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+The Project Gutenberg EBook of On The Affinities of Leptarctus primus of
+Leidy, by J. L. Wortman
+
+This eBook is for the use of anyone anywhere at no cost and with
+almost no restrictions whatsoever. You may copy it, give it away or
+re-use it under the terms of the Project Gutenberg License included
+with this eBook or online at www.gutenberg.org
+
+
+Title: On The Affinities of Leptarctus primus of Leidy
+ American Museum of Natural History, Vol. VI, Article VIII, pp. 229-331.
+
+Author: J. L. Wortman
+
+Release Date: September 5, 2008 [EBook #26542]
+
+Language: English
+
+Character set encoding: ISO-8859-1
+
+*** START OF THIS PROJECT GUTENBERG EBOOK LEPTARCTUS PRIMUS OF LEIDY ***
+
+
+
+
+Produced by Gerard Arthus, Louise Blyton and the Online
+Distributed Proofreading Team at https://www.pgdp.net
+
+
+
+
+
+
+
+
+
+_On the Affinities of Leptarctus primus of Leidy._
+
+By J. L. WORTMAN.
+
+
+
+_AUTHOR'S EDITION, extracted from_
+
+BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY,
+
+VOL. VI, ARTICLE VIII, pp. 229-231.
+
+
+_New York, July 30, 1894._
+
+
+
+
+Article VIII.--ON THE AFFINITIES OF LEPTARCTUS PRIMUS OF LEIDY.
+
+By J. L. Wortman.
+
+
+Up to the present time but very little has been known of the existence
+of the peculiarly American family Procyonidæ in any deposits older
+than the very latest Quaternary. Leidy has described and figured[1] an
+isolated last upper tooth, from the Loup Fork deposits of Nebraska,
+under the name of _Leptarctus primus_, which has been referred to this
+family. The Museum Expedition of last year into this region was
+successful in obtaining additional material, which we provisionally
+refer to Leidy's species.
+
+
+=Leptarctus primus= _Leidy_.
+
+The specimen consists of the right ramus of the lower jaw, carrying
+the third and fourth premolars and the canine. The condyle is broken
+away, but the coronoid process and the angle are preserved. The
+specimen is from a young individual in which the last premolar had
+just cut the gum. The alveoli of all the other teeth are present and
+in a good state of preservation.
+
+The dental formula is as follows: I._3, C._1, Pm._3, M._2. The
+incisors are not preserved, but their alveoli indicate that they were
+much crowded, the outside one being placed almost directly in front of
+the canine, and the middle one pushed back considerably out of
+position. This series is in marked contrast with that of the Raccoon,
+in which the crowns of the incisors form almost a straight line across
+the jaw, and the middle one is crowded backwards to a very slight
+extent. The canine is peculiar and differs markedly from that of the
+Raccoon. It is rather robust, very much recurved and grooved by a deep
+vertical sulcus upon its antero-internal face. This sulcus is but
+faintly indicated in the Raccoon. The postero-external face of the
+crown is marked by a sharp ridge which becomes more prominent near the
+apex. The first premolar is not preserved, but its alveolus indicates
+that it was a single-rooted tooth, placed behind the canine after the
+intervention of a very short diastema. The second premolar is
+bifanged; its crown is composed of a principal cusp, to which is added
+behind a small though very distinct second cusp. There is in addition
+to these cusps a distinct basal cingulum, most prominent in the region
+of the heel. The third premolar, like the second, is double rooted;
+its crown moreover is made up of two cusps, the posterior being almost
+as large as the principal one. These cusps do not stand in the line of
+the long axis of the jaw, but are placed very obliquely to it. The
+heel is not very prominent, but the basal cingulum is well developed,
+both in front and behind. As compared with the Raccoon, the second
+premolar is more complex in that it has two cusps instead of one. In
+the third premolar the posterior cusp is much better developed, and
+placed more obliquely than in the corresponding tooth of _Procyon_;
+the heel is moreover not so broad.
+
+The first molar is not preserved, but judging from the size of its
+roots it was decidedly the longest tooth of the series. The second
+molar was likewise bifanged but much smaller; it was placed close
+against the base of the coronoid.
