> + +<p class="indent"> +FOSSILISATION.—The term "fossilisation" is applied to all those +processes through which the remains of organised beings may pass +in being converted into fossils. These processes are numerous +and varied; but there are three principal modes of fossilisation +which alone need be considered here. In the first instance, the +fossil is to all intents and purposes an actual portion of the +original organised being—such as a bone, a shell, or a piece +of wood. In some rare instances, as in the case of the body of +the Mammoth discovered embedded in ice at the mouth of the Lena +in Siberia, the fossil may be preserved +<a name="page_12"><span class="page">Page 12</span></a> +almost precisely in its original condition, and even with its soft +parts uninjured. More commonly, certain changes have taken place +in the fossil, the principal being the more or less total removal +of the organic matter originally present. Thus bones become light +and porous by the removal of their gelatine, so as to cleave to +the tongue on being applied to that organ; whilst shells become +fragile, and lose their primitive colours. In other cases, though +practically the real body it represents, all the cavities of the +fossil, down to its minutest recesses, may have become infiltrated +with mineral matter. It need hardly be added, that it is in the +more modern rocks that we find the fossils, as a rule, least +changed from their former condition; but the original structure +is often more or less completely retained in some of the fossils +from even the most ancient formations. +</p> + +<p class="indent"> +In the second place, we very frequently meet with fossils in +the state of "casts" or moulds of the original organic body. +What occurs in this case will be readily understood if we imagine +any common bivalve shell, as an Oyster, or Mussel, or Cockle, +embedded in clay or mud. If the clay were sufficiently soft and +fluid, the first thing would be that it would gain access to the +interior of the shell, and would completely fill up the space +between the valves. The pressure, also, of the surrounding matter +would insure that the clay would everywhere adhere closely to +the exterior of the shell. If now we suppose the clay to be in +any way hardened so as to be converted into stone, and if we were +to break up the stone, we should obviously have the following +state of parts. The clay which filled the shell would form an +accurate cast of the <i>interior</i> of the shell, and the clay +outside would give us an exact impression or cast of +the <i>exterior</i> of the shell (fig. 1). We should have, then, + +<span style="float: left; margin: 4px; width: 260px; + font-size: smaller; text-align: justify;"> +<img src="images/fig001.jpg" width="252" height="179" alt="Fig. 1" /> +<br /> +Fig. 1.—<i>Trigonia longa</i>, showing casts of the exterior +and interior of the shell.—Cretaceous (Neocomian). +</span> + +two casts, an interior and an exterior, and the two would be +very different to one another, since the inside of a shell is +very unlike the outside. In the case, in fact, of many univalve +shells, the interior cast or "mould" is so unlike the exterior +cast, or unlike the shell itself, that it may be difficult to +determine the true origin of the former. +</p> + +<p class="indent"> +It only remains to add that there is sometimes a further +complication. If the rock be very porous and permeable by +<a name="page_13"><span class="page">Page 13</span></a> +water, it may happen that the original shell is +entirely dissolved away, leaving the interior cast loose, +like the kernel of a nut, within the case formed by the exterior +cast. Or it may happen that subsequent to the attainment of this +state of things, the space thus left vacant between the interior +and exterior cast—the space, that is, formerly occupied by +the shell itself—may be filled up by some foreign mineral +deposited there by the infiltration of water. In this last case +the splitting open of the rock would reveal an interior cast, +an exterior cast, and finally a body which would have the exact +form of the original shell, but which would be really a much +later formation, and which would not exhibit under the microscope +the minute structure of shell. +</p> + +<p class="indent"> +In the third class of cases we have fossils which present with +the greatest accuracy the external form, and even sometimes the +internal minute structure, of the original organic body, but +which, nevertheless, are not themselves truly organic, but have +been formed by a "replacement" of the particles of the primitive +organism by some mineral substance. The most elegant example of +this is afforded by fossil wood which has been "silicified" or +converted into flint (<i>silex</i>). In such cases we have fossil +wood which presents the rings of growth and fibrous structure of +recent wood, and which under the microscope exhibits the minutest +vessels which characterise ligneous tissue, together with the even +more minute markings of the vessels (fig. 2). The whole, however, + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 268px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig002.jpg" width="260" height="260" alt="Fig. 2" /> +<br /> +Fig. 2.—Microscopic section of the silicified +wood of a Conifer (<i>Sequoia</i>) cut in the long direction +of the fibres. Post-tertiary? Colorado. (Original.) +</span> + +<span style="width: 271px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig003.jpg" width="263" height="260" alt="Fig. 3" /> +<br /> +Fig. 3.—Microscopic section of the wood of the +common Larch (<i>Abies larix</i>), cut in the long direction +of the fibres. In both the fresh and the fossil wood (fig. 2) +are seen the discs characteristic of coniferous wood. (Original.) +</span> + +</span> + +instead of being composed of the original carbonaceous matter of +the wood, is now converted into flint. The only explanation that +can be given +<a name="page_14"><span class="page">Page 14</span></a> +of this by no means rare phenomenon, is that the wood must have +undergone a slow process of decay in water charged with silica or +flint in solution. As each successive particle of wood was removed +by decay, its place was taken by a particle of flint deposited +from the surrounding water, till ultimately the entire wood was +silicified. The process, therefore, resembles what would take +place if we were to pull down a house built of brick by successive +bricks, replacing each brick as removed by a piece of stone of +precisely the same size and form. The result of this would be that +the house would retain its primitive size, shape, and outline, but +it would finally have been converted from a house of brick into a +house of stone. Many other fossils besides wood—such as shells, +corals, sponges, &c.—are often found silicified; and this +may be regarded as the commonest form of fossilisation by replacement. +In other cases, however, though the principle of the process is the +same, the replacing substance may be iron pyrites, oxide of iron, +sulphur, malachite, magnesite, talc, &c.; but it is rarely that +the replacement with these minerals is so perfect as to preserve the +more delicate details of internal structure. +</p> + +<h3>CHAPTER II.</h3> + +<p class="subtitle"> +THE FOSSILIFEROUS ROCKS. +</p> + +<p class="indent"> +Fossils are found in rocks, though not universally or promiscuously; +and it is therefore necessary that the palæontologist should +possess some acquaintance with, at any rate, those rocks which +yield organic remains, and which are therefore said to be +"<i>fossiliferous</i>." In geological language, all the materials +which enter into the composition of the solid crust of the earth, +be their texture what it may—from the most impalpable mud to +the hardest granite—are termed "rocks;" and for our present +purpose we may divide these into two great groups. In the first +division are the <i>Igneous Rocks</i>—such as the lavas and +ashes of volcanoes—which are formed within the body of the +earth itself, and which owe their structure and origin to the +action of heat. The Igneous Rocks are formed primarily below the +surface of the earth, which they only reach as the result of +volcanic action; they are generally destitute of distinct +"stratification," or arrangement in successive layers; and they +do not contain fossils, except in the comparatively +<a name="page_15"><span class="page">Page 15</span></a> +rare instances where volcanic ashes have enveloped +animals or plants which were living in the sea or on the land in +the immediate vicinity of the volcanic focus. The second great +division of rocks is that of the <i>Fossiliferous, Aqueous</i>, +or <i>Sedimentary</i> Rocks. These are formed at the surface of +the earth, and, as implied by one of their names, are invariably +deposited in water. They are produced by vital or chemical action, +or are formed from the "sediment" produced by the disintegration +and reconstruction of previously existing rocks, without previous +solution; they mostly contain fossils; and they are arranged in +distinct layers or "strata." The so-called "aerial" rocks which, +like beds of blown sand, have been formed by the action of the +atmosphere, may also contain fossils; but they are not of such +importance as to require special notice here. +</p> + +<p class="indent"> +For all practical purposes, we may consider that the Aqueous +Rocks are the natural cemetery of the animals and plants of bygone +ages; and it is therefore essential that the palæontological +student should be acquainted with some of the principal facts as +to their physical characters, their minute structure and mode of +origin, their chief varieties, and their historical succession. +</p> + +<p class="indent"> +The Sedimentary or Fossiliferous Rocks form the greater portion of +that part of the earth's crust which is open to our examination, and +are distinguished by the fact that they are regularly "stratified" or +arranged in distinct and definite layers or "strata." These layers +may consist of a single material, as in a block of sandstone, or +they may consist of different materials. When examined on a large +scale, they are always found to consist of alternations of layers +of different mineral composition. We may examine any given area, +and find in it nothing but one kind of rock—sandstone, perhaps, +or limestone. In all cases, however, if we extend our examination +sufficiently far, we shall ultimately come upon different rocks; +and, as a general rule, the thickness of any particular set of +beds is comparatively small, so that different kinds of rock +alternate with one another in comparatively small spaces. +</p> + +<p class="indent"> +As regards the origin of the Sedimentary Rocks, they are for +the most part "derivative" rocks, being derived from the wear +and tear of pre-existent rocks. Sometimes, however, they owe +their origin to chemical or vital action, when they would more +properly be spoken of simply as Aqueous Rocks. As to their mode +of deposition, we are enabled to infer that the materials which +compose them have formerly been spread out by the action of water, +from what we see going on every day +<a name="page_16"><span class="page">Page 16</span></a> +at the mouths of our great +rivers, and on a smaller scale wherever there is running water. +Every stream, where it runs into a lake or into the sea, carries + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 575px; + font-size: smaller; text-align: center;"> +<img src="images/fig004.jpg" width="571" height="429" alt="Fig. 4" /> +<br /> +Fig. 4.—Sketch of Carboniferous strata at Kinghorn, in Fife, +showing stratified beds (limestone and shales) surmounted by an +unstratified mass of trap. (Original.) +</span> +</span> + +with it a burden of mud, sand, and rounded pebbles, derived from +the waste of the rocks which form its bed and banks. When these +materials cease to be impelled by the force of the moving water, +they sink to the bottom, the heaviest pebbles, of course, sinking +first, the smaller pebbles and sand next, and the finest mud +last. Ultimately, therefore, as might have been inferred upon +theoretical grounds, and as is proved by practical experience, +every lake becomes a receptacle for a series of stratified rocks +produced by the streams flowing into it. These deposits may vary +in different parts of the lake, according as one stream brought +down one kind of material and another stream contributed another +material; but in all cases the materials will bear ample evidence +that they were produced, sorted, and deposited by running water. +The finer beds of clay or sand will all be arranged in thicker or +thinner layers or laminæ; and if there are any beds of pebbles +these will all be rounded or smooth, just like the water-worn +pebbles of any brook-course. In all probability, also, we should +find in some of the beds the remains +<a name="page_17"><span class="page">Page 17</span></a> +of fresh-water shells or plants or other organisms which inhabited +the lake at the time these beds were being deposited. +</p> + +<p class="indent"> +In the same way large rivers—such as the Ganges or +Mississippi—deposit all the materials which they bring down +at their mouths, forming in this way their "deltas." Whenever +such a delta is cut through, either by man or by some channel of +the river altering its course, we find that it is composed of a +succession of horizontal layers or strata of sand or mud, varying +in mineral composition, in structure, or in grain, according to +the nature of the materials brought down by the river at different +periods. Such deltas, also, will contain the remains of animals +which inhabit the river, with fragments of the plants which grew +on its banks, or bones of the animals which lived in its basin. +</p> + +<p class="indent"> +Nor is this action confined, of course, to large rivers only, +though naturally most conspicuous in the greatest bodies of water. +On the contrary, all streams, of whatever size, are engaged in +the work of wearing down the dry land, and of transporting the +materials thus derived from higher to lower levels, never resting +in this work till they reach the sea. +</p> + +<p class="indent"> +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 575px; + font-size: smaller; text-align: center;"> +<img src="images/fig005.jpg" width="571" height="385" alt="Fig. 5" /> +Fig. 5.—Diagram to illustrate the formation of sedimentary +deposits at the point where a river debouches into the sea. +</span> +</span> +Lastly, the sea itself—irrespective of the materials +delivered into it by rivers—is constantly preparing fresh +<a name="page_18"><span class="page">Page 18</span></a> +by its own action. +Upon every coast-line the sea is constantly eating back into +the land and reducing its component rocks to form the shingle +and sand which we see upon every shore. The materials thus +produced are not, however, lost, but are ultimately deposited +elsewhere in the form of new stratified accumulations, in which +are buried the remains of animals inhabiting the sea at the time. +</p> + +<p class="indent"> +Whenever, then, we find anywhere in the interior of the land +any series of beds having these characters—composed, that is, +of distinct layers, the particles of which, both large and small, +show distinct traces of the wearing action of water—whenever +and wherever we find such rocks, we are justified in assuming that +they have been deposited by water in the manner above mentioned. +Either they were laid down in some former lake by the combined +action of the streams which flowed into it; or they were deposited +at the mouth of some ancient river, forming its delta; or they +were laid down at the bottom of the ocean. In the first two cases, +any fossils which the beds might contain would be the remains +of fresh-water or terrestrial organisms. In the last case, the +majority, at any rate, of the fossils would be the remains of +marine animals. +</p> + +<p class="indent"> +The term "formation" is employed by geologists to express "any +group of rocks which have some character in common, whether of +origin, age, or composition" (Lyell); so that we may speak of +stratified and unstratified formations, aqueous or igneous +formations, fresh-water or marine formations, and so on. +</p> + +<h4>CHIEF DIVISIONS OF THE AQUEOUS ROCKS.</h4> + +<p class="indent"> +The Aqueous Rocks may be divided into two great sections, the +Mechanically-formed and the Chemically-formed, including under +the last head all rocks which owe their origin to vital action, +as well as those produced by ordinary chemical agencies. +</p> + +<p class="indent"> +A. MECHANICALLY-FORMED ROCKS.—These are all those Aqueous +Rocks of which we can obtain proofs that their particles have +been mechanically transported to their present situation. Thus, +if we examine a piece of <i>conglomerate</i> or puddingstone, we +find it to be composed of a number of rounded pebbles embedded +in an enveloping matrix or paste, which is usually of a sandy +nature, but may be composed of carbonate of lime (when the rock +is said to be a "calcareous conglomerate"). The pebbles in all +conglomerates are worn and rounded by the action of water in motion, +and thus show +<a name="page_19"><span class="page">Page 19</span></a> +that they have been subjected +to much mechanical attrition, whilst they have been mechanically +transported for a greater or less distance from the rock of which +they originally formed part. The analogue of the old conglomerates +at the present day is to be found in the great beds of shingle +and gravel which are formed by the action of the sea on every +coast-line, and which are composed of water-worn and well-rounded +pebbles of different sizes. A <i>breccia</i> is a mechanically-formed +rock, very similar to a conglomerate, and consisting of larger or +smaller fragments of rock embedded in a common matrix. The fragments, +however, are in this case all more or less angular, and are not +worn or rounded. The fragments in breccias may be of large size, +or they may be comparatively small (fig. 6); and the matrix may + +<span style="float: right; margin: 4px; width: 268px; + font-size: smaller; text-align: justify;"> +<img src="images/fig006.jpg" width="260" height="258" alt="Fig. 6" /> +<br /> +Fig. 6.—Microscopic section of a calcareous breccia in the +Lower Silurian (Coniston Limestone) of Shap Wells, Westmoreland. +The fragments are all of small size, and consist of angular +pieces of transparent quartz, volcanic ashes, and limestone +embedded in a matrix of crystalline limestone. (Original.) +</span> + +be composed of sand (arenaceous) or of carbonate of lime +(calcareous). In the case of an ordinary sandstone, again, we +have a rock which may be regarded as simply a very fine-grained +conglomerate or breccia, being composed of small grains of sand +(silica), sometimes rounded, sometimes more or less angular, +cemented together by some such substance as oxide of iron, silicate +of iron, or carbonate of lime. A sandstone, therefore, like a +conglomerate is a mechanically-formed rock, its component grams +being equally the result of mechanical attrition and having equally +been transported from a distance; and the same is true of the +ordinary sand of the sea-shore, which is nothing more than an +unconsolidated sandstone. Other so-called sands and sandstones, +though equally mechanical in their origin, are truly calcareous in +their nature, and are more or less entirely composed of carbonate +of lime. Of this kind are the shell-sand so common on our coasts, +and the coral-sand which is so largely formed in the neighbourhood +of coral-reefs. In these cases the rock is composed of fragments +of the skeletons of shellfish, and numerous other marine animals, +together, in many instances, with the remains of certain sea-weeds +(<i>Corallines</i>, <i>Nullipores</i>, &c,) which are endowed +with the power of secreting +<a name="page_20"><span class="page">Page 20</span></a> +carbonate of lime from the sea-water. Lastly, in certain rocks +still finer in their texture than sandstones, such as the various +mud-rocks and shales, we can still recognise a mechanical source +and origin. If slices of any of these rocks sufficiently thin to +be transparent are examined under the microscope, it will be found +that they are composed of minute grains of different sizes, which +are all more or less worn and rounded, and which clearly show, +therefore, that they have been subjected to mechanical attrition. +</p> + +<p class="indent"> +All the above-mentioned rocks, then, are <i>mechanically-formed</i> +rocks; and they are often spoken of as "Derivative Rocks," in +consequence of the fact that their particles can be shown to have +been mechanically <i>derived</i> from other pre-existent rocks. +It follows from this that every bed of any mechanically-formed +rock is the measure and equivalent of a corresponding amount of +destruction of some older rock. It is not necessary to enter +here into a minute account of the subdivisions of these rocks, but +it may be mentioned that they may be divided into two principal +groups, according to their chemical composition. In the one group +we have the so-called <i>Arenaceous</i> (Lat. <i>arena</i>, sand) +or <i>Siliceous</i> Rocks, which are essentially composed of +larger or smaller grains of flint or silica. In this group are +comprised ordinary sand, the varieties of sandstone and grit, and +most conglomerates and breccias. We shall, however, afterwards +see that some siliceous rocks are of organic origin. In the second +group are the so-called <i>Argillaceous</i> (Lat. <i>argilla</i>, +clay) Rocks, which contain a larger or smaller amount of clay or +hydrated silicate of alumina in their composition. Under this +head come clays, shales, marls, marl-slate, clay-slates, and +most flags and flagstones. +</p> + +<p class="indent"> +B. CHEMICALLY-FORMED ROCKS.—In this section are comprised all +those Aqueous or Sedimentary Rocks which have been formed by +chemical agencies. As many of these chemical agencies, however, +are exerted through the medium of living beings, whether animals +or plants, we get into this section a number of what may be called +"<i>organically-formed rocks</i>." These are of the greatest +possible importance to the palæontologist, as being to a +greater or less extent composed of the actual remains of animals or +vegetables, and it will therefore be necessary to consider their +character and structure in some detail. +</p> + +<p class="indent"> +By far the most important of the chemically-formed rocks are +the so-called <i>Calcareous Rocks</i> (Lat. <i>calx</i>, lime), +comprising all those which contain a large proportion of carbonate +<a name="page_21"><span class="page">Page 21</span></a> +of lime, or are wholly composed of this substance. +Carbonate of lime is soluble in water holding a certain amount +of carbonic acid gas in solution; and it is, therefore, found in +larger or smaller quantity dissolved in all natural waters, both +fresh and salt, since these waters are always to some extent +charged with the above-mentioned solvent gas. A great number of +aquatic animals, however, together with some aquatic plants, are +endowed with the power of separating the lime thus held in +solution in the water, and of reducing it again to its solid +condition. In this way shell-fish, crustaceans, sea-urchins, +corals, and an immense number of other animals, are enabled to +construct their skeletons; whilst some plants form hard structures +within their tissues in a precisely similar manner. We do meet +with some calcareous deposits, such as the "stalactites" and +"stalagmites" of caves, the "calcareous tufa" and "travertine" +of some hot springs, and the spongy calcareous deposits of +so-called "petrifying springs," which are purely chemical in +their origin, and owe nothing to the operation of living beings. +Such deposits are formed simply by the precipitation of carbonate +of lime from water, in consequence of the evaporation from the +water of the carbonic acid gas which formerly held the lime in +solution; but, though sometimes forming masses of considerable +thickness and of geological importance, they do not concern us +here. Almost all the limestones which occur in the series of the +stratified rocks are, primarily at any rate, of <i>organic</i> +origin, and have been, directly or indirectly, produced by the +action of certain lime-making animals or plants, or both combined. +The presumption as to all the calcareous rocks, which cannot be +clearly shown to have been otherwise produced, is that they are +thus organically formed; and in many cases this presumption can +be readily reduced to a certainty. There are many varieties of +the calcareous rocks, but the following are those which are of +the greatest importance:— +</p> + +<p class="indent"> +<i>Chalk</i> is a calcareous rock of a generally soft and +pulverulent texture, and with an earthy fracture. It varies in +its purity, being sometimes almost wholly composed of carbonate +of lime, and at other times more or less intermixed with foreign +matter. Though usually soft and readily reducible to powder, +chalk is occasionally, as in the north of Ireland, tolerably +hard and compact; but it never assumes the crystalline aspect +and stony density of limestone, except it be in immediate contact +with some mass of igneous rock. By means of the microscope, the +true nature and mode of formation of chalk can be determined +with the greatest ease. In the case of the harder varieties, the +examination can be conducted by means of +<a name="page_22"><span class="page">Page 22</span></a> +slices ground down to a thinness sufficient to render them +transparent; but in the softer kinds the rock must be disintegrated +under water, and the <i>dĂ©bris</i> examined microscopically. +When investigated by either of these methods, chalk is found to be +a genuine organic rock, being composed of the shells or hard parts +of innumerable marine animals of different kinds, some entire, +some fragmentary, cemented together by a matrix of very finely +granular carbonate of lime. Foremost amongst the animal remains +which so largely compose chalk are the shells of the minute +creatures which will be subsequently spoken of under the name of +<i>Foraminifera</i> (fig. 7), and which, in spite of their + +<span style="float: left; margin: 4px; width: 271px; + font-size: smaller; text-align: justify;"> +<img src="images/fig007.jpg" width="263" height="261" alt="Fig. 7" /> +<br /> +Fig. 7.—Section of Gravesend Chalk, examined by transmitted +light and highly magnified. Besides the entire shells of +<i>Globigerina</i>, <i>Rotalia</i>, and <i>Textularia</i>, +numerous detached chambers of <i>Globigerina</i> are seen. +(Original.) +</span> + +microscopic dimensions, play a more important part in the process +of lime-making than perhaps any other of the larger inhabitants of +the ocean. +</p> + +<p class="indent"> +As chalk is found in beds of hundreds of feet in thickness, +and of great purity, there was long felt much difficulty in +satisfactorily accounting for its mode of formation and origin. +By the researches of Carpenter, Wyville Thomson, Huxley, Wallich, +and others, it has, however, been shown that there is now forming, +in the profound depths of our great oceans, a deposit which is +in all essential respects identical with chalk, and which is +generally known as the "Atlantic ooze," from its having been first +discovered in that sea. This ooze is found at great depths (5000 +to over 15,000 feet) in both the Atlantic and Pacific, covering +enormously large areas of the sea-bottom, and it presents itself +as a whitish-brown, sticky, impalpable mud, very like greyish +chalk when dried. Chemical examination shows that the ooze is +composed almost wholly of carbonate of lime, and microscopical +examination proves it to be of organic origin, and to be made up +of the remains of living beings. The principal forms of these +belong to the <i>Foraminifera</i>, and the commonest of these +are the irregularly-chambered shells of <i>Globigerina</i>, +absolutely indistinguishable from the <i>GlobigerinĹ“</i> +which are so largely present in the chalk (fig. 8). Along with +these occur fragments of the skeletons of other larger creatures, +<a name="page_23"><span class="page">Page 23</span></a> +and a certain proportion of the flinty cases of minute animal +and vegetable organisms (<i>Polycystina</i> and <i>Diatoms</i>). + +<span style="float: right; margin: 4px; width: 275px; + font-size: smaller; text-align: justify;"> +<img src="images/fig008.jpg" width="263" height="258" alt="Fig. 8" /> +<br /> +Fig. 8.—Organisms in the Atlantic Ooze, chiefly +<i>Foraminifera</i> (<i>Globigerina</i> and <i>Textularia</i>), +with <i>Polycystina</i> and sponge-spicules; highly magnified. +(Original.) +</span> + +Though many of the minute animals, the hard parts of which form +the ooze, undoubtedly live at or near the surface of the sea, +others, probably, really live near the bottom; and the ooze itself +forms a congenial home for numerous sponges, sea-lilies, and +other marine animals which flourish at great depths in the sea. +There is thus established an intimate and most interesting +parallelism between the chalk and the ooze of modern oceans. +Both are formed essentially in the same way, and the latter only +requires consolidation to become actually converted into chalk. Both +are fundamentally organic deposits, apparently requiring a great +depth of water for their accumulation, and mainly composed of the +remains of <i>Foraminifera</i>, together with the entire or broken +skeletons of other marine animals of greater dimensions. It is to be +remembered, however, that the ooze, though strictly representative +of the chalk, cannot be said in any proper sense to be actually +<i>identical</i> with the formation so called by geologists. A +great lapse of time separates the two, and though composed of +the remains of representative classes or groups of animals, it +is only in the case of the lowly-organised <i>GlobigerinĹ“</i>, +and of some other organisms of little higher grade, that we find +absolutely the same kinds or species of animals in both. +</p> + +<p class="indent"> +<i>Limestone</i>, like chalk, is composed of carbonate of lime, +sometimes almost pure, but more commonly with a greater or less +intermixture of some foreign material, such as alumina or silica. +The varieties of limestone are almost innumerable, but the great +majority can be clearly proved to agree with chalk in being +essentially of organic origin, and in being more or less largely +composed of the remains of living beings. In many instances the +organic remains which compose limestone are so large as to be +readily visible to the naked eye, and the rock is at once seen to +be nothing more than an agglomeration of the skeletons, generally +fragmentary, of certain marine animals, cemented together by a +matrix of carbonate of lime. +<a name="page_24"><span class="page">Page 24</span></a> +This is the +case, for example, with the so-called "Crinoidal Limestones" and +"Encrinital Marbles" with which the geologist is so familiar, +especially as occurring in great beds amongst the older formations +of the earth's crust. These are seen, on weathered or broken +surfaces, or still better in polished slabs (fig. 9), to be + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 493px; + font-size: smaller; text-align: justify;"> +<img src="images/fig009.jpg" width="485" height="360" alt="Fig. 9" /> +<br /> +Fig. 9.—Slab of Crinoidal marble, from the Carboniferous +limestone of Dent, in Yorkshire, of the natural size. The polished +surface intersects the columns of the Crinoids at different angles, +and thus gives rise to varying appearances. (Original.) +</span> +</span> + +composed more or less exclusively of the +broken stems and detached plates of sea-lilies (<i>Crinoids</i>). +Similarly, other limestones are composed almost entirely of the +skeletons of corals; and such old coralline limestones can readily +be paralleled by formations which we can find in actual course of +production at the present day. We only need to transport ourselves +to the islands of the Pacific, to the West Indies, or to the Indian +Ocean, to find great masses of lime formed similarly by living +corals, and well known to everyone under the name of "coral-reefs." +Such reefs are often of vast extent, both superficially and in +vertical thickness, and they fully equal in this respect any of +the coralline limestones of bygone ages. Again, we find other +limestones—such as the celebrated "Nummulitic Limestone" +(fig. 10), which sometimes attains a thickness of some thousands +of feet—which are almost entirely made up of the shells of +<i>Foraminifera</i>. In the case of the "Nummulitic Limestone," +just mentioned, these shells are of large size, varying from the +up to that of a +<a name="page_25"><span class="page">Page 25</span></a> +florin. There are, however, as we shall see, many other limestones, +which are likewise largely made up of <i>Foraminifera</i>, but in + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 414px; + font-size: smaller; text-align: center;"> +<img src="images/fig010.jpg" width="406" height="319" alt="Fig. 10" /> +<br/> +Fig. 10.—Piece of Nummulitic Limestone from the Great Pyramid. +Of the natural size. (Original.) +</span> +</span> + +which the shells are very much more minute, and would hardly be +seen at all without the microscope. +</p> + +<p class="indent"> +We may, in fact, consider that the great agents in the production +of limestones in past ages have been animals belonging to the +<i>Crinoids</i>, the <i>Corals</i>, and the <i>Foraminifera</i>. +At the present day, the Crinoids have been nearly extinguished, +and the few known survivors seem to have retired to great depths +in the ocean; but the two latter still actively carry on the +work of lime-making, the former being very largely helped in +their operations by certain lime-producing marine plants +(<i>Nullipores</i> and <i>Corallines</i>). We have to remember, +however, that though the limestones, both ancient and modern, +that we have just spoken of, are truly organic, they are not +necessarily formed out of the remains of animals which actually +lived on the precise spot where we now find the limestone itself. +We may find a crinoidal limestone, which we can show to have +been actually formed by the successive growth of generations +of sea-lilies <i>in place</i>; but we shall find many others in +which the rock is made up of innumerable fragments of the skeletons +of these creatures, which have been clearly worn and rubbed by +the sea-waves, and which have been mechanically transported to +their present site. In the same way, a limestone may be shown +to have been an actual coral-reef, by the fact that we find in +it great masses of coral, growing in their natural position, and +<a name="page_26"><span class="page">Page 26</span></a> +exhibiting plain proofs that they were +simply quietly buried by the calcareous sediment as they grew; but +other limestones may contain only numerous rolled and water-worn +fragments of corals. This is precisely paralleled by what we can +observe in our existing coral-reefs. Parts of the modern +coral-islands and coral-reefs are really made up of corals, dead +or alive, which actually grew on the spot where we now find them; +but other parts are composed of a limestone-rock ("coral-rock"), +or of a loose sand ("coral-sand"), which is organic in the sense +that it is composed of lime formed by living beings, but which, +in truth, is composed of fragments of the skeletons of these living +beings, mechanically transported and heaped together by the sea. To +take another example nearer home, we may find great accumulations +of calcareous matter formed <i>in place</i>, by the growth of +shell-fish, such as oysters or mussels; but we can also find +equally great accumulations on many of our shores in the form of +"shell-sand," which is equally composed of the shells of molluscs, +but which is formed by the trituration of these shells by the +mechanical power of the sea-waves. We thus see that though all +these limestones are primarily organic, they not uncommonly become +"mechanically-formed" rocks in a secondary sense, the materials +of which they are composed being formed by living beings, but +having been mechanically transported to the place where we now +find them. +</p> + +<p class="indent"> +Many limestones, as we have seen, are composed of large and +conspicuous organic remains, such as strike the eye at once. +Many others, however, which at first sight appear compact, more +or less crystalline, and nearly devoid of traces of life, are +found, when properly examined, to be also composed of the remains +of various organisms. All the commoner limestones, in fact, from +the Lower Silurian period onwards, can be easily proved to be thus +<i>organic</i> rocks, if we investigate weathered or polished +surfaces with a lens, or, still better, if we cut thin slices of +the rock and grind these down till they are transparent. When +thus examined, the rock is usually found to be composed of +innumerable entire or fragmentary fossils, cemented together +by a granular or crystalline matrix of carbonate of lime (figs. +11 and 12). When the matrix is granular, the rock is precisely +similar to chalk, except that it is harder and less earthy in +texture, whilst the fossils are only occasionally referable to +the <i>Foraminifera</i>. In other cases, the matrix is more or +less crystalline, and when this crystallisation has been carried +to a great extent, the original organic nature of the rock may +be greatly or completely obscured +<a name="page_27"><span class="page">Page 27</span></a> +thereby. Thus, in limestones +which have been greatly altered or "metamorphosed" by the combined +action of heat and pressure, all traces of organic remains become + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 268px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig011.jpg" width="260" height="258" alt="Fig. 11" /> +<br /> +Fig. 11.—Section of Carboniferous Limestone from Spergen Hill, +Indiana, U.S., showing numerous large-sized <i>Foraminifera</i> +(<i>Endothyra</i>) and a few oolitic grains; magnified. +(Original.) +</span> + +<span style="width: 268px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig012.jpg" width="263" height="258" alt="Fig. 12" /> +<br /> +Fig 12.—Section of Coniston Limestone (Lower Silurian) from +Keisler, Westmoreland; magnified. The matrix is very coarsely +crystalline, and the included organic remains are chiefly stems +of Crinoids. (Original.) +</span> + +</span> + +annihilated, and the rock becomes completely crystalline throughout. +This, for example, is the case with the ordinary white "statuary +marble," slices of which exhibit under the microscope nothing but +an aggregate of beautifully transparent crystals of carbonate +of lime, without the smallest traces of fossils. There are also +other cases, where the limestone is not necessarily highly +crystalline, and where no metamorphic action in the strict sense +has taken place, in which, nevertheless, the microscope fails +to reveal any evidence that the rock is organic. Such cases are +somewhat obscure, and doubtless depend on different causes in +different instances; but they do not affect the important +generalisation that limestones are fundamentally the product +of the operation of living beings. This fact remains certain; +and when we consider the vast superficial extent occupied by +calcareous deposits, and the enormous collective thickness of +these, the mind cannot fail to be impressed with the immensity of +the period demanded for the formation of these by the agency of +such humble and often microscopic creatures as Corals, Sea-lilies, +Foraminifers, and Shell-fish. +</p> + +<p class="indent"> +Amongst the numerous varieties of limestone, a few are of such +interest as to deserve a brief notice. <i>Magnesian limestone</i> +or <i>dolomite</i>, differs from ordinary limestone in containing +a certain proportion of carbonate of magnesia along with the +carbonate of lime. The typical dolomites contain a large proportion +<a name="page_28"><span class="page">Page 28</span></a> +of carbonate of magnesia, and are highly +crystalline. The ordinary magnesian limestones (such as those of +Durham in the Permian series, and the Guelph Limestones of North +America in the Silurian series) are generally of a yellowish, +buff, or brown colour, with a crystalline or pearly aspect, +effervescing with acid much less freely than ordinary limestone, +exhibiting numerous cavities from which fossils have been dissolved +out, and often assuming the most varied and singular forms in +consequence of what is called "concretionary action." Examination +with the microscope shows that these limestones are composed of an +aggregate of minute but perfectly distinct crystals, but that minute +organisms of different kinds, or fragments of larger fossils, are +often present as well. Other magnesian limestones, again, exhibit +no striking external peculiarities by which the presence of magnesia +would be readily recognised, and though the base of the rock is +crystalline, they are replete with the remains of organised beings. +Thus many of the magnesian limestones of the Carboniferous series +of the North of England are very like ordinary limestone to look +at, though effervescing less freely with acids, and the microscope +proves them to be charged with the remains of <i>Foraminifera</i> +and other minute organisms. +</p> + +<p class="indent"> +<i>Marbles</i> are of various kinds, all limestones which are +sufficiently hard and compact to take a high polish going by +this name. Statuary marble, and most of the celebrated foreign +marbles, are "metamorphic" rocks, of a highly crystalline nature, +and having all traces of their primitive organic structure +obliterated. Many other marbles, however, differ from ordinary +limestone simply in the matter of density. Thus, many marbles +(such as Derbyshire marble) are simply "crinoidal limestones" +(fig. 9); whilst various other British marbles exhibit innumerable +organic remains under the microscope. Black marbles owe their +colour to the presence of very minute particles of carbonaceous +matter, in some cases at any rate; and they may either be +metamorphic, or they may be charged with minute fossils such as +<i>Foraminifera</i> (<i>e.g.</i>, the black limestones of Ireland, +and the black marble of Dent, in Yorkshire). +</p> + +<p class="indent"> +"<i>Oolitic</i>" <i>limestones</i>, or "<i>oolites</i>," as they +are often called, are of interest both to the palæontologist +and geologist. The peculiar structure to which they owe their name +is that the rock is more or less entirely composed of spheroidal +or oval grains, which vary in size from the head of a small pin or +less up to the size of a pea, and which may be in almost immediate +contact with one another, or may be cemented together by a +<a name="page_29"><span class="page">Page 29</span></a> +more or less abundant calcareous matrix. When the +grains are pretty nearly spherical and are in tolerably close +contact, the rock looks very like the roe of a fish, and the name +of "oolite" or "egg-stone" is in allusion to this. When the grains +are of the size of peas or upwards, the rock is often called a +"pisolite" (Lat. <i>pisum</i>, a pea). Limestones having this +peculiar structure are especially abundant in the Jurassic formation, +which is often called the "Oolitic series" for this reason; but +essentially similar limestones occur not uncommonly in the Silurian, +Devonian, and Carboniferous formations, and, indeed, in almost all +rock-groups in which limestones are largely developed. Whatever may +be the age of the formation in which they occur, and whatever may +be the size of their component "eggs," the structure of oolitic +limestones is fundamentally the same. All the ordinary oolitic +limestones, namely, consist of little spherical or ovoid +"concretions," as they are termed, cemented together by a larger +or smaller amount of crystalline carbonate of lime, together, in +many instances, with numerous organic remains of different kinds + +<span style="float: right; margin: 4px; width: 263px; + font-size: smaller; text-align: justify;"> +<img src="images/fig013.jpg" width="255" height="253" alt="Fig. 13" /> +<br /> +Fig. 13.—Slice of oolitic limestone from the Jurassic series +(Coral Rag) of Weymouth; magnified. (Original.) +</span> + +(fig. 13). When examined in polished slabs, or in thin sections +prepared for the microscope, each of these little concretions is seen +to consist of numerous concentric coats of carbonate of lime, which +sometimes simply surround an imaginary centre, but which, more +commonly, have been successively deposited round some foreign body, +such as a little crystal of quartz, a cluster of sand-grains, or a +minute shell. In other cases, as in some of the beds of the Carboniferous +limestone in the North of England, where the limestone is highly +"arenaceous," there is a modification of the oolitic structure. +Microscopic sections of these sandy limestones (fig. 14) show +numerous generally angular or oval grains of silica or flint, each +of which is commonly surrounded by a thin coating of carbonate of +lime, or sometimes by several such coats, the whole being cemented +together along with the shells of <i>Foraminifera</i> and other +minute fossils by a matrix of crystalline calcite. As compared +with typical oolites, the concretions in these limestones are +usually much more irregular in shape, +<a name="page_30"><span class="page">Page 30</span></a> +often lengthened out and +almost cylindrical, at other times angular, the central nucleus + +<span style="float: left; margin: 4px; width: 254px; + font-size: smaller; text-align: justify;"> +<img src="images/fig014.jpg" width="252" height="248" alt="Fig. 14" /> +<br /> +Fig. 14.—Slice of arenaceous and oolitic limestone +from the Carboniferous series of Shap, Westmoreland; magnified. +The section also exhibit <i>Foraminifera</i> and other minute +fossils. (Original.) +</span> + +being of large size, and the surrounding envelope of lime being +very thin, and often exhibiting no concentric structure. In both +these and the ordinary oolites, the structure is fundamentally +the same. Both have been formed in a sea, probably of no great +depth, the waters of which were charged with carbonate of lime +in solution, whilst the bottom was formed of sand intermixed with +minute shells and fragments of the skeletons of larger marine +animals. The excess of lime in the sea-water was precipitated +round the sand-grams, or round the smaller shells, as so many +nuclei, and this precipitation must often have taken place time +after time, so as to give rise to the concentric structure so +characteristic of oolitic concretions. Finally, the oolitic grains +thus produced were cemented together by a further precipitation +of crystalline carbonate of lime from the waters of the ocean. +</p> + +<p class="indent"> +<i>Phosphate of Lime</i> is another lime-salt, which is of interest +to the palæontologist. It does not occur largely in the +stratified series, but it is found in considerable beds [4] in +the Laurentian formation, and less abundantly in some later +rock-groups, whilst it occurs abundantly in the form of nodules +in parts of the Cretaceous (Upper Greensand) and Tertiary deposits. +Phosphate of lime forms the larger proportion of the earthy matters +of the bones of Vertebrate animals, and also occurs in less amount +in the skeletons of certain of the Invertebrates (<i>e.g.</i>, +<i>Crustacea</i>). It is, indeed, perhaps more distinctively than +carbonate of lime, an organic compound; and though the formation +of many known deposits of phosphate of +<a name="page_31"><span class="page">Page 31</span></a> +lime cannot be positively shown to be connected with the previous +operation of living beings, there is room for doubt whether this +salt is not in reality always primarily a product of vital action. +The phosphatic nodules of the Upper Greensand are erroneously +called "coprolites," from the belief originally entertained that +they were the droppings or fossilised excrements of extinct +animals; and though this is not the case, there can be little +doubt but that the phosphate of lime which they contain is in +this instance of organic origin.[5] It appears, in fact, that +decaying animal matter has a singular power of determining the +precipitation around it of mineral salts dissolved in water. +Thus, when any animal bodies are undergoing decay at the bottom +of the sea, they have a tendency to cause the precipitation +from the surrounding water of any mineral matters which may be +dissolved in it; and the organic body thus becomes a centre +round which the mineral matters in question are deposited in +the form of a "concretion" or "nodule." The phosphatic nodules +in question were formed in a sea in which phosphate of lime, +derived from the destruction of animal skeletons, was held largely +in solution; and a precipitation of it took place round any body, +such as a decaying animal substance, which happened to be lying on +the sea-bottom, and which offered itself as a favourable nucleus. +In the same way we may explain the formation of the calcareous +nodules, known as "septaria" or "cement stones," which occur so +commonly in the London Clay and Kimmeridge Clay, and in which the +principal ingredient is carbonate of lime. A similar origin is to +be ascribed to the nodules of clay iron-stone (impure carbonate of +iron) which occur so abundantly in the shales of the Carboniferous +series and in other argillaceous deposits; and a parallel modern +example is to be found in the nodules of manganese, which were +found by Sir Wyville Thomson, in the Challenger, to be so numerously +scattered over the floor of the Pacific at great depths. In +accordance with this mode of origin, it is exceedingly common +to find in the centre of all these nodules, both old and new, +some organic body, such as a bone, a shell, or a tooth, which +acted as the original nucleus of precipitation, and +<a name="page_32"><span class="page">Page 32</span></a> +was thus preserved in a shroud of mineral matter. +Many nodules, it is true, show no such nucleus; but it has been +affirmed that all of them can be shown, by appropriate +microscopical investigation, to have been formed round an original +organic body to begin with (Hawkins Johnson). +</p> + +<p class="footnote"> +[Footnote 4: Apart from the occurrence or phosphate of lime in +actual beds in the stratified rocks, as in the Laurentian and +Silurian series, this salt may also occur disseminated through +the rock, when it can only be detected by chemical analysis. It +is interesting to note that Dr Hicks has recently proved the +occurrence of phosphate of lime in this disseminated form in +rocks as old as the Cambrian, and that in quantity quite equal to +what is generally found to be present in the later fossiliferous +rocks. This affords a chemical proof that animal life flourished +abundantly in the Cambrian seas.] +</p> + +<p class="footnote"> +[Footnote 5: It has been maintained, indeed, that the phosphatic +nodules so largely worked for agricultural purposes, are in +themselves actual organic bodies or true fossils. In a few cases +this admits of demonstration, as it can be shown that the nodule +is simply an organism (such as a sponge) infiltrated with phosphate +of lime (Sollas); but there are many other cases in which no actual +structure has yet been shown to exist, and as to the true origin +of which it would be hazardous to offer a positive opinion.] +</p> + +<p class="indent"> +The last lime-salt which need be mentioned is <i>gypsum</i>, +or <i>sulphate of lime</i>. This substance, apart from other +modes of occurrence, is not uncommonly found interstratified +with the ordinary sedimentary rocks, in the form of more or less +irregular beds; and in these cases it has a palæontological +importance, as occasionally yielding well-preserved fossils. Whilst +its exact mode of origin is uncertain, it cannot be regarded as +in itself an organic rock, though clearly the product of chemical +action. To look at, it is usually a whitish or yellowish-white +rock, as coarsely crystalline as loaf-sugar, or more so; and +the microscope shows it to be composed entirely of crystals of +sulphate of lime. +</p> + +<p class="indent"> +We have seen that the <i>calcareous</i> or lime-containing rocks +are the most important of the group of organic deposits; whilst +the <i>siliceous</i> or flint-containing rocks may be regarded as +the most important, most typical, and most generally distributed +of the mechanically-formed rocks. We have, however, now briefly +to consider certain deposits which are more or less completely +formed of flint; but which, nevertheless, are essentially organic +in their origin. +</p> + +<p class="indent"> +Flint or silex, hard and intractable as it is, is nevertheless +capable of solution in water to a certain extent, and even of +assuming, under certain circumstances, a gelatinous or viscous +condition. Hence, some hot-springs are impregnated with silica +to a considerable extent; it is present in small quantity in +sea-water; and there is reason to believe that a minute proportion +must very generally be present in all bodies of fresh water as +well. It is from this silica dissolved in the water that many +animals and some plants are enabled to construct for themselves +flinty skeletons; and we find that these animals and plants are and +have been sufficiently numerous to give rise to very considerable +deposits of siliceous matter by the mere accumulation of their +skeletons. Amongst the animals which require special mention in +this connection are the microscopic organisms which are known +to the naturalist as <i>Polycystina</i>. These little creatures +are of the lowest possible grade of organisation, very closely +related to the animals which we have previously spoken of as +<i>Foraminifera</i>, but differing in the fact that they secrete +a shell or skeleton composed of flint instead of lime. The +<i>Polycystina</i> occur abundantly in our present seas; +<a name="page_33"><span class="page">Page 33</span></a> +and their shells are present in some numbers +in the ooze which is found at great depths in the Atlantic and +Pacific oceans, being easily recognised by their exquisite +shape, their glassy transparency, the general presence of longer +or shorter spines, and the sieve-like perforations in the walls. +Both in Barbadoes and in the Nicobar islands occur geological +formations which are composed of the flinty skeletons of these +microscopic animals; the deposit in the former locality +attaining a great thickness, and having been long known to +workers with the microscope under the name of "Barbadoes earth" +(fig. 15). +</p> + +<p class="indent"> +In addition to flint-producing animals, we have also the great +group of fresh-water and marine microscopic plants known as + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 266px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig015.jpg" width="258" height="253" alt="Fig. 15" /> +<br /> +Fig. 15.—Shells of <i>Polycystina</i> from "Barbadoes +earth;" greatly magnified. (Original.) +</span> + +<span style="width: 266px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig016.jpg" width="258" height="252" alt="Fig. 16" /> +<br /> +Fig 16.—Cases of Diatoms in the Richmond "Infusorial +earth;" highly magnified. (Original.) +</span> + +</span> + +<i>Diatoms</i>, which likewise secrete a siliceous skeleton, often +of great beauty. The skeletons of Diatoms are found abundantly at +the present day in lake-deposits, guano, the silt of estuaries, +and in the mud which covers many parts of the sea-bottom; they +have been detected in strata of great age; and in spite of their +microscopic dimensions, they have not uncommonly accumulated to +form deposits of great thickness, and of considerable superficial +extent. Thus the celebrated deposit of "tripoli" ("Polir-schiefer") +of Bohemia, largely worked as polishing-powder, is composed wholly, +or almost wholly, of the flinty cases of Diatoms, of which it +is calculated that no less than forty-one thousand millions go +to make up a single cubic inch of the stone. Another celebrated +deposit is the so-called "Infusorial earth" of Richmond in Virginia, +where there is a stratum in places thirty feet thick, composed +almost entirely of the microscopic shells of Diatoms. +</p> + +<p class="indent"> +Nodules or layers of <i>flint</i>, or the impure variety of flint +<a name="page_34"><span class="page">Page 34</span></a> +known as <i>chert</i>, are found in limestones +of almost all ages from the Silurian upwards; but they are +especially abundant in the chalk. When these flints are examined +in thin and transparent slices under the microscope, or in polished +sections, they are found to contain an abundance of minute organic +bodies—such as <i>Foraminifera</i>, sponge-spicules, +&c.—embedded in a siliceous basis. In many instances the +flint contains larger organisms—such as a Sponge or a +Sea-urchin. As the flint has completely surrounded +and infiltrated the fossils which it contains, it is obvious +that it must have been deposited from sea-water in a gelatinous +condition, and subsequently have hardened. That silica is capable +of assuming this viscous and soluble condition is known; and +the formation of flint may therefore be regarded as due to the +separation of silica from the sea-water and its deposition round +some organic body in a state of chemical change or decay, just as +nodules of phosphate of lime or carbonate of iron are produced. +The existence of numerous organic bodies in flint has long been +known; but it should be added that a recent observer (Mr Hawkins +Johnson) asserts that the existence of an organic structure can +be demonstrated by suitable methods of treatment, even in the +actual matrix or basis of the flint.[6] +</p> + +<p class="footnote"> +[Footnote 6: It has been asserted that the flints of the chalk +are merely fossil sponges. No explanation of the origin of flint, +however, can be satisfactory, unless it embraces the origin of +chert in almost all great limestones from the Silurian upwards, +as well as the common phenomenon of the silicification of organic +bodies (such as corals and shells) which are known with certainty +to have been originally calcareous.] +</p> + +<p class="indent"> +In addition to deposits formed of flint itself, there are other +siliceous deposits formed by certain <i>silicates</i>, and also of +organic origin. It has been shown, namely—by observations +carried out in our present seas—that the shells of +<i>Foraminifera</i> are liable to become completely infiltrated +by silicates (such as "glauconite," or silicate of iron and +potash). Should the actual calcareous shell become dissolved +away subsequent to this infiltration—as is also liable +to occur—then, in place of the shells of the +<i>Foraminifera</i>, we get a corresponding number of green +sandy grains of glauconite, each grain being the <i>cast</i> +of a single shell. It has thus been shown that the green sand +found covering the sea-bottom in certain localities (as found by +the Challenger expedition along the line of the Agulhas current) +is really organic, and is composed of casts of the shells of +<i>Foraminifera</i>. Long before these observations had been +made, it had been shown by Professor Ehrenberg that the green +sands of various geological formations are composed mainly of +the internal casts of the shells of <i>Foraminifera</i>, and +<a name="page_35"><span class="page">Page 35</span></a> +we have thus another and a very +interesting example how rock-deposits of considerable extent +and of geological importance can be built up by the operation +of the minutest living beings. +</p> + +<p class="indent"> +As regards <i>argillaceous</i> deposits, containing <i>alumina</i> +or <i>clay</i> as their essential ingredient, it cannot be said that +any of these have been actually shown to be of organic origin. A +recent observation by Sir Wyville Thomson would, however, render it +not improbable that some of the great argillaceous accumulations of +past geological periods may be really organic. This distinguished +observer, during the cruise of the Challenger, showed that the +calcareous ooze which has been already spoken of as covering +large areas of the floor of the Atlantic and Pacific at great +depths, and which consists almost wholly of the shells of +<i>Foraminifera</i>, gave place at still greater depths to a +red ooze consisting of impalpable clayey mud, coloured by oxide +of iron, and devoid of traces of organic bodies. As the existence +of this widely-diffused red ooze, in mid-ocean, and at such great +depths, cannot be explained on the supposition that it is a sediment +brought down into the sea by rivers, Sir Wyville Thomson came to +the conclusion that it was probably formed by the action of the +sea-water upon the shells of <i>Foraminifera</i>. These shells, +though mainly consisting of lime, also contain a certain proportion +of alumina, the former being soluble in the carbonic acid dissolved +in the sea-water, whilst the latter is insoluble. There would +further appear to be grounds for believing that the solvent power +of the sea-water over lime is considerably increased at great +depths. If, therefore, we suppose the shells of <i>Foraminifera</i> +to be in course of deposition over the floor of the Pacific, at +certain depths they would remain unchanged, and would accumulate +to form a calcareous ooze; but at greater depths they would be +acted upon by the water, their lime would be dissolved out, their +form would disappear, and we should simply have left the small +amount of alumina which they previously contained. In process +of time this alumina would accumulate to form a bed of clay; and +as this clay had been directly derived from the decomposition +of the shells of animals, it would be fairly entitled to be +considered an organic deposit. Though not finally established, +the hypothesis of Sir Wyville Thomson on this subject is of the +greatest interest to the palæontologist, as possibly serving to +explain the occurrence, especially in the older formations, of +great deposits of argillaceous matter which are entirely destitute +of traces of life. +</p> + +<p class="indent"> +It only remains, in this connection, to shortly consider the +rock-deposits in which <i>carbon</i> is found to be present in +<a name="page_36"><span class="page">Page 36</span></a> +greater or less quantity. In the great +majority of cases where rocks are found to contain carbon or +carbonaceous matter, it can be stated with certainty that this +substance is of organic origin, though it is not necessarily +derived from vegetables. Carbon derived from the decomposition +of animal bodies is not uncommon; though it never occurs in +such quantity from this source as it may do when it is derived +from plants. Thus, many limestones are more or less highly +bituminous; the celebrated siliceous flags or so-called +"bituminous schists" of Caithness are impregnated with oily +matter apparently derived from the decomposition of the +numerous fishes embedded in them; Silurian shales containing +Graptolites, but destitute of plants, are not uncommonly +"anthracitic," and contain a small percentage of carbon derived +from the decay of these zoophytes; whilst the petroleum so largely +worked in North America has not improbably an animal origin. +That the fatty compounds present in animal bodies should more or +less extensively impregnate fossiliferous rock-masses, is only +what might be expected; but the great bulk of the carbon which +exists stored up in the earth's crust is derived from plants; +and the form in which it principally presents itself is that of +coal. We shall have to speak again, and at greater length, of +coal, and it is sufficient to say here that all the true coals, +anthracites, and lignites, are of organic origin, and consist +principally of the remains of plants in a more or less altered +condition. The bituminous shales which are found so commonly +associated with beds of coal also derive their carbon primarily +from plants; and the same is certainly, or probably, the case +with similar shales which are known to occur in formations younger +than the Carboniferous. Lastly, carbon may occur as a conspicuous +constituent of rock-masses in the form of <i>graphite</i> or +<i>black-lead</i>. In this form, it occurs in the shape of detached +scales, of veins or strings, or sometimes of regular layers;[7] +and there can be little doubt that in many instances it has an +organic origin, though this is not capable of direct proof. When +present, at any rate, in quantity, and in the form of layers +associated with stratified rocks, as is often the case in the +Laurentian formation, there can be little hesitation in regarding +it as of vegetable origin, and as an altered coal. +</p> + +<p class="footnote"> +[Footnote 7: In the Huronian formation at Steel River, on the +north shore of Lake Superior, there exists a bed of carbonaceous +matter which is regularly interstratified with the surrounding +rocks, and has a thickness of from 30 to 40 feet. This bed is +shown by chemical analysis to contain about 50 per cent of carbon, +partly in the form of graphite, partly in the form of anthracite; +and there can be little doubt but that it is really a stratum +of "metamorphic" coal.] +</p> + +<h3> +<a name="page_37"><span class="page">Page 37</span></a> +CHAPTER III.</h3> + +<p class="subtitle"> +CHRONOLOGICAL SUCCESSION OF THE FOSSILIFEROUS ROCKS. +</p> + +<p class="indent"> +The physical geologist, who deals with rocks simply as rocks, +and who does not necessarily trouble himself about what fossils +they may contain, finds that the stratified deposits which form +so large a portion of the visible part of the earth's crust are +not promiscuously heaped together, but that they have a certain +definite arrangement. In each country that he examines, he finds +that certain groups of strata lie above certain other groups; +and in comparing different countries with one another, he finds +that, in the main, the same groups of rocks are always found in the +same relative position to each other. It is possible, therefore, +for the physical geologist to arrange the known stratified rocks +into a successive series of groups, or "formations," having a +certain definite order. The establishment of this physical order +amongst the rocks introduces, however, at once the element of +<i>time</i>, and the physical succession of the strata can be +converted directly into a historical or <i>chronological</i> +succession. This is obvious, when we reflect that any bed or +set of beds of sedimentary origin is clearly and necessarily +younger than all the strata upon which it rests, and older than +all those by which it is surmounted. +</p> + +<p class="indent"> +It is possible, then, by an appeal to the rocks alone, to determine +in each country the general physical succession of the strata, and +this "stratigraphical" arrangement, when once determined, gives us +the <i>relative</i> ages of the successive groups. The task, +however, of the physical geologist in this matter is immensely +lightened when he calls in palæontology to his aid, and +studies the evidence of the fossils embedded in the rocks. Not only +is it thus much easier to determine the order of succession of the +strata in any given region, but it becomes now for the first time +possible to compare, with certainty and precision, the order of +succession in one region with that which exists in other regions +far distant. The value of fossils as tests of the relative ages +of the sedimentary rocks depends on the fact that they are not +indefinitely or promiscuously scattered through the crust of the +earth,—as it is conceivable that they might be. On the +contrary, the first and most firmly established law of +Palæontology is, that <i>particular kinds of fossils are +confined to particular rocks</i>, and <i>particular groups of +fossils are confined to particular groups of rocks</i>. +<a name="page_38"><span class="page">Page 38</span></a> +Fossils, then, are distinctive +of the rocks in which they are found—much more distinctive, +in fact, than the mere mineral character of the rock can be, for +<i>that</i> commonly changes as a formation is traced from one +region to another, whilst the fossils remain unaltered. It would +therefore be quite possible for the palæontologist, by an +appeal to the fossils alone, to arrange the series of sedimentary +deposits into a pile of strata having a certain definite order. +Not only would this be possible, but it would be found—if +sufficient knowledge had been brought to bear on both +sides—that the palæontological arrangement of the +strata would coincide in its details with the stratigraphical +or physical arrangement. +</p> + +<p class="indent"> +Happily for science, there is no such division between the +palæontologist and the physical geologist as here supposed; but +by the combined researches of the two, it has been found possible +to divide the entire series of stratified deposits into a number +of definite <i>rock-groups</i> or <i>formations</i>, which have a +recognised order of succession, and each of which is characterised +by possessing an assemblage of organic remains which do not occur +in association in any other formation. Such an <i>assemblage of +fossils</i>, characteristic of any given formation, represents +the <i>life</i> of the particular <i>period</i> in which the +formation was deposited. In this way the past history of the earth +becomes divided into a series of successive <i>life-periods</i>, +each of which corresponds with the deposition of a particular +<i>formation</i> or group of strata. +</p> + +<p class="indent"> +Whilst particular <i>assemblages</i> of organic forms characterise +particular <i>groups</i> of rocks, it may be further said that, +in a general way, each subdivision of each formation has its own +peculiar fossils, by which it may be recognised by a skilled +worker in Palæontology. Whenever, for instance, we meet with +examples of the fossils which are known as <i>Graptolites</i>, +we may be sure that we are dealing with <i>Silurian</i> rocks +(leaving out of sight one or two forms doubtfully referred to +this family). We may, however, go much farther than this with +perfect safety. If the Graptolites belong to certain genera, +we may be quite certain that we are dealing with <i>Lower</i> +Silurian rocks. Furthermore, if certain special forms are present, +we may be even able to say to what exact subdivision of the Lower +Silurian series they belong. +</p> + +<p class="indent"> +As regards particular fossils, however, or even particular classes +of fossils, conclusions of this nature require to be accompanied +by a tacit but well-understood reservation. So far as +<a name="page_39"><span class="page">Page 39</span></a> +our present observation goes, none of the undoubted +Graptolites have ever been discovered in rocks later than those +known upon other grounds to be Silurian; but it is possible that +they might at any time be detected in younger deposits. Similarly, +the species and genera which we now regard as characteristic of the +Lower Silurian, may at some future time be found to have survived +into the Upper Silurian period. We should not forget, therefore, +in determining the age of strata by palæontological evidence, +that we are always reasoning upon generalisations which are the +result of experience alone, and which are liable to be vitiated +by further and additional discoveries. +</p> + +<p class="indent"> +When the palæontological evidence as to the age of any given +set of strata is corroborated by the physical evidence, our +conclusions may be regarded as almost certain; but there are +certain limitations and fallacies in the palæontological method +of inquiry which deserve a passing mention. In the first place, +fossils are not always present in the stratified rocks; many +aqueous rocks are unfossiliferous, through a thickness of hundreds +or even thousands of feet of little-altered sediments; and even +amongst beds which do contain fossils, we often meet with strata +of many feet or yards in thickness which are wholly destitute +of any traces of fossils. There are, therefore, to begin with, +many cases in which there is no palæontological evidence extant +or available as to the age of a given group of strata. In the +second place, palæontological observers in different parts of +the world are liable to give different names to the same fossil, +and in all parts of the world they are occasionally liable to group +together different fossils under the same title. Both these sources +of fallacy require to be guarded against in reasoning as to the age +of strata from their fossil remains. Thirdly, the mere fact of +fossils being found in beds which are known by physical evidence +to be of different ages, has commonly led palæontologists +to describe them as different species. Thus, the same fossil, +occurring in successive groups of strata, and with the merely +trivial and varietal differences due to the gradual change in its +environment, has been repeatedly described as a distinct species, +with a distinct name, in every bed in which it was found. We know, +however, that many fossils range vertically through many groups +of strata, and there are some which even pass through several +formations. The mere fact of a difference of physical position +ought never to be taken into account at all in considering and +determining the true affinities of a fossil. Fourthly, the results +of experience, instead of being an assistance, are sometimes +liable to operate as a source of error. When once, +<a name="page_40"><span class="page">Page 40</span></a> +namely, a generalisation has been established +that certain fossils occur in strata of a certain age, +palæontologists are apt to infer that <i>all</i> beds +containing similar fossils must be of the same age. There is a +presumption, of course, that this inference would be correct; +but it is not a conclusion resting upon absolute necessity, and +there might be physical evidence to disprove it. Fifthly, the +physical geologist may lead the palæontologist astray by +asserting that the physical evidence as to the age and position +of a given group of beds is clear and unequivocal, when such +evidence may be, in reality, very slight and doubtful. In this +way, the observer may be readily led into wrong conclusions as +to the nature of the organic remains—often obscure and +fragmentary—which it is his business to examine, or he may +be led erroneously to think that previous generalisations as to +the age of certain kinds of fossils are premature and incorrect. +Lastly, there are cases in which, owing to the limited exposure +of the beds, to their being merely of local development, or to +other causes, the physical evidence as to the age of a given +group of strata may be entirely uncertain and unreliable, and +in which, therefore, the observer has to rely wholly upon the +fossils which he may meet with. +</p> + +<p class="indent"> +In spite of the above limitations and fallacies, there can be +no doubt as to the enormous value of palæontology in enabling +us to work out the historical succession of the sedimentary rocks. +It may even be said that in any case where there should appear +to be a clear and decisive discordance between the physical and +the palæontological evidence as to the age of a given series +of beds, it is the former that is to be distrusted rather than +the latter. The records of geological science contain not a few +cases in which apparently clear physical evidence of superposition +has been demonstrated to have been wrongly interpreted; but the +evidence of palæontology, when in any way sufficient, has +rarely been upset by subsequent investigations. Should we find +strata containing plants of the Coal-measures apparently resting +upon other strata with Ammonites and Belemnites, we may be sure +that the physical evidence is delusive; and though the above is an +extreme case, the presumption in all such instances is rather that +the physical succession has been misunderstood or misconstrued, +than that there has been a subversion of the recognised succession +of life-forms. +</p> + +<p class="indent"> +We have seen, then, that as the collective result of observations +made upon the superposition of rocks in different localities, from +their mineral characters, and from their included +<a name="page_41"><span class="page">Page 41</span></a> +fossils, geologists have been able to divide the entire +stratified series into a number of different divisions or +formations, each characterised by a <i>general</i> uniformity of +mineral composition, and by a special and peculiar <i>assemblage</i> +of organic forms. Each of these primary groups is in turn divided +into a series of smaller divisions, characterised and distinguished +in the same way. It is not pretended for a moment that all these +primary rock-groups can anywhere be seen surmounting one another +regularly.[8] There is no region upon the earth where all the +stratified formations can be seen together; and, even when most +of them occur in the same country, they can nowhere be seen all +succeeding each other in their regular and uninterrupted succession. +The reason of this is obvious. There are many places—to take +a single example—where one may see the the Silurian rocks, +the Devonian, and the Carboniferous rocks succeeding one another +regularly, and in their proper order. This is because the +particular region where this occurs was always submerged beneath +the sea while these formations were being deposited. There are, +however, many more localities in which one would find the +Carboniferous rocks resting unconformably upon the Silurians +without the intervention of any strata which could be referred +to the Devonian period. This might arise from one of two causes: +1. The Silurians might have been elevated above the sea +immediately after their deposition, so as to form dry land +during the whole of the Devonian period, in which case, of course, +no strata of the latter age could possibly be deposited in that +area. 2. The Devonian might have been deposited upon the Silurian, +and then the whole might have been elevated above the sea, and +subjected to an amount of denudation sufficient to remove the +Devonian strata entirely. In this case, when the land was again +submerged, the Carboniferous rocks, or any younger formation, +might be deposited directly upon Silurian strata. From one or +other of these causes, then, or from subsequent disturbances +and denudations, it happens that we can +<a name="page_42"><span class="page">Page 42</span></a> +rarely find many of the primary formations following one another +consecutively and in their regular order. +</p> + +<p class="footnote"> +[Footnote 8: As we have every reason to believe that dry land +and sea have existed, at any rate from the commencement of the +Laurentian period to the present day, it is quite obvious that +no one of the great formations can ever, under any circumstances, +have extended over the entire globe. In other words, no one of +the formations can ever have had a greater geographical extent +than that of the seas of the period in which the formation was +deposited. Nor is there any reason for thinking that the proportion +of dry land to ocean has ever been materially different to what +it is at present, however greatly the areas of sea and land may +have changed as regards their place. It follows from the above, +that there is no sufficient basis for the view that the crust of +the earth is composed of a succession of concentric layers, like +the coats of an onion, each layer representing one formation.] +</p> + +<p class="indent"> +In no case, however, do we ever find the Devonian resting upon +the Carboniferous, or the Silurian rocks reposing on the +Devonian. We have therefore, by a comparison of many different +areas, an established order of succession of the stratified +formations, as shown in the subjoined ideal section of the crust +of the earth (fig. 17). +</p> + +<p class="indent"> +The main subdivisions of the stratified rocks are known by the +following names:— +</p> + +<div class="center"> +<table summary="" style=""> + <tr><td class="right">1.</td> + <td colspan="2" class="left">Laurentian.</td></tr> + <tr><td class="right">2.</td> + <td colspan="2" class="left">Cambrian (with + Huronian?).</td></tr> + <tr><td class="right">3.</td> + <td colspan="2" class="left">Silurian.</td></tr> + <tr><td class="right">4.</td> + <td colspan="2" class="left">Devonian or Old Red + Sandstone.</td></tr> + <tr><td class="right">5.</td> + <td colspan="2" class="left">Carboniferous.</td></tr> + <tr><td class="right">6.</td> + <td class="left">Permian</td> + <td class="left" rowspan="2"><span class="big">}</span> + New Red Sandstone.</td></tr> + <tr><td class="right">7.</td> + <td colspan="2" class="left">Triassic</td></tr> + <tr><td class="right">8.</td> + <td colspan="2" class="left">Jurassic or Oolitic.</td></tr> + <tr><td class="right">9.</td> + <td colspan="2" class="left">Cretaceous.</td></tr> + <tr><td class="right">10.</td> + <td colspan="2" class="left">Eocene.</td></tr> + <tr><td class="right">11.</td> + <td colspan="2" class="left">Miocene.</td></tr> + <tr><td class="right">12.</td> + <td colspan="2" class="left">Pliocene.</td></tr> + <tr><td class="right">13.</td> + <td colspan="2" class="left">Post-tertiary.</td></tr> +</table> +</div> + +<p> </p> + +<div class="center"> +<table summary="" style=""> +<tr><td class="center"><span class="image"> +<a name="page_43"><span class="page">Page 43</span></a> +IDEAL SECTION OF THE CRUST OF THE EARTH. +</span></td></tr> +<tr><td class="center"><span class="image">Fig. 17. +</span></td></tr> +<tr><td> +<img src="images/fig017.jpg" width="518" height="848" alt="Fig. 17" /> +</td></tr> +</table> +</div> + +<p class="indent"> +<a name="page_44"><span class="page">Page 44</span></a> +Of these primary rock divisions, the Laurentian, Cambrian, Silurian, +Devonian, Carboniferous, and Permian are collectively grouped +together under the name of the Primary or <i>PalĹ“ozoic</i> +rocks (Gr. <i>palaios</i>, ancient; <i>zoe</i>, life). Not only do +they constitute the oldest stratified accumulations, but from the +extreme divergence between their animals and plants and those now +in existence, they may appropriately be considered as belonging to +an "Old-Life" period of the world's history. The Triassic, Jurassic, +and Cretaceous systems are grouped together as the +<i>Secondary</i> or <i>Mesozoic</i> formations (Gr. <i>mesos</i>, +intermediate; <i>zoe</i>, life); the organic remains of this +"Middle-Life" period being, on the whole, intermediate in their +characters between those of the palæozoic epoch and those of +more modern strata. Lastly, the Eocene, Miocene, and Pliocene +formations are grouped together as the <i>Tertiary</i> or +<i>Kainozoic</i> rocks (Gr. <i>kainos</i>, new; <i>zoe</i>, life); +because they constitute a "New-Life" period, in which the organic +remains approximate in character to those now existing upon the +globe. The so-called <i>Post-Tertiary</i> deposits are placed +with the Kainozoic, or may be considered as forming a separate +<i>Quaternary</i> system. +</p> + +<h3>CHAPTER IV.</h3> + +<p class="subtitle"> +THE BREAKS IN THE GEOLOGICAL AND PALÆONTOLOGICAL +RECORD. +</p> + +<p class="indent"> +The term "contemporaneous" is usually applied by geologists to +groups of strata in different regions which contain the same +fossils, or an assemblage of fossils in which many identical +forms are present. That is to say, beds which contain identical, +or nearly identical, fossils, however widely separated they may +be from one another in point of actual distance, are ordinarily +believed to have been deposited during the same period of the +earth's history. This belief, indeed, constitutes the keystone +of the entire system of determining the age of strata by their +fossil contents; and if we take the word "contemporaneous" in a +general and strictly geological sense, this belief can be accepted +as proved beyond denial. We must, however, guard ourselves against +too literal an interpretation of the word "contemporaneous," and +we must bear in mind the enormously-prolonged periods of time with +which the geologist has to deal. When we say that two groups of +<a name="page_45"><span class="page">Page 45</span></a> +strata in different regions are "contemporaneous," we simply mean +that they were formed during the same geological period, and +perhaps at different stages of that period, and we do not mean to +imply that they were formed at precisely the same instant of time. +</p> + +<p class="indent"> +A moment's consideration will show us that it is only in the +former sense that we can properly speak of strata being +"contemporaneous;" and that, in point of fact, beds containing +the same fossils, if occurring in widely distant areas, can hardly +be "contemporaneous" in any literal sense; but that the very +identity of their fossils is proof that they were deposited one +after the other. If we find strata containing identical fossils +within the limits of a single geographical region—say in +Europe—then there is a reasonable probability that these beds +are strictly contemporaneous, in the sense that they were deposited +at the same time. There is a reasonable probability of this, +because there is no improbability involved in the idea of an +ocean occupying the whole area of Europe, and peopled throughout +by many of the same species of marine animals. At the present +day, for example, many identical species of animals are found +living on the western coasts of Britain and the eastern coasts +of North America, and beds now in course of deposition off the +shores of Ireland and the seaboard of the state of New York would +necessarily contain many of the same fossils. Such beds would be +both literally and geologically contemporaneous; but the case is +different if the distance between the areas where the strata occur +be greatly increased. We find, for example, beds containing identical +fossils (the Quebec or Skiddaw beds) in Sweden, in the north of +England, in Canada, and in Australia. Now, if all these beds were +contemporaneous, in the literal sense of the term, we should have +to suppose that the ocean at one time extended uninterruptedly +between all these points, and was peopled throughout the vast area +thus indicated by many of the same animals. Nothing, however, +that we see at the present day would justify us in imagining an +ocean of such enormous extent, and at the same time so uniform +in its depth, temperature, and other conditions of marine life, +as to allow the same animals to flourish in it from end to end; +and the example chosen is only one of a long and ever-recurring +series. It is therefore much more reasonable to explain this, +and all similar cases, as owing to the <i>migration</i> of the +fauna, in whole or in part, from one marine area to another. +Thus, we may suppose an ocean to cover what is now the European +area, and to be peopled by certain species of animals. Beds of +sediment—clay, sands, and limestones—will be deposited +over the sea-bottom, and +<a name="page_46"><span class="page">Page 46</span></a> +will entomb the +remains of the animals as fossils. After this has lasted for a +certain length of time, the European area may undergo elevation, +or may become otherwise unsuitable for the perpetuation of its +fauna; the result of which would be that some or all of the +marine animals of the area would migrate to some more suitable +region. Sediments would then be accumulated in the new area to +which they had betaken themselves, and they would then appear, +for the second time, as fossils in a set of beds widely +separated from Europe. The second set of beds would, however, +obviously not be strictly or literally contemporaneous with the +first, but would be separated from them by the period of time +required for the migration of the animals from the one area +into the other. It is only in a wide and comprehensive sense +that such strata can be said to be contemporaneous. +</p> + +<p class="indent"> +It is impossible to enter further into this subject here; but it +may be taken as certain that beds in widely remote geographical +areas can only come to contain the same fossils by reason of a +migration having taken place of the animals of the one area to +the other. That such migrations can and do take place is quite +certain, and this is a much more reasonable explanation of the +observed facts than the hypothesis that in former periods the +conditions of life were much more uniform than they are at present, +and that, consequently, the same organisms were able to range over +the entire globe at the same time. It need only be added, that +taking the evidence of the present as explaining the phenomena +of the past—the only safe method of reasoning in geological +matters—we have abundant proof that deposits which <i>are</i> +actually contemporaneous, in the strict sense of the term, <i>do +not contain the same fossils, if far removed from one another in +point of distance</i>. Thus, deposits of various kinds are now +in process of formation in our existing seas, as, for example, in +the Arctic Ocean, the Atlantic, and the Pacific, and many of these +deposits are known to us by actual examination and observation +with the sounding-lead and dredge. But it is hardly necessary to +add that the animal remains contained in these deposits—the +fossils of some future period—instead of being identical, are +widely different from one another in their characters. +</p> + +<p class="indent"> +We have seen, then, that the entire stratified series is capable of +subdivision into a number of definite rock-groups or "formations," +each possessing a peculiar and characteristic assemblage of fossils, +representing the "life" of the "period" in which the formation +was deposited. We have still to inquire shortly how it came to +pass that two successive formations <i>should</i> +<a name="page_47"><span class="page">Page 47</span></a> +thus be broadly distinguished by their life-forms, and why +they should not rather possess at any rate a majority of identical +fossils. It was originally supposed that this could be explained by +the hypothesis that the close of each formation was accompanied by +a general destruction of all the living beings of the period, and +that the commencement of each new formation was signalised by the +creation of a number of brand-new organisms, destined to figure +as the characteristic fossils of the same. This theory, however, +ignores the fact that each formation—as to which we have +any sufficient evidence—contains a few, at least, of the +life-forms which existed in the preceding period; and it invokes +forces and processes of which we know nothing, and for the supposed +action of which we cannot account. The problem is an undeniably +difficult one, and it will not be possible here to give more than +a mere outline of the modern views upon the subject. Without +entering into the at present inscrutable question as to the manner +in which new life-forms are introduced upon the earth, it may be +stated that almost all modern geologists hold that the living beings +of any given formation are in the main modified forms of others +which have preceded them. It is not believed that any general or +universal destruction of life took place at the termination of +each geological period, or that a general introduction of new +forms took place at the commencement of a new period. It is, on +the contrary, believed that the animals and plants of any given +period are for the most part (or exclusively) the lineal but +modified descendants of the animals and plants of the immediately +preceding period, and that some of them, at any rate, are +continued into the next succeeding period, either unchanged, or +so far altered as to appear as new species. To discuss these +views in detail would lead us altogether too far, but there is +one very obvious consideration which may advantageously receive +some attention. It is obvious, namely, that the great +discordance which is found to subsist between the animal life of +any given formation and that of the next succeeding formation, +and which no one denies, would be a fatal blow to the views just +alluded to, unless admitting of some satisfactory explanation. +Nor is this discordance one purely of life-forms, for there is +often a physical break in the successions of strata as well. +Let us therefore briefly consider how far these interruptions +and breaks in the geological and palæontological record +can be accounted for, and still allow us to believe in some +theory of continuity as opposed to the doctrine of intermittent +and occasional action. +</p> + +<p class="indent"> +<a name="page_48"><span class="page">Page 48</span></a> +In the first place, it is perfectly clear that if we admit the +conception above mentioned of a continuity of life from the +Laurentian period to the present day, we could never <i>prove</i> +our view to be correct, unless we could produce in evidence +fossil examples of <i>all</i> the kinds of animals and plants +that have lived and died during that period. In order to do +this, we should require, to begin with, to have access to an +absolutely unbroken and perfect succession of all the deposits +which have ever been laid down since the beginning. If, however, +we ask the physical geologist if he is in possession of any such +uninterrupted series, he will at once answer in the negative. +So far from the geological series being a perfect one, it is +interrupted by numerous gaps of unknown length, many of which we +can never expect to fill up. Nor are the proofs of this far to +seek. Apart from the facts that we have hitherto examined only +a limited portion of the dry land, that nearly two-thirds of +the entire area of the globe is inaccessible to geological +investigation in consequence of its being covered by the sea, +that many deposits can be shown to have been more or less +completely destroyed subsequent to their deposition, and that +there may be many areas in which living beings exist where no +rock is in process of formation, we have the broad fact that +rock-deposition only goes on to any extent in water, and that +the earth must have always consisted partly of dry land and +partly of water—at any rate, so far as any period of +which we have geological knowledge is concerned. There +<i>must</i>, therefore, always have existed, at some part or +another of the earth's surface, areas where no deposition of +rock was going on, and the proof of this is to be found in the +well-known phenomenon of "<i>unconformability</i>." Whenever, +namely, deposition of sediment is continuously going on within +the limits of a single ocean, the beds which are laid down succeed +one another in uninterrupted and regular sequence. Such beds are +said to be "conformable," and there are many rock-groups known +where one may pass through fifteen or twenty thousand feet of strata +without a break—indicating that the beds had been deposited +in an area which remained continuously covered by the sea. On +the other hand, we commonly find that there is no such regular +succession when we pass from one great formation to another, but +that, on the contrary, the younger formation rests "unconformably," +as it is called, either upon the formation immediately preceding +it in point of time, or upon some still older one. The essential +physical feature of this unconformability is that the beds of the +younger formation rest upon a worn and eroded surface formed by the +<a name="page_49"><span class="page">Page 49</span></a> +beds of the older series (fig. 18); and a moment's +consideration will show us what this indicates. It indicates, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 528px; + font-size: smaller; text-align: center;"> +<img src="images/fig018.jpg" width="520" height="253" alt="Fig. 18" /> +<br /> +Fig. 18.—Section showing strata of Tertiary age (<i>a</i>) +resting upon a worn and eroded surface of White Chalk +(<i>b</i>), the stratification of which is marked by lines of +flint. +</span> +</span> + +beyond the possibility of misconception, that there was an interval +between the deposition of the older series and that of the newer +series of strata; and that during this interval the older beds +were raised above the sea-level, so as to form dry land, and were +subsequently depressed again beneath the waters, to receive upon +their worn and wasted upper surface the sediments of the later +group. During the interval thus indicated, the deposition of rock +must of necessity have been proceeding more or less actively +in other areas. Every unconformity, therefore, indicates that +at the spot where it occurs, a more or less extensive series +of beds must be actually missing; and though we may sometimes +be able to point to these missing strata in other areas, there +yet remains a number of unconformities for which we cannot at +present supply the deficiency even in a partial manner. +</p> + +<p class="indent"> +It follows from the above that the series of stratified deposits +is to a greater or less extent irremediably imperfect; and in +this imperfection we have one great cause why we can never obtain +a perfect series of all the animals and plants that have lived +upon the globe. Wherever one of these great physical gaps occurs, +we find, as we might expect, a corresponding break in the series +of life-forms. In other words, whenever we find two formations +to be unconformable, we shall always find at the same time that +there is a great difference in their fossils, and that many of +the fossils of the older formation do not survive into the newer, +whilst many of those in the newer are not known to occur in the +older. The cause of this is, obviously, +<a name="page_50"><span class="page">Page 50</span></a> +that the lapse of time, indicated by the unconformability, has +been sufficiently great to allow of the dying out or modification +of many of the older forms of life, and the introduction of new +ones by immigration. +</p> + +<p class="indent"> +Apart, however, altogether, from these great physical breaks +and their corresponding breaks in life, there are other reasons +why we can never become more than partially acquainted with the +former denizens of the globe. Foremost amongst these is the fact +that an enormous number of animals possess no hard parts of the +nature of a skeleton, and are therefore incapable, under any +ordinary circumstances, of leaving behind them any traces of +their existence. It is true that there are cases in which animals +in themselves completely soft-bodied are nevertheless able to leave +marks by which their former presence can be detected: Thus every +geologist is familiar with the winding and twisting "trails" formed +on the surface of the strata by sea-worms; and the impressions +left by the stranded carcases of Jelly-fishes on the fine-grained +lithographic slates of Solenhofen supply us with an example of how +a creature which is little more than "organised sea-water" may +still make an abiding mark upon the sands of time. As a general +rule, however, animals which have no skeletons are incapable of +being preserved as fossils, and hence there must always have +been a vast number of different kinds of marine animals of which +we have absolutely no record whatever. Again, almost all the +fossiliferous rocks have been laid down in water; and it is a +necessary result of this that the great majority of fossils are +the remains of aquatic animals. The remains of air-breathing +animals, whether of the inhabitants of the land or of the air +itself, are comparatively rare as fossils, and the record of +the past existence of these is much more imperfect than is the +case with animals living in water. Moreover, the fossiliferous +deposits are not only almost exclusively aqueous formations, but +the great majority are marine, and only a comparatively small +number have been formed by lakes and rivers. It follows from the +foregoing that the palæontological record is fullest and most +complete so far as sea-animals are concerned, though even here we +find enormous gaps, owing to the absence of hard structures in +many great groups; of animals inhabiting fresh waters our knowledge +is rendered still further incomplete by the small proportion +that fluviatile and lacustrine deposits bear to marine; whilst +we have only a fragmentary acquaintance with the air-breathing +animals which inhabited the earth during past ages. +</p> + +<p class="indent"> +Lastly, the imperfection of the palæontological record, due +<a name="page_51"><span class="page">Page 51</span></a> +to the causes above enumerated, is greatly aggravated, especially +as regards the earlier portion of the earth's history, by the fact +that many rocks which contained fossils when deposited have since +been rendered barren of organic remains. The principal cause of +this common phenomenon is what is known as "metamorphism"—that +is, the subjection of the rock to a sufficient amount of heat to +cause a rearrangement of its particles. When at all of a pronounced +character, the result of metamorphic action is invariably the +obliteration of any fossils which might have been originally +present in the rock. Metamorphism may affect rocks of any age, +though naturally more prevalent in the older rocks, and to this +cause must be set down an irreparable loss of much fossil evidence. +The most striking example which is to be found of this is the great +Laurentian series, which comprises some 30,000 feet of +highly-metamorphosed sediments, but which, with one not wholly +undisputed exception, has as yet yielded no remains of living +beings, though there is strong evidence of the former existence +in it of fossils. +</p> + +<p class="indent"> +Upon the whole, then, we cannot doubt that the earth's crust, so +far as yet deciphered by us, presents us with but a very imperfect +record of the past. Whether the known and admitted imperfections +of the geological and palæontological records are sufficiently +serious to account satisfactorily for the deficiency of direct +evidence recognisable in some modern hypotheses, may be a matter +of individual opinion. There can, however, be little doubt that +they are sufficiently extensive to throw the balance of evidence +decisively in favour of some theory of <i>continuity</i>, as +opposed to any theory of intermittent and occasional action. The +apparent breaks which divide the great series of the stratified +rocks into a number of isolated formations, are not marks of mighty +and general convulsions of nature, but are simply indications of +the imperfection of our knowledge. Never, in all probability, +shall we be able to point to a complete series of deposits, or a +complete succession of life linking one great geological period +to another. Nevertheless, we may well feel sure that such deposits +and such an unbroken succession must have existed at one time. +We are compelled to believe that nowhere in the long series of +the fossiliferous rocks has there been a total break, but that +there must have been a complete continuity of life, and a more +or less complete continuity of sedimentation, from the Laurentian +period to the present day. One generation hands on the lamp of +life to the next, and each system of rocks is the direct offspring +of those which preceded it in time. Though there +<a name="page_52"><span class="page">Page 52</span></a> +has not been continuity in any given area, still the +geological chain could never have been snapped at one point, and +taken up again at a totally different one. Thus we arrive at the +conviction that <i>continuity</i> is the fundamental law of +geology, as it is of the other sciences, and that the lines of +demarcation between the great formations are but gaps in our own +knowledge. +</p> + +<h3>CHAPTER V.</h3> + +<p class="subtitle"> +CONCLUSIONS TO BE DRAWN FROM FOSSILS. +</p> + +<p class="indent"> +We have already seen that geologists have been led by the study +of fossils to the all-important generalisation that the vast +series of the Fossiliferous or Sedimentary Rocks may be divided +into a number of definite groups or "formations," each of which is +characterised by its organic remains. It may simply be repeated here +that these formations are not properly and strictly characterised +by the occurrence in them of any one particular fossil. It may be +that a formation contains some particular fossil or fossils not +occurring out of that formation, and that in this way an observer +may identify a given group with tolerable certainty. It very +often happens, indeed, that some particular stratum, or sub-group +of a series, contains peculiar fossils, by which its existence +may be determined in various localities. As before remarked, +however, the great formations are characterised properly by the +association of certain fossils, by the predominance of certain +families or orders, or by an <i>assemblage</i> of fossil remains +representing the "life" of the period in which the formation +was deposited. +</p> + +<p class="indent"> +Fossils, then, enable us to determine the <i>age</i> of the deposits +in which they occur. Fossils further enable us to come to very +important conclusions as to the mode in which the fossiliferous +bed was deposited, and thus as to the condition of the particular +district or region occupied by the fossiliferous bed at the time +of the formation of the latter. If, in the first place, the bed +contain the remains of animals such as now inhabit rivers, we +know that it is "fluviatile" in its origin, and that it must at +one time have either formed an actual riverbed, or been deposited +by the overflowing of an ancient stream. Secondly, if the bed +contain the remains of shellfish, minute crustaceans, or fish, +such as now inhabit lakes, +<a name="page_53"><span class="page">Page 53</span></a> +we know that it is "lacustrine," and was deposited beneath the +waters of a former lake. Thirdly, if the bed contain the remains +of animals such as now people the ocean, we know that it is +"marine" in its origin, and that it is a fragment of an old +sea-bottom. +</p> + +<p class="indent"> +We can, however, often determine the conditions under which a bed +was deposited with greater accuracy than this. If, for example, the +fossils are of kinds resembling the marine animals now inhabiting +shallow waters, if they are accompanied by the detached relics +of terrestrial organisms, or if they are partially rolled and +broken, we may conclude that the fossiliferous deposit was laid +down in a shallow sea, in the immediate vicinity of a coast-line, +or as an actual shore-deposit. If, again, the remains are those +of animals such as now live in the deeper parts of the ocean, +and there is a very sparing intermixture of extraneous fossils +(such as the bones of birds or quadrupeds, or the remains of +plants), we may presume that the deposit is one of deep water. +In other cases, we may find, scattered through the rock, and +still in their natural position, the valves of shells such as +we know at the present day as living buried in the sand or mud +of the sea-shore or of estuaries. In other cases, the bed may +obviously have been an ancient coral-reef, or an accumulation of +social shells, like Oysters. Lastly, if we find the deposit to +contain the remains of marine shells, but that these are dwarfed +of their fair proportions and distorted in figure, we may conclude +that it was laid down in a brackish sea, such as the Baltic, in +which the proper saltness was wanting, owing to its receiving +an excessive supply of fresh water. +</p> + +<p class="indent"> +In the preceding, we have been dealing simply with the remains +of aquatic animals, and we have seen that certain conclusions +can be accurately reached by an examination of these. As regards +the determination of the conditions of deposition from the remains +of aerial and terrestrial animals, or from plants, there is not +such an absolute certainty. The remains of land-animals would, +of course, occur in "sub-aerial" deposits—that is, in beds, +like blown sand, accumulated upon the land. Most of the remains +of land-animals, however, are found in deposits which have been +laid down in water, and they owe their present position to the +fact that their former owners were drowned in rivers or lakes, +or carried out to sea by streams. Birds, Flying Reptiles, and +Flying Mammals might also similarly find their way into aqueous +deposits; but it is to be remembered that many birds and mammals +habitually spend a great part of their time in the water, and +that these might therefore be naturally expected to present +themselves as fossils in +<a name="page_54"><span class="page">Page 54</span></a> +Sedimentary Rocks. Plants, again, even when undoubtedly such as +must have grown on land, do not prove that the bed in which they +occur was formed on land. Many of the remains of plants known to +us are extraneous to the bed in which they are now found, having +reached their present site by falling into lakes or rivers, or +being carried out to sea by floods or gales of wind. There are, +however, many cases in which plants have undoubtedly grown on +the very spot where we now find them. Thus it is now generally +admitted that the great coal-fields of the Carboniferous age +are the result of the growth <i>in situ</i> +of the plants which compose coal, and that these grew on vast + +<span style="float: left; margin: 4px; width: 284px; + font-size: smaller; text-align: justify;"> +<img src="images/fig019.jpg" width="276" height="429" alt="Fig. 19" /> +<br /> +Fig. 19.—Erect Tree containing Reptilian remains. +Coal-measures, Nova Scotia. (After Dawson.) +</span> + +marshy or partially submerged tracts of level alluvial land. We +have, however, distinct evidence of old land-surfaces, both in +the Coal-measures and in other cases (as, for instance, in the +well-known "dirt-bed" of the Purbeck series). When, for example, +we find the erect stumps of trees standing at right angles to +the surrounding strata, we know that the surface through which +these send their roots was at one time the surface of the dry +land, or, in other words, was an ancient soil (fig. 19). +</p> + +<p class="indent"> +In many cases fossils enable us to come to important conclusions +as to the climate of the period in which they lived but only a +few instances of this can be here adduced. As fossils in the +majority of instances are the remains of marine animals, it is +mostly the temperature of the sea which can alone be determined +in this way; and it is important to remember that, owing to the +existence of heated currents, the marine climate of a given area +does not necessarily imply a correspondingly warm climate in +the neighbouring land. Land-climates can only be determined by +the remains of land-animals or land-plants, and these are +comparatively rare as fossils. It is also important to remember +that all conclusions on this +<a name="page_55"><span class="page">Page 55</span></a> +head are really based upon the present distribution of animal +and vegetable life on the globe, and are therefore liable to be +vitiated by the following considerations:— +</p> + +<p class="indent"> +<i>a</i>. Most fossils are extinct, and it is not certain that +the habits and requirements of any extinct animal were exactly +similar to those of its nearest living relative. +</p> + +<p class="indent"> +<i>b</i>. When we get very far back in time, we meet with groups +of organisms so unlike anything we know at the present day as to +render all conjectures as to climate founded upon their supposed +habits more or less uncertain and unsafe. +</p> + +<p class="indent"> +<i>c</i>. In the case of marine animals, we are as yet very far +from knowing the exact limits of distribution of many species within +our present seas; so that conclusions drawn from living forms as +to extinct species are apt to prove incorrect. For instance, it +has recently been shown that many shells formerly believed to +be confined to the Arctic Seas have, by reason of the extension +of Polar currents, a wide range to the south; and this has thrown +doubt upon the conclusions drawn from fossil shells as to the +Arctic conditions under which certain beds were supposed to have +been deposited. +</p> + +<p class="indent"> +<i>d</i>. The distribution of animals at the present day is certainly +dependent upon other conditions beside climate alone; and the causes +which now limit the range of given animals are certainly such as +belong to the existing order of things. But the establishment of +the present order of things does not date back in many cases to +the introduction of the present species of animals. Even in the +case, therefore, of existing species of animals, it can often +be shown that the past distribution of the species was different +formerly to what it is now, not necessarily because the climate +has changed, but because of the alteration of other conditions +essential to the life of the species or conducing to its extension. +</p> + +<p class="indent"> +Still, we are in many cases able to draw completely reliable +conclusions as to the climate of a given geological period, by +an examination of the fossils belonging to that period. Among +the more striking examples of how the past climate of a region +may be deduced from the study of the organic remains contained in +its rocks, the following may be mentioned: It has been shown that +in Eocene times, or at the commencement of the Tertiary period, +the climate of what is now Western Europe was of a tropical or +sub-tropical character. Thus the Eocene beds are found to contain +the remains of shells such as now inhabit tropical seas, as, for +example, Cowries and Volutes; and with these are the fruits of +palms, and the remains of other tropical plants. It has been shown, +<a name="page_56"><span class="page">Page 56</span></a> +again, that in Miocene times, or +about the middle of the Tertiary period, Central Europe was +peopled with a luxuriant flora resembling that of the warmer +parts of the United States, and leading to the conclusion that +the mean annual temperature must have been at least 30° +hotter than it is at present. It has been shown that, at the +same time, Greenland, now buried beneath a vast ice-shroud, was +warm enough to support a large number of trees, shrubs, and other +plants, such as inhabit temperate regions of the globe. Lastly, +it has been shown upon physical as well as palæontological +evidence, that the greater part of the North Temperate Zone, at +a comparatively recent geological period, has been visited with +all the rigours of an Arctic climate, resembling that of +Greenland at the present day. This is indicated by the occurrence +of Arctic shells in the superficial deposits of this period, +whilst the Musk-ox and the Reindeer roamed far south of their +present limits. +</p> + +<p class="indent"> +Lastly, it was from the study of fossils that geologists learnt +originally to comprehend a fact which may be regarded as of cardinal +importance in all modern geological theories and +speculations—namely, that the crust of the earth is liable +to local elevations and subsidences. For long after the remains of +shells and other marine animals were for the first time observed +in the solid rocks forming the dry land, and at great heights +above the sea-level, attempts were made to explain this almost +unintelligible phenomenon upon the hypothesis that the fossils in +question were not really the objects they represented, but were +in truth mere <i>lusus naturĹ“</i>, due to some "plastic virtue +latent in the earth." The common-sense of scientific men, however, +soon rejected this idea, and it was agreed by universal consent +that these bodies really were remains of animals which formerly +lived in the sea. When once this was admitted, the further steps +were comparatively easy, and at the present day no geological +doctrine stands on a firmer basis than that which teaches us +that our present continents and islands, fixed and immovable as +they appear, have been repeatedly sunk beneath the ocean. +</p> + +<h3> +<a name="page_57"><span class="page">Page 57</span></a> +CHAPTER VI.</h3> + +<p class="subtitle"> +THE BIOLOGICAL RELATIONS OF FOSSILS. +</p> + +<p class="indent"> +Not only have fossils, as we have seen, a most important bearing +upon the sciences of Geology and Physical Geography, but they +have relations of the most complicated and weighty character with +the numerous problems connected with the study of living beings, +or in other words, with the science of Biology. To such an extent +is this the case, that no adequate comprehension of Zoology and +Botany, in their modern form, is so much as possible without +some acquaintance with the types of animals and plants which have +passed away. There are also numerous speculative questions in +the domain of vital science, which, if soluble at all, can only +hope to find their key in researches carried out on extinct +organisms. To discuss fully the biological relations of fossils +would, therefore, afford matter for a separate treatise; and all +that can be done here is to indicate very cursorily the principal +points to which the attention of the palæontological student +ought to be directed. +</p> + +<p class="indent"> +In the first place, the great majority of fossil animals and +plants are "extinct"—that is to say, they belong to species +which are no longer in existence at the present day. So far, +however, from there being any truth in the old view that there +were periodic destructions of all the living beings in existence +upon the earth, followed by a corresponding number of new creations +of animals and plants, the actual facts of the case show that +the extinction of old forms and the introduction of new forms +have been processes constantly going on throughout the whole +of geological time. Every species seems to come into being at +a certain definite point of time, and to finally disappear at +another definite point; though there are few instances indeed, +if there are any, in which our present knowledge would permit +us safely to fix with precision the times of entrance and exit. +There are, moreover, marked differences in the actual time during +which different species remained in existence, and therefore +corresponding differences in their "vertical range," or, in other +words, in the actual amount and thickness of strata through which +they present themselves as fossils. Some species are found to +range through two or even three formations, and a few have an even +more extended life. More commonly the species which begin in the +<a name="page_58"><span class="page">Page 58</span></a> +commencement of a great formation die +out at or before its close, whilst those which are introduced for +the first time near the middle or end of the formation may either +become extinct, or may pass on into the next succeeding formation. +As a general rule, it is the animals which have the lowest and +simplest organisation that have the longest range in time, and +the additional possession of microscopic or minute dimensions +seems also to favour longevity. Thus some of the <i>Foraminifera</i> +appear to have survived, with little or no perceptible alteration, +from the Silurian period to the present day; whereas large and +highly-organised animals, though long-lived as <i>individuals</i>, +rarely seem to live long <i>specifically</i>, and have, therefore, +usually a restricted vertical range. Exceptions to this, however, +are occasionally to be found in some "persistent types," which +extend through a succession of geological periods with very +little modification. Thus the existing Lampshells of the genus +<i>Lingula</i> are little changed from the <i>LingulĹ“</i> +which swarmed in the Lower Silurian seas; and the existing Pearly +Nautilus is the last descendant of a clan nearly as ancient. On +the other hand, some forms are singularly restricted in their +limits, and seem to have enjoyed a comparatively brief lease +of life. An example of this is to be found in many of the +<i>Ammonites</i>—close allies of the Nautilus—which +are often confined strictly to certain zones of strata, in some +cases of very insignificant thickness. +</p> + +<p class="indent"> +Of the <i>causes</i> of extinction amongst fossil animals and +plants, we know little or nothing. All we can say is, that the +attributes which constitute a <i>species</i> do not seem to be +intrinsically endowed with permanence, any more than the attributes +which constitute an <i>individual</i>, though the former may +endure whilst many successive generations of the latter have +disappeared. Each species appears to have its own life-period, +its commencement, its culmination, and its gradual decay; and +the life-periods of different species may be of very different +duration. +</p> + +<p class="indent"> +From what has been said above, it may be gathered that our existing +species of animals and plants are, for the most part, quite of modern +origin, using the term "modern" in its geological acceptation. +Measured by human standards, the majority of existing animals +(which are capable of being preserved as fossils) are known to +have a high antiquity; and some of them can boast of a pedigree +which even the geologist may regard with respect. Not a few of +our shellfish are known to have commenced their existence at some +point of the Tertiary period; one Lampshell +<a name="page_59"><span class="page">Page 59</span></a> +(<i>Terebratulina caput-serpentis</i>) is believed to have +survived since the Chalk; and some of the <i>Foraminifera</i> date, +at any rate, from the Carboniferous period. We learn from this +the additional fact that our existing animals and plants do not +constitute an assemblage of organic forms which were introduced +into the world collectively and simultaneously, but that they +commenced their existence at very different periods, some being +extremely old, whilst others may be regarded as comparatively +recent animals. And this introduction of the existing fauna and +flora was a slow and <i>gradual</i> process, as shown admirably +by the study of the fossil shells of the Tertiary period. Thus, +in the earlier Tertiary period, we find about 95 per cent of the +known fossil shells to be species that are no longer in existence, +the remaining 5 per cent being forms which are known to live in +our present seas. In the middle of the Tertiary period we find +many more recent and still existing species of shells, and the +extinct types are much fewer in number; and this gradual +introduction of forms now living goes on steadily, till, at the +close of the Tertiary period, the proportions with which we +started may be reversed, as many as 90 or 95 per cent of the +fossil shells being forms still alive, while not more than 5 +per cent may have disappeared. +</p> + +<p class="indent"> +All known animals at the present day may be divided into some +five or six primary divisions, which are known technically as +"<i>sub-kingdoms</i>." Each of these sub-kingdoms [9] may be +regarded as representing a certain type or plan of structure, +and all the animals comprised in each are merely modified forms +of this common type. Not only are all known living animals thus +reducible to some five or six fundamental plans of structure, +but amongst the vast series of fossil forms no one has yet been +found—however unlike any existing animal—to possess +peculiarities which would entitle it to be placed in a new +sub-kingdom. All fossil animals, therefore, are capable of +being referred to one or other of the primary divisions of the +animal kingdom. Many fossil groups have no closely-related group +now in existence; but in no case do we meet with any grand +structural type which has not survived to the present day. +</p> + +<p class="footnote"> +[Footnote 9: In the Appendix a brief definition is given of the +sub-kingdoms, and the chief divisions of each are enumerated.] +</p> + +<p class="indent"> +The old types of life differ in many respects from those now +upon the earth; and the further back we pass in time, the more +marked does this divergence become. Thus, if we were to compare +the animals which lived in the Silurian seas with +<a name="page_60"><span class="page">Page 60</span></a> +those inhabiting our present oceans, we should in most +instances find differences so great as almost to place us in +another world. This divergence is the most marked in the +Palæozoic forms of life, less so in those of the Mesozoic +period, and less still in the Tertiary period. Each successive +formation has therefore presented us with animals becoming +gradually more and more like those now in existence; and though +there is an immense and striking difference between the Silurian +animals and those of to-day, this difference is greatly reduced +if we compare the Silurian fauna with the Devonian; <i>that</i> +again with the Carboniferous; and so on till we reach the present. +</p> + +<p class="indent"> +It follows from the above that the animals of any given formation +are more like those of the next formation below, and of the next +formation above, than they are to any others; and this fact of +itself is an almost inexplicable one, unless we believe that +the animals of any given formation are, in part at any rate, the +lineal descendants of the animals of the preceding formation, +and the progenitors, also in part at least, of the animals of the +succeeding formation. In fact, the palæontologist is so +commonly confronted with the phenomenon of closely-allied forms of +animal life succeeding one another in point of time, that he is +compelled to believe that such forms have been developed from +some common ancestral type by some process of "<i>evolution</i>." +On the other hand, there are many phenomena, such as the apparently +sudden introduction of new forms throughout all past time, and +the common occurrence of wholly isolated types, which cannot +be explained in this way. Whilst it seems certain, therefore, +that many of the phenomena of the succession of animal life in +past periods can only be explained by some law of evolution, it +seems at the same time certain that there has always been some +other deeper and higher law at work, on the nature of which it +would be futile to speculate at present. +</p> + +<p class="indent"> +Not only do we find that the animals of each successive formation +become gradually more and more like those now existing upon the +globe, as we pass from the older rocks into the newer, but we also +find that there has been a gradual progression and development in +the <i>types</i> of animal life which characterise the geological +ages. If we take the earliest-known and oldest examples of any +given group of animals, it can sometimes be shown that these +primitive forms, though in themselves highly organised, possessed +certain characters such as are now only seen in the <i>young</i> +of their existing representatives. In technical language, the +early forms of life in some +<a name="page_61"><span class="page">Page 61</span></a> +instances possess +"<i>embryonic</i>" characters, though this does not prevent them +often attaining a size much more gigantic than their nearest living +relatives. Moreover, the ancient forms of life are often what is +called "comprehensive types"—that is to say, they possess +characters in combination such as we nowadays only find separately +developed in different, groups of animals. Now, this permanent +retention of embryonic characters and this "comprehensiveness" of +structural type are signs of what a zoologist considers to be a +comparatively low grade of organisation; and the prevalence of +these features in the earlier forms of animals is a very striking +phenomenon, though they are none the less perfectly organised so +far as their own type is concerned. As we pass upwards in the +geological scale, we find that these features gradually disappear, +higher and ever higher forms are introduced, and "specialisation" +of type takes the place of the former comprehensiveness. We shall +have occasion to notice many of the facts on which these views are +based at a later period, and in connection with actual examples. +In the meanwhile, it is sufficient to state, as a widely-accepted +generalisation of palæontology, that there has been in the past +a general progression of organic types, and that the appearance +of the lower forms of life has in the main preceded that of the +higher forms in point of time. +</p> + +<h2><a name="page_63"><span class="page">Page 63</span> +PART II.</a></h2> + +<hr /> + +<p class="part"> +HISTORICAL PALÆONTOLOGY. +</p> + +<h3> +<a name="page_65"><span class="page">Page 65</span></a> +CHAPTER VII.</h3> + +<p class="subtitle"> +THE LAURENTIAN AND HURONIAN PERIODS. +</p> + +<p class="indent"> +The <i>Laurentian Rocks</i> constitute the base of the entire +stratified series, and are, therefore, the oldest sediments of +which we have as yet any knowledge. They are more largely and more +typically developed in North America, and especially in Canada, +than in any known part of the world, and they derive their title +from the range of hills which the old French geographers named +the "Laurentides." These hills are composed of Laurentian Rocks, +and form the watershed between the valley of the St Lawrence river +on the one hand, and the great plains which stretch northwards +to Hudson Bay on the other hand. The main area of these ancient +deposits forms a great belt of rugged and undulating country, +which extends from Labrador westwards to Lake Superior, and then +bends northwards towards the Arctic Sea. Throughout this extensive +area the Laurentian Rocks for the most part present themselves +in the form of low, rounded, ice-worn hills, which, if generally +wanting in actual sublimity, have a certain geological grandeur +from the fact that they "have endured the battles and the storms +of time longer than any other mountains" (Dawson). In some places, +however, the Laurentian Rocks produce scenery of the most magnificent +character, as in the great gorge cut through them by the river +Saguenay, where they rise at times into vertical precipices 1500 +feet in height. In the famous group of the Adirondack mountains, +also, in the state of New York, they form elevations no less than +6000 feet above the level of the sea. As a general rule, the +character of the Laurentian region is that of a rugged, rocky, +rolling country, often +<a name="page_66"><span class="page">Page 66</span></a> +densely timbered, but rarely well fitted for agriculture, and +chiefly attractive to the hunter and the miner. +</p> + +<p class="indent"> +As regards its mineral characters, the Laurentian series is composed +throughout of metamorphic and highly crystalline rocks, which are +in a high degree crumpled, folded, and faulted. By the late Sir +William Logan the entire series was divided into two great groups, +the <i>Lower Laurentian</i> and the <i>Upper Laurentian</i>, of +which the latter rests unconformably upon the truncated edges +of the former, and is in turn unconformably overlaid by strata +of Huronian and Cambrian age (fig. 20). +</p> + +<p class="indent"> +The <i>Lower Laurentian</i> series attains the enormous thickness + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 597px; + font-size: smaller; text-align: justify;"> +<img src="images/fig020.jpg" width="589" height="79" alt="Fig. 20" /> +<br /> +Fig. 20.—Diagrammatic section of the Laurentian +Rocks in Lower Canada. <i>a</i> Lower Laurentian; <i>b</i> Upper +Laurentian, resting unconformably upon the lower series; <i>c</i> +Cambrian strata (Potsdam Sandstone), resting unconformably on +the Upper Laurentian. +</span> +</span> + +of over 20,000 feet, and is composed mainly of great beds of gneiss, +altered sandstones (quartzites), mica-schist, hornblende-schist, +magnetic iron-ore, and hæmatite, together with masses of +limestone. The limestones are especially interesting, and have an +extraordinary development—three principal beds being known, of +which one is not less than 1500 feet thick; the collective +thickness of the whole being about 3500 feet. +</p> + +<p class="indent"> +The <i>Upper Laurentian</i> series, as before said, reposes +unconformably upon the Lower Laurentian, and attains a thickness of +at least 10,000 feet. Like the preceding, it is wholly metamorphic, +and is composed partly of masses of gneiss and quartzite; but it +is especially distinguished by the possession of great beds of +felspathic rock, consisting principally of "Labrador felspar." +</p> + +<p class="indent"> +Though typically developed in the great Canadian area already +spoken of, the Laurentian Rocks occur in other localities, both +in America and in the Old World. In Britain, the so-called +"fundamental gneiss" of the Hebrides and of Sutherlandshire is +probably of Lower Laurentian age, and the "hypersthene rocks" +of the Isle of Skye may, with great probability, be regarded +as referable to the Upper Laurentian. In other localities in +Great Britain (as in St David's, South Wales; the Malvern Hills; +and the North of Ireland) occur ancient metamorphic deposits +which also are probably referable to the Laurentian series. The +so-called "primitive gneiss" of Norway appears to belong to the +Laurentian, and the +<a name="page_67"><span class="page">Page 67</span></a> +ancient metamorphic rocks of Bohemia and Bavaria may be regarded +as being approximately of the same age. +</p> + +<p class="indent"> +By some geological writers the ancient and highly metamorphosed +sediments of the Laurentian and the succeeding Huronian series +have been spoken of as the "Azoic rocks" (Gr. <i>a</i>, without; +<i>zoe</i>, life); but even if we were wholly destitute of any +evidence of life during these periods, this name would be +objectionable upon theoretical grounds. If a general name be +needed, that of "Eozoic" (Gr. <i>eos</i>, dawn; <i>zoe</i>, life), +proposed by Principal Dawson, is the most appropriate. Owing +to their metamorphic condition, geologists long despaired of +ever detecting any traces of life in the vast pile of strata +which constitute the Laurentian System. Even before any direct +traces were discovered, it was, however, pointed out that there +were good reasons for believing that the Laurentian seas had been +tenanted by an abundance of living beings. These reasons are +briefly as follows:—(1) Firstly, the Laurentian series consists, +beyond question, of marine sediments which originally differed +in no essential respect from those which were subsequently laid +down in the Cambrian or Silurian periods. (2) In all formations +later than the Laurentian, any limestones which are present can +be shown, with few exceptions, to be <i>organic</i> rocks, and to +be more or less largely made up of the comminuted debris of marine +or fresh-water animals. The Laurentian limestones, in consequence +of the metamorphism to which they have been subjected, are so +highly crystalline (fig. 21) that the microscope fails to detect + +<span style="float: right; margin: 4px; width: 263px; + font-size: smaller; text-align: justify;"> +<img src="images/fig021.jpg" width="255" height="255" alt="Fig. 21" /> +<br /> +Fig. 21.—Section of Lower Laurentian Limestone from Hull, +Ottawa; enlarged five diameters. The rock is very highly +crystalline, and contains mica and other minerals. The irregular +black masses in it are graphite. (Original.) +</span> + +any organic structure in the rock, and no fossils beyond those +which will be spoken of immediately have as yet been discovered in +them. We know, however, of numerous cases in which limestones, +of later age, and undoubtedly organic to begin with, have been +rendered so intensely crystalline by metamorphic action that +all traces of organic structure have been obliterated. We have +therefore, by analogy, the strongest possible ground for believing +that the vast beds of Laurentian limestone have been originally +organic in their origin, and primitively composed, in the main, +of the calcareous skeletons +<a name="page_68"><span class="page">Page 68</span></a> +of marine +animals. It would, in fact, be a matter of great difficulty to +account for the formation of these great calcareous masses on any +other hypothesis. (3) The occurrence of phosphate of lime in the +Laurentian Rocks in great abundance, and sometimes in the form of +irregular beds, may very possibly be connected with the former +existence in the strata of the remains of marine animals of whose +skeleton this mineral is a constituent. (4) The Laurentian Rocks +contain a vast amount of carbon in the form of black-lead or +<i>graphite</i>. This mineral is especially abundant in the +limestones, occurring in regular beds, in veins or strings, or +disseminated through the body of the limestone in the shape of +crystals, scales, or irregular masses. The amount of graphite in +some parts of the Lower Laurentian is so great that it has been +calculated as equal to the quantity of carbon present in an equal +thickness of the Coal-measures. The general source of solid +carbon in the crust of the earth is, however, plant-life; and it +seems impossible to account for the Laurentian graphite, except +upon the supposition that it is metamorphosed vegetable matter. +(5) Lastly, the great beds of iron-ore (peroxide and magnetic +oxide) which occur in the Laurentian series interstratified with +the other rocks, point with great probability to the action of +vegetable life; since similar deposits in later formations can +commonly be shown to have been formed by the deoxidising power +of vegetable matter in a state of decay. +</p> + +<p class="indent"> +In the words of Principal Dawson, "anyone of these reasons might, +in itself, be held insufficient to prove so great and, at first +sight, unlikely a conclusion as that of the existence of abundant +animal and vegetable life in the Laurentian; but the concurrence +of the whole in a series of deposits unquestionably marine, forms +a chain of evidence so powerful that it might command belief +even if no fragment of any organic or living form or structure +had ever been recognised in these ancient rocks." Of late years, +however, there have been discovered in the Laurentian Rocks certain +bodies which are believed to be truly the remains of animals, and +of which by far the most important is the structure known under +the now celebrated name of <i>Eozoön</i>. If truly organic, +a very special and exceptional interest attaches itself to +<i>Eozoön</i>, as being the most ancient fossil animal of which +we have any knowledge; but there are some who regard it really +a peculiar form of mineral structure, and a severe, protracted, +and still unfinished controversy has been carried on as to its +nature. Into this controversy it is wholly unnecessary to enter +here; and it will be sufficient to briefly explain the structure +of <i>Eozoön</i>, as elucidated by the elaborate and masterly +investigations of Carpenter +<a name="page_69"><span class="page">Page 69</span></a> +and Dawson, from the standpoint that it is a genuine +organism—the balance of evidence up to this moment inclining +decisively to this view. +</p> + +<p class="indent"> +The structure known as <i>Eozoön</i> is found in various +localities in the Lower Laurentian limestones of Canada, in the form +of isolated masses or spreading layers, which are composed of thin +alternating laminæ, arranged more or less concentrically (fig. +22). The laminæ of these masses are usually of different colours + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 488px; + font-size: smaller; text-align: center;"> +<img src="images/fig022.jpg" width="480" height="255" alt="Fig. 22" /> +<br /> +Fig. 22.—Fragment of <i>Eozoön</i>, of the natural size, +showing alternate laminæ of loganite and dolomite. (After +Dawson.) +</span> +</span> + +and composition; one series being white, and composed of carbonate +of lime—whilst the laminæ of the second series alternate +with the preceding, are green in colour, and are found by chemical +analysis to consist of some silicate, generally serpentine or the +closely-related "loganite." In some instances, however, all the +laminæ are calcareous, the concentric arrangement still remaining +visible in consequence of the fact that the laminæ are composed +alternately of lighter and darker coloured limestone. +</p> + +<p class="indent"> +When first discovered, the masses of <i>Eozoön</i> were supposed +to be of a mineral nature; but their striking general resemblance +to the undoubted fossils which will be subsequently spoken of under +the name of <i>Stromatopora</i> was recognised by Sir William +Logan, and specimens were submitted for minute examination, first +to Principal Dawson, and subsequently to Dr W. B. Carpenter. +After a careful microscopic examination, these two distinguished +observers came to the conclusion that <i>Eozoön</i> was truly +organic, and in this opinion they were afterwards corroborated +by other high authorities (Mr W. K. Parker, Professor Rupert +Jones, Mr H. B. Brady, Professor Gümbel, &c.) Stated briefly, +the structure of <i>Eozoön</i>, as exhibited by the microscope, +is as follows:— +</p> + +<p class="indent"> +<a name="page_70"><span class="page">Page 70</span></a> +The concentrically-laminated mass of <i>Eozoön</i> is composed +of numerous calcareous layers, representing the original skeleton +of the organism (fig. 23, <i>b</i>). These calcareous layers + +<span style="float: left; margin: 4px; width: 342px; + font-size: smaller; text-align: justify;"> +<img src="images/fig023.jpg" width="334" height="273" alt="Fig. 23" /> +<br /> +Fig. 23.—Diagram of a portion of <i>Eozoön</i> +cut vertically. A, B, C, Three tiers of chambers communicating +with one another by slightly constricted apertures: <i>a a</i>, +The true shell-wall, perforated by numerous delicate tubes; <i>b +b</i>. The main calcareous skeleton ("intermediate skeleton"); +<i>c</i>, Passage of communication ("stolon-passage") from one +tier of chambers to another; <i>d</i>, Ramifying tubes in the +calcareous skeleton. (After Carpenter.) +</span> + +serve to separate and define a series of chambers arranged in +successive tiers, one above the other (fig. 23, A, B, C); and +they are perforated not only by passages (fig. 23, <i>c</i>), +which serve to place successive tiers of chambers in communication, +but also by a system of delicate branching canals (fig. 23, +<i>d</i>). Moreover, the central and principal portion of each +calcareous layer, with the ramified canal-system just spoken +of, is bounded both above and below by a thin lamina which has +a structure of its own, and which may be regarded as the proper +shell-wall (fig. 23, <i>a a</i>). This proper wall forms the +actual lining of the chambers, as well as the outer surface of +the whole mass; and it is perforated with numerous fine vertical +tubes (fig. 24, <i>a a</i>), opening into the chambers and on to +the surface by corresponding fine pores. From the resemblance +of this tubulated layer to similar structures in the shell of +the Nummulite, it is often spoken of as the "Nummuline layer." +The chambers are sometimes piled up one above the other in an +irregular manner; but they are more commonly arranged in regular +tiers, the separate chambers being marked off from one another +by projections of the wall in the form of partitions, which are +so far imperfect as to allow of a free communication between +contiguous chambers. In the original condition of the organism, +all these chambers, of course, must have been filled with +living-matter; but they are found in the present state of the +fossil to be generally filled with some silicate, such as serpentine, +which not only fills the actual chambers, but has also penetrated +the minute tubes of the proper wall and the branching canals of +the intermediate skeleton. In some cases +<a name="page_71"><span class="page">Page 71</span></a> +the chambers are simply filled with crystalline carbonate of lime. +When the originally porous fossil has been permeated by a silicate, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 396px; + font-size: smaller; text-align: justify;"> +<img src="images/fig024.jpg" width="388" height="355" alt="Fig. 24" /> +<br /> +Fig. 24.—The animal of <i>Nonionina</i>, one of the +<i>Foraminifera</i>, after the shell has been removed by +a weak acid; <i>b, Gromia</i>, a single-chambered Foraminifer +(after Schultze), showing the shell surrounded by a network of +filaments derived from the body substance. +</span> +</span> + +it is possible to dissolve away the whole of the calcareous skeleton +by means of acids, leaving an accurate and beautiful cast of the +chambers and the tubes connected with them in the insoluble silicate. +</p> + +<p class="indent"> +The above are the actual appearances presented by <i>Eozoön</i> +when examined microscopically, and it remains to see how far they +enable us to decide upon its true position in the animal kingdom. +Those who wish to study this interesting subject in detail must +consult the admirable memoirs by Dr W. B. Carpenter and Principal +Dawson: it will be enough here to indicate the results which +have been arrived at. The only animals at the present day which +possess a continuous calcareous skeleton, perforated by pores and +penetrated by canals, are certain organisms belonging to the +group of the <i>Foraminifera</i>. We have had occasion before +to speak of these animals, and as they are not conspicuous or +commonly-known forms of life, it may be well to say a few words as +to the structure of the living representatives of the group. The +<i>Foraminifera</i> are all inhabitants of the sea, and are mostly +of small or even microscopic dimensions. Their bodies are composed +<a name="page_72"><span class="page">Page 72</span></a> +of an apparently structureless animal +substance of an albuminous nature ("sarcode"), of a gelatinous +consistence, transparent, and exhibiting numerous minute granules +or rounded particles. The body-substance cannot be said in itself +to possess any definite form, except in so far as it may be bounded +by a shell; but it has the power, wherever it may be exposed, of +emitting long thread-like filaments ("pseudopodia"), which interlace +with one another to form a network (fig. 25, <i>b</i>). These + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 370px; + font-size: smaller; text-align: justify;"> +<img src="images/fig025.jpg" width="362" height="557" alt="Fig. 25" /> +<br /> +Fig. 25.—The animal of <i>Nonionina</i>, one of the +<i>Foraminifera</i>, after the shell has been removed by +a weak acid; <i>b, Gromia</i>, a single-chambered Foraminifer +(after Schultze), showing the shell surrounded by a network of +filaments derived from the body substance. +</span> +</span> + +filaments can be thrown out at will, and to considerable distances, +and can be again retracted into the soft mass of the general +body-substance, and they are the agents by which the animal obtains +its food. The soft bodies of the <i>Foraminifera</i> are protected +by a shell, which is usually calcareous, but may be composed of +sand-grains cemented +<a name="page_73"><span class="page">Page 73</span></a> +together; and it may consist +of a single chamber (fig. 26, <i>a</i>), or of many chambers arranged +in different ways (fig. 26, <i>b-f</i>). Sometimes the shell has but + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 564px; + font-size: smaller; text-align: justify;"> +<img src="images/fig026.jpg" width="556" height="485" alt="Fig. 26" /> +<br /> +Fig. 26.—Shells of living <i>Foraminifera</i>. +<i>a, Orbulina universa</i>, in its perfect condition, showing +the tubular spines which radiate from the surface of the shell; +<i>b, Globigerina bulloides</i>, in its ordinary condition, the +thin hollow spines which are attached to the shell when perfect +having been broken off; <i>c, Textularia variabilis; d, Peneroplis +planatus; e, Rotalia concamerata; f, Cristellaria subarcuatula.</i> +[Fig. <i>a</i> is after Wyville Thomson; the others are after +Williamson. All the figures are greatly enlarged. +</span> +</span> + +one large opening into it—the mouth; and then it is from this +aperture that the animal protrudes the delicate net of filaments +with which it seeks its food. In other cases the entire shell +is perforated with minute pores (fig. 26, <i>e</i>), through +which the soft body-substance gains the exterior, covering the +whole shell with a gelatinous film of animal matter, from which +filaments can be emitted at any point. When the shell consists +of many chambers, all of these are placed in direct communication +with one another, and the actual substance of the shell is often +traversed by minute canals filled with living matter (<i>e.g.</i>, +in <i>Calcarina</i> and <i>Nummulina</i>). The shell, therefore, +may be regarded, in such cases, as a more or less completely +porous calcareous structure, +<a name="page_74"><span class="page">Page 74</span></a> +filled to its minutest internal recesses with the substance of the +living animal, and covered externally with a layer of the same +substance, giving off a network of interlacing filaments. +</p> + +<p class="indent"> +Such, in brief, is the structure of the living <i>Foraminifera</i>; +and it is believed that in <i>Eozoön</i> we have an extinct +example of the same group, not only of special interest from its +immemorial antiquity, but hardly less striking from its gigantic +dimensions. In its original condition, the entire chamber-system +of <i>Eozoön</i> is believed to have been filled with soft +structureless living matter, which passed from chamber to chamber +through the wide apertures connecting these cavities, and from +tier to tier by means of the tubuli in the shell-wall and the +branching canals in the intermediate skeleton. Through the +perforated shell-wall covering the outer surface the soft +body-substance flowed out, forming a gelatinous investment, from +every point of which radiated an interlacing net of delicate +filaments, providing nourishment for the entire colony. In its +present state, as before said, all the cavities originally +occupied by the body-substance have been filled with some mineral +substance, generally with one of the silicates of magnesia; and +it has been asserted that this fact militates strongly against +the organic nature of <i>Eozoön</i>, if not absolutely +disproving it. As a matter of fact, however—as previously +noticed—it is by no means very uncommon at the present day +to find the shells of living species of <i>Foraminifera</i> in +which all the cavities primitively occupied by the body-substance, +down to the minutest pores and canals, have been similarly +injected by some analogous silicate, such as glauconite. +</p> + +<p class="indent"> +Those, then, whose opinions on such a subject deservedly carry the +greatest weight, are decisively of opinion that we are presented +in the <i>Eozoön</i> of the Laurentian Rocks of Canada with an +ancient, colossal, and in some respects abnormal type of the +<i>Foraminifera</i>. In the words of Dr Carpenter, it is not +pretended that "the doctrine of the Foraminiferal nature of +<i>Eozoön</i> can be <i>proved</i> in the demonstrative sense;" +but it may be affirmed "that the <i>convergence of a number of +separate and independent probabilities</i>, all accordant with +that hypothesis, while a separate explanation must be invented +for each of them on any other hypothesis, gives it that <i>high +probability</i> on which we rest in the ordinary affairs of life, +in the verdicts of juries, and in the interpretation of geological +phenomena generally." +</p> + +<p class="indent"> +It only remains to be added, that whilst <i>Eozoön</i> is by far +the most important organic body hitherto found in the Laurentian, +and has been here treated at proportionate length, other +<a name="page_75"><span class="page">Page 75</span></a> +traces of life have been detected, which may subsequently prove of +great interest and importance. Thus, Principal Dawson has recently +described under the name of <i>ArchĹ“osphĹ“rinĹ“</i> +certain singular rounded bodies which he has discovered in the +Laurentian limestones, and which he believes to be casts of the +shells of <i>Foraminifera</i> possibly somewhat allied to the +existing <i>GlobigerinĹ“</i>. The same eminent +palæontologist has also described undoubted worm-burrows +from rocks probably of Laurentian age. Further and more extended +researches, we may reasonably hope, will probably bring to light +other actual remains of organisms in these ancient deposits. +</p> + +<h4>THE HURONIAN PERIOD.</h4> + +<p class="indent"> +The so-called <i>Huronian Rocks</i>, like the Laurentian, have +their typical development in Canada, and derive their name from +the fact that they occupy an extensive area on the borders of +Lake Huron. They are wholly metamorphic, and consist principally +of altered sandstones or quartzites, siliceous, felspathic, or +talcose slates, conglomerates, and limestones. They are largely +developed on the north shore of Lake Superior, and give rise +to a broken and hilly country, very like that occupied by the +Laurentians, with an abundance of timber, but rarely with sufficient +soil of good quality for agricultural purposes. They are, however, +largely intersected by mineral veins, containing silver, gold, +and other metals, and they will ultimately doubtless yield a rich +harvest to the miner. The Huronian Rocks have been identified, +with greater or less certainty, in other parts of North America, +and also in the Old World. +</p> + +<p class="indent"> +The total thickness of the Huronian Rocks in Canada is estimated +as being not less than 18,000 feet, but there is considerable +doubt as to their precise geological position. In their typical +area they rest unconformably on the edges of strata of <i>Lower</i> +Laurentian age; but they have never been seen in direct contact +with the <i>Upper</i> Laurentian, and their exact relations to +this series are therefore doubtful. It is thus open to question +whether the Huronian Rocks constitute a distinct formation, to +be intercalated in point of time between the Laurentian and the +Cambrian groups; or whether, rather, they should not be considered +as the metamorphosed representatives of the Lower Cambrian Rocks +of other regions. +</p> + +<p class="indent"> +As regards the fossils of the Huronian Rocks, little can be said. +Some of the specimens of <i>Eozoön Canadense</i> which have +<a name="page_76"><span class="page">Page 76</span></a> +been discovered in Canada are thought to come +from rocks which are probably of Huronian age. In Bavaria, +Dr Gümbel has described a species of <i>Eozoön</i> under +the name of <i>Eozoön Bavaricum</i>, from certain metamorphic +limestones which he refers to the Huronian formation. Lastly, the +late Mr Billings described, from rocks in Newfoundland apparently +referable to the Huronian, certain problematical limpet-shaped +fossils, to which he gave the name of <i>Aspidella</i>. +</p> + +<h4>LITERATURE.</h4> + +<p class="indent"> +Amongst the works and memoirs which the student may consult with +regard to the Laurentian and Huronian deposits may be mentioned +the following:[10]— +</p> + +<table summary="" style=""> +<tr><td class="right" valign="top">(1)</td> + <td>'Report of Progress of the Geological Survey of Canada + from its Commencement to 1863,' pp. 38-49, and pp. 50-66.</td> +</tr><tr><td class="right" valign="top">(2)</td> + <td>'Manual of Geology.' Dana. 2d Ed. 1875.</td> +</tr><tr><td class="right" valign="top">(3)</td> + <td>'The Dawn of Life.' J. W, Dawson. 1876.</td> +</tr><tr><td class="right" valign="top">(4)</td> + <td>"On the Occurrence of Organic Remains in the Laurentian + Rocks of Canada." Sir W. E. Logan. 'Quart. Journ. Geol.' + Soc.,' xxi. 45-50.'</td> +</tr><tr><td class="right" valign="top">(5)</td> + <td>"On the Structure of Certain Organic Remains in the + Laurentian Limestones of Canada." J. W. Dawson. 'Quart. + Journ. Geol. Soc.,' xxi. 51-59.</td> +</tr><tr><td class="right" valign="top">(6)</td> + <td>"Additional Note on the Structure and Affinities of + Eozoön Canadense." W. B, Carpenter. 'Quart. Journ. + Geol. Soc.,' xxi. 59-66.</td> +</tr><tr><td class="right" valign="top">(7)</td> + <td>"Supplemental Notes on the Structure and Affinities of + Eozoön' Canadense," W. B. Carpenter, 'Quart. Journ. + Geol. Soc.,' xxii. 219-228.</td> +</tr><tr><td class="right" valign="top">(8)</td> + <td>"On the So-Called Eozoönal Rocks." King & Rowney. + 'Quart. Journ. Geol. Soc.,' xxii. 185-218.</td> +</tr><tr><td class="right" valign="top">(9)</td> + <td>'Chemical and Geological Essays.' Sterry Hunt.</td> +</tr> +</table> + +<p class="indent"> +The above list only includes some of the more important memoirs +which may be consulted as to the geological and chemical features +of the Laurentian and Huronian Rocks, and as to the true nature +of <i>Eozoön</i>. Those who are desirous of studying the later +phases of the controversy with regard to <i>Eozoön</i> must consult +the papers of Carpenter, Carter, Dawson, King & Rowney, Hahn, +and others, in the 'Quart. Journ. of the Geological Society,' +the 'Proceedings of the Royal Irish Academy,' the 'Annals of +Natural History,' the 'Geological Magazine,' &c. Dr Carpenter's +'Introduction to the Study of the Foraminifera' should also be +consulted. +</p> + +<p class="footnote"> +[Footnote 10: In this and in all subsequently following +bibliographical lists, not only is the selection of works and +memoirs quoted necessarily extremely limited; but only such have, +as a general rule, been chosen for mention as are easily accessible +to students who are in the position of being able to refer to a good +library. Exceptions, however, are occasionally made to this rule, +in favour of memoirs or works of special historical interest. It +is also unnecessary to add that it has not been thought requisite +to insert in these lists the well-known handbooks of geological +and palæontological science; except in such instances as where +they contain special information on special points.] +</p> + +<h3> +<a name="page_77"><span class="page">Page 77</span></a> +CHAPTER VIII.</h3> + +<p class="subtitle"> +THE CAMBRIAN PERIOD. +</p> + +<p class="indent"> +The traces of life in the Laurentian period, as we have seen, are +but scanty; but the <i>Cambrian Rocks</i>—so called from their +occurrence in North Wales and its borders ("Cambria ")—have +yielded numerous remains of animals and some dubious plants. The +Cambrian deposits have thus a special interest as being the oldest +rocks in which occur any number of well-preserved and unquestionable +organisms. We have here the remains of the first <i>fauna</i>, or +assemblage of animals, of which we have at present knowledge. +As regards their geographical distribution, the Cambrian Rocks +have been recognised in many parts of the world, but there is +some question as to the precise limits of the formation, and +we may consider that their most typical area is in South Wales, +where they have been carefully worked out, chiefly by Dr Henry +Hicks. In this region, in the neighbourhood of the promontory +of St David's, the Cambrian Rocks are largely developed, resting +upon an ancient ridge of Pre-Cambrian (Laurentian?) strata, and +overlaid by the lowest beds of the Lower Silurian. The subjoined +sketch-section (fig. 27) exhibits in a general manner the succession +of strata in this locality. +</p> + +<p class="indent"> +From this section it will be seen that the Cambrian Rocks in Wales +are divided in the first place into a lower and an upper group. +The <i>Lower Cambrian</i> is constituted at the base by a great +series of grits, sandstones, conglomerates, and slates, which +are known as the "Longmynd group," from their vast development in +the Longmynd Hills in Shropshire, and which attain in North Wales +a thickness of 8000 feet or more. The Longmynd beds are succeeded +by the so-called "Menevian group," a series of sandstones, flags, +and grits, about 600 feet in thickness, and containing a +considerable number of fossils. The <i>Upper Cambrian</i> series +consists in its lower portion of nearly 5000 feet of strata, +principally shaly and slaty, which are known as the "Lingula +Flags," from the great abundance in them of a shell referable +to the genus <i>Lingula</i>. These are followed by 1000 feet +of dark shales and flaggy sandstones, which are known as the +"Tremadoc slates," from their occurrence near Tremadoc in North +Wales; and these in turn are surmounted, apparently quite +conformably, by the basement beds of the Lower Silurian. +</p> + +<p class="indent"> +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 551px; + font-size: smaller; text-align: center;"> +<a name="page_78"><span class="page">Page 78</span></a> +GENERALIZED SECTION OF THE CAMBRIAN ROCKS IN WALES +<br /> +Fig. 27. +<br /> +<img src="images/fig027.jpg" width="543" height="700" alt="Fig. 27" /> +</span> +</span> +The above may be regarded as giving a typical series of the Cambrian +Rocks in a typical locality; but strata of Cambrian age are known in +many other regions, of which it is only possible here to allude to +a few of the most important. In Scandinavia occurs a well-developed +series of Cambrian deposits, representing both the lower and upper +parts of the +<a name="page_79"><span class="page">Page 79</span></a> +formation. In Bohemia, the +Upper Cambrian, in particular, is largely developed, and constitutes +the so-called "Primordial zone" of Barrande. Lastly, in North +America, whilst the Lower Cambrian is only imperfectly developed, +or is represented by the Huronian, the Upper Cambrian formation has +a wide extension, containing fossils similar in character to the +analogous strata in Europe, and known as the "Potsdam Sandstone." +The subjoined table shows the chief areas where Cambrian Rocks are +developed, and their general equivalency: +</p> + +<div class="center"> +<table border="1" cellspacing="0" width="90%"> + +<tr> + <td colspan="4" class="center">TABULAR VIEW OF THE CAMBRIAN + FORMATION.</td> +</tr> + +<tr> + <td> </td> + <td class="center"><i>Britain.</i></td> + <td class="center"><i>Europe.</i></td> + <td class="center"><i>America.</i></td> +</tr> + +<tr> + <td rowspan="2" class="center">Upper Cambrian.</td> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>a</i>.</td> + <td valign="top" class="left">Tremadoc Slates.</td></tr> + </table></td> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>a</i>.</td> + <td valign="top" class="left">Primordial zone of + Bohemia.</td></tr> + </table></td> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>a</i>.</td> + <td valign="top" class="left">Potsdam Sandstone.</td></tr> + </table></td> +</tr> + +<tr> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>b</i>.</td> + <td valign="top" class="left">Lingula Flags.</td></tr> + </table></td> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>b</i>.</td> + <td valign="top" class="left">Paradoxides Schists, + Olenus Schists, and Dictyonema schists of + Sweden.</td></tr> + </table></td> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>b</i>.</td> + <td valign="top" class="left">Acadian group of + New Brunswick.</td></tr> + </table></td> +</tr> + +<tr> + <td rowspan="6" class="center">Lower Cambrian.</td> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>a</i>.</td> + <td valign="top" class="left">Longmynd Beds.</td></tr> + </table></td> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>a</i>.</td> + <td valign="top" class="left">Fucoidal Sandstone of + Sweden.</td></tr> + </table></td> + <td valign="top" class="left">Huronian + Formation?</td> +</tr> + +<tr> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>b</i>.</td> + <td valign="top" class="left">Llanberis Slates.</td></tr> + </table></td> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>b</i>.</td> + <td valign="top" class="left"><i>Eophyton</i> Sandstone + of Sweden.</td></tr> + </table></td> + <td> </td> +</tr> + +<tr> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>c</i>.</td> + <td valign="top" class="left">Harlech Grits.</td></tr> + </table></td> + <td> </td> + <td> </td> +</tr> + +<tr> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>d</i>.</td> + <td valign="top" class="left"><i>Oldhamia</i> Slates + of Ireland.</td></tr> + </table></td> + <td> </td> + <td> </td> +</tr> + +<tr> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>e</i>.</td> + <td valign="top" class="left">Conglomerates and + Sandstones of Sutherlandshire?</td></tr> + </table></td> + <td> </td> + <td> </td> +</tr> + +<tr> + <td valign="top" class="left"> + <table border="0" cellspacing="0" cellpadding="1"> + <tr><td valign="top" class="left"><i>f</i>.</td> + <td valign="top" class="left">Menevian + Beds.</td></tr> + </table></td> + <td> </td> + <td> </td> +</tr> + +</table> +</div> + +<p class="indent"> +Like all the older Palæozoic deposits, the Cambrian Rocks, +though by no means necessarily what would be called actually +"metamorphic," have been highly cleaved, and otherwise altered +from their original condition. Owing partly to their indurated +state, and partly to their great antiquity, they are usually +found in the heart of mountainous districts, which have undergone +great disturbance, and have been subjected to an enormous amount +of denudation. In some cases, as in the Longmynd Hills in +Shropshire, they form low rounded elevations, largely covered by +pasture, and with few or no elements of sublimity. In other cases, +however, they rise into bold and rugged mountains, girded by +precipitous cliffs. Industrially, the Cambrian Rocks are of +interest, if only for the reason that the celebrated Welsh slates +of Llanberis are derived from highly-cleaved beds of this age. +Taken as a whole, the Cambrian formation is essentially composed +of arenaceous and +<a name="page_80"><span class="page">Page 80</span></a> +muddy sediments, the +latter being sometimes red, but more commonly nearly black in +colour. It has often been supposed that the Cambrians are a +deep-sea deposit, and that we may thus account for the few +fossils contained in them; but the paucity of fossils is to a +large extent imaginary, and some of the Lower Cambrian beds +of the Longmynd Hills would appear to have been laid down in +shallow water; as they exhibit rain-prints, sun-cracks, and +ripple-marks—incontrovertible evidence of their having been +a shore-deposit. The occurrence, of innumerable worm-tracks and +burrows in many Cambrian strata is also a proof of shallow-water +conditions; and the general absence of limestones, coupled with +the coarse mechanical nature of many of the sediments of the +Lower Cambrian, maybe taken as pointing in the same direction. +</p> + +<p class="indent"> +The <i>life</i> of the Cambrian, though not so rich as in the +succeeding Silurian period, nevertheless consists of representatives +of most of the great classes of invertebrate animals. The coarse +sandy deposits of the formation, which abound more particularly +towards its lower part, naturally are to a large extent barren +of fossils; but the muddy sediments, when not too highly cleaved, +and especially towards the summit of the group, are replete with +organic remains. This is also the case, in many localities at any +rate, with the finer beds of the Potsdam Sandstone in America. +Limestones are known to occur in only a few areas (chiefly in +America), and this may account for the apparent total absence +of corals. It is, however, interesting to note that, with this +exception, almost all the other leading groups of Invertebrates +are known to have come into existence during the Cambrian period. +</p> + +<p class="indent"> +Of the land-surfaces of the Cambrian period we know nothing; +and there is, therefore, nothing surprising in the fact that +our acquaintance with the Cambrian vegetation is confined to +some marine plants or sea-weeds, often of a very obscure and +problematical nature. The "Fucoidal Sandstone" of Sweden, and the +"Potsdam Sandstone" of North America, have both yielded numerous +remains which have been regarded as markings left by sea-weeds or +"Fucoids;" but these are highly enigmatical in their characters, +and would, in many instances, seem to be rather referable to the +tracks and burrows of marine worms. The first-mentioned of these +formations has also yielded the curious, furrowed and striated +stems which have been described as a kind of land-plant under the +name of <i>Eopkyton</i> (fig. 28). It cannot be said, however, +that the vegetable origin of these singular bodies has been +satisfactorily proved. Lastly, there are found in certain green +<a name="page_81"><span class="page">Page 81</span></a> +and purple beds of Lower Cambrian age at Bray Head, Wicklow, +Ireland, some very remarkable fossils, which are well known under + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 436px; + font-size: smaller; text-align: center;"> +<img src="images/fig028.jpg" width="428" height="536" alt="Fig. 28" /> +<br /> +Fig. 28.—Fragment of <i>Eophyton Linneanum</i>, a supposed +land-plant. Lower Cambrian, Sweden, of the natural size. +</span> +</span> + +the name of <i>Oldhamia</i>, but the true nature of which is +very doubtful. The commonest form of <i>Oldhamia</i> (fig. 29) +consists of a thread-like stem or axis, from which spring at +regular intervals bundles of short filamentous branches in a +fan-like manner. In the locality where it occurs, the fronds +of <i>Oldhamia</i> are very abundant, and are spread over the +surfaces of the strata in tangled layers. That it is organic +is certain, and that it is a calcareous sea-weed is probable; +but it may possibly belong to the sea-mosses (<i>Polyzoa</i>), +or to the sea-firs (<i>Sertularians</i>). +</p> + +<p class="indent"> +Amongst the lower forms of animal life (<i>Protozoa</i>), we +find the Sponges represented by the curious bodies, composed +of netted fibres, to which the name of <i>Protospongia</i> has +been given (fig. 32, <i>a</i>); and the comparatively gigantic, +conical, or +<a name="page_82"><span class="page">Page 82</span></a> +cylindrical fossils termed <i>ArchĹ“ocyathus</i> by Mr +Billings are certainly referable either to the <i>Foraminifera</i> + +<span style="float: left; margin: 4px; width: 243px; + font-size: smaller; text-align: justify;"> +<img src="images/fig029.jpg" width="235" height="330" alt="Fig. 29" /> +<br /> +Fig. 29.—A portion of <i>Oldhamia antiqua</i>, Lower +Cambrian, Wicklow, Ireland, of the natural size. (After Salter.) +</span> + +or to the Sponges. The almost total absence of limestones in +the formation may be regarded as a sufficient explanation of +the fact that the <i>Foraminifera</i> are not more largely and +unequivocally represented; though the existence of greensands +in the Cambrian beds of Wisconsin and Tennessee may be taken as +an indication that this class of animals was by no means wholly +wanting. The same fact may explain the total absence of corals, +so far as at present known. +</p> + +<p class="indent"> +The group of the <i>Echinodermata</i> (Sea-lilies, Sea-urchins, +and their allies) is represented by a few forms, which are +principally of interest as being the earliest-known examples of +the class. It is also worthy of note that these precursors of a +group which subsequently attains such geological importance, are +referable to no less than three distinct <i>orders</i>—the +Crinoids or Sea-lilies, represented by a species of +<i>Dendrocrinus</i>; the Cystideans by <i>Protocystites</i>; and +the Star-fishes by <i>Palasterina</i> and some other forms. Only +the last of these groups, however, appears to occur in the Lower +Cambrian. +</p> + +<p class="indent"> +The Ringed-worms (<i>Annelida</i>), if rightly credited with all +the remains usually referred to them, appear to have swarmed in +the Cambrian seas. Being soft-bodied, we do not find the actual +worms themselves in the fossil condition, but we have, nevertheless, +abundant traces of their existence. In some cases we find vertical +burrows of greater or less depth, often expanded towards their +apertures, in which the worm must have actually lived (fig. 30), +as various species do at the present day. In these cases, the +tube must have been rendered more or less permanent by receiving +a coating of mucus, or perhaps a genuine membranous secretion, +from the body of the animal; and it may be found quite empty, or +occupied by a cast of sand or mud. Of this nature are the burrows +which have been described under the names of <i>Scolithus</i> and +<i>Scolecoderma</i>, and probably the <i>Histioderma</i> of the +Lower Cambrian +<a name="page_83"><span class="page">Page 83</span></a> +of Ireland. In other cases, as in <i>Arenicolites</i> +(fig. 32, <i>b</i>), the worm seems to have inhabited a double + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 380px; + font-size: smaller; text-align: center;"> +<img src="images/fig030.jpg" width="372" height="306" alt="Fig. 30" /> +<br /> +Fig. 30.—Annelide-burrows (<i>Scolithus linearus</i>) +from the Potsdam Sandstone of Canada, of the natural size. (After +Billings.) +</span> +</span> + +burrow, shaped like the letter U, and having two openings placed +close together on the surface of the stratum. Thousands of these +twin-burrows occur in some of the strata of the Longmynd, and +it is supposed that the worm used one opening to the burrow as +an aperture of entrance, and the other as one of exit. In other +cases, again, we find simply the meandering trails caused by the +worm dragging its body over the surface of the mud. Markings of +this kind are commoner in the Silurian Rocks, and it is generally +more or less doubtful whether they may not have been caused by +other marine animals, such as shellfish, whilst some of them +have certainly nothing whatever to do with the worms. Lastly, +the Cambrian beds often show twining cylindrical bodies, commonly +more or less matted together, and not confined to the surfaces +of the strata, but passing through them. These have often been +regarded as the remains of sea-weeds, but it is more probable +that they represent casts of the underground burrows of worms +of similar habits to the common lob-worm (<i>Arenicola</i>) of +the present day. +</p> + +<p class="indent"> +The <i>Articulate</i> animals are numerously represented in the +Cambrian deposits, but exclusively by the class of +<i>Crustaceans</i>. Some of these are little double-shelled +creatures, resembling our living water-fleas (<i>Ostracoda</i>). A +few are larger forms, and belong to the same group as the existing +brine-shrimps and fairy-shrimps (<i>Phyllopoda</i>). One of the +most characteristic +<a name="page_84"><span class="page">Page 84</span></a> +of these is the +<i>Hymenocaris vermicauda</i> of the Lingula Flags (fig. +32, <i>d</i>). By far the larger number of the Cambrian +<i>Crustacea</i> belong, however, to the remarkable and wholly +extinct group of the <i>Trilobites</i>. These extraordinary +animals must have literally swarmed in the seas of the later +portion of this and the whole of the succeeding period; and they +survived in greatly diminished numbers till the earlier portion +of the Carboniferous period. They died out, however, wholly before +the close of the Palæozoic epoch, and we have no Crustaceans +at the present day which can be considered as their direct +representatives. They have, however, relationships of a more or +less intimate character with the existing groups of the Phyllopods, +the King-crabs (<i>Limulus</i>), and the Isopods ("Slaters," +Wood-lice, &c.) Indeed, one member of the last-mentioned +order, namely, the <i>Serolis</i> of the coasts of Patagonia, has +been regarded as the nearest living ally of the Trilobites. Be +this as it may, the Trilobites possessed a skeleton which, though +capable of undergoing almost endless variations, was wonderfully +constant in its pattern of structure, and we may briefly describe +here the chief features of this. +</p> + +<p class="indent"> +The upper surface of the body of a Trilobite was defended by a +strong shell or "crust," partly horny and partly calcareous in +its composition. This shell (fig. 31) generally exhibits a very +distinct "trilobation" or division into three longitudinal lobes, +one central and two lateral. It also exhibits a more important and +more fundamental division into three transverse portions, which +are so loosely connected with one another as very commonly to be +found separate. The first and most anterior of these divisions +is a shield or buckler which covers the head; the second or middle +portion is composed of movable rings covering the trunk ("thorax +"); and the third is a shield which covers the tailor "abdomen." +The head-shield (fig. 31, <i>e</i>) is generally more or less +semicircular in shape; and its central portion, covering the +stomach of the animal, is usually strongly elevated, and generally +marked by lateral furrows. A little on each side of the head are +placed the eyes, which are generally crescentic in shape, and +resemble the eyes of insects and many existing Crustaceans in being +"compound," or made up of numerous simple eyes aggregated together. +So excellent is the state of preservation of many specimens of +Trilobites, that the numerous individual lenses of the eyes have +been uninjured, and as many as four hundred have been counted in +each eye of some forms. The eyes may be supported upon prominences, +but they are never carried on movable stalks (as they are in +the existing lobsters and crabs); and +<a name="page_85"><span class="page">Page 85</span></a> +in some of the Cambrian +Trilobites, such as the little <i>Agnosti</i> (fig. 31 <i>g</i>), +the animal was blind. The lateral portions of the head-shield +are usually separated from the central portion by a peculiar + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 575px; + font-size: smaller; text-align: justify;"> +<img src="images/fig031.jpg" width="569" height="444" alt="Fig. 31" /> +<br /> +Fig. 31.—Cambrian Trilobites: <i>a, Paradoxides +Bohemicus</i>, reduced in size; <i>b, Ellipsocephalus Hoffi; c, +Sao hirsuta; d, Conocorypke Sultzeri</i> (all the above, together +with fig. <i>g</i>, are from the Upper Cambrian or "Primordial +Zone" of Bohemia); <i>e</i>, Head-shield of <i>Dikellocephalus +Celticus</i>, from the Lingula Flags of Wales; <i>f</i>, Head-shield +of <i>Conocoryphe Matthewi</i>, from the Upper Cambrian (Acadian +Group) of New Brunswick; <i>g, Agnostus rex</i>, Bohemia; <i>h</i>, +Tail-shield of <i>Dikellocephalus Minnesotensis</i>, from the +Upper Cambrian (Potsdam Sandstone) of Minnesota. (After Barrande, +Dawson, Salter, and Dale Owen.) +</span> +</span> + +line of division (the so-called "facial suture") on each side; +but this is also wanting in some of the Cambrian species. The +backward angles of the head-shield, also, are often prolonged +into spines, which sometimes reach a great length. Following +the head-shield behind, we have a portion of the body which is +composed of movable segments or "body-rings," and which is +technically called the "thorax," Ordinarily, this region is +strongly trilobed, and each ring consists of a central convex +portion, and of two flatter side-lobes. The number of body-rings +in the thorax is very variable (from two to twenty-six), but is +fixed for the adult forms of each group of the Trilobites. The +young forms have much fewer rings than the full-grown ones; and +it is curious to find that the Cambrian +<a name="page_86"><span class="page">Page 86</span></a> +Trilobites very commonly have either a great many rings (as in +<i>Paradoxides</i>, fig. 31, <i>a</i>), or else very few (as in +<i>Agnostus</i>, fig. 31, <i>g</i>). In some instances, the +body-rings do not seem to have been so constructed as to allow +of much movement, but in other cases this region of the body is +so flexible that the animal possessed the power of rolling itself +up completely, like a hedgehog; and many individuals have been +permanently preserved as fossils in this defensive condition. +Finally, the body of the Trilobite was completed by a tail-shield +(technically termed the "pygidium"), which varies much in size +and form, and is composed of a greater or less number of rings, +similar to those which form the thorax, but immovably +amalgamated with one another (fig. 31, <i>h</i>). +</p> + +<p class="indent"> +The under surface of the body in the Trilobites appears to have +been more or less entirely destitute of hard structures, with the +exception of a well-developed upper lip, in the form of a plate +attached to the inferior side of the head-shield in front. There +is no reason to doubt that the animal possessed legs; but these +structures seem to have resembled those of many living Crustaceans +in being quite soft and membranous. This, at any rate, seems to +have been generally the case; though structures which have been +regarded as legs have been detected on the under surface of one +of the larger species of Trilobites. There is also, at present, +no direct evidence that the Trilobites possessed the two pairs +of jointed feelers ("antennæ") which are so characteristic +of recent Crustaceans. +</p> + +<p class="indent"> +The Trilobites vary much in size, and the Cambrian formation +presents examples of both the largest and the smallest members +of the order. Some of the young forms may be little bigger than +a millet-seed, and some adult examples of the smaller species +(such as <i>Agnostus</i>) may be only a few lines in length; +whilst such giants of the order as <i>Paradoxides</i> and +<i>Asaphus</i> may reach a length of from one to two feet. Judging +from what we actually know as to the structure of the Trilobites, +and also from analogous recent forms, it would seem that these +ancient Crustaceans were mud-haunting creatures, denizens of +shallow seas, and affecting the soft silt of the bottom rather +than the clear water above. Whenever muddy sediments are found +in the Cambrian and Silurian formations, there we are tolerably +sure to find Trilobites, though they are by no means absolutely +wanting in limestones. They appear to have crawled out upon the +sea-bottom, or burrowed in the yielding mud, with the soft under +surface directed downwards; and it is probable that they really +derived their nutriment from the organic matter contained in the +ooze amongst which they +<a name="page_87"><span class="page">Page 87</span></a> +lived. The vital +organs seem to have occupied the central lobe of the skeleton, +by which they were protected; and a series of delicate leaf-like +paddles, which probably served as respiratory organs, would appear +to have been carried on the under surface of the thorax. That +they had their enemies may be regarded as certain; but we have no +evidence that they were furnished with any offensive weapons, or, +indeed, with any means of defence beyond their hard crust, and +the power, possessed by so many of them, of rolling themselves +into a ball. An additional proof of the fact that they for the +most part crawled along the sea-bottom is found in the occurrence +of tracks and markings of various kinds, which can hardly be +ascribed to any other creatures with any show of probability. +That this is the true nature of some of the markings in question +cannot be doubted at all; and in other cases no explanation so +probable has yet been suggested. If, however, the tracks which +have been described from the Potsdam Sandstone of North America +under the name of <i>Protichnites</i> are really due to the +peregrinations of some Trilobite, they must have been produced +by one of the largest examples of the order. +</p> + +<p class="indent"> +As already said, the Cambrian Rocks are very rich in the remains +of Trilobites. In the lowest beds of the series (Longmynd Rocks), +representatives of some half-dozen genera have now been detected, +including the dwarf <i>Agnostus</i> and the giant <i>Paradoxides</i>. +In the higher beds, the number both of genera and species is +largely increased; and from the great comparative abundance of +individuals, the Trilobites have every right to be considered as +the most characteristic fossils of the Cambrian period,—the +more so as the Cambrian species belong to peculiar types, which, +for the most part, died out before the commencement of the Silurian +epoch. +</p> + +<p class="indent"> +All the remaining Cambrian fossils which demand any notice here +are members of one or other division of the great class of the +<i>Mollusca</i>, or "Shell-fish" properly so called. In the Lower +Cambrian Rocks the Lamp-shells (<i>Brachiopoda</i>) are the principal +or sole representatives of the class, and appear chiefly in three +interesting and important types—namely, <i>Lingulella, +Discina,</i> and <i>Obolella</i>. Of these the last (fig. 32, +<i>i</i>) is highly characteristic of these ancient deposits; +whilst <i>Discina</i> is one of those remarkable persistent types +which, commencing at this early period, has continued to be +represented by varying forms through all the intervening geological +formations up to the present day. <i>Lingulella</i> (fig. 32, +<i>c</i>), again, is closely allied to the existing "Goose-bill" +Lamp-shell (<i>Lingula anatina</i>), and thus presents us with +another example of an extremely long-lived +<a name="page_88"><span class="page">Page 88</span></a> +type. The <i>LingulellĹ“</i> and their successors; the +<i>LingulĹ“</i>, are singular in possessing a shell which +is of a horny texture, and contains but a small proportion of +calcareous matter. In the Upper Cambrian Rocks, the +<i>LingulellĹ“</i> become much more abundant, the broad +satchel-shaped species known as <i>L. Davisii</i> (fig. 32, <i>e</i>) +being so abundant that one of the great divisions of the Cambrian +is termed the "Lingula Flags." Here, also, we meet for the first +time with examples of the genus Orthis (fig. 32, <i>f, k, l</i>) + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 505px; + font-size: smaller; text-align: justify;"> +<img src="images/fig032.jpg" width="497" height="355" alt="Fig. 32" /> +<br /> +Fig. 32.—Cambrian Fossils: <i>a, Protospongia +fenestrata</i>, Menevian Group; <i>b, Arenicolites didymus</i>, +Longmynd Group; <i>c, Lingulella ferruginea</i>, Longmynd and +Menevian, enlarged; <i>d, Hymenocaris vermicauda</i>, Lingula +Flags; <i>e, Lingulella Davisii</i>, Lingula Flags; <i>f, Orthis +lenticularis</i>, Lingula Flags; <i>g, Theca Davidii</i>, Tremadoc +Slates; <i>h, Modiolopsis Solvensis</i>, Tremadoc Slates; <i>i, +Obolela sagittalis</i>, interior of valve, Menevian; <i>j,</i> +Exterior of the same; <i>k, Orthis Hicksii</i>, Menevian; <i>l</i>, +Cast of the same; <i>m, Olenus micrurus</i>, Lingula Flags. (Alter +Salter, Hicks, and Davidson.) +</span> +</span> + +a characteristic Palæozoic type of the Brachiopods, which is +destined to undergo a vast extension in later ages. +</p> + +<p class="indent"> +Of the higher groups of the <i>Mollusca</i> the record is as yet +but scanty. In the Lower Cambrian, we have but the thin, fragile, +dagger-shaped shells of the free-swimming oceanic Molluscs or +"Winged-snails" (<i>Pteropoda</i>), of which the most characteristic +is the genus <i>Theca</i> (fig. 32, <i>g</i>). In the Upper Cambrian, +in addition to these, we have a few Univalves (<i>Gasteropoda</i>), +and, thanks to the researches of Dr Hicks, quite a small assemblage +of Bivalves (<i>Lamellibranchiata</i>), though these are mostly +of no great dimensions (fig. 32, <i>h</i>). Of the chambered +<i>Cephalopoda</i> (Cuttle-fishes and their allies), we have but +<a name="page_89"><span class="page">Page 89</span></a> +few traces; and these wholly confined to the higher beds of the +formation. We meet, however, with examples of the wonderful genus + +<span style="float: right; margin: 4px; width: 128px; + font-size: smaller; text-align: justify;"> +<img src="images/fig033.jpg" width="120" height="240" alt="Fig. 33" /> +<br /> +Fig. 33.—Fragment of <i>Dictyonema sociale</i>, +considerably enlarged, showing the horny branches, with their +connecting cross-bars, and with a row of cells on each side. +(Original.) +</span> + +<i>Orthoceras</i>, with its straight, partitioned shell, which we +shall find in an immense variety of forms in the Silurian rocks. +Lastly, it is worthy of note that the lowest of all the groups of +the <i>Mollusca</i>—namely, that of the Sea-mats, Sea-mosses, +and Lace-corals (<i>Polyzoa</i>)—is only doubtfully known to +have any representatives in the Cambrian, though undergoing a +large and varied development in the Silurian deposits. +</p> + +<p class="indent"> +An exception, however, may with much probability be made to this +statement in favour of the singular genus <i>Dictyonema</i> (fig. +33), which is highly characteristic of the highest Cambrian beds +(Tremadoc Slates). This curious fossil occurs in the form of +fan-like or funnel-shaped expansions, composed of slightly-diverging +horny branches, which are united in a net-like manner by numerous +delicate cross-bars, and exhibit a row of little cups or cells, in +which the animals were contained, on each side. <i>Dictyonema</i> +has generally been referred to the <i>Graptolites</i>; but it +has a much greater affinity with the plant-like Sea-firs +(<i>Sertularians</i>) or the Sea-mosses (<i>Polyzoa</i>), and +the balance of evidence is perhaps in favour of placing it with +the latter. +</p> + +<h4 style="clear: right;">LITERATURE.</h4> + +<p class="indent"> +The following are the more important and accessible works and +memoirs which may be consulted in studying the stratigraphical +and palæontological relations of the Cambrian Rocks:— +</p> + +<table border="0" cellspacing="0"> +<tr><td class="right" valign="top">(1)</td> + <td>'Siluria.' Sir Roderick Murchison. 5th ed., pp. 21-46.</td></tr> +<tr><td class="right" valign="top">(2)</td> + <td>'Synopsis of the Classification of the British Palæozoic +Rocks.' Sedgwick. Introduction to the 3d Fasciculus of the +'Descriptions of British Palæozoic Fossils in the Woodwardian +Museum,' by F. M'Coy, pp. i-xcviii, 1855.</td></tr> +<tr><td class="right" valign="top">(3)</td> + <td>'Catalogue of the Cambrian and Silurian Fossils in the Geological +Museum of the University of Cambridge.' Salter. With a Preface +by Prof. Sedgwick. 1873.</td></tr> +<tr><td class="right" valign="top">(4)</td> + <td>'Thesaurus Siluricus.' Bigsby. 1868.</td></tr> +<tr><td class="right" valign="top">(5)</td> + <td>"History of the Names Cambrian and Silurian." Sterry +Hunt.—'Geological Magazine.' 1873.</td></tr> +<tr><td class="right" valign="top">(6)</td> + <td>'Système Silurien du Centre de la BohĂŞme.' Barrande. + Vol. I.</td></tr> +<tr><td class="right" valign="top">(7)</td> + <td>'Report of Progress of the Geological Survey of Canada, from +its Commencement to 1863,' pp. 87-109.</td></tr> +<tr><td class="right" valign="top">(8) +<a name="page_90"><span class="page">Page 90</span></a></td> + <td>'Acadian Geology.' Dawson. Pp. 641-657.</td></tr> +<tr><td class="right" valign="top">(9)</td> + <td>"Guide to the Geology of New York," Lincklaen; and "Contributions +to the Palæontology of New York," James Hall.—'Fourteenth +Report on the State Cabinet.' 1861.</td></tr> +<tr><td class="right" valign="top">(10)</td> + <td>'Palæozoic Fossils of Canada.' Billings. 1865.</td></tr> +<tr><td class="right" valign="top">(11)</td> + <td>'Manual of Geology.' Dana. Pp. 166-182. 2d ed. 1875.</td></tr> +<tr><td class="right" valign="top">(12)</td> + <td>"Geology of North Wales," Ramsay; with Appendix on the +Fossils, Salter.—'Memoirs of the Geological Survey of Great +Britain,' vol. iii. 1866.</td></tr> +<tr><td class="right" valign="top">(13)</td> + <td>"On the Ancient Rocks of the St David's Promontory, South +Wales, and their Fossil Contents." Harkness and Hicks.—'Quart. +Journ. Geol. Soc.,' xxvii. 384-402. 1871.</td></tr> +<tr><td class="right" valign="top">(14)</td> + <td>"On the Tremadoc Rocks in the Neighbourhood of St David's, +South Wales, and their Fossil Contents." Hicks.—'Quart. +Journ. Geol. Soc.,' xxix. 39-52. 1873.</td></tr> +</table> + +<p class="indent"> +In the above list, allusion has necessarily been omitted to numerous +works and memoirs on the Cambrian deposits of Sweden and Norway, +Central Europe, Russia, Spain, and various parts of North America, +as well as to a number of important papers on the British Cambrian +strata by various well-known observers. Amongst these latter +may be mentioned memoirs by Prof. Phillips, and Messrs Salter, +Hicks, Belt, Plant, Homfray, Ash, Holl, &c. +</p> + +<h3>CHAPTER IX.</h3> + +<p class="subtitle"> +THE LOWER SILURIAN PERIOD. +</p> + +<p class="indent"> +The great system of deposits to which Sir Roderick Murchison +applied the name of "Silurian Rocks" reposes directly upon the +highest Cambrian beds, apparently without any marked unconformity, +though with a considerable change in the nature of the fossils. The +name "Silurian" was originally proposed by the eminent geologist +just alluded to for a great series of strata lying below the Old +Red Sandstone, and occupying districts in Wales and its borders +which were at one time inhabited by the "Silures," a tribe of +ancient Britons. Deposits of a corresponding age are now known +to be largely developed in other parts of England, in Scotland, +and in Ireland, in North America, in Australia, in India, in +Bohemia, Saxony, Bavaria, Russia, Sweden and Norway, Spain, and +in various other regions of less note. In some regions, as in +the neighbourhood of St Petersburg, the Silurian strata are found +not only to have preserved their original horizontality, but +also to have retained almost unaltered their primitive soft and +incoherent nature. In other regions, as in Scandinavia and many +<a name="page_91"><span class="page">Page 91</span></a> +parts of North America, similar strata, now +consolidated into shales, sandstones, and limestones, may be +found resting with a very slight inclination on still older +sediments. In a great many regions, however, the Silurian +deposits are found to have undergone more or less folding, +crumpling, and dislocation, accompanied by induration and +"cleavage" of the finer and softer sediments; whilst in some +regions, as in the Highlands of Scotland, actual "metamorphism" +has taken place. In consequence of the above, Silurian districts +usually present the bold, rugged, and picturesque outlines which +are characteristic of the older "Primitive" rocks of the earth's +crust in general. In many instances, we find Silurian strata +rising into mountain-chains of great grandeur and sublimity, +exhibiting the utmost diversity of which rock-scenery is capable, +and delighting the artist with endless changes of valley, lake, +and cliff. Such districts are little suitable for agriculture, +though this is often compensated for by the valuable mineral +products contained in the rocks. On the other hand, when the +rocks are tolerably soft and uniform in their nature, or when +few disturbances of the crust of the earth have taken place, we +may find Silurian areas to be covered with an abundant pasturage +or to be heavily timbered. +</p> + +<p class="indent"> +Under the head of "Silurian Rocks," Sir Roderick Murchison included +all the strata between the summit of the "Longmynd." beds and +the Old Red Sandstone, and he divided these into the two great +groups of the <i>Lower</i> Silurian and <i>Upper</i> Silurian. It +is, however, now generally admitted that a considerable portion +of the basement beds of Murchison's Silurian series must be +transferred—if only upon palæontological grounds—to +the Upper Cambrian, as has here been done; and much controversy has +been carried on as to the proper nomenclature of the Upper Silurian +and of the remaining portion of Murchison's Lower Silurian. Thus, +some would confine the name "Silurian" exclusively to the Upper +Silurian, and would apply the name of "Cambro-Silurian" to the +Lower Silurian, or would include all beds of the latter age in the +"Cambrian" series of Sedgwick. It is not necessary to enter into +the merits of these conflicting views. For our present purpose, +it is sufficient to recognise that there exist two great groups +of rocks between the highest Cambrian beds, as here defined, and +the base of the Devonian or Old Red Sandstone. These two great +groups are so closely allied to one another, both physically +and palæontologically, that many authorities have established +a third or intermediate group (the "Middle Silurian"), by which a +<a name="page_92"><span class="page">Page 92</span></a> +passage is made from one into the other. This +method of procedure involves disadvantages which appear to outweigh +its advantages; and the two groups in question are not only +generally capable of very distinct stratigraphical separation, +but at the same time exhibit, together with the alliances above +spoken of, so many and such important palæontological +differences, that it is best to consider them separately. We +shall therefore follow this course in the present instance; and +pending the final solution of the controversy as to Cambrian and +Silurian nomenclature, we shall distinguish these two groups of +strata as the "Lower Silurian" and the "Upper Silurian." +</p> + +<p class="indent"> +The <i>Lower Silurian Rocks</i> are known already to be developed +in various regions; and though their <i>general</i> succession +in these areas is approximately the same, each area exhibits +peculiarities of its own, whilst the subdivisions of each are +known by special names. All, therefore, that can be attempted +here, is to select two typical areas—such as Wales and +North America and to briefly consider the grouping and divisions +of the Lower Silurian in each. +</p> + +<p class="indent"> +In Wales, the line between the Cambrian and Lower Silurian is +somewhat ill-defined, and is certainly not marked by any strong +unconformity. There are, however; grounds for accepting the line +proposed, for palæontological reasons, by Dr Hicks, in accordance +with which the Tremadoc Slates ("Lower Tremadoc" of Salter) become +the highest of the Cambrian deposits of Britain. If we take this +view, the Lower Silurian rocks of Wales and adjoining districts +are found to have the following <i>general</i> succession from +below upwards (fig. 34):— +</p> + +<p class="indent"> +1. The <i>Arenig Group</i>.—This group derives its name from +the Arenig mountains, where it is extensively developed. It consists +of about 4000 feet of slates, shales, and flags, and is divisible +into a lower, middle, and upper division, of which the former +is often regarded as Cambrian under the name of "Upper Tremadoc +Slates." +</p> + +<p class="indent"> +2. The <i>Llandeilo Group</i>.—The thickness of this group +varies from about 4000 to as much as 10,000 feet; but in this +latter case a great amount of the thickness is made up of volcanic +ashes and interbedded traps. The sedimentary beds of this group +are principally slates and flags, the latter occasionally with +calcareous bands; and the whole series can be divided into a +lower, middle, and upper Llandeilo division, of which the last +is the most important. The name of "Llandeilo" is derived from +the town of the same name in Wales, where strata of this age +were described by Murchison. +</p> + +<p class="indent"> +<a name="page_93"><span class="page">Page 93</span></a> +3. The <i>Caradoc</i> or <i>Bala Group</i>.—The alternative +names of this group are also of local origin, and are derived, +the one from Caer Caradoc in Shropshire, the other from Bala +in Wales, strata of this age occurring in both localities. The +series is divided into a lower and upper group, the latter chiefly +composed of shales and flags, and the former of sandstones and +shales, together with the important and interesting calcareous +band known as the "Bala Limestone." The thickness of the entire +series varies from 4000 to as much as 12,000 feet, according +as it contains more or less of interstratified igneous rocks. +</p> + +<p class="indent"> +4. The <i>Llandovery Group</i> (Lower Llandovery of +Murchison).—This series, as developed near the town of +Llandovery, in Caermarthenshire, consists of less than 1000 feet +of conglomerates, sandstones, and shales. It is probable, however, +that the little calcareous band known as the "Hirnant Limestone," +together with certain pale-coloured slates which lie above the +Bala Limestone, though usually referred to the Caradoc series, +should in reality be regarded as belonging to the Llandovery group. +</p> + +<p class="indent"> +The general succession of the Lower Silurian strata of Wales +and its borders, attaining a maximum thickness (along with +contemporaneous igneous matter) of nearly 30,000 feet, is +diagramatically represented in the annexed sketch-section (fig. +34):— +</p> + +<p class="indent"> +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 549px; + font-size: smaller; text-align: center;"> +<a name="page_94"><span class="page">Page 94</span></a> +GENERALIZED SECTION OF THE LOWER SILURIAN ROCKS OF WALES. +<br /> +Fig. 34. +<br /> +<img src="images/fig034.jpg" width="541" height="761" alt="Fig. 34" /> +</span> +</span> +In North America, both in the United States and in Canada, the +Silurian rocks are very largely developed, and may be +<a name="page_95"><span class="page">Page 95</span></a> +regarded as constituting an exceedingly full and +typical series of the deposits of this period. The chief groups +of the Silurian rocks of North America are as follows, beginning, +as before, with the lowest strata, and proceeding upwards (fig. +35):— +</p> + +<p class="indent"> +1. <i>Quebec Group</i>.—This group is typically developed in +the vicinity of Quebec, where it consists of about 5000 feet of +strata, chiefly variously-coloured shales, together with some +sandstones and a few calcareous bands. It contains a number of +peculiar Graptolites, by which it can be identified without +question with the Arenig group of Wales and the corresponding +Skiddaw Slates of the North of England. It is also to be noted +that numerous Trilobites of a distinct Cambrian <i>facies</i> +have been obtained in the limestones of the Quebec group, near +Quebec. These fossils, however, have been exclusively obtained +from the limestones of the group; and as these limestones are +principally calcareous breccias or conglomerates, there is room +for believing that these primordial fossils are really derived, +in part at any rate, from fragments of an upper Cambrian +limestone. In the State of New York, the Graptolitic shales of +Quebec are wanting; and the base of the Silurian is constituted +by the so-called "Calciferous Sand-rock" and "Chazy +Limestone."[11] The first of these is essentially and typically +calcareous, and the second is a genuine limestone. +</p> + +<p class="footnote"> +[Footnote 11: The precise relations of the Quebec shales with +Graptolites (Levis Formation) to the Calciferous and Chazy beds +are still obscure, though there seems little doubt but that the +Quebec Shales are superior to the Calciferous Sand-rock.] +</p> + +<p class="indent"> +2. The <i>Trenton Group</i>.—This is an essentially calcareous +group, the various limestones of which it is composed being known +as the "Bird's-eye," "Black River," and "Trenton" Limestones, of +which the last is the thickest and most important. The thickness +of this group is variable, and the bands of limestone in it are +often separated by beds of shale. +</p> + +<p class="indent"> +3. The <i>Cincinnati Group</i> (Hudson River +Formation[12]).—This group consists essentially of a lower +series of shales, often black in colour and highly charged with +bituminous matter (the "Utica Slates "), and of an upper series +of shales, sandstones, +<a name="page_96"><span class="page">Page 96</span></a> +and limestones (the +"Cincinnati" rocks proper). The exact parallelism of the +Trenton and Cincinnati groups with the subdivisions of the Welsh +Silurian series can hardly be stated positively. Probably no +precise equivalency exists; but there can be no doubt but that +the Trenton and Cincinnati groups correspond, as a whole, with +the Llandeilo and Caradoc groups of Britain. The subjoined +diagrammatic section (fig. 35) gives a general idea of the +succession of the Lower Silurian rocks of North America:— + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 575px; + font-size: smaller; text-align: center;"> +GENERALIZED SECTION OF THE LOWER SILURIAN ROCKS OF NORTH AMERICA. +<br /> +Fig. 35. +<br /> +<img src="images/fig035.jpg" width="549" height="661" alt="Fig. 35" /> +</span> +</span> +</p> + +<p class="footnote"> +[Footnote 12: There is some difficulty about the precise nomenclature +of this group. It was originally called the "Hudson River Formation;" +but this name is inappropriate, as rocks of this age hardly touch +anywhere the actual Hudson River itself, the rocks so called +formerly being now known to be of more ancient date. There is +also some want of propriety in the name of "Cincinnati Group," +since the rocks which are known under this name in the vicinity of +Cincinnati itself are the representatives of the Trenton Limestone, +Utica Slates, and the old Hudson River group, inseparably united +in what used to be called the "Blue Limestone Series."]. +</p> + +<p class="indent"> +<a name="page_97"><span class="page">Page 97</span></a> +Of the <i>life</i> of the Lower Silurian period we have record +in a vast number of fossils, showing that the seas of this period +were abundantly furnished with living denizens. We have, however, +in the meanwhile, no knowledge of the land-surfaces of the period. +We have therefore no means of speculating as to the nature of +the terrestrial animals of this ancient age, nor is anything +known with certainty of any land-plants which may have existed. +The only relics of vegetation upon which a positive opinion can +be expressed belong to the obscure group of the "Fucoids," and +are supposed to be the remains of sea-weeds. Some of the fossils +usually placed under this head are probably not of a vegetable + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 412px; + font-size: smaller; text-align: center;"> +<img src="images/fig036.jpg" width="406" height="539" alt="Fig. 36" /> +<br /> +Fig. 36.—<i>Licrophycus Ottawaensis</i> a "Fucoid," from the +Trenton Limestone (Lower Silurian) of Canada. (After Billings.) +</span> +</span> + +nature at all, but others (fig. 36) appear to be unquestionable +plants. The true affinities of these, however, are extremely +dubious. All that can be said is, that remains which appear to +be certainly vegetable, +<a name="page_98"><span class="page">Page 98</span></a> +and which are most +probably due to marine plants, have been recognised nearly at +the base of the Lower Silurian (Arenig), and that they are found +throughout the series whenever suitable conditions recur. +</p> + +<p class="indent"> +The Protozoans appear to have flourished extensively in the Lower +Silurian seas, though to a large extent under forms which are +still little understood. We have here for the first time the +appearance of Foraminifera of the ordinary type—one of the +most interesting observations in this collection being that made +by Ehrenberg, who showed that the Lower Silurian sandstones of +the neighbourhood of St Petersburg contained casts in glauconite +of Foraminiferous shells, some of which are referable to the +existing genera <i>Rotalia</i> and <i>Texularia</i>. True +<i>Sponges</i>, belonging to that section of the group in which +the skeleton is calcareous, are also not unknown, one of the + +<span style="float: left; margin: 4px; width: 263px; + font-size: smaller; text-align: justify;"> +<img src="images/fig037.jpg" width="255" height="253" alt="Fig. 37" /> +<br /> +Fig. 37.—<i>Astylospongia prĹ“morsa</i>, cut +vertically so as to exhibit the canal-system in the interior. +Lower Silurian, Tennessee. (After Ferdinand RĹ“mer.) +</span> + +most characteristic genera being <i>Astylospongia</i> (fig. 37). +In this genus are included more or less globular, often lobed +sponges, which are believed not to have been attached to foreign +bodies. In the form here figured there is a funnel-shaped cavity +at the summit; and the entire mass of the sponge is perforated, +as in living examples, by a system of canals which convey the +sea-water to all parts of the organism. The canals by which the +sea-water gains entrance open on the exterior of the sphere, +and those by which it again escapes from the sponge open into +the cup-shaped depression at the summit. +</p> + +<p class="indent"> +The most abundant, and at the same time the least understood, +of Lower Silurian Protozoans belong, however, to the genera +<i>Stromatopora</i> and <i>Receptaculites</i>, the structure +of which can merely be alluded to here. The specimens of +<i>Stromatopora</i> (fig. 38) occur as hemispherical, pear-shaped, +globular, or irregular masses, often of very considerable size, +and sometimes demonstrably attached to foreign bodies. In their +structure these masses consist of numerous thin calcareous +laminæ, usually arranged concentrically, and separated by +narrow interspaces. These interspaces are generally crossed by +numerous vertical calcareous pillars, giving the +<a name="page_99"><span class="page">Page 99</span></a> +vertical section of the fossil a +lattice-like appearance. There are also usually minute pores in the +concentric laminæ, by which the successive interspaces are + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 454px; + font-size: smaller; text-align: center;"> +<img src="images/fig038.jpg" width="446" height="408" alt="Fig. 38" /> +<br /> +Fig. 38.—A small and perfect specimen of <i>Stromatopora +rugosa</i>, of the natural size, from the Trenton Limestone of +Canada. (After Billings.) +</span> +</span> + +placed in communication; and sometimes the surface presents large +rounded openings, which appear to correspond with the water-canals +of the Sponges. Upon the whole, though presenting some curious +affinities to the calcareous Sponges, <i>Stromatopora</i> is +perhaps more properly regarded as a gigantic <i>Foraminifer</i>. If +this view be correct, it is of special interest as being probably the +nearest ally of <i>Eozoön</i>, the general appearance of the two +being strikingly similar, though their minute structure is not at +all the same. Lastly, in the fossils known as <i>Receptaculites</i> +and <i>Ischadites</i> we are also presented with certain singular +Lower Silurian Protozoans, which may with great probability be +regarded as gigantic <i>Foraminifera</i>. Their structure is very +complex; but fragments are easily recognised by the fact that the +exterior is covered with numerous rhomboidal calcareous plates, +closely fitting together, and arranged in peculiar intersecting +curves, presenting very much the appearance of the engine-turned +case of a watch. +</p> + +<p class="indent"> +Passing next to the sub-kingdom of <i>CĹ“lenterate</i> animals +(Zoophytes, Corals, &c.), we find that this great group, almost +or wholly absent in the Cambrian, is represented in Lower +<a name="page_100"><span class="page">Page 100</span></a> +Silurian deposits by a great number of forms belonging +on the one hand to the true Corals, and en the other hand to the +singular family of the <i>Graptolites</i>. If we except certain +plant-like fossils which probably belong rather to the Sertularians +or the Polyzoans (e.g., <i>Dictyonema, Dendrograptus</i>, &c.), +the family of the <i>Graptolites</i> may be regarded as exclusively +Silurian in its distribution. Not only is this the case, but it +attained its maximum development almost upon its first appearance, +in the Arenig Rocks; and whilst represented by a great variety of +types in the Lower Silurian; it only exists in the Upper Silurian +in a much diminished form. The <i>Graptolites</i> (Gr. <i>grapho</i>, +I write; <i>lithos</i>, stone) were so named by Linnæus, from +the resemblance of some of them to written or pencilled marks upon +the stone, though the great naturalist himself did not believe +them to be true fossils at all. They occur as linear or leaf-like +bodies, sometimes simple, sometimes compound and branched; and no +doubt whatever can be entertained as to their being the skeletons +of composite organisms, or colonies of semi-independent animals +united together by a common fleshy trunk, similar to what is +observed in the colonies of the existing Sea-firs (Sertularians). +This fleshy trunk or common stem of the colony was protected +by a delicate horny sheath, and it gave origin to the little +flower-like "polypites," which constituted the active element of +the whole assemblage. These semi-independent beings were, in turn, +protected each by a little horny cup or cell, directly connected +with the common sheath below, and terminating above in an opening +through which the polypite could protrude its tentacled head or +could again withdraw itself for safety. The entire skeleton, +again, was usually, if not universally, supported by a delicate +horny rod or "axis," which appears to have been hollow, and which +often protrudes to a greater or less extent beyond one or both +of the extremities of the actual colony. +</p> + +<p class="indent"> +The above gives the elementary constitution of any <i>Graptolite</i>, +but there are considerable differences as to the manner in which +these elements are arranged and combined. In some forms the common +stem of the colony gives origin to but a single row of cells +on one side. If the common stem is a simple, straight, or +slightly-curved linear body, then we have the simplest form of +Graptolite known (the genus <i>Monograptus</i>); and it is worthy +of note that these simple types do not come into existence till +comparatively late (Llandeilo), and last nearly to the very close +of the Upper Silurian. In other cases, whilst there is still but +a single row of cells, the colony may consist of two of these +simple stems springing from a +<a name="page_101"><span class="page">Page 101</span></a> +common point, as in the so-called +"twin Graptolites" (<i>Didymograptus</i>, fig. 40). This type is + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 551px; + font-size: smaller; text-align: center;"> +<img src="images/fig039.jpg" width="543" height="610" alt="Fig. 39" /> +<br /> +Fig. 39.—<i>Dichograptus octobrachiatus</i>, a branched, +"unicellular" Graptolite from the Skiddaw and Quebec Groups +(Arenig). (After Hall.) +</span> +</span> + +entirely confined to the earlier portion of the Lower Silurian period +(Arenig and Llandeilo). In other cases, again, there may be four of +such stems springing from a central point (<i>Tetragraptus</i>). +Lastly, there are numerous complex forms (such as <i>Dichograptus, +Loganograptus</i>, &c.) in which there are eight or more of these +simple branches, all arising from a common centre (fig. 39), +which is sometimes furnished with a singular horny disc. These +complicated branching forms, as well as the <i>Tetragrapti</i>, +are characteristic of the horizon of the Arenig group. Similar +forms, often specifically identical, are found at this horizon +in Wales, in the great series of the Skiddaw Slates of the north +of England, in the Quebec group in Canada, in equivalent beds in +Sweden, and in certain gold-bearing slates of the same age in +Victoria in Australia. +</p> + +<p class="indent"> +In another great group of Graptolites (including the genera +<i>Diplograptus, Dicranograptus, Climacograptus</i>, &c.) the +common stem of the colony gives origin, over part or the whole or +its length, to <i>two</i> rows of cells, one on each side (fig. 41). +These "double-celled" Graptolites are highly characteristic of the +Lower Silurian deposits; and, with an exception more apparent than +<a name="page_102"><span class="page">Page 102</span></a> +real in Bohemia, they are exclusively confined to strata of Lower +Silurian age, and are not known to occur in the Upper Silurian. + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 582px; + font-size: smaller; text-align: center;"> +<img src="images/fig040.jpg" width="574" height="217" alt="Fig. 40" /> +<br /> +Fig. 40.—Central portion of the colony of <i>Didymegraptus +divaricatus</i>, Upper Llandeilo, Dumfresshire. (Original.) +</span> +</span> + +Lastly, there is a group of Graptolites (<i>Phyllograptus</i>, fig. +42) in which the colony is leaf-like in form, and is composed + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 235px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig041.jpg" width="227" height="460" alt="Fig. 41" /> +<br /> +Fig. 41.—Examples of <i>Diplograptus pristis</i>, +showing variations in the appendages at the base. Upper Llandeilo, +Dumfriesshire. (Original.) +</span> + +<span style="width: 276px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig042.jpg" width="268" height="383" alt="Fig. 42" /> +<br /> +Fig. 42.—Group of individuals of <i>Phyllograptus +typus</i>, from the Quebec group of Canada. (After Hall.) One +of the four rows of cells is hidden on the under surface. +</span> + +</span> + +of <i>four</i> rows of cells springing in a cross-like +<a name="page_103"><span class="page">Page 103</span></a> +manner from the common stem. These forms are highly +characteristic of the Arenig group. +</p> + +<p class="indent"> +The Graptolites are usually found in dark-coloured, often black +shales, which sometimes contain so much carbon as to become +"anthracitic." They may be simply carbonaceous; but they are +more commonly converted into iron-pyrites, when they glitter +with the brilliant lustre of silver as they lie scattered on the +surface of the rock, fully deserving in their metallic tracery +the name of "written stones." They constitute one of the most +important groups of Silurian fossils, and are of the greatest +value in determining the precise stratigraphical position of +the beds in which they occur. They present, however, special +difficulties in their study; and it is still a moot point as +to their precise position in the zoological scale. The balance +of evidence is in favour of regarding them as an ancient and +peculiar group of the Sea-firs (Hydroid Zoophytes), but some +regard them as belonging rather to the Sea-mosses (<i>Polyzoa</i>). +Under any circumstances, they cannot be directly compared either +with the ordinary Sea-firs or the ordinary Sea-mosses; for these +two groups consist of fixed organisms, whereas the Graptolites +were certainly free-floating creatures, living at large in the +open sea. The only Hydroid Zoophytes or Polyzoans which have +a similar free mode of existence, have either no skeleton at +all, or have hard structures quite unlike the horny sheaths of +the Graptolites. +</p> + +<p class="indent"> +The second great group of CĹ“lenterate animals +(<i>Actinozoa</i>) is represented in the Lower Silurian rocks +by numerous Corals. These, for obvious reasons, are much more +abundant in regions where the Lower Silurian series is largely +calcareous (as in North America) than in districts like Wales, +where limestones are very feebly developed. The Lower Silurian +Corals, though the first of their class, and presenting certain +peculiarities, may be regarded as essentially similar in nature +to existing Corals. These, as is well known, are the calcareous +skeletons of animals—the so-called +"Coral-Zoophytes"—closely allied to the common Sea-anemones +in structure and habit. A <i>simple</i> coral (fig. 43) consists +of a calcareous cup embedded in the soft tissues of the +flower-like polype, and having at its summit a more or less deep +depression (the "calice") in which the digestive organs are +contained. The space within the coral is divided into compartments +by numerous vertical calcareous plates (the "septa"), which spring +from the inside of the wall of the cup, and of which some generally +reach the centre. <i>Compound</i> corals, again (fig. 44), consist +of a greater or less number of structures similar in structure +to the above, +<a name="page_104"><span class="page">Page 104</span></a> +but united together in different +ways into a common mass. <i>Simple</i> corals, therefore, are the + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 294px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig043.jpg" width="286" height="332" alt="Fig. 43" /> +<br /> +Fig. 43.—<i>Zaphrentis Stokesi</i>, a simple "cup-coral," +Upper Silurian, Canada. (After Billings.) +</span> + +<span style="width: 265px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig044.jpg" width="257" height="202" alt="Fig. 44" /> +<br /> +Fig. 44.—Upper surface of a mass of <i>Strombodes +pentagonus</i>. Upper Silurian, Canada. (After Billings.) +</span> + +</span> + +skeletons of <i>single</i> and independent polypes; whilst +<i>compound</i> corals are the skeletons of assemblages or +colonies of similar polypes, living united with one another +another as an organic community. +</p> + +<p class="indent"> +In the general details of their structure, the Lower Silurian +Corals do not differ from the ordinary Corals of the present +day. The latter, however, have the vertical calcareous plates of +the coral ("septa") arranged in multiples of six or five; whereas +the former have these structures arranged in multiples of four, +and often showing a cross-like disposition. For this reason, the +common Lower Silurian Corals are separated to form a distinct +group under the name of <i>Rugose</i> Corals or <i>Rugosa</i>. +They are further distinguished by the fact that the cavity of +the coral ("visceral chamber") is usually subdivided by more +or less numerous horizontal calcareous plates or partitions, +which divide the coral into so many tiers or storeys, and which +are known as the "tabulæ" (fig. 45). +</p> + +<p class="indent"> +In addition to the Rugose Corals, the Lower Silurian rocks contain +a number of curious compound corals, the tubes of which have either +no septa at all or merely rudimentary ones, but which have the +transverse partitions or "tabulæ" very highly developed. These +are known as the <i>Tabulate Corals</i>; and recent researches on +some of their existing allies (such as <i>Heliopora</i>) have shown +that they are really allied to +<a name="page_105"><span class="page">Page 105</span></a> +the modern Sea-pens, Organ-pipe +Corals, and Red Coral, rather than to the typical stony Corals. +Amongst the characteristic Rugose Corals of the Lower Silurian + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 381px; + font-size: smaller; text-align: justify;"> +<img src="images/fig045.jpg" width="373" height="260" alt="Fig. 45" /> +<br /> +Fig. 45.—<i>Columnaria alveolata</i>, a Rugose compound +coral, with imperfect septa, but having the corallites partitioned +off into storeys by "tabulæ." Lower Silurian, Canada. (After +Billings.) +</span> +</span> + +may be mentioned species belonging to the genera <i>Columnaria, +Favistella, Streptelasma</i>, and <i>Zaphrentis</i>; whilst amongst +the "Tabulate" Corals, the principal forms belong to the genera +<i>ChĹ“tetes, Halysites</i> (the Chain-coral), <i>Constellaria</i>, +and <i>Heliolites</i>. These groups of the Corals, however, attain +a greater development at a later period, and they will be noticed +more particularly hereafter. +</p> + +<p class="footnote"> +[Footnote 13: The genus <i>Caryocrinus</i> is sometimes regarded +as properly belonging to the <i>Crinoids</i>, but there seem to +be good reasons for rather considering it as an abnormal form +of <i>Cystidean</i>.] +</p> + +<p class="indent"> +Passing onto higher animals, we find that the class of the +<i>Echinodermata</i> is represented by examples of the Star-fishes +(<i>Asteroidea</i>), the Sea-lilies (<i>Crinoidea</i>), and the +peculiar extinct group of the Cystideans (<i>Cystoidea</i>), with +one or two of the Brittle-stars (<i>Ophiuroidea</i>)—the +Sea-urchins (<i>Echinoidea</i>) being still wanting. The Crinoids, +though in some places extremely numerous, have not the varied +development that they possess in the Upper Silurian, in connection +with which their structure will be more fully spoken of. In the +meanwhile, it is sufficient to note that many of the calcareous +deposits of the Lower Silurian are strictly entitled to the name +of "Crinoidal limestones," being composed in great part of the +detached joints, and plates, and broken stems, of these beautiful +but fragile organisms (see fig. 12). Allied to the Crinoids are +the singular creatures which are known as <i>Cystideans</i> (fig. +46). These are generally composed of a globular or ovate body +(the "calyx"), supported upon a short stalk (the "column"), by +which the organism was usually attached to some foreign body. The +body was enclosed by closely-fitting calcareous plates, accurately +<a name="page_106"><span class="page">Page 106</span></a> +jointed together; and the stem was made up of numerous distinct +pieces or joints, flexibly united to each other by membrane. The + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 503px; + font-size: smaller; text-align: justify;"> +<img src="images/fig046.jpg" width="495" height="419" alt="Fig. 46" /> +<br /> +Fig. 46.—Group of Cystideans. A, <i>Caryocrinus +ornatus</i>,[13] Upper Silurian, America; B, <i>Pleurocystites +squamosus</i>, showing two short "arms," Lower Silurian, Canada; +C, <i>Pseudocrinus bifasciatus</i>, Upper Silurian, England; D, +<i>Lepadocrinus Gebhartii</i>, Upper Silurian, America. (After +Hall, Billings, and Salter.) +</span> +</span> + +chief distinction which strikes one in comparing the Cystideans +with the Crinoids is, that the latter are always furnished, as +will be subsequently seen, with a beautiful crown of branched +and feathery appendages, springing from the summit of the calyx, +and which are composed of innumerable calcareous plates or joints, +and are known as the "arms." In the Cystideans, on the other hand, +there are either no "arms" at all, or merely short, unbranched, +rudimentary arms. The Cystideans are principally, and indeed +nearly exclusively, Silurian fossils; and though occurring in +the Upper Silurian in no small numbers, they are pre-eminently +characteristic of the Llandeilo-Caradoc period of Lower Silurian +time. They commenced their existence, so far as known, in the +Upper Cambrian; and though examples are not absolutely unknown +<a name="page_107"><span class="page">Page 107</span></a> +in later periods, they are pre-eminently characteristic of the +earlier portion of the Palæozoic +epoch. +</p> + +<p class="indent"> +The Ringed Worms (<i>Annelides</i>) are abundantly represented +in the Lower Silurian, but principally by tracks and burrows +similar in essential respects to those which occur so commonly +in the Cambrian formation, and calling for no special comment. +Much more important are the <i>Articulate</i> animals, represented +as heretofore, wholly by the remains of the aquatic group of the + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 575px; + font-size: smaller; text-align: justify;"> +<img src="images/fig047.jpg" width="551" height="585" alt="Fig. 47" /> +<br /> +Fig. 47.—Lower Silurian Crustaceans. <i>a, Asaphus +tyrannus</i>, Upper Llandeilo; <i>b. Ogygia Buchii</i>, Upper +Llandeilo; <i>c, Trinucleus concentricus</i>, Caradoc; <i>d, +Caryocaris Wrightii</i>, Arenig (Skiddaw Slates); <i>e, Beyrichia +complicata</i>, natural size and enlarged, Upper Llandeilo and +Caradoc; <i>f, Primitia strangulata</i>, Caradoc: <i>g.</i> +Head-shield of <i>Calymene Blumenbachii</i>, var. +<i>brevicapitata</i>, Caradoc; <i>h</i>, Head-shield of <i>Triarthrus +Becki</i> (Utica Slates), United States: <i>i</i>, Shield of +<i>Leperditia Canadensis</i>, var. <i>Josephiana</i>, of the +natural size, Trenton Limestone, Canada; <i>j</i>, The same, +viewed from the front. (After Salter, M'Coy, Rupert Jones, and +Dana.) +</span> +</span> + +<i>Crustaceans</i>. Amongst these are numerous little bivalved +forms—such as species of <i>Primitia</i> (fig. 47, <i>f</i>), +<a name="page_108"><span class="page">Page 108</span></a> +<i>Beyrichia</i> (fig. 47, <i>e</i>), and +<i>Leperditia</i> (fig. 47, <i>i</i> and <i>j</i>). Most of these +are very small, varying from the size of a pin's head up to that +of a hemp seed; but they are sometimes as large as a small bean +(fig. 47, <i>i</i>), and they are commonly found in myriads +together in the rock. As before said, they belong to the same +great group as the living Water-fleas (<i>Ostracoda</i>). Besides +these, we find the pod-shaped head-shields of the shrimp-like +Phyllopods—such as <i>Caryocaris</i> (fig. 47, <i>d</i>) +and <i>Ceratiocaris</i>. More important, however, than any of +these are the <i>Trilobites</i>, which may be considered as +attaining their maximum development in the Lower Silurian. The +huge <i>Paradoxides</i> of the Cambrian have now disappeared, +and with them almost all the principal and characteristic +"primordial" genera, save <i>Olenus</i> and <i>Agnostus</i>. In +their place we have a great number of new forms—some of them, +like the great <i>Asaphus tyrannus</i> of the Upper Llandeilo +(fig. 47, <i>a</i>), attaining a length of a foot or more, and +thus hardly yielding in the matter of size to their ancient rivals. +Almost every subdivision of the Lower Silurian series has its own +special and characteristic species of Trilobites; and the study +of these is therefore of great importance to the geologist. A +few widely-dispersed and characteristic species have been here +figured (fig. 47); and the following may be considered as the +principal Lower Silurian genera—<i>Asaphus, Ogygia, Cheirurus, +Ampyx, Caiymene, Trinucleus, Lichas, IllĹ“nus, Æglina, +Harpes, Remopleurides, Phacops, Acidaspis</i>, and +<i>Homalonotus</i>, a few of them passing upwards under new forms +into the Upper Silurian. +</p> + +<p class="indent"> +Coming next to the <i>Mollusca</i>, we find the group of the +Sea-mosses and Sea-mats (<i>Polyzoa</i>) represented now by quite a +number of forms. Amongst these are examples of the true Lace-corals +(<i>Retepora</i> and <i>Fenestella</i>), with their netted fan-like +or funnel-shaped fronds; and along with these are numerous delicate +encrusting forms, which grew parasitically attached to shells +and corals (<i>Hippothoa, Alecto</i>, &c.); but perhaps the most +characteristic forms belong to the genus <i>Ptilodictya</i> (figs. +48 and 49). In this group the frond is flattened, with thin striated +edges, sometimes sword-like or scimitar-shaped, but often more or +less branched; and it consists of two layers of cells, separated +by a delicate membrane, and opening upon opposite sides. Each of +these little chambers or "cells" was originally tenanted by a +minute animal, and the whole thus constituted a compound organism +or colony. +</p> + +<p class="indent"> +The Lamp-shells or <i>Brachiopods</i> are so numerous, and present +such varied types, both in this and the succeeding period of +the Upper Silurian, that the name of "Age of Brachiopods" +<a name="page_109"><span class="page">Page 109</span></a> +has with justice been applied to the Silurian period as a whole. +It would be impossible here to enter into details as to the many + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 268px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig048.jpg" width="230" height="232" alt="Fig. 48" /> +<br /> +Fig. 48.—<i>Ptilodictya falciformis</i>. <i>a</i>, +Small specimen of the natural size; <i>b</i>, Cross-section, +showing the shape of the frond; <i>c</i>, Portion of the surface, +enlarged. Trenton Limestone and Cincinnati Group, America. +(Original.) +</span> + +<span style="width: 271px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig049.jpg" width="276" height="227" alt="Fig. 49" /> +<br /> +Fig. 49.—A, <i>Ptilodictya acuta</i>; B. <i>Ptilodictya +Schafferi</i>. <i>a</i>, Fragment, of the natural size; <i>b</i>, +Portion, enlarged to show the cells. Cincinnati Group of Ohio +and Canada. (Original.) +</span> + +</span> + +different forms of Brachiopods which present themselves in the +Lower Silurian deposits; but we may select the three genera +<i>Orthis, Strophomena</i>, and <i>LeptĹ“na</i> for +illustration, as being specially characteristic of this period, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 603px; + font-size: smaller; text-align: justify;"> +<img src="images/fig050.jpg" width="595" height="337" alt="Fig. 50" /> +<br /> +Fig. 50.—Lower Silurian Brachiopods. <i>a</i> +and <i>a'</i>, <i>Orthis biforata</i>, Llandeilo-Caradoc, Britain +and America: <i>b, Orthis flabellulum</i>, Caradoc, Britain: <i>c, +Orthis subquadrata</i>, Cincinnati Group, America; <i>c'</i>, +Interior of the dorsal valve of the same: <i>d, Strophomena +deltoidea</i>, Llandeilo-Caradoc, Britain and America. (After +Meek, Hall, and Salter.) +</span> +</span> + +though not exclusively confined to it. The numerous shells which +belong to the extensive and cosmopolitan genus <i>Orthis</i> (fig. +50, <i>a, b, c,</i> +<a name="page_110"><span class="page">Page 110</span></a> +and fig. 51, <i>c</i> and <i>d</i>), are +usually more or less transversely-oblong or subquadrate, the two +valves (as more or less in all the Brachiopods) of unequal sizes, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 625px; + font-size: smaller; text-align: justify;"> +<img src="images/fig051.jpg" width="617" height="283" alt="Fig. 51" /> +<br /> +Fig. 51.—Lower Silurian Brachiopods, <i>a, +Strophomena alternata</i>, Cincinnati Group, America; <i>b, +Strophomena filitexta</i>, Trenton and Cincinnati Groups, America; +<i>c, Orthis testudinaria</i>, Caradoc, Europe, and America; +<i>d, d', Orthis plicateila</i>, Cincinnati Group, America; <i>e, +e', e'', LeptĹ“na sericea</i>, Llandeilo and Caradoc, Europe +and America. (After Meek, Hall, and the Author.) +</span> +</span> + +generally more or less convex, and marked with radiating ribs or +lines. The valves of the shell are united to one another by teeth +and sockets, and there is a straight hinge-line. The beaks are +also separated by a distinct space ("hinge-area"), formed in part +by each valve, which is perforated by a triangular opening, through +which, in the living condition, passed a muscular cord attaching +the shell to some foreign object. The genus <i>Strophomena</i> +(fig. 50, <i>d</i>, and 51, <i>a</i> and <i>b</i>) is very like +<i>Orthis</i> in general character; but the shell is usually much +flatter, one or other valve often being concave, the hinge-line is +longer, and the aperture for the emission of the stalk of attachment +is partially closed by a calcareous plate. In <i>LeptĹ“na</i>, +again (fig. 51, <i>e</i>), the shell is like <i>Strophomena</i> +in many respects, but generally comparatively longer, often +completely semicircular, and having one valve convex and the +other valve concave. Amongst other genera of Brachiopods which are +largely represented in the Lower Silurian rocks may be mentioned +<i>Lingula, Crania, Discina, Trematis, Siphonotreta, Acrotreta, +Rhynchonella</i>, and <i>Athyris</i>; but none of these can claim +the importance to which the three previously-mentioned groups +are entitled. +</p> + +<p class="indent"> +The remaining Lower Silurian groups of <i>Mollusca</i> can be but +briefly glanced at here. The Bivalves (<i>Lamellibranchiata</i>) +find numerous representatives, belonging to such genera as +<a name="page_111"><span class="page">Page 111</span></a> +<i>Modiolopsis, Ctenodonta, Orthonota, PalĹ“arca, Lyrodesma, + +<span style="float: right; margin: 4px; width: 115px; + font-size: smaller; text-align: justify;"> +<img src="images/fig052.jpg" width="107" height="273" alt="Fig. 52" /> +<br /> +Fig. 52.—<i>Murchisonia gracilis</i>, Trenton Limestone, +America. (After Billings.) +</span> + +Ambonychia</i>,and <i>Cleidophorus</i>. The Univalves +(<i>Gasteropoda</i>) are also very numerous, the two most important +genera being <i>Murchisonia</i> (fig. 52) and <i>Pleurotomaria</i>. +In both these groups the outer lip of the shell is notched; but +the shell in the former is elongated and turreted, whilst in +the latter it is depressed. The curious oceanic Univalves known +as the <i>Heteropods</i> are also very abundant, the principal +forms belonging to <i>Bellerophon</i> and <i>Maclurea</i>. In the +former (fig. 53) there is a symmetrical convoluted shell, like +that of the Pearly Nautilus in shape, but without any internal +partitions, and having the aperture often expanded and notched +behind. The species of <i>Maclurea</i> (fig. 54) are found both +in North America and in Scotland, and are exclusively confined +to the Lower Silurian period, so far as known. They have the +shell coiled into a flat spiral, the mouth being furnished with +a very curious, thick, and solid lid or "operculum." The Lower +Silurian <i>Pteropods</i>, or "Winged snails," are numerous, +and belong principally to the genera <i>Theca, Conularia</i>, +and <i>Tentaculites</i>, the last-mentioned of these often being +extremely abundant in certain strata. +</p> + +<p class="indent"> +Lastly, the Lower Silurian Rocks have yielded a vast number of +chambered shells, referable to animals which belong to the same +great division as the Cuttle-fishes (the <i>Cephalopoda</i>), and +of which the Pearly Nautilus is the only living representative +at the present day. In this group of <i>Cephalopods</i> the animal + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 368px; + font-size: smaller; text-align: center;"> +<img src="images/fig053.jpg" width="360" height="184" alt="Fig. 53" /> +<br /> +Fig. 53.—Different views of <i>Bellerophon Argo</i>, +Trenton Limestone, Canada. (After Billings.) +</span> +</span> + +possesses a well-developed external shell, which is divided into +chambers by shelly partitions ("septa"). The animal lives in +the last-formed and largest chamber of the shell, to which +<a name="page_112"><span class="page">Page 112</span></a> +it is organically connected by muscular attachments. The head is +furnished with long muscular processes or "arms," and can be + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 556px; + font-size: smaller; text-align: center;"> +<img src="images/fig054.jpg" width="548" height="196" alt="Fig. 54" /> +<br /> +Fig. 54.—Different views of <i>Maclurea crenulata</i>, +Quebec Group, Newfoundland. (After Billings.) +</span> +</span> + +protruded from the mouth of the shell at will, or again withdrawn +within it. We learn, also, from the Pearly Nautilus, that these +animals must have possessed two pairs of breathing organs or +"gills;" hence all these forms are grouped together under the +name of the "Tetrabranchiate" Cephalopods (Gr. <i>tetra</i>, +four; <i>bragchia</i>, gill). On the other hand, the ordinary +Cuttle-fishes and Calamaries either possess an internal skeleton, +or if they have an external shell, it is not chambered; their +"arms" are furnished with powerful organs of adhesion in the form +of suckers; and they possess only a single pair of gills. For +this last reason they are termed the "Dibranchiate" Cephalopods +(Gr. <i>dis</i>, twice; <i>bragchia</i>, gill). No trace of the +true Cuttle-fishes has yet been found in Lower Silurian deposits; +but the Tetrabranchiate group is represented by a great number +of forms, sometimes of great size. The principal Lower Silurian +genus is the well-known and widely-distributed <i>Orthoceras</i> +(fig. 55). The shell in this genus agrees with that of the existing +<i>Pearly Nautilus</i>, in consisting of numerous chambers separated +by shelly partitions (or septa), the latter being perforated by +a tube which runs the whole length of the shell after the last +chamber, and is known as the "siphuncle" (fig. 56, <i>s</i>). +The last chamber formed is the largest, and in it the animal +lives. The chambers behind this are apparently filled with some +gas secreted by the animal itself; and these are supposed to +act as a kind of float, enabling the creature to move with ease +under the weight of its shell. The various air-chambers, though +the siphuncle passes through them, have no direct connection +with one another; and it is believed that the animal has the +power of slightly altering its specific gravity, and thus of +rising or sinking in the water by driving additional fluid into +the siphuncle or partially emptying it. The <i>Orthoceras</i> +<a name="page_113"><span class="page">Page 113</span></a> +further agrees with the Pearly Nautilus in the fact that +the partitions or septa separating the different air-chambers are + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 268px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig055.jpg" width="227" height="448" alt="Fig. 55" /> +<br /> +Fig. 55.—Fragment of <i>Orthoceras crebriseptum</i>, +Cincinnati Group, North America, of the natural size. The lower +figure section showing the air-chambers, and the form and position +of the siphuncle. (After Billings.) +</span> + +<span style="width: 271px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig056.jpg" width="235" height="475" alt="Fig. 56" /> +<br /> +Fig. 56.—[14] Restoration of Orthoceras, the shell +being supposed to be divided vertically, and only its upper part +being shown. <i>a</i>, Arms; <i>f</i>, Muscular tube ("funnel") +by which water is expelled from the mantle-chamber; <i>c</i>, +Air-chambers; <i>s</i>, Siphuncle. +</span> + +</span> + +simple and smooth, concave in front and convex behind, and devoid +of the elaborate lobation which they exhibit in the Ammonites; +whilst the siphuncle pierces the septa either in the centre or +near it. In the Nautilus, however, the shell is coiled into a +flat spiral; whereas in <i>Orthoceras</i> the shell is a straight, +longer or shorter cone, tapering behind, and gradually expanding +towards its mouth in front. The chief objections to the belief +that the animal of the <i>Orthoceras</i> was essentially like +that of the Pearly Nautilus are—the comparatively small size +of the body-chamber, the often contracted aperture of the mouth, +and the enormous size of some specimens of +<a name="page_114"><span class="page">Page 114</span></a> +the shell. Thus, some <i>Orthocerata</i> have been discovered +measuring ten or twelve feet in length, with a diameter of a foot +at the larger extremity. These colossal dimensions certainly make +it difficult to imagine that the comparatively small body-chamber +could have held an animal large enough to move a load so ponderous +as its own shell. To some, this difficulty has appeared so great +that they prefer to believe that the <i>Orthoceras</i> did not +live in its shell at all, but that its shell was an internal +skeleton similar to what we shall find to exist in many of the +true Cuttle-fishes. There is something to be said in favour of +this view, but it would compel us to believe in the existence in +Lower Silurian times of Cuttle-fishes fully equal in size to the +giant "Kraken" of fable. It need only be added in this connection +that the Lower Silurian rocks have yielded the remains of many +other Tetrabranchiate Cephalopods besides <i>Orthoceras</i>. Some +of these belong to <i>Cyrtoceras</i>, which only differs from +<i>Orthoceras</i> in the bow-shaped form of the shell; others +belong to <i>Phragmoceras</i>, <i>Lituites</i>, &c.; and, +lastly; we have true <i>Nautili</i>, with their spiral shells, +closely resembling the existing Pearly Nautilus. +</p> + +<p class="footnote"> +[Footnote 14: This illustration is taken from a rough sketch +made by the author many years ago, but he is unable to say from +what original source it was copied.] +</p> + +<p class="indent"> +Whilst all the sub-kingdoms of the Invertebrate animals are +represented in the Lower Silurian rocks, no traces of Vertebrate +animals have ever been discovered in these ancient deposits, +unless the so-called "Conodonts" found by Pander in vast numbers +in strata of this age [15] in Russia should prove to be really +of this nature. These problematical bodies are of microscopic +size, and have the form of minute, conical, tooth-shaped spines, +with sharp edges, and hollow at the base. Their original discoverer +regarded them as the horny teeth of fishes allied to the Lampreys; +but Owen came to the conclusion that they probably belonged to +Invertebrates. The recent investigation of a vast number of similar +but slightly larger bodies, of very various forms, in the +Carboniferous rocks of Ohio, has led Professor Newberry to the +conclusion that these singular fossils really are, as Pander +thought, the teeth of Cyclostomatous fishes. The whole of this +difficult question has thus been reopened, and we may yet have +to record the first advent of Vertebrate animals in the Lower +Silurian. +</p> + +<p class="footnote"> +[Footnote 15: According to Pander, the "Conodonts" are found not +only in the Lower Silurian beds, but also in the "Ungulite Grit" +(Upper Cambrian), as well as in the Devonian and Carboniferous +deposits of Russia. Should the Conodonts prove to be truly the +remains of fishes, we should thus have to transfer the first +appearance of vertebrates to, at any rate, as early a period as +the Upper Cambrian.] +</p> + +<h3> +<a name="page_115"><span class="page">Page 115</span></a> +CHAPTER X.</h3> + +<p class="subtitle"> +THE UPPER SILURIAN PERIOD. +</p> + +<p class="indent"> +Having now treated of the Lower Silurian period at considerable +length, it will not be necessary to discuss the succeeding group of +the <i>Upper Silurian</i> in the same detail—the more so, as +with a general change of <i>species</i> the Upper Silurian animals +belong for the most part to the same great types as those which +distinguish the Lower Silurian. As compared, also, as regards +the total bulk of strata concerned, the thickness of the Upper +Silurian is generally very much below that of the Lower Silurian, +indicating that they represent a proportionately shorter period of +time. In considering the general succession of the Upper Silurian +beds, we shall, as before, select Wales and America as being two +regions where these deposits are typically developed. +</p> + +<p class="indent"> +In Wales and its borders the general succession of the Upper +Silurian rocks may be taken to be as follows, in ascending order +(fig. 57):— +</p> + +<p class="indent"> +(1) The base of the Upper Silurian series is constituted by a +series of arenaceous beds, to which the name of "May Hill Sandstone" +was applied by Sedgwick. These are succeeded by a series of +greenish-grey or pale-grey slates ("Tarannon Shales"), sometimes +of great thickness; and these two groups of beds together form +what may be termed the "<i>May Hill Group</i>" (Upper Llandovery +of Murchison). Though not very extensively developed in Britain, +this zone is one very well marked by its fossils; and it corresponds +with the "Clinton Group" of North America, in which similar fossils +occur. In South Wales this group is clearly unconformable to the +highest member of the subjacent Lower Silurian (the Llandovery +group); and there is reason to believe that a similar, though +less conspicuous, physical break occurs very generally between +the base of the Upper and the summit of the Lower Silurian. +</p> + +<p class="indent"> +(2) The <i>Wenlock Group</i> succeeds the May Hill group, and +constitutes the middle member of the Upper Silurian. At its base +it may have an irregular limestone ("Woolhope Limestone"), and +its summit may be formed by a similar but thicker calcareous +deposit ("Wenlock Limestone"); but the bulk of the group is made +up of the argillaceous and shaly strata known as the "Wenlock +Shale." In North Wales the Wenlock group is, represented by a +great accumulation of flaggy and gritty strata (the "Denbighshire +Flags and Grits"), and similar beds (the +<a name="page_116"><span class="page">Page 116</span></a> +"Coniston Flags" and "Coniston Grits") take the same place in +the north of England. +</p> + +<p class="indent"> +(3) The <i>Ludlow Group</i> is the highest member of the Upper +Silurian, and consists typically of a lower arenaceous and shaly +series (the "Lower Ludlow Rock") a middle calcareous member (the +"Aymestry Limestone"), and an upper shaly and sandy series (the +"Upper Ludlow Rock" and "Downton Sandstone"). At the summit, or +close to the summit, of the Upper Ludlow, is a singular stratum +only a few inches thick (varying from an inch to a foot), which +contains numerous remains of crustaceans and fishes, and is well +known under the name of the "bone-bed." Finally, the Upper Ludlow +rock graduates invariably into a series of red sandy deposits, +which, when of a flaggy character, are known locally as the +"Tile-stones." These beds are probably to be regarded as the +highest member of the Upper Silurian; but they are sometimes +looked upon as passage-beds into the Old Red Sandstone, or as +the base of this formation. It is, in fact, apparently impossible +to draw any actual line of demarcation between the Upper Silurian +and the overlying deposits of the Devonian or Old Red Sandstone +series. Both in Britain and in America the Lower Devonian beds +repose with perfect conformity upon the highest Silurian beds, and +the two formations appear to pass into one another by a gradual +and imperceptible transition. +</p> + +<p class="indent"> +The Upper Silurian strata of Britain vary from perhaps 3000 or +4000 feet in thickness up to 8000 or 10,000 feet. In North America +the corresponding series, though also variable, is generally of +much smaller thickness, and may be under 1000 feet. The general +succession of the Upper Silurian deposits of North America is +as follows:— +</p> + +<p class="indent"> +(1) <i>Medina Sandstone</i>.—This constitutes the base of the +Upper Silurian, and consists of sandy strata, singularly devoid +of life, and passing below in some localities into a conglomerate +("Oneida Conglomerate"), which is stated to contain pebbles derived +from the older beds, and which would thus indicate an unconformity +between the Upper and Lower Silurian. +</p> + +<p class="indent"> +(2) <i>Clinton Group</i>.—Above the Medina sandstone are beds +of sandstone and shale, sometimes with calcareous bands, which +constitute what is known as the "Clinton Group." The Medina and +Clinton groups are undoubtedly the equivalent of the "May Hill +Group" of Britain, as shown by the identity of their fossils. +</p> + +<p class="indent"> +<span style="width: 100%; text-align: center;"> +<span style="margin: 4px; width: 575px; + font-size: smaller; text-align: center;"> +<a name="page_117"><span class="page">Page 117</span></a> +GENERALIZED SECTION OF THE UPPER SILURIAN STRATA OF WALES AND +SHROPSHIRE. +<br /> +Fig. 57. +<br /> +<img src="images/fig057.jpg" width="546" height="505" alt="Fig. 57" /> +</span> +</span> + +(3) <i>Niagara Group</i>.—This group consists typically of a +series of argillaceous beds ("Niagara Shale") capped by limestones +("Niagara Limestone"); and the name of the group is derived from +the fact that it is over limestones of this age that the Niagara +river is precipitated to form the great Falls. In places the +Niagara group is wholly calcareous, and it is continued upwards +into a series of marls and sandstones, with beds of salt and masses +of gypsum (the "Salina Group"), or into a series of magnesian +limestones ("Guelph Limestones"). The Niagara group, as a whole, +corresponds unequivocally with the Wenlock group of Britain. +</p> + +<p class="indent"> +(4) <i>Lower Helderberg Group</i>.—The Upper Silurian period +in North America was terminated by the deposition of a series of +calcareous beds, which derive the name of "Lower Helderberg" from +the Helderberg mountains, south of Albany, and +<a name="page_118"><span class="page">Page 118</span></a> +which are divided into several zones, capable of recognition +by their fossils, and known by local names (Tentaculite Limestone, +Water-lime, Lower Pentamerus Limestone, Delthyris Shaly Limestone, +and Upper Pentamerus Limestone). As a whole, this series may be +regarded as the equivalent of the Ludlow group of Britain, though +it is difficult to establish any precise parallelism. The summit +of the Lower Heiderberg group is constituted by a coarse-grained +sandstone (the "Oriskany Sandstone"), replete with organic +remains, which have to a large extent a Silurian <i>facies</i>. +Opinions differ as to whether this sandstone is to be regarded +as the highest bed of the Upper Silurian or the base of the +Devonian. We thus see that in America, as in Britain, no other +line than an artificial one can be drawn between the Upper +Silurian and the overlying Devonian. +</p> + +<p class="indent"> +As regards the <i>life</i> of the Upper Silurian period, we have, +as before, a number of so-called "Fucoids," the true vegetable +nature of which is in many instances beyond doubt. In addition +to these, however, we meet for the first time, in deposits of +this age, with the remains of genuine land-plants, though our +knowledge of these is still too scanty to enable us to construct +any detailed picture of the terrestrial vegetation of the period. +Some of these remains indicate the existence of the remarkable +genus <i>Lepidodendron</i>—a genus which played a part of +great importance in the forests of the Devonian and Carboniferous +periods, and which may be regarded as a gigantic and extinct type of +the Club-mosses (<i>LycopodiaceĹ“</i>). Near the summit of the +Ludlow formation in Britain there have also been found beds charged +with numerous small globular bodies, which Dr Hooker has shown +to be the seed-vessels or "sporangia" of Club-mosses. Principal +Dawson further states that he has seen in the same formation +fragments of wood with the structure of the singular Devonian +Conifer known as <i>Prototaxites</i>. Lastly, the same distinguished +observer has described from the Upper Silurian of North America +the remains of the singular land-plants belonging to the genus +<i>Psilophyton</i>, which will be referred to at greater length +hereafter. +</p> + +<p class="indent"> +The marine life of the Upper Silurian is in the main constituted by +types of animals similar to those characterising the Lower Silurian, +though for the most part belonging to different species. The +<i>Protozoans</i> are represented principally by <i>Stromatopora</i> +and <i>Ischadites</i>, along with a number of undoubted sponges +(such as <i>Amphispongia, AstrĹ“ospongia, Astylospongia</i>, +and <i>PalĹ“omanon</i>). +</p> + +<p class="indent"> +Amongst the <i>CĹ“lenterates</i>, we find the old group of +<i>Graptolites</i> now verging on extinction. Individuals still +<a name="page_119"><span class="page">Page 119</span></a> +remain numerous, but the variety of generic +and specific types has now become greatly reduced. All the +branching and complex forms of the Arenig, the twin-Graptolites + +<span style="float: right; margin: 4px; width: 199px; + font-size: smaller; text-align: justify;"> +<img src="images/fig058.jpg" width="191" height="424" alt="Fig. 1" /> +<br /> +Fig. 58.—A, <i>Monograptus priodon</i>, slightly enlarged. +B, Fragment of the same viewed from behind. C, Fragment of the +same viewed in front, showing the mouths of the cellules. D, +Cross-section of the same. From the Wenlock Group (Coniston +Flags of the North of England). (Original.) +</span> + +and <i>Dicranograpti</i> of the Llandeilo, and the double-celled +<i>Diplograpti</i> and <i>Climacograpti</i> of the Bala group, +have now disappeared. In their place we have the singular +<i>Retiolites</i>, with its curiously-reticulated skeleton; and +several species of the single-celled genus <i>Monograptus</i>, +of which a characteristic species (<i>M. Priodon</i>) is here +figured. If we remove from this group the plant-like +<i>DictyonemĹ“</i>, which are still present, and which +survive into the Devonian, no known species of <i>Graptolite</i> +has hitherto been detected in strata higher in geological position +than the Ludlow. This, therefore, presents us with the first +instance we have as yet met with of the total disappearance and +extinction of a great and important series of organic forms. +</p> + +<p class="indent"> +The <i>Corals</i> are very numerously represented in the Upper +Silurian rocks some of the limestones (such as the Wenlock Limestone) +being often largely composed of the skeletons of these animals. +Almost all the known forms of this period belong to the two great +divisions of the Rugose and Tabulate corals, the former being +represented by species of <i>Zaphrentis, Omphyma, Cystiphyllum, +Strombodes, Acervularia, Cyathophyllum</i>, &c.; whilst the latter +belong principally to the genera <i>Favosites, ChĹ“tetes, +Halysites, Syringopora, Heliolites</i>, and <i>Plasmopora</i>. +Amongst the <i>Rugosa</i>, the first appearance of the great and +important genus <i>Cyathophyllum</i>, so characteristic of the +Palæozoic period, is to be noted; and amongst the +<i>Tabulata</i> we have similarly the first appearance, in force +at any rate, of the widely-spread genus <i>Favosites</i>—the +"Honeycomb-corals." The "Chain-corals" (<i>Halysites</i>), figured +below (fig. 59), are also very common examples of the Tabulate +corals during this period, though they occur likewise in the Lower +Silurian. +</p> + +<p class="indent"> +<a name="page_120"><span class="page">Page 120</span></a> +Amongst the <i>Echinodermata</i>, all those orders which have hard +parts capable of ready preservation are more or less largely + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 508px; + font-size: smaller; text-align: justify;"> +<img src="images/fig059.jpg" width="500" height="383" alt="Fig. 59" /> +<br /> +Fig. 59.—<i>a, Halysites catenularia</i>, small variety, of +the natural size; <i>b,</i> Fragment of a large variety of the +same, of the natural size; <i>c</i>, Fragment of limestone with +the tubes of <i>Halysites agglomerata</i>, of the natural size; +<i>d,</i> Vertical section of two tubes of the same, showing the +tabulæ, enlarged. Niagara Limestone (Wenlock), Canada. +(Original.) +</span> +</span> + +represented. We have no trace of the Holothurians or Sea-cucumbers; +but this is not surprising, as the record of the past is throughout +almost silent as to the former existence of these soft-bodied +creatures, the scattered plates and spicules in their skin offering +a very uncertain chance of preservation in the fossil condition. +The Sea-urchins (<i>Echinoids</i>) are said to be represented by +examples of the old genus <i>PalĹ“chinus</i>. The Star-fishes +(<i>Asteroids</i>) and the Brittle-stars (<i>Ophiuroids</i>) are, +comparatively speaking, largely represented; the former by species +of <i>Palasterina</i> (fig. 60), <i>PalĹ“aster</i> (fig. 60), +<i>PalĹ“ocoma</i> (fig. 60), <i>Petraster, Glyptaster</i>, and +<i>Lepidaster</i>—and the latter by species of <i>Protaster</i> +(fig. 61), <i>PalĹ“odiscus, Acroura</i>, and <i>Eucladia</i>. +The singular <i>Cystideans</i>, or "Globe Crinoids," with their +globular or ovate, tesselated bodies (fig. 46, A, C, D,), are also +not uncommon in the Upper Silurian; and if they do not become finally +extinct here, they certainly survive the close of this period by +but a very brief time. By far the most important, however, of the +Upper Silurian Echinodenns, are the Sea-lilies or <i>Crinoids</i>. +The limestones of this period are often largely composed of the +fragmentary columns and detached +<a name="page_121"><span class="page">Page 121</span></a> +plates of these creatures, and some +of them (such as the Wenlock Limestone of Dudley) have yielded + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 513px; + font-size: smaller; text-align: justify;"> +<img src="images/fig060.jpg" width="505" height="173" alt="Fig. 60" /> +<br /> +Fig. 60.—Upper Silurian Star-fishes. 1, <i>Palasterina +primĹ“va</i>, Lower Ludlow; 2, <i>Paloeaster Ruthveni</i>, +Lower Ludlow; 3, <i>PalĹ“ocoma Colvini</i>, Lower Ludlow. +(After Salter.) +</span> +</span> + +perhaps the most exquisitely-preserved examples of this group +with which we are as yet acquainted. However varied in their +forms, these beautiful organisms consist of a globular, ovate, or + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 534px; + font-size: smaller; text-align: justify;"> +<img src="images/fig061.jpg" width="526" height="316" alt="Fig. 61" /> +<br /> +Fig. 61.—A, <i>Protaster Sedgwickii</i>, showing the disc and +bases of the arms; B, Portion of an arm, greatly enlarged. +Lower Ludlow. (After Salter.) +</span> +</span> + +pear-shaped body (the "calyx"), supported upon a longer or shorter +jointed stem (or "column"). The body is covered externally with +an armour of closely-fitting calcareous plates (fig. 62), and +its upper surface is protected by similar but smaller plates +more loosely connected by a leathery integument. From the upper +surface of the body, round its margin, springs a series of longer +or shorter flexible processes, composed of innumerable calcareous +joints or pieces, movably united with one +<a name="page_122"><span class="page">Page 122</span></a> +another. The arms are +typically five in number; but they generally subdivide at least +once, sometimes twice, and they are furnished with similar but + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 533px; + font-size: smaller; text-align: justify;"> +<img src="images/fig062.jpg" width="525" height="352" alt="Fig. 62" /> +<br /> +Fig. 62.—Upper Silurian Crinoids. <i>a</i>, Calyx and arms +of <i>Eucalyptocrinus polydactylus</i>, Wenlock Limestone; <i>b, +Ichthyocrinus lĹ“vis</i>, Niagara Limestone, America; <i>c, +Taxocrinus tuberculatus</i>, Wenlock Limestone. (After M'Coy +and Hall.) +</span> +</span> + +more slender lateral branches or "pinnules," thus giving rise +to a crown of delicate feathery plumes. The "column" is the stem +by which the animal is attached permanently to the bottom of the +sea; and it is composed of numerous separate plates, so jointed +together that whilst the amount of movement between any two pieces +must be very limited, the entire column acquires more or less +flexibility, allowing the organism as a whole to wave backwards +and forwards on its stalk. Into the exquisite <i>minutiĹ“</i> +of structure by which the innumerable parts entering into the +composition of a single Crinoid are adapted for their proper +purposes in the economy of the animal, it is impossible to enter +here. No period, as before said, has yielded examples of greater +beauty than the Upper Silurian, the principal genera represented +being <i>Cyathocrinus, Platycrinus, Marsupiocrinus, Taxocrinus, +Eucalyptocrinus, Ichthyocrinus, Mariacrinus, Periechocrinus, +Glyptocrinus, Crotalocrinus</i>, and <i>Edriocrinus</i>. +</p> + +<p class="indent"> +The tracks and burrows of <i>Annelides</i> are as abundant in +the Upper Silurian strata as in older deposits, and have just +as commonly been regarded as plants. The most abundant forms +are the cylindrical, twisted bodies (Planolites), which are +<a name="page_123"><span class="page">Page 123</span></a> +so frequently found on the surfaces of sandy +beds, and which have been described as the stems of sea-weeds. +These fossils (fig. 63), however, can be nothing more, in most + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 531px; + font-size: smaller; text-align: center;"> +<img src="images/fig063.jpg" width="523" height="421" alt="Fig. 63" /> +<br /> +Fig. 63.—<i>Planolites vulgaris</i>, the filled-up burrows +of a marine worm. Upper Silurian (Clinton Group), Canada. +(Original.) +</span> +</span> + +cases, than the filled-up +burrows of marine worms resembling the living Lob-worms. There +are also various remains which belong to the group of the +tube-inhabiting Annelides (<i>Tubicola</i>). Of this nature are +the tubes of <i>Serpulites</i> and <i>Cornultites</i>, and the +little spiral discs of <i>Spirorbis Lewisii</i>. +</p> + +<p class="indent"> +Amongst the <i>Articulates</i>, we still meet only with the remains +of <i>Crustaceans</i>. Besides the little bivalved +<i>Ostracoda</i>—which here are occasionally found of the size +of beans—and various <i>Phyllopods</i> of different kinds, we +have an abundance of <i>Trilobites</i>. These last-mentioned ancient +types, however, are now beginning to show signs of decadence; and +though still individually numerous, there is a great diminution +in the number of generic types. Many of the old genera, which +flourished so abundantly in Lower Silurian seas, have now died out; +and the group is represented chiefly by species of <i>Cheirurus, +Encrinurus, Harpes, Proetus, Lichas, Acidaspis, IllĹ“nus, +Calymene, Homalonotus</i>, and <i>Phacops</i>—the last of +these, one of the +<a name="page_124"><span class="page">Page 124</span></a> +highest and most beautiful of the groups of Trilobites, attaining +here its maximum of development. In the annexed illustration +(fig. 64) some of the characteristic Upper Silurian Trilobites are + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 518px; + font-size: smaller; text-align: justify;"> +<img src="images/fig064.jpg" width="510" height="383" alt="Fig. 64" /> +<br /> +Fig. 64.—Upper Silurian Trilobites. <i>a, Cheirurus +bimucronatus</i>, Wenlock and Caradoc; <i>b, Phacops +longicaudatus</i>, Wenlock, Britain, and America; <i>c, Phacops +DowningiĹ“</i>, Wenlock and Ludlow; <i>d, Harpes ungula</i>, +Upper Silurian, Bohemia. (After Salter and Barrande.) +</span> +</span> + +represented—all, however, belonging to genera which have their +commencement in the Lower Silurian period. In addition to the above, +the Ludlow rocks of Britain and the Lower Helderberg beds of North +America have yielded the remains of certain singular Crustaceans +belonging to the extinct order of the <i>Eurypterida</i>. Some +of these wonderful forms are not remarkable for their size; but +others, such as <i>Pterygotus Anglicus</i> (fig. 65), attain a +length of six feet or more, and may fairly be considered as the +giants of their class. The Eurypterids are most nearly allied +to the existing King-crabs (<i>Limuli</i>), and have the anterior +end of the body covered with a great head-shield, carrying two +pairs of eyes, the one simple and the other compound. The feelers +are converted into pincers, whilst the last pair of limbs have +their bases covered with spiny teeth so as to act as jaws, and +are flattened and widened out towards their extremities so as +to officiate as swimming-paddles. The hinder extremity of the +body is composed of thirteen rings, which have no legs attached +to them; and the last segment of the tail is either a flattened +plate or a +<a name="page_125"><span class="page">Page 125</span></a> +narrow, sword-shaped spine. Fragments of the skeleton +are easily recognised by the peculiar scale-like markings with + +<span style="float: right; margin: 4px; width: 254px; + font-size: smaller; text-align: justify;"> +<img src="images/fig065.jpg" width="235" height="442" alt="Fig. 65" /> +<br /> +Fig. 65.