+
+The whole jaw has, relatively, a greater depth than that of the
+Raccoon, and is remarkably straight upon its lower border, whereas in
+the recent genus it is considerably curved. The condyle is not
+preserved, and the angle is somewhat damaged, but it was apparently
+not so strongly inflected as in the Raccoon. The masseteric fossa is
+deep and prominent, and the coronoid is high and broad. The inferior
+dental canal is placed higher than it is in the Raccoon, being
+slightly above the tooth line. The symphysis is relatively deeper and
+more robust than in _Procyon_, and the chin is heavier and more
+abruptly rounded.
+
+The jaw of _Leptarctus_ differs from that of _Cercoleptes_ in the
+following characters: the coronoid is broader and of less vertical
+extent; the condyle is not placed so high; the angle is elevated above
+the lower border of the ramus, which is straight and not concave as it
+is in _Cercoleptes_. In the depth of the symphysis and abrupt rounding
+of the chin the two genera are similar. _Cercoleptes_, moreover, has a
+moderately deep groove upon the antero-internal face of the canine,
+but differs from that of _Leptarctus_ in having an external groove as
+well. _Cercoleptes_ again resembles _Leptarctus_ in having only three
+premolars in the lower jaw; the middle one, however, has only a single
+cusp upon the crown, whereas _Leptarctus_ has two.
+
+As compared with _Bassaricyon_,[2] the jaw is more robust, shorter and
+deeper, with a more prominent chin. The two genera differ again in the
+number of premolars.
+
+Altogether, _Leptarctus_ appears to offer a number of transitional
+characters between the more typical Procyonidæ and the aberrant
+_Cercoleptes_. This is especially to be seen in the proportions of the
+jaw, the reduction of the number of premolars, the reduction in size
+of the last molar, as well as the depth of the mandibular symphysis.
+
+
+FOOTNOTES:
+
+[1] Extinct Fauna of Dakota.
+
+[2] See J. A. Allen's paper, Proc. Phil. Acad., 1876, p. 21.
+
+
+ Transcriber's Note:
+
+ "Quartenary" was amended to "Quaternary" in the first paragraph.
+
+
+
+
+
+End of the Project Gutenberg EBook of On The Affinities of Leptarctus primus
+of Leidy, by J. L. Wortman
+
+*** END OF THIS PROJECT GUTENBERG EBOOK LEPTARCTUS PRIMUS OF LEIDY ***
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+<pre>
+
+The Project Gutenberg EBook of On The Affinities of Leptarctus primus of
+Leidy, by J. L. Wortman
+
+This eBook is for the use of anyone anywhere at no cost and with
+almost no restrictions whatsoever. You may copy it, give it away or
+re-use it under the terms of the Project Gutenberg License included
+with this eBook or online at www.gutenberg.org
+
+
+Title: On The Affinities of Leptarctus primus of Leidy
+ American Museum of Natural History, Vol. VI, Article VIII, pp. 229-331.
+
+Author: J. L. Wortman
+
+Release Date: September 5, 2008 [EBook #26542]
+
+Language: English
+
+Character set encoding: ISO-8859-1
+
+*** START OF THIS PROJECT GUTENBERG EBOOK LEPTARCTUS PRIMUS OF LEIDY ***
+
+
+
+
+Produced by Gerard Arthus, Louise Blyton and the Online
+Distributed Proofreading Team at https://www.pgdp.net
+
+
+
+
+
+
+</pre>
+
+
+<hr />
+
+<h3><i>On the Affinities of Leptarctus primus of Leidy.</i></h3>
+
+<h4>By <span class="smcap">J. L. Wortman</span>.</h4>
+
+<hr style='width: 22%;' />
+
+<div class="center">
+
+<i>AUTHOR&#8217;S EDITION, extracted from</i>
+<br /><br />
+Bulletin of the American Museum of Natural History,
+<br /><br />
+<span class="smcap">Vol. VI, Article VIII</span>, pp. 229&#8211;231.
+<br /><br /><br />
+<i>New York, July 30, 1894.</i>
+
+</div>
+
+<hr />
+
+
+<div class="center" style="padding-top: 2em;">
+<b>Article VIII.</b>&mdash;ON THE AFFINITIES OF LEPTARCTUS PRIMUS OF LEIDY.
+<span class='pagenum'>[Pg&nbsp;229]</span>
+<br /><br /><br />
+By <span class="smcap">J. L. Wortman</span>.