—<i>Pterygotus Anglicus</i>, viewed +from the under side, reduced in size, and restored. <i>c c</i>, +The feelers (antennæ), terminating in nipping-claws; <i>o +o</i>, Eyes; <i>m m</i>, Three pairs of jointed limbs, with +pointed extremities; <i>n n</i>, Swimming-paddles, the bases of +which are spiny and act as jaws. Upper Silurian, Lanarkshire. +(After Henry Woodward.) +</span> + +which the surface is adorned, and which look not at all unlike +the scales of a fish. The most famous locality for these great +Crustaceans is Lesmahagow, in Lanarkshire, where many different +species have been found. The true King-crabs (<i>Limuli</i>) of +existing seas also appear to have been represented by at least +one form (<i>Neolimulus</i>) in the Upper Silurian. +</p> + +<p class="indent"> +Coming to the <i>Mollusca</i>, we note the occurrence of the same +great groups as in the Lower Silurian. Amongst the Sea-mosses +(<i>Polyzoa</i>), we have the ancient Lace-corals (<i>Fenestella</i> +and <i>Retepora</i>), with the nearly-allied <i>Glauconome</i>, +and species of <i>Ptilodictya</i> (fig. 66); whilst many forms +often referred here may probably have to be transferred to the +Corals, just as some so-called Corals will ultimately be removed +to the present group. +</p> + +<p class="indent"> +The Brachiopods continued to flourish during the Upper Silurian +Period in immense numbers and under a greatly increased variety +of forms. The three prominent Lower Silurian genera <i>Orthis, +Strophomena</i>, and <i>LeptĹ“na</i> are still well represented, +though they have lost their former preeminence. Amongst the numerous +types which have now come upon the scene for the first time, or +which have now a special development, are <i>Spirifera</i> and +<i>Pentamerus</i>. In the first of these (fig. 69. <i>b, c</i>), +one of the valves of the shell (the dorsal) is furnished in its +interior with a pair of great calcareous spires, which served +for the support of the long and fringed fleshy processes or "arms" +which were attached to the sides of the mouth.[16] In the genus +<i>Pentamerus</i> (fig. 70) the +<a name="page_126"><span class="page">Page 126</span></a> +shell is curiously subdivided in its +interior by calcareous plates. The <i>Pentameri</i> commenced their +existence at the very close of the Lower Silurian (Llandovery), and + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 475px; + font-size: smaller; text-align: justify;"> +<img src="images/fig066.jpg" width="467" height="220" alt="Fig. 66" /> +<br /> +Fig. 66.—Upper Silurian Polyzoa. <i>1</i>, Fan-shaped +frond of <i>Rhinopora verrucosa</i>; 1<i>a</i>, Portion of the +surface of the same, enlarged; 2 and 2<i>a</i>, <i>PhĹ“nopora +ensiformis</i>, of the natural size and enlarged; 3 and 3<i>a</i>, +<i>Helopora fragilis</i>, of the natural size and enlarged; 4 +and 4<i>a</i>, <i>Ptilodictya raripora</i>, of the natural size +and enlarged. The specimens are all from the Clinton Formation +(May Hill Group) of Canada. (Original.) +</span> +</span> + +survived to the close of the Upper Silurian; but they are specially +characteristic of the May Hill and Wenlock groups, both in Britain +and in other regions. One species, <i>Pentamerus galeatus</i>, is +common to Sweden, Britain, and America. Amongst the remaining Upper +Silurian Brachiopods are the extraordinary +<a name="page_127"><span class="page">Page 127</span></a> +<i>Trimerellids</i>; the old and at the same time modern +<i>LingulĹ“, DiscinĹ“</i>, and <i>CraniĹ“</i>; +together with many species of <i>Atrypa</i> (fig. 68, <i>e</i>), + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 523px; + font-size: smaller; text-align: justify;"> +<img src="images/fig068.jpg" width="515" height="225" alt="Fig. 68" /> +<br /> +Fig. 68.—Upper +Silurian Brachiopods. <i>a a'</i>, <i>LeptocĹ“lia +plano-convexa</i>, Clinton Group, America; <i>b b'</i>, +<i>Rhynchonella neglecta</i>, Clinton Group, America; <i>c, +Rhynchonella cuneata</i>, Niagara Group, America, and Wenlock +Group, Britain; <i>d d', Orthis elelgantula</i>, Llandeilo to +Ludlow, America and Europe; <i>e e', Atrypa hemispherica</i>, +Clinton Group, America, and Llandovery and May Hill Groups, +Britain; <i>f f', Atrypa congesta</i>, Clinton Group, America; +<i>g g', Orthis Davidsoni</i>, Clinton Group, America. (After +Hall, Billings, and the Author.) +</span> +</span> + +<i>LeptocĹ“lia</i> (fig. 68, <i>a</i>), +<i>Rhynchonella</i> (fig. 68, <i>b, c</i>), <i>Meristella</i> +(fig. 69, <i>a, e, f</i>), <i>Athyris, Retzia, Chonetes</i>, +&c. +</p> + +<p class="footnote"> +[Footnote 16: In all the Lamp-shells the mouth is provided with +two long fleshy organs, which carry delicate filaments on their +sides, and which are usually coiled into a spiral. These organs +are known as the "arms," and it is from their presence that the +name of "<i>Brachiopoda</i>" is derived (Gr. <i>brachion</i>, arm; +<i>podes</i>, feet). In some cases the arms are merely coiled away +within the shell, without any support; but in other cases they are +carried upon a more or less elaborate shelly loop, often spoken +of as the "carriage-spring apparatus." In the <i>Spirifers</i>, +and in other ancient genera, this apparatus is coiled up into a +complicated spiral (fig. 67). It is these "arms," with or without + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 518px; text-align: center;"> +<img src="images/fig067.jpg" width="510" height="161" alt="Fig. 67" /> +<br /> +Fig. 67.—<i>Spirifera hysterica</i>. The right-hand figure +shows the interior of the dorsal valve with the calcareous +spires for the support of the arms. +</span> +</span> + +the supporting loops or spires, which serve as one of the special +characters distinguishing the <i>Brachiopods</i> from the true +Bivalves (<i>Lamellibranchiata</i>).] +</p> + +<div class="center"> +<table border="0" width="519"> +<tr><td class="center"> +<img src="images/fig069.jpg" width="515" height="358" alt="Fig. 69" /> +</td></tr> +<tr><td class="left"><span class="image"> +Fig. 69.—<i>a a', Meristella intermedia</i>, Niagara Group, America; +<i>b, Spirifera Niagarensis</i>, Niagara Group, America; <i>c c', +Spirifera crispa</i>, May Hill to Ludlow, Britain, and Niagara +Group, America; <i>d, Strophomena (Streptorhynchus) subplana</i>, +Niagara Group, America; <i>e, Meristella naviformis</i>, Niagara +Group, America; <i>f, Meristella cylindrica</i>, Niagara Group, +America. (After Hall, Billings, and the Author.) +</span></td></tr> +</table> +</div> + +<p class="indent"> +The higher groups of the <i>Mollusca</i> are also largely represented +in the Upper Silurian. Apart from some singular types, +<a name="page_128"><span class="page">Page 128</span></a> +such as the +huge and thick-shelled <i>Megalomi</i> of the American Wenlock +formation, the Bivalves (<i>Lamellibranchiata</i>) present little of + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 539px; + font-size: smaller; text-align: center;"> +<img src="images/fig070.jpg" width="531" height="199" alt="Fig. 70" /> +<br /> +Fig. 70.—<i>Pentamerus Knightii</i>. Wenlock and Ludlow. The +right-hand figure shows the internal partitions of the shell. +</span> +</span> + +special interest; for though sufficiently numerous, they are rarely +well preserved, and their true affinities are often uncertain. +Amongst the most characteristic genera of this period may be +mentioned <i>Cardiola</i> (fig. 71, A and C) and <i>Pterinea</i> + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 529px; + font-size: smaller; text-align: justify;"> +<img src="images/fig071.jpg" width="521" height="176" alt="Fig. 71" /> +<br /> +Fig. 71.—Upper Silurian Bivalves. A, <i>Cardiola +interrupta</i>, Wenlock and Ludlow; B, <i>Pterinea subfalcata</i>, +Wenlock; C, <i>Cardiola fibrosa</i>, Ludlow. (After Salter and +M'Coy.) +</span> +</span> + +(fig. 71, B), though the latter survives to a much later date. +The Univalves (<i>Gasteropoda</i>) are very numerous, and a +few characteristic forms are here figured (fig. 72). Of these, +no genus is perhaps more characteristic than <i>Euomphalus</i> +(fig. 72, <i>b</i>), with its flat discoidal shell, coiled up +into an oblique spiral, and deeply hollowed out on one side; +but examples of this group are both of older and of more modern +date. Another very extensive genus, especially in America, is +Platyceras (fig. 72, <i>a</i> and <i>f</i>), with its thin fragile +shell—often hardly coiled up at all—its minute spire, +and its widely-expanded, often sinuated mouth. The British +<i>AcroculiĹ“</i> should probably be placed here, and the +group has with reason been regarded as allied to the Violet-snails +(<i>Ianthina</i>) of the open Atlantic. The +<a name="page_129"><span class="page">Page 129</span></a> +species of <i>Platyostoma</i> (fig. 72, <i>h</i>) also belong +to the same family; and the entire group is continued throughout +the Devonian into the Carboniferous. Amongst other well-known +Upper Silurian Gasteropods are species of the genera <i>Holopea</i> +(fig. 72, <i>g</i>), <i>Holopella</i> (fig. 72. <i>e</i>), + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 541px; + font-size: smaller; text-align: justify;"> +<img src="images/fig072.jpg" width="533" height="339" alt="Fig. 72" /> +<br /> +Fig. 72.—Upper Silurian Gasteropods. <i>a, Platyceras +ventricosum</i>, Lower Helderberg, America; <i>b, Euomphalus +discors</i>, Wenlock, Britain; <i>c, Holopella obsoleta</i> Ludlow, +Britain; <i>d, Platyschisma helicites</i>, Upper Ludlow, Britain; +<i>e, Holopella gracilior</i>, Wenlock, Britain; <i>f, Platyceras +multisinuatum</i>, Lower Helderberg, America; <i>g, Holopea +subconica</i>, Lower Helderberg, America; <i>h, h', Platyostoma +Niagarense</i>, Niagara Group, America. (After Hall, M'Coy, and +Salter.) +</span> +</span> + +<i>Platyschisma</i> (fig. 72, <i>d</i>), <i>Cyclonema, +Pleurotomaria, Murchisonia, Trochonema</i>, &c. The oceanic + +<span style="float: right; margin: 4px; width: 59px; + font-size: smaller; text-align: justify;"> +<img src="images/fig073.jpg" width="51" height="207" alt="Fig. 73" /> +<br /> +Fig. 73.—<i>Tentaculites ornatus</i>. Upper Silurian of Europe +and North America. +</span> + +Univalves (<i>Heteropods</i>) are represented mainly by species of +<i>Bellerophon</i>; and the Winged Snails, or <i>Pteropods</i>, can +still boast of the gigantic <i>ThecĹ“</i> and +<i>ConulariĹ“</i>, which characterise yet older deposits. The +commonest genus of <i>Pteropoda</i>, however, is <i>Tentaculites</i> +(fig. 73), which clearly belongs here, though it has commonly been +regarded as the tube of an Annelide. The shell in this group +is a conical tube, usually adorned with prominent transverse +rings, and often with finer transverse or longitudinal striæ +as well; and many beds of the Upper Silurian exhibit myriads of +such tubes scattered promiscuously over their surfaces. +</p> + +<p class="indent"> +<a name="page_130"><span class="page">Page 130</span></a> +The last and highest group of the <i>Mollusca</i>—that +of the <i>Cephalopoda</i>—is still represented only by +<i>Tetrabranchiate</i> forms; but the abundance and variety of +these is almost beyond belief. Many hundreds of different species +are known, chiefly belonging to the straight <i>Orthoceratites</i>, +but the slightly-curved <i>Cyrtoceras</i> is only little less +common. There are also numerous forms of the genera <i>Phragmoceras, +Ascoceras, Gyroteras, Lituites</i>, and <i>Nautilus</i>. Here, also, +are the first-known species of the genus <i>Goniatites</i>—a +group which attains considerable importance in later deposits, and +which is to be regarded as the precursor of the <i>Ammonites</i> of +the Secondary period. +</p> + +<p class="indent"> +Finally, we find ourselves for the first time called upon to +consider the remains of undoubted vertebrate animals, in the + +<span style="float: left; margin: 4px; width: 164px; + font-size: smaller; text-align: justify;"> +<img src="images/fig074.jpg" width="156" height="197" alt="Fig. 74" /> +<br /> +Fig. 74.—Head-shield of <i>Pteraspis Banksii</i>, Ludlow +rocks. (After Murchison.) +</span> + +form of <i>Fishes</i>. The oldest of these remains, so far as +yet known, are found in the Lower Ludlow rocks, and they consist +of the bony head-shields or bucklers of certain singular armoured +fishes belonging to the group of the <i>Ganoids</i>, represented at +the present day by the Sturgeons, the Gar-pikes of North America, +and a few other less familiar forms. The principal Upper Silurian +genus of these is <i>Pteraspis</i>, and the annexed illustration +(fig. 74) will give some idea of the extraordinary form of the +shield covering the head in these ancient fishes. The remarkable +stratum near the top of the Ludlow formation known as the "bone-bed" +has also yielded the remains of shark-like fishes. Some of these, +for which the name of <i>Onchus</i> has been proposed, are in +the form of compressed, slightly-curved spines (fig. 75, A), +which would appear to be of the nature of the strong defensive +spines implanted in front of certain of the fins in many living +fishes. Besides these, have been found fragments of prickly skin + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 462px; + font-size: smaller; text-align: center;"> +<img src="images/fig075.jpg" width="454" height="66" alt="Fig. 75" /> +<br /> +Fig. 75.—A, Spine of <i>Onchus tenuistriatus</i>; B, +Shagreen-scales of <i>Thelodus</i>. Both from the "bone-bed" +of the Upper Ludlow rocks. (After Murchison.) +</span> +</span> + +or shagreen (<i>Sphagodus</i>), along with minute cushion-shaped +bodies (<i>Thelodus</i>, fig. 75, B), which +<a name="page_131"><span class="page">Page 131</span></a> +are doubtless the bony scales of some fish resembling the modern +Dog-fishes. As the above mentioned remains belong to two distinct, +and at the same time highly-organised, groups of the fishes, it +is hardly likely that we are really presented here with the first +examples of this great class. On the contrary, whether the +so-called "Conodonts" should prove to be the teeth of fishes or +not, we are justified in expecting that unequivocal remains of +this group of animals will still be found in the Lower Silurian. +It is interesting, also, to note that the first appearance of +fishes—the lowest class of vertebrate animals—so far +as known to us at present, does not take place until after all +the great sub-kingdoms of invertebrates have been long in +existence; and there is no reason for thinking that future +discoveries will materially affect the <i>relative</i> order of +succession thus indicated. +</p> + +<h4>LITERATURE.</h4> + +<p class="indent"> +From the vast and daily-increasing mass of Silurian literature, it +is impossible to do more than select a small number of works which +have a classical and historical interest to the English-speaking +geologist, or which embody researches on special groups of Silurian +animals—anything like an enumeration of all the works and +papers on this subject being wholly out of the question. Apart, +therefore, from numerous and in many cases extremely important +memoirs, by various well-known observers, both at home and abroad, +the following are some of the more weighty works to which the +student may refer in investigating the physical characters and +succession of the Silurian strata and their fossil contents:— +</p> + +<table border="0" cellspacing="0"> +<tr><td class="right" valign="top">(1)</td> + <td>'Siluria.' Sir Roderick Murchison.</td></tr> +<tr><td class="right" valign="top">(2)</td> + <td>'Geology of Russia in Europe.' Murchison (with M. de Verneuil + and Count von Keyserling).</td></tr> +<tr><td class="right" valign="top">(3)</td> + <td>'Bassin Silurien de BohĂŞme Centrale.' Barrande.</td></tr> +<tr><td class="right" valign="top">(4)</td> + <td>'Introduction to the Catalogue of British Palæozoic + Fossils in the Woodwardian Museum of Cambridge.' + Sedgwick.</td></tr> +<tr><td class="right" valign="top">(5)</td> + <td>'Die Urwelt Russlands.' Eichwald.</td></tr> +<tr><td class="right" valign="top">(6)</td> + <td>'Report on the Geology of Londonderry, Tyrone,' &c. + Portlock.</td></tr> +<tr><td class="right" valign="top">(7)</td> + <td>"Geology of North Wales"—'Mem. Geol. Survey of Great + Britain,' vol. iii. Ramsay.</td></tr> +<tr><td class="right" valign="top">(8)</td> + <td>'Geology of Canada,' 1863. Sir W. E. Logan; and the + 'Reports of Progress of the Geological Survey' since + 1863.</td></tr> +<tr><td class="right" valign="top">(9)</td> + <td>'Memoirs of the Geological Survey of Great + Britain.'</td></tr> +<tr><td class="right" valign="top">(10)</td> + <td>'Reports of the Geological Surveys of the States of New + York, Illinois, Ohio, Iowa, Michigan, Vermont, Wisconsin, + Minnesota,' &c. By Emmons, Hall, Worthen, Meek, Newberry, + Orton, Winchell, Dale Owen, &c.</td></tr> +<tr><td class="right" valign="top">(11)</td> + <td>'Thesaurus Siluricus.' Bigsby.</td></tr> +<tr><td class="right" valign="top">(12)</td> + <td>'British Palæozoic Fossils.' M'Coy.</td></tr> +<tr><td class="right" valign="top">(13)</td> + <td>'Synopsis of the Silurian Fossils of Ireland,' + M'Coy.</td></tr> +<tr><td class="right" valign="top">(14)</td> + <td>"Appendix to the Geology of North Wales"—'Mem. Geol. + Survey,' vol. iii. Salter.</td></tr> +<tr><td class="right" valign="top"> +<a name="page_132"><span class="page">Page 132</span></a> + (15)</td> + <td>'Catalogue of the Cambrian and Silurian Fossils in the + Woodwardian Museum of Cambridge.' Salter.</td></tr> +<tr><td class="right" valign="top">(16)</td> + <td>'Characteristic British Fossils.' Baily.</td></tr> +<tr><td class="right" valign="top">(17)</td> + <td>'Catalogue of British Fossils.' Morris.</td></tr> +<tr><td class="right" valign="top">(18)</td> + <td>'Palæozoic Fossils of Canada.' Billings.</td></tr> +<tr><td class="right" valign="top">(19)</td> + <td>'Decades of the Geological Survey of Canada.' Billings, + Salter, Rupert Jones.</td></tr> +<tr><td class="right" valign="top">(20)</td> + <td>'Decades of the Geological Survey of Great Britain.' Salter, + Edward, Forbes.</td></tr> +<tr><td class="right" valign="top">(21)</td> + <td>'Palæontology of New York,' vols. i.-iii. + Hall.</td></tr> +<tr><td class="right" valign="top">(22)</td> + <td>'Palæontology of Illinois.' Meek and + Worthen.</td></tr> +<tr><td class="right" valign="top">(23)</td> + <td>'Palæontology of Ohio.' Meek, Hall, Whitfield, + Nicholson.</td></tr> +<tr><td class="right" valign="top">(24)</td> + <td>'Silurian Fauna of West Tennessee' (Silurische Fauna des + Westlichen Tennessee). Ferdinand RĹ“mer.</td></tr> +<tr><td class="right" valign="top">(25)</td> + <td>'Reports on the State Cabinet of New York.' + Hall.</td></tr> +<tr><td class="right" valign="top">(26)</td> + <td>'Lethæa Geognostica.' Bronn.</td></tr> +<tr><td class="right" valign="top">(27)</td> + <td>'Index Palæontologicus.' Bronn.</td></tr> +<tr><td class="right" valign="top">(28)</td> + <td>'Lethæa Rossica.' Eichwald.</td></tr> +<tr><td class="right" valign="top">(29)</td> + <td>'Lethæa Suecica.' Hisinger.</td></tr> +<tr><td class="right" valign="top">(30)</td> + <td>'Palæontologica Suecica.' Angelin.</td></tr> +<tr><td class="right" valign="top">(31)</td> + <td>'Petrefacta Germaniæ.' Goldfuss.</td></tr> +<tr><td class="right" valign="top">(32)</td> + <td>'Versteinerungen der Grauwacken-Formation in Sachsen.' + Geinitz.</td></tr> +<tr><td class="right" valign="top">(33)</td> + <td>'Organisation of Trilobites' (Ray Society). + Burmeister.</td></tr> +<tr><td class="right" valign="top">(34)</td> + <td>'Monograph of the British Trilobites' + (Palæontographical Society). Salter.</td></tr> +<tr><td class="right" valign="top">(35)</td> + <td>'Monograph of the British Merostomata' + (Palæontographical Society). Henry Woodward.</td></tr> +<tr><td class="right" valign="top">(36)</td> + <td>'Monograph of British Brachiopoda' (Palæontographical + Society). Thomas Davidson.</td></tr> +<tr><td class="right" valign="top">(37)</td> + <td>'Graptolites of the Quebec Group.' James Hall.</td></tr> +<tr><td class="right" valign="top">(38)</td> + <td>'Monograph of the British Graptolitidæ.' + Nicholson.</td></tr> +<tr><td class="right" valign="top">(39)</td> + <td>'Monographs on the Trilobites. Pteropods, Cephalopods, + Graptolites,' &c. Extracted from the 'Système Silurien + du Centre de la BohĂŞme.' Barrande.</td></tr> +<tr><td class="right" valign="top">(40)</td> + <td>'Polypiers Fossiles des Terrains Paleozoiques,' and 'Monograph + of the British Corals' (Palæontographical Society). Milne + Edwards and Jules Haime.</td></tr> +</table> + +<h3>CHAPTER XI.</h3> + +<p class="subtitle"> +THE DEVONIAN AND OLD RED SANDSTONE PERIOD. +</p> + +<p class="indent"> +Between the summit of the Ludlow formation and the strata which +are universally admitted to belong to the Carboniferous series +<a name="page_133"><span class="page">Page 133</span></a> +is a great system of deposits, to which the +name of "Old Red Sandstone" was originally applied, to distinguish +them from certain arenaceous strata which lie above the coal ("New +Red Sandstone"). The Old Red Sandstone, properly so called, was +originally described and investigated as occurring in Scotland +and in South Wales and its borders; and similar strata occur in +the south of Ireland. Subsequently it was discovered that sediments +of a different mineral nature, and containing different organic +remains, intervened between the Silurian and the Carboniferous +rocks on the continent of Europe, and strata with similar +palæontological characters to these were found occupying +a considerable area in Devonshire. The name of "Devonian" was +applied to these deposits; and this title, by common usage, has +come to be regarded as synonymous with the name of "Old Red +Sandstone." Lastly, a magnificent series of deposits, containing +marine fossils, and undoubtedly equivalent to the true "Devonian" +of Devonshire, Rhenish Prussia, Belgium, and France, is found to +intervene in North America between the summit of the Silurian +and the base of the Carboniferous rocks. +</p> + +<p class="indent"> +Much difficulty has been felt in correlating the true "Devonian +Rocks" with the typical "Old Red Sandstone"—this difficulty +arising from the fact that though both formations are fossiliferous, +the peculiar fossils of each have only been rarely and partially +found associated together. The characteristic crustaceans and many +of the characteristic fishes of the Old Red are wanting in the +Devonian; whilst the corals and marine shells of the latter do +not occur in the former. It is impossible here to enter into any +discussion as to the merits of the controversy to which this +difficulty has given origin. No one, however, can doubt the +importance and reality of the Devonian series as an independent +system of rocks to be intercalated in point of time between the +Silurian and the Carboniferous. The want of agreement, both +lithologically and palæontologically, between the Devonian +and the Old Red, can be explained by supposing that these two +formations, though wholly or in great part <i>contemporaneous</i>, +and therefore strict equivalents, represent deposits in two +different geographical areas, laid down under different conditions. +On this view, the typical Devonian rocks of Europe, Britain, and +North America are the deep-sea deposits of the Devonian period, +or, at any rate, are genuine marine sediments formed far from +land. On the other hand, the "Old Red Sandstone" of Britain and +the corresponding "GaspĂ© Group" of Eastern +<a name="page_134"><span class="page">Page 134</span></a> +Canada represent the shallow-water shore-deposits +of the same period. In fact, the former of these last-mentioned +deposits contains no fossils which can be asserted positively +to be <i>marine</i> (unless the Eurypterids be considered so); +and it is even conceivable that it represents the sediments of +an inland sea. Accepting this explanation in the meanwhile, we +may very briefly consider the general succession of the deposits +of this period in Scotland, in Devonshire, and in North America. +</p> + +<p class="indent"> +In Scotland the "Old Red" forms a great series of arenaceous and +conglomeratic strata, attaining a thickness of many thousands +of feet, and divisible into three groups. Of these, the <i>Lower +Old Red Sandstone</i> reposes with perfect conformity upon the +highest beds of the Upper Silurian, the two formations being almost +inseparably united by an intermediate series of "passage-beds." +In mineral nature this group consists principally of massive +conglomerates, sandstones, shales, and concretionary limestones; +and its fossils consist chiefly of large crustaceans belonging +to the family of the <i>Eurypterids</i>, fishes, and plants. The +<i>Middle Old Red Sandstone</i> consists of flagstones, bituminous +shales, and conglomerates, sometimes with irregular calcareous +bands; and its fossils are principally fishes and plants. It +may be wholly wanting, when the <i>Upper Old Red</i> seems to +repose unconformably upon the lower division of the series. The +<i>Upper Old Red Sandstone</i> consists of conglomerates and +grits, along with a great series of red and yellow +sandstones—the fossils, as before, being fishes and remains +of plants. The Upper Old Red graduates upwards conformably into +the Carboniferous series. +</p> + +<p class="indent"> +The Devonian rocks of Devonshire are likewise divisible into +a lower, middle, and upper division. The <i>Lower Devonian</i> +or <i>Lynton Group</i> consists of red and purple sandstones, +with marine fossils, corresponding to the "Spirifer Sandstein" of +Germany, and to the arenaceous deposits (Schoharie and Cauda-Galli +Grits) at the base of the American Devonian. The <i>Middle +Devonian</i> or <i>Ilfracombe Group</i> consists of sandstones +and flags, with calcareous slates and crystalline limestones, +containing many corals. It corresponds with the great "Eifel +Limestone" of the Continent, and, in a general way, with the +Corniferous Limestone and Hamilton group of North America. The +<i>Upper Devonian</i> or <i>Pilton Group</i>, lastly, consists +of sandstones and calcareous shales which correspond with the +"Clymenia Limestone" and "Cypridina Shales" of the Continent, +and with the Chemung and Portage groups of +<a name="page_135"><span class="page">Page 135</span></a> +North America. It seems quite possible, also, that the +so-called "Carboniferous Slates" of Ireland correspond with +this group, and that the former would be more properly regarded +as forming the summit of the Devonian than the base of the +Carboniferous. +</p> + +<p class="indent"> +In no country in the world, probably, is there a finer or more +complete exposition of the strata intervening between the Silurian +and Carboniferous deposits than in the United States. The following +are the main subdivisions of the Devonian rocks in the State of +New York, where the series may be regarded as being typically +developed (fig. 67):— +</p> + +<p class="indent"> +(1) <i>Cauda-Galli Grit</i> and <i>Schoharie +Grit</i>.—Considering +the "Oriskany Sandstone" as the summit of the Upper Silurian, the +base of the Devonian is constituted by the arenaceous deposits +known by the above names, which rest quite conformably upon the +Silurian, and which represent the Lower Devonian of Devonshire. +The <i>Cauda-Galli Grit</i> is so called from the abundance of +a peculiar spiral fossil (<i>Spirophyton cauda-Galli</i>), which +is of common occurrence in the Carboniferous rocks of Britain, +and is supposed to be the remains of a sea-weed. +</p> + +<p class="indent"> +(2) The <i>Corniferous</i> or <i>Upper Helderberg +Limestone</i>.—A +series of limestones usually charged with considerable quantities +of siliceous matter in the shape of hornstone or chert (Lat. +<i>cornu</i>, horn). The thickness of this group rarely exceeds +300 feet; but it is replete with fossils, more especially with the +remains of corals. The Corniferous Limestone is the equivalent of +the coral-bearing limestones of the Middle Devonian of Devonshire +and the great "Eifel Limestone" of Germany. +</p> + +<p class="indent"> +(3) The <i>Hamilton Group</i>—consisting of shales at the +base ("Marcellus shales"); flags, shales, and impure limestones +("Hamilton beds") in the middle; and again a series of shales +("Genesee Slates") at the top. The thickness of this group varies +from 200 to 1200 feet, and it is richly charged with marine +fossils. +</p> + +<p class="indent"> +(4) The <i>Portage Group</i>.—A great series of shales, +flags, and shaly sandstones, with few fossils. +</p> + +<p class="indent"> +(5) The <i>Chemung Group</i>.—Another great series of +sandstones and shales, but with many fossils. The Portage and +Chemung groups may be regarded as corresponding with the Upper +Devonian of Devonshire. The Chemung beds are succeeded by a +great series of red sandstones and shales—the +<a name="page_136"><span class="page">Page 136</span></a> +"Catskill Group"—which pass conformably +upwards into the Carboniferous, and which may perhaps be +regarded as the equivalent of the great sandstones of the Upper +Old Red in Scotland. +</p> + +<p class="indent"> +Throughout the entire series of Devonian deposits in North America +no unconformability or physical break of any kind has hitherto been +detected; nor is there any marked interruption to the current of +life, though each subdivision of the series has its own fossils. +No completely natural line can thus be indicated, dividing the +Devonian in this region from the Silurian on the one hand, and the +Carboniferous on the other hand. At the same time, there is the +most ample evidence, both stratigraphical and palæontological, +as to the complete independence of the American Devonian series +as a distinct life-system between the older Silurian and the +later Carboniferous. The subjoined section (fig. 76) shows +diagrammatically the general succession of the Devonian rocks +of North America. +</p> + +<p class="indent"> +As regards the <i>life</i> of the Devonian period, we are now +acquainted with a large and abundant terrestrial +<i>flora</i>—this being the first time that we have met +with a land vegetation capable of reconstruction in any +fulness. By the researches of GĹ“ppert, Unger, Dawson, +Carruthers, and other botanists, a knowledge has been acquired +of a large number of Devonian plants, only a few of which can +be noticed here. As might have been anticipated, the greater +number of the vegetable remains of this period have been +obtained from such shallow-water deposits as the Old Red +Sandstone proper and the Gaspè series of North America, and +few traces of plant-life occur in the strictly marine sediments. +Apart from numerous remains, mostly of a problematical nature, +referred to the comprehensive group of the Sea-weeds, a large +number of Ferns have now been recognised, some being, of the +ordinary plant-like type (<i>Pecopteris, Neuropteris, Alethopteris, +Sphenopteris</i>, &c.), whilst others belong to the gigantic +group of the "Tree-ferns" (<i>Psaronius, Caulopteris</i>, &c.) +Besides these there is an abundant development of the singular extinct +types of the <i>Lepidodendroids</i>, the <i>Sigillarioids</i>, +and the <i>Calamites</i>, all of which attained their maximum in the +Carboniferous. Of these, the <i>Lepidodendra</i> may be regarded as +gigantic, tree-like Club-mosses (<i>LycopodiaceĹ“</i>); the +<i>Calamites</i> are equally gigantic Horse-tails +(<i>EquisetaceĹ“</i>); and the <i>Sigillarioids</i>, equally +huge in size, in some respects hold a position intermediate between +the Club-mosses and the Pines (Conifers). The Devonian rocks have + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 500px; + font-size: smaller; text-align: center;"> +<a name="page_137"><span class="page">Page 137</span></a> +GENERALIZED SECTION OF THE DEVONIAN ROCKS OF NORTH AMERICA. +<br /> +Fig. 76. +<br /> +<img src="images/fig076.jpg" width="482" height="544" alt="Fig. 76" /> +</span> +</span> + +also yielded traces of many other plants (such as <i>Annularia, +Asterophyllites, Cardiocarpon</i>, &c.), which acquire a +greater pre-dominance in the Carboniferous period, and which +will be spoken of in discussing the structure of the plants of the +Coal-measures. Upon the whole, the one plant which may be considered +as specially characteristic of the Devonian (though not confined +to this series) is the <i>Psilophyton</i> (fig. 77) of Dr Dawson. +These singular plants have slender branching stems, with sparse +needle-shaped leaves, the young stems being at first coiled up, +crosier-fashion, like the young fronds of ferns, whilst the old +branches carry numerous spore-cases. The +<a name="page_138"><span class="page">Page 138</span></a> +stems and branches seem +to have attained a height of two or three feet; and they sprang +from prostrate "root-stocks" or creeping stems. Upon the whole, + +<span style="float: left; margin: 4px; width: 228px; + font-size: smaller; text-align: justify;"> +<img src="images/fig077.jpg" width="220" height="638" alt="Fig. 77" /> +<br /> +Fig. 77.—Restoration of <i>Psilophyton princeps</i>. Devonian, +Canada. (After Dawson.) +</span> + +Principal Dawson is disposed to regard <i>Psilophyton</i> as +a "generalised type" of plants intermediate between the Ferns and +the Club-mosses. Lastly, the Devonian deposits have yielded the +remains of the first actual <i>trees</i> with which we are as yet +acquainted. About the nature of some of these (<i>Ormoxylon</i> +and <i>Dadoxylon</i>) no doubt can be entertained, since their +trunks not only show the concentric rings of growth characteristic +of exogenous trees in general, but their woody tissue exhibits +under the microscope the "discs" which are characteristic of +the wood of the Pines and Firs (see fig. 2). The singular genus +<i>Prototaxites</i>, however, which occurs in an older portion +of the Devonian series than the above, is not in an absolutely +unchallenged position. By Principal Dawson it is regarded as the +trunk of an ancient <i>Conifer</i>—the most ancient known; +but Mr Carruthers regards it as more probably the stem of a +gigantic sea-weed. The trunks of <i>Prototaxites</i> (fig. 78, A) +vary from one to three feet in diameter, and exhibit concentric +rings of growth; but its woody fibres have not hitherto been +clearly demonstrated to possess discs. Before leaving the Devonian +vegetation, it may be mentioned that the hornstone or chert so +abundant in the Corniferous limestone of North America has been +shown to contain the remains of various microscopic plants +(<i>Diatoms</i> and <i>Desmids</i>). We find also in the same +siliceous material the singular spherical bodies, with radiating +spines, which occur so abundantly in the chalk flints, and which +are termed <i>Xanthidia</i>. These may be regarded +<a name="page_139"><span class="page">Page 139</span></a> +as probably the spore-cases of the minute plants known as +<i>DesmidiĹ“</i>. +</p> + +<div class="center"> +<table border="0" width="563"> +<tr><td> +<img src="images/fig078.jpg" width="559" height="416" alt="Fig. 78" /> +</td></tr> +<tr><td class="left"><span class="image">Fig. 78.—A, Trunk +of <i>Prototaxites Logani</i>, eighteen inches in diameter, as seen +in the cliff near L'Anse Brehaut, GaspĂ©; B, Two wood-cells +showing spiral fibres and obscure pores, highly magnified. Lower +Devonian, Canada. (After Dawson) +</span></td></tr> +</table> +</div> + +<p class="indent"> +The Devonian <i>Protozoans</i> have still to be fully investigated. +True Sponges (such as <i>AstrtĹ“ospongia, SphĹ“rospongia</i>, +&c.) are not unknown; but by far the commonest representatives of +this sub-kingdom in the Devonian strata are <i>Stromatopora</i> and its +allies. These singular organisms (fig. 79) are not only very abundant +in some of the Devonian limestones—both in the Old World and +the New—but they often attain very large dimensions. However +much they may differ in minor details, the general structure of +these bodies is that of numerous, concentrically-arranged, thin, +calcareous laminæ, separated by narrow interspaces, which in +turn are crossed by numerous delicate vertical pillars, giving the +whole mass a cellular structure, and dividing it into innumerable +minute quadrangular compartments. Many of the Devonian +<i>StromatoporĹ“</i> also exhibit on their surface the rounded +openings of canals, which can hardly have served any other purpose +than that of permitting the sea-water to gain ready access to every +part of the organism. +</p> + +<p class="indent"> +No true <i>Graptolites</i> have ever been detected in strata of +<a name="page_140"><span class="page">Page 140</span></a> +of Devonian age; and the whole of this group has become +extinguished—unless we refer here the still surviving +<i>DictyonemĹ“</i>. The <i>CĹ“lenterates</i>, however, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 411px; + font-size: smaller; text-align: justify;"> +<img src="images/fig079.jpg" width="403" height="439" alt="Fig. 79" /> +<br /> +Fig. 79.—<i>a</i>, +Part of the under surface of <i>Stromatopora tuberculata</i>, +showing the wrinkled basement membrane and the openings of +water-canals, of the natural size; <i>b</i>, Portion of the +upper surface of the same, enlarged; <i>c</i>, Vertical section +of a fragment, magnified to show the internal structure. +Corniferous Limestone, Canada. (Original.) +</span> +</span> + +are represented by a vast number of <i>Corals</i>, of beautiful +forms and very varied types. The marbles of Devonshire, the +Devonian limestones of the Eifel and of France, and the calcareous +strata of the Corniferous and Hamilton groups of America, are +often replete with the skeletons of these organisms—so much so +as to sometimes entitle the rock to be considered as representing an +ancient coral-reef. In some instances the Corals have preserved +their primitive calcareous composition; and if they are embedded +in soft shales, they may weather out of the rock in almost all +their original perfection. In other cases, as in the marbles of +Devonshire, the matrix is so compact and crystalline that the +included corals can only be satisfactorily studied by means of +polished sections. In other cases, again, the corals have been +more or less completely converted into flint, as in the Corniferous +limestone of North America. When this is the case, they often come, +by the action of the weather, to stand out from +<a name="page_141"><span class="page">Page 141</span></a> +the enclosing +rock in the boldest relief, exhibiting to the observer the most +minute details of their organization. As before, the principal + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 312px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig080.jpg" width="304" height="690" alt="Fig. 80" /> +<br /> +Fig. 80.—<i>Cystiphyllum vesiculosum</i>, showing a +succession of cups produces by budding from the original coral. +Of the natural size. Devonian, America and Europe. (Original.) +</span> + +<span style="width: 281px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig081.jpg" width="273" height="508" alt="Fig. 81" /> +<br /> +Fig. 81—<i>Zaphrentis cornicula</i>, of the natural size. +Devonian, America. (Original.) +<br /> +<img src="images/fig082.jpg" width="260" height="135" alt="Fig. 82" /> +<br /> +Fig. 82—<i>Heliophyllum exiguum</i>, viewed from in front and +behind. Of the natural size. Devonian, Canada. (Original.) +</span> + +</span> + +representatives of the Corals are still referable to the groups of +the <i>Rugosa</i> and <i>Tabulata</i>. Amongst the Rugose group +we find a vast number of simple "cup-corals," generally known by +the quarrymen as "horns," from their shape. Of +<a name="page_142"><span class="page">Page 142</span></a> +the many forms of these, the species of <i>Cyathophyllum, +Heliophyllum</i> (fig. 82), <i>Zaphrentis</i> (fig. 81), and +<i>Cystiphyllum</i> (fig. 80), are perhaps those most abundantly +represented—none of these genera, however, except +<i>Heliophyllum</i>, being peculiar to the Devonian period. There +are also numerous compound Rugose corals, such as species of +<i>Eridophyllum, Diphyphyllum, Syringopora, +PhillipsastrĹ“a</i>, and some of the forms of +<i>Cyathophyllum</i> and <i>Crepidophyllum</i> (fig. 83). Some +of these compound corals attain a very large size, and form of + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 383px; + font-size: smaller; text-align: center;"> +<img src="images/fig083.jpg" width="375" height="329" alt="Fig. 83" /> +<br /> +Fig. 83.—Portion of a mass of <i>Crepidophyllum Archiaci</i>, +of the natural size. Hamilton Formation, Canada. (After +Billings.) +</span> +</span> + +themselves regular beds, which have an analogy, at any rate, with +existing coral-reefs, though there are grounds for believing that +these ancient types differed from the modern reef-builders in +being inhabitants of deep water. The "Tabulate Corals" are hardly +less abundant in the Devonian rocks than the <i>Rugosa</i>; and +being invariably compound, they hardly yield to the latter in the +dimensions of the aggregations which they sometimes form. +</p> + +<p class="indent"> +The commonest, and at the same time the largest, of these are +the "honeycomb corals," forming the genus <i>Favosites</i> (figs. +84, 85), which derive both their vernacular and their technical +names from their great likeness to masses of petrified honeycomb. +The most abundant species are <i>Favosites Gothlandica</i> and +<i>F. Hemispherica</i>, both here figured, which form masses +sometimes not less than two or three feet in diameter. Whilst +<i>Favosites</i> has acquired a popular name by its honey-combed +appearance, the resemblance of <i>Michelinia</i> to a fossilised +<a name="page_143"><span class="page">Page 143</span></a> +wasp's nest with the comb exposed is hardly less striking, and +has earned for it a similar recognition from the non-scientific + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 286px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig084.jpg" width="278" height="235" alt="Fig. 84" /> +<br /> +Fig. 84.—Portion of a mass of <i>Favosites Gothlandica</i>, +of the natural size. Upper Silurian and Devonian of Europe +and America. (Original.) Billings. +</span> + +<span style="width: 291px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig085.jpg" width="283" height="232" alt="Fig. 85" /> +<br /> +Fig. 85.—Fragment of <i>Favosites hemispherica</i>, of +the natural size. Upper Silurian and Devonian of America. +(After Billings.) +</span> + +</span> + +public. In addition to these, there are numerous branching or +plant-like Tabulate Corals, often of the most graceful form, +which are distinctive of the Devonian in all parts of the world. +</p> + +<p class="indent"> +The <i>Echinoderms</i> of the Devonian period call for little +special notice. Many of the Devonian limestones are "crinoidal;" +and the <i>Crinoids</i> are the most abundant and widely-distributed +representatives of their class in the deposits of this period. +</p> + +<p class="indent"> +The <i>Cystideans</i>, with doubtful exceptions, have not been +recognised in the Devonian; and their place is taken by the allied +group of the "Pentremites," which will be further spoken of as +occurring in the Carboniferous rocks. On the other hand, the +Star-fishes, Brittle-stars, and Sea-urchins are all continued +by types more or less closely allied to those of the preceding +Upper Silurian. +</p> + +<p class="indent"> +Of the remains of Ringed-worms (<i>Annelides</i>), the most numerous +and the most interesting are the calcareous envelopes of some +small tube-inhabiting species. No one who has visited the seaside +can have failed to notice the little spiral tubes of the existing +<i>Spirorbis</i> growing attached to shells, or covering the fronds +of the commoner Sea weeds (especially <i>Fucus serratus</i>). +These tubes are inhabited by a small Annelide, and structures of +a similar character occur not uncommonly from the Upper Silurian +upwards. In the Devonian rocks, <i>Spirorbis</i> is an extremely +common fossil, growing in hundreds attached to the outer surface +of corals and shells, and appearing +<a name="page_144"><span class="page">Page 144</span></a> +in many specific forms (figs. 86 and 87); but almost all the known + +<!-- Note: The book incorrectly numbered the next two plates as + Fig. 87 and Fig. 88. As a result there is no Fig. 86, but + there are two Fig. 88 illustrations. --> + +<span style="float: left; margin: 4px; width: 314px; + font-size: smaller; text-align: justify;"> +<img src="images/fig086.jpg" width="301" height="125" alt="Fig. 86" /> +<br /> +Fig. 87.—<i>a, Spirobois omphalodes</i>, natural size and +enlarged. Devonian, Europe and America; <i>b, Spirorbis +Arkonensis</i>, of the natural size and enlarged; <i>c</i>, The +same, with the tube twisted in the reverse direction. Devonian, +America. (Original.) +<br /> +<img src="images/fig087.jpg" width="306" height="102" alt="Fig. 87" /> +Fig. 88.—<i>a b, Spirorbis laxus</i>, enlarged, Upper +Silurian, America; <i>c, Spirorbis spinulifera</i>, of the +natural size and enlarged, Devonian, Canada. (After Hall and +the Author.) +</span> + +examples are of small size, and are liable to escape a cursory +examination. +</p> + +<p class="indent"> +The <i>Crustaceans</i> of the Devonian are principally +<i>Eurypterids</i> and <i>Trilobites</i>. Some of the former +attain gigantic dimensions, and the quarrymen in the Scotch Old +Red give them the name of "seraphim" from their singular scale-like +ornamentation. The <i>Trilobites</i>, though still sufficiently +abundant in some localites, have undergone a yet further diminution +since the close of the Upper Silurian. In both America and Europe +quite a number of generic types have survived from the Silurian, +but few or no new ones make their appearance during this period + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 528px; + font-size: smaller; text-align: justify;"> +<img src="images/fig088.jpg" width="520" height="317" alt="Fig. 88" /> +<br /> +Fig. 88.—Devonian Trilobites; <i>a, Phacops latifrons</i>, +Devonian of Britain, the Continent of Europe, and South America; +<i>b, Homalonotus armatus</i>, Europe; <i>c, Phacops +(Trimerocephalus) lĹ“vis</i>, Europe; <i>d</i>, Head-shield +of <i>Phacops (Portlockia) granulatus</i>, Europe. (After Salter +and Burmeister.) +</span> +</span> + +in either the Old World or the New. The <i>species</i>, however, +are distinct; and the +<a name="page_145"><span class="page">Page 145</span></a> +principal forms belong to the genera +<i>Phacops</i> (fig. 88, <i>a, c, d</i>), <i>Homalonotus</i> +(fig. 88, <i>b</i>), <i>Proetus</i>, and <i>Bronteus</i>. The +species figured above under the name of <i>Phacops latifrons</i> +(fig. 88, <i>a</i>), has an almost world-wide distribution, being +found in the Devonian of Britain, Belgium, France, Germany, Russia, +Spain, and South America; whilst its place is taken in North +America by the closely-allied <i>Phacops rana</i>. In addition +to the <i>Trilobites</i>, the Devonian deposits have yielded +the remains of a number of the minute <i>Ostracoda</i>, such as +<i>Entomis</i> ("<i>Cypridina</i>"), <i>Leperditia</i>, &c., +which sometimes occur in vast numbers, as in the so-called +"<i>Cypridina</i> Slates" of the German Devonian. There are also +a few forms of <i>Phyllopods</i> (<i>Estheria</i>). Taken as +a whole, the Crustacean fauna of the Devonian period presents +many alliances with that of the Upper Silurian, but has only +slight relationships with that of the Lower Carboniferous. +</p> + +<p class="indent"> +Besides <i>Crustaceans</i>, we meet here for the first time with +the remains of <i>air-breathing Articulates</i>, in the shape of +<i>Insects</i>. So far, these have only been obtained from the +Devonian rocks of North America, and they indicate the existence +of at least four generic types, all more or less allied to the +existing May-flies (<i>EphemeridĹ“</i>). One of these +interesting primitive insects, namely, <i>Platephemera antiqua</i> +(fig. 89), appears to have measured five inches in expanse of wing; + +<span style="float: left; margin: 4px; width: 273px; + font-size: smaller; text-align: center;"> +<img src="images/fig089.jpg" width="265" height="173" alt="Fig. 89" /> +<br /> +Fig. 89.—Wing of <i>Platephemera antiqua</i> Devonian, +America. (After Dawson.) +</span> + +and another (<i>Xelloneura antiquorum</i>) has attached to its wing +the remains of a "stridulating-organ" similar to that possessed +by the modern Grasshoppers—the instrument, as Principal +Dawson remarks, of "the first music of living things that Geology +as yet reveals to us." +</p> + +<p class="indent"> +Amongst the <i>Mollusca</i>, the Devonian rocks have yielded a +great number of the remains of Sea-mosses (<i>Polyzoa</i>). Some +of these belong to the ancient type <i>Ptilodictya</i>, which +seems to disappear here, or to the allied <i>Clathropora</i> +(fig. 90), with its fenestrated and reticulated fronds. We meet +also with the graceful and delicate stems of <i>Ceriopora</i> +(fig. 91). +</p> + +<p class="indent"> +The majority of the Devonian <i>Polyzoa</i> belong, however, +to the great and important Palæozoic group of the Lace-corals +(<i>Fenestella</i>, figs. 92 and 94, <i>Retepora</i>, fig. 93, +<i>Polypora</i>, and their allies). In all these forms there is +a horny skeleton, of a +<a name="page_146"><span class="page">Page 146</span></a> +fan-like or funnel-shaped form, which grew +attached by its base to some foreign body. The frond consists +of slightly-diverging or nearly parallel branches, which are + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 368px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig090.jpg" width="360" height="199" alt="Fig. 90" /> +<br /> +Fig. 90.—Fragment of <i>Clathropora intertexta</i>, of the +natural size and enlarged. Devonian, Canada. (Original.) +</span> + +<span style="width: 162px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig091.jpg" width="158" height="171" alt="Fig. 91" /> +<br /> +Fig. 91.—Fragment of <i>Ceriopora Hamiltonensis</i>, of the +natural size and enlarged. Devonian, Canada. (Original.) +</span> + +</span> + +either united by delicate cross-bars, or which bend alternately +from side to side, and become directly united with one another +at short intervals—in either case giving origin to numerous +oval or oblong perforations, which communicate to the whole + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 299px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig092.jpg" width="291" height="268" alt="Fig. 92" /> +<br /> +Fig. 92.—Fragment of <i>Fenestella magnifica</i>, of the +natural size and enlarged. Devonian, Canada. (Original.) +</span> + +<span style="width: 242px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig093.jpg" width="234" height="122" alt="Fig. 93" /> +<br /> +Fig. 93.—Fragment of <i>Retepora Phillipsi</i>, of the +natural size and enlarged. Devonian, Canada. (Original.) +<br /> +<img src="images/fig094.jpg" width="232" height="158" alt="Fig. 94" /> +<br /> +Fig. 94.—Fragment of <i>Fenestella cribrosa</i>, of the +natural size and enlarged. Dovonian, Canada. (Original.) +</span> + +</span> + +plant-like colony a characteristic netted and lace-like appearance. +On one of its surfaces—sometimes the internal, sometimes the +external—the frond carries a number of minute chambers or +<a name="page_147"><span class="page">Page 147</span></a> +"cells," which are generally borne in rows on the branches, and of +which each originally contained a minute animal. +</p> + +<p class="indent"> +The <i>Brachiopods</i> still continue to be represented in great +force through all the Devonian deposits, though not occurring in +the true Old Red Sandstone. Besides such old types as <i>Orthis, +Strophomena, Lingula, Athyris</i>, and <i>Rhynchonella</i>, we find +some entirely new ones; whilst various types which only commenced +their existence in the Upper Silurian, now undergo a great expansion +and development. This last is especially the case with the two +families of the <i>SpiriferidĹ“</i> and the +<i>ProduclidĹ“</i>. The <i>Spirifers</i>, in particular, are +especially characteristic of the Devonian, both in the Old and New +Worlds—some of the most typical forms, such as <i>Spirifera +mucronata</i> (fig. 96), having the shell "winged," or with the + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 161px; vertical-align:top; text-align: justify; + margin: 4px;"> +<img src="images/fig095.jpg" width="153" height="92" alt="Fig. 95" /> +<br /> +Fig. 95.—<i>Spirifera sculptilis</i>. Devonian, +Canada. (After Billings.) +</span> + +<span style="width: 383px; vertical-align:top; text-align: center; + margin: 4px;"> +<img src="images/fig096.jpg" width="375" height="112" alt="Fig. 96" /> +<br /> +Fig. 96.—<i>Spirifera mucronata</i>. Devonian, America. +(After Billings.) +</span> + +</span> + +lateral angles prolonged to such an extent as to have earned for +them the popular name of "fossil-butterflies." The closely-allied +<i>Spirifera disjunda</i> occurs in Britain, France, Spain, +Belgium, Germany, Russia, and China. The family of the +<i>ProductidĹ“</i> commenced to exist in the Upper Silurian, +in the genus <i>Chonetes</i>, and we shall hereafter find it +culminating in the Carboniferous in many forms of the great genus +<i>Producta</i>[17] itself. In the Devonian period, there is an +intermediate state of things, the genus <i>Chonetes</i> being +continued in new and varied types, and the Carboniferous +<i>ProdudĹ“</i> being represented by many forms of the +allied group <i>Productella</i>. Amongst other well-known +Devonian Brachiopods may be mentioned the two long-lived and +persistent types <i>Atrypa reticularis</i> (fig. 97) and +<i>Strophomena rhomboidalis</i> (fig. 98). The former of these +commences in the Upper Silurian, but is more abundantly developed +in the Devonian, having a geographical range that is nothing less +than world-wide; whilst the latter commences in the Lower Silurian, +<a name="page_148"><span class="page">Page 148</span></a> +and, with an almost equally cosmopolitan range, survives into the +Carboniferous period. +</p> + +<p class="footnote"> +[Footnote 17: The name of this genus is often written +<i>Productus</i>, just as <i>Spirifera</i> is often given in the +masculine gender as <i>Spirifer</i> (the name originally given +to it). The masculine termination to these names is, however, +grammatically incorrect, as the feminine noun <i>cochlea</i> +(shell) is in these cases <i>understood</i>.] +</p> + +<div class="center"> +<table border="0" width="489"> +<tr><td> +<img src="images/fig097.jpg" width="485" height="158" alt="Fig. 97" /> +</td></tr> +<tr><td class="center"><span class="image"> +Fig. 97.—<i>Atrypa reticularis</i>. Upper Silurian +and Devonian of Europe and America. (After Billings.) +</span></td></tr> +</table> +</div> + +<p class="indent"> +The Bivalves (<i>Lamellibranchiata</i>) of the Devonian call +for no special comment, the genera <i>Pterinea</i> and +<i>Megalodon</i> being, perhaps, the most noticeable. The Univalves + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 577px; + font-size: smaller; text-align: center;"> +<img src="images/fig098.jpg" width="569" height="171" alt="Fig. 98" /> +<br /> +Fig. 98.—<i>Strophomena rhomboidalis</i>. Lower Silurian, +Upper Silurian, and Devonian of Europe and America. +</span> +</span> + +(<i>Gasteropods</i>), also, need not be discussed in detail, +though many interesting forms of this group are known. The type +most abundantly represented, especially in America, is +<i>Platyceras</i> (fig. 99), comprising thin, wide-mouthed shells, + +<span style="float: left; margin: 4px; width: 291px; + font-size: smaller; text-align: justify;"> +<img src="images/fig099.jpg" width="283" height="173" alt="Fig. 99" /> +<br /> +Fig. 99.—Different views of <i>Platyceras dumosum</i>, of +the natural size. Devonian, Canada. (Original.) +</span> + +probably most nearly allied to the existing "Bonnet-limpets," +and sometimes attaining very considerable dimensions. We may +also note the continuance of the genus <i>Euomphalus</i>, with +its discoidal spiral shell. Amongst the <i>Heteropods</i>, the +survival of <i>Bellerophon</i> is to be recorded; and in the +"Winged-snails," or <i>Pteropods</i>, we find new forms of the +old genera <i>Tentaculites</i> and <i>Conularia</i> +<a name="page_149"><span class="page">Page 149</span></a> +(fig. 100). The latter, with its fragile, conical, and often +beautifully ornamented shell, is especially noticeable. +</p> + +<p class="indent"> +The remains of <i>Cephalopoda</i> are far from uncommon in the + +<span style="float: right; margin: 4px; width: 179px; + font-size: smaller; text-align: justify;"> +<img src="images/fig100.jpg" width="171" height="273" alt="Fig. 100" /> +<br /> +Fig. 100.—<i>Conularia ornata</i>, of the natural size. +Devonian, Europe. +</span> + +Devonian deposits, all the known forms being still Tetrabranchiate. +Besides the ancient types <i>Orthoceras</i> and <i>Cyrtoceras</i>, +we have now a predominance of the spirally-coiled chambered shells +of <i>Goniatites</i> and <i>Clymenia</i>. In the former of these the +shell is shaped like that of the <i>Nautilus</i>; but the partitions +between the chambers ("septa") are more or less lobed, folded, or +angulated, and the "siphuncle" runs along the <i>back</i> or convex +side of the shell—these being characters which approximate +<i>Goniatites</i> to the true Ammonites of the later rocks. In +<i>Clymenia</i>, on the other hand, whilst the shell (fig. 101) +is coiled into a flat spiral, and the partitions or septa are +simple or only slightly lobed, there is still this difference, as +compared with the <i>Nautilus</i>, that the tube of the siphuncle +is placed on the <i>inner</i> or concave side of the shell. The + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 322px; + font-size: smaller; text-align: center;"> +<img src="images/fig101.jpg" width="314" height="388" alt="Fig. 101" /> +<br /> +Fig. 101.—<i>Clymenia Sedgwickii</i>. Devonian, Europe. +</span> +</span> + +species of <i>Clymenia</i> are exclusively Devonian in +<a name="page_150"><span class="page">Page 150</span></a> +their range; and some of the limestones of this period in Germany +are so richly charged with fossils of this genus as to have received +the name of "Clymenien-kalk." +</p> + +<p class="indent"> +The sub-kingdom of the <i>Vertebrates</i> is still represented +by <i>Fishes</i> only; but these are so abundant, and belong to +such varied types, that the Devonian period has been appropriately +called the "Age of Fishes." Amongst the existing fishes there are +three great groups which are of special geological importance, +as being more or less extensively represented in past time. These +groups are: (1) The <i>Bony Fishes</i> (<i>Teleostei</i>), comprising +most existing fishes, in which the skeleton is more or less +completely converted into bone; the tail is symmetrically lobed +or divided into equal moieties; and the scales are usually thin, +horny, flexible plates, which overlap one another to a greater +or less extent. (2) The <i>Ganoid Fishes</i> (<i>Ganoidei</i>), +comprising the modern Gar-pikes, Sturgeons, &c., in which the +skeleton usually more or less completely retains its primitive +soft and cartilaginous condition; the tail is generally markedly +unsymmetrical, being divided into two unequal lobes; and the +scales (when present) have the form of plates of bone, usually +covered by a layer of shining enamel. These scales may overlap; +or they may be rhomboidal plates, placed edge to edge in oblique +rows; or they have the form of large-sized bony plates, which +are commonly united in the region of the head to form a regular +buckler. (3) The <i>Placoid Fishes</i>, or <i>Elasmobranchii</i>, +comprising the Sharks, Rays, and <i>ChimĹ“rĹ“</i> of +the present day, in which the skeleton is cartilaginous; the tail +is unsymmetrically lobed; and the scales have the form of detached +bony plates of variable size, scattered in the integument. +</p> + +<p class="indent"> +It is to the two last of these groups that the Devonian fishes +belong, and they are more specially referable to the <i>Ganoids</i>. +The order of the Ganoid fishes at the present day comprises but +some seven or eight genera, the species of which principally or +exclusively inhabit fresh waters, and all of which are confined +to the northern hemisphere. As compared, therefore, with the Bony +fishes, which constitute the great majority of existing forms, +the Ganoids form but an extremely small and limited group. It was +far otherwise, however, in Devonian times. At this period, the +bony fishes are not known to have come into existence at all, and +the Ganoids held almost undisputed possession of the waters. To +what extent the Devonian Ganoids were confined to fresh waters +remains yet to be proved; and that many of them lived in the sea +is certain. It was formerly supposed that the Old Red Sandstone +of Scotland and Ireland, with its abundant fish-remains, might +perhaps be a fresh-water deposit, since the habitat of its fishes is +<a name="page_151"><span class="page">Page 151</span></a> +uncertain, and it contains no indubitable +marine fossils. It has been now shown, however, that the marine +Devonian strata of Devonshire and the continent of Europe contain +some of the most characteristic of the Old Red Sandstone fishes of +Scotland; whilst the undoubted marine deposit of the Corniferous +limestone of North America contains numerous shark-like and Ganoid +fishes, including such a characteristic Old Red genus as +<i>Coccosleus</i>. There can be little doubt, therefore, but that +the majority of the Devonian fishes were truly marine in their +habits, though it is probable that many of them lived in shallow +water, in the immediate neighbourhood of the shore, or in +estuaries. +</p> + +<p class="indent"> +The Devonian Galloids belong to a number of groups; and it is + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 561px; + font-size: smaller; text-align: justify;"> +<img src="images/fig102.jpg" width="553" height="544" alt="Fig. 102" /> +<br /> +Fig. 102.—Fishes of the Devonian rocks of America. <i>a</i>, +Diagram of the jaws and teeth of <i>Dinichthys Hertzeri</i>, +viewed from the front, and greatly reduced; <i>b</i>, Diagram +of the skull of <i>Macropetalichthys Sullivanti</i>, reduced +in size; <i>c</i>, A portion of the enamelled surface of the +skull of the same, magnified; <i>d</i>, One of the scales of +<i>Onychodus sigmoides</i>, of the natural size; <i>e</i>, One +of the front teeth of the lower jaw of the same, of the natural +size: <i>f</i>, Fin-spine of <i>MachĹ“racanthus major</i>, +a shark-like fish, reduced in size. (After Newberry.)] +</span> +</span> + +only possible to notice a few of the most important forms here. +The modern group of the Sturgeons is represented, +<a name="page_152"><span class="page">Page 152</span></a> +more or less +remotely, by a few Devonian fishes—such as <i>Asterosteus</i>; +and the great <i>Macropetalichthys</i> of the Corniferous limestone +of North America is believed by Newberry to belong to this group. +In this fish (fig. 102, <i>b</i>) the skull was of large size, +its outer surface being covered with a tuberculated enamel; and, +as in the existing Sturgeons, the mouth seems to have been wholly +destitute of teeth. Somewhat allied, also, to the Sturgeons, is a +singular group of armoured fishes, which is highly characteristic +of the Devonian of Britain and Europe, and less so of that of +America. In these curious forms the head and front extremity of +the body were protected by a buckler composed of large enamelled +plates, more or less firmly united to one another; whilst the +hinder end of the body was naked, or was protected with small +scales. Some forms of this group—such as <i>Pteraspis</i> +and <i>Coccosteus</i>—date from the Upper Silurian; but +they attain their maximum in the Devonian, and none of them are +known to pass upwards into the overlying Carboniferous rocks. +Amongst the most characteristic forms of this group may be +mentioned <i>Cephalaspis</i> (fig. 103) and <i>Pterichthys</i> +(fig. 104). In the former of these the head-shield is of a + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 516px; + font-size: smaller; text-align: justify;"> +<img src="images/fig103.jpg" width="508" height="219" alt="Fig. 103" /> +<br /> +Fig. 103.—<i>Cephalaspis Lyellii</i>. Old Red Sandstone, +Scotland. (After Page.) +</span> +</span> + +crescentic shape, having its hinder angles produced backwards +into long "horns," giving it the shape of a "saddler's knife." +No teeth have been discovered; but the body was covered with small +ganoid scales, and there was an unsymmetrical tail-fin. In +<i>Pterichthys</i>—which, like the preceding, was first +brought to light by the labours of Hugh Miller—the whole +of the head and the front part of the body were defended by a +buckler of firmly-united enamelled plates, whilst the rest of the +body was covered with small scales. The form of the "pectoral +fins" was quite unique—these having the shape of two long, +curved spines, somewhat like wings, covered by finely-tuberculated +ganoid plates. All the preceding forms +<a name="page_153"><span class="page">Page 153</span></a> +of this group are of small size; but few fishes, living or extinct, +could rival the proportions of the great <i>Dinichthys</i>, referred + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 490px; + font-size: smaller; text-align: center;"> +<img src="images/fig104.jpg" width="482" height="189" alt="Fig. 104" /> +<br /> +Fig. 104.—<i>Pterichthys cornutus</i>. Old Red Sandstone, +Scotland. (After Agassiz.) +</span> +</span> + +to this family by Newberry. In this huge fish (fig. 102, <i>a</i>) +the head alone is over three feet in length, and the body is +supposed to have been twenty-five or thirty feet long. The head +was protected by a massive cuirass of bony plates firmly articulated +together, but the hinder end of the body seems to have been simply +enveloped in a leathery skin. The teeth are of the most formidable +description, consisting in both jaws of serrated dental plates +behind, and in front of enormous conical tusks (fig. 102, <i>a</i>). +Though immensely larger, the teeth of <i>Dinichthys</i> present +a curious resemblance to those of the existing Mud-fishes +(<i>Lepidosiren</i>). +</p> + +<p class="indent"> +In another great group of Devonian Ganoids, we meet with fishes +more or less closely allied to the living <i>Polypteri</i> (fig. +105) of the Nile and Senegal. In this group (fig. 106) the pectoral +fins consist of a central scaly lobe carrying the fin-rays on both +sides, the scales being sometimes rounded and overlapping (fig. +106), or more commonly rhomboidal and placed edge to edge (fig. 105, +A). Numerous forms of these "Fringe-finned" Ganoids occur in the +Devonian strata, such as <i>Holoptychius, GlyotolĹ“mus, +Osteolepis, Phaneropleuron</i>, &c. To this group is also to be +ascribed the huge <i>Onychodus</i> (fig. 102, <i>d</i> and <i>e</i>), +with its large, rounded, overlapping scales, an inch in diameter, +and its powerful pointed teeth. It is to be remembered, however, +that some of these "Fringe-finned" Ganoids are probably referable +to the small but singular group of the "Mud-fishes" (<i>Dipnoi</i>), +represented at the present day by the singular <i>Lepidosiren</i> +of South America and Africa, and the <i>Ceratodus</i> of the +rivers of Queensland. +</p> + +<p class="indent"> +Leaving the Ganoid fishes, it still remains to be noticed that +the Devonian deposits have yielded the remains of a number of +fishes more or less closely allied to the existing Sharks, +<a name="page_154"><span class="page">Page 154</span></a> +Rays, and <i>ChimĹ“rĹ“</i> (the <i>Elasmobranchii</i>). +The majority of the forms here alluded to are allied not to the true +Sharks and Dog-fishes, but to the more peaceable "Port Jackson + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 564px; + font-size: smaller; text-align: center;"> +<img src="images/fig105.jpg" width="556" height="316" alt="Fig. 105" /> +<br /> +Fig. 105.—A, <i>Polypterus</i>, a recent Ganoid fish; B, +<i>Osteolepis</i>, a Devonian Ganoid; <i>a a</i>, Pectoral +fins, showing the fin-rays arranged round a central lobe. +</span> +</span> + +Sharks," with their blunt teeth, adapted for crushing the shells of +Molluscs. The collective name of "Cestracionts" is applied to these; +and we have evidence of their past existence in the Devonian seas + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 518px; + font-size: smaller; text-align: center;"> +<img src="images/fig106.jpg" width="510" height="243" alt="Fig. 106" /> +<br /> +Fig. 106.—<i>Holoptychius nobilissimus</i>, restored. Old +Red Sandstone, Scotland. A, Scale of the same. +</span> +</span> + +both by their teeth, and by the defensive spines which were implanted +in front of a greater or less number of the fins. These are bony +spines, often variously grooved, serrated, or ornamented, with +hollow bases, implanted in the integument, and capable of being +erected or depressed at will. +<a name="page_155"><span class="page">Page 155</span></a> +Many of these "fin-spines" have been preserved to us in the fossil +condition, and the Devonian rocks have yielded examples belonging to +many genera. As some of the true Sharks and Dog-fishes, some of the +Ganoids, and even some Bony Fishes, possess similar defences, it is +often a matter of some uncertainty to what group a given spine is +to be referred. One of these spines, belonging to the genus +<i>MachĹ“racanthus</i>, from the Devonian rocks of America, has +been figured in a previous illustration (fig. 102, <i>f</i>). +</p> + +<p class="indent"> +In conclusion, a very few words may be said as to the validity of +the Devonian series as an independent system of rocks, preserving +in its successive strata the record of an independent system +of life. Some high authorities have been inclined to the view +that the Devonian formation has in nature no actual existence, +but that it is made up partly of beds which should be referred +to the summit of the Upper Silurian, and partly of beds which +properly belong to the base of the Carboniferous. This view seems +to have been arrived at in consequence of a too exclusive study +of the Devonian series of the British Isles, where the physical +succession is not wholly clear, and where there is a striking +discrepancy between the organic remains of those two members +of the series which are known as the "Old Red Sandstone" and +the "Devonian" rocks proper. This discrepancy, however, is not +complete; and, as we have seen, can be readily explained on the +supposition that the one group of rocks presents us with the +shallow water and littoral deposits of the period, while in the +other we are introduced to the deep-sea accumulations of the +same period. Nor can the problem at issue be solved by an appeal +to the phenomena of the British area alone, be the testimony of +these what it may. As a matter of fact, there is at present no +sufficient ground for believing that there is any irreconcilable +discordance between the succession of rocks and of life in Britain +during the period which elapsed between the deposition of the +Upper Ludlow and the formation of the Carboniferous Limestone, +and the order of the same phenomena during the same period in +other regions. Some of the Devonian types of life, as is the +case with all great formations, have descended unchanged from +older types; others pass upwards unchanged to the succeeding +period: but the fauna and flora of the Devonian period are, as +a whole, quite distinct from those of the preceding Silurian or +the succeeding Carboniferous; and they correspond to an equally +distinct rock-system, which in point of time holds an intermediate +position between the two great groups just mentioned. As +<a name="page_156"><span class="page">Page 156</span></a> +before remarked, this conclusion may be regarded +as sufficiently proved even by the phenomena of the British area; +but it maybe said to be rendered a certainty by the study of the +Devonian deposits of the continent of Europe—or, still more, +by the investigation of the vast, for the most part uninterrupted +and continuous series of sediments which commenced to be laid +down in North America at the beginning of the Upper Silurian, +and did not cease till, at any rate, the close of the Carboniferous. +</p> + +<h4>LITERATURE.</h4> + +<p class="indent"> +The following list comprises the more important works and memoirs +to which the student of Devonian rocks and fossils may +refer:— +</p> + +<table border="0" cellspacing="0"> +<tr><td class="right" valign="top">(1)</td> + <td>'Siluria.' Sir Roderick Murchison.</td></tr> +<tr><td class="right" valign="top">(2)</td> + <td>'Geology of Russia in Europe.' Murchison (together with De + Verneuil and Count von Keyserling).</td></tr> +<tr><td class="right" valign="top">(3)</td> + <td>"Classification of the Older Rocks of Devon and + Cornwall"—'Proc. Geol. Soc.,' vol. iii., 1839. Sedgwick + and Murchison.</td></tr> +<tr><td class="right" valign="top">(4)</td> + <td>"On the Physical Structure of Devonshire;" and on the + "Classification of the Older Stratified Rocks of Devonshire + and Cornwall"—'Trans. Geol. Soc.,' vol. v., 1840. Sedgwick + and Murchison.</td></tr> +<tr><td class="right" valign="top">(5)</td> + <td>"On the Distribution and Classification of the Older or + Palæozoic Rocks of North Germany and Belgium"—'Geol. + Trans.,' 2d ser., vol. vi., 1842. Sedgwick and + Murchison.</td></tr> +<tr><td class="right" valign="top">(6)</td> + <td>'Report on the Geology of Cornwall, Devon, and West + Somerset.' De la Beche.</td></tr> +<tr><td class="right" valign="top">(7)</td> + <td>'Memoirs of the Geological Survey of Ireland and + Scotland.' Jukes and Geikie.</td></tr> +<tr><td class="right" valign="top">(8)</td> + <td>"On the Carboniferous Slate (or Devonian Rocks) and the + Old Red Sandstone of South Ireland and North Devon"—'Quart. + Journ. Geol. Soc.,' vol. xxii. Jukes.</td></tr> +<tr><td class="right" valign="top">(9)</td> + <td>"On the Physical Structure of West Somerset and North + Devon;" and on the "Palæontological Value of Devonian + Fossils"—'Quart. Journ. Geol. Soc.,' vol. iii. + Etheridge.</td></tr> +<tr><td class="right" valign="top">(10)</td> + <td>"On the Connection of the Lower, Middle, and Upper Old Red + Sandstone of Scotland"—'Trans. Edin. Geol. Soc.,' vol. i. + part ii. Powrie.</td></tr> +<tr><td class="right" valign="top">(11)</td> + <td>'The Old Red Sandstone,' 'The Testimony of the Rocks,' and + 'Footprints of the Creator.' Hugh Miller.</td></tr> +<tr><td class="right" valign="top">(12)</td> + <td>"Report on the 4th Geological District"—'Geology of + New York,' vol. iv. James Hall.</td></tr> +<tr><td class="right" valign="top">(13)</td> + <td>'Geology of Canada,' 1863. Sir W. E. Logan.</td></tr> +<tr><td class="right" valign="top">(14)</td> + <td>'Acadian Geology.' Dawson.</td></tr> +<tr><td class="right" valign="top">(15)</td> + <td>'Manual of Geology.' Dana.</td></tr> +<tr><td class="right" valign="top">(16)</td> + <td>'Geological Survey of Ohio,' vol. i.</td></tr> +<tr><td class="right" valign="top">(17)</td> + <td>'Geological Survey of Illinois,' vol. i.</td></tr> +<tr><td class="right" valign="top">(18)</td> + <td>'Palæozoic Fossils of Cornwall, Devon, and West + Somerset.' Phillips.</td></tr> +<tr><td class="right" valign="top">(19)</td> + <td>'Recherches sur les Poissons Fossiles.' Agassiz.</td></tr> +<tr><td class="right" valign="top">(20)</td> + <td>'Poissous de l'Old Red.' Agassiz.</td></tr> +<tr><td class="right" valign="top">(21)</td> + <td>"On the Classification of Devonian Fishes"—'Mem. Geol. + Survey of Great Britain,' Decade X. Huxley.</td></tr> +<tr><td class="right" valign="top"> +<a name="page_157"><span class="page">Page 157</span></a> + (22)</td> + <td>'Monograph of the Fishes of the Old Red Sandstone of + Britain' (Palæontographical Society). Powrie and + Lankester.</td></tr> +<tr><td class="right" valign="top">(23)</td> + <td>'Fishes of the Devonian System, Palæontology of + Ohio.' Newberry.</td></tr> +<tr><td class="right" valign="top">(24)</td> + <td>'Monograph of British Trilobites' + (Palæontographical Society); Salter.</td></tr> +<tr><td class="right" valign="top">(25)</td> + <td>'Monograph of British Merostomata' + (Palæontographical Society). Henry Woodward.</td></tr> +<tr><td class="right" valign="top">(26)</td> + <td>'Monograph of British Brachiopoda' + (Palæontographical Society). Davidson.</td></tr> +<tr><td class="right" valign="top">(27)</td> + <td>'Monograph of British Fossil Corals' + (Palæontographical Society). Milne-Edwards and + Haime.</td></tr> +<tr><td class="right" valign="top">(28)</td> + <td>'Polypiers Foss. des Terrains PalĂ©ozoiques.' + Milne-Edwards and Jules Haime.</td></tr> +<tr><td class="right" valign="top">(29)</td> + <td>"Devonian Fossils of Canada West"—'Canadian + Journal,' new ser., vols. iv.-vi. Billings.</td></tr> +<tr><td class="right" valign="top">(30)</td> + <td>'Palæontology of New York,' vol. iv. James + Hall.</td></tr> +<tr><td class="right" valign="top">(31)</td> + <td>'Thirteenth, Fifteenth, and Twenty-third Annual Reports + on the State Cabinet.' James Hall.</td></tr> +<tr><td class="right" valign="top">(32)</td> + <td>'Palæozoic Fossils of Canada,' vol. ii. + Billings.</td></tr> +<tr><td class="right" valign="top">(33)</td> + <td>'Reports on the Palæontology of the Province of + Ontario for 1874 and 1875.' Nicholson.</td></tr> +<tr><td class="right" valign="top">(34)</td> + <td>"The Fossil Plants of the Devonian and Upper Silurian + Formations of Canada"—'Geol. Survey of Canada.' + Dawson.</td></tr> +<tr><td class="right" valign="top">(35)</td> + <td>'Petrefacta Germaniæ.' Goldfuss.</td></tr> +<tr><td class="right" valign="top">(36)</td> + <td>'Versteinerungen der Grauwacken-formation.' &c. + Geinitz.</td></tr> +<tr><td class="right" valign="top">(37)</td> + <td>'Beitrag zur Palæontologie des Thüringer-Waldes.' + Richter and Unger.</td></tr> +<tr><td class="right" valign="top">(38)</td> + <td>'Ueber die Placodermen der Devonischen System.' + Pander.</td></tr> +<tr><td class="right" valign="top">(39)</td> + <td>'Die Gattungen der Fossilen Pflanzen.' + GĹ“ppert.</td></tr> +<tr><td class="right" valign="top">(40)</td> + <td>'Genera et Species Plantarum Fossilium.' + Unger.</td></tr> +</table> + +<h3>CHAPTER XII.</h3> + +<p class="subtitle"> +THE CARBONIFEROUS PERIOD. +</p> + +<p class="indent"> +Overlying the Devonian formation is the great and important series +of the <i>Carboniferous Rocks</i>, so called because workable +beds of coal are more commonly and more largely developed in this +formation than in any other. Workable coal-seams, however, occur +in various other formations (Jurassic, Cretaceous, Tertiary), so +that coal is not an exclusively Carboniferous product; whilst +even in the Coal-measures themselves the coal bears but a very +small proportion to the total thickness of strata, occurring +only in comparatively thin beds intercalated in a great series +of sandstones, shales, and other genuine aqueous sediments. +</p> + +<p class="indent"> +<a name="page_158"><span class="page">Page 158</span></a> +Stratigraphically, the Carboniferous rocks usually repose +conformably upon the highest Devonian beds, so that the line of +demarcation between the Carboniferous and Devonian formations is +principally a palæontological one, founded on the observed +differences in the fossils of the two groups. On the other hand, +the close of the Carboniferous period seems to have been generally, +though not universally, signalised by movements of the crust of +the earth, so that the succeeding Permian beds often lie +unconformably upon the Carboniferous sediments. +</p> + +<p class="indent"> +Strata of Carboniferous age have been discovered in almost every +large land-area which has been sufficiently investigated; but +they are especially largely developed in Britain, in various +parts of the continent of Europe, and in North America. Their +general composition, however, is, comparatively speaking, so +uniform, that it will suffice to take a comprehensive view of +the formation without considering any one area in detail, though +in each region the subdivisions of the formation are known by +distinctive local names. Taking such a comprehensive view, it +is found that the Carboniferous series is generally divisible +into a <i>Lower</i> and essentially calcareous group (the +"Sub-Carboniferous" or "Carboniferous Limestone"); a <i>Middle</i> +and principally arenaceous group (the "Millstone Grit"); and an +Upper group, of alternating shales and sandstones, with workable +seams of coal (the "Coal-measures"). +</p> + +<p class="indent"> +I. The <i>Carboniferous, Sub-Carboniferous</i>, or <i>Mountain +Limestone Series</i> constitutes the general base of the +Carboniferous system. As typically developed in Britain, the +Carboniferous Limestone is essentially a calcareous formation, +sometimes consisting of a mass of nearly pure limestone from +1000 to 2000 feet in thickness, or at other times of successive +great beds of limestone with subordinate sandstones and shales. +In the north of England the base of the series consists of pebbly +conglomerates and coarse sandstones; and in Scotland generally, +the group is composed of massive sandstones with a comparatively +feeble development of the calcareous element. In Ireland, again, +the base of the Carboniferous Limestone is usually considered +to be formed by a locally-developed group of grits and shales +(the "Coomhola Grits" and "Carboniferous Slate"), which attain +the thickness of about 5000 feet, and contain an intermixture +of Devonian with Carboniferous types of fossils. Seeing that the +Devonian formation is generally conformable to the Carboniferous, +we need feel no surprise at this intermixture of forms; nor does it +<a name="page_159"><span class="page">Page 159</span></a> +appear to be of great moment whether these +strata be referred to the former or to the latter series. Perhaps +the most satisfactory course is to regard the Coomhola Grits and +Carboniferous Slates as "passage-beds" between the Devonian and +Carboniferous; but any view that may be taken as to the position +of these beds, really leaves unaffected the integrity of the +Devonian series as a distinct life-system, which, on the whole, +is more closely allied to the Silurian than to the Carboniferous. +In North America, lastly, the Sub-Carboniferous series is never +purely calcareous, though in the interior of the continent it +becomes mainly so. In other regions, however, it consists +principally of shales and sandstones, with subordinate beds of +limestone, and sometimes with this beds of coal or deposits of +clay-ironstone. +</p> + +<p class="indent"> +II. <i>The Millstone Grit</i>.—The highest beds of the +Carboniferous Limestone series are succeeded, generally with +perfect conformity, by a series of arenaceous beds, usually +known as the <i>Millstone Grit</i>. As typically developed in +Britain, this group consists of hard quartzose sandstones, +often so large-grained and coarse in texture as to properly +constitute fine conglomerates. In other cases there are regular +conglomerates, sometimes with shales, limestones, and thin beds +of coal—the thickness of the whole series, when well +developed, varying from 1000 to 5000 feet. In North America, +the Millstone Grit rarely reaches 1000 feet in thickness; and, +like its British equivalent, consists of coarse sandstones +and grits, sometimes with regular conglomerates. Whilst the +Carboniferous Limestone was undoubtedly deposited in a tranquil +ocean of considerable depth, the coarse mechanical sediments +of the Millstone Grit indicate the progressive shallowing of +the Carboniferous seas, and the consequent supervention of +shore-conditions. +</p> + +<p class="indent"> +III. <i>The Coal-measures</i>.—The Coal-measures properly +so called rest conformably upon the Millstone Grit, and usually +consist of a vast series of sandstones, shales, grits, and coals, +sometimes with beds of limestone, attaining in some regions a total +thickness of from 7000 to nearly 14,000 feet. Beds of workable +coal are by no means unknown in some areas in the inferior group +of the Sub-Carboniferous; but the general statement is true, +that coal is mostly obtained from the true Coal-measures—the +largest known, and at present most productive coal-fields of the +world being in Great Britain, North America, and Belgium. Wherever +they are found, with limited exceptions, the Coal-measures present +a singular <i>general</i> uniformity of mineral composition. They +<a name="page_160"><span class="page">Page 160</span></a> +consist, namely, of an indefinite alternation +of beds of sandstone, shale, and coal, sometimes with bands of +clay-ironstone or beds of limestone, repeated in no constant +order, but sometimes attaining the enormous aggregate thickness +of 14,000 feet, or little short of 3 miles. The beds of coal +differ in number and thickness in different areas, but they +seldom or never exceed one-fiftieth part of the total bulk of +the formation in thickness. The characters of the coal itself, +and the way in which the coal-beds were deposited, will be +briefly alluded to in speaking of the vegetable life of the +period. In Britain, and in the Old World generally, the +Coal-measures are composed partly of genuine terrestrial +deposits—such as the coal—and partly of sediments +accumulated in the fresh or brackish waters of vast lagoons, +estuaries, and marshes. The fossils of the Coal-measures in these +regions are therefore necessarily the remains either of terrestrial +plants and animals, or of such forms of life as inhabit fresh or +brackish waters, the occurrence of strata with marine fossils being +quite a local and occasional phenomenon. In various parts of North +America, on the other hand, the Coal-measures, in addition to +sandstones, shales, coal-seams, and bands of clay-ironstone, +commonly include beds of limestone, charged with marine remains, +and indicating marine conditions. The subjoined section (fig. 107) +gives, in a generalised form, the succession of the Carboniferous +strata in such a British area as the north of England, where +the series is developed in a typical form. +</p> + +<p class="indent"> +As regards the <i>life</i> of the Carboniferous period, we naturally +find, as has been previously noticed, great differences in different +parts of the entire series, corresponding to the different mode of +origin of the beds. Speaking generally, the Lower Carboniferous +(or the Sub-Carboniferous) is characterised by the remains of +marine animals; whilst the Upper Carboniferous (or Coal-measures) +is characterised by the remains of plants and terrestrial animals. +In all those cases, however, in which marine beds are found in +the series of the Coal-measures, as is common in America, then +we find that the fossils agree in their general characters with +those of the older marine deposits of the period. +</p> + +<p class="indent"> +Owing to the fact that coal is simply compressed and otherwise +altered vegetable matter, and that it is of the highest economic +value to man, the Coal-measures have been more thoroughly explored +than any other group of strata of equivalent thickness in the +entire geological series. Hence we have already a very extensive +acquaintance with the <i>plants</i> of the Carboniferous period; +and our knowledge on this subject is +<a name="page_161"><span class="page">Page 161</span></a> +daily undergoing increase. It is not to be supposed, however, +that the remains of plants are found solely in Coal-measures; + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 544px; + font-size: smaller; text-align: center;"> +GENERALIZED SECTION OF THE CARBONIFEROUS STRATA OF THE NORTH OF +ENGLAND. +<br /> +Fig. 107. +<br /> +<img src="images/fig107.jpg" width="536" height="697" alt="Fig. 107" /> +</span> +</span> + +for though most abundant towards the summit, they are found in +less numbers in all parts of the series. Wherever found, they +belong to the same great types of +<a name="page_162"><span class="page">Page 162</span></a> +vegetation; but, before reviewing these, a few words must be +said as to the origin and mode of formation of <i>coal</i>. +</p> + +<p class="indent"> +The coal-beds, as before mentioned, occur interstratified with +shales, sandstones, and sometimes limestones; and there may, +within the limits of a single coal-field, be as many as 80 or +100 of such beds, placed one above the other at different levels, +and varying in thickness from a few inches up to 20 or 30 feet. +As a general rule, each bed of coal rests upon a bed of shale or +clay, which is termed the "under-clay," and in which are found +numerous roots of plants; whilst the strata immediately on the +top of the coal may be shaly or sandy, but in either case are +generally charged with the leaves and stems of plants, and often +have upright trunks passing vertically through them. When we +add to this that the coal itself is, chemically, nearly wholly +composed of carbon, and that its microscopic structure shows it +to be composed almost entirely of fragments of stems, leaves, +bark, seeds, and vegetable <i>dĂ©bris</i> derived from +<i>land-plants</i>, we are readily enabled to understand how the +coal was formed. The "<i>under-clay</i>" immediately beneath the +coal-bed represents an old land-surface—sometimes, perhaps, +the bottom of a swamp or marsh, covered with a luxuriant vegetation; +the <i>coal bed</i> itself represents the slow accumulation, +through long periods, of the leaves, seeds, fruits, stems, and +fallen trunks of this vegetation, now hardened and compressed +into a fraction of its original bulk by the pressure of the +superincumbent rocks; and the strata of sand or shale above the +coal-bed—the so-called "roof" of the coal—represent +sediments quietly deposited as the land, after a long period of +repose, commenced to sink beneath the sea. On this view, the rank +and long-continued vegetation which gave rise to each coal-bed +was ultimately terminated by a slow depression of the surface on +which the plants grew. The land-surface then became covered by +the water, and aqueous sediments were accumulated to a greater +or less thickness upon the dense mass of decaying vegetation +below, enveloping any trunks of trees which might still be in an +erect position, and preserving between their layers the leaves +and branches of plants brought down from the neighbouring land by +streams, or blown into the wafer by the wind. Finally, there set in +a slow movement of elevation,—the old land again reappeared +above the water; a new and equally luxuriant vegetation flourished +upon the new land-surface; and another coal-bed was accumulated, +to be preserved ultimately in a similar fashion. Some few beds +of coal may have been formed by drifted vegetable matter brought +down into the ocean by rivers, +<a name="page_163"><span class="page">Page 163</span></a> +and deposited +directly on the bottom of the sea; but in the majority of cases +the coal is undeniably the result of the slow growth and decay of +plants <i>in situ</i>: and as the plants of the coal are not +<i>marine</i> plants, it is necessary to adopt some such theory +as the above to account for the formation of coal-seams. By this +theory, as is obvious, we are compelled to suppose that the vast +alluvial and marshy flats upon which the coal-plants grew were +liable to constantly-recurring oscillations of level, the successive +land-surfaces represented by the successive coal-beds of any +coal-field being thus successively buried beneath accumulations +of mud or sand. We have no need, however, to suppose that these +oscillations affected large areas at the same time; and geology +teaches us that local elevations and depressions of the land +have been matters of constant occurrence throughout the whole +of past time. +</p> + +<p class="indent"> +All the varieties of coal (bituminous coal, anthracite; cannel-coal, +&c.) show a more or less distinct "lamination"—that is to +say, they are more or less obviously composed of successive thin layers, +differing slightly in colour and texture. All the varieties of coal, +also, consist chemically of <i>carbon</i>, with varying proportions +of certain gaseous constituents and a small amount of incombustible +mineral or "ash." By cutting thin and transparent slices of coal, +we are further enabled, by means of the microscope, to ascertain +precisely not only that the carbon of the coal is derived from +vegetables, but also, in many cases, what kinds of plants, and what +parts of these, enter into the formation of coal. When examined +in this way, all coals are found to consist more or less entirely +of vegetable matter; but there is considerable difference in +different coals as to the exact nature of this. By Professor +Huxley it has been shown that many of the English coals consist +largely of accumulations of rounded discoidal sacs or bags, which +are unquestionably the seed-vessels or "spore-cases" of certain +of the commoner coal-plants (such as the <i>Lepidodendra</i>). +The best bituminous coals seem to be most largely composed of +these spore-cases; whilst inferior kinds possess a progressively +increasing amount of the dull carbonaceous substance which is +known as "mineral charcoal," and which is undoubtedly composed +of "the stems and leaves of plants reduced to little more than +their carbon." On the other hand, Principal Dawson finds that +the American coals only occasionally exhibit spore-cases to any +extent, but consist principally of the cells, vessels, and fibres +of the bark, integumentary coverings, and woody portions of the +Carboniferous plants. +</p> + +<p class="indent"> +The number of plants already known to have existed +<a name="page_164"><span class="page">Page 164</span></a> +during the Carboniferous period is so great, that +nothing more can be done here than to notice briefly the typical +and characteristic <i>groups</i> of these—such as the +Ferns, the Calamites, the Lepidodendroids, the Sigillarioids, +and the Conifers. +</p> + +<p class="indent"> +In accordance with M. Brongniart's generalisation, that the +Palæozoic period is, botanically speaking, the "Age of +Acrogens," we find the Carboniferous plants to be still mainly +referable to the Flowerless or "Cryptogamous" division of the +vegetable kingdom. The flowering or "Phanerogamous" plants, +which form the bulk of our existing vegetation, are hardly known, +with certainty, to have existed at all in the Carboniferous era, +except as represented by trees related to the existing Pines and + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 449px; + font-size: smaller; text-align: center;"> +<img src="images/fig108.jpg" width="441" height="347" alt="Fig. 108" /> +<br /> +Fig. 108.—<i>Odontopteris Schlotheimii</i>. Carboniferous, +Europe and North America. +</span> +</span> + +Firs, and possibly by the Cycads or "false palms."[18] Amongst +the "Cryptogams," there is no more striking or beautiful group of +Carboniferous plants than the <i>Ferns</i>. Remains of these are +found all through the Carboniferous, but in exceptional numbers +in the Coal-measures, and include both herbaceous forms like the +majority of existing species, and arborescent forms resembling +the living Tree-ferns of New Zealand. Amongst the latter, together +with some new types, are examples of the genera <i>Psaronius</i> +and <i>Caulopteris</i>, both of +<a name="page_165"><span class="page">Page 165</span></a> +which date from the Devonian. The simply herbaceous ferns are +extremely numerous, and belong to such widely-distributed and + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 477px; + font-size: smaller; text-align: center;"> +<img src="images/fig109.jpg" width="469" height="746" alt="Fig. 109" /> +<br /> +Fig. 109.—<i>Calamites cannĹ“formis</i>. Carboniferous +Rocks, Europe and North America. +</span> +</span> + +largely-represented genera as <i>Neuropteris, Odontopteris</i> +(fig. 108), <i>Alethopteris, Pecopteris, Sphenopteris, +Hymenophyllites</i>, &c. +</p> + +<p class="footnote"> +[Footnote 18: Whilst the vegetation of the Coal-period was mainly +a terrestrial one, aquatic plants are not unknown. Sea-weeds (such +as the <i>Spirophyton cauda-Galli</i>) are common in some of the +marine strata; whilst coal, according to the researches of the +AbbĂ© Castracane, is asserted commonly to contain the +siliceous envelopes of Diatoms.] +</p> + +<p class="indent"> +The fossils known as <i>Calamites</i> (fig. 109) are very common +<a name="page_166"><span class="page">Page 166</span></a> +in the Carboniferous deposits, and have given +occasion to an abundance of research and speculation. They present +themselves as prostrate and flattened striated stems, or as similar +uncompressed stems growing in an erect position, and sometimes +attaining a length of twenty feet or more. Externally, the stems +are longitudinally ribbed, with transverse joints at regular +intervals, these joints giving origin to a whorl or branchlets, +which mayor may not give origin to similar whorls of smaller +branchlets still. The stems, further, were hollow, with transverse +partitions at the joints, and having neither true wood nor bark, +but only a thin external fibrous shell. There can be little doubt +but that the <i>Calamites</i> are properly regarded as colossal +representatives of the little Horse-tails (<i>EquisetaceĹ“</i>) +of the present day. They agree with these not only in the general +details of their organisation, but also in the fact that the fruit +was a species of cone, bearing "spore-cases" under scales. +According to Principal Dawson, the <i>Calamites</i> "grew in +dense brakes on the sandy and muddy flats, subject to inundation, +or perhaps even in water; and they had the power of budding out +from the base of the stem, so as to form clumps of plants, and +also of securing their foothold by numerous cord-like roots +proceeding from various heights on the lower part of the stem." +</p> + +<p class="indent"> +The <i>Lepidodendroids</i>, represented mainly by the genus +<i>Lepidodendron</i> itself (fig. 110), were large tree-like +plants, which attain their maximum in the Carboniferous period, +but which appear to commence in the Upper Silurian, are well +represented in the Devonian, and survive in a diminished form into +the Permian. The trunks of the larger species of <i>Lepidodendron</i> +at times reach a length of fifty feet and upwards, giving off +branches in a regular bifurcating manner. The bark is marked +with numerous rhombic or oval scars, arranged in quincunx order, +and indicating the points where the long, needle-shaped leaves +were formerly attached. The fruit consisted of cones or spikes, +carried at the ends of the branches, and consisting of a central +axis surrounded by overlapping scales, each of which supports +a "spore-case" or seed-vessel. These cones have commonly been +described under the name of <i>Lepidostrobi</i>. In the structure +of the trunk there is nothing comparable to what is found in +existing trees, there being a thick bark surrounding a zone +principally composed of "scalariform" vessels, this in turn +enclosing a large central pith. In their general appearance the +<i>Lepidodendra</i> bring to mind the existing Araucarian Pines; +but they are true "Cryptogams," and are to be regarded as a +gigantic extinct type of the +<a name="page_167"><span class="page">Page 167</span></a> +modern Club-mosses (<i>LycopodiaceĹ“</i>). They are amongst +the commonest and most characteristic of the Carboniferous + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 505px; + font-size: smaller; text-align: justify;"> +<img src="images/fig110.jpg" width="497" height="738" alt="Fig. 110" /> +<br /> +Fig. 110.—<i>Lepidodendron Sternbergii</i>, Carboniferous, +Europe. The central figure represents a portion of the trunk with +its branches, much reduced in size. The right-hand figure is a +portion of a branch with the leaves partially attached to it; and +the left-hand figure represents the end of a branch bearing a cone +of fructification. +</span> +</span> + +plants; and the majority of the "spore-cases" so commonly found +in the coal appear to have been derived from the cones of +Lepidodendroids. +</p> + +<p class="indent"> +<a name="page_168"><span class="page">Page 168</span></a> +The so-called <i>Sigillanoids</i>, represented +mainly by <i>Sigillaria</i> itself (fig. 111), were no less +abundant and characteristic of the Carboniferous forests than the +<i>Lepidodendra</i>. They commence their existence, so far as +known, in the Devonian period, but they attain their maximum in the +Carboniferous; and—unlike the Lepidodendroids—they are +not known to occur in the Permian period. They are comparatively +gigantic in size, often attaining a height of from thirty to +fifty feet or more; but though abundant and well preserved, great +divergence of opinion prevails as to their true affinities. The +<i>name</i> of Sigillarioids (Lat. <i>sigilla</i>, little seals +or images) is derived from the fact that the bark is marked with +seal-like impressions or leaf-scars (fig. 111). +</p> + +<p class="indent"> +Externally, the trunks of <i>Sigillaria</i> present strong +longitudinal ridges, with vertical alternating rows of oval +leaf-scars indicating the points where the leaves were originally + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 373px; + font-size: smaller; text-align: justify;"> +<img src="images/fig111.jpg" width="365" height="337" alt="Fig. 111" /> +<br /> +Fig. 111.—Fragment of the external surface of <i>Sigillaria +GrĹ“seri</i>, showing the ribs and leaf-scars. The left-hand +figure represents a small portion enlarged. Carboniferous, +Europe. +</span> +</span> + +attached. The trunk was furnished with a large central pith, a thick +outer bark, and an intermediate woody zone,—composed, according +to Dawson, partly of the disc-bearing fibres so characteristic of +Conifers; but, according to Carruthers, entirely made up of the +"scalariform" vessels characteristic of Cryptogams. The size of the +pith was very great, and the bark seems to have been the most +durable portion of the trunk. Thus we have evidence that in many +cases the stumps and "stools" of <i>SigillariĹ“</i>, standing +<a name="page_169"><span class="page">Page 169</span></a> +upright in the old Carboniferous swamps, were +completely hollowed out by internal decay, till nothing but an +exterior shell of bark was left. Often these hollow stumps became +ultimately filled up with sediment, sometimes enclosing the remains +of galley-worms, land-snails, or Amphibians, which formerly found in +the cavity of the trunk a congenial home; and from the sandstone or +shale now filling such trunks some of the most interesting fossils +of the Coal-period have been obtained. There is little certainty as +to either the leaves or fruits of <i>Sigillaria</i>, and there +is equally little certainty as to the true botanical position of +these plants. By Principal Dawson they are regarded as being +probably flowering plants allied to the existing "false palms" +or "<i>Cycads</i>," but the high authority of Mr Carruthers is to +be quoted in support of the belief that they are Cryptogamic, +and most nearly allied to the Club-mosses. +</p> + +<p class="indent"> +Leaving the botanical position of <i>Sigillaria</i> thus undecided, +we find that it is now almost universally conceded that the fossils +originally described under the name of <i>Stigmaria</i> are the +<i>roots</i> of <i>Sigillaria</i>, the actual connection between +the two having been in numerous instances demonstrated in an +unmistakable manner. The <i>StigmariĹ“</i> (fig. 112) ordinarily +present themselves in the form of long, compressed or rounded + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 386px; + font-size: smaller; text-align: center;"> +<img src="images/fig112.jpg" width="378" height="227" alt="Fig. 112" /> +<br /> +Fig. 112.—<i>Stigmaria ficoides</i>. Quarter natural size. +Carboniferous. +</span> +</span> + +fragments, the external surface of which is covered with rounded +pits or shallow tubercles, each of which has a little pit or +depression in its centre. From each of these pits there proceeds, +in perfect examples, a long cylindrical rootlet; but in many cases +these have altogether disappeared. In their internal structure, +<i>Stigmaria</i> exhibits a central pith surrounded by a sheath of +scalariform vessels, the whole enclosed in a cellular envelope. +The <i>StigmariĹ“</i> are generally found ramifying in +<a name="page_170"><span class="page">Page 170</span></a> +the "under-clay," which forms the floor of a +bed of coal, and which represents the ancient soil upon which the +<i>SigillariĹ“</i> +grew. +</p> + +<p class="indent"> +The <i>Lepidodendroids</i> and <i>Sigillaroids</i>, though the first +were certainly, and the second possibly, Cryptogamic or flowerless +plants, must have constituted the main mass of the forests of +the Coal period; but we are not without evidence of the existence +at the same time of genuine "trees," in the technical sense of +this term—namely, flowering plants with large woody stems. So +far as is certainly known, all the true trees of the Carboniferous +formation were <i>Conifers</i>, allied to the existing Pines and +Firs. They are recognised by the great size and concentric woody +rings of their prostrate, rarely erect trunks, and by the presence +of disc-bearing fibres in their wood, as demonstrated by the +microscope; and the principal genera which have been recognised are +<i>Dadoxylon, PalĹ“oxylon, Araucarioxylon</i>, and <i>Pinites</i>. +Their fruit is not known with absolute certainty, unless it be +represented, as often conjectured, by <i>Trigonocarpon</i> (fig. +113). The fruits known under this name are nut-like, often of + +<span style="float: left; margin: 4px; width: 187px; + font-size: smaller; text-align: justify;"> +<img src="images/fig113.jpg" width="179" height="117" alt="Fig. 113" /> +<br /> +Fig. 113.—<i>Trigonocarpon ovatum</i>. Coal-measures, Britain. +(After Liudley and Hutton.) +</span> + +considerable size, and commonly three- or six-angled. They probably +originally possessed a fleshy envelope; and if truly referable +to the <i>Conifers</i>, they would indicate that these ancient +evergreens produced berries instead of cones, and thus resembled +the modern Yews rather than Pines. It seems, further, that the +great group of the <i>Cycads</i>, which are nearly allied to the +<i>Conifers</i>, and which attained such a striking prominence in +the Secondary period, probably commenced its existence during the +Coal period; but these anticipatory forms are comparatively few +in number, and for the most part of somewhat dubious affinities. +</p> + +<h3>CHAPTER XIII.</h3> + +<p class="subtitle"> +THE CARBONIFEROUS PERIOD—<i>Continued</i>. +</p> + +<p class="center"> +ANIMAL LIFE OF THE CARBONIFEROUS. +</p> + +<p class="indent"> +We have seen that there exists a great difference as to the mode +of origin of the Carboniferous sediments, some being purely marine, +whilst others are terrestrial; and others, again, +<a name="page_171"><span class="page">Page 171</span></a> +have been formed in inland swamps and morasses, or in +brackish-water lagoons, creeks, or estuaries. A corresponding +difference exists necessarily in the animal remains of these +deposits, and in many regions this difference is extremely well +marked and striking. The great marine limestones which +characterise the lower portion of the Carboniferous series in +Britain, Europe, and the eastern portion of America, and the +calcareous beds which are found high up in the Carboniferous in +the western States of America, may, and do, often contain the +remains of drifted plants; but they are essentially characterised +by marine fossils; and, moreover, they can be demonstrated by +the microscope to be almost wholly composed of the remains of +animals which formerly inhabited the ocean. On the other hand, +the animal remains of the beds accompanying the coal are typically +the remains of air-breathing, terrestrial, amphibious, or aerial +animals, together with those which inhabit fresh or brackish +waters. Marine fossils may be found in the Coal-measures, but +they are invariably confined to special horizons in the strata, +and they indicate temporary depressions of the land beneath the +sea. Whilst the distinction here mentioned is one which cannot +fail to strike the observer, it is convenient to consider the +animal life of the Carboniferous as a whole: and it is simply +necessary, in so doing, to remember that the marine fossils are +in general derived from the inferior portion of the system; +whilst the air-breathing, fresh-water, and brackish-water forms +are almost exclusively derived from the superior portion of the +same. +</p> + +<p class="indent"> +The Carboniferous <i>Protozoans</i> consist mainly of +<i>Foraminifera</i> and <i>Sponges</i>. The latter are still +very insufficiently known, but the former are very abundant, +and belong to very varied types. Thin slices of the limestones +of the period, when examined by the microscope, very commonly +exhibit the shells of <i>Foraminifera</i> in greater or less +plenty. Some limestones, indeed, are made up of little else than +these minute and elegant shells, often belonging to types, such +as the Textularians and Rotalians, differing little or not at +all from those now in existence. This is the case, for example, +with the Carboniferous Limestone of Spergen Hill in Indiana (fig. +114), which is almost wholly made up of the spiral shells of a +species of <i>Endothyra</i>. In the same way, though to a less +extent, the black Carboniferous marbles of Ireland, and the similar +marbles of Yorkshire, the limestones of the west of England and +of Derbyshire, and the great "Scar Limestones" of the north of +England, contain great numbers of Foraminiferous shells; whilst +similar organisms commonly occur in the shale-beds associated +<a name="page_172"><span class="page">Page 172</span></a> +with the limestones throughout the Lower Carboniferous series. +One of the most interesting of the British Carboniferous forms + +<span style="float: left; margin: 4px; width: 268px; + font-size: smaller; text-align: justify;"> +<img src="images/fig114.jpg" width="260" height="258" alt="Fig. 114" /> +<br /> +Fig. 114.—Transparent slice of Carboniferous Limestone, +from Spergen Hill, Indiana, U.S., showing numerous shells of +<i>Endothyra</i> (<i>Rotalia</i>), <i>Baiteyi</i> slightly +enlarged. (Original.) +</span> + +is the <i>Saccammina</i> of Mr Henry Brady, which is sometimes +present in considerable numbers in the limestones of Northumberland, +Cumberland, and the west of Scotland, and which is conspicuous +for the comparatively large size of its spheroidal or pear-shaped +shell (reaching from an eighth to a fifth of an inch in size). +More widely distributed are the generally spindle-shaped shells +of <i>Fusulina</i> (fig. 115), which occur in vast numbers in +the Carboniferous Limestone of Russia, Armenia, the Southern +Alps, and Spain, similar forms occurring in equal profusion in +the higher limestones which are found in the Coal-measures of the +United States, in Ohio, Illinois, Indiana, Missouri, &c. Mr +Henry Brady, lastly, has shown that we have in the <i>Nummulina +Pristina</i> of the Carboniferous Limestone of Namur a genuine + +<span style="float: left; margin: 4px; width: 289px; + font-size: smaller; text-align: center;"> +<img src="images/fig115.jpg" width="281" height="76" alt="Fig. 115" /> +<br /> +Fig. 115.—<i>Fusulina cylindrica</i>, Carboniferous +Limestone, Russia. +</span> + +<i>Nummulite</i>, precursor of the great and important family +of the Tertiary Nummulites. +</p> + +<p class="indent"> +The sub-kingdom of the <i>CĹ“lenterates</i>, so far as certainly +known, is represented only by <i>Corals</i>;[19] but the remains of +these are so abundant in many of the limestones of the Carboniferous +formation as to constitute a feature little or not at all less +conspicuous than that afforded by the Crinoids. As is the case in +the preceding period, the Corals belong, almost exclusively, to +the groups of the <i>Rugosa</i> and <i>Tabulata</i>; and there is +a general and striking resemblance and relationship between the +coral-fauna of the Devonian as a whole, and that +<a name="page_173"><span class="page">Page 173</span></a> +of the Carboniferous. Nevertheless, there is an equally decided and +striking amount of difference between these successive faunas, due +to the fact that the great majority of the Carboniferous <i>species</i> +are new; whilst some of the most characteristic Devonian <i>genera</i> +have nearly or quite disappeared, and several new genera now make +their appearance for the first time. Thus, the characteristic Devonian +types <i>Heliophyllum, Pachyphyllum, Chonophyllum, Acervularia, +Spongophyllum, Smithia, Endophyllum</i>, and <i>Cystiphyllum</i>, +have now disappeared; and the great masses of <i>Favosites</i> +which are such a striking feature in the Devonian limestones, +are represented but by one or two degenerate and puny successors. +On the other hand, we meet in the Carboniferous rocks not only with +entirely new genera—such as <i>Axophyllum, Lophophyllum</i>, +and <i>Londsdaleia</i>—but we have an enormous expansion of +certain types which had just begun to exist in the preceding +period. This is especially well seen in the Case of the genus +<i>Lithostrotion</i> (fig. 116, <i>b</i>), which more than any +other may be considered as the predominant Carboniferous group +of Corals. All the species of <i>Lithostrotion</i> are compound, +consisting either of bundles of loosely-approximated cylindrical +stems, or of similar "coral-lites" closely aggregated together into +astræiform colonies, and rendered polygonal by mutual pressure. +This genus has a historical interest, as having been noticed as +early as in the year 1699 by Edward Lhwyd; and it is geologically +important from its wide distribution in the Carboniferous rocks +of both the Old and New Worlds. Many species are known, and whole +beds of limestone are often found to be composed of little else +than the skeletons of these ancient corals, still standing upright +as they grew. Hardly less characteristic of the Carboniferous +than the above is the great group of simple "cup-corals," of +which <i>Clisiophyllum</i> is the central type. Amongst types +which commenced in the Silurian and Devonian, but which are still +well represented here, may be mentioned <i>Syringopora</i> (fig. +116, <i>e</i>), with its colonies of delicate cylindrical tubes +united at intervals by cross-bars; <i>Zaphrentis</i> (fig. 116, +<i>d</i>), with its cup-shaped skeleton and the well-marked +depression (or "fossula") on one side of the calice; <i>Amplexus</i> +(fig. 116, <i>c</i>), with its cylindrical, often irregularly +swollen coral and short septa; <i>Cyathophyllum</i> (fig. 116, +<i>a</i>), sometimes simple, sometimes forming great masses of +star-like corallites; and <i>ChĹ“tetes</i>, with its branched +stems, and its minute, "tabulate" tubes (fig. 116, <i>f</i>). +The above, together with other and hardly less characteristic +forms, combine to constitute a coral-fauna which is not only in +itself perfectly distinctive, but which is of especial interest, +<a name="page_174"><span class="page">Page 174</span></a> +from the fact that almost all the varied types of which it is +composed disappeared utterly before the close of the Carboniferous + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 557px; + font-size: smaller; text-align: justify;"> +<img src="images/fig116.jpg" width="549" height="593" alt="Fig. 116" /> +<br /> +Fig. 116—Corals of the Carboniferous Limestone. <i>a. +Cyathophyllum paracida</i>, showing young corallites budded forth +from the disc of the old one; <i>a'</i>, One of the corallites of +the same, seen in cross-section; <i>b</i>, Fragment of a mass of +<i>Lithostrotion irregulare; b'</i>, One of the corallites of the +same, divided transversely; <i>c</i>, Portion of the simple +cylindrical coral of <i>Amplexus coralloides; c'</i>, Transverse +section of the same species; <i>d, Zaphrentis vermicularis</i>, +showing the depression or "fossula" on one side of the cup; +<i>e</i>, Fragrent of a mass of <i>Syringopora ramulosa; f</i>, +Fragment of <i>CĹ“tetes tumidus; f'</i>, Portion of the same +of the same, enlarged. From the Carboniferous Limestone of Britain +and Belgium. (After Thomson, De Koninck, Milne-Edwards and Haime, +and the Author.) +</span> +</span> + +period. In the first marine sediments of a calcareous nature +which succeeded to the Coal-measures (the magnesian limestones +of the Permian), the great group of the <i>Rugose corals</i>, +which flourished so largely throughout the Silurian, Devonian, +and Carboniferous periods, is found to have all but +<a name="page_175"><span class="page">Page 175</span></a> +disappeared, and it is never again represented save +sporadically and by isolated forms. +</p> + +<p class="footnote"> +[Footnote 19: A singular fossil has been described by Professor +Martin Duncan and Mr Jenkins from the Carboniferous rocks under +the name of <i>PalĹ“ocoryne</i>, and has been referred to +the Hydroid Zoophytes (<i>Corynida</i>). Doubt, however, has been +thrown by other observers on the correctness of this reference.] +</p> + +<p class="indent"> +Amongst the <i>Echinoderms</i>, by far the most important forms +are the Sea-lilies and the Sea-urchins—the former from their +great abundance, and the latter from their singular structure; but +the little group of the "Pentremites" also requires to be noticed. +The Sea-lilies are so abundant in the Carboniferous rocks, that it +has been proposed to call the earlier portion of the period the +"Age of Crinoids." Vast masses of the limestones of the period are +"crinoidal," being more or less extensively composed of the broken +columns, and detached plates and joints of Sea-lilies, whilst +perfect "heads" may be exceedingly rare and difficult to procure. +In North America the remains of Crinoids are even more abundant +at this horizon than in Britain, and the specimens found seem +to be commonly more perfect. The commonest of the Carboniferous +Crinoids belong to the genera <i>Cyathocrinus, Actinocrinus, +Platycrinus</i>, (fig. 117), <i>Poteriocrinus, Zeacrinus</i>, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 536px; + font-size: smaller; text-align: justify;"> +<img src="images/fig117.jpg" width="528" height="419" alt="Fig. 117" /> +<br /> +Fig. 117.—<i>Platycrinus tricontadactylus</i>, Lower +Carboniferous. The left-hand figure shows the calyx, arms, and +upper part of the stem; and the figure next this shows the surface +of one of the joints of the column. The right-hand figure shows +the proboscis. (After M'Coy.) +</span> +</span> + +and <i>Forbesiocrinus</i>. Closely allied to the Crinoids, or +forming a kind of transition +<a name="page_176"><span class="page">Page 176</span></a> +between these and the Cystideans, is the +little group of the "Pentremites," or <i>Blastoids</i> (fig. 118). +This group is first known to have commenced its existence in + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 485px; + font-size: smaller; text-align: justify;"> +<img src="images/fig118.jpg" width="477" height="432" alt="Fig. 118" /> +<br /> +Fig. 118.—A, <i>Pentremites pyriformis</i>, side-view of the +body ("calyx"); B, The same viewed from below, showing the +arrangement of the plates; C, Body of <i>Pentremites conoideus</i>, +viewed from above. Carboniferous. +</span> +</span> + +the Upper Silurian, and it increased considerably in numbers in +the Devonian; but it was in the seas of the Carboniferous period +that it attained its maximum, and no certain representative of the +family has been detected in any later deposits. The "Pentremites" +resemble the Crinoids in having a cup-shaped body (fig. 118, A) +enclosed by closely-fitting calcareous plates, and supported on +a short stem or "column," composed of numerous calcareous pieces +flexibly articulated together. They differ from the Crinoids, +however, in the fact that the upper surface of the body does +not support the crown of branched feathery "arms," which are +so characteristic of the latter. On the contrary, the summit of +the cup is closed up in the fashion of a flower-bud, whence the +technical name of <i>Blastoidea</i> applied to the group (Gr. +<i>blastos</i>, a bud; <i>eidos</i>, form). From the top of the +cup radiate five broad, transversely-striated areas (fig. 118, C), +each with a longitudinal groove down its middle; and along each +side of each of +<a name="page_177"><span class="page">Page 177</span></a> +these grooves there seems to have been attached a row of short +jointed calcareous filaments or "pinnules." +</p> + +<p class="indent"> +A few Star-fishes and Brittle-stars are known to occur in the +Carboniferous rocks; but the only other Echinodemls of this period +which need be noticed are the Sea-urchins (<i>Echinoids</i>). +Detached plates and spines of these are far from rare in the +Carboniferous deposits; but anything like perfect specimens are +exceedingly scarce. The Carboniferous Sea-urchins agree with those +of the present day in having the body enclosed in a shell formed +by an enormous number of calcareous plates articulated together. +The shell may be regarded as, typically, nearly spherical in +shape, with the mouth in the centre of the base, and the excretory +opening or vent at its summit. In both the ancient forms and the +recent ones, the plates of the shell are arranged in ten zones + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 513px; + font-size: smaller; text-align: justify;"> +<img src="images/fig119.jpg" width="505" height="240" alt="Fig. 119" /> +<br /> +Fig. 119.—<i>PalĹ“chinus ellipticus</i>, one of the +Carboniferous Sea-urchins. The left-hand figure shows one of the +"ambulacral areas" enlarged, exhibiting the perforated plates. +The right-land figure exhibits a single plate from one of the +"inter-ambulacral areas." (After M'Coy.) +</span> +</span> + +which generally radiate from the summit to the centre of the base. In +five of these zones—termed the "ambulacral areas"—the +plates are perforated by minute apertures or "pores," through which +the animal can protrude the little water-tubes ("tube-feet") by which +its locomotion is carried on. In the other five zones—the +so-called "inter-ambulacral areas"—the plates are of larger +size, and are not perforated by any apertures. In all the modern +Sea-urchins each of these ten zones, whether perforate or imperforate, +is composed of two rows of plates; and there are thus twenty rows of +plates in all. In the Palæozoic Sea-urchins, on the other hand, +the "ambulacral areas" are often like those of recent forms, in +consisting of <i>two</i> rows of perforated plates (fig. 119); but +the "inter-ambulacral areas" are always quite +<a name="page_178"><span class="page">Page 178</span></a> +peculiar in consisting each of three, four, five, or more rows of large +imperforate plates, whilst there are sometimes four or ten rows of +plates in the "ambulacral areas" also: so that there are many more than +twenty rows of plates in the entire shell. Some of the Palæozoic +Sea-urchins, also, exhibit a very peculiar singularity of structure +which is only known to exist in a very few recently-discovered +modern forms (viz., <i>Calveria</i> and <i>Phormosoma</i>). The +plates of the inter-ambulacral areas, namely, overlap one another +in an imbricating manner, so as to communicate a certain amount +of flexibility to the shell; whereas in the ordinary living forms +these plates are firmly articulated together by their edges, +and the shell forms a rigid immovable box. The Carboniferous +Sea-urchins which exhibit this extraordinary peculiarity belong +to the genera <i>Lepidechinus</i> and <i>Lepidesthes</i>, and +it seems tolerably certain that a similar flexibility of the +shell existed to a less degree in the much more abundant genus +<i>ArchĹ“ocidaris</i>. The Carboniferous Sea-urchins, like +the modern ones, possessed movable spines of greater or less +length, articulated to the exterior of the shell; and these +structures are of very common occurrence in a detached condition. +The most abundant genera are <i>ArchĹ“ocidaris</i> and +<i>PalĹ“chinus</i>; but the characteristic American forms +belong principally to <i>Melonites, Oligoporus</i>, and +<i>Lepidechinus</i>. +</p> + +<p class="indent"> +Amongst the <i>Annelides</i> it is only necessary to notice the +little spiral tubes of <i>Spirorbis Carbonarius</i> (fig. 120), + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 332px; + font-size: smaller; text-align: center;"> +<img src="images/fig120.jpg" width="324" height="191" alt="Fig. 120" /> +<br /> +Fig. 120.—<i>Spirorbis (Microconchus) Carbonarius</i>, of +the natural size, attached to a fossil plant, and magnified. +Carboniferous Britain and North America. (After Dawson.) +</span> +</span> + +which are commonly found attached to the leaves or stems of the +Coal-plants. This fact shows that though the modern species of +<i>Spirorbis</i> are inhabitants of the sea, these old +representatives of the genus must have been capable of living +in the brackish waters of lagoons and estuaries. +</p> + +<p class="indent"> +The <i>Crustaceans</i> of the Carboniferous rocks are numerous, +<a name="page_179"><span class="page">Page 179</span></a> +and belong partly to structural types with which we are already +familiar, and partly to higher groups which come into existence +here for the first time. The gigantic <i>Eurypterids</i> of the +Upper Silurian and Devonian are but feebly represented, and make +their final exit here from the scene of life. Their place, however, +is taken by peculiar forms belonging to the allied group of the +<i>Xiphosura</i>, represented at the present day by the King-crabs +or "Horse-shoe Crabs" (<i>Limulus</i>). Characteristic forms of this +group appear in the Coal-measures both of Europe and America; and +though constituting three distinct genera (<i>Prestwichia, Belinurus</i>, +and <i>Euproöps</i>), they are all nearly related to one +another. The best known of them, perhaps, is the <i>Prestwichia +rotundala</i> of Coalbrookdale, here figured (fig. 121). The ancient + +<span style="float: right; margin: 4px; width: 304px; + font-size: smaller; text-align: justify;"> +<img src="images/fig121.jpg" width="296" height="321" alt="Fig. 121" /> +<br /> +Fig. 121.—<i>Prestwichia rotundata</i>, a Limuloid Crustacean. +Coal-measures, Britain. (After Henry Woodward.) +</span> + +and formerly powerful order of the <i>Trilobites</i> also undergoes +its final extinction here, not surviving the deposition of the +Carboniferous Limestone series in Europe, but extending its range +in America into the Coal-measures. All the known Carboniferous forms +are small in size and degraded in point of structure, and they are +referable to but three genera (<i>Phillipsia, Griffithides</i>, +and <i>Brachymetopus</i>), belonging to a single family. The +<i>Phillipsia seminifera</i> here figured (fig. 122, <i>a</i>) +is a characteristic species in the Old World. The Water-fleas +(<i>Ostracoaa</i>) are extremely abundant in the Carboniferous +rocks, whole strata being often made up of little else than the +little bivalved shells of these Crustaceans. Many of them are +extremely small, averaging about the size of a millet-seed; but +a few forms, such as <i>Entomoconchus Scouleni</i> (fig. 122, +<i>c</i>), may attain a length of from one to three quarters of +an inch. The old group of the <i>Phyllopods</i> is is likewise +still represented in some abundance, partly by tailed forms of a +shrimp-like appearance, such as <i>Dithyrocaris</i> (fig. 122, +<i>d</i>), and partly by the curious striated <i>EstheriĹ“</i> +and their allies, which present a curious +<a name="page_180"><span class="page">Page 180</span></a> +resemblance to the true Bivalve Molluscs (fig. 122, <i>b</i>). +Lastly, we meet for the first time in the Carboniferous rocks +with the remains of the highest of all the groups of +<i>Crustaceans</i>—namely, the so-called "Decapods," in +which there are five pairs of walking-limbs, and the hinder end +of the body ("abdomen") is composed of separate rings, whilst +the anterior end is covered by a head-shield or "carapace." All +the Carboniferous Decapods hitherto discovered resemble the +existing Lobsters, Prawns, and + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 531px; + font-size: smaller; text-align: justify;"> +<img src="images/fig122.jpg" width="523" height="455" alt="Fig. 122" /> +<br /> +Fig. 122.—Crustaceans of the Carboniferous Rocks. <i>a, +Phillipsia seminifera</i>, of the natural size—Mountain +Limestone, Europe; <i>b</i>, One valve of the shell of <i>Estheria +tenella</i>, of the natural size and enlarged—Coal-measures, +Europe; <i>c</i>, Bivalved shell of <i>Entomoconchus Scouleri</i>, +of the natural size—Mountain Limestone, Europe; <i>d, +Dithyrocaris Scouleri</i>, reduced in size—Mountain Limestone, +Ireland; <i>e, PalĹ“ocaris typus</i>, slightly +enlarged—Coal-measures, North America; <i>f, +AnthrapalĹ“mon gracilis</i>, of the natural +size—Coal-measures, North America. (After De Koninck, M'Coy, +Rupert Jones, and Meek and Worthen.) +</span> +</span> + +Shrimps (the <i>Macrura</i>), in having a long and well-developed +abdomen terminated by an expanded tail-fin. The <i>PalĹ“ocaris +typus</i> (fig. 122, <i>e</i>) and the <i>AnthrapalĹ“mon +gracilis</i> (fig. 122, <i>f</i>), from the Coal-measures of +Illinois, are two of the best understood and most perfectly +preserved of the few known representatives of the "Long-tailed" +Decapods in the Carboniferous series. The group of the Crabs or +"Short-tailed" +<a name="page_181"><span class="page">Page 181</span></a> +Decapods (<i>Brachyura</i>), in which the abdomen is short, not +terminated by a tail-fin, and tucked away out of sight beneath +the body, is at present not known to be represented at all in +the Carboniferous deposits. +</p> + +<p class="indent"> +In addition to the water-inhabiting group of the Crustaceans, we +find the articulate animals to be represented by members belonging +to the air-breathing classes of the <i>Arachnida, Myriapoda</i>, +and <i>Insecta</i>. The remains of these, as might have been +expected, are not known to occur in the marine limestones of the +Carboniferous series, but are exclusively found in beds associated +with the Coal, which have been deposited in lagoons, estuaries, or +marshes, in the immediate vicinity of the land, and which actually +represent an old land-surface. The <i>Arachnids</i> are at present +the oldest known of their class, and are represented both by true +Spiders and Scorpions. Remains of the latter (fig. 123) have been +found both in the Old and New Worlds, and indicate the existence + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 575px; + font-size: smaller; text-align: center;"> +<img src="images/fig123.jpg" width="523" height="447" alt="Fig. 123" /> +<br /> +Fig. 123.—<i>Cyclophthalmus senior</i>. A fossil Scorpion +from the Coal-measures of Bohemia. +</span> +</span> + +in the Carboniferous period of Scorpions differing but very little +from existing forms. The group of the <i>Myriapoda</i>, including +the recent Centipedes and Galley-worms, is likewise represented +in the Carboniferous strata, +<a name="page_182"><span class="page">Page 182</span></a> +but by forms in many respects +very unlike any that are known to exist at the present day. +The most interesting of these were obtained by Principal Dawson, +along with the bones of Amphibians and the shells of Land-snails, +in the sediment filling the hollow trunks of <i>Sigillaria</i>, +and they belong to the genera <i>Xylobius</i> (fig. 124) and +Archiulus. Lastly, the true <i>insects</i> are represented by + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 296px; + font-size: smaller; text-align: justify;"> +<img src="images/fig124.jpg" width="288" height="237" alt="Fig. 124" /> +<br /> +Fig. 124.—<i>Xylobius SigillariĹ“</i>, a Carboniferous +Myriapod. <i>a</i>, A specimen, of the natural size; <i>b</i>, +Anterior portion of the same, enlarged; <i>c</i>, Posterior +portion, enlarged. From the Coal-measures of Nova Scotia. +(After Dawson.) +</span> +</span> + +various forms of Beetles (<i>Coleoptera</i>), <i>Orthoptera</i> +(such as Cockroaches), and <i>Neuropterous</i> insects resembling +those which we have seen to have existed towards the close of + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 613px; + font-size: smaller; text-align: center;"> +<img src="images/fig125.jpg" width="605" height="316" alt="Fig. 125" /> +<br /> +Fig. 125—<i>Haplophlebium Barnesi</i>, a Carboniferous +insect, from the Coal-meastures of Nova Scotia. (After Dawson.) +</span> +</span> + +the Devonian period. One of the most remarkable of the latter +is a huge May-fly (<i>Haplophlebium Barnesi</i>, fig. 125), with +<a name="page_183"><span class="page">Page 183</span></a> +netted wings attaining an expanse of fully seven inches, and +therefore much exceeding any existing Ephemerid in point of size. +</p> + +<p class="indent"> +The lower groups of the <i>Mollusca</i> are abundantly represented +in the marine strata of the Carboniferous series by <i>Polyzoans</i> + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 487px; + font-size: smaller; text-align: justify;"> +<img src="images/fig126.jpg" width="479" height="570" alt="Fig. 126" /> +<br /> +Fig. 126.—Carboniferous <i>Polyzoa</i>. <i>a</i>, Fragment of +<i>Polypora dendroides</i>, of the natural size, Ireland; <i>a'</i> +Small portion of the same, enlarged to show the cells; <i>b, +Glauconome pulcherrima</i>, a fragment, of the natural size, +Ireland; <i>b'</i>, Portion of the same, enlarged; <i>c</i>, The +central screw-like axis of <i>Archimedes Wortheni</i>, of the +natural size—Carboniferous, America; <i>c'</i>, Portion of +the exterior of the frond of the same, enlarged; <i>c''</i>, +Portion of the interior of the frond of the same showing the +mouths of the cells, enlarged. (After M'Coy and Hall.)] +</span> +</span> + +and <i>Brachiopods</i>. Amongst the former, although a variety +of other types are known, the majority still belong to the old +group of the "Lace-corals" (<i>FenestellidĹ“</i>), some of +the characteristic forms of which are here figured (fig. 126). +The graceful +<a name="page_184"><span class="page">Page 184</span></a> +netted fronds of <i>Fenestella, +Retepora</i>, and <i>Polypora</i> (fig. 126, <i>a</i>) are highly +characteristic, as are the slender toothed branches of +<i>Glauconome</i> (fig. 126, <i>b</i>). A more singular form, +however, is the curious <i>Archimedes</i> (fig. 126, <i>c</i>), +which is so characteristic of the Carboniferous formation of +North America. In this remarkable type, the colony consists of +a succession of funnel-shaped fronds, essentially similar to +<i>Fenestella</i> in their structure, springing in a continuous +spiral from a strong screw-like vertical axis. The outside of +the fronds is simply striated; but the branches exhibit on the +interior the mouths of the little cells in which the +semi-independent beings composing the colony originally lived. +</p> + +<p class="indent"> +The <i>Brachiopods</i> are extremely abundant, and for the most +part belong to types which are exclusively or principally +Palæozoic in their range. The old genera <i>Strophomena, +Orthis</i> (fig. 127, <i>c</i>), <i>Athyris</i> (fig. 127, +<i>e</i>), <i>Rhynchonella</i> (fig. 127, <i>g</i>), and +<i>Spirifera</i> (fig. 127, <i>h</i>), are still well +represented—the latter, in particular, existing under +numerous specific forms, conspicuous by their abundance and +sometimes by their size. Along with these ancient groups, we +have representatives—for the first time in any plenty—of +the great genus <i>Terebratula</i> (fig. 127, <i>d</i>), which +underwent a great expansion during later periods, and still exists +at the present day. The most characteristic Carboniferous +Brachiopods, however, belong to the family of the +<i>ProductidĹ“</i>, of which the principal genus is +<i>Producta</i> itself. This family commenced its existence in +the Upper Silurian with the genus <i>Chonetes</i>, distinguished +by its spinose hinge-margin. This genus lived through the +Devonian, and flourished in the Carboniferous (fig. 127, +<i>f</i>). The genus <i>Producta</i> itself, represented in the +Devonian by the nearly allied <i>Productella</i>, appeared first +in the Carboniferous, at any rate, in force, and survived into +the Permian; but no member of this extensive family has yet been +shown to have over-lived the Palæozoic period. The +<i>ProductĹ“</i> of the Carboniferous are not only +exceedingly abundant, but they have in many instances a most +extensive geographical range, and some species attain what may +fairly be considered-gigantic dimensions. The shell (fig. 127, +<i>a</i> and <i>b</i>) is generally more or less semicircular, +with a straight hinge-margin, and having its lateral angles +produced into larger or smaller ears (hence its generic +name—"<i>cochlea producta</i>"). One valve (the ventral) is +usually strongly convex, whilst the other (the dorsal) is flat +or concave, the surface of both being adorned with radiating +ribs, and with hollow tubular spines, often of great length. +The valves are not locked together by teeth, and there is no +sign in the +<a name="page_185"><span class="page">Page 185</span></a> +fully-grown shell of an opening in or between the valves for +the emission of a muscular stalk for the attachment of the shell +to foreign objects. It is probable, therefore, that the +<i>ProductĹ“</i>, unlike the ordinary Lamp-shells, lived +an independent existence, their long spines apparently serving +to anchor them firmly in the mud or ooze of the sea-bottom; but +Mr Robert Etheridge, jun.; has recently shown that in one species + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 498px; + font-size: smaller; text-align: justify;"> +<img src="images/fig127.jpg" width="490" height="488" alt="Fig. 127" /> +<br /> +Fig. 127.—Carboniferous <i>Braciopoda. a, Producta +semireticulata</i>, showing the slightly concave dorsal valve; +<i>a'</i> Side view of the same, showing the convex ventral valve; +<i>b, Producta longispina; c, Orthis resupinata; d, Terebratula +hastata; e, Athyris subtilita; f, Chonetes Hardrensis; g, +Rhynchonella pleurodon; h, Spirifera trigonalis</i>. Most of +these forms are widely distributed in the Carboniferous Limestone +of Britain, Europe, America, &c. All the figures are of the +natural size. (After Davidson, De Koninck, and Meek.) +</span> +</span> + +the spines were actually employed as organs of adhesion, whereby +the shell was permanently attached to some extraneous object, +such as the stem of a Crinoid. The two species here figured are +interesting for their extraordinarily extensive geographical +range—<i>Producta semireticulata</i> (fig. 127, <i>a</i>) +being found in the Carboniferous rocks of Britain, the continent +of Europe, Central Asia, China, India, Australia, Spitzbergen, +and North +<a name="page_186"><span class="page">Page 186</span></a> +and South America; whilst <i>P. Longispina</i> (fig. 127, +<i>b</i>) has a distribution little if at all less wide. +</p> + +<p class="indent"> +The higher <i>Mollusca</i> are abundantly represented in the +Carboniferous rocks by Bivalves (<i>Lamellibranchs</i>), Univalves +(<i>Gasteropoda</i>), Winged-snails (<i>Pteropoda</i>), and +<i>Cephalopods</i>. Amongst the Bivalves we may note the great +abundance of Scallops (<i>Aviculopecten</i> and other allied forms), +together with numerous other types—some of ancient origin, +others represented here for the first time. Amongst the Gasteropods, +we find the characteristically Palæozoic genera +<i>Macrocheilus</i> and <i>Loxonema</i>, the almost exclusively +Palæozoic <i>Euomphalus</i>, and the persistent, genus +<i>Pleurotomaria</i>; whilst the free-swimming Univalves +(<i>Heteropoda</i>)are represented by <i>Bellerophon</i> and +<i>Porcellia</i>, and the <i>Pteropoda</i> by the old genus +<i>Conularia</i>. With regard to the Carboniferous Univalves, +it is also of interest to note here the first appearance of true +air-breathing or terrestrial Molluscs, as discovered by +Dawson and Bradley in the Coal-measures of Nova Scotia and Illinois. +Some of these (<i>Conulus priscus</i>) are true Land-snails, +resembling the existing <i>Zonites</i>; whilst others (<i>Pupa +vetusta</i>, fig. 128) appear to be generically inseparable from + +<span style="float: left; margin: 4px; width: 217px; + font-size: smaller; text-align: justify;"> +<img src="images/fig128.jpg" width="209" height="324" alt="Fig. 128" /> +<br /> +Fig. 128.—<i>Pupa (Dendropupa) vetusta</i>, a Carboniferous +Land-snail from the Coal-measures of Nova Scotia. <i>a</i>, The +shell, of the natural size; <i>b</i>, The same, magnified; +<i>c</i>, Apex of the shell, enlarged; <i>d</i>, Portion of the +surface, enlarged. (After Dawson.) +</span> + +the "Chrysalis-shells" (<i>Pupa</i>) of the present day. All the +known forms—three in number—are of small size, and appear +to have been local in their distribution or in their preservation. +More important, however, than any of the preceding, are the +<i>Cephalopoda</i>, represented, as before, exclusively by the +chambered shells of the Tetrabranchiates. The older and simpler +type of these, with simple plain septa, and mostly a central +siphuncle, is represented by the straight conical shells of the +ancient genus Orthoceras, and the bow-shaped shells of the equally +ancient <i>Cyrtoceras</i>—some of the former attaining a +great size. The spirally-curved discoidal shells of the persistent +genus <i>Nautilus</i> are also not unknown, and some of these +likewise exhibit very considerable dimensions. Lastly, the more +complex family of the <i>AmmonitidĹ“</i>, +<a name="page_187"><span class="page">Page 187</span></a> +with lobed or angulated septa, and a dorsally-placed +siphuncle (situated on the convex side of the curved shells), now +for the first time commences to acquire a considerable prominence. +The principal representative of this group is the genus +<i>Goniatites</i> (fig. 129), which +commenced its existence in the Upper Silurian, is well represented +in the Devonian, and attains its maximum here. In this genus, +the shell is spirally curved, the septa are strongly lobed or +angulated, though not elaborately frilled as in the Ammonites, +and the siphuncle is dorsal. In addition to <i>Goniatites</i>, +the shells of true <i>Ammonites</i>, so characteristic of the +Secondary period, have been described by Dr Waagen as occurring +in the Carboniferous rocks of India. +</p> + +<div class="center"> +<table border="0" width="555"> +<tr><td> +<img src="images/fig129.jpg" width="551" height="590" alt="Fig. 129" /> +</td></tr> +<tr><td class="center"><span class="image"> +Fig. 129.—<i>Goniatites (Aganides) FossĹ“</i>. +Carboniferous Limestone. +</span></td></tr> +</table> +</div> + +<p class="indent"> +Coming finally to the <i>Vertebrata</i>, we have in the first +place to very briefly consider the Carboniferous <i>fishes</i>. +These are numerous; but, with the exception of the still dubious +"Conodonts," belong wholly to the groups of the <i>Ganoids</i> and +the <i>Placoids</i> (including under the former head remains which +perhaps are truly referable to the group of the <i>Dipnoi</i> or +Mud-fishes). Amongst the <i>Ganoids</i>, the singular buckler-headed +fishes of the Upper Silurian and Devonian (<i>CephalaspidĹ“</i>) +<a name="page_188"><span class="page">Page 188</span></a> +have apparently disappeared; and the principal +types of the Carboniferous belong to the groups respectively +represented at the present day by the Gar pike (<i>Lepidosteus</i>) +of the North American lakes, and the <i>Polypterus</i> of the rivers +of Africa. Of the former, the genera <i>PalĹ“oniscus</i> and +<i>Amblypterus</i> (fig. 130), with their small rhomboidal and + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 518px; + font-size: smaller; text-align: center;"> +<img src="images/fig130.jpg" width="510" height="225" alt="Fig. 130" /> +<br /> +Fig. 130.—<i>Amblypterus macropterus</i>. +</span> +</span> + +enamelled scales, and their strongly unsymmetrical tails, are +perhaps the most abundant. Of the latter, the most important are +species belonging to the genera <i>Megalichthys</i> and +<i>Rhizodus</i>, comprising large fishes, with rhomboidal scales, +unsymmetrical ("heterocercal") tails, and powerful conical teeth. +These fishes are sometimes said to be "sauroid," from their +presenting some Reptilian features in their organisation, and +they must have been the scourges of the Carboniferous seas. The +remains of <i>Placoid</i> fishes in the Carboniferous strata are +very numerous, but consist wholly of teeth and fin-spines, +referable to forms more or less closely allied to our existing +Port Jackson Sharks, Dog-fishes, and Rays. The teeth are of very +various shapes and sizes,—some with sharp, cutting edges +(<i>Petalodus, Cladodus</i>, &c.); others in the form of +broad crushing plates, adapted, like the teeth of the existing +Port Jackson Shark (<i>Cestracion Philippi</i>), for breaking +down the hard shells of Molluscs and Crustaceans. Amongst the +many kinds of these latter, the teeth of <i>Psammodus</i> and +<i>Cochliodus</i> (fig. 131) may be mentioned as specially +characteristic. The fin-spines are mostly similar to those so +common in the Devonian deposits, consisting of hollow defensive +spines implanted in front of the pectoral or other fins, usually +slightly curved, often superficially ribbed or sculptured, and +not uncommonly serrated or toothed. The genera <i>Ctenacanthus, +Gyracanthus, Homacanthus</i>, &c., have been founded for the +reception of these defensive weapons, some of which indicate +fishes of great size and predaceous habits. +</p> + +<p class="indent"> +<a name="page_189"><span class="page">Page 189</span></a> +In the Devonian rocks we meet with no +other remains of Vertebrated animals save fishes only; but the +Carboniferous deposits have yielded remains of the higher group + +<span style="float: right; margin: 4px; width: 263px; + font-size: smaller; text-align: center;"> +<img src="images/fig131.jpg" width="255" height="179" alt="Fig. 131" /> +<br /> +Fig. 131.—Teeth of <i>Cochliodus contortus</i>. +Carboniferous Limestone, Britain. +</span> + +of the <i>Amphibians</i>. This class, comprising our existing +Frogs, Toads, and Newts, stands to some extent in a position +midway between the class of the fishes and that of the true +reptiles, being distinguished from the latter by the fact +that its members invariably possess gills in their early +condition, if not throughout life; whilst they are separated +from the former by always possessing true lungs when adult, and +by the fact that the limbs (when present at all) are never in +the form of fins. The Amphibians, therefore, are all +water-breathers when young, and have respiratory organs adapted +for an aquatic mode of life; whereas, when grown up, they +develop lungs, and with these the capacity for breathing air +directly. Some of them, like the Frogs and Newts, lose their +gills altogether on attaining the adult condition; but others, +such as the living <i>Proteus</i> and <i>Menobranchus</i>, retain +their gills even after acquiring their lungs, and are thus fitted +indifferently for an aquatic or terrestrial existence. The name +of "Amphibia," though applied to the whole class, is thus not +precisely appropriate except to these last-mentioned forms +(Gr. <i>amphi</i>, both; <i>bios</i>, life). The Amphibians also +differ amongst themselves according as to whether they keep +permanently the long tail which they all possess when young +(as do the Newts and Salamanders), or lose this appendage when +grown up (as do the Frogs and Toads). Most of them have naked +skins, but a few living and many extinct forms have hard +structures in the shape of scales developed in the integument. +All of them have well-ossified skeletons, though some fossil +types are partially deficient in this respect; and all of them +which possess limbs at all have these appendages supported by +bones essentially similar to those found in the limbs of the +higher Vertebrates. All the Carboniferous Amphibians belong to +a group which has now wholly passed away—namely, that of +the <i>Labyrinthodonts</i>. In the marine strata which form the +base of the Carboniferous series these creatures have only been +recognised by their curious hand-shaped footprints, similar +<a name="page_190"><span class="page">Page 190</span></a> +in character to those which occur in the +Triassic rocks, and which will be subsequently spoken of under +the name of <i>Cheirotherium</i>. In the Coal-measures of +Britain, the continent of Europe, and North America, however, +many bones of these animals have been found, and we are now +tolerably well acquainted with a considerable number of forms. +All of them seem to have belonged to the division of Amphibians +in which the long tail of the young is permanently retained; and +there is evidence that some of them kept the gills also throughout +life. The skull is of the characteristic Amphibian type (fig. 132, +<i>a</i>), with two occipital condyles, and having its surface + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 531px; + font-size: smaller; text-align: justify;"> +<img src="images/fig132.jpg" width="523" height="436" alt="Fig. 132" /> +<br /> +Fig. 132.—<i>a</i>, Upper surface of the skull of +<i>Anthracosaurus Russelli</i>, one-sixth of the natural size: +<i>b</i>, Part of one of the teeth cut across, and highly magnified +to show the characteristic labyrinthine structure; <i>c</i>, One +of the integumentary shields or scales, one-half of the natural +size. Coal-measures, Northumberland. (After Atthey.) +</span> +</span> + +singularly pitted and sculptured; and the vertebræ are +hollowed out at both ends. The lower surface of the body was +defended by an armour of singular integumentary shields or scales +(fig. 132, <i>c</i>); and an extremely characteristic feature +(from which the entire group derives its name) is, that the walls +of the teeth are deeply folded, so as to give rise to an +extraordinary "labyrinthine" pattern when they are cut across +(fig. 132, <i>b</i>). Many of the Carboniferous Labyrinthodonts +are of no great size, some of them +<a name="page_191"><span class="page">Page 191</span></a> +very small, but others attain comparatively gigantic dimensions, +though all fall short in this respect of the huge examples of +this group which occur in the Trias. One of the largest, and +at the same time most characteristic, forms of the Carboniferous +series, is the genus <i>Anthracosaurus</i>, the skull of which +is here figured. +</p> + +<p class="indent"> +No remains of true Reptiles, Birds, or Quadrupeds have as yet +been certainly detected in the Carboniferous deposits in any part +of the world. It should, however, be mentioned, that Professor +Marsh, one of the highest authorities on the subject, has described +from the Coal-formation of Nova Scotia certain vertebræ which +he believes to have belonged to a marine reptile (<i>Eosaurus +Acadianus</i>), allied to the great <i>Ichthyosauri</i> of the +Lias. Up to this time no confirmation of this determination has +been obtained by the discovery of other and more unquestionable +remains, and it therefore remains doubtful whether these bones of +<i>Eosaurus</i> may not really belong to large Labyrinthodonts. +</p> + +<h4>LITERATURE.</h4> + +<p class="indent"> +The following list contains some of the more important of the +original sources of information to which the student of Carboniferous +rocks and fossils may refer:— +</p> + +<table border="0" cellspacing="0"> +<tr><td class="right" valign="top">(1)</td> + <td>'Geology of Yorkshire,' vol. ii.; 'The Mountain Limestone + District.' John Phillips.</td></tr> +<tr><td class="right" valign="top">(2)</td> + <td>'Siluria.' Sir Roderick Murchison.</td></tr> +<tr><td class="right" valign="top">(3)</td> + <td>'Memoirs of the Geological Survey of Great Britain and + Ireland.'</td></tr> +<tr><td class="right" valign="top">(4)</td> + <td>'Geological Report on Londonderry,' &c. Portlock.</td></tr> +<tr><td class="right" valign="top">(5)</td> + <td>'Acadian Geology.' Dawson.</td></tr> +<tr><td class="right" valign="top">(6)</td> + <td>'Geology of Iowa,' vol. i. James Hall.</td></tr> +<tr><td class="right" valign="top">(7)</td> + <td>'Reports of the Geological Survey of Illinois' (Geology and + Palæontology). Meek, Worthen, &c.</td></tr> +<tr><td class="right" valign="top">(8)</td> + <td>'Reports of the Geological Survey of Ohio' (Geology and + Palæontology). Newberry, Cope, Meek, Hall, &c.</td></tr> +<tr><td class="right" valign="top">(9)</td> + <td>'Description des Animaux fossiles qui se trouvent dans le + Terrain Carbonifère de la Belgique,' 1843; with subsequent + monographs on the genera <i>Productus</i> and <i>Chonetes</i>, + on <i>Crinoids</i>, on <i>Corals</i>, &c. De Koninck.</td></tr> +<tr><td class="right" valign="top">(10)</td> + <td>'Synopsis of the Carboniferous Fossils of Ireland.' + M'Coy.</td></tr> +<tr><td class="right" valign="top">(11)</td> + <td>'British Palæozoic Fossils.' M'Coy.</td></tr> +<tr><td class="right" valign="top">(12)</td> + <td>'Figures of Characteristic British Fossils.' Baily.</td></tr> +<tr><td class="right" valign="top">(13)</td> + <td>'Catalogue of British Fossils.' Morris.</td></tr> +<tr><td class="right" valign="top">(14)</td> + <td>'Monograph of the Carboniferous Brachiopoda of Britain' + (Palæontographical Society). Davidson.</td></tr> +<tr><td class="right" valign="top">(15)</td> + <td>'Monograph of the British Carboniferous Corals' + (Palæontographical Society). Milne-Edwards and + Haime.</td></tr> +<tr><td class="right" valign="top">(16)</td> + <td>'Monograph of the Carboniferous Bivalve Entomostraca of + Britain' (Palæontographical Society). Rupert Jones, + Kirkby, and George S. Brady.</td></tr> +<tr><td class="right" valign="top"> +<a name="page_192"><span class="page">Page 192</span></a> + (17)</td> + <td>'Monograph of the Carboniferous Foraminifera of Britain' + (Palæontographical Society). H. B. Brady.</td></tr> +<tr><td class="right" valign="top">(18)</td> + <td>"On the Carboniferous Fossils of the West of + Scotland"—'Trans. Geol. Soc.,' of Glasgow, vol. iii., + Supplement. Young and Armstrong.</td></tr> +<tr><td class="right" valign="top">(19)</td> + <td>'Poissons Fossiles.' Agassiz.</td></tr> +<tr><td class="right" valign="top">(20)</td> + <td>"Report on the Labyrinthodonts of the + Coal-measures"—'British Association Report,' 1873. L. C. + Miall.</td></tr> +<tr><td class="right" valign="top">(21)</td> + <td>'Introduction to the Study of Palæontological + Botany.' John Hutton Balfour.</td></tr> +<tr><td class="right" valign="top">(22)</td> + <td>'TraitĂ© de PalĂ©ontologie + VĂ©gĂ©tale.' Schimper.</td></tr> +<tr><td class="right" valign="top">(23)</td> + <td>'Fossil Flora.' Lindley and Hutton.</td></tr> +<tr><td class="right" valign="top">(24)</td> + <td>'Histoire des VĂ©gĂ©taux Fossiles.' + Brongniart.</td></tr> +<tr><td class="right" valign="top">(25)</td> + <td>'On Calamites and Calamodendron' (Monographs of the + Palæontographical Society). Binney.</td></tr> +<tr><td class="right" valign="top">(26)</td> + <td>'On the Structure of Fossil Plants found in the + Carboniferous Strata' (Palæontographical Society). + Binney.</td></tr> +</table> + +<p class="indent"> +Also numerous memoirs by Huxley, Davidson, Martin Duncan, Professor +Young, John Young, R. Etheridge, jun., Baily, Carruthers, Dawson, +Binney, Williamson, Hooker, Jukes, Geikie, Rupert Jones, Salter, +and many other British and foreign observers. +</p> + +<h3>CHAPTER XIV.</h3> + +<p class="subtitle"> +THE PERMIAN PERIOD. +</p> + +<p class="indent"> +The Permian formation closes the long series of the Palæozoic +deposits, and may in some respects be considered as a kind of +appendix to the Carboniferous system, to which it cannot be compared +in importance, either as regards the actual bulk of its sediments +or the interest and variety of its life-record. Consisting, as it +does, largely of red rocks—sandstones and marls—for the +most part singularly destitute of organic remains, the Permian +rocks have been regarded as a lacustrine or fluviatile deposit; +but the presence of well-developed limestones with indubitable +marine remains entirely negatives this view. It is, however, +not improbable that we are presented in the Permian formation, +as known to us at present, with a series of sediments laid down +in inland seas of great extent, due to the subsidence over large +areas of the vast land-surfaces of the Coal-measures. This view, +at any rate, would explain some of the more puzzling physical +characters of the formation, and would not be definitely negatived +by any of its fossils. +</p> + +<p class="indent"> +A large portion of the Permian series, as already remarked, consists +of sandstones and marls, deeply reddened by peroxide +<a name="page_193"><span class="page">Page 193</span></a> +of iron, and often accompanied by beds of gypsum or deposits +of salt. In strata of this nature few or no fossils are found; but +their shallow-water origin is sufficiently proved by the presence of +the footprints of terrestrial animals, accompanied in some cases by +well-defined "ripple-marks." Along with these are occasionally found +massive breccias, holding larger or smaller blocks derived from the +older formations; and these have been supposed to represent an old +"boulder-clay," and thus to indicate the prevalence of an arctic +climate. Beds of this nature must also have been deposited in +shallow water. In all regions, however, where the Permian formation +is well developed, one of its most characteristic members is a +Magnesian limestone, often highly and fantastically concretionary, +but containing numerous remains of genuine marine animals, and +clearly indicating that it was deposited beneath a moderate depth +of salt water. +</p> + +<p class="indent"> +It is not necessary to consider here whether this formation can +be retained as a distinct division of the geological series. The +name of <i>Permian</i> was given to it by Sir Roderick Murchison, +from the province of Perm in Russia, where rocks of this age are +extensively developed. Formerly these rocks were grouped with +the succeeding formation of the Trias under the common name of +"New Red Sandstone." This name was given them because they contain +a good deal of red sandstone, and because they are superior to the +Carboniferous rocks, while the Old Red Sandstone is inferior. +Nowadays, however, the term "New Red Sandstone" is rarely employed, +unless it be for red sandstones and associated rocks, which are +seen to overlie the Coal-measures, but which contain no fossils by +which their exact age may be made out. Under these circumstances, +it is sometimes convenient to employ the term "New Red Sandstone." +The New Red, however, of the older geologists, is now broken up +into the two formations of the Permian and Triassic rocks—the +former being usually considered as the top of the Palæozoic +series, and the latter constituting the base of the Mesozoic. +</p> + +<p class="indent"> +In many instances, the Permian rocks are seen to repose unconformably +upon the underlying Carboniferous, from which they can in addition +be readily separated by their lithological characters. In other +instances, however, the Coal-measures terminate upwards in red +rocks, not distinguishable by their mineral characters from the +Permian; and in other cases no physical discordance between the +Carboniferous and Permian strata can be detected. As a general +rule, also, the Permian rocks appear to pass upwards conformably +into the +<a name="page_194"><span class="page">Page 194</span></a> +Trias. The division, therefore, +between the Permian and Triassic rocks, and consequently between +the Palæozoic and Mesozoic series, is not founded upon any +conspicuous or universal physical break, but upon the difference +in life which is observed in comparing the marine animals of the +Carboniferous and Permian with those of the Trias. It is to be +observed, however, that this difference can be solely due to the +fact that the Magnesian Limestone of the Permian series presents +us with only a small, and not a typical, portion of the marine +deposits which must have been accumulated in some area at present +unknown to us during the period which elapsed between the +formation of the great marine limestones of the Lower +Carboniferous and the open-sea and likewise calcareous sediments +of the Middle Trias. +</p> + +<p class="indent"> +The Permian rocks exhibit their most typical features in Russia +and Germany, though they are very well developed in parts of +Britain, and they occur in North America. When well developed, +they exhibit three main divisions: a lower set of sandstones, +a middle group, generally calcareous, and an upper series of +sandstones, constituting respectively the Lower, Middle, and Upper +Permians. +</p> + +<p class="indent"> +In Russia, Germany, and Britain, the Permian rocks consist of +the following members:— +</p> + +<p class="indent"> +1. The <i>Lower Permians</i>, consisting mainly of a great series +of sandstones, of different colours, but usually red. The base +of this series is often constituted by massive breccias with +included fragments of the older rocks, upon which they may happen +to repose; and similar breccias sometimes occur in the upper +portion of the series as well. The thickness of this group varies +a good deal, but may amount to 3000 or 4000 feet. +</p> + +<p class="indent"> +2. The <i>Middle Permians</i>, consisting, in their typical +development, of laminated marls, or "marl-slate," surmounted +by beds of magnesian limestone (the "Zechstein" of the German +geologists). Sometimes the limestones are degenerate or wholly +deficient, and the series may consist of sandy shales and gypsiferous +clays. The magnesian limestone, however, of the Middle Permians +is, as a rule, so well marked a feature that it was long spoken +of as <i>the</i> Magnesian Limestone. +</p> + +<p class="indent"> +3. The <i>Upper Permians</i>, consisting of a series of sandstones +and shales, or of red or mottled marls, often gypsiferous, and +sometimes including beds of limestone. +</p> + +<p class="indent"> +In North America, the Permian rocks appear to be confined to the +region west of the Mississippi, being especially well developed +in Kansas. Their exact limits have not as yet been +<a name="page_195"><span class="page">Page 195</span></a> +made out, and their total thickness is not more than +a few hundred feet. They consist of sandstones, conglomerates, +limestones, marls, and beds of gypsum. +</p> + +<p class="indent"> +The following diagrammatic section shows the general sequence of +the Permian deposits in the north of England, where the series +is extensively developed (fig. 133):— +</p> + +<div class="center"> +<table border="0" width="527"> +<tr><td class="center"><span class="image"> +GENERALISED SECTION OF THE PERMIAN ROCKS IN THE NORTH OF ENGLAND. +</span></td></tr> +<tr><td class="center"><span class="image"> +Fig. 133. +</span></td></tr> +<tr><td> +<img src="images/fig133.jpg" width="523" height="600" alt="Fig. 133" /> +</td></tr> +</table> +</div> + +<p class="indent"> +The record of the <i>life</i> of the Permian period is but a scanty +one, owing doubtless to the special peculiarities of such of the +<a name="page_196"><span class="page">Page 196</span></a> +deposits of this age with which we are as yet +acquainted. Red rocks are, as a general rule, more or less completely +unfossiliferous, and sediments of this nature are highly +characteristic of the Permian. Similarly, magnesian limestones are +rarely as highly charged with organic remains as is the case with +normal calcareous deposits, especially when they have been subjected +to concretionary action, as is observable to such a marked extent in +the Permian limestones. Nevertheless, much interest is attached to +the organic remains, as marking a kind of transition-period between +the Palæozoic and Mesozoic epochs. +</p> + +<p class="indent"> +The <i>plants</i> of the Permian period, as a whole, have a +distinctly Palæozoic aspect, and are far more nearly allied to +those of the Coal-measures than they are to those of the earlier +Secondary rocks; though the Permian <i>species</i> are mostly +distinct from the Carboniferous, and there are some new genera. +Thus, we find species of <i>Lepidodendron, Calamites, Equisetites, +Asterophyllites, Annularia</i>, and other highly characteristic +Carboniferous genera. On the other hand, the <i>Sigillariods</i> +of the Coal seem to have finally disappeared at the close of the +Carboniferous period. Ferns are abundant in the Permian rocks, +and belong for the most part to the well-known Carboniferous +genera <i>Alethopteris, Neuropteris, Sphenopteris</i>, and +<i>Pecopteris</i>. There are also Tree-ferns referable to the +ancient genus <i>Psaronius</i>. The <i>Conifers</i> of the Permian +period are numerous, and belong in part to Carboniferous genera. +A characteristic genus, however, is <i>Walchia</i> (fig. 134), + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 493px; + font-size: smaller; text-align: center;"> +<img src="images/fig134.jpg" width="485" height="240" alt="Fig. 134" /> +<br /> +Fig. 134.—<i>Walchia piniformis</i>, from the Permian of +Saxony, <i>a</i>, Branch; <i>b</i>, Twig, (After Gutbier.) +</span> +</span> + +distinguished by its lax short leaves. This genus, though not +exclusively Permian, is mainly so, the best-known species being +the <i>W. Piniformis</i>. Here, also, we meet with Conifers which +produce true cones, and which differ, therefore, in an important +degree from the +<a name="page_197"><span class="page">Page 197</span></a> +Taxoid Conifers of the +Coal-measures. Besides <i>Walchia</i>, a characteristic form of +these is the <i>Ullmania selaginoides</i>, which occurs in the +Magnesian Limestone of Durham, the Middle Permian of Westmorland, +and the "Kupfer-schiefer" of Germany. The group of the +<i>Cycads</i>, which we shall subsequently find to be so +characteristic of the vegetation of the Secondary period, is, on +the other hand, only doubtfully represented in the Permian +deposits by the singular genus <i>NĹ“ggerathia</i>. +</p> + +<p class="indent"> +The <i>Protozoans</i> of the Permian rocks are few in number, and +for the most part imperfectly known. A few <i>Foraminifera</i> +have been obtained from the Magnesian Limestone of England, and +the same formation has yielded some ill-understood Sponges. It +does not seem, however, altogether impossible that some of the +singular "concretions" of this formation may ultimately prove to +have an organic structure, though others would appear to be clearly +of purely inorganic origin. From the Permian of Saxony, Professor +Geinitz has described two species of <i>Spongillopsis</i>, which +he believes to be most nearly allied to the existing fresh-water +Sponges (<i>Spongilla</i>). This observation has an interest as +bearing upon the mode of deposition and origin of the Permian +sediments. +</p> + +<p class="indent"> +The <i>CĹ“lenterates</i> are represented in the Permian by +but a few Corals. These belong partly to the <i>Tabulate</i> and +partly to the <i>Rugose</i> division; but the latter great group, +so abundantly represented in Silurian, Devonian, and Carboniferous +seas, is now extraordinarily reduced in numbers, the British +strata of this age yielding only species of the single genus +<i>PolycĹ“lia</i>. So far, therefore, as at present known, all +the characteristic genera of the Rugose Corals of the Carboniferous +had become extinct before the deposition of the limestones of +the Middle Permian. +</p> + +<p class="indent"> +The <i>Echinoderms</i> are represented by a few <i>Crinoids</i>, +and by a Sea-urchin belonging to the genus <i>Eocidaris</i>. The +latter genus is nearly allied to the <i>ArchĹ“ocidaris</i> +of the Carboniferous, so that this Permian form belongs to a +characteristically Palæozoic type. +</p> + +<p class="indent"> +A few <i>Annelides</i> (<i>Spirorbis, Vermilia</i>, &c.) have +been described, but are of no special importance. Amongst the +<i>Crustaceans</i>, however, we have to note the total absence +of the great Palæozoic group of the <i>Trilobites</i>; whilst +the little <i>Ostracoda</i> and <i>Phyllopods</i> still continue +to be represented. We have also to note the first appearance +here of the "Short-tailed" Decapods or Crabs (<i>Brachyura</i>), +the highest of all the groups of <i>Crustacea</i>, in the person +of <i>Hemitrochiscus paradoxus</i>, an extremely minute Crab +from the Permian of Germany. +</p> + +<p class="indent"> +<a name="page_198"><span class="page">Page 198</span></a> +Amongst the <i>Mollusca</i>, the remains of <i>Polyzoa</i> may +fairly be said to be amongst the most abundant of all the fossils +of the Permian formation, The principal forms of these are the +fronds of the Lace-corals (<i>Fenestella, Retepora</i>, and +<i>Synocladia</i>), which are very abundant in the Magnesian +Limestone of the north of England, and belong to various highly +characteristic species (such as <i>Fenestella retiformis, Retepora +Ehrenbergi</i>, and <i>Synocladia virgulacea</i>). The +<i>Brachiopoda</i> are also represented in moderate numbers in the +Permian. Along with species of the persistent genera <i>Discina, +Crania</i>, and <i>Lingula</i>, we still meet with representatives of +the old groups <i>Spirifera, Athyris</i>, and <i>Streptorhynchus</i>; +and the Carboniferous <i>ProductĹ“</i> yet survive under +well-marked and characteristic types, though in much-diminished +numbers. The species of Brachiopods here figured (fig. 135) are +characteristic of the Magnesian Limestone in Britain and of the + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 495px; + font-size: smaller; text-align: center;"> +<img src="images/fig135.jpg" width="487" height="232" alt="Fig. 135" /> +<br /> +Fig. 135.—Brachiopods of the Permian formation. <i>a, +Producta horrida; b, Lingula Credneri; c, Terebratula elongata; +d</i> and <i>e, Camarophoria globulina</i>. (After King.) +</span> +</span> + +corresponding strata on the Continent. Upon the whole, the most +characteristic Permian <i>Brachiopods</i> belong to the genera +<i>Producta, Strophalosia</i>, and <i>Camarophoria</i>. +</p> + +<p class="indent"> +The <i>Bivalves</i> (<i>Lamellibranchiata</i>) have a tolerably +varied development in the Permian rocks; but nearly all the old +types, except some of those which occur in the Carboniferous, have +now disappeared. The principal Permian Bivalves belong to the groups +of the Pearl Oysters (<i>AviculidĹ“</i>) and the +<i>TrigoniadĹ“</i>, represented by genera such as <i>Bakewellia</i> +and <i>Schizodus</i>; the true Mussels (<i>MytilidĹ“</i>), +represented by species which have been referred to <i>Mytilus</i> +itself; and the Arks (<i>ArcadĹ“</i>), represented by species +of the genera <i>Arca</i> (fig. 136) and <i>Byssoarca</i>. The first +and last of these three families have a very ancient origin; but +the family of the <i>TrigoniadĹ“</i>, though +<a name="page_199"><span class="page">Page 199</span></a> +feebly represented at the present day, is one which attained +its maximum development in the Mesozoic period. +</p> + +<p class="indent"> +The <i>Univalves</i> (<i>Gasteropoda</i>) are rare, and do not +demand special notice. It may be observed, however, that the + +<span style="float: right; margin: 4px; width: 258px; + font-size: smaller; text-align: center;"> +<img src="images/fig136.jpg" width="250" height="214" alt="Fig. 136" /> +<br /> +Fig. 136.—<i>Arca antiqua</i>. Permian. +</span> + +Palæozoic genera <i>Euomphalus, Murchisonia, Loxonema</i>, +and <i>Macrocheilus</i> are still in existence, together with the +persistent genus <i>Pleurotomaria</i>. <i>Pteropods</i> of the +old genera <i>Theca</i> and <i>Conularia</i> have been discovered; +but the first of these characteristically Palæozoic types +finally dies out here, and the second only survives but a short time +longer. Lastly, a few <i>Cephalopods</i> have been found, still +wholly referable to the Tetrabranchiate group, and belonging +to the old genera <i>Orthoceras</i> and <i>Cyrtoceras</i> and +the long-lived <i>Nautilus</i>. +</p> + +<p class="indent"> +Amongst <i>Vertebrates</i>, we meet in the Permian period not +only with the remains of Fishes and Amphibians, but also, for +the first time, with true Reptiles. The <i>Fishes</i> are mainly +<i>Ganoids</i>, though there are also remains of a few Cestraciont + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 375px; + font-size: smaller; text-align: center;"> +<img src="images/fig137.jpg" width="367" height="250" alt="Fig. 137" /> +<br /> +Fig. 137.—<i>Platysomus gibbosus</i>, a "heterocercal" +Ganoid, from the Middle Permian of Russia. +</span> +</span> + +Sharks. Not only are the <i>Ganoids</i> still the predominant group +of Fishes, but all the known forms possess the unsymmetrical +("heterocercal") tail which is so characteristic of the +Palæozoic Ganoids. Most of the remains of the Permian Fishes +have been obtained from the "Marl-slate" of Durham and the +corresponding "Kupfer-schiefer" of Germany, on the horizon +<a name="page_200"><span class="page">Page 200</span></a> +of the Middle Permian; and the principal genera of the Ganoids +are <i>PalĹ“oniscus</i> and <i>Platysomus</i> (fig. 137). +</p> + +<p class="indent"> +The <i>Amphibians</i> of the Permian period belong principally +to the order of the <i>Labyrinthodonts</i>, which commenced to +be represented in the Carboniferous, and has a large development +in the Trias. Under the name, however, of <i>PalĹ“osiren +Beinerti</i>, Professor Geinitz has described an Amphibian from +the Lower Permian of Germany, which he believes to be most nearly +allied to the existing "Mud-eel" (<i>Siren lacertina</i>) of +North America, and therefore to be related to the Newts and +Salamanders (<i>Urodela</i>). +</p> + +<p class="indent"> +Finally, we meet in the Permian deposits with the first undoubted +remains of true <i>Reptiles</i>. These are distinguished, as a +class, from the <i>Amphibians</i>, by the fact that they are +air-breathers throughout the whole of their life, and therefore +are at no time provided with gills; whilst they are exempt from +that metamorphosis which all the <i>Amphibia</i> undergo in early +life, consequent upon their transition from an aquatic to a more +or less purely aerial mode of respiration. Their skeleton is well +ossified; they usually have horny or bony plates, singly or in +combination, developed in the skin; and their limbs (when present) +are never either in the form of <i>fins</i> or <i>wings</i>, +though sometimes capable of acting in either of these capacities, +and liable to great modifications of form and structure. Though +there can be no doubt whatever as to the occurrence of genuine +Reptiles in deposits of unquestionable Permian age, there is +still uncertainty as to the precise number of types which may +have existed at this period. This uncertainty arises partly from +the difficulty of deciding in all cases. whether a given bone +be truely Labyrinthodont or Reptilian, but more especially from +the confusion which exists at present between the Permian and +the overlying Triassic deposits. Thus there are various deposits +in different regions which have yielded the remains of Reptiles, +and which cannot in the meanwhile be definitely referred either +to the Permian series or to the Trias by clear stratigraphical +or palæontological evidence. All that can be done in such +cases is to be guided by the characters of the Reptiles themselves, +and to judge by their affinities to remains from known Triassic +or Permian rocks to which of these formations the beds containing +them should be referred; but it is obvious that this method of +procedure is seriously liable to lead to error. In accordance, +however, with this, the only available mode of determination +in some cases, the remains of <i>Thecodontosaurus</i> and +<i>Palæosaurus</i> discovered in the dolomitic conglomerates +<a name="page_201"><span class="page">Page 201</span></a> +near Bristol will be considered as Triassic, thus leaving +<i>Protorosaurus</i>[20] as the principal and most important + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 552px; + font-size: smaller; text-align: center;"> +<img src="images/fig138.jpg" width="546" height="608" alt="Fig. 138" /> +<br /> +Fig. 138.—<i>Protorosaurus Speneri</i>, Middle Permian, +Thuringia, reduced in size. (After Von Meyer.) [Copied from +Dana. +</span> +</span> + +representative of the Permian Reptiles.[21] The type-species +of the genus <i>Protorusaurus</i> is the <i>P. Speneri</i>(fig. +138) of the "Kupfer-schiefer" of +<a name="page_202"><span class="page">Page 202</span></a> +Thuringia, but other allied +species have been detected in the Middle Permian of Germany and +the north of England. This Reptile attained a length of from +three to four feet; and it has been generally referred to the +group of the Lizards (<i>Lacertilia</i>), to which it is most +nearly allied in its general structure, at the same time that it +differs from all existing members of this group in the fact that +its numerous conical and pointed teeth were implanted in distinct +sockets in the jaws—this being a Crocodilian character. In +other respects, however, <i>Protorosaurus</i> approximates closely +to the living Monitors (<i>VaranidĹ“</i>); and the fact that +the bodies of the vertebræ are slightly cupped or hollowed +out at the ends would lead to the belief that the animal was aquatic +in its habits. At the same time, the structure of the hind-limbs +and their bony supports proves clearly that it must have also +possessed the power of progression upon the land. Various other +Reptilian bones have been described from the Permian formation, of +which some are probably really referable to Labyrinthodonts, whilst +others are regarded by Professor Owen as referable to the order of +the "Theriodonts," in which the teeth are implanted in sockets, +and resemble those of carnivorous quadrupeds in consisting of +three groups in each jaw (namely, incisors, canines, and molars). +Lastly, in red sandstones of Permian age in Dumfriesshire have +been discovered the tracks of what would appear to have been +<i>Chelonians</i> (Tortoises and Turtles); but it would not be +safe to accept this conclusion as certain upon the evidence of +footprints alone. The <i>Chelichnus Duncani</i>, however, described +by Sir William Jardine in his magnificent work on the 'Ichnology +of Annandale,' bears a great resemblance to the track of a Turtle. +</p> + +<p class="footnote"> +[Footnote 20: Though commonly spelt as above, it is probable +that the name of this Lizard was really intended to have been +<i>Proterosaurus</i>—from the Greek <i>proteros</i>, first; +and <i>saura</i>, lizard: and this spelling is followed by many +writers.] +</p> + +<p class="footnote"> +[Footnote 21: In an extremely able paper upon the subject (Quart. +Journ. Geol. Soc., vol. xxvi.), Mr Etheridge has shown that there +are good physical grounds for regarding the dolomitie conglomerate +of Bristol as of Triassic age, and as probably corresponding in +time with the Muschelkalk of the Continent.] +</p> + +<p class="indent"> +No remains of Birds or Quadrupeds have hitherto been detected +in deposits of Permian age. +</p> + +<h4>LITERATURE.</h4> + +<p class="indent"> +The following works may be consulted by the student with regard +to the Permian formation and its fossils:— +</p> + +<table border="0" cellspacing="0"> +<tr><td class="right" valign="top">(1)</td> + <td>"On the Geological Relations and Internal Structure of the + Magnesian Limestone and the Lower Portions of the New Red + Sandstone Series, &c."—'Trans. Geol. Soc.,' ser. 2, + vol. iii. Sedgwick.</td></tr> +<tr><td class="right" valign="top">(2)</td> + <td>'The Geology of Russia in Europe.' Murchison, De Verneuil, + and Von Keyserling.</td></tr> +<tr><td class="right" valign="top">(3)</td> + <td>'Siluria,' Murchison.</td></tr> +<tr><td class="right" valign="top">(4)</td> + <td>'Permische System in Sachsen.' Geinitz and + Gutbier.</td></tr> +<tr><td class="right" valign="top">(5)</td> + <td>'Die Versteinerungen des Deutschen Zechsteingebirges,' + Geinitz.</td></tr> +<tr><td class="right" valign="top">(6)</td> + <td>'Die Animalischen Ueberreste der Dyas.' + Geinitz.</td></tr> +<tr><td class="right" valign="top"> +<a name="page_203"><span class="page">Page 203</span></a> + (7)</td> + <td>'Monograph of the Permian Fossils of England' + (Palæontographical Society). King.</td></tr> +<tr><td class="right" valign="top">(8)</td> + <td>'Monograph of the Permian Brachiopoda of Britain' +(Palæontographical Society). Davidson.</td></tr> +<tr><td class="right" valign="top">(9)</td> + <td>"On the Permian Rocks of the North-West of England and + their Extension into Scotland"—'Quart. Journ. Geol. + Soc.,' vol. xx. Murchison and Harkness.</td></tr> +<tr><td class="right" valign="top">(10)</td> + <td>'Catalogue of the Fossils of the Permian System of the + Counties of Northumberland and Durham.' Howse.</td></tr> +<tr><td class="right" valign="top">(11)</td> + <td>'Petrefacta Germaniæ.' Goldfuss.</td></tr> +<tr><td class="right" valign="top">(12)</td> + <td>'Beiträge zur Petrefaktenkunde.' Munster.</td></tr> +<tr><td class="right" valign="top">(13)</td> + <td>'Ein Beitrag zur Palæontologie des Deutschen + Zechsteingebirges.' Von Schauroth.</td></tr> +<tr><td class="right" valign="top">(14)</td> + <td>'Saurier aus dem Kupfer-schiefer der + Zechstein-formation.' Von Meyer.</td></tr> +<tr><td class="right" valign="top">(15)</td> + <td>'Manual of Palæontology.' Owen.</td></tr> +<tr><td class="right" valign="top">(16)</td> + <td>'Recherches sur les Poissons Fossiles.' + Agassiz.</td></tr> +<tr><td class="right" valign="top">(17)</td> + <td>'Ichnology of Annandale.' Sir William Jardine.</td></tr> +<tr><td class="right" valign="top">(18)</td> + <td>'Die Fossile Flora der Permischen Formation.' + GĹ“ppert.</td></tr> +<tr><td class="right" valign="top">(19)</td> + <td>'Genera et Species Plantarum Fossilium.' Unger.</td></tr> +<tr><td class="right" valign="top">(20)</td> + <td>"On the Red Rocks of England of older Date than the + Trias"—'Quart. Journ. Geol. Soc.,' vol. xxvii. + Ramsay.</td></tr> +</table> + +<h3>CHAPTER XV.</h3> + +<p class="subtitle"> +THE TRIASSIC PERIOD. +</p> + +<p class="indent"> +We come now to the consideration of the great <i>Mesozoic</i>, or +Secondary series of formations, consisting, in ascending order, +of the Triassic, Jurassic, and Cretaceous systems. The Triassic +group forms the base of the Mesozoic series, and corresponds +with the higher portion of the New Red Sandstone of the older +geologists. Like the Permian rocks, and as implied by its name, +the <i>Trias</i> admits of a subdivision into three groups—a +Lower, Middle, and Upper Trias. Of these sub-divisions the middle +one is wanting in Britain; and all have received German names, +being more largely and typically developed in Germany than in any +other country. Thus, the Lower Trias is known as the <i>Bunter +Sandstein</i>; the Middle Trias is called the <i>Muschelkalk</i>; +and the Upper Trias is known as the <i>Keuper</i>. +</p> + +<p class="indent"> +I. The lowest division of the Trias is known as the <i>Bunter +Sandstein</i> (the <i>Grès bigarrĂ©</i> of the French), +from the generally variegated colours of the beds which compose it +(German, <i>bunt</i>, variegated). The Bunter Sandstein of the +continent of Europe consists of red and white sandstones, with red +<a name="page_204"><span class="page">Page 204</span></a> +clays, and thin limestones, the whole +attaining a thickness of about 1500 feet. The term "marl" is very +generally employed to designate the clays of the Lower and Upper +Trias; but the term is inappropriate, as they may contain no lime, +and are therefore not always genuine marls. In Britain the Bunter +Sandstein consists of red and mottled sandstones, with +unconsolidated conglomerates, or "pebble-beds," the whole having a +thickness of 1000 to 2000 feet. The Bunter Sandstein, as a rule, +is very barren of fossils. +</p> + +<p class="indent"> +II. The Middle Trias is not developed in Britain, but it is largely +developed in Germany, where it constitutes what is known as the +<i>Muschelkalk</i> (Germ. <i>Muschel</i>, mussel; <i>kalk</i>, +limestone), from the abundance of fossil shells which it contains. +The Muschelkalk (the <i>Calcaire coquillier</i> of the French) +consists of compact grey or yellowish limestones, sometimes +dolomitic, and including occasional beds of gypsum and rock-salt. +</p> + +<p class="indent"> +III. The Upper Trias, or <i>Keuper</i> (the <i>Marnes +irisĂ©es</i> of the French), as it is generally called, +occurs in England; but is not so well developed as it is in +Germany. In Britain, the Keuper is 1000 feet or more in thickness, +and consists of white and brown sandstones, with red marls, the +whole topped by red clays with rock-salt and gypsum. +</p> + +<p class="indent"> +The Keuper in Britain is extremely unfossiliferous; but it passes +upwards with perfect conformity into a very remarkable group of +beds, at one time classed with the Lias, and now known under +the names of the Penarth beds (from Penarth, in Glamorganshire), +the Rhætic beds (from the Rhætic Alps), or the <i>Avicula +contorta</i> beds (from the occurrence in them of great numbers +of this peculiar Bivalve). These singular beds have been variously +regarded as the highest beds of the Trias, or the lowest beds of +the Lias, or as an intermediate group. The phenomena observed +on the Continent, however, render it best to consider them as +Triassic, as they certainly agree with the so-called Upper St +Cassian or Kössen beds which form the top of the Trias in the +Austrian Alps. +</p> + +<p class="indent"> +The Penarth beds occur in Glamorganshire, Gloucestershire, +Warwickshire, Staffordshire, and the north of Ireland; and they +generally consist of a small thickness of grey marls, white +limestones, and black shales, surmounted conformably by the lowest +beds of the Lias. The most characteristic fossils which they +contain are the three Bivalves <i>Cardium RhĹ“ticum, Avicula +contorta</i>, and <i>Pecten Valoniensis</i>; but they have yielded +many other fossils, amongst which the most important are the +remains of Fishes and small Mammals (<i>Microlestes</i>). +</p> + +<p class="indent"> +In the Austrian Alps the Trias terminates upwards in an +<a name="page_205"><span class="page">Page 205</span></a> +extraordinary series of fossiliferous beds, replete with +marine fossils. Sir Charles Lyell gives the following table of +these remarkable deposits:— +</p> + +<p class="center"> +<i>Strata below the Lias in the Austrian Alps, in descending +order.</i> +</p> +<table border="1" cellspacing="0" cellpadding="4"> +<tr><td> + <table border="0" cellspacing="0"> + <tr><td valign="top">1.</td> + <td valign="top">Koessen beds.<br/>(Synonyms, Upper St Cassian + beds of Escher and Merian.</td></tr> + </table> +</td><td colspan="2"> + Grey and black limestone, with calcareous marls having a thickness + of about 50 feet. Among the fossils, Brachiopoda very numerous; + some few species common to the genuine Lias; many peculiar. + <i>Avicula contorta, Pecten Valoniensis, Cardium RhĹ“ticum, + Avicula inĹ“quivalvis, Spirifer Münsteri</i>, Dav. + Strata containing the above fossils alternate with the Dachstein + beds, lying next below. +</td></tr> + +<tr><td> + <table border="0" cellspacing="0"> + <tr><td valign="top">2.</td> + <td valign="top">Dachstein beds.</td></tr> + </table> +</td><td colspan="2"> + White or greyish limestone, often in beds three or four feet + thick. Total thickness of the formation above 2000 feet. Upper + part fossiliferous, with some strata composed of corals + (<i>Lithodendron</i>.) Lower portion without fossils. Among the + characteristic shells are <i>Hemicardium</i> <i>Wulfeni, + Megalodon triqueler</i>, and other large bivalves. +</td></tr> + +<tr><td> + <table border="0" cellspacing="0"> + <tr><td valign="top">3.</td> + <td valign="top">Hallstadt beds<br/>(or St Cassian)</td></tr> + </table> +</td><td colspan="2"> + Red, pink, or white marbles, from 800 to 1000 feet in thickness, + containing more than 800 species of marine fossils, for the most + part mollusca. Many species of <i>Orthoceras</i>. True + <i>Ammonites</i>, besides <i>Ceratites</i> and <i>Goniatites, + Belemnites</i> (rare), <i>Porcellia, Pleurotomania, Trochus, + Monotis salinaria</i>, &c. +</td></tr> + +<tr><td> + <table border="0" cellspacing="0"> + <tr><td valign="top" rowspan="2">4.</td> + <td valign="top"><i>A</i>.</td> + <td valign="top">Guttenstein beds.</td></tr> + <tr><td valign="top"><i>B</i>.</td> + <td valign="top">Werfen beds, base of Upper Trias?<br/>Lower + Trias of some geologists.</td></tr> + </table> +</td><td> + <table border="0" cellspacing="0"> + <tr><td valign="top"><i>A</i>.</td> + <td valign="top">Black and grey limestone 150 feet thick, + alternating with the underlying Werfen beds.</td></tr> + <tr><td valign="top"><i>B</i>.</td> + <td valign="top">Red and green shale and sandstone, with salt + and gypsum.</td></tr> + </table> +</td><td> + Among the fossils are <i>Ceratites cassianus, Myacites fassaensis, + Naticella costata</i>, &c. +</td></tr> +</table> + +<p class="indent"> +In the United States, rocks of Triassic age occur in several +areas between the Appalachians and the Atlantic seaboard; but +they show no such triple division as in Germany, and their exact +place in the system is uncertain. The rocks of these areas consist +of red sandstones, sometimes shaly or conglomeratic, occasionally +with beds of impure limestone. Other more extensive areas where +Triassic rocks appear at the surface, are found west of the +Mississippi, on the slopes of the Rocky Mountains, where the beds +consist of sandstones and gypsiferous +<a name="page_206"><span class="page">Page 206</span></a> +marls. The American Trias is chiefly remarkable for having yielded +the remains of a small Marsupial (<i>Dromatherium</i>), and numerous +footprints, which have generally been referred to Birds +(<i>Brontozoum</i>), along with the tracks of undoubted Reptiles +(<i>Otozoum, Anisopus</i>, &c.) +</p> + +<p class="indent"> +The subjoined section (fig. 139) expresses, in a diagrammatic +manner, the general sequence of the Triassic rocks when fully +developed, as, for example, in the Bavarian Alps:— +</p> + +<div class="center"> +<table border="0" width="550"> +<tr><td class="center"><span class="image"> +GENERALIZED SECTION OF THE TRIASSIC ROCKS OF CENTRAL EUROPE. +</span></td></tr> +<tr><td class="center"><span class="image">Fig. + 139.</span></td></tr> +<tr><td> +<img src="images/fig139.jpg" width="546" height="628" alt="Fig. 139" /> +</td></tr> +</table> +</div> + +<p class="indent"> +With regard to the <i>life</i> of the Triassic period, we have to +<a name="page_207"><span class="page">Page 207</span></a> +notice a difference as concerns the +different members of the group similar to that which has been +already mentioned in connection with the Permian formation. The +arenaceous deposits of the series, namely, resemble those of the +Permian, not only in being commonly red or variegated in their +colour, but also in their conspicuous paucity of organic remains. +They for the most part are either wholly unfossiliferous, or they +contain the remains of plants or the bones of reptiles, such as +may easily have been drifted from some neighbouring shore. The +few fossils which may be considered as properly belonging to +these deposits are chiefly Crustaceans (<i>Estheria</i>) or +Fishes, which may well have lived in the waters of estuaries or +vast inland seas. We may therefore conclude, with considerable +probability, that the barren sandy and marly accumulations of +the Bunter Sandstein and Lower Keuper were not laid down in an +open sea, but are probably brackish-water deposits, formed in +estuaries or land-locked bodies of salt water. This at any rate +would appear to be the case as regards these members of the +series as developed in Britain and in their typical areas on +the continent of Europe; and the origin of most of the North +American Trias would appear to be much the same. Whether this +view be correct or not, it is certain that the beds in question +were laid down in <i>shallow</i> water, and in the immediate +vicinity of <i>land</i>, as shown by the numerous drifted plants +which they contain and the common occurrence in them of the +footprints of air-breathing animals (Birds, Reptiles, and +Amphibians). On the other hand, the middle and highest members +of the Trias are largely calcareous, and are replete with the +remains of undoubted marine animals. There cannot, therefore, +be the smallest doubt but that the Muschelkalk and the Rhætic +or Kössen beds were slowly accumulated in an open sea, of at +least a moderate depth; and they have preserved for us a very +considerable selection from the marine fauna of the Triassic +period. +</p> + +<p class="indent"> +The <i>plants</i> of the Trias are, on the whole, as distinctively +Mesozoic in their aspect as those of the Permian are Palæozoic. +In spite, therefore, of the great difficulty which is experienced +in effecting a satisfactory stratigraphical separation between the +Permian and the Trias, we have in this fact a proof that the two +formations were divided by an interval of time sufficient to allow +of enormous changes in the terrestrial vegetation of the world. The +<i>Lepidodendroids, Asterophyllites</i>, and <i>AnnulariĹ“</i>, +of the Coal and Permian formations, have now apparently wholly +disappeared: and the Triassic flora consists mainly of Ferns, +Cycads, and Conifers, of which only the two +<a name="page_208"><span class="page">Page 208</span></a> +last need special +notice. The <i>Cycads</i> (fig. 140) are true exogenous plants, +which in general form and habit of growth present considerable + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 370px; + font-size: smaller; text-align: center;"> +<img src="images/fig140.jpg" width="370" height="250" alt="Fig. 140" /> +<br /> +Fig. 140.—<i>Zamia spiralis</i>, a living Cycad. Australia. +</span> +</span> + +resemblance to young Palms, but which in reality are most nearly +related to the Pines and Firs (<i>ConiferĹ“</i>). The trunk +is unbranched, often much shortened, and bears a crown of feathery +pinnate fronds. The leaves are usually "circinate"—they unroll +in expanding, like the fronds of ferns. The seeds are not protected +by a seed-vessel, but are borne upon the edge of altered leaves, +or are carried on the scales of a cone. All the living species of +Cycads are natives of warm countries, such as South America, the +West Indies, Japan, Australia, Southern Asia, and South Africa. +The remains of Cycads, as we have seen, are not known to occur +in the Coal formation, or only to a very limited extent towards +its close; nor are they known with certainty as occurring in +Permian deposits. In the Triassic period, however, the remains +of Cycads belonging to such genera as <i>Pterophyllum</i> (fig. +141, <i>b</i>), <i>Zamites</i>, and <i>Podozamites</i> (fig. 141, +<i>c</i>), are sufficiently abundant to constitute quite a marked +feature in the vegetation; and they continue to be abundantly +represented throughout the whole Mesozoic series. The name "Age +of Cycads," as applied to the Secondary epoch, is therefore, +from a botanical point of view, an extremely appropriate one. +The <i>Conifers</i> of the Trias are not uncommon, the principal +form being <i>Veltzia</i> (fig. 141, <i>a</i>), which possesses +some peculiar characters, but would appear to be most nearly +related to the recent Cypresses. +</p> + +<p class="indent"> +As regards the <i>Invertebrate animals</i> of the Trias, our +knowledge is still principally derived from the calcareous beds +which constitute the centre of the system (the Muschelkalk) +<a name="page_209"><span class="page">Page 209</span></a> +on the continent +of Europe, and from the St Cassain and Rhætic beds +still higher in the series; whilst some of the Triassic strata + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 500px; + font-size: smaller; text-align: justify;"> +<img src="images/fig141.jpg" width="500" height="541" alt="Fig. 141" /> +<br /> +Fig. 141.—Triassic Conifers and Cycads. <i>a, Voltzia</i> +(<i>Schizoneura</i>) <i>heterophylla</i>, portion of a branch, +Europe and America; <i>b</i>, Part of the frond of +<i>Pterophyllum JĹ“geri</i>, Europe; <i>c</i>, Part of the +frond of <i>Podozamites lanceolatus</i>, America. +</span> +</span> + +of California and Nevada have likewise yielded numerous +remains of marine Invertebrates. The <i>Protozoans</i> are +represented by <i>Foraminifera</i> and <i>Sponges</i>, and the +<i>CĹ“lenterates</i> by a small number of <i>Corals</i>; but +these require no special notice. It may be mentioned, however, +that the great Palæozoic group of the <i>Rugose</i> corals +has no known representative here, its place being taken by corals +of Secondary type (such as <i>Montlivaltia, SynastĹ“a</i>, +&c.) +</p> + +<p class="indent"> +The <i>Echinoderms</i> are represented principally by +<i>Crinoids</i>, the remains of which are extremely abundant +in some of the limestones. The best-known species is the famous +"Lily-Encrinite" (<i>Encrinus liliiformis</i>, fig. 142), which +is characteristic +<a name="page_210"><span class="page">Page 210</span></a> +of the Muschelkalk. In this beautiful species, +the flower-like head is supported upon a rounded stem, the joints + +<span style="float: left; margin: 4px; width: 148px; + font-size: smaller; text-align: justify;"> +<img src="images/fig142.jpg" width="148" height="526" alt="Fig. 142" /> +<br /> +Fig. 142.—Head and upper part of the column of <i>Encrinus +liliiformis</i>. The lower figure shows the articulating surface +of one of the joints of the column. Muschelkalk, Germany. +</span> + +of which are elaborately articulated with one another; and the +fringed arms are composed each of a double series of alternating +calcareous pieces. The Palæozoic Urchins, with their +supernumerary rows of plates, the Cystideans, and the Pentremites +have finally disappeared; but both Star-fishes and Brittle-stars +continue to be represented. One of the latter—namely, the + +<span style="float: right; margin: 4px; width: 365px; + font-size: smaller; text-align: center;"> +<img src="images/fig143.jpg" width="365" height="151" alt="Fig. 143" /> +<br /> +Fig. 143.—<i>Aspidura loricata</i>, a Triassic +Ophiuroid. Muschelkalk, Germany. +</span> + +<i>Aspidura loricata</i> of Goldfuss (fig. 143)—is highly +characteristic of the Muschelkalk. +</p> + +<p class="indent"> +The remains of <i>Articulate Animals</i> are not very abundant in +the Trias, if we except the bivalved cases of the little Water-fleas +(<i>Ostracoda</i>), which are occasionally very plentiful. There +are also many species of the horny, concentrically-striated valves +of the <i>EstheriĹ“</i> (see fig. 122, <i>b</i>), which might +easily be taken for small Bivalve Molluscs. The "Long-tailed" +Decapods of the type of the Lobster, are not without examples +but they become much more numerous in the succeeding Jurassic +period. Remains of insects have also been discovered. +</p> + +<p class="indent"> +Amongst the <i>Mollusca</i> we have to note the disappearance, +amongst the lower groups, of many characteristic Palæozoic +types. Amongst the <i>Polyzoans</i>, the characteristic "Lace-corals," +<i>Fenestella, Retepora</i>,[22] <i>Synocladia, Polypora</i>, &c., +<a name="page_211"><span class="page">Page 211</span></a> +have become apparently extinct. The same is true of many of the ancient +types of <i>Brachiopods</i>, and conspicuously so of the great family +of the <i>ProductidĹ“</i>, which played such an important part in +the seas of the Carboniferous and Permian periods. +</p> + +<p class="footnote"> +[Footnote 22: The genus <i>Retefora</i> is really a recent one, +represented by living forms; and the so-called <i>ReteporĹ“</i> +of the Palæozoic rocks should properly receive another name +(<i>Phyllopora</i>), as being of a different nature. The name +<i>Retepora</i> has been here retained for these old forms simply +in accordance with general usage.] +</p> + +<p class="indent"> +<i>Bivalves</i> (<i>Lamellibranchiata</i>) and <i>Univalves</i> +(<i>Gasteropoda</i>) are well represented in the marine beds of +the Trias, and some of the former are particularly characteristic +either of the formation as a whole or of minor subdivisions of it. A +few of these characteristic species are figured in the accompanying +illustration (fig. 144). Bivalve shells of the genera <i>Daonella</i> +(fig. 144, <i>a</i>) and <i>Halobia</i> (<i>Monotis</i>) are very + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 464px; + font-size: smaller; text-align: justify;"> +<img src="images/fig144.jpg" width="464" height="363" alt="Fig. 144" /> +<br /> +Fig. 144. Triassic Lamellibranchs. <i>a, Daonella</i> (<i>Halobia</i>) +<i>Lommelli; b, Pecten Valoniensis; c, Myophoria lineata; d. Cardium +RhĹ“ticum; e. Avicula contorta; f. Avicula socialis</i>. +</span> +</span> + +abundant, and are found in the Triassic strata of almost all +regions. These groups belong to the family of the Pearl-oysters +(<i>AviculidĹ“</i>), and are singular from the striking +resemblance borne by some of their included forms to the +<i>StrophomenĹ“</i> amongst the Lamp-shells, though, of course, +no real relation exists between the two. The little Pearl-oyster, +<i>Avicula socialis</i> (fig. 144, <i>f</i>), is found throughout +the greater part of the Triassic series, and is especially abundant +in the Muschelkalk. The genus <i>Myophoria</i> (fig. 144, <i>c</i>), +belonging to the <i>TrigoniadĹ“</i>, and related therefore to +the Permian <i>Schizodus</i>, is characteristically Triassic, many +species of the genus being known in deposits of this age. Lastly, +the so-called "Rhætic" or "Kössen" beds are +<a name="page_212"><span class="page">Page 212</span></a> +characterised by the occurrence in them of the Scallop, +<i>Pecten Valoniensis</i> (fig. 144, <i>b</i>); the small Cockle, +<i>Cardium RhĹ“ticum</i> (fig. 144, <i>d</i>); and the +curiously-twisted Pearl-oyster, <i>Avicula contorta</i> (fig. 144, +<i>e</i>)—this last Bivalve being so abundant that the strata +in question are often spoken of as the "Avicula contorta beds." +</p> + +<p class="indent"> +Passing over the groups of the <i>Heteropods</i> and +<i>Pteropods</i>, we have to notice the <i>Cephalopoda</i>, which +are represented in the Trias not only by the chambered shells +of <i>Tetrabranchiates</i>, but also, for the first time, by +the internal skeletons of <i>Dibranchiate</i> forms. The Trias, +therefore, marks the first recognised appearance of true +Cuttle-fishes. All the known examples of these belong to the great +Mesozoic group of the <i>BelemnitidĹ“</i>; and as this family +is much more largely developed in the succeeding Jurassic period, +the consideration of its characters will be deferred till that +formation is treated of. Amongst the chambered <i>Cephalopods</i> +we find quite a number of the Palæozoic <i>Orthoceratites</i>, +some of them of considerable size, along with the ancient +<i>Cyrtoceras</i> and <i>Goniatites</i>; and these old types, +singularly enough, occur in the higher portion of the Trias (St +Cassian beds), but have, for some unexplained reason, not yet +been recognised in the lower and equally fossiliferous formation +of the Muschelkalk. Along with these we meet for the first time +with true <i>Ammonites</i>, which fill such an extensive place + +<span style="float: left; margin: 4px; width: 332px; + font-size: smaller; text-align: center;"> +<img src="images/fig145.jpg" width="322" height="281" alt="Fig. 145" /> +<br /> +Fig. 145.—<i>Ceratites nodosus</i>, viewed from the side and +from behind. Muschelkalk. +</span> + +in the Jurassic seas, and which will be spoken of hereafter. The +form, however, which is most characteristic of the Trias is +<i>Ceratites</i> (fig. 145). In this genus the shell is curved +into a flat spiral, the volutions of which are in contact; and it +further agrees with both <i>Goniatites</i> and <i>Ammonites</i> +in the fact that the septa or partitions between the air-chambers +are not simple and plain (as in the <i>Nautilus</i> and its allies), +but are folded and bent as they approach the outer wall of the +shell. In the <i>Goniatite</i> these foldings of the septa are +of a simply lobed or angulated nature, and in the <i>Ammonite</i> +<a name="page_213"><span class="page">Page 213</span></a> +they are extremely complex; whilst in the +<i>Ceratite</i> there is an intermediate state of things, the +special feature of which is, that those foldings which are turned +towards the mouth of the shell are merely rounded, whereas those +which are turned away from the mouth are characteristically +toothed. The genus <i>Ceratites</i>, though principally Triassic, +has recently been recognised in strata of Carboniferous age in +India. +</p> + +<p class="indent"> +From the foregoing it will be gathered that one of the most important +points in connection with the Triassic <i>Mollusca</i> is the +remarkable intermixture of Palæozoic and Mesozoic types which +they exhibit. It is to be remembered, also, that this intermixture +has hitherto been recognised, not in the Middle Triassic limestones +of the Muschelkalk, in which—as the oldest Triassic beds with +marine fossils—we should naturally expect to find it, but in +the St Cassian beds, the age of which is considerably later than +that of the Muschelkalk. The intermingling of old and new types of +Shell-fish in the Upper Trias is well brought out in the annexed +table, given by Sir Charles Lyell in his 'Student's Elements of +Geology' (some of the less important forms in the table being +omitted here):— +</p> + +<p class="center"> +GENERA OF FOSSIL MOLLUSCA IN THE ST CASSIAN AND HALLSTADT BEDS. +</p> + +<table border="0" cellspacing="0" cellpadding="4" width="100%"> +<tr><td valign="top" class="genera"> + Common to Older Rocks. +</td><td valign="top" class="genera"> + Characteristic of Triassic Rocks. +</td><td valign="top" class="genera"> + Common to Newer Rocks. +</td></tr> + +<tr><td valign="top" class="genera"> + <table border="0" cellspacing="0" cellpadding="0"> + <tr><td class="left">Orthoceras.</td></tr> + <tr><td class="left">Bactrites.</td></tr> + <tr><td class="left">Macrocheilus.</td></tr> + <tr><td class="left">Loxonema.</td></tr> + <tr><td class="left">Holopella.</td></tr> + <tr><td class="left">Murchisonia.</td></tr> + <tr><td class="left">Porcellia.</td></tr> + <tr><td class="left">Athyris.</td></tr> + <tr><td class="left">Retzia.</td></tr> + <tr><td class="left">Cyrtina.</td></tr> + <tr><td class="left">Euomphalus.</td></tr> + </table> +</td><td valign="top" class="genera"> + <table border="0" cellspacing="0" cellpadding="0"> + <tr><td class="left">Ceratites.</td></tr> + <tr><td class="left">Cochloceras.</td></tr> + <tr><td class="left">Rhabdoceras.</td></tr> + <tr><td class="left">Aulacoceras.</td></tr> + <tr><td class="left">Naticella.</td></tr> + <tr><td class="left">Platystoma.</td></tr> + <tr><td class="left">Halobia.</td></tr> + <tr><td class="left">Hörnesia.</td></tr> + <tr><td class="left">Koninckia.</td></tr> + <tr><td class="left">Scoliostoma.</td></tr> + <tr><td class="left">Myophoria.</td></tr> + </table> + (The last two are principally but not exclusively Triassic.) +</td><td valign="top" class="genera"> + <table border="0" cellspacing="0" cellpadding="0"> + <tr><td class="left">Ammonites.</td></tr> + <tr><td class="left">Chemnitzia.</td></tr> + <tr><td class="left">Cerithium.</td></tr> + <tr><td class="left">Monodonta.</td></tr> + <tr><td class="left">SphĹ“ra.</td></tr> + <tr><td class="left">Cardita.</td></tr> + <tr><td class="left">Myoconcha.</td></tr> + <tr><td class="left">Hinnites.</td></tr> + <tr><td class="left">Monotis.</td></tr> + <tr><td class="left">Plicatula.</td></tr> + <tr><td class="left">Pachyrisma.</td></tr> + <tr><td class="left">Thecidium.</td></tr> + </table></td></tr> +</table> + +<p class="indent"> +Thus, to emphasise the more important points alone, the Trias +has yielded, amongst the Gasteropods, the characteristically +Palæozoic <i>Loxonema, Holopella, Murchisonia, Euomphalus</i>, +and <i>Porcellia</i>, along with typically Triassic forms like +<i>Platystoma</i> and <i>Scoliostoma</i>, and the great modern +groups <i>Chemnitzia</i> and <i>Cerithium</i>. Amongst the Bivalves +we find the Palæozoic <i>Megalodon</i> side by side with the +Triassic <i>Halobia</i> and <i>Myophoria</i>, these being associated +with the <i>CarditĹ“, Hinnites, PlicatulĹ“</i>, and +<i>TrigoniĹ“</i> of later deposits. The Brachiopods +<a name="page_214"><span class="page">Page 214</span></a> +exhibit the Palæozoic <i>Athyris, Retzia</i>, and +<i>Cyrtina</i>, with the Triassic <i>Koninckia</i> and the modern +<i>Thecidium</i>. Finally, it is here that the ancient genera +<i>Orthoceras, Cyrtoceras</i>, and <i>Goniatites</i> make their +last appearance upon the scene of life, the place of the last of +these being taken by the more complex and almost exclusively +Triassic <i>Ceratites</i>, whilst the still more complex genus +<i>Ammonites</i> first appears here in force, and is never again +wanting till we reach the close of the Mesozoic period. The first +representatives of the great Secondary family of the +<i>Belemnites</i> are also recorded from this horizon. +</p> + +<p class="indent"> +Amongst the <i>Vertebrate Animals</i> of the Trias, the <i>Fishes</i> +are represented by numerous forms belonging to the <i>Ganoids</i> and +the <i>Placoids</i>. The Ganoids of the period are still all provided +with unsymmetrical ("heterocercal") tails, and belong principally +to such genera as <i>PalĹ“oniscus</i> and <i>Catopterus</i>. +The remains of Placoids are in the form of teeth and spines, the +two principal genera being the two important Secondary groups +<i>Acrodus</i> and <i>Hybodus</i>. Very nearly at the summit +of the Trias in England, in the Rhætic series, is a singular +stratum, which is well known as the "bone-bed," from the number +of fish-remains which it contains. More interesting, however, +than the above, are the curious palate-teeth of the Trias, upon +which Agassiz founded the genus <i>Ceratodus</i>. The teeth of +Ceratodus (fig. 146) are singular flattened plates, composed + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 441px; + font-size: smaller; text-align: center;"> +<img src="images/fig146.jpg" width="441" height="151" alt="Fig. 146" /> +<br /> +Fig. 146.—<i>a</i>, Dental plate of <i>Ceratodus serratus</i>, +Keuper; <i>b</i>, Dental plate of <i>Ceratodus altus</i>, +Keuper; (After Agassiz.) +</span> +</span> + +of spongy bone beneath, covered superficially with a layer of +enamel. Each plate is approximately triangular, one margin (which +we now know to be the outer one) being prolonged into prongs or +conical prominences, whilst the surface is more or less regularly +undulated. Until recently, though the master-mind of Agassiz +recognised that these singular bodies were undoubtedly the teeth +of fishes, we were entirely ignorant as to their precise relation +to the animal, or as to the exact affinities of the fish thus +armed. Lately, however, there has been discovered in the rivers +of Queensland (Australia) a living species of <i>Ceratodus</i> +(<i>C. Fosteri</i>, fig. 147), +<a name="page_215"><span class="page">Page 215</span></a> +with teeth precisely similar to those +of its Triassic predecessor; and we thus have become acquainted +with the use of these structures and the manner in which they + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 528px; + font-size: smaller; text-align: center;"> +<img src="images/fig147.jpg" width="528" height="107" alt="Fig. 147" /> +<br /> +Fig. 147.—<i>Ceratodus Fosteri</i>, the Australian Mud-fish, +reduced in size. +</span> +</span> + +were implanted in the mouth. The palate carries two of these +plates, with their longer straight sides turned towards each other, +their sharply-sinuated sides turned outwards, and their short +straight sides or bases directed backwards. Two similar plates in +the lower jaw correspond to the upper, their undulated surfaces +fitting exactly to those of the opposite teeth. There are also +two sharp-edged front teeth, which are placed in the front of +the mouth in the upper jaw; but these have not been recognised +in the fossil specimens. The living <i>Ceratodus</i> feeds on +vegetable matters, which are taken up or tom off from plants by +the sharp front teeth, and then partially crushed between the +undulated surfaces of the back teeth (Günther); and there need +be little doubt but that the Triassic <i>Ceratodi</i> followed +a similar mode of existence. From the study of the living +<i>Ceratodus</i>, it is certain that the genus belongs to the +same group as the existing Mud-fishes (<i>Dipnoi</i>); and we +therefore learn that this, the highest, group of the entire class +of Fishes existed in Triassic times under forms little or not +at all different from species now alive; whilst it has become +probable that the order can be traced back into the Devonian +period. +</p> + +<p class="indent"> +The <i>Amphibians</i> of the Trias all belong to the old order +of the <i>Labyrinthodonts</i>, and some of them are remarkable +for their gigantic dimensions. They were first known by their +footprints, which were found to occur plentifully in the Triassic +sandstones of Britain and the continent of Europe, and which +consisted of a double series of alternately-placed pairs of +hand-shaped impressions, the hinder print of each pair being +much larger than the one in front (fig. 148). So like were these +impressions to the shape of the human hand, that the at that +time unknown animal which produced them was at once christened +<i>Cheirotherium</i>, or "Hand-beast." Further discoveries, however, +soon showed that the footprints of <i>Cheirotherium</i> were really +produced by species of Amphibians which, like the existing Frogs, +possessed hind-feet of a much larger size than the fore-feet, +<a name="page_216"><span class="page">Page 216</span></a> + +<span style="float:left; text-align: justify; width: 232px; + margin: 4px; font-size: smaller;"> +<img src="images/fig148a.jpg" width="232" height="312" alt="Fig. 148a" /> +</span> + +<span style="float: left; width: 100%; text-align: center; + clear: left;"> + +<span style="margin: 4px; text-align: justify; width: 505px; + font-size: smaller;"> +<img src="images/fig148b.jpg" width="505" height="516" alt="Fig. 148b" /> +<br /> +Fig. 148.—Footprints of a Labyrinthodont (<i>Cheirotherium</i>), +from the Triassic Sandstones of Hessberg, near Hildburghausen, +Germany, reduced one-eighth. The lower figure shows a slab, with +several prints, and traversed by reticulated sun-cracks: the upper +figure shows the impression of one of the hind-feet, one-half of +the natural size. (After Sickler.) +</span> + +</span> + +and to which the name of <i>Labyrinthodonts</i> was applied in +consequence of the complex microscopic structure of the teeth +(fig. 149). In the essential details of their structure, the +Triassic Labyrinthodonts did not differ materially from their +predecessors in the Coal-measures and Permian rocks. They possessed +the same frog-like skulls (fig. 150), with a lizard-like body, a +long tail, and comparatively feeble limbs. The hind-limbs were +stronger and longer than the fore-limbs, and the lower +<a name="page_217"><span class="page">Page 217</span></a> +surface of the body was protected by an armour of bony +plates. Some of the Triassic Labyrinthodonts must have attained +dimensions utterly unapproached amongst existing Amphibians, the +skull of <i>Labyrinthodon JĹ“geri</i> (fig. 150) being upwards + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 345px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig149.jpg" width="345" height="317" alt="Fig. 149" /> +<br /> +Fig. 149.—Section of the tooth of <i>Labryinthodon +(Mastodonsaurus) JĹ“geri</i>, showing the microscopic +structure. Greatly enlarged. Trias. +</span> + +<span style="width: 173px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig150.jpg" width="173" height="283" alt="Fig. 150" /> +<br /> +Fig. 150.—<i>a</i>, Skull of <i>Labyrinthodon JĹ“geri</i>, +much reduced in size; <i>b</i>, Tooth of the same. Trias +Württemberg. +</span> + +</span> + +of three feet in length and two feet in breadth. Restorations of +some of these extraordinary creatures have been attempted in the +guise of colossal Frogs; but they must in reality have more +closely resembled huge Newts. +</p> + +<p class="indent"> +Remains of <i>Reptiles</i> are very abundant in Triassic deposits, +and belong to very varied types. The most marked feature, in fact, +connected with the Vertebrate fauna of the Trias, and of the +Secondary rocks in general, is the great abundance of Reptilian +life. Hence the Secondary period is often spoken of as the "Age +of Reptiles." Many of the Triassic reptiles depart widely in +their structure from any with which we are acquainted as existing +on the earth at the present day, and it is only possible here to +briefly note some of the more important of these ancient forms. +Amongst the group of the Lizards (<i>Lacertilia</i>), represented +by <i>Protorosaurus</i> in the older Permian strata, three types +more or less certainly referable to this order may be mentioned. +One of these is a small reptile which was found many years ago +in sandstones near Elgin, in Scotland, and which excited special +interest at the time in consequence of the fact that the strata +in question were believed to belong to the Old Red Sandstone +formation. It is, however, +<a name="page_218"><span class="page">Page 218</span></a> +now certain that +the Elgin sandstones which contain <i>Telerpeton Elginense</i>, as +this reptile is termed, are really to be regarded as of Triassic +age. By Professor Huxley, <i>Telerpeton</i> is regarded as a +Lizard, which cannot be considered as "in any sense a less +perfectly-organised creature than the Gecko, whose swift and +noiseless run over walls and ceilings surprises the traveller +in climates warmer than our own." The "Elgin Sandstones" have +also yielded another Lizard, which was originally described by +Professor Huxley under the name of <i>Hyperodapedon</i>, the +remains of the same genus having been subsequently discovered +in Triassic strata in India and South Africa. The Lizards of +this group must therefore have at one time enjoyed a very wide +distribution over the globe; and the living <i>Sphenodon</i> of +New Zealand is believed by Professor Huxley to be the nearest +living ally of this family. The <i>Hyperodapedon</i> of the +Elgin Sandstones was about six feet in length, with limbs adapted +for terrestrial progression, but with the bodies of the +vertebræ slightly biconcave, and having two rows of palatal +teeth, which become worn down to the bone in old age. Lastly, the +curious <i>Rhynchosaurus</i> of the Trias is also referred, by +the eminent comparative anatomist above mentioned, to the order +of the Lizards. In this singular reptile + +<span style="float: left; margin: 4px; width: 254px; + font-size: smaller; text-align: center;"> +<img src="images/fig151.jpg" width="237" height="125" alt="Fig. 151" /> +<br /> +Fig. 151.—Skull of <i>Rhynchosaurus articeps</i>. Trias. +(After Owen.) +</span> + +(fig. 151) the skull is somewhat bird-like, and the jaws appear +to have been destitute of teeth, and to have been encased in a +horny sheath like the beak of a Turtle or a Bird. It is possible, +however, that the palate was furnished with teeth. +</p> + +<p class="indent"> +The group of the Crocodiles and Alligators (<i>Crocadilia</i>), +distinguished by the fact that the teeth are implanted in distinct +sockets and the skin more or less extensively provided with bony +plates, is represented in the Triassic rocks by the +<i>Stagonolepis</i> of the Elgin Sandstones. The so-called +"Thecodont" reptiles (such as <i>Belodon, Thecodontosaurus</i>, +and <i>PalĹ“osaurus</i>, fig. 152, <i>c, d, e</i>) are also +nearly related to the Crocodiles, though it is doubtful if they +should be absolutely referred to this group. In these reptiles, the +teeth are implanted in distinct sockets in the jaws, their crowns +being more or less compressed and pointed, "with trenchant and +finely serrate margins" (Owen). The bodies of the vertebræ are +hollowed out at both ends, but the limbs appear to be adapted +for progression on the land. The genus <i>Belodon</i> (fig. 152, +<i>c</i>) is known to occur in the Keuper of Germany and in America; +<a name="page_219"><span class="page">Page 219</span></a> +and <i>PalĹ“osaurus</i> (fig. 153. <i>e</i>) has also been +found in the Trias of the same region. Teeth of the latter, however, +are found, along with remains of <i>Thecodontosaurus</i> (fig. 153, +<i>d</i>), in a singular magnesian conglomerate near Bristol, which +was originally believed to be of Permian age, but which appears to +be undoubtedly Triassic. +</p> + +<div class="center"> +<table border="0" width="509"> +<tr><td> +<img src="images/fig152.jpg" width="505" height="258" alt="Fig. 152" /> +</td></tr> +<tr><td class="left"><span class="image"> +Fig. 152.—Triassic Reptiles. <i>a</i>, Skull of <i>Nothosaurus +mirabilis</i>, reduced in size—Muschelkalk, Germany; <i>b</i>, +Tooth of <i>Simosaurus Gaillardoti</i>, of the natural +size—Muschelkalk, Germany; <i>c</i>, Tooth of <i>Beladon +Carolinensis</i>—Trias, America; <i>d</i>, Tooth of +<i>Thecodontosaurus antiquus</i>, slightly enlarged—Britain; +<i>e</i>, Tooth of <i>PalĹ“osaurus platyodon</i>, of the +natural size—Britain. +</span></td></tr> +</table> +</div> + +<p class="indent"> +The Trias has also yielded the remains of the great marine reptiles +which are often spoken of collectively as the "Enaliosaurians" +or "Sea-lizards," and which will be more particularly spoken +of in treating of the Jurassic period, of which they are more +especially characteristic. In all these reptiles the limbs are +flattened out, the digits being enclosed in a continuous skin, +thus forming powerful swimming-paddles, resembling the "flippers" +of the Whales and Dolphins both in their general structure and +in function. The tail is also long, and adapted to act as a +swimming-organ; and there can be no doubt but that these +extraordinary and often colossal reptiles frequented the sea, and +only occasionally came to the land. The Triassic Enaliosaurs belong +to a group of which the later genus <i>Plesiosaurus</i> is the type +(the <i>Sauropterygia</i>). One of the best known of the Triassic +genera is <i>Nothosaurus</i> (fig. 152, <i>a</i>), in which the +neck was long and bird-like, the jaws being immensely elongated, +and carrying numerous powerful conical teeth implanted in distinct +sockets. The teeth in <i>Simosaurus</i> (152, <i>b</i>) are of a +similar nature; but the orbits are of enormous size, indicating +eyes of corresponding dimensions, and perhaps pointing to the +nocturnal habits of the animal. In the singular +<a name="page_220"><span class="page">Page 220</span></a> +<i>Placodus</i>, +again, the teeth are in distinct sockets, but resemble those +of many fishes in being rounded and obtuse (fig. 153), forming + +<span style="float: left; margin: 4px; width: 255px; + font-size: smaller; text-align: justify;"> +<img src="images/fig153.jpg" width="255" height="219" alt="Fig. 153" /> +<br /> +Fig. 153.—Under surface of the upper jaw and palate of +<i>Placodus gigas</i>. Muschelkalk, Germany. +</span> + +broad crushing plates adapted for the comminution of shell-fish. +There is a row of these teeth all round the upper jaw proper, +and a double series on the palate, but the lower jaw has only a +single row of teeth. <i>Placodus</i> is found in the Muschelkalk, +and the characters of its dental apparatus indicate that it was +much more peaceful in its habits than its associates the Nothosaur +and Simosaur. +</p> + +<p class="indent"> +The Triassic rocks of South Africa and India have yielded the +remains of some extraordinary Reptiles, which have been placed +by Professor Owen in a separate order under the name of +<i>Anomodontia</i>. The two principal genera of this group are +<i>Dicynodon</i> and <i>Oudenodon</i>, both of which appear to +have been large Reptiles, with well-developed limbs, organised +for progression upon the dry land. In <i>Oudenodon</i> (fig. +154, B) the jaws seem to have been wholly destitute of teeth, and +must have been encased in a horny sheath, similar to that with +which we are familiar in the beak of a Turtle. In <i>Dicynodon</i> +(fig. 154, A), on the other hand, the front of the upper jaw +and the whole of the lower jaw were destitute of teeth, and the +front of the mouth must have constituted a kind of beak; but +the upper jaw possessed on each side a single huge conical tusk, +which is directed downwards, and must have continued to grow +during the life of the animal. +</p> + +<p class="indent"> +It may be mentioned that the above-mentioned Triassic sandstones of +South Africa have recently yielded to the researches of Professor +Owen a new and unexpected type of Reptile, which exhibits some +of the structural peculiarities which we have been accustomed +to regard as characteristic of the Carnivorous quadrupeds. The +Reptile in question has been named <i>Cyanodraco</i>, and it is +looked upon by its distinguished discoverer as the type of a new +order, to which he has given the name of <i>Theriodontia</i>. The +teeth of this singular form agree with those of the Carnivorous +quadrupeds in consisting of three distinct groups—namely, +front teeth or <i>incisors</i>, eye teeth or <i>canines</i>, and +back teeth or <i>molars</i>. The canines +<a name="page_221"><span class="page">Page 221</span></a> +also are long and pointed, +very much compressed, and having their lateral margins finely +serrated, thus presenting a singular resemblance to the teeth + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 347px; + font-size: smaller; text-align: justify;"> +<img src="images/fig154.jpg" width="347" height="416" alt="Fig. 154" /> +<br /> +Fig. 154.—Triassic Anomodont Reptiles. A, Skull of +<i>Dicynodon lacerticeps</i>, showing one of the great maxillary +tusks; B, Skull of <i>Oudenodon Bainii</i>, showing the toothless, +beak-like jaws. From the Trias of South Africa. (After Owen.) +</span> +</span> + +of the extinct "Sabre-toothed Tiger" (<i>Machairodus</i>). The +bone of the upper arm (humerus) further shows some remarkable +resemblances to the same bone in the Carnivorous Mammals. As +has been previously noticed, Professor Owen is of opinion that +some of the Reptilian remains of the Permian deposits will also +be found to belong to this group of the "Theriodonts." +</p> + +<p class="indent"> +Lastly, we find in the Triassic rocks the remains of Reptiles +belonging to the great Mesozoic order of the <i>Deinosauria</i>. +This order attains its maximum at a later period, and will be +spoken of when the Jurassic and Cretaceous deposits come to be +considered. The chief interest of the Triassic Reptiles of this +group arises from the fact that they are known by their footprints +as well as by their bones; and a question has arisen whether the +supposed footprints of <i>birds</i> which occur in the Trias +have not really been produced by Deinosaurs. This leads us, +therefore, to speak at the same time as to the evidence which we +have of the existence of the class of Birds during the Triassic +period. No actual bones of any bird have as +<a name="page_222"><span class="page">Page 222</span></a> +yet been detected in any Triassic deposit; but we have tolerably +clear evidence of their existence at this time in the form of +<i>footprints</i>. The impressions in question are found in +considerable numbers in certain red sandstones of the age of the +Trias in the valley of the Connecticut River, in the United States. +They vary much in size, and have evidently been produced by many +different animals walking over long stretches of estuarine mud +and sand exposed at low water. The footprints now under +consideration form a double series of <i>single</i> prints, and +therefore, beyond all question, are the tracks of a +<i>biped</i>—that is, of an animal which walked upon two +legs. No living animals, save Man and the Birds, walk habitually +on two legs; and there is, therefore, a <i>primâ facie</i> +presumption that the authors of these prints were Birds. Moreover, +each impression consists of the marks of three toes turned +forwards (fig. 155), and therefore are precisely such as might be + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 436px; + font-size: smaller; text-align: center;"> +<img src="images/fig155.jpg" width="436" height="327" alt="Fig. 155" /> +<br /> +Fig. 155.—Supposed footprint of a Bird, from the Triassic +Sandstones of the Connecticut River. The slab shows also numerous +"rain-prints." +</span> +</span> + +produced by Wading or Cursorial Birds. Further, the impressions +of the toes show exactly the same numerical progression in the +number of the joints as is observable in living Birds—that is +to say, the innermost of the three toes consists of three joints, +the middle one of four, and the outer one of five joints. Taking +this evidence collectively, it would have seemed, until lately, +quite certain that these tracks could only have been formed by +Birds. It has, however, been shown that the Deinosaurian Reptiles +possess, in some cases at any rate, some singularly bird-like +characters, amongst +<a name="page_223"><span class="page">Page 223</span></a> +which is the fact that +the animal possessed the power of walking, temporarily at least, +on its hind-legs, which were much longer and stronger than the +fore-limbs, and which were sometimes furnished with no more than +three toes. As the bones and teeth of Deinosaurs have been found +in the Triassic deposits of North America, it may be regarded as +certain that <i>some</i> of the bipedal tracks originally ascribed +to Birds must have really been produced by these Reptiles. It seems +at the same time almost a certainty that others of the three-toed +impressions of the Connecticut sandstones were in truth produced +by Birds, since it is doubtful if the bipedal mode of progression +was more than an occasional thing amongst the Deinosaurs, and the +greater number of the many known tracks exhibit no impressions +of fore-feet. Upon the whole, therefore, we may, with much +probability, conclude that the great class of Birds (<i>Aves</i>) +was in existence in the Triassic period. If this be so, not only +must there have been quite a number of different forms, but some +of them must have been of very large size. Thus the largest +footprints hitherto discovered in the Connecticut sandstones are +22 inches long and 12 inches wide, with a proportionate length +of stride. These measurements indicate a foot four times as large +as that of the African Ostrich; and the animal which produced +them—whether a Bird or a Deinosaur—must have been of +colossal dimensions. +</p> + +<p class="indent"> +Finally, the Trias completes the tale of the great classes of +the Vertebrate sub-kingdom by presenting us with remains of the +first known of the true Quadrupeds or <i>Mammalia</i>. These +are at present only known by their teeth, or, in one instance, +by one of the halves of the lower jaw; and these indicate minute +Quadrupeds, which present greater affinities with the little Banded +Anteater (<i>Myrmecobius fasciatus</i>, fig. 158) of Australia +than with any other living form. If this conjecture be correct, + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 283px; vertical-align:bottom; text-align: center; + margin: 4px;"> +<img src="images/fig156.jpg" width="283" height="92" alt="Fig. 156" /> +<br /> +Fig. 156.—Lower jaw of <i>Dromatherium sylvestre</i>. Trias, +North Carolina. (After Emmons.) +</span> + +<span style="width: 186px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig157.jpg" width="186" height="107" alt="Fig. 157" /> +<br /> +Fig. 157.—<i>a</i>, Molar tooth of <i>Micro estes antiquus</i>, +magnified; <i>b</i>, Crown of the same, magnified still further. +Trias, Germany. +</span> + +</span> + +these ancient Mammals belonged to the order of the Marsupials or +Pouched Quadrupeds (<i>Marsupialia</i>), which +<a name="page_224"><span class="page">Page 224</span></a> +are now exclusively confined +to the Australian province, South America, and the southern + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 524px; + font-size: smaller; text-align: center;"> +<img src="images/fig158.jpg" width="524" height="278" alt="Fig. 158" /> +<br /> +Fig. 158.—The Banded Ant-eater (<i>Myrmecobius fasciatus</i>) +of Australia. +</span> +</span> + +portion of North America. In the Old World, the only known Triassic +Mammals belong to the genus <i>Microlestes</i>, and to the probably +identical <i>Hypsiprymnopsis</i> of Professor Boyd Dawkins. The +teeth of <i>Microlestes</i> (fig. 157) were originally discovered +by Plieninger in 1847 in the "bone-bed" which is characteristic +of the summit of the Rhætic series both in Britain and on the +continent of Europe; and the known remains indicate two species. +In Britain, teeth of <i>Microlestes</i> have been discovered +by Mr Charles Moore in deposits of Upper Triassic age, filling +a fissure in the Carboniferous limestone near Frome, in Somersetshire; +and a molar tooth of <i>Hypsiprymnopsis</i> was found by Professor +Boyd Dawkins in Rhætic marls below the "bone-bed" at Watchet, +also in Somersetshire. In North America, lastly, there has been +found in strata of Triassic age one of the branches of the lower +jaw of a small Mammal, which has been described under the name of +<i>Dromatherium sylvestre</i> (fig. 156). The fossil exhibits ten +small molars placed side by side, one canine, and three incisors, +separated by small intervals, and it indicates a small insectivorous +animal, probably most nearly related to the existing +<i>Myrmecobius</i>. +</p> + +<h4>LITERATURE.</h4> + +<p class="indent"> +The following list comprises a few of the more important sources of +information as to the Triassic strata and their fossil +contents:— +</p> + +<table border="0" cellspacing="0"> +<tr><td class="right" valign="top">(1)</td> + <td>'Geology of Oxford and the Valley of the Thames.' + Phillips.</td></tr> +<tr><td class="right" valign="top">(2)</td> + <td>'Memoirs of the Geological Survey of Great Britain and + Ireland.'</td></tr> +<tr><td class="right" valign="top">(3)</td> + <td>'Report on the Geology of Londonderry,' &c. Portlock.</td></tr> +<tr><td class="right" valign="top"> + <a name="page_225"><span class="page">Page 225</span></a> + (4)</td> + <td>"On the Zone of Avicula contorta," &c.—'Quart. Journ. + Geol. Soc.,' vol. xvi., 1860. Dr Thomas Wright.</td></tr> +<tr><td class="right" valign="top">(5)</td> + <td>"On the Zones of the Lower Lias and the Avicula contorta + Zone"—'Quart. Journ. Geol. Soc.,' vol. xvii., 1861. Charles + Moore.</td></tr> +<tr><td class="right" valign="top">(6)</td> + <td>"On Abnormal Conditions of Secondary Deposits," &c.—'Quart. + Journ. Geol. Soc.,' vol. xxiii., 1876-77. Charles Moore.</td></tr> +<tr><td class="right" valign="top">(7)</td> + <td>'Geognostische Beschreibung des Bayerischen Alpengebirges.' + Gümbel.</td></tr> +<tr><td class="right" valign="top">(8)</td> + <td>'Lethæa Rossica.' Pander.</td></tr> +<tr><td class="right" valign="top">(9)</td> + <td>'Lethæa Geognostica.' Bronn.</td></tr> +<tr><td class="right" valign="top">(10)</td> + <td>'Petrefacta Germaniæ.' Goldfuss.</td></tr> +<tr><td class="right" valign="top">(11)</td> + <td>'Petrefaktenkunde.' Quenstedt.</td></tr> +<tr><td class="right" valign="top">(12)</td> + <td>'Monograph of the Fossil Estheriæ' + (Palæontographical Society). Rupert Jones.</td></tr> +<tr><td class="right" valign="top">(13)</td> + <td>"Fossil Remains of Three Distinct Saurian Animals, + recently discovered in the Magnesian Conglomerate near + Bristol"—'Trans. Geol. Soc.,' ser. 2, vol. v., 1840. Riley + and Stutchbury.</td></tr> +<tr><td class="right" valign="top">(14)</td> + <td>'Die Saurier des Muschekalkes.' Von Meyer.</td></tr> +<tr><td class="right" valign="top">(15)</td> + <td>'Beiträge zur Palæontologie Württembergs.' Von + Meyer and Plieninger.</td></tr> +<tr><td class="right" valign="top">(16)</td> + <td>'Manual of Palæontology.' Owen.</td></tr> +<tr><td class="right" valign="top">(17)</td> + <td>'Odontography:' Owen.</td></tr> +<tr><td class="right" valign="top">(18)</td> + <td>'Report on Fossil Reptiles' (British Association, 1841). + Owen.</td></tr> +<tr><td class="right" valign="top">(19)</td> + <td>"On Dicynodon"—'Trans. Geol. Soc.,' vol. iii., 1845. + Owen.</td></tr> +<tr><td class="right" valign="top">(20)</td> + <td>'Descriptive Catalogue of Fossil Reptilia and Fishes in the + Museum of the Royal College of Surgeons, England.' + Owen.</td></tr> +<tr><td class="right" valign="top">(21)</td> + <td>"On Species of Labyrinthodon from Warwickshire"—'Trans. + Geol. Soc.,' ser. 2, vol. vi. Owen.</td></tr> +<tr><td class="right" valign="top">(22)</td> + <td>"On a Carnivorous Reptile" (Cynodraco major), + &c.—'Quart. Journ. Geol. Soc.,' vol. xxxii., 1876. + Owen.</td></tr> +<tr><td class="right" valign="top">(23)</td> + <td>"On Evidences of Theriodonts in Permian Deposits," + &c.—'Quart. Journ. Geol. Soc.,' vol. xxxii., 1876. + Owen.</td></tr> +<tr><td class="right" valign="top">(24)</td> + <td>"On the Stagonolepis Robertsoni," &c.—'Quart. Journ. + Geol. Soc.,' vol. xv., 1859. Huxley.</td></tr> +<tr><td class="right" valign="top">(25)</td> + <td>"On a New Specimen of Telerpeton Elginense"—'Quart. Journ. + Geol. Soc.,' vol. xxiii., 1866. Huxley.</td></tr> +<tr><td class="right" valign="top">(26)</td> + <td>"On Hyperodapedon"—'Quart. Journ. Geol. Soc.,' vol. xxv., + 1869. Huxley.</td></tr> +<tr><td class="right" valign="top">(27)</td> + <td>"On the Affinities between the Deinosaurian Reptiles and + Birds"—'Quart. Journ. Geol. Soc.,' vol. xxvi., 1870. + Huxley.</td></tr> +<tr><td class="right" valign="top">(28)</td> + <td>"On the Classification of the Deinosauria," + &c.—'Quart. Journ. Geol. Soc.,' vol. xxvi., 1870. + Huxley.</td></tr> +<tr><td class="right" valign="top">(29)</td> + <td>"Palæontologica Indica"—'Memoirs of the Geol. + Survey of India.'</td></tr> +<tr><td class="right" valign="top">(30)</td> + <td>"On the Geological Position and Geographical Distribution of + the Dolomitic Conglomerate of the Bristol Area"—'Quart. + Journ. Geol. Soc.,' vol. xxvi., 1870. R. Etheridge, sen.</td></tr> +<tr><td class="right" valign="top">(31)</td> + <td>"Remains of Labyrinthodonta from the Keuper Sandstone of + Warwick"—'Quart. Journ. Geol. Soc.,' vol. xxx., 1874 + Miall.</td></tr> +<tr><td class="right" valign="top">(32)</td> + <td>'Manual of Geology.' Dana.</td></tr> +<tr><td class="right" valign="top">(33)</td> + <td>'Synopsis of Extinct Batrachia and Reptilia of North America.' + Cope.</td></tr> +<tr><td class="right" valign="top">(34)</td> + <td>'Fossil Footmarks.' Hitchcock.</td></tr> +<tr><td class="right" valign="top">(35)</td> + <td>'Ichnology of New England.' Hitchcock.</td></tr> +<tr><td class="right" valign="top">(36)</td> + <td>'TraitĂ© de PalĂ©ontologie + VĂ©gĂ©tale.' Schimper.</td></tr> +<tr><td class="right" valign="top">(37)</td> + <td>'Histoire des VĂ©gĂ©taux Fossiles.' + Brongniart.</td></tr> +<tr><td class="right" valign="top">(38)</td> + <td>'Monographie der Fossilen Coniferen.' GĹ“ppert.</td></tr> +</table> + +<h3> +<a name="page_226"><span class="page">Page 226</span></a> +CHAPTER XVI.</h3> + +<p class="subtitle"> +THE JURASSIC PERIOD. +</p> + +<p class="indent"> +Resting upon the Trias, with perfect conformity, and with an almost +undeterminable junction, we have the great series of deposits which +are known as the <i>Oolitic Rocks</i>, from the common occurrence +in them of oolitic limestones, or as the <i>Jurassic Rocks</i>, +from their being largely developed in the mountain-range of the +Jura, on the western borders of Switzerland. Sediments of this +series occupy extensive areas in Great Britain, on the continent +of Europe, and in India. In North America, limestones and marls +of this age have been detected in "the Black Hills, the Laramie +range, and other eastern ridges of the Rocky Mountains; also over +the Pacific slope, in the Uintah, Wahsatch, and Humboldt Mountains, +and in the Sierra Nevada" (Dana); but in these regions their +extent is still unknown, and their precise subdivisions have +not been determined. Strata belonging to the Jurassic period +are also known to occur in South America, in Australia, and in +the Arctic zone. When fully developed, the Jurassic series is +capable of subdivision into a number of minor groups, of which +some are clearly distinguished by their mineral characters, whilst +others are separated with equal certainty by the differences of +the fossils that they contain. It will be sufficient for our +present purpose, without entering into the more minute subdivisions +of the series, to give here a very brief and general account of the +main sub-groups of the Jurassic rocks, as developed in +Britain—the arrangement of the Jura-formation of the continent +of Europe agreeing in the main with that of England. +</p> + +<p class="indent"> +I. THE LIAS.—The base of the Jurassic series of Britain is +formed by the great calcareo-argillaceous deposit of the "Lias," +which usually rests conformably and almost inseparably upon the +Rhætic beds (the so-called "White Lias"), and passes up, +generally conformably, into the calcareous sandstones of the +Inferior Oolite. The Lias is divisible into the three principal +groups of the Lower, Middle, and Upper Lias, as under, and these +in turn contain many well-marked "zones;" so that the Lias has +some claims to be considered as an independent formation, +equivalent to all the remaining Oolitic rocks. The <i>Lower +Lias</i> (<i>Terrain Sinemurien</i> of D'Orbigny) sometimes +attains a thickness of as much as 600 feet, and consists of a +great series of bluish or greyish laminated clays, +<a name="page_227"><span class="page">Page 227</span></a> +alternating with thin bands of blue or grey +limestone—the whole, when seen in quarries or cliffs from a +little distance, assuming a characteristically striped and banded +appearance. By means of particular species of <i>Ammonites</i>, +taken along with other fossils which are confined to particular +zones, the Lower Lias may be subdivided into several well-marked +horizons. The <i>Middle Lias</i>, or <i>Marlstone Series</i> +(<i>Terrain Liasien</i> of D'Orbigny), may reach a thickness +of 200 feet, and consists of sands, arenaceous marls, and +argillaceous limestones, sometimes with ferruginous beds. The +<i>Upper Lias</i> (<i>Terrain Toarcien</i> of D'Orbigny) attains +a thickness of 300 feet, and consists principally of shales below, +passing upwards into arenaceous strata. +</p> + +<p class="indent"> +II. THE LOWER OOLITES.—Above the Lias comes a complex series +of partly arenaceous and argillaceous, but principally calcareous +strata, of which the following are the more important groups: +<i>a</i>, The <i>Inferior Oolite</i> (<i>Terrain Bajocien</i> +of D'Orbigny), consisting of more than 200 feet of oolitic +limestones, sometimes more or less sandy; <i>b</i>, The <i>Fuller's +Earth</i>, a series of shales, clays, and marls, about 120 feet in +thickness; <i>c</i>, The <i>Great Oolite</i> or <i>Bath Oolite</i> +(<i>Terrain Bathonien</i> of D'Orbigny), consisting principally +of oolitic limestones, and attaining a thickness of about 130 +feet. The well-known "Stonesfield Slates" belong to this horizon; +and the locally developed "Bradford Clay," "Corn brash," and +"Forest-marble" may be regarded as constituting the summit of +this group. +</p> + +<p class="indent"> +III. THE MIDDLE OOLITES.—The central portion of the Jurassic +series of Britain is formed by a great argillaceous deposit, +capped by calcareous strata, as follows: <i>a</i>, The <i>Oxford +Clay</i> (<i>Terrain Callovien</i> and <i>Terrain Oxfordien</i> of +D'Orbigny), consisting of dark-coloured laminated clays, sometimes +reaching a thickness of 700 feet, and in places having its lower +portion developed into a hard calcareous sandstone ("Kelloway +Rock"); <i>b</i>, The Coral-Rag (<i>Terrain Corallien</i> of +D'Orbigny, "Nerinean Limestone" of the Jura, "Diceras Limestone" +of the Alps), consisting, when typically developed, of a central +mass of oolitic limestone, underlaid and surmounted by calcareous +grits. +</p> + +<p class="indent"> +IV. THE UPPER OOLITES.—<i>a</i>, The base of the Upper Oolites +of Britain is constituted by a great thickness (600 feet or more) +of laminated, sometimes carbonaceous or bituminous clays, which are +known as the <i>Kimmeridge Clay</i> (<i>Terrain KimmĂ©ridgien</i> +of D'Orbigny); <i>b</i>, The <i>Portland Beds</i> (<i>Terrain +Portlandien</i> of D'Orbigny) succeed the Kimmeridge clay, and +consist inferiorly of sandy beds surmounted by oolitic limestones +<a name="page_228"><span class="page">Page 228</span></a> +("Portland Stone"), the whole series attaining +a thickness of 150 feet or more, and containing marine fossils; +<i>c</i>, The <i>Purbeck</i> Beds are apparently peculiar to Great +Britain, where they form the summit of the entire Oolitic series, +attaining a total thickness of from 150 to 200 feet. The Purbeck +beds consist of arenaceous, argillaceous, and calcareous strata, +which can be shown by their fossils to consist of a most remarkable +alternation of fresh-water, brackish-water, and purely marine +sediments, together with old land-surfaces, or vegetable soils, +which contain the upright stems of trees, and are locally known as +"Dirt-beds." +</p> + +<p class="indent"> +One of the most important of the Jurassic deposits of the continent +of Europe, which is believed to be on the horizon of the Coral-rag +or of the lower part of the Upper Oolites, is the "<i>Solenhofen +Slate</i>" of Bavaria, an exceedingly fine-grained limestone, +which is largely used in lithography, and is celebrated for the +number and beauty of its organic remains, and especially for +those of Vertebrate animals. +</p> + +<p class="indent"> +The subjoined sketch-section (fig. 159) exhibits in a diagrammatic +form the general succession of the Jurassic rocks of Britain. +</p> + +<p class="indent"> +Regarded as a whole, the Jurassic formation is essentially marine; +and though remains of drifted plants, and of insects and other +air-breathing animals, are not uncommon, the fossils of the formation +are in the main marine. In the Purbeck series of Britain, +anticipatory of the great river-deposit of the Wealden, there are +fresh-water, brackish-water, and even terrestrial strata, indicating +that the floor of the Oolitic ocean was undergoing upheaval, and +that the marine conditions which had formerly prevailed were nearly +at an end. In places also, as in Yorkshire and Sutherlandshire, +are found actual beds of coal: but the great bulk of the formation +is an indubitable sea-deposit; and its limestones, oolitic as +they commonly are, nevertheless are composed largely of the +comminuted skeletons of marine animals. Owing to the enormous +number and variety of the organic remains which have been yielded +by the richly fossiliferous strata of the Oolitic series, it will +not be possible here to do more than to give an outline-sketch +of the principal forms of life which characterise the Jurassic +period as a whole. It is to be remembered, however, that every +minor group of the Jurassic formation has its own peculiar fossils, +and that by the labours of such eminent observers as Quenstedt, +Oppel, D'Orbigny, Wright, De la Beche, Tate, and others, the +entire series of Jurassic sediments admits of a more complete +and more elaborate subdivision into zones +<a name="page_229"><span class="page">Page 229</span></a> +characterised by special life-forms than has as yet been found +practicable in the case of any other rock-series. +</p> + +<div class="center"> +<table border="0" width="545"> +<tr><td><span class="image">GENERALIZED SECTION OF THE JURASSIC +ROCKS OF ENGLAND.</span></td></tr> +<tr><td><span class="image">Fig. 159.</span></td></tr> +<tr><td> +<img src="images/fig159.jpg" width="541" height="708" alt="Fig. 159" /> +</td></tr> +</table> +</div> + +<p class="indent"> +The <i>plants</i> of the Jurassic period consist principally of +Ferns, Cycads, and Conifers—agreeing in this respect, +<a name="page_230"><span class="page">Page 230</span></a> +therefore, with those of the preceding +Triassic formation. The <i>Ferns</i> are very abundant, and +belong partly to old and partly to new genera. The <i>Cycads</i> +are also very abundant, and, on the whole, constitute the most +marked feature of the Jurassic vegetation, many genera of this +group being known (<i>Pterophyllum, Otozamites, Zamites, +Crossozamia, Williamsonia, Bucklandia,</i> &c.) The so-called +"dirt-bed" of the Purbeck series consists of an ancient soil, +in which stand erect the trunks of Conifers and the silicified +stools of Cycads of the genus <i>Mantellia</i> (fig.160). The +<i>ConiferĹ“</i> of the Jurassic are represented by various + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 301px; + font-size: smaller; text-align: center;"> +<img src="images/fig160.jpg" width="301" height="204" alt="Fig. 160" /> +<br /> +Fig. 160.—<i>Mantellia</i> (<i>Cycadeoidea</i>) +<i>megalophylla</i>, a Cycad from the Purbeck "dirt-bed." Upper +Oolites, England. +</span> +</span> + +forms more or less nearly allied to the existing +<i>AraucariĹ“</i>; and these are known not only by their +stems or branches, but also in some cases by their cones. We meet, +also, with the remains of undoubted Endogenous plants, the most +important of which are the fruits of forms allied to the existing +Screw-pines (<i>PandaneĹ“</i>), such as <i>Podocarya</i> and +<i>Kaidacarpum</i>. So far, however, no remains of Palms have been +found; nor are we acquainted with any Jurassic plants which could +be certainly referred to the great "Angiospermous" group of the +Exogens, including the majority of our ordinary plants and trees. +</p> + +<p class="indent"> +Amongst animals, the <i>Protozoans</i> are well represented in +the Jurassic deposits by numerous <i>Foraminifers</i> and +<i>Sponges</i>; as are the <i>CĹ“lenterates</i> by numerous +<i>Corals</i>. Remains of these last-mentioned organisms are +extremely abundant in some of the limestones of the formation, +such as the "Coral-rag" and the Great Oolite; and the former of +these may fairly be considered as an ancient "reef." The <i>Rugose +Corals</i> have not hitherto been detected in the Jurassic rocks; +and the "<i>Tabulate Corals</i>," so-called, are represented +only by examples of the modern genus <i>Millepora</i>. With this +<a name="page_231"><span class="page">Page 231</span></a> +exception, all the Jurassic Corals belong to +the great group which predominates in recent seas (<i>Zoantharia +sclerodermata</i>); and the majority belong to the important +reef-building family of the "Star-corals" (<i>AstrĹ“idoe</i>). +The form here figured (<i>Thecosmilia annularis</i>, fig. 161) is +one of the characteristic species of the Coral-rag. +</p> + +<div class="center"> +<table border="0" width="435"> +<tr><td> +<img src="images/fig161.jpg" width="431" height="342" alt="Fig. 161" /> +</td></tr> +<tr><td class="center"><span class="image"> +Fig. 161.—<i>Thecosmilia annularis</i>, Coral-rag, England. +</span></td></tr> +</table> +</div> + +<p class="indent"> +The <i>Echinoderms</i> are very numerous and abundant fossils +in the Jurassic series, and are represented by Sea-lilies, +Sea-urchins, Star-fishes, and Brittle-stars. The <i>Crinoids</i> +are still common, and some of the limestones of the series are +largely composed of the <i>dĂ©bris</i> of these organisms. +Most of the Jurassic forms resemble those with which we are already +familiar, in having the body permanently attached to some foreign +object by means of a longer or shorter jointed stalk or "column." +One of the most characteristic Jurassic genera of these "stalked" +Crinoids (though not exclusively confined to this period) is +<i>Pentacrinus</i> (fig. 162). In this genus, the column is +five-sided, with whorls of "side-arms;" and the arms are long, +slender, and branched. The genus is represented at the present +day by the beautiful "Medusa-head Pentacrinite" (<i>Pentacrinus +caput-medusĹ“</i>). Another characteristic Oolitic genus is +<i>Apiocrinus</i>, comprising the so-called "Pear Encrinites." In +this group the column is long and rounded, with a dilated base, +and having its uppermost joints expanded so as to form, with the +cup itself, a pear-shaped mass, from the summit of which spring +the comparatively short arms. Besides the +<a name="page_232"><span class="page">Page 232</span></a> +"stalked" Crinoids, the +Jurassic rocks have yielded the remains of the higher group of the + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 426px; + font-size: smaller; text-align: justify;"> +<img src="images/fig162.jpg" width="426" height="743" alt="Fig. 162" /> +<br /> +Fig. 162.—<i>Pentacrinus fasciculos</i>, Lias. The left-hand +figure shows a few or the joints of the column; the middle figure +shows the arms, and the summit of the column with its side-arms; +and the right-hand figure shows the articulating surface of one +of the column-joints. +</span> +</span> + +"free" Crinoids, such as <i>Saccosoma</i>. These forms resemble the +existing "Feather-stars" (<i>Comatula</i>) in being attached when +<a name="page_233"><span class="page">Page 233</span></a> +young to some foreign body by means of a +jointed stem, from which they detach themselves when fully grown +to lead an independent existence. In this later stage of their +life, therefore, they closely resemble the Brittle-stars in +appearance. True Star-fishes (<i>Asteroids</i>) and Brittle-stars +(<i>Ophiuroids</i>) are abundant in the Jurassic rocks, and the +Sea-urchins (<i>Echinoids</i>) are so numerous and so well +preserved as to constitute quite a marked feature of some beds +of the series. All the Oolitic urchins agree with the modern +<i>Echinoids</i> in having the shell composed of no more than +twenty rows of plates. Many different genera are known, and a +characteristic species of the Middle Oolites (<i>Hemicidaris +crenularis</i>, fig. 163) is here figured. +</p> + +<div class="center"> +<table border="0" width="535"> +<tr><td> +<img src="images/fig163.jpg" width="531" height="222" alt="Fig. 163" /> +</td></tr> +<tr><td class="center"><span class="image"> +Fig. 163.—<i>Hemicidaris crenularis</i>, showing the great +tubercles on which the spines were supported. Middle Oolites. +</span></td></tr> +</table> +</div> + +<p class="indent"> +Passing over the <i>Annelides</i>, which, though not uncommon, +are of little special interest, we come to the <i>Articulates</i>, +which also require little notice. Amongst the <i>Crustaceans</i>, +whilst the little Water-fleas (<i>Ostracoda</i>) are still abundant, +the most marked feature is the predominance which is now assumed +by the <i>Decapods</i>—the highest of the known groups of +the class. True Crabs (<i>Brachyura</i>) are by no means unknown; +but the principal Oolitic Decapods belonged to the "Long-tailed" +group (<i>Macrura</i>), of which the existing Lobsters, Prawns, +and Shrimps are members. The fine-grained lithographic slates of +Solenhofen are especially famous as a depot for the remains of +these Crustaceans, and a characteristic species from this locality +(<i>Eryon arctiformis</i>, fig. 164) is here represented. Amongst +the air-breathing <i>Articulates</i>, we meet in the Oolitic +rocks with the remains of Spiders (<i>Arachnida</i>), Centipedes +(<i>Myriapoda</i>), and numerous true Insects (<i>Insecta</i>). +In connection with the last-mentioned of these groups, it is of +interest to note the occurrence of the oldest known fossil +Butterfly—the <i>PalĹ“ontina Oolitica</i> of the +Stonesfield slate—the +<a name="page_234"><span class="page">Page 234</span></a> +relationships of which appear to be with some of the living +Butterflies of Tropical America. +</p> + +<p class="indent"> +Coming to the <i>Mollusca</i>, the <i>Polyzoans</i>, numerous + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 321px; + font-size: smaller; text-align: center;"> +<img src="images/fig164.jpg" width="321" height="452" alt="Fig. 164" /> +<br /> +Fig. 164.—<i>Eryon arctiformis</i>, a "Long-tailed Decapod," +from the Middle Oolites (Solenhofen Slate). +</span> +</span> + +and beautiful as they are, must be at once dismissed; but the +<i>Brachiopods</i> deserve a moment's attention. The Jurassic +Lamp-shells (fig. 165) do not fill by any means such a predominant +place in the marine fauna of the period, as in many Palæozoic +deposits, but they are still individually numerous. The two ancient +genera <i>LeptĹ“na</i> (fig. 165, <i>a</i>) and <i>Spirifera</i> +(fig. 165, <i>b</i>), dating the one from the Lower and the other +from the Upper Silurian, appear here for the last time upon the +scene, but they have not hitherto been recognised in deposits later +than the Lias. The great majority of the Jurassic <i>Brachiopods</i>, +however, belong to the genera <i>Terebratula</i> (fig. 165, <i>c, +e, f</i>) and <i>Rhynchonella</i> (fig. 165. <i>d</i>), both of +which are represented by living forms at the present day. The +<i>TerebratulĹ“</i>, in particular, are very abundant, and +the species are often confined to special horizons in the series. +</p> + +<p class="indent"> +Remains of <i>Bivalves</i> (<i>Lamellibranchiata</i>) are very +<a name="page_235"><span class="page">Page 235</span></a> +numerous in the +Jurassic deposits, and in many cases highly characteristic. +In the marine beds of the Oolites, which constitute + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 520px; + font-size: smaller; text-align: justify;"> +<img src="images/fig165.jpg" width="520" height="370" alt="Fig. 165" /> +<br /> +Fig. 165.—Jurassic +Brachiopod. <i>a. LeptĹ“na Liassica</i>, enlarged, the small +cross below the figure indicating the true size of the +shell—Lias; <i>b, Spirifera rostrata</i>, Lias; +<i>c, Terebratula quadrifida</i>, Lias; <i>d, d', Rhynchonella +varians</i>, Fulter's Earth and Kelloway Rock; <i>e, Terebratula +sphĹ“roidalis</i>, Inferior Oolite; <i>f, Terebratula +digona</i>, Bradford Clay, Forest-marble, and Great Oolite. +(After Davidson). +</span> +</span> + +by far the greater portion of the whole formation, the Bivalyes +are of course marine, and belong to such genera as <i>Trigonia, +Lima, Pholadomya, Cardinia, Avicula, Hippopodium</i>, &c.; +but in the Purbeck beds, at the summit of the series, we find +bands of Oysters alternating with strata containing fresh-water +or brackish-water Bivalves, such as <i>CyrenĹ“</i> and +<i>CorbulĹ“</i>. The predominant Bivalves of the Jurassic, +however, are the <i>Oysters</i>, which occur under many forms, +and often in vast numbers, particular species being commonly +restricted to particular horizons. Thus of the true Oysters, +<i>Ostrea distorta</i> is characteristic of the Purbeck series, +where it forms a bed twelve feet in thickness, known locally as +the "Cinder-bed;" <i>Ostrea expansa</i> abounds in the Portland +beds; <i>Ostrea deltoidea</i> is characteristic of the Kimmeridge +clay; <i>Ostrea gregaria</i> predominates in the Coral-rag; +<i>Ostrea acuminata</i> characterises the small group of the +Fuller's Earth; whilst the plaited <i>Ostrea Marshii</i> (fig. +166) is a common shell in the Lower and Middle Oolites. Besides +the more typical Oysters, the Oolitic rocks abound in examples +of the singularly unsymmetrical forms belonging +<a name="page_236"><span class="page">Page 236</span></a> +to the genera <i>Exogyra</i> and <i>GryphĹ“a</i> (fig. 167). +In the former of these are included Oysters with the beaks + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 228px; vertical-align:bottom; text-align: center; + margin: 4px;"> +<img src="images/fig166.jpg" width="228" height="206" alt="Fig. 166" /> +<br /> +Fig. 166.—<i>Ostrea Marshii</i>. Middle and Lower Oolites. +</span> + +<span style="width: 255px; vertical-align:bottom; text-align: center; + margin: 4px;"> +<img src="images/fig167.jpg" width="255" height="194" alt="Fig. 167" /> +<br /> +Fig. 167.—<i>GryphĹ“a incurva</i>. Lias. +</span> + +</span> + +"reversed"—that is to say, turned towards the hinder part +of the shell; whilst in the latter are Oysters in which the lower +valve of the shell is much the largest, and has a large incurved +beak, whilst the upper valve is small and concave. One of the +most characteristic <i>ExogyrĹ“</i> is the <i>E. +Virgula</i> of the Oxford Clay, and of the same horizon on the +Continent; and the <i>GryphĹ“a incurva</i> (fig. 167) is +equally abundant in, and characteristic of, the formation of +the Lias. Lastly, we may notice the extraordinary shells +belonging to the genus <i>Diceras</i> (fig. 168), which are + +<span style="float: left; margin: 4px; width: 254px; + font-size: smaller; text-align: center;"> +<img src="images/fig168.jpg" width="242" height="215" alt="Fig. 168" /> +<br /> +Fig. 168.—<i>Diceras arietina</i>. Middle Oolite. +</span> + +exclusively confined to the Middle Oolites. In this formation +in the Alps they occur in such abundance as to give rise to the +name of "Calcaire à Dicerates," applied to beds of the +same age as the Coral-rag of Britain. The genus <i>Diceras</i> +belongs to the same family as the "Thorny Clams" (Chama) of the +present day—the shell being composed of nearly equally-sized +valves, the beaks of which are extremely prominent and twisted +into a spiral. The shell was attached to some foreign body by +the beak of one of its valves. +</p> + +<p class="indent"> +Amongst the Jurassic Univalves (<i>Gasteropoda</i>) there are +many examples of the ancient and long-lived <i>Pleurotomaria</i>; +but on the whole the Univalves begin to have a modern aspect. +The round-mouthed ("holostomatous"), vegetable-eating +<a name="page_237"><span class="page">Page 237</span></a> +Sea-snails, such as the Limpets (<i>PatellidĹ“</i>), the Nerites +(<i>Nerita</i>), the <i>TurritellĹ“, ChemnitziĹ“</i>, &c., +still hold a predominant place. The two most noticeable genera of +this group are <i>Cerithium</i> and <i>NerinĹ“a</i>—the +former of these attaining great importance in the Tertiary and Recent +seas, whilst the latter (fig. 169) is highly characteristic of the +Jurassic series, though not exclusively confined to it. One of the + +<span style="float: right; margin: 4px; width: 254px; + font-size: smaller; text-align: justify;"> +<img src="images/fig169.jpg" width="209" height="245" alt="Fig. 169" /> +<br /> +Fig. 169.—<i>NerinĹ“a Goodhallii</i>, one-fourth of the +natural size. The left-hand figure shows the appearance presented +by the shell when vertically divided. Coral-rag, England. +</span> + +limestones of the Jura, believed to be of the age of the Coral-rag +(Middle Oolite) of Britain, abounds to such an extent in the turreted +shells of <i>NerinĹ“a</i> as to have gained the name of "Calcaire +à NĂ©rinĂ©es." In addition to forms such as the +preceding, we now for the first time meet, in any force, with the +Carnivorous Univalves, in which the mouth of the shell is notched or +produced into a canal, giving rise to the technical name of +"siphonostomatous" applied to the shell. Some of the carnivorous +forms belong to extinct types, such as the <i>Purpuroidea</i> of the +Great Oolite; but others are referable to well-known existing genera. +Thus we meet here with species of the familiar groups of the Whelks +(<i>Buccinum</i>), the Spindle-shells (<i>Fusus</i>), the Spider-shells +(<i>Pteroceras</i>), <i>Murex, Rostellaria</i>, and others which are +not at present known to occur in any earlier formation. +</p> + +<p class="indent"> +Amongst the Wing-shells (<i>Pteropoda</i>), it is sufficient +to mark the final appearance in the Lias of the ancient genus +<i>Conularia</i>. +</p> + +<p class="indent"> +Lastly, the order of the <i>Cephalopoda</i>, in both its +Tetrabranchiate and Dibranchiate sections, undergoes a vast +development in the Jurassic period. The old and comparatively +simple genus <i>Nautilus</i> is still well represented, one +species being very similar to the living Pearly Nautilus (<i>N. +Pompilius</i>); but the <i>Orthocerata</i> and <i>Goniatites</i> +of the Trias have finally disappeared; and the great majority +of the Tetrabranchiate forms are referable to the comprehensive +genus <i>Ammonites</i>, with its many sub-genera and its hundreds +of recorded species. The shell in <i>Ammonites</i> is in the +form of a flat spiral, all the coils of which are in contact +(figs. 170 and 171). The innermost whorls of the shell are more +or less concealed; and the body-chamber is elongated and narrow, +rather than expanded towards the mouth. The tube or siphuncle +<a name="page_238"><span class="page">Page 238</span></a> +which runs through the air-chambers is placed on the dorsal or +<i>convex</i> side of the shell; but the principal character +which distinguishes <i>Ammonites</i> from <i>Goniatites</i> and + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 480px; + font-size: smaller; text-align: center;"> +<img src="images/fig170.jpg" width="480" height="312" alt="Fig. 170" /> +<br /> +Fig. 170.—<i>Ammonites Humphresianus</i>. Inferior Oolite. +</span> +</span> + +<i>Ceratites</i> is the wonderfully complex manner in which the +<i>septa</i>, or partitions between the air-chambers, are folded and +undulated. To such an extent does this take place, that the edges +of the septa, when exposed by the removal of the shell-substance, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 403px; + font-size: smaller; text-align: center;"> +<img src="images/fig171.jpg" width="403" height="288" alt="Fig. 171" /> +<br /> +Fig. 171.—<i>Ammonites bifrons</i>. Lias. +</span> +</span> + +present in an exaggerated manner the appearance exhibited by +an elaborately-dressed shirt-frill when viewed edgewise. The +species of <i>Ammonites</i> range from the Carboniferous to the +Chalk; but they have not been found in +<a name="page_239"><span class="page">Page 239</span></a> +deposits older than the Secondary, in any region except India; +and they are therefore to be regarded as essentially Mesozoic +fossils. Within these limits, each formation is characterised +by particular species, the number of individuals being often +very great, and the size which is sometimes attained being +nothing short of gigantic. In the Lias, particular species of +<i>Ammonites</i> may succeed one another regularly, each having +a more or less definite horizon, which it does not transgress. +It is thus possible to distinguish a certain number of zones, +each characterised by a particular Ammonite, together with +other associated fossils. Some of these zones are very +persistent and extend over very wide areas, thus affording +valuable aid to the geologist in his determination of rocks. +It is to be remembered, however, that there are other species +which are not thus restricted in their vertical range, even +in the same formations in which definite zones occur. +</p> + +<p class="indent"> +The Cuttle-fishes or <i>Dibranchiate Cephalopods</i> constitute a +feature in the life of the Jurassic period little less conspicuous +and striking than that afforded by the multitudinous and varied +chambered shells of the <i>AmmonitidĹ“</i>. The remains by +which these animals are recognised are necessarily less perfect, as +a rule, than those of the latter, as no external shell is present +(except in rare and more modern groups), and the internal skeleton +is not necessarily calcareous. Nevertheless,we have an ample +record of the Cuttle-fishes of the Jurassic period, in the shape +of the fossilised jaws or beak, the ink-bag, and, most commonly +of all, the horny or calcareous structure which is embedded in +the soft tissues, and is variously known as the "pen" or "bone." +The beaks of Cuttle-fishes, though not abundant, are sufficiently +plentiful to have earned for themselves the general title of +"Rhyncholites;" and in their form and function they resemble the +horny, parrot-like beak of the existing Cephalopods. The ink-bag +or leathery sac in which the Cuttle-fishes store up the black +pigment with which they obscure the water when attacked, owes its +preservation to the fact that the colouring-matter which it contains +is finely-divided carbon, and therefore nearly indestructible except +by heat. Many of these ink-bags have been found in the Lias; and +the colouring-matter is sometimes so well preserved that it has +been, as an experiment, employed in painting as a fossil "sepia." +The "pens" of the Cuttle-fishes are not commonly preserved, owing +to their horny consistence, but they are not unknown. The form +here figured (<i>Beloteuthis subcostata</i>, fig. 172) belonged +to an old type essentially similar to our modern Calamaries, the +skeleton of which consists of a horny shaft +<a name="page_240"><span class="page">Page 240</span></a> +and two lateral wings, +somewhat like a feather in general shape. When, on the other + +<span style="float: left; margin: 4px; width: 123px; + font-size: smaller; text-align: justify;"> +<img src="images/fig172.jpg" width="123" height="306" alt="Fig. 172" /> +<br /> +Fig. 172.