+</div>
+
+
+<p style="padding-top: 2em;">Up to the present time but very little has been known of the
+existence of the peculiarly American family Procyonid&aelig; in any
+deposits older than the very latest Quaternary. Leidy has
+described and figured<a name="FNanchor_1_1" id="FNanchor_1_1"></a><a href="#Footnote_1_1" class="fnanchor">[1]</a> an isolated last upper tooth, from the
+Loup Fork deposits of Nebraska, under the name of <i>Leptarctus
+primus</i>, which has been referred to this family. The Museum
+Expedition of last year into this region was successful in obtaining
+additional material, which we provisionally refer to Leidy&#8217;s
+species.</p>
+
+<div class="center" style="padding-top: 0.5em;">
+<b>Leptarctus primus</b> <i>Leidy</i>.
+</div>
+
+<p style="padding-top: 1.0em;">The specimen consists of the right ramus of the lower jaw,
+carrying the third and fourth premolars and the canine. The
+condyle is broken away, but the coronoid process and the angle
+are preserved. The specimen is from a young individual in which
+the last premolar had just cut the gum. The alveoli of all the
+other teeth are present and in a good state of preservation.</p>
+
+<p>The dental formula is as follows: I.<sub>3</sub>, C.<sub>1</sub>, Pm.<sub>3</sub>, M.<sub>2</sub>. The
+incisors are not preserved, but their alveoli indicate that they
+were much crowded, the outside one being placed almost directly
+in front of the canine, and the middle one pushed back considerably
+out of position. This series is in marked contrast with
+that of the Raccoon, in which the crowns of the incisors form
+almost a straight line across the jaw, and the middle one is
+crowded backwards to a very slight extent. The canine is peculiar
+and differs markedly from that of the Raccoon. It is rather
+robust, very much recurved and grooved by a deep vertical sulcus
+upon its antero-internal face. This sulcus is but faintly indicated
+in the Raccoon. The postero-external face of the crown is
+marked by a sharp ridge which becomes more prominent near
+<span class='pagenum'>[Pg&nbsp;230]</span>the apex. The first premolar is not preserved, but its alveolus
+indicates that it was a single-rooted tooth, placed behind the
+canine after the intervention of a very short diastema. The
+second premolar is bifanged; its crown is composed of a principal
+cusp, to which is added behind a small though very distinct
+second cusp. There is in addition to these cusps a distinct basal
+cingulum, most prominent in the region of the heel. The third
+premolar, like the second, is double rooted; its crown moreover
+is made up of two cusps, the posterior being almost as large as
+the principal one. These cusps do not stand in the line of the
+long axis of the jaw, but are placed very obliquely to it. The
+heel is not very prominent, but the basal cingulum is well developed,
+both in front and behind. As compared with the Raccoon,
+the second premolar is more complex in that it has two cusps
+instead of one. In the third premolar the posterior cusp is
+much better developed, and placed more obliquely than in the
+corresponding tooth of <i>Procyon</i>; the heel is moreover not so
+broad.</p>
+
+<p>The first molar is not preserved, but judging from the size of
+its roots it was decidedly the longest tooth of the series. The
+second molar was likewise bifanged but much smaller; it was
+placed close against the base of the coronoid.</p>
+
+<p>The whole jaw has, relatively, a greater depth than that of the
+Raccoon, and is remarkably straight upon its lower border,
+whereas in the recent genus it is considerably curved. The condyle
+is not preserved, and the angle is somewhat damaged, but it
+was apparently not so strongly inflected as in the Raccoon. The
+masseteric fossa is deep and prominent, and the coronoid is high
+and broad. The inferior dental canal is placed higher than it is
+in the Raccoon, being slightly above the tooth line. The symphysis
+is relatively deeper and more robust than in <i>Procyon</i>, and
+the chin is heavier and more abruptly rounded.</p>
+
+<p>The jaw of <i>Leptarctus</i> differs from that of <i>Cercoleptes</i> in the
+following characters: the coronoid is broader and of less vertical