—<i>Beloteuthis subcostata</i> Jurassic (Lias). +</span> + +hand, the internal skeleton is calcareous, then it is very easily +preserved in a fossil condition; and the abundance of remains of +this nature in the Secondary rocks, combined with their apparent +total absence in Palæozoic strata, is a strong presumption in +favour of the view that the order of the Cuttle-fishes did not +come into existence till the commencement of the Mesozoic period. +The great majority of the skeletons of this kind which are found +in the Jurassic rocks belong to the great extinct family of the +"Belemnites" (<i>Belemnitidoa</i>), which, so far as known, is +entirely confined to rocks of Secondary age. From its pointed, +generally cylindro-conical form, the skeleton of the Belemnite is +popularly known as a "thunderbolt". (fig. 173, C). In its perfect +condition—in which it is, however, rarely obtainable—the +skeleton consists of a chambered conical shell (the "phragmacone"), +the partitions between the chambers of which are pierced by a +marginal tube or "siphuncle." This conical shell—curiously +similar in its structure to the <i>external</i> shell of the +Nautilus—is extended forwards into a horny "pen," and is sunk +in a corresponding conical pit (fig. 173, B), excavated in the +substance of a nearly cylindrical fibrous body or "guard," which +projects backwards for a longer or shorter distance, and is the +part most usually found in a fossil condition. Many different +kinds of <i>Belemnites</i> are known, and their guards literally +swarm in many parts of the Jurassic series, whilst some specimens +attain very considerable dimensions. Not only is the internal +skeleton known, but specimens of <i>Belemnites</i> and the nearly +allied <i>Belemnoteuthis</i> have been found in some of the +fine-grained sediments of the Jurassic formation, from which much +has been learnt even as to the anatomy of the soft parts of the +animal. Thus we know that the Belemnites were in many respects +comparable with the existing Calamaries or Squids, the body being +furnished with lateral fins, and the head carrying a circle of ten +"arms," two of which were longer than the others (fig. 173, A). +The suckers on the arms were provided, further, with horny hooks; +there was a large ink-sac; and the mouth was armed with horny +mandibles resembling in shape the beak of a parrot. +</p> + +<p class="indent"> +Coming next to the <i>Vertebrates</i>, we find that the Jurassic +<a name="page_241"><span class="page">Page 241</span></a> +<i>Fishes</i> are still represented by <i>Ganoids</i> and +<i>Placoids</i>. The Ganoids, however, unlike the old forms, now + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 485px; + font-size: smaller; text-align: justify;"> +<img src="images/fig173.jpg" width="485" height="434" alt="Fig. 173" /> +<br /> +Fig. 173.—A, Restoration of the animal of the Belemnite; B, +Diagram showing the complete skeleton of a Belemnite, consisting +of the chambered phragmacone (<i>a</i>), the guard (<i>b</i>), +and the horny pen (<i>c</i>); C, Specimen of <i>Belemnites +canaliculatus</i>, from the Inferior Oolite. (After Phillips.) +</span> +</span> + +for the most part possess nearly or quite symmetrical ("homocercal") +tails. A characteristic genus is <i>Tetragonolepis</i> (fig. 174), + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 575px; + font-size: smaller; text-align: center;"> +<img src="images/fig174.jpg" width="515" height="220" alt="Fig. 174" /> +<br /> +Fig. 174.—<i>Tetragonolepis</i> (restored), and scales of the +same. Lias. +</span> +</span> + +with its deep compressed body, its rhomboidal, closely-fitting +scales, and its single long dorsal fin. Amongst the <i>Placoids</i> +<a name="page_242"><span class="page">Page 242</span></a> +the teeth of true Sharks (<i>Notidanus</i>) occur for the first +time; but by far the greater number of remains referable to this +group are still the fin-spines and teeth of "Cestracionts," +resembling the living Port-Jackson Shark. Some of these teeth are +pointed (<i>Hybodus</i>); but others are rounded, and are adapted +for crushing shell-fish. Of these latter, the commonest are the +teeth of <i>Acrodus</i> (fig. 175), of which the hinder ones are + +<span style="float: left; margin: 4px; width: 254px; + font-size: smaller; text-align: center;"> +<img src="images/fig175.jpg" width="278" height="107" alt="Fig. 175" /> +<br /> +Fig. 175.—Tooth of <i>Acrodus nobilis</i>. Lias. +</span> + +of an elongated form, with a rounded surface, covered with fine +transverse striæ proceeding from a central longitudinal line. +From their general form and striation, and their dark colour, these +teeth are commonly called "fossil leeches" by the quarrymen. +</p> + +<p class="indent"> +The Amphibian group of the <i>Labyrinthodonts</i>, which was +so extensively developed in the Trias, appears to have become +extinct, no representative of the order having hitherto been +detected in rocks of Jurassic age. +</p> + +<p class="indent"> +Much more important than the Fishes of the Jurassic series are +the <i>Reptiles</i>, which are both very numerous, and belong to +a great variety of types, some of these being very extraordinary +in their anatomical structure. The predominant group is that +of the "Enaliosaurs" or "Sea-lizards," divided into two great +orders, represented respectively by the <i>Ichthyosaurus</i> +and the <i>Plesiosaurus</i>. +</p> + +<p class="indent"> +The <i>Ichthyosauri</i> or "Fish-Lizards" are exclusively Mesozoic +in their distribution, ranging from the Lias to the Chalk, but +abounding especially in the former. They were huge Reptiles, of +a fish-like form, with a hardly conspicuous neck (fig. 176), +and probably possessing a simply smooth or wrinkled skin, since + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 498px; + font-size: smaller; text-align: center;"> +<img src="images/fig176.jpg" width="498" height="122" alt="Fig. 176" /> +<br /> +Fig. 176.—<i>Ichthyosaurus communis</i>. Lias. +</span> +</span> + +no traces of scales or bony integumentary plates have ever been +discovered. The tail was long, and was probably furnished at its +extremity with a powerful expansion of the skin, constituting a +tail-fin similar to that possessed by the Whales. The limbs are +also like those of +<a name="page_243"><span class="page">Page 243</span></a> +Whales in the essentials +of their structure, and in their being adapted to act as +swimming-paddles. Unlike the Whales, however, the Ichthyosaurs +possessed the hind-limbs as well as the fore-limbs, both pairs +having the bones flattened out and the fingers completely +enclosed in the skin, the arm and leg being at the same time +greatly shortened. The limbs are thus converted into efficient +"flippers," adapting the animal for an active existence in the +sea. The different joints of the backbone (vertebræ) also +show the same adaptation to an aquatic mode of life, being +hollowed out at both ends, like the biconcave vertebræ +of Fishes. The spinal column in this way was endowed with the +flexibility necessary for an animal intended to pass the +greater part of its time in water. Though the <i>Ichthyosaurs</i> +are undoubtedly marine animals, there is, however, reason to +believe that they occasionally came on shore, as they possess +a strong bony arch, supporting the fore-limbs, such as would +permit of partial, if laborious, terrestrial progression. The +head is of enormous size, with greatly prolonged jaws, holding +numerous powerful conical teeth lodged in a common groove. The +nature of the dental apparatus is such as to leave no doubt as to +the rapacious and predatory habits of the Ichthyosaurs—an +inference which is further borne out by the examination of their +petrified droppings, which are known to geologists as "coprolites," +and which contain numerous fragments of the bones and scales +of the Ganoid fishes which inhabited the same seas. The orbits +are of huge size; and as the eyeball was protected, like that +of birds, by a ring of bony plates in its outer coat, we even +know that the pupils of the eyes were of correspondingly large +dimensions. As these bony plates have the function of protecting the +eye from injury under sudden changes of pressure in the surrounding +medium, it has been inferred, with great probability, that the +Ichthyosaurs were in the habit of diving to considerable depths +in the sea. Some of the larger specimens of <i>Ichthyosaurus</i> +which have been discovered in the Lias indicate an animal of +from 20 to nearly 40 feet in length; and many species are known +to have existed, whilst fragmentary remains of their skeletons +are very abundant in some localities. We may therefore safely +conclude that these colossal Reptiles were amongst the most +formidable of the many tyrants of the Jurassic seas. +</p> + +<p class="indent"> +The <i>Plesiosaurus</i> (fig. 177) is another famous Oolitic +Reptile, and, like the preceding, must have lived mainly or +exclusively in the sea. It agrees with the Ichthyosaur in some +important features of its organisation, especially in the fact +that both pairs of limbs are converted into "flippers" or +<a name="page_244"><span class="page">Page 244</span></a> +swimming-paddles, +whilst the skin seems to have been equally destitute +of any scaly or bony investiture. Unlike the <i>Ichthyosaur</i>, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 575px; + font-size: smaller; text-align: center;"> +<img src="images/fig177.jpg" width="508" height="299" alt="Fig. 177" /> +<br /> +Fig. 177.—<i>Plesiosaurus dolichodeirus</i>, restored. Lias. +</span> +</span> + +however, the Plesiosaur had the paddles placed +far back, the tail being extremely short, and the neck greatly +lengthened out, and composed of from twenty to forty vertebræ. +The bodies of the vertebræ, also, are not deeply biconcave, but +are flat, or only slightly cupped. The head is of relatively small +size, with smaller orbits than those of the <i>Ichthyosaur</i>, +and with a snout less elongated. The jaws, however, were armed +with numerous conical teeth, inserted in distinct sockets. As +regards the habits of the Plesiosaur, Dr Conybeare arrives at +the following conclusions: "That it was aquatic is evident from +the form of its paddles; that it was marine is almost equally so +from the remains with which it is universally associated; that +it may have occasionally visited the shore, the resemblance of +its extremities to those of the Turtles may lead us to conjecture: +its movements, however, must have been very awkward on land; +and its long neck must have impeded its progress through the +water, presenting a strong contrast to the organisation which so +admirably fits the <i>Ichthyosaurus</i> to cut through the waves." +As its respiratory organs were such that it must of necessity +have required to obtain air frequently, we may conclude "that +it swam upon or near the surface, arching back its long neck +like a swan, and occasionally darting it down at the fish which +happened to float within its reach. It may perhaps have lurked +in shoal water along the coast, concealed amongst the sea-weed; +and raising its nostrils to a level with +<a name="page_245"><span class="page">Page 245</span></a> +the surface from a considerable depth, may have found a secure +retreat from the assaults of powerful enemies; while the length +and flexibility of its neck may have compensated for the want +of strength in its jaws, and its incapacity for swift-motion +through the water." +</p> + +<p class="indent"> +About twenty species of <i>Plesiosaurus</i> are known, ranging +from the Lias to the Chalk, and specimens have been found indicating +a length of from eighteen to twenty feet. The nearly related +"<i>Pliosaurs</i>," however, with their huge heads and short +necks, must have occasionally reached a length of at least forty +feet—the skull in some species being eight, and the paddles +six or seven feet long, whilst the teeth are a foot in length. +</p> + +<p class="indent"> +Another extraordinary group of Jurassic Reptiles is that of the +"Winged Lizards" or <i>Pterosauria</i>. These are often spoken +of collectively as "Pterodactyles," from <i>Pterodactylus</i>, +the type-genus of the group. As now restricted, however, the +genus <i>Pterodactylus</i> is more Cretaceous than Jurassic, and +it is associated in the Oolitic rocks with the closely allied +genera <i>Dimorphodon</i> and <i>Rhamphorhynchus</i>. In all +three of these genera we have the same general structural +organisation, involving a marvellous combination of characters, +which we are in the habit of regarding as peculiar to Birds on +the one hand, to Reptiles on another hand, and to the Flying +Mammals or Bats in a third direction. The "Pterosaurs" are "Flying" +Reptiles, in the true sense of the term, since they were indubitably +possessed of the power of active locomotion in the air, after the +manner of Birds. The so-called "Flying" Reptiles of the present +day, such as the little <i>Draco volans</i> of the East Indies +and Indian Archipelago, possess, on the other hand, no power +of genuine flight, being merely able to sustain themselves in +the air through the extensive leaps which they take from tree +to tree, the wing-like expansions of the skin simply exercising +the mechanical function of a parachute. The apparatus of flight +in the "Pterosaurs" is of the most remarkable character, and most +resembles the "wing" of a Bat, though very different in some +important particulars. The "wing" of the Pterosaurs is like that +of Bats, namely, in consisting of a thin leathery expansion of the +skin which is attached to the sides of the body, and stretches +between the fore and hind limbs, being mainly supported by an +enormous elongation of certain of the digits of the hand. In +the Bats, it is the four outer fingers which are thus lengthened +out; but in the Pterosaurs, the wing-membrane is borne by a single +immensely-extended finger (fig. 178). No trace of the actual +wing-membrane itself has, of course, +<a name="page_246"><span class="page">Page 246</span></a> +been found fossilised; but +we could determine that the "Pterodactyles" possessed the power +of flight, quite apart from the extraordinary conformation of + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 431px; + font-size: smaller; text-align: justify;"> +<img src="images/fig178.jpg" width="431" height="376" alt="Fig. 178" /> +<br /> +Fig. 178.—<i>Pterodactylus crassirostis</i>. From the +Lithographic Slates of Solenhofen (Middle Oolite). The figure is +"restored," and it seems certain that the restoration is incorrect +in the comparatively unimportant particular, that the hand should +consist of no more than four fingers, three short and one long, +instead of five, as represented. +</span> +</span> + +the hand. The proofs of this are to be found partly in the fact +that the breast-bone was furnished with an elevated ridge or +keel, serving for the attachment of the great muscles of flight, +and still more in the fact that the bones were hollow and were +filled with air—a peculiarity wholly confined amongst living +animals to Birds only. The skull of the Pterosaurs is long, light, +and singularly bird-like in appearance—a resemblance which is +further increased by the comparative length of the neck and the size +of the vertebræ of this region (fig. 178). The jaws, however, +unlike those of any existing Bird, were, with one exception to be +noticed hereafter, furnished with conical teeth sunk in distinct +sockets; and there was always a longer or shorter tail composed +of distinct vertebræ; whereas in all existing Birds the tail +is abbreviated, and the terminal vertebræ are amalgamated to +form a single bone, which generally supports the great feathers +of the tail. +</p> + +<p class="indent"> +Modern naturalists have been pretty generally agreed that the +<i>Pterosaurs</i> should be regarded as a peculiar group of the +Reptiles; though they have been and are still regarded by high +authorities, like Professor Seeley, as being really referable +<a name="page_247"><span class="page">Page 247</span></a> +to the Birds, or as forming a class by +themselves. The chief points which separate them from Birds, as +a class, are the character of the apparatus of flight, the +entirely different structure of the fore-limb, the absence of +feathers, the composition of the tail out of distinct +vertebræ, and the general presence of conical teeth sunk +in distinct sockets in the jaws. The gap between the Pterosaurs +and the Birds has, however, been greatly lessened of late by +the discovery of fossil animals (<i>Ichthyornis</i> and +<i>Hesperornis</i>) with the skeleton proper to Birds combined +with the presence of teeth in the jaws, and by the still more +recent discovery of other fossil animals (<i>Pteranodon</i>) +with a Pterosaurian skeleton, but without teeth; whilst the +undoubtedly feathered <i>ArchĹ“opteryx</i> possessed a long +tail composed of separate vertebræ. Upon the whole, therefore, +the relationships of the Pterosaurs cannot be regarded as absolutely +settled. It seems certain, however, that they did not possess +feathers—this implying that they were cold-blooded animals; +and their affinities with Reptiles in this, as in other characters, +are too strong to be overlooked. +</p> + +<p class="indent"> +The <i>Pterosaurs</i> are wholly Mesozoic, ranging from the Lias +to the Chalk inclusive; and the fine-grained Lithographic Slate +of Solenhofen has proved to be singularly rich in their remains. +The genus <i>Pterodactylus</i> itself has the jaws toothed to the +extremities with equal-sized conical teeth, and its species range +from the Middle Oolites to the Cretaceous series, in connection +with which they will be again noticed, together with the toothless +genus <i>Pteranodon</i>. The genus <i>Dimorphodon</i> is Liassic, +and is characterised by having the front teeth long and pointed, +whilst the hinder teeth are small and lancet-shaped. Lastly, +the singular genus <i>Rhamphorhynchus</i>, also from the Lower +Oolites, is distinguished by the fact that there are teeth present +in the hinder portions of both jaws; but the front portions are +toothless, and may have constituted a horny beak. Like most of +the other Jurassic Pterosaurs, <i>Rhamphorhynchus</i> (fig. 179) +does not seem to have been much bigger than a pigeon, in this +respect falling far below the giant "Dragons" of the Cretaceous +period. It differed from its relatives, not only in the armature of +the mouth, but also in the fact that the tail was of considerable +length. With regard to its habits and mode of life, Professor +Phillips remarks that, "gifted with ample means of flight, able +at least to perch on rocks and scuffle along the shore, perhaps +competent to dive, though not so well as a Palmiped bird, many +fishes must have yielded to the cruel beak and sharp teeth of +Rhamphorhynchus. If we ask to which of the many families of Birds +the analogy of +<a name="page_248"><span class="page">Page 248</span></a> +structure and probable way of life would lead us +to assimilate Rhamphorhynchus, the answer must point to the +swimming races with long wings, clawed feet, hooked beak, and + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 428px; + font-size: smaller; text-align: center;"> +<img src="images/fig179.jpg" width="428" height="176" alt="Fig. 179" /> +<br /> +Fig. 179—<i>Rhamphorhynchus Bucklandi</i>, restored. Bath +Oolite, England. (After the late Professor Phillips.) +</span> +</span> + +habits or violence and voracity; and for preference, the shortness +of the legs, and other circumstances, may be held to claim for +the Stonesfield fossil a more than fanciful similitude to the +groups of Cormorants, and other marine divers, which constitute +an effective part of the picturesque army of robbers of the sea." +</p> + +<p class="indent"> +Another extraordinary and interesting group of the Mesozoic Reptiles +is constituted by the <i>Deinosauria</i>, comprising a series of +mostly gigantic forms, which range from the Trias to the Chalk. +All the "Deinosaurs" are possessed of the two pairs of limbs +proper to Vertebrate animals, and these organs are in the main +adapted for walking on the dry land. Thus, whilst the Mesozoic +seas swarmed with the huge Ichthyosaurs and Plesiosaurs, and +whilst the air was tenanted by the Dragon-like Pterosaurs, the +land-surfaces of the Secondary period were peopled by numerous +forms of Deinosaurs, some of them of even more gigantic dimensions +than their marine brethren. The limbs of the <i>Deinosaurs</i> +are, as just said, adapted for progression on the land; but in +some cases, at any rate, the hind-limbs were much longer and +stronger than the fore-limbs; and there seems to be no reason to +doubt that many of these forms possessed the power of walking, +temporarily or permanently, on their hind-legs, thus presenting +a singular resemblance to Birds. Some very curious and striking +points connected with the structure of the skeleton have also +been shown to connect these strange Reptiles with the true Birds; +and such high authorities as Professors Huxley and Cope are of +opinion that the Deinosaurs are distinctly related to this class, +being in some respects intermediate between the proper Reptiles +and the great wingless Birds, like the Ostrich and Cassowary. +On the other hand, Professor Owen has shown that the Deinosaurs +<a name="page_249"><span class="page">Page 249</span></a> +possess some weighty points of relationship +with the so-called "Pachydermatous" Quadrupeds, such as the +Rhinoceros and Hippopotamus. The most important Jurassic genera +of <i>Deinosauria</i> are <i>Megalosaurus</i> and +<i>Cetiosaurus</i>, both of which extend their range into the +Cretaceous period, in which flourished, as we shall see, some +other well-known members of this order. +</p> + +<p class="indent"> +<i>Megalosaurus</i> attained gigantic dimensions, its thigh and +shank bones measuring each about three feet in length, and its +total length, including the tail, being estimated at from forty +to fifty feet. As the head of the thigh-bone is set on nearly at +right angles with the shaft, whilst all the long bones of the +skeleton are hollowed out internally for the reception of the +marrow, there can be no doubt as to the terrestrial habits of +the animal. The skull (fig. 180) was of large size, four or five + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 492px; + font-size: smaller; text-align: center;"> +<img src="images/fig180.jpg" width="492" height="299" alt="Fig. 180" /> +<br /> +Fig. 180.—Skull of <i>Megalosaurus</i>, on a scale +one-tenth of nature. Restored. (After Professor Phillips.) +</span> +</span> + +feet in length, and the jaws were armed with a series of powerful +pointed teeth. The teeth are conical in shape, but are strongly +compressed towards their summits, their lateral edges being finely +serrated. In their form and their saw-like edges, they resemble +the teeth of the "Sabre-toothed Tiger" (<i>Machairodus</i>), and +they render it certain that the Megalosaur was in the highest +degree destructive and carnivorous in its habits. So far as is +known, the skin was not furnished with any armour of scales or bony +plates; and the fore-limbs are so disproportionately small as +compared with the hind-limbs, that this huge Reptile—like +the equally huge Iguanodon—may be +<a name="page_250"><span class="page">Page 250</span></a> +conjectured to have commonly supported itself on its hind-legs +only. +</p> + +<p class="indent"> +The <i>Cetiosaur</i> attained dimensions even greater than those +of the Megalosaur, one of the largest thigh-bones measuring over +five feet in length and a foot in diameter in the middle, and the +total length of the animal being probably not less than fifty +feet. It was originally regarded as a gigantic Crocodile, but +it has been shown to be a true Deinosaur. Having obtained a +magnificent series of remains of this reptile, Professor Phillips +has been able to determine many very interesting points as to +the anatomy and habits of this colossal animal, the total length +of which he estimates as being probably not less than sixty or +seventy feet. As to its mode of life, this accomplished writer +remarks:— +</p> + +<p class="indent"> +"Probably when 'standing at ease' not less than ten feet in height, +and of a bulk in proportion, this creature was unmatched in magnitude +and physical strength by any of the largest inhabitants of the +Mesozoic land or sea. Did it live in the sea, in fresh waters, +or on the land? This question cannot be answered, as in the case +of Ichthyosaurus, by appeal to the accompanying organic remains; +for some of the bones lie in marine deposits, others in situations +marked by estuarine conditions, and, out of the Oxfordshire district, +in Sussex, in fluviatile accumulations. Was it fitted to live +exclusively in water? Such an idea was at one time entertained, +in consequence of the biconcave character of the caudal vertebræ, +and it is often suggested by the mere magnitude of the creature, +which would seem to have an easier life while floating in water, +than when painfully lifting its huge bulk, and moving with slow +steps along the ground. But neither of these arguments is valid. The +ancient earth was trodden by larger quadrupeds than our elephant; +and the biconcave character of vertebræ, which is not uniform +along the column in Cetiosaurus, is perhaps as much a character +of a geological period as of a mechanical function of life. Good +evidence of continual life in water is yielded in the case of +Ichthyosaurus and other Enaliosaurs, by the articulating surfaces +of their limb-bones, for these, all of them, to the last phalanx, +have that slight and indefinite adjustment of the bones, with much +intervening cartilage, which fits the leg to be both a flexible +and forcible instrument of natation, much superior to the ordinary +oar-blade of the boatman. On the contrary, in Cetiosaur, as well as +in Megalosaur and Iguanodon, all the articulations are definite, +and made so as to correspond to determinate movements in particular +directions, and these are such as to be suited +<a name="page_251"><span class="page">Page 251</span></a> +for walking. In particular, the femur, by its head projecting +freely from the acetabulum, seems to claim a movement of free +stepping more parallel to the line of the body, and more +approaching to the vertical than the sprawling gait of the +crocodile. The large claws concur in this indication of +terrestrial habits. But, on the other hand, these characters are +not contrary to the belief that the animal may have been +amphibious; and the great vertical height of the anterior part +of the tail seems to support this explanation, but it does not +go further.... We have therefore a marsh-loving or river-side +animal, dwelling amidst filicine, cycadaceous, and coniferous +shrubs and trees full of insects and small mammalia. What was +its usual diet? If <i>ex ungue leonem</i>, surely <i>ex dente +cibum</i>. We have indeed but one tooth, and that small and +incomplete. It resembles more the tooth of Iguanodon than that +of any other reptile; for this reason it seems probable that +the animal was nourished by similar vegetable food which +abounded in the vicinity, and was not obliged to contend with +Megalosaurus for a scanty supply of more stimulating diet." +</p> + +<p class="indent"> +All the groups of Jurassic Reptiles which we have hitherto been +considering are wholly unrepresented at the present day, and +do not even pass upwards into the Tertiary period. It may be +mentioned, however, that the Oolitic deposits have also yielded +the remains of Reptiles belonging to three of the existing orders +of the class-namely, the Lizards (<i>Lacertilia</i>), the Turtles +(<i>Chelonia</i>), and the Crocodiles (<i>Crocodilia</i>). The +Lizards occur both in the marine strata of the Middle Oolites and +also in the fresh-water beds of the Purbeck series; and they are of +such a nature that their affinities with the typical Lacertilians +of the present day cannot be disputed. The Chelonians, up to +this point only known by the doubtful evidence of footprints +in the Permian and Triassic sandstones, are here represented by +unquestionable remains, indicating the existence of marine Turtles +(the <i>Chelone planiceps</i> of the Portland Stone). No remains +of Serpents (<i>Ophidians</i>) have as yet been detected in the +Jurassic; but strata of this age have yielded the remains of +numerous <i>Crocodilians</i>, which probably inhabited the sea. +The most important member of this group is <i>Teleosaurus</i>, +which attained a length of over thirty feet, and is in some +respects allied to the living Gavials of India. +</p> + +<p class="indent"> +The great class of the Birds, as we have seen, is represented +in rocks earlier than the Oolites simply by the not absolutely +certain evidence of the three-toed footprints of the Connecticut +Trias. In the Lithographic Slate of Solenhofen (Middle +<a name="page_252"><span class="page">Page 252</span></a> +Oolite), there has been discovered, +however, the at present unique skeleton of a Bird well known +under the name of the <i>ArchĹ“opteryx macrura</i> (figs. +181, 182). The only known specimen—now in the British + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 520px; + font-size: smaller; text-align: center;"> +<img src="images/fig181.jpg" width="520" height="311" alt="Fig. 181" /> +<br /> +Fig. 181.—<i>ArchĹ“opteryx macrura</i>, showing +tail and tail-feathers, with detached bones. Reduced. From the +Lithographic Slate of Solenhofen. +</span> +</span> + +Museum—unfortunately does not exhibit the skull; but the +fine-grained matrix has preserved a number of the other bones + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 551px; + font-size: smaller; text-align: center;"> +<img src="images/fig182.jpg" width="551" height="299" alt="Fig. 182" /> +<br /> +Fig. 182.—Restoration of <i>ArchĹ“opteryx macrura</i>. +(After Owen.) +</span> +</span> + +of the skeleton, along with the impressions of the tail and wing +feathers. From these remains we know that <i>ArchĹ“opteryx</i> +differed in some remarkable +<a name="page_253"><span class="page">Page 253</span></a> +peculiarities of its structure from all existing members of the +class of Birds. This extraordinary Bird (fig. 182) appears to +have been about as big as a Rook—the tail being long and +extremely slender, and composed of separate vertebræ, each +of which supports a single pair of quill-feathers. In the flying +Birds of the present day, as before mentioned, the terminal +vertebræ of the tail are amalgamated to form a single bone +("ploughshare-bone"), which supports a cluster of tail-feathers; +and the tail itself is short. In the embryos of existing Birds +the tail is long, and is made up of separate vertebræ, and the +same character is observed in many existing Reptiles. The tail +of <i>ArchĹ“opteryx</i>, therefore, is to be regarded as the +permanent retention of an embryonic type of structure, or as an +approximation to the characters of the Reptiles. Another remarkable +point in connection with <i>ArchĹ“opteryx</i>, in which it +differs from all known Birds, is, that the wing was furnished with two +free claws. From the presence of feathers, <i>ArchĹ“opteryx</i> +may be inferred to have been hot-blooded; and this character, +taken along with the structure of the skeleton of the wing, may +be held as sufficient to justify its being considered as belonging +to the class of Birds. In the structure of the tail, however, +it is singularly Reptilian; and there is reason to believe that +its jaws were furnished with teeth sunk in distinct sockets, +as is the case in no existing Bird. This conclusion, at any rate, +is rendered highly probable by the recent discovery of "Toothed +Birds" (<i>Odonturnithes</i>) in the Cretaceous rocks of North +America. +</p> + +<p class="indent"> +The <i>Mammals</i> of the Jurassic period are known to us by +a number of small forms which occur in the "Stonesfield Slate" +(Great Oolite) and in the Purbeck beds (Upper Oolite). The remains +of these are almost exclusively separated halves of the lower +jaw, and they indicate the existence during the Oolitic period in +Europe of a number of small "Pouched animals" (<i>Marsupials</i>). +In the horizon of the Stonesfield Slate four genera of these +little Quadrupeds have been described—viz., <i>Amphilestes, +Amphitherium, Phascolotherium</i>, and <i>Stereognathus</i>. In +<i>Amphitherium</i> (fig. 183), the molar teeth are furnished with +small pointed eminences or "cusps;" and the animal was doubtless +insectivorous. By Professor Owen, the highest living authority +on the subject, <i>Amphitherium</i> is believed to be a small +Marsupial, most nearly allied to the living Banded Ant-eater +(<i>Myrmecobius</i>) of Australia (fig. 158). <i>Amphilestes</i> +and <i>Phascolotherium</i> (fig. 184) are also believed by the +same distinguished anatomist and palæontologist to have been +insect-eating Marsupials, and +<a name="page_254"><span class="page">Page 254</span></a> +the latter is supposed to +find its nearest living ally in the Opossums (<i>Didelphys</i>) +of America. Lastly, the <i>Stereognathus</i> of the Stonesfield + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 293px; + font-size: smaller; text-align: center;"> +<img src="images/fig183.jpg" width="293" height="110" alt="Fig. 183" /> +<br /> +Fig. 183.—Lower jaw of <i>Amphitherium</i> +(<i>Thylacotherium</i>) <i>Prevostii</i>. Stonesfield Slate +(Great Oolite.) +</span> +</span> + +Slate is in a dubious position. It may have been a Marsupial; +but, upon the whole, Professor Owen is inclined to believe that +it must have been a hoofed and herbivorous Quadruped belonging +to the series of the higher Mammals (<i>Placentalia</i>). In the +Middle Purbeck beds, near to the close of the Oolitic period, we +have also evidence of the existence of a number of small Mammals, +all of which are probably Marsupials. Fourteen species are known, +all of small size, the largest being no bigger than a Polecat +or Hedgehog. The genera to which these little quadrupeds have +been referred are <i>Plagiaulax, Spalacotherium, Triconodon</i>, +and <i>Galestes</i>. The first of these (fig. 184, 4) is believed + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 449px; + font-size: smaller; text-align: justify;"> +<img src="images/fig184.jpg" width="449" height="199" alt="Fig. 184" /> +<br /> +Fig. 184. Oolitic Mammals.—1, Lower jaw and teeth of +<i>Phascolotherium</i>, Stonesfield Slate; 2, Lower jaw and +teeth of <i>Amphitherium</i>, Stonesfield Slate; 3, Lower jaw +and teeth of <i>Triconodon</i>, Purbeck beds; 4, Lower jaw and +teeth of <i>Plagiaulax</i>, Purbeck beds. All the figures are +of the natural size. +</span> +</span> + +by Professor Owen to have been carnivorous in its habits; but +other authorities maintain that it was most nearly allied to +the living Kangaroo-rats (<i>Hypsiprymnus</i>) of Australia, +and that it was essentially herbivorous. The remaining three +genera appear to have been certainly insectivorous, and find +their nearest living representatives in the Australian Phalangers +and the American Opossums. +</p> + +<p class="indent"> +Finally, it is interesting to notice in how many respects the +<a name="page_255"><span class="page">Page 255</span></a> +Jurassic fauna of Western Europe approached +to that now inhabiting Australia. At the present day, Australia +is almost wholly tenanted by Marsupials; upon its land-surface +flourish <i>AraucariĹ“</i> and Cycadaceous plants, and in +its seas swims the Port-Jackson Shark (<i>Cestracion +Philippi</i>); whilst the Molluscan genus <i>Trigonia</i> is +nowadays exclusively confined to the Australian coasts. In +England, at the time of the deposition of the Jurassic rocks, +we must have had a fauna and flora very closely resembling what +we now see in Australia. The small Marsupials, <i>Amphitherium, +Phascolotherium</i>, and others, prove that the Mammals were +the same in order; cones of Araucarian pines, with tree-ferns +and fronds of Cycads, occur throughout the Oolitic series; +spine-bearing fishes, like the Port-Jackson Shark, are abundantly +represented by genera such as <i>Acrodus</i> and <i>Strophodus</i>; +and lastly, the genus <i>Trigonia</i>, now exclusively Australian, +is represented in the Oolites by species which differ little +from those now existing. Moreover, the discovery during recent +years of the singular Mud-fish, the <i>Ceratodus Fosteri</i> in +the rivers of Queensland, has added another and a very striking +point of resemblance to those already mentioned; since this genus +of Fishes, though preeminently Triassic, nevertheless extended +its range into the Jurassic. Upon the whole, therefore, there +is reason to conclude that Australia has undergone since the +close of the Jurassic period fewer changes and vicissitudes than +any other known region of the globe; and that this wonderful +continent has therefore succeeded in preserving a greater number +of the characteristic life-features of the Oolites than any other +country with which we are acquainted. +</p> + +<h4>LITERATURE.</h4> + +<p class="indent"> +The following list comprises some of the more important sources +of information as to the rocks and fossils of the Jurassic +series:— +</p> + +<table border="0" cellspacing="0"> +<tr><td class="right" valign="top">(1)</td> + <td>'Geology of Oxford and the Thames Valley.' Phillips.</td></tr> +<tr><td class="right" valign="top">(2)</td> + <td>'Geology of Yorkshire,' vol. ii. Phillips.</td></tr> +<tr><td class="right" valign="top">(3)</td> + <td>'Memoirs of the Geological Survey of Great Britain.'</td></tr> +<tr><td class="right" valign="top">(4)</td> + <td>'Geology of Cheltenham.' Murchison, 2d ed. Buckman.</td></tr> +<tr><td class="right" valign="top">(5)</td> + <td>'Introduction to the Monograph of the Oolitic Asteriadæ' +(Palæontographical Society). Wright.</td></tr> +<tr><td class="right" valign="top">(6)</td> + <td>"Zone of Avicula contorta and the Lower Lias of the South of + England"—'Quart. Journ. Geol. Soc.,' vol. xvi., 1860. + Wright.</td></tr> +<tr><td class="right" valign="top">(7)</td> + <td>"Oolites of Northamptonshire"—'Quart. Journ. Geol. Soc.,' + vols. Xxvi. and xxix. Sharp.</td></tr> +<tr><td class="right" valign="top">(8)</td> + <td>'Manual of Geology.' Dana.</td></tr> +<tr><td class="right" valign="top">(9)</td> + <td>'Der Jura.' Quenstedt.</td></tr> +<tr><td class="right" valign="top">(10)</td> + <td>'Das Flötzgebirge Württembergs.' + Quenstedt.</td></tr> +<tr><td class="right" valign="top">(11)</td> + <td>'Jura Formation.' Oppel.</td></tr> +<tr><td class="right" valign="top"> +<a name="page_256"><span class="page">Page 256</span></a> + (12)</td> + <td>'PalĂ©ontologie du DĂ©partement de la Moselle.' + Terquem.</td></tr> +<tr><td class="right" valign="top">(13)</td> + <td>'Cours Ă©lĂ©mentaire de PalĂ©ontologie.' + D'Orbigny.</td></tr> +<tr><td class="right" valign="top">(14)</td> + <td>'PalĂ©ontologie Française.' D'Orbigny.</td></tr> +<tr><td class="right" valign="top">(15)</td> + <td>'Fossil Echinodermata of the Oolitic Formation' +(Palæontographical Society). Wright.</td></tr> +<tr><td class="right" valign="top">(16)</td> + <td>'Brachiopoda of the Oolitic Formation' + (Palæontographical Society). Davidson.</td></tr> +<tr><td class="right" valign="top">(17)</td> + <td>'Mollusca of the Great Oolite' (Palæontographical + Society). Morris and Lycett.</td></tr> +<tr><td class="right" valign="top">(18)</td> + <td>'Monograph of the Fossil Trigoniæ' + (Palæontographical Society). Lycett.</td></tr> +<tr><td class="right" valign="top">(19)</td> + <td>'Corals of the Oolitic Formation' (Palæontographical + Society). Edwards and Haime.</td></tr> +<tr><td class="right" valign="top">(20)</td> + <td>'Supplement to the Corals of the Oolitic Formation' +(Palæontographical Society). Martin Duncan.</td></tr> +<tr><td class="right" valign="top">(21)</td> + <td>'Monograph of the Belemnitidæ' + (Palæontographical Society). Phillips.</td></tr> +<tr><td class="right" valign="top">(22)</td> + <td>'Structure of the Belemnitidæ' (Mem. Geol. Survey). + Huxley.</td></tr> +<tr><td class="right" valign="top">(23)</td> + <td>'Sur les Belemnites.' Blainville.</td></tr> +<tr><td class="right" valign="top">(24)</td> + <td>'Cephalopoden.' Quenstedt.</td></tr> +<tr><td class="right" valign="top">(25)</td> + <td>'Mineral Conchology.' Sowerby.</td></tr> +<tr><td class="right" valign="top">(26)</td> + <td>'Jurassic Cephalopoda' (Palæontologica Indica). + Waagen.</td></tr> +<tr><td class="right" valign="top">(27)</td> + <td>'Manual of the Mollusca.' Woodward.</td></tr> +<tr><td class="right" valign="top">(28)</td> + <td>'Petrefaktenkunde.' Schlotheim.</td></tr> +<tr><td class="right" valign="top">(29)</td> + <td>'Bridgewater Treatise.' Buckland.</td></tr> +<tr><td class="right" valign="top">(30)</td> + <td>'Versteinerungen des Oolithengebirges.' Roemer.</td></tr> +<tr><td class="right" valign="top">(31)</td> + <td>'Catalogue of British Fossils.' Morris.</td></tr> +<tr><td class="right" valign="top">(32)</td> + <td>'Catalogue of Fossils in the Museum of Practical Geology.' + Etheridge.</td></tr> +<tr><td class="right" valign="top">(33)</td> + <td>'Beiträge zur Petrefaktenkunde.' + Münster.</td></tr> +<tr><td class="right" valign="top">(34)</td> + <td>'Petrefacta Germaniæ.' Goldfuss.</td></tr> +<tr><td class="right" valign="top">(35)</td> + <td>'Lethæa Rossica.' Eichwald.</td></tr> +<tr><td class="right" valign="top">(36)</td> + <td>'Fossil Fishes' (Decades of the Geol. Survey). Sir Philip + Egerton.</td></tr> +<tr><td class="right" valign="top">(37)</td> + <td>'Manual of Palæontology.' Owen.</td></tr> +<tr><td class="right" valign="top">(38)</td> + <td>'British Fossil Mammals and Birds.' Owen.</td></tr> +<tr><td class="right" valign="top">(39)</td> + <td>'Monographs of the Fossil Reptiles of the Oolitic + Formation' (Palæontographical Society). Owen.</td></tr> +<tr><td class="right" valign="top">(40)</td> + <td>'Fossil Mammals of the Mesozoic Formations' + (Palæontographical Society). Owen.</td></tr> +<tr><td class="right" valign="top">(41)</td> + <td>'Catalogue of Ornithosauria.' Seeley.</td></tr> +<tr><td class="right" valign="top">(42)</td> + <td>"Classification of the Deinosauria"—'Quart. Journ. + Geol. Soc.,' vol. xxvi., 1870. Huxley.</td></tr> +</table> + +<h3>CHAPTER XVII.</h3> + +<p class="subtitle"> +THE CRETACEOUS PERIOD. +</p> + +<p class="indent"> +The next series of rocks in ascending order is the great and +important series of the Cretaceous Rocks, so called from the +general occurrence in the system of chalk (Lat. <i>creta</i>, +<a name="page_257"><span class="page">Page 257</span></a> +chalk). As developed in Britain and Europe generally, the +following leading subdivisions may be recognised in the +Cretaceous series:— +</p> + +<div class="center"> +<table border="1" cellspacing="0" cellpadding="3"> + <tr><td class="left"> + 1. Wealden,<br/> + 2. Lower Greensand or Neocomian, + </td><td class="center">Lower Cretaceous.</td></tr> + + <tr><td class="left"> + 3. Gault,<br/> + 4. Upper Greensand,<br/> + 5. Chalk,<br/> + 6. Maestricht beds,<br/> + </td><td class="center">Upper Cretaceous.</td></tr> +</table> +</div> + +<p class="indent"> +I. <i>Wealden</i>.—The <i>Wealden</i> formation, though of +considerable importance, is a local group, and is confined to +the southeast of England, France, and some other parts of Europe. +Its name is derived from the <i>Weald</i>, a district comprising +parts of Surrey, Sussex, and Kent, where it is largely developed. +Its lower portion, for a thickness of from 500 to 1000 feet, +is arenaceous, and is known as the Hastings Sands. Its Upper +portion, for a thickness of 150 to nearly 300 feet, is chiefly +argillaceous, consisting of clays with sandy layers, and occasionally +courses of limestone. The geological importance of the Wealden +formation is very great, as it is undoubtedly the delta of an +ancient river, being composed almost wholly of fresh-water beds, +with a few brackish-water and even marine strata, intercalated +in the lower portion. Its geographical extent, though uncertain, +owing to the enormous denudation to which it has been subjected, +is nevertheless great, since it extends from Dorsetshire to France, +and occurs also in North Germany. Still, even if it were continuous +between all these points, it would not be larger than the delta +of such a modern river as the Ganges. The river which produced +the Wealden series must have flowed from an ancient continent +occupying what is now the Atlantic Ocean; and the time occupied +in the formation of the Wealden must have been very great, though +we have, of course, no data by which we can accurately calculate +its duration. +</p> + +<p class="indent"> +The fossils of the Wealden series are, naturally, mostly the +remains of such animals as we know at the present day as inhabiting +rivers. We have, namely, fresh-water Mussels (<i>Unio</i>), +River-snails (<i>Paludina</i>), and other fresh-water shells, +with numerous little bivalved Crustaceans, and some fishes. +</p> + +<p class="indent"> +II. <i>Lower Greensand</i> (<i>NĂ©ocomien</i> of +D'Orbigny).—The Wealden beds pass upward, often by insensible +gradations, into the Lower Greensand. The name Lower Greensand is +not an appropriate one, for green sands only occur sparingly and +occasionally, and are found in other formations. For this +<a name="page_258"><span class="page">Page 258</span></a> +reason it has been proposed to substitute for Lower Greensand the +name <i>Neocomian</i>, derived from the town of +Neufchâtel—anciently called <i>Neocomum</i>—in +Switzerland. If this name were adopted, as it ought to be, the +Wealden beds would be called the Lower Neocomian. +</p> + +<p class="indent"> +The Lower Greensand or Neocomian of Britain has a thickness of +about 850 feet, and consists of alternations of sands, sandstones, +and clays, with occasional calcareous bands. The general colour +of the series is dark brown, sometimes red; and the sands are +occasionally green, from the presence of silicate of iron. +</p> + +<p class="indent"> +The fossils of the Lower Greensand are purely marine, and among +the most characteristic are the shells of <i>Cephalopods</i>. +</p> + +<p class="indent"> +The most remarkable point, however, about the fossils of the +Lower Cretaceous series, is their marked divergence from the +fossils of the Upper Cretaceous rocks. Of 280 species of fossils +in the Lower Cretaceous series, only 51, or about 18 per cent, pass +on into the Upper Cretaceous. This break in the life of the two +periods is accompanied by a decided physical break as well; for the +Gault is often, if not always, unconformably superimposed on the +Lower Greensand. At the same time, the Lower and Upper Cretaceous +groups form a closely-connected and inseparable series, as shown +by a comparison of their fossils with those of the underlying +Jurassic rocks and the overlying Tertiary beds. Thus, in Britain +no marine fossil is known to be common to the marine beds of +the Upper Oolites and the Lower Greensand; and of more than 500 +species of fossils in the Upper Cretaceous rocks, almost everyone +died out before the formation of the lowest Tertiary strata, the +only survivors being one Brachiopod and a few <i>Foraminifera</i>. +</p> + +<p class="indent"> +III. <i>Gault</i> (<i>Aptien</i> of D'Orbigny).—The lowest +member of the Upper Cretaceous series is a stiff, dark-grey, blue, +or brown clay, often worked for brick-making, and known as the +<i>Gault</i>, from a provincial English term. It occurs chiefly +in the south-east of England, but can be traced through France +to the flanks of the Alps and Bavaria. It never exceeds 100 feet +in thickness; but it contains many fossils, usually in a state +of beautiful preservation. +</p> + +<p class="indent"> +IV. <i>Upper Greensand</i> (<i>Albien</i> of D'Orbigny; +<i>Unterquader</i> and <i>Lower Plänerkalk</i> of +Germany).—The Gault is succeeded upward by the <i>Upper +Greensand</i>, which varies in thickness from 3 up to 100 feet, +and which derives its name from the occasional occurrence in it +of green sands. These, however, are local and sometimes wanting, +and the name "Upper +<a name="page_259"><span class="page">Page 259</span></a> +Greensand" is to be +regarded as a <i>name</i> and not a description. The group +consists, in Britain, of sands and clays, sometimes with bands +of calcareous grit or siliceous limestone, and occasionally +containing concretions of phosphate of lime, which are largely +worked for agricultural purposes. +</p> + +<p class="indent"> +V. <i>White Chalk</i>.—The top of the Upper Greensand becomes +argillaceous, and passes up gradually into the base of the great +formation known as the true <i>Chalk</i>, divided into the three +subdivisions of the chalk-marl, white chalk without flints, and +white chalk with flints. The first of these is simply argillaceous +chalk, and passes up into a great mass of obscurely-stratified white +chalk in which there are no flints (<i>Turonien</i> of D'Orbigny; +<i>Mittelquader</i> of Germany). This, in turn, passes up into a +great mass of white chalk, in which the stratification is marked +by nodules of black flint arranged in layers (<i>SĂ©nonien</i> +of D'Orbigny; <i>Oberquader</i> of Germany). The thickness of +these three subdivisions taken together is sometimes over 1000 +feet, and their geographical extent is very great. White Chalk, +with its characteristic appearance, may be traced from the north +of Ireland to the Crimea, a distance of about 1140 geographical +miles; and, in an opposite direction, from the south of Sweden +to Bordeaux, a distance of about 840 geographical miles. +</p> + +<p class="indent"> +VI. In Britain there occur no beds containing Chalk fossils, or +in any way referable to the Cretaceous period, above the true +White Chalk with flints. On the banks of the Maes, however, near +Maestricht in Holland, there occurs a series of yellowish limestones, +of about 100 feet in thickness, and undoubtedly superior to the +White Chalk. These <i>Maestricht beds</i> (<i>Danien</i> of +D'Orbigny) contain a remarkable series of fossils, the characters +of which are partly Cretaceous and partly Tertiary. Thus, with +the characteristic Chalk fossils, <i>Belemnites, Baculites</i>, +Sea-Urchins, &c., are numerous Univalve Molluscs, such as +Cowries and Volutes, which are otherwise exclusively Tertiary or +Recent. +</p> + +<p class="indent"> +Holding a similar position to the Maestricht beds, and showing +a similar intermixture of Cretaceous forms with later types, are +certain beds which occur in the island of Seeland, in Denmark, +and which are known as the <i>Faxöe Limestone</i>. +</p> + +<p class="indent"> +Of a somewhat later date than the Maestricht beds is the <i>Pisolitic +Limestone</i> of France, which rests unconformably on the White +Chalk, and contains a large number of Tertiary fossils along +with some characteristic Cretaceous types. +</p> + +<p class="indent"> +The subjoined sketch-section exhibits the general succession of +the Cretaceous deposits in Britain:— +</p> + +<p class="indent"> +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 575px; + font-size: smaller; text-align: center;"> +<a name="page_260"><span class="page">Page 260</span></a> +GENERALIZED SECTION OF THE CRETACEOUS SERIES OF BRITAIN. +<br /> +Fig. 185. +<br /> +<img src="images/fig185.jpg" width="538" height="651" alt="Fig. 185" /> +</span> +</span> + +In North America, strata of Lower Cretaceous age are well represented +in Missouri, Wyoming, Utah, and in some other areas; but the greater +portion of the American deposits of this period are referable +to the Upper Cretaceous. The rocks of this series are mostly +sands, clays, and limestones—<i>Chalk</i> itself being unknown +except in Western Arkansas. Amongst the sandy accumulations, one +of the most important is the +<a name="page_261"><span class="page">Page 261</span></a> +so-called "marl" +of New Jersey, which is truly a "Greensand," and contains a large +proportion of glauconite (silicate of iron and potash). It also +contains a little phosphate of lime, and is largely worked for +agricultural purposes. The greatest thickness attained by the +Cretaceous rocks of North America is about 9000 feet, as in Wyoming, +Utah, and Colorado. According to Dana, the Cretaceous rocks of the +Rocky Mountain territories pass upwards "without interruption into +a coal-bearing formation, several thousand feet thick, on which the +following Tertiary strata lie unconformably." The lower portion of +this "Lignitic formation" appears to be Cretaceous, and contains +one or more beds of Coal; but the upper part of it perhaps belongs +to the Lower Tertiary. In America, therefore, the lowest Tertiary +strata appear to rest conformably upon the highest Cretaceous; +whereas in Europe, the succession at this point is invariably an +unconformable one. Owing, however, to the fact that the American +"Lignitic formation" is a shallow-water formation, it can hardly +be expected to yield much material whereby to bridge over the +great palæontological gap between the White Chalk and Eocene +in the Old World. +</p> + +<p class="indent"> +Owing to the fact that so large a portion of the Cretaceous formation +has been deposited in the sea, much of it in deep water, the +<i>plants</i> of the period have for the most part been found +special members of the series, such as the Wealden beds, the +Aix-la-Chapelle sands, and the Lignitic beds of North America. Even +the purely marine strata, however, have yielded plant-remains, and +some of these are peculiar and proper to the deep-sea deposits of +the series. Thus the little calcareous discs termed "coccoliths," +which are known to be of the nature of calcareous sea-weeds +(<i>AlgĹ“</i>) have been detected in the White Chalk; and the +flints of the same formation commonly contain the spore-cases of +the microscopic <i>Desmids</i> (the so-called Xanthidia), along +with the siliceous cases of the equally diminutive <i>Diatoms</i>. +</p> + +<p class="indent"> +The plant-remains of the Lower Cretaceous greatly resemble those +of the Jurassic period, consisting mainly of Ferns, Cycads, and +Conifers. The Upper Cretaceous rocks, however, both in Europe and +in North America, have yielded an abundant flora which resembles +the existing vegetation of the globe in consisting mainly of +Angiospermous Exogens and of Monocotyledons.[23] In Europe the +plant-remains in question have +<a name="page_262"><span class="page">Page 262</span></a> +been found chiefly in certain sands in the neighbourhood of +Aix-la-Chapelle, and they consist of numerous Ferns, Conifers +(such as <i>Cycadopteris</i>), Screw Pines (<i>Pandanus</i>), Oaks +(<i>Quercus</i>), Walnut (<i>Juglans</i>), Fig (<i>Ficus</i>), and +many <i>ProteaceĹ“</i>, some of which are referred to existing +genera (<i>Dryandra, Banksia, Grevillea</i>, &c.) +</p> + +<p class="footnote"> +[Footnote 23: The "Flowering plants" are divided into the two +great groups of the Endogens and Exogens. The <i>Endogens</i> +(such as Grasses, Palms, Lilies, &c.) have no true bark, nor +rings of growth, and the stem is said to be "endogenous;" the young +plant also possesses but a single seed-leaf or "cotyledon." Hence +these plants are often simply called "<i>Monocotyledons</i>." The +<i>Exogens</i>, on the other hand, have a true bark; and the stem +increases by annual additions to the outside, so that rings of growth +are produced. The young plant has two seed-leaves or "cotyledons," +and these plants are therefore called "<i>Dicotyledons</i>." +Amongst the Exogens, the Pines (<i>Conifers</i>) and the Cycads +have seeds which are unprotected by a seed-vessel, and they are +therefore called "<i>Gymnosperms</i>." All the other Exogens, +including the ordinary trees, shrubs, and flowering plants, have +the seeds enclosed in a seed-vessel, and are therefore called +"<i>Angiosperms</i>." The derivation of these terms will be found +in the Glossary at the end of the volume.] +</p> + +<p class="indent"> +In North America, the Cretaceous strata of New Jersey, Alabama, +Nebraska, Kansas, &c., have yielded the remains of numerous +plants, many of which belong to existing genera. Amongst these +may be mentioned Tulip-trees (<i>Liriodendron</i>), Sassafras (fig. +186), Oaks (<i>Quercus</i>), Beeches (<i>Fagus</i>), Plane-trees +(<i>Platanus</i>), Alders (<i>Alnus</i>), Dog-wood (<i>Cornus</i>), +Willows (<i>Salix</i>), Poplars (<i>Populus</i>), Cypresses +(<i>Cupressus</i>), Bald Cypresses (<i>Taxodium</i>), Magnolias, +&c. Besides these, however, there occur other forms which have +now entirely disappeared from North America—as, for example, +species of <i>Cinnamomum</i> and <i>Araucaria</i>. +</p> + +<p class="indent"> +It follows from the above, that the Lower and Upper Cretaceous +rocks are, from a botanical point of view, sharply separated +from one another. The Palæozoic period, as we have seen, is +characterised by the prevalance of "Flowerless" plants +(<i>Cryptogams</i>), its higher vegetation consisting almost +exclusively of Conifers. The Mesozoic period, as a whole, is +characterised by the prevalence of the Cryptogamic group of the +Ferns, and the Gymnospermic groups of the Conifers and the Cycads. +Up to the close of the Lower Cretaceous, no Angiospermous Exogens +are certainly known to have existed, and Monocotyledonous plants +or Endogens are very poorly represented. With the Upper Cretaceous, +however, a new era of plant-life, of which our present is but +the culmination, commenced, with a great and apparently sudden +development of new forms. In place of the Ferns, Cycads, and +Conifers of the earlier Mesozoic deposits, we have now an +astonishingly large number of true Angiospermous Exogens, many +of them belonging to existing types; and along with these are +various Monocotyledonous plants, including the first examples of +the great and important +<a name="page_263"><span class="page">Page 263</span></a> +group of the Palms. It is thus a matter +of interest to reflect that plants closely related to those now + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 510px; + font-size: smaller; text-align: center;"> +<img src="images/fig186.jpg" width="510" height="495" alt="Fig. 186" /> +<br /> +Fig. 186.—Cretaceous Angiosperms. <i>a. Sassafras Cretaceum; +b, Liriodendron Meekii; c, Leguminosites Marcouanus; d, Salix +Meekii</i>. (After Dana.) +</span> +</span> + +inhabiting the earth, were in existence at a time when the ocean +was tenanted by Ammonites and Belemnites, and when land and sea +and air were peopled by the extraordinary extinct Reptiles of +the Mesozoic period. +</p> + +<p class="indent"> +As regards animal life, the <i>Protozoans</i> of the Cretaceous +period are exceedingly numerous, and are represented by +<i>Foraminifera</i> and <i>Sponges</i>. As we have already seen, +the White Chalk itself is a deep-sea deposit, almost entirely +composed of the microscopic shells of <i>Foraminifers</i>, along +with Sponge-spicules, and organic <i>dĂ©bris</i> of different +kinds (see fig. 7). The green grains which are so abundant in several +minor subdivisions of the Cretaceous, are also in many instances +really casts in glauconite of the chambered shells of these minute +organisms. A great many species of <i>Foraminifera</i> have been +recognised in the Chalk; but the three principal genera are +<a name="page_264"><span class="page">Page 264</span></a> +<i>Globigerina, Rotalia</i> (fig. 187), and <i>Textularia</i>—groups +which are likewise characteristic of the "ooze" of the Atlantic and + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 472px; + font-size: smaller; text-align: center;"> +<img src="images/fig187.jpg" width="472" height="161" alt="Fig. 187" /> +<br /> +Fig. 187—<i>Kotalia Boueana</i>. +</span> +</span> + +Pacific Oceans at great depths. The flints of the Chalk also commonly +contain the shells of <i>Foraminifera</i>. The Upper Greensand +has yielded in considerable numbers the huge <i>Foraminifera</i> +described by Dr Carpenter under the name of <i>Parkeria</i>, the +spherical shells of which are composed of sand-grains agglutinated +together, and sometimes attain a diameter of two and a quarter +inches. The Cretaceous Sponges are extremely numerous, and occur +under a great number of varieties of shape and structure; but +the two most characteristic genera are <i>Siphonia</i> and +<i>Ventriculites</i>, both of which are exclusively confined to +strata of this age. The <i>SiphoniĹ“</i> (fig. 188) consist of a +pear-shaped, sometimes lobed head, supported by a longer or shorter +stern, which breaks up at its base into a number of root-like +processes of attachment. The water gained access to the interior +of the Sponge by a number of minute openings covering the surface, +and ultimately escaped by a single, large, chimney-shaped aperture +at the summit. In some respects these sponges present a singular +resemblance to the beautiful "Vitreous Sponges" (<i>Holtenia</i> +or <i>Pheronema</i>) of the deep Atlantic; and, like these, they +were probably denizens of a deep sea, The <i>Ventriculites</i> +of the Chalk (fig. 189) is, however, a genus still more closely +allied to the wonderful flinty Sponges, which have been shown, +by the researches of the Porcupine, Lightning, and Challenger +expeditions, to live half buried in the Calcareous ooze of the +abysses of our great oceans. Many forms of this genus are known, +having "usually the form of graceful vases, tubes, or funnels, +variously ridged or grooved, or otherwise ornamented on the surface, +frequently expanded above into a cup-like lip, and continued +below into a bundle of fibrous roots. The minute structure of +these bodies shows an extremely delicate tracery of fine tubes, +sometimes empty, sometimes filled with loose calcareous matter +<a name="page_265"><span class="page">Page 265</span></a> +dyed with peroxide of iron."—(Sir Wyville Thomson.) Many of +the Chalk sponges, originally calcareous, have been converted into +flint subsequently; but the Ventriculites are really composed + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 125px; vertical-align:top; text-align: center; + margin: 4px;"> +<img src="images/fig188.jpg" width="125" height="424" alt="Fig. 188" /> +<br /> +Fig. 188.—<i>Siphonia ficus</i>. Upper Greensand. Europe. +</span> + +<span style="width: 222px; vertical-align:top; text-align: center; + margin: 4px;"> +<img src="images/fig189.jpg" width="222" height="424" alt="Fig. 189" /> +<br /> +Fig. 189.—<i>Ventriculites simplex</i>. White Chalk. Britain. +</span> + +</span> + +of this substance, and are therefore genuine "Siliceous Sponges," +like the existing Venus's Flower-Basket (<i>Euplectella</i>). +Like the latter, the skeleton was doubtless originally composed, +in the young state, of disconnected six-rayed spicules, which +ultimately become fixed together to constitute a continuous +frame-work. The sea-water, as in the recent forms, must have +been admitted to the interior of the Sponge by numerous apertures +on its exterior, subsequently escaping by a single large opening +at its summit. +</p> + +<p class="indent"> +Amongst the <i>CĹ“lenterates</i>, the "Hydroid Zoophytes" are +represented by a species of the encrusting genus <i>Hydractinia</i>, +the horny polypary of which is so commonly found at the present +day adhering to the exterior of shells. The occurrence of this +genus is of interest, because it is the first known instance in +the entire geological series of the occurrence of an unquestionable +Hydroid of a modern type, though many of the existing forms of +these animals possess structures which are +<a name="page_266"><span class="page">Page 266</span></a> +perfectly fitted for +preservation in the fossil condition. The corals of the Cretaceous +series are not very numerous, and for the most part are referable +to types such as <i>Trochocyathus, Stephanophyllia, Parasmilia, +Synhelia</i> (fig. 190), &c., which belong to the same great +group of corals as the majority of existing forms. We have also + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 480px; + font-size: smaller; text-align: center;"> +<img src="images/fig190.jpg" width="480" height="319" alt="Fig. 190" /> +<br /> +Fig. 190.—<i>Synhelia Sharpeana</i>. Chalk, England. +</span> +</span> + +a few "Tabulate Corals" (<i>Polytremacis</i>), hardly, if at all, +generically separable from very ancient forms (<i>Heliolites</i>); +and the Lower Greensand has yielded the remains of the little +<i>Holocystis elegans</i>, long believed to be the last of the +great Palæozoic group of the <i>Rugosa</i>. +</p> + +<p class="indent"> +As regards the <i>Echinoderms</i>, the group of the <i>Crinoids</i> +now exhibits a marked decrease in the number and variety of its +types. The "stalked" forms are represented by <i>Pentacrinus</i> +and <i>Bourgueticrinus</i>, and the free forms by Feather-stars +like our existing <i>ComatulĹ“</i>; whilst a link between the +stalked and free groups is constituted by the curious "Tortoise +Encrinite (<i>Marsupites</i>). By far the most abundant Cretaceous +Echinoderms, however, are Sea-urchins (<i>Echinoids</i>); though +several Star-fishes are known as well. The remains of Sea-urchins +are so abundant in various parts of the Cretaceous series, especially +in the White Chalk, and are often so beautifully preserved, that +they constitute one of the most marked features of the fauna +of the period. From the many genera of Sea-urchins which occur +in strata of this age, it is difficult to select characteristic +types; but the genera <i>Galerites</i> (fig. 191), <i>Discoidea</i> +(fig. 192), <i>Micraster, Ananchytes, Diadema, Salenia</i>, and +<a name="page_267"><span class="page">Page 267</span></a> +<i>Cidaris</i>, may be mentioned as being all important Cretaceous +groups. +</p> + +<p class="indent"> +Coming to the <i>Annulose Animals</i> of the Cretaceous period, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 533px; + font-size: smaller; text-align: center;"> +<img src="images/fig191.jpg" width="533" height="181" alt="Fig. 191" /> +<br /> +Fig. 191.—<i>Galerites albogalerus</i>, viewed from below, +from the side, and from above. White Chalk. +</span> +</span> + +there is little special to remark. The <i>Crustaceans</i> belong +for the most part to the highly-organised groups of the Lobsters + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 531px; + font-size: smaller; text-align: center;"> +<img src="images/fig192.jpg" width="531" height="168" alt="Fig. 192" /> +<br /> +Fig. 192.—<i>Discoidea cylindrica</i>; under, side, and +upper aspect. Upper Greensand. +</span> +</span> + +and the Crabs (the Macrurous and Brachyurous Decapods); but there +are also numerous little <i>Ostracodes</i>, especially in the +fresh-water strata of the Wealden. It should further be noted +that there occurs here a great development of the singular +<i>Crustaceous</i> family of the Barnacles (<i>LepadidĹ“</i>), +whilst the allied family of the equally singular Acorn-shells +(<i>BalanidĹ“</i>) is feebly represented as well. +</p> + +<p class="indent"> +Passing on to the <i>Mollusca</i>, the class of the Sea-mats +and Sea-mosses (<i>Polyzoa</i>) is immensely developed in the +Cretaceous period, nearly two hundred species being known to +occur in the Chalk. Most of the Cretaceous forms belong to the +family of the <i>EscharidĹ“</i>, the genera <i>Eschara</i> and +<i>Escharina</i> (fig. 193) being particularly well represented. +Most of the Cretaceous <i>Polyzoans</i> are of small size, but +some attain considerable dimensions, and many simulate Corals +in their general form and appearance. +</p> + +<p class="indent"> +<a name="page_268"><span class="page">Page 268</span></a> +The Lamp-shells (<i>Brachiopods</i>) have now reached a further +stage of the progressive decline, which they have been undergoing + +<span style="float: left; margin: 4px; width: 237px; + font-size: smaller; text-align: justify;"> +<img src="images/fig193.jpg" width="237" height="194" alt="Fig. 193" /> +<br /> +Fig. 193.—A small fragment of <i>Escharina Oceani</i>, +of the natural size; and a portion of the same enlarged. Upper +Greensand. +</span> + +ever since the close of the Palæozoic period. Though +individually not rare, especially in certain minor subdivisions of +the series, the number of generic types has now become distinctly +diminished, the principal forms belonging to the genera +<i>Terebratula, Terebratella</i> (fig. 194), <i>Terebratulina, +Rhynchonella</i>, and <i>Crania</i> (fig. 195). In the last +mentioned of these, the shell is attached to foreign bodies by +the substance of one of the valves (the ventral), whilst the other +or free valve is more or less limpet-shaped. All the above-mentioned + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 575px; + font-size: smaller; text-align: center;"> +<img src="images/fig194.jpg" width="531" height="209" alt="Fig. 194" /> +<br /> +Fig. 194.—<i>Terebratella Astieriana</i>. Gault. +</span> +</span> + +genera are in existence at the present day; and one +<i>species</i>—namely, <i>Terebratulina +striata</i>—appears to be undistinguishable from one now +living—the <i>Terebratulina caputserpentis</i>. +</p> + +<p class="indent"> +Whilst the Lamp-shells are slowly declining, the Bivalves +(<i>Limellibranchs</i>) are greatly developed, and are amongst +the most abundant and characteristic fossils of the Cretaceous +period. In the great river-deposit of the Wealden, the Bivalves +are forms proper to fresh water, belonging to the existing +River-mussels (<i>Unio</i>), <i>Cyrena</i> and <i>Cyclas</i>; +but most of the Cretaceous Lamellibranchs are marine. Some of +the most abundant and characteristic of these belong to the great +family of the Oysters (<i>OstreidĹ“</i>). Amongst these are +the genera <i>GryphtĹ“a</i> and <i>Exogyra</i>, both of which +we have seen to occur +<a name="page_269"><span class="page">Page 269</span></a> +abundantly in the Jurassic; and there are +also numerous true Oysters (<i>Ostrea</i>, fig. 196) and Thorny +Oysters (<i>Spondylus</i>, fig. 197). The genus <i>Trigonia</i>, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 579px; + font-size: smaller; text-align: justify;"> +<img src="images/fig195.jpg" width="579" height="212" alt="Fig. 195" /> +<br /> +Fig. 195.—<i>Crania Ignabergensis</i>. The left-hand +figure shows the perfect shell, attached by its ventral valve +to a foreign body; the middle figure shows the exterior of the +limpet-shaped dorsal valve; and the right-hand figure represents +the interior of the attached valve. White Chalk. +</span> +</span> + +so characteristic of the Mesozoic deposits in general, is likewise +well represented in the Cretaceous strata. No single genus of + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 575px; + font-size: smaller; text-align: center;"> +<img src="images/fig196.jpg" width="525" height="268" alt="Fig. 196" /> +<br /> +Fig. 196.—<i>Ostrea Couloni</i>. Lower Greensand. +</span> +</span> + +Bivalves is, however, so highly characteristic of the Cretaceous +period as <i>Inoceramus</i>, a group belonging to the family of the +Pearl-mussels (<i>AviculidĹ“</i>). The shells of this genus +(fig. 198) have the valves unequal in size, the larger valve often +being much twisted, and both valves being marked with radiating +ribs or concentric furrows. The hinge-line is long and straight, +with numerous pits for the attachment of the ligament which serves +to open the shell. Some of the <i>Inocerami</i> attain a length +of two or three feet, and fragments of the shell are often found +perforated by boring +<a name="page_270"><span class="page">Page 270</span></a> +Sponges. Another extraordinary family of +Bivalves, which is exclusively confined to the Cretaceous rocks, is +that of the <i>HippuritidĹ“</i>. All the members of this group + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 497px; + font-size: smaller; text-align: center;"> +<img src="images/fig197.jpg" width="497" height="217" alt="Fig. 197" /> +<br /> +Fig. 197.—<i>Spondylus spinosus</i>. White Chalk. +</span> +</span> + +(fig. 199) were attached to foreign objects, and lived associated +in beds, like Oysters. The two valves of the shell are always + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 497px; + font-size: smaller; text-align: center;"> +<img src="images/fig198.jpg" width="459" height="163" alt="Fig. 198" /> +<br /> +Fig. 198.—<i>Inoceramus sulcatus</i>. Gault. +</span> +</span> + +altogether unlike in sculpturing, appearance, shape, and size; +and the cast of the interior of the shell is often extremely +unlike the form of the outer surface. The type-genus of the family +is <i>Hippurites</i> itself (fig. 199), in which the shell is in +the shape of a straight or slightly-twisted horn, sometimes a +foot or more in length, constituted by the attached lower valve, +and closed above by a small lid-like free upper valve. About +a hundred species of the family of the <i>HippuritidĹ“</i> +are known, all of these being Cretaceous, and occurring in Britain +(one species only), in Southern Europe, the West Indies, North +America, Algeria, and Egypt. Species of this family occur in +such numbers in certain compact marbles in the south of Europe, +of the age of the Upper Cretaceous (Lower Chalk), as to have +given origin to the name of "Hippurite Limestones," applied to +these strata. +</p> + +<p class="indent"> +<a name="page_271"><span class="page">Page 271</span></a> +The Univalves (<i>Gasteropods</i>) of the Cretaceous period are +not very numerous, nor particularly remarkable. Along with species +of the persistent genus <i>Pleurotomaria</i> and the Mesozoic + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 230px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig199.jpg" width="230" height="442" alt="Fig. 199" /> +<br /> +Fig. 199.—<i>Hippurites Toucasiana</i>. A large individual, +with two smaller ones attached to it. Upper Cretaceous, South +of Europe. +</span> + +<span style="width: 168px; vertical-align:bottom; text-align: center; + margin: 4px;"> +<img src="images/fig200.jpg" width="168" height="534" alt="Fig. 200" /> +<br /> +Fig. 200.—<i>Voluta elongata</i>. White Chalk. +</span> + +</span> + +<i>NerinĹ“a</i>, we meet with examples of such modern types +as <i>Turritella</i> and <i>Natica</i>, the Staircase-shells +(<i>Solarium</i>), the Wentle-traps (<i>Scalaria</i>), the +Carrier-shells (<i>Phorus</i>), &c. Towards the close of the +Cretaceous period, and especially in such transitional strata +as the Maestricht beds, the Faxöe Limestone, and the Pisolitic +Limestone of France, we meet with a number of carnivorous +("siphonostomatous") Univalves, in which the mouth of the shell is +notched or produced into a canal. Amongst these it is interesting +to recognise examples of such existing genera as the Volutes +(<i>Voluta</i>, fig. 200), the Cowries (<i>CyprĹ“a</i>), the +Mitre-shells (<i>Mitra</i>), the Wing - shells (<i>Strombus</i>), +the Scorpion-shells (<i>Pteroceras</i>), &c. +</p> + +<p class="indent"> +<a name="page_272"><span class="page">Page 272</span></a> +Upon the whole, the most characteristic of all the Cretaceous +Molluscs are the <i>Cephalopods</i>, represented by the remains +of both <i>Tetrabranchiate</i> and <i>Dibranchiate</i> forms. +Amongst the former, the long-lived genus <i>Nautilus</i> (fig. +201) again reappears, with its involute shell, its capacious + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 495px; + font-size: smaller; text-align: center;"> +<img src="images/fig201.jpg" width="495" height="245" alt="Fig. 201" /> +<br /> +Fig. 201.—Different views of <i>Nautilus Danicus</i>. +Faxöe Limestone (Upper Cretaceous), Denmark. +</span> +</span> + +body-chamber, its simple septa between the air-chambers, and its +nearly or quite central siphuncle. The majority of the chambered +<i>Cephalopods</i> of the Cretaceous belong, however, to the +complex and beautiful family of the <i>AmmonitidĹ“</i>, with +their elaborately folded and lobed septa and dorsally-placed +siphuncle. This family disappears wholly at the close of the +Cretaceous period; but its approaching extinction, so far from +being signalised by any slow decrease and diminution in the number +of specific or generic types, seems to have been attended by the +development of whole series of new forms. The genus <i>Ammonites</i> +itself, dating from the Carboniferous, has certainly passed its +prime, but it is still represented by many species, and some of +these attained enormous dimensions (two or three feet in diameter). +The genus <i>Ancyloceras</i> (fig. 202), though likewise of more +ancient origin (Jurassic), is nevertheless very characteristic +of the Cretaceous. In this genus the first portion of the shell +is in the form of a flat spiral, the coils of which are not in +contact; and its last portion is produced at a tangent, becoming +ultimately bent back in the form of a crosier. Besides these +pre-existent types, the Cretaceous rocks have yielded a great +number of entirely new forms of the <i>AmmonitidĹ“</i>, which +are not known in any deposits of earlier or later date. Amongst the +more important of these may be mentioned <i>Crioceras, Turrilites, +Scaphites, Hamites,</i> +<a name="page_273"><span class="page">Page 273</span></a> +<i>Ptychoceras</i>, and <i>Baulites</i>. In the +genus <i>Crioceras</i> (fig. 204, <i>d</i>), the shell consists +of an open spiral, the volutions of which are not in contact, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 528px; + font-size: smaller; text-align: center;"> +<img src="images/fig202.jpg" width="528" height="176" alt="Fig. 202" /> +<br /> +Fig. 202.—<i>Ancyloceras Matheronianus</i>. Gault. +</span> +</span> + +thus resembling a partially-unrolled <i>Ammonite</i> or the inner +portion of an <i>Ancyloceras</i>. In <i>Turrilites</i> (fig. +203), the shell is precisely like that of the <i>Ammonite</i> +in its structure; but instead of forming a flat spiral, it is +coiled into an elevated turreted shell, the whorls of which are +in contact with one another. In the genus <i>Scaphites</i> (fig. +204, <i>e</i>), the shell resembles that of <i>Ancyloceras</i> in +consisting of a series of volutions coiled into a flat spiral, +the last being detached from the others, produced, and ultimately +bent back in the form of a crosier; but the whorls of the enrolled +part of the shell are in contact, instead of being separate as +in the latter. In the genus <i>Hamites</i> (fig. 204, <i>f</i>), +the shell is an extremely elongated cone, which is bent upon +itself more than once, in a hook-like manner, all the volutions +being separate. The genus <i>Ptychoteras</i> (fig. 204, <i>a</i>) +is very like <i>Hamites</i>, except that the shell is only bent +once; and the two portions thus bent are in contact with one +another. Lastly, in the genus <i>Baculites</i> (fig. 204, <i>b</i> +and <i>c</i>) the shell is simply a straight elongated cone, not +bent in any way, but possessing the folded septa which characterise +the whole Ammonite family. The <i>Baculite</i> is the simplest of all +the forms of the <i>AmmonitidĹ“</i>; and all the other forms, +however complex, may be regarded as being simply produced by the +bending or folding of such a conical septate shell in different +ways. The <i>Baculite</i>, therefore, corresponds, in the series of +the <i>AmmonitidĹ“</i>, to the <i>Orthoceras</i> in the series +of the <i>NautilidĹ“</i>. All the above-mentioned genera are +characteristically, or exclusively, Cretaceous, and they are +accompanied by a number of other allied forms, which cannot be +noticed here. Not a single one of these genera, further, has hitherto +been detected in any strata higher than the Cretaceous. We may +therefore consider that these wonderful, varied, and elaborate +<a name="page_274"><span class="page">Page 274</span></a> +forms of <i>AmmonitidĹ“</i> constitute one of the most +conspicuous features in the life of the Chalk period. +</p> + +<p class="indent"> +The <i>Dibranchiate Cephalopods</i> are represented partly by + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 181px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig203.jpg" width="181" height="674" alt="Fig. 203" /> +<br /> +Fig. 203.—<i>Turrilites catenatus</i>. The lower figure +represents the entire shell; the upper figure represents the base +of the shell seen from below. Gault. +</span> + +<span style="width: 319px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig204.jpg" width="319" height="674" alt="Fig. 204" /> +<br /> +Fig. 204.—<i>a, Ptychoceras Emericianum</i>, +reduced—Lower Greensand; <i>b, Baculites anceps</i>, +reduced—Chalk; <i>c</i>, Portion of the same, showing the +folded edges of the septa; <i>d, Crioceras cristatum</i>, +reduced—Gault; <i>e, Scaphites Ĺ“qualis</i>, natural +size—Chalk; <i>f, Hamites rotundus</i>, restored—Gault. +</span> + +</span> + +the beak-like jaws of unknown species of Cuttle-fishes and partly +by the internal skeletons of Belemnites. Amongst the latter, the +genus <i>Belemnites</i> itself holds its place in the lower part +<a name="page_275"><span class="page">Page 275</span></a> +of the Cretaceous series; but it disappears in the upper portion +of the series, and its place is taken by the nearly-allied genus +<i>Belemnitella</i> (fig. 205), distinguished by the possession +of a straight fissure in the upper end of the guard. This also + +<span style="float: right; margin: 4px; width: 56px; + font-size: smaller; text-align: justify;"> +<img src="images/fig205.jpg" width="56" height="289" alt="Fig. 205" /> +<br /> +Fig. 205.—Guard of <i>Belemnitella mucronata</i>. +</span> + +disappears at the close of the Cretaceous period; and no member of +the great Mesozoic family of the <i>BelemnitidĹ“</i> has +hitherto been discovered in any Tertiary deposit, or is known to +exist at the present day. +</p> + +<p class="indent"> +Passing on next to the <i>Vertebrate Animals</i> of the Cretaceous +period, we find the <i>Fishes</i> represented as before by the +Ganoids and the Placoids, to which, however, we can now add the +first known examples of the great group of the <i>Bony Fishes</i> +or <i>Teleosteans</i>, comprising the great majority of existing +forms. The <i>Ganoid</i> fishes of the Cretaceous (<i>Lepidotus, +Pycnodus</i>, &c.) present no features of special interest. +Little, also, need be said about the <i>Placoid</i> fishes of this +period. As in the Jurassic deposits, the remains of these consist +partly of the teeth of genuine Sharks (<i>Lamna, Odontaspis</i>, +&c.) and partly of the teeth and defensive spines of Cestracionts, +such as the living Port-Jackson Shark. The pointed and sharp-edged +teeth of true Sharks are very abundant in some beds, such as +the Upper Greensand, and are beautifully preserved. The teeth +of some forms (<i>Carcharias</i>, &c.) attain occasionally a +length of three or four inches, and indicate the existence in the +Cretaceous seas of huge predaceous fishes, probably larger than +any existing Sharks. The remains of <i>Cestracionts</i> consist +partly of the flattened teeth of genera such as <i>Acrodus</i> +and <i>Ptychodus</i> (the latter confined to rocks of this age), +and partly of the pointed teeth of <i>Hybodus</i>, a genus which +dates from the Trias. In this genus the teeth (fig. 206) consist +of a principal central cone, flanked by minor lateral cones; and + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 94px; vertical-align:bottom; text-align: center; + margin: 4px;"> +<img src="images/fig206.jpg" width="94" height="125" alt="Fig. 206" /> +<br /> +Fig. 206.—Tooth of <i>Hybodus</i>. +</span> + +<span style="width: 423px; vertical-align:bottom; text-align: center; + margin: 4px;"> +<img src="images/fig207.jpg" width="423" height="56" alt="Fig. 207" /> +<br /> +Fig. 207.—Fin-spine of <i>Hybodus</i>. Lower Greensand. +</span> + +</span> + +the fin-spines (fig. 207) are longitudinally grooved, and carry +a series of small spines on their hinder or concave margin. Lastly, +<a name="page_276"><span class="page">Page 276</span></a> +the great modern +order of the Bony Fishes or <i>Teleosteans</i> makes its first +appearance in the Upper Cretaceous rocks, where it is represented by +forms belonging to no less than three existing groups—namely, +the Salmon family (<i>SalmonidĹ“</i>), the Herring family +(<i>ClupeidĹ“</i>), and the Perch family (<i>PercidĹ“</i>). +All these fishes have thin, horny, overlapping scales, symmetrical + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 490px; + font-size: smaller; text-align: center;"> +<img src="images/fig208.jpg" width="490" height="434" alt="Fig. 208" /> +<br /> +Fig. 208.—1, <i>Beryx Lewesiensis</i>, a Percoid fish from +the Chalk; 2, <i>Osmeroides Mantelli</i>, a Salmonoid fish from +the Chalk. +</span> +</span> + +("homocercal") tails, and bony skeletons. The genus <i>Beryx</i> +(fig. 208, 1) is one represented by existing species at the present +day, and belongs to the Perch family. The genus <i>Osmeroides</i>, +again (fig. 208, 2), is supposed to be related to the living Smelts +(<i>Osmerus</i>), and, therefore, to belong to the Salmon tribe. +</p> + +<p class="indent"> +No remains of <i>Amphibians</i> have hitherto been detected in any +part of the Cretaceous series; but <i>Reptiles</i> are extremely +numerous, and belong to very varied types. As regards the great +extinct groups of Reptiles which characterise the Mesozoic period +as a whole, the huge "Enaliosaurs" or "Sea-Lizards" are still +represented by the <i>Ichthyosaur</i> and the <i>Plesiosaur</i>. +Nearly allied to the latter of these is the <i>Elasmosaurus</i> +of the American Cretaceous, which combined +<a name="page_277"><span class="page">Page 277</span></a> +the long tail of the Ichthyosaur with the long neck of the +Plesiosaur. The length of this monstrous Reptile could not have +been less than fifty feet, the neck consisting of over sixty +vertebræ and measuring over twenty feet in length. The +extraordinary Flying Reptiles of the Jurassic are likewise well +represented in the Cretaceous rocks by species of the genus +<i>Pterodactylus</i> itself, and these later forms are much more +gigantic in their dimensions than their predecessors. Thus some of +the Cretaceous Pterosaurs seem to have had a spread of wing of from +twenty to twenty-five feet, more than realising the "Dragons" of +fable in point of size. The most remarkable, however, of the +Cretaceous <i>Pterosaurs</i> are the forms which have recently +been described by Professor Marsh under the generic title of +<i>Pteranodon</i>. In these singular forms—so far only known +as American—the animal possessed a skeleton in all respects +similar to that of the typical Pterodactyles, except that the jaws +are completely destitute of teeth. There is, therefore, the +strongest probability that the jaws were encased in a horny sheath, +thus coming to resemble the beak of a Bird. Some of the recognised +species of <i>Pteranodon</i> are very small; but the skull of one +species (<i>P. Longiceps</i>) is not less than a yard in length, +and there are portions of the skull of another species which would +indicate a length of four feet for the cranium. These measurements +would point to dimensions larger than those of any other known +Pterosaurs. +</p> + +<p class="indent"> +The great Mesozoic order of the <i>Deinosaurs</i> is largely +represented in the Cretaceous rocks, partly by genera which +previously existed in the Jurassic period, and partly by entirely new +types. The great delta-deposit of the Wealden, in the Old World, has +yielded the remains of various of these huge terrestrial Reptiles, +and very many others have been found in the Cretaceous deposits +of North America. One of the most celebrated of the Cretaceous +Deinosaurs is the <i>Iguanodon</i>, so called from the curious +resemblance of its teeth to those of the existing but comparatively +diminutive <i>Iguana</i>. The teeth (fig. 209) are soldered to the +inner face of the jaw, instead of being sunk in distinct sockets; +and they have the form of somewhat flattened prisms, longitudinally +ridged on the outer surface, with an obtusely triangular crown, +and having the enamel crenated on one or both sides. They present +the extraordinary feature that the crowns became worn down flat +by mastication, showing that the <i>Iguanodon</i> employed its +teeth in actually chewing and triturating the vegetable matter +on which it fed. There can therefore be no doubt but that the +<i>Iguanodon</i>, in spite of its immense bulk, was an herbivorous +Reptile, and +<a name="page_278"><span class="page">Page 278</span></a> +lived principally on the foliage of the Cretaceous forests +amongst which it dwelt. Its size has been variously estimated + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 426px; + font-size: smaller; text-align: center;"> +<img src="images/fig209.jpg" width="426" height="347" alt="Fig. 209" /> +<br /> +Fig. 209.—Teeth of Iguanodon Mantellii. Wealden, Britain. +</span> +</span> + +at from thirty to fifty feet, the thigh-bone in large examples +measuring nearly five feet in length, with a circumference of +twenty-two inches in its smallest part. With the strong and massive +hind-limbs are associated comparatively weak and small fore-limbs; +and there seems little reason to doubt that the <i>Iguanodon</i> +must have walked temporarily or permanently upon its hind-limbs, +after the manner of a Bird. This conjecture is further supported +by the occurrence in the strata which contain the bones of the +<i>Iguanodon</i> of gigantic three-toed foot-prints, disposed +<i>singly</i> in a double track. These prints have undoubtedly +been produced by some animal walking on two legs; and they can +hardly, with any probability, be ascribed to any other than this +enormous Reptile. Closely allied to the <i>Iguanodon</i> is the +<i>Hadrosaurus</i> of the American Cretaceous, the length of +which is estimated at twenty-eight feet. <i>Iguanodon</i> does +not appear to have possessed any integumentary skeleton; but the +great <i>HylĹ“osaurus</i> of the Wealden seems to have been +furnished with a longitudinal crest of large spines running down +the back, similar to that which is found in the comparatively +small Iguanas of the present day. The <i>Megalosaurus</i> of +the Oolites continued to exist in the Cretaceous period; and, +as we have previously seen, it was carnivorous in its habits. +The American <i>LĹ“laps</i> was also carnivorous, and, like +the Megalosaur, +<a name="page_279"><span class="page">Page 279</span></a> +which it very closely resembles, appears to have walked upon its +hind-legs, the fore-limbs being disproportionately small. +</p> + +<p class="indent"> +Another remarkable group of Reptiles, exclusively confined to +the Cretaceous series, is that of the <i>Mosasauroids</i>, so +called from the type-genus <i>Mosasaurus</i>. The first species +of <i>Mosasaurus</i> known to science was the <i>M. Camperi</i> +(fig. 210), the skull of which—six feet in length—was + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 508px; + font-size: smaller; text-align: center;"> +<img src="images/fig210.jpg" width="508" height="314" alt="Fig. 210" /> +<br /> +Fig. 210.—Skull of <i>Mosasaurus Camperi</i>, greatly +reduced. Maestricht Chalk. +</span> +</span> + +discovered in 1780 in the Maestricht Chalk at Maestricht. As this +town stands on the river Meuse, the name of <i>Mosasaurus</i> +("Lizard of the Meuse") was applied to this immense Reptile. Of +late years the remains of a large number of Reptiles more or less +closely related to <i>Mosasaurus</i>, or absolutely belonging to +it, have been discovered in the Cretaceous deposits of North +America, and have been described by Professors Cope and Marsh. +All the known forms of this group appear to have been of large +size—one of them, <i>Mosasaurus princeps</i>, attaining +the length of seventy-five or eighty feet, and thus rivalling +the largest of existing Whales in its dimensions. The teeth in +the "Mosasauroids" are long, pointed, and slightly curved; and +instead of being sunk in distinct sockets, they are firmly +amalgamated with the jaws, as in modern Lizards. The palate +also carried teeth, and the lower jaw was so constructed as to +allow of the mouth being opened to an immense width, somewhat +as in the living Serpents. The body was long and snake-like, +with a very long tail, which is laterally compressed, and must +have served as a powerful swimming-apparatus. In addition to +this, both pairs of limbs have the bones connecting them with +<a name="page_280"><span class="page">Page 280</span></a> +the trunk greatly shortened; whilst the digits were enclosed in +the integuments, and constituted paddles, closely resembling in +structure the "flippers" of Whales and Dolphins. The neck is +sometimes moderately long, but oftener very short, as the great +size and weight of the head would have led one to anticipate. +Bony plates seem in some species to have formed an at any rate +partial covering to the skin; but it is not certain that these +integumentary appendages were present in all. Upon the whole, +there can be no doubt but that the Mosasauroid Reptiles—the +true "Sea-serpents" of the Cretaceous period—were +essentially aquatic in their habits, frequenting the sea, and +only occasionally coming to the land. +</p> + +<p class="indent"> +The "Mosasauroids" have generally been regarded as a greatly +modified group of the Lizards (<i>Lacertilia</i>). Whether this +reference be correct or not—and recent investigations render +it dubious—the Cretaceous rocks have yielded the remains of +small Lizards not widely removed from existing forms. The recent +order of the <i>Chelonians</i> is also represented in the + +<span style="float: left; margin: 4px; width: 311px; + font-size: smaller; text-align: center;"> +<img src="images/fig211.jpg" width="311" height="429" alt="Fig. 211" /> +<br /> +Fig. 211.—Carapace of <i>Chelone Benstedi</i>. Lower Chalk. +(After Owen.) +</span> + +Cretaceous rocks, by forms closely resembling living types. Thus +the fresh-water deposits of the Wealden have yielded examples of +the "Terrapins" or "Mud-Turtles" (<i>Emys</i>); and the marine +Cretaceous strata have been found to contain the remains of various +species of Turtles, one of which is here figured (fig. 211). No +true Serpents (<i>Ophidia</i>) have as yet been detected in the +Cretaceous rocks; and this order does not appear to have come +into existence till the Tertiary period. Lastly, true Crocodiles +are known to have existed in considerable numbers in the Cretaceous +period. The oldest of these occur in the fresh-water deposit of the +Wealden; and they differ from +<a name="page_281"><span class="page">Page 281</span></a> +the existing forms of +the group in the fact that the bodies of the vertebræ, like +those of the Jurassic Crocodiles, are bi-concave, or hollowed out +at both ends. In the Greensand of North America, however, occur +the remains of Crocodiles which agree with all the living species +in having the bodies of the vertebræ in the region of the +back hollowed out in front and convex behind. +</p> + +<p class="indent"> +<i>Birds</i> have not hitherto been shown, with certainty, to +have existed in Europe during the Cretaceous period, except in +a few instances in which fragmentary remains belonging to this +class have been discovered. The Cretaceous deposits of North +America have, however, been shown by Professor Marsh to contain +a considerable number of the remains of Birds, often in a state +of excellent preservation. Some of these belong to Swimming or +Wading Birds, differing in no point of special interest from +modern birds of similar habits. Others, however, exhibit such +extraordinary peculiarities that they merit more than a passing +notice. One of the forms in question constitutes the genus +<i>Ichthyornis</i> of Marsh, the type-species of which (<i>I. +Dispar</i>) was about as large as a Pigeon. In two remarkable +respects, this singular Bird differs from all known living members +of the class. One of these respects concerns the jaws, both of +which exhibit the Reptilian character of being armed with numerous +small pointed <i>teeth</i> (fig. 212, <i>a</i>), sunk in distinct +sockets. No existing bird possesses teeth; and this character +forcibly recalls the Bird-like Pterosaurs, with their toothed +jaws. <i>Ichthyornis</i>, however, possessed fore-limbs constructed +strictly on the type of the "wing" of the living Birds; and it +cannot, therefore, be separated from this class. Another +extraordinary peculiarity of <i>Ichthyornis</i> is, that the +bodies of the <i>vertebrie</i> (fig. 212, <i>c</i>) were +<i>bi-concave</i>, as is the case with many extinct Reptiles and +almost all Fishes, but as does not occur in any living Bird. There +can be little doubt that <i>Ichthyornis</i> was aquatic in its +habits, and that it lived principally upon fishes; but its powerful +wings at the same time indicate that it was capable of prolonged +flight. The tail of <i>Ichthyornis</i> has, unfortunately, not +been discovered; and it is at present impossible to say whether +this resembled the tail of existing Birds, or whether it was +elongated and composed of separate vertebræ, as in the +Jurassic <i>ArchĹ“opteryx</i>. +</p> + +<p class="indent"> +Still more wonderful than <i>Ichthyornis</i> is the marvellous +bird described by Marsh under the name of <i>Hesperornis +regalis</i>. This presents us with a gigantic diving bird, +somewhat resembling the existing "Loons" (<i>Colymbus</i>), but +agreeing with <i>Ichthyornis</i> in having the jaws furnished +<a name="page_282"><span class="page">Page 282</span></a> +with conical, recurved, pointed teeth (fig. 212, <i>b</i>). +Hence these forms are grouped together in a new sub-class, under +the name of <i>Odontornithes</i> or "Toothed Birds." The teeth +of <i>Hesperornis</i> (fig. 212, <i>d</i>) resemble those of +<i>Ichthyornis</i> in their general form; but instead of being + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 525px; + font-size: smaller; text-align: justify;"> +<img src="images/fig212.jpg" width="525" height="373" alt="Fig. 212" /> +<br /> +Fig. 212.—Toothed Birds (<i>Odontornithes</i>) of the +Cretaceous Rocks of America. <i>a</i>. Left lower jaw of +<i>Ichthyornis dispar</i>, slightly enlarged; <i>b</i>, Left +lower jaw of <i>Hesperornis regalis</i>, reduced to nearly +one-fourth of the natural size; <i>c</i>. Cervical vertebra +of <i>Ichthyornis dispar</i>, front view, twice the natural +size; <i>c'</i>, Side view of the same; <i>d</i>, Tooth of +<i>Hesperornis regalis</i>, enlarged to twice the natural +size. (After Marsh.) +</span> +</span> + +sunk in distinct sockets, they are simply implanted in a deep +continuous groove in the bony substance of the jaw. The front of +the upper jaw does not carry teeth, and was probably encased in +a horny beak. The breast-bone is entirely destitute of a central +ridge or keel, and the wings are minute and quite rudimentary; +so that <i>Hesperornis</i>, unlike <i>Ichthyornis</i>, must have +been wholly deprived of the power of flight, in this respect +approaching the existing Penguins. The tail consists of about +twelve vertebræ, of which the last three or four are amalgamated +to form a flat terminal mass, there being at the same time clear +indications that the tail was capable of up and down movement +in a vertical plane, this probably fitting it to serve as a +swimming-paddle or rudder. The legs were powerfully constructed, +and the feet were adapted to assist the bird in rapid motion through +the water. The known remains of <i>Hesperornis regalis</i> prove +it to have been a swimming and diving bird, of larger dimensions +<a name="page_283"><span class="page">Page 283</span></a> +than any of the aquatic members of the class of Birds with which +we are acquainted at the present day. It appears to have stood +between five and six feet high, and its inability to fly is fully +compensated for by the numerous adaptations of its structure to a +watery life. Its teeth prove it to have been carnivorous in its +habits, and it probably lived upon fishes. It is a curious fact +that two Birds agreeing with one another in the wholly abnormal +character of possessing teeth, and in other respects so entirely +different, should, like <i>Ichthyornis</i> and <i>Hesperornis</i>, +have lived not only in the same geological period, but also in the +same geographical area; and it is equally curious that the area +inhabited by these toothed Birds should at the same time have been +tenanted by winged and bird-like Reptiles belonging to the toothed +genus <i>Pterodactylus</i> and the toothless genus <i>Pteranodon</i>. +</p> + +<p class="indent"> +No remains of <i>Mammals</i>, finally, have as yet been detected +in any sedimentary accumulations of Cretaceous age. +</p> + +<h4>LITERATURE.</h4> + +<p class="indent"> +The following list comprises some of the more important works and +memoirs which may be consulted with reference to the Cretaceous +strata and their fossil contents:— +</p> + +<table border="0" cellspacing="0"> +<tr><td class="right" valign="top">(1)</td> + <td>'Memoirs of the Geological Survey of Great Britain.'</td></tr> +<tr><td class="right" valign="top">(2)</td> + <td>'Geology of England and Wales.' Conybeare and + Phillips.</td></tr> +<tr><td class="right" valign="top">(3)</td> + <td>'Geology of Yorkshire,' vol. ii. Phillips.</td></tr> +<tr><td class="right" valign="top">(4)</td> + <td>'Geology of Oxford and the Thames Valley.' Phillips.</td></tr> +<tr><td class="right" valign="top">(5)</td> + <td>'Geological Excursions through the Isle of Wight.' + Mantell.</td></tr> +<tr><td class="right" valign="top">(6)</td> + <td>'Geology of Sussex.' Mantell.</td></tr> +<tr><td class="right" valign="top">(7)</td> + <td>'Report on Londonderry,' &c. Portlock.</td></tr> +<tr><td class="right" valign="top">(8)</td> + <td>'Recherches sur le Terrain CrĂ©tacĂ© + SupĂ©rieur de l'Angleterre et de l'Irlande.' + Barrois.</td></tr> +<tr><td class="right" valign="top">(9)</td> + <td>"Geological Survey of Canada"—'Report of Progress, + 1872-73.'</td></tr> +<tr><td class="right" valign="top">(10)</td> + <td>'Geological Survey of California.' Whitney.</td></tr> +<tr><td class="right" valign="top">(11)</td> + <td>'Geological Survey of Montana, Idaho, Wyoming, and Utah.' + Hayden and Meek.</td></tr> +<tr><td class="right" valign="top">(12)</td> + <td>'Report on Geology,' &c. (British North American + Boundary Commission). G. M. Dawson.</td></tr> +<tr><td class="right" valign="top">(13)</td> + <td>'Manual of Geology.' Dana.</td></tr> +<tr><td class="right" valign="top">(14)</td> + <td>'Lethæa Rossica.' Eichwald.</td></tr> +<tr><td class="right" valign="top">(15)</td> + <td>'Petrefacta Germaniæ.' Goldfuss.</td></tr> +<tr><td class="right" valign="top">(16)</td> + <td>'Fossils of the South Downs.' Mantell.</td></tr> +<tr><td class="right" valign="top">(17)</td> + <td>'Medals of Creation.' Mantell.</td></tr> +<tr><td class="right" valign="top">(18)</td> + <td>'Mineral Conchology.' Sowerby.</td></tr> +<tr><td class="right" valign="top">(19)</td> + <td>'Lethæa Geognostica.' Bronn.</td></tr> +<tr><td class="right" valign="top">(20)</td> + <td>'Malacostracous Crustacea of the British Cretaceous + Formation' (Palæontographical Society). Bell.</td></tr> +<tr><td class="right" valign="top">(21)</td> + <td>'Brachiopoda of the Cretaceous Formation' + (Palæontographical Society). Davidson.</td></tr> +<tr><td class="right" valign="top">(22)</td> + <td>'Corals of the Cretaceous Formation' + (Palæontographical Society). Milne-Edwards and + Haime.</td></tr> +<tr><td class="right" valign="top"> +<a name="page_284"><span class="page">Page 284</span></a> + (23)</td> + <td>'Supplement to the Fossil Corals' (Palæontographical + Society). Martin Duncan.</td></tr> +<tr><td class="right" valign="top">(24)</td> + <td>'Echinodermata or the Cretaceous Formation' + (Palæontographical Society). Wright.</td></tr> +<tr><td class="right" valign="top">(25)</td> + <td>'Monograph of the Belemnitidæ' (Palæontographical + Society). Phillips.</td></tr> +<tr><td class="right" valign="top">(26)</td> + <td>'Monograph of the Trigoniæ' (Palæontographical + Society). Lycett.</td></tr> +<tr><td class="right" valign="top">(27)</td> + <td>'Fossil Cirripedes' (Palæontographical Society). + Darwin.</td></tr> +<tr><td class="right" valign="top">(28)</td> + <td>'Fossil Mollusca of the Chalk of Britain' + (Palæontographical Society). Sharpe.</td></tr> +<tr><td class="right" valign="top">(29)</td> + <td>'Entomostraca of the Cretaceous Formation' + (Palæontographical Society). Rupert Jones.</td></tr> +<tr><td class="right" valign="top">(30)</td> + <td>'Monograph of the Fossil Reptiles of the Cretaceous + Formation' (Palæontographical Society). Owen.</td></tr> +<tr><td class="right" valign="top">(31)</td> + <td>'Manual of Palæontology.' Owen.</td></tr> +<tr><td class="right" valign="top">(32)</td> + <td>'Synopsis of Extinct Batrachia and Reptilia.' Cope.</td></tr> +<tr><td class="right" valign="top">(33)</td> + <td>"Structure of the Skull and Limbs in Mosasauroid + Reptiles"—'American Journ. Sci. and Arts, 1872.' + Marsh.</td></tr> +<tr><td class="right" valign="top">(34)</td> + <td>"On Odontornithes"—'American Journ. Sci. and Arts, + 1875.' Marsh.</td></tr> +<tr><td class="right" valign="top">(35)</td> + <td>'Ossemens Fossiles.' Cuvier.</td></tr> +<tr><td class="right" valign="top">(36)</td> + <td>'Catalogue of Ornithosauria.' Seeley.</td></tr> +<tr><td class="right" valign="top">(37)</td> + <td>'PalĂ©ontologie Française.' D'Orbigny.</td></tr> +<tr><td class="right" valign="top">(38)</td> + <td>'Synopsis des Echinides fossiles.' Desor.</td></tr> +<tr><td class="right" valign="top">(39)</td> + <td>'Cat. RaisonnĂ© des Echinides.' Agassiz and + Desor.</td></tr> +<tr><td class="right" valign="top">(40)</td> + <td>"Echinoids"—'Decades of the Geol. Survey of Britain.' + E. Forbes.</td></tr> +<tr><td class="right" valign="top">(41)</td> + <td>'PalĂ©ontologie Française.' Cotteau.</td></tr> +<tr><td class="right" valign="top">(42)</td> + <td>'Versteinerungen der Böhmischen Kreide-formation.' + Reuss.</td></tr> +<tr><td class="right" valign="top">(43)</td> + <td>"Cephalopoda, Gasteropoda, Pelecypoda, Brachiopoda; &c., + of the Cretaceous Rocks of India"—'Palæontologica + Indica,' ser. i., iii., v., vi., viii. Stoliczka.</td></tr> +<tr><td class="right" valign="top">(44)</td> + <td>"Cretaceous Reptiles of the United States"—'Smithsonian + Contributions to Knowledge,' vol. xiv. Leidy.</td></tr> +<tr><td class="right" valign="top">(45)</td> + <td>'Invertebrate Cretaceous, and Tertiary Fossils of the Upper + Missouri Country,' 1876. Meek.</td></tr> +</table> + +<h3>CHAPTER XVIII.</h3> + +<p class="subtitle"> +THE EOCENE PERIOD. +</p> + +<p class="indent"> +Before commencing the study of the subdivisions of the Kainozoic +series, there are some general considerations to be noted. In +the first place, there is in the Old World a complete and entire +physical break between the rocks of the Mesozoic and Kainozoic +periods. In no instance in Europe are Tertiary strata to be found +resting conformably upon any Secondary rock. The Chalk has invariably +suffered much erosion and denudation before the lowest Tertiary +strata were deposited upon it. This is shown by the fact that the +<a name="page_285"><span class="page">Page 285</span></a> +actually eroded surface of the Chalk can often +be seen; or, failing this, that we can point to the presence of the +chalk-flints in the Tertiary strata. This last, of course, affords +unquestionable proof that the Chalk must have been subjected to +enormous denudation prior to the formation of the Tertiary beds, +all the chalk itself having been removed, and nothing left but the +flints, while these are all rolled and rounded. In the continent +of North America, on the other hand, the lowest Tertiary strata +have been shown to graduate downwards conformably with the highest +Cretaceous beds, it being a matter of difficulty to draw a precise +line of demarcation between the two formations. +</p> + +<p class="indent"> +In the second place, there is a marked break in the <i>life</i> +of the Mesozoic and Kainozoic periods. With the exception of a +few <i>Foraminifera</i>, and one <i>Brachiopod</i> (the latter +doubtful), no Cretaceous species is known to have survived the +Cretaceous period; while several characteristic <i>families</i>, +such as the <i>AmmonitidĹ“, BelemnitidĹ“</i>, and +<i>HippuritidĹ“</i>, died out entirely with the close of the +Cretaceous rocks. In the Tertiary rocks, on the other hand, not +only are all the animals and plants more or less like existing +types, but we meet with a constantly-increasing number of <i>living +species</i> as we pass from the bottom of the Kainozoic series to +the top. Upon this last fact is founded the modern classification +of the Kainozoic rocks, propounded by Sil Charles Lyell. +</p> + +<p class="indent"> +The absence in strata of Tertiary age of the chambered Cephalopods, +the Belemnites, the <i>Hippurites</i>, the <i>Inocerami</i>, and +the diversified types of Reptiles which form such conspicuous +features in the Cretaceous fauna, render the palæontological +break between the Chalk and the Eocene one far too serious to +be overlooked. At the same time, it is to be remembered that +the evidence afforded by the explorations carried out of late +years as to the animal life of the deep sea, renders it certain +that the extinction of marine forms of life at the close of the +Cretaceous period was far less extensive than had been previously +assumed. It is tolerably certain, in fact, that we may look upon +some of the inhabitants of the depths of our existing oceans +as the direct, if modified, descendants of animals which were +in existence when the Chalk was deposited. +</p> + +<p class="indent"> +It follows from the general want of conformity between the Cretaceous +and Tertiary rocks, and still more from the great difference in +life, that the Cretaceous and Tertiary periods are separated, in +the Old World at any rate, by an enormous lapse of unrepresented +time. How long this interval may have been, we have no means of +judging exactly, but it very possibly was as long as the whole +Kainozoic epoch itself. Some day we shall +<a name="page_286"><span class="page">Page 286</span></a> +doubtless find, at some part of the earth's surface, marine strata +which were deposited during this period, and which will contain +fossils intermediate in character between the organic remains +which respectively characterise the Secondary and Tertiary periods. +At present, we have only slight traces of such deposits—as, +for instance, the Maestricht beds, the Faxöe Limestone, and +the Pisolitic Limestone of France. +</p> + +<p class="indent"> +CLASSIFICATION OF THE TERTIARY ROCKS.—The classification +of the Tertiary rocks is a matter of unusual difficulty, in +consequence of their occurring in disconnected basins, forming +a series of detached areas, which hold no relations of +superposition to one another. The order, therefore, of the +Tertiaries in point of time, can only be determined by an appeal +to fossils; and in such determination Sir Charles Lyell proposed +to take as the basis of classification the <i>proportion of +living or existing species of Mollusca which occurs in each +stratum or group of strata</i>. Acting upon this principle, +Sir Charles Lyell divides the Tertiary series into four +groups:— +</p> + +<p class="indent"> +I. The <i>Eocene</i> formation (Gr. <i>eos</i>, dawn; <i>kainos</i>, +new), containing the smallest proportion of existing species, and +being, therefore, the oldest division. In this classification, +only the <i>Mollusca</i> are taken into account; and it was found +that of these about three and a half per cent were identical +with existing species. +</p> + +<p class="indent"> +II. The <i>Miocene</i> formation (Gr. <i>meion</i>, less; +<i>kainos</i>, new), with more recent species than the Eocene, +but <i>less</i> than the succeeding formation, and less than +one-half the total number in the formation. As before, only the +<i>Mollusca</i> are taken into account, and about 17 per cent +of these agree with existing species. +</p> + +<p class="indent"> +III. The <i>Pliocene</i> formation (Gr. <i>pleion</i>, more; +<i>kainos</i>, new), with generally <i>more</i> than half the +species of shells identical with existing species—the +proportion of these varying from 35 to 50 per cent in the lower +beds of this division, up to 90 or 95 per cent in its higher +portion. +</p> + +<p class="indent"> +IV. The <i>Post-Tertiary Formations</i>, in which all <i>the +shells belong to existing species</i>. This, in turn, is divided +into two minor groups—the <i>Post-Pliocene</i> and <i>Recent +Formations</i>. In the <i>Post-Pliocene</i> formations, while +all the <i>Mollusca</i> belong to existing species, most of the +<i>Mammals</i> belong to extinct species. In the Recent period, +the quadrupeds, as well as the shells, belong to living species. +</p> + +<p class="indent"> +The above, with some modifications, was the original classification +proposed by Sir Charles Lyell for the Tertiary rocks, and now +universally accepted. More recent researches, it is true, have +somewhat altered the proportions of existing species +<a name="page_287"><span class="page">Page 287</span></a> +to extinct, as stated above. The general principle, however, of +an increase in the number of living species, still holds good; and +this is as yet the only satisfactory basis upon which it has been +proposed to arrange the Tertiary deposits. +</p> + +<p class="center">EOCENE FORMATION.</p> + +<p class="indent"> +The Eocene rocks are the lowest of the Tertiary series, and comprise +all those Tertiary deposits in which there is only a small proportion +of existing <i>Mollusca</i>—from three and a half to five per +cent. The Eocene rocks occur in several basins in Britain, France, +the Netherlands, and other parts of Europe, and in the United +States. The subdivisions which have been established are extremely +numerous, and it is often impossible to parallel those of one +basin with those of another. It will be sufficient, therefore, +to accept the division of the Eocene formation into three great +groups—Lower, Middle, and Upper Eocene—and to consider +some of the more important beds comprised under these heads in +Europe and in North America. +</p> + +<p class="indent"> +I. EOCENE OF BRITAIN. (1.) LOWER EOCENE.—The base of the Eocene +series in Britain is constituted by about 90 feet of light-coloured, +sometimes argillaceous sands (<i>Thanet Sands</i>), which are of +marine origin. Above these, or forming the base of the formation +where these are wanting, come mottled clays and sands with lignite +(<i>Woolwich and Reading series</i>), which are estuarine or +fluvio-marine in origin. The highest member of the Lower Eocene +of Britain is the "London Clay," consisting of a great mass of +dark-brown or blue clay, sometimes with sandy beds, or with layers +of "septaria," the whole attaining a thickness of from 200 to as +much as 500 feet. The London Clay is a purely marine deposit, +containing many marine fossils, with the remains of terrestrial +animals and plants; all of which indicate a high temperature of +the sea and tropical or sub-tropical conditions of the land. +</p> + +<p class="indent"> +(2.) MIDDLE EOCENE.—The inferior portion of the Middle +Eocene of Britain consists of marine beds, chiefly consisting +of sand, clays, and gravels, and attaining a very considerable +thickness (<i>Bag-shot and Bracklesham beds</i>). The superior +portion of the Middle Eocene of Britain, on the other hand, +consists of deposits which are almost exclusively fresh-water +or brackish-water in origin (<i>Headon and Osborne series</i>). +</p> + +<p class="indent"> +The chief Continental formations of Middle Eocene age are the +"Calcaire grossier" of the Paris basin, and the "Nummulitic +Limestone" of the Alps. +</p> + +<p class="indent"> +(3.) UPPER EOCENE.—If the Headon and Osborne beds of +<a name="page_288"><span class="page">Page 288</span></a> +the Isle of Wight be placed in the Middle Eocene, the only +British representatives of the Upper Eocene are the <i>Bembridge +beds</i>. These strata consist of limestones, clays, and marls, +which have for the most part been deposited in fresh or brackish +water. +</p> + +<p class="indent"> +II. EOCENE BEDS OF THE PARIS BASIN.—The Eocene strata are +very well developed in the neighbourhood of Paris, where they +occupy a large area or basin scooped out of the Chalk. The beds +of this area are partly marine, partly freshwater in origin; and +the following table (after Sir Charles Lyell) shows their +subdivisions and their parallelism with the English series:— +</p> + +<p class="center">GENERAL TABLE OF FRENCH EOCENE STRATA.</p> + +<table border="0" width="100%"> +<tr><td colspan="5" class="center">UPPER EOCENE.</td></tr> +<tr><td> </td> + <td colspan="2" class="center"><i>French Subdivisions.</i></td> + <td colspan="2" class="center"><i>English Equivalents.</i></td></tr> + +<tr><td valign="top">A.</td> + <td valign="top">1.</td> + <td valign="top">Gypseous series of Mont + Montmartre.</td> + <td valign="top">1.</td> + <td valign="top">Bembridge series.</td></tr> + +<tr><td valign="top">A.</td> + <td valign="top">2.</td> + <td valign="top">Calcaire silicieux, or Travertin + InfĂ©rieur.</td> + <td valign="top">2.</td> + <td valign="top">Osborne and Headon series.</td></tr> + +<tr><td valign="top">A.</td> + <td valign="top">3.</td> + <td valign="top">Grès de Beauchamp, or Sables Moyens.</td> + <td valign="top">3.</td> + <td valign="top">White sand and clay of Barton Cliff, + Hants.</td></tr> + +<tr><td colspan="5" class="center">MIDDLE EOCENE.</td></tr> + +<tr><td valign="top">B.</td> + <td valign="top">1.</td> + <td valign="top">Calcaire Grossier.</td> + <td valign="top">1.</td> + <td valign="top">Bagshot and Bracklesham beds.</td></tr> + +<tr><td valign="top">B.</td> + <td valign="top">2.</td> + <td valign="top">Soissonnais Sands, or Lits Coquilliers.</td> + <td valign="top">2.</td> + <td valign="top">Wanting.</td></tr> + +<tr><td colspan="5" class="center">LOWER EOCENE.</td></tr> + +<tr><td valign="top">C.</td> + <td valign="top">1.</td> + <td valign="top">Argile de Londres at base of Hill of Cassel, + near Dunkirk.</td> + <td valign="top">1.</td> + <td valign="top">London clay.</td></tr> + +<tr><td valign="top">C.</td> + <td valign="top">2.</td> + <td valign="top">Argile plastique and lignite.</td> + <td valign="top">2.</td> + <td valign="top">Plastic clay and sand with lignite (Woolwich + and Reading series).</td></tr> + +<tr><td valign="top">C.</td> + <td valign="top">3.</td> + <td valign="top">Stables de Bracheux.</td> + <td valign="top">3.</td> + <td valign="top">Thanet sands.</td></tr> + +</table> + +<p class="indent"> +III. EOCENE STRATA OF THE UNITED STATES.—The lowest member of +the Eocene deposits of North America is the so-called "<i>Lignitic +Formation</i>," which is largely developed in Mississippi, Tennessee, +Arkansas, Wyoming, Utah, Colorado, and California, and sometimes +attains a thickness of several thousand feet. Stratigraphically, +this formation exhibits the interesting point that it graduates +downwards insensibly and conformably into the Cretaceous, whilst +it is succeeded <i>uncomformably</i> by strata of Middle Eocene +age. Lithologically, the series consists principally of sands +and clays, with beds of lignite and coal, and its organic remains +show that it is principally of fresh-water origin with a partial +intermixture of marine beds. +<a name="page_289"><span class="page">Page 289</span></a> +These marine +strata of the "Lignitic formation" are of special interest, as +showing such a commingling of Cretaceous and Tertiary types of +life, that it is impossible to draw any rigid line in this region +between the Mesozoic and Kainozoic systems. Thus the marine beds +of the Lignitic series contain such characteristic Cretaceous +forms as <i>Inoceramus</i> and <i>Ammonites</i>, along with a great +number of Univalves of a distinctly Tertiary type (Cones, Cowries, +&c.) Upon the whole, therefore, we must regard this series of +deposits as affording a kind of transition between the Cretaceous +and the Eocene, holding in some respects a position which may be +compared with that held by the Purbeck beds in Britain as regards +the Jurassic and Cretaceous. +</p> + +<p class="indent"> +The Middle Eocene of the United States is represented by the +<i>Claiborne</i> and <i>Jackson</i> beds. The <i>Claiborne series</i> +is extensively developed at Claiborne, Alabama, and consists of +sands, clays, lignites, marls, and impure limestones, containing +marine fossils along with numerous plant-remains. The <i>Jackson +series</i> is represented by lignitic clays and marls which occur +at Jackson, Mississippi. Amongst the more remarkable fossils of +this series are the teeth and bones of Cetaceans of the genus +<i>Zeuglodon</i>. +</p> + +<p class="indent"> +Strata of Upper Eocene age occur in North America at Vicksburg, +Mississippi, and are known as the <i>Vicksburg series</i>. They +consist of lignites, clays, marls, and limestones. Freshwater +deposits of Eocene age are also largely developed in parts of +the Rocky Mountain region. The most remarkable fossils of these +beds are Mammals, of which a large number of species have been +already determined. +</p> + +<p class="center">LIFE OF THE EOCENE PERIOD.</p> + +<p class="indent"> +The fossils of the Eocene deposits are so numerous that nothing +more can be attempted here than to give a brief and general sketch +of the life of the period, special attention being directed to some +of the more prominent and interesting types, amongst which—as +throughout the Tertiary series—the Mammals hold the first place. +It is not uncommon, indeed, to speak of the Tertiary period as a +whole under the name of the "Age of Mammals," a title at least +as well deserved as that of "Age of Reptiles" applied to the +Mesozoic, or "Age of Molluscs" applied to the Palæozoic epoch. +</p> + +<p class="indent"> +As regards the <i>plants</i> of the Eocene, the chief point to +be noticed is, that the conditions which had already set in with +the commencement of the Upper Cretaceous, are here continued, +<a name="page_290"><span class="page">Page 290</span></a> +and still further enforced. The +<i>Cycads</i> of the Secondary period, if they have not totally +disappeared, are exceedingly rare; and the <i>Conifers</i>, +losing the predominance which they enjoyed in the Mesozoic, are +now relegated to a subordinate though well-defined place in the +terrestrial vegetation. The great majority of the Eocene plants +are referable to the groups of the Angiospermous Exogens and the +Monocotyledons; and the vegetation of the period, upon the whole, +approximates closely to that now existing upon the earth. The +plants of the European Eocene are, however, in the main most +closely allied to forms which are now characteristic of tropical +or sub-tropical regions. Thus, in the London Clay are found +numerous fruits of Palms (<i>Napdites</i>, fig. 213), along with +various other plants, + +<span style="float: left; margin: 4px; width: 176px; + font-size: smaller; text-align: justify;"> +<img src="images/fig213.jpg" width="176" height="222" alt="Fig. 213" /> +<br /> +Fig. 213.—<i>Napadites ellipticus</i>, the fruit of a fossil +Palm. London Clay, Isle of Sheppey. +</span> + +most of which indicate a warm climate as prevailing in the south +of England at the commencement of the Eocene period. In the Eocene +strata of North America occur numerous plants belonging to existing +types—such as Palms, Conifers, the Magnolia, Cinnamon, Fig. +Dog-wood, Maple, Hickory, Poplar, Plane, &c. Taken as a whole, +the Eocene flora of North America is nearly related to that of +the Miocene strata of Europe, as well as to that now existing +in the American area. We conclude, therefore, that "the forests +of the American Eocene resembled those of the European Miocene, +and even of modern America" (Dana). +</p> + +<p class="indent"> +As regards the <i>animals</i> of the Eocene period, the +<i>Protozoans</i> are represented by numerous <i>Foraminifera</i>, +which reach here their maximum of development, both as regards +the size of individuals and the number of generic types. Many +of the Eocene Foraminifers are of small size; but even these not +uncommonly form whole rock-masses. Thus, the so-called "Miliolite +Limestone" of the Paris basin, largely used as a building-stone, +is almost wholly composed of the shells of a small species of +<i>Miliola</i>. The most remarkable, however, of the many members +of this group of animals which flourished in Eocene times, are the +"Nummulites" (<i>Nummulina</i>), so called from their resemblance +in shape to coins (Lat. <i>nummus</i>, a coin). The Nummulites are +amongst the largest of all known <i>Foraminifera</i>, sometimes +attaining a size of three inches in circumference; and their +internal structure is very complex (fig. 214). +<a name="page_291"><span class="page">Page 291</span></a> +Many species are +known, and they are particularly characteristic of the Middle and +Upper of these periods—their place being sometimes taken + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 441px; + font-size: smaller; text-align: center;"> +<img src="images/fig214.jpg" width="441" height="176" alt="Fig. 214" /> +<br /> +Fig. 214.—<i>Nummulina lĹ“vigata</i>. Middle Eocene. +</span> +</span> + +by <i>Orbitoides</i>, a form very similar to the Nummulite in +external appearance, but differing in its internal details. In +the Middle Eocene, the remains of Nummulites are found in vast +numbers in a very widely-spread and easily-recognised formation +known as the "Nummulitic Limestone" (fig. 10). According to Sir +Charles Lyell, "the Nummulitic Limestone of the Swiss Alps rises +to more than 10,000 feet above the level of the sea, and attains +here and in other mountain-chains a thickness of several thousand +feet. It may be said to play a far more conspicuous part than +any other Tertiary group in the solid framework of the earth's +crust, whether in Europe, Asia, or Africa. It occurs in Algeria +and Morocco, and has been traced from Egypt, where it was largely +quarried of old for the building of the Pyramids, into Asia Minor, +and across Persia by Bagdad to the mouths of the Indus. It has +been observed not only in Cutch, but in the mountain-ranges which +separate Scinde from Persia, and which form the passes leading +to Cabul; and it has been followed still further eastward into +India, as far as Eastern Bengal and the frontiers of China." The +shells of Nummulites have been found at an elevation of 16,500 +feet above the level of the sea in Western Thibet; and the +distinguished and philosophical geologist just quoted, further +remarks, that "when we have once arrived at the conviction that +the Nummulitic formation occupies a middle and upper place in the +Eocene series, we are struck with the comparatively modern date to +which some of the greatest revolutions in the physical geography +of Europe, Asia, and Northern Africa must be referred. All the +mountain-chains—such as the Alps, Pyrenees, Carpathians, and +Himalayas—into the composition of whose central and loftiest +parts the Nummulitic +<a name="page_292"><span class="page">Page 292</span></a> +strata enter bodily, could have had no existence till after the +Middle Eocene period. During that period, the sea prevailed where +these chains now rise; for Nummulites and their accompanying +Testacea were unquestionably inhabitants of salt water." +</p> + +<p class="indent"> +The <i>CĹ“lenterates</i> of the Eocene are represented +principally by <i>Corals</i>, mostly of types identical with or +nearly allied to those now in existence. Perhaps the most +characteristic group of these is that of the <i>TurbinolidĹ“</i>, +comprising a number of simple "cup-corals," which probably lived in +moderately deep water. One of the forms belonging to this family is +here figured (fig. 215). Besides true Corals, the Eocene deposits have + +<span style="float: left; margin: 4px; width: 156px; + font-size: smaller; text-align: justify;"> +<img src="images/fig215.jpg" width="156" height="490" alt="Fig. 215" /> +<br /> +Fig. 215.—<i>Turbinolia sulcata</i>, viewed from one side, +and from above. Eocene. +</span> + +yielded the remains of the "Sea-pens" (<i>PennatulidĹ“</i>) and +the branched skeletons of the "Sea-shrubs" (<i>GorgontidĹ“</i>). +</p> + +<p class="indent"> +The <i>Echinoderms</i> are represented principally by Sea-urchins, +and demand nothing more than mention. It is to be observed, however, +that the great group of the Sea-lilies (<i>Crinoids</i>) is now +verging on extinction, and is but very feebly represented. +</p> + +<p class="indent"> +Amongst the <i>Mollusca</i>, the <i>Polyzoans</i> and +<i>Brachiopods</i> also require no special mention, beyond the +fact that the latter are greatly reduced in numbers, and belong +principally to the existing genera <i>Terebratula</i> and +<i>Rhynchonella</i>. The Bivalves (<i>Lamellibranchs</i>) and +the Univalves (<i>Gasteropods</i>) are exceedingly numerous, and +almost all the principal existing genera are now represented; +though less than five percent of the Eocene <i>species</i> are +identical with those now living. It is difficult to make any +selection from the many Bivalves which are known in deposits of +this age; but species of <i>Cardita, Crassatella, Leda, Cyrena, +Mactra, Cardium, Psammobia</i>, &c., may be mentioned as very +characteristic. The <i>Caradita planicosta</i> here figured (fig. +216) is not only very abundant in the Middle Eocene, but is very +widely distributed, ranging from Europe to the Pacific coast of +North America. The <i>Univalves</i> of the Eocene are extremely +numerous, and generally beautifully preserved. The majority of them +belong to that great section of the <i>Gasteropods</i> in which the +mouth of the shell is notched or produced into +<a name="page_293"><span class="page">Page 293</span></a> +a canal (when the +shell is said to be "siphonostomatous")—this section including +the carnivorous and most highly-organized groups of the class. Not + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 530px; + font-size: smaller; text-align: center;"> +<img src="images/fig216.jpg" width="530" height="209" alt="Fig. 216" /> +<br /> +Fig. 216.—<i>Cardita planicosta</i>. Middle Eocene. +</span> +</span> + +only is this the case, but a large number of the Eocene Univalves +belong to types which now attain their maximum of development in the +warmer regions of the globe. Thus we find numerous species of Cones +(<i>Conus</i>), Volutes (<i>Voluta</i>), Cowries (<i>CyprĹ“a</i>, + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 281px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig217.jpg" width="281" height="212" alt="Fig. 217" /> +<br /> +Fig. 217.—<i>Typhis tubifer</i>, a "siphonostomatous" Univalve. +Eocene. +</span> + +<span style="width: 156px; vertical-align:bottom; text-align: justify; + margin: 4px;"> +<img src="images/fig218.jpg" width="156" height="240" alt="Fig. 218" /> +<br /> +Fig. 218.—CyprĹ“a elegans. Eocene. +</span> + +</span> + +fig. 218), Olives and Rice-shells (<i>Oliva</i>), Mitre-shells +(<i>Mitra</i>), Trumpet-shells (<i>Triton</i>), Auger-shells +(<i>Terebra</i>), and Fig-shells (<i>Pyrula</i>). Along with these +are many forms of <i>Pleurotoma, Rostellaria</i>, Spindle-shells +(<i>Fusus</i>), Dog-whelks (<i>Nassa</i>), <i>Murices</i>, and +many round-mouthed ("holostomatous") species, belonging to such +genera as <i>Turritella, Nerita, Natica, Scalaria</i>, &c. +The genus <i>Cerithium</i> (fig. 219), most of the living forms +of which are found in warm regions, inhabiting fresh or brackish +waters, undergoes a vast development in the Eocene period, where +<a name="page_294"><span class="page">Page 294</span></a> +it is represented by an immense number of specific forms, some +of which attain very large dimensions. In the Eocene strata of + +<span style="float: left; margin: 4px; width: 138px; + font-size: smaller; text-align: justify;"> +<img src="images/fig219.jpg" width="138" height="266" alt="Fig. 219" /> +<br /> +Fig. 219.—<i>Cerithium hexagonum</i>. Eocene. +</span> + +the Paris basin alone, nearly one hundred and fifty species of +this genus have been detected. The more strictly fresh-water +deposits of the Eocene period have also yielded numerous remains +of Univalves such as are now proper to rivers and lakes, together +with the shells of true Land-snails. Amongst these may be mentioned +numerous species of <i>LimnĹ“a</i> (fig. 220), <i>Physa</i> +(fig. 221), <i>Melania, Paludina, Planorbis, Helix, Bulimus</i>, +and <i>Cyclostoma</i> (fig. 222). +</p> + +<p class="indent"> +With regard to the <i>Cephalopods</i>, the chief point to be +noticed is, that all the beautiful and complex forms which peculiarly +characterised the Cretaceous period have here disappeared. We no +longer meet with a single example of the Turrilite, the Baculite, +the Hamite, the Scaphite, or the Ammonite. The only exception +to this statement is the occurrence of one species of Ammonite + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 92px; vertical-align:bottom; text-align: center; + margin: 12px;"> +<img src="images/fig220.jpg" width="92" height="186" alt="Fig. 220" /> +<br /> +Fig. 220.—<i>LimnĹ“a pyramidalis</i>. Eocene. +</span> + +<span style="width: 64px; vertical-align:bottom; text-align: center; + margin: 12px;"> +<img src="images/fig221.jpg" width="64" height="194" alt="Fig. 221" /> +<br /> +Fig. 221.—<i>Physa columnaris</i>. Eocene. +</span> + +<span style="width: 95px; vertical-align:bottom; text-align: center; + margin: 12px;"> +<img src="images/fig222.jpg" width="95" height="156" alt="Fig. 222" /> +<br /> +Fig. 222.—<i>Cyclostoma Arnoudii</i>. Eocene. +</span> + +</span> + +in the so-called "Lignitic Formation" of North America; but the +beds containing this may possibly be rather referable to the +Cretaceous—and this exception does not affect the fact +that the <i>AmmonitidĹ“</i>, as a family, had become +extinct before the Eocene strata were deposited. The ancient +genus <i>Nautilus</i> still survives, the sole representative +of the once mighty order of the Tetrabranchiate Cephalopods. +In the order of the <i>Dibranchiates</i>, we have a like +phenomenon to observe in the total extinction of the great +family of the "Belemnites." No form referable to this group +<a name="page_295"><span class="page">Page 295</span></a> +has hitherto been found in any Tertiary stratum; but the +internal skeletons of Cuttle-fishes (such as <i>Belosepia</i>) +are not unknown. +</p> + +<p class="indent"> +Remains of <i>Fishes</i> are very abundant in strata of Eocene +age, especially in certain localities. The most famous depot for +the fossil fishes of this period is the limestone of Monte Bolca, +near Verona, which is interstratified with beds of volcanic ashes, +the whole being referable to the Middle Eocene. The fishes here +seem to have been suddenly destroyed by a volcanic eruption, +and are found in vast numbers. Agassiz has described over one +hundred and thirty species of Fishes from this locality, belonging +to seventy-seven genera. All the <i>species</i> are extinct; but +about one-half of the <i>genera</i> are represented by living +forms. The great majority of the Eocene Fishes belong to the + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 500px; + font-size: smaller; text-align: center;"> +<img src="images/fig223.jpg" width="500" height="291" alt="Fig. 223" /> +<br /> +Fig. 223.—<i>Rhombus minimus</i>, a small fossil Turbot from +the Eocene Tertiary, Monte Bolca. +</span> +</span> + +order of the "Bony Fishes" (<i>Teleosteans</i>), so that in the +main the forms of Fishes characterising the Eocene are similar +to those which predominate in existing seas. In addition to the +above, a few <i>Ganoids</i> and a large number of <i>Placoids</i> +are known to occur in the Eocene rocks. Amongst the latter are +found numerous teeth of true Sharks, such as <i>Otodus</i> (fig. +224) and <i>Carcharodon</i>. The pointed and serrated teeth of the +latter sometimes attain a length of over half a foot, indicating +that these predaceous fishes attained gigantic dimensions; and it +is interesting to note that teeth, in external appearance very +similar to those of the early Tertiary genus <i>Carcharodon</i>, +have been dredged from great depths during the recent expedition +of the Challenger. There also occur not uncommonly the flattened +<a name="page_296"><span class="page">Page 296</span></a> +teeth of Rays (fig. 225), consisting of flat bony pieces placed +close together, and forming "a kind of mosaic pavement on both +the upper and lower jaws" (Owen). +</p> + +<p class="indent"> +In the class of the <i>Reptiles</i>, the disappearance of the + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 130px; vertical-align:bottom; + text-align: center; margin: 4px;"> +<img src="images/fig224.jpg" width="130" height="168" alt="Fig. 224" /> +<br /> +Fig. 224.—Tooth of <i>Otodus obliquus</i>. Eocene. +</span> + +<span style="width: 240px; vertical-align:bottom; + text-align: center; margin: 4px;"> +<img src="images/fig225.jpg" width="240" height="153" alt="Fig. 225" /> +<br /> +Fig. 225.—Flattened dental plates of a Ray (<i>Myliobatis +Edwardsii</i>). Eocene. +</span> + +</span> + +characteristic Mesozoic types is as marked a phenomenon as the +introduction of new forms. The Ichthyosaurs, the Plesiosaurs, +the Pterosaurs, and the Mosasaurs of the Mesozoic, find no +representatives in the Eocene Tertiary; and the same is true +of the Deinosaurs, if we except a few remains from the +doubtfully-situated "Lignitic formation" of the United States, +On the other hand, all the modern orders of Reptiles are known to +have existed during the Eocene period. The <i>Chelonians</i> are +represented by true marine Turtles, by "Terrapins" +(<i>EmydidĹ“</i>), and by "Soft Tortoises" +(<i>TrionycidĹ“</i>). The order of the Snakes and Serpents +(<i>Ophidia</i>) makes its appearance here, for the first time +under several forms—all of which, however, are referable to +the non-venomous group of the "Constricting Serpents" +(<i>BoidĹ“</i>). The oldest of these is the <i>PalĹ“ophis +toliapicus</i> of the London Clay of Sheppey, first made known to +science by the researches of Professor Owen. The nearly-allied +<i>PalĹ“ophis typhĹ“us</i> of the Eocene beds of +Bracklesham appears to have been a Boa-constrictor-like Snake of +about twenty feet in length. Similar Python-like Snakes +(<i>PalĹ“ophis, Dinophis</i>, &c.) have been described +from the Eocene deposits of the United States. True Lizards +(<i>Lacertilians</i>) are found in some abundance in the Eocene +deposits,—some being small terrestrial forms, like the +common European lizards of the present day; whilst others equal or +exceed the living Monitors in size. Lastly, the modern order of +the <i>Crocodilia</i> is largely represented in Eocene times, by +species belonging to all the existing genera, together with +others referable to extinct types. As pointed out by Owen, it +is an interesting fact that in the Eocene rocks of the south-west +<a name="page_297"><span class="page">Page 297</span></a> +of England, there occur fossil remains of all the three living +types of Crocodilians—namely, the Gavials, the true +Crocodiles, and the Alligators (fig. 226)—though at the + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 502px; + font-size: smaller; text-align: center;"> +<img src="images/fig226.jpg" width="502" height="263" alt="Fig. 226" /> +<br /> +Fig. 226.—Upper jaw of Alligator. Eocene Tertiary, Isle of +Wight. +</span> +</span> + +present day these forms are all geographically restricted in +their range, and are never associated together. +</p> + +<p class="indent"> +Almost all the existing orders of <i>Birds</i>, if not all, are +represented in the Eocene deposits by remains often very closely +allied to existing types. Thus, amongst the Swimming Birds +(<i>Natatores</i>) we find examples of forms allied to the living +Pelicans and Mergansers; amongst the Waders (<i>Grallatores</i>) +we have birds resembling the Ibis (the <i>Numenius gypsorum</i> of +the Paris basin); amongst the Running Birds (<i>Cursores</i>) we +meet with the great <i>Gastornis Parisiensis</i>, which equalled +the African Ostrich in height, and the still more gigantic +<i>Dasornis Londinensis</i>; remains of a Partridge represent +the Scratching Birds (<i>Rasores</i>); the American Eocene has +yielded the bones of one of the Climbing Birds (<i>Scansores</i>), +apparently referable to the Woodpeckers; the <i>Protornis +Glarisiensis</i> of the Eocene Schists of Glaris is the oldest +known example of the Perching Birds (<i>Insessores</i>); and +the Birds of Prey (<i>Raptores</i>) are represented by Vultures, +Owls, and Hawks. The toothed Birds of the Upper Cretaceous are no +longer known to exist; but Professor Owen has recently described +from the London Clay the skull of a very remarkable Bird, in +which there is, at any rate, an approximation to the structure of +<i>Ichthyornis</i> and <i>Hesperornis</i>. The bird in question +has been named the <i>Odontopteryx totiapicus</i>, its generic +title being derived from the very remarkable characters of its +jaws. In this singular form (fig. 227) the margins of both jaws +<a name="page_298"><span class="page">Page 298</span></a> +are furnished with tooth-like denticulations, which differ from +true teeth in being actually portions of the bony substance of + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 559px; + font-size: smaller; text-align: center;"> +<img src="images/fig227.jpg" width="559" height="202" alt="Fig. 227" /> +<br /> +Fig. 227.—Skull of <i>Odontopteryx toliapicus</i> restored. +(After Owen.) +</span> +</span> + +the jaw itself, with which they are continuous, and which were +probably encased by extensions of the horny sheath of the bill. +These tooth-like processes are of two sizes, the larger ones +being comparable to canines; and they are all directed forwards, +and have a triangular or compressed conical form. From a careful +consideration of all the discovered remains of this bird, Professor +Owen concludes that "<i>Odontopteryx</i> was a warm-blooded +feathered biped, with wings; and further, that it was web-footed +and a fish-eater, and that in the catching of its slippery prey +it was assisted by this Pterosauroid armature of its jaws." Upon +the whole, <i>Odontopteryx</i> would appear to be most nearly +related to the family of the Geese (<i>AnserinĹ“</i>) or Ducks +(<i>AnatidĹ“</i>); but the extension of the bony substance of +the jaws into tooth-like processes is an entirely unique character, +in which it stands quite alone. +</p> + +<p class="indent"> +The known <i>Mammals</i> of the Mesozoic period, as we have seen, +are all of small size; and with one not unequivocal exception, +they appear to be referable to the order of the Pouched Quadrupeds +(<i>Marsupials</i>), almost the lowest group of the whole class +of the Mammalia. In the Eocene rocks, on the other hand, numerous +remains of Quadrupeds have been brought to light, representing +most of the great Mammalian orders now in existence upon the +earth, and in many cases indicating animals of very considerable +dimensions. We are, in fact, in a position to assert that the +majority of the great groups of Quadrupeds with which we are +familiar at the present day were already in existence in the +Eocene period, and that their ancient root-stocks were even in +this early time separated by most of the fundamental differences +of structure +<a name="page_299"><span class="page">Page 299</span></a> +which distinguish their living representatives. At the same time, +there are some amongst the Eocene quadrupeds which have a +"generalised" character, and which may be regarded as structural +types standing midway between groups now sharply separated from +one another. +</p> + +<p class="indent"> +The order of the <i>Marsupials</i>—including the existing +Kangaroos, Wombats, Opossums, Phalangers, &c.—is poorly +represented in deposits of Eocene age. The most celebrated +example of this group is the <i>Didelphys gypsorum</i> of the +Gypseous beds of Montmartre, near Paris, an Opossum very nearly +allied to the living Opossums of North and South America. +</p> + +<p class="indent"> +No member of the <i>Edenates</i> (Sloths, Ant-eaters, and Armadillos) +has hitherto been detected in any Eocene deposit. The aquatic +order of the <i>Sirenians</i> (Dugongs and Manatees), with their +fish-like bodies and tails, paddle-shaped forelimbs, and wholly +deficient hind-limbs, are represented in strata of this age by +remains of the ancient "Sea-Cows," to which the name of +<i>Halitherium</i> has been applied. Nearly allied to the preceding +is the likewise aquatic order of the Whales and Dolphins +(<i>Cetaceans</i>), in which the body is also fish-like, the +hind-limbs are wanting, the fore-limbs are converted into powerful +"flippers" or swimming-paddles, and the terminal extremity of +the body is furnished with a horizontal, tail-fin. Many existing +Cetaceans (such as the Whalebone Whales) have no true teeth; +but others (Dolphins, Porpoises, Sperm Whales) possess simple + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 510px; + font-size: smaller; text-align: center;"> +<img src="images/fig228.jpg" width="510" height="248" alt="Fig. 228" /> +<br /> +Fig. 228.—<i>Zeuglodon cetoides</i>. A, Molar +tooth of the natural size; B, Vertebra, reduced in size. From the +Middle Eocene of the United States. (After Lyell.) +</span> +</span> + +conical teeth. In strata of Eocene age, however, we find a singular +group of Whales, constituting the genus <i>Zeuglodon</i> (fig. 228), +<a name="page_300"><span class="page">Page 300</span></a> +in which the teeth differed from those of all +existing forms in being of two kinds,—the front ones being +conical incisors, whilst the back teeth or molars have serrated +triangular crowns, and are inserted in the jaw by two roots. Each +molar (fig. 228, A) looks as if it were composed of two separate +teeth united on one side by their crowns; and it is this peculiarity +which is expressed by the generic name (Gr. <i>zeugle</i>, a yoke; +<i>odous</i>, tooth). The best-known species of the genus is +the <i>Zeuglodon cetoides</i> of Owen, which attained a length +of seventy feet. Remains of these gigantic Whales are very common +in the "Jackson Beds" of the Southern United States. So common +are they that, according to Dana, "the large vertebræ, some of +them a foot and a half long and a foot in diameter, were formerly +so abundant over the country, in Alabama, that they were used +for making walls, or were burned to rid the fields of them." +</p> + +<p class="indent"> +The great and important order of the Hoofed Quadrupeds +(<i>Ungulata</i>) is represented in the Eocene by examples of both +of its two principal sections—namely, those with an uneven +number of toes (one or three) on the foot (<i>Perissodactyle +Ungulates</i>), and those with an even number of toes (two or four) +to each foot (<i>Artiodactyle Ungulates</i>). Amongst the Odd-toed +Ungulates, the living family of the Tapirs (<i>TapirdĹ“</i>) is +represented by the genus <i>Coryphodon</i> of Owen. Nearly related +to the preceding are the species of <i>PalĹ“otherium</i>, +which have a historical interest as being amongst the first of the +Tertiary Mammals investigated by the illustrious Cuvier. Several +species of <i>PalĹ“othere</i> are known, varying greatly in +size, the smallest being little bigger than a hare, whilst the +largest must have equalled a good-sized horse in its dimensions. The +species of <i>PalĹ“otherium</i> appear to have agreed with the +existing Tapirs in possessing a lengthened and flexible nose, which +formed a short proboscis or trunk (fig. 229), suitable as an +instrument for stripping off the foliage of trees—the characters +of the molar teeth showing them to have been strictly herbivorous +in their habits. They differ, however, from the Tapirs, amongst +other characters, in the fact that both the fore and the hind feet +possessed three toes each; whereas in the latter there are four +toes on each fore-foot, and the hind-feet alone are three-toed. The +remains of <i>PalĹ“otheria</i> have been found in such abundance +in certain localities as to show that these animals roamed in great +herds over the fertile plains of France and the south of England +during the later portion of the Eocene period. The accompanying +illustration (fig. 229) represents the notion which the great +Cuvier was induced by +<a name="page_301"><span class="page">Page 301</span></a> +his researches to form as to the outward +appearance of <i>PalĹ“otherium magnum</i>. Recent discoveries, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 487px; + font-size: smaller; text-align: center;"> +<img src="images/fig229.jpg" width="487" height="294" alt="Fig. 229" /> +<br /> +Fig. 229.—Outline of <i>PalĹ“otherium magnum</i>, +restored. Upper Eocene, Europe. (After Cuvier.) +</span> +</span> + +however, have rendered it probable that this restoration is in some +important respects inaccurate. Instead of being bulky, massive, +and more or less resembling the living Tapirs in form, it would +rather appear that <i>PalĹ“otherium magnum</i> was in reality a +slender, graceful, and long-necked animal, more closely resembling +in general figure a Llama, or certain of the Antelopes. +</p> + +<p class="indent"> +The singular genus <i>Anchitherium</i> forms a kind of transition +between the <i>PalĹ“otheria</i> and the true Horses +(<i>EquidĹ“</i>). The Horse (fig. 230, D) possesses but one +fully-developed toe to each foot, this being terminated by a single +broad hoof, and representing the <i>middle</i> toe—the +<i>third</i> of the typical five-fingered or five-toed limb of +Quadrupeds in general. In addition, however, to this fully-developed +toe, each foot in the horse carries two rudimentary toes which are +concealed beneath the skin, and are known as the "splint-bones." +These are respectively the <i>second</i> and <i>fourth</i> toes, +in an aborted condition; and the first and fifth toes are wholly +wanting. In <i>Hipparion</i> (fig. 230, C), the foot is essentially +like that of the modern Horses, except that the second and fourth +toes no longer are mere "splint-bones," hidden beneath the skin; +but have now little hoofs, and hang freely, but uselessly, by the +side of the great middle toe, not being sufficiently developed to +reach the ground. In <i>Anchitherium</i>, again (fig. 230, B), +the foot is three-toed, like that of <i>Hipparion</i>; but the +two lateral toes (the second and fourth) are so far +<a name="page_302"><span class="page">Page 302</span></a> +developed that they now reach the ground. +The <i>first</i> digit (thumb or great toe) is still wanting; +as also is the <i>fifth</i> digit (little finger or little toe). + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 518px; + font-size: smaller; text-align: justify;"> +<img src="images/fig230.jpg" width="518" height="283" alt="Fig. 230" /> +<br /> +Fig. 230.—Skeleton of the foot in various forms belonging to +the family of the <i>EquidĹ“</i>. A, Foot of <i>Orohippus</i>, +Eocene; B, Foot of <i>Anchitherium</i>, Upper Eocene and Lower +Miocene; C, Foot of <i>Hipparion</i>, Upper Miocene and Pliocene: +D, Foot of Horse (<i>Equus</i>), Pliocene and Recent. The figures +indicate the numbers of the digits in the typical five-fingered +hand of Mammals. (After Marsh.) +</span> +</span> + +Lastly, the Eocene rocks have yielded in North America the remains +of a small Equine quadruped, to which Marsh has given the name of +<i>Orohippus</i>. In this singular form—which was not larger +than a fox—the foot (fig. 230, A) carries <i>four</i> toes, +all of which are hoofed and touch the ground, but of which the +<i>third</i> toe is still the largest. The <i>first</i> toe +(thumb or great toe) is still wanting; but in this ancient +representative of the Horses, the <i>fifth</i> or "little" toe +appears for the first time. As all the above-mentioned forms +succeed one another in point of time, it may be regarded as +probable that we shall yet be able to point, with some certainty, +to some still older example of the <i>EquidĹ“</i>, in which +the first digit is developed, and the foot assumes its typical +five-fingered condition. +</p> + +<p class="indent"> +Passing on to the Even-toed or <i>Artiodactyle Ungulates</i>, no +representative of the <i>Hippotamus</i> seems yet to have existed, +but there are several forms (<i>ChĹ“ropotamus, Hyopotamus</i>, +&c.) more or less closely allied to the Pigs (<i>Suida</i>); and +the singular group of the <i>AnoplotheridĹ“</i> may be regarded +as forming a kind of transition between the Swine and the Ruminants. +The <i>Anoplotheria</i> (fig. 231) were slender in form, the +largest not exceeding a donkey in size, with long tails, and +having the feet terminated by two hoofed toes each, sometimes +with a pair of small accessory hoofs as well. The teeth exhibit +<a name="page_303"><span class="page">Page 303</span></a> +the peculiarity that they are arranged in a continuous series, +without any gap or interval between the molars and the canines; + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 495px; + font-size: smaller; text-align: center;"> +<img src="images/fig231.jpg" width="495" height="240" alt="Fig. 231" /> +<br /> +Fig. 231.—<i>Anoplotherium commune</i>. Eocene Tertiary, +France. (After Cuvier.) +</span> +</span> + +and the back teeth, like those of all the Ungulates, are adapted +for grinding vegetable food, their crowns resembling in form +those of the true Ruminants. The genera <i>Dichobune</i> and +<i>Xiphodon</i>, of the Middle and Upper Eocene, are closely related +to <i>Anoplotherium</i>, but are more slender and deer-like in +form. No example of the great Ruminant group of the Ungulate +Quadrupeds has as yet been detected in deposits of Eocene age. +</p> + +<p class="indent"> +Whilst true Ruminants appear to be unknown, the Eocene strata +of North America have yielded to the researches of Professor +Marsh examples of an extraordinary group (<i>Dinocerata</i>), +which may be considered as in some respects intermediate between +the Ungulates and the Proboscideans. In <i>Dinoceras</i> itself +(fig. 232) we have a large animal, equal in dimensions to the +living Elephants, which it further resembles in the structure +of the massive limbs, except that there are only four toes to +each foot. The upper jaw was devoid of front teeth, but there +were two very large canine teeth, in the form of tusks directed +perpendicularly downwards; and there was also a series of six small +molars on each. Each upper jaw-bone carried a bony projection, which +was probably of the nature of a "horn-core," and was originally +sheathed in horn. Two similar, but smaller, horn-cores are carried +on the nasal bones; and two much larger projections, also probably +of the nature of horn-cores, were carried upon the forehead. We +may thus infer that <i>Dinoceras</i> possessed three pairs of +horns, all of which resembled the horns of the Sheep and Oxen +in consisting of a central bony "core," surrounded by a horny +<a name="page_304"><span class="page">Page 304</span></a> +sheath. The nose was not prolonged into a proboscis or "trunk," +as in the existing Elephants; and the tail was short and slender. + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 538px; + font-size: smaller; text-align: center;"> +<img src="images/fig232.jpg" width="538" height="411" alt="Fig. 232" /> +<br /> +Fig. 232.—Skull of <i>Dinoceras mirabilis</i>, greatly +reduced. Eocene, North America. (After Marsh.) +</span> +</span> + +Many forms of the <i>Dinocerata</i> are known; but all these +singular and gigantic quadrupeds appear to have been confined +to the North American continent, and to be restricted to the +Eocene period. +</p> + +<p class="indent"> +The important order of the Elephants (<i>Proboscidea</i>) is +also not known to have come into existence during the Eocene +period. On the other hand, the great order of the Beasts of Prey +(<i>Carnivora</i>) is represented in Eocene strata by several +forms belonging to different types. Thus the <i>Ardocyon</i> +presents us with an Eocene Carnivore more or less closely allied +to the existing Racoons; the <i>PalĹ“onyctis</i> appears to +be related to the recent Civet-cats; the genus <i>HyĹ“nodon</i> +is in some respects comparable to the living Hyænas; and the +<i>Canis Parisiensis</i> of the gypsum-bearing beds of Montmartre +may perhaps be allied to the Foxes. +</p> + +<p class="indent"> +The order of the Bats (<i>Cheiroptera</i>) is represented in Eocene +strata of the Paris basin (Gypseous series of Montmartre) by the +<i>Vespertilio Parisiensis</i> (fig. 233), an insect-eating Bat +very similar to some of the existing European forms. Lastly, the +Eocene deposits have yielded more or less satisfactory evidence of +<a name="page_305"><span class="page">Page 305</span></a> +the existence in Europe at this period of examples of the orders + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 451px; + font-size: smaller; text-align: center;"> +<img src="images/fig233.jpg" width="451" height="370" alt="Fig. 233" /> +<br /> +Fig. 233.—Portion of the skeleton of <i>Vespertilio +Parisienis</i>. Eocene Tertiary, France. +</span> +</span> + +of the Gnawing Mammals (<i>Rodentia</i>), the Insect-eating Mammals +(<i>Insectivora</i>), and the Monkeys (<i>Quadrumana</i>).[24] +</p> + +<p class="footnote"> +[Footnote 24: A short list of the more important works relating to +the Eocene rocks and fossils will be given after all the Tertiary +deposits have been treated of.] +</p> + +<h3>CHAPTER XIX.</h3> + +<p class="subtitle"> +THE MIOCENE PERIOD. +</p> + +<p class="indent"> +The Miocene rocks comprise those Tertiary deposits which contain +less than about 35 per cent of existing species of shells +(<i>Mollusca</i>), and more than 5 per cent—or those deposits +in which the proportion of living shells is less than of extinct +species. They are divisible into a <i>Lower Miocene</i> +(<i>Oligocene</i>) and an <i>Upper Miocene</i> series. +</p> + +<p class="indent"> +In <i>Britain</i>, the Miocene rocks are very poorly developed, one +of their leading developments being at Bovey Tracy in Devonshire, +where there occur sands, clays, and beds of lignite +<a name="page_306"><span class="page">Page 306</span></a> +or imperfect coal. These strata contain numerous plants, +amongst which are Vines, Figs, the Cinnamon-tree, Palms, and many +Conifers, especially those belonging to the genus Sequoia (the +"Red-Foods"). These Bovey Tracy lignites are of Lower Miocene age, +and they are lacustrine in origin. Also of Lower Miocene age are +the so-called "Hempstead Beds" of Yarmouth in the Isle of Wight. +These attain a thickness of less than 200 feet, and are shown by +their numerous fossils to be principally a true marine formation. +Lastly, the Duke of Argyll, in 1851, showed that there existed at +Ardtun, in the island of Mull, certain Tertiary strata containing +numerous remains of plants; and these also are now regarded as +belonging to the Lower Miocene. +</p> + +<p class="indent"> +In <i>France</i>, the Lower Miocene is represented in Auvergne, +Cantal, and Velay, by a great thickness of nearly horizontal +strata of sands, sandstone, clays, marls, and limestones, the +whole of fresh-water origin. The principal fossils of these +lacustrine deposits are <i>Mammalia</i>, of which the remains +occur in great abundance. In the valley of the Loire occur the +typical European deposits of Upper Miocene age. These are known +as the "Faluns," from a provincial term applied to shelly sands, +employed to spread upon soils which are deficient in lime; and +the Upper Miocene is hence sometimes spoken of as the "Falunian" +formation. The Faluns occur in scattered patches, which are rarely +more than 50 feet in thickness, and consist of sands and marls. +The fossils are chiefly marine; but there occur also land and +fresh-water shells, together with the remains of numerous Mammals. +About 25 per cent of the shells of the Faluns are identical with +existing species. The sands, limestones, and marls of the Department +of Gers, near the base of the Pyrenees, rendered famous by the +number or Mammalian remains exhumed from them by M. Lartet, also +belong to the age of the Faluns. +</p> + +<p class="indent"> +In <i>Switzerland</i>, between the Alps and the Jura, there occurs +a great series of Miocene deposits, known collectively as the +"Molasse," from the soft nature of a greenish sandstone, which +constitutes one of its chief members. It attains a thickness of +many thousands of feet, and rises into lofty mountains, some of +which—as the Rigi—are more than 6000 feet in height. +The middle portion of the Molasse is of marine origin, and is +shown by its fossils to be of the age of the Faluns; but the +lower and upper portions of the formation are mainly or entirely +of fresh-water origin. The Lower Molasse (of Lower Miocene age) +has yielded about 500 species of plants, mostly of tropical or +sub-tropical forms. The Upper +<a name="page_307"><span class="page">Page 307</span></a> +Molasse has yielded about the same number of plants, with about +900 species of Insects, such as wood-eating Beetles Water-beetles, +White Ants, Dragon-flies, &c. +</p> + +<p class="indent"> +In <i>Belgium</i>, strata of both Lower and Upper Miocene age are +known,—the former (<i>Rupelian Clays</i>) containing numerous +marine fossils; whilst the latter (<i>Bolderberg Sands</i>) have +yielded numerous shells corresponding with those of the Faluns. +</p> + +<p class="indent"> +In <i>Austria</i>, Miocene strata are largely developed, marine +beds belonging to both the Lower and Upper division of the formation +occurring extensively in the Vienna basin. The well-known Brown +Coals of Radaboj, in Croatia, with numerous plants and insects, +are also of Lower Miocene age. +</p> + +<p class="indent"> +In <i>Germany</i>, deposits belonging to both the Lower and Upper +division of the Miocene formation are extensively developed. To +the former belong the marine strata of the Mayence basin, and +the marine <i>Rupelian Clay</i> near Berlin; whilst a celebrated +group of strata belonging to the Upper Miocene occurs near +Epplesheim, in Hesse-Darmstadt, and is well known for the number +of its Mammalian remains. +</p> + +<p class="indent"> +In <i>Greece</i>, at PikermĂ©, near Athens, there occurs a +celebrated deposit of Upper Miocene age, well known to +palæontologists through the researches of M. M. Wagner, Roth, +and Gaudry upon the numerous Mammalia which it contains. In +<i>Italy</i>, also, strata of both Lower and Upper Miocene age are +well developed in the neighbourhood of Turin. +</p> + +<p class="indent"> +In the <i>Siwâlik Hills</i>, in India, at the southern foot +of the Himalayas, occurs a series of Upper Miocene strata, which +have become widely celebrated through the researches of Dr Falconer +and Sir Proby Cautley upon the numerous remains of Mammals and +Reptiles which they contain. Beds of corresponding age, with +similar fossils, are known to occur in the island of Perim in +the Gulf of Cambay. +</p> + +<p class="indent"> +Lastly, Miocene deposits are found in <i>North America</i>, in +New Jersey, Maryland, Virginia, Missouri, California, Oregon, +&c., attaining a thickness of 1500 feet or more. They consist +principally of clays, sands, and sandstones, sometimes of marine +and sometimes of fresh-water origin. Near Richmond, in Virginia, +there occurs a remarkable stratum, wrongly called "Infusorial +Earth," which is occasionally 30 feet in thickness, and consists +almost wholly of the siliceous envelopes of certain low forms of +plants (Diatoms), along with the spicules of Sponges and other +siliceous organisms (see fig. 16). The <i>White River Group</i> of +Hayden occurs in the Upper Missouri region, and is largely exposed +over the barren and desolate +<a name="page_308"><span class="page">Page 308</span></a> +district known as the "Mauvaises Terres." They have a thickness of +1000 feet or more, and contain numerous remains of Mammals. They +are of lacustrine origin, and are believed to be of the age of the +Lower Miocene. Upon the whole, about from 15 to 30 per cent of the +<i>Mollusca</i> of the American Miocene are identical with +existing species. +</p> + +<p class="indent"> +In addition to the regions previously enumerated, Miocene strata +are known to be developed in <i>Greenland, Iceland, Spitzbergen</i>, +and in other areas of less importance. +</p> + +<p class="indent"> +The <i>life</i> of the Miocene period is extremely abundant, +and, from the nature of the deposits of this age, also extremely +varied in its character. The marine beds of the formation have +yielded numerous remains of both Vertebrate and Invertebrate +sea-animals; whilst the fresh-water deposits contain the skeletons +of such shells, fishes, &c., as now inhabit rivers or lakes. +Both the marine and the lacustrine beds have been shown to contain +an enormous number of plants, the latter more particularly; whilst +the Brown Coals of the formation are made up of vegetable matter +little altered from its original condition. The remains of +air-breathing animals, such as Insects, Reptiles, Birds, and +Mammals, are also abundantly found, more especially in the +fresh-water beds. +</p> + +<p class="indent"> +The <i>plants</i> of the Miocene period are extraordinarily numerous, +and only some of the general features of the vegetation of this +epoch can be indicated here. Our chief sources of information as +to the Miocene plants are the Brown Coals of Germany and Austria, +the Lower and Upper Molasse of Switzerland, and the Miocene strata +of the Arctic regions. The lignites of Austria have yielded very +numerous plants, chiefly of a tropical character—one of the +most noticeable forms being a Palm of the genus <i>Sabal</i> +(fig. 234, B), now found in America. The plants of the Lower +Miocene of Switzerland are also mostly of a tropical character, +but include several forms now found in North America, such as a +Tulip-tree (<i>Liriodendron</i>) and a Cypress (<i>Taxodium</i>). +Amongst the more remarkable forms from these beds may be mentioned +Fan-Palms (<i>ChamĹ“rops</i>, fig. 234, A), numerous tropical +ferns, and two species of Cinnamon. The plant-remains of the +Upper Molasse of Switzerland indicate an extraordinarily rank and +luxuriant vegetation, composed mainly of plants which now live +in warm countries. Among the commoner plants of this formation may +be enumerated many species of Maple (<i>Acer</i>), Plane-trees +(<i>Platanus</i> fig. 235), Cinnamon-trees (fig. 236), and other +members of the <i>LauraceĹ“</i>, many species of +<i>ProteaccĹ“</i> (<i>Banksia, Grevillea</i>, &c.), +several species of Sarsaparilla (<i>Smilax</i>), Palms, Cypresses, +&c. +</p> + +<p class="indent"> +<a name="page_309"><span class="page">Page 309</span></a> +In Britain, the Lower Miocene strata of Bovey Tracy have yielded +remains of Ferns, Vines, Fig, Cinnamon, <i>ProteaccĹ“</i>, &c., + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 513px; + font-size: smaller; text-align: center;"> +<img src="images/fig234.jpg" width="513" height="253" alt="Fig. 234" /> +<br /> +Fig. 234.—Miocene Palms A, <i>ChamĹ“rops Helvetica</i>; B, +<i>Sabal major</i>. Lower Miocene of Switzerland and France. +</span> +</span> + +along with numerous Conifers. The most abundant of these last is a +gigantic pine—the <i>Sequoia CouttsiĹ“</i>—which is + +<span style="float: left; width: 100%; + text-align: center; font-size: smaller;"> + +<span style="width: 217px; vertical-align:bottom; text-align: justify; + margin: 12px;"> +<img src="images/fig235.jpg" width="217" height="217" alt="Fig. 235" /> +<br /> +Fig. 235.—<i>Platanus aceroides</i>, an Upper Miocene +Plane-tree. <i>a</i>, Leaf; <i>b</i>, The core of a bundle +of fruits; <i>c</i>, A single fruit. +</span> + +<span style="width: 168px; vertical-align:bottom; text-align: center; + margin: 12px;"> +<img src="images/fig236.jpg" width="128" height="248" alt="Fig. 236" /> +<br /> +Fig. 236.—<i>Cinnamomum polymorphum</i>. <i>a</i>, Leaf; +<i>b</i>, Flower. Upper Miocene. +</span> + +</span> + +very nearly allied to the huge <i>Sequoia</i> (<i>Wellingtonia</i>) +<i>gigantea</i> of California. A nearly-allied form (<i>Sequoia +Langsdorffi</i>) has been detected in the leaf-bed of Ardtun, +in the Hebrides. +</p> + +<p class="indent"> +In Greenland, as well as in other parts of the Arctic regions, +Miocene strata have been discovered which have yielded a great +number of plants, many of which are identical with species found +in the European Miocene. Amongst these +<a name="page_310"><span class="page">Page 310</span></a> +plants are found many trees, such as Conifers, Beeches, Oaks, +Maples, Plane-trees, Walnuts, Magnolias, &c., with numerous +shrubs, ferns, and other smaller plants. With regard to the Miocene +flora of the Arctic regions, Sir Charles Lyell remarks that "more +than thirty species of Coniferæ have been found, including +several Sequoias (allied to the gigantic Wellingtonia of California), +with species of <i>Thujopsis</i> and <i>Salisburia</i>, now peculiar +to Japan. There are also beeches, oaks, planes, poplars, maples, +walnuts, limes, and even a magnolia, two cones of which have +recently been obtained, proving that this splendid evergreen not +only lived but ripened its fruit within the Arctic circle. Many +of the limes, planes, and oaks were large-leaved species; and both +flowers and fruits, besides immense quantities of leaves, are in +many cases preserved. Among the shrubs are many evergreens, as +<i>Andromeda</i>, and two extinct genera, <i>Daphnogene</i> and +<i>M'Clintockia</i>, with fine leathery leaves, together with +hazel, blackthorn, holly, logwood, and hawthorn. A species of +Zamia (<i>Zimites</i>) grew in the swamps, with <i>Potamogeton, +Sparganium</i>, and <i>Menyanthes</i>; while ivy and villes twined +around the forest-trees, and broad-leaved ferns grew beneath their +shade. Even in Spitzbergen, as far north as lat. 78° 56', no +less than ninety-five species of fossil plants have been obtained, +including <i>Taxodium</i> of two species, hazel, poplar, alder, +beech, plane-tree, and lime. Such a vigorous growth of trees within +12° of the pole, where now a dwarf willow and a few herbaceous +plants form the only vegetation, and where the ground is covered +with almost perpetual snow and ice, is truly remarkable." +</p> + +<p class="indent"> +Taking the Miocene flora as a whole, Dr Heer concludes from his +study of about 3000 plants contained in the European Miocene +alone, that the Miocene plants indicate tropical or sub-tropical +conditions, but that there is a striking inter-mixture of forms +which are at present found in countries widely removed from one +another. It is impossible to state with certainty how many of the +Miocene plants belong to existing species, but it appears that +the larger number are extinct. According to Heer, the American +types of plants are most largely represented in the Miocene flora, +next those of Europe and Asia, next those of Africa, and lastly +those of Australia. Upon the whole, however, the Miocene flora +of Europe is mostly nearly allied to the plants which we now +find inhabiting the warmer parts of the United States; and this +has led to the suggestion that in Miocene times the Atlantic +Ocean was dry land, and that a migration of +<a name="page_311"><span class="page">Page 311</span></a> +American plants to Europe was thus permitted. This view is +borne out by the fact that the Miocene plants of Europe are most +nearly allied to the living plants of the eastern or Atlantic +seaboard of the United States, and also by the occurrence of a +rich Miocene flora in Greenland. As regards Greenland, Dr Heer +has determined that the Miocene plants indicate a temperate +climate in that country, with a mean annual temperature at least +30° warmer than it is at present. +</p> + +<p class="indent"> +The present limit of trees is the isothermal which gives the +mean temperature of 500 Fahr. in July, or about the parallel of +67° N. latitude. In Miocene times, however, the Limes, Cypresses, +and Plane-trees reach the 79th degree of latitude, and the Pines +and Poplars must have ranged even further north than this. +</p> + +<p class="indent"> +The <i>Invertebrate Animals</i> of the Miocene period are very +numerous, but they belong for the most part to existing types, +and they can only receive scanty consideration here. The little +shells of <i>Foraminifera</i> are extremely abundant in some beds, +the genera being in many cases such as now flourish abundantly in +our seas. The principal forms belong to the genera <i>Textularia</i> +(fig. 237), <i>Robulina, Glandulina, Polystomella, Amplistegina</i>, + +<span style="float: right; margin: 4px; width: 268px; + font-size: smaller; text-align: center;"> +<img src="images/fig237.jpg" width="268" height="176" alt="Fig. 237" /> +<br /> +Fig. 237.—<i>Textularia Meyeriana</i>, greatly +enlarged. Miocene Tertiary. +</span> + +&c. Corals are very abundant, in many instances forming regular +"reefs;" but all the more important groups are in existence at +the present day. The Red Coral (<i>Corallium</i>), so largely +sought after as an ornamental material, appears for the first +time in deposits of this age. Amongst the <i>Echinoderms</i>, +we meet with Heart-Urchins (<i>Spatangus</i>), Cake-Urchins +(<i>Scutella</i>; fig. 238), and various other forms, the majority +of which are closely allied to forms now in existence. +</p> + +<p class="indent"> +Numerous Crabs and Lobsters represent the <i>Crustacea</i>; but +the most important of the Miocene Articulate Animals are the +<i>Insects</i>. Of these, more than thirteen hundred species +have been determined by Dr Heer from the Miocene strata of +Switzerland alone. They include almost all the existing orders +of insects, such as numerous and varied forms of Beetles +(<i>Coleoptera</i>), Forest-bugs (<i>Hemiptera</i>), Ants +(<i>Hymenoptera</i>), Flies (<i>Diptera</i>), Termites and +Dragon-flies (<i>Neuroptera</i>), Grasshoppers (<i>Orthoptera</i>), +and Butterflies (<i>Lepidoptera</i>). +<a name="page_312"><span class="page">Page 312</span></a> +One of the latter, +the well-known <i>Vanessa Pluto</i> of the Brown Coals of Croatia, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 528px; + font-size: smaller; text-align: center;"> +<img src="images/fig238.jpg" width="528" height="158" alt="Fig. 238" /> +<br /> +Fig. 238.—Different views of <i>Scutella subrotunda</i>, +a Miocene "Cake-Urchin" from the south of France. +</span> +</span> + +even exhibits the pattern of the wing, and to some extent its +original coloration; whilst the more durably-constructed insects +are often in a state of exquisite preservation. +</p> + +<p class="indent"> +The <i>Mollusca</i> of the Miocene period are very numerous, +but call for little special comment. Upon the whole, they are +generically very similar to the Shell-fish of the present day; +whilst, as before stated, from fifteen to thirty per cent of +the <i>species</i> are identical with those now in existence. +So far as the European area is concerned, the Molluscs indicate +a decidedly hotter climate than the present one, though they +have not such a distinctly tropical character as is the case +with the Eocene shells. Thus we meet with many Cones, Volutes, +Cowries, Olive-shells, Fig-shells, and the like, which are +decidedly indicative of a high temperature of the sea. +<i>Polyzoans</i> are abundant, and often attain considerable +dimensions; whilst <i>Brachiopods</i>, on the other hand, are +few in number. Bivalves and <i>Univalves</i> are extremely +plentiful; and we meet here with the shells of Winged-Snails +(<i>Pteropods</i>), belonging to such existing genera as +<i>Hyalea</i> (fig. 239) and <i>Cleodora</i>. Lastly, the + +<span style="float: left; margin: 4px; width: 245px; + font-size: smaller; text-align: justify;"> +<img src="images/fig239.jpg" width="245" height="79" alt="Fig. 239" /> +<br /> +Fig. 239.—Different views of the shell of <i>Hyalea +Orbignyana</i>, a Miocene Pteropod. +</span> + +<i>Cephalopods</i> are represented both by the chambered shells +of <i>Nautili</i> and by the internal skeletons of Cuttle-fishes +(<i>Spirulirostra</i>.) +</p> + +<p class="indent"> +The <i>Fishes</i> of the Miocene Period are very abundant but of +little special importance. Besides the remains of Bony Fishes, we +meet in the marine deposits of this age with numerous pointed +teeth belonging to different kinds of Sharks. Some of the genera of +these—such as <i>Carcharodon</i> (fig. 241), <i>Oxyrhina</i> +(fig. 240), <i>Lamna</i>, and <i>Galeocerdo</i>—are very +widely distributed, ranging through +<a name="page_313"><span class="page">Page 313</span></a> +both the Old and New Worlds; and some of the species attain gigantic +dimensions. +</p> + +<table border="0" class="right"> +<tr><td align="center" valign="bottom"> + +<table border="0" width="121"> +<tr><td> +<img src="images/fig240.jpg" width="117" height="189" alt="Fig. 240" /> +</td></tr><tr><td class="left"><span class="image"> +Fig. 240.—Tooth of <i>Oxyrhina xiphodon</i>. Miocene. +</span></td></tr> +</table> + +</td><td align="center" valign="bottom"> + +<table border="0" width="119"> +<tr><td> +<img src="images/fig241.jpg" width="115" height="127" alt="Fig. 241" /> +</td></tr><tr><td class="left"><span class="image"> +Fig. 241.—Tooth of <i>Carcharodon productus</i>. Miocene. +</span></td></tr> +</table> + +</td></tr> +</table> + +<p class="indent"> +Amongst the <i>Amphibians</i> we meet with distinctly modern +types, such as Frogs (<i>Rana</i>) and Newts or Salamanders. +The most celebrated of the latter is the famous <i>Andrias +Scheuchzeri</i> (fig. 242), discovered in the year 1725 in the +fresh-water Miocene deposits of Ĺ’ningen, in Switzerland. +The skeleton indicates an animal nearly five feet in length; and +it was originally described by Scheuchzer, a Swiss physician, in +a dissertation published in 1731, as the remains of one of the +human beings who were in existence at the time of the Noachian +Deluge. Hence he applied to it the name of <i>Homo diluvii +testis</i>. In reality, however, as shown by Cuvier, we have here +the skeleton of a huge Newt, very closely allied to the Giant +Salamander (<i>Menopoma maxima</i>) of Java. +</p> + +<p class="indent"> +The remains of <i>Reptiles</i> are far from uncommon in the Miocene +rocks, consisting principally of Chelonians and Crocodilians. The +Land-tortoises (<i>TestudinidĹ“</i>) make their first appearance +during this period. The most remarkable form of this group is the +huge <i>Colossochelys Atlas</i> of the Upper Miocene deposits of +the Siwâlik Hills in India, described by Dr Falconer and Sir +Proby Cautley. Far exceeding any living Tortoise in its dimensions, +this enormous animal is estimated as having had a length of about +twenty feet, measured from the tip of the snout to the extremity +of the tail, and to have stood upwards of seven feet high. All the +details of its organisation, however, prove that it must have been +"strictly a land animal, with herbivorous habits, and probably of +the most inoffensive nature." The accomplished palæontologist +just quoted, shows further that some of the traditions of the +Hindoos would render it not improbable that this colossal Tortoise +had survived into the earlier portion of the human period. +</p> + +<p class="indent"> +Of the <i>Birds</i> of the Miocene period it is sufficient to +remark that though specifically distinct, they belong, so far +as known, wholly to existing groups, and therefore present no +points of special palæontological interest. +</p> + +<p class="indent"> +The <i>Mammals</i> of the Miocene are very numerous, and only +<a name="page_314"><span class="page">Page 314</span></a> + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 462px; + font-size: smaller; text-align: center;"> +<img src="images/fig242.jpg" width="462" height="858" alt="Fig. 242" /> +<br /> +Fig. 242.—Front portion of the skeleton of <i>Andrias +Scheuchzeri</i>, a Giant Salamander from the Miocene Tertiary of +Ĺ’ningen, in Switzerland. Reduced in size. +</span> +</span> + +<a name="page_315"> +the more important forms can be here alluded to. Amongst the</a> +<i>Marsupials</i>, the Old World still continued to possess species +<span class="page">Page 315</span> +of Opossum (<i>Didephys</i>), allied to the existing American +forms. The <i>Edentates</i> (Sloths, Armadillos, and Ant-eaters), +at the present day mainly South American, are represented by two +large European forms. One of these is the large <i>Macrotherium +giganteum</i> of the Upper Miocene of Gers in Southern France, +which appears to hare been in many respects allied to the existing +Scaly Ant-eaters or Pangolins, at the same time that the +disproportionately long fore-limbs would indicate that it possessed +the climbing habits of the Sloths. The other is the still more +gigantic <i>Ancylotherium Pentelici</i> of the Upper Miocene of +PikermĂ©, which seems to have been as large as, or larger than, +the Rhinoceros, and which must have been terrestrial in its habits. +This conclusion is further borne out by the comparative equality +of length which subsists between the fore and hind limbs, and is +not affected by the curvature and crookedness of the claws, this +latter feature being well marked in such existing terrestrial +Edentates as the Great Ant-eater. +</p> + +<p class="indent"> +The aquatic <i>Sirenians</i> and <i>Cetaceans</i> are represented +in Miocene times by various forms of no special importance. Amongst +the former, the previously existing genus <i>Halitherium</i> +continued to survive, and amongst the latter we meet with remains +of Dolphins and of Whales of the "Zeuglodont" family. We may +also note here the first appearance of true "Whalebone Whales," +two species of which, resembling the living "Right Whale" of +Arctic seas, and belonging to the same genus (<i>BalĹ“na</i>), +have been detected in the Miocene beds of North America. +</p> + +<p class="indent"> +The great order of the <i>Ungulates</i> or Hoofed Quadrupeds +is very largely developed in strata of Miocene age, various new +types of this group making their appearance here for the first +time, whilst some of the characteristic genera of the preceding +period are still represented under new shapes. Amongst the Odd-toed +or "Perissodactyle" Ungulates, we meet for the first time with +representatives of the family <i>RhinoceridĹ“</i> comprising +only the existing Rhinoceroses. In India in the Upper Miocene beds +of the Siwâlik Hills, and in North America, several species +of Rhinoceros have been detected, agreeing with the existing forms +in possessing three toes to each foot, and in having one or two +solid fibrous "horns" carried upon the front of the head. On the +other hand, the forms of this group which distinguish the Miocene +deposits of Europe appear to have been for the most part hornless, +and to have resembled the Tapirs in having three-toed hind-feet, +but four-toed fore-feet. +<a name="page_316"><span class="page">Page 316</span></a> +The family of the Tapirs is represented, both in the Old and New +Worlds, by species of the genus <i>Lophiodon</i>, some of which +were quite diminutive in point of size, whilst others attained +the dimensions of a horse. Nearly allied to this family, also, +is the singular group of quadrupeds which Marsh has described +from the Miocene strata of the United States under the name of +<i>BrontotheridĹ“</i>. These extraordinary animals, typified +by <i>Brontotherium</i> (fig. 243) itself, agree with the existing + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 457px; + font-size: smaller; text-align: center;"> +<img src="images/fig243.jpg" width="457" height="238" alt="Fig. 243" /> +<br /> +Fig. 243.—Skull of <i>Brontotherium ingens</i>. Miocene +Tertiary, United States. (After Marsh.) +</span> +</span> + +Tapirs of South America and the Indian Archipelago in having the +fore-feet four-toed, whilst the hind-feet are three-toed; and +a further point of resemblance is found in the fact (as shown by +the form of the nasal bones) that the nose was long and flexible, +forming a short movable proboscis or trunk, by means of which the +animal was enabled to browse on shrubs or trees. They differ, +however, from the Tapirs, not only in the apparent presence of +a long tail, but also in the possession of a pair of very large +"horn-cores," carried upon the nasal bones, indicating that the +animal possessed horns of a similar structure to those of the +"Hollow-horned" Ruminants (<i>e.g.</i>, Sheep and Oxen). +<i>Brontotherium gigas</i> is said to be nearly as large as an +Elephant, whilst <i>B. Ingens</i> appears to have attained dimensions +still more gigantic. The well-known genus <i>Titanotherium</i> of +the American Miocene would also appear to belong to this group. +</p> + +<p class="indent"> +The family of the Horses (<i>EquidĹ“</i>) appears under +various forms in the Miocene, but the most important and best +known of these is <i>Hipparion</i>. In this genus the general +conformation of the skeleton is extremely similar to that of the +existing Horses, and the external appearance of the animal must +have been very much the same. The foot of <i>Hipparion</i>, +<a name="page_317"><span class="page">Page 317</span></a> +however, as has been previously mentioned, +differed from that of the Horse in the fact that whilst both +possess the middle toe greatly developed and enclosed in a +broad hoof, the former, in addition, possessed two lateral +toes, which were sufficiently developed to carry hoofs, but +were so far rudimentary that they hung idly by the side of the +central toe without touching the ground (see fig. 230). In the +Horse, on the other hand, these lateral toes, though present, +are not only functionally useless, but are concealed beneath +the skin. Remains of the <i>Hipparion</i> have been found in +various regions in Europe and in India; and from the immense +quantities of their bones found in certain localities, it may +be safely inferred that these Middle Tertiary ancestors of the +Horses lived, like their modern representatives, in great +herds, and in open grassy plains or prairies. +</p> + +<p class="indent"> +Amongst the Even-toed or <i>Artiodactyle</i> Ungulates, we for +the first time meet with examples of the <i>Hippopotamus</i>, with +its four-toed feet, its massive body, and huge tusk-like lower +canine teeth. The Miocene deposits of Europe have not hitherto +yielded any remains of <i>Hippopotamus</i>; but several species +have been detected in the Upper Miocene of the Siwâlik Hills +by Dr Falconer and Sir Proby Cautley. These ancient Indian forms, +however, differ from the existing <i>Hippopotamus amphibius</i> +of Africa in the fact that they possessed six incisor teeth in +each jaw (fig. 244), whereas the latter has only four. +</p> + +<p class="indent"> +Amongst the other Even-toed Ungulates, the family of the Pigs +(<i>Suida</i>) is represented by true Swine (<i>Sus Erymanthius</i>), +Peccaries (<i>Dicotyles antiquus</i>), and by forms which, like +the great <i>Elotherium</i> of the American Miocene, have no +representative at the present day. The Upper Miocene of India has +yielded examples of the Camels. Small Musk-deer (<i>Amphitragulus</i> +and <i>Dremotherium</i>) are known to have existed in France and +Greece; and the true Deer (<i>CervidĹ“</i>), with their solid +bony antlers, appear for the first time here in the person of +species allied to the living Stags (<i>Cervus</i>), accompanied +by the extinct genus <i>Dorcatherium</i>. The Giraffes +(<i>CamelopardalidĹ“</i>), now confined to Africa, are known to +have lived in India and Greece; and the allied <i>Helladotherium</i>, +in some respects intermediate between the Giraffes and the Antelopes, +ranged over Southern Europe from Attica to France. The great group +of the "Hollow-horned" Ruminants (<i>Cavicornia</i>), lastly, +came into existence in the Miocene period; and though the typical +families of the Sheep and Oxen are apparently wanting, there are +true Antelopes, together with forms which, if systematically +referable to the <i>AntilopidĹ“</i>, nevertheless are more or +less clearly transitional between this and the family of the Sheep +<a name="page_318"><span class="page">Page 318</span></a> +and Goats. Thus the <i>PalĹ“oreas</i> of +the Upper Miocene of Greece may be regarded as a genuine Antelope; +but the <i>Tragoceras</i> of the same deposit is intermediate in + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 521px; + font-size: smaller; text-align: justify;"> +<img src="images/fig244.jpg" width="521" height="447" alt="Fig. 244" /> +<br /> +Fig. 244.—<i>a</i>, Skull of <i>Hippopotamus Sivalensis</i>, +viewed from below, one-eighth of the natural size; <i>b</i>, Molar +tooth of the same, showing the surface of the crown, one-half of +the natural size: <i>c</i>, Front of the lower jaw of the same, +showing the six incisors and the tusk-like canines, one-eighth of +the natural size. Upper Miocene, Siwâlik Hills; (After +Falconer and Cautley.) +</span> +</span> + +its characters between the typical Antelopes and the Goats. Perhaps +the most remarkable, however, of these Miocene Ruminants is the +<i>Sivatherium giganteum</i> (fig. 245) of the Siwâlik Hills, +in India. In this extraordinary animal there were two pairs of horns, +supported by bony "horn-cores," so that there can be no hesitation +in referring <i>Sivatherium</i> to the Cavicorn Ruminants. If all +these horns had been simple, there would have been no difficulty +in considering <i>Sivatherium</i> as simply a gigantic four-horned +Antelope, essentially similar to the living <i>Antilope</i> +(<i>Tetraceros</i>) <i>quadricornis</i> of India. The hinder pair +of horns, however, is not only much larger than the front pair, +but each possesses two branches or snags—a peculiarity +not to be paralleled amongst any existing Antelope, save the +abnormal Prongbuck (<i>Antilocapra</i>) of North America. Dr +Murie, however, in an admirable memoir on the structure and +relationships +<a name="page_319"><span class="page">Page 319</span></a> +of <i>Sivatherium</i>, has drawn attention +to the fact that the Prongbuck sheds the <i>sheath</i> of its + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 477px; + font-size: smaller; text-align: center;"> +<img src="images/fig245.jpg" width="477" height="442" alt="Fig. 245" /> +<br /> +Fig. 245.—Skull of <i>Sivatherium giganteum</i>, reduced +in size. Miocene, India. (After Murie.) +</span> +</span> + +horns annually, and has suggested that this may also have been the +case with the extinct form. This conjecture is rendered probable, +amongst other reasons, by the fact that no traces of a horny +sheath surrounding the horn-cores of the Indian fossil have been +as yet detected. Upon the whole, therefore, we may regard the +elephantine <i>Sivatherium</i> as being most nearly allied to +the Prongbuck of Western America, and thus as belonging to the +family of the Antelopes. +</p> + +<p class="indent"> +It is to the Miocene period, again, to which we must refer the +first appearance of the important order of the Elephants and their +allies (<i>Proboscideans</i>), all of which are characterised by +their elongated trunk-like noses, the possession of five toes to +the foot, the absence of canine teeth, the development of two or +more of the incisor teeth into long tusks, and the adaptation of +the molar teeth to a vegetable diet. Only three generic groups of +this order are known-namely, the extinct <i>Deinotherium</i>, the +equally extinct <i>Mastodons</i>, and the <i>Elephants</i>; and all +these three types are known to have been in existence as +<a name="page_320"><span class="page">Page 320</span></a> +early as the Miocene period, the first of them being exclusively +confined to deposits of this age. Of the three, the genus +<i>Deinotherium</i> is much the most abnormal in its characters; so +much so, that good authorities regard it as really being one of the +Sea-cows (<i>Sirenia</i>)—though this view has been rendered +untenable by the discovery of limb-bones which can hardly belong +to any other animal, and which are distinctly Proboscidean in +type. The most celebrated skull of the Deinothere (fig. 246) is one + +<span style="float: left; margin: 4px; width: 232px; + font-size: smaller; text-align: justify;"> +<img src="images/fig246.jpg" width="232" height="263" alt="Fig. 246" /> +<br /> +Fig. 246.—Skull of <i>Deinotherium giganteum</i>, greatly +reduced. From the Upper Micene of Germany. +</span> + +which was exhumed from the Upper Miocene deposits of Epplesheim, +in Hesse-Darmstadt, in the year 1836. This skull was four and a +half feet in length, and indicated an animal larger than any +existing species of Elephant. The upper jaw is destitute of incisor +or canine teeth, but is furnished on each side with five molars, +which are opposed to a corresponding series of grinding teeth in +the lower jaw. No canines are present in the lower jaw; but the +front portion of the jaw is abruptly bent downwards, and carries +two huge tusk-like incisor teeth, which are curved downwards and +backwards, and the use of which is rather problematical. Not +only does the Deinothere occur in Europe, but remains belonging +to this genus have also been detected in the Siwâlik Hills, in +India. +</p> + +<p class="indent"> +The true Elephants (<i>Elephas</i>) do not appear to have existed +during the Miocene period in Europe, but several species have been +detected in the Upper Miocene deposits of the Siwâlik Hills, +in India. The fossil forms, though in all cases specifically, and +in some cases even sub-generically, distinct, agree with those +now in existence in the general conformation of their skeleton, +and in the principal characters of their dentition. In all, the +canine teeth are wanting in both jaws; and there are no incisor +teeth in the lower jaw, whilst there are two incisors in the +front of the upper jaw, which are developed into two huge "tusks." +There are six molar teeth on each side of both the upper and lower +jaw, but only one, or at most a part of two, is in actual use +at any given time; and as this becomes worn away, it is pushed +forward and replaced by its successor behind it. The molars are of +<a name="page_321"><span class="page">Page 321</span></a> +very large size, and are each composed of a number of transverse + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 479px; + font-size: smaller; text-align: justify;"> +<img src="images/fig247.jpg" width="479" height="534" alt="Fig. 247" /> +<br /> +Fig. 247.—A, Molar tooth of <i>Elephas planifrons</i>, +one-third of the natural size, showing the grinding +surface—from the Upper Miocene of India; B, Profile view +of the last upper molar of <i>Mastodon Sivalensis</i>, one-third +of the natural size—from the Upper Miocene of India. (After +Falconer.) +</span> +</span> + +plates of enamel united together by ivory; and by the process +of mastication, the teeth become worn down to a flat surface, +crossed by the enamel-ridges in varying patterns; These patterns +are different in the different species of Elephants, though constant +for each; and they constitute one of the most readily available +means of separating the fossil forms from one another. Of the +seven Miocene Elephants of India, as judged by the characters of +the molar, teeth, two are allied to the existing Indian Elephant, +one is related to the living African Elephant, and the remaining +four are in some respects intermediate between the true Elephants +and the Mastodons. +</p> + +<p class="indent"> +The <i>Mastodons</i>, lastly, though quite elephantine in their +<a name="page_322"><span class="page">Page 322</span></a> +general characters, possess molar teeth +which have their crowns furnished with conical eminences or +tubercles placed in pairs (fig. 247, B), instead of having the +approximately flat surface characteristic of the grinders of the +Elephants. As in the latter, there are two upper incisor teeth, +which grow permanently during the life of the animal, and which +constitute great tusks; but the Mastodons, in addition, often +possess two lower incisors, which in some cases likewise grow +into small tusks. Three species of <i>Mastodon</i> are known to +occur in the Upper Miocene of the Siwâlik Hills of India; +and the Miocene deposits of the European area have yielded the +remains of four species, of which the best known are the <i>M. +Longirostris</i> and the <i>M. Angustidens</i>. +</p> + +<p class="indent"> +Whilst herbivorous Quadrupeds, as we have seen, were extremely +abundant during Miocene times, and often attained gigantic +dimensions, Beasts of Prey (<i>Carnivora</i>) were by no means +wanting, most of the principal existing families of the order +being represented in deposits of this age. Thus, we find aquatic +Carnivores belonging to both the living groups of the Seals and +Walruses; true Bears are wanting, but their place is filled by the +closely-allied genus <i>Amphicyon</i>, of which various species +are known; Weasels and Otters were not unknown, and the +<i>HyĹ“nictis</i> and <i>Iditherium</i> of the Upper Miocene +of Greece are apparently intermediate between the Civet-cats and +the Hyænas; whilst the great Cats of subsequent periods are +more than adequately represented by the huge "Sabre-toothed Tiger" +(<i>Machairodus</i>), with its immense trenchant and serrated +canine teeth. +</p> + +<p class="indent"> +Amongst the <i>Rodent</i> Mammals, the Miocene rocks have yielded +remains of Rabbits, Porcupines (such as the <i>Hystrix +primigenius</i> of Greece), Beavers, Mice, Jerboas, Squirrels, +and Marmots. All the principal living groups of this order were +therefore differentiated in Middle Tertiary times. +</p> + +<p class="indent"> +The <i>Cheiroptera</i> are represented by small insect-eating +Bats; and the order of the Insectivorous Mammals is represented +by Moles, Shrew-mice, and Hedgehogs. +</p> + +<p class="indent"> +Lastly, the Monkeys (<i>Quadrumana</i>) appear to have existed +during the Miocene period under a variety of forms, remains of +these animals having been found both in Europe and in India; but +no member of this order has as yet been detected in the Miocene +Tertiary of the North American continent. Amongst the Old World +Monkeys of the Miocene, the two most interesting are the +<i>Pliopithecus</i> and <i>Dryopithecus</i> of France. The former +of these (fig. 248) is supposed to have been most nearly related +to the living <i>Semnopitheci</i> of Southern Asia, in +<a name="page_323"><span class="page">Page 323</span></a> +which case it must have possessed a long tail. The <i>Mesopithecus</i> +of the Upper Miocene of Greece is also one of the lower Monkeys, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 334px; + font-size: smaller; text-align: center;"> +<img src="images/fig248.jpg" width="334" height="268" alt="Fig. 248" /> +<br /> +Fig. 248.—Lower jaw of <i>Pliopithcus antiquus</i>. Upper +Miocene, France. +</span> +</span> + +as it is most closely allied to the existing Macaques. On the +other hand, the <i>Dryopithecus</i> of the French Upper Miocene +is referable to the group of the "Anthropoid Apes," and is most +nearly related to the Gibbons of the present day, in which the tail +is rudimentary and there are no cheek-pouches. <i>Dryopithecus</i> +was, also, of large size, equalling Man in stature, and apparently +living amongst the trees and feeding upon fruits. +</p> + +<h3>CHAPTER XX.</h3> + +<p class="subtitle"> +THE PLIOCENE PERIOD. +</p> + +<p class="indent"> +The highest division of the Tertiary deposits is termed the +<i>Pliocene</i> formation, in accordance with the classification +proposed by Sir Charles Lyell. The Pliocene formations contain +from 40 to 95 per cent of existing species of <i>Mollusca</i>, +the remainders belonging to extinct species. They are divided +by Sir Charles Lyell into two divisions, the Older Pliocene and +Newer Pliocene. +</p> + +<p class="indent"> +The Pliocene deposits of Britain occur in Suffolk, and are known +by the name of "Crags," this being a local term used for certain +shelly sands, which are employed in agriculture. Two of these +Crags are referable to the Older Pliocene, viz., +<a name="page_324"><span class="page">Page 324</span></a> +the White and Red Crags,—and one belongs to the Newer +Pliocene, viz., the Norwich Crag. +</p> + +<p class="indent"> +The <i>White or Coralline Crag</i> of Suffolk is the oldest of +the Pliocene deposits of Britain, and is an exceedingly local +formation, occurring in but a single small area, and having a +maximum thickness of not more than 50 feet. It consists of soft +sands, with occasional intercalations of flaggy limestone. Though +of small extent and thickness, the Coralline Crag is of importance +from the number of fossils which it contains. The name "Coralline" +is a misnomer; since there are few true Corals, and the so-called +"Corals" of the formation are really <i>Polyzoa</i>, often of +very singular forms. The shells of the Coralline Crag are mostly +such as inhabit the seas of temperate regions; but there occur +some forms usually looked upon as indicating a warm climate. +</p> + +<p class="indent"> +The <i>Upper</i> or <i>Red Crag</i> of Suffolk—like the +Coralline Crag—has a limited geographical extent and a +small thickness, rarely exceeding 40 feet. It consists of +quartzose sands, usually deep red or brown in colour, and +charged with numerous fossils. +</p> + +<p class="indent"> +Altogether more than 200 species of shells are known from the +Red Crag, of which 60 per cent are referable to existing species. +The shells indicate, upon the whole, a temperate or even cold +climate, decidedly less warm than that indicated by the organic +remains of the Coralline Crag. It appears, therefore, that a +gradual refrigeration was going on during the Pliocene period, +commencing in the Coralline Crag, becoming intensified in the Red +Crag, being still more severe in the Norwich Crag, and finally +culminating in the Arctic cold of the Glacial period. +</p> + +<p class="indent"> +Besides the <i>Mollusca</i>, the Red Crag contains the ear-bones of +Whales, the teeth of Sharks and Rays, and remains of the Mastodon, +Rhinoceros, and Tapir. +</p> + +<p class="indent"> +The <i>Newer Pliocene</i> deposits are represented in Britain +by the <i>Norwich Crag</i>, a local formation occurring near +Norwich. It consists of incoherent sands, loams, and gravels, +resting in detached patches, from 2 to 20 feet in thickness, +upon an eroded surface of Chalk. The Norwich Crag contains a +mixture of marine, land, and fresh-water shells, with remains of +fishes and bones of mammals; so that it must have been deposited +as a local sea-deposit near the mouth of an ancient river. It +contains altogether more than 100 marine shells, of which 89 +per cent belong to existing species. Of the Mammals, the two +most important are an Elephant (<i>Elephas meridionalis</i>), +and the characteristic Pliocene Mastodon +<a name="page_325"><span class="page">Page 325</span></a> +(<i>M. Arvernensis</i>), which is hitherto the only Mastodon +found in Britain. +</p> + +<p class="indent"> +According to the most recent views of high authorities, certain +deposits—such as the so-called "Bridlington Crag" of +Yorkshire, and the "Chillesford beds" of Suffolk—are to +be also included in the Newer Pliocene, upon the ground that +they contain a small proportion of extinct shells. Our knowledge, +however, of the existing Molluscan fauna, is still so far +incomplete, that it may reasonably be doubted if these supposed +extinct forms have actually made their final disappearance, +whilst the strata in question have a strong natural connection +with the "Glacial deposits," as shown by the number of Arctic +Mollusca which they contain. Here, therefore, these beds will +be included in the Post-Pliocene series, in spite of the fact +that some of their species of shells are not known to exist at +the present day. +</p> + +<p class="indent"> +The following are the more important Pliocene deposits which have +been hitherto recognised out of Britain:— +</p> + +<p class="indent"> +1. In the neighbourhood of Antwerp occur certain "crags," which +are the equivalent of the White and Red Crag in part. The lowest +of these contains less than 50 per cent, and the highest 60 per +cent, of existing species of shells, the remainder being extinct. +</p> + +<p class="indent"> +2. Bordering the chain of the Apennines, in Italy, on both sides +is a series of low hills made up of Tertiary strata, which are +known as the Sub-Apennine beds. Part of these is of Miocene age, +part is Older Pliocene, and a portion is Newer Pliocene. The +Older Pliocene portion of the Sub-Apennines consists of blue or +brown marls, which sometimes attain a thickness of 2000 feet. +</p> + +<p class="indent"> +3. In the valley of the Arno, above Florence, are both Older +and Newer Pliocene strata. The former consist of blue clays and +lignites, with an abundance of plants. The latter consist of sands +and conglomerates, with remains of large Carnivorous Mammals, +Mastodon, Elephant, Rhinoceros, Hippopotamus, &c. +</p> + +<p class="indent"> +4. In Sicily, Newer Pliocene strata are probably more largely +developed than anywhere else in the world, rising sometimes to a +height of 3000 feet above the sea. The series consists of clays, +marls, sands, and conglomerates, capped by a compact limestone, +which attains a thickness of from 700 to 800 feet. The fossils of +these beds belong almost entirely to living species, one of the +commonest being the Great Scallop of the Mediterranean (<i>Pecten +JacobĹ“us</i>). +</p> + +<p class="indent"> +5. Occupying an extensive area round the Caspian, Aral, +<a name="page_326"><span class="page">Page 326</span></a> +and Azof Seas, are Pliocene deposits known +as the "Aralo-Caspian" beds. The fossils in these beds are partly +freshwater, partly marine, and partly intermediate in character, +and they are in great part identical with species now inhabiting +the Caspian. The entire formation appears to indicate the former +existence of a great sheet of brackish water, forming an inland +sea, like the Caspian, but as large as, or larger than, the +Mediterranean. +</p> + +<p class="indent"> +6. In the United States, strata of Pliocene age are found in +North and South Carolina. They consist of sands and clays, with +numerous fossils, chiefly <i>Molluscs</i> and <i>Echinoderms</i>. +From 40 to 60 per cent of the fossils belong to existing species. +On the Loup Fork of the river Platte, in the Upper Missouri region, +are strata which are also believed to be referable to the Pliocene +period, and probably to its upper division. They are from 300 to +400 feet thick, and contain land-shells, with the bones of numerous +Mammals, such as Camels, Rhinoceroses, Mastodons, Elephants, the +Horse, Stag, &c. +</p> + +<p class="indent"> +As regards the <i>life</i> of the Pliocene period, there are +only two classes of organisms to which our attention need be +directed—namely, the Shell-fish and the Mammals. So far as +the former are concerned, we have to note in the first place that +the introduction of new species of animals upon the globe went +on rapidly during this period. In the Older Pliocene deposits, +the number of shells of existing species is only from 40 to 60 +per cent; but in the Newer Pliocene the proportion of living +forms rises to as much as from 80 to 95 per cent. Whilst the +Molluscs thus become rapidly modernised, the Mammals still all +belong to extinct species, though modern generic types gradually +supersede the more antiquated forms of the Miocene. In the second +place, there is good evidence to show that the Pliocene period +was one in which the climate of the northern hemisphere underwent +a gradual refrigeration. In the Miocene period, there is evidence +to show that Europe possessed a climate very similar to that +now enjoyed by the Southern United States, and certainly very +much warmer than it is at present. The presence of Palm-trees +upon the land, and of numerous large Cowries, Cones, and other +shells of warm regions in the sea, sufficiently proves this. In +the Older Pliocene deposits, on the other hand, northern forms +predominate amongst the Shells, though some of the types of hotter +regions still survive. In the Newer Pliocene, again, the Molluscs +are such as almost exclusively inhabit the seas of temperate +or even cold regions; whilst if we regard deposits like the +"Bridlington Crag" and +<a name="page_327"><span class="page">Page 327</span></a> +"Chillesford beds" +as truly referable to this period, we meet at the close of this +period with shells such as nowadays are distinctively +characteristic of high latitudes. It might be thought that the +occurrence of Quadrupeds such as the Elephant, Rhinoceros, and +Hippopotamus, would militate against this generalisation, and +would rather support the view that the climate of Europe and +the United States must have been a hot one during the later +portion of the Pliocene period. We have, however, reason to +believe that many of these extinct Mammals were more abundantly +furnished with hair, and more adapted to withstand a cool +temperature, than any of their living congeners. We have also +to recollect that many of these large herbivorous quadrupeds +may have been, and indeed probably were, more or less migratory +in their habits; and that whilst the winters of the later portion +of the Pliocene period were cold, the summers might have been +very hot. This would allow of a northward migration of such +terrestrial animals during the summer-time, when there would be +an ample supply of food and a suitably high temperature, and a +southward recession towards the approach of winter. +</p> + +<p class="indent"> +The chief palæontological interests of the Pliocene deposits, +as of the succeeding Post-Pliocene, centre round the Mammals of the +period; and amongst the many forms of these we may restrict our +attention to the orders of the Hoofed Quadrupeds (<i>Ungulates</i>), +the <i>Proboscideans</i>, the <i>Carnivora</i>, and the +<i>Quadrumana</i>. Almost all the other Mammalian orders are +more or less fully represented in Pliocene times, but none of them +attains any special interest till we enter upon the Post-Pliocene. +</p> + +<p class="indent"> +Amongst the Odd-toed Ungulates, in addition to the remains of +true Tapirs (<i>Tapirus Arvernensis</i>), we meet with the bones +of several species of Rhinoceros, of which the <i>Rhinoceros +Etruscus</i> and <i>R. Megarhinus</i> (fig. 249) are the most +important. The former of these (fig. 249, A) derives its specific +name from its abundance in the Pliocene deposits of the Val d'Arno, +near Florence, and though principally Pliocene in its distribution, +it survived into the earlier portion of the Post-Pliocene period. +<i>Rhinoceros Etruscus</i> agreed with the existing African forms +in having two horns placed one behind the other, the front one +being the longest; but it was comparatively slight and slender in +its build, whilst the nostrils were separated by an incomplete bony +partition. In the <i>Rhinoceros megarhinus</i> (fig. 249, B), on +the other hand, no such partition exists between the nostrils, and +the nasal bones are greatly developed in size. It was a two-horned +form, and is found associated with <i>Elephas meridionalis</i> and +<i>E. Antiquus</i> in the Pliocene deposits of the +<a name="page_328"><span class="page">Page 328</span></a> +Val d'Arno, near Florence. Like the preceding, it survived, in +diminished numbers, into the earlier portion of the Post-Pliocene +period. +</p> + +<p class="indent"> +The Horses (<i>EquidĹ“</i>) are represented, both in Europe + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 492px; + font-size: smaller; text-align: justify;"> +<img src="images/fig249.jpg" width="492" height="518" alt="Fig. 249" /> +<br /> +Fig. 249.—A. Under surface of the skull of <i>Rhinoceros +Etruscus</i>, one-seventh of the natural size—Pliocene, +Italy.; B, Crowns of the three true molars of the upper jaw, +left side, of <i>Rhinoceros megarhinus</i> (<i>R. Leptorhinus</i>, +Falconer), one-half of the natural size—Pliocene, France. +(After Falconer.) +</span> +</span> + +and America, by the three-toed Hipparions, which survive from the +Miocene, but are now verging upon extinction. For the first time, +also, we meet with genuine Horses (<i>Equus</i>), in which each +foot is provided with a single complete toe only, encased in a +single broad hoof. One of the American species of this period +(the <i>Equus excelsus</i>) quite equalled the modern Horse in +stature; and it is interesting to note the occurrence of indigenous +horses in America at such a comparatively late geological epoch, +seeing that this continent certainly possessed none of these +animals when first discovered by the Spaniards. +</p> + +<p class="indent"> +Amongst the Even-toed Ungulates, we may note the occurrence of +<a name="page_329"><span class="page">Page 329</span></a> +Swine (<i>Suida</i>), of forms allied to +the Camels (<i>CamelidĹ“</i>), and of various kinds of Deer +(<i>CervidĹ“</i>); but the most interesting Pliocene Mammal +belonging to this section is the great <i>Hippopotamus major</i> +of Britain and Europe. This well-known species is very closely +allied to the living <i>Hippopotamus amphibius</i> of Africa, +from which it is separated only by its larger dimensions, and by +certain points connected with the conformation of the skeleton. +It is found very abundantly in the Pliocene deposits of Italy +and France, associated with the remains of the Elephant, Mastodon, +and Rhinoceros, and it survived into the earlier portion of the +Post-Pliocene period. During this last-mentioned period, it +extended its range northwards, and is found associated with the +Reindeer, the Bison, and other northern animals. From this fact +it has been inferred, with great probability, that the +<i>Hippotamus major</i> was furnished with a long coat of hair +and fur, thus differing from its nearly hairless modern +representative, and resembling its associates, the Mammoth and +the Woolly Rhinoceros. +</p> + +<p class="indent"> +Passing on to the Pliocene Proboscideans, we find that the great +<i>Deinotheria</i> of the Miocene have now wholly disappeared, +and the sole representatives of the order are Mastodons and +Elephants. The most important member of the former group is the +<i>Mastodon Arvernensis</i> (fig. 250), which ranged widely over + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 436px; + font-size: smaller; text-align: center;"> +<img src="images/fig250.jpg" width="436" height="276" alt="Fig. 250" /> +<br /> +Fig. 250.—Third milk-molar of the left side of the upper +jaw of <i>Mastodon Arvernensis</i>, showing the grinding surface. +Pliocene. +</span> +</span> + +Southern Europe and England, being generally associated with +remains of the <i>Elephas meridionalis, E. antiquus, Rhinoceros +megarhinus</i>, and <i>Hippopotamus major</i>. The lower jaw +seems to have been destitute of incisor teeth; but the upper +incisors are developed into great tusks, which sometimes reach a +<a name="page_330"><span class="page">Page 330</span></a> +length of nine feet, and which have the simple curvature of the +tusks of the existing Elephants. Amongst the Pliocene Elephants +the two most important are the <i>Elephas meridionalis</i> and the +<i>Elephas antiquus</i>. Of these, the <i>Elephas meridionalis</i> +(fig. 251) is found abundantly in the Pliocene deposits of Southern + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 472px; + font-size: smaller; text-align: center;"> +<img src="images/fig251.jpg" width="472" height="230" alt="Fig. 251" /> +<br /> +Fig. 251.—Molar tooth of <i>Elephas meridionalis</i>, +one-third of the natural size. Pliocene and Post-Pliocene. +</span> +</span> + +Europe and England, and also survived into the earlier portion +of the Post-Pliocene period. Its molar teeth are of the type of +those of the existing African Elephant, the spaces enclosed by the +transverse enamel-plates being more or less lozenge-shaped, whilst +the curvature of the tusks is simple. The <i>Elephas antiquus</i> +(fig. 252) is very generally associated with the preceding, and + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 421px; + font-size: smaller; text-align: center;"> +<img src="images/fig252.jpg" width="421" height="148" alt="Fig. 252" /> +<br /> +Fig. 252.—Molar tooth of <i>Elephas antiquus</i>, one-third +of the natural size. Pliocene and Post-Pliocene. +</span> +</span> + +it survived to an even later stage of the Post-Pliocene period. +The molar teeth are of the type of the existing Indian Elephant, +with comparatively thin enamel-ridges, placed closer together +than in the African type; whilst the tusks were nearly straight. +</p> + +<p class="indent"> +Amongst the Pliocene <i>Carnivores</i>, we meet with true Bears +(<i>Ursus Arvernensis</i>), Hyænas (such as <i>HyĹ“na +Hipparionum</i>), and genuine Lions (such as the <i>Felis +angustus</i> of North America); but the most remarkable of the +beasts of prey of +<a name="page_331"><span class="page">Page 331</span></a> +this period is the great "Sabre-toothed Tiger" (<i>Machairodus</i>), +species of which existed in the earlier Miocene, +and survived to the later Post-Pliocene. In this remarkable +form we are presented with perhaps the most highly carnivorous +type of all known beasts of prey. Not only are the jaws shorter +in proportion even than those of the great Cats of the present +day, but the canine teeth (fig. 253) are of enormous size, greatly + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 538px; + font-size: smaller; text-align: center;"> +<img src="images/fig253.jpg" width="538" height="378" alt="Fig. 253" /> +<br /> +Fig. 253.—A, Skull of <i>Machairodus cultridens</i>, +without the lower jaw, reduced in size; B, Canine tooth of the +same, one-half the natural size. Pliocene, France. +</span> +</span> + +flattened so as to assume the form of a poignard, and having +their margins finely serrated. A part from the characters of the +skull, the remainder of the skeleton, so far as known, exhibits +proofs that the Sabre-toothed Tiger was extraordinarily muscular +and powerful, and in the highest degree adapted for a life of +rapine. Species of <i>Machairodus</i> must have been as large +as the existing Lion; and the genus is not only European, but +is represented both in South America and in India, so that the +geographical range of these predaceous beasts must have been +very extensive. +</p> + +<p class="indent"> +Lastly, we may note that the Pliocene deposits of Europe have +yielded the remains of Monkeys (<i>Quadrumana</i>), allied to +the existing <i>Semnopitheci</i> and Macaques. +</p> + +<h4> +<a name="page_332"><span class="page">Page 332</span></a> +LITERATURE.</h4> + +<p class="indent"> +The following list comprises a small selection of some of the +more important and readily accessible works and memoirs relating +to the Tertiary rocks and their fossils. With few exceptions, +foreign works relating to the Tertiary strata of the continent +of Europe or their organic remains have been omitted:— +</p> + +<table border="0" cellspacing="0"> +<tr><td class="right" valign="top">(1)</td> + <td>'Elements of Geology.' Lyell.</td> +</tr><tr><td class="right" valign="top">(2)</td> + <td>'Students' Elements of Geology.' Lyell.</td> +</tr><tr><td class="right" valign="top">(3)</td> + <td>'Manual of Palæontology.' Owen.</td> +</tr><tr><td class="right" valign="top">(4)</td> + <td>'British Fossil Mammals and Birds.' Owen.</td> +</tr><tr><td class="right" valign="top">(5)</td> + <td>'TraitĂ© de PalĂ©ontologie.' Pictet.</td> +</tr><tr><td class="right" valign="top">(6)</td> + <td>'Cours ElĂ©mentaire de PalĂ©ontologie.' D'Orbigny.</td> +</tr><tr><td class="right" valign="top">(7)</td> + <td>"Probable Age of the London Clay," &c.—'Quart. + Journ. Geol. Soc.,' vol. iii. Prestwich.</td> +</tr><tr><td class="right" valign="top">(8)</td> + <td>'Structure and Probable Age of the Bagshot + Sands'—Ibid., vol. iii. Prestwich.</td> +</tr><tr><td class="right" valign="top">(9)</td> + <td>'Tertiary Formations of the Isle of Wight'—Ibid., + vol. ii. Prestwich.</td> +</tr><tr><td class="right" valign="top">(10)</td> + <td>'Structure of the Strata between the London Clay and the + Chalk,' &c.—Ibid., vols. vi., viii., and x. + Prestwich.</td> +</tr><tr><td class="right" valign="top">(11)</td> + <td>'Correlation of the Eocene Tertiaries of England, France, + and Belgium'—Ibid., vol. xxvii. Prestwich.</td> +</tr><tr><td class="right" valign="top">(12)</td> + <td>'On the Fluvio-marine Formations of the Isle of + Wight'—Ibid., vol. ix. Edward Forbes.</td> +</tr><tr><td class="right" valign="top">(13)</td> + <td>'Newer Tertiary Deposits of the Sussex Coast'—Ibid., + vol. xiii. Godwin-Austen.</td> +</tr><tr><td class="right" valign="top">(14)</td> + <td>'Kainozoic Formations of Belgium'—Ibid., vol. xxii. + Godwin-Austen.</td> +</tr><tr><td class="right" valign="top">(15)</td> + <td>'Tertiary Strata of Belgium and French + Flanders'—Ibid., vol. viii. Lyell.</td> +</tr><tr><td class="right" valign="top">(16)</td> + <td>'On Tertiary Leaf-beds in the Isle of Mull'—Ibid., + vol. vii. The Duke of Argyll.</td> +</tr><tr><td class="right" valign="top">(17)</td> + <td>'Newer Tertiaries of Suffolk and their Fauna'—Ibid., + vol. xxvi. Ray Lankester.</td> +</tr><tr><td class="right" valign="top">(18)</td> + <td>'Lower London Tertiaries of Kent'—Ibid., vol. xxii. + Whitaker.</td> +</tr><tr><td class="right" valign="top">(19)</td> + <td>"Guide to the Geology of London"—'Mem. Geol. Survey.' + Whitaker.</td> +</tr><tr><td class="right" valign="top">(20)</td> + <td>'Memoirs of the Geological Survey of Great Britain.'</td> +</tr><tr><td class="right" valign="top">(21)</td> + <td>'Introductory Outline of the Geology of the Crag District' + (Supplement to Crag Mollusca, Palæontographical + Society). S. V. Wood, jun., and F. w. Harmer.</td> +</tr><tr><td class="right" valign="top">(22)</td> + <td>"Tertiary Fluvio-marine Deposits of the Isle of Wight." + Edward Forbes. Edited by Godwin-Austen; with Descriptions of + the Fossils by Morris, Salter, and Rupert Jones—'Memoirs + of the Geological Survey.'</td> +</tr><tr><td class="right" valign="top">(23)</td> + <td>'Geological Excursions round the Isle of Wight.' + Mantell.</td> +</tr><tr><td class="right" valign="top">(24)</td> + <td>'Catalogue of British Fossils.' Morris.</td> +</tr><tr><td class="right" valign="top">(25)</td> + <td>'Catalogue of Fossils in the Museum of Practical Geology.' + Etheridge.</td> +</tr><tr><td class="right" valign="top">(26)</td> + <td>'Monograph of the Crag Polyzoa' (Palæontographical + Society). Busk.</td> +</tr><tr><td class="right" valign="top">(27)</td> + <td>'Monograph of the Tertiary Brachiopoda' (Ibid.) + Davidson.</td> +</tr><tr><td class="right" valign="top">(28)</td> + <td>'Monograph of the Tertiary Malacostracous Crustacea' + (Ibid.) Bell.</td> +</tr><tr><td class="right" valign="top"> +<a name="page_333"><span class="page">Page 333</span></a> + (29)</td> + <td>'Monograph of the Tertiary Corals' (Ibid.) Milne-Edwards + and Haime.</td> +</tr><tr><td class="right" valign="top">(30)</td> + <td>'Supplement to the Tertiary Corals' (Ibid.) Martin + Duncan.</td> +</tr><tr><td class="right" valign="top">(31)</td> + <td>'Monograph of the Eocene Mollusca' (Ibid.) Fred. E. + Edwards.</td> +</tr><tr><td class="right" valign="top">(32)</td> + <td>'Monograph of the Eocene Mollusca' (Ibid.) Searles V. + Wood.</td> +</tr><tr><td class="right" valign="top">(33)</td> + <td>'Monograph of the Crag Mollusca' (Ibid.) Searles V. + Wood.</td> +</tr><tr><td class="right" valign="top">(34)</td> + <td>'Monograph of the Tertiary Entomostraca' (Ibid.) Rupert + Jones.</td> +</tr><tr><td class="right" valign="top">(35)</td> + <td>'Monograph of the Foraminifera of the Crag' (Ibid.) Rupert + Jones, Parker, and H. B. Brady.</td> +</tr><tr><td class="right" valign="top">(36)</td> + <td>'Monograph of the Radiaria of the London Clay' (Ibid.) + Edward Forbes.</td> +</tr><tr><td class="right" valign="top">(37)</td> + <td>'Monograph of the Cetacea of the Red Crag' (Ibid.) + Owen.</td> +</tr><tr><td class="right" valign="top">(38)</td> + <td>'Monograph of the Fossil Reptiles of the London Clay' + (Ibid.) Owen and Bell.</td> +</tr><tr><td class="right" valign="top">(39)</td> + <td>"On the Skull of a Dentigerous Bird from the London Clay + of Sheppey"—'Quart. Journ. Geol. Soc.,' vol. xxix. + Owen.</td> +</tr><tr><td class="right" valign="top">(40)</td> + <td>'Ossemens Fossiles.' Cuvier.</td> +</tr><tr><td class="right" valign="top">(41)</td> + <td>'Fauna Antiqua Sivalensis.' Falconer and Sir Proby + Cautley.</td> +</tr><tr><td class="right" valign="top">(42)</td> + <td>'Palæontological Memoirs.' Falconer.</td> +</tr><tr><td class="right" valign="top">(43)</td> + <td>'Animaux Fossiles et GĂ©ologie de l'Attique.' + Gaudry.</td> +</tr><tr><td class="right" valign="top">(44)</td> + <td>"Principal Characters of the Dinocerata"—'American + Journ. of Science and Arts,' vol. xi. Marsh.</td> +</tr><tr><td class="right" valign="top">(45)</td> + <td>'Principal Characters of the Brontotheridæ' (Ibid.) + Marsh.</td> +</tr><tr><td class="right" valign="top">(46)</td> + <td>'Principal Characters of the Tillodontia' (Ibid.) + Marsh.</td> +</tr><tr><td class="right" valign="top">(47)</td> + <td>"Extinct Vertebrata of the Eocene of + Wyoming"—'Geological Survey of Montana,' &c., + 1872. Cope.</td> +</tr><tr><td class="right" valign="top">(48)</td> + <td>"Ancient Fauna of Nebraska"—'Smithsonian + Contributions to Knowledge,' vol. vi. Leidy.</td> +</tr><tr><td class="right" valign="top">(49)</td> + <td>'Manual of Geology.' Dana.</td> +</tr><tr><td class="right" valign="top">(50)</td> + <td>"Palæontology and Evolution" (Presidential Address + to the Geological Society of London, 1870)—'Quart. + Journ. Geol. Soc.,' vol. xxvi. Huxley.'</td> +</tr><tr><td class="right" valign="top">(51)</td> + <td>'Mineral Conchology.' Sowerby.</td> +</tr><tr><td class="right" valign="top">(52)</td> + <td>'Description des Coquilles Fossiles,' &c. + Deshayes.</td> +</tr><tr><td class="right" valign="top">(53)</td> + <td>'Description des Coquilles Tertiaires de Belgique.' + Nyst.</td> +</tr><tr><td class="right" valign="top">(54)</td> + <td>'Fossilen Polypen des Wiener Tertiär-beckens.' + Reuss.</td> +</tr><tr><td class="right" valign="top">(55)</td> + <td>'Palæontologische Studien über die älteren + Tertiär-schichten der Alpen.' Reuss.</td> +</tr><tr><td class="right" valign="top">(56)</td> + <td>'Land und Süss-wasser Conchylien der Vorwelt.' + Sandberger.</td> +</tr><tr><td class="right" valign="top">(57)</td> + <td>'Flora Tertiaria Helvetica.' Heer.</td> +</tr><tr><td class="right" valign="top">(58)</td> + <td>'Flora Fossilis Arctica.' Heer.</td> +</tr><tr><td class="right" valign="top">(59)</td> + <td>'Recherches sur le Climat et la VĂ©gĂ©tation + du Pays Tertiaire.' Heer.</td> +</tr><tr><td class="right" valign="top">(60)</td> + <td>'Fossil Flora of Great Britain.' Lindley and Hutton.</td> +</tr><tr><td class="right" valign="top">(61)</td> + <td>'Fossil Fruits and Seeds of the London Clay.' + Bowerbank.</td> +</tr><tr><td class="right" valign="top">(62)</td> + <td>"Tertiary Leaf-beds of the Isle of Mull"—'Quart. + Journ. Geol. Soc.,' vol. vii. Edward Forbes.</td> +</tr><tr><td class="right" valign="top">(63)</td> + <td>'The Geology of England and Wales.' Horace B. + Woodward.[25]</td></tr> +</table> + +<p class="footnote"> +[Footnote 25: This work—published whilst these sheets were +going through the press—gives to the student a detailed +view of all the strata of England and Wales, with their various +sub-divisions, from the base of the Palæozoic to the top +of the Tertiary.] +</p> + +<h3> +<a name="page_334"><span class="page">Page 334</span></a> +CHAPTER XXI.</h3> + +<p class="subtitle"> +THE QUATERNARY PERIOD. +</p> + +<p class="center"> +THE POST-PLIOCENE PERIOD. +</p> + +<p class="indent"> +Later than any of the Tertiary formations are various detached and +more or less superficial accumulations, which are generally spoken +of as the <i>Post-Tertiary formations</i>, in accordance with the +nomenclature of Sir Charles Lyell—or as the <i>Quaternary +formations</i>, in accordance with the general usage of Continental +geologists. In all these formations we meet with no <i>Mollusca</i> +except such as are now alive—with the partial and very limited +exception of some of the oldest deposits of this period, in which +a few of the shells occasionally belong to species not known to +be in existence at the present day. Whilst the <i>Shell-fish</i> +of the Quaternary deposits are, generally speaking, identical +with existing forms, the <i>Mammals</i> are sometimes referable +to living, sometimes to extinct species. In accordance with this, +the Quaternary formations are divided into two groups: (1) The +<i>Post-Pliocene</i>, in which the shells are almost invariably +referable to existing species, but some of the <i>Mammals are +extinct</i>; and (2) the <i>Recent</i>, in which <i>the shells +and the Mammals alike belong to existing species</i>. The +Post-Pliocene deposits are often spoken of as the Pleistocene +formations (Gr. <i>pleistos</i>, most; <i>kainos</i>, new or +recent), in allusion to the fact that the great majority of the +living beings of this period belong to the species characteristic +of the "new" or Recent period. +</p> + +<p class="indent"> +The <i>Recent</i> deposits, though of the highest possible interest, +do not properly concern the palæontologist strictly so-called, +but the zoologist, since they contain the remains of none but +existing animals. They are "Pre-historic," but they belong entirely +to the existing terrestrial order. The <i>Post-Pliocene</i> deposits, +on the other hand, contain the remains of various extinct Mammals; +and though Man undoubtedly existed in, at any rate, the later +portion of this period, if not throughout the whole of it, they +properly form part of the domain of the palæontologist. +</p> + +<p class="indent"> +The Post-Pliocene deposits are extremely varied, and very widely +distributed; and owing to the mode of their occurrence, the ordinary +geological tests of age are in their case but very partially +available. The subject of the classification of these deposits +<a name="page_335"><span class="page">Page 335</span></a> +is therefore an extremely complicated one; +and as regards the age of even some of the most important of them, +there still exists considerable difference of opinion. For our +present purpose, it will be convenient to adopt a classification +of the Post-Pliocene deposits founded on the relations which they +bear in time to the great "Ice-age" or "Glacial period;" though +it is not pretended that our present knowledge is sufficient to +render such a classification more than a provisional one. +</p> + +<p class="indent"> +In the early Tertiary period, as we have seen, the climate of the +northern hemisphere, as shown by the Eocene animals and plants, was +very much hotter than it is at present—partaking, indeed, of +a sub-tropical character. In the Middle Tertiary or Miocene period, +the temperature, though not so high, was still much warmer than +that now enjoyed by the northern hemisphere; and we know that the +plants of temperate regions at this time flourished within the +Arctic circle. In the later Tertiary or Pliocene period, again, +there is evidence that the northern hemisphere underwent a further +progressive diminution of temperature; though the climate of Europe +generally seems at the close of the Tertiary period to have been +if anything warmer, or at any rate not colder, than it is at +the present day. With the commencement of the Quaternary period, +however, this diminution of temperature became more decided; and +beginning with a temperate climate, we find the greater portion +of the northern hemisphere to become gradually subjected to all +the rigours of intense Arctic cold. All the mountainous regions +of Northern and Central Europe, of Britain, and of North America, +became the nurseries of huge ice-streams, and large areas of the +land appear to have been covered with a continuous ice-sheet. +The Arctic conditions of this, the well-known "Glacial period," +relaxed more than once, and were more than once re-established +with lesser intensity. Finally, a gradual but steadily progressive +amelioration of temperature took place; the ice slowly gave way, +and ultimately disappeared altogether; and the climate once more +became temperate, except in high northern latitudes. +</p> + +<p class="indent"> +The changes of temperature sketched out above took place slowly +and gradually, and occupied the whole of the Post-Pliocene period. +In each of the three periods marked out by these changes—in +the early temperate, the central cold, and the later temperate +period—certain deposits were laid down over the surface of +the northern hemisphere; and these deposits collectively constitute +the Post-Pliocene formations. Hence we may conveniently classify +all the accumulations of +<a name="page_336"><span class="page">Page 336</span></a> +this age under the +heads of (1) <i>Pre-Glacial</i> deposits, (2) <i>Glacial</i> +deposits, and (3) <i>Post-Glacial</i> deposits, according as they +were formed before, during, or after the "Glacial period." It cannot +by any means be asserted that we can definitely fix the precise +relations in time of all the Post-Pliocene deposits to the Glacial +period. On the contrary, there are some which hold a very disputed +position as regards this point; and there are others which do not +admit of definite allocation in this manner at all, in consequence +of their occurrence in regions where no "Glacial Period" is known +to have been established. For our present purpose, however, dealing +as we shall have to do principally with the northern hemisphere, +the above classification, with all its defects, has greater +advantages than any other that has been yet proposed. +</p> + +<p class="indent"> +I. PRE-GLACIAL DEPOSITS.—The chief pre-glacial deposit of +Britain is found on the Norfolk coast, reposing upon the Newer +Pliocene (Norwich Crag), and consists of an ancient land-surface +which is known as the "Cromer Forest-bed." +</p> + +<p class="indent"> +This consists of an ancient soil, having embedded in it the stumps +of many trees, still in an erect position, with remains of living +plants, and the bones of recent and extinct quadrupeds. It is +overlaid by fresh-water and marine beds, all the shells of which +belong to existing species, and it is finally surmounted by true +"glacial drift." While all the shells and plants of the Cromer +Forest-bed and its associated strata belong to existing species, +the Mammals are partly living, partly extinct. Thus we find the +existing Wolf (<i>Canis lupus</i>), Red Deer (<i>Cervus elaphus</i>), +Roebuck (<i>Cervus capreolus</i>), Mole (<i>Talpa EuroptĹ“a</i>), +and Beaver (<i>Castor fiber</i>), living in western England side +by side with the <i>Hippopotamus major, Elephas antiquus, Elephas +meridionalis, Rhinoceros Etruscus</i>, and <i>R. Megarhinus</i> +of the Pliocene period, which are not only extinct, but imply +an at any rate moderately warm climate. Besides the above, the +Forest-bed has yielded the remains of several extinct species of +Deer, of the great extinct Beaver (<i>Trogontherium Cuvieri</i>), +of the Caledonian Bull or "Urus" (<i>Bos primigenius</i>), and of +a Horse (<i>Equus fossilis</i>), little if at all distinguishable +from the existing form. +</p> + +<p class="indent"> +The so-called "Bridlington Crag" of Yorkshire, and the "Chillesford +Beds" of Suffolk, are probably to be regarded as also belonging +to this period; though many of the shells which they contain +are of an Arctic character, and would indicate that they were +deposited in the commencement of the Glacial period itself. Owing, +however, to the fact that a few of the shells of these deposits +are not known to occur in a living +<a name="page_337"><span class="page">Page 337</span></a> +condition, these, and some other similar accumulations, are +sometimes considered as referable to the Pliocene period. +</p> + +<p class="indent"> +II. GLACIAL DEPOSITS.—Under this head is included a great +series of deposits which are widely spread over both Europe and +America, and which were formed at a time when the climate of +these countries was very much colder than it is at present, and +approached more or less closely to what we see at the present +day in the Arctic regions. These deposits are known by the +general name of the <i>Glacial deposits</i>, or by the more +specialised names of the Drift, the Northern Drift, the +Boulder-clay, the Till, &c. +</p> + +<p class="indent"> +These glacial deposits are found in Britain as far south as the +Thames, over the whole of Northern Europe, in all the more elevated +portions of Southern and Central Europe, and over the whole of +North America, as far south as the 39th parallel. They generally +occur as sands, clays, and gravels, spread in widely-extended +sheets over all the geological formations alike, except the most +recent, and are commonly spoken of under the general term of +"Glacial drift." They vary much in their exact nature in different +districts, but they universally consist of one, or all, of the +following members:— +</p> + +<p class="indent"> +1. <i>Unstratified</i> clays, or loams, containing numerous angular +or sub-angular blocks of stone, which have often been transported +for a greater or less distance from their parent rock, and which +often exhibit polished, grooved, or striated surfaces. These +beds are what is called <i>Boulder-clay</i>, or <i>Till</i>. +</p> + +<p class="indent"> +2. Sands, gravels, and clays, often more or less regularly +<i>stratified</i>, but containing erratic blocks, often of large +size, and with their edges <i>unworn</i>, derived from considerable +distances from the place where they are now found. In these beds it +is not at all uncommon to find fossil shells; and these, though of +existing species, are generally of an Arctic character, comprising +a greater or less number of forms which are now exclusively found +in the icy waters of the Arctic seas. These beds are often spoken +of as "Stratified Drift." +</p> + +<p class="indent"> +3. <i>Stratified</i> sands and gravels, in which the pebbles +are <i>worn</i> and rounded, and which have been produced by a +rearrangement of ordinary glacial beds by the sea. These beds +are commonly known as "Drift-gravels," or "Regenerated Drift". +</p> + +<p class="indent"> +Some of the last-mentioned of these are doubtless post-glacial; +but, in the absence of fossils, it is often impossible to arrive at +a positive opinion as to the precise age of superficial accumulations +of this nature. It is also the opinion of high authorities that +a considerable number of the so-called "cave-deposits," +<a name="page_338"><span class="page">Page 338</span></a> +with the bones of extinct Mammals, truly belong +to the Glacial period, being formed during warm intervals when +the severity of the Arctic cold had become relaxed. It is further +believed that some, at any rate, of the so-called "high-level" +river-gravels and "brick-earths" have likewise been deposited during +mild or warm intervals in the great age of ice; and in two or three +instances this has apparently been demonstrated—deposits of +this nature, with the bones of extinct animals and the implements +of man, having been shown to be overlaid by true Boulder-clay. +</p> + +<p class="indent"> +The fossils of the undoubted Glacial deposits are principally +shells, which are found in great numbers in certain localities, +sometimes with <i>Foraminifera</i>, the bivalved cases of Ostracode +Crustaceans, &c. Whilst some of the shells of the "Drift" are +such as now live in the seas of temperate regions, others, as +previously remarked, are such as are now only known to live in the +seas of high latitudes; and these therefore afford unquestionable +evidence of cold conditions. Amongst these Arctic forms of shells +which characterise the Glacial beds may be mentioned <i>Pecten +Islandicus</i> (fig. 254), + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 345px; + font-size: smaller; text-align: center;"> +<img src="images/fig254.jpg" width="345" height="370" alt="Fig. 254" /> +<br /> +Fig. 254.—Left valve of <i>Pecten Islandicus</i>, Glacial +and Recent. +</span> +</span> + +<i>Pecten GrĹ“nlandicus, Scalaria GrĹ“nlandica, Leda +truncata, Astarte borealis, Tellina proxima, Nattra clausa</i>, +&c. +</p> + +<p class="indent"> +III. POST-GLACIAL DEPOSITS.—As the intense cold of the +Glacial period became gradually mitigated, and temperate +<a name="page_339"><span class="page">Page 339</span></a> +conditions of climate were once more +re-established, various deposits were formed in the northern +hemisphere, which are found to contain the remains of extinct +Mammals, and which, therefore, are clearly of Post-Pliocene +age. To these deposits the general name of <i>Post-Glacial</i> +formations is given; but it is obvious that, from the nature +of the case, and with our present limited knowledge, we cannot +draw a rigid line of demarcation between the deposits formed +towards the close of the Glacial period, or during warm +"interglacial" periods, and those laid down after the ice had +fairly disappeared. Indeed it is extremely improbable that any +such rigid line of demarcation should ever have existed; and it +is far more likely that the Glacial and Post-Glacial periods, +and their corresponding deposits, shade into one another by an +imperceptible gradation. Accepting this reservation, we may group +together, under the general head of "Post-Glacial Deposits," +most of the so-called "Valley-gravels," "Brick-earths," and +"Cave-deposits," together with some "raised beaches" and various +deposits of peat. Though not strictly within the compass of this +work, a few words may be said here as to the origin and mode of +formation of the Brick-earths, Valley-gravels, and Cave-deposits, +as the subject will thus be rendered more clearly intelligible. +</p> + +<p class="indent"> +Every river produces at the present day beds of fine mud and +loam, and accumulations of gravel, which it deposits at various +parts of its course—the gravel generally occupying the +lowest position, and the finer sands and mud coming above. Numerous +deposits of a similar nature are found in most countries in various +localities, and at various heights above the present channels of +our rivers. Many of these fluviatile (Lat. <i>fluvius</i>, a +river) deposits consist of fine loam, worked for brick-making, +and known as "Brick-earths;" and they have yielded the remains +of numerous extinct Mammals, of which the Mammoth (<i>Elephas +primigenius</i>) is the most abundant. In the valley of the Rhine +these fluviatile loams (known as "Loess") attain a thickness of +several hundred feet, and contain land and fresh-water shells of +existing species. With these occur the remains of Mammals, such +as the Mammoth and Woolly Rhinoceros. Many of these Brick-earths +are undoubtedly Post-Glacial, but others seem to be clearly +"inter-glacial;" and instances have recently been brought forward +in which deposits of Brick-earth containing bones and shells of +fresh-water Molluscs have been found to be overlaid by regular +unstratified boulder-clay. +</p> + +<p class="indent"> +The so-called "Valley-gravels," like the Brick-earths, are +<a name="page_340"><span class="page">Page 340</span></a> +fluviatile deposits, but are of a coarser nature, consisting of +sands and gravels. Every river gives origin to deposits of this kind +at different points along the course of its valley; and it is not +uncommon to find that there exist in the valley of a single river +two or more sets of these gravel-beds, formed by the river itself, +but formed at times when the river ran at different levels, and +therefore formed at different periods. These different accumulations +are known as the "high-level" and "low-level" gravels; and +a reference to the accompanying diagram will explain the origin +and nature of these deposits (fig. 255). When a river begins + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 556px; + font-size: smaller; text-align: justify;"> +<img src="images/fig255.jpg" width="556" height="184" alt="Fig. 255" /> +<br /> +Fig. 255.—Recent and Post-Pliocene Alluvial Deposits. 1, +Peat of the recent period; 2, Gravel of the modern river: 2', +Loam of the modern river; 3. Lower-level valley-gravel with +bones of extinct Mammals (Post-Pliocene); 3', Loam of the same +age as 3; 4. Higher-level valley-gravel (Post-Pliocene); 4', +Loam of the same age as 4; 5. Upland gravels of various kinds +(often glacial drift); 6, Older rock. (After Sir Charles Lyell.) +</span> +</span> + +to occupy a particular line of drainage, and to form its own +channel, it will deposit fluviatile sands and gravels along its +sides. As it goes on deepening the bed or valley through which +it flows, it will deposit other fluviatile strata at a lower +level beside its new bed. In this way have arisen the terms +"high-level" and "low-level" gravels. We find, for instance, a +modern river flowing through a valley which it has to a great +extent or entirely formed itself; by the side of its immediate +channel we may find gravels, sand, and loam (fig. 255, 2 2') +deposited by the river flowing in its present bed. These are +<i>recent</i> fluviatile or alluvial deposits. At some distance +from the present bed of the river, and at a higher level, we +may find other sands and gravels, quite like the recent ones +in character and origin, but formed at a time when the stream +flowed at a higher level, and before it had excavated its valley +to its present depth. These (fig. 255, 3 3') are the so-called +"<i>low-level</i> gravels" of a river. At a still higher level, +and still farther removed from the present bed of the river, we +may find another terrace, composed of just the same materials +as the lower one, but formed at a still earlier period, when the +<a name="page_341"><span class="page">Page 341</span></a> +excavation of the valley had proceeded to a much less extent. +These (fig. 255, 4 4') are the so-called "<i>high-level</i> +gravels" of a river, and there may be one or more terraces of these. +</p> + +<p class="indent"> +The important fact to remember about these fluviatile deposits is +this—that here the ordinary geological rule is reversed. The +high-level gravels are, of course, the highest, so far as their +actual elevation above the sea is concerned; but geologically the +lowest, since they are obviously much older than the low-level +gravels, as these are than the recent gravels. How much older +the high-level gravels may be than the low-level ones, it is +impossible to say. They occur at heights varying from 10 to 100 +feet above the present river-channels, and they are therefore +older than the recent gravels by the time required by the river +to dig out its own bed to this depth. How long this period may +be, our data do not enable us to determine accurately; but if +we are to calculate from the observed rate of erosion of the +actually existing rivers, the period between the different +valley-gravels must be a very long one. +</p> + +<p class="indent"> +The lowest or recent fluviatile deposits which occur beside the +bed of the present river, are referable to the Recent period, +as they contain the remains of none but living Mammals. The two +other sets of gravels are Post-Pliocene, as they contain the +bones of extinct Mammals, mixed with land and fresh-water shells +of existing species. Among the more important extinct Mammals of +the low-level and high-level valley-gravels may be mentioned the +<i>Elephas antiquus</i>, the Mammoth (<i>Elephas primigenius</i>), +the Woolly Rhinoceros (<i>R. Tichorhinus</i>), the Hippopotamus, +the Cave-lion, and the Cave-bear. Along with these are found +unquestionable traces of the existence of Man, in the form of +rude flint implements of undoubted human workmanship. +</p> + +<p class="indent"> +The so-called "Cave-deposits," again, though exhibiting +peculiarities due to the fact of their occurrence in caverns +or fissures in the rocks, are in many respects essentially +similar to the older valley-gravels. Caves, in the great +majority of instances, occur in limestone. When this is not +the case, it will generally be found that they occur along +lines of sea-coast, or along lines which can be shown to have +anciently formed the coast-line. There are many caves, however, +in the making of which it can be shown that the sea has had no +hand; and these are most of the caves of limestone districts. +These owe their origin to the solvent action upon lime of water +holding carbonic acid in solution. The rain which falls upon a +limestone district absorbs a certain amount of carbonic acid from +<a name="page_342"><span class="page">Page 342</span></a> +the air, or from the soil. It then +percolates through the rock, generally along the lines of jointing +so characteristic of limestones, and in its progress it dissolves +and carries off a certain quantity of carbonate of lime. In this +way, the natural joints and fissures in the rock are widened, as +can be seen at the present day in any or all limestone districts. +By a continuance of this action for a sufficient length of time, +caves may ultimately be produced. Nothing, also, is commoner in +a limestone district than for the natural drainage to take the +line of some fissure, dissolving the rock in its course. In this +way we constantly meet in limestone districts with springs issuing +from the limestone rock—sometimes as large rivers—the +waters of which are charged with carbonate of lime, obtained by +the solution of the sides of the fissure through which the waters +have flowed. By these and similar actions, every district in which +limestones are extensively developed will be found to exhibit +a number of natural caves, rents, or fissures. The first element, +therefore, in the production of cave-deposits, is the existence +of a period in which limestone rocks were largely dissolved, and +caves were formed in consequence of the then existing drainage +taking the line of some fissure. +</p> + +<p class="indent"> +Secondly, there must have been a period in which various deposits +were accumulated in the caves thus formed. These cavern-deposits +are of very various nature, consisting of mud, loam, gravel, +or breccias of different kinds. In all cases, these materials +have been introduced into the cave at some period subsequent to, +or contemporaneous with, the formation of the cave. Sometimes +the cave communicates with the surface by a fissure through which +sand, gravel, &c., may be washed by rains or by floods from +some neighbouring river. Sometimes the cave has been the bed of an +ancient stream, and the deposits have been formed as are fluviatile +deposits at the surface. Or, again, the river has formerly flowed +at a greater elevation than it does at present, and the cave +has been filled with fluviatile deposits by the river at a time +prior to the excavation of its bed to the present depth (fig. +256). In this last case, the cave-deposits obviously bear exactly +the same relation in point of antiquity to recent deposits, as +do the low-level and high-level valley-gravels to recent +river-gravels. In any case, it is necessary for the physical +geography of the district to change to some extent, in order +that the cave-deposits should be preserved. If the materials +have been introduced by a fissure, the cave will probably become +ultimately filled to the roof, and the aperture of admission +thus blocked up. If a river has flowed through the cave, the +surface configuration +<a name="page_343"><span class="page">Page 343</span></a> +of the district must be altered so far as to divert the river +into a new channel. And if the cave is placed in the side of +a river-valley, as in fig. 256, the river must have excavated + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 571px; + font-size: smaller; text-align: justify;"> +<img src="images/fig256.jpg" width="571" height="163" alt="Fig. 256" /> +<br /> +Fig. 256.—Diagrammatic section across a river-valley and +cave. <i>a a</i>, Recent valley-gravels near the channel +(<i>b</i>) of the existing river; <i>c</i>, Cavern, partly filled +with cave-earth; <i>d d</i>, High-level gravels, filling fissures +in the limestone, which perhaps communicate in some instances +with the cave, and form a channel by which materials of various +kinds were introduced into it; <i>e e</i>, Inclined beds of +limestone. +</span> +</span> + +its channel to such a depth that it can no longer wash out the +contents of the cave even in high floods. +</p> + +<p class="indent"> +If the cave be entirely filled, the included deposits generally +get more or less completely cemented together by the percolation +through them of water holding carbonate of lime in solution. If +the cave is only partially filled, the dropping of water from +the roof holding lime in solution, and its subsequent evaporation, +would lead to the formation over the deposits below of a layer of +stalagmite, perhaps several inches, or even feet, in thickness. +In this way cave-deposits, with their contained remains, may +be hermetically sealed up and preserved without injury for an +altogether indefinite period of time. +</p> + +<p class="indent"> +In all caves in limestone in which deposits containing bones are +found, we have then evidence of three principal sets of changes. +(1.) A period during which the cave was slowly hollowed out by +the percolation of acidulated water; (2.) A period in which the +cave became the channel of an engulfed river, or otherwise came +to form part of the general drainage-system of the district; (3.) +A period in which the cave was inhabited by various animals. +</p> + +<p class="indent"> +As a typical example of a cave with fossiliferous Post-Pliocene +deposits, we may take Kent's Cavern, near Torquay, in which a +systematic and careful examination has revealed the following +sequence of accumulations in descending order:— +</p> + +<p class="indent"> +(<i>a</i>) Large blocks of limestone, which lie on the floor of +the cave, having fallen from the roof, and which are sometimes +cemented together by stalagmite. +</p> + +<p class="indent"> +(<i>b</i>) A layer of black mould, from three to twelve inches +thick, with human bones, fragments of pottery, stone and +<a name="page_344"><span class="page">Page 344</span></a> +bronze implements, and the bones of animals now living in Britain. +This, therefore, is a <i>recent</i> deposit. +</p> + +<p class="indent"> +(<i>c</i>) A layer of stalagmite, from sixteen to twenty inches +thick, but sometimes as much as five feet, containing the bones +of Man, together with those of extinct Post-Pliocene Mammals. +</p> + +<p class="indent"> +(<i>d</i>) A bed of red cave-earth, sometimes four feet in thickness, +with numerous bones of extinct Mammals (Mammoth, Cave-bear, &c.), +together with human implements of flint and horn. +</p> + +<p class="indent"> +(<i>e</i>) A second bed of stalagmite, in places twelve feet in +thickness, with bones of the Cave-bear. +</p> + +<p class="indent"> +(<i>f</i>) A red-loam and cave-breccia, with remains of the Cave-bear +and human implements. +</p> + +<p class="indent"> +The most important Mammals which are found in cave-deposits in +Europe generally, are the Cave-bear, the Cave-lion, the Cave-hyæna, +the Reindeer, the Musk-ox, the Glutton, and the Lemming—of which +the first three are probably identical with existing forms, and +the remainder are certainly so—together with the Mammoth and +the Woolly Rhinoceros, which are undoubtedly extinct. Along with +these are found the implements, and in some cases the bones, of +Man himself, in such a manner as to render it absolutely certain +that an early race of men was truly contemporaneous in Western +Europe with the animals above mentioned. +</p> + +<p class="indent"> +IV. UNCLASSIFIED POST-PLIOCENE DEPOSITS.—Apart from any +of the afore mentioned deposits, there occur other +accumulations—sometimes superficial, sometimes in +caves—which are found in regions where a "Glacial period" +has not been fully demonstrated, or where such did not take +place; and which, therefore, are not amenable to the above +classification. The most important of these are known to occur +in South America and Australia; and though their numerous +extinct Mammalia place their reference to the Post-Pliocene +period beyond doubt, their relations to the glacial period and +its deposits in the northern hemisphere have not been precisely +determined. +</p> + +<h3>CHAPTER XXII.</h3> + +<p class="subtitle"> +THE POST-PLIOCENE PERIOD—<i>Continued.</i> +</p> + +<p class="indent"> +As regards the <i>life</i> of the Post-Pliocene period, we have, +in the first place, to notice the effect produced throughout the +<a name="page_345"><span class="page">Page 345</span></a> +northern hemisphere by the gradual +supervention of the Glacial period. Previous to this the climate +must have been temperate or warm-temperate; but as the cold +gradually came on, two results were produced as regards the +living beings of the area thus affected. In the first place, +all those Mammals which, like the Mammoth, the Woolly Rhinoceros, +the Lion, the Hyæna, and the Hippopotamus, require, at +any rate, moderately warm conditions, would be forced to +migrate southwards to regions not affected by the new state +of things. In the second place, Mammals previously inhabiting +higher latitudes, such as the Reindeer, the Musk-ox, and the +Lemming, would be enabled by the increasing cold to migrate +southwards, and to invade provinces previously occupied by the +Elephant and the Rhinoceros. A precisely similar, but more +slowly-executed process, must have taken place in the sea, the +northern Mollusca moving southwards as the arctic conditions of +the Glacial period became established, whilst the forms proper +to temperate seas receded. As regards the readily locomotive +Mammals, also, it is probable that this process was carried on +repeatedly in a partial manner, the southern and northern forms +alternately fluctuating backwards and forwards over the same +area, in accordance with the fluctuations of temperature which +have been shown by Mr James Geikie to have characterised the +Glacial period as a whole. We can thus readily account for the +intermixture which is sometimes found of northern and southern +types of Mammalia in the same deposits, or in deposits apparently +synchronous, and within a single district. Lastly, at the final close +of the arctic cold of the Glacial period, and the re-establishment +of temperate conditions over the northern hemisphere, a reversal of +the original process took place—the northern Mammals retiring +within their ancient limits, and the southern forms pressing +northwards and reoccupying their original domains. +</p> + +<p class="indent"> +The <i>Invertebrate</i> animals of the Post-Pliocene deposits +require no further mention—all the known forms, except a +few of the shells in the lowest beds of the formation, being +identical with species now in existence upon the globe. The +only point of importance in this connection has been previously +noticed—namely, that in the true Glacial deposits +themselves a considerable number of the shells belong to +northern or Arctic types. +</p> + +<p class="indent"> +As regards the <i>Vertebrate</i> animals of the period, no extinct +forms of Fishes, Amphibians, or Reptiles are known to occur, but we +meet with both extinct Birds and extinct Mammals. The remains of +the former are of great interest, as indicating +<a name="page_346"><span class="page">Page 346</span></a> +the existence during Post-Pliocene times, at widely remote +points of the southern hemisphere, of various wingless, and for the +most part gigantic, Birds. All the great wingless Birds of the order +<i>Cursores</i> which are known as existing at the present day upon +the globe, are restricted to regions which are either wholly or +in great part south of the equator. Thus the true Ostriches are +African; the Rheas are South American; the Emeus are Australian; +the Cassowaries are confined to Northern Australia, Papua, and the +Indian Archipelago; the species of <i>Apteryx</i> are natives of New +Zealand; and the Dodo and Solitaire (wingless, though probably not +true <i>Cursores</i>), both of which have been exterminated within +historical times, were inhabitants of the islands of Mauritius +and Rodriguez, in the Indian Ocean. In view of these facts, it +is noteworthy that, so far as known, all the Cursorial Birds +of the Post-Pliocene period should have been confined to the +same hemisphere as that inhabited by the living representatives +of the order. It is still further interesting to notice that +the extinct forms in question are only found in geographical +provinces which are now, or have been within historical times, +inhabited by similar types. The greater number of the remains of +these have been discovered in New Zealand, where there now live +several species of the curious wingless genus <i>Apteryx</i>; +and they have been referred by Professor Owen to several generic +groups, of which <i>Dinornis</i> is the most important (fig. +257). Fourteen species of <i>Dinornis</i> have been described by +the distinguished palæontologist just mentioned, all of them +being large wingless birds of the type of the existing Ostrich, +having enormously powerful hind-limbs adapted for running, but +with the wings wholly rudimentary, and the breast-bone devoid +of the keel or ridge which characterises this bone in all birds +which fly. The largest species is the <i>Dinornis giganteus</i>, +one of the most gigantic of living or fossil birds, the shank +(tibia) measuring a yard in length, and the total height being at +least ten feet. Another species, the <i>Dinornis Elephantopus</i> +(fig. 257), though not standing more than about six feet in height, +was of an even more ponderous construction—"the framework of +the skeleton being the most massive of any in the whole class of +Birds," whilst "the toe-bones almost rival those of the Elephant" +(Owen). The feet in <i>Dinornis</i> were furnished with three +toes, and are of interest as presenting us with an undoubted +Bird big enough to produce the largest of the foot-prints of +the Triassic Sandstones of Connecticut. New Zealand has now been +so far explored, that it seems questionable if it can retain in +its recesses any living example of <i>Dinornis</i>; but it +<a name="page_347"><span class="page">Page 347</span></a> +is certain that species of this genus were alive during the human +period, and survived up to quite a recent date. Not only are the +bones very numerous in certain localities, but they are found in + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 383px; + font-size: smaller; text-align: center;"> +<img src="images/fig257.jpg" width="383" height="585" alt="Fig. 257" /> +<br /> +Fig. 257.—Skeleton of <i>Dinornis elephantopus</i>, greatly +reduced. Post-Pliocene, New Zealand. (After Owen.) +</span> +</span> + +the most recent and superficial deposits, and they still contain a +considerable proportion of animal matter; whilst in some instances +bones have been found with the feathers attached, or with the horny +skin of the legs still adhering to them. Charred bones have been +found in connection with native "ovens;" and the traditions of +the Maories contain circumstantial accounts of gigantic wingless +Birds, the "Moas," which were hunted both for their flesh and +their plumage. Upon the whole, therefore, there can be no doubt +<a name="page_348"><span class="page">Page 348</span></a> +but that the Moas of New Zealand have been exterminated at quite +a recent period—perhaps within the last century—by the +unrelenting pursuit of Man,—a pursuit which their wingless +condition rendered them unable to evade. +</p> + +<p class="indent"> +In Madagascar, bones have been discovered of another huge wingless +Bird, which must have been as large as, or larger than, the +<i>Dinornis giganteus</i>, and which has been described under the +name of <i>Æpiornis maximus</i>. With the bones have been found +eggs measuring from thirteen to fourteen inches in diameter, and +computed to have the capacity of three Ostrich eggs. At least two +other smaller species of <i>Æpiornis</i> have been described by +Grandidier and Milne-Edwards as occurring in Madagascar; and they +consider the genus to be so closely allied to the <i>Dinornis</i> +of New Zealand, as to prove that these regions, now so remote, +were at one time united by land. Unlike New Zealand, where there +is the <i>Apteryx</i>, Madagascar is not known to possess any +living wingless Birds; but in the neighbouring island of Mauritius +the wingless Dodo (<i>Didus ineptus</i>) has been exterminated +less than three hundred years ago; and the little island of +Rodriguez, in the same geographical province, has in a similar +period lost the equally wingless Solitaire (<i>Pezophaps</i>), both +of these, however, being generally referred to the <i>Rasores</i>. +</p> + +<p class="indent"> +The <i>Mammals</i> of the Post-Pliocene period are so numerous, +that in spite of the many points of interest which they present, +only a few of the more important forms can be noticed here, and +that but briefly. The first order that claims our attention is +that of the <i>Marsupials</i>, the headquarters of which at the +present day is the Australian province. In Oolitic times Europe +possessed its small Marsupials, and similar forms existed in the +same area in the Eocene and Miocene periods; but if size be any +criterion, the culminating point in the history of the order was +attained during the Post-Pliocene period in + +<span style="float: left; margin: 4px; width: 265px; + font-size: smaller; text-align: justify;"> +<img src="images/fig258.jpg" width="265" height="161" alt="Fig. 258" /> +<br /> +Fig. 258.