+extent; the condyle is not placed so high; the angle is
+elevated above the lower border of the ramus, which is straight
+and not concave as it is in <i>Cercoleptes</i>. In the depth of the
+symphysis and abrupt rounding of the chin the two genera are
+<span class='pagenum'>[Pg&nbsp;231]</span>similar. <i>Cercoleptes</i>, moreover, has a moderately deep groove
+upon the antero-internal face of the canine, but differs from that
+of <i>Leptarctus</i> in having an external groove as well. <i>Cercoleptes</i>
+again resembles <i>Leptarctus</i> in having only three premolars in the
+lower jaw; the middle one, however, has only a single cusp upon
+the crown, whereas <i>Leptarctus</i> has two.</p>
+
+<p>As compared with <i>Bassaricyon</i>,<a name="FNanchor_2_2" id="FNanchor_2_2"></a><a href="#Footnote_2_2" class="fnanchor">[2]</a> the jaw is more robust, shorter
+and deeper, with a more prominent chin. The two genera differ
+again in the number of premolars.</p>
+
+<p>Altogether, <i>Leptarctus</i> appears to offer a number of transitional
+characters between the more typical Procyonid&aelig; and the aberrant
+<i>Cercoleptes</i>. This is especially to be seen in the proportions of
+the jaw, the reduction of the number of premolars, the reduction
+in size of the last molar, as well as the depth of the mandibular
+symphysis.</p>
+
+<div class="footnote"><p><a name="Footnote_1_1" id="Footnote_1_1"></a><a href="#FNanchor_1_1"><span class="label">[1]</span></a> Extinct Fauna of Dakota.</p></div>
+
+<div class="footnote"><p><a name="Footnote_2_2" id="Footnote_2_2"></a><a href="#FNanchor_2_2"><span class="label">[2]</span></a> See J. A. Allen&#8217;s paper, Proc. Phil. Acad., 1876, p. 21.</p></div>
+
+<div class="transnote">
+<p><b>Transcriber&#8217;s Note:</b></p>
+
+<p style="padding-bottom: 1.0em;">&#8220;Quartenary&#8221; was amended to &#8220;Quaternary&#8221; in the first paragraph.</p>
+</div>
+
+
+
+
+
+
+
+
+<pre>
+
+
+
+
+
+End of the Project Gutenberg EBook of On The Affinities of Leptarctus primus
+of Leidy, by J. L. Wortman
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+The Project Gutenberg EBook of On The Affinities of Leptarctus primus of
+Leidy, by J. L. Wortman
+
+This eBook is for the use of anyone anywhere at no cost and with
+almost no restrictions whatsoever. You may copy it, give it away or
+re-use it under the terms of the Project Gutenberg License included
+with this eBook or online at www.gutenberg.org
+
+
+Title: On The Affinities of Leptarctus primus of Leidy
+ American Museum of Natural History, Vol. VI, Article VIII, pp. 229-331.
+
+Author: J. L. Wortman
+
+Release Date: September 5, 2008 [EBook #26542]
+
+Language: English
+
+Character set encoding: ASCII
+
+*** START OF THIS PROJECT GUTENBERG EBOOK LEPTARCTUS PRIMUS OF LEIDY ***
+
+
+
+
+Produced by Gerard Arthus, Louise Blyton and the Online
+Distributed Proofreading Team at https://www.pgdp.net
+
+
+
+
+
+
+
+
+
+_On the Affinities of Leptarctus primus of Leidy._
+
+By J. L. WORTMAN.
+
+
+
+_AUTHOR'S EDITION, extracted from_
+
+BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY,
+
+VOL. VI, ARTICLE VIII, pp. 229-231.
+
+
+_New York, July 30, 1894._
+
+
+
+
+Article VIII.--ON THE AFFINITIES OF LEPTARCTUS PRIMUS OF LEIDY.
+
+By J. L. Wortman.
+
+
+Up to the present time but very little has been known of the existence
+of the peculiarly American family Procyonidae in any deposits older
+than the very latest Quaternary. Leidy has described and figured[1] an
+isolated last upper tooth, from the Loup Fork deposits of Nebraska,
+under the name of _Leptarctus primus_, which has been referred to this
+family. The Museum Expedition of last year into this region was
+successful in obtaining additional material, which we provisionally
+refer to Leidy's species.
+
+
+=Leptarctus primus= _Leidy_.
+
+The specimen consists of the right ramus of the lower jaw, carrying
+the third and fourth premolars and the canine. The condyle is broken
+away, but the coronoid process and the angle are preserved. The
+specimen is from a young individual in which the last premolar had
+just cut the gum. The alveoli of all the other teeth are present and
+in a good state of preservation.