—Skull of <i>Diprotodon Australis</i>, greatly +reduced. Post-Pliocene, Australia. +</span> + +Australia. From deposits of this age there has been disentombed +a whole series of remains of extinct, and for the most part +gigantic, examples of this group of Quadrupeds. Not to speak of +Wombats and Phalangers, two forms stand out prominently as +representatives of the Post-Pliocene animals of Australia. One +of these is <i>Diprotodon</i> (fig. 258), representing, with +many differences, the well-known modern group of the Kangaroos. +<a name="page_349"><span class="page">Page 349</span></a> +In its teeth, <i>Diprotodon</i> shows +itself to be closely allied to the living, grass-eating Kangaroos; +but the hind-limbs were not so disproportionately long. In size, +also, <i>Diprotodon</i> must have many times exceeded the +dimensions of the largest of its living successors, since the +skull measures no less than three feet in length. The other form +in question is <i>Thylacoleo</i> (fig. 259), which is believed by +Professor Owen to belong to the same group as the existing "Native +Devil" (<i>Dasyurus</i>) of Van Diemen's Land, and therefore to +have been flesh-eating and rapacious in its habits, though this +view is not accepted by others. The principal feature in the skull +of <i>Thylacoleo</i> is the presence, on each side of each jaw, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 536px; + font-size: smaller; text-align: center;"> +<img src="images/fig259.jpg" width="536" height="378" alt="Fig. 259" /> +<br /> +Fig. 259.—Skull of <i>Thylacoleo</i>. Post-Pliocene, +Australia. Greatly reduced. (After Flower.) +</span> +</span> + +of a single huge tooth, which is greatly compressed, and has a +cutting edge. This tooth is regarded by Owen as corresponding to +the great cutting tooth of the jaw of the typical Carnivores, +but Professor Flower considers that <i>Thylacoleo</i> is rather +related to the Kangaroo-rats. The size of the crown of the tooth +in question is not less than two inches and a quarter; and whether +carnivorous or not, it indicates an animal of a size exceeding +that of the largest of existing Lions. +</p> + +<p class="indent"> +The order of the <i>Edentates</i>, comprising the existing Sloths, +Ant-eaters, and Armadillos, and entirely restricted at the present +day to South America, Southern Asia, and Africa, is one alike +<a name="page_350"><span class="page">Page 350</span></a> +singular for the limited geographical range +of its members, their curious habits of life, and the well-marked +peculiarities of their anatomical structure. South America is the +metropolis of the existing forms; and it is an interesting fact +that there flourished within Post-Pliocene times in this continent, +and to some extent in North America also, a marvellous group of +extinct Edentates, representing the living Sloths and Armadillos, +but of gigantic size. The most celebrated of these is the huge +<i>Megatherium Cuvieri</i> (fig. 260) of the South American Pampas. + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 543px; + font-size: smaller; text-align: center;"> +<img src="images/fig260.jpg" width="543" height="301" alt="Fig. 260" /> +<br /> +Fig. 260.—<i>Megatherium Cuvieri</i>. Post-Pliocene, South +America. +</span> +</span> + +The Megathere was a colossal Sloth-like animal which attained a +length of from twelve to eighteen feet, with bones more massive +than those of the Elephant. Thus the thigh-bone is nearly thrice +the thickness of the same bone in the largest of existing Elephants, +its circumference at its narrowest point nearly equalling its +total length; the massive bones of the shank (tibia and fibula) +are amalgamated at their extremities; the heel-bone (calcaneum) +is nearly half a yard in length; the haunch-bones (ilia) are +from four to five feet across at their crests; and the bodies of +the vertebræ at the root of the tail are from five to seven +inches in diameter, from which it has been computed that the +circumference of the tail at this part might have been from five +to six feet. The length of the fore-foot is about a yard, and +the toes are armed with powerful curved claws. It is known now +that the Megathere, in spite of its enormous weight and ponderous +construction, walked, like the existing Ant-eaters and Sloths, +upon the outside edge of the fore-feet, with the claws more or +less bent inwards +<a name="page_351"><span class="page">Page 351</span></a> +towards the palm of the +hand. As in the great majority of the Edentate order, incisor and +canine teeth are entirely wanting, the front of the jaws being +toothless. The jaws, however, are furnished with five upper and +four lower molar teeth on each side. These grinding teeth are from +seven to eight inches in length, in the form of four-sided prisms, +the crowns of which are provided with well-marked transverse ridges; +and they continue to grow during the whole life of the animal. +There are indications that the snout was prolonged, and more or +less flexible; and the tongue was probably prehensile. From the +characters of the molar teeth it is certain that the Megathere was +purely herbivorous in its habits; and from the enormous size and +weight of the body, it is equally certain that it could not have +imitated its modern allies, the Sloths, in the feat of climbing, +back downwards, amongst the trees. It is clear, therefore, that +the Megathere sought its sustenance upon the ground; and it was +originally supposed to have lived upon roots. By a masterly piece +of deductive reasoning, however, Professor Owen showed that this +great "Ground-Sloth" must have truly lived upon the foliage of +trees, like the existing Sloths—but with this difference, +that instead of climbing amongst the branches, it actually +uprooted the tree bodily. In this <i>tour de force</i>, the animal +sat upon its huge haunches and mighty tail, as on a tripod, and +then grasping the trunk with its powerful arms, either wrenched it +up by the roots or broke it short off above the ground. Marvellous +as this may seem, it can be shown that every detail in the skeleton +of the Megathere accords with the supposition that it obtained +its food in this way. Similar habits were followed by the allied +<i>Mylodon</i> (fig. 261), another of the great "Ground-Sloths," +which inhabited South America during the Post-Pliocene period. In +most respects, the <i>Mylodon</i> is very like the Megathere; but +the crowns of the molar teeth are flat instead of being ridged. +The nearly-related genus <i>Megalonyx</i>, unlike the Megathere, +but like the Mylodon, extended its range northwards as far as +the United States. +</p> + +<p class="indent"> +Just as the Sloths of the present day were formerly represented +in the same geographical area by the gigantic Megatheroids, so +the little banded and cuirassed Armadillos of South America were +formerly represented by gigantic species, constituting the genus +<i>Glyptodon</i>. The <i>Glyptodons</i> (fig. 262) differed from +the living Armadillos in having no bands in their armour, so that +they must have been unable to roll themselves up. It is rare +at the present day to meet with any +<a name="page_352"><span class="page">Page 352</span></a> +Armadillo over two or three feet in length; but the length of the +<i>Glyptodon clavipes</i>, from the tip of the snout to the end +of the tail, was more than nine feet. +</p> + +<p class="indent"> +There are no canine or incisor teeth in the <i>Glyptodon</i>, but + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 382px; + font-size: smaller; text-align: center;"> +<img src="images/fig261.jpg" width="382" height="434" alt="Fig. 261" /> +<br /> +Fig. 261.—Skeleton of <i>Mylodon robustus</i>. Post-Pliocene, +South America. +</span> +</span> + +there are eight molars on each side of each jaw, and the crowns +of these are fluted and almost trilobed. The head is covered + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 472px; + font-size: smaller; text-align: center;"> +<img src="images/fig262.jpg" width="472" height="222" alt="Fig. 262" /> +<br /> +Fig. 262.—Skeleton of <i>Glyptodon clavipes</i>. +Post-Pliocene, South America. +</span> +</span> + +by a helmet of bony plates, and the trunk was defended by an +armour of almost hexagonal bony pieces united by sutures, and +<a name="page_353"><span class="page">Page 353</span></a> +exhibiting special patterns of sculpturing +in each species. The tail was also defended by a similar armour, +and the vertebræ were mostly fused together so as to form +a cylindrical bony rod. In addition to the above-mentioned forms, +a number of other Edentate animals have been discovered by the +researches of M. Lund in the Post-Pliocene deposits of the +Brazilian bone-caves. Amongst these are true Ant-eaters, +Armadillos, and Sloths, many of them of gigantic size, and +all specifically or generically distinct from existing forms. +</p> + +<p class="indent"> +Passing over the aquatic orders of the <i>Sirenians</i> and +<i>Cetaceans</i>, we come next to the great group of the Hoofed +Quadrupeds, the remains of which are very abundant in Post-Pliocene +deposits both in Europe and North America. Amongst the Odd-toed +Ungulates the most important are the Rhinoceroses, of which three +species are known to have existed in Europe during the Post-Pliocene +period. Two of these are the well-known Pliocene forms, the +<i>Rhinoceros Etruscus</i> and the <i>R. Megarhinus</i> still +surviving in diminished numbers; but the most famous is the +<i>Rhinoceros tichorhinus</i> (fig. 263), or so-called "Woolly + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 525px; + font-size: smaller; text-align: center;"> +<img src="images/fig263.jpg" width="525" height="281" alt="Fig. 263" /> +<br /> +Fig. 263.—Skull of the Tichorhine Rhinoceros, the horns +being wanting. One-tenth of the natural size. Post-Pliocene +deposits of Europe and Asia. +</span> +</span> + +Rhinoceros." This species is known not only by innumerable bones, +but also by a carcass, at the time of its discovery complete, +which was found embedded in the frozen soil of Siberia towards +the close of last century, and which was partly saved from +destruction by the exertions of the naturalist Pallas. From this, +we know that the Tichorhine Rhinoceros, like its associate the +Mammoth, was provided with a coating of hair, and therefore was +enabled to endure a more severe climate than any existing +<a name="page_354"><span class="page">Page 354</span></a> +species. The skin was not thrown into the folds +which characterise most of the existing forms; and the technical +name of the species refers to the fact that the nostrils were +completely separated by a bony partition. The head carried two +horns, placed one behind the other, the front one being unusually +large. As regards its geographical range, the Woolly Rhinoceros +is found in Europe in vast numbers north of the Alps and Pyrenees, +and it also abounded in Siberia; so that it would appear to be a +distinctly northern form, and to have been adapted for a temperate +climate. It is not known to occur in Pliocene deposits, but it +makes its first appearance in the Pre-Glacial deposits, surviving +the Glacial period, and being found in abundance in Post-Glacial +accumulations. It was undoubtedly a contemporary of the earlier +races of men in Western Europe; and it may perhaps be regarded as +being the actual substantial kernel of some of the "Dragons" of +fable. +</p> + +<p class="indent"> +The only other Odd-toed Ungulate which needs notice is the so-called +<i>Equus fossilis</i> of the Post-Pliocene of Europe. This made +its appearance before the Glacial period, and appears to be in +reality identical with the existing Horse (<i>Equus caballus</i>). +True Horses also occur in the Post-Pliocene of North America; +but, from some cause or another, they must have been exterminated +before historic times. +</p> + +<p class="indent"> +Amongst the Even-toed Ungulates, the great <i>Hippopotamus major</i> +of the Pliocene still continued to exist in Post-Pliocene times in +Western Europe; and the existing Wild Boar (<i>Sus scrofa</i>), +the parent of our domestic breeds of Pigs, appeared for the first +time. The Old World possessed extinct representatives of its +existing Camels, and lost types of the living Llamas inhabited +South America. Amongst the Deer, the Post-Pliocene accumulations +have yielded the remains of various living species, such as the +Red Deer (<i>Cervus elaphus</i>), the Reindeer (<i>Cervus +tarandus</i>), the Moose or Elk (<i>Alces malchis</i>), and the +Roebuck (<i>Cervus capreolus</i>), together with a number of +extinct forms. Among the latter, the great "Irish Elk" (<i>Cervus +megaceros</i>) is justly celebrated both for its size and for +the number and excellent preservation of its discovered remains. +This extinct species (fig. 264) has been found principally in +peat-mosses and Post-Pliocene lake-deposits, and is remarkable +for the enormous size of the spreading antlers, which are widened +out towards their extremities, and attain an expanse of over +ten feet from tip to tip. It is not a genuine Elk, but is +intermediate between the Reindeer and the Fallow-deer. Among the +existing Deer +<a name="page_355"><span class="page">Page 355</span></a> +of the Post-Pliocene, the most noticeable is the +Reindeer, an essentially northern type, existing at the present +day in Northern Europe, and also (under the name of the "Caribou") + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 576px; + font-size: smaller; text-align: center;"> +<img src="images/fig264.jpg" width="576" height="618" alt="Fig. 264" /> +<br /> +Fig. 264—Skeleton of the "Irish Elk" (<i>Cervus +megaceros</i>). Post-Pliocene, Britain. +</span> +</span> + +in North America. When the cold of the Glacial period became +established, this boreal species was enabled to invade Central +and Western Europe in great herds, and its remains are found +abundantly in cave-earths and other Post-Pliocene deposits as +far south as the Pyrenees. +</p> + +<p class="indent"> +In addition to the above, the Post-Pliocene deposits of Europe +and North America have yielded the remains of various Sheep and +Oxen. One of the most interesting of the latter is +<a name="page_356"><span class="page">Page 356</span></a> +the "Urus" or Wild Bull (<i>Bos primigenius</i>, fig. 265), which, +though much larger than any of the existing fossils, is believed to + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 513px; + font-size: smaller; text-align: center;"> +<img src="images/fig265.jpg" width="513" height="322" alt="Fig. 265" /> +<br /> +Fig. 265.—Skull of the Urns (<i>Bos primigenius</i>). +Post-Pliocene and Recent. (After Owen.) +</span> +</span> + +be specifically undistinguishable from the domestic Ox (<i>Bos +taurus</i>), and to be possibly the ancestor of some of the larger +European varieties of oxen. In the earlier part of its existence +the Urus ranged over Europe and Britain in company with the Woolly +Rhinoceros and the Mammoth; but it long survived these, and does +not appear to have been finally exterminated till about the twelfth +century. Another remarkable member of the Post-Pliocene Cattle, +also to begin with an associate of the Mammoth and Rhinoceros, +is the European Bison or "Aurochs" (<i>Bison priscus</i>). This +"maned" ox formerly abounded in Europe in Post-Glacial times, +and was not rare even in the later periods of the Roman empire, +though much diminished in numbers, and driven back into the wilder +and more inaccessible parts of the country. At present this fine +species has been so nearly exterminated that it no longer exists in +Europe save in Lithuania, where its preservation has been secured +by rigid protective laws. Lastly, the Post-Pliocene deposits have +yielded the remains of the singular living animal which is known +as the Musk-ox or Musk-sheep (<i>Ovibos moschatus</i>). At the +present day, the Musk-ox is an inhabitant of the "barren grounds" +of Arctic America, and it is remarkable for the great length of +its hair. It is, like the Reindeer, a +<a name="page_357"><span class="page">Page 357</span></a> +distinctively northern animal; but it enjoyed during the Glacial +period a much wider range than it has at the present day, the +conditions suitable for its existence being then extended over +a considerable portion of the northern hemisphere. Thus remains +of the Musk-Ox are found in greater or less abundance in +Post-Pliocene deposits over a great part of Europe, extending +even to the south of France; and closely-related forms are found +in similar deposits in the United States. +</p> + +<p class="indent"> +Coming to the <i>Proboscideans</i>, we find that the +<i>Mastodons</i> seem to have disappeared in Europe at the +close of the Pliocene period, or at the very commencement +of the Post-Pliocene. In the New World, on the other hand, +a species of Mastodon (<i>M. Americanus</i> or <i>M. +Ohioticus</i>) is found abundantly in deposits of Post-Pliocene +age, from Canada to Texas. Very perfect skeletons of this +species have been exhumed from morasses and swamps, and large +individuals attained a length (exclusive of the tusks) of +seventeen feet and a height of eleven feet, the tusks being +twelve feet in length. Remains of <i>Elephants</i> are also +abundant in the Post-Pliocene deposits of both the Old and the +New World. Amongst these, we find in Europe the two familiar +Pliocene species <i>E. Meridionales</i> and <i>E. Antiquus</i> +still surviving, but in diminished numbers. With these are found +in vast abundance the remains of the characteristic Elephant +of the Post-Pliocene, the well-known "Mammoth" (<i>Elephas +primigenius</i>), which is accompanied in North America by the +nearly-allied, but more southern species, the <i>Elephas +Americanus</i>. The Mammoth (fig. 266) is considerably larger +than the largest of the living Elephants, the skeleton being +over sixteen feet in length, exclusive of the tusks, and over +nine feet in height. The tusks are bent almost into a circle, +and are sometimes twelve feet in length, measured along their +curvature. In the frozen soil of Siberia several carcasses of +the Mammoth have been discovered with the flesh and skin still +attached to the bones, the most celebrated of these being a Mammoth +which was discovered at the beginning of this century at the +mouth of the Lena, on the borders of the Frozen Sea, and the +skeleton of which is now preserved at St Petersburg (fig. 266). +From the occurrence of the remains of the Mammoth in vast numbers +in Siberia, it might have been safely inferred that this ancient +Elephant was able to endure a far more rigorous climate than its +existing congeners. This inference has, however, been rendered +a certainty by the specimens just referred to, which show that +the Mammoth was protected against the cold by a thick coat of +reddish-brown wool, some nine or ten inches long, interspersed +with strong, coarse black +<a name="page_358"><span class="page">Page 358</span></a> +hair more than a foot in length. The +teeth of the Mammoth (fig.267) are of the type of those of the +existing Indian Elephant, and are found in immense numbers in +certain localities. The Mammoth was essentially northern in its + +<!-- The original image description was displayed sideways --> +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 487px; + font-size: smaller; text-align: center;"> +<img src="images/fig266.jpg" width="487" height="749" alt="Fig. 266" /> +<br /> +Fig. 266.—Skeleton of the Mammoth (<i>Elephas +primigenius</i>). Portions of the integument still adhere +to the head, and the thick skin of the soles is still +attached to the feet. Post-Pliocene. +</span> +</span> + +distribution, never passing south of a line drawn through the +Pyrenees, the Alps, the northern shores of the Caspian, Lake +<a name="page_359"><span class="page">Page 359</span></a> +Baikal, Kamschatka, and the Stanovi Mountains (Dawkins). It +occurs in the Pre-Glacial forest-bed of Cromer in Norfolk, + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 418px; + font-size: smaller; text-align: center;"> +<img src="images/fig267.jpg" width="418" height="324" alt="Fig. 267" /> +<br /> +Fig. 267.—Molar tooth of the Mammoth (<i>Elephas +primigenius</i>), upper jaw, right side, one-third of the natural +size. <i>a</i>, Grinding surface; <i>b</i>, Side view. +Post-Pliocene. +</span> +</span> + +survived the Glacial period, and is found abundantly in +Post-Glacial deposits in France, Germany, Britain, Russia in +Europe, Asia, and North America, being often associated with the +Reindeer, Lemming, and Musk-ox. That it survived into the earlier +portion of the human period is unquestionable, its remains having +been found in a great number of instances associated with +implements of human manufacture; whilst in one instance a +recognisable portrait of it has been discovered, carved on bone. +</p> + +<p class="indent"> +Amongst other Elephants which occur in Post-Pliocene deposits +may be mentioned, as of special interest, the pigmy Elephants of +Malta. One of these—the <i>Elephas Melitensis</i>, or +so-called "Donkey-Elephant"—was not more than four and a +half feet in height. The other—the <i>Elephas Falconeri</i>, +of Busk—was still smaller, its average height at the +withers not exceeding two and a half to three feet. +</p> + +<p class="indent"> +Whilst herbivorous animals abounded during the Post-Pliocene, +we have ample evidence of the coexistence with them of a number +of Carnivorous forms, both in the New and the Old World. The +Bears are represented in Europe by at least three species, two +of which—namely, the great Grizzly Bear (<i>Ursus ferox</i>) +and the smaller Brown Bear (<i>Ursus arctos</i>)—are in +existence at the present day. The third species is the +<a name="page_360"><span class="page">Page 360</span></a> +celebrated Cave-bear +(<i>Ursus spelĹ“us</i>, fig. 268), which is now extinct. The +Cave-bear exceeded in its dimensions the largest of modern Bears; + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 497px; + font-size: smaller; text-align: center;"> +<img src="images/fig268.jpg" width="497" height="263" alt="Fig. 268" /> +<br /> +Fig. 268.—Skull of <i>Ursus spelpeus</i>. Post-Pliocene, +Europe. One-sixth of the natural size. +</span> +</span> + +and its remains, as its name implies; have been found mainly in +cavern-deposits. Enormous numbers of this large and ferocious +species must have lived in Europe in Post-Glacial times; and that +they survived into the human period, is clearly shown by the common +association of their bones with the implements of man. They are +occasionally accompanied by the remains of a Glutton (the <i>Gulo +spelĹ“us</i>), which does not appear to be really separable from +the existing Wolverine or Glutton of northern regions (the <i>Gulo +luscus</i>). In addition, we meet with the bones of the Wolf, Fox, +Weasel, Otter, Badger, Wild Cat, Panther, Hyæna, and Lion, +&c., together with the extinct <i>Machairodus</i> or "Sabre-toothed +Tiger." The only two of these that deserve further mention are +the Hyæna and the Lion. The Cave-hyæna (<i>HyĹ“na +spelĹ“a</i>, fig. 269) is regarded by high authorities as nothing +more than a variety of the living Spotted Hyæna (<i>H. +Crocuta</i>) of South Africa. This well-known species inhabited Britain +and a considerable portion of Europe during a large part of the +Post-Pliocene period; and its remains often occur in great abundance. +Indeed, some caves, such as the Kirkdale Cavern in Yorkshire, were +dens inhabited during long periods by these animals, and thus contain +the remains of numerous individuals and of successive generations of +Hyænas, together with innumerable gnawed and bitten bones of +their prey. That the Cave-hyæna was a contemporary with Man in +Western Europe during Post-Glacial times is shown beyond a doubt by +the common association of its bones with human implements. +</p> + +<p class="indent"> +<a name="page_361"><span class="page">Page 361</span></a> +Lastly, the so-called Cave-lion (<i>Felis spelĹ“a</i>), long +supposed to be a distinct species, has been shown to be nothing + +<span style="float: left; width: 100%; text-align: center;"> +<span style="margin: 4px; width: 454px; + font-size: smaller; text-align: center;"> +<img src="images/fig269.jpg" width="454" height="245" alt="Fig. 269" /> +<br /> +Fig. 269.—Skull of <i>HyĹ“na spelĹ“a</i>, one-fourth +of the natural size. Post-Phocene, Europe. +</span> +</span> + +more than a large variety of the existing Lion (<i>Felis leo</i>). +This animal inhabited Britain and Western Europe in times posterior +to the Glacial period, and was a contemporary of the Cave-hyæna, +Cave-bear, Woolly Rhinoceros, and Mammoth. The Cave-lion also +unquestionably survived into the earlier portion of the human +period in Europe. +</p> + +<p class="indent"> +The Post-Pliocene deposits of Europe have further yielded the remains +of numerous <i>Rodents</i>—such as the Beaver, the Northern +Lemming, Marmots, Mice, Voles, Rabbits, &c.—together with +the gigantic extinct Beaver known as the <i>Trogontherium Cuvieri</i> +(fig. 270). The great <i>Castoroides Ohioensis</i> of the + +<span style="float: right; margin: 4px; width: 258px; + font-size: smaller; text-align: justify;"> +<img src="images/fig270.jpg" width="258" height="148" alt="Fig. 270" /> +<br /> +Fig. 270.—Lower jaw of <i>Trogontherium Cuvieri</i>, one-fourth +of the natural size. Post-Pliocene, Britain. +</span> + +Post-Pliocene of North America is also a great extinct Beaver, +which reached a length of about five feet. Lastly, the Brazilian +bone-caves have yielded the remains of numerous Rodents of types +now characteristic of South America, such as Guinea-pigs, Capybaras, +tree-inhabiting Porcupines, and Coypus. +</p> + +<p class="indent"> +The deposits just alluded to have further yielded the remains of +various Monkeys, such as Howling Monkeys, Squirrel Monkeys, and +Marmosets, all of which belong to the group of <i>Quadrumana</i> +which is now exclusively confined to the +<a name="page_362"><span class="page">Page 362</span></a> +South American continent—namely, the "Platyrhine" Monkeys. +</p> + +<p class="indent"> +We still have very briefly to consider the occurrence of Man +in Post-Pliocene deposits; but before doing so, it will be well +to draw attention to the evidence afforded by the Post-Pliocene +Mammals as to the climate of Western Europe at this period. The +chief point which we have to notice is, that a considerable +revolution of opinion has taken place on this point. It was +originally believed that the presence of such animals as Elephants, +Lions, the Rhinoceros, and the Hippopotamus afforded an irrefragable +proof that the climate of Europe must have been a warm one, at any +rate during Post-Glacial times. The existence, also, of numbers +of Mammoths in Siberia, was further supposed to indicate that +this high temperature extended itself very far north. Upon the +whole, however, the evidence is against this view. Not only is +there great difficulty in supposing that the Arctic conditions of +the Glacial period were immediately followed by anything warmer +than a cold-temperate climate; but there is nothing in the nature +of the Mammals themselves which would absolutely forbid their +living in a temperate climate. The <i>Hippopotamus major</i>, +though probably clad in hair, offers some difficulty—since, +as pointed out by Professor Busk, it must have required a climate +sufficiently warm to insure that the rivers were not frozen over +in the winter; but it was probably a migratory animal, and its +occurrence may be accounted for by this. The Woolly Rhinoceros +and the Mammoth are known with certainty to have been protected +with a thick covering of wool and hair; and their extension +northwards need not necessarily have been limited by anything +except the absence of a sufficiently luxuriant vegetation to +afford them food. The great American Mastodon, though not certainly +known to have possessed a hairy covering, has been shown to have +lived upon the shoots of Spruce and Firs, trees characteristic of +temperate regions—as shown by the undigested food which has +been found with its skeleton, occupying the place of the stomach. +The Lions and Hyænas, again, as shown by Professor Boyd Dawkins, +do not indicate necessarily a warm climate. Wherever a sufficiency +of herbivorous animals to supply them with food can live, there +they can live also; and they have therefore no special bearing +upon the question of climate. After a review of the whole evidence, +Professor Dawkins concludes that the nearest approach at the +present day to the Post-Pliocene climate of Western Europe is +to be found in the climate of the great Siberian plains which +stretch from the Altai Mountains to the Frozen Sea. "Covered +<a name="page_363"><span class="page">Page 363</span></a> +by impenetrable forests, for the most part +of Birch, Poplar, Larch, and Pines, and low creeping dwarf Cedars, +they present every gradation in climate from the temperate to that +in which the cold is too severe to admit of the growth of trees, +which decrease in size as the traveller advances northwards, and +are replaced by the grey mosses and lichens that cover the low +marshy 'tundras.' The maximum winter cold, registered by Admiral +Von Wrangel at Nishne Kolymsk, on the banks of the Kolyma, +is—65° in January. 'Then breathing becomes difficult; +the Reindeer, that citizen of the Polar region, withdraws to the +deepest thicket of the forest, and stands there motionless as +if deprived of life;' and trees burst asunder with the cold. +Throughout this area roam Elks, Black Bears, Foxes, Sables, and +Wolves, that afford subsistence to the Jakutian and Tungusian +fur-hunters. In the northern part countless herds of Reindeer, +Elks, Foxes, and Wolverines make up for the poverty of vegetation +by the rich abundance of animal life. 'Enormous flights of Swans, +Geese, and Ducks arrive in the spring, and seek deserts where +they may moult and build their nests in safety. Ptarmigans run in +troops amongst the bushes; little Snipes are busy along the brooks +and in the morasses; the social Crows seek the neighbourhood of +new habitations; and when the sun shines in spring, one may even +sometimes hear the cheerful note of the Finch, and in autumn +that of the Thrush.' Throughout this region of woods, a hardy, +middle-sized breed of horses lives under the mastership and care +of man, and is eminently adapted to bear the severity of the +climate.... The only limit to their northern range is the difficulty +of obtaining food. The severity of the winter through the southern +portion of this vast wooded area is almost compensated for by the +summer heat and its marvellous effect on +vegetation."—(Dawkins, 'Monograph of Pleistocene Mammalia.') +</p> + +<p class="indent"> +Finally, a few words must be said as to the occurrence of the +remains of Man in Post-Pliocene deposits. That Man existed in +Western Europe and in Britain during the Post-Pliocene period, is +placed beyond a doubt by the occurrence of his bones in deposits +of this age, along with the much more frequent occurrence of +implements of human manufacture. At what precise point of time +during the Post-Pliocene period he first made his appearance is +still a matter of conjecture. Recent researches would render +it probable that the early inhabitants of Britain and Western +Europe were witnesses of the stupendous phenomena of the Glacial +period; but this cannot be said to have been demonstrated. That +Man existed in these +<a name="page_364"><span class="page">Page 364</span></a> +regions during the +Post-Glacial division of Post-Pliocene time cannot be doubted +for a moment. As to the physical peculiarities of the ancient +races that lived with the Mammoth and the Woolly Rhinoceros, +little is known compared with what we may some day hope to know. +Such information as we have, however, based principally on the +skulls of the Engis, Neanderthal, Cro-Magnon, and Bruniquel +caverns, would lead to the conclusion that Post-Pliocene Man +was in no respect inferior in his organisation to, or less +highly developed than, many existing races. All the known +skulls of this period, with the single exception of the +Neanderthal cranium, are in all respects average and normal in +their characters; and even the Neanderthal skull possessed a +cubic capacity at least equal to that of some existing races. The +implements of Post-Pliocene Man are exclusively of stone or bone; +and the former are invariably of rude shape and <i>undressed</i>. +These "palæolithic" tools (Gr. <i>palaios</i>; ancient; +<i>lithos</i>, stone) point to a very early condition of the +arts; since the men of the earlier portion of the Recent period, +though likewise unacquainted with the metals, were in the habit +of polishing or dressing the stone implements which they +fabricated. +</p> + +<p class="indent"> +It is impossible here to enter further into this subject; and +it would be useless to do so without entering as well into a +consideration of the human remains of the Recent period—a +period which lies outside the province of the present work. So +far as Post-Pliocene Man is concerned, the chief points which +the palæontological student has to remember have been +elsewhere summarised by the author as follows:— +</p> + +<p class="indent"> +1. Man unquestionably existed during the later portion of what +Sir Charles Lyell has termed the "Post-Pliocene" period. In other +words, Man's existence dates back to a time when several remarkable +Mammals, previously mentioned, had not yet become extinct; but he +does not date back to a time anterior to the present <i>Molluscan</i> +fauna. +</p> + +<p class="indent"> +2. The antiquity of the so-called Post-Pliocene period is a matter +which must be mainly settled by the evidence of Geology proper, +and need not be discussed here. +</p> + +<p class="indent"> +3. The extinct Mammals with which man coexisted in Western Europe +are mostly of large size, the most important being the Mammoth +(<i>Elephas primogenius</i>), the Woolly Rhinoceros (<i>Rhinoceros +tichorhinus</i>), the Cave-lion (<i>Felis spelĹ“a</i>), the +Cave-hyæna(<i>HyĹ“na spelĹ“a</i>), and the Cave-bear +(<i>Ursus spelĹ“us</i>). We do not know the causes which led +to the extinction of these Mammals; but we know that hardly any +Mammalian species has become extinct during the historical period. +</p> + +<p class="indent"> +<a name="page_365"><span class="page">Page 365</span></a> +4. The extinct Mammals with which man coexisted are referable in +many cases to species which presumably required a very different +climate to that now prevailing in Western Europe. How long a period, +however, has been consumed in the bringing about of the climatic +changes thus indicated, we have no means of calculating with any +approach to accuracy. +</p> + +<p class="indent"> +5. Some of the deposits in which the remains of man have been +found associated with the bones of extinct Mammals, are such as +to show incontestably that great changes in the physical geography +and surface-configuration of Western Europe have taken place +since the period of their accumulation. We have, however, no +means at present of judging of the lapse of time thus indicated +except by analogies and comparisons which may be disputed. +</p> + +<p class="indent"> +6. The human implements which are associated with the remains +of extinct Mammals, themselves bear evidence of an exceedingly +barbarous condition of the human species. Post-Pliocene or +"Palæolithic" Man was clearly unacquainted with the use +of any of the metals. Not only so, but the workmanship of these +ancient races was much inferior to that of the later tribes, who +were also ignorant of the metals, and who also used nothing but +weapons and tools of stone, bone, &c. +</p> + +<p class="indent"> +7. Lastly, it is only with the human remains of the Post-Pliocene +period that the palæontologist proper has to deal. When we +enter the "Recent" period, in which the remains of Man are +associated with those of <i>existing species of Mammals</i>, we +pass out of the region of pure palæontology into the domain +of the Archæologist and the Ethnologist. +</p> + +<h4>LITERATURE.</h4> + +<p class="indent"> +The following are some of the principal works and memoirs to which +the student may refer for information as to the Post-Pliocene +deposits and the remains which they contain, as well as to the +primitive races of mankind:— +</p> + +<table border="0" cellspacing="0"> +<tr><td class="right" valign="top">(1)</td> + <td>'Elements of Geology.' Lyell.</td> +</tr><tr><td class="right" valign="top">(2)</td> + <td>'Antiquity of Man.' Lyell.</td> +</tr><tr><td class="right" valign="top">(3)</td> + <td>'Palæontological Memoirs.' Falconer.</td> +</tr><tr><td class="right" valign="top">(4)</td> + <td>'The Great Ice-age.' James Geikie.</td> +</tr><tr><td class="right" valign="top">(5)</td> + <td>'Manual of Palæontology.' Owen.</td> +</tr><tr><td class="right" valign="top">(6)</td> + <td>'British Fossil Mammals and Birds.' Owen.</td> +</tr><tr><td class="right" valign="top">(7)</td> + <td>'Cave-Hunting.' Boyd Dawkins.</td> +</tr><tr><td class="right" valign="top">(8)</td> + <td>'Prehistoric Times.' Lubbock.</td> +</tr><tr><td class="right" valign="top">(9)</td> + <td>'Ancient Stone Implements.' Evans.</td> +</tr><tr><td class="right" valign="top">(10)</td> + <td>'Prehistoric Man.' Daniel Wilson.</td> +</tr><tr><td class="right" valign="top">(11)</td> + <td>'Prehistoric Races of the United States.' Foster.</td> +</tr><tr><td class="right" valign="top">(12)</td> + <td>'Manual of Geology.' Dana.</td> +</tr><tr><td class="right" valign="top"> +<a name="page_366"><span class="page">Page 366</span></a> + (13)</td> + <td>'Monograph of Pleistocene Mammalia' (Palæontographical + Society). Boyd Dawkins and Sanford.</td> +</tr><tr><td class="right" valign="top">(14)</td> + <td>'Monograph of the Post-Tertiary Entomostraca of Scotland, + &c., with an Introduction on the Post-Tertiary Deposits + of Scotland' (Ibid.) G. S. Brady, H. W. Crosskey, and D. + Robertson.</td> +</tr><tr><td class="right" valign="top">(15)</td> + <td>"Reports on Kent's Cavern"—'British Association + Reports.' Pengelly.</td> +</tr><tr><td class="right" valign="top">(16)</td> + <td>"Reports on the Victoria Cavern, Settle"—'British + Association Reports.' Tiddeman.</td> +</tr><tr><td class="right" valign="top">(17)</td> + <td>'Ossemens Fossiles.' Cuvier.</td> +</tr><tr><td class="right" valign="top">(18)</td> + <td>'Reliquiæ Diluvianæ.' Buckland.</td> +</tr><tr><td class="right" valign="top">(19)</td> + <td>"Fossil Mammalia"—'Zoology of the Voyage of the + Beagle.' Owen.</td> +</tr><tr><td class="right" valign="top">(20)</td> + <td>'Description of the Tooth and Part of the Skeleton of + the <i>Glyptodon</i>.' Owen.</td> +</tr><tr><td class="right" valign="top">(21)</td> + <td>"Memoir on the Extinct Sloth Tribe of North + America"—'Smithsonian Contributions to Knowledge.' + Leidy.</td> +</tr><tr><td class="right" valign="top">(22)</td> + <td>"Report on Extinct Mammals of Australia"—'British + Association,' 1844. Owen.</td> +</tr><tr><td class="right" valign="top">(23)</td> + <td>'Description of the Skeleton of an Extinct Gigantic + Sloth (<i>Mylodon robtutus</i>).' Owen.</td> +</tr><tr><td class="right" valign="top">(24)</td> + <td>"Affinities and Probable Habits of Thylacoleo"—'Quart. + Journ. Geol. Soc.,' vol. xxiv. Flower.</td> +</tr><tr><td class="right" valign="top">(25)</td> + <td>'Prodromus of the Palæontology of Victoria.' + M'Coy.</td> +</tr><tr><td class="right" valign="top">(26)</td> + <td>'Les Ossemens Fossiles des Cavernes de Liège.' + Schmerling.</td> +</tr><tr><td class="right" valign="top">(27)</td> + <td>'Die Fauna der Pfahlbauten in der Schweiz.' + Rütimeyer.</td> +</tr><tr><td class="right" valign="top">(28)</td> + <td>"Extinct and Existing Bovine Animals of + Scandinavia"—'Annals of Natural History,' ser. 2, vol. + iv., 1849. Nilsson.</td> +</tr><tr><td class="right" valign="top">(29)</td> + <td>'Man's Place in Nature.' Huxley.</td> +</tr><tr><td class="right" valign="top">(30)</td> + <td>'Les Temps AntĂ©historiques en Belgique.' + Dupont.</td> +</tr><tr><td class="right" valign="top">(31)</td> + <td>"Classification of the Pleistocene Strata of Britain and + the Continent"—'Quart. Journ. Geol. Soc.,' vol. xxviii. + Boyd Dawkins.</td> +</tr><tr><td class="right" valign="top">(32)</td> + <td>'Distribution of the Post-Glacial Mammalia' (Ibid.), vol. + xxv. Boyd Dawkins.</td> +</tr><tr><td class="right" valign="top">(33)</td> + <td>'On British Fossil Oxen' (Ibid.), vols. xxii. and xxiii. + Boyd Dawkins.</td> +</tr><tr><td class="right" valign="top">(34)</td> + <td>'British Prehistoric Mammals' (Congress of Prehistoric + Archæology, 1868). Boyd Dawkins.</td> +</tr><tr><td class="right" valign="top">(35)</td> + <td>'Reliquiæ Aquitanicæ.' Lartet and + Christy.</td> +</tr><tr><td class="right" valign="top">(36)</td> + <td>'Zoologie et PalĂ©ontologie Françaises.' + Gervais.</td> +</tr><tr><td class="right" valign="top">(37)</td> + <td>'Notes on the Post-Pliocene Geology of Canada.' Dawson.</td> +</tr><tr><td class="right" valign="top">(38)</td> + <td>"On the Connection between the existing Fauna and Flora of + Great Britain and certain Geological Changes"—'Mem. Geol. + Survey.' Edward Forbes.</td> +</tr><tr><td class="right" valign="top">(39)</td> + <td>'Cavern-Researches.' M'Enery. Edited by Vivian.</td> +</tr><tr><td class="right" valign="top">(40)</td> + <td>"Quaternary Gravels"—'Quart. Journ. Geol. Soc.,' vol. + xxv. Tylor.</td></tr> +</table> + +<h3> +<a name="page_367"><span class="page">Page 367</span></a> +CHAPTER XXIII.</h3> + +<p class="subtitle"> +THE SUCCESSION OF LIFE UPON THE GLOBE. +</p> + +<p class="indent"> +In conclusion, it may not be out of place if we attempt to summarise, +in the briefest possible manner, some of the principal results +which may be deduced as to the succession of life upon the earth +from the facts which have in the preceding portion of this work +been passed in review. That there was a time when the earth was +void of life is universally admitted, though it may be that the +geological record gives us no direct evidence of this. That the +globe of to-day is peopled with innumerable forms of life whose +term of existence has been, for the most part, but as it were +of yesterday, is likewise an assertion beyond dispute. Can we +in any way connect the present with the remote past, and can we +indicate even imperfectly the conditions and laws under which the +existing order was brought about? The long series of fossiliferous +deposits, with their almost countless organic remains, is the +link between what has been and what is; and if any answer to the +above question can be arrived at, it will be by the careful and +conscientious study of the facts of Palæontology. In the present +state of our knowledge, it may be safely said that anything like +a dogmatic or positive opinion as to the precise sequence of +living forms upon the globe, and still more as to the manner in +which this sequence may have been brought about, is incapable of +scientific proof. There are, however, certain general deductions +from the known facts which may be regarded as certainly established. +</p> + +<p class="indent"> +In the first place, it is certain that there has been a +<i>succession</i> of life upon the earth, different specific and +generic types succeeding one another in successive periods. It +follows from this, that the animals and plants with which we are +familiar as living, were not always upon the earth, but that they +have been preceded by numerous races more or less differing from +them. What is true of the species of animals and plants, is true +also of the higher zoological divisions; and it is, in the second +place, quite certain that there has been a similar <i>succession</i> +in the order of appearance of the primary groups ("sub-kingdoms," +"classes," &c.) of animals and vegetables. These great groups did +not all come into existence at once, but they made their appearance +successively. It is true that we cannot be said to be certainly +acquainted with the first <i>absolute</i> +<a name="page_368"><span class="page">Page 368</span></a> +appearance of any great group of animals. No one dare assert +positively that the apparent first appearance of Fishes in the +Upper Silurian is really their first introduction upon the earth: +indeed, there is a strong probability against any such supposition. +To whatever extent, however, future discoveries may push back the +first advent of any or of all of the great groups of life, there is +no likelihood that anything will be found out which will materially +alter the <i>relative</i> succession of these groups as at present +known to us. It is not likely, for example, that the future has +in store for us any discovery by which it would be shown that +Fishes were in existence before Molluscs, or that Mammals made their +appearance before Fishes. The sub-kingdoms of Invertebrate animals +were all represented in Cambrian times—and it might therefore +be inferred that <i>these</i> had all come simultaneously into +existence; but it is clear that this inference, though incapable +of actual disproof, is in the last degree improbable. Anterior +to the Cambrian is the great series of the Laurentian, which, +owing to the metamorphism to which it has been subjected, has so +far yielded but the singular <i>Eozoön</i>. We may be certain, +however, that others of the Invertebrate sub-kingdoms besides +the Protozoa were in existence in the Laurentian period; and we +may infer from known analogies that they appeared successively, +and not simultaneously. +</p> + +<p class="indent"> +When we come to smaller divisions than the sub-kingdoms—such +as classes, orders, and families—a similar succession of groups +is observable. The different classes of any given sub-kingdom, or +the different orders of any given class, do not make their +appearance together and all at once, but they are introduced upon +the earth in <i>succession</i>. More than this, the different classes +of a sub-kingdom, or the different orders of a class, <i>in the +main succeed one another in the relative order of their zoological +rank—the lower groups appearing first and the higher groups +last</i>. It is true that in the Cambrian formation—the +earliest series of sediments in which fossils are abundant—we +find numerous groups, some very low, others very high, in the +zoological scale, which <i>appear</i> to have simultaneously +flashed into existence. For reasons stated above, however, we +cannot accept this appearance as real; and we must believe that +many of the Cambrian groups of animals really came into being long +before the commencement of the Cambrian period. At any rate, in the +long series of fossiliferous deposits of later date than the Cambrian +the above-stated rule holds good as a broad generalisation—that +the lower groups, namely, precede the higher in point of time; +<a name="page_369"><span class="page">Page 369</span></a> +and though there are apparent exceptions to the rule, there are none +of such a nature as not to admit of explanation. Some of the leading +facts upon which this generalisarion is founded will be enumerated +immediately; but it will be well, in the first place, to consider +briefly what we precisely mean when we speak of "higher" and "lower" +groups. +</p> + +<p class="indent"> +It is well known that naturalists are in the habit of "classifying" +the innumerable animals which now exist upon the globe; or, in +other words, of systematically arranging them into groups. The +precise arrangement adopted by one naturalist may differ in minor +details from that adopted by another; but all are agreed as to the +fundamental points of classification, and all, therefore, agree in +placing certain groups in a certain sequence. What, then, is the +principle upon which this sequence is based? Why, for example, are +the Sponges placed below the Corals; these below the Sea-urchins; +and these, again, below the Shell-fish? Without entering into +a discussion of the principles of zoological classification, +which would here be out of place, it must be sufficient to say +that the sequence in question is based upon the <i>relative type +of organisation</i> of the groups of animals classified. The +Corals are placed above the Sponges upon the ground that, regarded +as a whole, the <i>plan or type of structure</i> of a Coral is +more complex than that of a Sponge. It is not in the slightest +degree that the Sponge is in any respect less highly organised +or less perfect, as a Sponge, than is the Coral as a Coral. Each +is equally perfect in its own way; but the structural pattern +of the Coral is the highest, and therefore it occupies a higher +place in the zoological scale. It is upon this principle, then, +that the primary subdivisions of the animal kingdom (the so-called +"sub-kingdoms") are arranged in a certain order. Coming, again, to +the minor subdivisions (classes, orders, &c.) of each sub-kingdom, +we find a different but entirely analogous principle employed as +a means of classification. The numerous animals belonging to +any given sub-kingdom are formed upon the same fundamental plan +of structure; but they nevertheless admit of being arranged in +a regular series of groups. All the Shell-fish, for example, +are built upon a common plan, this plan representing the ideal +Mollusc; but there are at the same time various groups of the +<i>Mollusca</i>, and these groups admit of an arrangement in a +given sequence. The principle adopted in this case is simply of +<i>the relative elaboration of the common type</i>. The Oyster +is built upon the same ground-plan as the Cuttle-fish; but this +plan is carried out with much greater elaboration, and with many +more complexities, in the latter than in the former: and +<a name="page_370"><span class="page">Page 370</span></a> +in accordance with this, the <i>Cephalopoda</i> constitute a +higher group than the Bivalve Shell-fish. As in the case of +superiority of structural type, so in this case also, it is not +in the least that the Oyster is an <i>imperfect</i> animal. On +the contrary, it is just as perfectly adapted by its +organisation to fill its own sphere and to meet the exigencies +of its own existence as is the Cuttle-fish; but the latter lives +a life which is, physiologically, higher than the former, and its +organisation is correspondingly increased in complexity. +</p> + +<p class="indent"> +This being understood, it may be repeated that, in the main, +the succession of life upon the globe in point of <i>time</i> +has corresponded with the relative order of succession of the +great groups of animals in <i>zoological rank</i>; and some of +the more striking examples of this may be here alluded to. Amongst +the <i>Echinoderms</i>, for instance, the two orders generally +admitted to be the "lowest" in the zoological scale—namely, +the <i>Crinoids</i> and the <i>Cystoids</i>—are likewise the +oldest, both, appearing in the Cambrian, the former slowly dying +out as we approach the Recent period, and the latter disappearing +wholly before the close of the Palæozoic period. Amongst the +<i>Crustaceans</i>, the ancient groups of the Trilobites, Ostracodes, +Phyllopods, Eurypterids, and Limuloids, some of which exist at the +present day, are all "low" types; whereas the highly-organised +Decapods do not make their appearance till near the close of the +Palæozoic epoch, and they do not become abundant till we reach +Mesozoic times. Amongst the <i>Mollusca</i>, those Bivalves which +possess breathing-tubes (the "siphonate" Bivalves) are generally +admitted to be higher than those which are destitute of these +organs (the "asiphonate" Bivalves); and the latter are especially +characteristic of the Palæozoic period, whilst the former abound +in Mesozoic and Kainozoic formations. Similarly, the Univalves +with breathing-tubes and a corresponding notch in the mouth of the +shell ("siphonostomatous" Univalves) are regarded as higher in the +scale than the round-mouthed vegetable-eating Sea-snails, in which +no respiratory siphons exist ("holostomatous" Univalves); but the +latter abound in the Palæozoic rocks—whereas the former +do not make their appearance till the Jurassic period, and their +higher groups do not seem to have existed till the close of the +Cretaceous. The <i>Cephalopods</i>, again—the highest of all +the groups of Mollusca—are represented in the Palæozoic +rocks exclusively by Tetrabranchiate forms, which constitute the +lowest of the two orders of this class; whereas the more highly +specialised Dibranchiates do not make their appearance till the +commencement of the Mesozoic. The Palæozoic +<a name="page_371"><span class="page">Page 371</span></a> +Tetrabranchiates, also, are of a much simpler type than the highly +complex <i>AmmonitidĹ“</i> of the Mesozoic. +</p> + +<p class="indent"> +Similar facts are observable amongst the <i>Vertebrate animals</i>. +The Fishes are the lowest class of Vertebrates, and they are the +first to appear, their first certain occurrence being in the +Upper Silurian; whilst, even if the Lower Silurian and Upper +Cambrian "Conodonts" were shown to be the teeth of Fishes, there +would still remain the enormously long periods of the Laurentian +and Lower Cambrian, during which there were Invertebrates, but no +Vertebrates. The <i>Amphibians</i>, the next class in zoological +order, appears later than the Fishes, and is not represented +till the Carboniferous; whilst its highest group (that of the +Frogs and Toads) does not make its entrance upon the scene till +Tertiary times are reached. The class of the <i>Reptiles</i>, +again, the next in order, does not appear till the Permian, and +therefore not till after Amphibians of very varied forms had +been in existence for a protracted period. The <i>Birds</i> seem +to be undoubtedly later than the Reptiles; but, owing to the +uncertainty as to the exact point of their first appearance, it +cannot be positively asserted that they preceded Mammals, as they +should have done. Finally, the Mesozoic types of <i>Mammals</i> are +mainly, if not exclusively, referable to the <i>Marsupials</i>, +one of the lowest orders of the class; whilst the higher orders +of the "Placental" Quadrupeds are not with certainty known to +have existed prior to the commencement of the Tertiary period. +</p> + +<p class="indent"> +Facts of a very similar nature are offered by the succession of +Plants upon the globe. Thus the vegetation of the Palæozoic +period consisted principally of the lowly-organised groups of +the Cryptogamous or Flowerless plants. The Mesozoic formations, +up to the Chalk, are especially characterised by the naked-seeded +Flowering plants—the Conifers and the Cycads; whilst the +higher groups of the Angiospermous Exogens and Monocotyledons +characterise the Upper Cretaceous and Tertiary rocks. +</p> + +<p class="indent"> +Facts of the above nature—and they could be greatly +multiplied—seem to point clearly to the existence of some +law of progression, though we certainly are not yet in a position +to formulate this law, or to indicate the precise manner in which +it has operated. Two considerations, also, must not be overlooked. +In the first place, there are various groups, some of them highly +organised, which make their appearance at an extremely ancient +date, but which continue throughout geological time almost +unchanged, and certainly unprogressive. Many of these "persistent +types" are known—such as various of the +<a name="page_372"><span class="page">Page 372</span></a> +<i>Foraminifera</i>, the <i>LingulĹ“</i>, the <i>Nautili</i>, +&c.; and they indicate that under given conditions, at present +unknown to us, it is possible for a life-form to subsist for an +almost indefinite period without any important modification of +its structure. In the second place, whilst the facts above +mentioned point to some general law of progression of the great +zoological groups, it cannot be asserted that the primeval types +<i>of any given group</i> are necessarily "lower," zoologically +speaking, than their modern representatives. Nor does this seem to +be at all necessary for the establishment of the law in question. +It cannot be asserted, for example, that the Ganoid and Placoid +Fishes of the Upper Silurian are in themselves less highly organised +than their existing representatives; nor can it even be asserted +that the Ganoid and Placoid orders are low <i>groups</i> of the +class <i>Pisces</i>. On the contrary, they are high groups; but +then it must be remembered that these are probably not really +the first Fishes, and that if we meet with Fishes at some future +time in the Lower Silurian or Cambrian, these may easily prove +to be representatives of the lower orders of the class. This +question cannot be further entered into here, as its discussion +could be carried out to an almost unlimited length; but whilst +there are facts pointing both ways, it appears that at present +we are not justified in asserting that the earlier types of each +group—so far as these are known to us, or really are without +predecessors—are <i>necessarily</i> or <i>invariably</i> more +"degraded" or "embryonic" in their structure than their more +modern representatives. +</p> + +<p class="indent"> +It remains to consider very briefly how far Palæontology +supports the doctrine of "Evolution," as it is called; and this, +too, is a question of almost infinite dimensions, which can but +be glanced at here. Does Palæontology teach us that the +almost innumerable kinds of animals and plants which we know to +have successively flourished upon the earth in past times were +produced separately and wholly independently of each other, at +successive periods? or does it point to the theory that a large +number of these supposed distinct forms, have been in reality +produced by the slow modification of a comparatively small number +of primitive types? Upon the whole, it must be unhesitatingly +replied that the evidence of Palæontology is in favour of +the view that the succession of life-forms upon the globe has +been to a large extent regulated by some orderly and +constantly-acting law of modification and evolution. Upon no +other theory can we comprehend how the fauna of any given +formation is more closely related to that of the formation next +below in the series, and to that of +<a name="page_373"><span class="page">Page 373</span></a> +the formation next above, than to that of any other series of +deposits. Upon no other view can we comprehend why the +Post-Tertiary Mammals of South America should consist +principally of Edentates, Llamas, Tapirs, Peccaries, Platyrhine +Monkeys, and other forms now characterising this continent; +whilst those of Australia should be wholly referable to the +order of Marsupials. On no other view can we explain the common +occurrence of "intermediate" or "transitional" forms of life, +filling in the gaps between groups now widely distinct. +</p> + +<p class="indent"> +On the other hand, there are facts which point clearly to the +existence of some law other than that of evolution, and probably +of a deeper and more far-reaching character. Upon no theory of +evolution can we find a satisfactory explanation for the constant +introduction throughout geological time of new forms of life, which +do not appear to have been preceded by pre-existent allied types; +The Graptolites and Trilobites have no known predecessors, and leave +no known successors. The Insects appear suddenly in the Devonian, +and the Arachnides and Myriapods in the Carboniferous, under +well-differentiated and highly-specialised types. The Dibranchiate +Cephalopods appear with equal apparent suddenness in the older +Mesozoic deposits, and no known type of the Palæozoic period can +be pointed to as a possible ancestor. The <i>HippuritidĹ“</i> +of the Cretaceous burst into a varied life to all appearance +almost immediately after their first introduction into existence. +The wonderful Dicotyledonous flora of the Upper Cretaceous period +similarly surprises us without any prophetic annunciation from +the older Jurassic. +</p> + +<p class="indent"> +Many other instances could be given; but enough has been said +to show that there is a good deal to be said on both sides, and +that the problem is one environed with profound difficulties. +One point only seems now to be universally conceded, and that +is, that the record of life in past time is not interrupted by +gaps other than those due to the necessary imperfections of the +fossiliferous series, to the fact that many animals are incapable +of preservation in a fossil condition, or to other causes of a +like nature. All those who are entitled to speak on this head +are agreed that the introduction of new and the destruction of +old species have been slow and gradual processes, in no sense of +the term "catastrophistic." Most are also willing to admit that +"Evolution" has taken place in the past, to a greater or less +extent, and that a greater or less number of so-called species of +fossil animals are really the modified descendants of pre-existent +forms. <i>How</i> this process of evolution has +<a name="page_374"><span class="page">Page 374</span></a> +been effected, to what extent it has taken place, under what +conditions and laws it has been carried out, and how far it may +be regarded as merely auxiliary and supplemental to some deeper +law of change and progress, are questions to which, in spite of +the brilliant generalisations of Darwin, no satisfactory answer +can as yet be given. In the successful solution of this +problem—if soluble with the materials available to our +hands—will lie the greatest triumph that Palæontology +can hope to attain; and there is reason to think that, thanks to +the guiding-clue afforded by the genius of the author of the +'Origin of Species,' we are at least on the road to a sure, +though it may be a far-distant, victory. +</p> + +<h3> +<a name="page_375"><span class="page">Page 375</span></a> +APPENDIX.</h3> + +<hr /> + +<p class="center"> +TABULAR VIEW OF THE CHIEF DIVISIONS OF THE ANIMAL KINGDOM. +</p> + +<p class="center"> +(Extinct groups are marked with an asterisk. Groups not represented +at all as fossils are marked with two asterisks.) +</p> + +<h4>INVERTEBRATE ANIMALS.</h4> + +<p class="center">SUB-KINGDOM I.—PROTOZOA.</p> + +<p class="indent"> +Animal simple or compound; body composed of "sarcode," not definitely +segmented; no nervous system; and no digestive apparatus, beyond +occasionally a mouth and gullet. +</p> + +<table border="0" cellspacing="0"> +<tr> + <td>CLASS I.</td><td>GREGARINIDÆ.**</td> +</tr><tr> + <td>CLASS II.</td><td>RHIZOPODA.</td> +</tr><tr> + <td> </td> + <td> + <table border="0" cellspacing="0"> + <tr> + <td valign="top" class="center"><i>Order</i></td> + <td>1. <i>Monera</i>.**</td> + </tr><tr> + <td valign="top" class="center"><i>"</i></td> + <td>2. <i>AmĹ“bea</i>.**</td> + </tr><tr> + <td valign="top" class="center"><i>"</i></td> + <td>3. <i>Foraminifera</i>.</td> + </tr><tr> + <td valign="top" class="center"><i>"</i></td> + <td>4. <i>Radiolaria</i> (Polycystines, &c.)</td> + </tr><tr> + <td valign="top" class="center"><i>"</i></td> + <td>5. <i>Spongida</i> (Sponges).</td> + </tr> + </table> + </td> +</tr><tr> + <td>CLASS III.</td><td>INFUSORIA.**</td> +</tr> +</table> + +<p class="center">SUB-KINGDOM II.—CĹ’LENTERATA.</p> + +<p class="indent"> +Animal simple or compound; body-wall composed of two principal +layers; digestive canal freely communicating with the general +cavity of the body; no circulating organs, and no nervous system +or a rudimentary one; mouth surrounded by tentacles, arranged, +like the internal organs, in a "radiate" or star-like manner. +</p> + +<table border="0" cellspacing="0"> +<tr> + <td>CLASS I.</td><td>HYDROZOA.</td> +</tr><tr> + <td> </td> + <td> + <table border="0" cellspacing="0"> + <tr><td valign="top"><i>Sub-class</i></td> + <td>1. <i>Hydroida</i> ("Hydroid Zoophytes"). <i>Ex.</i> + Fresh-water Polypes,** Pipe-corallines (<i>Tubularia</i>), + Sea-Firs (<i>Sertularia</i>).</td></tr> + <tr><td valign="top"><i>Sub-class</i></td> + <td>2. <i>Siphonophora</i>** ("Oceanic Hydrozoa"). + <i>Ex</i>. Portuguese Man-of-war (<i>Physalia</i>).</td></tr> + <tr><td valign="top"> +<a name="page_376"><span class="page">Page 376</span></a> + <i>Sub-class</i></td> + <td>3. <i>Discophora</i> ("Jelly-fishes"). Only known as + fossils by impressions of their stranded carcasses.</td></tr> + <tr><td valign="top"><i>Sub-class</i></td> + <td>4. <i>Lucernarida</i> ("Sea-blubbers"). Also only known as + fossils by impressions left in fine-grained strata.</td></tr> + <tr><td valign="top"><i>Sub-class</i></td> + <td>5. <i>GraptolitidĹ“</i>* ("Graptolites").</td></tr> + </table> + </td> +</tr><tr> + <td>CLASS II.</td><td>ACTINOZOA.</td> +</tr><tr> + <td> </td> + <td> + <table border="0" cellspacing="0"> + <tr><td valign="top" class="center"><i>Order</i></td> + <td>1. <i>Zoantharia</i>. <i>Ex</i>. Sea-anemones** + (<i>ActinidĹ“</i>), Star-corals + (<i>AstrĹ“idĹ“</i>).</td></tr> + <tr><td valign="top" class="center"><i>Order</i></td> + <td>2. <i>Alcyonaria</i>. <i>Ex</i>. Sea-pens + (<i>Pennatula</i>), Organ-pipe Coral (<i>Tubipora</i>), + Red Coral (<i>Corallium</i>).</td></tr> + <tr><td valign="top" class="center"><i>Order</i></td> + <td>3. <i>Rugosa</i> ("Rugose Corals").</td></tr> + <tr><td valign="top" class="center"><i>"</i></td> + <td>4. <i>Ctenophora</i>.** <i>Ex</i>. Venus's Girdle + (<i>Cestum</i>).</td></tr> + </table> + </td> +</tr> +</table> + +<p class="center">SUB-KINGDOM III.—ANNULOIDA.</p> + +<p class="indent"> +Animals in which the digestive canal is completely shut off from +the cavity of the body; a distinct nervous system; a system of +branched "water-vessels," which usually communicate with the +exterior. Body of the adult often "radiate," and never composed +of a succession of definite rings. +</p> + +<table border="0" cellspacing="0"> +<tr> + <td>CLASS I.</td><td>ECHINODERMATA.</td> +</tr><tr> + <td> </td> + <td> + <table border="0" cellspacing="0"> + <tr><td valign="top"><i>Order</i></td> + <td>1. <i>Crinoidea</i> ("Sea-lilies"). <i>Ex</i>. + Feather-star (<i>Comatula</i>), Stone-lily + (<i>Encrinus</i>*).</td></tr> + <tr><td valign="top"><i>Order</i></td> + <td>2. <i>Blastoidea</i>* ("Pentremites").</td></tr> + <tr><td valign="top" class="center"><i>"</i></td> + <td>3. <i>Cystoidea</i>* ("Globe-lilies").</td></tr> + <tr><td valign="top" class="center"><i>"</i></td> + <td>4. <i>Ophiuroidea</i> ("Brittle-stars"). + <i>Ex</i>.</td></tr> + <tr><td valign="top" class="center"><i>"</i></td> + <td>Sand-stars (<i>Ophiura</i>), Brittle-stars + (<i>Ophiocoma</i>).</td></tr> + <tr><td valign="top"><i>Order</i></td> + <td>5. <i>Asteroidea</i> ("Star-fishes"). Ex. Cross-fish + (<i>Uraster</i>), Sun-star (<i>Solaster</i>).</td></tr> + <tr><td valign="top"><i>Order</i></td> + <td>6. <i>Echinoidea</i> ("Sea-urchins"). Ex. Sea-eggs + (<i>Echinus</i>), Heart-urchins (<i>Spatangus</i>).</td></tr> + <tr><td valign="top"><i>Order</i></td> + <td>7. <i>Holothuroidea</i> ("Sea-cucumbers"). <i>Ex</i>. + Trepangs (<i>Holothuria</i>).</td></tr> + </table> + </td> +</tr><tr> + <td>CLASS II.</td><td>SCOLECIDA** (Intestinal Worms, Wheel + Animalcules, &c.)</td> +</tr> +</table> + +<p class="center">SUB-KINGDOM IV.—ANNULOSA.</p> + +<p class="indent"> +Animal composed of numerous definite segments placed one behind +the other; nervous system forming a knotted cord placed along +the lower (ventral) surface of the body. +</p> + +<p class="center"> +<i>Division A. Anarthropoda</i>. No jointed limbs. +</p> + +<table border="0" cellspacing="0"> +<tr> + <td>CLASS I.</td><td>GEPHYREA** ("Spoon-worms").</td> +</tr><tr> + <td>CLASS II.</td><td>ANNELIDA. ("Ringed-worms").</td> +</tr><tr> + <td> </td> + <td><i>Ex</i>. Leeches** (<i>Hirudinea</i>), Earthworms** + (<i>OligochĹ“ta</i>), Tube-worms (<i>Tubicola</i>), + Sea-worms and Sea-centipedes (<i>Errantia</i>).</td> +</tr><tr> + <td>CLASS III.</td><td>CHÆTOGNATHA** + ("Arrow-worms").</td> +</tr> +</table> + +<p class="center"> +<i>Division B. Arthropoda or Articulata</i>. Limbs jointed to +the body. +</p> + + +<table border="0" cellspacing="0"> +<tr> + <td>CLASS I.</td><td>CRUSTACEA ("Crustaceans").</td> +</tr><tr> + <td> </td> + <td><i>Ex</i>. Barnacles and Acorn-shells (<i>Cirripedia</i>), + Water-fleas (<i>Ostracoda</i>), Brine-shrimps and Fairy-shrimps + (<i>Phyllopoda</i>), Trilobites* (<i>Trilobita</i>), King-crabs + and Eurypterids* (<i>Merostomata</i>), Wood-lice and Slaters + (<i>Isopoda</i>), Sand-hoppers (<i>Amphipoda</i>), Lobsters, + Shrimps, Hermit-crabs, and Crabs (<i>Decapoda</i>).</td> +</tr><tr> + <td>CLASS II.</td><td> +<a name="page_377"><span class="page">Page 377</span></a> + ARACHNIDA.</td> +</tr><tr> + <td> </td> + <td><i>Ex.</i> Mites (<i>Acarina</i>), Scorpions + (<i>Pedipalpi</i>), Spiders (<i>Araneida</i>).</td> +</tr><tr> + <td>CLASS III.</td><td>MYRIAPODA.</td> +</tr><tr> + <td> </td> + <td><i>Ex.</i> Centipedes (<i>Chilopoda</i>), Millipedes and + Galley-worms (<i>Chilignatha</i>).</td> +</tr><tr> + <td>CLASS IV.</td><td>INSECTA ("Insects").</td> +</tr><tr> + <td> </td> + <td><i>Ex</i>. Field-bugs (<i>Hemiptera</i>); Crickets, + Grasshoppers, &c. (<i>Orthoptera</i>); Dragon-flies and + May-flies (<i>Neuroptera</i>); Goats and House-flies + (<i>Diptera</i>); Butterflies and Moths (<i>Lepidoptera</i>); + Bees, Wasps, and Ants (<i>Hymenoptera</i>); Beetles + (<i>Coleoptera</i>).</td> +</tr> +</table> + +<p class="center">SUB-KINGDOM V.—MOLLUSCA.</p> + +<p class="indent"> +Animal soft-bodied, generally with a hard covering or shell; no +distinct segmentation of the body; nervous system of scattered +masses. +</p> + +<table border="0" cellspacing="0"> +<tr> + <td>CLASS I.</td><td>POLYZOA ("Sea-Mosses").</td> +</tr><tr> + <td> </td> + <td><i>Ex</i>. Sea-mats (<i>Flustra</i>), Lace-corals + (<i>FenestellidĹ“</i>*).</td> +</tr><tr> + <td>CLASS II.</td><td>TUNICATA** ("Tunicaries").</td> +</tr><tr> + <td> </td> + <td><i>Ex</i>. Sea-squirts (<i>Ascidia</i>).</td> +</tr><tr> + <td>CLASS III.</td><td>BRACHIOPODA ("Lamp-shells").</td> +</tr><tr> + <td> </td> + <td><i>Ex</i>. Goose-bill Lamp-shell (<i>Lingula</i>).</td> +</tr><tr> + <td>CLASS IV.</td><td>LAMELLIBRANCHIATA ("Bivalves").</td> +</tr><tr> + <td> </td> + <td><i>Ex</i>. Oyster (<i>Ostrea</i>), Mussel (<i>Mytilus</i>), + Scallop (<i>Pecten</i>), Cockle (<i>Cardium</i>).</td> +</tr><tr> + <td>CLASS V.</td><td>GASTEROPODA ("Univalves").</td> +</tr><tr> + <td> </td> + <td><i>Ex</i>. Whelks (<i>Buccinum</i>), Limpets + (<i>Patella</i>), Sea-slugs** (<i>Doris</i>), Land-snails + (<i>Helix</i>).</td> +</tr><tr> + <td>CLASS VI.</td><td>PTEROPODA ("Winged Snails").</td> +</tr><tr> + <td> </td> + <td>Ex. <i>Hyalea, Cleodora</i>.</td> +</tr><tr> + <td>CLASS VII.</td><td>CEPHALOPODA ("Cuttle-fishes").</td> +</tr><tr> + <td> </td> + <td><i>Ex</i>. Calamary (<i>Loligo</i>), Poulpe (<i>Octopus</i>), + Paper Nautilus (<i>Arganauta</i>), Pearly Nautilus + (<i>Nautilus</i>), Belemnites,* Orthoceratites,* + Ammonites.*</td> +</tr> +</table> + +<h4>VERTEBRATE ANIMALS.</h4> + +<p class="center">SUB-KINGDOM VI.—VERTEBRATA.</p> + +<p class="indent"> +Body composed of definite segments arranged longitudinally one +behind the other; main masses of the nervous system placed dorsally; +a backbone or "vertebral column" in the majority. +</p> + +<table border="0" cellspacing="0"> +<tr> + <td>CLASS I.</td><td>PISCES ("Fishes").</td> +</tr><tr> + <td> </td> + <td><i>Ex</i>. Lancelet** (<i>Amphioxus</i>); Lampreys and Hag-fishes + (<i>Marsipobranchii</i>**); Herring, Salmon, Perch, &c. + (<i>Teleostei</i> or "Bony Fishes"); Gar-pike, Sturgeon, &c. + (<i>Ganoidei</i>); Sharks, Dog-fishes, Rays, &c. + (<i>Elasmobranchii</i> or "Placoids").</td> +</tr><tr> + <td>CLASS II.</td><td>AMPHIBIA ("Amphibians").</td> +</tr><tr> + <td> </td> + <td>Ex. <i>Labyrinthodontia</i>,* Cæcilians,** Newts and + Salamanders (<i>Urodela</i>), Frogs and Toads (<i>Anoura</i>).</td> +</tr><tr> + <td>CLASS III.</td><td>REPTILIA ("Reptiles").</td> +</tr><tr> + <td> </td> + <td>Ex. <i>Deinosauria</i>,* <i>Pterosauria</i>,* <i>Anomodontia</i>,* + Plesiosaurs (<i>Sauropterygia</i>*), Ichthyosaurs + (<i>Ichthyopterygia</i>*), Tortoises and Turtles (<i>Chelonia</i>), + Snakes (<i>Ophidia</i>), Lizards (<i>Lacertilia</i>), Crocodiles + (<i>Crocodilia</i>).</td> +</tr><tr> + <td>CLASS IV.</td><td>AVES ("Birds").</td> +</tr><tr> + <td> </td> + <td><i>Ex</i>. Toothed Birds (<i>Odontornithes</i>*); Lizard-tailed + Birds (<i>ArchĹ“opteryx</i>*); Ducks, Geese, Gulls, &c. + (<i>Natatores</i>); Storks, Herons, Snipes, Plovers, &c. + (<i>Grallatores</i>); Ostrich, Emeu, Cassowary, Dinornis,* + Æpiornis,* &c. (<i>Cursores</i>); Fowls, Game Birds, and + Doves (<i>Rasores</i>); Cuckoos, Woodpeckers, Parrots, &c. + (<i>Scansores</i>); Crows, Starlings, Finches, Hummingbirds, + Swallows, &c. (<i>Insessores</i>); Owls, Hawks, Eagles, Vultures + (<i>Raptores</i>).</td> +</tr><tr> + <td>CLASS V.</td><td> +<a name="page_378"><span class="page">Page 378</span></a> + MAMMALIA ("Quadrupeds").</td> +</tr><tr> + <td> </td> + <td><i>Ex</i>. Duck-mole and Spiny Ant-eater + (<i>Monotremata</i>**); Kangaroos, Phalangers, Opossums, + Tasmanian Devil, &c. (<i>Marsupialia</i>); Sloths, Ant-eaters, + Armadillos (<i>Edentata</i>); Manatees and Dugongs + (<i>Sirenia</i>); Whales, Dolphins, Porpoises (<i>Cetacea</i>); + Rhinoceros, Tapir, Horses, Hippopotamus, Pigs, Camels and + Llamas, Giraffes, Deer, Antelopes, Sheep, Goats, Oxen + (<i>Ungulata</i>); Hyrax (<i>Hyracoidea</i>**); Elephants, + Mastodon,* Deinotherium* (<i>Proboscidea</i>); Seals, + Walrus, Bears, Dogs, Wolves, Cats, Lions, Tigers, &c. + (<i>Carnivora</i>); Hares, Rabbits, Porcupines, Beavers, + Rats, Mice, Lemmings, Squirrels, Marmots, &c. (<i>Rodentia</i>); + Bats (<i>Cheiroptera</i>); Moles, Shrew-mice, Hedgehogs + (<i>Insectivora</i>); Lemurs, Spider-monkeys, Macaques, + Baboons, Apes (<i>Quadrumana</i>); Man (<i>Bimana</i>).</td> +</tr> +</table> + +<h3> +<a name="page_379"><span class="page">Page 379</span></a> +GLOSSARY.</h3> + +<hr /> + +<p> +ABDOMEN (Lat. <i>abdo</i>, I conceal). The posterior cavity of +the body, containing the intestines and others of the viscera. In +many Invertebrates there is no separation of the body-cavity into +thorax and abdomen, and it is only in the higher <i>Annulosa</i> +that a distinct abdomen can be said to exist.<br/> +ABERRANT (Lat. <i>aberro</i>, I wander away). Departing from the +regular type.<br/> +ABNORMAL (Lat. <i>ab</i>, from; <i>norma</i>, a rule). Irregular; +deviating from the ordinary standard.<br/> +ACRODUS (Gr. <i>akros</i>, high; <i>odous</i>, tooth). A genus +of the Cestraciont fishes, so called from the elevated teeth.<br/> +ACROGENS (Gr. <i>akros</i>, high; <i>gennao</i>, I produce). +Plants which increase in height by additions made to the summit +of the stem by the union of the bases of the leaves.<br/> +ACROTRETA (Gr. <i>akros</i>, high; <i>tretos</i>, pierced). A +genus of Brachiopods, so called from the presence of a foramen +at the summit of the shell.<br/> +ACTINOCRINUS (Gr. <i>aktin</i>, a ray; <i>krinon</i>, a lily). +A genus of Crinoids.<br/> +ACTINOZOA (Gr. <i>aktin</i>, a ray; and <i>zoön</i>, an animal). +That division of the <i>CĹ“lenterata</i> of which the +Sea-anemones may be taken as the type.<br/> +ÆGLINA (<i>ÆglĂ©</i>, a sea-nymph). A genus of +Trilobites.<br/> +ÆPIORNIS (Gr. <i>aipus</i>, huge; <i>ornis</i>, bird). A genus +of gigantic Cursorial birds.<br/> +AGNOSTUS (Gr. <i>a</i>, not; <i>gignosko</i>, I know). A genus +of Trilobites.<br/> +ALCES (Lat. <i>alces</i>, elk). The European Elk or Moose.<br/> +ALECTO (the proper name of one of the Furies). A genus of +<i>Polyzoa</i>.<br/> +ALETHOPTERIS (Gr. <i>alethes</i>, true; <i>pteris</i>, fern). +A genus of Ferns.<br/> +ALGÆ. (Lat. <i>alga</i>, a marine plant). The order of plants +comprising the Sea-weeds and many fresh-water plants.<br/> +ALVEOLUS (Lat. <i>alvus</i>, belly). Applied to the sockets of +the teeth.<br/> +AMBLYPTERUS (Gr. <i>amblus</i>, blunt; <i>pteron</i>, fin). An +order of Ganoid Fishes.<br/> +AMBONYCHIA (Gr. <i>ambon</i>, a boss; <i>onux</i>, claw). A genus +of Palæozoic Bivalves.<br/> +AMBULACRA (Lat. <i>ambulacrum</i>, a place for walking). The +perforated spaces or "avenues" through which are protruded the +tube-feet, by means of which locomotion is effected in the +<i>Echinodermata</i>.<br/> +AMMONITIDÆ. A family of Tetrabranchiate Cephalopods, so called +from the resemblance of the shell of the type-genus, <i>Ammonites</i>, +to the horns of the Egyptian God, Jupiter-Ammon.<br/> +AMORPHOZOA (Gr. <i>a</i>, without; <i>morphe</i>, shape; +<i>zoön</i>, animal). A name sometimes used to designate the +<i>Sponges</i>.<br/> +AMPHIBIA (Gr. <i>amphi</i>, both; <i>bios</i>, life). The Frogs, +Newts, and the like, which have gills when young, but can always +breathe air directly when adult.<br/> +AMPHICYON (Gr. <i>amphi</i>, both—implying doubt; <i>kuon</i>, +dog). An extinct genus of <i>Carnivora</i>.<br/> +<a name="page_380"><span class="page">Page 380</span></a> +AMPHILESTES (Gr. <i>amphi</i>, both; <i>lestes</i>, a thief). A +genus of Jurassic Mammals.<br/> +AMPHISPONGIA (Gr. <i>amphi</i>, both; <i>spoggos</i>, sponge). +A genus of Silurian sponges.<br/> +AMPHISTEGINA (Gr. <i>amphi</i>, both; <i>stegĂ©</i>, roof). A +genus of <i>Foraminifera</i>.<br/> +AMPHITHERIUM (Gr. <i>amphi</i>, both; <i>therion</i>, beast). +A genus of Jurassic Mammals.<br/> +AMPHITRAGULUS (Gr. <i>amphi</i>, both; dim. of <i>tragos</i>, +goat). An extinct genus related to the living Musk-deer.<br/> +AMPLEXUS (Lat. an Ambrace). A genus of Rugose Corals.<br/> +AMPYX (Gr. <i>ampux</i>, a wreath or wheel). A genus of +Trilobites.<br/> +ANARTHROPODA (Gr. <i>a</i>. without; <i>arthros</i>, a joint; +<i>pous</i>, foot). That division of <i>Annulose</i> animals +in which there are no articulated appendages.<br/> +ANCHITHERIUM (Gr. <i>agchi</i>, near; <i>therion</i>, beast). An +extinct genus of Mammals.<br/> +ANCYLOCERAS (Gr. <i>agkulos</i>, crooked; <i>ceras</i>, horn). +A genus of <i>AmmonitidĹ“</i>.<br/> +ANCYLOTHERIUM (Gr. <i>agkulos</i>, crooked; <i>therion</i>, beast). +An extinct genus of Edentate Mammals.<br/> +ANDRIAS (Gr. <i>andrias</i>, image of man). An extinct genus of +tailed Amphibians.<br/> +ANGIOSPERMS (Gr. <i>angeion</i>, a vessel; <i>sperma</i>, seed). +Plants which have their seeds enclosed in a seed-vessel.<br/> +ANNELIDA (a Gallicised form of <i>Annulata</i>). The Ringed Worms, +which form one of the divisions of the <i>Anarthropoda</i>.<br/> +ANNULARIA (Lat. <i>annulus</i>, a ring). A genus of Palæozoic +plants, with leaves in whorls.<br/> +ANNULOSA (Lat. <i>annulus</i>). The sub-kingdom comprising the +<i>Anarthropoda</i> and the <i>Arthropoda</i> or <i>Articulata</i>, +in all of which the body is more or less evidently composed of +a succession of rings.<br/> +ANOMODONTIA (Gr. <i>anomos</i>, irregular; <i>odous</i>, tooth). +An extinct order of Reptiles, often called <i>Dicynodontia</i>.<br/> +ANOMURA (Gr. <i>anomos</i>, irregular; <i>oura</i>, tail). A +tribe of Decapod <i>Crustacea</i>, of which the Hermit-crab is +the type.<br/> +ANOPLOTHERIDÆ (Gr. <i>anoplos</i>, unarmed; <i>ther</i>, beast). +A family of Tertiary Ungulates.<br/> +ANOURA (Gr. <i>a</i>, without; <i>oura</i>, tail). The order +of <i>Amphibia</i> comprising the Frogs and Toads, in which the +adult is destitute of a tail. Often, called <i>Batrachia</i>.<br/> +ANTENNÆ (Lat. <i>antenna</i>, a yard-arm). The jointed horns +or feelers possessed by the majority of the <i>Articulata</i>.<br/> +ANTENNULES (dim. of <i>AntennĹ“</i>). Applied to the smaller +pair of antennæ in the <i>Crustacea</i>.<br/> +ANTHRACOSAURUS (Gr. <i>anthrax</i>, coal; <i>saura</i>, lizard). +A genus of Labyrinthodont Amphibians.<br/> +ANTHRAPALÆMON (Gr. <i>anthrax</i>, coal; <i>palĹ“mon</i>, a +prawn—originally a proper name). A genus of long-tailed +Crustaceans from the Coal-measures.<br/> +ANTLERS. Properly the branches of the horns of the Deer tribe +(<i>CervidĹ“</i>), but generally applied to the entire horns.<br/> +APIOCRINIDÆ (Gr. <i>apion</i>, a pear; <i>krinon</i>, lily). A +family of Crinoids—the "Pear-encrinites."<br/> +APTERYX (Gr. <i>a</i>, without; <i>pterux</i>, a wing). A wingless +bird of New Zealand, belong to the order <i>Cursores</i>.<br/> +AQUEOUS (Lat. <i>aqua</i>, water). Formed in or by water.<br/> +ARACHNIDA (Gr. <i>arachne</i>, a spider). A class of the +<i>Articulata</i>, comprising Spiders, Scorpions, and allied +animals.<br/> +ARBORESCENT. Branched like a tree.<br/> +ARCHÆOCIDARIS (Gr. <i>archaios</i>, ancient; Lat. <i>cidaris</i>, +a diadem). A Palæozoic genus of Sea-urchins, related to the +existing <i>Cidaris</i>.<br/> +ARCHÆOCYATHUS (Gr. <i>archaios</i>, ancient; <i>kuathos</i>, cup). +A genus of Palæozoic fossils allied to the Sponges.<br/> +ARCHÆOPTERYX (Gr. <i>archaios</i>, ancient; <i>pterux</i>, a +wing). The singular fossil bird which alone constitutes the order +of the <i>SaururĹ“</i>.<br/> +<a name="page_381"><span class="page">Page 381</span></a> +ARCTOCYON (Gr. <i>arctos</i>, bear; <i>kuon</i>, dog). An extinct +genus of Carnivora.<br/> +ARENACEOUS. Sandy, or composed of grains of sand.<br/> +ARENICOLITES (Lat. <i>arena</i>, sand; <i>colo</i>, I inhabit). A +genus founded on burrows supposed to be formed by worms resembling +the living Lobworms (<i>Arenicola</i>).<br/> +ARTICULATA (Lat. <i>articulus</i>, a joint). A division of the +animal kingdom, comprising Insects, Centipedes, Spiders, and +Crustaceans, characterised by the possession of jointed bodies +or jointed limbs. The term <i>Arthropoda</i> is now more usually +employed.<br/> +ARTIODACTYLA (Gr. <i>artios</i>, even; <i>daktulos</i>, a finger +or toe). A division of the hoofed quadrupeds (<i>Ungulata</i>) +in which each foot has an even number of toes (two or four).<br/> +ASAPHUS (Gr. <i>Asaphes</i>, obscure). A genus of Trilobites.<br/> +ASCOCERAS (Gr. <i>askos</i>, a leather bottle; <i>keras</i>, horn). +A genus of Tetrabranchiate Cephalopods.<br/> +ASIPHONATE. Not possessing a respiratory tube or siphon. (Applied +to a division of the <i>Lamellibranchiate</i> Molluscs.)<br/> +ASTEROID (Gr. <i>aster</i>, a star; and <i>eidos</i>, form). +Star-shaped, or possessing radiating lobes or rays like a +star-fish.<br/> +ASTEROIDEA. An order of <i>Echinodermata</i>, comprising the +Star-fishes, characterised by their rayed form.<br/> +ASTEROPHYLLITES (Gr. <i>aster</i>, a star; <i>phullon</i>, leaf). +A genus of Palæozoic plants, with leaves in whorls.<br/> +ASTRÆIDÆ (Gr. <i>AstrĹ“a</i>, a proper name). +The family of the Star-corals.<br/> +ASTYLOSPONGIA (Gr. <i>a</i>, without; <i>stulos</i>, a column; +<i>spoggos</i>, a sponge). A genus of Silurian Sponges.<br/> +ATHYRIS (Gr. <i>a</i>, without; <i>thura</i>, door). A genus of +Brachiopods.<br/> +ATRYPA (Gr. <i>a</i>, without; <i>trupa</i>, a hole). A genus +of Brachiopods.<br/> +AVES (Lat. <i>avis</i>, a bird). The class of the Birds.<br/> +AVICULA (Lat. a little bird). The genus of Bivalve Molluscs +comprising the Pearl-oysters.<br/> +AXOPHYLLUM (Gr. <i>axon</i>, a pivot; <i>phullon</i>, a leaf). +A genus of Rugose Corals.<br/> +AZOIC (Gr. <i>a</i>, without; <i>zoĂ©</i>, life). Destitute of +traces of living beings.<br/> +</p> + +<p> +BACULITES (Lat. <i>baculum</i>, a staff). A genus of the +<i>AmmonitidĹ“</i>.<br/> +BALÆNA (Lat. a whale). The genus of the Whalebone Whales.<br/> +BALANIDÆ (Gr. <i>balanos</i>, an acorn). A family of sessile +<i>Cirripedes</i>, commonly called "Acorn-shells."<br/> +BATRACHIA (Gr. <i>batrachos</i>, a frog). Often loosely applied +to any of the <i>Amphibia</i>, but sometimes restricted to the +Amphibians as a class, or to the single order of the +<i>Anoura</i>.<br/> +BELEMNITIDÆ (Gr. <i>belemnon</i>, a dart). An extinct group +of Dibranchiate Cephalopods, comprising the Belemnites and their +allies.<br/> +BELEMNOTEUTHIS (Gr. <i>belemnon</i>, a dart; <i>teuthis</i>, a +cuttle-fish). A genus allied to the Belemnites proper.<br/> +BELINURUS (Gr. <i>belos</i>, a dart; <i>oura</i>, tail). A genus +of fossil King-crabs.<br/> +BELLEROPHON (Gr. proper name). A genus of oceanic Univalves +(<i>Heteropoda</i>).<br/> +BELOTEUTHIS (Gr. <i>belos</i>, a dart; <i>teuthis</i>, a +cuttle-fish). An extinct genus of Dibranchiate Cephalopods.<br/> +BEYRICHIA (named after Prof. Beyrich). A genus of Ostracode +Crustaceans.<br/> +BILATERAL. Having two symmetrical sides.<br/> +BIMANA (Lat. <i>Bis</i>, twice; <i>manus</i>, a hand). The order +of <i>Mammalia</i> comprising man alone.<br/> +BIPEDAL (Lat. <i>bis</i>, twice; <i>pes</i>, foot). Walking upon +two legs.<br/> +BIVALVE (Lat. <i>bis</i>, twice; <i>valvĹ“</i>, folding-doors). +Composed of two plates or valves; applied to the shell of the +<i>Lamellibranchiata</i> and <i>Brachiopoda</i>, and to the +carapace of certain <i>Crustacea</i>.<br/> +BLASTOIDEA (Gr. <i>blastos</i>, a bud; and <i>eidos</i>, form). +An extinct order of <i>Echinodermata</i>, often called +<i>Pentremites</i>.<br/> +BRACHIOPODA (Gr. <i>brachion</i>, an arm; <i>pous</i>, the foot). +<a name="page_382"><span class="page">Page 382</span></a> +A class or the <i>Molluscoida</i>, often called "Lamp-shells," +characterised by possessing two fleshy arms continued from the sides +of the mouth.<br/> +BRACHYURA (Gr. <i>brachus</i>, short; <i>oura</i>, tail). A tribe +of the Decapod <i>Crustaceans</i> with short tails (<i>i.e.</i>, +the Crabs).<br/> +BRADYPODIDÆ. (Gr. <i>bradus</i>, slow; <i>podes</i>, feet). The +family of <i>Edentata</i> comprising the Sloths.<br/> +BRANCHIA (Gr. <i>bragchia</i>, the gill of a fish). A respiratory +organ adapted to breathe air dissolved in water.<br/> +BRANCHIATE. Possessing gills or branchiæ.<br/> +BRONTEUS (Gr. <i>bronĂ©</i>, thunder—an epithet of Jupiter +the Thunderer). A genus of Trilobites.<br/> +BRONTOTHERIUM (Gr. <i>brontĂ©</i>, thunder; <i>therion</i> beast). +An extinct genus of Ungulate Quadrupeds.<br/> +BRONTOZOUM (Gr. <i>brontĂ©</i>, thunder; <i>zoön</i>, animal). +A genus founded on the largest footprints of the Triassic Sandstones +of Connecticut.<br/> +BUCCINUM (Lat. <i>buccinun</i>, a trumpet). The genus of Univalves +comprising the Whelks. +</p> + +<p> +CAINOZOIC (<i>See</i> Kainozoic.)<br/> +CALAMITES (Lat. <i>calamus</i>, a reed). Extinct plants with +reed-like stems, believed to be gigantic representatives of the +<i>EquisetaceĹ“</i>.<br/> +CALCAREOUS (Lat. <i>calx</i>, lime). Composed of carbonate of +lime.<br/> +CALICE. The little cup in which the polype of a coralligenous +Zoophyte (<i>Actinozoön</i>) is contained.<br/> +CALYMENE (Gr. <i>kalumenĂ©</i>, concealed). A genus of +Trilobites.<br/> +CALYX (Lat. a cup). Applied to the cup-shaped body of a +<i>Crinoid</i> (<i>Echinodermata</i>).<br/> +CAMAROPHORIA (Gr. <i>kamara</i>, a chamber; <i>phero</i>, I +carry). A genus of Brachiopods.<br/> +CAMELOPARDALIDÆ. (Lat. <i>camelus</i>, a camel; +<i>pardalis</i>, a panther). The family of the Giraffes.<br/> +CANINE (Lat. <i>canis</i>, a dog). The eye-tooth of Mammals, +or the tooth which is placed at or close to the præmaxillary +suture in the upper jaw, and the corresponding tooth in the lower +jaw.<br/> +CARAPACE. A protective shield. Applied to the upper shell of +Crabs, Lobsters, and many other <i>Crustacea</i>. Also the upper +half of the immovable case in which the body of a Chelonian is +protected.<br/> +CARCHARODON (Gr. <i>karcharos</i>. rough; <i>odous</i>, tooth). +A genus of Sharks.<br/> +CARDIOCARPON (Gr. <i>kardia</i>, the heart; <i>karpos</i>, fruit). +A genus of fossil fruit from the Coal-measures.<br/> +CARDIUM (Gr. <i>kardia</i>, the heart). The genus of Bivalve +Molluscs comprising the Cockles. <i>Cardinia, Cardiola</i>, and +<i>Cardita</i> have the same derivation.<br/> +CARNIVORA (Lat. <i>caro</i>, flesh; <i>voro</i>, I devour). An +order of the <i>Mammalia</i>. The "Beasts of Prey."<br/> +CARNIVOROUS (Lat. <i>caro</i>, flesh; <i>voro</i>, I devour). +Feeding upon flesh.<br/> +CARYOCARIS (Gr. <i>karua</i>, a nut; <i>karis</i>, a shrimp). +A genus of Phyllopod Crustaceans.<br/> +CARYOCRINUS (Gr. <i>karua</i>, a nut; <i>krinon</i>, a lily). +A genus of Cystideans.<br/> +CAUDAL (Lat. <i>cauda</i>, the tail). Belonging to the +tail.<br/> +CAVICORNIA (Lat. <i>cavus</i>, hollow; <i>cornu</i>, a horn). +The "hollow-horned" Ruminants, in which the horn consists of +a central bony "horn-core" surrounded by a horny sheath.<br/> +CENTRUM (Gr. <i>kentron</i>, the point round which a circle is +described by a pair of compasses). The central portion or "body" +of a vertebra.<br/> +CEPHALASPIDÆ. (Gr. <i>kephale</i>, head; <i>aspis</i>, +shield). A family of fossil fishes.<br/> +CEPHALIC (Gr. <i>kephale</i>, head). Belonging to the head.<br/> +CEPHALOPODA (Gr. <i>kephale</i>; and <i>podes</i>, feet). A class +of the <i>Mollusca</i>, comprising the Cuttle-fishes and their +allies, in which there is a series of arms ranged round the +head.<br/> +CERATIOCARIS (Gr. <i>keras</i>, a horn; <i>karis</i>, a shrimp). +A genus of Phyllopod Crustaceans.<br/> +<a name="page_383"><span class="page">Page 383</span></a> +CERATITES (Gr. <i>keras</i>, a horn). A genus +of <i>AmmonitidĹ“</i>.<br/> +CERATODUS (Gr. <i>keras</i>, a horn; <i>odous</i>, tooth). A +genus of Dipnoous fishes.<br/> +CERVICAL (Lat. <i>cervix</i>, the neck). Connected with or +belonging to the region of the neck.<br/> +CERVIDÆ (Lat. <i>cervus</i>, a stag). The family of the +Deer.<br/> +CESTRAPHORI (Gr. <i>kestra</i>, a weapon; <i>phero</i>, I carry). +The group of the "Cestraciont Fishes," represented at the present +day by the Port-Jackson Shark; so called from their defensive +spines.<br/> +CETACEA (Gr. <i>ketos</i>, a whale). The order of Mammals +comprising the Whales and the Dolphins.<br/> +CETIOSAURUS (Gr. <i>ketos</i>, whale; <i>saura</i>, lizard). A +genus of Deinosaurian Reptiles.<br/> +CHEIROPTERA (Gr. <i>cheir</i>, hand; <i>pteron</i>, wing). The +Mammalian order of the Bats.<br/> +CHEIROTHERIUM (Gr. <i>cheir</i>, hand; <i>therion</i>, beast). +The generic name applied originally to the hand-shaped footprints +of Labyrinthodonts.<br/> +CHEIRURUS (Gr. <i>cheir</i>, hand; <i>oura</i>, tail). A genus +of Trilobites.<br/> +CHELONIA (Gr. <i>chelonĂ©</i>, a tortoise). The Reptilian +order of the Tortoises and Turtles.<br/> +CHONETES (Gr. <i>chonĂ©</i> or <i>choanĂ©</i>, a +chamber or box). A genus of Brachiopods.<br/> +CIDARIS (Lat. a diadem). A genus of Sea-urchins.<br/> +CLADODUS (Gr. <i>klados</i>, branch; <i>odous</i>, tooth). A genus +of Fishes.<br/> +CLATHROPORA (Lat. <i>clathti</i>, a trellis; <i>porus</i>, a pore). +A genus of Lace-corals (<i>Polyzoa</i>).<br/> +CLISIOPHYLLUM (Gr. <i>klision</i>, a hut; <i>phullon</i>, leaf). +A genus of Rugose Corals.<br/> +CLYMENIA (<i>Clumene</i>, a proper name). A genus of +Tetrabranchiate Cephalopods.<br/> +COCCOSTEUS (Gr. <i>kokkos</i>, berry; <i>osteon</i>, bone). A +genus of Ganoid Fishes.<br/> +COCHLIODUS (Gr. <i>kochlion</i>, a snail-shell; <i>odous</i>, +tooth). A genus of Cestraciont Fishes.<br/> +CĹ’LENTERATA (Gr. <i>koilos</i>, hollow; <i>enteron</i>, the +bowel). The sub-kingdom which comprises the <i>Hydrozoa</i> and +<i>Actinozoa</i>. Proposed by Frey and Leuckhart in place of the +old term <i>Radiata</i>, which included other animals as +well.<br/> +COLEOPTERA (Gr. <i>koleos</i>, a sheath; <i>pteron</i>, wing). +The order of Insects (Beetles) in which the anterior pair of wings +are hardened, and serve as protective cases for the posterior +pair of membranous wings.<br/> +COLOSSOCHELYS (Gr. <i>kolossos</i>, a gigantic statue; <i>chelus</i>, +a tortoise). A huge extinct Land-tortoise.<br/> +COMATULA (Gr. <i>koma</i>, the hair). The Feather-star, so called +in allusion to its tress-like arms.<br/> +CONDYLE (Gr. <i>kondulos</i>, a knuckle). The surface by which +one bone articulates with another. Applied especially to the +articular surface or surfaces by which the skull articulates with +the vertebral column.<br/> +CONIFERÆ (Lat. <i>conus</i>, a cone; <i>fero</i>, I carry). +The order of the Firs, Pines, and their allies, in which the +fruit is generally a "cone" or "fir-apple."<br/> +CONULARIA (Lat. <i>conulus</i>, a little-cone). An extinct genus +of Pteropods.<br/> +COPRALITES (Gr. <i>kopros</i>, dung; <i>lithos</i>, stone). Properly +applied to the fossilised excrements of animals; but often employed +to designate phosphatic concretions which are not of this +nature.<br/> +CORALLITE. The corallum secreted by an <i>Actinozoön</i> which +consists of a single polype; or the portion of a composite corallum +which belongs to, and is secreted by, an individual polype.<br/> +CORALLUM (from the Latin for Red Coral). The hard structures +deposited in, or by the tissues of an +<i>Actinozoön</i>,—commonly called a "coral."<br/> +CORIACEOUS (Lat. <i>corium</i>. hide). Leathery.<br/> +CORYPHODON (Gr. <i>korus</i>, helmet; <i>odous</i>, tooth). An +extinct genus of Mammals, allied to the Tapirs.<br/> +CRANIUM (Gr. <i>kranion</i>, the skull). The bony or cartilaginous +case in which the brain is contained.<br/> +CRETACEOUS (Lat. <i>creta</i>, chalk). The formation which in +Europe contains white chalk as one of its most conspicuous +members.<br/> +<a name="page_384"><span class="page">Page 384</span></a> +CRINOIDEA (Gr. <i>krinon</i>, a lily; <i>eidos</i>, form). An order +of <i>Echinodermata</i>, comprising forms which are usually stalked, +and sometimes resemble lilies in shape.<br/> +CRIOCERAS (Gr. <i>krios</i>, a ram; <i>keras</i>, a horn). A genus +of <i>AmmonitidĹ“</i>.<br/> +CROCODILIA (Gr. <i>krokodeilos</i>, a crocodile). An order of +Reptiles.<br/> +CROSSOPTERYGIDÆ. (Gr. <i>krossotos</i>, a fringe; +<i>pterux</i>, a fin). A sub-order of Ganoids in which the paired +fins possess a central lobe.<br/> +CRUSTACEA (Lat. <i>crusta</i>, a crust). A class of Articulate +animals, comprising Crabs, Lobsters, &c., characterised by the +possession of a hard shell or crust, which they cast +periodically.<br/> +CRYPTOGAMS (Gr. <i>kruptos</i>, concealed; <i>gamos</i>, marriage). +A division of plants in which the organs of reproduction are +obscure and there are no true flowers.<br/> +CTENACANTHUS (Gr. <i>kteis</i>, a comb; <i>akantha</i>, a thorn). +A genus of fossil fishes, named from its fin-spines.<br/> +CTENOID (Gr. <i>kteis</i>, a comb; <i>eidos</i>, form). Applied +to those scales of fishes the hinder margins of which are fringed +with spines or comb-like projections.<br/> +CURSORES (Lat. <i>curro</i>, I run). An order of <i>Aves</i>, +comprising birds destitute of the power of flight, but formed +for running vigorously (<i>e.g.</i>, the Ostrich and Emeu).<br/> +CUSPIDATE. Furnished with small pointed eminences or "cusps."<br/> +CYATHOCRINUS (Gr. <i>kuathos</i>, a cup; <i>krinon</i>, a lily). +A genus of Crinoids.<br/> +CYATHOPHYLLUM (Gr. <i>kuathos</i>, a cup; <i>phullon</i>, a leaf). +A genus of Rugose Corals.<br/> +CYCLOID (Gr. <i>kuklos</i>, a circle; <i>eidos</i>, form). Applied +to those scales of fishes which have a regularly circular or +elliptical outline with an even margin.<br/> +CYCLOPHTHALMUS (Gr. <i>kuklos</i>, a circle; <i>ophthalmos</i>, +eye). A genus of fossil Scorpions.<br/> +CYCLOSTOMI (Gr. <i>kuklos</i>, and <i>stoma</i>, mouth). Sometimes +used to designate the Hag-fishes and Lampreys, forming the order +<i>Marsipobranchii</i>.<br/> +CYPRÆA (a name of Venus). The genus of Univalve Molluscs +comprising the Cowries.<br/> +CYRTOCERAS (Gr. <i>kurtos</i>. crooked; <i>keras</i>, horn). A +genus of Tetrabranchiate Cephalopods.<br/> +CYSTIPHYLLUM (Gr. <i>kustis</i>, a bladder; <i>phullon</i>, a +leaf). A genus of Rugose Corals.<br/> +CYSTOIDEA (Gr. <i>kustis</i>, a bladder; <i>eidos</i>, form). +The "Globe-crinoids," an extinct order of +<i>Echinodermata</i>. +</p> + +<p> +DADOXYLON (Gr. <i>dadion</i>, a torch; <i>xulon</i>, wood). An +extinct genus of Coniferous trees.<br/> +DECAPODA (Gr. <i>deka</i>, ten; <i>podes</i>, feet). The division +of <i>Crustacea</i> which have ten feet; also the family of +Cuttle-fishes, in which there are ten arms or cephalic +processes.<br/> +DECIDUOUS (Lat. <i>decido</i>, I fall off). Applied to parts which +fall off or are shed during the life of the animal.<br/> +DEINOSAURIA (Gr. <i>deinos</i>, terrible; <i>saura</i>, lizard). +An extinct order of Reptiles.<br/> +DEINOTHERIUM (Gr. <i>deinos</i>, terrible; <i>therion</i>, beast). +An extinct genus of Proboscidean Mammals.<br/> +DENDROGRAPTUS (Gr. <i>dendron</i>, tree; <i>grapho</i>, I write). +A genus of Graptolites.<br/> +DESMIDIÆ. Minute fresh-water plants, of a green colour, without +a siliceous epidermis.<br/> +DIATOMACEÆ (Gr. <i>diatemno</i>, I sever). An order of minute +plants which are provided with siliceous envelopes.<br/> +DIBRANCHIATA (Gr. <i>dis</i>; twice; <i>bragchia</i>, gill). +The order of <i>Cephalopoda</i> (comprising the Cuttle-fishes, +&c.) in which only two gills are present.<br/> +DICERAS (Gr. <i>dis</i>, twice; <i>keras</i>, horn). An extinct +genus of Bivalve Molluscs.<br/> +DICTYONEMA (Gr. <i>diktuon</i>, a net; <i>nema</i>, thread). An +extinct genus of <i>Polyzoa</i>.<br/> +<a name="page_385"><span class="page">Page 385</span></a> +DICYNODONTIA (Gr. <i>dis</i>, twice; <i>kuon</i>, dog; <i>odous</i>, +tooth). An extinct order of Reptiles.<br/> +DIDYMOGRAPTUS (Gr. <i>didumos</i>, twin; <i>grapho</i>, I write). +A genus of Graptolites.<br/> +DIMORPHODON (Gr. <i>dis</i>, twice; <i>morphĂ©</i>, shape; +<i>oduos</i>, tooth). A genus of Pterosaurian reptiles.<br/> +DINICHTHYS (Gr. <i>deinos</i>, terrible; <i>ichthus</i>, fish). +An extinct genus of Fishes.<br/> +DINOCERAS (Gr. <i>deinos</i>, terrible; <i>keras</i>, horn). An +extinct genus of Mammals.<br/> +DINOPHIS (Gr. <i>deinos</i>, terrible; <i>ophis</i>, snake). An +extinct genus of Snakes.<br/> +DINORNIS (Gr. <i>deinos</i>, terrible; <i>ornis</i>, bird). An +extinct genus of Birds.<br/> +DIPLOGRAPTUS (Gr. <i>diplos</i>, double; <i>grapho</i>, I write). +A genus of Graptolites.<br/> +DIPNOI (Gr. <i>dis</i>, twice; <i>pnoĂ©</i>, breath). An order +of Fishes, comprising the Mud-fishes, so called in allusion to +their double mode of respiration.<br/> +DIPROTODON (Gr. <i>dis</i>, twice; <i>protos</i>, first; +<i>odous</i>, tooth). A genus of extinct Marsupials.<br/> +DIPTERA (Gr. <i>dis</i>, twice; <i>pteron</i>, wing). An order +of Insects characterised by the possession of two wings.<br/> +DISCOID (Gr. <i>diskos</i>, a quoit; <i>eidos</i>, form). Shaped +like a round plate or quoit.<br/> +DOLOMITE (named after M. Dolomieu). Magnesian limestone.<br/> +DORSAL (Lat. <i>dorsum</i>, the back). Connected with or placed +upon the back.<br/> +DROMATHERIUM (Gr. <i>dromaios</i>, nimble; <i>therion</i>, beast). +A genus of Triassic Mammals.<br/> +DRYOPITHECUS (Gr. <i>drus</i>, an oak; <i>pithekos</i>, an ape). +An extinct genus of Monkeys. +</p> + +<p> +ECHINODERMATA (Gr. <i>echinos</i>; and <i>derma</i>, skin). A +class of animals comprising the Sea-urchins, Star-fishes, and +others, most of which have spiny skins.<br/> +ECHINOIDEA (Gr. <i>echinos</i>; and <i>eidos</i>, form). An order +of <i>Echinodermata</i>, comprising the Sea-urchins.<br/> +EDENTATA (Lat. <i>e</i>, without; <i>dens</i>, tooth). An order +of <i>Mammalia</i> often called <i>Bruta</i>.<br/> +EDENTULOUS. Toothless, without any dental apparatus. Applied to +the mouth of any animal, or to the hinge of the Bivalve +Molluscs.<br/> +ELASMOBRANCHII (Gr. <i>elasma</i>, a plate; <i>bragchia</i>, gill). +An order of Fishes, including the Sharks and Rays.<br/> +ENALIOSAURIA (Gr. <i>enalios</i>, marine; <i>saura</i>, lizard), +Sometimes employed as a common term to designate the extinct +Reptilian orders of the <i>Ichthyosauria</i> and +<i>Plesiosauria</i>.<br/> +EOCENE (Gr. <i>eos</i>, dawn; <i>kainos</i>, new or recent). +The lowest division of the Tertiary rocks, in which species of +existing shells are to a small extent represented.<br/> +EOPHYTON (Gr. <i>eos</i>, dawn; <i>phuton</i>, a plant). A genus +of Cambrian fossils, supposed to be of a vegetable nature.<br/> +EOZOÖN (Gr. <i>eos</i>, dawn; <i>zoön</i>, animal). A +genus of chambered calcareous organisms found in the Laurentian and +Huronian formations.<br/> +EQUILATERAL (Lat. <i>Ĺ“quus</i>, equal; <i>latus</i>, side). +Having its sides equal. Usually applied to the shells of the +<i>Brachiopoda</i>. When applied to the spiral shells of the +<i>Foraminifera</i>, it means that all the convolutions of the +shell lie in the same plane.<br/> +EQUISETACEÆ (Lat. <i>equus</i>, horse; <i>seta</i>, bristle). A +group of Cryptogamous plants, commonly known as "Horse-tails."<br/> +EQUIVALVE (Lat. <i>Ĺ“quus</i>, equal; <i>valvĹ“</i>, +folding-doors). Applied to shells which are composed of two equal +pieces or valves.<br/> +ERRANTIA (Lat. <i>erro</i>, I wander). An order of <i>Annelida</i>, +often called <i>Nereidea</i>, distinguished by their great locomotive +powers.<br/> +EUOMPHALUS (Gr. <i>eu</i>, well; <i>omphalos</i>, navel). An extinct +genus of Univalve Molluscs.<br/> +EURYPTERIDA (Gr. <i>eurus</i>, broad; <i>pteron</i>, wing). An +extinct sub-order of <i>Crustacea</i>.<br/> +EXOGYRA (Gr. <i>exo</i>, outside; <i>guros</i>, circle). An extinct +genus of Oysters. +</p> + +<p> +<a name="page_386"><span class="page">Page 386</span></a> +FAUNA (Lat. <i>Fauni</i>, the rural deities of the Romans). The +general assemblage of the animals of any region or district.<br/> +FAVOSITES (Lat. <i>favus</i>, a honeycomb). A genus of Tabulate +Corals.<br/> +FENESTELLIDÆ. (Lat. <i>fenestella</i>, a little window). The +"Lace-corals," a group of Palæozoic Polyzoans.<br/> +FILICES (Lat. <i>filix</i>, a fern). The order of Cryptogamic +plants comprising the Ferns.<br/> +FILIFORM (Lat. <i>filum</i>, a thread; <i>forma</i>, shape). +Thread-shaped.<br/> +FLORA (Lat. <i>Flora</i>, the goddess of flowers). The general +assemblage of the plants of any region or district.<br/> +FORAMINIFERA (Lat. <i>foramen</i>, an aperture; <i>fero</i>, +I carry). An order of Protozoa, usually characterised by the +possession of a shell perforated by numerous pseudopodial +apertures.<br/> +FRUGIVOROUS (Lat. <i>frux</i>, fruit; <i>voro</i>, I devour). +Living upon fruits.<br/> +FUCOIDS (Lat. <i>fucus</i>, sea-weed; Gr. <i>eidos</i>, likeness). +Fossils, often of an obscure nature, believed to be the remains +of sea-weeds.<br/> +FUSULINA (Lat. <i>fusus</i>, a spindle). An extinct genus of +<i>Foraminifera</i>.<br/> +</p> + +<p> +GANOID (Gr, <i>ganos</i>, splendour, brightness). Applied to +those scales or plates which are composed of an inferior layer +of true bone covered by a superior layer of polished enamel.<br/> +GANOIDEI. An order of Fishes.<br/> +GASTEROPODA (Gr. <i>gaster</i>, stomach; <i>pous</i>, foot). +The class of the Mollusca comprising the ordinary Univalves, +in which locomotion is usually effected by a muscular expansion +of the under surface of the body (the "foot").<br/> +GLOBIGERINA (Lat. <i>globus</i>, a globe; <i>gero</i>, I carry). +A genus of <i>Foraminifera</i>.<br/> +GLYPTODON (Gr. <i>glupho</i>, I engrave; <i>odous</i>, tooth). An +extinct genus of Armadillos, so named in allusion to the fluted +teeth.<br/> +GONIATITES (Gr. <i>gonia</i>, angle). A genus of Tetrabranchiate +Cephalopods.<br/> +GRALLATORES (Lat. <i>grallĹ“</i>, stilts). The order of the +long-legged Wading Birds.<br/> +GRAPTOLITIDÆ. (Gr. <i>grapho</i>, I write; <i>lithos</i>, +stone). An extinct sub-class of the <i>Hydrozoa</i>.<br/> +GYMNOSPERMS (Gr. <i>gumnos</i>, naked; <i>sperma</i>, seed). The +Conifers and Cycads, in which the seed is not protected within +a seed-vessel. +</p> + +<p> +HALITHERIUM (Gr. <i>hals</i>, sea; <i>therion</i>, beast). An +extinct genus of Sea-cows (<i>Sirenia</i>).<br/> +HAMITES (Lat. <i>hamus</i>, a hook). A genus of the +<i>AmmonitidĹ“</i>.<br/> +HELIOPHYLLUM (Gr. <i>helios</i>, the sun; <i>phullon</i>, leaf). +A genus of Rugose Corals.<br/> +HELLADOTHERIUM (Gr. <i>Hellas</i>, Greece; <i>therion</i>, beast). +An extinct genus of Ungulate Mammals.<br/> +HEMIPTERA (Gr. <i>hemi</i>, and <i>pteron</i>, wing). An order +of Insects in which the anterior wings are sometimes +"hemelytra."<br/> +HESPERORNIS (Gr. <i>Hesperos</i>, the evening star; <i>ornis</i>, +bird). An extinct genus of Birds.<br/> +HETEROCERCAL (Gr. <i>heteros</i>, diverse; <i>kerkos</i>, tail). +Applied to the tail of Fishes when it is unsymmetrical, or composed +of two unequal lobes.<br/> +HETEROPODA (Gr. <i>heteros</i>, diverse; <i>podes</i>, feet). An +aberrant group of the Gasteropods, in which the foot is modified +so as to form a swimming organ.<br/> +HIPPARION (Gr. <i>hipparion</i>, a little horse). An extinct genus +of <i>EquidĹ“</i>.<br/> +HIPPOPOTAMUS (Gr. <i>hippos</i>, horse; <i>potamos</i>, river). +A genus of Hoofed Quadrupeds—the "River-horses."<br/> +HIPPURITIDÆ. (Gr. <i>hippos</i>, horse; <i>oura</i>, tail). An +extinct family of Bivalve Molluscs.<br/> +HOLOPTYCHIUS (Gr. <i>holos</i>, whole; <i>ptucĂ©</i>, wrinkle). +An extinct genus of Ganoid Fishes.<br/> +HOLOSTOMATA (Gr. <i>holos</i>, whole; <i>stoma</i>, mouth). A +division of <i>Gasteropodous Molluscs</i>, in which the aperture +of the shell is rounded, or "entire."<br/> +HOLOTHUROIDEA (Gr. <i>holothourion</i>, and <i>eidos</i>, form). +An order of <i>Echinodermata</i> comprising the Trepangs.<br/> +<a name="page_387"><span class="page">Page 387</span></a> +HOMOCERCAL (Gr. <i>homos</i>, same; <i>kerkol</i>, tail). Applied +to the tail of Fishes when it is symmetrical, or composed of two +equal lobes.<br/> +HYBODUNTS (Gr. <i>hubos</i>, curved; <i>odous</i>, tooth). A group +of Fishes of which <i>Hybodus</i> is the type-genus.<br/> +HYDROIDA (Gr. <i>hudra</i>; and <i>eidos</i>, form). The sub-class +of the <i>Hydrozoa</i>, which comprises the animals most nearly +allied to the Hydra.<br/> +HYDROZOA (Gr. <i>hudra</i>; and <i>zoön</i>, animal). The class +of the <i>CĹ“lenterata</i> which comprises animals constructed +after the type of the Hydra.<br/> +HYMENOPTERA (Gr. <i>humen</i>, a membrane; <i>pteron</i>, a wing). +An order of Insects (comprising Bees, Ants, &c.) characterised +by the possession of four membranous wings. +</p> + +<p> +ICHTHYODORULITE (Gr. <i>ichthus</i>, fish; <i>dorus</i>, spear; +<i>lithos</i>, stone). The fossil fin-spine of Fishes.<br/> +ICHTHYOPTERYGIA (Gr. <i>ichthus</i>; <i>pterux</i>, wing). An +extinct order of Reptiles.<br/> +ICHTHYORNIS (Gr. <i>ichthus</i>, fish; <i>ornis</i>, bird). An +extinct genus of Birds.<br/> +ICHTHYOSAURIA (Gr. <i>ichthus</i>; <i>saura</i>, lizard). Synonymous +with <i>Ichthyopterygia</i>.<br/> +IGUANODON (<i>Iguana</i>, a living lizard; Gr. <i>odous</i>, tooth). +A genus of Deinosaurian Reptiles.<br/> +INCISOR (Lat. <i>incido</i>, I cut). The cutting teeth fixed +in the intermaxillary bones of the <i>Mammalia</i>, and the +corresponding teeth in the lower jaw.<br/> +INEQUILATERAL. Having the two sides unequal, as in the case of +the shells of the ordinary bivalves (<i>Lamellibranchiata</i>). +When applied to the shells of the <i>Foraminifera</i>, it implies +that the convolutions of the shell do not lie in the same plane, +but are obliquely wound round an axis.<br/> +INEQUIVALVE. Composed of two unequal pieces or valves.<br/> +INOCERAMUS (Gr. <i>is</i>, a fibre; <i>keramos</i>, an earthen +vessel). An extinct genus of Bivalve Molluscs.<br/> +INSECTA (Lat. <i>inseco</i>, I cut into). The class of articulate +animals commonly known as Insects.<br/> +INSECTIVORA (Lat. <i>insectum</i>, an insect; <i>voro</i>, I +devour). An order of Mammals.<br/> +INSECTIVOROUS. Living upon Insects.<br/> +INSESSORES (Lat. <i>insedeo</i>, I sit upon). The order of the +Perching Birds, often called <i>Passeres</i>.<br/> +INTERAMBULACRA. The rows of plates in an <i>Echinoid</i> which +are not perforated for the emission of the "tube-feet."<br/> +INTERMMAXILLÆ or PRÆMAXILLÆ. The two bones +which are situated between the two superior maxillæ in +<i>Vertebrata</i>. In man, and some monkeys, the +præmaxillæ anchylose with the maxillæ, so +as to be irrecognisable in the adult.<br/> +INVERTEBRATA (Lat. <i>in</i>, without; <i>vertebra</i>, a bone +of the back). Animals without a spinal column or backbone.<br/> +ISOPODA. (Gr. <i>isos</i>, equal; <i>podes</i>, feet). An order +of <i>Crustacea</i> in which the feet are like one another and +equal. +</p> + +<p> +KAINOZOIC (Gr. <i>kainos</i>, recent; <i>zoe</i>, life). The +Tertiary period in Geology comprising those formations in which +the organic remains approximate more or less closely to the existing +fauna and flora. +</p> + +<p> +LABYRINTHODONTIA (Gr. <i>laburinthos</i>, a labyrinth; <i>odous</i>, +tooth). An extinct order of <i>Amphibia</i>, so called from the +complex microscopic structure of the teeth.<br/> +LACERTILIA (Lat. <i>lacerta</i>, a lizard). An order of +<i>Reptilia</i> comprising the Lizards and Slow-worms.<br/> +LAMELLIBRANCHIATA (Lat. <i>lamella</i>, a plate; Gr. <i>bragchia</i>, +gill). The class of <i>Mollusca</i> comprising the ordinary bivalves, +characterised by the possession of lamellar gills.<br/> +LEPIDODENDRON (Gr. <i>lepis</i>, a scale; <i>dendron</i>, a tree). +A genus of extinct plants, so named from the scale-like scars +upon the stem left by the falling off of the leaves.<br/> +<a name="page_388"><span class="page">Page 388</span></a> +LEPIDOPTERA (Gr. <i>lepis</i>, a scale; <i>pteron</i>, a wing). An +order of Insects, comprising Butterflies and Moths, characterised by +possessing four wings which are usually covered with minute scales.<br/> +LEPIDOSIREN (Gr. <i>lepis</i>, a scale; <i>seiren</i>, a +siren—the generic name of the Mud-eel or <i>Siren +lacertina</i>). A genus of Dipnoous fishes, comprising the +"Mud-fishes."<br/> +LEPIDOSTROBUS (Gr. <i>lepis</i>, a scale; <i>strobilos</i>, a +fir-cone). A genus founded on the cones of <i>Lepidodendron</i>.<br/> +LEPTÆNA (Gr. <i>leptos</i>. slender). A genus of +Brachiopods.<br/> +LINGULA (Lat. <i>lingula</i>, a little tongue). A genus of +Brachiopods.<br/> +LYCOPODIACEÆ (Gr. <i>lupos</i>, a wolf; <i>pous</i>, foot). The +group of Cryptogamic plants generally known as "Club-mosses." +</p> + +<p> +MACHÆRACANTHUS (Gr. <i>machaira</i>, a sabre; <i>acantha</i>, +thorn or spine). An extinct genus of Fishes.<br/> +MACHAIRODUS (Gr. <i>machaira</i>, a sabre; <i>odous</i>, tooth). +An extinct genus of Carnivora.<br/> +MACROTHERIUM (Gr. <i>makros</i>, long; <i>therion</i>. beast). +An extinct genus of Edentata.<br/> +MACRURA (Gr. <i>makros</i>, long; <i>oura</i>, tail). A tribe of +Decapod <i>Crustaceans</i> with long tails (e.g., the Lobster, +Shrimp, &c.)<br/> +MAMMALIA (Lat. <i>mamma</i>, the breast). The class of Vertebrate +animals which suckle their young.<br/> +MANDIBLE (Lat. <i>mandibulum</i>, a jaw). The upper pair of jaws +in Insects; also applied to one of the pairs of jaws in +<i>Crustacea</i> and Spiders, to the beak of Cephalopods, the +lower jaw of Vertebrates, &c.<br/> +MANTLE. The external integument of most of the Mollusca, which +is largely developed, and forms a cloak in which the viscera +are protected. Technically called the "pallium."<br/> +MANUS (Lat. the hand). The hand of the higher Vertebrates.<br/> +MARSIPOBRANCHII (Gr. <i>marsipos</i>, a pouch; <i>bragchia</i>, +gill). The order of Fishes comprising the Hag-fishes and Lampreys, +with pouch-like gills.<br/> +MARSUPIALIA (Lat. <i>marsupium</i>, a pouch). An order of Mammals +in which the females mostly have an abdominal pouch in which +the young are carried.<br/> +MASTODON (Gr. <i>mastos</i>, nipple; <i>odous</i>, tooth). An +extinct genus of Elephantine Mammals.<br/> +MEGALONYX (Gr. <i>megas</i>, great; <i>onux</i>, nail). An extinct +genus of Edentate Mammals.<br/> +MEGALOSAURUS (Gr. <i>megas</i>, great; <i>saura</i>, lizard). +A genus of Deinosaurian Reptiles.<br/> +MEGATHERIUM (Gr. <i>megas</i>, great; <i>therion</i>, beast). +An extinct genus of Edentata.<br/> +MESOZOIC (Gr. <i>mesos</i>, middle; and <i>zoe</i>, life). The +Secondary period in Geology.<br/> +MICROLESTES (Gr. <i>mikros</i>, little; <i>lestes</i>, thief). +An extinct genus of Triassic Mammals.<br/> +MILLEPORA (Lat. <i>mille</i>, one thousand; <i>porus</i>, a pore). +A genus of "Tabulate Corals."<br/> +MIOCENE (Gr. <i>meion</i>, less; <i>kainol</i>, new). The Middle +Tertiary period.<br/> +MOLARS (Lat. <i>mola</i>, a mill). The "grinders" in man, or the +teeth in diphyodont Mammals which are not preceded by +milk-teeth.<br/> +MOLLUSCA (Lat. <i>mollis</i>, soft). The sub-kingdom which includes +the Shell-fish proper, the <i>Polyzoa</i>, the <i>Tunicata</i>, +and the Lamp-shells; so called from the generally soft nature +of their bodies.<br/> +MOLLUSCOIDA (<i>Mollusca</i>; Gr. <i>eidos</i>, form). The lower +division of the <i>Mollusca</i>, comprising the <i>Polyzoa, +Tunicata</i>, and <i>Brachiopoda</i>.<br/> +MONOGRAPTUS (Gr. <i>monos</i>, single; <i>grapho</i>, I write). +A genus of Graptolites.<br/> +MYLODON (Gr. <i>mulos</i>, a mill; <i>odous</i>, tooth). An extinct +genus of Edentate Mammals.<br/> +MYRIAPODA or MYRIOPODA (Gr. <i>murios</i>, ten thousand; +<i>podes</i>, feet). A class of <i>Arthropoda</i> comprising +the Centipedes and their allies, characterised by their numerous +feet. +</p> + +<p> +<a name="page_389"><span class="page">Page 389</span></a> +NATATORES (Lat. <i>nare</i>, to swim). The order of the Swimming +Birds.<br/> +NATATORY (Lat. <i>nare</i>, to swim). Formed for swimming.<br/> +NAUTILOID. Resembling the shell of the <i>Nautilus</i> in +shape.<br/> +NERVURES (Lat. <i>nervus</i>, a sinew). The ribs which support +the membranous wings of insects.<br/> +NEUROPTERA (Gr. <i>neuron</i>, a nerve; <i>pteron</i>, a wing). +An order of Insects characterised by four membranous wings with +numerous reticulated nervures (<i>e.g.</i>, Dragon-flies).<br/> +NEUROPTERIS (Gr. <i>neuron</i>, a nerve; <i>pteris</i>, a fern). +An extinct genus of Ferns.<br/> +NOTHOSAURUS (Gr. <i>nothos</i>, spurious; <i>saura</i>, lizard). +A genus of <i>Plesiosaurian</i> Reptiles.<br/> +NOTOCHORD (Gr. <i>notos</i>, back; <i>chorde</i>, string). A +cellular rod which is developed in the embryo of Vertebrates +immediately beneath the spinal cord, and which is usually replaced +in the adult by the vertebral column. Often it is spoken of as +the "chorda dorsalis."<br/> +NUDIBRANCHIATA (Lat. <i>nudus</i>, naked; and Gr. <i>bragchia</i>, +gill). An order of the <i>Gasteropoda</i> in which the gills +are naked.<br/> +NUMMULINA (Lat. <i>nummus</i>, a coin). A genus of +<i>Foraminifera</i>, comprising the coin-shaped "Nummulites." +</p> + +<p> +OBOLELLA (Lat. dim. of <i>obolus</i>, a small coin). An extinct +genus of Brachiopods.<br/> +OCCIPITAL. Connected with the <i>occiput</i>, or the back part +of the head.<br/> +OCEANIC. Applied to animals which inhabit the open ocean (= +pelagic).<br/> +ODONTOPTERYX (Gr. <i>oduos</i>, tooth; <i>pterux</i>, wing). An +extinct genus of Birds.<br/> +ODONTORNITHES (Gr. <i>oduos</i>, tooth; <i>ornis</i>, bird). +The extinct order of Birds, comprising forms with distinct teeth +in sockets.<br/> +OLIGOCENE (Gr. <i>oligos</i>, few; <i>kainos</i>, new). A name +used by many Continental geologists as synonymous with the Lower +Miocene.<br/> +OPHIDIA (Gr. <i>ophis</i>, a serpent). The order of Reptiles +comprising the Snakes.<br/> +OPHIUROIDEA (Gr. <i>ophis</i>, snake; <i>oura</i>, tail; +<i>eidos</i>, form). An order of <i>Echinodermata</i>, comprising +the Brittle-stars and Sand-stars.<br/> +ORNITHOSCELIDA (Gr. <i>ornis</i>, bird; <i>skelos</i>, leg). +Applied by Huxley to the Deinosaurian Reptiles, together with the +genus <i>Compsognathus</i>, on account of the bird-like character +of their hind-limbs.<br/> +ORTHIS (Gr. <i>orthos</i>, straight). A genus of Brachiopods, +named in allusion to the straight hinge-line.<br/> +ORTHOCERATIDÆ (Gr. <i>orthos</i>, straight; <i>keras</i>, +horn). A family of the <i>NautilidĹ“</i>, in which the shell +is straight, or nearly so.<br/> +ORTHOPTERA (Gr. <i>orthos</i>, straight; <i>pteron</i>, wing). +An order of Insects.<br/> +OSTEOLEPIS (Gr. <i>osteon</i>, bone; <i>lepis</i>, scale). An +extinct genus of Ganoid Fishes.<br/> +OSTRACODA (Gr. <i>ostrakon</i>, a shell). An order of small +Crustaceans which are enclosed in bivalve shells.<br/> +OTODUS (Gr. <i>ota</i>, ears; <i>odous</i>, tooth). An extinct +genus of Sharks.<br/> +OUDENODON (Gr. <i>ouden</i>, none; <i>odous</i>, tooth). A genus +of Dicynodont Reptiles.<br/> +OVIBUS (Lat. <i>ovis</i>, sheep; <i>bos</i>, ox). The genus +comprising the Musk-ox. +</p> + +<p> +PACHYDERMATA (Gr. <i>pachus</i>, thick; <i>derma</i>, skin). +An old Mammalian order constituted by Cuvier for the reception +of the Rhinoceros, Hippopotamus, Elephant, &c.<br/> +PALÆASTER (Gr. <i>palaios</i>, ancient; <i>aster</i>, star). +An extinct genus of Star-fishes.<br/> +PALÆOCARIS (Gr. <i>palaios</i>, ancient; <i>karis</i>, shrimp). +An extinct genus of Decapod Crustaceans.<br/> +PALÆOLITHIC (Gr. <i>palaios</i>, ancient; <i>lithos</i>, +stone). Applied to the rude stone implements of the earliest known +races of men, to the men who made these implements, or to the +period at which they were made.<br/> +<a name="page_390"><span class="page">Page 390</span></a> +PALÆONTOLOGY (Gr. <i>palaios</i>, ancient; and <i>logos</i>, +discourse). The science of fossil remains or of extinct organised +beings.<br/> +PALÆOPHIS (Gr. <i>palaios</i>, ancient; <i>ophis</i>, serpent). +An extinct genus of Snakes.<br/> +PALÆOSAURUS (Gr. <i>palaios</i>, ancient; <i>saura</i>, lizard). +A genus of Thecodont Reptiles.<br/> +PALÆOTHERIDÆ. (Gr. <i>palaios</i>, ancient; <i>ther</i>, +beast). A group of Tertiary Ungulates.<br/> +PALÆOZOIC (Gr. <i>palaios</i>, ancient; and <i>zoe</i>, life). +Applied to the oldest of the great geological epochs.<br/> +PARADOXIDES (Lat. <i>paradoxus</i>, marvellous). A genus of +Trilobites.<br/> +PATAGIUM (Lat. the border of a dress). Applied to the expansion +of the integument by which Bats, Flying Squirrels, and other +animals support themselves in the air.<br/> +PECOPTERIS (Gr. <i>peko</i>, I comb; <i>pteris</i>, a fern). An +extinct genus of Ferns.<br/> +PECTEN (Lat. a comb). The genus of Bivalve Molluscs comprising +the Scallops.<br/> +PECTORAL (Lat. <i>pectus</i>, chest). Connected with, or placed +upon, the chest.<br/> +PENTACRINUS (Gr. <i>penta</i>, five; <i>krinon</i>, lily). A genus +of Crinoids in which the column is five-sided.<br/> +PENTAMERUS (Gr. <i>penta</i>, five; <i>meros</i>, part). An extinct +genus of Brachiopods.<br/> +PENTREMITES (Gr. <i>penta</i>, five; <i>trema</i>, aperture). A +genus of <i>Blastoidea</i>, so named in allusion to the apertures +at the summit of the calyx.<br/> +PERENNIBRANCHIATA (Lat. <i>perennis</i>, perpetual; Gr. +<i>bragchia</i>, gill). Applied to those Amphibia in which the +gills are permanently retained throughout life.<br/> +PERISSODACTYLA (Gr. <i>perissos</i>, uneven; <i>daktulos</i>, +finger). Applied to those Hoofed Quadrupeds (<i>Ungulata</i>) +in which the feet have an uneven number of toes.<br/> +PETALOID. Shaped like the petal of a flower.<br/> +PHACOPS (Gr. <i>phakĂ©</i>, a lentil; <i>ops</i>, the eye). +A genus of Trilobites.<br/> +PHALANGES (Gr. <i>phalanx</i>, a row). The small bones composing +the digits of the higher <i>Vertebrata</i>. Normally each digit +has three phalanges.<br/> +PHANEROGAMS (Gr. <i>phaneros</i>, visible; <i>gamos</i>, marriage). +Plants which have the organs of reproduction conspicuous, and +which bear true flowers.<br/> +PHARYNGOBRANCHII (Gr. <i>pharugx</i>, pharynx; <i>bragchia</i>, +gill). The order of Fishes comprising only the Lancelet.<br/> +PHASCOLOTHERIUM (Gr. <i>phaskolos</i>, a pouch; <i>therion</i>, +a beast). A genus of Oolitic Mammals.<br/> +PHRAGMACONE (Gr. <i>phragma</i>, a partition; and <i>konos</i>, +a cone). The chambered portion of the internal shell of a +<i>Belemnite</i>.<br/> +PHYLLOPODA (Gr. <i>phullon</i>, leaf; and <i>pous</i>, foot). +An order of <i>Crustacea</i>.<br/> +PINNATE (Lat. <i>pinna</i>, a feather). Feather-shaped; or possessing +lateral processes.<br/> +PINNIGRADA (Lat. <i>pinna</i>, a feather; <i>gradior</i>, I walk). +The group of <i>Carnivora</i>, comprising the Seals and Walruses, +adapted for an aquatic life. Often called <i>Pinnipedia</i>.<br/> +PINNULÆ. (Lat. dim. of <i>pinna</i>). The lateral processes of +the arms of <i>Crinoids</i>.<br/> +PISCES (Lat. <i>piscis</i>, a fish). The class of Vertebrates +comprising the Fishes.<br/> +PLACOID (Gr. <i>plax</i>, a plate; <i>eidos</i>, form). Applied +to the irregular bony plates, grains, or spines which are found +in the skin of various fishes (<i>Elasmobranchii</i>).<br/> +PLAGIOSTOMI (Gr. <i>plagios</i>, transverse; <i>stoma</i>, mouth). +The Sharks and Rays, in which the mouth is transverse, and is +placed on the under surface of the head.<br/> +PLATYCERAS (Gr. <i>platus</i>, broad; <i>keras</i>, horn). A genus +of Univalve Molluscs.<br/> +PLATYCRINUS (Gr. <i>platus</i>, broad; <i>krinom</i>, lily). A +genus of Crinoidea.<br/> +PLATYRHINA (Gr. <i>platus</i>, broad; <i>rhines</i>, nostrils). +A group of the <i>Quadrumana</i>.<br/> +PLATYSOMUS (Gr. <i>platus</i>, wide; <i>soma</i>, body). A genus +of Ganoid Fishes.<br/> +PLEISTOCENE (Gr. <i>pleistos</i>, most; <i>kainos</i>, new). Often +used as synonymous with "Post-Pliocene."<br/> +<a name="page_391"><span class="page">Page 391</span></a> +PLEUROTOMARIA (Gr. <i>pleura</i>, the side; <i>tomĂ©</i>, +notch). A genus of Univalve shells.<br/> +PLIOCENE (Gr. <i>pleion</i>, more; <i>kainos</i>, new). The later +Tertiary period.<br/> +PLIOPITHECUS (Gr. <i>pleion</i>, more; <i>pithekos</i>, ape). +An extinct genus of monkeys.<br/> +PLIOSAURUS (Gr. <i>pleion</i>, more; <i>saura</i>, lizard). A +genus of Plesiosaurian Reptiles.<br/> +POLYCYSTINA (Gr. <i>polus</i>, many; and <i>kustis</i>, a cyst). +An order of <i>Protozoa</i> with foraminated siliceous shells.<br/> +POLYPARY. The hard chitinous covering secreted by many of the +<i>Hydrozoa</i>.<br/> +POLYPE (Gr. <i>polus</i>, many; <i>pous</i>, foot). Restricted +to the single individual of a simple <i>Actinozoön</i>, such +as a Sea-anemone, or to the separate zooids of a compound +<i>Actinozoön</i>. Often applied indiscriminately to any of +the <i>CĹ“lenterata</i>, or even to the Polyzoa.<br/> +POLYPORA (Gr. <i>polus</i>, many; <i>poros</i>, a passage). A +genus of Lace-corals (<i>FenestellidĹ“</i>).<br/> +POLYTHALAMOUS (Gr. <i>polus</i>; and <i>thalamos</i>, chamber). +Having many chambers; applied to the shells of <i>Foraminifera</i> +and <i>Cephalopoda</i>.<br/> +POLYZOA (Gr. <i>polus</i>; and <i>zoön</i>, animal). A division +of the <i>Molluscoida</i> comprising compound animals, such as +the Sea-mat—sometimes called <i>Bryozoa</i>.<br/> +PORIFERA (Lat. <i>porus</i>, pore; and <i>fero</i>, I carry). +Sometimes used to designate the <i>Foraminifera</i>, or the +<i>Sponges</i>.<br/> +PRÆMOLARS (Lat. <i>prĹ“</i>, before; <i>molares</i>, the +grinders). The molar teeth of Mammals which succeed the molars of the +milk-set of teeth. In man, the bicuspid teeth.<br/> +PROBOSCIDEA (Lat. <i>proboscis</i>, the snout). The order of Mammals +comprising the Elephants.<br/> +PROCĹ’LOUS (Gr. <i>pro</i>, before; <i>koilos</i>, hollow). +Applied to vertebræ the bodies of which are hollow or concave +in front.<br/> +PRODUCTA (Lat. <i>productus</i>, drawn out or extended). An extinct +genus of Brachiopods, in which the shell is "eared," or has its +lateral angles drawn out.<br/> +PROTICHNITES (Gr. <i>protos</i>, first; <i>ichnos</i>, footprint). +Applied to certain impressions in the Potsdam sandstone of North +America, believed to have been produced by large Crustaceans.<br/> +PROTOPHYTA (Gr. <i>protos</i>; and <i>phuton</i>, plant). The +lowest division of plants.<br/> +PROTOPLASM (Gr. <i>protos</i>; and <i>plasso</i> I mould). The +elementary basis of organised tissues. Sometimes used synonymously +for the "sarcode" of the <i>Protozoa</i>.<br/> +PROTOROSAURUS or PROTEROSAURUS (Gr. <i>protos</i>, first; +<i>orao</i>, I see or discover; <i>saura</i>, lizard: or +<i>proteros</i>, earlier; <i>saura</i>, lizard). A genus of Permian +lizards.<br/> +PROTOZOA (Gr. <i>protos</i>; and <i>zoön</i>, animal). The +lowest division of the animal kingdom.<br/> +PSAMMODUS (Gr. <i>psammos</i>, sand; <i>odous</i>, tooth). An +extinct genus of Cestraciont Sharks.<br/> +PSEUDOPODIA (Gr. <i>pseudos</i>, falsity; and <i>pous</i>, foot). +The extensions of the body-substance which are put forth by the +<i>Rhizopoda</i> at will, and which serve for locomotion and +prehension.<br/> +PSILOPHYTON (Gr. <i>psilos</i>, bare; <i>phuton</i>, plant). An +extinct genus of Lycopodiaceous plants.<br/> +PTERANODON (Gr. <i>pteron</i>, wing; <i>a</i>, without; <i>odous</i>, +tooth). A genus of Pterosaurian Reptiles.<br/> +PTERASPIS (Gr. <i>pteron</i>, wing; <i>aspis</i>, shield). A genus +of Ganoid Fishes.<br/> +PTERICHTHYS (Gr. <i>pteron</i>, wing; <i>ichthus</i>, fish). A +genus of Ganoid Fishes.<br/> +PTERODACTYLUS (Gr. <i>pteron</i>, wing; <i>daktulos</i>, finger). +A genus of Pterosaurian Reptiles.<br/> +PTEROPODA (Gr. <i>pteron</i>, wing; and <i>pous</i>, foot). A +class of the <i>Mollusca</i> which swim by means of fins attached +near the head.<br/> +PTEROSAURIA (Gr. <i>pteron</i>, wing; <i>saura</i>, lizard). An +extinct order of Reptiles.<br/> +PTILODICTYA (Gr. <i>ptilon</i>, a feather; <i>diktuon</i>, a net). +An extinct genus of <i>Polyzoa</i>.<br/> +<a name="page_392"><span class="page">Page 392</span></a> +PTYCHOCERAS (Gr. <i>ptucĂ©</i>, a fold; <i>keras</i>, a +horn). A genus of <i>AmmonitidĹ“</i>.<br/> +PULMONATE. Possessing lungs.<br/> +PYRIFORM (Lat. <i>pyrus</i>, a pear; and <i>forma</i>, form). +Pear-shaped. +</p> + +<p> +QUADRUMANA (Lat. <i>quatuor</i>, four; <i>manus</i>, hand). The +order of Mammals comprising the Apes, Monkeys, Baboons, Lemurs, +&c. +</p> + +<p> +RADIATA (Lat. <i>radius</i>, a ray). Formerly applied to a large +number of animals which are now placed in separate sub-kingdoms +(e.g., the <i>CĹ“lenterata</i>, the <i>Echinodermata</i>, the +<i>Infusoria</i>, &c.)<br/> +RADIOLARIA (Lat. <i>radius</i>, a ray). A division of +<i>Protozoa</i>.<br/> +RAMUS (Lat. a branch). Applied to each half or branch of the lower +jaw, or mandible, of Vertebrates.<br/> +RAPTORES (Lat. <i>rapto</i>, I plunder). The order of the Birds +of Prey.<br/> +RASORES (Lat. <i>rado</i>, I scratch). The order of the Scratching +Birds (Fowls. Pigeons, &c.)<br/> +RECEPTACULITES (Lat. <i>receptaculum</i>, a storehouse). An extinct +genus of Protozoa.<br/> +REPTILIA (Lat. <i>repto</i>, I crawl). The class of the +<i>Vertebrata</i> comprising the Tortoises, Snakes, Lizards, +Crocodiles, &c.<br/> +RETEPORA (Lat. <i>retĂ©</i>, a net; <i>porus</i>, a pore). A +genus of Lace-corals (<i>Polyzoa</i>).<br/> +RHAMPHORHYNCHUS (Gr. <i>rhamphos</i>, beak; <i>rhugchos</i>, nose). +A genus of Pterosaurian Reptiles.<br/> +RHINOCEROS (Gr. <i>rhis</i>, the nose; <i>keras</i>, horn). A +genus of Hoofed Quadrupeds.<br/> +RHIZOPODA (Gr. <i>rhiza</i>, a root; and <i>pous</i>, foot). +The division of <i>Protozoa</i> comprising all those which are +capable of emitting pseudopodia.<br/> +RHYNCHOLITES (Gr. <i>rhugchos</i>, beak; and <i>lithos</i>, stone). +Beak-shaped fossils consisting of the mandibles of +<i>Cephalopoda</i>.<br/> +RHYNCHONELLA (Gr. <i>rhugchos</i>, nose or beak). A genus of +Brachiopods.<br/> +RODENTIA (Lat. <i>rodo</i>, I gnaw). An order of the Mammals; +often called <i>Glires</i> (Lat. <i>glis</i>, a dormouse).<br/> +ROTALIA (Lat. <i>rota</i>, a wheel). A genus of +<i>Foraminifera</i>.<br/> +RUGOSA (Lat. <i>rugosus</i>, wrinkled). An order of Corals.<br/> +RUMINANTIA (Lat. <i>ruminor</i>, I chew the cud). The group of +Hoofed Quadrupeds (<i>Ungulata</i>) which "ruminate" or chew +the cud.<br/> +</p> + +<p> +SARCODE (Gr. <i>sarx</i>, flesh; <i>eidos</i>, form). The jelly-like +substance of which the bodies of the <i>Protozoa</i> are composed. +It is an albuminous body containing oil-granules, and is sometimes +called "animal protoplasm."<br/> +SAURIA (Gr. <i>saura</i>, a lizard). Any lizard-like Reptile is +often spoken of as a "Saurian;" but the term is sometimes restricted +to the Crocodiles alone, or to the Crocodiles and Lacertilians.<br/> +SAUROPTERYGIA (Gr. <i>sauro</i>; <i>pterux</i>, wing). An extinct +order of Reptiles, called by Huxley <i>Plesiosauria</i>, from +the typical genus <i>Plesiosaurus</i>.<br/> +SAURURÆ (Gr. <i>saura</i>; <i>oura</i>, tail). The extinct order +of Birds comprising only the <i>ArchĹ“opteryx</i>.<br/> +SCANSORES (Lat. <i>scando</i>, I climb). The order of the Climbing +Birds (Parrots, Woodpeckers, &c.)<br/> +SCAPHITES (Lat. <i>scapha</i>, a boat). A genus of the +<i>AmmonitidĹ“</i>.<br/> +SCOLITHUS (Gr. <i>skolex</i>, a worm; <i>lithos</i>, a stone). +The vertical burrows of sea-worms in rocks.<br/> +SCUTA (Lat. <i>scutum</i>, a shield). Applied to any shield-like +plates; especially to those which are developed in the integument +of many Reptiles.<br/> +SELACHIA or SELACHII (Gr. <i>selachos</i>, a cartilaginous fish, +probably a shark). The sub-order of <i>Elasmobranchii</i> comprising +the Sharks and Dog-fishes.<br/> +SEPIOSTAIRE. The internal shell of the Sepia, commonly known as +the "cuttle-bone."<br/> +SEPTA. Partitions.<br/> +SERPENTIFORM. Resembling a serpent in shape.<br/> +SERTULARIDA (Lat. <i>sertum</i>, a wreath). An order of +<i>Hydrozoa</i>.<br/> +<a name="page_393"><span class="page">Page 393</span></a> +SESSILE (Lat. <i>sedo</i>, I sit). Not supported upon a stalk or +peduncle; attached by a base.<br/> +SETHÆ (Lat. bristles). Bristles or long stiff hairs.<br/> +SIGILLARIOIDS (Lat. <i>sigilla</i>, little images). A group of +extinct plants of which <i>Sigillaria</i> is the type, so called +from the seal-like markings on the bark.<br/> +SILICEOUS (Lat. <i>silex</i>, flint). Composed of flint.<br/> +SINISTRAL (Lat. <i>sinistra</i>, the left hand). Left-handed; +applied to the direction of the spiral in certain shells, which +are said to be "reversed."<br/> +SIPHON (Gr. a tube). Applied to the respiratory tubes in the +<i>Mollusca</i>; also to other tubes of different functions.<br/> +SIPHONIA (Gr. <i>siphon</i>, a tube). A genus of fossil Sponges.<br/> +SIPHONOSTOMATA (Gr. <i>siphon</i>; and <i>stoma</i>, mouth). The +division of <i>Gasteropodous Molluscs</i> in which the aperture +of the shell is not "entire," but possesses a notch or tube for +the emission of the respiratory siphon.<br/> +SIPHUNCLE (Lat. <i>siphunculus</i>, a little tube). The tube +which connects together the various chambers of the shell of +certain <i>Cephalopoda</i> (<i>e.g.</i>, the Pearly Nautilus).<br/> +SIRENIA (Gr. <i>seiren</i>. a mermaid). The order of <i>Mammalia</i> +comprising the Dugongs and Manatees.<br/> +SIVATHERIUM (<i>Siva</i>, a Hindoo deity; Gr. <i>therion</i>, +beast). An extinct genus of Hoofed Quadrupeds.<br/> +SOLIDUNGULA (Lat. <i>solidus</i>, solid; <i>ungula</i>, a hoof). +The group of Hoofed Quadrupeds comprising the Horse, Ass, and +Zebra, in which each foot has only a single solid hoof. Often +called <i>Solipedia</i>.<br/> +SPHENOPTERIS (Gr. <i>sphen</i>, a wedge; <i>pteris</i>, a fern). +An extinct genus of ferns.<br/> +SPICULA (Lat. <i>spicidum</i>, a point). Pointed needle-shaped +bodies.<br/> +SPIRIFERA (Lat. <i>spira</i>, a spire or coil; <i>fero</i>, I +carry). An extinct genus of Brachiopods, with large spiral supports +for the "arms."<br/> +SPIRORBIS (Lat. <i>spira</i>, a spire; <i>orbis</i>, a circle). +A genus of tube-inhabiting Annelides, in which the shelly tube +is coiled into a spiral disc.<br/> +SPONGIDA (Gr. <i>spoggos</i>, a sponge). The division of +<i>Protozoa</i> commonly known as sponges.<br/> +STALACTITES (Gr. <i>stalasso</i>, I drop). Icicle-like encrustations +and deposits of lime, which hang from the roof of caverns in +limestone.<br/> +STALAGMITE (Gr. <i>stalagma</i>, a drop). Encrustations of lime +formed on the floor of caverns which are hollowed out of +limestone.<br/> +STIGMARIA (Gr. <i>stigma</i>, a mark made with a pointed instrument). +A genus founded on the roots of various species of +<i>Sigillaria</i>.<br/> +STRATUM (Lat. <i>stratus</i>, spread out; or <i>stratum</i>, a +thing spread out). A layer of rock.<br/> +STROMATOPORA (Gr. <i>stroma</i>, a thing spread out; <i>paras</i>, +a passage or pore). A Palæozoic genus of <i>Protozoa</i>.<br/> +STROPHOHENA (Gr. <i>strophao</i>, I twist; <i>menĂ©</i>, moon). +An extinct genus of Brachiopods.<br/> +SUB-CALCAREOUS. Somewhat calcareous.<br/> +SUB-CENTRAL. Nearly central, but not quite.<br/> +SUTURE (Lat. <i>suo</i>, I sew). The line of junction of two +parts which are immovably connected together. Applied to the +line where the whorls of a univalve shell join one another; also +to the lines made upon the exterior of the shell of a chambered +<i>Cephalopod</i> by the margins of the septa.<br/> +SYRINGOPORA (Gr. <i>surigx</i>, a pipe; <i>poros</i>, a pore). +A genus of Tabulate Corals. +</p> + +<p> +TABULÆ. (Lat. <i>tabula</i>, a tablet). Horizontal plates or +floors found in some Corals, extending across the cavity of the +"theca" from side to side.<br/> +TEGUMENTARY (Lat. <i>tegumentum</i>, a covering). Connected with +the integument or skin.<br/> +TELEOSAURUS (Gr. <i>teleios</i>, perfect; <i>saura</i>, lizard). +An extinct genus of Crocodilian Reptiles.<br/> +TELEOSTEI (Gr. <i>teleios</i>, perfect; <i>osteon</i>, bone). +The order of the "Bony Fishes."<br/> +<a name="page_394"><span class="page">Page 394</span></a> +TELSON (Gr. a limit). The last joint in the abdomen of +<i>Crustacea</i>; variously regarded as a segment without +appendages, or as an azygous appendage.<br/> +TENTACULITES (Lat. <i>tentaculum</i>, a feeler). A genus of +<i>Pteropoda</i>.<br/> +TEREBRATULA (Lat. <i>terebratus</i>, bored or pierced). A genus +of <i>Brachiopoda</i>, so called in allusion to the perforated +beak of the ventral valve.<br/> +TEST (Lat. <i>testa</i>, shell). The shell of <i>Mollusca</i>, +which are for this reason sometimes called "<i>Testacea</i>;" +also, the calcareous case of <i>Echinoderms</i>; also, the thick +leathery outer tunic in the <i>Tunicata</i>.<br/> +TESTACEOUS. Provided with a shell or hard covering.<br/> +TESTUDINIDÆ (Lat. <i>testudo</i>, a tortoise). The family of +the Tortoises.<br/> +TETRABRANCHIATA (Gr. <i>tetra</i>, four; <i>bragchia</i>, gill). +The order of <i>Cephalopoda</i> characterised by the possession +of four gills.