+
+The dental formula is as follows: I._3, C._1, Pm._3, M._2. The
+incisors are not preserved, but their alveoli indicate that they were
+much crowded, the outside one being placed almost directly in front of
+the canine, and the middle one pushed back considerably out of
+position. This series is in marked contrast with that of the Raccoon,
+in which the crowns of the incisors form almost a straight line across
+the jaw, and the middle one is crowded backwards to a very slight
+extent. The canine is peculiar and differs markedly from that of the
+Raccoon. It is rather robust, very much recurved and grooved by a deep
+vertical sulcus upon its antero-internal face. This sulcus is but
+faintly indicated in the Raccoon. The postero-external face of the
+crown is marked by a sharp ridge which becomes more prominent near the
+apex. The first premolar is not preserved, but its alveolus indicates
+that it was a single-rooted tooth, placed behind the canine after the
+intervention of a very short diastema. The second premolar is
+bifanged; its crown is composed of a principal cusp, to which is added
+behind a small though very distinct second cusp. There is in addition
+to these cusps a distinct basal cingulum, most prominent in the region
+of the heel. The third premolar, like the second, is double rooted;
+its crown moreover is made up of two cusps, the posterior being almost
+as large as the principal one. These cusps do not stand in the line of
+the long axis of the jaw, but are placed very obliquely to it. The
+heel is not very prominent, but the basal cingulum is well developed,
+both in front and behind. As compared with the Raccoon, the second
+premolar is more complex in that it has two cusps instead of one. In
+the third premolar the posterior cusp is much better developed, and
+placed more obliquely than in the corresponding tooth of _Procyon_;
+the heel is moreover not so broad.
+
+The first molar is not preserved, but judging from the size of its
+roots it was decidedly the longest tooth of the series. The second
+molar was likewise bifanged but much smaller; it was placed close
+against the base of the coronoid.
+
+The whole jaw has, relatively, a greater depth than that of the
+Raccoon, and is remarkably straight upon its lower border, whereas in
+the recent genus it is considerably curved. The condyle is not
+preserved, and the angle is somewhat damaged, but it was apparently
+not so strongly inflected as in the Raccoon. The masseteric fossa is
+deep and prominent, and the coronoid is high and broad. The inferior
+dental canal is placed higher than it is in the Raccoon, being
+slightly above the tooth line. The symphysis is relatively deeper and
+more robust than in _Procyon_, and the chin is heavier and more
+abruptly rounded.
+
+The jaw of _Leptarctus_ differs from that of _Cercoleptes_ in the
+following characters: the coronoid is broader and of less vertical
+extent; the condyle is not placed so high; the angle is elevated above
+the lower border of the ramus, which is straight and not concave as it
+is in _Cercoleptes_. In the depth of the symphysis and abrupt rounding
+of the chin the two genera are similar. _Cercoleptes_, moreover, has a
+moderately deep groove upon the antero-internal face of the canine,
+but differs from that of _Leptarctus_ in having an external groove as
+well. _Cercoleptes_ again resembles _Leptarctus_ in having only three
+premolars in the lower jaw; the middle one, however, has only a single
+cusp upon the crown, whereas _Leptarctus_ has two.
+
+As compared with _Bassaricyon_,[2] the jaw is more robust, shorter and
+deeper, with a more prominent chin. The two genera differ again in the
+number of premolars.
+
+Altogether, _Leptarctus_ appears to offer a number of transitional
+characters between the more typical Procyonidae and the aberrant
+_Cercoleptes_. This is especially to be seen in the proportions of the
+jaw, the reduction of the number of premolars, the reduction in size
+of the last molar, as well as the depth of the mandibular symphysis.
+
+
+FOOTNOTES:
+
+[1] Extinct Fauna of Dakota.
+
+[2] See J. A. Allen's paper, Proc. Phil. Acad., 1876, p. 21.
+
+
+ Transcriber's Note:
+
+ "Quartenary" was amended to "Quaternary" in the first paragraph.
+
+
+
+
+
+End of the Project Gutenberg EBook of On The Affinities of Leptarctus primus
+of Leidy, by J. L. Wortman
+
+*** END OF THIS PROJECT GUTENBERG EBOOK LEPTARCTUS PRIMUS OF LEIDY ***
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