<br/> +TEXTULARIA. (Lat. <i>textilis</i>, woven). A genus of +<i>Foraminifera</i>.<br/> +THECA (Gr. <i>thekĂ©</i>, a sheath). A genus of Pteropods.<br/> +THECODONTOSAURUS (Gr. <i>thekĂ©</i>, a sheath; <i>odous</i>, +tooth; <i>saura</i>, lizard). A genus of "Thecodont" Reptiles, so +named in allusion to the fact that the teeth are sunk in distinct +sockets.<br/> +THERIODONT (Gr. <i>therion</i>, a beast; <i>odous</i>, tooth). A +group of Reptiles so named by Owen in allusion to the Mammalian +character of their teeth.<br/> +THORAX (Gr. a breastplate). The region of the chest.<br/> +THYLACOLEO (Gr. <i>thulakos</i>, a pouch; <i>leo</i>, a lion). +An extinct genus of Marsupials.<br/> +TRIGONIA (Gr. <i>treis</i>, three; <i>gonia</i>, angle). A genus +of Bivalve Molluscs.<br/> +TRIGONOCARPON (Gr. <i>treis</i>, three; <i>gonia</i>. angle; +<i>karpos</i>, fruit). A genus founded on fossil fruits of a +three-angled form.<br/> +TRILOBITA (Gr. <i>treis</i>, three; <i>lobos</i>, a lobe). An +extinct order of <i>Crustaceans</i>.<br/> +TRINUCLEUS (Lat. <i>tris</i>, three; <i>nucleus</i>, a kernel). +A genus of Trilobites.<br/> +TROGONTHERIUM (Gr. <i>trogo</i>, I gnaw; <i>therion</i>, beast). +An extinct genus of Beavers.<br/> +TUBICOLA (Lat. <i>tuba</i>, a tube; and <i>colo</i>, I inhabit). +The order of <i>Annelida</i> which construct a tubular case in +which they protect themselves.<br/> +TUBICOLOUS. Inhabiting a tube.<br/> +TUNICATA (Lat. <i>tunica</i>, a cloak). A class of <i>Molluscoida</i> +which are enveloped in a tough leathery case or "test."<br/> +TURBINATED (Lat. <i>turbo</i>, a top). Top-shaped; conical with +a round base.<br/> +TURRILITES (Lat, <i>turris</i>, a tower). A genus of the +<i>AmmonitidĹ“</i>.<br/> +</p> + +<p> +UMBO (Lat. the boss of a shield). The beak of a bivalve shell.<br/> +UNGUICULATE (Lat. <i>unguis</i>, nail). Furnished with claws.<br/> +UNGULATA (Lat. <i>ungula</i>, hoof). The order of Mammals comprising +the Hoofed Quadrupeds.<br/> +UNGULATE. Furnished with expanded nails constituting hoofs.<br/> +UNILOCULAR (Lat. <i>unus</i>, one; and <i>loculus</i>. a little +purse). Possessing a single cavity or chamber. Applied to the +shells of <i>Foraminifera</i> and <i>Mollusca</i>.<br/> +UNIVALVE (Lat. <i>unus</i>, one; <i>valvĹ“</i>, folding-doors). +A shell composed of a single piece or valve.<br/> +URODELA (Gr. <i>oura</i>, tail; <i>delos</i>, visible). The order +of the Tailed Amphibians (Newts, &c.) +</p> + +<p> +VENTRAL (Lat. <i>venter</i>, the stomach). Relating to the inferior +surface of the body.<br/> +VENTRICULITES (Lat. <i>ventriculum</i>, a little stomach). A genus +of siliceous Sponges.<br/> +VERMIFORM (Lat. <i>vermis</i>, worm; and <i>forma</i>, form). +Worm-like.<br/> +VERTEBRA (Lat. <i>verto</i>, I turn). One of the bony segments +of the vertebral column or backbone.<br/> +VERTEBRATA (Lat. <i>vertebra</i>, a bone of the back, from +<i>vertere</i>, to turn). The division of the Animal Kingdom +roughly characterised by the possession of a backbone.<br/> +VESICLE (Lat. <i>vesica</i>, a bladder). A little sac or cyst. +</p> + +<p> +<a name="page_395"><span class="page">Page 395</span></a> +WHORL. The spiral turn of a univalve shell. +</p> + +<p> +XIPHOSURA (Gr. <i>xiphos</i>, a sworn; and <i>oura</i>, tail). +An order of <i>Crustacea</i>, comprising the <i>Limuli</i> or +King-Crabs, characterised by their long sword-like tails.<br/> +XYLOBIUS (Gr. <i>xulon</i>, wood; <i>bios</i>, life). An extinct +genus of Myriapods, named in allusion to the fact that the animal +lived on decaying wood. +</p> + +<p> +ZAPHRENTIS (proper name). A genus of Rugose Corals.<br/> +ZEUGLODONTIDÆ. (Gr. <i>zeuglĂ©</i>, a yoke; +<i>odous</i>, a tooth). An extinct family of Cetaceans, in which +the molar teeth are two-fanged, and look as if composed of two +parts united by a neck.<br/> +ZOOPHYTE (Gr. <i>zoön</i>, animal; <i>phuton</i>, plant). +Loosely applied to many plant-like animals, such as Sponges, +Corals, Sea-anemones, Sea-mats, &c. +</p> + +<h3> +<a name="page_396"><span class="page">Page 396</span></a> +INDEX.</h3> + +<hr /> + +<!-- The original index was arranged in two columns. --> + +<p class="index"> +Acadian Group, <a href="#page_79">79</a>.<br/> +<i>Acer</i>, <a href="#page_308">308</a>.<br/> +<i>Acervularia</i>, <a href="#page_119">119</a>, + <a href="#page_173">173</a>.<br/> +<i>Acidaspis</i>, <a href="#page_123">123</a>.<br/> +Acorn-shells, <a href="#page_267">267</a>.<br/> +<i>Acroculia</i>, <a href="#page_128">128</a>.<br/> +<i>Acrodus</i>, <a href="#page_214">214</a>, + <a href="#page_242">242</a>, <a href="#page_275">275</a>; + <i>nobilis</i>, <a href="#page_242">242</a>.<br/> +<i>Acrotreta</i>, <a href="#page_110">110</a>.<br/> +<i>Acroura</i>, <a href="#page_120">120</a>.<br/> +<i>Actinocrinus</i>, <a href="#page_175">175</a>.<br/> +<i>Æglina</i>, <a href="#page_108">108</a>.<br/> +<i>Æpiornis</i>, <a href="#page_348">348</a>.<br/> +<i>Agnostus</i>, <a href="#page_85">85-87</a>, + <a href="#page_108">108</a>; <i>rex</i>, + <a href="#page_85">85</a>.<br/> +<i>Alces malchis</i>, <a href="#page_354">354</a>.<br/> +<i>Alecto</i>, <a href="#page_108">108</a>.<br/> +<i>Alethopteris</i>, <a href="#page_136">136</a>, + <a href="#page_165">165</a>, <a href="#page_196">196</a>.<br/> +<i>AlgĹ“</i> (<i>see</i> Sea-weeds).<br/> +Alligators, <a href="#page_218">218</a>, + <a href="#page_297">297</a>.<br/> +<i>Alnus</i>, <a href="#page_262">262</a>.<br/> +<i>Amblypterus</i>, <a href="#page_188">188</a>; + <i>macropterus</i>, <a href="#page_188">188</a>.<br/> +<i>Ambonychia</i>, <a href="#page_111">111</a>.<br/> +<i>Ammonites</i>, <a href="#page_187">187</a>, + <a href="#page_212">212-214</a>, <a href="#page_237">237-239</a>, + <a href="#page_272">272</a>; + <i>Humpresianus</i>, <a href="#page_238">238</a>; + <i>bifrons</i>, <a href="#page_238">238</a>.<br/> +<i>AmmonitidĹ“</i>, <a href="#page_239">239</a>, + <a href="#page_272">272</a>, <a href="#page_285">285</a>, + <a href="#page_294">294</a>.<br/> +<i>Amphibia</i>, <a href="#page_189">189</a>; of the Carboniferous, + <a href="#page_189">189-191</a>; of the Permian, + <a href="#page_200">200</a>; of the Trias, + <a href="#page_215">215-217</a>; + of the Jurassic, <a href="#page_242">242</a>; + of the Miocene, <a href="#page_313">313</a>.<br/> +<i>Amphicyon</i>, <a href="#page_322">322</a>.<br/> +<i>Amphilestes</i>, <a href="#page_253">253</a>.<br/> +<i>Amphispongia</i>, <a href="#page_118">118</a>.<br/> +<i>Amphistegina</i>, <a href="#page_311">311</a>.<br/> +<i>Amphitherium</i>, <a href="#page_253">253</a>, + <a href="#page_255">255</a>; <i>Prevostii</i>, + <a href="#page_254">254</a>.<br/> +<i>Amphitragulus</i>, <a href="#page_317">317</a>.<br/> +<i>Amplexus</i>, <a href="#page_173">173</a>; + <i>coralloides</i>, <a href="#page_174">174</a>.<br/> +<i>Ampyx</i>, <a href="#page_108">108</a>.<br/> +<i>Anachytes</i>, <a href="#page_266">266</a>.<br/> +<i>Anchitherium</i>, <a href="#page_301">301-302</a>.<br/> +<i>Ancyloceras</i>, <a href="#page_272">272</a>, + <a href="#page_273">273</a>; <i>Matheronianus</i>, + <a href="#page_273">273</a>.<br/> +<i>Ancylotherium Pentelici</i>, <a href="#page_315">315</a>.<br/> +<i>Andrias Scheuchzeri</i>, <a href="#page_313">313</a>, + <a href="#page_314">314</a>.<br/> +<i>Angiosperms</i>, <a href="#page_261">261</a>, + <a href="#page_262">262</a>.<br/> +Animal Kingdom, divisions of, <a href="#page_375">375-378</a>.<br/> +<i>Anisopus</i>, <a href="#page_206">206</a>.<br/> +<i>Annelida</i>, of the Cambrian period, <a href="#page_82">82</a>, + <a href="#page_83">83</a>; of the Lower Silurian, + <a href="#page_107">107</a>; of the Upper Silurian, + <a href="#page_122">122</a>, <a href="#page_123">123</a>; + of the Devonian, <a href="#page_143">143</a>, + <a href="#page_144">144</a>; of the Carboniferous, + <a href="#page_178">178</a>.<br/> +<i>Annularia</i>, <a href="#page_137">137</a>, + <a href="#page_196">196</a>, <a href="#page_207">207</a>.<br/> +<i>Anomodontia</i>, <a href="#page_220">220</a>.<br/> +<i>AnoplotheridĹ“</i>, <a href="#page_302">302</a>.<br/> +<i>Anoplotherium</i>, <a href="#page_302">302</a>, + <a href="#page_303">303</a>; <i>commune</i>, + <a href="#page_303">303</a>.<br/> +Ant-eaters, <a href="#page_299">299</a>, + <a href="#page_315">315</a>, <a href="#page_349">349</a>, + <a href="#page_350">350</a>, <a href="#page_353">353</a>.<br/> +Antelopes, <a href="#page_317">317</a>.<br/> +<i>Anthracosaurus Russelli</i>, <a href="#page_190">190</a>.<br/> +<i>AnthrapalĹ“mon gracilis</i>, + <a href="#page_180">180</a>.<br/> +<i>Antilocapra</i>, <a href="#page_318">318</a>.<br/> +<i>Antilope quadricornis</i>, <a href="#page_318">318</a>.<br/> +Antwerp Crag, <a href="#page_325">325</a>.<br/> +Apes, <a href="#page_323">323</a>.<br/> +<i>Apiocrinus</i>, <a href="#page_231">231</a>.<br/> +<i>Apteryx</i>, <a href="#page_346">346</a>, + <a href="#page_348">348</a>.<br/> +Aqueous rocks, <a href="#page_15">15</a>.<br/> +<i>Arachnida</i> of the Coal-measures, + <a href="#page_181">181</a>.<br/> +Aralo-Caspian Beds, <a href="#page_326">326</a>.<br/> +<i>Araucaria</i>, <a href="#page_262">262</a>.<br/> +<i>Araucarioxylon</i>, <a href="#page_170">170</a>.<br/> +<i>Arca</i>, <a href="#page_198">198</a>; + <i>antiqua</i>, <a href="#page_199">199</a>.<br/> +<i>ArchĹ“ocidaris</i>, <a href="#page_178">178</a>.<br/> +<i>ArchĹ“ocyathus</i>, <a href="#page_82">82</a>.<br/> +<i>ArchĹ“opteryx</i>, <a href="#page_252">252</a>, + <a href="#page_281">281</a>; <i>macrura</i>, + <a href="#page_252">252</a>, <a href="#page_253">253</a>.<br/> +<i>ArchĹ“ospĹ“rinĹ“</i>, + <a href="#page_75">75</a>.<br/> +<i>Archimedes</i>, <a href="#page_184">184</a>; + <i>Wortheni</i>, <a href="#page_183">183</a>.<br/> +<i>Archiulus</i>, <a href="#page_182">182</a>.<br/> +Arctic regions, Miocene flora of, + <a href="#page_310">310</a>.<br/> +<i>Arctocyon</i>, <a href="#page_304">304</a>.<br/> +Arenaceous rocks, <a href="#page_20">20</a>.<br/> +<i>Arenicolites</i>, <a href="#page_83">83</a>; + <i>didymus</i>, <a href="#page_88">88</a>.<br/> +Arenig rocks, <a href="#page_92">92</a>, + <a href="#page_94">94</a>.<br/> +Argillaceous rocks, <a href="#page_20">20</a>.<br/> +Armadillos, <a href="#page_299">299</a>, + <a href="#page_351">351</a>, <a href="#page_353">353</a>.<br/> +<i>Artiodactyle Ungulates</i>, <a href="#page_300">300</a>.<br/> +<i>Asaphus</i>, <a href="#page_108">108</a>; + <i>tyrannus</i>, <a href="#page_107">107</a>, + <a href="#page_108">108</a>.<br/> +<i>Ascoceras</i>, <a href="#page_130">130</a>.<br/> +<i>Aspidella</i>, <a href="#page_76">76</a>.<br/> +<i>Aspidura loricata</i>, <a href="#page_210">210</a>.<br/> +<i>Astarte borealis</i>, <a href="#page_338">338</a>.<br/> +<i>Asterophyllites</i>, <a href="#page_137">137</a>, + <a href="#page_196">196</a>.<br/> +<i>Asterosteus</i>, <a href="#page_152">152</a>.<br/> +<i>AstrĹ“idĹ“</i>, <a href="#page_231">231</a>.<br/> +<i>AstrĹ“ospongia</i>, <a href="#page_118">118</a>, + <a href="#page_139">139</a>.<br/> +<i>Astylospongia</i>, <a href="#page_98">98</a>; + <i>prĹ“morsa</i>, <a href="#page_139">139</a>.<br/> +<i>Athyris</i>, <a href="#page_110">110</a>, + <a href="#page_127">127</a>, <a href="#page_147">147</a>, + <a href="#page_198">198</a>; <i>subtilita</i>, + <a href="#page_185">185</a>.<br/> +Atlantic Ooze, <a href="#page_22">22</a>, + <a href="#page_23">23</a>.<br/> +<i>Atrypa</i>, <a href="#page_127">127</a>; <i>congesta</i>, + <a href="#page_127">127</a>; <i>hemispĹ“rica</i>, + <a href="#page_127">127</a>; <i>reticularis</i>, + <a href="#page_147">147</a>, <a href="#page_148">148</a>.<br/> +Auger-shells, <a href="#page_293">293</a>.<br/> +Aurochs, <a href="#page_356">356</a>.<br/> +Aves (<i>see</i> Birds).<br/> +<a name="page_397"><span class="page">Page 397</span></a> +<i>Avicula</i>, <a href="#page_235">235</a>; <i>cantorta</i>, + <a href="#page_211">211</a>, <a href="#page_212">212</a>; + <i>socialis</i>, <a href="#page_211">211</a>.<br/> +"Avicula contorta Beds", <a href="#page_204">204</a>, + <a href="#page_212">212</a>.<br/> +<i>AviculidĹ“</i>, <a href="#page_198">198</a>, + <a href="#page_269">269</a>.<br/> +<i>Aviculopecten</i>, <a href="#page_186">186</a>.<br/> +<i>Axophyllum</i>, <a href="#page_173">173</a>.<br/> +Aymestry Limestone, <a href="#page_116">116</a>, + <a href="#page_117">117</a>.<br/> +Azoic rocks, <a href="#page_67">67</a>. +</p> + +<p class="index"> +<i>Baculites</i>, <a href="#page_273">273</a>; + <i>anceps</i>, <a href="#page_274">274</a>.<br/> +Bagshot and Bracklesham Beds, <a href="#page_287">287</a>.<br/> +<i>Bakewellia</i>, <a href="#page_198">198</a>.<br/> +<i>BalĹ“na</i>, <a href="#page_315">315</a>.<br/> +Bala Group, <a href="#page_93">93</a>, + <a href="#page_94">94</a>.<br/> +Bala Limestone, <a href="#page_93">93</a>.<br/> +<i>BalanidĹ“</i>, <a href="#page_267">267</a>.<br/> +<i>Banksia</i>, <a href="#page_262">262</a>, + <a href="#page_308">308</a>.<br/> +Barbadoes Earth, <a href="#page_33">33</a>.<br/> +Barnacles, <a href="#page_267">267</a>.<br/> +Bath Oolite, <a href="#page_227">227</a>.<br/> +Bats, <a href="#page_304">304</a>, <a href="#page_322">322</a>.<br/> +Bears, <a href="#page_330">330</a>, <a href="#page_359">359</a>.<br/> +Beaver, <a href="#page_322">322</a>, <a href="#page_336">336</a>.<br/> +Beetles, <a href="#page_182">182</a>, <a href="#page_311">311</a>.<br/> +<i>Belemnitella mucronata</i>, <a href="#page_275">275</a>.<br/> +<i>Beleminites</i>, <a href="#page_214">214</a>, + <a href="#page_240">240</a>, <a href="#page_274">274</a>; + <i>canaliculatus</i>, <a href="#page_241">241</a>.<br/> +<i>BelemnitidĹ“</i>, <a href="#page_240">240</a>, + <a href="#page_285">285</a>.<br/> +<i>Belemnoteuthis</i>, <a href="#page_240">240</a>.<br/> +<i>Belinurus</i>, <a href="#page_179">179</a>.<br/> +<i>Bellerophon</i>, <a href="#page_111">111</a>, + <a href="#page_129">129</a>, <a href="#page_148">148</a>, + <a href="#page_186">186</a>; <i>Argo</i>, + <a href="#page_111">111</a>.<br/> +<i>Belodon</i>, <a href="#page_218">218</a>; + <i>Carolinensis</i>, <a href="#page_219">219</a>.<br/> +<i>Belosepia</i>, <a href="#page_295">295</a>.<br/> +<i>Beloteuthis subcostata</i>, <a href="#page_239">239</a>, + <a href="#page_240">240</a>.<br/> +Bembridge Beds, <a href="#page_288">288</a>.<br/> +<i>Beryx</i>, <a href="#page_276">276</a>; + <i>Lewesiensis</i>, <a href="#page_276">276</a>.<br/> +<i>Beyrichia</i>, <a href="#page_107">107</a>; + <i>complicata</i>, <a href="#page_107">107</a>.<br/> +Bird's-eye Limestone, <a href="#page_95">95</a>, + <a href="#page_96">96</a>.<br/> +Birds, of the Trias, <a href="#page_222">222</a>; + of the Jurassic, <a href="#page_251">251-253</a>; + of the Cretaceous, <a href="#page_281">281</a>, + <a href="#page_282">282</a>; of the Eocene, + <a href="#page_297">297</a>; of the Post-Pliocene, + <a href="#page_345">345-348</a>.<br/> +<i>Bison priscus</i>, <a href="#page_356">356</a>.<br/> +Bituminous Schists of Caithness, <a href="#page_36">36</a>.<br/> +Bivalves (<i>see</i> Lamellibranchiata).<br/> +Black-lead (<i>see</i> Graphite).<br/> +Black-River Limestone, <a href="#page_95">95</a>, + <a href="#page_96">96</a>.<br/> +<i>Blastoidea</i>, <a href="#page_176">176</a>; + of the Devonian, <a href="#page_143">143</a>; + of the Carboniferous, <a href="#page_176">176</a>.<br/> +<i>BoidĹ“</i>, <a href="#page_296">296</a>.<br/> +Bolderberg Beds, <a href="#page_307">307</a>.<br/> +Bone-bed, of the Upper Ludlow, <a href="#page_116">116</a>; + of the Trias, <a href="#page_224">224</a>.<br/> +Bony Fishes (<i>see</i> Teleostean Fishes).<br/> +<i>Bos primigenius</i>, <a href="#page_356">356</a>; + <i>taurus</i>, <a href="#page_356">356</a>.<br/> +Boulder-clay, <a href="#page_337">337</a>.<br/> +<i>Bourgueticrinus</i>, <a href="#page_266">266</a>.<br/> +Bovey-Tracy Beds, <a href="#page_305">305</a>, + <a href="#page_309">309</a>.<br/> +<i>Brachiopoda</i>, <a href="#page_125">125</a>; + of the Cambrian rocks, <a href="#page_87">87</a>; + of the Lower Silurian, <a href="#page_108">108-110</a>; + of the Upper Silurian, <a href="#page_125">125-128</a>; + of the Devonian, <a href="#page_147">147</a>, + <a href="#page_148">148</a>; of the Carboniferous, + <a href="#page_184">184-186</a>; of the Permian, + <a href="#page_198">198</a>; of the Trias, + <a href="#page_211">211</a>; of the Jurassic, + <a href="#page_234">234</a>; of the Cretaceous, + <a href="#page_268">268</a>; of the Eocene, + <a href="#page_292">292</a>.<br/> +<i>Brachymetopus</i>, <a href="#page_179">179</a>.<br/> +Brachyurous Crustaceans, <a href="#page_180">180</a>, + <a href="#page_197">197</a>.<br/> +Bradford Clay, <a href="#page_227">227</a>.<br/> +Breaks in the Geological and Palæontological record, + <a href="#page_44">44-52</a>.<br/> +Breccia, <a href="#page_19">19</a>.<br/> +Brick-earths, <a href="#page_339">339</a>.<br/> +Bridlington Crag, <a href="#page_325">325</a>, + <a href="#page_326">326</a>, <a href="#page_336">336</a>.<br/> +Brittle-stars (<i>see</i> Ophiuroidea).<br/> +<i>Bronteus</i>, <a href="#page_145">145</a>.<br/> +<i>BrontotheridĹ“</i>, <a href="#page_316">316</a>.<br/> +<i>Brontotherium ingens</i>, <a href="#page_316">316</a>.<br/> +<i>Brontozoum</i>, <a href="#page_206">206</a>.<br/> +<i>Buccinum</i>, <a href="#page_237">237</a>.<br/> +<i>Bucklandia</i>, <a href="#page_230">230</a>.<br/> +<i>Bulimus</i>, <a href="#page_294">294</a>.<br/> +Bunter Sandstein, <a href="#page_203">203</a>, + <a href="#page_204">204</a>, <a href="#page_206">206</a>.<br/> +Butterflies, <a href="#page_233">233</a>, + <a href="#page_311">311</a>.<br/> +<i>Byssoarca</i>, <a href="#page_198">198</a>. +</p> + +<p class="index"> +Cainozoic (<i>see</i> Kainozoic).<br/> +Calamaries, <a href="#page_239">239</a>.<br/> +<i>Calamites</i>, <a href="#page_165">165</a>, + <a href="#page_166">166</a>, <a href="#page_196">196</a>; + <i>cannĹ“formis</i>, <a href="#page_166">166</a>.<br/> +Calcaire Grossier, <a href="#page_287">287</a>, + <a href="#page_288">288</a>.<br/> +Calcareous rocks, <a href="#page_20">20-32</a>; Tufa, + <a href="#page_21">21</a>.<br/> +Calciferous Sand-rock, <a href="#page_95">95</a>, + <a href="#page_96">96</a>.<br/> +<i>Calveria</i>, <a href="#page_178">178</a>.<br/> +<i>Calymene</i>, <a href="#page_108">108</a>, + <a href="#page_123">123</a>; <i>Blumenbachii</i>, + <a href="#page_107">107</a>.<br/> +<i>Camarophoria globulina</i>, <a href="#page_198">198</a>.<br/> +Cambrian period, <a href="#page_77">77-90</a>; + rocks of, in Britain, <a href="#page_77">77</a>, + <a href="#page_78">78</a>; in Bohemia, <a href="#page_79">79</a>; + in North America, <a href="#page_79">79</a>; life of, + <a href="#page_80">80-90</a>.<br/> +<i>CamelopardalidĹ“</i>, <a href="#page_317">317</a>.<br/> +Camels, <a href="#page_317">317</a>, <a href="#page_354">354</a>.<br/> +<i>Canis lupus</i>, <a href="#page_336">336</a>; + <i>Parisiensis</i>, <a href="#page_304">304</a>.<br/> +Caradoc rocks, <a href="#page_93">93</a>, + <a href="#page_94">94</a>, <a href="#page_96">96</a>.<br/> +Carbon, origin of, <a href="#page_36">36</a>.<br/> +Carboniferous Limestone, <a href="#page_157">157</a>, + <a href="#page_158">158</a>.<br/> +Carboniferous period, <a href="#page_157">157-192</a>; + rocks of, <a href="#page_157">157-160</a>; life of, + <a href="#page_160">160-191</a>.<br/> +Carboniferous Slates of Ireland, <a href="#page_135">135</a>, + <a href="#page_158">158</a>, <a href="#page_159">159</a>.<br/> +<i>Carcharias</i>, <a href="#page_275">275</a>.<br/> +<i>Carcharodon</i>, <a href="#page_295">295</a>, + <a href="#page_312">312</a>; <i>productus</i>, + <a href="#page_313">313</a>.<br/> +<i>Cardinia</i>, <a href="#page_235">235</a>.<br/> +<i>Cardiocarpon</i>, <a href="#page_137">137</a>.<br/> +<i>Cardiola</i>, <a href="#page_128">128</a>; + <i>fibrosa</i>, <a href="#page_128">128</a>; + <i>interrupta</i>, <a href="#page_128">128</a>.<br/> +<i>Cardita</i>, <a href="#page_213">213</a>, + <a href="#page_292">292</a>; <i>planicosta</i>, + <a href="#page_292">292</a>, <a href="#page_293">293</a>.<br/> +<i>Cardium</i>, <a href="#page_292">292</a>; + <i>RhĹ“ticum</i>, <a href="#page_211">211</a>, + <a href="#page_212">212</a>.<br/> +Caribou, <a href="#page_355">355</a>.<br/> +<i>Carnivora</i>, of the Eocene, <a href="#page_304">304</a>; + of the Miocene, <a href="#page_322">322</a>; of the Pliocene, + <a href="#page_330">330</a>, <a href="#page_331">331</a>; + of the Post-Pliocene, <a href="#page_359">359-361</a>.<br/> +<i>Caryocaris</i>, <a href="#page_107">107</a>, + <a href="#page_108">108</a>.<br/> +<i>Caryocrinus ornatus</i>, <a href="#page_106">106</a>.<br/> +<i>Castor fiber</i>, <a href="#page_336">336</a>.<br/> +<i>Castoroides Ohioensis</i>, <a href="#page_361">361</a>.<br/> +Catastrophism, theory of, <a href="#page_3">3</a>.<br/> +<i>Catopterus</i>, <a href="#page_214">214</a>.<br/> +Cauda-Galli Grit, <a href="#page_135">135</a>, + <a href="#page_137">137</a>.<br/> +<i>Caulopteris</i>, <a href="#page_136">136</a>, + <a href="#page_164">164</a>.<br/> +Cave-bear, <a href="#page_360">360</a>.<br/> +Cave-deposits, <a href="#page_337">337</a>, + <a href="#page_339">339</a>, <a href="#page_341">341-344</a>.<br/> +Cave-hyæna, <a href="#page_360">360</a>.<br/> +Cave-lion, <a href="#page_361">361</a>.<br/> +Caves, formation of, <a href="#page_341">341</a>; deposits in, + <a href="#page_342">342</a>.<br/> +<i>Cavicornia</i>, <a href="#page_317">317</a>.<br/> +Cement-stones, <a href="#page_31">31</a>.<br/> +<i>Cephalaspis</i>, <a href="#page_152">152</a>.<br/> +<i>Cephalopoda</i>, of the Cambrian period, + <a href="#page_88">88</a>; of the Lower Silurian, + <a href="#page_111">111-114</a>; of the Upper Silurian, + <a href="#page_130">130</a>; of the Devonian, + <a href="#page_149">149</a>; of the Carboniferous, + <a href="#page_186">186</a>, <a href="#page_187">187</a>; + of the Permian, <a href="#page_199">199</a>; of the Trias, + <a href="#page_212">212</a>; of the Jurassic, + <a href="#page_237">237-240</a>; of the Cretaceous, + <a href="#page_272">272-275</a>; of the Eocene, + <a href="#page_294">294</a>; of the Miocene, + <a href="#page_312">312</a>.<br/> +<a name="page_398"><span class="page">Page 398</span></a> +<i>Ceratiocaris</i>, <a href="#page_108">108</a>.<br/> +<i>Ceratites</i>, <a href="#page_212">212-214</a>; + <i>nodosus</i>, <a href="#page_212">212</a>.<br/> +<i>Ceratodus</i>, <a href="#page_214">214</a>; + <i>altus</i>, <a href="#page_214">214</a>; + <i>Fosteri</i>, <a href="#page_214">214</a>; + <i>serratus</i>, <a href="#page_214">214</a>.<br/> +<i>Ceriopora</i>, <a href="#page_145">145</a>; + <i>Hamiltonensis</i>, <a href="#page_146">146</a>.<br/> +<i>Cerithium</i>, <a href="#page_213">213</a>, + <a href="#page_293">293</a>; <i>hexagonum</i>, + <a href="#page_294">294</a>.<br/> +<i>CervidĹ“</i>, of the Miocene period, + <a href="#page_314">314</a>; of the Pliocene, + <a href="#page_329">329</a>; of the Post-Pliocene, + <a href="#page_354">354</a>, <a href="#page_355">355</a>.<br/> +<i>Cervus</i>, <a href="#page_317">317</a>; + <i>capreolus</i>, <a href="#page_336">336</a>, + <a href="#page_354">354</a>; <i>elaphus</i>, + <a href="#page_336">336</a>, <a href="#page_355">355</a>; + <i>megaceros</i>, <a href="#page_354">354</a>, + <a href="#page_355">355</a>; <i>tarandus</i>, + <a href="#page_354">354</a>.<br/> +<i>Cestracion Philippi</i>, <a href="#page_188">188</a>, + <a href="#page_255">255</a>.<br/> +Cestracionts, of the Devonian, <a href="#page_154">154</a>; + of the Carboniferous, <a href="#page_188">188</a>; + of the Permian, <a href="#page_199">199</a>; + of the Trias, <a href="#page_214">214</a>; + of the Jurassic, <a href="#page_242">242</a>; + of the Cretaceous, <a href="#page_275">275</a>.<br/> +<i>Cetacea</i>, <a href="#page_299">299</a>; + of the Eocene, <a href="#page_299">299</a>; + of the Miocene, <a href="#page_315">315</a>.<br/> +<i>Cetiosaurus</i>, <a href="#page_249">249</a>, + <a href="#page_25">25</a>.<br/> +<i>ChĹ“ropotamus</i>, <a href="#page_302">302</a>.<br/> +<i>ChĹ“tetes</i>, <a href="#page_105">105</a>, + <a href="#page_173">173</a>; <i>tumidus</i>, + <a href="#page_174">174</a>.<br/> +Chain-coral, <a href="#page_119">119</a>.<br/> +Chalk, <a href="#page_259">259</a>; structure of, + <a href="#page_21">21-23</a>; Foraminifera of, + <a href="#page_22">22</a>, <a href="#page_263">263</a>; + origin of, <a href="#page_23">23</a>; with flints, + <a href="#page_253">253</a>; without flints, + <a href="#page_259">259</a>.<br/> +<i>Chama</i>, <a href="#page_236">236</a>.<br/> +<i>ChamĹ“rops</i>, <a href="#page_308">308</a>; + <i>Helvetica</i>, <a href="#page_309">309</a>.<br/> +Chazy Limestone, <a href="#page_95">95</a>, + <a href="#page_96">96</a>.<br/> +<i>Cheiroptera</i>, of the Eocene, <a href="#page_304">304</a>, + <a href="#page_305">305</a>; of the Miocene, + <a href="#page_322">322</a>.<br/> +<i>Cheirotherium</i>, <a href="#page_215">215</a>, + <a href="#page_216">216</a>.<br/> +<i>Cheirurus</i>, <a href="#page_108">108</a>, + <a href="#page_123">123</a>; <i>bimucronatus</i>, + <a href="#page_124">124</a>.<br/> +<i>Chelichnus Duncani</i>, <a href="#page_202">202</a>.<br/> +<i>Chelone Benstedi</i>, <a href="#page_280">280</a>; + <i>planiceps</i>, <a href="#page_251">251</a>.<br/> +<i>Chelonia</i>, of the Permian, <a href="#page_202">202</a>; + of the Jurassic, <a href="#page_251">251</a>; + of the Cretaceous, <a href="#page_280">280</a>; + of the Eocene, <a href="#page_296">296</a>; + of the Miocene, <a href="#page_213">213</a>.<br/> +<i>Chemnitzia</i>, <a href="#page_213">213</a>.<br/> +Chemung Group, <a href="#page_135">135</a>, + <a href="#page_136">136</a>, <a href="#page_137">137</a>.<br/> +Chert, <a href="#page_34">34</a>.<br/> +Chillesford Beds, <a href="#page_325">325</a>, + <a href="#page_326">326</a>, <a href="#page_336">336</a>.<br/> +<i>Chonetes</i>, <a href="#page_127">127</a>, + <a href="#page_147">147</a>, <a href="#page_184">184</a>; + <i>Hardrensis</i>, <a href="#page_185">185</a>.<br/> +<i>Chonophyllum</i>, <a href="#page_173">173</a>.<br/> +<i>Cidaris</i>, <a href="#page_266">266</a>.<br/> +Cincinnati Group, <a href="#page_95">95</a>, + <a href="#page_96">96</a>.<br/> +<i>Cinnamomum polymorphum</i>, <a href="#page_309">309</a>.<br/> +Cinnamon-trees, <a href="#page_262">262</a>, + <a href="#page_290">290</a>, <a href="#page_306">306</a>, + <a href="#page_308">308</a>, <a href="#page_309">309</a>.<br/> +<i>Cladodus</i>, <a href="#page_188">188</a>.<br/> +Claiborne Beds, <a href="#page_289">289</a>.<br/> +<i>Clathropora</i>, <a href="#page_145">145</a>; + <i>intertexta</i>, <a href="#page_146">146</a>.<br/> +Clay, <a href="#page_20">20</a>; Red, origin of, + <a href="#page_35">35</a>.<br/> +Clay-ironstone, nodules of, <a href="#page_31">31</a>.<br/> +<i>Cleidophorus</i>, <a href="#page_111">111</a>.<br/> +<i>Cleodora</i>, <a href="#page_312">312</a>.<br/> +<i>Climacograptus</i>, <a href="#page_101">101</a>, + <a href="#page_119">119</a>.<br/> +Clinton Formation, <a href="#page_116">116</a>, + <a href="#page_117">117</a>.<br/> +<i>Clisiophyllum</i>, <a href="#page_173">173</a>.<br/> +<i>ClupeidĹ“</i>, <a href="#page_276">276</a>.<br/> +<i>Clymenia</i>, <a href="#page_149">149</a>; + <i>Sedgwickii</i>, <a href="#page_149">149</a>.<br/> +Coal, <a href="#page_36">36</a>; structure of, + <a href="#page_163">163</a>; mode of formation of, + <a href="#page_162">162</a>.<br/> +Coal-measures, <a href="#page_159">159</a>, + <a href="#page_160">160</a>; mineral characters of, + <a href="#page_159">159</a>; mode of formation of, + <a href="#page_160">160</a>, <a href="#page_162">162</a>; + plants of, <a href="#page_162">162-170</a>.<br/> +Coccoliths, <a href="#page_261">261</a>.<br/> +<i>Coccosteus</i>, <a href="#page_151">151</a>, + <a href="#page_152">152</a>.<br/> +<i>Cochliodus</i>, <a href="#page_188">188</a>; + <i>cantortus</i>, <a href="#page_189">189</a>.<br/> +<i>Coleoptera</i>, <a href="#page_182">182</a>, + <a href="#page_311">311</a>.<br/> +<i>Colossochelys Atlas</i>, <a href="#page_313">313</a>.<br/> +<i>Columnaria</i>, <a href="#page_105">105</a>; + <i>alveolata</i>, <a href="#page_105">105</a>.<br/> +<i>Comatula</i>, <a href="#page_232">232</a>, + <a href="#page_266">266</a>.<br/> +Conclusions to be drawn from Fossils, + <a href="#page_52">52-56</a>.<br/> +Concretions, calcareous, <a href="#page_29">29</a>; + phosphatic, <a href="#page_31">31</a>; of clay-ironstone, + <a href="#page_31">31</a>; of manganese, + <a href="#page_31">31</a>.<br/> +Conglomerate, <a href="#page_18">18</a>.<br/> +<i>ConiferĹ“</i>, <a href="#page_262">262</a>; + wood of, <a href="#page_13">13</a>; of Devonian period, + <a href="#page_138">138</a>; of the Carboniferous, + <a href="#page_170">170</a>; of the Permian, + <a href="#page_196">196</a>; of the Trias, + <a href="#page_208">208</a>; of the Jurassic period, + <a href="#page_230">230</a>.<br/> +Coniston Flags and Grits, <a href="#page_116">116</a>.<br/> +Connecticut Sandstones, footprints of, + <a href="#page_222">222</a>, <a href="#page_346">346</a>.<br/> +<i>Conocoryphe Mathewi</i>, <a href="#page_85">85</a>; + <i>Sultzeri</i>, <a href="#page_85">85</a>.<br/> +Conodonts, <a href="#page_114">114</a>, + <a href="#page_131">131</a>.<br/> +<i>Constellaria</i>, <a href="#page_105">105</a>.<br/> +Constricting serpents of the Eocene, + <a href="#page_296">296</a>.<br/> +Contemporaneity of strata, <a href="#page_44">44-46</a>.<br/> +Continuity, theory of, <a href="#page_5">5-7</a>.<br/> +<i>Conularia</i>, <a href="#page_111">111</a>, + <a href="#page_129">129</a>, <a href="#page_148">148</a>, + <a href="#page_186">186</a>, <a href="#page_199">199</a>, + <a href="#page_237">237</a>; <i>ornata</i>, + <a href="#page_149">149</a>.<br/> +<i>Conulus</i>, <a href="#page_186">186</a>.<br/> +<i>Conus</i>, <a href="#page_293">293</a>.<br/> +Coomhola Grits, <a href="#page_158">158</a>, + <a href="#page_159">159</a>.<br/> +Coprolites, <a href="#page_31">31</a>, + <a href="#page_243">243</a>.<br/> +Coralline Crag, <a href="#page_324">324</a>.<br/> +Corallines, <a href="#page_25">25</a>.<br/> +<i>Corallium</i>, <a href="#page_311">311</a>.<br/> +Coral-rag, <a href="#page_227">227</a>, + <a href="#page_229">229</a>, <a href="#page_230">230</a>.<br/> +Coral-reefs, <a href="#page_24">24-26</a>.<br/> +Coral-rock, <a href="#page_26">26</a>.<br/> +Coral-sand, <a href="#page_19">19</a>, + <a href="#page_26">26</a>.<br/> +Corals, <a href="#page_103">103</a>; of the Lower Silurian, + <a href="#page_104">104</a>; of the Upper Silurian, + <a href="#page_119">119</a>; of the Devonian, + <a href="#page_140">140-143</a>; of the Carboniferous, + <a href="#page_172">172-175</a>; of the Permian, + <a href="#page_197">197</a>; of the Trias, + <a href="#page_209">209</a>; of the Jurassic, + <a href="#page_230">230</a>, <a href="#page_231">231</a>; + of the Cretaceous, <a href="#page_266">266</a>; + of the Eocene, <a href="#page_292">292</a>; of the + Miocene, <a href="#page_311">311</a>.<br/> +<i>Corbula</i>, <a href="#page_235">235</a>.<br/> +Cornbrash, <a href="#page_227">227</a>, + <a href="#page_229">229</a>.<br/> +Corniferous Limestone, <a href="#page_135">135</a>, + <a href="#page_137">137</a>.<br/> +<i>Cornulites</i>, <a href="#page_123">123</a>.<br/> +Cornus, <a href="#page_262">262</a>.<br/> +<i>Coryphodon</i>, <a href="#page_300">300</a>.<br/> +Cowries, <a href="#page_259">259</a>, + <a href="#page_271">271</a>, <a href="#page_293">293</a>.<br/> +Crabs, <a href="#page_180">180</a>, <a href="#page_197">197</a>, + <a href="#page_233">233</a>, <a href="#page_267">267</a>.<br/> +Crag, Red, <a href="#page_324">324</a>; White, + <a href="#page_324">324</a>; Norwich, + <a href="#page_324">324</a>; Antwerp, + <a href="#page_325">325</a>; Bridlington, + <a href="#page_325">325</a>; Coralline, + <a href="#page_324">324</a>.<br/> +<i>Crania</i>, <a href="#page_110">110</a>, + <a href="#page_127">127</a>, <a href="#page_198">198</a>, + <a href="#page_269">269</a>; <i>Ignabergensis</i>, + <a href="#page_269">269</a>.<br/> +<i>Crassatella</i>, <a href="#page_292">292</a>.<br/> +<i>Crepidophyllum</i>, <a href="#page_142">142</a>; + <i>Archiaci</i>, <a href="#page_142">142</a>.<br/> +Cretaceous period, <a href="#page_256">256-283</a>; + rocks of, in Britain, <a href="#page_257">257-259</a>; + in North America, <a href="#page_260">260</a>, + <a href="#page_261">261</a>; life of, + <a href="#page_261">261-283</a>.<br/> +Crinoidal Limestone, <a href="#page_24">24</a>, + <a href="#page_25">25</a>.<br/> +<i>Crinoidea</i>, <a href="#page_120">120</a>; of the Cambrian, + <a href="#page_82">82</a>; of the Lower Silurian, + <a href="#page_105">105</a>; of the Upper Silurian, + <a href="#page_120">120-122</a>; of the Devonian, + <a href="#page_143">143</a>; of the Carboniferous, + <a href="#page_175">175</a>; of the Permian, + <a href="#page_197">197</a>; of the Triss, + <a href="#page_209">209</a>; of the Jurassic, + <a href="#page_231">231</a>; of the Cretaceous, + <a href="#page_266">266</a>; of the Eocene, + <a href="#page_292">292</a>.<br/> +<i>Crioceras</i>, <a href="#page_273">273</a>; + <i>cristatum</i>, <a href="#page_274">274</a>.<br/> +<i>Crocodilia</i>, <a href="#page_218">218</a>; of the Trias, + <a href="#page_218">218</a>; of the Jurassic, + <a href="#page_251">251</a>; of the Cretaceous, + <a href="#page_280">280</a>; of the Eocene, + <a href="#page_296">296</a>, <a href="#page_297">297</a>.<br/> +Cromer Forest-bed, <a href="#page_336">336</a>.<br/> +<i>Crossozamites</i>, <a href="#page_230">230</a>.<br/> +<i>Crotalocrinus</i>, <a href="#page_122">122</a>.<br/> +<a name="page_399"><span class="page">Page 399</span></a> +<i>Crustacea</i>, of the Cambrian, <a href="#page_83">83-87</a>; + of the Lower Silurian, <a href="#page_107">107</a>, + <a href="#page_108">108</a>; of the Upper Silurian, + <a href="#page_123">123-125</a>; of the Devonian, + <a href="#page_144">144</a>; of the Carboniferous, + <a href="#page_178">178-181</a>; of the Permian, + <a href="#page_197">197</a>; of the Trias, + <a href="#page_210">210</a>; of the Jurassic, + <a href="#page_233">233</a>; of the Cretaceous, + <a href="#page_267">267</a>.<br/> +Cryptogams, <a href="#page_164">164</a>, + <a href="#page_262">262</a>.<br/> +<i>Ctenacanthus</i>, <a href="#page_188">188</a>.<br/> +<i>Ctenodonta</i>, <a href="#page_111">111</a>.<br/> +<i>Cupressus</i>, <a href="#page_262">262</a>.<br/> +Cursores, <a href="#page_297">297</a>, + <a href="#page_346">346</a>.<br/> +Cuttle-fishes (<i>see</i> Dibranchiate Cephalopods).<br/> +<i>Cyathocrinus</i>, <a href="#page_175">175</a>.<br/> +<i>Cyathophyllum</i>, <a href="#page_119">119</a>, + <a href="#page_142">142</a>, <a href="#page_173">173</a>.<br/> +<i>Cycadopteris</i>, <a href="#page_262">262</a>.<br/> +Cycads, <a href="#page_208">208</a>; of the Carboniferous, + <a href="#page_170">170</a>; of the Permian, + <a href="#page_197">197</a>; of the Trias, + <a href="#page_208">208</a>; of the Jurassic, + <a href="#page_230">230</a>; of the Cretaceous, + <a href="#page_261">261</a>.<br/> +<i>Cyclas</i>, <a href="#page_268">268</a>.<br/> +<i>Cyclonema</i>, <a href="#page_129">129</a>.<br/> +<i>Cyclophthalmus senior</i>, <a href="#page_181">181</a>.<br/> +<i>Cyclostoma</i>, <a href="#page_294">294</a>; + <i>Arnoudii</i>, <a href="#page_294">294</a>.<br/> +<i>Cynodraco</i>, <a href="#page_220">220</a>.<br/> +<i>CyprĹ“a</i>, <a href="#page_271">271</a>, + <a href="#page_293">293</a>; <i>elegans</i>, + <a href="#page_393">393</a>.<br/> +Cypress, <a href="#page_262">262</a>, + <a href="#page_308">308</a>, <a href="#page_311">311</a>.<br/> +<i>Cypridina</i>, <a href="#page_145">145</a>.<br/> +Cypridina Slates, <a href="#page_145">145</a>.<br/> +<i>Cyrena</i>, <a href="#page_235">235</a>, + <a href="#page_268">268</a>, <a href="#page_292">292</a>.<br/> +<i>Cyrtina</i>, <a href="#page_213">213</a>, + <a href="#page_214">214</a>.<br/> +<i>Cyrtoceras</i>, <a href="#page_114">114</a>.<br/> +<i>Cystiphyllum</i>, <a href="#page_119">119</a>, + <a href="#page_142">142</a>, <a href="#page_173">173</a>; + <i>vesiculosum</i>, <a href="#page_141">141</a>.<br/> +<i>Cystoidea</i>, <a href="#page_105">105-107</a>; of the + Cambrian, <a href="#page_82">82</a>; of the Lower Silurian, + <a href="#page_106">106</a>; of the Upper Silurian, + <a href="#page_120">120</a>. +</p> + +<p class="index"> +Dachstein Beds, <a href="#page_205">205</a>, + <a href="#page_206">206</a>.<br/> +<i>Dadoxylon</i>, <a href="#page_138">138</a>, + <a href="#page_170">170</a>.<br/> +<i>Daonella</i>, <a href="#page_211">211</a>; + <i>Lommelli</i>, <a href="#page_211">211</a>.<br/> +<i>Dasornis Londinensis</i>, <a href="#page_297">297</a>.<br/> +Decapod Crustaceans, <a href="#page_180">180</a>.<br/> +Deer, <a href="#page_317">317</a>, + <a href="#page_329">329</a>, <a href="#page_354">354</a>.<br/> +<i>Deinosauria</i>, <a href="#page_248">248</a>; of the Trias, + <a href="#page_221">221</a>; of the Jurassic, + <a href="#page_248">248-251</a>; of the Cretaceous, + <a href="#page_277">277-279</a>.<br/> +<i>Deinotherium</i>, <a href="#page_319">319</a>, + <a href="#page_320">320</a>; <i>giganteum</i>, + <a href="#page_320">320</a>.<br/> +Denbighshire Flags and Grits, <a href="#page_115">115</a>.<br/> +<i>Dendrocrinus</i>, <a href="#page_82">82</a>.<br/> +<i>Dendrograptus</i>, <a href="#page_100">100</a>.<br/> +Desmids, <a href="#page_138">138</a>, + <a href="#page_262">262</a>.<br/> +Devonian Formation, <a href="#page_133">133-136</a>; origin of + name, <a href="#page_133">133</a>; relation to Old Red + Sandstone, <a href="#page_133">133</a>, + <a href="#page_134">134</a>; of Devonshire, + <a href="#page_134">134</a>; of North America, + <a href="#page_135">135</a>, <a href="#page_136">136</a>; + life of, <a href="#page_136">136-156</a>.<br/> +<i>Diadema</i>, <a href="#page_266">266</a>.<br/> +Diatoms, <a href="#page_33">33</a>; of the Devonian, + <a href="#page_138">138</a>; of the Carboniferous, + <a href="#page_164">164</a>; of flints, + <a href="#page_261">261</a>; of Richmond Earth, + <a href="#page_33">33</a>, <a href="#page_307">307</a>.<br/> +Dibranchiate Cephalopods, <a href="#page_112">112</a>; + of the Trias, <a href="#page_212">212</a>; of the Jurassic, + <a href="#page_239">239-241</a>; of the Cretaceous, + <a href="#page_274">274</a>, <a href="#page_275">275</a>; + of the Eocene, <a href="#page_294">294</a>; of the Miocene, + <a href="#page_312">312</a>.<br/> +<i>Diceras</i>, <a href="#page_236">236</a>; + <i>arietina</i>, <a href="#page_236">236</a>.<br/> +Diceras Limestone, <a href="#page_227">227</a>, + <a href="#page_236">236</a>.<br/> +<i>Dichobune</i>, <a href="#page_303">303</a>.<br/> +<i>Dichograptus</i>, <a href="#page_101">101</a>; + <i>octobrachiatus</i>, <a href="#page_101">101</a>.<br/> +Dicotyledonous plants, <a href="#page_262">262</a>.<br/> +<i>Dicotyles antiquus</i>, <a href="#page_317">317</a>.<br/> +<i>Dicranograptus</i>, <a href="#page_101">101</a>, + <a href="#page_119">119</a>.<br/> +<i>Dictyonema</i>, <a href="#page_89">89</a>, + <a href="#page_100">100</a>, <a href="#page_119">119</a>; + <i>sociale</i>, <a href="#page_89">89</a>.<br/> +<i>Dicynodon</i>, <a href="#page_220">220</a>; + <i>lacerticeps</i>, <a href="#page_221">221</a>.<br/> +<i>Didelphys</i>, <a href="#page_254">254</a>, + <a href="#page_315">315</a>; <i>gypsorum</i>, + <a href="#page_299">299</a>.<br/> +<i>Didus ineptus</i>, <a href="#page_348">348</a>.<br/> +<i>Didymograptus</i>, <a href="#page_101">101</a>; + <i>divaricatus</i>, <a href="#page_102">102</a>.<br/> +<i>Dikellocephalus Celticus</i>, <a href="#page_84">84</a>; + <i>Minnesotensis</i>, <a href="#page_84">84</a>.<br/> +<i>Dimorphodon</i>, <a href="#page_247">247</a>.<br/> +<i>Dinichthys</i>, <a href="#page_153">153</a>; + <i>Hertzeri</i>, <a href="#page_151">151</a>.<br/> +<i>Ditoceras</i>, <a href="#page_303">303</a>; + <i>mirabilis</i>, <a href="#page_304">304</a>.<br/> +<i>Dinocerata</i>, <a href="#page_303">303</a>, + <a href="#page_304">304</a>.<br/> +<i>Dinophis</i>, <a href="#page_296">296</a>.<br/> +<i>Dinornis</i>, <a href="#page_346">346</a>, + <a href="#page_348">348</a>; <i>elephantopus</i>, + <a href="#page_346">346</a>; <i>giganteus</i>, + <a href="#page_346">346</a>.<br/> +<i>Dinosauria</i> (see <i>Deinosauria</i>).<br/> +<i>Dinotherium</i> (see <i>Deinotherium</i>).<br/> +<i>Diphyphyllum</i>, <a href="#page_142">142</a>.<br/> +<i>Diplograptus</i>, <a href="#page_101">101</a>, + <a href="#page_119">119</a>; <i>pristis</i>, + <a href="#page_102">102</a>.<br/> +<i>Dipnoi</i>, <a href="#page_153">153</a>, + <a href="#page_187">187</a>, <a href="#page_215">215</a>.<br/> +<i>Diprotodon</i>, <a href="#page_348">348</a>, + <a href="#page_349">349</a>; <i>australis</i>, + <a href="#page_348">348</a>.<br/> +<i>Diptera</i>, <a href="#page_311">311</a>.<br/> +<i>Discina</i>, <a href="#page_87">87</a>, + <a href="#page_110">110</a>, <a href="#page_127">127</a>, + <a href="#page_198">198</a>.<br/> +<i>Discoidea</i>, <a href="#page_266">266</a>; + <i>cylindrica</i>, <a href="#page_267">267</a>.<br/> +<i>Dithyrocaris</i>, <a href="#page_179">179</a>; + <i>Scouleri</i>, <a href="#page_180">180</a>.<br/> +Dodo, <a href="#page_348">348</a>.<br/> +Dog whelks.<br/> +Dolomite, <a href="#page_293">293</a>.<br/> +Dolomitic Couglomerate of Bristol, <a href="#page_201">201</a>, + <a href="#page_219">219</a>.<br/> +Dolphins, <a href="#page_299">299</a>, + <a href="#page_315">315</a>.<br/> +<i>Dorcatherium</i>, <a href="#page_317">317</a>.<br/> +Downton Sandstone, <a href="#page_116">116</a>.<br/> +<i>Draco volans</i>, <a href="#page_245">245</a>.<br/> +Dragon-flies, <a href="#page_311">311</a>.<br/> +Drift, Glacial, <a href="#page_337">337</a>.<br/> +<i>Dremotherium</i>, <a href="#page_317">317</a>.<br/> +<i>Dromatherium sylvestre</i>, <a href="#page_223">223</a>, + <a href="#page_224">224</a>.<br/> +<i>Dryandra</i>, <a href="#page_262">262</a>.<br/> +<i>Dryopithecus</i>, <a href="#page_323">323</a>.<br/> +Dugougs, <a href="#page_299">299</a>. +</p> + +<p class="index"> +<i>Echinodermata</i>, of the Cambrian, + <a href="#page_82">82</a>; of the Lower Silurian, + <a href="#page_105">105</a>; of the Upper Silurian, + <a href="#page_120">120</a>; of the Devonian, + <a href="#page_143">143</a>; of the Carboniferous, + <a href="#page_175">175</a>; of the Permian, + <a href="#page_197">197</a>; of the Trias, + <a href="#page_209">209</a>; of the Jurassic, + <a href="#page_231">231</a>; of the Cretaceous, + <a href="#page_266">266</a>; of the Eocene, + <a href="#page_292">292</a>.<br/> +<i>Echinoidea</i>, <a href="#page_177">177</a>; of the Upper + Silurian, <a href="#page_120">120</a>; or the Devonian, + <a href="#page_143">143</a>; of the Carboniferous, + <a href="#page_177">177</a>; of the Permian, + <a href="#page_197">197</a>; of the Jurassic, + <a href="#page_233">233</a>; of the Cretaceous, + <a href="#page_266">266</a>.<br/> +<i>Edentata</i>, <a href="#page_349">349</a>; + of the Eocene, <a href="#page_299">299</a>; + of the Miocene, <a href="#page_315">315</a>; + of the Post-Pliocene, <a href="#page_349">349-353</a>.<br/> +<i>Edriocrinus</i>, <a href="#page_122">122</a>.<br/> +Eifel Limostone, <a href="#page_135">135</a>.<br/> +<i>Elasmobranchii</i> (<i>See</i> Placoid Fishes).<br/> +<i>Elasmosaurus</i>, <a href="#page_276">276</a>.<br/> +Elephants, <a href="#page_319">319</a>, + <a href="#page_320">320</a>, <a href="#page_330">330</a>.<br/> +<i>Elphas</i>, <a href="#page_320">320</a>; + <i>Americanus</i>, <a href="#page_357">357</a>; + <i>antiquus</i>, <a href="#page_329">329</a>, + <a href="#page_330">330</a>, <a href="#page_336">336</a>, + <a href="#page_341">341</a>, <a href="#page_357">357</a>; + <i>Falconeri</i>, <a href="#page_359">359</a>; + <i>Melitensis</i>, <a href="#page_359">359</a>; + <i>meridionalis</i>, <a href="#page_329">329</a>, + <a href="#page_330">330</a>, <a href="#page_336">336</a>, + <a href="#page_357">357</a>; <i>planifrons</i>, + <a href="#page_321">321</a>; <i>primigenius</i>, + <a href="#page_339">339</a>, <a href="#page_341">341</a>, + <a href="#page_357">357</a>, <a href="#page_358">358</a>.<br/> +Elk, <a href="#page_354">354</a>; Irish, + <a href="#page_354">354</a>, <a href="#page_355">355</a>.<br/> +<i>Ellipsocephalus Hoffi</i>, <a href="#page_84">84</a>.<br/> +<i>Elotherium</i>, <a href="#page_317">317</a>.<br/> +<i>EmydidĹ“</i>, <a href="#page_296">296</a>.<br/> +<i>Emys</i>, <a href="#page_280">280</a>.<br/> +Enaliosaurians, <a href="#page_219">219</a>, + <a href="#page_242">242</a>, <a href="#page_276">276</a>.<br/> +Encrinital warble, <a href="#page_24">24</a>.<br/> +<a name="page_400"><span class="page">Page 400</span></a> +<i>Encrinurus</i>, <a href="#page_123">123</a>.<br/> +<i>Encrinus liliiformis</i>, <a href="#page_209">209</a>, + <a href="#page_210">210</a>.<br/> +Endogenous plants, <a href="#page_261">261</a>.<br/> +<i>Endophyllum</i>, <a href="#page_173">173</a>.<br/> +<i>Endothyra</i>, <a href="#page_171">171</a>; + <i>Bailyi</i>, <a href="#page_172">172</a>.<br/> +Engis skull, <a href="#page_364">364</a>.<br/> +<i>Entomis</i>, <a href="#page_145">145</a>.<br/> +<i>Entomoconchus Scouleri</i>, <a href="#page_179">179</a>, + <a href="#page_180">180</a>.<br/> +Eocene period, <a href="#page_284">284</a>; rocks of, in + Britain, <a href="#page_287">287</a>, + <a href="#page_288">288</a>; in France, + <a href="#page_288">288</a>; in North America, + <a href="#page_288">288</a>, <a href="#page_289">289</a>; + life of, <a href="#page_289">289, 305</a>.<br/> +<i>Eocidaris</i>, <a href="#page_197">197</a>.<br/> +<i>Eophyton</i>, <a href="#page_80">80</a>; + <i>Linneanum</i>, <a href="#page_81">81</a>.<br/> +Eophyton Sandstone, <a href="#page_79">79</a>.<br/> +<i>Eosaurus Acadianus</i>, <a href="#page_191">191</a>.<br/> +Eozoic rocks, <a href="#page_67">67</a>.<br/> +<i>Eozoön Bavaricum</i>, <a href="#page_76">76</a>.<br/> +<i>Eozoön Canadense</i>, <a href="#page_68">68</a>, + <a href="#page_76">76</a>; appearance of, in mass, + <a href="#page_69">69</a>; minute structure of, + <a href="#page_70">70</a>, <a href="#page_71">71</a>; + affinities of, with <i>Foraminifera</i>, + <a href="#page_71">71-74</a>.<br/> +<i>EphemeridĹ“</i>, <a href="#page_145">145</a>, + <a href="#page_183">183</a>.<br/> +<i>EquisetaceĹ“</i>, <a href="#page_166">166</a>.<br/> +<i>Equisetites</i>, <a href="#page_196">196</a>.<br/> +<i>EquidĹ“</i>, <a href="#page_301">301</a>, + <a href="#page_302">302</a>, <a href="#page_316">316</a>, + <a href="#page_328">328</a>.<br/> +<i>Equus</i>, <a href="#page_302">302</a>; <i>caballus</i>, + <a href="#page_354">354</a>; <i>excelsus</i>, + <a href="#page_328">328</a>; <i>fossilis</i>, + <a href="#page_336">336</a>, <a href="#page_354">354</a>.<br/> +<i>Eridophyllum</i>, <a href="#page_142">142</a>.<br/> +<i>Eryon arctiformis</i>, <a href="#page_233">233</a>, + <a href="#page_234">234</a>.<br/> +<i>Eschara</i>, <a href="#page_267">267</a>.<br/> +<i>EscharidĹ“</i>, <a href="#page_267">267</a>.<br/> +<i>Escharina</i>, <a href="#page_267">267</a>; + <i>Oceani</i>, <a href="#page_268">268</a>.<br/> +<i>Estheria</i>, <a href="#page_145">145</a>, + <a href="#page_179">179</a>, <a href="#page_210">210</a>; + <i>tenella</i>, <a href="#page_180">180</a>.<br/> +<i>Eucalyptocrinus</i>, <a href="#page_122">122</a>; + <i>polydactylus</i>, <a href="#page_122">122</a>.<br/> +<i>Eucladia</i>, <a href="#page_120">120</a>.<br/> +<i>Euomphalus</i>, <a href="#page_128">128</a>, + <a href="#page_148">148</a>, <a href="#page_186">186</a>, + <a href="#page_199">199</a>, <a href="#page_213">213</a>; + <i>discors</i>, <a href="#page_129">129</a>.<br/> +<i>Euplectella</i>, <a href="#page_265">265</a>.<br/> +<i>Euproöps</i>, <a href="#page_179">179</a>.<br/> +European Bison, <a href="#page_356">356</a>.<br/> +<i>Eurypterida</i>, <a href="#page_124">124</a>, + <a href="#page_179">179</a>; of the Upper Silurian, + <a href="#page_124">124</a>; of the Devonian, + <a href="#page_144">144</a>.<br/> +Even-toed Ungulates, <a href="#page_300">300</a>, + <a href="#page_317">317</a>, <a href="#page_354">354</a>.<br/> +Exogenous plants, <a href="#page_266">266</a>.<br/> +<i>Exogyra</i>, <a href="#page_236">236</a>; + <i>virgula</i>, <a href="#page_236">236</a>.<br/> +Extinction of species, <a href="#page_57">57</a>, + <a href="#page_58">58</a>. +</p> + +<p class="index"> +<i>Fagus</i>, <a href="#page_262">262</a>.<br/> +Faluns, <a href="#page_306">306</a>.<br/> +Fan-palms, <a href="#page_308">308</a>.<br/> +<i>Favistella</i>, <a href="#page_105">105</a>.<br/> +<i>Favostites</i>, <a href="#page_119">119</a>, + <a href="#page_142">142</a>; <i>Gothlandica</i>, + <a href="#page_143">143</a>; <i>hemisphĹ“rica</i>, + <a href="#page_143">143</a>.<br/> +Faxöe Limestone, <a href="#page_259">259</a>, + <a href="#page_286">286</a>.<br/> +<i>Felis angustus</i>, <a href="#page_330">330</a>; + <i>leo</i>, <a href="#page_361">361</a>; <i>spelĹ“a</i>, + <a href="#page_361">361</a>.<br/> +<i>Fenestella</i>, <a href="#page_108">108</a>, + <a href="#page_125">125</a>, <a href="#page_145">145</a>, + <a href="#page_184">184</a>, <a href="#page_198">198</a>, + <a href="#page_210">210</a>; <i>cribrosa</i>, + <a href="#page_146">146</a>; <i>magnifica</i>, + <a href="#page_146">146</a>; <i>retiformis</i>, + <a href="#page_198">198</a>.<br/> +<i>FenestellidĹ“</i>, <a href="#page_183">183</a>.<br/> +Ferns, of the Devonian, <a href="#page_136">136</a>; + of the Carboniferous, <a href="#page_164">164</a>; + of the Permian, <a href="#page_196">196</a>; of the Trias, + <a href="#page_207">207</a>; of the Jurassic, + <a href="#page_229">229</a>; of the Cretaceous, + <a href="#page_261">261</a>.<br/> +Fig-shells, <a href="#page_293">293</a>.<br/> +Fishes, <a href="#page_150">150</a>; of the Upper Silurian, + <a href="#page_130">130</a>, <a href="#page_131">131</a>; + of the Devonian, <a href="#page_150">150-155</a>; of the + Carboniferous, <a href="#page_187">187</a>, + <a href="#page_188">188</a>; of the Permian, + <a href="#page_199">199</a>, <a href="#page_200">200</a>; + of the Trias, <a href="#page_214">214</a>, + <a href="#page_215">215</a>; of the Jurassic, + <a href="#page_240">240-242</a>; of the Cretaceous, + <a href="#page_275">275</a>, <a href="#page_276">276</a>; of + the Eocene, <a href="#page_295">295</a>, + <a href="#page_296">296</a>; of the Miocene, + <a href="#page_312">312</a>, <a href="#page_313">313</a>.<br/> +Flint, <a href="#page_33">33</a>; structure of, + <a href="#page_34">34</a>; origin of, <a href="#page_34">34</a>; + organisms of, <a href="#page_34">34</a>, + <a href="#page_138">138</a>, <a href="#page_263">263</a>; + of Chalk, <a href="#page_34">34</a>, <a href="#page_259">259</a>, + <a href="#page_261">261</a>. + Human implements associated with bones of extinct Mammals, + <a href="#page_363">363</a>, <a href="#page_364">364</a>.<br/> +Flora (<i>see</i> Plants).<br/> +Footprints of <i>Cheirotherium</i>, <a href="#page_215">215</a>, + <a href="#page_216">216</a>; of the Triassic sandstones of + Connecticut, <a href="#page_222">222</a>.<br/> +<i>Foraminifera</i>, <a href="#page_22">22-24</a>, + <a href="#page_71">71-74</a>; of the Cambrian, + <a href="#page_82">82</a>; of the Lower Silurian, + <a href="#page_98">98</a>; of the Carboniferous, + <a href="#page_171">171</a>, <a href="#page_172">172</a>; + of the Permian, <a href="#page_197">197</a>; of the Trias, + <a href="#page_209">209</a>; of the Jurassic, + <a href="#page_230">230</a>; of the Cretaceous, + <a href="#page_21">21</a>, <a href="#page_22">22</a>, + <a href="#page_263">263</a>; of the Eocene, + <a href="#page_290">290</a>; of the Miocene, + <a href="#page_311">311</a>; of the Post-Pliocene, + <a href="#page_338">338</a>; of Atlantic ooze, + <a href="#page_22">22</a>, <a href="#page_23">23</a>; as + builders of limestone, <a href="#page_24">24</a>, + <a href="#page_25">25</a>, <a href="#page_28">28</a>; + as forming green sands, <a href="#page_34">34</a>.<br/> +<i>Forbesiocrinus</i>, <a href="#page_175">175</a>.<br/> +Forest-bed of Cromer, <a href="#page_336">336</a>.<br/> +Forest-bugs, <a href="#page_311">311</a>.<br/> +Forest-marble, <a href="#page_227">227</a>.<br/> +Formation, definition of, <a href="#page_18">18</a>; + succession of, <a href="#page_42">42</a>.<br/> +Fossiliferous rocks, <a href="#page_14">14-37</a>; + chronological succession of, <a href="#page_37">37-44</a>.<br/> +Fossilisation, processes of, <a href="#page_11">11-14</a>.<br/> +Fossils, definition of, <a href="#page_11">11</a>; distinctive, + of rock-groups, <a href="#page_38">38</a>; conclusions + to be drawn from, <a href="#page_52">52-56</a>; biological + relations of, <a href="#page_57">57-61</a>.<br/> +Foxes, <a href="#page_304">304</a>.<br/> +Fringe-finned Ganoids, <a href="#page_153">153</a>.<br/> +Fucoidal Sandstone, <a href="#page_79">79</a>, + <a href="#page_80">80</a>.<br/> +Fucoids, <a href="#page_80">80</a>, <a href="#page_97">97</a>.<br/> +Fuller's Earth, <a href="#page_227">227</a>, + <a href="#page_229">229</a>.<br/> +<i>Fusulina</i>, <a href="#page_172">172</a>; <i>cylindrica</i>, + <a href="#page_172">172</a>.<br/> +<i>Fusus</i>, <a href="#page_237">237</a>, + <a href="#page_293">293</a>. +</p> + +<p class="index"> +<i>Galeocerdo</i>, <a href="#page_312">312</a>.<br/> +<i>Galerites</i>, <a href="#page_266">266</a>; + <i>albo-galerus</i>, <a href="#page_267">267</a>.<br/> +<i>Galestes</i>, <a href="#page_254">254</a>.<br/> +Ganoid Fishes, <a href="#page_150">150</a>; of the Upper + Silurian, <a href="#page_130">130</a>; of the Devonian, + <a href="#page_150">150-153</a>; of the Carboniferous, + <a href="#page_187">187</a>, <a href="#page_188">188</a>; + of the Permian, <a href="#page_199">199</a>; of the Trias, + <a href="#page_214">214</a>; of the Jurassic, + <a href="#page_241">241</a>; of the Cretaceous, + <a href="#page_275">275</a>; of the Eocene, + <a href="#page_292">292</a>, <a href="#page_293">293</a>.<br/> +GaspĂ© Beds, <a href="#page_134">134</a>.<br/> +<i>Gasteropoda</i>, of the Cambrian, <a href="#page_88">88</a>; + of the Lower Silurian, <a href="#page_111">111</a>; of the + Upper Silurian, <a href="#page_128">128</a>, + <a href="#page_129">129</a>; of the Devonian, + <a href="#page_148">148</a>; of the Carboniferous, + <a href="#page_186">186</a>; of the Permian, + <a href="#page_199">199</a>; of the Trias, + <a href="#page_213">213</a>; of the Jurassic, + <a href="#page_236">236</a>, <a href="#page_237">237</a>; + of the Cretaceous, <a href="#page_271">271</a>; of the + Eocene, <a href="#page_292">292</a>, + <a href="#page_293">293</a>.<br/> +<i>Gastornis Parisiensis</i>, <a href="#page_297">297</a>.<br/> +Gault, <a href="#page_257">257</a>, <a href="#page_258">258</a>.<br/> +Gavial, <a href="#page_251">251</a>, <a href="#page_297">297</a>.<br/> +Genesee Slates, <a href="#page_135">135</a>.<br/> +Geological record, breaks in the, <a href="#page_47">47-52</a>.<br/> +Giraffes, <a href="#page_317">317</a>.<br/> +Glacial period, <a href="#page_335">335</a>; deposits of, + <a href="#page_337">337</a>, <a href="#page_338">338</a>.<br/> +<i>Glandulina</i>, <a href="#page_311">311</a>.<br/> +Glauconite, <a href="#page_34">34</a>, <a href="#page_74">74</a>, + <a href="#page_98">98</a>, <a href="#page_263">263</a>.<br/> +<i>Glauconome</i>, <a href="#page_126">126</a>, + <a href="#page_184">184</a>; <i>pulcherrima</i>, + <a href="#page_183">183</a>.<br/> +Globe Crinoids (<i>see</i> Cystoidea).<br/> +<i>Globigerina</i>, <a href="#page_22">22</a>, + <a href="#page_23">23</a>, <a href="#page_264">264</a>.<br/> +Glutton, <a href="#page_360">360</a>.<br/> +<i>Glyptaster</i>, <a href="#page_120">120</a>.<br/> +<i>Glyptocrinus</i>, <a href="#page_122">122</a>.<br/> +<i>Glyptodon</i>, <a href="#page_351">351</a>, + <a href="#page_352">352</a>; <i>clavipes</i>, + <a href="#page_352">352</a>.<br/> +<i>GlyptolĹ“mus</i>, <a href="#page_153">153</a>.<br/> +Goats, <a href="#page_318">318</a>.<br/> +<i>Goniatites</i>, <a href="#page_130">130</a>, + <a href="#page_149">149</a>, <a href="#page_187">187</a>, + <a href="#page_214">214</a>; <i>JossĹ“</i>, + <a href="#page_187">187</a>.<br/> +<a name="page_401"><span class="page">Page 401</span></a> +<i>GorgonidĹ“</i>, <a href="#page_292">292</a>.<br/> +<i>Grallatores</i>, <a href="#page_297">297</a>.<br/> +Graphite, <a href="#page_36">36</a>; mode of occurrence of, + <a href="#page_36">36</a>, <a href="#page_68">68</a>; origin of, + <a href="#page_36">36</a>.<br/> +<i>Graptolites</i>, <a href="#page_89">89</a>, + <a href="#page_100">100</a>; structure of, + <a href="#page_100">100</a>; of the Lower Silurian, + <a href="#page_100">100-103</a>; of the Upper Silurian, + <a href="#page_118">118</a>, <a href="#page_119">119</a>.<br/> +Great Oolite, <a href="#page_227">227</a>, + <a href="#page_229">229</a>; Upper, <a href="#page_257">257</a>, + <a href="#page_258">258</a>, <a href="#page_260">260</a>.<br/> +Greenland. Miocene plants of, <a href="#page_311">311</a>.<br/> +Greensand, Lower, <a href="#page_257">257</a>.<br/> +Green sands, origin of, <a href="#page_44">44</a>, + <a href="#page_263">263</a>.<br/> +<i>Grevillea</i>, <a href="#page_262">262</a>, + <a href="#page_308">308</a>.<br/> +<i>Griffithides</i>, <a href="#page_197">197</a>.<br/> +Grizzly Bear, <a href="#page_359">359</a>.<br/> +Groond Sloths, <a href="#page_351">351</a>.<br/> +<i>GryphĹ“a</i>, <a href="#page_236">236</a>; + <i>incurva</i>, <a href="#page_236">236</a>.<br/> +Guelph Limestone, <a href="#page_117">117</a>.<br/> +<i>Gulo luscus</i>, <a href="#page_360">360</a>; + <i>spelĹ“us</i>, <a href="#page_360">360</a>.<br/> +Guttenstein Beds, <a href="#page_205">205</a>, + <a href="#page_206">206</a>.<br/> +Gymnospermous Exogens, <a href="#page_262">262</a>.<br/> +Gypsum, <a href="#page_32">32</a>, <a href="#page_193">193</a>, + <a href="#page_204">204</a>.<br/> +<i>Gyracanthus</i>, <a href="#page_188">188</a>.<br/> +<i>Gyroceras</i>, <a href="#page_130">130</a>. +</p> + +<p class="index"> +<i>Hadrosaurus</i>, <a href="#page_278">278</a>.<br/> +<i>Halitherium</i>, <a href="#page_299">299</a>.<br/> +Hallstadt Beds, <a href="#page_205">205</a>, + <a href="#page_206">206</a>.<br/> +<i>Halobia</i>, <a href="#page_211">211</a>.<br/> +<i>Halysites</i>, <a href="#page_119">119</a>; + <i>agglomerata</i>, <a href="#page_120">120</a>; + <i>catenularia</i>, <a href="#page_120">120</a>.<br/> +Hamilton formation, <a href="#page_135">135</a>, + <a href="#page_137">137</a>.<br/> +<i>Hamites</i>, <a href="#page_273">273</a>; + <i>rotundus</i>, <a href="#page_274">274</a>.<br/> +<i>Haplophlebium Barnesi</i>, <a href="#page_182">182</a>.<br/> +Harlech Grits, <a href="#page_78">78</a>, + <a href="#page_79">79</a>.<br/> +<i>Harpes</i>, <a href="#page_108">108</a>, + <a href="#page_123">123</a>; <i>ungula</i>, + <a href="#page_124">124</a>.<br/> +Hastings Sands, <a href="#page_257">257</a>.<br/> +Headon and Osborne series, <a href="#page_287">287</a>, + <a href="#page_288">288</a>.<br/> +Heart-urchins, <a href="#page_311">311</a>.<br/> +<i>Heliolites</i>, <a href="#page_105">105</a>, + <a href="#page_119">119</a>, <a href="#page_266">266</a>.<br/> +<i>Heliophyllum</i>, <a href="#page_142">142</a>, + <a href="#page_173">173</a>; <i>exiguum</i>, + <a href="#page_141">141</a>.<br/> +<i>Helix</i>, <a href="#page_294">294</a>.<br/> +<i>Helladotherium</i>, <a href="#page_317">317</a>.<br/> +<i>Helopora fragilis</i>, <a href="#page_126">126</a>.<br/> +<i>Hemicidaris crenularis</i>, <a href="#page_233">233</a>.<br/> +<i>Hemiptera</i>, <a href="#page_311">311</a>.<br/> +<i>Hemitrochiscus paradoxus</i>, <a href="#page_197">197</a>.<br/> +Hempstead Beds, <a href="#page_306">306</a>.<br/> +<i>Hesperornis</i>, <a href="#page_281">281</a>, + <a href="#page_282">282</a>; <i>regalis</i>, + <a href="#page_282">282</a>.<br/> +<i>Heteropoda</i>, <a href="#page_111">111</a>; + of the Lower Silurian, <a href="#page_111">111</a>; + of the Upper Silurian, <a href="#page_129">129</a>; + of the Devonian, <a href="#page_148">148</a>; + of the Carboniferous, <a href="#page_186">186</a>.<br/> +<i>Hinnites</i>, <a href="#page_213">213</a>.<br/> +<i>Hipparion</i>, <a href="#page_301">301</a>, + <a href="#page_302">302</a>, <a href="#page_316">316</a>, + <a href="#page_317">317</a>, <a href="#page_328">328</a>.<br/> +<i>Hippopodium</i>, <a href="#page_235">235</a>.<br/> +<i>Hippopotamus</i>, <a href="#page_302">302</a>; + <i>amphibus</i>, <a href="#page_317">317</a>, + <a href="#page_329">329</a>; <i>major</i>, + <a href="#page_329">329</a>, <a href="#page_336">336</a>, + <a href="#page_354">354</a>; <i>Sivalensis</i>, + <a href="#page_318">318</a>.<br/> +<i>Hippothoa</i>, <a href="#page_108">108</a>.<br/> +Hippurite Marble, <a href="#page_270">270</a>.<br/> +<i>Hippurites</i>, <a href="#page_270">270</a>; + <i>Toucasiana</i>, <a href="#page_271">271</a>.<br/> +<i>HippuritidĹ“</i>, <a href="#page_270">270</a>, + <a href="#page_285">285</a>.<br/> +<i>Histioderma</i>, <a href="#page_82">82</a>.<br/> +Hollow-horned Ruminants, <a href="#page_317">317</a>.<br/> +<i>Holocystis elegan</i>, <a href="#page_266">266</a>.<br/> +<i>Holopea</i>, <a href="#page_129">129</a>; + <i>Subconica</i>, <a href="#page_129">129</a>.<br/> +<i>Holopella</i>, <a href="#page_129">129</a>, + <a href="#page_213">213</a>; <i>obsoleta</i>, + <a href="#page_129">129</a>.<br/> +<i>Holoptychius</i>, <a href="#page_153">153</a>; + <i>nobilissimus</i>, <a href="#page_154">154</a>.<br/> +Holostomatous Univalves, <a href="#page_236">236</a>, + <a href="#page_293">293</a>.<br/> +Holothurians, <a href="#page_120">120</a>.<br/> +<i>Holtenia</i>, <a href="#page_264">264</a>.<br/> +<i>Homacanthus</i>, <a href="#page_188">188</a>.<br/> +<i>Homalonotus</i>, <a href="#page_123">123</a>, + <a href="#page_145">145</a>; <i>armatus</i>, + <a href="#page_144">144</a>.<br/> +<i>Homo diluvii testis</i>, <a href="#page_313">313</a>.<br/> +Honeycomb Corals, <a href="#page_142">142</a>.<br/> +Hoofed Quadrupeds, <a href="#page_300">300</a>.<br/> +Hudson River Group, <a href="#page_95">95</a>.<br/> +Huronian Period, <a href="#page_75">75</a>, + <a href="#page_76">76</a>; rocks of, + <a href="#page_75">75</a>.<br/> +<i>HyĹ“na crocuta</i>, <a href="#page_360">360</a>; + <i>spelĹ“a</i>, <a href="#page_360">360</a>; + <i>Hipparionum</i>, <a href="#page_330">330</a>.<br/> +<i>HyĹ“nictis</i>, <a href="#page_322">322</a>.<br/> +<i>HyĹ“nodon</i>, <a href="#page_304">304</a>.<br/> +<i>Hyalea D'Orbignyana</i>, <a href="#page_312">312</a>.<br/> +<i>Hybodus</i>, <a href="#page_214">214</a>, + <a href="#page_242">242</a>, <a href="#page_275">275</a>.<br/> +<i>Hydractinia</i>, <a href="#page_265">265</a>.<br/> +Hydroid Zoophytes, <a href="#page_103">103</a>, + <a href="#page_265">265</a>.<br/> +<i>Hymenocaris vermicauda</i>, <a href="#page_84">84</a>, + <a href="#page_88">88</a>.<br/> +<i>Hymenophyllites</i>, <a href="#page_165">165</a>.<br/> +<i>Hymenoptera</i>, <a href="#page_311">311</a>.<br/> +<i>Hyopotamus</i>, <a href="#page_302">302</a>.<br/> +<i>Hyperodapedon</i>, <a href="#page_218">218</a>.<br/> +<i>Hypsiprymnopsis</i>, <a href="#page_224">224</a>.<br/> +<i>Hystrix primigenius</i>, <a href="#page_322">322</a>. +</p> + +<p class="index"> +<i>Ichthyocrinus lĹ“vis</i>, <a href="#page_122">122</a>.<br/> +<i>Ichthyornis</i>, <a href="#page_281">281</a>, + <a href="#page_282">282</a>; <i>dispar</i>, + <a href="#page_281">281</a>, <a href="#page_282">282</a>.<br/> +<i>Ichthyosaurus</i>, <a href="#page_242">242</a>, + <a href="#page_243">243</a>, <a href="#page_276">276</a>; + <i>communis</i>, <a href="#page_242">242</a>.<br/> +<i>Ictitherium</i>, <a href="#page_322">322</a>.<br/> +<i>Iguana</i>, <a href="#page_277">277</a>.<br/> +<i>Iguanodon</i>, <a href="#page_277">277</a>, + <a href="#page_278">278</a>; <i>Mantelli</i>, + <a href="#page_278">278</a>.<br/> +Ilfracombe Group, <a href="#page_134">134</a>.<br/> +<i>IllĹ“nus</i>, <a href="#page_108">108</a>, + <a href="#page_123">123</a>.<br/> +Imperfection of the palæontological record, + <a href="#page_50">50</a>, <a href="#page_51">51</a>.<br/> +Inferior Oolite, <a href="#page_227">227</a>, + <a href="#page_229">229</a>.<br/> +Infusorial Earth, <a href="#page_33">33</a>.<br/> +<i>Inoceramus</i>, <a href="#page_269">269</a>; <i>sulcatus</i>, + <a href="#page_270">270</a>.<br/> +<i>Insectivora</i>, of the Eocene, <a href="#page_305">305</a>; + of the Miocene, <a href="#page_322">322</a>.<br/> +Insects, of the Devonian, <a href="#page_145">145</a>; of the + Carboniferous, <a href="#page_182">182</a>; of the Jurassic, + <a href="#page_233">233</a>; of the Miocene, + <a href="#page_311">311</a>, <a href="#page_312">312</a>.<br/> +Irish Elk, <a href="#page_354">354</a>, + <a href="#page_355">355</a>.<br/> +<i>Ischadites</i>, <a href="#page_99">99</a>, + <a href="#page_118">118</a>.<br/> +Isopod Crustaceans, <a href="#page_84">84</a>. +</p> + +<p class="index"> +Jackson Beds, <a href="#page_289">289</a>.<br/> +Jurassic period, <a href="#page_226">226</a>; rocks of, + <a href="#page_226">226-229</a>; life of, + <a href="#page_229">229-255</a>. +</p> + +<p class="index"> +<i>Kaidacarpum</i>, <a href="#page_230">230</a>.<br/> +Kainozoic period, <a href="#page_44">44</a>, + <a href="#page_284">284-287</a>.<br/> +Kangaroo, <a href="#page_348">348</a>.<br/> +Kelloway Rock, <a href="#page_227">227</a>.<br/> +Kent's Cavern, deposits in, <a href="#page_343">343</a>.<br/> +Keuper, <a href="#page_204">204</a>, + <a href="#page_206">206</a>.<br/> +Kimmeridge Clay, <a href="#page_227">227</a>, + <a href="#page_229">229</a>.<br/> +King-crabs, <a href="#page_84">84</a>, + <a href="#page_124">124</a>, <a href="#page_125">125</a>, + <a href="#page_179">179</a>.<br/> +<i>Koninckia</i>, <a href="#page_213">213</a>, + <a href="#page_214">214</a>.<br/> +Kössen Beds, <a href="#page_205">205</a>, + <a href="#page_206">206</a>. +</p> + +<p class="index"> +<i>Labyrinthodon JĹ“geri</i>, <a href="#page_217">217</a>.<br/> +<i>Labyrinthodontia</i>, <a href="#page_290">290</a>; of the + Carboniferous, <a href="#page_189">189-191</a>; of the Permian, + <a href="#page_200">200</a>; of the Trias, + <a href="#page_215">215-217</a>.<br/> +Lace-corals, <a href="#page_108">108</a>, + <a href="#page_125">125</a>, <a href="#page_145">145</a>, + <a href="#page_183">183</a>, <a href="#page_198">198</a>, + <a href="#page_210">210</a>.<br/> +<i>Lacertilia</i>, <a href="#page_202">202</a>; of the Permian, + <a href="#page_201">201</a>, <a href="#page_202">202</a>; + of the Trias, <a href="#page_217">217</a>, + <a href="#page_218">218</a>; of the Jurassic, + <a href="#page_251">251</a>; of the Cretaceous, + <a href="#page_280">280</a>.<br/> +<i>LĹ“laps</i>, <a href="#page_278">278</a>.<br/> +<i>Lamellibranchiata</i>, of the Cambrian, + <a href="#page_88">88</a>; of the Lower Silurian, + <a href="#page_110">110</a>; of the Upper Silurian, + <a href="#page_128">128</a>; of the Devonian, +<a name="page_402"><span class="page">Page 402</span></a> + <a href="#page_148">148</a>; of the Carboniferous, + <a href="#page_186">186</a>; of the Permian, + <a href="#page_198">198</a>; of the Trias, + <a href="#page_211">211</a>; of the Jurassic, + <a href="#page_234">234-236</a>; of the Cretaceous, + <a href="#page_268">268-270</a>; of the Eocene, + <a href="#page_292">292</a>.<br/> +<i>Lamna</i>, <a href="#page_275">275</a>, + <a href="#page_312">312</a>.<br/> +Lamp-shells (see <i>Brachiopoda</i>).<br/> +Land-tortoises, <a href="#page_313">313</a>.<br/> +<i>LauraceĹ“</i>, <a href="#page_308">308</a>.<br/> +Laurentian period, <a href="#page_65">65</a>; rocks of, + <a href="#page_65">65</a>, <a href="#page_66">66</a>; + Lower Laurentian, <a href="#page_66">66</a>; Upper + Laurentian, <a href="#page_66">66</a>; areas occupied by + Laurentian rocks, <a href="#page_66">66</a>; limestones of, + <a href="#page_66">66</a>; iron-ores of, + <a href="#page_68">68</a>; phosphate of lime of, + <a href="#page_68">68</a>; graphite of, + <a href="#page_68">68</a>; life of, + <a href="#page_67">67-75</a>.<br/> +Leaf-beds of the Isle of Mull, <a href="#page_306">306</a>.<br/> +<i>Leda</i>, <a href="#page_292">292</a>; <i>truncata</i>, + <a href="#page_338">338</a>.<br/> +<i>Leguminosites Marcouanus</i>, <a href="#page_263">263</a>.<br/> +Lemming, <a href="#page_344">344</a>, + <a href="#page_345">345</a>.<br/> +<i>LepadidĹ“</i>, <a href="#page_267">267</a>.<br/> +<i>Lepadocrinus Gebhardi</i>, <a href="#page_106">106</a>.<br/> +<i>Leperditia</i>, <a href="#page_108">108</a>; + <i>canadensis</i>, <a href="#page_107">107</a>.<br/> +<i>Lepidaster</i>, <a href="#page_120">120</a>.<br/> +<i>Lepidechinus</i>, <a href="#page_178">178</a>.<br/> +<i>Lepidesthes</i>, <a href="#page_178">178</a>.<br/> +Lepidodendroids, <a href="#page_166">166</a>, + <a href="#page_167">167</a>, <a href="#page_207">207</a>.<br/> +<i>Lepidodendron</i>, <a href="#page_118">118</a>, + <a href="#page_136">136</a>, <a href="#page_166">166</a>, + <a href="#page_196">196</a>; <i>Sternberg</i>, + <a href="#page_167">167</a>.<br/> +<i>Lepidoptera</i>, <a href="#page_311">311</a>.<br/> +<i>Lepidosiren</i>, <a href="#page_153">153</a>.<br/> +<i>Lepidosteus</i>, <a href="#page_188">188</a>.<br/> +<i>Lepidostrobus</i>, <a href="#page_166">166</a>.<br/> +<i>Lepidotus</i>, <a href="#page_275">275</a>.<br/> +<i>LeptĹ“na</i>, <a href="#page_109">109</a>, + <a href="#page_110">110</a>, <a href="#page_125">125</a>, + <a href="#page_234">234</a>; <i>Liassica</i>, + <a href="#page_235">235</a>; <i>sericea</i>, + <a href="#page_110">110</a>.<br/> +<i>LeptocĹ“lia</i>, <a href="#page_127">127</a>; + <i>plano-convexa</i>, <a href="#page_127">127</a>.<br/> +Lias, <a href="#page_226">226</a>, <a href="#page_227">227</a>, + <a href="#page_229">229</a>.<br/> +Lichas, <a href="#page_108">108</a>.<br/> +<i>Licrophycus Ottawaensis</i>, <a href="#page_97">97</a>.<br/> +Lignitic Formation of North America, + <a href="#page_288">288</a>, <a href="#page_297">297</a>.<br/> +Lily-encrinite, <a href="#page_209">209</a>, + <a href="#page_210">210</a>.<br/> +<i>Lima</i>, <a href="#page_235">235</a>.<br/> +Lime, phosphate of, <a href="#page_30">30</a>, + <a href="#page_31">31</a>.<br/> +Limestone, <a href="#page_23">23-27</a>; varieties of, + <a href="#page_27">27-30</a>; origin of, + <a href="#page_21">21</a>; microscopical structure of, + <a href="#page_26">26</a>; Crinoidal, <a href="#page_24">24</a>; + Foraminiferal, <a href="#page_24">24</a>, + <a href="#page_26">26</a>; coralline, + <a href="#page_24">24</a>; magnesian, <a href="#page_27">27</a>; + metamorphic, <a href="#page_27">27</a>; oolitic, + <a href="#page_28">28-30</a>; pisolitic, + <a href="#page_29">29</a>; bituminous, <a href="#page_36">36</a>; + Laurentian, <a href="#page_67">67</a>.<br/> +<i>LimnĹ“a</i>, <a href="#page_294">294</a>; + <i>pyramidalis</i>, <a href="#page_294">294</a>.<br/> +<i>Limulus</i>, <a href="#page_84">84</a>, + <a href="#page_124">124</a>, <a href="#page_125">125</a>, + <a href="#page_179">179</a>.<br/> +<i>Lingula</i>, <a href="#page_87">87</a>, + <a href="#page_88">88</a>, <a href="#page_110">110</a>, + <a href="#page_127">127</a>, <a href="#page_147">147</a>, + <a href="#page_198">198</a>; <i>Credneri</i>, + <a href="#page_198">198</a>.<br/> +Lingula Flags, <a href="#page_77">77</a>, + <a href="#page_78">78</a>, <a href="#page_79">79</a>, + <a href="#page_88">88</a>.<br/> +<i>Lingulella</i>, <a href="#page_87">87</a>, + <a href="#page_88">88</a>; <i>Davisii</i>, + <a href="#page_88">88</a>; <i>ferruginea</i>, + <a href="#page_88">88</a>.<br/> +<i>Liriodendron</i>, <a href="#page_262">262</a>, + <a href="#page_308">308</a>; <i>Meeki</i>, + <a href="#page_263">263</a>.<br/> +<i>Lithostrotion</i>, <a href="#page_173">173</a>; + <i>irregulare</i>, <a href="#page_174">174</a>.<br/> +<i>Lituites</i>, <a href="#page_130">130</a>.<br/> +Lizards (see <i>Lacertilia</i>).<br/> +Llama, <a href="#page_354">354</a>.<br/> +Llanberis Slates, <a href="#page_79">79</a>.<br/> +Llandeilo rocks, <a href="#page_92">92</a>, + <a href="#page_94">94</a>, <a href="#page_96">96</a>.<br/> +Llandovery rocks, <a href="#page_93">93</a>; Lower, + <a href="#page_93">93</a>; Upper, + <a href="#page_115">115</a>.<br/> +Lobsters, <a href="#page_180">180</a>, + <a href="#page_210">210</a>, <a href="#page_233">233</a>, + <a href="#page_267">267</a>.<br/> +Loess, <a href="#page_339">339</a>.<br/> +London Clay, <a href="#page_287">287</a>, + <a href="#page_288">288</a>.<br/> +Longmynd rocks, <a href="#page_77">77-80</a>, + <a href="#page_83">83</a>.<br/> +<i>Lonsdaleia</i>, <a href="#page_173">173</a>.<br/> +<i>Lophiodon</i>, <a href="#page_316">316</a>.<br/> +<i>Lophophyllum</i>, <a href="#page_173">173</a>.<br/> +Lower Cambrian, <a href="#page_77">77-79</a>; Chalk, + <a href="#page_259">259</a>; Cretaceous, + <a href="#page_257">257</a>, <a href="#page_258">258</a>; + Devonian, <a href="#page_134">134</a>; Eocene, + <a href="#page_287">287</a>, <a href="#page_288">288</a>; + Greensand, <a href="#page_257">257</a>, + <a href="#page_258">258</a>; Helderberg, + <a href="#page_117">117</a>, <a href="#page_118">118</a>; + Laurentian rocks, <a href="#page_66">66</a>; Ludlow rock, + <a href="#page_116">116</a>; Miocene, + <a href="#page_305">305</a>; Old Red Sandstone, + <a href="#page_134">134</a>; Oolites, + <a href="#page_227">229</a>; Silurian period, + <a href="#page_90">90-114</a>; rocks of, in Britain, + <a href="#page_92">92-94</a>; in North America, + <a href="#page_94">94-96</a>; life of, + <a href="#page_97">97-114</a>.<br/> +<i>Loxonema</i>, <a href="#page_186">186</a>, + <a href="#page_199">199</a>, <a href="#page_213">213</a>.<br/> +Ludlow rock, <a href="#page_116">116</a>, + <a href="#page_117">117</a>.<br/> +<i>LycopodiaceĹ“</i>, <a href="#page_118">118</a>, + <a href="#page_136">136</a>, <a href="#page_167">167</a>.<br/> +Lynton Group, <a href="#page_134">134</a>.<br/> +<i>Lyrodesma</i>, <a href="#page_111">111</a>. +</p> + +<p class="index"> +Macaques, <a href="#page_323">323</a>, + <a href="#page_331">331</a>.<br/> +<i>MachĹ“racanthus major</i>, + <a href="#page_151">151</a>, <a href="#page_155">155</a>.<br/> +<i>Machairodus</i>, <a href="#page_221">221</a>, + <a href="#page_249">249</a>, <a href="#page_322">322</a>, + <a href="#page_331">331</a>, <a href="#page_360">360</a>; + <i>cultridens</i>, <a href="#page_331">331</a>.<br/> +<i>Maclurea</i>, <a href="#page_111">111</a>; <i>crenulata</i>, + <a href="#page_112">112</a>.<br/> +<i>Macrocheilus</i>, <a href="#page_186">186</a>, + <a href="#page_199">199</a>, <a href="#page_213">213</a>.<br/> +<i>Macropetalichthys</i>, <a href="#page_152">152</a>; + <i>Sullivanti</i>, <a href="#page_151">151</a>.<br/> +<i>Macrotherium giganteum</i>, <a href="#page_315">315</a>.<br/> +<i>Macrurous Crustaceans</i>, <a href="#page_180">180</a>.<br/> +<i>Mactra</i>, <a href="#page_292">292</a>.<br/> +Maestricht Chalk, <a href="#page_259">259</a>, + <a href="#page_279">279</a>, <a href="#page_286">286</a>.<br/> +Magnesian Limestone, <a href="#page_27">27</a>; nature and + structure of, <a href="#page_28">28</a>; of the Permian + series, <a href="#page_194">194</a>, <a href="#page_196">196</a>.<br/> +Magnolia, <a href="#page_262">262</a>, <a href="#page_290">290</a>, + <a href="#page_310">310</a>.<br/> +<i>Mammalia</i>, of the Trias, <a href="#page_223">223</a>, + <a href="#page_224">224</a>; of the Jurassic, + <a href="#page_253">253</a>, <a href="#page_254">254</a>; + of the Eocene, <a href="#page_299">299-305</a>; of the Miocene, + <a href="#page_313">313-323</a>; of the Pliocene, + <a href="#page_327">327-331</a>; of the Post-Pliocene, + <a href="#page_348">348-362</a>.<br/> +Mammoth, <a href="#page_339">339</a>, <a href="#page_341">341</a>, + <a href="#page_344">344</a>, <a href="#page_357">357-359</a>.<br/> +Man, remains of, in Post-Pliocene deposits, + <a href="#page_341">341</a>, <a href="#page_344">344</a>.<br/> +Manatee, <a href="#page_299">299</a>.<br/> +<i>Mantellia</i>, <a href="#page_230">230</a>; + <i>megalophylla</i>, <a href="#page_230">230</a>.<br/> +Maple, <a href="#page_290">290</a>, <a href="#page_308">308</a>, + <a href="#page_310">310</a>.<br/> +Marble, <a href="#page_28">28</a>; encrinital, + <a href="#page_24">24</a>; statuary, <a href="#page_27">27</a>.<br/> +Marcellus Shales, <a href="#page_135">135</a>.<br/> +<i>Mariacrinus</i>, <a href="#page_122">122</a>.<br/> +Marmots, <a href="#page_322">322</a>.<br/> +Marsupials, <a href="#page_299">299</a>; of the Trias, + <a href="#page_223">223</a>; of the Jurassic, + <a href="#page_253">253</a>, <a href="#page_254">254</a>; + of the Eocene, <a href="#page_299">299</a>; of the Miocene, + <a href="#page_315">315</a>; of the Post-Pliocene, + <a href="#page_348">348</a>, <a href="#page_319">319</a>.<br/> +<i>Marsupiocrinus</i>, <a href="#page_122">122</a>.<br/> +<i>Marsupites</i>, <a href="#page_266">266</a>.<br/> +<i>Mastodon</i>, <a href="#page_319">319</a>, + <a href="#page_321">321</a>, <a href="#page_322">322</a>; + <i>Americanus, angustidens</i>, <a href="#page_322">322</a>; + <i>Arvenensis</i>, <a href="#page_329">329</a>; + <i>longirostris</i>, <a href="#page_322">322</a>; + <i>Ohioticus</i>, <a href="#page_357">357</a>; + <i>Sivalensis</i>, <a href="#page_321">321</a>.<br/> +Medina Sandstone, <a href="#page_116">116</a>.<br/> +<i>Megalichthys</i>, <a href="#page_188">188</a>.<br/> +<i>Megalodon</i>, <a href="#page_148">148</a>.<br/> +<i>Megalomus</i>, <a href="#page_128">128</a>.<br/> +<i>Megalonyx</i>, <a href="#page_351">351</a>.<br/> +<i>Megalosaurus</i>, <a href="#page_249">249</a>, + <a href="#page_278">278</a>.<br/> +<i>Megatherium</i>, <a href="#page_350">350</a>, + <a href="#page_351">351</a>; <i>Cuvieri</i>, + <a href="#page_350">350</a>.<br/> +<i>Melania</i>, <a href="#page_294">294</a>.<br/> +<i>Melonites</i>, <a href="#page_178">178</a>.<br/> +Menevian Group, <a href="#page_77">77-79</a>.<br/> +<i>Menobranchus</i>, <a href="#page_189">189</a>.<br/> +<i>Meristella</i>, <a href="#page_127">127</a>; + <i>cylindrica</i>, <a href="#page_127">127</a>; + <i>intermedia</i>, <a href="#page_127">127</a>; + <i>naviformis</i>, <a href="#page_127">127</a>.<br/> +<i>Mesopithecus</i>, <a href="#page_323">323</a>.<br/> +Mesozoic Period, <a href="#page_44">44</a>.<br/> +<i>Michelinia</i>, <a href="#page_142">142</a>.<br/> +<i>Micraster</i>, <a href="#page_266">266</a>.<br/> +<i>Microlestes</i>, <a href="#page_224">224</a>; + <i>antiquus</i>, <a href="#page_223">223</a>.<br/> +<a name="page_403"><span class="page">Page 403</span></a> +Middle Devonian, <a href="#page_134">134</a>; Eocene, + <a href="#page_287">287</a>, <a href="#page_288">288</a>, + <a href="#page_289">289</a>; Oolites, + <a href="#page_227">227</a>; Silurian, + <a href="#page_91">91</a>.<br/> +Miliolite Limestone, <a href="#page_290">290</a>.<br/> +<i>Millepora</i>, <a href="#page_230">230</a>.<br/> +Millstone Grit, <a href="#page_159">159</a>, + <a href="#page_161">161</a>.<br/> +Miocene period, <a href="#page_305">305</a>; rocks of, + in Britain, <a href="#page_305">305</a>, + <a href="#page_306">306</a>; in France, + <a href="#page_306">306</a>; in Belgium, + <a href="#page_307">307</a>; in Switzerland, + <a href="#page_306">306</a>; in Austria, + <a href="#page_307">307</a>; in Germany, + <a href="#page_307">307</a>; in Italy, + <a href="#page_307">307</a>; in India, + <a href="#page_307">307</a>; in North America, + <a href="#page_307">307</a>; life of, + <a href="#page_308">308-323</a>.<br/> +Mitre-shells, <a href="#page_371">371</a>, + <a href="#page_293">293</a>.<br/> +<i>Mitra</i>, <a href="#page_271">271</a>, + <a href="#page_293">293</a>.<br/> +Moas of New Zealand, <a href="#page_346">346-348</a>.<br/> +<i>Modiolopsis</i>, <a href="#page_111">111</a>; + <i>Solvensis</i>, <a href="#page_88">88</a>.<br/> +Molasse, <a href="#page_306">306</a>.<br/> +Mole, <a href="#page_322">322</a>, + <a href="#page_336">336</a>.<br/> +Monkeys, <a href="#page_305">305</a>, + <a href="#page_331">331</a>.<br/> +Monocotyledonous plant, <a href="#page_262">262</a>.<br/> +<i>Monograptus</i>, <a href="#page_100">100</a>, + <a href="#page_119">119</a>; <i>priodon</i>, + <a href="#page_119">119</a>.<br/> +<i>Monotis</i>, <a href="#page_211">211</a>.<br/> +Monte Bolca, fishes of, <a href="#page_295">295</a>.<br/> +<i>Montlivaltia</i>, <a href="#page_209">209</a>.<br/> +Mosasauroids, <a href="#page_279">279</a>, + <a href="#page_280">280</a>.<br/> +<i>Mosasaurus</i>, <a href="#page_279">279</a>; + <i>Camperi</i>, <a href="#page_279">279</a>; + <i>princeps</i>, <a href="#page_279">279</a>.<br/> +Mountain Limestone, <a href="#page_158">158</a>, + <a href="#page_161">161</a>.<br/> +Mud-fishes, <a href="#page_153">153</a>, + <a href="#page_215">215</a>.<br/> +Mud-turtles, <a href="#page_280">280</a>.<br/> +Mull, Miocene strata of, <a href="#page_306">306</a>.<br/> +<i>Murchisonia</i>, <a href="#page_111">111</a>, + <a href="#page_129">129</a>, <a href="#page_199">199</a>, + <a href="#page_213">213</a>; <i>gracilis</i>, + <a href="#page_111">11</a>.<br/> +<i>Murex</i>, <a href="#page_237">237</a>, + <a href="#page_293">293</a>.<br/> +Muschelkalk, <a href="#page_203">203</a>, + <a href="#page_204">204</a>, <a href="#page_206">206</a>.<br/> +Musk-deer, <a href="#page_317">317</a>.<br/> +Musk-ox, <a href="#page_344">344</a>, + <a href="#page_345">345</a>, <a href="#page_356">356</a>.<br/> +Musk-sheep, <a href="#page_356">356</a>.<br/> +<i>Myliobatis Edwardsii</i>, <a href="#page_296">296</a>.<br/> +<i>Mylodon</i>, <a href="#page_351">351</a>; + <i>robustus</i>, <a href="#page_352">352</a>.<br/> +<i>Myophoria</i>, <a href="#page_211">211</a>; + <i>lineata</i>, <a href="#page_211">211</a>.<br/> +<i>Myriapoda</i> of the Coal, <a href="#page_181">181</a>, + <a href="#page_182">182</a>. +</p> + +<p class="index"> +<i>Nassa</i>, <a href="#page_293">293</a>.<br/> +<i>Natatores</i>, <a href="#page_297">297</a>.<br/> +<i>Natica</i>, <a href="#page_271">271</a>, + <a href="#page_293">293</a>.<br/> +<i>Nautilus</i>, <a href="#page_112">112-114</a>, + <a href="#page_130">130</a>, <a href="#page_149">149</a>, + <a href="#page_186">186</a>, <a href="#page_199">199</a>, + <a href="#page_237">237</a>, <a href="#page_272">272</a>, + <a href="#page_294">294</a>; <i>Danicus</i>, + <a href="#page_272">272</a>; <i>pompilius</i>, + <a href="#page_237">237</a>.<br/> +Neanderthal skull, <a href="#page_364">364</a>.<br/> +Neocomian series, <a href="#page_257">257</a>, + <a href="#page_260">260</a>.<br/> +<i>Neolimulus</i>, <a href="#page_125">125</a>.<br/> +<i>NerinĹ“a</i>, <a href="#page_237">237</a>, + <a href="#page_271">271</a>; <i>Goodhallii</i>, + <a href="#page_237">237</a>.<br/> +<i>Nerita</i>, <a href="#page_393">393</a>.<br/> +<i>Neuroptera</i>, <a href="#page_311">311</a>.<br/> +<i>Neuropteris</i>, <a href="#page_136">136</a>.<br/> +Newer Pliocene, <a href="#page_323">323</a>, + <a href="#page_324">324</a>.<br/> +New Red Sandstone, <a href="#page_193">193</a>, + <a href="#page_203">203</a>.<br/> +Newts, <a href="#page_189">189</a>, <a href="#page_200">200</a>, + <a href="#page_217">217</a>.<br/> +Niagara Limestone, <a href="#page_117">117</a>.<br/> +<i>Nipadites</i>, <a href="#page_290">290</a>; + <i>ellipticus</i>, <a href="#page_290">290</a>.<br/> +<i>NĹ“ggerathia</i>, <a href="#page_197">197</a>.<br/> +Norwich Crag, <a href="#page_324">324</a>.<br/> +<i>Nothosaurus</i>, <a href="#page_219">219</a>; + <i>mirabilis</i>, <a href="#page_219">219</a>.<br/> +<i>Notidanus</i>, <a href="#page_241">241</a>.<br/> +<i>Numenius gypsorum</i>, <a href="#page_297">297</a>.<br/> +<i>Nummulina</i>, <a href="#page_172">172</a>, + <a href="#page_290">290</a>; <i>lĹ“vigata</i>, + <a href="#page_290">290</a>; <i>pristina</i>, + <a href="#page_172">172</a>.<br/> +<i>Nummulitic Limestone</i>, <a href="#page_24">24</a>, + <a href="#page_287">287</a>, <a href="#page_291">291</a>. +</p> + +<p class="index"> +Oak, <a href="#page_262">262</a>, <a href="#page_310">310</a>.<br/> +<i>Obolella</i>, <a href="#page_87">87</a>; <i>sagittalis</i>, + <a href="#page_88">88</a>.<br/> +Odd-toed Ungulates, <a href="#page_300">300</a>, + <a href="#page_315">315</a>, <a href="#page_327">327</a>, + <a href="#page_353">353</a>.<br/> +<i>Odontaspis</i>, <a href="#page_275">275</a>.<br/> +<i>Odontopteris</i>, <a href="#page_165">165</a>; + <i>Schlotheimi</i>, <a href="#page_164">164</a>.<br/> +<i>Odontopteryx</i>, <a href="#page_297">297</a>; + <i>toliapicus</i>, <a href="#page_297">297</a>, + <a href="#page_298">298</a>.<br/> +<i>Odontornithes</i>, <a href="#page_282">282</a>.<br/> +<i>Ogygia</i>, <a href="#page_108">108</a>; <i>Buchii</i>, + <a href="#page_107">107</a>.<br/> +Older Pliocene, <a href="#page_323">323</a>, + <a href="#page_324">324</a>.<br/> +<i>Oldhamia</i>, <a href="#page_81">81</a>; <i>antiqua</i>, + <a href="#page_82">82</a>; slates of Ireland, + <a href="#page_79">79</a>, <a href="#page_80">80</a>.<br/> +Old Red Sandstone, <a href="#page_133">133</a>; origin of name, + <a href="#page_133">133</a>; of Scotland, + <a href="#page_134">134</a>; relations of, to Devonian, + <a href="#page_133">133</a>, <a href="#page_134">134</a>, + <a href="#page_155">155</a>.<br/> +<i>Olenus</i>, <a href="#page_108">108</a>; <i>micrurus</i>, + <a href="#page_88">88</a>.<br/> +Oligocene, <a href="#page_305">305</a>.<br/> +<i>Oligoporus</i>, <a href="#page_178">178</a>.<br/> +Olive-shells, <a href="#page_293">293</a>.<br/> +<i>Omphyma</i>, <a href="#page_119">119</a>.<br/> +<i>Onchus</i>, <a href="#page_130">130</a>; + <i>tenuistriatus</i>, <a href="#page_131">131</a>.<br/> +Oneida Conglomerate, <a href="#page_116">116</a>.<br/> +<i>Onychodus</i>, <a href="#page_153">153</a>; + <i>sigmoides</i>, <a href="#page_151">151</a>.<br/> +Oolitic limestone, structure of, <a href="#page_28">28</a>; + mode of formation of, <a href="#page_30">30</a>.<br/> +Oolitic rocks (<i>see</i> Jurassic).<br/> +Ooze, Atlantic, <a href="#page_22">22</a>, + <a href="#page_33">33</a>.<br/> +<i>Ophidia</i>, <a href="#page_251">251</a>; of the Eocene, + <a href="#page_296">296</a>.<br/> +<i>Ophiuroidea</i>, of the Lower Silurian, + <a href="#page_105">105</a>; of the Upper Silurian, + <a href="#page_120">120</a>; of the Carboniferous, + <a href="#page_177">177</a>; of the Trias, + <a href="#page_210">210</a>; of the Jurassic, + <a href="#page_233">233</a>.<br/> +Opossum, <a href="#page_299">299</a>, + <a href="#page_315">315</a>.<br/> +<i>Orbitoides</i>, <a href="#page_291">291</a>.<br/> +Oriskany Sandstone, <a href="#page_135">135</a>.<br/> +<i>Ormoxylon</i>, <a href="#page_138">138</a>.<br/> +<i>Orohippus</i>, <a href="#page_302">302</a>.<br/> +<i>Orthis</i>, <a href="#page_38">38</a>, + <a href="#page_109">109</a>, <a href="#page_125">125</a>, + <a href="#page_147">147</a>, <a href="#page_184">184</a>, + <a href="#page_199">199</a>; <i>biforata</i>, + <a href="#page_109">109</a>; <i>Davidsoni</i>, + <a href="#page_127">127</a>; <i>elegantula</i>, + <a href="#page_127">127</a>; <i>flabellulum</i>, + <a href="#page_109">109</a>; <i>Hicksii</i>, + <a href="#page_38">38</a>; <i>lenticularis</i>, + <a href="#page_38">38</a>; <i>plicatella</i>, + <a href="#page_110">110</a>; <i>resupinata</i>, + <a href="#page_185">185</a>; <i>subquadrala</i>, + <a href="#page_109">109</a>; <i>testudinaria</i>, + <a href="#page_110">110</a>.<br/> +<i>Orthoceras</i>, <a href="#page_89">89</a>, + <a href="#page_112">112</a>, <a href="#page_113">113</a>, + <a href="#page_130">130</a>, <a href="#page_149">149</a>, + <a href="#page_186">186</a>, <a href="#page_213">213</a>; + <i>crebriseptum</i>, <a href="#page_113">113</a>.<br/> +<i>Orthonota</i>, <a href="#page_111">111</a>.<br/> +<i>Orthoptera</i>, <a href="#page_182">182</a>, + <a href="#page_311">311</a>.<br/> +<i>Osmeroides</i>, <a href="#page_276">276</a>; + <i>Mantelli</i>, <a href="#page_276">276</a>.<br/> +<i>Osmerus</i>, <a href="#page_276">276</a>.<br/> +<i>Ostealepis</i>, <a href="#page_153">153</a>.<br/> +<i>Ostracode</i> Crustaceans of the Cambrian, + <a href="#page_83">83</a>; of the Lower Silurian, + <a href="#page_107">107</a>; of the Upper Silurian, + <a href="#page_123">123</a>; of the Devonian, + <a href="#page_145">145</a>; of the Carboniferous, + <a href="#page_179">179</a>; of the Permian, + <a href="#page_197">197</a>; of the Trias, + <a href="#page_210">210</a>; of the Jurassic, + <a href="#page_233">233</a>; of the Cretaceous, + <a href="#page_267">267</a>.<br/> +<i>Ostrea acuminata</i>, <a href="#page_235">235</a>; + <i>Couloni</i>, <a href="#page_269">269</a>; + <i>deltoidea</i>, <a href="#page_235">235</a>; + <i>distorta</i>, <a href="#page_235">235</a>; + <i>expansa, gregarea</i>, <a href="#page_235">235</a>; + <i>Marshii</i>, <a href="#page_235">235</a>, + <a href="#page_236">236</a>.<br/> +<i>Otodus</i>, <a href="#page_295">295</a>; + <i>obtiquus</i>, <a href="#page_296">296</a>.<br/> +<i>Otozamites</i>, <a href="#page_230">230</a>.<br/> +<i>Otozoum</i>, <a href="#page_206">206</a>.<br/> +<i>Oudenodon</i>, <a href="#page_220">220</a>; + <i>Bainii</i>, <a href="#page_221">221</a>.<br/> +<i>Ovibos moschatus</i>, <a href="#page_356">356</a>.<br/> +Oxford Clay, <a href="#page_227">227</a>, + <a href="#page_229">229</a>.<br/> +<i>Oxyrhina</i>, <a href="#page_312">312</a>; + <i>xiphodon</i>, <a href="#page_313">313</a>.<br/> +Oysters, <a href="#page_235">235</a>, + <a href="#page_236">236</a>, <a href="#page_269">269</a>. +</p> + +<p class="index"> +<i>Pachyphyllum</i>, <a href="#page_173">173</a>.<br/> +<i>PalĹ“arca</i>, <a href="#page_111">111</a>.<br/> +<i>PalĹ“aster</i>, <a href="#page_120">120</a>; + <i>Ruthveni</i>, <a href="#page_121">121</a>.<br/> +<i>Palasterina</i>, <a href="#page_120">120</a>; + <i>primĹ“va</i>, <a href="#page_121">121</a>.<br/> +<i>PalĹ“chinus</i>, <a href="#page_120">120</a>, + <a href="#page_178">178</a>; <i>ellipticus</i>, + <a href="#page_177">177</a>.<br/> +<i>PalĹ“ocaris</i>, <a href="#page_180">180</a>; + <i>typus</i>, <a href="#page_180">180</a>.<br/> +<i>PalĹ“ocoma</i>, <a href="#page_120">120</a>; + <i>Colvini</i>, <a href="#page_121">121</a>.<br/> +<i>PalĹ“ocoryne</i>, <a href="#page_172">172</a>.<br/> +Palæolithic man, remains of, + <a href="#page_363">363-365</a>.<br/> +<i>PalĹ“omanon</i>, <a href="#page_118">118</a>.<br/> +<i>PalĹ“oniscus</i>, <a href="#page_188">188</a>, + <a href="#page_200">200</a>.<br/> +<a name="page_404"><span class="page">Page 404</span></a> +<i>PalĹ“ontina Oolitica</i>, <a href="#page_233">233</a>.<br/> +Palæontological evidence as to Evolution, + <a href="#page_60">60</a>, <a href="#page_372">372-374</a>.<br/> +Palæontological record, imperfection of the, + <a href="#page_50">50</a>, <a href="#page_51">51</a>.<br/> +Palæontology, definition of, <a href="#page_10">10</a>.<br/> +<i>PalĹ“onyctis</i>, <a href="#page_304">304</a>.<br/> +<i>PalĹ“ophis</i>, <a href="#page_296">296</a>; + <i>toliapictus</i>, <a href="#page_296">296</a>; + <i>typhĹ“us</i>, <a href="#page_296">296</a>.<br/> +<i>PalĹ“oreas</i>, <a href="#page_318">318</a>.<br/> +<i>PalĹ“osaurus</i>, <a href="#page_200">200</a>, + <a href="#page_218">218</a>, <a href="#page_219">219</a>; + <i>platyodon</i>, <a href="#page_219">219</a>.<br/> +<i>PalĹ“osiren Beinerti</i>, <a href="#page_200">200</a>.<br/> +<i>PalĹ“otherium</i>, <a href="#page_300">300</a>; + <i>magnum</i>, <a href="#page_301">301</a>.<br/> +<i>PalĹ“oxylon</i>, <a href="#page_170">170</a>.<br/> +Palæozoic period, <a href="#page_44">44</a>.<br/> +Palms, <a href="#page_230">230</a>, <a href="#page_263">263</a>, + <a href="#page_290">290</a>, <a href="#page_308">308</a>, + <a href="#page_309">309</a>.<br/> +<i>Paludina</i>, <a href="#page_257">257</a>, + <a href="#page_294">294</a>.<br/> +<i>PandaneĹ“</i>, <a href="#page_230">230</a>.<br/> +<i>Pandanus</i>, <a href="#page_262">262</a>.<br/> +<i>Paradoxides</i>, <a href="#page_86">86</a>, + <a href="#page_87">87</a>, <a href="#page_108">108</a>; + <i>Bohemicus</i>, <a href="#page_85">85</a>.<br/> +<i>Parasmilia</i>, <a href="#page_266">266</a>.<br/> +<i>Parkeria</i>, <a href="#page_264">264</a>.<br/> +Pear Encrinite, <a href="#page_231">231</a>.<br/> +Pearly Nautilus, <a href="#page_58">58</a>, + <a href="#page_111">111</a>, <a href="#page_112">112</a>, + <a href="#page_237">237</a>.<br/> +Peccaries, <a href="#page_317">317</a>.<br/> +<i>Pecopteris</i>, <a href="#page_136">136</a>, + <a href="#page_165">165</a>, <a href="#page_196">196</a>.<br/> +<i>Pecten GrĹ“nlandicus</i>, <a href="#page_338">338</a>; + <i>Islandicus</i>, <a href="#page_338">338</a>; + <i>Valoniensis</i>, <a href="#page_211">211</a>, + <a href="#page_212">212</a>, <a href="#page_204">204</a>.<br/> +Penarth Beds, <a href="#page_204">204</a>.<br/> +<i>PennatulidĹ“</i>, <a href="#page_292">292</a>.<br/> +<i>Pentacrinus</i>, <a href="#page_231">231</a>; + <i>caput-medusĹ“</i>, <a href="#page_231">231</a>; + <i>fasciculosus</i>, <a href="#page_232">232</a>.<br/> +<i>Pentamerus</i>, <a href="#page_125">125</a>, + <a href="#page_126">126</a>; <i>galeatus</i>, + <a href="#page_126">126</a>; <i>Knightii</i>, + <a href="#page_128">128</a>.<br/> +<i>Pentremites</i> (<i>see</i> Blastoidea).<br/> +<i>Pentremites conoideus</i>, <a href="#page_176">176</a>; + <i>pyriformis</i>, <a href="#page_176">176</a>.<br/> +Perching Birds, <a href="#page_297">297</a>.<br/> +<i>PercidĹ“</i>, <a href="#page_276">276</a>.<br/> +<i>Periechocrinus</i>, <a href="#page_122">122</a>.<br/> +<i>Perissodactyle Ungulates</i>, <a href="#page_300">300</a>, + <a href="#page_315">315</a>, <a href="#page_327">327</a>.<br/> +Permian period, <a href="#page_192">192-202</a>; rocks of, in + Britain, <a href="#page_194">194</a>; in North America, + <a href="#page_194">194</a>; life of, + <a href="#page_195">195-302</a>.<br/> +Persistent types of life, <a href="#page_58">58</a>, + <a href="#page_371">371</a>.<br/> +<i>Petalodus</i>, <a href="#page_188">188</a>.<br/> +<i>Petraster</i>, <a href="#page_120">120</a>.<br/> +Petroleum, origin of, <a href="#page_36">36</a>.<br/> +Pezophaps, <a href="#page_348">348</a>.<br/> +<i>Phacops</i>, <a href="#page_108">108</a>, + <a href="#page_123">123</a>, <a href="#page_145">145</a>; + <i>DowningiĹ“</i>, <a href="#page_124">124</a>; + <i>granulatus</i>, <a href="#page_144">144</a>; + <i>lĹ“vis</i>, <a href="#page_144">144</a>; + <i>latifrons</i>, <a href="#page_144">144</a>, + <a href="#page_145">145</a>; <i>longicaudatus</i>, + <a href="#page_124">124</a>; <i>rana</i>, + <a href="#page_145">145</a>.<br/> +<i>PhĹ“nopora ensiformis</i>, <a href="#page_126">126</a>.<br/> +Phalangers, <a href="#page_348">348</a>.<br/> +Phanerogams, <a href="#page_164">164</a>.<br/> +<i>Phaneropleuron</i>, <a href="#page_153">153</a>.<br/> +<i>Phascolotherium</i>, <a href="#page_253">253</a>, + <a href="#page_254">254</a>.<br/> +<i>Pheronema</i>, <a href="#page_264">164</a>.<br/> +<i>PhillipsastrĹ“a</i>, <a href="#page_142">142</a>.<br/> +<i>Phillipsia</i>, <a href="#page_179">179</a>; + <i>seminifera</i>, <a href="#page_180">180</a>.<br/> +<i>Pholadomya</i>, <a href="#page_235">235</a>.<br/> +<i>Phormosoma</i>, <a href="#page_178">178</a>.<br/> +<i>Phorus</i>, <a href="#page_271">271</a>.<br/> +Phosphate of lime, concretions of, <a href="#page_30">30</a>; + disseminated in rocks, <a href="#page_30">30</a>; origin of, + <a href="#page_31">31</a>.<br/> +<i>Phyllograptus</i>, <a href="#page_102">102</a>; + <i>typus</i>, <a href="#page_102">102</a>.<br/> +<i>Phyllopoda</i>, of the Cambrian, <a href="#page_83">83</a>; + of the Lower Silurian, <a href="#page_108">108</a>; of the + Upper Silurian, <a href="#page_123">123</a>; of the Devonian, + <a href="#page_145">145</a>; of the Carboniferous, + <a href="#page_179">179</a>; of the Permian, + <a href="#page_197">197</a>; of the Trias, + <a href="#page_210">210</a>.<br/> +<i>Phyllopora</i>, <a href="#page_210">210</a>.<br/> +<i>Physa</i>, <a href="#page_294">294</a>; + <i>columnaris</i>, <a href="#page_294">294</a>.<br/> +Pigs, <a href="#page_302">302</a>, <a href="#page_317">317</a>, + <a href="#page_329">329</a>, <a href="#page_354">354</a>.<br/> +Pilton Group, <a href="#page_135">135</a>.<br/> +<i>Pinites</i>, <a href="#page_170">170</a>.<br/> +<i>Pisces</i> (<i>see</i> Fishes).<br/> +<i>Pisolite</i>, <a href="#page_29">29</a>.<br/> +Pisolitic Limestone of France, <a href="#page_259">259</a>, + <a href="#page_286">286</a>.<br/> +<i>Placodus</i>, <a href="#page_220">220</a>; <i>gigas</i>, + <a href="#page_220">220</a>.<br/> +Placoid Fishes, <a href="#page_150">150</a>; of the Upper + Silurian, <a href="#page_130">130</a>, <a href="#page_131">131</a>; + of the Devonian, <a href="#page_153">153-155</a>; of the + Carboniferous, <a href="#page_188">188</a>; of the Permian, + <a href="#page_199">199</a>; of the Trias, + <a href="#page_214">214</a>; of the Jurassic, + <a href="#page_241">241</a>; of the Cretaceous, + <a href="#page_275">275</a>; of the Eocene, + <a href="#page_295">295</a>; of the Miocene, + <a href="#page_312">312</a>.<br/> +<i>Plagiaulax</i>, <a href="#page_254">254</a>.<br/> +<i>Planolites</i>, <a href="#page_122">122</a>; + <i>vulgaris</i>, <a href="#page_123">123</a>.<br/> +<i>Planorbis</i>, <a href="#page_294">294</a>.<br/> +Plants, of the Cambrian, <a href="#page_80">80</a>, + <a href="#page_81">81</a>; of the Lower Silurian, + <a href="#page_97">97</a>, <a href="#page_98">98</a>; of the + Upper Silurian, <a href="#page_118">118</a>; of the Devonian, + <a href="#page_136">136-139</a>; of the Carboniferous, + <a href="#page_163">163-170</a>; of the Permian, + <a href="#page_196">196</a>; of the Trias, + <a href="#page_207">207</a>, <a href="#page_208">208</a>; + of the Jurassic, <a href="#page_229">229</a>, + <a href="#page_230">230</a>; of the Cretaceous, + <a href="#page_261">261-263</a>; of the Eocene, + <a href="#page_289">289</a>, <a href="#page_290">290</a>; + of the Miocene, <a href="#page_308">308-311</a>.<br/> +<i>Plasmopora</i>, <a href="#page_119">119</a>.<br/> +<i>Platanus</i>, <a href="#page_262">262</a>, + <a href="#page_308">308</a>; <i>aceroides</i>, + <a href="#page_309">309</a>.<br/> +<i>Platephemera antiqua</i>, <a href="#page_145">145</a>.<br/> +<i>Platyceras</i>, <a href="#page_128">128</a>, + <a href="#page_148">148</a>; <i>dumosum</i>, + <a href="#page_148">148</a>; <i>multisinuatum</i>, + <a href="#page_129">129</a>; <i>ventricosum</i>, + <a href="#page_129">129</a>.<br/> +<i>Platycrinus</i>, <a href="#page_122">122</a>, + <a href="#page_175">175</a>; <i>tricontadactylus</i>, + <a href="#page_175">175</a>.<br/> +<i>Platyostoma</i>, <a href="#page_129">129</a>; + <i>Niagarense</i>, <a href="#page_129">129</a>.<br/> +Platyrhine Monkeys, <a href="#page_362">362</a>.<br/> +<i>Platyschisma helicites</i>, <a href="#page_129">129</a>.<br/> +<i>Platysomus</i>, <a href="#page_200">200</a>; + <i>gibbosus</i>, <a href="#page_199">199</a>.<br/> +<i>Platystoma</i>, <a href="#page_213">213</a>.<br/> +Pleistocene period; climate of.<br/> +<i>Plesiosaurus</i>, <a href="#page_334">334</a>; + <i>dolichodeirus</i>, <a href="#page_244">244</a>.<br/> +<i>Pleurocystites squamosus</i>, <a href="#page_106">106</a>.<br/> +<i>Pleurotoma</i>, <a href="#page_293">293</a>.<br/> +<i>Pleurotomaria</i>, <a href="#page_111">111</a>, + <a href="#page_129">129</a>, <a href="#page_186">186</a>, + <a href="#page_199">199</a>, <a href="#page_236">236</a>, + <a href="#page_271">271</a>.<br/> +<i>Plicatula</i>, <a href="#page_213">213</a>.<br/> +Pliocene period, <a href="#page_323">323</a>; rocks of, in + Britain, <a href="#page_324">324</a>; in Belgium, + <a href="#page_325">325</a>; in Italy, + <a href="#page_325">325</a>; in North America, + <a href="#page_326">326</a>; life of, + <a href="#page_326">326-331</a>.<br/> +<i>Pliopithecus</i>, <a href="#page_322">322</a>; + <i>antiquus</i>, <a href="#page_323">323</a>.<br/> +<i>Pliosaurus</i>, <a href="#page_245">245</a>.<br/> +<i>Podocarya</i>, <a href="#page_230">230</a>.<br/> +<i>Podozamites</i>, <a href="#page_208">208</a>; + <i>lanceolatus</i>, <a href="#page_209">209</a>.<br/> +Polir-schiefer, <a href="#page_33">33</a>.<br/> +<i>Polycystina</i>, <a href="#page_32">32</a>; of + Barbadoes-earth, <a href="#page_33">33</a>.<br/> +<i>Polypora</i>, <a href="#page_145">145</a>, + <a href="#page_184">184</a>; <i>dendroides</i>, + <a href="#page_183">183</a>.<br/> +<i>Polypterus</i>, <a href="#page_153">153</a>, + <a href="#page_188">188</a>.<br/> +<i>Polystomella</i>, <a href="#page_311">311</a>.<br/> +<i>Polytremacis</i>, <a href="#page_266">266</a>.<br/> +<i>Polyzoa</i>, of the Cambrian, <a href="#page_81">81</a>, + <a href="#page_89">89</a>; of the Lower Silurian, + <a href="#page_108">108</a>; of the Upper Silurian, + <a href="#page_125">125</a>; of the Devonian, + <a href="#page_145">145</a>, <a href="#page_146">146</a>; + of the Carboniferous, <a href="#page_183">183</a>, + <a href="#page_184">184</a>; of the Permian, + <a href="#page_198">198</a>; of the Trias, + <a href="#page_210">210</a>; of the Cretaceous, + <a href="#page_267">267</a>; of the Miocene, + <a href="#page_312">312</a>.<br/> +<i>Populus</i>, <a href="#page_262">262</a>.<br/> +<i>Porcellia</i>, <a href="#page_186">186</a>.<br/> +Porcupines, <a href="#page_322">322</a>.<br/> +Portage Group, <a href="#page_135">135</a>.<br/> +Port-Jackson Shark, <a href="#page_154">154</a>.<br/> +Portland beds, <a href="#page_227">227</a>, + <a href="#page_229">229</a>.<br/> +Post-Glacial deposits, <a href="#page_336">336</a>, + <a href="#page_338">338</a>.<br/> +<a name="page_405"><span class="page">Page 405</span></a> +Post-Pliocene period, <a href="#page_334">334</a>.<br/> +Post-Tertiary period, <a href="#page_286">286</a>.<br/> +<i>Poteriocrinus</i>, <a href="#page_175">175</a>.<br/> +Potsdam Sandstone, <a href="#page_79">79</a>.<br/> +Pre-Glacial deposits, <a href="#page_336">336</a>.<br/> +<i>Prestwichia</i>, <a href="#page_179">179</a>; + <i>rotundata</i>, <a href="#page_179">179</a>.<br/> +<i>Primitia</i>, <a href="#page_107">107</a>; + <i>strangulata</i>, <a href="#page_107">107</a>.<br/> +Primordial Trilobites, <a href="#page_85">85</a>.<br/> +Primordial zone, <a href="#page_79">79</a>.<br/> +<i>Proboscidea</i>, of the Miocene, + <a href="#page_319">319</a>, <a href="#page_322">322</a>; + of the Pliocene, <a href="#page_329">329</a>, + <a href="#page_330">330</a>; of the Post-Pliocene, + <a href="#page_357">357-359</a>.<br/> +<i>Producta</i>, <a href="#page_147">147</a>, + <a href="#page_184">184</a>, <a href="#page_198">198</a>; + <i>horrida</i>, <a href="#page_198">198</a>; + <i>longispina</i>, <a href="#page_185">185</a>; + <i>semireticulata</i>, <a href="#page_185">185</a>.<br/> +<i>Productella</i>, <a href="#page_147">147</a>, + <a href="#page_184">184</a>.<br/> +<i>ProductidĹ“</i>, <a href="#page_147">147</a>, + <a href="#page_211">211</a>.<br/> +<i>Proëtus</i>, <a href="#page_123">123</a>.<br/> +Prong-buck, <a href="#page_318">318</a>.<br/> +<i>Protaster</i>, <a href="#page_120">120</a>; + <i>Sedgwickii</i>, <a href="#page_121">121</a>.<br/> +<i>ProteaceĹ“</i>, <a href="#page_262">262</a>, + <a href="#page_308">308</a>, <a href="#page_309">309</a>.<br/> +<i>Proteus</i>, <a href="#page_189">189</a>.<br/> +<i>Protichnites</i>, <a href="#page_87">87</a>.<br/> +<i>Protocystites</i>, <a href="#page_82">82</a>.<br/> +<i>Protornis Glarisiensis</i>, <a href="#page_279">279</a>.<br/> +<i>Protorosaurus</i>, <a href="#page_201">201</a>, + <a href="#page_202">202</a>; <i>Speneri</i>, + <a href="#page_201">201</a>.<br/> +<i>Protospongia</i>, <a href="#page_81">81</a>; + <i>fenestrata</i>, <a href="#page_88">88</a>.<br/> +<i>Prototaxites</i>, <a href="#page_118">118</a>, + <a href="#page_138">138</a>; <i>Logani</i>, + <a href="#page_139">139</a>.<br/> +<i>Psammobia</i>, <a href="#page_292">292</a>.<br/> +<i>Psammodus</i>, <a href="#page_188">188</a>.<br/> +<i>Psaronius</i>, <a href="#page_136">136</a>, + <a href="#page_164">164</a>.<br/> +<i>Pseudocrinus bifasciatus</i>, <a href="#page_106">106</a>.<br/> +<i>Psilophyton</i>, <a href="#page_118">118</a>, + <a href="#page_137">137</a>, <a href="#page_138">138</a>; + <i>princeps</i>, <a href="#page_138">138</a>.<br/> +<i>Pteranodon</i>, <a href="#page_247">247</a>, + <a href="#page_277">277</a>; <i>longiceps</i>, + <a href="#page_277">277</a>.<br/> +<i>Pteraspis</i>, <a href="#page_130">130</a>, + <a href="#page_152">152</a>; <i>Banksii</i>, + <a href="#page_130">130</a>.<br/> +<i>Pterichthys</i>, <a href="#page_152">152</a>; + <i>cornutus</i>, <a href="#page_153">153</a>.<br/> +<i>PterinĹ“a</i>, <a href="#page_128">128</a>; + <i>subfalcata</i>, <a href="#page_128">128</a>.<br/> +<i>Pteroceras</i>, <a href="#page_237">237</a>, + <a href="#page_271">271</a>.<br/> +<i>Pterodactylus</i>, <a href="#page_245">245</a>, + <a href="#page_277">277</a>; <i>crassirostris</i>, + <a href="#page_246">246</a>.<br/> +<i>Pterophyllum</i>, <a href="#page_208">208</a>, + <a href="#page_230">230</a>; <i>JĹ“geri</i>, + <a href="#page_209">209</a>.<br/> +<i>Pteropoda</i>, of the Cambrian, <a href="#page_88">88</a>; + of the Lower Silurian, <a href="#page_111">111</a>; of the + Upper Silurian, <a href="#page_129">129</a>; of the Devonian, + <a href="#page_148">148</a>; of the Carboniferous, + <a href="#page_186">186</a>; of the Permian, + <a href="#page_199">199</a>; of the Jurassic, + <a href="#page_237">237</a>.<br/> +<i>Pterosauria</i>, <a href="#page_245">245</a>; of the + Jurassic, <a href="#page_245">245-248</a>; of the + Cretaceous, <a href="#page_277">277</a>.<br/> +<i>Pterygotus Anglicus</i>, <a href="#page_124">124</a>, + <a href="#page_125">125</a>.<br/> +<i>Ptilodictya</i>, <a href="#page_108">108</a>, + <a href="#page_125">125</a>; <i>acuta</i>, + <a href="#page_109">109</a>; <i>falciformis</i>, + <a href="#page_109">109</a>; <i>raripora</i>, + <a href="#page_126">126</a>; <i>Schafferi</i>, + <a href="#page_109">109</a>.<br/> +<i>Ptychoceras</i>, <a href="#page_273">273</a>; + <i>Emericianum</i>, <a href="#page_274">274</a>.<br/> +<i>Ptychodus</i>, <a href="#page_275">275</a>.<br/> +<i>Pupa vetusta</i>, <a href="#page_186">186</a>.<br/> +Purbeck Beds, <a href="#page_228">228</a>; Mammals of, + <a href="#page_254">254</a>.<br/> +<i>Puryuroidea</i>, <a href="#page_237">237</a>.<br/> +<i>Pycnodus</i>, <a href="#page_275">275</a>.<br/> +<i>Pyrula</i>, <a href="#page_293">293</a>. +</p> + +<p class="index"> +<i>Quadrumana</i>, of the Eocene, <a href="#page_305">305</a>; + of the Miocene, <a href="#page_322">322</a>, + <a href="#page_323">323</a>; of the Pliocene, + <a href="#page_331">331</a>; of the Post-Pliocene, + <a href="#page_361">361</a>.<br/> +Quadrupeds (<i>see</i> Mammalia).<br/> +Quaternary period, <a href="#page_334">334</a>.<br/> +Quebec Group, <a href="#page_95">95</a>, + <a href="#page_96">96</a>.<br/> +<i>Quercus</i>, <a href="#page_262">262</a>. +</p> + +<p class="index"> +Rabbits, <a href="#page_322">322</a>.<br/> +<i>Rana</i>, <a href="#page_313">313</a>.<br/> +<i>Raptores</i>, <a href="#page_297">297</a>.<br/> +<i>Rasores</i>, <a href="#page_297">297</a>.<br/> +Recent period, <a href="#page_286">286</a>, + <a href="#page_334">334</a>.<br/> +<i>Reptaculites</i>, <a href="#page_99">99</a>.<br/> +Red clays, origin of, <a href="#page_35">35</a>.<br/> +Red Coral, <a href="#page_311">311</a>.<br/> +Red Crag, <a href="#page_324">324</a>.<br/> +Red Deer, <a href="#page_336">336</a>, + <a href="#page_354">354</a>.<br/> +Reindeer, <a href="#page_344">344</a>, + <a href="#page_345">345</a>, <a href="#page_354">354</a>, + <a href="#page_355">355</a>.<br/> +<i>Remopleurides</i>, <a href="#page_188">188</a>.<br/> +Reptiles, <a href="#page_200">200</a>; of the Permian, + <a href="#page_200">200-202</a>; of the Trias, + <a href="#page_217">217-221</a>; of the Jurassic, + <a href="#page_242">242-251</a>; of the Cretaceous, + <a href="#page_276">276-281</a>; of the Eocene, + <a href="#page_296">296</a>, <a href="#page_297">297</a>.<br/> +<i>Retepora</i>, <a href="#page_108">108</a>, + <a href="#page_125">125</a>, <a href="#page_145">145</a>, + <a href="#page_184">184</a>, <a href="#page_198">198</a>, + <a href="#page_210">210</a>; <i>Ehrenbergi</i>, + <a href="#page_198">198</a>; <i>Phillipsi</i>, + <a href="#page_146">146</a>.<br/> +<i>Retiolites</i>, <a href="#page_119">119</a>.<br/> +<i>Retzia</i>, <a href="#page_127">127</a>.<br/> +<i>Rhætic Beds</i>, <a href="#page_204">204-206</a>.<br/> +<i>Rhamphorhynchus</i>, <a href="#page_247">247</a>; + <i>Bucklandi</i>, <a href="#page_248">248</a>.<br/> +<i>RhinoceridĹ“</i>, <a href="#page_315">315</a>.<br/> +<i>Rhinoceros Etruscus</i>, <a href="#page_327">327</a>, + <a href="#page_328">328</a>, <a href="#page_336">336</a>, + <a href="#page_353">353</a>; <i>leptorhinus</i>, + <a href="#page_328">328</a>; <i>megarhinus</i>, + <a href="#page_327">327-329</a>, <a href="#page_336">336</a>, + <a href="#page_353">335</a>; <i>tichorhinus</i>, + <a href="#page_353">353</a>, <a href="#page_354">354</a>.<br/> +<i>Rhinopora verrucosa</i>, <a href="#page_126">126</a>.<br/> +<i>Rhizodus</i>, <a href="#page_188">188</a>.<br/> +<i>Rhombus minimus</i>, <a href="#page_295">295</a>.<br/> +Rhyncholites, <a href="#page_239">239</a>.<br/> +<i>Rhynchonella</i>, <a href="#page_110">110</a>, + <a href="#page_127">127</a>, <a href="#page_147">147</a>, + <a href="#page_184">184</a>, <a href="#page_234">234</a>, + <a href="#page_268">268</a>, <a href="#page_292">292</a>; + <i>cuneata</i>, <a href="#page_127">127</a>; <i>neglecta</i>, + <a href="#page_127">127</a>; <i>pleurodon</i>, + <a href="#page_185">185</a>; <i>varians</i>, + <a href="#page_235">235</a>.<br/> +<i>Rhynchosaurus</i>, <a href="#page_218">218</a>; + <i>articeps</i>, <a href="#page_218">218</a>.<br/> +Rice-shells, <a href="#page_293">293</a>.<br/> +Richmond Earth, <a href="#page_33">33</a>, + <a href="#page_307">307</a>.<br/> +Ringed Worms (<i>see</i> Annelida).<br/> +River-gravels, high-level and low-level, + <a href="#page_340">340</a>, <a href="#page_341">341</a>.<br/> +<i>Robulina</i>, <a href="#page_311">311</a>.<br/> +Rocks, definition of, <a href="#page_14">14</a>; divisions of, + <a href="#page_14">14</a>, <a href="#page_15">15</a>; igneous, + <a href="#page_14">14</a>; aqueous, + <a href="#page_15">15-18</a>; mechanically-formed, + <a href="#page_18">18-20</a>; chemically-formed, + <a href="#page_20">20</a>; organically-formed, + <a href="#page_20">20-37</a>; arenaceous, + <a href="#page_20">20</a>; argillaceous, + <a href="#page_20">20</a>; calcareous, + <a href="#page_20">20-32</a>; siliceous, + <a href="#page_20">20</a>, <a href="#page_32">32-34</a>.<br/> +<i>Rodentia</i>, of the Eocene, <a href="#page_305">305</a>; + of the Miocene, <a href="#page_322">322</a>; of the + Post-Pliocene, <a href="#page_361">361</a>.<br/> +Roebuck, <a href="#page_336">336</a>, + <a href="#page_354">354</a>.<br/> +<i>Rostellaria</i>, <a href="#page_237">237</a>, + <a href="#page_293">293</a>.<br/> +<i>Rotalia</i>, <a href="#page_22">22</a>, + <a href="#page_98">98</a>, <a href="#page_171">171</a>, + <a href="#page_264">264</a>; <i>Boueana</i>, + <a href="#page_264">264</a>.<br/> +Rugose Corals, <a href="#page_104">104</a>; of the Lower + Silurian, <a href="#page_104">104</a>, + <a href="#page_105">105</a>; of the Upper Silurian, + <a href="#page_119">119</a>; of the Devonian, + <a href="#page_141">141</a>; of the Carboniferous, + <a href="#page_172">172-174</a>; of the Permian, + <a href="#page_197">197</a>; of the Upper Greensand, + <a href="#page_266">266</a>.<br/> +Rupelian Clay, <a href="#page_307">307</a>. +</p> + +<p class="index"> +<i>Sabal major</i>, <a href="#page_309">309</a>.<br/> +Sabre-toothed Tiger, <a href="#page_322">322</a>, + <a href="#page_331">331</a>.<br/> +<i>Saccammina</i>, <a href="#page_172">172</a>.<br/> +<i>Saccosoma</i>, <a href="#page_232">232</a>.<br/> +Salamanders, <a href="#page_189">189</a>, + <a href="#page_313">313</a>.<br/> +Salina Group, <a href="#page_117">117</a>.<br/> +<i>Salix</i>, <a href="#page_262">262</a>; <i>Meeki</i>, + <a href="#page_263">263</a>.<br/> +<i>SalmonidĹ“</i>, <a href="#page_276">276</a>.<br/> +<i>Sao hirsuta</i>, <a href="#page_85">85</a>.<br/> +<i>Sassafras cretacea</i>, <a href="#page_263">263</a>.<br/> +<i>Sauropterygia</i>, <a href="#page_219">219</a>.<br/> +<i>Scalaria</i>, <a href="#page_271">271</a>, + <a href="#page_293">293</a>; <i>GrĹ“nlandica</i>, + <a href="#page_338">338</a>.<br/> +<i>Scaphites</i>, <a href="#page_272">272</a>, + <a href="#page_273">273</a>; <i>Ĺ“qualis</i>, + <a href="#page_274">274</a>.<br/> +<i>Schizodus</i>, <a href="#page_198">198</a>, + <a href="#page_211">211</a>.<br/> +Schoharie Grit, <a href="#page_135">135</a>, + <a href="#page_137">137</a>.<br/> +<i>Scolecoderma</i>, <a href="#page_82">82</a>.<br/> +<i>Scoliostoma</i>, <a href="#page_213">213</a>.<br/> +<i>Scolithus</i>, <a href="#page_82">82</a>; + <i>Canadensis</i>, <a href="#page_83">83</a>.<br/> +Scorpions of the Coal-measures, + <a href="#page_181">181</a>.<br/> +Scorpion-shells, <a href="#page_271">271</a>.<br/> +<a name="page_406"><span class="page">Page 406</span></a> +Screw-pines, <a href="#page_230">230</a>.<br/> +<i>Scutella</i>, <a href="#page_311">311</a>; + <i>subrotunda</i>, <a href="#page_312">312</a>.<br/> +Sea-cows (<i>see</i> Sirenia).<br/> +Sea-lilies (<i>see</i> Crinoidea).<br/> +Sea-lizards (<i>see</i> Enaliosaurians).<br/> +Seals, <a href="#page_322">322</a>.<br/> +Sea-mats and Sea-mosses (<i>see</i> Polyzoa).<br/> +Sea-shrubs (<i>see</i> Gorgonidæ).<br/> +Sea-urchins (<i>see</i> Echinoidea).<br/> +Sea-weeds, <a href="#page_80">80</a>, <a href="#page_81">81</a>, + <a href="#page_83">83</a>, <a href="#page_97">97</a>, + <a href="#page_136">136</a>, <a href="#page_164">164</a>, + <a href="#page_261">261</a>.<br/> +Secondary period, <a href="#page_44">44</a>.<br/> +Sedimentary rocks, <a href="#page_15">15</a>.<br/> +<i>Semnopithecus</i>, <a href="#page_322">322</a>, + <a href="#page_331">331</a>.<br/> +Septaria, <a href="#page_31">31</a>.<br/> +<i>Sequoia</i>, <a href="#page_306">306</a>, + <a href="#page_309">309</a>, <a href="#page_310">310</a>; + <i>CouttsiĹ“</i>, <a href="#page_309">309</a>; + <i>gigantea</i>, <a href="#page_309">309</a>; + <i>Langsdorffii</i>, <a href="#page_309">309</a>.<br/> +<i>Serolis</i>, <a href="#page_84">84</a>.<br/> +Serpents (<i>see</i> Ophidia).<br/> +<i>Serpulites</i>, <a href="#page_123">123</a>.<br/> +Sewâlik Hills (<i>see</i> Siwâlik Hills).<br/> +Sheep, <a href="#page_355">355</a>.<br/> +Shell-sands, <a href="#page_19">19</a>.<br/> +<i>Sigillaria</i>, <a href="#page_168">168</a>; + <i>GrĹ“seri</i>, <a href="#page_168">168</a>.<br/> +Sigillarioids, <a href="#page_136">136</a>, + <a href="#page_168">168</a>, <a href="#page_170">170</a>, + <a href="#page_196">196</a>.<br/> +Silicates, infiltration of the shells of Foraminifera by, + <a href="#page_34">34</a>, <a href="#page_74">74</a>.<br/> +Siliceous rocks, <a href="#page_20">20</a>, + <a href="#page_32">32</a>.<br/> +Siliceous Sponges, <a href="#page_265">265</a>.<br/> +Silicification, <a href="#page_13">13</a>, + <a href="#page_14">14</a>.<br/> +Silurian period (<i>see</i> Lower Silurian and Upper Silurian), + <a href="#page_90">90-114</a>, <a href="#page_115">115-132</a>.<br/> +<i>Simosaurus</i>, <a href="#page_219">219</a>; <i>Gaillardoti</i>, + <a href="#page_219">219</a>.<br/> +<i>Siphonia</i>, <a href="#page_264">264</a>; <i>ficus</i>, + <a href="#page_265">265</a>.<br/> +Siphonostomatous Univalves, <a href="#page_237">237</a>, + <a href="#page_271">271</a>, <a href="#page_293">293</a>.<br/> +<i>Siphonotreta</i>, <a href="#page_110">110</a>.<br/> +<i>Sirenia</i>, <a href="#page_299">299</a>, + <a href="#page_320">320</a>; of the Eocene, + <a href="#page_299">299</a>; of the Miocene, + <a href="#page_315">315</a>.<br/> +<i>Siren lacertina</i>, <a href="#page_200">200</a>.<br/> +<i>Sivatherium</i>, <a href="#page_318">318</a>; + <i>giganteum</i>, <a href="#page_319">319</a>.<br/> +Siwâlik Hills, Miocene strata of, + <a href="#page_307">307</a>.<br/> +Skiddaw Slates, <a href="#page_101">101</a>.<br/> +Sloths, <a href="#page_315">315</a>, + <a href="#page_349">349-351</a>.<br/> +<i>Smilax</i>, <a href="#page_308">308</a>.<br/> +<i>Smithia</i>, <a href="#page_173">173</a>.<br/> +Snakes (<i>see</i> Ophidia).<br/> +Soft Tortoises, <a href="#page_296">296</a>.<br/> +<i>Solarium</i>, <a href="#page_271">271</a>.<br/> +Solenhofen Slates, <a href="#page_228">228</a>.<br/> +Solitaire, <a href="#page_346">346</a>, + <a href="#page_348">348</a>.<br/> +<i>Spalacotherium</i>, <a href="#page_254">254</a>.<br/> +<i>Spatangus</i>, <a href="#page_311">311</a>.<br/> +<i>SphĹ“rospongia</i>, <a href="#page_139">139</a>.<br/> +<i>Sphagodus</i>, <a href="#page_130">130</a>.<br/> +<i>Sphenodon</i>, <a href="#page_218">218</a>.<br/> +<i>Sphenopteris</i>, <a href="#page_136">136</a>, + <a href="#page_165">165</a>, <a href="#page_196">196</a>.<br/> +Spiders of the Coal-measures, <a href="#page_181">181</a>.<br/> +Spider-shells, <a href="#page_237">237</a>.<br/> +Spindle-shells, <a href="#page_237">237</a>.<br/> +<i>Spirifera</i>, <a href="#page_125">125</a>, + <a href="#page_147">147</a>, <a href="#page_184">184</a>, + <a href="#page_198">198</a>, <a href="#page_234">234</a>; + <i>crispa</i>, <a href="#page_127">127</a>; + <i>disjuncta</i>, <a href="#page_147">147</a>; + <i>hysterica</i>, <a href="#page_126">126</a>; + <i>mucronata</i>, <a href="#page_147">147</a>; + <i>Niagarensis</i>, <a href="#page_127">127</a>; + <i>rostrata</i>, <a href="#page_235">235</a>; + <i>sculptilis</i>, <a href="#page_147">147</a>; + <i>trigonalis</i>, <a href="#page_185">185</a>.<br/> +<i>SpiriferidĹ“</i>, <a href="#page_147">147</a>.<br/> +<i>Spirophyton cauda-Galli</i>, <a href="#page_135">135</a>, + <a href="#page_164">164</a>.<br/> +<i>Spirorbis</i>, <a href="#page_123">123</a>, + <a href="#page_143">143</a>, <a href="#page_178">178</a>; + <i>Arkonensis</i>, <a href="#page_144">144</a>; + <i>Carbonarus</i>, <a href="#page_178">178</a>; + <i>laxus</i>, <a href="#page_144">144</a>; <i>Lewisii</i>, + <a href="#page_123">123</a>; <i>omphalodes</i>, + <a href="#page_144">144</a>; <i>spinulifera</i>, + <a href="#page_144">144</a>.<br/> +<i>Spirulirostra</i>, <a href="#page_312">312</a>.<br/> +<i>Spondylus</i>, <a href="#page_269">269</a>; + <i>spinosus</i>, <a href="#page_270">270</a>.<br/> +Sponges, of the Cambrian, <a href="#page_81">81</a>; of the + Lower Silurian, <a href="#page_98">98</a>; of the Upper + Silurian, <a href="#page_119">119</a>; of the Devonian, + <a href="#page_139">139</a>; of the Carboniferous, + <a href="#page_171">171</a>; of the Permian, + <a href="#page_197">197</a>; of the Trias, + <a href="#page_209">209</a>; of the Jurassic, + <a href="#page_230">230</a>; of the Cretaceous, + <a href="#page_264">264</a>, <a href="#page_265">265</a>.<br/> +<i>Spongilla</i>, <a href="#page_197">197</a>.<br/> +<i>Spongillopsis</i>, <a href="#page_197">197</a>.<br/> +<i>Spongophyllum</i>, <a href="#page_173">173</a>.<br/> +Spore-eases, of Cryptogams in the Ludlow rocks, + <a href="#page_118">118</a>; in the Coal, + <a href="#page_163">163</a>.<br/> +Squirrels, <a href="#page_322">322</a>.<br/> +<i>Stagonolepis</i>, <a href="#page_218">218</a>.<br/> +Staircase-shell, <a href="#page_271">271</a>.<br/> +Stalactite, <a href="#page_21">21</a>.<br/> +Stalagmite, <a href="#page_21">21</a>.<br/> +Star-corals, <a href="#page_231">231</a>.<br/> +Star-fishes, <a href="#page_105">105</a>, + <a href="#page_120">120</a>, <a href="#page_210">210</a>.<br/> +St Cassian Beds, <a href="#page_205">205</a>, + <a href="#page_206">206</a>.<br/> +<i>Stephanophyllia</i>, <a href="#page_266">266</a>.<br/> +<i>Stereognathus</i>, <a href="#page_253">253</a>, + <a href="#page_254">254</a>.<br/> +<i>Stigmaria</i>, <a href="#page_169">169</a>; + <i>ficoides</i>, <a href="#page_169">169</a>.<br/> +Stonesfield Slate, <a href="#page_227">227</a>; + Mammals of, <a href="#page_253">253</a>.<br/> +Strata, contemporaneity of, <a href="#page_44">44</a>.<br/> +Stratified rock, <a href="#page_15">15-18</a>.<br/> +<i>Streptelasma</i>, <a href="#page_105">105</a>.<br/> +<i>Streptorhynchus</i>, <a href="#page_198">198</a>.<br/> +<i>Stromatopora</i>, <a href="#page_98">98</a>, + <a href="#page_99">99</a>, <a href="#page_118">118</a>, + <a href="#page_139">139</a>; <i>rugosa</i>, + <a href="#page_99">99</a>; <i>tuberculata</i>, + <a href="#page_140">140</a>.<br/> +<i>Strombodes</i>, <a href="#page_119">119</a>; + <i>pentagonus</i>, <a href="#page_104">104</a>.<br/> +<i>Strombus</i>, <a href="#page_271">271</a>.<br/> +<i>Strophalosia</i>, <a href="#page_198">198</a>.<br/> +<i>Strophodus</i>, <a href="#page_255">255</a>.<br/> +<i>Strophomena</i>, <a href="#page_109">109</a>, + <a href="#page_110">110</a>; <i>alternata</i>, + <a href="#page_110">110</a>; <i>deltoidea</i>, + <a href="#page_109">109</a>; <i>filitexta</i>, + <a href="#page_110">110</a>; <i>rhomboidalis</i>, + <a href="#page_147">147</a>, <a href="#page_148">148</a>; + <i>Subplana</i>, <a href="#page_127">127</a>.<br/> +Sub-Apennine Beds, <a href="#page_325">325</a>.<br/> +Sub-Carboniferous rocks, <a href="#page_158">158</a>, + <a href="#page_161">161</a>.<br/> +Succession of life upon the globe, + <a href="#page_367">367-374</a>.<br/> +<i>Suida</i>, <a href="#page_302">302</a>, + <a href="#page_317">317</a>, <a href="#page_329">329</a>.<br/> +Sulphate of lime, <a href="#page_22">22</a>.<br/> +<i>Sus Erymanthius</i>, <a href="#page_317">317</a>; + <i>scrofa</i>, <a href="#page_354">354</a>.<br/> +<i>SynastrĹ“a</i>, <a href="#page_209">209</a>.<br/> +<i>Synhelia Sharpeana</i>, <a href="#page_266">266</a>.<br/> +<i>Synocladia</i>, <a href="#page_198">198</a>; + <i>virgulacea</i>, <a href="#page_198">198</a>.<br/> +<i>Syringopora</i>, <a href="#page_119">119</a>, + <a href="#page_173">173</a>; <i>ramulosa</i>, + <a href="#page_174">174</a>. +</p> + +<p class="index"> +Tabulate Corals, <a href="#page_104">104</a>; of the Lower + Silurian, <a href="#page_105">105</a>; of the Upper + Silurian, <a href="#page_142">142</a>; of the Devonian, + <a href="#page_142">142</a>; of the Carboniferous, + <a href="#page_172">172</a>; of the Permian, + <a href="#page_197">197</a>.<br/> +<i>Talpa EuropĹ“a</i>, <a href="#page_336">336</a>.<br/> +<i>TapiridĹ“</i>, <a href="#page_300">300</a>.<br/> +Tapirs, <a href="#page_300">300</a>.<br/> +<i>Tapirus Arvernensis</i>, <a href="#page_327">327</a>.<br/> +<i>Taxocrinus tuberculatus</i>, <a href="#page_122">122</a>.<br/> +<i>Taxodium</i>, <a href="#page_262">262</a>, + <a href="#page_308">308</a>, <a href="#page_310">310</a>.<br/> +<i>Teleosaurus</i>, <a href="#page_251">251</a>.<br/> +Teleostean Fishes, <a href="#page_150">150</a>; of the + Cretaceous, <a href="#page_276">276</a>.<br/> +<i>Telerpeton Elginense</i>, <a href="#page_218">218</a>.<br/> +<i>Tellina proxima</i>, <a href="#page_338">338</a>.<br/> +<i>Tentaculites</i>, <a href="#page_129">129</a>, + <a href="#page_148">148</a>; <i>ornatus</i>, + <a href="#page_129">129</a>.<br/> +<i>Terebra</i>, <a href="#page_293">293</a>.<br/> +<i>Terebratella</i>, <a href="#page_268">268</a>, + <i>Astleriana</i>, <a href="#page_268">268</a>.<br/> +<i>Terebratula</i>, <a href="#page_184">184</a>, + <a href="#page_234">234</a>; <i>digona</i>, + <a href="#page_235">235</a>; <i>elongata</i>, + <a href="#page_168">168</a>; <i>hastata</i>, + <a href="#page_185">185</a>; <i>quadrifida</i>, + <a href="#page_235">235</a>; <i>sphĹ“roidalis</i>, + <a href="#page_235">235</a>.<br/> +<i>Terebratulina</i>, <a href="#page_268">268</a>; + <i>caput-serpentis</i>, <a href="#page_268">268</a>; + <i>striata</i>, <a href="#page_268">268</a>.<br/> +Termites, <a href="#page_311">311</a>.<br/> +Terrapins, <a href="#page_280">280</a>, + <a href="#page_296">296</a>.<br/> +Tertiary period, <a href="#page_44">44</a>, + <a href="#page_284">284-287</a>.<br/> +Tertiary rocks, classification of, + <a href="#page_284">284-287</a>.<br/> +<a name="page_407"><span class="page">Page 407</span></a> +<i>TestudinidĹ“</i>, <a href="#page_313">313</a>.<br/> +Tetrabranchiate Cephalopods, <a href="#page_112">112</a>; of + the Cambrian, <a href="#page_89">89</a>; of the Lower + Silurian, <a href="#page_112">112-114</a>; of the Upper + Silurian, <a href="#page_130">130</a>; of the Devonian, + <a href="#page_149">149</a>; of the Carboniferous, + <a href="#page_186">186</a>, <a href="#page_187">187</a>; + of the Permian, <a href="#page_199">199</a>; of the Trias, + <a href="#page_212">212</a>; of the Jurassic, + <a href="#page_237">237-239</a>; of the Cretaceous, + <a href="#page_272">272-274</a>; of the Eocene, + <a href="#page_294">294</a>; of the Miocene, + <a href="#page_312">312</a>.<br/> +<i>Textularia</i>, <a href="#page_22">22</a>, + <a href="#page_264">264</a>, <a href="#page_311">311</a>; + <i>Meyeriana</i>, <a href="#page_311">311</a>.<br/> +Thanet Sands, <a href="#page_287">287</a>, + <a href="#page_288">288</a>.<br/> +<i>Theca</i>, <a href="#page_88">88</a>, + <a href="#page_111">111</a>, <a href="#page_129">129</a>.<br/> +<i>Theca Davidii</i>, <a href="#page_88">88</a>.<br/> +<i>Thecidium</i>, <a href="#page_213">213</a>.<br/> +Thecodont Reptiles, <a href="#page_218">218</a>.<br/> +<i>Thecodontosaurus</i>, <a href="#page_200">200</a>, + <a href="#page_218">218</a>; <i>antiquus</i>, + <a href="#page_219">219</a>.<br/> +<i>Thecosmilia annularis</i>, <a href="#page_231">231</a>.<br/> +<i>Thelodus</i>, <a href="#page_131">131</a>.<br/> +Theriodont Reptiles, <a href="#page_202">202</a>, + <a href="#page_220">220</a>.<br/> +<i>Thylacoleo</i>, <a href="#page_349">349</a>.<br/> +Tile-stones, <a href="#page_116">116</a>.<br/> +<i>Titanotherium</i>, <a href="#page_316">316</a>.<br/> +Toothed Birds, <a href="#page_281">281-283</a>.<br/> +Tortoises, <a href="#page_202">202</a>, + <a href="#page_296">296</a>.<br/> +<i>Tragoceras</i>, <a href="#page_318">318</a>.<br/> +Travertine, <a href="#page_21">21</a>.<br/> +Tree-Ferns, of the Devonian, <a href="#page_136">136</a>; + of the Coal-measures, <a href="#page_164">164</a>.<br/> +Tremadoc Slates, <a href="#page_77">77-79</a>.<br/> +<i>Trematis</i>, <a href="#page_110">110</a>.<br/> +Trenton Limestone, <a href="#page_95">95</a>, + <a href="#page_96">96</a>.<br/> +<i>Trianthrus Beckii</i>, <a href="#page_107">107</a>.<br/> +Triassic period, <a href="#page_203">203</a>; rocks of, in + Britain, <a href="#page_204">204</a>; in Germany, + <a href="#page_204">204</a>; in the Austrian Alps, + <a href="#page_205">205</a>; in North America, + <a href="#page_205">205</a>; life of, + <a href="#page_206">206-224</a>.<br/> +<i>Triconodon</i>, <a href="#page_254">254</a>.<br/> +<i>Trigonia</i>, <a href="#page_235">235</a>, + <a href="#page_255">255</a>, <a href="#page_269">269</a>.<br/> +<i>TrigoniadĹ“</i>, <a href="#page_198">198</a>, + <a href="#page_211">211</a>.<br/> +<i>Trigonocarpum</i>, <a href="#page_170">170</a>; + <i>ovatum</i>, <a href="#page_170">170</a>.<br/> +Trilobites, <a href="#page_84">84-87</a>; of the Cambrian, + <a href="#page_85">85</a>, <a href="#page_87">87</a>; of the + Lower Silurian, <a href="#page_107">107</a>, + <a href="#page_108">108</a>; of the Upper Silurian, + <a href="#page_123">123</a>, <a href="#page_124">124</a>; of + the Devonian, <a href="#page_144">144</a>, + <a href="#page_145">145</a>; of the Carboniferous, + <a href="#page_179">179</a>.<br/> +<i>TrimerellidĹ“</i>, <a href="#page_127">127</a>.<br/> +<i>Trinucleus</i>, <a href="#page_108">108</a>; + <i>concentricus</i>, <a href="#page_107">107</a>.<br/> +<i>TrionycidĹ“</i>, <a href="#page_296">296</a>.<br/> +<i>Triton</i>, <a href="#page_293">293</a>.<br/> +<i>Trochocyathus</i>, <a href="#page_266">266</a>.<br/> +<i>Trochonema</i>, <a href="#page_129">129</a>.<br/> +<i>Trogontherium</i>, <a href="#page_361">361</a>; + <i>Cuvieri</i>, <a href="#page_336">336</a>, + <a href="#page_361">361</a>.<br/> +Trumpet-shells, <a href="#page_293">293</a>.<br/> +Tulip-tree, <a href="#page_262">262</a>, + <a href="#page_308">308</a>.<br/> +<i>Turbinolia sulcata</i>, <a href="#page_292">292</a>.<br/> +<i>TurbinolidĹ“</i>, <a href="#page_292">292</a>.<br/> +<i>Turrilites</i>, <a href="#page_272">272</a>, + <a href="#page_273">273</a>; <i>catenulatus</i>, + <a href="#page_274">274</a>.<br/> +<i>Turritella</i>, <a href="#page_271">271</a>, + <a href="#page_293">293</a>.<br/> +Turtles, <a href="#page_202">202</a>, + <a href="#page_251">251</a>, <a href="#page_280">280</a>, + <a href="#page_296">296</a>.<br/> +<i>Typhis tubifer</i>, <a href="#page_293">293</a>. +</p> + +<p class="index"> +<i>Ullmania selaginoides</i>, <a href="#page_197">197</a>.<br/> +Unconformability of strata, <a href="#page_48">48</a>.<br/> +Under-clay of coal, <a href="#page_162">162</a>.<br/> +<i>Ungulata</i>, of the Eocene, <a href="#page_300">300-303</a>; + of the Miocene, <a href="#page_315">315-319</a>; of the + Pliocene, <a href="#page_327">327-329</a>; of the + Post-Pliocene, <a href="#page_353">353-357</a>.<br/> +Uniformity, doctrine of, <a href="#page_5">5-7</a>.<br/> +<i>Unio</i>, <a href="#page_250">250</a>.<br/> +Univalves (<i>see</i> Gasteropoda).<br/> +Upper Cambrian, <a href="#page_77">77-79</a>; Chalk, + <a href="#page_259">259</a>; Cretaceous, + <a href="#page_257">257</a>, <a href="#page_259">259</a>; + Devonian, <a href="#page_135">135</a>; Eocene, + <a href="#page_287">287</a>, <a href="#page_288">288</a>; + Greensand, <a href="#page_258">258</a>; Helderberg, + <a href="#page_135">135</a>; Laurentian, + <a href="#page_66">66</a>; Llandovery, + <a href="#page_115">115</a>; Ludlow rock, + <a href="#page_116">116</a>; Miocene, + <a href="#page_305">305</a>; Oolites, + <a href="#page_227">227</a>; Silurian period, + <a href="#page_115">115</a>; rocks of, in Britain, + <a href="#page_115">115</a>, <a href="#page_116">116</a>; + in North America, <a href="#page_116">116-118</a>; life of, + <a href="#page_118">118-131</a>.<br/> +<i>Ursus arctos</i>, <a href="#page_359">359</a>; + <i>Arvernensis</i>, <a href="#page_339">339</a>; + <i>ferox</i>, <a href="#page_359">359</a>; + <i>spelĹ“a</i>, <a href="#page_360">360</a>.<br/> +<i>Ursus</i>, <a href="#page_336">336</a>, + <a href="#page_356">356</a>. +</p> + +<p class="index"> +Valley-gravels, high-level and low-level, + <a href="#page_339">339-341</a>.<br/> +<i>Vanessa Pluto</i>, <a href="#page_312">312</a>.<br/> +<i>VaranidĹ“</i>, <a href="#page_202">202</a>.<br/> +Vegetation (<i>see</i> Plants).<br/> +<i>Ventriculites</i>, <a href="#page_264">264</a>, + <a href="#page_265">265</a>; <i>simplex</i>, + <a href="#page_265">265</a>.<br/> +Venus's Flower-basket, <a href="#page_265">265</a>.<br/> +<i>Vermilia</i>, <a href="#page_197">197</a>.<br/> +<i>Vespertilio Parisiensis</i>, + <a href="#page_304">304</a>, <a href="#page_305">305</a>.<br/> +Vicksburg Beds, <a href="#page_289">289</a>.<br/> +Vines, <a href="#page_306">306</a>, + <a href="#page_309">309</a>, <a href="#page_310">310</a>.<br/> +Vitreous Sponges, <a href="#page_264">264</a>.<br/> +<i>Voltzia</i>, <a href="#page_208">208</a>; + <i>heterophylla</i>, <a href="#page_209">209</a>.<br/> +<i>Voluta</i>, <a href="#page_271">271</a>, + <a href="#page_293">293</a>; <i>elongata</i>, + <a href="#page_271">271</a>.<br/> +Volutes, <a href="#page_271">271</a>, + <a href="#page_293">293</a>, <a href="#page_312">312</a>. +</p> + +<p class="index"> +<i>Walchia</i>, <a href="#page_196">196</a>, + <a href="#page_197">197</a>; <i>piniformis</i>, + <a href="#page_196">196</a>.<br/> +Walrus, <a href="#page_322">322</a>.<br/> +Wealden Beds, <a href="#page_257">257</a>.<br/> +<i>Wellingtonia</i>, <a href="#page_309">309</a>, + <a href="#page_310">310</a>.<br/> +Wenlock Beds, <a href="#page_115">115</a>, + <a href="#page_117">117</a>; Limestone, + <a href="#page_115">115</a>; Shale, + <a href="#page_115">115</a>.<br/> +Wentle-traps, <a href="#page_271">271</a>.<br/> +Werfen Beds, <a href="#page_205">205</a>, + <a href="#page_206">206</a>.<br/> +Whalebone Whales, <a href="#page_299">299</a>, + <a href="#page_315">315</a>.<br/> +Whales, <a href="#page_299">299</a>, + <a href="#page_315">315</a>.<br/> +Whelks, <a href="#page_237">237</a>.<br/> +White Chalk, <a href="#page_259">259</a>; structure of, + <a href="#page_21">21</a>, <a href="#page_22">22</a>; + origin of, <a href="#page_23">23</a>, + <a href="#page_263">263</a>.<br/> +White Crag, <a href="#page_324">324</a>.<br/> +White River Beds, <a href="#page_307">307</a>.<br/> +Wild Boar, <a href="#page_354">354</a>.<br/> +<i>Williamsonia</i>, <a href="#page_230">230</a>.<br/> +Winged Lizards (<i>see</i> Pterosauria).<br/> +Winged Snails (<i>see</i> Pteropods).<br/> +Wing-shells, <a href="#page_271">271</a>.<br/> +Wolf, <a href="#page_336">336</a>, + <a href="#page_360">360</a>.<br/> +Wolverine, <a href="#page_360">360</a>.<br/> +Wombats, <a href="#page_348">348</a>.<br/> +Woolhope Limestone, <a href="#page_115">115</a>.<br/> +Woolly Rhinoceros, <a href="#page_339">339</a>, + <a href="#page_341">341</a>, <a href="#page_344">344</a>, + <a href="#page_353">353</a>.<br/> +Woolwich and Reading Beds, <a href="#page_287">287</a>.<br/> +Worm-burrows, <a href="#page_82">82</a>, + <a href="#page_83">83</a>, <a href="#page_123">123</a>. +</p> + +<p class="index"> +<i>Xanthidia</i>, <a href="#page_138">138</a>, + <a href="#page_161">161</a>.<br/> +<i>Xenoneura antiquorum</i>, <a href="#page_145">145</a>.<br/> +<i>Xiphodon</i>, <a href="#page_303">303</a>.<br/> +<i>Xylobius</i>, <a href="#page_182">182</a>; + <i>SigillariĹ“</i>, <a href="#page_182">182</a>. +</p> + +<p class="index"> +<i>Zamia spiralis</i>, <a href="#page_208">208</a>.<br/> +<i>Zamites</i>, <a href="#page_208">208</a>, + <a href="#page_230">230</a>, <a href="#page_310">310</a>.<br/> +<i>Zaphrentis</i>, <a href="#page_105">105</a>, + <a href="#page_119">119</a>, <a href="#page_142">142</a>, + <a href="#page_173">173</a>; <i>cornicula</i>, + <a href="#page_141">141</a>; <i>Stokesi</i>, + <a href="#page_104">104</a>; <i>vermicularis</i>, + <a href="#page_174">174</a>.<br/> +<i>Zeacrinus</i>, <a href="#page_175">175</a>.<br/> +Zechstein, <a href="#page_194">194</a>.<br/> +<i>Zeuglodon</i>, <a href="#page_299">299</a>, + <a href="#page_315">315</a>; <i>cetoides</i>, + <a href="#page_299">200</a>, <a href="#page_300">300</a>. +</p> + +<div>*** END OF THE PROJECT GUTENBERG EBOOK 14279 ***</div> +</body> +</html> diff --git a/14279-h/images/fig001.jpg b/14279-h/images/fig001.jpg Binary files differnew file mode 100644 index 0000000..ed64bad --- /dev/null +++ b/14279-h/images/fig001.jpg diff --git a/14279-h/images/fig002.jpg b/14279-h/images/fig002.jpg Binary files differnew file mode 100644 index 0000000..42eacf6 --- /dev/null +++ b/14279-h/images/fig002.jpg diff --git a/14279-h/images/fig003.jpg b/14279-h/images/fig003.jpg Binary files differnew file mode 100644 index 0000000..385c7a8 --- /dev/null +++ b/14279-h/images/fig003.jpg diff --git a/14279-h/images/fig004.jpg b/14279-h/images/fig004.jpg Binary 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